Add updates up until 2024-08-08 19:55:34
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<mods:title id="57C41C09597999B612C9923D89289721">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
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<mods:title id="CDBD5B1C7AD29236CF106BBDCD4A08C8">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
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<mods:namePart id="CF57EAE80319E664ACD56D73D4CC9DDD">Harden, Curt W.</mods:namePart>
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<mods:nameIdentifier id="C8535E91FE508820552C78D7229A0ECD" type="ORCID">0000-0002-5226-8725</mods:nameIdentifier>
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<mods:affiliation id="455D1223431F2C0BBDBAA9C9DF3732E5">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:namePart id="A7434E8F0DAD42231922570A95CD83C5">Harden, Curt W.</mods:namePart>
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<mods:affiliation id="3B553D2E26CFB146243322C99C2F7C51">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:namePart id="9E73AC55AECCED7A20808EC9E5EE20C0">Caterino, Michael S.</mods:namePart>
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<mods:affiliation id="D009418BE06B1269B195A1E4D625AE37">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:namePart id="9803BDD706994BA5578FEAAFD28C06BD">Caterino, Michael S.</mods:namePart>
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<mods:affiliation id="F729B88952D3EC684A93DB33A7FDD938">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:part id="511012667467E843650CF167FFDD4B47">
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<mods:date id="E6F5E6CC3523DCBD56FB4B6839ABFE1B">2024</mods:date>
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<treatment id="01B6AA753C235D27BFF5921091C3AAB0" ID-DOI="http://doi.org/10.5281/zenodo.13270375" ID-Zenodo-Dep="13270375" ID-arpha="01B6AA75-3C23-5D27-BFF5-921091C3AAB0" ID-zoobank="https://zoobank.org/9968711C-1A42-426E-B295-23D834F53A73" LSID="urn:lsid:plazi:treatment:01B6AA753C235D27BFF5921091C3AAB0" LSID-ZBK="urn:lsid:zoobank.org:act:9968711C-1A42-426E-B295-23D834F53A73" httpUri="http://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0">
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<subSubSection id="827A98E20B480E9B743EA59FB2135658" type="nomenclature">
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@ -67,11 +67,11 @@
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<heading id="5AA267858F5C7E0C043A8781EAD8F782" reason="title">Type material.</heading>
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</paragraph>
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||||
<paragraph id="345C7639FAFAA45C5652FE5BE4FD710F">
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||||
<materialsCitation id="A6C21007563A8953974AC6717F43EA41" collectingDate="1989-10-28" collectionCode="NCSU" determinerName="C. W. Harden" specimenCode="ENT 00327997" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||||
<materialsCitation id="69520BB787B2EC19148B85EB6B58365C" collectingDate="1989-10-28" collectionCode="NCSU" determinerName="C. W. Harden" specimenCode="NCSU_ENT 00327997" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis id="094EA62334B7CF4FFDD1A73B457D277D" bold="true">
|
||||
<emphasis id="D55523B649873F21E277E4A33635EFC5" bold="true" italics="true">
|
||||
<typeStatus id="D873294AD4220D9345E8533EB602D0A4" type="holotype">Holotype</typeStatus>
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male
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<specimenCount id="EF4CE01724DD795314ECAD59B073BED2" type="male">male</specimenCount>
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||||
</emphasis>
|
||||
(
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<named-content id="276A1DF3126A95C28CD85F164AB22766" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/north-carolina-state-university-insect-museum" xlink_title="North Carolina State University">
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@ -88,24 +88,23 @@ JF & TADCornell ” “
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<emphasis id="F03D1AAD2EB8996A586D180CF255F624" italics="true">Serranillus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FBEF85DEBEB2821FA47C419E837F9FC4" rank="species">new species</taxonomicNameLabel>
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<specimenCount id="73DF8DB47DD9CD06CAF7F328D13AA774" count="1" type="male">♂</specimenCount>
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det.
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♂ det.
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<determinerName id="8057A93B73186796A7E6E719FABF5560">C. W. Harden</determinerName>
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||||
2021 ” “ [QR Code]
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||||
<collectionCode id="B36A7A59CC6F8C66EB74DCC56ADC2048" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
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_
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||||
<specimenCode id="02C179DDE89F25A7D2D6DCE6A14A3967" collectionCode="NCSU" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">ENT 00327997</specimenCode>
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” “
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<typeStatus id="AC98B6EEBFC6F9C67108157EF494CE28" type="holotype">HOLOTYPE</typeStatus>
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<specimenCode id="933E45355F7D48322967DFB3349AAD61">NCSU_ENT 00327997</specimenCode>
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” “ HOLOTYPE
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<taxonomicName id="D326B980525C8A42D9D729EBEB109394" authority="Harden & Caterino 2024" authorityName="Harden & Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monadnock">
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<originalDetermination id="374AD9FFC949CD066760A5180936DD47">
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<emphasis id="9C9992EA17C9F7958806A3061A34913F" italics="true">Serranillus monadnock</emphasis>
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||||
<bibRefCitation id="5675D45CAD64D4B6BA714882232273BC" DOI="10.1093/isd/ixae009" author="Caterino MS & Harden CW" refId="B15" refString="Caterino MS, Harden CW (2024) A review of Appalachian Dasycerus Brongniart and the recognition of cryptic diversity within Dasycerus carolinensis (Coleoptera: Staphylinidae: Dasycerinae). Insect Systematics and Diversity 8 (2) [ixae 009]: 1 - 22. https: // doi. org / 10.1093 / isd / ixae 009" year="2024">Harden & Caterino 2024</bibRefCitation>
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||||
</originalDetermination>
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||||
</taxonomicName>
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||||
” [red cardstock].
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” [red cardstock]
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||||
</materialsCitation>
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||||
.
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</paragraph>
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<paragraph id="D0C2AABF03084097E4172B62E95C2615">
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<materialsCitation id="13248069306C88C2FFF343D5377268DD" collectionCode="NCSU" country="United States of America" county="Newberry Co." location="USA" specimenCount="1" stateProvince="South Carolina" typeStatus="paratype">
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<materialsCitation id="3847C9474F308FAD039BBEA4CDF5BA95" collectedFrom="Pine Stump Hole" collectingDate="2007-12-08" collectingMethod="sift and berlese litter 20 "" collectionCode="NCSU" collectorName="J. and S. Cornell" country="United States of America" county="Newberry Co." latitude="34.188" location="Little Mountain" longLatPrecision="71" longitude="-81.408" specimenCode="ENT 00327998, ENT 00327999" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="paratype">
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||||
<emphasis id="45167600A8DAC4474860988A2FEB8C1E" bold="true" italics="true">
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||||
<typeStatus id="1B03E90D892B4974B501D7FC5062A1C5" type="paratype">Paratypes</typeStatus>
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||||
</emphasis>
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@ -119,9 +118,7 @@ _
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<emphasis id="0CB4ED7AB33D7BEF12195F5A7E02E6C5" bold="true">
|
||||
<collectingCountry id="569AD53A44936AFE1E9B85382D0420E8" name="United States of America">USA</collectingCountry>
|
||||
</emphasis>
|
||||
</materialsCitation>
|
||||
•
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||||
<materialsCitation id="150AA4B8E2D1EF9A422E45D40E8C049D" collectingDate="2007-12-08" collectionCode="NCSU" collectorName="J. and S. Cornell & D Pine Stump Hole" country="United States of America" county="Newberry Co." latitude="34.188" location="Little Mountain" longLatPrecision="71" longitude="-81.408" specimenCode="ENT 00327998, ENT 00327999" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="paratype">
|
||||
<emphasis id="CC16837346E89CD6928199693E6B1F14" bold="true">
|
||||
<collectingRegion id="703573D60FE79A5302880117D4505581" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
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</emphasis>
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@ -147,8 +144,10 @@ _
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|||
</date>
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||||
;
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<collectorName id="CED25CFCE9195F1F0BE7A6E6807906D9">J. and S. Cornell</collectorName>
|
||||
leg.; sift and berlese litter 20 ”
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||||
<collectorName id="A7306F20EFAD61142F949FB6D1276B50">D Pine Stump Hole</collectorName>
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leg.;
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||||
<collectingMethod id="5229592A73CB9AF14041D55CA2B07DF6">sift and berlese litter 20 ”</collectingMethod>
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||||
D
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<collectedFrom id="184B289F8CC5C530933810AEC2842876">Pine Stump Hole</collectedFrom>
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||||
# 5;
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<collectionCode id="3FE4F5A0105269B8FBFDC7197B22A90C" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
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_
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@ -188,9 +187,7 @@ in which males have a flattened medial section of the second abdominal ventrite
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(
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<typeStatus id="EF5295478D6696CD109ED44F5B0AAEE7" type="paratype">paratypes</typeStatus>
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), the elytra and pronotum are broad, the head is relatively small (
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<abbrev id="ABBRID0ETNEK" xlink_title="Maximum head width">
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||||
<collectionCode id="6ECDD8026DB54FE1D2E14D1C06B18829">HW</collectionCode>
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||||
</abbrev>
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<abbrev id="ABBRID0ETNEK" xlink_title="Maximum head width">HW</abbrev>
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/
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<abbrev id="ABBRID0EXNEK" xlink_title="Maximum pronotal width">PW</abbrev>
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= 0.65–0.70), the pronotum has the posterior angles produced posteriorly (Fig.
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@ -232,13 +229,9 @@ median lobe. Scale bars: 1 mm (
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<paragraph id="8D0323BAACC0A3605D37369E8C96E1F5">
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<emphasis id="2E8B3659A32F57D4A0523B008D74B278" bold="true" italics="true">Habitus</emphasis>
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||||
Large (
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||||
<abbrev id="ABBRID0EUPEK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">
|
||||
<collectionCode id="047957EF6AF7F247544669859B2A3275" country="Netherlands" lsid="urn:lsid:biocol.org:col:12548" name="Adviesbureau voor Bryologie en Lichenologie" type="Herbarium">ABL</collectionCode>
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||||
</abbrev>
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||||
<abbrev id="ABBRID0EUPEK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
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= 2.79–2.87) and robust (
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<abbrev id="ABBRID0EYPEK" xlink_title="Maximum pronotal width">
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<collectionCode id="D56199D6E38C263E8E5672171B5FC7D5" country="Thailand" name="Paleontological Collections">PW</collectionCode>
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||||
</abbrev>
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||||
<abbrev id="ABBRID0EYPEK" xlink_title="Maximum pronotal width">PW</abbrev>
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/
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<abbrev id="ABBRID0E3PEK" xlink_title="Maximum elytral width">EW</abbrev>
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= 0.84–0.88,
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@ -246,9 +239,7 @@ Large (
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/
|
||||
<abbrev id="ABBRID0EEQEK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
|
||||
= 0.36–0.38), with relatively small head (
|
||||
<abbrev id="ABBRID0EIQEK" xlink_title="Maximum head width">
|
||||
<collectionCode id="B3EE81F7C3634F3A50D489C5FA8D6428">HW</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EIQEK" xlink_title="Maximum head width">HW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EMQEK" xlink_title="Maximum pronotal width">PW</abbrev>
|
||||
= 0.65–0.70).
|
||||
|
|
@ -260,9 +251,7 @@ Frontoclypeal horn well-developed and prominent. Ocular tubercle present on each
|
|||
). Antennae short, not reaching posterior margin of pronotum when bent backward.
|
||||
<emphasis id="8FADDE9B91D1A826E9D70A349C251B07" bold="true" italics="true">Pronotum</emphasis>
|
||||
Broad (
|
||||
<abbrev id="ABBRID0E4QEK" xlink_title="Maximum pronotal width">
|
||||
<collectionCode id="487A786B791DE32E3412779CAA53DCC2" country="Thailand" name="Paleontological Collections">PW</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0E4QEK" xlink_title="Maximum pronotal width">PW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EBREK" xlink_title="Maximum elytral width">EW</abbrev>
|
||||
= 0.84–0.88), margins not sinuate before posterior angles, which are slightly constricted (
|
||||
|
|
@ -272,9 +261,7 @@ Broad (
|
|||
= 0.69–0.71) and protrude beyond posterior pronotal margin.
|
||||
<emphasis id="C2B397491ECD1B0D7BA3B89FF977EFD4" bold="true" italics="true">Elytra</emphasis>
|
||||
Broad (
|
||||
<abbrev id="ABBRID0EQREK" xlink_title="Maximum elytral width">
|
||||
<collectionCode id="31C7810EE9D019F9C3BAED4291D8D6C7">EW</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EQREK" xlink_title="Maximum elytral width">EW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EUREK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
|
||||
= 0.36–0.38), disc flattened, with traces of five striae.
|
||||
|
|
@ -338,9 +325,7 @@ dorsal aspect of abdominal ventrites of
|
|||
<heading id="0A27D234268746EC46DA589B36D8C529" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="41BC199DE2236AE3B4C3924E024344EC">
|
||||
Known from two localities in
|
||||
<collectingRegion id="FACBE970590AC50C5E8B3769DC248093" country="United Kingdom" name="York">York</collectingRegion>
|
||||
and Newberry Counties (Fig.
|
||||
Known from two localities in York and Newberry Counties (Fig.
|
||||
<figureCitation id="6D6540F27369CE9B5BAC0B6CCCB51A2A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution map of Serranillus species that occur in South Carolina. The locality for Serranillus sp. “ South Carolina, Coon Branch ” is the same as the single South Carolina S. jeanneli occurrence, and is not shown. Data are from Harden (2024)." figureDoi="10.3897/zookeys.1209.125897.figure16" httpUri="https://binary.pensoft.net/fig/1110355">16</figureCitation>
|
||||
), both isolated monadnocks in the
|
||||
<collectingRegion id="8019BC1EED7534D58FE25E1FDC9DB899" country="Italy" name="Piemonte">Piedmont</collectingRegion>
|
||||
|
|
@ -444,7 +429,7 @@ Figs
|
|||
<heading id="DF85D1FBE4AD1D4929C68B300AAAA82A" reason="title">Material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FBB4127A6CEF0B18380CD99C7DB00B34">
|
||||
<materialsCitation id="36E8180AC7DD06CDAEB747B9B34263F1" collectingDate="2021-10-02" collectionCode="CUAC" collectorName="C. W. Harden & On" country="United States of America" county="Oconee Co." latitude="35.0251" location="Coon Branch Natural Area" longLatPrecision="6" longitude="-83.0053" specimenCode="CUAC 000169317" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina">
|
||||
<materialsCitation id="36E8180AC7DD06CDAEB747B9B34263F1" collectedFrom="On underside of embedded rock, steep north-facing slope" collectingDate="2021-10-02" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="35.0251" location="Coon Branch Natural Area" longLatPrecision="6" longitude="-83.0053" specimenCode="CUAC 000169317" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina">
|
||||
<emphasis id="C03371E95FC61A81DC799B71F6103D2A" bold="true">
|
||||
<collectingCountry id="A29D6386326AF1D6B864C2404F3CA156" name="United States of America">USA</collectingCountry>
|
||||
</emphasis>
|
||||
|
|
@ -473,8 +458,8 @@ Figs
|
|||
;
|
||||
<collectorName id="D4D7C17B163C9B5483B7F306AF0FE545">C. W. Harden</collectorName>
|
||||
leg.;
|
||||
<collectorName id="650ABE16268C7E116152D5F16390B3AF">On</collectorName>
|
||||
underside of embedded rock, steep north-facing slope;
|
||||
<collectedFrom id="D4D85B6E27449B882418117AB547CAFD">On underside of embedded rock, steep north-facing slope</collectedFrom>
|
||||
;
|
||||
<specimenCode id="119E48F22E39E431884E147A7FE6CAF0" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000169317</specimenCode>
|
||||
, CWH- 400;
|
||||
<named-content id="6CA57634A4BFD4AFF8D30F6836CC3B24" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
|
||||
|
|
@ -505,9 +490,7 @@ GenBank:
|
|||
<paragraph id="889F8131EEC542B3A96D5707CBB16A50">
|
||||
<emphasis id="60074B2DECD461FBBD31FFFF1117F1C3" bold="true">Diagnosis.</emphasis>
|
||||
In addition to being larger (
|
||||
<abbrev id="ABBRID0EM1EK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">
|
||||
<collectionCode id="1118BED3B48DE2BBB694CD298A748DF2" country="Netherlands" lsid="urn:lsid:biocol.org:col:12548" name="Adviesbureau voor Bryologie en Lichenologie" type="Herbarium">ABL</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EM1EK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
|
||||
=
|
||||
<quantity id="D79EB941298538065C8A88B05E24A8BD" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.70 mm</quantity>
|
||||
), this female specimen differs from the widespread
|
||||
|
|
@ -568,11 +551,11 @@ did not collect this species, nor did a soilwash sample taken in
|
|||
<paragraph id="8F65CB07668BC49BE3D8285CDAFEF30D">
|
||||
<emphasis id="5DBB923C84549EACB991755FAC8DDAD8" bold="true">Notes.</emphasis>
|
||||
Without associated males, the identity of this species is uncertain. This female from Coon Branch possibly belongs to one of the undescribed species known from adjacent parts of
|
||||
<collectingCountry id="27119A5958931D8347DC6845CE3DE280" name="Georgia">Georgia</collectingCountry>
|
||||
<collectingRegion id="8BC7DC8EED3F6830AB4C5275E997E9C8" country="United States of America" name="Georgia">Georgia</collectingRegion>
|
||||
and North Carolina that we lack DNA sequence data for. Males of both
|
||||
<emphasis id="0B970EFDFC686B4D48D886A492173CA5" italics="true">S.</emphasis>
|
||||
sp. “
|
||||
<collectingCountry id="84853EDE49360AF8B32ABA7BC88D6398" name="Georgia">Georgia</collectingCountry>
|
||||
<collectingRegion id="BFDF45EB1834DF6D9EEC6FBA619A321D" country="United States of America" name="Georgia">Georgia</collectingRegion>
|
||||
, Rabun Bald sp. 1 ” and
|
||||
<emphasis id="5D030A88EA2381F8C53FCC95CD373EEB" italics="true">S.</emphasis>
|
||||
sp. “ North Carolina, Riley Knob ” possess genitalia similar to
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue