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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1179.107587" ID-Pensoft-Pub="1313-2970-1179-35" ID-Pensoft-UUID="07CEAFC9849852A1AC00767783926142" ID-ZooBank="D9BD50CBB127487981E99421E4F059BA" ModsDocID="1313-2970-1179-35" checkinTime="1694247957851" checkinUser="pensoft" docAuthor="Gao, Biao, Han, Hui-Lin, Kononenko, Vladimir S. &amp; Pan, Zhao-Hui" docDate="2023" docId="89BC1B58008B5ECCA6D3CAC459469F1A" docLanguage="en" docName="ZooKeys 1179: 35-61" docOrigin="ZooKeys 1179" docPubDate="2023-09-08" docSource="http://dx.doi.org/10.3897/zookeys.1179.107587" docTitle="Hermonassa nigricans Gao, Han &amp; Kononenko 2023, sp. nov." docType="treatment" docUuid="51C58FF7-BBED-4279-B252-9C3E217B25C8" docUuidSource="ZooBank" docVersion="1" id="07CEAFC9849852A1AC00767783926142" lastPageNumber="35" masterDocId="07CEAFC9849852A1AC00767783926142" masterDocTitle="Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)" masterLastPageNumber="61" masterPageNumber="35" pageNumber="35" updateTime="1694247957851" updateUser="pensoft">
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<mods:title id="6D27AA4EC96E11C8DD8041FAD8B16CDF">Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)</mods:title>
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<mods:namePart id="8A6F6C844AE890D70A23B69425212DA0">Gao, Biao</mods:namePart>
<mods:affiliation id="C81A46F34CD22DAA2E8BAA6F420FDE2F">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="34D474C41BB5021B4531893E1E7F2B0F">Han, Hui-Lin</mods:namePart>
<mods:nameIdentifier id="9743E39D042A21900573A787DA491217" type="ORCID">https://orcid.org/0000-0002-2045-6182</mods:nameIdentifier>
<mods:affiliation id="B10F6774F488555E680597CF3095628E">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="B9FB8F99C6C2A29008929C0A74491798">Kononenko, Vladimir S.</mods:namePart>
<mods:nameIdentifier id="1D75352BF045CEECD43FFB100BC4C5F5" type="ORCID">https://orcid.org/0000-0001-6103-4800</mods:nameIdentifier>
<mods:affiliation id="D854F048B22A5AB4C63699B50B6908CD">Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 15004, China</mods:affiliation>
<mods:nameIdentifier id="55C6CE81888FA405979F2B52D57A14B7" type="email">vsk528217@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1EDFD9420126A1011AF4B44504063ACD">Pan, Zhao-Hui</mods:namePart>
<mods:affiliation id="4292E2C0A11942113468C7EC29F8F1E9">Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China</mods:affiliation>
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<mods:date id="AA60F01554ED9C90413C26202364B6ED">2023</mods:date>
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<treatment id="89BC1B58008B5ECCA6D3CAC459469F1A" LSID="urn:lsid:zoobank.org:act:51C58FF7-BBED-4279-B252-9C3E217B25C8" httpUri="http://treatment.plazi.org/id/89BC1B58008B5ECCA6D3CAC459469F1A" lastPageNumber="35" pageId="0" pageNumber="35">
<subSubSection id="27C7D5AA833D036BFA1D832223E5595D" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="71FCD286D7B394988A94D67E69923443" pageId="0" pageNumber="35">
<taxonomicName id="40ED749A0ECB56E18518E2477B045AC3" LSID="https://zoobank.org/51C58FF7-BBED-4279-B252-9C3E217B25C8" authority="Gao, Han &amp; Kononenko" authorityName="Gao, Han &amp; Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa nigricans" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="nigricans" status="sp. nov.">Hermonassa nigricans Gao, Han &amp; Kononenko</taxonomicName>
<taxonomicNameLabel id="F5251CD212FB95B87FDFA66B152EB779" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="09CAEB56E256512D4A805E1E81A78156" pageId="0" pageNumber="35" type="description">
<paragraph id="1A529D81BDABCAEE8113D5ED9C094A42" pageId="0" pageNumber="35">
<figureCitation id="63C8757FC78FB6309BCEDF0BC841C3E3" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">Figs 1</figureCitation>
<figureCitation id="75E65AEB3C9339CCF7D0FE7AEB3D791F" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">, 2</figureCitation>
<figureCitation id="CE1EA166882240CDEA03EAF64C31E420" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">, 11</figureCitation>
<figureCitation id="F66BD3A36525CD24CF4E3E9654F84015" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">, 12</figureCitation>
<figureCitation id="BDAA56EF2FD93395320BAA49147B0133" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7ED1D2428D05C60E32622B334047229" pageId="0" pageNumber="35" type="reference_group">
<paragraph id="96D47077B0D203B8A08B5343F0BC4F4D" pageId="0" pageNumber="35">
<taxonomicName id="B3CDF6ECDC28693C265D406AB47D5E8D" authority="Gao &amp; Han &amp; Kononenko &amp; Pan, 2023" authorityName="Gao &amp; Han &amp; Kononenko &amp; Pan" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa anthracina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="anthracina" status="sp. nov.">Hermonassa anthracina</taxonomicName>
:
<bibRefCitation id="A9650E4DAD601B99918CE7F6D396D3C3" author="Sugi, S" editor="Haruta, T" journalOrPublisher="Tinea 14 (suppl. 2)" pageId="0" pageNumber="35" pagination="90 - 109" refId="B25" refString="Sugi, S, 1995. Noctuidae: Noctuinae, Hermonassa. In: Haruta, T, Ed., Moths of Nepal IV. Tinea 14 (suppl. 2): 90 - 109" title="Noctuidae: Noctuinae, Hermonassa." volumeTitle="Moths of Nepal IV." year="1995">Sugi 1995</bibRefCitation>
: 90, pl. 117, fig. 3, genit. fig. 696 (♂), 732 (♀) (misidentification).
</paragraph>
</subSubSection>
<subSubSection id="02D07E93FE3C6A1E0EC73D265D7ED7FF" pageId="0" pageNumber="35" type="type material">
<paragraph id="F8FDE45404A207A263E9E3CD31CEACAA" pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph id="594FDE6535D26C84C49EDCBD2967F3FE" pageId="0" pageNumber="35">
<emphasis id="F790438894C7B36CC3AF35F73547AC84" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
: ♂, China, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, Cordyceps base, 4121 m, 9 Nov. 2016, H.L. Han leg., genit. prep. GB-101-1.
<emphasis id="9BE21A7B4A315D8743BBAFA1B634B626" bold="true" italics="true" pageId="0" pageNumber="35">Paratypes</emphasis>
: China: 2 ♀, Aut. Reg. Xizang, Linzhi City, Lulang town, Layue village, 2200 m, 14-15 Aug. 2014, H.L. Han leg., genit. prep. GB-20-2, GB-202-2; 1 ♂, 6 ♀, Aut. Reg. Xizang, Linzhi City, Lulang town, Mt. Sejila, Yaguo, 3650 m, 22 Aug. 2014, H.L. Han leg., genit. prep. GB-19-2, GB-26-2, GB-28-1, GB-197-2, GB-200-2, GB-201-2, GB-204-2; 1 ♂, Aut. Reg. Xizang, Linzhi City, Lulang town, Military depot, 3121 m, 3 Aug. 2015, H.L. Han leg., genit. prep. GB-119-1; 1 ♂, Aut. Reg. Xizanzhi City, Lulang town, 3506 m, 5 Aug. 2015, HL Han leg., genit. prep. GB-199-1; 1 ♀, Aut. Reg. Xizang, Linzhi City, Lulang town, Mt. Sejila, Yaguo, 3150 m, 24 Aug. 2015, H.L. Han leg., genit. prep. GB-25-2; 1 ♂, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, Cordyceps base, 4121 m, 9 Nov. 2016, H.L. Han leg., genit. prep. GB-101-1; 1 ♂, Aut. Reg. Xizang, Linzhi City, Zhangmai village, 13 Aug. 2017, H.L. Han leg., genit. prep. GB-120-1; 1 ♀, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, Cordyceps base, 17 Aug. 2017, H.L. Han leg., genit. prep. GB-108-2.
</paragraph>
</subSubSection>
<subSubSection id="6A90C8EE4910912499ACA56637199213" pageId="0" pageNumber="35" type="diagnosis">
<paragraph id="74711E0ADDD0B23F5EC55849502FA6F3" pageId="0" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="D478A39A94DB329B0BBF5957C88CA25C" pageId="0" pageNumber="35">
The new species is superficially very similar to
<taxonomicName id="37F24444C8BEFBD55AC831869927A0B5" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="B2DFAC3AFB61D348C92209EEBF10BB7F" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
(Figs
<figureCitation id="5D70E81AF48D054668DAD620183F45F5" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">3</figureCitation>
,
<figureCitation id="AB74161A7D345ACE804801B5456787B5" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">4</figureCitation>
,
<figureCitation id="E192A3550C3DE6794FEFBB0842ED8F7D" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">13-15</figureCitation>
,
<figureCitation id="121E845237D3E276841487E572AFEA3B" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">24</figureCitation>
), and
<taxonomicName id="E5DB789E06250DC147F3635E27B381A3" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="9E4AD65E25F1EE35B2D770B3279430A8" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
(Figs
<figureCitation id="1A0A84124D971861724FB7A9E05C21A5" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">5</figureCitation>
,
<figureCitation id="10D06F7001FAFDE20B83A4E7D2D1959B" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">6</figureCitation>
,
<figureCitation id="674B66550A344EAF0D93E0492DE0CFFF" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">16-18</figureCitation>
,
<figureCitation id="0BA97F69BC54E020F97A4399761EB497" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">25</figureCitation>
) and can barely be distinguished from these species by external characters. The new species is blackish-brown, somewhat darker than both other species; the antemedial line is somewhat broader than in both species, the postmedial line is doubled and more distinct than in both species. The hindwing is somewhat more whitish than in the related species. The main specific differences are in the genitalia of both sexes. In the male genitalia of
<taxonomicName id="87B7087553396808DD2851A77281A181" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="D2DBE9B5959AC2D5D253D6E19821B95A" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov. the uncus is more massive and gradually expands from the base to the top (in
<taxonomicName id="758B6DA462F95FD2E8FC4ED38A83AD47" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="8D111CD1DB49EA483208C0C556750799" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="67B847DCE03F8BC91B7F597802A6072E" authorityName="Kovacs, G. Ronkay &amp; L. Ronkay" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="kalamantra">
<emphasis id="88C0B28AD3576AF59DE3E7FDD1BAA99A" italics="true" pageId="0" pageNumber="35">Hermonassa kalamantra</emphasis>
</taxonomicName>
it is suddenly expands from 2/3 of the base); the tegumen is broader than in both species; the harpe is more massive and the club-like top of harpe is more swollen compared with
<taxonomicName id="55F5F2B57C759A7F8D106565FFCBA907" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="105AD3308B05BB826DE4BA2498F01101" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="9F1D015424C9F0B2326A5D2109F66678" authorityName="Kovacs, G. Ronkay &amp; L. Ronkay" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="kalamantra">
<emphasis id="5C477ED1C647C0E388CB5B77EA7A73D7" italics="true" pageId="0" pageNumber="35">Hermonassa kalamantra</emphasis>
</taxonomicName>
; the sclerotised flattened extension of sacculus is rounded and directed upward (in
<taxonomicName id="7B099437389E1071521AC2DDFF55643A" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="4B7E82CEBAF43B37CC582386B7EDE596" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
it is directed straight, in
<taxonomicName id="616C000F3CF239383EFBB2635447585E" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="7120A067A8EC4BB01A815AE956D41924" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
downward). The aedeagus of
<taxonomicName id="E5AA458B8CCDC11AD7B0C40E206C138C" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="F7146454CF4293718BAFB9F35E824BD8" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov. is larger than in related species, the carina is broader than in
<taxonomicName id="87B4DA2F86A3A6DC4B1F50B578ABD349" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="D59D2F513E696B7145F3285E32D2D97A" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="F9E44223B6C7DD19A1E896A2963BDD36" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="DFF7EFB4075CB55C79473E03BBFCF94A" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
. In the female genitalia: the apophyses posteriores are broader and stronger (in
<taxonomicName id="5BD0FB067ECFCE678CE23F804AB3E353" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="25F00BFB47C8C57A00C30488CB5423FA" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="EA2D37C1B9FF9A74F893C754410DB140" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="2A81FE659336E58828D77A17AD1FA2B8" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
they are slender); the antrum is more massive than in both species, the ductus bursae has two strong sclerotised bands (in
<taxonomicName id="A3A1ED58DEB48C2EE5AB44658BB3A283" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="6F6C217B3934F0A31A88550A6753C729" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
they are broader and longer, not strongly sclerotised; in
<taxonomicName id="62A4B0B91CCF48DE06801036FF8A61C2" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="D5D02DA9E88288256BA788A025684C72" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
they are 2
<normalizedToken id="5333272C2FB91E0CA9800D031401B519" originalValue="×">x</normalizedToken>
shorter than in
<taxonomicName id="EC53E596F91B3851A183D175DBFA710E" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="53C7E7CE2F3B6188540F0AAF511F6134" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7F0080CB57FB2D4707D85A57D84ADA91" pageId="0" pageNumber="35" type="description">
<paragraph id="7451BE2EE5BEA5F338BC66ED3FFAFB20" pageId="0" pageNumber="35">Description.</paragraph>
<paragraph id="043614EDD5B3ECF8CEA1C8888DFF9879" pageId="0" pageNumber="35">
<emphasis id="366F09BB8925DB1A1395555965BA9632" bold="true" italics="true" pageId="0" pageNumber="35">Adult</emphasis>
(Figs
<figureCitation id="159B8C25579A1AF8764A3378EC196D9A" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">1</figureCitation>
,
<figureCitation id="849C7CD787AF45FD4011D18D15A3557C" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">2</figureCitation>
). Wingspan 30-32 mm. Head dark brown to black; labial palps brown; thorax black to brownish black; abdomen dark brown. Forewing ground colour blackish brown to smoky black; basal line black, double; antemedial line wavy, double, black, distinctly excurved at inner margin; median line very indistinct; postmedial line double, brownish black, excurved; subterminal line wavy and slightly black; terminal line black, dotted line; orbicular spot irregular and deep black; reniform spot crescent-shaped. Hindwing pale greyish white; discal spot arc-shaped, indistinct; vein brown; terminal line same as in forewing; fringe grey.
<emphasis id="D582AEB6054E391D8EA00E48CBFF426C" bold="true" italics="true" pageId="0" pageNumber="35">Male genitalia</emphasis>
(Figs
<figureCitation id="C066E253D8BF7179DDD14AF2D1875C4D" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">11</figureCitation>
,
<figureCitation id="61B00038D3102600CA3E1D8DDCAF3493" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">12</figureCitation>
). Uncus flat, from 1/3 of shovel-shaped base apically. Tegumen broad, ~ 2
<normalizedToken id="30F7CD25780EBC503642D68085B28BC5" originalValue="×">x</normalizedToken>
the length of uncus. Juxta tongue-shaped anteriorly, large and duck palm-shaped posteriorly. Valva bifurcated, basally broad; sacculus strongly sclerotised, sacculus process flat, tongue-shaped, rounded, with fold; harpe club-shaped, strongly sclerotised, exceed costal margin; cucullus densely setose. Saccus long U-shaped, sclerotised. Aedeagus narrow from caecum to carina and strongly sclerotised; caecum curved, with knife-shaped process at dorsal part. Vesica membranous, multiple median diverticula extending with a small cornutus.
<emphasis id="ABF7A0FFF626E20F43E2B98572A84D92" bold="true" italics="true" pageId="0" pageNumber="35">Female genitalia</emphasis>
(Fig.
<figureCitation id="1CF7079897D02097C301002AEE1F2D78" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">23</figureCitation>
). Papillae anales broad, slightly sclerotised. Apophyses anteriores ~ 1/4
<normalizedToken id="382A54146C14AD7E91CCD8CAD5716106" originalValue="×">x</normalizedToken>
the length of apophyses posteriores. Ostium bursae broad, flat, antevaginal plate concave, strongly sclerotised. Ductus bursae flat, moderately sclerotised, with two strongly sclerotised irregular bands. Appendix bursae slightly sclerotised, wrinkled. Corpus bursae long, ~ 4
<normalizedToken id="62A33066AC499E934A81495B192CB58E" originalValue="×">x</normalizedToken>
length of ductus bursae, with longitudinal wrinkles, and two slender longitudinal signa bands.
</paragraph>
</subSubSection>
<subSubSection id="1553590FD9A0E227BFE0A64FC5832E42" pageId="0" pageNumber="35" type="etymology">
<paragraph id="E6BCA0D95873ACF4280887E5277D1984" pageId="0" pageNumber="35">Etymology.</paragraph>
<paragraph id="A9002B724EE1A45138801F3D32F8B4BB" pageId="0" pageNumber="35">
The name of the new species is derived from the Latin word
<taxonomicName id="3C9091D343C5E6F677D30965657CCA76" authorityName="Gao, Han &amp; Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa nigricans" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="E37D3AABF47041A0050653D4F5750F0B" italics="true" pageId="0" pageNumber="35">Hermonassa nigricans</emphasis>
</taxonomicName>
that means black.
</paragraph>
</subSubSection>
<subSubSection id="28EE46F188A9CE2C0F07C47B46DBB4E8" pageId="0" pageNumber="35" type="distribution">
<paragraph id="E5B4A27F4D35D8AC22D8A45BF247A507" pageId="0" pageNumber="35">Distribution and biology.</paragraph>
<paragraph id="22E6684392CF29CECA47CA801ACF4A84" pageId="0" pageNumber="35">
<taxonomicName id="20DBEA4A2F5A97439D9EFF73B06D6028" authorityName="Gao, Han &amp; Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa nigricans" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="D752BBF47E0F600B12A15B9F74BFBA14" italics="true" pageId="0" pageNumber="35">Hermonassa nigricans</emphasis>
</taxonomicName>
is distributed in southwest China (Aut. Reg. Xizang) and Nepal. The species is rather common in the grassland and shrubby areas in eastern spurs of the Tibet plateau (Aut. Reg. Xizang) at altitude 2200-4100 m. It occurs sympatrically with its related species
<taxonomicName id="D4518BE6879B757EC955F2CD7F1BAF5A" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="93520AE11C1EF9FE980009BC4B68BB2C" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="4DACC41575303228D61BC0475BB3B7C4" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra." order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra.">
<emphasis id="A509E6CC016BCA6DFFF4BA173793F6EE" italics="true" pageId="0" pageNumber="35">H. kalamantra.</emphasis>
</taxonomicName>
Collecting period range from early August to beginning of November.
</paragraph>
</subSubSection>
<subSubSection id="646CAE6B6A9B07A0921551A591460158" pageId="0" pageNumber="35" type="remarks">
<paragraph id="B979351A1DE9A96147C4113B74EBE7AA" pageId="0" pageNumber="35">Remark.</paragraph>
<paragraph id="31359D89C2579DCFCAD14C562D34D565" pageId="0" pageNumber="35">
The species was reported from Nepal due to a misidentification; the adult and its male genitalia were illustrated by
<bibRefCitation id="F4F75E4CD6F599347301F5E05081D7CA" author="Sugi, S" editor="Haruta, T" journalOrPublisher="Tinea 14 (suppl. 2)" pageId="0" pageNumber="35" pagination="90 - 109" refId="B25" refString="Sugi, S, 1995. Noctuidae: Noctuinae, Hermonassa. In: Haruta, T, Ed., Moths of Nepal IV. Tinea 14 (suppl. 2): 90 - 109" title="Noctuidae: Noctuinae, Hermonassa." volumeTitle="Moths of Nepal IV." year="1995">Sugi (1995)</bibRefCitation>
as
<taxonomicName id="FC4834FFE412A53EBB7C4B99766C318A" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="F607EF2BEF7CC6B60B81F89D997CE200" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
(reproduced in Fig.
<figureCitation id="FC9B5FC55F2F75889016EC21AEF50A0E" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">12</figureCitation>
).
</paragraph>
<caption id="2C117070F28BCD5FA4825F8198BDB513" doi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35" start="Figures 1–10" startId="F1">
<paragraph id="3D8AC862DDDEF1EF064927D0CCD90063" pageId="0" pageNumber="35">
<emphasis id="75742AA25FA6F9998B4D89A970CD25D3" bold="true" pageId="0" pageNumber="35">Figures 1-10.</emphasis>
<taxonomicName id="E76C6E4D49CB2ADE187D27C8B6DC74F4" authorityName="Walker" authorityYear="1865" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="1FB38BCEF950ED86C0A910F6AFF97347" italics="true" pageId="0" pageNumber="35">Hermonassa</emphasis>
</taxonomicName>
spp., adults
<emphasis id="9E724FBF1A25402EEDEB09F17D3964EB" bold="true" pageId="0" pageNumber="35">1</emphasis>
<taxonomicName id="9BB03D028013C0D026366ECDA32F6BFF" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="E73FB04784209FA07644C73309276806" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov., ♂, holotype, China
<emphasis id="6C324936BFB35922A25E7EF4F13E7446" bold="true" pageId="0" pageNumber="35">2</emphasis>
Ditto, ♀, paratype
<emphasis id="9D84CFD3E111B90C94D67A875BA6CD8B" bold="true" pageId="0" pageNumber="35">3</emphasis>
<taxonomicName id="E6E9B32E84B9F29AD8D58835A591D1A1" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="E1F5FC2944FDF01AD495E7819A6F1D44" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, ♂, holotype, China (ZFMK)
<emphasis id="E30AC70A985946C4FC74D8885681BA37" bold="true" pageId="0" pageNumber="35">4</emphasis>
<taxonomicName id="DDAB2BF0687AB1ADE26BC6B2DD7A1047" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="C84841D3ACB8EC84573ED6D2A2AF0CAB" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, ♀, China
<emphasis id="FD57671072EF62B1BD178D06D8D483EC" bold="true" pageId="0" pageNumber="35">5</emphasis>
<taxonomicName id="049B5C107C99C15A05E835887A81A6EC" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="AC8050E7595E03183B784F41D1D3B71C" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
, ♂, holotype, Nepal (after
<bibRefCitation id="318992CF73B9796F58AD3ADBE811A49A" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="7A9B8DB765B164B0A713328355E482FD" originalValue="Kovács">Kovacs</normalizedToken>
et al. 2018
</bibRefCitation>
)
<emphasis id="89FEB130DF33AAF556EBCCA71E337229" bold="true" pageId="0" pageNumber="35">6</emphasis>
<taxonomicName id="F80493F88E014E42B7660A2C270F68A0" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="FEC7A066BF602BAC126693B4256F9DE4" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
, ♂, China
<emphasis id="54CC0A65848824F6F6652B596F198098" bold="true" pageId="0" pageNumber="35">7</emphasis>
<taxonomicName id="23B45930D10A9913BA5802E4A0DED0D5" family="Noctuidae" kingdom="Animalia" lsidName="H. conusa" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="conusa">
<emphasis id="E7D6AC582074FA651E6F14022F3FD5DD" italics="true" pageId="0" pageNumber="35">H. conusa</emphasis>
</taxonomicName>
sp. nov., ♂, holotype, China
<emphasis id="6F72445A2523DA91C9042917E2BDA96C" bold="true" pageId="0" pageNumber="35">8</emphasis>
Ditto, ♀, paratype, China
<emphasis id="6934EDA92AC1E901466491DA022C09D7" bold="true" pageId="0" pageNumber="35">9</emphasis>
<taxonomicName id="4BFC39A0FE74AC7CC44A8062EBBC809D" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
<emphasis id="F1D6EEACED9C787F8035890747AE045B" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
</taxonomicName>
, ♂, holotype, China (NACR)
<emphasis id="E9B5007E0E0795E4E6CC3FA42A1CA646" bold="true" pageId="0" pageNumber="35">10</emphasis>
<taxonomicName id="799ADECBBAC538A24701291AC206551F" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
<emphasis id="3BC247A0283C523AA99185C7192B1EB8" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
</taxonomicName>
, ♂, holotype, India (NIAES).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

284
data/89/BC/4F/89BC4FB1F79A9AE4FCF4FECF16B116B5.xml Normal file
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.770.22470" ID-GBIF-Dataset="5abea0d1-6d9e-41b0-8839-9e7d84e7e61e" ID-PMC="PMC6041365" ID-Pensoft-Pub="1313-2970-770-117" ID-Pensoft-UUID="FF87AD70FFD3FFBBFFEFFFAEE805FFFD" ID-PubMed="30002592" ID-Zenodo-Dep="1310203" ID-ZooBank="1402C2E76F9448838F60E34CF9B3D348" ModsDocID="1313-2970-770-117" checkinTime="1530781158320" checkinUser="pensoft" docAuthor="Li, Bing, Zhao, Zhe, Zhang, Chuntian &amp; Li, Shuqiang" docDate="2018" docId="89BC4FB1F79A9AE4FCF4FECF16B116B5" docLanguage="en" docName="ZooKeys 770: 117-135" docOrigin="ZooKeys 770" docPubDate="2018-07-04" docSource="http://dx.doi.org/10.3897/zookeys.770.22470" docTitle="Sinodraconarius yui Z. Zhao &amp; S. Li 2018, sp. n." docType="treatment" docUuid="1BB98FE5-7F5A-4CA3-8DDF-1D1C79E52CD8" docUuidSource="ZooBank" docVersion="4" id="FF87AD70FFD3FFBBFFEFFFAEE805FFFD" lastPageNumber="117" masterDocId="FF87AD70FFD3FFBBFFEFFFAEE805FFFD" masterDocTitle="Sinodraconarius gen. n., a new genus of Coelotinae spiders from Southwest China (Araneae, Agelenidae)" masterLastPageNumber="135" masterPageNumber="117" pageNumber="117" updateTime="1668165945013" updateUser="ExternalLinkService">
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<mods:title>Sinodraconarius gen. n., a new genus of Coelotinae spiders from Southwest China (Araneae, Agelenidae)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Li, Bing</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang, Liaoning 110034, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Zhe</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhang, Chuntian</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang, Liaoning 110034, China</mods:affiliation>
<mods:nameIdentifier type="email">chuntianzhang@aliyun.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:number>2018-07-04</mods:number>
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<subSection pageId="0" pageNumber="117" type="taxonomy">
<treatment ID-GBIF-Taxon="182229667" LSID="urn:lsid:zoobank.org:act:1BB98FE5-7F5A-4CA3-8DDF-1D1C79E52CD8" httpUri="http://treatment.plazi.org/id/89BC4FB1F79A9AE4FCF4FECF16B116B5" lastPageNumber="117" pageId="0" pageNumber="117">
<subSubSection pageId="0" pageNumber="117" type="nomenclature">
<paragraph pageId="0" pageNumber="117">
<taxonomicName LSID="http://zoobank.org/1BB98FE5-7F5A-4CA3-8DDF-1D1C79E52CD8" authority="Z. Zhao &amp; S. Li" authorityName="Z. Zhao &amp; S. Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Sinodraconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinodraconarius yui" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="yui" status="sp. n.">Sinodraconarius yui Z. Zhao &amp; S. Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="117">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Left male palp of Sinodraconarius yui sp. n., holotype. A Prolateral view B Ventral view C Retrolateral view. Scale bar equal for A, B, C." figureDoi="10.3897/zookeys.770.22470.figure9" httpUri="https://binary.pensoft.net/fig/214755" pageId="0" pageNumber="117">Figs 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigyne and habitus of Sinodraconarius yui sp. n. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bar equal for D and E." figureDoi="10.3897/zookeys.770.22470.figure10" httpUri="https://binary.pensoft.net/fig/214756" pageId="0" pageNumber="117">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Localities of Sinodraconarius species in the Southwest China. 1 S. cawarongensis sp. n. 2 S. muruoensis sp. n. 3 S. patellabifidus 4 S. sangjiuensis sp. n. 5 S. yui sp. n." figureDoi="10.3897/zookeys.770.22470.figure11" httpUri="https://binary.pensoft.net/fig/214757" pageId="0" pageNumber="117">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="reference_group">
<paragraph pageId="0" pageNumber="117">
<taxonomicName authority="Li &amp; Zhao &amp; Zhang &amp; Li, 2018" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelotes himalayaensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="himalayaensis" status="sp. n.">Coelotes himalayaensis</taxonomicName>
<bibRefCitation author="Hu, JL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="117" refId="B3" refString="Hu, JL, 2001. Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp." title="Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp." year="2001">Hu 2001</bibRefCitation>
: 134, fig. 45.3-4 (♂ only, ♀ mismatched).
</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authority="Li &amp; Zhao &amp; Zhang &amp; Li, 2018" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Draconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Draconarius himalayaensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="himalayaensis" status="sp. n.">Draconarius himalayaensis</taxonomicName>
<bibRefCitation author="Wang, XP" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="117" pagination="499 - 662" refId="B8" refString="Wang, XP, 2003. Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences 54: 499 - 662" title="Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae)." volume="54" year="2003">Wang 2003</bibRefCitation>
: 534 (♂ only, ♀ mismatched);
<bibRefCitation author="Zhu, MS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="117" refId="B13" refString="Zhu, MS, Wang, XP, Zhang, ZS, 2017. Fauna Sinica: Invertebrata Vol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing" title="Fauna Sinica: Invertebrata Vol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae." year="2017">Zhu et al. 2017</bibRefCitation>
: 289, fig. 166C-D (♂ only, ♀ mismatched)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="type material">
<paragraph pageId="0" pageNumber="117">Type material.</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Holotype</emphasis>
♂ (IZCAS): China: Tibet: Nyingchi: Segula Mountain,
<geoCoordinate degrees="29.605017" direction="north" orientation="latitude" precision="1" value="29.605017">N29.605017°</geoCoordinate>
,
<geoCoordinate degrees="94.609117" direction="east" orientation="longitude" precision="1" value="94.609116">E94.609117°</geoCoordinate>
, 4184
<normalizedToken originalValue="±">+/-</normalizedToken>
4 m, 11.X.2010, Hao Yu leg.
<emphasis bold="true" pageId="0" pageNumber="117">Paratypes</emphasis>
: 2♂♂ (IZCAS): same data as holotype; 1♀ (IZCAS): China: Tibet: Nyingchi: Positive face of Segula Mountain,
<geoCoordinate degrees="29.607583" direction="north" orientation="latitude" precision="1" value="29.607582">N29.607583°</geoCoordinate>
,
<geoCoordinate degrees="94.608767" direction="east" orientation="longitude" precision="1" value="94.608765">E94.608767°</geoCoordinate>
, 4190
<normalizedToken originalValue="±">+/-</normalizedToken>
13 m, 12.VIII.2010, Hao Yu leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="etymology">
<paragraph pageId="0" pageNumber="117">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="117">The specific name is after Hao Yu, the collector of specimens used in this study; noun (name) in genitive case.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="diagnosis">
<paragraph pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="117">
The males are similar to these of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n. by the ventral branch of the patellar apophysis larger than dorsal one and the apex of the conductor pointed and bent retrolaterally, but can be easily distinguished from
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n. by the short palp (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Left male palp of Sinodraconarius sangjiuensis sp. n., holotype. A Prolateral view B Ventral view C Retrolateral view. Scale bar: equal for A, B, C." figureDoi="10.3897/zookeys.770.22470.figure1" httpUri="https://binary.pensoft.net/fig/214747" pageId="0" pageNumber="117">1A-C</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Left male palp of Sinodraconarius yui sp. n., holotype. A Prolateral view B Ventral view C Retrolateral view. Scale bar equal for A, B, C." figureDoi="10.3897/zookeys.770.22470.figure9" httpUri="https://binary.pensoft.net/fig/214755" pageId="0" pageNumber="117">9A-C</figureCitation>
). The females are similar to
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n. by having the apex of the septum tapering but can be easily distinguished from
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n. by the epigynal hoods located anterolaterally
<emphasis italics="true" pageId="0" pageNumber="117">vs.</emphasis>
mediolaterally in
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n. (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epigyne and habitus of Sinodraconarius sangjiuensis sp. n. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bar equal for D and E." figureDoi="10.3897/zookeys.770.22470.figure2" httpUri="https://binary.pensoft.net/fig/214748" pageId="0" pageNumber="117">2A-B</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigyne and habitus of Sinodraconarius yui sp. n. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bar equal for D and E." figureDoi="10.3897/zookeys.770.22470.figure10" httpUri="https://binary.pensoft.net/fig/214756" pageId="0" pageNumber="117">10A-B</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1310329" doi="10.3897/zookeys.770.22470.figure9" httpUri="https://binary.pensoft.net/fig/214755" pageId="0" pageNumber="117" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 9.</emphasis>
Left male palp of
<taxonomicName authorityName="Z. Zhao &amp; S. Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Sinodraconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinodraconarius yui" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="yui">
<emphasis italics="true" pageId="0" pageNumber="117">Sinodraconarius yui</emphasis>
</taxonomicName>
sp. n., holotype.
<emphasis bold="true" pageId="0" pageNumber="117">A</emphasis>
Prolateral view
<emphasis bold="true" pageId="0" pageNumber="117">B</emphasis>
Ventral view
<emphasis bold="true" pageId="0" pageNumber="117">C</emphasis>
Retrolateral view. Scale bar equal for
<emphasis bold="true" pageId="0" pageNumber="117">A, B, C</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="description">
<paragraph pageId="0" pageNumber="117">Description.</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Male (holotype).</emphasis>
Total length 7.59. Carapace 4.00 long, 2.60 wide. Abdomen 3.59 long, 2.56 wide. Eye sizes and interdistances: AME 0.09, ALE 0.20, PME 0.14, PLE 0.19; AME-AME 0.08, AME-ALE 0.05, AME-PME 0.16, ALE-PLE 0, PME-PME 0.09, PME-PLE 0.13. Leg measurements: I 12.15 (4.06, 4.00, 2.68, 1.41); II 10.22 (3.44, 3.50, 2.03, 1.25); III 9.99 (3.28, 3.00, 2.40, 1.31); IV 12.82 (4.06, 3.80, 3.40, 1.56). Palp: ventral branch of patellar apophysis larger than dorsal one; anterior 1/5 part of RTA extending beyond the tibia; LTA obvious; cymbial furrow less than 1/4 of cymbium length; the apex of conductor concave and spiral; median apophysis conspicuous, finger-like, the apex of the median apophysis expanded and open; dorsal conductor apophysis broad, the visible part (between conductor and tegulum) hidden by conductor in ventral view; embolus beginning at position 10
<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Left male palp of Sinodraconarius yui sp. n., holotype. A Prolateral view B Ventral view C Retrolateral view. Scale bar equal for A, B, C." figureDoi="10.3897/zookeys.770.22470.figure9" httpUri="https://binary.pensoft.net/fig/214755" pageId="0" pageNumber="117">9A-C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Female (paratype).</emphasis>
Total length 7.89. Carapace 3.40 long, 2.40 wide. Abdomen 4.49 long, 2.40 wide. Eye sizes and interdistances: AME 0.10, ALE 0.18, PME 0.15, PLE 0.15; AME-AME 0.11, AME-ALE 0.06, AME-PME 0.15, ALE-PLE 0, PME-PME 0.10, PME-PLE 0.15. Leg measurements: I 9.17 (3.58, 2.88, 1.75, 0.96); II 8.66 (3.07, 2.88, 1.75, 0.96); III 8.47 (3.20, 2.56, 1.75, 0.96); IV 11.05 (3.59, 3.52, 2.50, 1.44). Epigyne: apex of the V-shaped septum tapering; hoods located laterally; atrium two times wider than long, occupying approx. 1/4 of epigynal plate; receptacles spaced by the width of a receptacle; the head of the receptacles broad and short, located anteriorly (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigyne and habitus of Sinodraconarius yui sp. n. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bar equal for D and E." figureDoi="10.3897/zookeys.770.22470.figure10" httpUri="https://binary.pensoft.net/fig/214756" pageId="0" pageNumber="117">10A-B</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1310311" doi="10.3897/zookeys.770.22470.figure10" httpUri="https://binary.pensoft.net/fig/214756" pageId="0" pageNumber="117" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 10.</emphasis>
Epigyne and habitus of
<taxonomicName authorityName="Z. Zhao &amp; S. Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Sinodraconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinodraconarius yui" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="yui">
<emphasis italics="true" pageId="0" pageNumber="117">Sinodraconarius yui</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="117">A</emphasis>
Epigyne, ventral
<emphasis bold="true" pageId="0" pageNumber="117">B</emphasis>
Vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="117">C</emphasis>
Male habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="117">D</emphasis>
Female habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="117">E</emphasis>
Female habitus, ventral. Scale bar equal for
<emphasis bold="true" pageId="0" pageNumber="117">D</emphasis>
and
<emphasis bold="true" pageId="0" pageNumber="117">E</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="comments">
<paragraph pageId="0" pageNumber="117">Comments.</paragraph>
<paragraph pageId="0" pageNumber="117">
The male of the new species was first described by
<bibRefCitation author="Hu, JL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="117" refId="B3" refString="Hu, JL, 2001. Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp." title="Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp." year="2001">Hu (2001)</bibRefCitation>
as the allotype of
<taxonomicName authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelotes himalayaensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="himalayaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Coelotes himalayaensis</emphasis>
</taxonomicName>
and later transferred from
<taxonomicName authorityName="Blackwall" authorityYear="1841" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelotes" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Coelotes</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Ovtchinnikov" authorityYear="1999" class="Arachnida" family="Agelenidae" genus="Draconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Draconarius" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Draconarius</emphasis>
</taxonomicName>
by
<bibRefCitation author="Wang, XP" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="117" pagination="499 - 662" refId="B8" refString="Wang, XP, 2003. Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences 54: 499 - 662" title="Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae)." volume="54" year="2003">Wang (2003)</bibRefCitation>
. However, we found that the male and female of
<taxonomicName lsidName="D. himalayaensis" pageId="0" pageNumber="117" rank="species" species="himalayaensis">
<emphasis italics="true" pageId="0" pageNumber="117">D. himalayaensis</emphasis>
</taxonomicName>
were mismatched. Therefore, the male is established as a new species here, and both sexes are described.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="variation">
<paragraph pageId="0" pageNumber="117">Variation.</paragraph>
<paragraph pageId="0" pageNumber="117">Total length of males (n = 3) 7.05-7.59.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="distribution">
<paragraph pageId="0" pageNumber="117">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="117">
Known only from Nyingchi, Tibet (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Localities of Sinodraconarius species in the Southwest China. 1 S. cawarongensis sp. n. 2 S. muruoensis sp. n. 3 S. patellabifidus 4 S. sangjiuensis sp. n. 5 S. yui sp. n." figureDoi="10.3897/zookeys.770.22470.figure11" httpUri="https://binary.pensoft.net/fig/214757" pageId="0" pageNumber="117">11</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1310313" doi="10.3897/zookeys.770.22470.figure11" httpUri="https://binary.pensoft.net/fig/214757" pageId="0" pageNumber="117" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 11.</emphasis>
Localities of
<taxonomicName authorityName="Z. Zhao &amp; S. Li" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Sinodraconarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinodraconarius" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Sinodraconarius</emphasis>
</taxonomicName>
species in the Southwest China.
<emphasis bold="true" pageId="0" pageNumber="117">1</emphasis>
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. cawarongensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="cawarongensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. cawarongensis</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="117">2</emphasis>
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. muruoensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="muruoensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. muruoensis</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="117">3</emphasis>
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. patellabifidus" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="patellabifidus">
<emphasis italics="true" pageId="0" pageNumber="117">S. patellabifidus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">4</emphasis>
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. sangjiuensis" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="sangjiuensis">
<emphasis italics="true" pageId="0" pageNumber="117">S. sangjiuensis</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="117">5</emphasis>
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="S. yui" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="yui">
<emphasis italics="true" pageId="0" pageNumber="117">S. yui</emphasis>
</taxonomicName>
sp. n.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
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Girault, 1913
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Blood &amp; Kryger, 1928
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Novicki, 1934
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Girault, 1938
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Novicky, 1946
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De Santis, 1957
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De Santis, 1957
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