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<document id="8CA584769C4FDE0B8408971AD610511F" ID-DOI="10.11646/zootaxa.4743.2.14" ID-GBIF-Dataset="5006e921-fa87-45cb-9f17-b76b3e0ce7ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="3687752" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582709296776" checkinUser="plazi" docAuthor="Ponel, Philippe, Cascio, Pietro Lo &amp; Soldati, Fabien" docDate="2020" docId="CD4B034CA154677484898DF1391DFC45" docLanguage="en" docName="zootaxa.4743.2.14.pdf" docOrigin="Zootaxa 4743 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catomus aeolicus Ponel &amp; Cascio &amp; Soldati 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="298" masterDocId="31727B34A1556777841E8C2D3B5CFFC4" masterDocTitle="A new species of Catomus Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from the Aeolian archipelago (Sicily, Italy)" masterLastPageNumber="300" masterPageNumber="295" pageNumber="296" updateTime="1698721555727" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2A7F54A34EF91FA83D76E5AD1E41F473">A new species of Catomus Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from the Aeolian archipelago (Sicily, Italy)</mods:title>
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<mods:namePart id="3029CBE3AD58144BE81B4DDEA181CCB6">Ponel, Philippe</mods:namePart>
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<mods:namePart id="948B6655BDFF2764EE979A6260B5B09C">Cascio, Pietro Lo</mods:namePart>
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<mods:namePart id="DC4586231E18FD218474CC12CAC85C87">Soldati, Fabien</mods:namePart>
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<treatment id="CD4B034CA154677484898DF1391DFC45" ID-DOI="http://doi.org/10.5281/zenodo.3687916" ID-GBIF-Taxon="162267556" ID-Zenodo-Dep="3687916" LSID="urn:lsid:plazi:treatment:CD4B034CA154677484898DF1391DFC45" httpUri="http://treatment.plazi.org/id/CD4B034CA154677484898DF1391DFC45" lastPageId="3" lastPageNumber="298" pageId="1" pageNumber="296">
<subSubSection id="0DF8E1D1A154677684898DF139ECFE32" box="[151,688,476,503]" pageId="1" pageNumber="296" type="nomenclature">
<paragraph id="455DB25AA154677684898DF139ECFE32" blockId="1.[151,1437,440,2014]" box="[151,688,476,503]" pageId="1" pageNumber="296">
<heading id="1E150536A154677684898DF139ECFE32" bold="true" box="[151,688,476,503]" fontSize="11" level="1" pageId="1" pageNumber="296" reason="1">
<emphasis id="77966E48A154677684898DF139ECFE32" bold="true" box="[151,688,476,503]" pageId="1" pageNumber="296">
<taxonomicName id="82E2C9D9A154677684898DF13A31FE33" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[151,365,476,503]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A154677684898DF13A31FE33" bold="true" box="[151,365,476,503]" italics="true" pageId="1" pageNumber="296">Catomus aeolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CA5D333A1546776856B8DF13A8EFE32" box="[373,466,476,502]" pageId="1" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="DDD9AEDFA154677685FC8DF139FBFE32" box="[482,679,476,502]" captionStart="FIGURE 1" captionStartId="2.[151,244,1858,1880]" captionTargetBox="[160,1427,250,1824]" captionTargetId="figure@2.[151,1436,237,1832]" captionTargetPageId="2" captionText="FIGURE 1. Catomus aeolicus sp. nov. from Panarea,Aeolian archipelago, Sicily.A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by Catomus aeolicus sp. nov. (from Wikimedia, modified). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="1" pageNumber="296">Figs 1A, 1B, 1C</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DF8E1D1A154677684898E2F3F2FF88A" pageId="1" pageNumber="296" type="description">
<paragraph id="455DB25AA154677684898E2F381BFD42" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">
<emphasis id="77966E48A154677684898E2F3A7EFDDE" bold="true" box="[151,290,514,538]" pageId="1" pageNumber="296">Description.</emphasis>
Male (
<figureCitation id="DDD9AEDFA154677685708E2F3AE1FDDE" box="[366,445,514,538]" captionStart="FIGURE 1" captionStartId="2.[151,244,1858,1880]" captionTargetBox="[160,1427,250,1824]" captionTargetId="figure@2.[151,1436,237,1832]" captionTargetPageId="2" captionText="FIGURE 1. Catomus aeolicus sp. nov. from Panarea,Aeolian archipelago, Sicily.A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by Catomus aeolicus sp. nov. (from Wikimedia, modified). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="1" pageNumber="296">Fig 1A</figureCitation>
). Body length from epistome to elytral apex 4.0–
<quantity id="821A1FBFA154677687D18E2F3F7FFDDD" box="[975,1059,514,538]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="296" unit="mm" value="7.3">7.3 mm</quantity>
. Entirely black brown, without any metallic shine, antennae and legs paler; entirely glabrous on dorsal side, ventrites covered with very short, scattered, whitish setae, antennae and legs with dense, whitish pubescence; head and pronotum shiny although densely punctuate, elytra dull and alutaceous; body elongated, pterothorax oval; wingless.
</paragraph>
<paragraph id="455DB25AA154677684D98EBF39DFFC0E" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">Head. Strongly and densely punctuate, punctures as large as or larger than areas between them, elongated, sometimes almost confluent and forming short striae, subrounded on head sides and behind vertex, intervals with fine microreticulation visible at strong magnification (x50); epistome straight, truncate, separated from vertex by a transverse depression, eyes prominent, appearing rounded when viewed from above, oval in lateral view, not notched by epistome or by antennal insertion; without short vertical groove from bottom edge of the eyes; last maxillary palpomere strongly securiform; labial palpi with last palpomere enlarged but not securiform; antennae long, exceeding posteriad the middle of elytra by at least one antennomere, first antennomere strongly clavate, second transverse, third to seventh elongated, especially the third which is three times longer than broad, slightly enlarged from base to apex, eighth to eleventh distinctly flattened, enlarged from base to apex, last one reniform.</paragraph>
<paragraph id="455DB25AA154677684D98FFB39A1FBBA" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">Pronotum. 1.2 times broader than long, anterior angles rounded but marked, posterior angles completely obsolete, totally rounded, pronotum widest prior middle, puncturation coarse and dense on disc (similar to that of the head), less dense and coarse laterally; punctures elongated, sometimes almost confluent and forming short striae, space between them with fine microreticulation visible at higher magnification (x50), similar to that on head; pronotum entirely and finely rimmed, the middle third of front margin excepted.</paragraph>
<paragraph id="455DB25AA154677684D988A73A17FAF6" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">Elytra. Oval, looking more dull and alutaceous than head and pronotum due to a finer and denser microreticulation; humeri entirely rounded, humeral callus absent; all striae narrow and deep, deeper at base, punctuate, punctures slightly broader than striae, slightly notching interstriae; the latter regularly convex from base to apex; cuticle on interstriae surface apparently smooth but, at higher magnification (x50), covered with sparse superficial punctures; scutellum broad, dull and smooth.</paragraph>
<paragraph id="455DB25AA154677684D989133AEAFA59" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">Ventral side. Prohypomera dull, impunctuate, with longitudinal wrinkles; head, prosternum, mesoventrite and metaventrite punctuate, punctures stronger and denser on mesoventrite; all ventrites finely punctuate, each puncture bearing a fine and short whitish seta.</paragraph>
<paragraph id="455DB25AA154677684D9898739F0F996" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">Legs. All tibiae bent, mesotibiae stronger so; male protarsi strongly widened, first to third tarsomeres, taken together, two times longer than broad and much wider than apex of protibiae; mesotarsi less widened, first to third tarsomeres, taken together, almost three times longer than broad; metatarsi simple; all tarsomeres with sparse whitish setae on upper surface, first to fourth tarsomeres of protarsi and mesotarsi covered with a dense whitish pubescence on inferior surface, tarsomeres of metatarsi simple; onychium simple.</paragraph>
<paragraph id="455DB25AA154677684D98A733AEFF979" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">
Aedeagus (
<figureCitation id="DDD9AEDFA1546776855E8A733AD3F9B2" box="[320,399,1630,1654]" captionStart="FIGURE 1" captionStartId="2.[151,244,1858,1880]" captionTargetBox="[160,1427,250,1824]" captionTargetId="figure@2.[151,1436,237,1832]" captionTargetPageId="2" captionText="FIGURE 1. Catomus aeolicus sp. nov. from Panarea,Aeolian archipelago, Sicily.A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by Catomus aeolicus sp. nov. (from Wikimedia, modified). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="1" pageNumber="296">Fig 1C</figureCitation>
). Parameres four times shorter than phallobase, tapering from base to apex, acuminate, without constriction between phallobase and parameres; inner margin of parameres very slightly sinuate, phallus very parallel with simple rounded apex.
</paragraph>
<paragraph id="455DB25AA154677684D98AE73F2FF88A" blockId="1.[151,1437,440,2014]" pageId="1" pageNumber="296">
Female (
<figureCitation id="DDD9AEDFA1546776853B8AE73A2FF926" box="[293,371,1738,1762]" captionStart="FIGURE 1" captionStartId="2.[151,244,1858,1880]" captionTargetBox="[160,1427,250,1824]" captionTargetId="figure@2.[151,1436,237,1832]" captionTargetPageId="2" captionText="FIGURE 1. Catomus aeolicus sp. nov. from Panarea,Aeolian archipelago, Sicily.A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by Catomus aeolicus sp. nov. (from Wikimedia, modified). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="1" pageNumber="296">Fig 1B</figureCitation>
). Body length from epistome to elytral apex
<quantity id="821A1FBFA154677687418AE738B8F925" box="[863,996,1738,1762]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.2" metricValueMin="4.8" pageId="1" pageNumber="296" unit="mm" value="6.5" valueMax="8.2" valueMin="4.8">4.8–8.2 mm</quantity>
. Pronotum less shiny, more strongly microreticulate, punctures finer, punctation sparser; antennae shorter, hardly reaching the anterior fifth of elytra, eighth to eleventh antennomeres indistinctly flattened, slightly enlarged from base to apex; all tibiae bent as in males; pro-, meso- and metatarsi simple, not widened, lower surface without any thick whitish pubescence sole.
</paragraph>
</subSubSection>
<subSubSection id="0DF8E1D1A154677484D98B773E35FE85" lastPageId="3" lastPageNumber="298" pageId="1" pageNumber="296" type="diagnosis">
<paragraph id="455DB25AA154677584D98B773ABCFF12" blockId="1.[151,1437,440,2014]" lastBlockId="2.[151,1436,154,214]" lastPageId="2" lastPageNumber="297" pageId="1" pageNumber="296">
<emphasis id="77966E48A154677684D98B773A60F8B6" bold="true" box="[199,316,1882,1906]" pageId="1" pageNumber="296">Diagnosis.</emphasis>
<taxonomicName id="82E2C9D9A154677685588B773954F8B6" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[326,520,1882,1906]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A154677685588B773954F8B6" box="[326,520,1882,1906]" italics="true" pageId="1" pageNumber="296">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A1546776860D8B773930F8B6" bold="true" box="[531,620,1882,1906]" pageId="1" pageNumber="296">
<taxonomicNameLabel id="6CA5D333A1546776860D8B773930F8B6" box="[531,620,1882,1906]" pageId="1" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily separated from the other European
<taxonomicName id="82E2C9D9A154677680838B773FA0F8B5" authorityName="Allard" authorityYear="1876" box="[1181,1276,1882,1905]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="77966E48A154677680838B773FA0F8B5" box="[1181,1276,1882,1905]" italics="true" pageId="1" pageNumber="296">Catomus</emphasis>
</taxonomicName>
species by its strongly bent mesotibiae, unusually widened male protarsi, long male antennae and shape of its aedeagus (
<figureCitation id="DDD9AEDFA1546776810A8B533E34F852" box="[1300,1384,1918,1942]" captionStart="FIGURE 1" captionStartId="2.[151,244,1858,1880]" captionTargetBox="[160,1427,250,1824]" captionTargetId="figure@2.[151,1436,237,1832]" captionTargetPageId="2" captionText="FIGURE 1. Catomus aeolicus sp. nov. from Panarea,Aeolian archipelago, Sicily.A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by Catomus aeolicus sp. nov. (from Wikimedia, modified). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="1" pageNumber="296">Fig. 1C</figureCitation>
). By its long antennae this species most closely resembles
<taxonomicName id="82E2C9D9A154677686C88B8F38B3F87E" authority="(Vauloger, 1900)" baseAuthorityName="Vauloger" baseAuthorityYear="1900" box="[726,1007,1954,1978]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="77966E48A154677686C88B8F386FF87E" box="[726,819,1954,1978]" italics="true" pageId="1" pageNumber="296">C. fallax</emphasis>
(
<bibRefCitation id="2173CFABA1546776875F8B8F38BBF87E" author="Vauloger, M. de" box="[833,999,1954,1978]" pageId="1" pageNumber="296" pagination="669 - 722" refId="ref4484" refString="Vauloger, M. de (1900) Contribution au Catalogue des Coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe entomologique de France, 68, 669 - 722. [1899]" type="journal article" year="1900">Vauloger, 1900</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="3DF5F2CAA1546776802E8B8F3F23F87E" box="[1072,1151,1954,1978]" name="Tunisia" pageId="1" pageNumber="296">Tunisia</collectingCountry>
and
<collectingCountry id="3DF5F2CAA154677680AB8B8F3FAAF87E" box="[1205,1270,1954,1978]" name="Libya" pageId="1" pageNumber="296">Libya</collectingCountry>
, which belongs to the same group. But the latter differs by having less punctuate pronotum, smooth and hardly punctuate striae on elytra, more plane interstriae in both sexes, prohypomera with scarce and large punctures. Furthermore, male protarsi are not so enlarged as in the new species.
</paragraph>
<caption id="119DE2D2A157677584898B6F3A18F800" ID-DOI="http://doi.org/10.5281/zenodo.3687754" ID-Zenodo-Dep="3687754" httpUri="https://zenodo.org/record/3687754/files/figure.png" pageId="2" pageNumber="297" startId="2.[151,244,1858,1880]" targetBox="[160,1427,250,1824]" targetPageId="2">
<paragraph id="455DB25AA157677584898B6F3A18F800" blockId="2.[151,1437,1858,1988]" pageId="2" pageNumber="297">
<emphasis id="77966E48A157677584898B6F3A51F893" bold="true" box="[151,269,1858,1880]" pageId="2" pageNumber="297">FIGURE 1.</emphasis>
<taxonomicName id="82E2C9D9A1576775850C8B6E3AE1F893" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[274,445,1858,1880]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A1576775850C8B6E3AE1F893" box="[274,445,1858,1880]" italics="true" pageId="2" pageNumber="297">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A157677585DD8B6E3953F89C" bold="true" box="[451,527,1859,1880]" pageId="2" pageNumber="297">
<taxonomicNameLabel id="6CA5D333A157677585DD8B6E3953F89C" box="[451,527,1859,1880]" pageId="2" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Panarea, Aeolian archipelago, Sicily. A, habitus, male holotype, length: 5.3 mm. B, habitus, female allotype, length: 5.5 mm. C, aedeagus in lateral view (left) and ventral view from below (middle), right: phallus separated from the tegmen. D, the Aeolian archipelago, in black: islands occupied by
<taxonomicName id="82E2C9D9A157677587648BA63F7BF85B" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[890,1063,1930,1952]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A157677587648BA63F7BF85B" box="[890,1063,1930,1952]" italics="true" pageId="2" pageNumber="297">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A157677580308BA63F20F864" bold="true" box="[1070,1148,1931,1952]" pageId="2" pageNumber="297">
<taxonomicNameLabel id="6CA5D333A157677580308BA63F20F864" box="[1070,1148,1931,1952]" pageId="2" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(from Wikimedia, modified). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="455DB25AA156677484D98CB43E35FE85" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<emphasis id="77966E48A156677484D98CB43A14FF75" bold="true" box="[199,328,153,177]" pageId="3" pageNumber="298">Variability.</emphasis>
The new species displays well marked sexual dimorphism in morphology of antennae and legs. The size of specimens is extremely variable: in the studied material the smaller
<specimenCount id="53E479D3A156677487A38C93388DFF11" box="[957,977,190,213]" pageId="3" pageNumber="298" type="male">♂</specimenCount>
is only 4.0 mm long, whereas the larger
<specimenCount id="53E479D3A156677481968C933EC0FF11" box="[1416,1436,190,213]" pageId="3" pageNumber="298" type="female">♀</specimenCount>
reaches
<quantity id="821A1FBFA156677484F38CCC3A1EFF3D" box="[237,322,225,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="298" unit="mm" value="8.2">8.2 mm</quantity>
. The color ranges from very dark to reddish, the latter probably corresponding to teneral specimens. Furthermore, the average size in the specimens from Stromboli (
<specimenCount id="53E479D3A1566774873E8D283801FED9" box="[800,861,261,285]" pageId="3" pageNumber="298" type="male">9 ♂♂</specimenCount>
: m =
<quantity id="821A1FBFA156677487BF8D283F5FFED9" box="[929,1027,261,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.03" pageId="3" pageNumber="298" unit="mm" value="6.03">6.03 mm</quantity>
, s.d. = 0.95;
<specimenCount id="53E479D3A156677480958D283F9AFED9" box="[1163,1222,261,285]" pageId="3" pageNumber="298" type="female">6 ♀♀</specimenCount>
: m =
<quantity id="821A1FBFA156677481158D283E30FED9" box="[1291,1388,261,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.21" pageId="3" pageNumber="298" unit="mm" value="7.21">7.21 mm</quantity>
, s.d. = 0.72) is larger than in those from Panarea (
<specimenCount id="53E479D3A1566774866A8D0439E3FE85" box="[628,703,297,321]" pageId="3" pageNumber="298" type="male">28 ♂♂</specimenCount>
: m =
<quantity id="821A1FBFA1566774871B8D04383AFE85" box="[773,870,297,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="3" pageNumber="298" unit="mm" value="5.05">5.05 mm</quantity>
, s.d. = 0.69;
<specimenCount id="53E479D3A156677487F18D043F6BFE85" box="[1007,1079,297,321]" pageId="3" pageNumber="298" type="female">11 ♀♀</specimenCount>
: m =
<quantity id="821A1FBFA156677480638D043F82FE85" box="[1149,1246,297,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="3" pageNumber="298" unit="mm" value="6.05">6.05 mm</quantity>
, s.d. = 1.15).
</paragraph>
</subSubSection>
<subSubSection id="0DF8E1D1A156677484D98D603864FD09" pageId="3" pageNumber="298" type="materials_examined">
<paragraph id="455DB25AA156677484D98D603A19FE69" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<emphasis id="77966E48A156677484D98D603A32FEA1" bold="true" box="[199,366,333,357]" pageId="3" pageNumber="298">Type material.</emphasis>
<materialsCitation id="F58AB807A1566774856B8D603A19FE69" ID-GBIF-Occurrence="2571176118" collectingDate="2015-11-07" collectorName="P. Ponel &amp; P. Lo Cascio &amp; F. Soldati" pageId="3" pageNumber="298" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="9A590CF8A1566774856B8D603A84FEA1" box="[373,472,333,357]" pageId="3" pageNumber="298" type="holotype">Holotype</typeStatus>
<specimenCount id="53E479D3A156677485FE8D633AAAFEA1" box="[480,502,334,357]" pageId="3" pageNumber="298" type="male">♂</specimenCount>
, labelled: “SICILE-EOLIENNES, île Panarea,
<date id="315C949AA156677487EA8D603F3BFEA1" box="[1012,1127,333,357]" pageId="3" pageNumber="298" value="2015-11-07">
<collectingDate id="21186D72A156677487EA8D603F3BFEA1" box="[1012,1127,333,357]" pageId="3" pageNumber="298" value="2015-11-07">7-XI-2015</collectingDate>
</date>
,
<collectorName id="E817D78CA1566774806D8D633F96FEA1" box="[1139,1226,333,357]" pageId="3" pageNumber="298">P. Ponel</collectorName>
leg.” (white label); “
<typeStatus id="9A590CF8A156677484BF8D5C3A5BFE4D" box="[161,263,369,393]" pageId="3" pageNumber="298" type="holotype">Holotype</typeStatus>
<specimenCount id="53E479D3A156677485108D5F3A78FE4D" box="[270,292,370,393]" pageId="3" pageNumber="298" type="male">♂</specimenCount>
,
<taxonomicName id="82E2C9D9A1566774852F8D5F3AACFE4D" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[305,496,369,393]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A1566774852F8D5F3AACFE4D" box="[305,496,369,393]" italics="true" pageId="3" pageNumber="298">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A156677485E68D5C3913FE4D" bold="true" box="[504,591,369,393]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="6CA5D333A156677485E68D5C3913FE4D" box="[504,591,369,393]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<collectorName id="E817D78CA156677486438D5F39EAFE4D" box="[605,694,369,393]" pageId="3" pageNumber="298">P. Ponel</collectorName>
,
<collectorName id="E817D78CA156677486DC8D5F3817FE4D" box="[706,843,369,393]" pageId="3" pageNumber="298">P. Lo Cascio</collectorName>
&amp;
<collectorName id="E817D78CA156677487718D5F3884FE4D" box="[879,984,369,393]" pageId="3" pageNumber="298">F. Soldati</collectorName>
” (red label), Muséum National d’Histoire Naturelle, Paris.
</materialsCitation>
</paragraph>
<paragraph id="455DB25AA156677484D98D943ED9FE31" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<materialsCitation id="F58AB807A156677484D98D943ED9FE31" ID-GBIF-Occurrence="2571176117" collectingDate="2015-11-07" collectorName="P. Ponel &amp; P. Lo Cascio &amp; F. Soldati" pageId="3" pageNumber="298" specimenCount="2" specimenCount-female="2" typeStatus="allotype">
<typeStatus id="9A590CF8A156677484D98D943A78FE15" box="[199,292,441,465]" pageId="3" pageNumber="298" type="allotype">Allotype</typeStatus>
<specimenCount id="53E479D3A156677485358D973A63FE15" box="[299,319,442,465]" pageId="3" pageNumber="298" type="female">♀</specimenCount>
, labelled: “SICILE-EOLIENNES, île Panarea,
<date id="315C949AA156677487278D9438F0FE15" box="[825,940,441,465]" pageId="3" pageNumber="298" value="2015-11-07">
<collectingDate id="21186D72A156677487278D9438F0FE15" box="[825,940,441,465]" pageId="3" pageNumber="298" value="2015-11-07">7-XI-2015</collectingDate>
</date>
,
<collectorName id="E817D78CA156677487A98D973F52FE15" box="[951,1038,441,465]" pageId="3" pageNumber="298">P. Ponel</collectorName>
leg.” (white label); “
<typeStatus id="9A590CF8A156677480F38D943E11FE15" box="[1261,1357,441,465]" pageId="3" pageNumber="298" type="allotype">Allotype</typeStatus>
<specimenCount id="53E479D3A1566774814D8D973E3BFE15" box="[1363,1383,442,465]" pageId="3" pageNumber="298" type="female">♀</specimenCount>
,
<taxonomicName id="82E2C9D9A1566774816A8D973A6BFE31" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A1566774816A8D973A6BFE31" italics="true" pageId="3" pageNumber="298">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A156677485238DF03ACFFE31" bold="true" box="[317,403,477,501]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="6CA5D333A156677485238DF03ACFFE31" box="[317,403,477,501]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<collectorName id="E817D78CA156677485BE8DF33AA4FE31" box="[416,504,477,501]" pageId="3" pageNumber="298">P. Ponel</collectorName>
,
<collectorName id="E817D78CA1566774861D8DF339D5FE31" box="[515,649,477,501]" pageId="3" pageNumber="298">P. Lo Cascio</collectorName>
&amp;
<collectorName id="E817D78CA156677486B58DF3384FFE31" box="[683,787,477,501]" pageId="3" pageNumber="298">F. Soldati</collectorName>
” (red label), Muséum National d’Histoire Naturelle, Paris.
</materialsCitation>
</paragraph>
<paragraph id="455DB25AA156677484D98E2F3864FD09" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<materialsCitation id="F58AB807A156677484D98E2F39AFFDDD" ID-GBIF-Occurrence="2571176122" box="[199,755,513,538]" collectingDate="2015-11-07" collectorName="P. Ponel &amp; P. Lo Cascio &amp; F. Soldati" pageId="3" pageNumber="298" specimenCount="9" specimenCount-female="3" specimenCount-male="6" typeStatus="paratype">
<typeStatus id="9A590CF8A156677484D98E2F3A6FFDDE" box="[199,307,514,538]" pageId="3" pageNumber="298" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="53E479D3A156677485208E2C3A27FDDD" box="[318,379,513,537]" pageId="3" pageNumber="298" type="male">6 ♂♂</specimenCount>
and
<specimenCount id="53E479D3A156677485AF8E2C3AB3FDDD" box="[433,495,513,537]" pageId="3" pageNumber="298" type="female">3 ♀♀</specimenCount>
, same data as Holotype
</materialsCitation>
;
<materialsCitation id="F58AB807A156677486E18E2C3A31FDF9" ID-GBIF-Occurrence="2571176120" collectingDate="2015-11-08" collectorName="P. Ponel" pageId="3" pageNumber="298" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="53E479D3A156677486E18E2C3860FDDD" box="[767,828,513,537]" pageId="3" pageNumber="298" type="male">3 ♂♂</specimenCount>
and
<specimenCount id="53E479D3A1566774876C8E2C38ECFDDD" box="[882,944,513,537]" pageId="3" pageNumber="298" type="female">2 ♀♀</specimenCount>
, “SICILE-EOLIENNES, île Stromboli,
<date id="315C949AA1566774817F8E2C3B93FDF9" pageId="3" pageNumber="298" value="2015-11-08">
<collectingDate id="21186D72A1566774817F8E2C3B93FDF9" pageId="3" pageNumber="298" value="2015-11-08">8-XI- 2015</collectingDate>
</date>
,
<collectorName id="E817D78CA156677484C48E0B3A6DFDF9" box="[218,305,549,573]" pageId="3" pageNumber="298">P. Ponel</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="F58AB807A156677485678E083F81FDF9" ID-GBIF-Occurrence="2571176119" box="[377,1245,549,573]" collectingDate="2016-11-07" collectorName="P. Ponel" pageId="3" pageNumber="298" specimenCount="28" specimenCount-female="7" specimenCount-male="21" typeStatus="paratype">
<specimenCount id="53E479D3A156677485678E083A9EFDF9" box="[377,450,549,573]" pageId="3" pageNumber="298" type="male">21 ♂♂</specimenCount>
and
<specimenCount id="53E479D3A156677485E98E0B3968FDF9" box="[503,564,550,573]" pageId="3" pageNumber="298" type="female">7 ♀♀</specimenCount>
, “SICILE-EOLIENNES, île Panarea,
<date id="315C949AA156677487D28E083F63FDF9" box="[972,1087,549,573]" pageId="3" pageNumber="298" value="2016-11-07">
<collectingDate id="21186D72A156677487D28E083F63FDF9" box="[972,1087,549,573]" pageId="3" pageNumber="298" value="2016-11-07">7-XI-2016</collectingDate>
</date>
,
<collectorName id="E817D78CA156677480548E0B3FFDFDF9" box="[1098,1185,549,573]" pageId="3" pageNumber="298">P. Ponel</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="F58AB807A156677480F78E083F7AFD41" ID-GBIF-Occurrence="2571176123" collectingDate="2016-12" collectorName="P. Lo Cascio" location="All" pageId="3" pageNumber="298" specimenCount="12" specimenCount-female="5" specimenCount-male="7" typeStatus="paratype">
<specimenCount id="53E479D3A156677480F78E083E79FDF9" box="[1257,1317,549,573]" pageId="3" pageNumber="298" type="male">6 ♂♂</specimenCount>
and
<specimenCount id="53E479D3A156677481478E083ECBFDF9" box="[1369,1431,549,573]" pageId="3" pageNumber="298" type="female">4 ♀♀</specimenCount>
, “STROMBOLI: Nel Cannestrà, beating
<taxonomicName id="82E2C9D9A156677486568E673850FDA5" box="[584,780,586,609]" class="Magnoliopsida" family="Fabaceae" genus="Spartium" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="298" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="77966E48A156677486568E673850FDA5" box="[584,780,586,609]" italics="true" pageId="3" pageNumber="298">Spartium junceum</emphasis>
</taxonomicName>
,
<date id="315C949AA156677487048E643822FDA5" box="[794,894,585,609]" pageId="3" pageNumber="298" value="2016-12">
<collectingDate id="21186D72A156677487048E643822FDA5" box="[794,894,585,609]" pageId="3" pageNumber="298" value="2016-12">XII-2016</collectingDate>
</date>
leg.
<collectorName id="E817D78CA156677487AD8E673F1CFDA5" box="[947,1088,585,609]" pageId="3" pageNumber="298">P. Lo Cascio</collectorName>
”.
<location id="403DE481A1566774804D8E643F29FDA5" LSID="urn:lsid:plazi:treatment:CD4B034CA154677484898DF1391DFC45:403DE481A1566774804D8E643F29FDA5" box="[1107,1141,585,609]" name="All" pageId="3" pageNumber="298">All</location>
<typeStatus id="9A590CF8A156677480638E673FBFFDA6" box="[1149,1251,586,610]" pageId="3" pageNumber="298" type="paratype">paratypes</typeStatus>
have a red label: “
<typeStatus id="9A590CF8A156677484BF8E403A5DFD41" box="[161,257,621,645]" pageId="3" pageNumber="298" type="paratype">Paratype</typeStatus>
<specimenCount id="53E479D3A156677485168E433A40FD41" box="[264,284,622,645]" pageId="3" pageNumber="298" type="male">♂</specimenCount>
(or
<specimenCount id="53E479D3A156677485548E403A01FD41" box="[330,349,621,645]" pageId="3" pageNumber="298" type="female">♀</specimenCount>
),
<taxonomicName id="82E2C9D9A1566774856B8E433968FD41" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[373,564,621,645]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A1566774856B8E433968FD41" box="[373,564,621,645]" italics="true" pageId="3" pageNumber="298">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A156677486228E4039CEFD41" bold="true" box="[572,658,621,645]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="6CA5D333A156677486228E4039CEFD41" box="[572,658,621,645]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, P. Ponel, P. Lo Cascio &amp; F. Soldati”
</materialsCitation>
.
<materialsCitation id="F58AB807A1566774802F8E433869FD09" ID-GBIF-Occurrence="2571176121" collectorName="P. Ponel &amp; P. Lo Cascio &amp; F. Soldati" location="Universita degli Studi di Firenze" pageId="3" pageNumber="298" specimenCount="1" typeStatus="paratype">
<typeStatus id="9A590CF8A1566774802F8E433FC5FD42" box="[1073,1177,622,646]" pageId="3" pageNumber="298" type="paratype">Paratypes</typeStatus>
are deposited in the author’s collections
<collectorName id="E817D78CA1566774854A8EBF3AF0FD6D" box="[340,428,657,681]" pageId="3" pageNumber="298">P. Ponel</collectorName>
,
<collectorName id="E817D78CA156677485A88EBF3967FD6D" box="[438,571,657,681]" pageId="3" pageNumber="298">P. Lo Cascio</collectorName>
and
<collectorName id="E817D78CA156677486738EBF3988FD6D" box="[621,724,657,681]" pageId="3" pageNumber="298">F. Soldati</collectorName>
, as well as in the Piero Leo collection (Cagliari) and the Zoological Museum of the
<location id="403DE481A156677485218E9839C7FD09" LSID="urn:lsid:plazi:treatment:CD4B034CA154677484898DF1391DFC45:403DE481A156677485218E9839C7FD09" box="[319,667,693,717]" name="Universita degli Studi di Firenze" pageId="3" pageNumber="298">Università degli Studi di Firenze</location>
“La Specola”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF8E1D1A156677484D98EF4397DFCD1" pageId="3" pageNumber="298" type="etymology">
<paragraph id="455DB25AA156677484D98EF4397DFCD1" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<emphasis id="77966E48A156677484D98EF43A14FD35" bold="true" box="[199,328,729,753]" pageId="3" pageNumber="298">Etymology.</emphasis>
The name
<taxonomicName id="82E2C9D9A156677485DA8EF43941FD35" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[452,541,729,753]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="aeolicus">
<emphasis id="77966E48A156677485DA8EF43941FD35" box="[452,541,729,753]" italics="true" pageId="3" pageNumber="298">aeolicus</emphasis>
</taxonomicName>
is an adjective derived from this group of islands, named after Aeolus (in ancient Greek Αιολός), the god of the winds.
</paragraph>
</subSubSection>
<subSubSection id="0DF8E1D1A156677484D98F0C391DFC45" pageId="3" pageNumber="298" type="biology_ecology">
<paragraph id="455DB25AA156677484D98F0C391DFC45" blockId="3.[151,1437,153,897]" pageId="3" pageNumber="298">
<emphasis id="77966E48A156677484D98F0C3A7BFCFD" bold="true" box="[199,295,801,825]" pageId="3" pageNumber="298">Ecology.</emphasis>
All the specimens of
<taxonomicName id="82E2C9D9A156677486138F0F3996FCFD" authority="Ponel &amp; Cascio &amp; Soldati, 2020" authorityName="Ponel &amp; Cascio &amp; Soldati" authorityYear="2020" box="[525,714,801,825]" class="Insecta" family="Tenebrionidae" genus="Catomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="aeolicus" status="sp. nov.">
<emphasis id="77966E48A156677486138F0F3996FCFD" box="[525,714,801,825]" italics="true" pageId="3" pageNumber="298">Catomus aeolicus</emphasis>
</taxonomicName>
<emphasis id="77966E48A156677486CE8F0C3879FCFD" bold="true" box="[720,805,801,825]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="6CA5D333A156677486CE8F0C3879FCFD" box="[720,805,801,825]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected by beating old woody shrubs with dehiscent bark, especially the Spanish broom
<taxonomicName id="82E2C9D9A1566774860B8F6B3984FC99" box="[533,728,838,861]" class="Magnoliopsida" family="Fabaceae" genus="Spartium" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="298" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis id="77966E48A1566774860B8F6B3984FC99" box="[533,728,838,861]" italics="true" pageId="3" pageNumber="298">Spartium junceum</emphasis>
</taxonomicName>
L. and other
<taxonomicName id="82E2C9D9A156677487768F683892FC99" box="[872,974,837,861]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="298" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
, but the species has been found also on old branches of lentisk
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<emphasis id="77966E48A156677485798F47397EFC45" box="[359,546,873,897]" italics="true" pageId="3" pageNumber="298">Pistacia lentiscus</emphasis>
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L.
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132
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Lamotte
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(
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<emphasis id="02F939338F172087772B79988427A990" italics="true" pageId="null" pageNumber="580">A. selengensis</emphasis>
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auct.)
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Ostasiatischer
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Unterscheidet sich von
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<emphasis id="0B3C0A6DE24364545F44E0AAD768A945" italics="true" pageId="null" pageNumber="580">A. vulgaris</emphasis>
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(Nr. 1) durch folgende Merkmale:
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Mit langen
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;
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30-200 cm hoch;
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mit aromatischem Geruch;
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-
<emphasis id="DCFB37355CDEAC8C7CD75DA627476F93" italics="true" pageId="null" pageNumber="580">12mal so lang wie breit</emphasis>
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Herbst.
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<emphasis id="2F88D80D5A0FFE206E1F41525B6CB1D1" italics="true" pageId="null" pageNumber="580">Zytologische Angaben. 2n</emphasis>
=
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Material aus Bologna, Pollenmeiose normal (Vignoli 1945a), von der Insel Sachalin (Sokolovskaya 1960), aus botanischen
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in Europa (Kawatani und Ohno 1964).
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Kollin und montan.
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wie
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<emphasis id="8838A1BBF7A7DA8513631F2B9947A918" italics="true" pageId="null" pageNumber="580">A. vulgaris</emphasis>
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(Nr. 1).
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<emphasis id="0E72B11F47DC497619638ADBA5AE61CC" italics="true" pageId="null" pageNumber="580">Verbreitung.</emphasis>
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<emphasis id="82320744DB2C5BF5D6BDE483D47157DF" italics="true" pageId="null" pageNumber="580">ostasiatische Pflanze</emphasis>
(vgl. Pampani 1929); heute
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ganz Eurasien verschleppt. - Im Gebiet ziemlich verbreitet und oft
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<emphasis id="1D36C830A73C1A02EBEC8CC30B777D48" italics="true" pageId="null" pageNumber="580">Bemerkungen.</emphasis>
Trat im Gebiet erstmals 1920 im Bodenseegebiet auf.
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56
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4.
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. Pl. VI. figs. 6, 7. B.M.
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, Trans. Ent. Soc. Lond. iii. new ser.
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<taxonomicName id="2031E457EE2FC3A5862093B6DAF01763" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138081" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" lsidName-HNS="Atta quadriglumis Haliday" order="Hymenoptera" pageId="151" pageNumber="151" phylum="Arthropoda" rank="species" species="quadriglumis">Atta quadriglumis</taxonomicName>
, Halid. Trans. Linn. Soc. xiv. 328.
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<paragraph id="D4CF6C4D1E196B69EDE8FF7FCA638C2D" pageId="151" pageNumber="151">Hab. Brazil (Para; Santarem; Ega).</paragraph>
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This species is readily distinguished by its having the head, thorax, legs and nodes of the abdomen opake black, and the abdomen pale reddish yellow. Mr. Bates says, &quot;This is the largest species of the genus that I have seen; its columns are narrower than those of the other species, in fact they generally run along in single file, and forage in the same way. I have found them carrying off great numbers of cocoons of a species of
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; I have searched the columns long and repeatedly without finding any individuals with large heads and long mandibles.&quot;
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Length ~ 15 mm, width 1-2 mm, 31-35 chaetigers, prostomium subpentagonal, three pairs of branchiae, two or three spinous notochaetae, one or two composite neurochaetae, short lateral cirri. Body papillae short, with sediment particles. Large ventral papillae absent.</paragraph>
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<paragraph pageId="0" pageNumber="1">Records.</paragraph>
<paragraph pageId="0" pageNumber="1">12 specimens. Suppl. material 1: ops. 31, 33, 46, 54, 87 (AM).</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
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sp. nov. 2. (AM W.52561)
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sp. nov. 3. (AM W.52560)
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sp. nov. 4. (AM W.52828)
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sp. (AM W.52552)
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Gunton &amp; Kupriyanova &amp; Alvestad &amp; Avery &amp; Blake &amp; Biriukova &amp; Böggemann &amp; Borisova &amp; Budaeva &amp; Burghardt &amp; Capa &amp; Georgieva &amp; Glasby &amp; Hsueh &amp; Hutchings &amp; Jimi &amp; Kongsrud &amp; Langeneck &amp; Meißner &amp; Murray &amp; Nikolic &amp; Paxton &amp; Ramos &amp; Schulze &amp; Sobczyk &amp; Watson &amp; Wiklund &amp; Wilson &amp; Zhadan &amp; Zhang" authorityYear="2021" family="Acrocirridae" kingdom="Animalia" lsidName="" order="Phyllodocida" pageId="0" pageNumber="1" rank="family">Acrocirridae</taxonomicName>
gen. sp. 2 (AM W.52833). Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
); 500
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B-F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="CD4BC127516A23585779A3BDE1DC90B1" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Agonum fossiger Dejean 1828" docType="treatment" docVersion="9" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="758" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="757" updateTime="1668154792637" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bousquet, Yves</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bousquety1@yahoo.com</mods:nameIdentifier>
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<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-11-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>245</mods:number>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.245.3416</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.245.3416</mods:identifier>
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<treatment ID-GBIF-Taxon="127869252" LSID="urn:lsid:plazi:treatment:CD4BC127516A23585779A3BDE1DC90B1" httpUri="http://treatment.plazi.org/id/CD4BC127516A23585779A3BDE1DC90B1" lastPageId="757" lastPageNumber="758" pageId="756" pageNumber="757">
<subSubSection pageId="756" pageNumber="757" type="nomenclature">
<paragraph pageId="756" pageNumber="757">
<taxonomicName LSID="CD4BC127-516A-2358-5779-A3BDE1DC90B1" authority="Dejean, 1828" authorityName="Dejean" authorityYear="1828" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agonum fossiger" order="Coleoptera" pageId="756" pageNumber="757" phylum="Arthropoda" rank="species" species="fossiger">Agonum fossiger Dejean, 1828</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="757" lastPageNumber="758" pageId="756" pageNumber="757" type="reference_group">
<paragraph lastPageId="757" lastPageNumber="758" pageId="756" pageNumber="757">
<taxonomicName authorityName="Dejean" authorityYear="1828" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="756" pageNumber="757" phylum="Arthropoda" rank="species" species="fossiger">Agonum fossiger</taxonomicName>
Dejean, 1828: 160. Type locality:
<normalizedToken originalValue="«Californie»">&quot;Californie&quot;</normalizedToken>
(original citation), herein restricted to Redwood Creek, Humboldt County (see Casey 1920: 121, as
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="756" pageNumber="757" phylum="Arthropoda" rank="species" species="tumidulum">
<emphasis italics="true" pageId="756" pageNumber="757">Agonum tumidulum</emphasis>
</taxonomicName>
). Holotype [by monotypy] in MHNP (Lindroth 1955b: 21). Note. Because Dejean (1828: 160) used the spelling
<emphasis italics="true" pageId="756" pageNumber="757">fossiger</emphasis>
in combination with the genus name
<taxonomicName authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="756" pageNumber="757" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="756" pageNumber="757">Agonum</emphasis>
</taxonomicName>
, he obviously regarded the specific name as a noun in apposi
<pageBreakToken pageId="757" pageNumber="758" start="start">tion</pageBreakToken>
, not as an adjective. In such case, the noun need not agree in gender with the generic name with which it is combined (ICZN 1999: Articles 31.2.1 and 34.2.1).
<taxonomicName authorityName="Dejean" authorityYear="1828" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="fossiger">
<emphasis italics="true" pageId="757" pageNumber="758">Agonum fossiger</emphasis>
</taxonomicName>
, not
<taxonomicName authorityName="Dejean" authorityYear="1828" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="fossigerum">
<emphasis italics="true" pageId="757" pageNumber="758">Agonum fossigerum</emphasis>
</taxonomicName>
, is the correct spelling for this species.
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="famelicum">Agonum famelicum</taxonomicName>
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
, 1843: 58. Type locality:
<normalizedToken originalValue="«Californie»">&quot;Californie&quot;</normalizedToken>
(original citation). Lectotype (♀), designated by Lindroth (1966: 596), in ZMH. Synonymy established by LeConte (1863b: 7), confirmed by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Agonothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="robustus">Agonothorax robustus</taxonomicName>
Motschulsky, 1859a: 158. Type locality: California (inferred from title of the paper). Two syntypes in ZMMU (Keleinikova 1976: 214) and one in MCZ (collection LeConte) (Lindroth 1966: 596). Synonymy established by LeConte (1863b: 7), confirmed by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Notman" authorityYear="1919" class="Hexapoda" family="Carabidae" genus="Platynus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="foveiceps">Platynus foveiceps</taxonomicName>
Notman, 1919b: 233. Type locality: &quot;Franktown [Washoe County], Nevada&quot; (original citation). Holotype [by monotypy] (♀) in USNM. Synonymy established by Liebherr (1991a: 121).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="breviusculum">Agonum breviusculum</taxonomicName>
Casey, 1920: 119. Type locality: &quot;Lake Tahoe [Placer County], California&quot; (original citation). Lectotype (♀), designated by Lindroth (1975: 127), in USNM [# 47485]. Synonymy established by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="pertinax">Agonum pertinax</taxonomicName>
Casey, 1920: 119. Type locality: &quot;Reno [Washoe County], Nevada&quot; (original citation). Holotype [by monotypy] (♂) in USNM [# 47479]. Synonymy established by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="atromicans">Agonum atromicans</taxonomicName>
Casey, 1920: 120. Type locality: &quot;probably Colorado&quot; (original citation). Holotype [by monotypy] (♂) in USNM [# 47484]. Synonymy established by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="vegetum">Agonum vegetum</taxonomicName>
Casey, 1920: 121. Type locality: &quot;S[an]ta Cruz [Santa Cruz County], California&quot; (original citation). Lectotype (♀), designated by Lindroth (1975: 127), in USNM [# 47487]. Synonymy established by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="columbicum">Agonum columbicum</taxonomicName>
Casey, 1920: 121. Type locality: &quot;The Dalles [Wasco County], Oregon&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 127), in USNM [# 47488]. Synonymy established by Hatch (1953: 143), confirmed by Lindroth (1966: 596).
</paragraph>
<paragraph pageId="757" pageNumber="758">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="species" species="tumidulum">Agonum tumidulum</taxonomicName>
Casey, 1920: 121. Type locality: &quot;Redwood Creek, Humboldt Co[unty], California&quot; (original citation for the lectotype). Lectotype (♀), designated by Lindroth (1975: 127), in USNM [# 47489]. Synonymy established by Lindroth (1966: 596).
</paragraph>
</subSubSection>
<subSubSection pageId="757" pageNumber="758" type="distribution">
<paragraph pageId="757" pageNumber="758">Distribution.</paragraph>
<paragraph pageId="757" pageNumber="758">
This species is found west of the Rocky Mountains from
<normalizedToken originalValue="“Washington”">&quot;Washington&quot;</normalizedToken>
(Hatch 1953: 143) and western Idaho (Nez Perce County, CNC) to southern California (Fall 1901a: 46). The record from Baja California (Horn 1894: 309) needs confirmation since the species is not listed in
<normalizedToken originalValue="Liebherr’s">Liebherr's</normalizedToken>
(1994) review of the Mexican
<taxonomicName authorityName="Bonelli" authorityYear="1810" class="Hexapoda" family="Carabidae" genus="Agonum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="757" pageNumber="758" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="757" pageNumber="758">Agonum</emphasis>
</taxonomicName>
; those from southwestern British Columbia (Hatch 1953: 143) and Colorado (LeConte 1858a: 28) are probably in error.
</paragraph>
</subSubSection>
<subSubSection pageId="757" pageNumber="758" type="records">
<paragraph pageId="757" pageNumber="758">Records.</paragraph>
<paragraph pageId="757" pageNumber="758">
<emphasis bold="true" pageId="757" pageNumber="758">USA</emphasis>
: CA, ID, NV, OR, WA
</paragraph>
</subSubSection>
</treatment>
</document>