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<document ENCODING="UTF8" ID-DOI="http://doi.org/10.5281/zenodo.6294087" ID-GBIF-Dataset="3fc50716-3f1c-4903-b672-28bcaeaee724" ID-GBIF-Taxon="125136287" ID-Zenodo-Dep="6294087" checkinTime="1439764468610" checkinUser="donat" docAuthor="Goode, G. B. &amp; Bean, T. H." docDate="1883" docId="895C6AB1F2AFAF874EBFBA249F1AF8E5" docLanguage="en" docName="goode_Bean_1883_188042100200024991.htm.xml" docOrigin="Bulletin of the Museum of Comparative Zoology at Harvard College 10 (5)" docTitle="Laemonema barbatula" docType="treatment" docVersion="6" lastPageNumber="206" masterDocId="5B7296A6F34C923B9F5FBB9D7E3D1B3E" masterDocTitle="Reports on the results of dredging under the supervision of Alexander Agassiz, on the east coast of the United States, during the summer of 1880, by the U. S. coast survey steamer “ Blake, ” Commander J. R. Bartlett, U. S. N., commanding." masterLastPageNumber="226" masterPageNumber="183" pageNumber="204" updateTime="1645849994371" updateUser="ExternalLinkService">
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<mods:title>Reports on the results of dredging under the supervision of Alexander Agassiz, on the east coast of the United States, during the summer of 1880, by the U. S. coast survey steamer “ Blake, ” Commander J. R. Bartlett, U. S. N., commanding.</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Goode, G. B.</mods:namePart>
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<mods:namePart>Bean, T. H.</mods:namePart>
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<mods:title>Bulletin of the Museum of Comparative Zoology at Harvard College</mods:title>
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<mods:part>
<mods:date>1883</mods:date>
<mods:detail type="volume">
<mods:number>10</mods:number>
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<mods:detail type="issue">
<mods:number>5</mods:number>
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<mods:start>183</mods:start>
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<subSubSection type="nomenclature">
<paragraph pageId="21" pageNumber="204" type="mainText">
18.
<taxonomicName class="Actinopterygii" family="Moridae" genus="Laemonema" kingdom="Animalia" order="Gadiformes" pageId="21" pageNumber="204" phylum="Chordata" rank="species" species="barbatula">Laemonema barbatula</taxonomicName>
, new specics.
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageId="21" pageNumber="204" type="mainText">Head contained in body (without caudal) 4?- times; height of body 4.\ times. Diameter of orbit in length of head three times, upper jaw a little more than twice. Barbel half as loner as the diameter of the eve. Vent situated under</paragraph>
<paragraph lastPageId="22" lastPageNumber="205" pageId="21" pageNumber="204" type="mainText">
O * the sixth or seventh rav of second dorsal. Distance of first dorsal from snout
<pageBreakToken pageId="22" pageNumber="205" start="start">equal</pageBreakToken>
to one fourth standard length of body. The base of the first dorsal is half as long as the middle caudal rays; that of the second, slightly more than three times the length of the head. The first dorsal is composed of live rays, the first of which is elongate, three times as long as the middle caudal rays: it extends to the base of the twenty-fourth ray of the second dorsal. Anal fin inserted at a distance from the tip of the snout equal to twice the length of the head, its distance from the insertion of the ventrals being equal to the length of the head. The length of the ventrals is equal to that of the pectorals, their tips not extending to the vent.
</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">Scales small, very thin, deciduous, crowded anteriorly. Lateral line not well defined on the posterior part of the body.</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">Radial formula: 13. VII; D. 5-63; A. 59; F. 19; V. 2.</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">Scales 13-140-31.</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">
Color similar to that of the various species of
<taxonomicName class="Actinopterygii" family="Gadidae" genus="Phycis" kingdom="Animalia" order="Gadiformes" pageId="22" pageNumber="205" phylum="Chordata" rank="genus">Phycis</taxonomicName>
; the dorsal and anal fins have narrow black margins.
</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">The length of the first dorsal ray is very variable in individuals, being shorter in younger specimens.</paragraph>
</subSubSection>
<subSubSection type="diagnosis">
<paragraph pageId="22" pageNumber="205" type="mainText">
This species differs from L. Yctrrellii by its much smaller scales, and from
<taxonomicName class="Actinopterygii" family="Moridae" genus="Laemonema" kingdom="Animalia" order="Gadiformes" pageId="22" pageNumber="205" phylum="Chordata" rank="species" species="robustum">L. robustum</taxonomicName>
by the greater number of rays in the dorsal and anal fins, and its much shorter ventrals.
</paragraph>
</subSubSection>
<subSubSection type="materials_examined">
<paragraph pageId="22" pageNumber="205" type="mainText">Specimens of this species were obtained at the following stations of the steamer &quot; Blake.&quot;</paragraph>
<paragraph pageId="22" pageNumber="205" type="table">Station. 321 315 316 N. Lat. 32° 43' 25&quot; 32° 18' 20&quot; 32° 7' vr. Long. 77° 20' 30&quot; 78° 43' 78° 37' 30&quot; Faihoms. Specimens. 233 1 and part of I aJ - '/ 229 2 3</paragraph>
<paragraph pageId="22" pageNumber="205" type="mainText">A single specimen was also obtained by the Fish Commission steamer &quot; Fish Ilawk&quot; in Lat. 38° 35' N., Long. 73° 13' W., at a depth of 312 fathoms, Oct. 10, 1881 (U. S. Nat. Mus., No. 29*046).</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageId="22" pageNumber="205" type="mainText">MEASUREMENTS.</paragraph>
<paragraph pageId="22" pageNumber="205" type="table">Extreme length ..... Length to base of middle caudal rays Body. Greatest height Greatest width. Height at ventrals Least height of tail. Head. Greatest length Greatest width. Width of interorbital area. Length of snout. Length of upper jaw. Length of mandible Millimeters.. 178 160. 36 18. 32 4. 33 20 7 hr 7, 15 17 20 C</paragraph>
<paragraph pageId="23" pageNumber="206" type="table">
<table border="1" pageId="23" pageNumber="206">
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">
<pageBreakToken pageId="23" pageNumber="206" start="start">Distance</pageBreakToken>
from snout to orbit
</td>
<td pageId="23" pageNumber="206">9</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Diameter of orbit.....</td>
<td pageId="23" pageNumber="206">11</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Dorsal (first). Distance from snout.</td>
<td pageId="23" pageNumber="206">. 40</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of base......</td>
<td pageId="23" pageNumber="206">9</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of longest ray.....</td>
<td pageId="23" pageNumber="206">. 54</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">(second). Length of base.</td>
<td pageId="23" pageNumber="206">105</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of first ray .....</td>
<td pageId="23" pageNumber="206">. 14</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of longest ray .....</td>
<td pageId="23" pageNumber="206">17</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Anal. Distance from snout ....</td>
<td pageId="23" pageNumber="206">. 65</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of base ......</td>
<td pageId="23" pageNumber="206">86</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of first ray .....</td>
<td pageId="23" pageNumber="206">7</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of longest ray.</td>
<td pageId="23" pageNumber="206">16</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length of last ray .....</td>
<td pageId="23" pageNumber="206">3</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Caudal. Length of middle rays.</td>
<td pageId="23" pageNumber="206">18</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Pectoral. Distance from snout ....</td>
<td pageId="23" pageNumber="206">. 36</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length ........</td>
<td pageId="23" pageNumber="206">25</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Ventral. Distance from snout ....</td>
<td pageId="23" pageNumber="206">. 30</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Length ........</td>
<td pageId="23" pageNumber="206">25</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Branchiostegals.......</td>
<td pageId="23" pageNumber="206">. VII</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Dorsal .........</td>
<td pageId="23" pageNumber="206">. 5-63</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Anal .........</td>
<td pageId="23" pageNumber="206">. 59</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Caudal .........</td>
<td isEmpty="true" pageId="23" pageNumber="206">-</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Pectoral ........</td>
<td pageId="23" pageNumber="206">19</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Ventral .........</td>
<td pageId="23" pageNumber="206">2</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Number of scales in lateral line ....</td>
<td pageId="23" pageNumber="206">ca. 140</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Number of transverse rows above lateral line</td>
<td pageId="23" pageNumber="206">. 13</td>
</tr>
<tr pageId="23" pageNumber="206">
<td pageId="23" pageNumber="206">Number of transverse rows below lateral line</td>
<td pageId="23" pageNumber="206">31</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:affiliation>Museum fuer Naturkunde - Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115, Berlin, Germany &amp; Research Centre for Deep-sea, Indonesian Institute of Science (LIPI), Jl Y Syaranamual, Poka, Tlk. Ambon, Kota Ambon, Maluku, Indonesia &amp; Institute of Geological Sciences, Freie Universitaet Berlin, Malteserstrasse 74 - 100 Building C and D, 12249, Berlin, Germany</mods:affiliation>
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<mods:namePart>Jones, Diana S.</mods:namePart>
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<treatment ID-GBIF-Taxon="168143945" LSID="urn:lsid:plazi:treatment:895C82623F6451E1B57AE17155EF4EEC" httpUri="http://treatment.plazi.org/id/895C82623F6451E1B57AE17155EF4EEC" lastPageNumber="649" pageId="0" pageNumber="649">
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<taxonomicName LSID="895C8262-3F64-51E1-B57A-E17155EF4EEC" authority="(Hoek, 1883)" baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Hexanauplia" family="Scalpellidae" genus="Diotascalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Diotascalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">Diotascalpellum rubrum (Hoek, 1883)</taxonomicName>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diotascalpellum rubrum (Hoek, 1883) (MNHN-IU- 2019 - 4925). a. left lateral view showing the capitulum and peduncle; b. right lateral view showing the capitulum and peduncle; c. rostral view showing the capitulum and peduncle; d. carinal view. Scale bar: 5 mm (a-d)." figureDoi="10.3897/zse.96.55733.figure9" httpUri="https://binary.pensoft.net/fig/457263" pageId="0" pageNumber="649">Figure 9</figureCitation>
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<taxonomicName authorityName="Hoek" authorityYear="1883" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">Scalpellum rubrum</taxonomicName>
Hoek, 1883: 91, pl. IV fig. 18. -
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Bulletin of the United States Bureau of Fisheries, Washington" pageId="0" pageNumber="649" refId="B131" refString="Pilsbry, HA, 1911. Barnacles of Japan and Bering Sea. Bulletin of the United States Bureau of Fisheries, Washington, XXIX, 1909, 739: 61-84. [pls. VIII-XVII]" title="Barnacles of Japan and Bering Sea. Bulletin of the United States Bureau of Fisheries, Washington, XXIX, 1909, 739: 61 - 84. [pls. VIII-XVII]" year="1911">Pilsbry 1911</bibRefCitation>
: 62, text-fig. 1, pl. VIII, figs 1-4. -
<bibRefCitation DOI="https://doi.org/10.1080/00222931808562293" author="Calman, WT" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="649" pagination="96 - 124" refId="B36" refString="Calman, WT, 1918b. On barnacles of the genus Scalpellum from deep-sea telegraph cables. Annals and Magazine of Natural History 9 (1): 96 - 124, DOI: https://doi.org/10.1080/00222931808562293" title="On barnacles of the genus Scalpellum from deep-sea telegraph cables." url="https://doi.org/10.1080/00222931808562293" volume="9" year="1918 b">Calman 1918b</bibRefCitation>
: 122-123. -
<bibRefCitation author="Broch, H" journalOrPublisher="Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening (Copenhagen)" pageId="0" pageNumber="649" pagination="215 - 358" refId="B21" refString="Broch, H, 1922. Studies on Pacific Cirripedes. Papers from Dr Th. Mortensen's Pacific Expedition 1914-1916 (10). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening (Copenhagen) 73: 215 - 358" title="Studies on Pacific Cirripedes. Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916 (10)." volume="73" year="1922">Broch 1922</bibRefCitation>
: 237. -
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1927.tb07432.x" author="Nilsson-Cantell, CA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="649" pagination="743 - 790" refId="B116" refString="Nilsson-Cantell, CA, 1927. Some barnacles in the British Museum (Nat. Hist.). Proceedings of the Zoological Society of London 1927 (3): 743 - 790, DOI: https://doi.org/10.1111/j.1469-7998.1927.tb07432.x" title="Some barnacles in the British Museum (Nat. Hist.)." url="https://doi.org/10.1111/j.1469-7998.1927.tb07432.x" volume="1927" year="1927">Nilsson-Cantell 1927</bibRefCitation>
: 745-747, text-fig. 2. - 1931: 2.
</paragraph>
<paragraph pageId="0" pageNumber="649">
<taxonomicName baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Maxillopoda" family="Scalpellidae" genus="Trianguloscalpellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trianguloscalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">Trianguloscalpellum rubrum</taxonomicName>
. -
<bibRefCitation author="Zevina, GB" journalOrPublisher="Fauna SSSR" pageId="0" pageNumber="649" pagination="1 - 398" refId="B187" refString="Zevina, GB, 1981. Barnacles of the Suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. I: Family Scalpellidae. Fauna SSSR 127: 1 - 398" title="Barnacles of the Suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. I: Family Scalpellidae." volume="127" year="1981">Zevina 1981</bibRefCitation>
: 316-317, fig. 240. -
<bibRefCitation author="Rosell, NC" journalOrPublisher="Memoires ORSTOM" pageId="0" pageNumber="649" refId="B148" refString="Rosell, NC, 1986. Barnacles. Guide to the Flora and Fauna of the Philippines (Vol. 7). National Resources Management Center, Ministry of Natural Resources, The Philippines, 79-164." title="Barnacles. Guide to the Flora and Fauna of the Philippines (Vol. 7). National Resources Management Center, Ministry of Natural Resources, The Philippines, 79 - 164." year="1986">Rosell 1986</bibRefCitation>
: 89, text-fig. 1a. - 1991: 22, fig. 1f, g. -
<bibRefCitation author="Huang, ZG" editor="Huang, ZG" journalOrPublisher="China Ocean Press, Beijing" pageId="0" pageNumber="649" pagination="516 - 523" refId="B83" refString="Huang, ZG, 1994. Cirripedia. In: Huang, ZG, Ed., Marine Species and their Distribution in China's Seas. China Ocean Press, Beijing: 516 - 523" title="Cirripedia." volumeTitle="Marine Species and their Distribution in China's Seas." year="1994">Huang 1994</bibRefCitation>
: 517. - 2001: 318. -
<bibRefCitation author="Jones, DS" journalOrPublisher="National University of Singapore" pageId="0" pageNumber="649" pagination="233 - 308" refId="B90" refString="Jones, DS, Hewitt, MA, Sampey, A, 2001. A checklist of the Cirripedia of the South China Sea. Raffles Bulletin of Zoology. National University of Singapore 48: 233 - 308" title="A checklist of the Cirripedia of the South China Sea. Raffles Bulletin of Zoology." volume="48" year="2001">Jones et al. 2001</bibRefCitation>
: 254. -
<bibRefCitation DOI="https://doi.org/10.1163/20021975-99990147" author="Young, PS" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="649" pagination="456 - 468" refId="B177" refString="Young, PS, 2001. Redescription of Scalpellopsis striatociliata Broch, 1922, with a discussion on its phylogenetic position and the peduncular plate pattern in scalpellids. Journal of Crustacean Biology 21 (2): 456 - 468, DOI: https://doi.org/10.1163/20021975-99990147" title="Redescription of Scalpellopsis striatociliata Broch, 1922, with a discussion on its phylogenetic position and the peduncular plate pattern in scalpellids." url="https://doi.org/10.1163/20021975-99990147" volume="21" year="2001">Young 2001</bibRefCitation>
: 464, fig. 7. -
<bibRefCitation author="Jones, DS" editor="Payri, CE" journalOrPublisher="Institut de Recherche pour le Developpement, Noumea" pageId="0" pageNumber="649" pagination="289 - 294" refId="B87" refString="Jones, DS, 2007. The Cirripedia of New Caledonia. In: Payri, CE, Richer de Forges, B, Eds., Compendium of Marine Species from New Caledonia. Institut de Recherche pour le Developpement, Noumea: 289 - 294" title="The Cirripedia of New Caledonia." volumeTitle="Compendium of Marine Species from New Caledonia." year="2007">Jones 2007</bibRefCitation>
: 292. -
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3804.1.1" author="Shalaeva, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="649" pagination="1 - 63" refId="B154" refString="Shalaeva, K, Boxshall, G, 2014. An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS &quot;Challenger&quot; expedition and deposited in the Natural History Museum, London. Zootaxa 3804 (1): 1 - 63, DOI: https://doi.org/10.11646/zootaxa.3804.1.1" title="An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS &quot; Challenger &quot; expedition and deposited in the Natural History Museum, London." url="https://doi.org/10.11646/zootaxa.3804.1.1" volume="3804" year="2014">Shalaeva and Boxshall 2014</bibRefCitation>
: 49, fig. 34.
</paragraph>
<paragraph pageId="0" pageNumber="649">
<taxonomicName baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Hexanauplia" family="Scalpellidae" genus="Diotascalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Diotascalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">Diotascalpellum rubrum</taxonomicName>
. -
<bibRefCitation DOI="https://doi.org/10.1111/zoj.12321" author="Gale, AS" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="649" pagination="266 - 304" refId="B59" refString="Gale, AS, 2016. Phylogeny of the deep-sea cirripede family Scalpellidae (Crustacea, Thoracica) based on shell capitular plate morphology. Zoological Journal of the Linnean Society 176: 266 - 304, DOI: https://doi.org/10.1111/zoj.12321" title="Phylogeny of the deep-sea cirripede family Scalpellidae (Crustacea, Thoracica) based on shell capitular plate morphology." url="https://doi.org/10.1111/zoj.12321" volume="176" year="2016">Gale 2016</bibRefCitation>
: 297, figs 2A, 6U-Y, 11Q-T, 13K-M, 18A-G.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="materials_examined">
<paragraph pageId="0" pageNumber="649">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="649">
- Kei Islands: 2 specimens, MNHN-IU-2019-4925, Stn. CP 09,
<geoCoordinate degrees="05" direction="south" minutes="23" orientation="latitude" precision="925" value="-5.383333">05°23'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="29" orientation="longitude" precision="925" value="132.48334">132°29'E</geoCoordinate>
, 368-389 m depth, 23 October 1991; 1 specimen, MNHN-IU-2019-4926, Stn. CP 16,
<geoCoordinate degrees="05" direction="south" minutes="17" orientation="latitude" precision="925" value="-5.2833333">05°17'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="50" orientation="longitude" precision="925" value="132.83333">132°50'E</geoCoordinate>
, 330-350 m depth, 24 October 1991, attached to gorgonian.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="diagnosis">
<paragraph pageId="0" pageNumber="649">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="649">Capitulum with 14 plates, smooth, white and reddish coloured; carina simply, strongly bowed, roof slightly convex with umbo at apex; rostrum triangular, distinct; upper latus quadrangular, large; infra-median latus triangular.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="description">
<paragraph pageId="0" pageNumber="649">Description.</paragraph>
<paragraph pageId="0" pageNumber="649">Capitulum flat, rather broad, not covered by distinct membrane. Scutum with occludent margin arched, forming with tergal margin a triangular portion projecting over tergum. Tergum surpassing scutal area with occludent margin almost straight. Upper latus quadrangular, angle at apex between scutal and tergal margins distinctly projecting over scutum. Rostrum small, triangular; rostrolatus very low, quadrangular; infra-median latus small, triangular, umbo at apex; carinal latus larger than other latera with carinal margin arched. Cirrus I with anterior and posterior rami almost same length; cirrus VI with long caudal appendages. Maxillule not notched, with large spine on upper side, cutting edge almost straight; mandible with three large teeth excluding inferior angle. Measurements of two specimens: height of capitulum 12.51-20.57 mm, width 7.39-10.55 mm, thickness 4.09-6.51 mm; length of peduncle 5.10-7.76 mm, width 4.36-6.62 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="distribution">
<paragraph pageId="0" pageNumber="649">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="649">
Java Sea, Indonesia. Pacific: western central and northwest; Philippines; Kagoshima Sea and Sagami Bay, Japan; 133-551 m depth (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3804.1.1" author="Shalaeva, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="649" pagination="1 - 63" refId="B154" refString="Shalaeva, K, Boxshall, G, 2014. An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS &quot;Challenger&quot; expedition and deposited in the Natural History Museum, London. Zootaxa 3804 (1): 1 - 63, DOI: https://doi.org/10.11646/zootaxa.3804.1.1" title="An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS &quot; Challenger &quot; expedition and deposited in the Natural History Museum, London." url="https://doi.org/10.11646/zootaxa.3804.1.1" volume="3804" year="2014">Shalaeva and Boxshall 2014</bibRefCitation>
). In this study,
<taxonomicName baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Hexanauplia" family="Scalpellidae" genus="Diotascalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Diotascalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">
<emphasis italics="true" pageId="0" pageNumber="649">Diotascalpellum rubrum</emphasis>
</taxonomicName>
was found at Kei Islands, Indonesia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="type locality">
<paragraph pageId="0" pageNumber="649">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="649">
Philippines,
<geoCoordinate degrees="12" direction="north" minutes="43" orientation="latitude" precision="925" value="12.716666">12°43'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="10" orientation="longitude" precision="925" value="122.166664">122°10'E</geoCoordinate>
; depth, 180 m and 207 m; bottom, mud (near Luzon) (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12873" author="Hoek, PPC" journalOrPublisher="Challenger, Zoology" pageId="0" pageNumber="649" pagination="1 - 169" refId="B78" refString="Hoek, PPC, 1883. Report on the Cirripedia collected by H.M.S. Challenger during the years 1873-76. Report of the Scientific Results from the Exploratory Voyages of H.M.S. Challenger, Zoology 8: 1 - 169, DOI: https://doi.org/10.5962/bhl.title.12873" title="Report on the Cirripedia collected by H. M. S. Challenger during the years 1873 - 76. Report of the Scientific Results from the Exploratory Voyages of H. M. S." url="https://doi.org/10.5962/bhl.title.12873" volume="8" year="1883">Hoek 1883</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.55733.figure9" httpUri="https://binary.pensoft.net/fig/457263" pageId="0" pageNumber="649" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="649">
<emphasis bold="true" pageId="0" pageNumber="649">Figure 9.</emphasis>
<taxonomicName baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Hexanauplia" family="Scalpellidae" genus="Diotascalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Diotascalpellum rubrum" order="Scalpelliformes" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="Rubrum">
<emphasis italics="true" pageId="0" pageNumber="649">Diotascalpellum rubrum</emphasis>
</taxonomicName>
(Hoek, 1883) (MNHN-IU-2019-4925).
<emphasis bold="true" pageId="0" pageNumber="649">
<normalizedToken originalValue="а">a</normalizedToken>
</emphasis>
. left lateral view showing the capitulum and peduncle;
<emphasis bold="true" pageId="0" pageNumber="649">b.</emphasis>
right lateral view showing the capitulum and peduncle;
<emphasis bold="true" pageId="0" pageNumber="649">c.</emphasis>
rostral view showing the capitulum and peduncle;
<emphasis bold="true" pageId="0" pageNumber="649">d.</emphasis>
carinal view. Scale bar: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="649">a-d</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<paragraph id="014A36BFE101AF00FF0C38C8FE16FF0F" blockId="9.[151,442,151,178]" box="[151,442,151,178]" pageId="9" pageNumber="82">
<heading id="5A0281D3E101AF00FF0C38C8FE16FF0F" bold="true" box="[151,442,151,178]" fontSize="11" level="1" pageId="9" pageNumber="82" reason="1">
<emphasis id="3381EAADE101AF00FF0C38C8FE16FF0F" bold="true" box="[151,442,151,178]" pageId="9" pageNumber="82">
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<emphasis id="3381EAADE101AF00FF0C38C8FEC5FF0C" bold="true" box="[151,361,151,178]" italics="true" pageId="9" pageNumber="82">Dysdera aculeata</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
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The
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<emphasis id="3381EAADE101AF00FF5E3883FEB0FF4A" box="[197,284,220,244]" italics="true" pageId="9" pageNumber="82">aculeata</emphasis>
</taxonomicName>
group,
<emphasis id="3381EAADE101AF00FEF13883FE0EFF4D" box="[362,418,220,243]" italics="true" pageId="9" pageNumber="82">sensu</emphasis>
<treatmentCitation id="805410AEE101AF00FE323883FCD0FF4A" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[425,892,220,244]" page="231" pageId="9" pageNumber="82" year="1988">
<bibRefCitation id="65644B4EE101AF00FE323883FCD0FF4A" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[425,892,220,244]" pageId="9" pageNumber="82" pagination="141 - 269" refId="ref7612" refString="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterranee- nnes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman (1988: 231)</bibRefCitation>
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<emphasis id="3381EAADE101AF00FF0C397EFEBEFE85" bold="true" box="[151,274,289,315]" pageId="9" pageNumber="82">Diagnosis.</emphasis>
The group can be distinguished by the following characters: hexagonal carapace, leg femora equipped with spines, psembolus distinctly longer than tegulum, psembolus with a median crest and acuminate apex (
<bibRefCitation id="65644B4EE101AF00FACB3919FDBCFE3A" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." pageId="9" pageNumber="82" pagination="141 - 269" refId="ref7612" refString="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterranee- nnes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988</bibRefCitation>
).
</paragraph>
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<emphasis id="3381EAADE101AF00FF5C39D2FECBFE19" bold="true" box="[199,359,397,423]" pageId="9" pageNumber="82">Composition.</emphasis>
Eleven species are assigned to this species group:
<taxonomicName id="C6F54D3CE101AF00FC3739D0FB5BFE17" authority="Kroneberg, 1875" authorityName="Kroneberg" authorityYear="1875" box="[940,1271,398,425]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="3381EAADE101AF00FC3739D0FB83FE16" box="[940,1071,398,424]" italics="true" pageId="9" pageNumber="82">D. aculeata</emphasis>
<bibRefCitation id="65644B4EE101AF00FBA339D1FB5BFE17" author="Kroneberg, A." box="[1080,1271,398,425]" pageId="9" pageNumber="82" pagination="1 - 58" refId="ref7864" refString="Kroneberg, A. (1875) Araneae. In: Fedtschenko, A. P. (ed.), Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau, 19 (3), 1 - 58, pl. 1 - 5. [includes 3 maps by Fedtschenko] [in Russian]" type="book chapter" year="1875">Kroneberg, 1875</bibRefCitation>
</taxonomicName>
(Central Asia, probably
<collectingCountry id="79E2762FE101AF00FE9A39EDFE9DFE72" box="[257,305,434,460]" name="Iran" pageId="9" pageNumber="82">Iran</collectingCountry>
, introduced to
<collectingCountry id="79E2762FE101AF00FE4E39EDFD80FE73" box="[469,556,434,461]" name="Croatia" pageId="9" pageNumber="82">Croatia</collectingCountry>
),
<taxonomicName id="C6F54D3CE101AF00FDA539ECFC41FE73" authority="Denis, 1958" authorityName="Denis" authorityYear="1958" box="[574,1005,434,461]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="afghana">
<emphasis id="3381EAADE101AF00FDA539ECFD17FE72" box="[574,699,434,460]" italics="true" pageId="9" pageNumber="82">D. afghana</emphasis>
Denis, 1958 (
<collectingCountry id="79E2762FE101AF00FCCD39EDFC49FE73" box="[854,997,434,461]" name="Afghanistan" pageId="9" pageNumber="82">Afghanistan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FC6C39ECFF5DFE4E" authority="Deeleman-Reinhold, 1988" authorityName="Deeleman-Reinhold" authorityYear="1988" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="3381EAADE101AF00FC6C39ECFBC3FE72" box="[1015,1135,434,460]" italics="true" pageId="9" pageNumber="82">D. arabica</emphasis>
<bibRefCitation id="65644B4EE101AF00FBED39EDFA30FE72" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[1142,1436,434,460]" pageId="9" pageNumber="82" pagination="141 - 269" refId="ref7612" refString="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterranee- nnes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold, 1988</bibRefCitation>
(
<collectingCountry id="79E2762FE101AF00FF043989FF45FE4E" box="[159,233,470,496]" name="Oman" pageId="9" pageNumber="82">Oman</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FF673988FD6BFE4F" authority="L. Koch, 1878" authorityName="L. Koch" authorityYear="1878" box="[252,711,470,497]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="3381EAADE101AF00FF673988FE2AFE4E" box="[252,390,471,496]" italics="true" pageId="9" pageNumber="82">D. concinna</emphasis>
L. Koch, 1878 (
<collectingCountry id="79E2762FE101AF00FDA63989FD12FE4F" box="[573,702,470,497]" name="Azerbaijan" pageId="9" pageNumber="82">Azerbaijan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FD493988FBCCFE4E" authority="Lin, Chang et Li, 2020" authorityName="Lin, Chang et Li" authorityYear="2020" box="[722,1120,470,496]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE101AF00FD493988FCF5FE4E" box="[722,857,470,496]" italics="true" pageId="9" pageNumber="82">D. dushengi</emphasis>
<bibRefCitation id="65644B4EE101AF00FCFB3989FBCCFE4E" author="Lin, Y. J. &amp; Chang, W. - J. &amp; Li, S. Q." box="[864,1120,470,496]" pageId="9" pageNumber="82" pagination="97 - 103" refId="ref7933" refString="Lin, Y. J., Chang, W. - J. &amp; Li, S. Q. (2020) Dysdera dushengi sp. nov., the easternmost species of the spider family Dysderidae (Arachnida: Araneae). Zoological Systematics, 45 (2), 97 - 103. https: // doi. org / 10.11865 / zs. 202014" type="journal article" year="2020">Lin, Chang et Li, 2020</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="79E2762FE101AF00FBF43989FB5FFE4E" box="[1135,1267,470,496]" name="Kazakhstan" pageId="9" pageNumber="82">Kazakhstan</collectingCountry>
and Xinjiang),
<taxonomicName id="C6F54D3CE101AF00FF0C39A4FD34FDAB" authority="(Caporiacco, 1947) (Ethiopia)" baseAuthorityName="Caporiacco" baseAuthorityYear="1947" box="[151,664,506,533]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="dysderoides">
<emphasis id="3381EAADE101AF00FF0C39A4FE92FDAA" box="[151,318,506,532]" italics="true" pageId="9" pageNumber="82">D. dysderoides</emphasis>
(Caporiacco, 1947) (
<collectingCountry id="79E2762FE101AF00FDB039A5FD3CFDAB" box="[555,656,506,533]" name="Ethiopia" pageId="9" pageNumber="82">Ethiopia</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FD3839A4FBD4FDAB" authority="Dunin, 1985" authorityName="Dunin" authorityYear="1985" box="[675,1144,506,533]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="limitanea">
<emphasis id="3381EAADE101AF00FD3839A4FC83FDAA" box="[675,815,506,532]" italics="true" pageId="9" pageNumber="82">D. limitanea</emphasis>
<bibRefCitation id="65644B4EE101AF00FCAE39A5FC6DFDAB" author="Dunin, P. M." box="[821,961,506,533]" pageId="9" pageNumber="82" pagination="114 - 120" refId="ref7702" refString="Dunin, P. M. (1985) The spider family Dysderidae (Aranei, Haplogynae) in the Soviet Central Asia. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 139, 114 - 120. [in Russian with English summary]" type="journal article" year="1985">Dunin, 1985</bibRefCitation>
(
<collectingCountry id="79E2762FE101AF00FC5439A5FBDCFDAB" box="[975,1136,506,533]" name="Turkmenistan" pageId="9" pageNumber="82">Turkmenistan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FB1839A4FEB6FD87" authority="Dunin, 1992" authorityName="Dunin" authorityYear="1992" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE101AF00FB1839A4FAA6FDAA" box="[1155,1290,507,532]" italics="true" pageId="9" pageNumber="82">D. pamirica</emphasis>
<bibRefCitation id="65644B4EE101AF00FA8B39A5FA30FDAA" author="Dunin, P. M." box="[1296,1436,506,532]" pageId="9" pageNumber="82" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin, 1992</bibRefCitation>
(
<collectingCountry id="79E2762FE101AF00FF043A41FEB8FD87" box="[159,276,542,569]" name="Tajikistan" pageId="9" pageNumber="82">Tajikistan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="C6F54D3CE101AF00FEBD3A40FD9FFD87" authority="Dunin, 1985" authorityName="Dunin" authorityYear="1985" box="[294,563,542,569]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="pococki">
<emphasis id="3381EAADE101AF00FEBD3A40FE0CFD86" box="[294,416,542,568]" italics="true" pageId="9" pageNumber="82">D. pococki</emphasis>
<bibRefCitation id="65644B4EE101AF00FE3C3A41FD9FFD87" author="Dunin, P. M." box="[423,563,542,569]" pageId="9" pageNumber="82" pagination="114 - 120" refId="ref7702" refString="Dunin, P. M. (1985) The spider family Dysderidae (Aranei, Haplogynae) in the Soviet Central Asia. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 139, 114 - 120. [in Russian with English summary]" type="journal article" year="1985">Dunin, 1985</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="79E2762FE101AF00FDD93A41FDDDFD86" box="[578,625,542,568]" name="Iran" pageId="9" pageNumber="82">Iran</collectingCountry>
and
<collectingCountry id="79E2762FE101AF00FD3C3A41FCEAFD87" box="[679,838,542,569]" name="Turkmenistan" pageId="9" pageNumber="82">Turkmenistan</collectingCountry>
),
<taxonomicName id="C6F54D3CE101AF00FCC13A40FB09FD87" authority="Kroneberg, 1875" authorityName="Kroneberg" authorityYear="1875" box="[858,1189,542,569]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="3381EAADE101AF00FCC13A40FC4CFD86" box="[858,992,543,568]" italics="true" pageId="9" pageNumber="82">D. tartarica</emphasis>
<bibRefCitation id="65644B4EE101AF00FC7C3A41FB09FD87" author="Kroneberg, A." box="[999,1189,542,569]" pageId="9" pageNumber="82" pagination="1 - 58" refId="ref7864" refString="Kroneberg, A. (1875) Araneae. In: Fedtschenko, A. P. (ed.), Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau, 19 (3), 1 - 58, pl. 1 - 5. [includes 3 maps by Fedtschenko] [in Russian]" type="book chapter" year="1875">Kroneberg, 1875</bibRefCitation>
</taxonomicName>
(Central Asia) and
<taxonomicName id="C6F54D3CE101AF00FA1A3A40FE6EFDE3" authority="Charitonov, 1956" authorityName="Charitonov" authorityYear="1956" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="3381EAADE101AF00FA1A3A40FF55FDE2" italics="true" pageId="9" pageNumber="82">D. zarudnyi</emphasis>
<bibRefCitation id="65644B4EE101AF00FE9B3A1DFE6EFDE3" author="Charitonov, D. E." box="[256,450,578,605]" pageId="9" pageNumber="82" pagination="17 - 39" refId="ref7559" refString="Charitonov, D. E. (1956) Obzor paukov semeistva Dysderidae fauny SSSR [A survey of the family Dysderidae from the USSR fauna]. Uchenye Zapiski Molotovskogo Gosudarstvennogo Universiteta Imeni A. M. Gor'kogo, 10 (1), 17 - 39. [in Russian]" type="journal article" year="1956">Charitonov, 1956</bibRefCitation>
</taxonomicName>
(Central Asia and
<collectingCountry id="79E2762FE101AF00FD083A1DFC8DFDE3" box="[659,801,578,605]" name="Afghanistan" pageId="9" pageNumber="82">Afghanistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E101AF00FF5C3A3AFB6FF938" pageId="9" pageNumber="82" type="discussion">
<paragraph id="014A36BFE101AF00FF5C3A3AFC11FD52" blockId="9.[151,1437,289,748]" pageId="9" pageNumber="82">
<emphasis id="3381EAADE101AF00FF5C3A3AFEE3FDC1" bold="true" box="[199,335,613,639]" pageId="9" pageNumber="82">Comments.</emphasis>
<bibRefCitation id="65644B4EE101AF00FEC33A39FC88FD3E" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[344,804,614,640]" pageId="9" pageNumber="82" pagination="141 - 269" refId="ref7612" refString="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterranee- nnes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman (1988)</bibRefCitation>
did not examine
<taxonomicName id="C6F54D3CE101AF00FC773A38FBD8FD3E" authorityName="Dunin" authorityYear="1992" box="[1004,1140,615,640]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE101AF00FC773A38FBD8FD3E" box="[1004,1140,615,640]" italics="true" pageId="9" pageNumber="82">D. pamirica</emphasis>
</taxonomicName>
and therefore this species was not assigned to any of the species groups proposed by them.
<taxonomicName id="C6F54D3CE101AF00FCE53AD4FBAAFD1A" authorityName="Dunin" authorityYear="1992" box="[894,1030,651,676]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE101AF00FCE53AD4FBAAFD1A" box="[894,1030,651,676]" italics="true" pageId="9" pageNumber="82">D. pamirica</emphasis>
</taxonomicName>
is here considered a member of the
<taxonomicName id="C6F54D3CE101AF00FF0C3AF1FF5BFD76" authorityName="Kroneberg" authorityYear="1875" box="[151,247,686,712]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="3381EAADE101AF00FF0C3AF1FF5BFD76" box="[151,247,686,712]" italics="true" pageId="9" pageNumber="82">aculeata</emphasis>
</taxonomicName>
group based on the proportions and the shape of its bulb: viz., psembolus significantly longer than tegulum, having a median crest and a pointed apex; see figs
<quantity id="C60D9B5AE101AF00FD4B3A8DFCBBFD52" box="[720,791,722,748]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="9" pageNumber="82" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7–9 in</quantity>
<bibRefCitation id="65644B4EE101AF00FC853A8DFC15FD52" author="Dunin, P. M." box="[798,953,722,748]" pageId="9" pageNumber="82" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="014A36BFE101AF00FF0C3E15FB6FF938" blockId="9.[151,1436,1610,1671]" pageId="9" pageNumber="82">
<emphasis id="3381EAADE101AF00FF0C3E15FEF9F9DC" bold="true" box="[151,341,1610,1634]" pageId="9" pageNumber="82">FIGURES 37–40.</emphasis>
Bulb (37–38) and psembolus (39–40) of
<taxonomicName id="C6F54D3CE101AF00FD663E14FC4FF9DD" authorityName=": Marusik" authorityYear="2017" box="[765,995,1611,1635]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="subcylindrica">
<emphasis id="3381EAADE101AF00FD663E14FC4FF9DD" box="[765,995,1611,1635]" italics="true" pageId="9" pageNumber="82">Dysdera subcylindrica</emphasis>
</taxonomicName>
(37, 39) and
<taxonomicName id="C6F54D3CE101AF00FBF63E14FB73F9DD" authorityName="Charitonov" authorityYear="1956" box="[1133,1247,1611,1635]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="arnoldii">
<emphasis id="3381EAADE101AF00FBF63E14FB73F9DD" box="[1133,1247,1611,1635]" italics="true" pageId="9" pageNumber="82">D. arnoldii</emphasis>
</taxonomicName>
(38, 40). 37–40— posterior. Scale bars: 37–38=
<quantity id="C60D9B5AE101AF00FE253E30FDA3F938" box="[446,527,1647,1671]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="82" unit="mm" value="0.2">0.2 mm</quantity>
, 39–40=
<quantity id="C60D9B5AE101AF00FDFD3E31FD68F938" box="[614,708,1646,1670]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
. Abbreviations:
<emphasis id="3381EAADE101AF00FCF23E30FC29F938" box="[873,901,1647,1670]" italics="true" pageId="9" pageNumber="82">Ap</emphasis>
–posterior apophysis,
<emphasis id="3381EAADE101AF00FBFF3E30FBD3F938" box="[1124,1151,1647,1670]" italics="true" pageId="9" pageNumber="82">Ax</emphasis>
–apex.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="CBFAFE2F70CA58EBF5F6ED69C1CF10D6" type="DOI">10.11646/zootaxa.5006.1.10</mods:identifier>
<mods:identifier id="56263C616891C696E0CF2167A9FE4818" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0997FBC6707E83BBF75FBD27E3A9A55" type="Zenodo-Dep">5157005</mods:identifier>
<mods:identifier id="322861BED48315B9AED313DA6CDEEA27" type="ZooBank">251E38AC-1DDD-4AAD-8673-BCC7B0A179F8</mods:identifier>
</mods:mods>
<treatment id="895C87A9E101AF02FF0C3E87FB1AFCAB" LSID="urn:lsid:plazi:treatment:895C87A9E101AF02FF0C3E87FB1AFCAB" httpUri="http://treatment.plazi.org/id/895C87A9E101AF02FF0C3E87FB1AFCAB" lastPageId="11" lastPageNumber="84" pageId="9" pageNumber="82">
<subSubSection id="49EF6534E101AF00FF0C3E87FD3DF94D" box="[151,657,1752,1779]" pageId="9" pageNumber="82" type="nomenclature">
<paragraph id="014A36BFE101AF00FF0C3E87FD3DF94D" blockId="9.[151,718,1752,1816]" box="[151,657,1752,1779]" pageId="9" pageNumber="82">
<heading id="5A0281D3E101AF00FF0C3E87FD3DF94D" bold="true" box="[151,657,1752,1779]" fontSize="11" level="1" pageId="9" pageNumber="82" reason="1">
<taxonomicName id="C6F54D3CE101AF00FF0C3E87FD3DF94D" ID-CoL="98YVF" authority="Lin, Chang et Li, 2020" authorityName="Lin, Chang et Li" authorityYear="2020" box="[151,657,1752,1779]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE101AF00FF0C3E87FD3DF94D" bold="true" box="[151,657,1752,1779]" pageId="9" pageNumber="82">
<emphasis id="3381EAADE101AF00FF0C3E87FEDDF94D" bold="true" box="[151,369,1752,1779]" italics="true" pageId="9" pageNumber="82">Dysdera dushengi</emphasis>
<bibRefCitation id="65644B4EE101AF00FEE33E86FD3DF94D" author="Lin, Y. J. &amp; Chang, W. - J. &amp; Li, S. Q." box="[376,657,1753,1779]" pageId="9" pageNumber="82" pagination="97 - 103" refId="ref7933" refString="Lin, Y. J., Chang, W. - J. &amp; Li, S. Q. (2020) Dysdera dushengi sp. nov., the easternmost species of the spider family Dysderidae (Arachnida: Araneae). Zoological Systematics, 45 (2), 97 - 103. https: // doi. org / 10.11865 / zs. 202014" type="journal article" year="2020">Lin, Chang et Li, 2020</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E101AF00FF0C3EA2FD62F8A6" box="[151,718,1789,1816]" pageId="9" pageNumber="82" type="description">
<paragraph id="014A36BFE101AF00FF0C3EA2FD62F8A6" blockId="9.[151,718,1752,1816]" box="[151,718,1789,1816]" pageId="9" pageNumber="82">
<figureCitation id="99CE2A3AE101AF00FF0C3EA2FF57F8A9" box="[151,251,1789,1815]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="9" pageNumber="82">Figs 3–4</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FE9D3EA2FEE3F8A9" box="[262,335,1789,1815]" captionStart="FIGURES 13–18" captionStartId="4.[151,264,1703,1727]" captionTargetBox="[151,1435,181,1679]" captionTargetId="figure-25@4.[151,1435,181,1679]" captionTargetPageId="4" captionText="FIGURES 13–18. Prosoma of Dysdera dushengi male (13–15) and female (16–18). 13, 16—dorsal; 14, 17—lateral; 15, 18— ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157079" httpUri="https://zenodo.org/record/5157079/files/figure.png" pageId="9" pageNumber="82">13–18</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FEC13EA2FED4F8A9" box="[346,376,1789,1815]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[184,1408,193,1920]" captionTargetId="figure-25@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 25–30. Male palp (25–27) and bulb (28–30) of Dysdera subcylindrica (25, 28–30), D. dushengi (26) and D. mikhailovi sp. nov. (27). 25–28—prolateral; 29—anterior; 30—retrolateral. Scale bars: 25–27=0.5 mm, 28–30=0.2 mm. Ab- breviations: Ap–posterior apophysis, Ax–apex, C–crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157024" httpUri="https://zenodo.org/record/5157024/files/figure.png" pageId="9" pageNumber="82">26</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FE1F3EA2FE62F8A9" box="[388,462,1789,1815]" captionStart="FIGURES 31–36" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1407,181,1919]" captionTargetId="figure-25@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 31–36. Psembolus of Dysdera subcylindrica (31–33) and bulb of D. dushengi (34–36). 31, 34—prolateral; 32, 35— anterior; 33, 36—retrolateral. Scale bars: 31–33=0.15 mm, 34–36=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, C–crest, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157028" httpUri="https://zenodo.org/record/5157028/files/figure.png" pageId="9" pageNumber="82">34–36</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FE433EA2FD8DF8A9" box="[472,545,1789,1815]" captionStart="FIGURES 41–46" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1434,419,1866]" captionTargetId="figure-177@10.[151,1436,417,1867]" captionTargetPageId="10" captionText="FIGURES 41–46. Psembolus of Dysdera dushengi (41–43) and bulb of D. mikhailovi sp. nov. (44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="9" pageNumber="82">41–43</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FDB73EA2FDD9F8A6" box="[556,629,1789,1816]" captionStart="FIGURES 50–57" captionStartId="12.[151,264,1524,1548]" captionTargetBox="[153,1436,273,1500]" captionTargetId="figure-73@12.[151,1436,273,1500]" captionTargetPageId="12" captionText="FIGURES 50–57. Endogyne of D. subcylindrica (50–51), D. dushengi (52–53), D. mikhailovi sp. nov. (54–55) and D. arnoldii (56–57). 50, 52, 54, 56—dorsal; 51, 53, 55, 57—ventral. Scale bars=0.2 mm.Abbreviations: Dd–dorsal arch of the diverticulum, La–lateral arm of the spermatheca, Pd–posterior diverticulum, Sk–stalk of the spermatheca, St–spermateca, Tb–transverse bar of the posterior diverticulum." figureDoi="http://doi.org/10.5281/zenodo.5157040" httpUri="https://zenodo.org/record/5157040/files/figure.png" pageId="9" pageNumber="82">52–53</figureCitation>
,
<figureCitation id="99CE2A3AE101AF00FD1B3EA2FD62F8A6" box="[640,718,1789,1816]" captionStart="FIGURES 58–60" captionStartId="13.[151,264,1657,1681]" captionTargetBox="[151,1437,669,1634]" captionTargetId="figure-289@13.[151,1437,669,1634]" captionTargetPageId="13" captionText="FIGURES 58–60. Collecting localities of the studied Dysdera species (58–59) and published records of other Central Asian Dysdera species, except for D. crocata (60). Triangle—Dysdera arnoldii, diamond—D. mikhailovi sp. nov., circle—D. dushengi, square—D. subcylindrica. Black symbols refer to the studied material, white symbols refer to literature-derived data. Some symbols could refer to more than one closely situated localities. The frame of Fig. 59 refers to the content of Fig. 58. The numbers in Fig. 60 refer to the following species: 1—D. aculeata; 2—D. arnoldii; 3—D. brignolii; 4—D. dushengi; 5—D. fedtschenkoi Dunin, 1992; 6—D. ferghanica Dunin, 1985; 7—D. kronebergi Dunin, 1992; 8—D. kugitangica; 9—D. kusnetsovi; 10—D. limitanea; 11—D. nenilini; 12—D. pamirica, 13—D. pococki; 14—D. subcylindrica; 15—D. tartarica; 16—D. tystshenkoi Dunin, 1989; 17—D. zarudnyi; 18—D. zonsteini Dimitrov, 2021; 19—D. mikhailovi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5157042" httpUri="https://zenodo.org/record/5157042/files/figure.png" pageId="9" pageNumber="82">58–60.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E101AF00FF0C3F1DFC07F8E7" box="[151,939,1857,1881]" pageId="9" pageNumber="82" type="reference_group">
<paragraph id="014A36BFE101AF00FF0C3F1DFC07F8E7" blockId="9.[151,939,1857,1881]" box="[151,939,1857,1881]" pageId="9" pageNumber="82">
<treatmentCitationGroup id="21E51191E101AF00FF0C3F1DFC07F8E7" box="[151,939,1857,1881]" pageId="9" pageNumber="82">
<treatmentCitation id="805410AEE101AF00FF0C3F1DFD98F8E7" ID-DOI="http://doi.org/10.5281/zenodo.5105805" ID-Zenodo-Dep="5105805" author="Lin, Y. J. &amp; Chang, W. - J. &amp; Li, S. Q." box="[151,564,1857,1881]" httpUri="http://treatment.plazi.org/id/03EE87CEFFF33B56DC99FE12FB0B1738" page="100" pageId="9" pageNumber="82" year="2020">
<taxonomicName id="C6F54D3CE101AF00FF0C3F1DFD98F8E7" ID-CoL="98YVF" authority="Lin, Chang et Li, 2020: 100" authorityName="Lin, Chang et Li" authorityPageNumber="100" authorityYear="2020" box="[151,564,1857,1881]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE101AF00FF0C3F1DFEBDF8E7" box="[151,273,1857,1881]" italics="true" pageId="9" pageNumber="82">D. dushengi</emphasis>
<bibRefCitation id="65644B4EE101AF00FE833F1EFD98F8E7" author="Lin, Y. J. &amp; Chang, W. - J. &amp; Li, S. Q." box="[280,564,1857,1881]" pageId="9" pageNumber="82" pagination="97 - 103" refId="ref7933" refString="Lin, Y. J., Chang, W. - J. &amp; Li, S. Q. (2020) Dysdera dushengi sp. nov., the easternmost species of the spider family Dysderidae (Arachnida: Araneae). Zoological Systematics, 45 (2), 97 - 103. https: // doi. org / 10.11865 / zs. 202014" type="journal article" year="2020">Lin, Chang et Li, 2020: 100</bibRefCitation>
</taxonomicName>
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, f. 1A–D, 2A–B, 3, 4A–C, 5C (
<specimenCount id="17F3FD36E101AF00FCEC3F1EFC22F8E7" box="[887,910,1857,1881]" pageId="9" pageNumber="82" type="male">♂</specimenCount>
<specimenCount id="17F3FD36E101AF00FC153F1EFC32F8E7" box="[910,926,1857,1881]" pageId="9" pageNumber="82" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E101AF00FF0C3FD8FBC5F87B" pageId="9" pageNumber="82" type="materials_examined">
<paragraph id="014A36BFE101AF00FF0C3FD8FBC5F87B" blockId="9.[151,1436,1927,2026]" pageId="9" pageNumber="82">
<emphasis id="3381EAADE101AF00FF0C3FD8FEAAF81F" bold="true" box="[151,262,1927,1953]" pageId="9" pageNumber="82">Material.</emphasis>
<materialsCitation id="B19D3CE2E101AF00FE963FD7FBC9F87B" collectingDate="2016-04-22" collectionCode="ISEA" collectorName="R. Yu. Dudko" country="Kazakhstan" elevation="900" latitude="43.883335" location="Natural Boundary" longLatPrecision="1140" longitude="75.5" municipality="Karabastau Village" pageId="9" pageNumber="82" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy">
<collectingCountry id="79E2762FE101AF00FE963FD7FE63F81C" box="[269,463,1928,1954]" name="Kazakhstan" pageId="9" pageNumber="82">KAZAKHSTAN</collectingCountry>
:
<emphasis id="3381EAADE101AF00FE433FD8FD2FF81F" bold="true" box="[472,643,1927,1954]" italics="true" pageId="9" pageNumber="82">
<collectingRegion id="C331F85DE101AF00FE433FD8FD2FF81F" box="[472,643,1927,1954]" country="Kazakhstan" name="Almaty Oblysy" pageId="9" pageNumber="82">Almaty Region</collectingRegion>
</emphasis>
:
<specimenCount id="17F3FD36E101AF00FD093FD7FD0BF81C" box="[658,679,1928,1954]" pageId="9" pageNumber="82" type="male">♂</specimenCount>
<specimenCount id="17F3FD36E101AF00FD353FD7FD6FF81C" box="[686,707,1928,1954]" pageId="9" pageNumber="82" type="female">♀</specimenCount>
(
<collectionCode id="67E4AE7AE101AF00FD4A3FD7FCBFF81C" box="[721,787,1928,1954]" pageId="9" pageNumber="82">ISEA</collectionCode>
, 001.8644), c.
<quantity id="C60D9B5AE101AF00FC233FD8FC53F81F" box="[952,1023,1927,1954]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="82" unit="km" value="15.0">15 km</quantity>
NNW from
<collectingMunicipality id="E12EACC5E101AF00FB103FD8FACFF81F" box="[1163,1379,1927,1953]" pageId="9" pageNumber="82">Karabastau Village</collectingMunicipality>
, Tyr- nakty
<location id="042A6064E101AF00FF473FF4FE0BF87B" LSID="urn:lsid:plazi:treatment:895C87A9E101AF02FF0C3E87FB1AFCAB:042A6064E101AF00FF473FF4FE0BF87B" box="[220,423,1963,1989]" country="Kazakhstan" latitude="43.883335" longLatPrecision="1140" longitude="75.5" municipality="Karabastau Village" name="Natural Boundary" pageId="9" pageNumber="82" stateProvince="Almaty Oblysy">Natural Boundary</location>
,
<geoCoordinate id="64C15078E101AF00FE293FF4FDBFF878" box="[434,531,1963,1990]" degrees="43" direction="north" minutes="53" orientation="latitude" pageId="9" pageNumber="82" precision="925" value="43.883335">43°53’N</geoCoordinate>
<geoCoordinate id="64C15078E101AF00FD813FF4FDD6F878" box="[538,634,1963,1990]" degrees="75" direction="east" minutes="30" orientation="longitude" pageId="9" pageNumber="82" precision="925" value="75.5">75°30’E</geoCoordinate>
,
<quantity id="C60D9B5AE101AF00FD1E3FF4FCAAF878" box="[645,774,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="9" pageNumber="82" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">
<elevation id="8AD8D18CE101AF00FD1E3FF4FCAAF878" box="[645,774,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="9" pageNumber="82" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">850–950 m</elevation>
</quantity>
, 20–
<date id="754B107FE101AF00FCA73FF4FC11F87B" box="[828,957,1963,1989]" pageId="9" pageNumber="82" value="2016-04-22">
<collectingDate id="650FE997E101AF00FCA73FF4FC11F87B" box="[828,957,1963,1989]" pageId="9" pageNumber="82" value="2016-04-22">22.04.2016</collectingDate>
</date>
,
<collectorName id="AC005369E101AF00FC533FF3FBC9F87B" box="[968,1125,1963,1990]" pageId="9" pageNumber="82">R. Yu. Dudko</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E101AF03FF5C3F90FB0CFE37" lastPageId="10" lastPageNumber="83" pageId="9" pageNumber="82" type="diagnosis">
<paragraph id="014A36BFE101AF00FF5C3F90FA30F857" blockId="9.[151,1436,1927,2026]" box="[199,1436,1999,2026]" pageId="9" pageNumber="82">
<emphasis id="3381EAADE101AF00FF5C3F90FEEEF857" bold="true" box="[199,322,1999,2025]" pageId="9" pageNumber="82">Diagnosis.</emphasis>
The male of
<taxonomicName id="C6F54D3CE101AF00FE433F8FFDCCF857" authorityName="Lin, Chang et Li" authorityYear="2020" box="[472,608,1999,2025]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE101AF00FE433F8FFDCCF857" box="[472,608,1999,2025]" italics="true" pageId="9" pageNumber="82">D. dushengi</emphasis>
</taxonomicName>
is most similar to that of
<taxonomicName id="C6F54D3CE101AF00FC193F8FFBA5F857" authorityName="Dunin" authorityYear="1992" box="[898,1033,2000,2025]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE101AF00FC193F8FFBA5F857" box="[898,1033,2000,2025]" italics="true" pageId="9" pageNumber="82">D. pamirica</emphasis>
</taxonomicName>
from
<collectingCountry id="79E2762FE101AF00FBD53F90FB10F857" box="[1102,1212,1999,2025]" name="Tajikistan" pageId="9" pageNumber="82">Tajikistan</collectingCountry>
in having the semi-
</paragraph>
<paragraph id="014A36BFE102AF03FF0C38C8FB0CFE37" blockId="10.[151,1437,151,393]" pageId="10" pageNumber="83">
elliptical median crest (
<emphasis id="3381EAADE102AF03FE0538C8FE69FF0F" box="[414,453,151,177]" italics="true" pageId="10" pageNumber="83">Cm</emphasis>
), the abducted retrolateral crest (
<emphasis id="3381EAADE102AF03FCAC38C8FCF9FF0F" box="[823,853,151,177]" italics="true" pageId="10" pageNumber="83">Cr</emphasis>
) and the psembolus being two times longer than the tegulum (
<emphasis id="3381EAADE102AF03FE9838E4FEB0FF6A" box="[259,284,187,212]" italics="true" pageId="10" pageNumber="83">Te</emphasis>
), but can be distinguished from it by having the tegulum and the psembolus being stretched in almost a straight line (compared to a 130° angle between them in
<taxonomicName id="C6F54D3CE102AF03FC923880FC23FF46" authorityName="Dunin" authorityYear="1992" box="[777,911,223,248]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE102AF03FC923880FC23FF46" box="[777,911,223,248]" italics="true" pageId="10" pageNumber="83">D. pamirica</emphasis>
</taxonomicName>
), the median crest being two times shorter than the tegulum (equal in length in
<taxonomicName id="C6F54D3CE102AF03FE60395CFD2FFEA2" authorityName="Dunin" authorityYear="1992" box="[507,643,259,284]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE102AF03FE60395CFD2FFEA2" box="[507,643,259,284]" italics="true" pageId="10" pageNumber="83">D. pamirica</emphasis>
</taxonomicName>
) and the hooked apex (
<emphasis id="3381EAADE102AF03FC15395CFC00FEA3" box="[910,940,259,285]" italics="true" pageId="10" pageNumber="83">Ax</emphasis>
) of psembolus (straight in
<taxonomicName id="C6F54D3CE102AF03FB46395CFAC9FEA2" authorityName="Dunin" authorityYear="1992" box="[1245,1381,259,284]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE102AF03FB46395CFAC9FEA2" box="[1245,1381,259,284]" italics="true" pageId="10" pageNumber="83">D. pamirica</emphasis>
</taxonomicName>
) (cf.
<figureCitation id="99CE2A3AE102AF03FF0C3978FEB4FEFF" box="[151,280,295,321]" captionStart="FIGURES 31–36" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1407,181,1919]" captionTargetId="figure-25@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 31–36. Psembolus of Dysdera subcylindrica (31–33) and bulb of D. dushengi (34–36). 31, 34—prolateral; 32, 35— anterior; 33, 36—retrolateral. Scale bars: 31–33=0.15 mm, 34–36=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, C–crest, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157028" httpUri="https://zenodo.org/record/5157028/files/figure.png" pageId="10" pageNumber="83">Figs 34–36</figureCitation>
,
<figureCitation id="99CE2A3AE102AF03FEB83978FEC5FEFF" box="[291,361,295,321]" captionStart="FIGURES 41–46" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1434,419,1866]" captionTargetId="figure-177@10.[151,1436,417,1867]" captionTargetPageId="10" captionText="FIGURES 41–46. Psembolus of Dysdera dushengi (41–43) and bulb of D. mikhailovi sp. nov. (44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="10" pageNumber="83">41–43</figureCitation>
and figs
<quantity id="C60D9B5AE102AF03FE493978FDB5FEFF" box="[466,537,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="10" pageNumber="83" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7–9 in</quantity>
<bibRefCitation id="65644B4EE102AF03FDBB3978FD69FEFF" author="Dunin, P. M." box="[544,709,295,321]" pageId="10" pageNumber="83" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin (1992))</bibRefCitation>
. The female of
<taxonomicName id="C6F54D3CE102AF03FCEE3978FC50FEFF" authorityName="Lin, Chang et Li" authorityYear="2020" box="[885,1020,295,321]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE102AF03FCEE3978FC50FEFF" box="[885,1020,295,321]" italics="true" pageId="10" pageNumber="83">D. dushengi</emphasis>
</taxonomicName>
is also similar to that of
<taxonomicName id="C6F54D3CE102AF03FA8E3978FA30FEFE" authorityName="Dunin" authorityYear="1992" box="[1301,1436,295,320]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE102AF03FA8E3978FA30FEFE" box="[1301,1436,295,320]" italics="true" pageId="10" pageNumber="83">D. pamirica</emphasis>
</taxonomicName>
in the shape of trapezoidal dorsal arch of the diverticulum (
<emphasis id="3381EAADE102AF03FCDE3914FCCBFEDB" box="[837,871,331,357]" italics="true" pageId="10" pageNumber="83">Dd</emphasis>
), but differs in having a distinct stalk (
<emphasis id="3381EAADE102AF03FAB63914FAEBFEDB" box="[1325,1351,331,357]" italics="true" pageId="10" pageNumber="83">Sk</emphasis>
) of the spermatheca which is absent from
<taxonomicName id="C6F54D3CE102AF03FD803930FD0EFE36" authorityName="Dunin" authorityYear="1992" box="[539,674,367,392]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE102AF03FD803930FD0EFE36" box="[539,674,367,392]" italics="true" pageId="10" pageNumber="83">D. pamirica</emphasis>
</taxonomicName>
(cf.
<figureCitation id="99CE2A3AE102AF03FD4C3930FCF8FE37" box="[727,852,366,393]" captionStart="FIGURES 50–57" captionStartId="12.[151,264,1524,1548]" captionTargetBox="[153,1436,273,1500]" captionTargetId="figure-73@12.[151,1436,273,1500]" captionTargetPageId="12" captionText="FIGURES 50–57. Endogyne of D. subcylindrica (50–51), D. dushengi (52–53), D. mikhailovi sp. nov. (54–55) and D. arnoldii (56–57). 50, 52, 54, 56—dorsal; 51, 53, 55, 57—ventral. Scale bars=0.2 mm.Abbreviations: Dd–dorsal arch of the diverticulum, La–lateral arm of the spermatheca, Pd–posterior diverticulum, Sk–stalk of the spermatheca, St–spermateca, Tb–transverse bar of the posterior diverticulum." figureDoi="http://doi.org/10.5281/zenodo.5157040" httpUri="https://zenodo.org/record/5157040/files/figure.png" pageId="10" pageNumber="83">Figs 52–53</figureCitation>
and fig.
<quantity id="C60D9B5AE102AF03FC2C3930FC5CFE37" box="[951,1008,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="83" unit="in" value="10.0">10 in</quantity>
<bibRefCitation id="65644B4EE102AF03FC6C3930FB30FE37" author="Dunin, P. M." box="[1015,1180,367,393]" pageId="10" pageNumber="83" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin (1992))</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="558A6637E102AF03FF0C3F3DFB5EF87C" ID-DOI="http://doi.org/10.5281/zenodo.5157033" ID-Zenodo-Dep="5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="10" pageNumber="83" startId="10.[151,264,1890,1914]" targetBox="[151,1434,419,1866]" targetPageId="10">
<paragraph id="014A36BFE102AF03FF0C3F3DFB5EF87C" blockId="10.[151,1437,1890,1986]" pageId="10" pageNumber="83">
<emphasis id="3381EAADE102AF03FF0C3F3DFEFAF8C4" bold="true" box="[151,342,1890,1914]" pageId="10" pageNumber="83">FIGURES 41–46.</emphasis>
Psembolus of
<taxonomicName id="C6F54D3CE102AF03FE693F3DFD00F8C4" authorityName="Lin, Chang et Li" authorityYear="2020" box="[498,684,1890,1914]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE102AF03FE693F3DFD00F8C4" box="[498,684,1890,1914]" italics="true" pageId="10" pageNumber="83">Dysdera dushengi</emphasis>
</taxonomicName>
(41–43) and bulb of
<taxonomicName id="C6F54D3CE102AF03FC163F3CFBB5F8C4" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[909,1049,1890,1914]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE102AF03FC163F3CFBB5F8C4" box="[909,1049,1890,1914]" italics="true" pageId="10" pageNumber="83">D. mikhailovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28B257D6E102AF03FBB93F3CFBDFF8C4" box="[1058,1139,1891,1914]" pageId="10" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
(44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations:
<emphasis id="3381EAADE102AF03FB3E3FD8FB6DF820" box="[1189,1217,1927,1950]" italics="true" pageId="10" pageNumber="83">Ap</emphasis>
–posterior apophysis,
<emphasis id="3381EAADE102AF03FF0C3FF4FF1EF87C" box="[151,178,1963,1986]" italics="true" pageId="10" pageNumber="83">Ax</emphasis>
–apex,
<emphasis id="3381EAADE102AF03FF603FF4FEB2F87C" box="[251,286,1963,1986]" italics="true" pageId="10" pageNumber="83">Cm</emphasis>
–median crest,
<emphasis id="3381EAADE102AF03FE233FF4FE7AF87C" box="[440,470,1963,1986]" italics="true" pageId="10" pageNumber="83">Cp</emphasis>
–prolateral crest,
<emphasis id="3381EAADE102AF03FD1D3FF4FD0DF87C" box="[646,673,1963,1986]" italics="true" pageId="10" pageNumber="83">Cr</emphasis>
–retrolateral crest,
<emphasis id="3381EAADE102AF03FCFB3FF4FCD4F87C" box="[864,888,1963,1986]" italics="true" pageId="10" pageNumber="83">Se</emphasis>
–stem of the psembolus,
<emphasis id="3381EAADE102AF03FBEF3FF4FB27F87C" box="[1140,1163,1963,1986]" italics="true" pageId="10" pageNumber="83">Te</emphasis>
–tegulum.
</paragraph>
</caption>
<subSubSection id="49EF6534E103AF02FF5C38C9FD58FD17" pageId="11" pageNumber="84" type="description">
<paragraph id="014A36BFE103AF02FF5C38C9FDACFE13" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="84">
<emphasis id="3381EAADE103AF02FF5C38C9FEF5FF0E" bold="true" box="[199,345,150,176]" pageId="11" pageNumber="84">Description.</emphasis>
Male. Total length 7.4. Carapace: 3.15 long, 2.45 wide. Abdomen: 4.25 long, 1.95 wide. AME 0.17, PME 0.14, PLE 0.14. General appearance as in
<figureCitation id="99CE2A3AE103AF02FD7038E5FC9DFF6A" box="[747,817,186,212]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="11" pageNumber="84">Figs 3</figureCitation>
,
<figureCitation id="99CE2A3AE103AF02FCA538E5FC2BFF6B" box="[830,903,186,213]" captionStart="FIGURES 13–18" captionStartId="4.[151,264,1703,1727]" captionTargetBox="[151,1435,181,1679]" captionTargetId="figure-25@4.[151,1435,181,1679]" captionTargetPageId="4" captionText="FIGURES 13–18. Prosoma of Dysdera dushengi male (13–15) and female (16–18). 13, 16—dorsal; 14, 17—lateral; 15, 18— ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157079" httpUri="https://zenodo.org/record/5157079/files/figure.png" pageId="11" pageNumber="84">13–15</figureCitation>
. Prosoma and chelicerae orange. Palps and legs yellow. Abdomen and spinnerets beige. Leg measurements: I: 2.85, 1.75, 2.55, 2.7, 0.7 (10.55). II: 2.65, 1.55, 2.3, 2.55, 0.65 (9.7). III: 2.05, 1.15, 1.35, 2.1, 0.55 (7.2). IV: 2.65, 1.35, 2.1, 2.6, 0.65 (9.35). Leg spination: I: Fe p3. II: Fe p3. III: Fe p3 r1; Ti p5 r3 v6; Mt p6 r3 v2. IV: Fe d8 r1; Ti p4 r5 v6; Mt p4 r3 v5. Bulb as in
<figureCitation id="99CE2A3AE103AF02FB763979FAEFFEFE" box="[1261,1347,294,320]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[184,1408,193,1920]" captionTargetId="figure-25@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 25–30. Male palp (25–27) and bulb (28–30) of Dysdera subcylindrica (25, 28–30), D. dushengi (26) and D. mikhailovi sp. nov. (27). 25–28—prolateral; 29—anterior; 30—retrolateral. Scale bars: 25–27=0.5 mm, 28–30=0.2 mm. Ab- breviations: Ap–posterior apophysis, Ax–apex, C–crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157024" httpUri="https://zenodo.org/record/5157024/files/figure.png" pageId="11" pageNumber="84">Figs 26</figureCitation>
,
<figureCitation id="99CE2A3AE103AF02FAD43979FA34FEFE" box="[1359,1432,294,320]" captionStart="FIGURES 31–36" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[178,1407,181,1919]" captionTargetId="figure-25@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 31–36. Psembolus of Dysdera subcylindrica (31–33) and bulb of D. dushengi (34–36). 31, 34—prolateral; 32, 35— anterior; 33, 36—retrolateral. Scale bars: 31–33=0.15 mm, 34–36=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, C–crest, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157028" httpUri="https://zenodo.org/record/5157028/files/figure.png" pageId="11" pageNumber="84">34–36</figureCitation>
,
<figureCitation id="99CE2A3AE103AF02FF0C3915FF4CFEDA" box="[151,224,330,356]" captionStart="FIGURES 41–46" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1434,419,1866]" captionTargetId="figure-177@10.[151,1436,417,1867]" captionTargetPageId="10" captionText="FIGURES 41–46. Psembolus of Dysdera dushengi (41–43) and bulb of D. mikhailovi sp. nov. (44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="11" pageNumber="84">41–43</figureCitation>
. Length/width ratio 3.2. Posterior apophysis (
<emphasis id="3381EAADE103AF02FD7B3914FD53FEDB" box="[736,767,331,357]" italics="true" pageId="11" pageNumber="84">Ap</emphasis>
) lanceolate in lateral view. Psembolus with three crests that differ in shapes: obovate median crest, semi-oval prolateral crest (
<emphasis id="3381EAADE103AF02FC1E3930FC0AFE37" box="[901,934,367,393]" italics="true" pageId="11" pageNumber="84">Cp</emphasis>
) and triangular retrolateral crest. Psembolus apex bent towards median crest.
</paragraph>
<paragraph id="014A36BFE103AF02FF5C39E9FD58FD17" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="84">
Female. Total length unknown (specimen with damaged abdomen). Carapace: 3.2 long 2.45 wide. AME 0.17, PME 0.14, PLE 0.16. Coloration as in male (
<figureCitation id="99CE2A3AE103AF02FD173985FD78FE4A" box="[652,724,474,500]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="11" pageNumber="84">Figs 4</figureCitation>
,
<figureCitation id="99CE2A3AE103AF02FD7B3985FC86FE4B" box="[736,810,474,501]" captionStart="FIGURES 13–18" captionStartId="4.[151,264,1703,1727]" captionTargetBox="[151,1435,181,1679]" captionTargetId="figure-25@4.[151,1435,181,1679]" captionTargetPageId="4" captionText="FIGURES 13–18. Prosoma of Dysdera dushengi male (13–15) and female (16–18). 13, 16—dorsal; 14, 17—lateral; 15, 18— ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157079" httpUri="https://zenodo.org/record/5157079/files/figure.png" pageId="11" pageNumber="84">16–18</figureCitation>
). Leg measurements: I: 2.7, 1.65, 2.3, 2.35, 0.6 (9.6). II: 2.4, 1.5, 2.0, 2.2, 0.6 (8.7). III: 1.95, 1.1, 1.3, 1.9, 0.55 (6.8). IV: 2.5, 1.4, 2.0, 2.45, 0.65 (9.0). Leg spination: I: Fe p3. II: Fe p2. III: Fe d1 p3 r1; Ti p4 r3 v6; Mt p6 r4 v3. IV: Fe d6 r1; Ti p4 r3 v6; Mt p6 r4 v5. Endogyne as in
<figureCitation id="99CE2A3AE103AF02FAF73A7DFF4CFDDF" captionStart="FIGURES 50–57" captionStartId="12.[151,264,1524,1548]" captionTargetBox="[153,1436,273,1500]" captionTargetId="figure-73@12.[151,1436,273,1500]" captionTargetPageId="12" captionText="FIGURES 50–57. Endogyne of D. subcylindrica (50–51), D. dushengi (52–53), D. mikhailovi sp. nov. (54–55) and D. arnoldii (56–57). 50, 52, 54, 56—dorsal; 51, 53, 55, 57—ventral. Scale bars=0.2 mm.Abbreviations: Dd–dorsal arch of the diverticulum, La–lateral arm of the spermatheca, Pd–posterior diverticulum, Sk–stalk of the spermatheca, St–spermateca, Tb–transverse bar of the posterior diverticulum." figureDoi="http://doi.org/10.5281/zenodo.5157040" httpUri="https://zenodo.org/record/5157040/files/figure.png" pageId="11" pageNumber="84">Figs 52–53</figureCitation>
. Spermatheca (
<emphasis id="3381EAADE103AF02FE103A18FE0DFDDF" box="[395,417,583,609]" italics="true" pageId="11" pageNumber="84">St</emphasis>
) fusiform, with pointed lateral arms (
<emphasis id="3381EAADE103AF02FCDD3A18FCC8FDDE" box="[838,868,583,608]" italics="true" pageId="11" pageNumber="84">La</emphasis>
) and clearly visible stalk. Dorsal arch of diverticu- lum trapezoidal, twice as long as wide. Transverse bar of the posterior diverticulum (
<emphasis id="3381EAADE103AF02FBCD3A35FBD8FD3A" box="[1110,1140,618,644]" italics="true" pageId="11" pageNumber="84">Tb</emphasis>
) straight, two times wider than spermatheca. Posterior diverticulum (
<emphasis id="3381EAADE103AF02FDEF3AD1FD3FFD16" box="[628,659,654,680]" italics="true" pageId="11" pageNumber="84">Pd</emphasis>
) square.
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E103AF02FF5C3AEDFB1AFCAB" pageId="11" pageNumber="84" type="distribution">
<paragraph id="014A36BFE103AF02FF5C3AEDFB1AFCAB" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="84">
<emphasis id="3381EAADE103AF02FF5C3AEDFECDFD72" bold="true" box="[199,353,690,716]" pageId="11" pageNumber="84">Distribution.</emphasis>
Until now, the species has been known only from the
<typeStatus id="DE4E881DE103AF02FC233AECFC44FD73" box="[952,1000,691,717]" pageId="11" pageNumber="84">type</typeStatus>
locality, the westernmost part of Xinji- ang Province of
<collectingCountry id="79E2762FE103AF02FED73A89FE3EFD4E" box="[332,402,726,752]" name="China" pageId="11" pageNumber="84">China</collectingCountry>
, neighbouring
<collectingCountry id="79E2762FE103AF02FDA33A89FD17FD4E" box="[568,699,726,752]" name="Kazakhstan" pageId="11" pageNumber="84">Kazakhstan</collectingCountry>
(
<figureCitation id="99CE2A3AE103AF02FD533A89FCE7FD4F" box="[712,843,726,753]" captionStart="FIGURES 58–60" captionStartId="13.[151,264,1657,1681]" captionTargetBox="[151,1437,669,1634]" captionTargetId="figure-289@13.[151,1437,669,1634]" captionTargetPageId="13" captionText="FIGURES 58–60. Collecting localities of the studied Dysdera species (58–59) and published records of other Central Asian Dysdera species, except for D. crocata (60). Triangle—Dysdera arnoldii, diamond—D. mikhailovi sp. nov., circle—D. dushengi, square—D. subcylindrica. Black symbols refer to the studied material, white symbols refer to literature-derived data. Some symbols could refer to more than one closely situated localities. The frame of Fig. 59 refers to the content of Fig. 58. The numbers in Fig. 60 refer to the following species: 1—D. aculeata; 2—D. arnoldii; 3—D. brignolii; 4—D. dushengi; 5—D. fedtschenkoi Dunin, 1992; 6—D. ferghanica Dunin, 1985; 7—D. kronebergi Dunin, 1992; 8—D. kugitangica; 9—D. kusnetsovi; 10—D. limitanea; 11—D. nenilini; 12—D. pamirica, 13—D. pococki; 14—D. subcylindrica; 15—D. tartarica; 16—D. tystshenkoi Dunin, 1989; 17—D. zarudnyi; 18—D. zonsteini Dimitrov, 2021; 19—D. mikhailovi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5157042" httpUri="https://zenodo.org/record/5157042/files/figure.png" pageId="11" pageNumber="84">Figs 58–59</figureCitation>
). The present record in
<collectingRegion id="C331F85DE103AF02FBD03A89FB5AFD4E" box="[1099,1270,726,752]" country="Kazakhstan" name="Almaty Oblysy" pageId="11" pageNumber="84">Almaty Region</collectingRegion>
of
<collectingCountry id="79E2762FE103AF02FA813A89FA31FD4E" box="[1306,1437,726,752]" name="Kazakhstan" pageId="11" pageNumber="84">Kazakhstan</collectingCountry>
represents the westernmost limit of the species range. Both localities lie in the Ili River valley.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="5B68FAE0B9DCA8E5BD504D3A194780C3" ID-DOI="10.11646/zootaxa.5006.1.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="5157005" ID-ZooBank="251E38AC-1DDD-4AAD-8673-BCC7B0A179F8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628037152584" checkinUser="plazi" docAuthor="Fomichev, Alexander A. &amp; Marusik, Yuri M." docDate="2021" docId="895C87A9E103AF05FF0C3E41FBDAFF47" docLanguage="en" docName="zootaxa.5006.1.10.pdf" docOrigin="Zootaxa 5006 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysdera brignolii Dunin 1989" docType="treatment" docVersion="3" lastPageNumber="84" masterDocId="7565FFD1E108AF09FF9B385FFFACFFBE" masterDocTitle="Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia" masterLastPageNumber="89" masterPageNumber="73" pageNumber="84" updateTime="1699033588743" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C4794CEC9CF4F36C0E16FC0091A40F7B">Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia</mods:title>
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<mods:namePart id="77FD0465D57BD9CEE15944D9A7BC5B80">Fomichev, Alexander A.</mods:namePart>
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<mods:affiliation id="29152512555AB895D808D4F86A35BA1A">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia &amp; a. fomichov @ mail. ru; https: // orcid. org / 0000 - 0001 - 9268 - 622 X</mods:affiliation>
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<mods:namePart id="510FD88A93E8009946ACBFC1FB473E55">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="2C5EBEEB327BC93F4AC0FB36992BB3FC">Institute for Biological Problems of the North, Portovaya Street 18, Magadan 685000, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa &amp; yurmar @ mail. ru; https: // orcid. org / 0000 - 0002 - 4499 - 5148</mods:affiliation>
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<mods:date id="65C2FA88D00F29E4EC4B5C9E3B0B5B96">2021</mods:date>
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<subSubSection id="49EF6534E103AF02FF0C3E41FE10F986" box="[151,444,1566,1593]" pageId="11" pageNumber="84" type="nomenclature">
<paragraph id="014A36BFE103AF02FF0C3E41FE10F986" blockId="11.[151,444,1566,1593]" box="[151,444,1566,1593]" pageId="11" pageNumber="84">
<heading id="5A0281D3E103AF02FF0C3E41FE10F986" bold="true" box="[151,444,1566,1593]" fontSize="11" level="1" pageId="11" pageNumber="84" reason="1">
<emphasis id="3381EAADE103AF02FF0C3E41FE10F986" bold="true" box="[151,444,1566,1593]" pageId="11" pageNumber="84">
<taxonomicName id="C6F54D3CE103AF02FF0C3E41FEC6F987" ID-CoL="388Z5" authorityName="Dunin" authorityYear="1989" box="[151,362,1566,1593]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FF0C3E41FEC6F987" bold="true" box="[151,362,1566,1593]" italics="true" pageId="11" pageNumber="84">Dysdera brignolii</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E103AF02FF0C3E3CFDC2F9C5" box="[151,622,1635,1659]" pageId="11" pageNumber="84" type="reference_group">
<paragraph id="014A36BFE103AF02FF0C3E3CFDC2F9C5" blockId="11.[151,622,1635,1659]" box="[151,622,1635,1659]" pageId="11" pageNumber="84">
<treatmentCitationGroup id="21E51191E103AF02FF0C3E3CFDC2F9C5" box="[151,622,1635,1659]" pageId="11" pageNumber="84">
The
<taxonomicName id="C6F54D3CE103AF02FF5E3E3CFEB2F9C5" ID-CoL="388Z5" authorityName="Dunin" authorityYear="1989" box="[197,286,1635,1659]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FF5E3E3CFEB2F9C5" box="[197,286,1635,1659]" italics="true" pageId="11" pageNumber="84">brignolii</emphasis>
</taxonomicName>
group,
<emphasis id="3381EAADE103AF02FEF73E3BFE08F9C5" box="[364,420,1636,1659]" italics="true" pageId="11" pageNumber="84">sensu</emphasis>
<treatmentCitation id="805410AEE103AF02FE313E3CFDC6F9C5" author="Dunin, P. M." box="[426,618,1635,1659]" page="137" pageId="11" pageNumber="84" year="1992">
<bibRefCitation id="65644B4EE103AF02FE313E3CFDC6F9C5" author="Dunin, P. M." box="[426,618,1635,1659]" pageId="11" pageNumber="84" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin (1992: 137)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="49EF6534E103AF02FF0C3EF6FAA3F8EA" pageId="11" pageNumber="84" type="diagnosis">
<paragraph id="014A36BFE103AF02FF0C3EF6FE49F8B5" blockId="11.[151,1437,1705,2020]" pageId="11" pageNumber="84">
<emphasis id="3381EAADE103AF02FF0C3EF6FEBEF97D" bold="true" box="[151,274,1705,1731]" pageId="11" pageNumber="84">Diagnosis.</emphasis>
The group can be recognized by the combination of the following characters: the strongly prominent semi-oval median crest, the psembolus 1.2–1.6 times longer than the tegulum, and the posterior apophysis situated in the middle part of the bulb.
</paragraph>
<paragraph id="014A36BFE103AF02FF5C3F4AFAA3F8EA" blockId="11.[151,1437,1705,2020]" pageId="11" pageNumber="84">
<emphasis id="3381EAADE103AF02FF5C3F4AFECBF891" bold="true" box="[199,359,1813,1839]" pageId="11" pageNumber="84">Composition.</emphasis>
Five species are included:
<taxonomicName id="C6F54D3CE103AF02FD0C3F49FC1EF891" authority="Dunin, 1989" authorityName="Dunin" authorityYear="1989" box="[663,946,1813,1839]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FD0C3F49FCB0F891" box="[663,796,1813,1839]" italics="true" pageId="11" pageNumber="84">D. brignolii</emphasis>
<bibRefCitation id="65644B4EE103AF02FCB83F4AFC1EF891" author="Dunin, P. M." box="[803,946,1813,1839]" pageId="11" pageNumber="84" pagination="865 - 874" refId="ref7748" refString="Dunin, P. M. (1989) Material on the fauna of spiders of the family Dysderidae (Aranei) from Turkmenistan. Entomologicheskoe Obozrenie, 68 (4), 865 - 874. [in Russian]" type="journal article" year="1989">Dunin, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FC263F49FB56F891" authority="Dunin, 1992" authorityName="Dunin" authorityYear="1992" box="[957,1274,1813,1839]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE103AF02FC263F49FBC8F891" box="[957,1124,1813,1839]" italics="true" pageId="11" pageNumber="84">D. kugitangica</emphasis>
<bibRefCitation id="65644B4EE103AF02FBF03F4AFB56F891" author="Dunin, P. M." box="[1131,1274,1813,1839]" pageId="11" pageNumber="84" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin, 1992</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FA9E3F49FE8AF8ED" authority="Dunin, 1989" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="kusnetsovi">
<emphasis id="3381EAADE103AF02FA9E3F49FA31F891" box="[1285,1437,1813,1839]" italics="true" pageId="11" pageNumber="84">D. kusnetsovi</emphasis>
<bibRefCitation id="65644B4EE103AF02FF0C3F66FE8AF8ED" author="Dunin, P. M." box="[151,294,1849,1875]" pageId="11" pageNumber="84" pagination="865 - 874" refId="ref7748" refString="Dunin, P. M. (1989) Material on the fauna of spiders of the family Dysderidae (Aranei) from Turkmenistan. Entomologicheskoe Obozrenie, 68 (4), 865 - 874. [in Russian]" type="journal article" year="1989">Dunin, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FEAA3F65FD90F8ED" authority="Dunin, 1989" authorityName="Dunin" authorityYear="1989" box="[305,572,1849,1875]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE103AF02FEAA3F65FE05F8ED" box="[305,425,1849,1875]" italics="true" pageId="11" pageNumber="84">D. nenilini</emphasis>
<bibRefCitation id="65644B4EE103AF02FE2B3F66FD90F8ED" author="Dunin, P. M." box="[432,572,1849,1875]" pageId="11" pageNumber="84" pagination="865 - 874" refId="ref7748" refString="Dunin, P. M. (1989) Material on the fauna of spiders of the family Dysderidae (Aranei) from Turkmenistan. Entomologicheskoe Obozrenie, 68 (4), 865 - 874. [in Russian]" type="journal article" year="1989">Dunin, 1989</bibRefCitation>
</taxonomicName>
(all from
<collectingCountry id="79E2762FE103AF02FD373F66FCE5F8EA" box="[684,841,1849,1876]" name="Turkmenistan" pageId="11" pageNumber="84">Turkmenistan</collectingCountry>
) and
<taxonomicName id="C6F54D3CE103AF02FC1D3F65FBB3F8ED" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[902,1055,1849,1875]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE103AF02FC1D3F65FBB3F8ED" box="[902,1055,1849,1875]" italics="true" pageId="11" pageNumber="84">D. mikhailovi</emphasis>
</taxonomicName>
<emphasis id="3381EAADE103AF02FBBD3F66FBD3F8ED" bold="true" box="[1062,1151,1849,1875]" pageId="11" pageNumber="84">
<taxonomicNameLabel id="28B257D6E103AF02FBBD3F66FBD3F8ED" box="[1062,1151,1849,1875]" pageId="11" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="79E2762FE103AF02FB153F66FAAFF8EA" box="[1166,1283,1849,1876]" name="Tajikistan" pageId="11" pageNumber="84">Tajikistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E103AF05FF5C3F02FBDAFF47" lastPageId="12" lastPageNumber="85" pageId="11" pageNumber="84" type="discussion">
<paragraph id="014A36BFE103AF05FF5C3F02FBDAFF47" blockId="11.[151,1437,1705,2020]" lastBlockId="12.[151,1437,151,250]" lastPageId="12" lastPageNumber="85" pageId="11" pageNumber="84">
<emphasis id="3381EAADE103AF02FF5C3F02FEE3F8C9" bold="true" box="[199,335,1885,1911]" pageId="11" pageNumber="84">Comments.</emphasis>
<bibRefCitation id="65644B4EE103AF02FEC33F02FE5EF8C9" author="Dunin, P. M." box="[344,498,1885,1911]" pageId="11" pageNumber="84" pagination="865 - 874" refId="ref7748" refString="Dunin, P. M. (1989) Material on the fauna of spiders of the family Dysderidae (Aranei) from Turkmenistan. Entomologicheskoe Obozrenie, 68 (4), 865 - 874. [in Russian]" type="journal article" year="1989">Dunin (1989)</bibRefCitation>
assigned representatives of the
<taxonomicName id="C6F54D3CE103AF02FCFA3F02FC6FF8C9" authorityName="Dunin" authorityYear="1989" box="[865,963,1885,1911]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FCFA3F02FC6FF8C9" box="[865,963,1885,1911]" italics="true" pageId="11" pageNumber="84">brignolii</emphasis>
</taxonomicName>
group, i.e.
<taxonomicName id="C6F54D3CE103AF02FBD33F01FB4DF8C9" authorityName="Dunin" authorityYear="1989" box="[1096,1249,1885,1911]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="kusnetsovi">
<emphasis id="3381EAADE103AF02FBD33F01FB4DF8C9" box="[1096,1249,1885,1911]" italics="true" pageId="11" pageNumber="84">D. kusnetsovi</emphasis>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FB6A3F01FAC7F8C9" authorityName="Dunin" authorityYear="1989" box="[1265,1387,1885,1911]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE103AF02FB6A3F01FAC7F8C9" box="[1265,1387,1885,1911]" italics="true" pageId="11" pageNumber="84">D. nenilini</emphasis>
</taxonomicName>
and
<taxonomicName id="C6F54D3CE103AF02FF0C3FDDFEB1F825" authorityName="Dunin" authorityYear="1989" box="[151,285,1921,1947]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FF0C3FDDFEB1F825" box="[151,285,1921,1947]" italics="true" pageId="11" pageNumber="84">D. brignolii</emphasis>
</taxonomicName>
, together with some species of the
<taxonomicName id="C6F54D3CE103AF02FD283FDEFCBFF825" authorityName="Kroneberg" authorityYear="1875" box="[691,787,1921,1947]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="3381EAADE103AF02FD283FDEFCBFF825" box="[691,787,1921,1947]" italics="true" pageId="11" pageNumber="84">aculeata</emphasis>
</taxonomicName>
group, viz.
<taxonomicName id="C6F54D3CE103AF02FC053FDDFB8AF825" authorityName="Kroneberg" authorityYear="1875" box="[926,1062,1922,1947]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="3381EAADE103AF02FC053FDDFB8AF825" box="[926,1062,1922,1947]" italics="true" pageId="11" pageNumber="84">D. tartarica</emphasis>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FBAE3FDDFB1CF825" authorityName="Dunin" authorityYear="1985" box="[1077,1200,1921,1947]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="pococki">
<emphasis id="3381EAADE103AF02FBAE3FDDFB1CF825" box="[1077,1200,1921,1947]" italics="true" pageId="11" pageNumber="84">D. pococki</emphasis>
</taxonomicName>
and
<taxonomicName id="C6F54D3CE103AF02FB723FDDFADBF825" authorityName="Dunin" authorityYear="1985" box="[1257,1399,1921,1947]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="limitanea">
<emphasis id="3381EAADE103AF02FB723FDDFADBF825" box="[1257,1399,1921,1947]" italics="true" pageId="11" pageNumber="84">D. limitanea</emphasis>
</taxonomicName>
, to the
<taxonomicName id="C6F54D3CE103AF02FF5B3FF9FE88F801" authorityName="Kroneberg" authorityYear="1875" box="[192,292,1958,1983]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="3381EAADE103AF02FF5B3FF9FE88F801" box="[192,292,1958,1983]" italics="true" pageId="11" pageNumber="84">tartarica</emphasis>
</taxonomicName>
group based on a characteristic leg spination, such as: coxae and patellae without spines, femora of all legs with spines. Later,
<bibRefCitation id="65644B4EE103AF02FE053F96FD99F85D" author="Dunin, P. M." box="[414,565,1993,2019]" pageId="11" pageNumber="84" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin (1992)</bibRefCitation>
put
<taxonomicName id="C6F54D3CE103AF02FDFD3F95FD46F85D" authorityName="Dunin" authorityYear="1989" box="[614,746,1993,2019]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FDFD3F95FD46F85D" box="[614,746,1993,2019]" italics="true" pageId="11" pageNumber="84">D. brignolii</emphasis>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FD633F95FC31F85D" authorityName="Dunin" authorityYear="1992" box="[760,925,1993,2019]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE103AF02FD633F95FC31F85D" box="[760,925,1993,2019]" italics="true" pageId="11" pageNumber="84">D. kugitangica</emphasis>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE103AF02FC303F95FBEEF85D" authorityName="Dunin" authorityYear="1989" box="[939,1090,1993,2019]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="kusnetsovi">
<emphasis id="3381EAADE103AF02FC303F95FBEEF85D" box="[939,1090,1993,2019]" italics="true" pageId="11" pageNumber="84">D. kusnetsovi</emphasis>
</taxonomicName>
and
<taxonomicName id="C6F54D3CE103AF02FBEC3F95FB43F85D" authorityName="Dunin" authorityYear="1989" box="[1143,1263,1993,2019]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE103AF02FBEC3F95FB43F85D" box="[1143,1263,1993,2019]" italics="true" pageId="11" pageNumber="84">D. nenilini</emphasis>
</taxonomicName>
to the
<taxonomicName id="C6F54D3CE103AF02FAA13F96FA30F85D" authorityName="Dunin" authorityYear="1989" box="[1338,1436,1993,2019]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE103AF02FAA13F96FA30F85D" box="[1338,1436,1993,2019]" italics="true" pageId="11" pageNumber="84">brignolii</emphasis>
</taxonomicName>
group without providing a diagnosis, which is given here (see above). The
<taxonomicName id="C6F54D3CE104AF05FC7C38C8FBE5FF0F" authorityName="Dunin" authorityYear="1989" box="[999,1097,151,177]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE104AF05FC7C38C8FBE5FF0F" box="[999,1097,151,177]" italics="true" pageId="12" pageNumber="85">brignolii</emphasis>
</taxonomicName>
species group has a very limited range, being known only from
<collectingCountry id="79E2762FE104AF05FDB038E4FD6AFF6B" box="[555,710,187,213]" name="Turkmenistan" pageId="12" pageNumber="85">Turkmenistan</collectingCountry>
(Kugitangtau and Kopet Dag Mt. Ranges, Karakum Desert and Sarykamysh Hollow) and the neighboring western part of
<collectingCountry id="79E2762FE104AF05FCB93880FC3CFF47" box="[802,912,223,249]" name="Tajikistan" pageId="12" pageNumber="85">Tajikistan</collectingCountry>
(Hissar Mt. Range).
</paragraph>
</subSubSection>
</treatment>
</document>

417
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<mods:title id="B6BC07529A30A431A902319ADB02469B">Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia</mods:title>
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<mods:affiliation id="D4F048171303DF0568AA010D424A1DDA">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia &amp; a. fomichov @ mail. ru; https: // orcid. org / 0000 - 0001 - 9268 - 622 X</mods:affiliation>
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<mods:affiliation id="DD7E9018180F9E4483144EFDDDB53962">Institute for Biological Problems of the North, Portovaya Street 18, Magadan 685000, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa &amp; yurmar @ mail. ru; https: // orcid. org / 0000 - 0002 - 4499 - 5148</mods:affiliation>
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<paragraph id="014A36BFE104AF05FF0C3E96FE44F95A" blockId="12.[151,718,1737,1801]" box="[151,488,1737,1764]" pageId="12" pageNumber="85">
<heading id="5A0281D3E104AF05FF0C3E96FE44F95A" bold="true" box="[151,488,1737,1764]" fontSize="11" level="1" pageId="12" pageNumber="85" reason="1">
<emphasis id="3381EAADE104AF05FF0C3E96FE44F95A" bold="true" box="[151,488,1737,1764]" pageId="12" pageNumber="85">
<taxonomicName id="C6F54D3CE104AF05FF0C3E96FE2FF95A" ID-CoL="99B43" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[151,387,1737,1764]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE104AF05FF0C3E96FE2FF95A" bold="true" box="[151,387,1737,1764]" italics="true" pageId="12" pageNumber="85">Dysdera mikhailovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28B257D6E104AF05FE113E95FE44F95A" box="[394,488,1738,1764]" pageId="12" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="49EF6534E104AF05FF0C3EB1FC1BF826" pageId="12" pageNumber="85" type="description">
<paragraph id="014A36BFE104AF05FF0C3EB1FD62F8B7" blockId="12.[151,718,1737,1801]" box="[151,718,1774,1801]" pageId="12" pageNumber="85">
<figureCitation id="99CE2A3AE104AF05FF0C3EB1FF56F8B7" box="[151,250,1774,1801]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="12" pageNumber="85">Figs 5–6</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FE9D3EB1FEE3F8B6" box="[262,335,1774,1800]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1791,1815]" captionTargetBox="[151,1435,181,1767]" captionTargetId="figure-19@5.[151,1435,181,1767]" captionTargetPageId="5" captionText="FIGURES 19–24. Prosoma of Dysdera mikhailovi sp. nov. male (19–21) and female (22–24). 19, 22—dorsal; 20, 23—lateral; 21, 24—ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157020" httpUri="https://zenodo.org/record/5157020/files/figure.png" pageId="12" pageNumber="85">19–24</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FEC13EB1FED4F8B6" box="[346,376,1774,1800]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[184,1408,193,1920]" captionTargetId="figure-25@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 25–30. Male palp (25–27) and bulb (28–30) of Dysdera subcylindrica (25, 28–30), D. dushengi (26) and D. mikhailovi sp. nov. (27). 25–28—prolateral; 29—anterior; 30—retrolateral. Scale bars: 25–27=0.5 mm, 28–30=0.2 mm. Ab- breviations: Ap–posterior apophysis, Ax–apex, C–crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157024" httpUri="https://zenodo.org/record/5157024/files/figure.png" pageId="12" pageNumber="85">27</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FE1F3EB1FE61F8B6" box="[388,461,1774,1800]" captionStart="FIGURES 41–46" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1434,419,1866]" captionTargetId="figure-177@10.[151,1436,417,1867]" captionTargetPageId="10" captionText="FIGURES 41–46. Psembolus of Dysdera dushengi (41–43) and bulb of D. mikhailovi sp. nov. (44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="12" pageNumber="85">44–46</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FE433EB1FD8DF8B6" box="[472,545,1774,1800]" captionStart="FIGURES 47–49" captionStartId="11.[151,264,1424,1448]" captionTargetBox="[151,1436,812,1399]" captionTargetId="figure-363@11.[151,1436,812,1400]" captionTargetPageId="11" captionText="FIGURES 47–49. Psembolus of Dysdera mikhailovi sp. nov. 47—prolateral; 48—anterior; 49—retrolateral. Scale bars=0.15 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest." figureDoi="http://doi.org/10.5281/zenodo.5157035" httpUri="https://zenodo.org/record/5157035/files/figure.png" pageId="12" pageNumber="85">47–49</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FDB73EB1FDD9F8B7" box="[556,629,1774,1801]" captionStart="FIGURES 50–57" captionStartId="12.[151,264,1524,1548]" captionTargetBox="[153,1436,273,1500]" captionTargetId="figure-73@12.[151,1436,273,1500]" captionTargetPageId="12" captionText="FIGURES 50–57. Endogyne of D. subcylindrica (50–51), D. dushengi (52–53), D. mikhailovi sp. nov. (54–55) and D. arnoldii (56–57). 50, 52, 54, 56—dorsal; 51, 53, 55, 57—ventral. Scale bars=0.2 mm.Abbreviations: Dd–dorsal arch of the diverticulum, La–lateral arm of the spermatheca, Pd–posterior diverticulum, Sk–stalk of the spermatheca, St–spermateca, Tb–transverse bar of the posterior diverticulum." figureDoi="http://doi.org/10.5281/zenodo.5157040" httpUri="https://zenodo.org/record/5157040/files/figure.png" pageId="12" pageNumber="85">54–55</figureCitation>
,
<figureCitation id="99CE2A3AE104AF05FD1B3EB1FD66F8B7" box="[640,714,1774,1801]" captionStart="FIGURES 58–60" captionStartId="13.[151,264,1657,1681]" captionTargetBox="[151,1437,669,1634]" captionTargetId="figure-289@13.[151,1437,669,1634]" captionTargetPageId="13" captionText="FIGURES 58–60. Collecting localities of the studied Dysdera species (58–59) and published records of other Central Asian Dysdera species, except for D. crocata (60). Triangle—Dysdera arnoldii, diamond—D. mikhailovi sp. nov., circle—D. dushengi, square—D. subcylindrica. Black symbols refer to the studied material, white symbols refer to literature-derived data. Some symbols could refer to more than one closely situated localities. The frame of Fig. 59 refers to the content of Fig. 58. The numbers in Fig. 60 refer to the following species: 1—D. aculeata; 2—D. arnoldii; 3—D. brignolii; 4—D. dushengi; 5—D. fedtschenkoi Dunin, 1992; 6—D. ferghanica Dunin, 1985; 7—D. kronebergi Dunin, 1992; 8—D. kugitangica; 9—D. kusnetsovi; 10—D. limitanea; 11—D. nenilini; 12—D. pamirica, 13—D. pococki; 14—D. subcylindrica; 15—D. tartarica; 16—D. tystshenkoi Dunin, 1989; 17—D. zarudnyi; 18—D. zonsteini Dimitrov, 2021; 19—D. mikhailovi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5157042" httpUri="https://zenodo.org/record/5157042/files/figure.png" pageId="12" pageNumber="85">58–60</figureCitation>
.
</paragraph>
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<emphasis id="3381EAADE104AF05FF0C3F69FF48F8EE" bold="true" box="[151,228,1846,1872]" pageId="12" pageNumber="85">
<collectingRegion id="C331F85DE104AF05FF0C3F69FF48F8EE" box="[151,228,1846,1872]" pageId="12" pageNumber="85">Types.</collectingRegion>
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,
<emphasis id="3381EAADE104AF05FE0C3F69FCFFF8EE" bold="true" box="[407,851,1845,1872]" italics="true" pageId="12" pageNumber="85">
<collectingRegion id="C331F85DE104AF05FE0C3F69FCFFF8EE" box="[407,851,1845,1872]" pageId="12" pageNumber="85">Districts of Republican Subordination</collectingRegion>
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:
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<specimenCount id="17F3FD36E104AF05FC563F68FC4EF8EF" box="[973,994,1847,1873]" pageId="12" pageNumber="85" type="male">♂</specimenCount>
(
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<typeStatus id="DE4E881DE104AF05FB163F68FB54F8EF" box="[1165,1272,1847,1873]" pageId="12" pageNumber="85" type="paratype">paratypes</typeStatus>
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(
<collectionCode id="67E4AE7AE104AF05FAA83F69FA3CF8EE" box="[1331,1424,1846,1872]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="85" type="Museum">ZMMU</collectionCode>
), nr.
<collectingRegion id="C331F85DE104AF05FF273F05FE83F8CA" box="[188,303,1882,1908]" country="Tajikistan" name="Dushanbe" pageId="12" pageNumber="85">Dushanbe</collectingRegion>
,
<collectingMunicipality id="E12EACC5E104AF05FEA73F05FDAAF8CB" box="[316,518,1882,1909]" pageId="12" pageNumber="85">Hissar Mt. Range</collectingMunicipality>
, 48th km of
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,
<geoCoordinate id="64C15078E104AF05FCBC3F05FC15F8CB" box="[807,953,1882,1909]" degrees="38" direction="north" minutes="55.531" orientation="latitude" pageId="12" pageNumber="85" precision="1" value="38.925518">38°55.531’N</geoCoordinate>
<geoCoordinate id="64C15078E104AF05FC5A3F05FBFEF8CA" box="[961,1106,1882,1908]" degrees="68" direction="east" minutes="48.292" orientation="longitude" pageId="12" pageNumber="85" precision="1" value="68.80486">68°48.292’E</geoCoordinate>
, southern slope with
<taxonomicName id="C6F54D3CE104AF05FADE3F05FA30F8CA" authorityName="Linnaeus" authorityYear="1753" box="[1349,1436,1882,1908]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="3381EAADE104AF05FADE3F05FA30F8CA" box="[1349,1436,1882,1908]" italics="true" pageId="12" pageNumber="85">Juglans</emphasis>
</taxonomicName>
litter and under stones,
<quantity id="C60D9B5AE104AF05FE063F21FE58F827" box="[413,500,1918,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="12" pageNumber="85" unit="m" value="1530.0">
<elevation id="8AD8D18CE104AF05FE063F21FE58F827" box="[413,500,1918,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="12" pageNumber="85" unit="m" value="1530.0">1530 m</elevation>
</quantity>
,
<date id="754B107FE104AF05FD9B3F21FDDEF827" box="[512,626,1918,1945]" pageId="12" pageNumber="85" value="2015-05-07">
<collectingDate id="650FE997E104AF05FD9B3F21FDDEF827" box="[512,626,1918,1945]" pageId="12" pageNumber="85" value="2015-05-07">7.05.2015</collectingDate>
</date>
,
<collectorName id="AC005369E104AF05FDE63F20FCF4F827" box="[637,856,1918,1945]" pageId="12" pageNumber="85">Yu. M. Marusik, M</collectorName>
. Saidov
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E104AF04FF5C3FFDFE2BFF47" lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85" type="etymology">
<paragraph id="014A36BFE104AF04FF5C3FFDFE2BFF47" blockId="12.[151,1437,1845,2017]" lastBlockId="13.[151,1437,151,645]" lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85">
<emphasis id="3381EAADE104AF05FF5C3FFDFEE2F802" bold="true" box="[199,334,1954,1980]" pageId="12" pageNumber="85">Etymology.</emphasis>
The new species is dedicated to our friend and colleague, Dr Kirill G. Mikhailov (
<collectingRegion id="C331F85DE104AF05FB633FFDFAF7F802" box="[1272,1371,1954,1980]" country="Russia" name="Moscow" pageId="12" pageNumber="85">Moscow</collectingRegion>
, Rus- sia), on the occasion of his 60th birthday. Kirill has made a great contribution to the taxonomy and the catalouging of spiders of the former Soviet Union, and an unprecedented contribution to biological sciences by establishing the publishing house ‘KMK Scientific Press’ (in 1992) which produces a set of academic biological/taxonomic journals and numerous books.
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E105AF04FF5C395DFB16FD3B" pageId="13" pageNumber="86" type="diagnosis">
<paragraph id="014A36BFE105AF04FF5C395DFB16FD3B" blockId="13.[151,1437,151,645]" pageId="13" pageNumber="86">
<emphasis id="3381EAADE105AF04FF5C395DFEEEFEA2" bold="true" box="[199,322,258,284]" pageId="13" pageNumber="86">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="C6F54D3CE105AF04FD38395CFC85FEA3" authorityName="Dunin" authorityYear="1989" box="[675,809,259,285]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FD38395CFC85FEA3" box="[675,809,259,285]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
,
<taxonomicName id="C6F54D3CE105AF04FCA1395CFC4EFEA3" authorityName="Dunin" authorityYear="1992" box="[826,994,259,285]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE105AF04FCA1395CFC4EFEA3" box="[826,994,259,285]" italics="true" pageId="13" pageNumber="86">D. kugitangica</emphasis>
</taxonomicName>
and
<taxonomicName id="C6F54D3CE105AF04FB86395CFB34FEA3" authorityName="Dunin" authorityYear="1989" box="[1053,1176,259,285]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FB86395CFB34FEA3" box="[1053,1176,259,285]" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
in having the strongly prominent semi-oval median crest (
<emphasis id="3381EAADE105AF04FDBE3978FDE0FEFF" box="[549,588,295,321]" italics="true" pageId="13" pageNumber="86">Cm</emphasis>
) and similar proportions of the bulb: viz., the psembolus 1.2–1.6 times lon- ger than tegulum (
<emphasis id="3381EAADE105AF04FEF33914FE2DFEDA" box="[360,385,331,356]" italics="true" pageId="13" pageNumber="86">Te</emphasis>
). The male of
<taxonomicName id="C6F54D3CE105AF04FDB23914FD6EFEDB" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[553,706,331,357]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE105AF04FDB23914FD6EFEDB" box="[553,706,331,357]" italics="true" pageId="13" pageNumber="86">D. mikhailovi</emphasis>
</taxonomicName>
<emphasis id="3381EAADE105AF04FD513914FC88FEDB" bold="true" box="[714,804,331,357]" pageId="13" pageNumber="86">
<taxonomicNameLabel id="28B257D6E105AF04FD513914FC88FEDB" box="[714,804,331,357]" pageId="13" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from that of
<taxonomicName id="C6F54D3CE105AF04FBE63914FAAEFEDB" authorityName="Dunin" authorityYear="1989" box="[1149,1282,331,357]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FBE63914FAAEFEDB" box="[1149,1282,331,357]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
by the psembolus stem (
<emphasis id="3381EAADE105AF04FEBB3930FE96FE37" box="[288,314,367,393]" italics="true" pageId="13" pageNumber="86">Se</emphasis>
) being two times thinner than the tegulum (equal in
<taxonomicName id="C6F54D3CE105AF04FC1C3930FBA7FE37" authorityName="Dunin" authorityYear="1989" box="[903,1035,367,393]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FC1C3930FBA7FE37" box="[903,1035,367,393]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
) and the median crest length/height ratio 3.3 (
<quantity id="C60D9B5AE105AF04FE9F39CCFEEAFE13" box="[260,326,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="13" pageNumber="86" unit="in" value="2.3">2.3 in</quantity>
<taxonomicName id="C6F54D3CE105AF04FED639CCFE7DFE13" authorityName="Dunin" authorityYear="1989" box="[333,465,403,429]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FED639CCFE7DFE13" box="[333,465,403,429]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
). The male of the new species can be distinguished from that of
<taxonomicName id="C6F54D3CE105AF04FB3239CCFAE3FE13" authorityName="Dunin" authorityYear="1992" box="[1193,1359,403,429]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE105AF04FB3239CCFAE3FE13" box="[1193,1359,403,429]" italics="true" pageId="13" pageNumber="86">D. kugitangica</emphasis>
</taxonomicName>
by the rounded proximal part of the median crest and the retrolateral crest (
<emphasis id="3381EAADE105AF04FC1439E8FC01FE6F" box="[911,941,439,465]" italics="true" pageId="13" pageNumber="86">Cr</emphasis>
) divided into two lobes (solid in
<taxonomicName id="C6F54D3CE105AF04FA8639E8FF6BFE4A" authorityName="Dunin" authorityYear="1992" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE105AF04FA8639E8FF6BFE4A" italics="true" pageId="13" pageNumber="86">D. kugitangica</emphasis>
</taxonomicName>
). From
<taxonomicName id="C6F54D3CE105AF04FEB83984FE30FE4B" authorityName="Dunin" authorityYear="1989" box="[291,412,475,501]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FEB83984FE30FE4B" box="[291,412,475,501]" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
the male can be distinguished by the rounded distal part of the median crest (pointed in
<taxonomicName id="C6F54D3CE105AF04FA1A3984FF41FDA7" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FA1A3984FF41FDA7" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
) and the curved bulb, forming an angle of 130° between the tegulum and the psembolus (tegulum and psem- bolus stretched out in almost a straight line in
<taxonomicName id="C6F54D3CE105AF04FD3E3A7CFCB3FD83" authorityName="Dunin" authorityYear="1989" box="[677,799,547,573]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FD3E3A7CFCB3FD83" box="[677,799,547,573]" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
). The female of
<taxonomicName id="C6F54D3CE105AF04FC463A7CFBDAFD83" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[989,1142,547,573]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE105AF04FC463A7CFBDAFD83" box="[989,1142,547,573]" italics="true" pageId="13" pageNumber="86">D. mikhailovi</emphasis>
</taxonomicName>
<emphasis id="3381EAADE105AF04FBE43A7CFB75FD83" bold="true" box="[1151,1241,547,573]" pageId="13" pageNumber="86">
<taxonomicNameLabel id="28B257D6E105AF04FBE43A7CFB75FD83" box="[1151,1241,547,573]" pageId="13" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from that of
<taxonomicName id="C6F54D3CE105AF04FEBF3A18FE66FDDF" authorityName="Dunin" authorityYear="1992" box="[292,458,583,609]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE105AF04FEBF3A18FE66FDDF" box="[292,458,583,609]" italics="true" pageId="13" pageNumber="86">D. kugitangica</emphasis>
</taxonomicName>
by the absence of the spermatheca stalk; from that of
<taxonomicName id="C6F54D3CE105AF04FBBD3A18FB32FDDF" authorityName="Dunin" authorityYear="1989" box="[1062,1182,583,609]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FBBD3A18FB32FDDF" box="[1062,1182,583,609]" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
it differs in having the rectangular rather than triangular spermatheca (
<emphasis id="3381EAADE105AF04FD373A34FD6EFD3B" box="[684,706,619,645]" italics="true" pageId="13" pageNumber="86">St</emphasis>
). The female of
<taxonomicName id="C6F54D3CE105AF04FCE73A34FBADFD3B" authorityName="Dunin" authorityYear="1989" box="[892,1025,619,645]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FCE73A34FBADFD3B" box="[892,1025,619,645]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
is yet unknown.
</paragraph>
</subSubSection>
<caption id="558A6637E105AF04FF0C3E26FB11F833" ID-DOI="http://doi.org/10.5281/zenodo.5157042" ID-Zenodo-Dep="5157042" httpUri="https://zenodo.org/record/5157042/files/figure.png" pageId="13" pageNumber="86" startId="13.[151,264,1657,1681]" targetBox="[151,1437,669,1634]" targetPageId="13">
<paragraph id="014A36BFE105AF04FF0C3E26FB11F833" blockId="13.[151,1437,1657,1934]" pageId="13" pageNumber="86">
<emphasis id="3381EAADE105AF04FF0C3E26FEFAF92F" bold="true" box="[151,342,1657,1681]" pageId="13" pageNumber="86">FIGURES 58–60.</emphasis>
Collecting localities of the studied
<taxonomicName id="C6F54D3CE105AF04FD5C3E26FCC2F92F" box="[711,878,1657,1681]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3381EAADE105AF04FD5C3E26FCB1F92F" box="[711,797,1657,1681]" italics="true" pageId="13" pageNumber="86">Dysdera</emphasis>
species
</taxonomicName>
(58–59) and published records of other Central Asian
<taxonomicName id="C6F54D3CE105AF04FF0C3EC2FEEDF90B" box="[151,321,1693,1717]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3381EAADE105AF04FF0C3EC2FF41F90B" box="[151,237,1693,1717]" italics="true" pageId="13" pageNumber="86">Dysdera</emphasis>
species
</taxonomicName>
, except for
<taxonomicName id="C6F54D3CE105AF04FE213EC1FD8AF90B" authorityName="C. L. Koch" authorityYear="1838" box="[442,550,1694,1717]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="crocata">
<emphasis id="3381EAADE105AF04FE213EC1FD8AF90B" box="[442,550,1694,1717]" italics="true" pageId="13" pageNumber="86">D. crocata</emphasis>
</taxonomicName>
(60). Triangle—
<taxonomicName id="C6F54D3CE105AF04FD4F3EC2FC2FF90B" authorityName="Charitonov" authorityYear="1956" box="[724,899,1693,1717]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="arnoldii">
<emphasis id="3381EAADE105AF04FD4F3EC2FC2FF90B" box="[724,899,1693,1717]" italics="true" pageId="13" pageNumber="86">Dysdera arnoldii</emphasis>
</taxonomicName>
, diamond—
<taxonomicName id="C6F54D3CE105AF04FB983EC1FB22F90B" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[1027,1166,1693,1717]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE105AF04FB983EC1FB22F90B" box="[1027,1166,1693,1717]" italics="true" pageId="13" pageNumber="86">D. mikhailovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28B257D6E105AF04FB0E3EC1FB4BF90B" box="[1173,1255,1694,1717]" pageId="13" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, circle—
<taxonomicName id="C6F54D3CE105AF04FADF3EC1FF6FF967" authorityName="Lin, Chang et Li" authorityYear="2020" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE105AF04FADF3EC1FF6FF967" italics="true" pageId="13" pageNumber="86">D. dushengi</emphasis>
</taxonomicName>
, square—
<taxonomicName id="C6F54D3CE105AF04FEB53E9DFE76F967" authorityName=": Marusik" authorityYear="2017" box="[302,474,1729,1753]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="subcylindrica">
<emphasis id="3381EAADE105AF04FEB53E9DFE76F967" box="[302,474,1729,1753]" italics="true" pageId="13" pageNumber="86">D. subcylindrica</emphasis>
</taxonomicName>
. Black symbols refer to the studied material, white symbols refer to literature-derived data. Some symbols could refer to more than one closely situated localities. The frame of Fig. 59 refers to the content of Fig. 58. The numbers in Fig. 60 refer to the following species: 1—
<taxonomicName id="C6F54D3CE105AF04FD573F55FCE8F89F" authorityName="Kroneberg" authorityYear="1875" box="[716,836,1801,1825]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="3381EAADE105AF04FD573F55FCE8F89F" box="[716,836,1801,1825]" italics="true" pageId="13" pageNumber="86">D. aculeata</emphasis>
</taxonomicName>
; 2—
<taxonomicName id="C6F54D3CE105AF04FCE13F55FC41F89F" authorityName="Charitonov" authorityYear="1956" box="[890,1005,1801,1825]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="arnoldii">
<emphasis id="3381EAADE105AF04FCE13F55FC41F89F" box="[890,1005,1801,1825]" italics="true" pageId="13" pageNumber="86">D. arnoldii</emphasis>
</taxonomicName>
; 3—
<taxonomicName id="C6F54D3CE105AF04FBB93F55FB30F89F" authorityName="Dunin" authorityYear="1989" box="[1058,1180,1801,1825]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="3381EAADE105AF04FBB93F55FB30F89F" box="[1058,1180,1801,1825]" italics="true" pageId="13" pageNumber="86">D. brignolii</emphasis>
</taxonomicName>
; 4—
<taxonomicName id="C6F54D3CE105AF04FB4A3F55FAE2F89F" authorityName="Lin, Chang et Li" authorityYear="2020" box="[1233,1358,1801,1825]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="dushengi">
<emphasis id="3381EAADE105AF04FB4A3F55FAE2F89F" box="[1233,1358,1801,1825]" italics="true" pageId="13" pageNumber="86">D. dushengi</emphasis>
</taxonomicName>
; 5—
<taxonomicName id="C6F54D3CE105AF04FA1F3F55FE0DF8F8" authority="Dunin, 1992" authorityName="Dunin" authorityYear="1992" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="3381EAADE105AF04FA1F3F55FEBAF8FB" italics="true" pageId="13" pageNumber="86">D. fedtschenkoi</emphasis>
<bibRefCitation id="65644B4EE105AF04FE863F72FE0DF8F8" author="Dunin, P. M." box="[285,417,1837,1862]" pageId="13" pageNumber="86" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin, 1992</bibRefCitation>
</taxonomicName>
; 6—
<taxonomicName id="C6F54D3CE105AF04FE493F71FD47F8FB" authority="Dunin, 1985" authorityName="Dunin" authorityYear="1985" box="[466,747,1837,1861]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="ferghanica">
<emphasis id="3381EAADE105AF04FE493F71FDCCF8FB" box="[466,608,1837,1861]" italics="true" pageId="13" pageNumber="86">D. ferghanica</emphasis>
<bibRefCitation id="65644B4EE105AF04FDFC3F72FD47F8FB" author="Dunin, P. M." box="[615,747,1837,1861]" pageId="13" pageNumber="86" pagination="114 - 120" refId="ref7702" refString="Dunin, P. M. (1985) The spider family Dysderidae (Aranei, Haplogynae) in the Soviet Central Asia. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 139, 114 - 120. [in Russian with English summary]" type="journal article" year="1985">Dunin, 1985</bibRefCitation>
</taxonomicName>
; 7—
<taxonomicName id="C6F54D3CE105AF04FC873F71FB99F8F8" authority="Dunin, 1992" authorityName="Dunin" authorityYear="1992" box="[796,1077,1837,1862]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kronebergi">
<emphasis id="3381EAADE105AF04FC873F71FC05F8FB" box="[796,937,1837,1861]" italics="true" pageId="13" pageNumber="86">D. kronebergi</emphasis>
<bibRefCitation id="65644B4EE105AF04FC2A3F72FB99F8F8" author="Dunin, P. M." box="[945,1077,1837,1862]" pageId="13" pageNumber="86" pagination="136 - 140" refId="ref7789" refString="Dunin, P. M. (1992) New species of Dysdera spiders (Aranei, Haplogynae, Dysderidae) from Middle Asia. Zoologicheskii Zhur- nal, 71 (2), 136 - 140. [in Russian with English summary]" type="journal article" year="1992">Dunin, 1992</bibRefCitation>
</taxonomicName>
; 8—
<taxonomicName id="C6F54D3CE105AF04FBFD3F71FB52F8FB" authorityName="Dunin" authorityYear="1992" box="[1126,1278,1837,1861]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kugitangica">
<emphasis id="3381EAADE105AF04FBFD3F71FB52F8FB" box="[1126,1278,1837,1861]" italics="true" pageId="13" pageNumber="86">D. kugitangica</emphasis>
</taxonomicName>
; 9—
<taxonomicName id="C6F54D3CE105AF04FAA83F71FF6CF8D7" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="kusnetsovi">
<emphasis id="3381EAADE105AF04FAA83F71FF6CF8D7" italics="true" pageId="13" pageNumber="86">D. kusnetsovi</emphasis>
</taxonomicName>
; 10—
<taxonomicName id="C6F54D3CE105AF04FE9A3F0DFE2DF8D7" authorityName="Dunin" authorityYear="1985" box="[257,385,1873,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="limitanea">
<emphasis id="3381EAADE105AF04FE9A3F0DFE2DF8D7" box="[257,385,1873,1897]" italics="true" pageId="13" pageNumber="86">D. limitanea</emphasis>
</taxonomicName>
; 11—
<taxonomicName id="C6F54D3CE105AF04FE5A3F0DFD83F8D7" authorityName="Dunin" authorityYear="1989" box="[449,559,1873,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="3381EAADE105AF04FE5A3F0DFD83F8D7" box="[449,559,1873,1897]" italics="true" pageId="13" pageNumber="86">D. nenilini</emphasis>
</taxonomicName>
; 12—
<taxonomicName id="C6F54D3CE105AF04FDEB3F0DFD47F8D7" authorityName="Dunin" authorityYear="1992" box="[624,747,1874,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="pamirica">
<emphasis id="3381EAADE105AF04FDEB3F0DFD47F8D7" box="[624,747,1874,1897]" italics="true" pageId="13" pageNumber="86">D. pamirica</emphasis>
</taxonomicName>
, 13—
<taxonomicName id="C6F54D3CE105AF04FCB63F0DFC30F8D7" authorityName="Dunin" authorityYear="1985" box="[813,924,1873,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="pococki">
<emphasis id="3381EAADE105AF04FCB63F0DFC30F8D7" box="[813,924,1873,1897]" italics="true" pageId="13" pageNumber="86">D. pococki</emphasis>
</taxonomicName>
; 14—
<taxonomicName id="C6F54D3CE105AF04FC463F0DFB2BF8D7" authorityName=": Marusik" authorityYear="2017" box="[989,1159,1873,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="subcylindrica">
<emphasis id="3381EAADE105AF04FC463F0DFB2BF8D7" box="[989,1159,1873,1897]" italics="true" pageId="13" pageNumber="86">D. subcylindrica</emphasis>
</taxonomicName>
; 15—
<taxonomicName id="C6F54D3CE105AF04FB533F0DFAEEF8D7" authorityName="Kroneberg" authorityYear="1875" box="[1224,1346,1874,1897]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="3381EAADE105AF04FB533F0DFAEEF8D7" box="[1224,1346,1874,1897]" italics="true" pageId="13" pageNumber="86">D. tartarica</emphasis>
</taxonomicName>
; 16—
<taxonomicName id="C6F54D3CE105AF04FA1F3F0DFE3CF830" authority="Dunin, 1989" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="tystshenkoi">
<emphasis id="3381EAADE105AF04FA1F3F0DFEA4F833" italics="true" pageId="13" pageNumber="86">D. tystshenkoi</emphasis>
<bibRefCitation id="65644B4EE105AF04FE953F2AFE3CF830" author="Dunin, P. M." box="[270,400,1909,1934]" pageId="13" pageNumber="86" pagination="865 - 874" refId="ref7748" refString="Dunin, P. M. (1989) Material on the fauna of spiders of the family Dysderidae (Aranei) from Turkmenistan. Entomologicheskoe Obozrenie, 68 (4), 865 - 874. [in Russian]" type="journal article" year="1989">Dunin, 1989</bibRefCitation>
</taxonomicName>
; 17—
<taxonomicName id="C6F54D3CE105AF04FE563F29FDE9F833" authorityName="Charitonov" authorityYear="1956" box="[461,581,1909,1933]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="3381EAADE105AF04FE563F29FDE9F833" box="[461,581,1909,1933]" italics="true" pageId="13" pageNumber="86">D. zarudnyi</emphasis>
</taxonomicName>
; 18—
<taxonomicName id="C6F54D3CE105AF04FD1F3F29FC0EF830" authority="Dimitrov, 2021" authorityName="Dimitrov" authorityYear="2021" box="[644,930,1909,1934]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="zonsteini">
<emphasis id="3381EAADE105AF04FD1F3F29FD51F833" box="[644,765,1910,1933]" italics="true" pageId="13" pageNumber="86">D. zonsteini</emphasis>
<bibRefCitation id="65644B4EE105AF04FC9F3F2AFC0EF830" author="Dimitrov, D." box="[772,930,1909,1934]" pageId="13" pageNumber="86" pagination="588 - 594" refId="ref7654" refString="Dimitrov, D. (2021) Description of Dysdera zonsteini n. sp. (Arachnida: Araneae: Dysderidae) from Turkmenistan. Zootaxa, 4938 (5), 588 - 594. https: // doi. org / 10.11646 / zootaxa. 4938.5.6" type="journal article" year="2021">Dimitrov, 2021</bibRefCitation>
</taxonomicName>
; 19—
<taxonomicName id="C6F54D3CE105AF04FC453F29FBC4F833" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[990,1128,1909,1933]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="mikhailovi" status="sp. nov.">
<emphasis id="3381EAADE105AF04FC453F29FBC4F833" box="[990,1128,1909,1933]" italics="true" pageId="13" pageNumber="86">D. mikhailovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28B257D6E105AF04FBF43F29FB11F833" box="[1135,1213,1910,1933]" pageId="13" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="558A6637E106AF07FF0C3A80FDF7FCA5" ID-DOI="http://doi.org/10.5281/zenodo.5157044" ID-Zenodo-Dep="5157044" httpUri="https://zenodo.org/record/5157044/files/figure.png" pageId="14" pageNumber="87" startId="14.[151,250,735,759]" targetBox="[151,1436,181,711]" targetPageId="14">
<paragraph id="014A36BFE106AF07FF0C3A80FDF7FCA5" blockId="14.[151,1436,735,795]" pageId="14" pageNumber="87">
<emphasis id="3381EAADE106AF07FF0C3A80FE8CFD49" bold="true" box="[151,288,735,759]" pageId="14" pageNumber="87">FIGURE 61.</emphasis>
Present centers of species diversity of
<taxonomicName id="C6F54D3CE106AF07FD363A80FCAFFD49" authorityName="Latreille" authorityYear="1804" box="[685,771,735,759]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="3381EAADE106AF07FD363A80FCAFFD49" box="[685,771,735,759]" italics="true" pageId="14" pageNumber="87">Dysdera</emphasis>
</taxonomicName>
in Central Asia. I—Kopet Dag Mts; II—Kugitangtau Mt. Range; III—Hissar Mt. Range; IV—Fergana Valley.
</paragraph>
</caption>
<subSubSection id="49EF6534E106AF07FF5C3B16FA34FA1D" pageId="14" pageNumber="87" type="description">
<paragraph id="014A36BFE106AF07FF5C3B16FB53FBE1" blockId="14.[151,1437,841,1623]" pageId="14" pageNumber="87">
<emphasis id="3381EAADE106AF07FF5C3B16FEF5FCDD" bold="true" box="[199,345,841,867]" pageId="14" pageNumber="87">Description.</emphasis>
Male. Total length 5.7. Carapace: 2.85 long, 2.05 wide. Abdomen: 2.95 long, 1.55 wide. AME 0.14, PME 0.11, PLE 0.11. General appearance as in
<figureCitation id="99CE2A3AE106AF07FD683B32FC90FC36" box="[755,828,877,904]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="14" pageNumber="87">Figs 5</figureCitation>
,
<figureCitation id="99CE2A3AE106AF07FCD23B32FC3EFC39" box="[841,914,877,903]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1791,1815]" captionTargetBox="[151,1435,181,1767]" captionTargetId="figure-19@5.[151,1435,181,1767]" captionTargetPageId="5" captionText="FIGURES 19–24. Prosoma of Dysdera mikhailovi sp. nov. male (19–21) and female (22–24). 19, 22—dorsal; 20, 23—lateral; 21, 24—ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157020" httpUri="https://zenodo.org/record/5157020/files/figure.png" pageId="14" pageNumber="87">19–21</figureCitation>
. Carapace pale-red. Sternum, labium and chelicerae orange. Maxillae and palps yellow-orange. Legs: I and II yellow-orange, III and IV yellow. Abdomen and spinnerets beige. Leg measurements: I: 2.3, 1.4, 1.95, 1.9, 0.5 (8.05). II: 2.05, 1.3, 1.75, 1.8, 0.5 (7.4). III: 1.65, 0.95, 1.15, 1.55, 0.8 (6.1). IV: 2.1, 1.1, 1.65, 1.95, 0.5 (7.3). Leg spination: I: Fe p2. II: Fe p1. III: Fe d1 p1; Ti p4 r2 v6; Mt p6 r2 v2. IV: Fe d5; Ti p3 r2 v3; Mt p4 r2 v5. Bulb as in
<figureCitation id="99CE2A3AE106AF07FCB13BA2FCD3FBA9" box="[810,895,1021,1047]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[184,1408,193,1920]" captionTargetId="figure-25@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 25–30. Male palp (25–27) and bulb (28–30) of Dysdera subcylindrica (25, 28–30), D. dushengi (26) and D. mikhailovi sp. nov. (27). 25–28—prolateral; 29—anterior; 30—retrolateral. Scale bars: 25–27=0.5 mm, 28–30=0.2 mm. Ab- breviations: Ap–posterior apophysis, Ax–apex, C–crest, Se–stem of the psembolus, Sp–spine, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157024" httpUri="https://zenodo.org/record/5157024/files/figure.png" pageId="14" pageNumber="87">Figs 27</figureCitation>
,
<figureCitation id="99CE2A3AE106AF07FC113BA2FC7FFBA9" box="[906,979,1021,1047]" captionStart="FIGURES 41–46" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1434,419,1866]" captionTargetId="figure-177@10.[151,1436,417,1867]" captionTargetPageId="10" captionText="FIGURES 41–46. Psembolus of Dysdera dushengi (41–43) and bulb of D. mikhailovi sp. nov. (44–46). 41, 44—prolateral; 42, 45—anterior; 43, 46—retrolateral. Scale bars: 41–43=0.15 mm, 44–46=0.2 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest, Se–stem of the psembolus, Te–tegulum." figureDoi="http://doi.org/10.5281/zenodo.5157033" httpUri="https://zenodo.org/record/5157033/files/figure.png" pageId="14" pageNumber="87">44–46</figureCitation>
,
<figureCitation id="99CE2A3AE106AF07FC463BA2FB8AFBA9" box="[989,1062,1021,1047]" captionStart="FIGURES 47–49" captionStartId="11.[151,264,1424,1448]" captionTargetBox="[151,1436,812,1399]" captionTargetId="figure-363@11.[151,1436,812,1400]" captionTargetPageId="11" captionText="FIGURES 47–49. Psembolus of Dysdera mikhailovi sp. nov. 47—prolateral; 48—anterior; 49—retrolateral. Scale bars=0.15 mm. Abbreviations: Ap–posterior apophysis, Ax–apex, Cm–median crest, Cp–prolateral crest, Cr–retrolateral crest." figureDoi="http://doi.org/10.5281/zenodo.5157035" httpUri="https://zenodo.org/record/5157035/files/figure.png" pageId="14" pageNumber="87">47–49</figureCitation>
. Length/width ratio 2.8. Posterior apophysis (
<emphasis id="3381EAADE106AF07FE8C3C7DFE9AFB82" box="[279,310,1058,1084]" italics="true" pageId="14" pageNumber="87">Ap</emphasis>
) leaf-shaped in lateral view. Psembolus with a prominent semi-oval median crest, fusiform prolateral crest (
<emphasis id="3381EAADE106AF07FF473C19FF51FBDE" box="[220,253,1094,1120]" italics="true" pageId="14" pageNumber="87">Cp</emphasis>
) and oblong retrolateral crest which is divided into two lobes. Psembolus apex (
<emphasis id="3381EAADE106AF07FB193C19FB0CFBDE" box="[1154,1184,1094,1120]" italics="true" pageId="14" pageNumber="87">Ax</emphasis>
) abrupt.
</paragraph>
<paragraph id="014A36BFE106AF07FF5C3C36FC37FAC1" blockId="14.[151,1437,841,1623]" pageId="14" pageNumber="87">
Female. Total length 11.0. Carapace: 4.7 long, 3.4 wide. Abdomen: 6.3 long, 3.45 wide. AME 0.21, PME 0.16, PLE 0.19. General appearance as in
<figureCitation id="99CE2A3AE106AF07FDA43CD2FD25FB19" box="[575,649,1165,1191]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1908,1932]" captionTargetBox="[160,1433,404,1873]" captionTargetId="figure-78@2.[151,1436,394,1883]" captionTargetPageId="2" captionText="FIGURES 1–6. Dorsal habitus of Dysdera subcylindrica (1–2), D. dushengi (3–4) and D. mikhailovi sp. nov. (5–6). 1, 3, 5—male; 2, 4, 6—female. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5157010" httpUri="https://zenodo.org/record/5157010/files/figure.png" pageId="14" pageNumber="87">Figs 6</figureCitation>
,
<figureCitation id="99CE2A3AE106AF07FD033CD2FD4DFB19" box="[664,737,1165,1191]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1791,1815]" captionTargetBox="[151,1435,181,1767]" captionTargetId="figure-19@5.[151,1435,181,1767]" captionTargetPageId="5" captionText="FIGURES 19–24. Prosoma of Dysdera mikhailovi sp. nov. male (19–21) and female (22–24). 19, 22—dorsal; 20, 23—lateral; 21, 24—ventral. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.5157020" httpUri="https://zenodo.org/record/5157020/files/figure.png" pageId="14" pageNumber="87">22–24</figureCitation>
. Carapace and chelicerae red. Sternum, labium and maxillae orange. Palps yellow-orange. Legs: I and II yellow-orange, III and IV yellow. Abdomen and spinnerets beige. Leg measurements: 3.75, 2.35, 3.25, 3.0, 0.65 (13.0). II: 3.55, 2.2, 3.05, 2.95, 0.65 (12.4). III: 2.9, 1.65, 2.0, 2.65, 0.6 (9.8). IV: 3.75, 2.0, 3.0, 3.6, 0.7 (13.05). Leg spination: III: Ti p4 r2 v3; Mt p6 r2 v3. IV: Fe d2; Ti p4 r2 v6; Mt p5 r3 v5. Endogyne as in
<figureCitation id="99CE2A3AE106AF07FEE93D42FE5EFA86" box="[370,498,1309,1336]" captionStart="FIGURES 50–57" captionStartId="12.[151,264,1524,1548]" captionTargetBox="[153,1436,273,1500]" captionTargetId="figure-73@12.[151,1436,273,1500]" captionTargetPageId="12" captionText="FIGURES 50–57. Endogyne of D. subcylindrica (50–51), D. dushengi (52–53), D. mikhailovi sp. nov. (54–55) and D. arnoldii (56–57). 50, 52, 54, 56—dorsal; 51, 53, 55, 57—ventral. Scale bars=0.2 mm.Abbreviations: Dd–dorsal arch of the diverticulum, La–lateral arm of the spermatheca, Pd–posterior diverticulum, Sk–stalk of the spermatheca, St–spermateca, Tb–transverse bar of the posterior diverticulum." figureDoi="http://doi.org/10.5281/zenodo.5157040" httpUri="https://zenodo.org/record/5157040/files/figure.png" pageId="14" pageNumber="87">Figs 54–55</figureCitation>
. Spermatheca rectangular, 3.4 times as wide as long. Dorsal arch of the diverticulum (
<emphasis id="3381EAADE106AF07FF3A3D1EFF6FFAE5" box="[161,195,1345,1371]" italics="true" pageId="14" pageNumber="87">Dd</emphasis>
) low partially overlaps by spermatheca. Transverse bar of posterior diverticulum (
<emphasis id="3381EAADE106AF07FBE33D1EFB3AFAE5" box="[1144,1174,1345,1371]" italics="true" pageId="14" pageNumber="87">Tb</emphasis>
) pentagonal, 1.3 times wider than spermatheca. Posterior diverticulum (
<emphasis id="3381EAADE106AF07FD203D3AFD76FAC1" box="[699,730,1381,1407]" italics="true" pageId="14" pageNumber="87">Pd</emphasis>
) with two peaks.
</paragraph>
<paragraph id="014A36BFE106AF07FF5C3DD6FA34FA1D" blockId="14.[151,1437,841,1623]" box="[199,1432,1417,1444]" pageId="14" pageNumber="87">
<emphasis id="3381EAADE106AF07FF5C3DD6FED1FA1D" bold="true" box="[199,381,1417,1443]" pageId="14" pageNumber="87">Size variations.</emphasis>
Females vary from 10.8 to 11.0 in body length, carapace 4.6–4.7 long, 3.35–3.45 wide (n=2).
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E106AF07FF5C3DF2FC26FA76" box="[199,906,1453,1480]" pageId="14" pageNumber="87" type="distribution">
<paragraph id="014A36BFE106AF07FF5C3DF2FC26FA76" blockId="14.[151,1437,841,1623]" box="[199,906,1453,1480]" pageId="14" pageNumber="87">
<emphasis id="3381EAADE106AF07FF5C3DF2FECDFA79" bold="true" box="[199,353,1453,1479]" pageId="14" pageNumber="87">Distribution.</emphasis>
Known only from the
<typeStatus id="DE4E881DE106AF07FDFB3DF1FD3CFA76" box="[608,656,1454,1480]" pageId="14" pageNumber="87">type</typeStatus>
locality (
<figureCitation id="99CE2A3AE106AF07FD613DF2FCD2FA76" box="[762,894,1453,1480]" captionStart="FIGURES 58–60" captionStartId="13.[151,264,1657,1681]" captionTargetBox="[151,1437,669,1634]" captionTargetId="figure-289@13.[151,1437,669,1634]" captionTargetPageId="13" captionText="FIGURES 58–60. Collecting localities of the studied Dysdera species (58–59) and published records of other Central Asian Dysdera species, except for D. crocata (60). Triangle—Dysdera arnoldii, diamond—D. mikhailovi sp. nov., circle—D. dushengi, square—D. subcylindrica. Black symbols refer to the studied material, white symbols refer to literature-derived data. Some symbols could refer to more than one closely situated localities. The frame of Fig. 59 refers to the content of Fig. 58. The numbers in Fig. 60 refer to the following species: 1—D. aculeata; 2—D. arnoldii; 3—D. brignolii; 4—D. dushengi; 5—D. fedtschenkoi Dunin, 1992; 6—D. ferghanica Dunin, 1985; 7—D. kronebergi Dunin, 1992; 8—D. kugitangica; 9—D. kusnetsovi; 10—D. limitanea; 11—D. nenilini; 12—D. pamirica, 13—D. pococki; 14—D. subcylindrica; 15—D. tartarica; 16—D. tystshenkoi Dunin, 1989; 17—D. zarudnyi; 18—D. zonsteini Dimitrov, 2021; 19—D. mikhailovi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5157042" httpUri="https://zenodo.org/record/5157042/files/figure.png" pageId="14" pageNumber="87">Figs 58–59</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="49EF6534E106AF07FF5C3D8EFF66F9E9" pageId="14" pageNumber="87" type="discussion">
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<emphasis id="3381EAADE106AF07FF5C3D8EFEE3FA55" bold="true" box="[199,335,1489,1515]" pageId="14" pageNumber="87">Comments.</emphasis>
Due to big differences in body sizes, there are doubts whether the
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male is correctly matched with the
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females. The carapace length in the female is 1.5 times longer than that in the male. Nonetheless, they were collected together and therefore were provisionally accepted as belonging to the same spe- cies.
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<emphasis id="3381EAADE109AF08FF0C3B98FF56FC5C" bold="true" box="[151,250,967,994]" italics="true" pageId="1" pageNumber="74">Dysdera</emphasis>
Latreille, 1804
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species:
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<emphasis id="3381EAADE109AF08FEAE3C4FFDAFFB94" box="[309,515,1040,1066]" italics="true" pageId="1" pageNumber="74">Dysdera erythrina</emphasis>
(Walckenaer, 1802)
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<taxonomicName id="E90FE0BCDB12FC8ADAFD034A22C3ECDD" ID-CoL="3RRSR" authority="Kühner" authorityName="Kühner" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" higherTaxonomySource="CoL" kingdom="Fungi" order="Russulales" pageId="0" pageNumber="102" phylum="Basidiomycota" rank="species" species="dryadophilus">Lactarius dryadophilus Kühner</taxonomicName>
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<taxonomicName id="9FC6EB0ED6492D816F921AB0D9D73B40" authority="Kühner" authorityName="Kühner" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" higherTaxonomySource="CoL" kingdom="Fungi" order="Russulales" pageId="0" pageNumber="102" phylum="Basidiomycota" rank="species" species="dryadophilus">Lactarius dryadophilus Kühner</taxonomicName>
, 1975a: 68. Holotype: Norway, Hardanger (LY). SYNONYM:
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Lactarius dryadophilus var. saliceticola
<authority id="ADE9ACA2E9831849C34F581715EB466F" pageId="0" pageNumber="102">Bon</authority>
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and Jamoni in Jamoni and Bon, 1992b: 21.
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A medium-sized to large, lilac staining
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with cream-coloured, viscid cap with a bearded margin; growing in arctic and alpine areas.
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<subSubSection id="B8FEB3AB7E9B5007D4379C3426116E6F" pageId="0" pageNumber="102" type="description">
<paragraph id="ABA09ABF63FBD609E4FFE497468A9579" box="[17,1490,594,1140]" pageId="0" pageNumber="102" type="mainText">DESCRIPTION: Cap 30-100(-150) mm, at first convex with a slightly depressed centre and decurved or inrolled margin, sometimes slightly umbonate; surface sticky, viscid, at the margin bearded and tomentose with up to 2 mm long hairs, whitish chrome to pale cream or warm buff but more brownish in the centre, azonate or slightly zonate. Gills adnate, medium broad, crowded, often forked, whitish to pale cream, later pinkish buff. Stem 20-30 x 15-20 mm, typically curved and tapering downwards; surface whitish to pale cream, sometimes yellowish at the base, very finely pitted or with ochraceous spots, especially towards the base. Flesh firm, becoming hollow in the stem, white, changing to lilac; smell fruity, sweetish; taste mild, like cedar wood to very slightly acrid. Milk white, changing to lilac in contact with the flesh. Spore deposit cream.</paragraph>
<paragraph id="0D6C52456121EE5E3A3FB40BEB718A7D" box="[9,1483,1142,1668]" pageId="0" pageNumber="102" type="mainText">Spores 9.3-11.8 x 7.2-9.2 um, av. 10.2-10.3 x 8.1-8.4 um, broadly ellipsoid to ellipsoid, Q = 1.15-1.40, av. 1.22-1.27; ornamentation up to 0.3 um high, of rather fine and narrow ridges, often aligned but not forming a reticulum or at most a very incomplete one; isolated warts often elongate; plage inamyloid. Basidia 60-70 x 12-15 um, subclavate, 4-spored. Pleuromacrocystidia moderately abundant, 80-120 x 10-15 um, fusiform, tapering to a mucronate apex. Gill edge sterile; cheilomacrocystidia 40-60 x 7-9 um, fusiform; paracystidia 10-25 x 4-6(-10) um; cylindric to tortuous or clavate. Pileipellis an ixocutis to an ixotrichoderm, 150-200 um thick; hyphae 1-5 um broad, very thin-walled and shrivelled, gelatinized, repent or ascending, some with conspicuous incrustations.</paragraph>
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<subSubSection id="20B3261DA6387CE52C9C2B4374DE9F1B" pageId="0" pageNumber="102" type="distribution">
<paragraph id="6AB4BC9432C076FB95393ED0C9A38766" box="[5,1474,1745,1913]" pageId="0" pageNumber="102" type="mainText">ECOLOGY AND DISTRIBUTION: Found in arctic and alpine vegetation types on rich, calcareous soils, often in Dryas-rich grasslands. Rather rare and known from Fennoscandia, the Alps, the Pyrenees and Greenland, and occuring from August to mid September.</paragraph>
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DISCUSSION: The cap is tomentose only to a short distance from the margin, unlike in
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, which also has more yellow colours, a longer stem and a different spore ornamentation.
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<paragraph id="33386844F3DCB59DA9FD52789F5E0260" box="[0,1469,2130,2301]" pageId="0" pageNumber="102" type="mainText">
The original description of
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was partly based on this species, but recently lectotypified as a synonym of
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(Knudsen nda Lamoure, 1993).
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