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<document id="024808FCED196CA0683465719E3138A7" ID-DOI="10.4202/app.01076.2023" ID-ISSN="1732-2421" ID-Zenodo-Dep="12198764" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718943727872" checkinUser="felipe" docAuthor="Bianucci, Giovanni, Sielfeld, Walter, Olguin, Nicole A. & Guzmán, Guillermo" docDate="2023" docId="039187881E02FA395B06FD06FC55FA0F" docLanguage="en" docName="ActaPalaeontolPol.68.3.477-491.pdf" docOrigin="Acta Palaeontologica Polonica 68 (3)" docSource="http://dx.doi.org/10.4202/app.01076.2023" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Ihlengesi changoensis Bianucci & Sielfeld & Olguin & Guzmán 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="484" masterDocId="FFA8FFF01E01FA3E5B64FFBFFFEBFFB2" masterDocTitle="A new diminutive fossil ziphiid from the deep-sea floor off northern Chile and some remarks on the body size evolution and palaeobiogeography of the beaked whales" masterLastPageNumber="491" masterPageNumber="477" pageNumber="480" updateTime="1744224479251" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="DC32061F18984CD0C706A622FEB1960B">
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<mods:title id="533E15A4094C01E2F4266ADFD2331961">A new diminutive fossil ziphiid from the deep-sea floor off northern Chile and some remarks on the body size evolution and palaeobiogeography of the beaked whales</mods:title>
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<mods:namePart id="98EE898E36BBCABD037DC532412BA616">Bianucci, Giovanni</mods:namePart>
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<mods:namePart id="3DBEA41DA1CC0C97DD329ADFD1404C23">Sielfeld, Walter</mods:namePart>
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<mods:namePart id="1A40295806A53EE5B02EBBF15FF74EA9">Olguin, Nicole A.</mods:namePart>
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<mods:namePart id="7B78C61CAF5E3A3FB015B9AF8CF5FD8E">Guzmán, Guillermo</mods:namePart>
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<mods:title id="777271B02B26F145D9818F4B79C4DA7B">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="48010A27C3F1D4AAC4DEAAA1A4466801">2023</mods:date>
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<mods:number id="96D90055E26E427D5B3A8B9836A466E0">2023-08-28</mods:number>
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<mods:url id="6B8C7A17B0685BF3F5E68D335E367CA5">http://dx.doi.org/10.4202/app.01076.2023</mods:url>
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<mods:identifier id="38290F3C44C3E7968A6E42BA1FB6BF0E" type="DOI">10.4202/app.01076.2023</mods:identifier>
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<mods:identifier id="0D889DCF8FFE4ABF6498AABCC4F59602" type="ISSN">1732-2421</mods:identifier>
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<mods:identifier id="0094F334B0A75E8A438A26239DFC3124" type="Zenodo-Dep">12198764</mods:identifier>
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<treatment id="039187881E02FA395B06FD06FC55FA0F" LSID="urn:lsid:plazi:treatment:039187881E02FA395B06FD06FC55FA0F" httpUri="http://treatment.plazi.org/id/039187881E02FA395B06FD06FC55FA0F" lastPageId="7" lastPageNumber="484" pageId="3" pageNumber="480">
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<subSubSection id="C32265151E02FA3D5B06FD06FE03FD67" box="[98,488,697,726]" pageId="3" pageNumber="480" type="nomenclature">
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<paragraph id="8B87369E1E02FA3D5B06FD06FE03FD67" blockId="3.[98,488,697,763]" box="[98,488,697,726]" pageId="3" pageNumber="480">
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<taxonomicName id="4C384D1D1E02FA3D5B06FD06FE96FD64" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[98,381,697,726]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
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||||
<emphasis id="B94CEA8C1E02FA3D5B06FD06FE96FD64" box="[98,381,697,726]" italics="true" pageId="3" pageNumber="480">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E02FA3D5AE1FD06FE03FD67" box="[389,488,697,725]" pageId="3" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="C32265151E02FA3D5B06FD5AFF14FC8C" pageId="3" pageNumber="480" type="description">
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<paragraph id="8B87369E1E02FA3D5B06FD5AFEF8FD48" blockId="3.[98,488,697,763]" box="[98,275,741,763]" pageId="3" pageNumber="480">
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<figureCitation id="13032A1B1E02FA3D5B06FD5AFF57FD49" box="[98,188,741,763]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[152,185,1511,1533]" captionStartId-1="5.[99,132,1696,1718]" captionStartId-2="6.[152,185,1191,1213]" captionStartId-3="7.[98,131,551,573]" captionTargetBox-0="[156,1531,170,1482]" captionTargetBox-1="[208,1371,159,1672]" captionTargetBox-2="[155,1529,161,1169]" captionTargetBox-3="[98,768,159,527]" captionTargetId-0="figure-241@4.[177,1531,160,1487]" captionTargetId-1="figure-103@5.[210,1371,160,1640]" captionTargetId-2="figure-340@6.[189,1529,169,1160]" captionTargetId-3="figure-819@7.[98,771,173,528]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 2. Holotype of the beaked whale Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A1), right lateral (A2), and ventral (A3) views." captionText-1="Fig.3. Comparison of the neurocrania and the posterior portion of the rostra in dorsal view in two beaked whales.A. The holotype of Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.B. The holotype of Ihlengesi saldanhae (SAM PQ 2792) from the sea floor off Saldanha Bay, South Africa; age unknown." captionText-2="Fig. 4. Comparison of the crania in anterior view of the holotype of three beaked whales. A. Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. B. The holotype of Ihlengesi saldanhae (SAM PQ 2792) from the sea floor off Saldanha Bay, South Africa; age unknown. C. The holotype Khoikhoicetus agulhasis (SAM PQ 2678) from the sea floor off Cape Agulhas, South Africa; age unknown. Diagonal lines represent broken surfaces." captionText-3="Fig. 5. Comparison of the rostra and posterior portion of the neurocrania in dorsal view of two beaked whales. A. The holotype of Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. B. The paratype of Ihlengesi saldanhae (SAM PQ 69673) from the sea floor off Saldanha Bay, South Africa; age unknown." figureDoi-0="http://doi.org/10.5281/zenodo.12198768" figureDoi-1="http://doi.org/10.5281/zenodo.12198770" figureDoi-2="http://doi.org/10.5281/zenodo.12198772" figureDoi-3="http://doi.org/10.5281/zenodo.12198774" httpUri-0="https://zenodo.org/record/12198768/files/figure.png" httpUri-1="https://zenodo.org/record/12198770/files/figure.png" httpUri-2="https://zenodo.org/record/12198772/files/figure.png" httpUri-3="https://zenodo.org/record/12198774/files/figure.png" pageId="3" pageNumber="480">Figs. 2–5</figureCitation>
|
||||
,
|
||||
<tableCitation id="C6BA03251E02FA3D5BA2FD5AFEE6FD48" box="[198,269,741,763]" captionStart="Table 1" captionStartId="3.[99,152,1100,1122]" captionTargetId="graphics-517@3.[129,1449,1169,1952]" captionTargetPageId="3" captionText="Table 1. Measurements (in mm) of the crania of Ihlengesi changoensis sp. nov.and Ihlengesi saldanhae (measurements from Bianucci et al. 2007). Abbreviations: e, estimate; –, no data." pageId="3" pageNumber="480">Table 1</tableCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E02FA3D5B06FCB3FF14FC8C" blockId="3.[98,771,780,1075]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D5B06FCB3FF07FC90" box="[98,236,780,802]" italics="true" pageId="3" pageNumber="480">Zoobank LSID</emphasis>
|
||||
:
|
||||
<uri id="FFA93A9C1E02FA3D5B9CFCB3FF14FC8C" pageId="3" pageNumber="480">
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||||
urn:lsid:zoobank.org:act:
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||||
<uuid id="FF9E0C4B1E02FA3D5ABAFCB3FF14FC8C" pageId="3" pageNumber="480">503E74DA-918C-4125-B419- C4A084E9F317</uuid>
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</uri>
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E02FA3D5B06FCF4FD8BFCCF" pageId="3" pageNumber="480" type="etymology">
|
||||
<paragraph id="8B87369E1E02FA3D5B06FCF4FD8BFCCF" blockId="3.[98,771,780,1075]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D5B06FCF4FF22FCD2" box="[98,201,843,864]" italics="true" pageId="3" pageNumber="480">Etymology</emphasis>
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||||
: From “Changos”, the indigenous people and fishermen, who originally inhabited the coast of Northern
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<collectingCountry id="F32F760E1E02FA3D5941FCD8FDB7FCCF" box="[549,604,871,893]" name="Chile" pageId="3" pageNumber="480">Chile</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E02FA3D5B06FC36FA2BFF06" pageId="3" pageNumber="480" type="materials_examined">
|
||||
<materialsCitation id="3B503CC31E02FA3D5B06FC36FF34FB80" collectingDate="2022-05" collectionCode="MUAP" collectorName="Dany Manzo" country="Chile" elevation="1000" latitude="-19.597223" location="Pisagua" longLatPrecision="21" longitude="-70.21333" municipality="Pisagua" pageId="3" pageNumber="480" specimenCode="MUAP(MM)-068" specimenCount="1" typeStatus="holotype">
|
||||
<paragraph id="8B87369E1E02FA3D5B06FC36FE75FC6A" blockId="3.[98,771,780,1075]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D5B06FC36FF51FC2C" box="[98,186,905,926]" italics="true" pageId="3" pageNumber="480">
|
||||
<typeStatus id="5483883C1E02FA3D5B06FC36FF51FC2C" box="[98,186,905,926]" pageId="3" pageNumber="480" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCode id="DB9E9EE51E02FA3D5BA2FC36FE9EFC2D" box="[198,373,905,927]" pageId="3" pageNumber="480">MUAP(MM)-068</specimenCode>
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||||
, a partial cranium including the complete rostrum, the medial facial area (premaxillary sac fossae and vertex) and part of both supraorbital regions.
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||||
</paragraph>
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||||
<paragraph id="8B87369E1E02FA3D5B06FC5AFF08FB80" blockId="3.[98,771,780,1075]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D5B06FC5AFF0AFC4B" box="[98,225,996,1018]" italics="true" pageId="3" pageNumber="480">Type locality</emphasis>
|
||||
: Off the port of
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<collectingMunicipality id="6BE3ACE41E02FA3D5AF2FC5BFE0AFC48" box="[406,481,996,1018]" pageId="3" pageNumber="480">Pisagua</collectingMunicipality>
|
||||
(
|
||||
<geoCoordinate id="EE0C50591E02FA3D5A97FC5BFD8CFC48" box="[499,615,996,1018]" degrees="19" direction="south" minutes="35" orientation="latitude" pageId="3" pageNumber="480" precision="15" seconds="50" value="-19.597223">19°35’50”S</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="EE0C50591E02FA3D5912FC5BFD1CFC4B" box="[630,759,996,1017]" degrees="70" direction="west" minutes="12" orientation="longitude" pageId="3" pageNumber="480" precision="15" seconds="48" value="-70.21333">70°12’48’’W</geoCoordinate>
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||||
), Northern
|
||||
<collectingCountry id="F32F760E1E02FA3D5BD9FBBEFF18FBA5" box="[189,243,1025,1047]" name="Chile" pageId="3" pageNumber="480">Chile</collectingCountry>
|
||||
, at a depth of
|
||||
<quantity id="4CC09B7B1E02FA3D5A17FBBEFE51FBA4" box="[371,442,1025,1046]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="480" unit="m" value="1000.0">
|
||||
<elevation id="0015D1AD1E02FA3D5A17FBBEFE51FBA4" box="[371,442,1025,1046]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="480" unit="m" value="1000.0">1000 m</elevation>
|
||||
</quantity>
|
||||
(
|
||||
<figureCitation id="13032A1B1E02FA3D5AA2FBBEFE17FBA4" box="[454,508,1025,1047]" captionStart="Fig" captionStartId="2.[152,185,778,800]" captionTargetBox="[258,1428,162,756]" captionTargetId="figure-509@2.[634,1343,161,737]" captionTargetPageId="2" captionText="Fig. 1. A. Schematic map providing the position of Pisagua in South America. B. Schematic map of Northern Chile coast around Pisagua and sea floor bathymetry showing approximative discovery locality of the holotype MUAP(MM)-068 cranium of the beaked whale Ihlengesi changoensis sp. nov. at a depth of 1000 m (star)." figureDoi="http://doi.org/10.5281/zenodo.12198766" httpUri="https://zenodo.org/record/12198766/files/figure.png" pageId="3" pageNumber="480">Fig. 1</figureCitation>
|
||||
), collected by
|
||||
<collectorName id="26CD53481E02FA3D59E3FBBEFCE8FBA4" box="[647,771,1025,1046]" pageId="3" pageNumber="480">Dany Manzo</collectorName>
|
||||
in
|
||||
<date id="FF86105E1E02FA3D5B1EFBA1FF34FB80" box="[122,223,1053,1075]" pageId="3" pageNumber="480" value="2022-05">
|
||||
<collectingDate id="EFC2E9B61E02FA3D5B1EFBA1FF34FB80" box="[122,223,1053,1075]" pageId="3" pageNumber="480" value="2022-05">May 2022</collectingDate>
|
||||
</date>
|
||||
.
|
||||
</paragraph>
|
||||
</materialsCitation>
|
||||
<paragraph id="8B87369E1E02FA3D584CFF1FFA2BFF06" blockId="3.[808,1472,159,181]" box="[808,1472,159,181]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFF1FFC4FFF06" box="[808,932,159,181]" italics="true" pageId="3" pageNumber="480">
|
||||
<typeStatus id="5483883C1E02FA3D584CFF1FFCBEFF07" box="[808,853,160,181]" pageId="3" pageNumber="480">Type</typeStatus>
|
||||
horizon
|
||||
</emphasis>
|
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: Iquique Basin, Plio-Pleistocene (Gonzáles et al. 2023).
|
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</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="C32265151E02FA3D584CFF78FBBCFD36" pageId="3" pageNumber="480" type="diagnosis">
|
||||
<paragraph id="8B87369E1E02FA3D584CFF78FBBCFD36" blockId="3.[808,1481,198,644]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFF78FC77FF52" box="[808,924,199,224]" italics="true" pageId="3" pageNumber="480">Diagnosis</emphasis>
|
||||
.—
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||||
<taxonomicName id="4C384D1D1E02FA3D58DBFF79FB29FF52" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[959,1218,198,224]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E02FA3D58DBFF79FB29FF52" box="[959,1218,198,224]" italics="true" pageId="3" pageNumber="480">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E02FA3D5FB4FF78FAC7FF53" box="[1232,1324,199,225]" pageId="3" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="4C384D1D1E02FA3D584CFF58FBF9FEB3" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[808,1042,231,257]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFF58FBF9FEB3" box="[808,1042,231,257]" italics="true" pageId="3" pageNumber="480">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
in the following characters: more elongated premaxillary sac fossa and consequently premaxillary foramen more anteriorly located (ratio between the longitudinal distance between the right premaxillary foramen and the rostrum base and the width of rostrum base equals 0.10 contra
|
||||
<quantity id="4CC09B7B1E02FA3D58D2FE37FBE3FE10" box="[950,1032,392,418]" metricMagnitude="-3" metricUnit="m" metricValue="8.128" pageId="3" pageNumber="480" unit="in" value="0.32">0.32 in</quantity>
|
||||
<taxonomicName id="4C384D1D1E02FA3D5F71FE36FB4EFE10" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[1045,1189,392,418]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5F71FE36FB4EFE10" box="[1045,1189,392,418]" italics="true" pageId="3" pageNumber="480">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
); dorsal margin of each premaxillary crest sloping markedly ventrolaterally and generating an acute dorsal profile of the vertex in anterior view (rounded profile in
|
||||
<taxonomicName id="4C384D1D1E02FA3D5F3CFE55FB03FDB1" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[1112,1256,489,515]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5F3CFE55FB03FDB1" box="[1112,1256,489,515]" italics="true" pageId="3" pageNumber="480">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
); less anterolateral extension of the right nasal forming part of the premaxillary crest; lateral margins of the nasals not anteriorly diverging but weakly convex; nasofrontal suture not convex posteriorly but anteriorly pointed.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E02FA39584CFD29FA21FCF2" lastPageId="7" lastPageNumber="484" pageId="3" pageNumber="480" type="description">
|
||||
<paragraph id="8B87369E1E02FA3D584CFD29FAB6FC41" blockId="3.[808,1481,661,1076]" pageId="3" pageNumber="480">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFD29FC40FD1D" box="[808,939,662,687]" italics="true" pageId="3" pageNumber="480">Description</emphasis>
|
||||
.—Judging from the size of the cranium (
|
||||
<tableCitation id="C6BA03251E02FA3D5E0EFD2AFA52FD02" box="[1386,1465,661,688]" captionStart="Table 1" captionStartId="3.[99,152,1100,1122]" captionTargetId="graphics-517@3.[129,1449,1169,1952]" captionTargetPageId="3" captionText="Table 1. Measurements (in mm) of the crania of Ihlengesi changoensis sp. nov.and Ihlengesi saldanhae (measurements from Bianucci et al. 2007). Abbreviations: e, estimate; –, no data." pageId="3" pageNumber="480">Table 1</tableCitation>
|
||||
),
|
||||
<taxonomicName id="4C384D1D1E02FA3D584CFD09FBCAFD62" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[808,1057,694,720]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFD09FBCAFD62" box="[808,1057,694,720]" italics="true" pageId="3" pageNumber="480">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
was a small ziphiid which, like
|
||||
<emphasis id="B94CEA8C1E02FA3D5EE1FD09FC36FD42" italics="true" pageId="3" pageNumber="480">
|
||||
<taxonomicName id="4C384D1D1E02FA3D5EE1FD09FC32FD42" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">Ihlengesi saldanhae</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
did not reach
|
||||
<quantity id="4CC09B7B1E02FA3D5FF7FD69FB28FD43" box="[1171,1219,726,753]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="480" unit="m" value="4.0">4 m</quantity>
|
||||
in length. Indeed, by roughly estimating the postorbital width of the cranium to
|
||||
<quantity id="4CC09B7B1E02FA3D5823FCA8FC4EFC83" box="[839,933,791,818]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="480" unit="mm" value="250.0">250 mm</quantity>
|
||||
and applying the aforementioned equation proposed by Bianucci et al. (2008), the body length results to be
|
||||
<quantity id="4CC09B7B1E02FA3D5829FCE8FC78FCC0" box="[845,915,855,882]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="480" unit="m" value="3.5">3.5 m</quantity>
|
||||
, a value close to that estimated for
|
||||
<taxonomicName id="4C384D1D1E02FA3D5E5EFCE7FA22FCC3" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[1338,1481,855,881]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5E5EFCE7FA22FCC3" box="[1338,1481,855,881]" italics="true" pageId="3" pageNumber="480">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<quantity id="4CC09B7B1E02FA3D5854FCC7FC9FFC20" box="[816,884,888,914]" metricMagnitude="0" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="480" unit="m" value="3.1">3.1 m</quantity>
|
||||
). Therefore, based on these estimations, species of
|
||||
<taxonomicName id="4C384D1D1E02FA3D584CFC27FC64FC00" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[808,911,920,946]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E02FA3D584CFC27FC64FC00" box="[808,911,920,946]" italics="true" pageId="3" pageNumber="480">Ihlengesi</emphasis>
|
||||
</taxonomicName>
|
||||
were smaller than all extant ziphiids with the exception of the pygmy beaked whale
|
||||
<taxonomicName id="4C384D1D1E02FA3D5FDCFC07FA23FC60" authorityName="Reyes, Mead & Van Waerebeek" authorityYear="1991" box="[1208,1480,952,978]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="peruvianus">
|
||||
<emphasis id="B94CEA8C1E02FA3D5FDCFC07FA23FC60" box="[1208,1480,952,978]" italics="true" pageId="3" pageNumber="480">Mesoplodon peruvianus</emphasis>
|
||||
</taxonomicName>
|
||||
(maximum body length
|
||||
<quantity id="4CC09B7B1E02FA3D5F5CFC66FB6DFC41" box="[1080,1158,985,1011]" metricMagnitude="0" metricUnit="m" metricValue="3.72" pageId="3" pageNumber="480" unit="m" value="3.72">3.72 m</quantity>
|
||||
;
|
||||
<bibRefCitation id="EFA94B6F1E02FA3D5FF6FC66FAB9FC41" author="Reyes" box="[1170,1362,985,1011]" etAl="et al." firstAuthor="Reyes" pageId="3" pageNumber="480" pagination="1 - 24" refId="ref13141" refString="Reyes, J. C., Mead, J. G., and Van Waerebeek, K. 1991. A new species of beaked whale Mesoplodon peruvianus sp. n. (Cetacea Ziphiidae) from Peru. Marine Mammal Science 7: 1 - 24." type="journal article" year="1991">Reyes et al. 1991</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E02FA3A582AFC46FD79F8F5" blockId="3.[808,1481,661,1076]" lastBlockId="4.[151,824,1581,1960]" lastPageId="4" lastPageNumber="481" pageId="3" pageNumber="480">
|
||||
The complete rostrum is moderately elongated and, similarly to the
|
||||
<typeStatus id="5483883C1E02FA3D58BBFBA5FBA9FB86" box="[991,1090,1050,1076]" pageId="3" pageNumber="480" type="paratype">paratype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C384D1D1E02FA3D5F0FFBA6FAA5FB81" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[1131,1358,1049,1075]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5F0FFBA6FAA5FB81" box="[1131,1358,1049,1075]" italics="true" pageId="3" pageNumber="480">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
(SAM PQ 69673), shows a strong lateral compression generating a narrow dorsal profile and a transverse section with a height significantly greater than its width (
|
||||
<figureCitation id="13032A1B1E05FA3A595FF9D2FD66F935" box="[571,653,1645,1671]" captionStart="Fig" captionStartId="4.[152,185,1511,1533]" captionTargetBox="[156,1531,170,1482]" captionTargetId="figure-241@4.[177,1531,160,1487]" captionTargetPageId="4" captionText="Fig. 2. Holotype of the beaked whale Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A1), right lateral (A2), and ventral (A3) views." figureDoi="http://doi.org/10.5281/zenodo.12198768" httpUri="https://zenodo.org/record/12198768/files/figure.png" pageId="4" pageNumber="481">Figs. 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13032A1B1E05FA3A59F8F9D2FD47F93A" box="[668,684,1645,1672]" captionStart="Fig" captionStartId="7.[98,131,551,573]" captionTargetBox="[98,768,159,527]" captionTargetId="figure-819@7.[98,771,173,528]" captionTargetPageId="7" captionText="Fig. 5. Comparison of the rostra and posterior portion of the neurocrania in dorsal view of two beaked whales. A. The holotype of Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. B. The paratype of Ihlengesi saldanhae (SAM PQ 69673) from the sea floor off Saldanha Bay, South Africa; age unknown." figureDoi="http://doi.org/10.5281/zenodo.12198774" httpUri="https://zenodo.org/record/12198774/files/figure.png" pageId="4" pageNumber="481">5</figureCitation>
|
||||
). Due to its narrow dorsal shape, the rostrum of
|
||||
<taxonomicName id="4C384D1D1E05FA3A592AF932FF5EF975" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E05FA3A592AF932FF5EF975" italics="true" pageId="4" pageNumber="481">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
clearly differs from
|
||||
<taxonomicName id="4C384D1D1E05FA3A5AC7F912FD97F97A" box="[419,636,1709,1736]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E05FA3A5AC7F912FDADF975" box="[419,582,1709,1735]" italics="true" pageId="4" pageNumber="481">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
Indeed, the rostrum of this latter genus is significantly transversely wider than in
|
||||
<taxonomicName id="4C384D1D1E05FA3A5B9DF951FE48F8B5" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[249,419,1773,1799]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E05FA3A5B9DF951FE48F8B5" box="[249,419,1773,1799]" italics="true" pageId="4" pageNumber="481">I. changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
, a feature particularly marked in
|
||||
<taxonomicName id="4C384D1D1E05FA3A5BF3F8B2FE52F895" authorityName="Lambert, de Muizon, Duhamel & van Der Plicht" authorityYear="2018" box="[151,441,1805,1831]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="kergueleni">
|
||||
<emphasis id="B94CEA8C1E05FA3A5BF3F8B2FE52F895" box="[151,441,1805,1831]" italics="true" pageId="4" pageNumber="481">Khoikhoicetus kergueleni</emphasis>
|
||||
</taxonomicName>
|
||||
having a triangular outline of the rostrum in dorsal view (
|
||||
<bibRefCitation id="EFA94B6F1E05FA3A5AC2F892FD6DF8F5" author="Lambert" box="[422,646,1837,1864]" etAl="et al." firstAuthor="Lambert" pageId="4" pageNumber="481" pagination="135 - 160" refId="ref11849" refString="Lambert, O., Muizon, C. de, Duhamel, G., and Van der Plicht, J. 2018. Neogene and Quaternary fossil remains of beaked whales (Cetacea, Odontoceti, Ziphiidae) from deep-sea deposits off Crozet and Kerguelen islands, Southern Ocean. Geodiversitas 40: 135 - 160." type="journal article" year="2018">Lambert et al. 2018</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF4766161E02FA3D5B07FBF3FE25FBCD" pageId="3" pageNumber="480" startId="3.[99,152,1100,1122]" targetBox="[137,1430,1177,1950]" targetIsTable="true" targetPageId="3" targetType="table">
|
||||
<paragraph id="8B87369E1E02FA3D5B07FBF3FE25FBCD" blockId="3.[98,1483,1100,1151]" pageId="3" pageNumber="480">
|
||||
Table 1. Measurements (in mm) of the crania of
|
||||
<taxonomicName id="4C384D1D1E02FA3D5943FBF3FD10FBD3" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[551,763,1100,1121]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E02FA3D5943FBF3FD10FBD3" box="[551,763,1100,1121]" italics="true" pageId="3" pageNumber="480">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E02FA3D599BFBF2FCA2FBD0" box="[767,841,1101,1122]" pageId="3" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C384D1D1E02FA3D5817FBF3FBD8FBD3" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[883,1075,1100,1121]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5817FBF3FBD8FBD3" box="[883,1075,1100,1121]" italics="true" pageId="3" pageNumber="480">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
(measurements from Bianucci et al. 2007). Abbreviations: e, estimate;
|
||||
<emphasis id="B94CEA8C1E02FA3D5A0DFBD5FE9EFBCD" bold="true" box="[361,373,1130,1151]" pageId="3" pageNumber="480">–</emphasis>
|
||||
, no data.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B87369E1E02FA3D59A8FB18FAD6F82F" pageId="3" pageNumber="480">
|
||||
<table id="F938C43E1E0205C15BEDFB26FA7DF82C" box="[137,1430,1177,1950]" gridcols="4" gridrows="22" pageId="3" pageNumber="480">
|
||||
<tr id="350834DC1E0205C15BEDFB26FA7DFB79" box="[137,1430,1177,1227]" gridrow="0" pageId="3" pageNumber="480" rowspan-0="1">
|
||||
<th id="76D95DA01E0205C159A8FB26FC1AFB79" box="[716,1009,1177,1227]" gridcol="1" gridrow="0" pageId="3" pageNumber="480">
|
||||
<taxonomicName id="4C384D1D1E02FA3D59A8FB18FC4AFB0E" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[716,929,1191,1212]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E02FA3D59A8FB18FC4AFB0E" box="[716,929,1191,1212]" italics="true" pageId="3" pageNumber="480">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E02FA3D58C2FB18FC1AFB0E" box="[934,1009,1191,1212]" pageId="3" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</th>
|
||||
<th id="76D95DA01E0205C15F72FB26FA7DFB79" box="[1046,1430,1177,1227]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="3" pageNumber="480">
|
||||
<taxonomicName id="4C384D1D1E02FA3D5F72FB26FAF7FB78" authority="Bianucci, Lambert, and Post, 2007 MUAP (MM)" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E02FA3D5F72FB26FB3CFB1C" box="[1046,1239,1177,1198]" italics="true" pageId="3" pageNumber="480">Ihlengesi saldanhae</emphasis>
|
||||
Bianucci, Lambert, and Post, 2007
|
||||
</taxonomicName>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFB6BFA7DFAB4" box="[137,1430,1236,1286]" gridrow="1" pageId="3" pageNumber="480" rowspan-0="1">
|
||||
<td id="76D95DA01E0205C159A8FB6BFC1AFAB4" box="[716,1009,1236,1286]" gridcol="1" gridrow="1" pageId="3" pageNumber="480">MUAP(MM)-068 (holotype)</td>
|
||||
<td id="76D95DA01E0205C15F72FB6BFB40FAB4" box="[1046,1195,1236,1286]" gridcol="2" gridrow="1" pageId="3" pageNumber="480">SAM PQ 2792 (holotype)</td>
|
||||
<td id="76D95DA01E0205C15F8EFB6BFA7DFAB4" box="[1258,1430,1236,1286]" gridcol="3" gridrow="1" pageId="3" pageNumber="480">SAM PQ 69673 (paratype)</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFAAFFA7DFA94" box="[137,1430,1296,1318]" gridrow="2" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFAAFFD45FA94" box="[137,686,1296,1318]" gridcol="0" gridrow="2" pageId="3" pageNumber="480">Length of rostrum</th>
|
||||
<td id="76D95DA01E0205C159A8FAAFFC1AFA94" box="[716,1009,1296,1318]" gridcol="1" gridrow="2" pageId="3" pageNumber="480">540</td>
|
||||
<td id="76D95DA01E0205C15F72FAAFFB40FA94" box="[1046,1195,1296,1318]" gridcol="2" gridrow="2" pageId="3" pageNumber="480">–</td>
|
||||
<td id="76D95DA01E0205C15F8EFAAFFA7DFA94" box="[1258,1430,1296,1318]" gridcol="3" gridrow="2" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFA8FFA7DFAF4" box="[137,1430,1328,1350]" gridrow="3" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFA8FFD45FAF4" box="[137,686,1328,1350]" gridcol="0" gridrow="3" pageId="3" pageNumber="480">Width of rostrum at mid-length</th>
|
||||
<td id="76D95DA01E0205C159A8FA8FFC1AFAF4" box="[716,1009,1328,1350]" gridcol="1" gridrow="3" pageId="3" pageNumber="480">38</td>
|
||||
<td id="76D95DA01E0205C15F72FA8FFB40FAF4" box="[1046,1195,1328,1350]" gridcol="2" gridrow="3" pageId="3" pageNumber="480">–</td>
|
||||
<td id="76D95DA01E0205C15F8EFA8FFA7DFAF4" box="[1258,1430,1328,1350]" gridcol="3" gridrow="3" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFAEFFA7DFAD4" box="[137,1430,1360,1382]" gridrow="4" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFAEFFD45FAD4" box="[137,686,1360,1382]" gridcol="0" gridrow="4" pageId="3" pageNumber="480">Width of rostrum base at prominental notch</th>
|
||||
<td id="76D95DA01E0205C159A8FAEFFC1AFAD4" box="[716,1009,1360,1382]" gridcol="1" gridrow="4" pageId="3" pageNumber="480">104</td>
|
||||
<td id="76D95DA01E0205C15F72FAEFFB40FAD4" box="[1046,1195,1360,1382]" gridcol="2" gridrow="4" pageId="3" pageNumber="480">108</td>
|
||||
<td id="76D95DA01E0205C15F8EFAEFFA7DFAD4" box="[1258,1430,1360,1382]" gridcol="3" gridrow="4" pageId="3" pageNumber="480">104</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFAD0FA7DFA37" box="[137,1430,1391,1413]" gridrow="5" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFAD0FD45FA37" box="[137,686,1391,1413]" gridcol="0" gridrow="5" pageId="3" pageNumber="480">Width of rostrum base at antorbital notch</th>
|
||||
<td id="76D95DA01E0205C159A8FAD0FC1AFA37" box="[716,1009,1391,1413]" gridcol="1" gridrow="5" pageId="3" pageNumber="480">180</td>
|
||||
<td id="76D95DA01E0205C15F72FAD0FB40FA37" box="[1046,1195,1391,1413]" gridcol="2" gridrow="5" pageId="3" pageNumber="480">e143</td>
|
||||
<td id="76D95DA01E0205C15F8EFAD0FA7DFA37" box="[1258,1430,1391,1413]" gridcol="3" gridrow="5" pageId="3" pageNumber="480">e142</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFA30FA7DFA17" box="[137,1430,1423,1445]" gridrow="6" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFA30FD45FA17" box="[137,686,1423,1445]" gridcol="0" gridrow="6" pageId="3" pageNumber="480">Width of premaxillae at rostrum base (anterior notch)</th>
|
||||
<td id="76D95DA01E0205C159A8FA30FC1AFA17" box="[716,1009,1423,1445]" gridcol="1" gridrow="6" pageId="3" pageNumber="480">55</td>
|
||||
<td id="76D95DA01E0205C15F72FA30FB40FA17" box="[1046,1195,1423,1445]" gridcol="2" gridrow="6" pageId="3" pageNumber="480">47</td>
|
||||
<td id="76D95DA01E0205C15F8EFA30FA7DFA17" box="[1258,1430,1423,1445]" gridcol="3" gridrow="6" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFA10FA7DFA77" box="[137,1430,1455,1477]" gridrow="7" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFA10FD45FA77" box="[137,686,1455,1477]" gridcol="0" gridrow="7" pageId="3" pageNumber="480">Distance rostrum base—anterior apex of palatine</th>
|
||||
<td id="76D95DA01E0205C159A8FA10FC1AFA77" box="[716,1009,1455,1477]" gridcol="1" gridrow="7" pageId="3" pageNumber="480">120</td>
|
||||
<td id="76D95DA01E0205C15F72FA10FB40FA77" box="[1046,1195,1455,1477]" gridcol="2" gridrow="7" pageId="3" pageNumber="480">–</td>
|
||||
<td id="76D95DA01E0205C15F8EFA10FA7DFA77" box="[1258,1430,1455,1477]" gridcol="3" gridrow="7" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFA70FA7DFA57" box="[137,1430,1487,1509]" gridrow="8" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFA70FD45FA57" box="[137,686,1487,1509]" gridcol="0" gridrow="8" pageId="3" pageNumber="480">Preorbital width of skull</th>
|
||||
<td id="76D95DA01E0205C159A8FA70FC1AFA57" box="[716,1009,1487,1509]" gridcol="1" gridrow="8" pageId="3" pageNumber="480">e230</td>
|
||||
<td id="76D95DA01E0205C15F72FA70FB40FA57" box="[1046,1195,1487,1509]" gridcol="2" gridrow="8" pageId="3" pageNumber="480">–</td>
|
||||
<td id="76D95DA01E0205C15F8EFA70FA7DFA57" box="[1258,1430,1487,1509]" gridcol="3" gridrow="8" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDFA51FA7DF992" box="[137,1430,1518,1568]" gridrow="9" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDFA51FD45F992" box="[137,686,1518,1568]" gridcol="0" gridrow="9" pageId="3" pageNumber="480">Longitudinal distance right premaxillary foramen —antero-median corner of naris</th>
|
||||
<td id="76D95DA01E0205C159A8FA51FC1AF992" box="[716,1009,1518,1568]" gridcol="1" gridrow="9" pageId="3" pageNumber="480">47</td>
|
||||
<td id="76D95DA01E0205C15F72FA51FB40F992" box="[1046,1195,1518,1568]" gridcol="2" gridrow="9" pageId="3" pageNumber="480">31</td>
|
||||
<td id="76D95DA01E0205C15F8EFA51FA7DF992" box="[1258,1430,1518,1568]" gridcol="3" gridrow="9" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF995FA7DF9F2" box="[137,1430,1578,1600]" gridrow="10" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF995FD45F9F2" box="[137,686,1578,1600]" gridcol="0" gridrow="10" pageId="3" pageNumber="480">Width of premaxillary sac fossae</th>
|
||||
<td id="76D95DA01E0205C159A8F995FC1AF9F2" box="[716,1009,1578,1600]" gridcol="1" gridrow="10" pageId="3" pageNumber="480">130</td>
|
||||
<td id="76D95DA01E0205C15F72F995FB40F9F2" box="[1046,1195,1578,1600]" gridcol="2" gridrow="10" pageId="3" pageNumber="480">105</td>
|
||||
<td id="76D95DA01E0205C15F8EF995FA7DF9F2" box="[1258,1430,1578,1600]" gridcol="3" gridrow="10" pageId="3" pageNumber="480">101</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF9F5FA7DF9D2" box="[137,1430,1610,1632]" gridrow="11" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF9F5FD45F9D2" box="[137,686,1610,1632]" gridcol="0" gridrow="11" pageId="3" pageNumber="480">Width of right premaxillary sac fossa</th>
|
||||
<td id="76D95DA01E0205C159A8F9F5FC1AF9D2" box="[716,1009,1610,1632]" gridcol="1" gridrow="11" pageId="3" pageNumber="480">69</td>
|
||||
<td id="76D95DA01E0205C15F72F9F5FB40F9D2" box="[1046,1195,1610,1632]" gridcol="2" gridrow="11" pageId="3" pageNumber="480">51</td>
|
||||
<td id="76D95DA01E0205C15F8EF9F5FA7DF9D2" box="[1258,1430,1610,1632]" gridcol="3" gridrow="11" pageId="3" pageNumber="480">e52</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF9D5FA7DF932" box="[137,1430,1642,1664]" gridrow="12" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF9D5FD45F932" box="[137,686,1642,1664]" gridcol="0" gridrow="12" pageId="3" pageNumber="480">Width of left premaxillary sac fossa</th>
|
||||
<td id="76D95DA01E0205C159A8F9D5FC1AF932" box="[716,1009,1642,1664]" gridcol="1" gridrow="12" pageId="3" pageNumber="480">55</td>
|
||||
<td id="76D95DA01E0205C15F72F9D5FB40F932" box="[1046,1195,1642,1664]" gridcol="2" gridrow="12" pageId="3" pageNumber="480">46</td>
|
||||
<td id="76D95DA01E0205C15F8EF9D5FA7DF932" box="[1258,1430,1642,1664]" gridcol="3" gridrow="12" pageId="3" pageNumber="480">44</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF936FA7DF912" box="[137,1430,1673,1696]" gridrow="13" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF936FD45F912" box="[137,686,1673,1696]" gridcol="0" gridrow="13" pageId="3" pageNumber="480">Width of bony nares</th>
|
||||
<td id="76D95DA01E0205C159A8F936FC1AF912" box="[716,1009,1673,1696]" gridcol="1" gridrow="13" pageId="3" pageNumber="480">45</td>
|
||||
<td id="76D95DA01E0205C15F72F936FB40F912" box="[1046,1195,1673,1696]" gridcol="2" gridrow="13" pageId="3" pageNumber="480">49</td>
|
||||
<td id="76D95DA01E0205C15F8EF936FA7DF912" box="[1258,1430,1673,1696]" gridcol="3" gridrow="13" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF916FA7DF90D" box="[137,1430,1705,1727]" gridrow="14" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF916FD45F90D" box="[137,686,1705,1727]" gridcol="0" gridrow="14" pageId="3" pageNumber="480">Minimum width of right ascending process of premaxilla</th>
|
||||
<td id="76D95DA01E0205C159A8F916FC1AF90D" box="[716,1009,1705,1727]" gridcol="1" gridrow="14" pageId="3" pageNumber="480">32</td>
|
||||
<td id="76D95DA01E0205C15F72F916FB40F90D" box="[1046,1195,1705,1727]" gridcol="2" gridrow="14" pageId="3" pageNumber="480">25</td>
|
||||
<td id="76D95DA01E0205C15F8EF916FA7DF90D" box="[1258,1430,1705,1727]" gridcol="3" gridrow="14" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF976FA7DF96D" box="[137,1430,1737,1759]" gridrow="15" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF976FD45F96D" box="[137,686,1737,1759]" gridcol="0" gridrow="15" pageId="3" pageNumber="480">Width of premaxillary crests</th>
|
||||
<td id="76D95DA01E0205C159A8F976FC1AF96D" box="[716,1009,1737,1759]" gridcol="1" gridrow="15" pageId="3" pageNumber="480">110</td>
|
||||
<td id="76D95DA01E0205C15F72F976FB40F96D" box="[1046,1195,1737,1759]" gridcol="2" gridrow="15" pageId="3" pageNumber="480">103</td>
|
||||
<td id="76D95DA01E0205C15F8EF976FA7DF96D" box="[1258,1430,1737,1759]" gridcol="3" gridrow="15" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF956FA7DF94D" box="[137,1430,1769,1791]" gridrow="16" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF956FD45F94D" box="[137,686,1769,1791]" gridcol="0" gridrow="16" pageId="3" pageNumber="480">Width of right premaxillary crest</th>
|
||||
<td id="76D95DA01E0205C159A8F956FC1AF94D" box="[716,1009,1769,1791]" gridcol="1" gridrow="16" pageId="3" pageNumber="480">59</td>
|
||||
<td id="76D95DA01E0205C15F72F956FB40F94D" box="[1046,1195,1769,1791]" gridcol="2" gridrow="16" pageId="3" pageNumber="480">47</td>
|
||||
<td id="76D95DA01E0205C15F8EF956FA7DF94D" box="[1258,1430,1769,1791]" gridcol="3" gridrow="16" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF8B6FA7DF8AD" box="[137,1430,1801,1823]" gridrow="17" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF8B6FD45F8AD" box="[137,686,1801,1823]" gridcol="0" gridrow="17" pageId="3" pageNumber="480">Width of left premaxillary crest</th>
|
||||
<td id="76D95DA01E0205C159A8F8B6FC1AF8AD" box="[716,1009,1801,1823]" gridcol="1" gridrow="17" pageId="3" pageNumber="480">42</td>
|
||||
<td id="76D95DA01E0205C15F72F8B6FB40F8AD" box="[1046,1195,1801,1823]" gridcol="2" gridrow="17" pageId="3" pageNumber="480">38</td>
|
||||
<td id="76D95DA01E0205C15F8EF8B6FA7DF8AD" box="[1258,1430,1801,1823]" gridcol="3" gridrow="17" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF897FA7DF88C" box="[137,1430,1832,1854]" gridrow="18" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF897FD45F88C" box="[137,686,1832,1854]" gridcol="0" gridrow="18" pageId="3" pageNumber="480">Minimum distance between premaxillary crests</th>
|
||||
<td id="76D95DA01E0205C159A8F897FC1AF88C" box="[716,1009,1832,1854]" gridcol="1" gridrow="18" pageId="3" pageNumber="480">34</td>
|
||||
<td id="76D95DA01E0205C15F72F897FB40F88C" box="[1046,1195,1832,1854]" gridcol="2" gridrow="18" pageId="3" pageNumber="480">34</td>
|
||||
<td id="76D95DA01E0205C15F8EF897FA7DF88C" box="[1258,1430,1832,1854]" gridcol="3" gridrow="18" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF8F7FA7DF8EC" box="[137,1430,1864,1886]" gridrow="19" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF8F7FD45F8EC" box="[137,686,1864,1886]" gridcol="0" gridrow="19" pageId="3" pageNumber="480">Maximum width of nasals</th>
|
||||
<td id="76D95DA01E0205C159A8F8F7FC1AF8EC" box="[716,1009,1864,1886]" gridcol="1" gridrow="19" pageId="3" pageNumber="480">41</td>
|
||||
<td id="76D95DA01E0205C15F72F8F7FB40F8EC" box="[1046,1195,1864,1886]" gridcol="2" gridrow="19" pageId="3" pageNumber="480">52</td>
|
||||
<td id="76D95DA01E0205C15F8EF8F7FA7DF8EC" box="[1258,1430,1864,1886]" gridcol="3" gridrow="19" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF8D7FA7DF8CC" box="[137,1430,1896,1918]" gridrow="20" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF8D7FD45F8CC" box="[137,686,1896,1918]" gridcol="0" gridrow="20" pageId="3" pageNumber="480">Length of medial suture of nasals on vertex</th>
|
||||
<td id="76D95DA01E0205C159A8F8D7FC1AF8CC" box="[716,1009,1896,1918]" gridcol="1" gridrow="20" pageId="3" pageNumber="480">45</td>
|
||||
<td id="76D95DA01E0205C15F72F8D7FB40F8CC" box="[1046,1195,1896,1918]" gridcol="2" gridrow="20" pageId="3" pageNumber="480">e45</td>
|
||||
<td id="76D95DA01E0205C15F8EF8D7FA7DF8CC" box="[1258,1430,1896,1918]" gridcol="3" gridrow="20" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
<tr id="350834DC1E0205C15BEDF837FA7DF82C" box="[137,1430,1928,1950]" gridrow="21" pageId="3" pageNumber="480">
|
||||
<th id="76D95DA01E0205C15BEDF837FD45F82C" box="[137,686,1928,1950]" gridcol="0" gridrow="21" pageId="3" pageNumber="480">Minimum posterior distance between maxillae</th>
|
||||
<td id="76D95DA01E0205C159A8F837FC1AF82C" box="[716,1009,1928,1950]" gridcol="1" gridrow="21" pageId="3" pageNumber="480">37</td>
|
||||
<td id="76D95DA01E0205C15F72F837FB40F82C" box="[1046,1195,1928,1950]" gridcol="2" gridrow="21" pageId="3" pageNumber="480">32</td>
|
||||
<td id="76D95DA01E0205C15F8EF837FA7DF82C" box="[1258,1430,1928,1950]" gridcol="3" gridrow="21" pageId="3" pageNumber="480">–</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<caption id="DF4766161E05FA3A5BFCFA58FD02F9AB" ID-DOI="http://doi.org/10.5281/zenodo.12198768" ID-Zenodo-Dep="12198768" httpUri="https://zenodo.org/record/12198768/files/figure.png" pageId="4" pageNumber="481" startId="4.[152,185,1511,1533]" targetBox="[156,1531,170,1482]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8B87369E1E05FA3A5BFCFA58FD02F9AB" blockId="4.[151,1536,1511,1563]" pageId="4" pageNumber="481">
|
||||
Fig. 2. Holotype of the beaked whale
|
||||
<taxonomicName id="4C384D1D1E05FA3A5A8FFA58FD58FA4E" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[491,691,1511,1532]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E05FA3A5A8FFA58FD58FA4E" box="[491,691,1511,1532]" italics="true" pageId="4" pageNumber="481">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E05FA3A59DDFA57FD14FA4F" box="[697,767,1512,1533]" pageId="4" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A
|
||||
<subScript id="17BC34DB1E05FA3A5A3CF9B2FE8BF9A9" attach="left" box="[344,352,1549,1563]" fontSize="6" pageId="4" pageNumber="481">1</subScript>
|
||||
), right lateral (A
|
||||
<subScript id="17BC34DB1E05FA3A5A9EF9B2FDE9F9A9" attach="right" box="[506,514,1549,1563]" fontSize="6" pageId="4" pageNumber="481">2</subScript>
|
||||
), and ventral (A
|
||||
<subScript id="17BC34DB1E05FA3A59FCF9B2FD4BF9A9" attach="right" box="[664,672,1549,1563]" fontSize="6" pageId="4" pageNumber="481">3</subScript>
|
||||
) views.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B87369E1E05FA3A5BD9F8F2FBADF8B5" blockId="4.[151,824,1581,1960]" lastBlockId="4.[862,1536,1581,1960]" pageId="4" pageNumber="481">
|
||||
The medial margins of the premaxillae on the rostrum never contact medially and the mesorostral groove is filled by the pachyosteoscleorotic vomer along its whole rostral portion (
|
||||
<figureCitation id="13032A1B1E05FA3A58DBF992FBF8F9F5" box="[959,1043,1581,1607]" captionStart="Fig" captionStartId="4.[152,185,1511,1533]" captionTargetBox="[156,1531,170,1482]" captionTargetId="figure-241@4.[177,1531,160,1487]" captionTargetPageId="4" captionText="Fig. 2. Holotype of the beaked whale Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A1), right lateral (A2), and ventral (A3) views." figureDoi="http://doi.org/10.5281/zenodo.12198768" httpUri="https://zenodo.org/record/12198768/files/figure.png" pageId="4" pageNumber="481">Fig. 2A</figureCitation>
|
||||
<figureCitation id="13032A1B1E05FA3A5F70F985FBF6F9F9" box="[1044,1053,1594,1611]" captionStart="Fig" captionStartId="2.[152,185,778,800]" captionTargetBox="[258,1428,162,756]" captionTargetId="figure-509@2.[634,1343,161,737]" captionTargetPageId="2" captionText="Fig. 1. A. Schematic map providing the position of Pisagua in South America. B. Schematic map of Northern Chile coast around Pisagua and sea floor bathymetry showing approximative discovery locality of the holotype MUAP(MM)-068 cranium of the beaked whale Ihlengesi changoensis sp. nov. at a depth of 1000 m (star)." figureDoi="http://doi.org/10.5281/zenodo.12198766" httpUri="https://zenodo.org/record/12198766/files/figure.png" pageId="4" pageNumber="481">
|
||||
<subScript id="17BC34DB1E05FA3A5F70F985FBF6F9F9" attach="right" box="[1044,1053,1594,1611]" fontSize="7" pageId="4" pageNumber="481">1</subScript>
|
||||
</figureCitation>
|
||||
), a feature that, at least in the extant ziphiids, is only observed in adult males (Bianucci et al. 2016b). The vomer greatly inflates towards the posterior portion of the rostrum as in the
|
||||
<typeStatus id="5483883C1E05FA3A5F2FF932FB45F915" box="[1099,1198,1677,1703]" pageId="4" pageNumber="481" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C384D1D1E05FA3A5FB5F932FC20F975" authority="(SAM PQ 2792)" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="481" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E05FA3A5FB5F932FA58F915" box="[1233,1459,1677,1703]" italics="true" pageId="4" pageNumber="481">Ihlengesi saldanhae</emphasis>
|
||||
(SAM PQ 2792)
|
||||
</taxonomicName>
|
||||
. A median longitudinal suture between the lateral walls of the vomer in the rostrum base area is visible as in all hyperoodontines.
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E05FA3A58E0F8B1FA14F8D5" blockId="4.[862,1536,1581,1960]" pageId="4" pageNumber="481">
|
||||
In ventral view the rostrum does not show a distinct alveolar groove whereas a clear maxilla-palatine suture extends about
|
||||
<quantity id="4CC09B7B1E05FA3A5880F8F2FBC3F8DA" box="[996,1064,1869,1896]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="481" unit="cm" value="12.0">12 cm</quantity>
|
||||
anterior to the antorbital notch (
|
||||
<figureCitation id="13032A1B1E05FA3A5EF4F8F2FA0EF8D5" box="[1424,1509,1869,1895]" captionStart="Fig" captionStartId="4.[152,185,1511,1533]" captionTargetBox="[156,1531,170,1482]" captionTargetId="figure-241@4.[177,1531,160,1487]" captionTargetPageId="4" captionText="Fig. 2. Holotype of the beaked whale Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A1), right lateral (A2), and ventral (A3) views." figureDoi="http://doi.org/10.5281/zenodo.12198768" httpUri="https://zenodo.org/record/12198768/files/figure.png" pageId="4" pageNumber="481">Fig. 2A</figureCitation>
|
||||
<figureCitation id="13032A1B1E05FA3A5E82F8E5FA04F8D9" box="[1510,1519,1882,1899]" captionStart="Fig" captionStartId="5.[99,132,1696,1718]" captionTargetBox="[208,1371,159,1672]" captionTargetId="figure-103@5.[210,1371,160,1640]" captionTargetPageId="5" captionText="Fig.3. Comparison of the neurocrania and the posterior portion of the rostra in dorsal view in two beaked whales.A. The holotype of Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.B. The holotype of Ihlengesi saldanhae (SAM PQ 2792) from the sea floor off Saldanha Bay, South Africa; age unknown." figureDoi="http://doi.org/10.5281/zenodo.12198770" httpUri="https://zenodo.org/record/12198770/files/figure.png" pageId="4" pageNumber="481">
|
||||
<subScript id="17BC34DB1E05FA3A5E82F8E5FA04F8D9" attach="right" box="[1510,1519,1882,1899]" fontSize="7" pageId="4" pageNumber="481">3</subScript>
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E05FA3858E0F8D2FE67FADE" blockId="4.[862,1536,1581,1960]" lastBlockId="6.[151,824,1328,1960]" lastPageId="6" lastPageNumber="483" pageId="4" pageNumber="481">
|
||||
Medial to the narrow antorbital notch, a distinct maxillary tubercle is visible. The position the suture between the maxilla and the missing lacrimojugal complex, well visible in the ventral surface of the left side of the neurocranium, supports that this tip, apparently laterally located to the rostrum base, is actually the maxillary tubercle. Indeed, this suture is located well lateral to the lateral margin of the rostrum, leaving exposed ventrally a wide portion of the maxilla that anteriorly bears what we interpret as maxillary tubercle. The presence of a well-defined maxillary tubercle separated from the lateral margin of the rostrum by a distinct prominental notch is a feature shared by most hyperoodontines including
|
||||
<taxonomicName id="4C384D1D1E07FA385AB7FA8FFD53FAF8" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[467,696,1328,1354]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385AB7FA8FFD53FAF8" box="[467,696,1328,1354]" italics="true" pageId="6" pageNumber="483">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
but absent in
|
||||
<taxonomicName id="4C384D1D1E07FA385BD1FAEEFE67FADE" box="[181,396,1361,1388]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E07FA385BD1FAEEFEB3FAD9" box="[181,344,1361,1387]" italics="true" pageId="6" pageNumber="483">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
<caption id="DF4766161E04FA3B5B07F91FFD44F95C" ID-DOI="http://doi.org/10.5281/zenodo.12198770" ID-Zenodo-Dep="12198770" httpUri="https://zenodo.org/record/12198770/files/figure.png" pageId="5" pageNumber="482" startId="5.[99,132,1696,1718]" targetBox="[208,1371,159,1672]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8B87369E1E04FA3B5B07F91FFD44F95C" blockId="5.[97,1481,1696,1774]" pageId="5" pageNumber="482">
|
||||
Fig. 3. Comparison of the neurocrania and the posterior portion of the rostra in dorsal view in two beaked whales.
|
||||
<emphasis id="B94CEA8C1E04FA3B5F06F91FFB99F907" bold="true" box="[1122,1138,1696,1717]" pageId="5" pageNumber="482">A</emphasis>
|
||||
. The holotype of
|
||||
<taxonomicName id="4C384D1D1E04FA3B5E75F91FFF92F960" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="482" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E04FA3B5E75F91FFF92F960" italics="true" pageId="5" pageNumber="482">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E04FA3B5B1BF903FF2EF963" box="[127,197,1724,1745]" pageId="5" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.
|
||||
<emphasis id="B94CEA8C1E04FA3B58ADF903FC33F963" bold="true" box="[969,984,1724,1745]" pageId="5" pageNumber="482">B</emphasis>
|
||||
. The holotype of
|
||||
<taxonomicName id="4C384D1D1E04FA3B5F1DF903FA22F963" authority="(SAM PQ 2792)" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1145,1481,1724,1745]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="482" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E04FA3B5F1DF903FAC5F963" box="[1145,1326,1724,1745]" italics="true" pageId="5" pageNumber="482">Ihlengesi saldanhae</emphasis>
|
||||
(SAM PQ 2792)
|
||||
</taxonomicName>
|
||||
from the sea floor off Saldanha Bay, South Africa; age unknown.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4766161E07FA385BFCFB18FDE4FAA3" ID-DOI="http://doi.org/10.5281/zenodo.12198772" ID-Zenodo-Dep="12198772" httpUri="https://zenodo.org/record/12198772/files/figure.png" pageId="6" pageNumber="483" startId="6.[152,185,1191,1213]" targetBox="[155,1529,161,1169]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8B87369E1E07FA385BFCFB18FDE4FAA3" blockId="6.[151,1536,1191,1297]" pageId="6" pageNumber="483">
|
||||
Fig. 4. Comparison of the crania in anterior view of the holotype of three beaked whales.
|
||||
<emphasis id="B94CEA8C1E07FA3858A1FB18FC3EFB0E" bold="true" box="[965,981,1191,1212]" pageId="6" pageNumber="483">A</emphasis>
|
||||
.
|
||||
<taxonomicName id="4C384D1D1E07FA385884FB18FB43FB0E" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[992,1192,1191,1212]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E07FA385884FB18FB43FB0E" box="[992,1192,1191,1212]" italics="true" pageId="6" pageNumber="483">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E07FA385FCAFB17FB1EFB0F" box="[1198,1269,1192,1213]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.
|
||||
<emphasis id="B94CEA8C1E07FA3859FEFB7BFD42FB6B" bold="true" box="[666,681,1220,1241]" pageId="6" pageNumber="483">B</emphasis>
|
||||
. The holotype of
|
||||
<taxonomicName id="4C384D1D1E07FA385834FB7CFB4CFB6B" authority="(SAM PQ 2792)" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[848,1191,1219,1241]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385834FB7CFBECFB6A" box="[848,1031,1219,1240]" italics="true" pageId="6" pageNumber="483">Ihlengesi saldanhae</emphasis>
|
||||
(SAM PQ 2792)
|
||||
</taxonomicName>
|
||||
from the sea floor off Saldanha Bay, South Africa; age unknown.
|
||||
<emphasis id="B94CEA8C1E07FA385AFEFB60FE41FB47" bold="true" box="[410,426,1247,1269]" pageId="6" pageNumber="483">C</emphasis>
|
||||
. The holotype
|
||||
<taxonomicName id="4C384D1D1E07FA385957FB60FC47FB47" authority="(SAM PQ 2678)" baseAuthorityName="SAM PQ" baseAuthorityYear="2678" box="[563,940,1247,1269]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="agulhasis">
|
||||
<emphasis id="B94CEA8C1E07FA385957FB60FCFAFB46" box="[563,785,1247,1268]" italics="true" pageId="6" pageNumber="483">Khoikhoicetus agulhasis</emphasis>
|
||||
(SAM PQ 2678)
|
||||
</taxonomicName>
|
||||
from the sea floor off Cape Agulhas, South Africa; age unknown. Diagonal lines represent broken surfaces.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B87369E1E07FA385BD9FACCFEC5F987" blockId="6.[151,824,1328,1960]" pageId="6" pageNumber="483">
|
||||
The prominental notch of the
|
||||
<taxonomicName id="4C384D1D1E07FA385974FACCFCE0FA3F" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[528,779,1395,1421]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E07FA385974FACCFCE0FA3F" box="[528,779,1395,1421]" italics="true" pageId="6" pageNumber="483">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="5483883C1E07FA385876FACCFF34FA1D" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
|
||||
is transversely wide, U-shaped and shallow as in the
|
||||
<typeStatus id="5483883C1E07FA385BA0FA08FECCFA63" box="[196,295,1463,1489]" pageId="6" pageNumber="483" type="paratype">paratype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C384D1D1E07FA385A30FA08FE09FA62" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[340,482,1462,1488]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385A30FA08FE09FA62" box="[340,482,1462,1488]" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
, whereas in the
|
||||
<taxonomicName id="4C384D1D1E07FA3859CDFA08FCD3FA62" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[681,824,1462,1488]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA3859CDFA08FCD3FA62" box="[681,824,1462,1488]" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="5483883C1E07FA385BF3FA67FF11FA40" box="[151,250,1496,1522]" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
|
||||
this notch is V-shaped and transversely shorter, suggesting that this character is subject to intraspecific variation (
|
||||
<figureCitation id="13032A1B1E07FA385BBAF9A4FECAF987" box="[222,289,1563,1589]" captionStart="Fig" captionStartId="5.[99,132,1696,1718]" captionTargetBox="[208,1371,159,1672]" captionTargetId="figure-103@5.[210,1371,160,1640]" captionTargetPageId="5" captionText="Fig.3. Comparison of the neurocrania and the posterior portion of the rostra in dorsal view in two beaked whales.A. The holotype of Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.B. The holotype of Ihlengesi saldanhae (SAM PQ 2792) from the sea floor off Saldanha Bay, South Africa; age unknown." figureDoi="http://doi.org/10.5281/zenodo.12198770" httpUri="https://zenodo.org/record/12198770/files/figure.png" pageId="6" pageNumber="483">Fig. 3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E07FA385BD9F982FDF9F96F" blockId="6.[151,824,1328,1960]" pageId="6" pageNumber="483">
|
||||
The dorsal surface of the preorbital area is roughly flat without a distinctive maxillary crest, as in
|
||||
<taxonomicName id="4C384D1D1E07FA3859ACF9E0FF2BF928" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA3859ACF9E0FF2BF928" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
, while in most of the other crown
|
||||
<taxonomicName id="4C384D1D1E07FA385936F93FFD2AF928" authorityName="Gray" authorityYear="1850" box="[594,705,1664,1690]" family="Ziphiidae" pageId="6" pageNumber="483" rank="family">Ziphiidae</taxonomicName>
|
||||
, including
|
||||
<taxonomicName id="4C384D1D1E07FA385BF3F91DFED2F90E" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[151,313,1698,1724]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E07FA385BF3F91DFED2F90E" box="[151,313,1698,1724]" italics="true" pageId="6" pageNumber="483">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
, the maxillary crest is present and generally elevated to form a peculiar dome.
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E07FA385BD9F95AFEB7F81A" blockId="6.[151,824,1328,1960]" pageId="6" pageNumber="483">
|
||||
Both the right and the left maxilla are pierced by a single dorsal infraorbital foramen near the prominental notch, 13- and 7-mm posterior to the rostrum base, respectively. Similar dorsal infraorbital foramina are observed in
|
||||
<taxonomicName id="4C384D1D1E07FA385BF3F8D3FE97F834" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[151,380,1900,1926]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385BF3F8D3FE97F834" box="[151,380,1900,1926]" italics="true" pageId="6" pageNumber="483">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
but more posteriorly located in
|
||||
<taxonomicName id="4C384D1D1E07FA385990F8D2FF01F815" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385990F8D2FF01F815" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="5483883C1E07FA385B95F831FEB3F81A" box="[241,344,1934,1960]" pageId="6" pageNumber="483" type="paratype">paratype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E07FA3958E0FA8FFD74FCF7" blockId="6.[862,1535,1328,1960]" lastBlockId="7.[98,771,714,1960]" lastPageId="7" lastPageNumber="484" pageId="6" pageNumber="483">
|
||||
The right and the left premaxillary foramina are distinctly posterior to the rostrum base (
|
||||
<quantity id="4CC09B7B1E07FA385E67FAEEFAAEFADE" box="[1283,1349,1361,1388]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="483" unit="mm" value="5.0">5 mm</quantity>
|
||||
) but less than in
|
||||
<taxonomicName id="4C384D1D1E07FA38583AFACCFBABFA3F" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[862,1088,1395,1421]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA38583AFACCFBABFA3F" box="[862,1088,1395,1421]" italics="true" pageId="6" pageNumber="483">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
as well quantified by the ratio between the longitudinal distance between the right premaxillary foramen and the rostrum base and the width of rostrum base, being this ratio equal to
|
||||
<quantity id="4CC09B7B1E07FA385FD6FA67FB17FA40" box="[1202,1276,1496,1522]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="483" unit="in" value="0.1">0.10 in</quantity>
|
||||
<taxonomicName id="4C384D1D1E07FA385E60FA66FA4CFA40" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[1284,1447,1496,1522]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E07FA385E60FA66FA4CFA40" box="[1284,1447,1496,1522]" italics="true" pageId="6" pageNumber="483">I. changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
against 0.34–
|
||||
<quantity id="4CC09B7B1E07FA3858F9FA45FC1AF9A6" box="[925,1009,1530,1556]" metricMagnitude="-3" metricUnit="m" metricValue="8.128" pageId="6" pageNumber="483" unit="in" value="0.32">0.32 in</quantity>
|
||||
<emphasis id="B94CEA8C1E07FA38589AFA45FB7CF9A6" box="[1022,1175,1530,1556]" italics="true" pageId="6" pageNumber="483">
|
||||
<taxonomicName id="4C384D1D1E07FA38589AFA45FB78F9A6" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1022,1171,1530,1556]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">I. saldanhae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
This significant difference in the position of the premaxillary foramina is related to the greater anteroposterior extension of the premaxillary sac fossae in
|
||||
<taxonomicName id="4C384D1D1E07FA3858AFF9E0FB84F9CB" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[971,1135,1631,1657]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E07FA3858AFF9E0FB84F9CB" box="[971,1135,1631,1657]" italics="true" pageId="6" pageNumber="483">I. changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
compared to
|
||||
<emphasis id="B94CEA8C1E07FA385E6AF9E0FA49F9CB" box="[1294,1442,1631,1657]" italics="true" pageId="6" pageNumber="483">
|
||||
<taxonomicName id="4C384D1D1E07FA385E6AF9E0FA75F9CB" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1294,1438,1631,1657]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">I. saldanhae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Indeed, the premaxillary sac fossae of
|
||||
<taxonomicName id="4C384D1D1E07FA385FADF93EFAB3F928" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1225,1368,1664,1690]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA385FADF93EFAB3F928" box="[1225,1368,1664,1690]" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
are unusually short so that this feature was previously considered an apomorphy of the
|
||||
<taxonomicName id="4C384D1D1E07FA385F64F97BFA8BF96C" authority="(Bianucci et al. 2007)" baseAuthorityName="Bianucci" baseAuthorityYear="2007" box="[1024,1376,1732,1758]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E07FA385F64F97BFB8CF96C" box="[1024,1127,1732,1758]" italics="true" pageId="6" pageNumber="483">Ihlengesi</emphasis>
|
||||
(Bianucci et al. 2007)
|
||||
</taxonomicName>
|
||||
. Interestingly, in
|
||||
<taxonomicName id="4C384D1D1E07FA3858E5F959FBF8F94D" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[897,1043,1765,1791]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E07FA3858E5F959FBF8F94D" box="[897,1043,1765,1791]" italics="true" pageId="6" pageNumber="483">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
the premaxillary foramina are distinctly posterior to the dorsal infraorbital foramina, an unusual condition in beaked whales. The premaxillary foramina of
|
||||
<taxonomicName id="4C384D1D1E07FA38583AF8F4FBEEF8D6" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[862,1029,1866,1892]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E07FA38583AF8F4FBEEF8D6" box="[862,1029,1866,1892]" italics="true" pageId="6" pageNumber="483">I. changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
are posteriorly located to the dorsal infraorbital foramina but to a lesser degree than
|
||||
<emphasis id="B94CEA8C1E07FA385E0CF8D2FA14F834" box="[1384,1535,1900,1926]" italics="true" pageId="6" pageNumber="483">
|
||||
<taxonomicName id="4C384D1D1E07FA385E0CF8D2FA10F834" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1384,1531,1900,1926]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="483" phylum="Chordata" rank="species" species="saldanhae">I. saldanhae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Regardless the position of the foramina with respect to the rostrum base, in all hyperoodontines, except some species of
|
||||
<taxonomicName id="4C384D1D1E06FA395BE7FD54FEE5FCB7" authorityName="Gervais" authorityYear="1850" box="[131,270,747,773]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395BE7FD54FEE5FCB7" box="[131,270,747,773]" italics="true" pageId="7" pageNumber="484">Mesoplodon</emphasis>
|
||||
</taxonomicName>
|
||||
(e.g.,
|
||||
<taxonomicName id="4C384D1D1E06FA395A3DFD54FD1CFCB6" authority="von Haast, 1876" authorityName="von Haast" authorityYear="1876" box="[345,759,746,773]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="grayi">
|
||||
<emphasis id="B94CEA8C1E06FA395A3DFD54FDC0FCB6" box="[345,555,747,773]" italics="true" pageId="7" pageNumber="484">Mesoplodon grayi</emphasis>
|
||||
<bibRefCitation id="EFA94B6F1E06FA395952FD54FD1CFCB6" author="von Haast" box="[566,759,746,773]" firstAuthor="von Haast" pageId="7" pageNumber="484" pagination="7 - 13" refId="ref11077" refString="Haast, J. von. 1876. On a new ziphioid whale. Proceedings of the Zoological Society, London 1876: 7 - 13." type="journal article" year="1876">von Haast, 1876</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), the premaxillary foramina are always roughly at the same level or weakly anterior to the maxillary foramina.
|
||||
</paragraph>
|
||||
<caption id="DF4766161E06FA395B06FD98FD1EFD1F" ID-DOI="http://doi.org/10.5281/zenodo.12198774" ID-Zenodo-Dep="12198774" httpUri="https://zenodo.org/record/12198774/files/figure.png" pageId="7" pageNumber="484" startId="7.[98,131,551,573]" targetBox="[98,768,159,527]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8B87369E1E06FA395B06FD98FD1EFD1F" blockId="7.[98,773,551,685]" pageId="7" pageNumber="484">
|
||||
Fig. 5. Comparison of the rostra and posterior portion of the neurocrania in dorsal view of two beaked whales.
|
||||
<emphasis id="B94CEA8C1E06FA395ACDFDFCFE52FDEA" bold="true" box="[425,441,579,600]" pageId="7" pageNumber="484">A</emphasis>
|
||||
. The holotype of
|
||||
<taxonomicName id="4C384D1D1E06FA395904FDFCFF64FDC7" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E06FA395904FDFCFF64FDC7" italics="true" pageId="7" pageNumber="484">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E06FA395BF1FDDFFF37FDC7" box="[149,220,608,629]" pageId="7" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene.
|
||||
<emphasis id="B94CEA8C1E06FA395A59FDC3FEA7FD23" bold="true" box="[317,332,636,657]" pageId="7" pageNumber="484">B</emphasis>
|
||||
. The paratype of
|
||||
<taxonomicName id="4C384D1D1E06FA395A89FDC4FD48FD22" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[493,675,635,656]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395A89FDC4FD48FD22" box="[493,675,635,656]" italics="true" pageId="7" pageNumber="484">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
(SAM PQ 69673) from the sea floor off Saldanha Bay, South Africa; age unknown.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B87369E1E06FA395BE3FCF4FEE8FC57" blockId="7.[98,771,714,1960]" pageId="7" pageNumber="484">
|
||||
The weakly concave premaxillary sac fossae of
|
||||
<taxonomicName id="4C384D1D1E06FA3959CEFCF3FF5CFC37" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E06FA3959CEFCF3FF5CFC37" italics="true" pageId="7" pageNumber="484">I. changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
are distinctly asymmetrical as in
|
||||
<emphasis id="B94CEA8C1E06FA395925FCD3FD3EFC37" box="[577,725,875,901]" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA395925FCD3FD3AFC37" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[577,721,875,901]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">I. saldanhae</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
being the right fossa transversally wider than the left fossa ratio between the widths of the left and right premaxillary fossae = 0.78).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA395BE3FC54FE3AFB74" blockId="7.[98,771,714,1960]" pageId="7" pageNumber="484">
|
||||
The ascending process of the premaxilla rises towards the vertex with its posterodorsal portion partly overhanging the bony nares (
|
||||
<figureCitation id="13032A1B1E06FA395A23FB94FE43FBFB" box="[327,424,1067,1097]" captionStart="Fig" captionStartId="4.[152,185,1511,1533]" captionTargetBox="[156,1531,170,1482]" captionTargetId="figure-241@4.[177,1531,160,1487]" captionTargetPageId="4" captionText="Fig. 2. Holotype of the beaked whale Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. Cranium in dorsal (A1), right lateral (A2), and ventral (A3) views." figureDoi="http://doi.org/10.5281/zenodo.12198768" httpUri="https://zenodo.org/record/12198768/files/figure.png" pageId="7" pageNumber="484">
|
||||
Fig. 2A
|
||||
<subScript id="17BC34DB1E06FA395AFBFB87FE43FBFB" attach="right" box="[415,424,1080,1097]" fontSize="7" pageId="7" pageNumber="484">2</subScript>
|
||||
</figureCitation>
|
||||
). Therefore, with the cranium in lateral view, the anterior profile of this portion of the ascending process of the premaxilla is clearly concave in
|
||||
<taxonomicName id="4C384D1D1E06FA395B06FB34FEB4FB17" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[98,351,1163,1189]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E06FA395B06FB34FEB4FB17" box="[98,351,1163,1189]" italics="true" pageId="7" pageNumber="484">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
as in
|
||||
<taxonomicName id="4C384D1D1E06FA395AC8FB33FDD0FB17" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[428,571,1163,1189]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395AC8FB33FDD0FB17" box="[428,571,1163,1189]" italics="true" pageId="7" pageNumber="484">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
, not straight and vertical as in
|
||||
<taxonomicName id="4C384D1D1E06FA395B9EFB14FE3AFB74" box="[250,465,1195,1222]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA395B9EFB14FE76FB77" box="[250,413,1195,1221]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA395BE3FB73FD70FA34" blockId="7.[98,771,714,1960]" pageId="7" pageNumber="484">
|
||||
The vertex is moderately elevated as in
|
||||
<taxonomicName id="4C384D1D1E06FA395902FB73FF5EFAB4" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395902FB73FF5EFAB4" italics="true" pageId="7" pageNumber="484">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C384D1D1E06FA395B95FB53FE20FAB4" box="[241,459,1260,1286]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA395B95FB53FE7FFAB4" box="[241,404,1260,1286]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
(ratio between the vertical distance between the dorsal margin of the maxilla at the rostrum base and the top of the vertex and the width of the premaxillary sac fossae between 0.70 and 1.0). All other hyperoodontines have a higher vertex (ratio> 1.0).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA395BE3FA33FE64F995" blockId="7.[98,771,714,1960]" pageId="7" pageNumber="484">The ascending process of the premaxilla displays a strong constriction between premaxillary sac fossa and premaxillary crest (ratio between the minimal width of ascending process of the right premaxilla and the width of right premaxillary crest = 0.56).</paragraph>
|
||||
<paragraph id="8B87369E1E06FA395BE3F992FD9FF8F5" blockId="7.[98,771,714,1960]" pageId="7" pageNumber="484">
|
||||
As in
|
||||
<taxonomicName id="4C384D1D1E06FA395BAFF993FE45F9F4" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[203,430,1580,1606]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395BAFF993FE45F9F4" box="[203,430,1580,1606]" italics="true" pageId="7" pageNumber="484">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<emphasis id="B94CEA8C1E06FA395A81F993FCE9F9F4" box="[485,770,1580,1606]" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA395A81F993FD15F9F4" baseAuthorityName="SAM PQ" baseAuthorityYear="2678" box="[485,766,1580,1606]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="agulhasis">Khoikhoicetus agulhasis</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
the posterolaterally directed premaxillary crests are transversally narrower than in the other hyperoodontines (ratio between the transverse width of the premaxillary crests and the width of premaxillary sac fossae = ca. 1.0), whereas the distance between the premaxillary crests is greater than in
|
||||
<taxonomicName id="4C384D1D1E06FA395B06F952FF06F8B5" authorityName="Gervais" authorityYear="1850" box="[98,237,1773,1799]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395B06F952FF06F8B5" box="[98,237,1773,1799]" italics="true" pageId="7" pageNumber="484">Mesoplodon</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C384D1D1E06FA395A42F952FE58F8B5" authorityName="Lacepede" authorityYear="1804" box="[294,435,1773,1799]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395A42F952FE58F8B5" box="[294,435,1773,1799]" italics="true" pageId="7" pageNumber="484">Hyperoodon</emphasis>
|
||||
</taxonomicName>
|
||||
(ratio between the minimum distance between the right and left premaxillary crests and the width of the premaxillary sac fossae = 0.3).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA395BE3F8F2FB28FF4B" blockId="7.[98,771,714,1960]" lastBlockId="7.[808,1482,158,832]" pageId="7" pageNumber="484">
|
||||
The dorsal margin of each premaxillary crest slopes markedly ventrolaterally generating an acute dorsal profile of the vertex in anterior view, similarly to species of
|
||||
<emphasis id="B94CEA8C1E06FA39584CFF21FBBBFF0A" box="[808,1104,158,184]" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFF21FC5DFF0A" authorityName="Gervais" authorityYear="1850" box="[808,950,158,184]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">Mesoplodon</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C384D1D1E06FA3958A7FF21FBBBFF0A" authorityName="Lacepede" authorityYear="1804" box="[963,1104,158,184]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">Hyperoodon</taxonomicName>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C384D1D1E06FA395F3BFF21FAE9FF0A" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[1119,1282,158,184]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395F3BFF21FAE9FF0A" box="[1119,1282,158,184]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
but not
|
||||
<emphasis id="B94CEA8C1E06FA395E06FF21FC4EFF6A" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA395E06FF21FC4BFF6A" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">Ihlengesi saldanhae</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
this latter having premaxillary crests with a rounded dorsolateral profile (
|
||||
<figureCitation id="13032A1B1E06FA395F16FF60FB5DFF4B" box="[1138,1206,223,249]" captionStart="Fig" captionStartId="6.[152,185,1191,1213]" captionTargetBox="[155,1529,161,1169]" captionTargetId="figure-340@6.[189,1529,169,1160]" captionTargetPageId="6" captionText="Fig. 4. Comparison of the crania in anterior view of the holotype of three beaked whales. A. Ihlengesi changoensis sp. nov. (MUAP(MM)-068) from the sea floor off Pisagua, Northern Chile; Plio-Pleistocene. B. The holotype of Ihlengesi saldanhae (SAM PQ 2792) from the sea floor off Saldanha Bay, South Africa; age unknown. C. The holotype Khoikhoicetus agulhasis (SAM PQ 2678) from the sea floor off Cape Agulhas, South Africa; age unknown. Diagonal lines represent broken surfaces." figureDoi="http://doi.org/10.5281/zenodo.12198772" httpUri="https://zenodo.org/record/12198772/files/figure.png" pageId="7" pageNumber="484">Fig. 4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA39582AFF40FC4DFD0C" blockId="7.[808,1482,158,832]" pageId="7" pageNumber="484">
|
||||
The elongated nasals (ratio between the length of medial suture of nasals on vertex and the maximum width of nasals = 1.1) show a deep anteromedial excavation on their joined dorsal surface as in the other hyperoodontines but
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFE3EFB16FE29" authority="(Longman, 1926)" baseAuthorityName="Longman" baseAuthorityYear="1926" box="[808,1277,385,411]" class="Mammalia" family="Hyperoodontidae" genus="Indopacetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="pacificus">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFE3EFBCDFE29" box="[808,1062,385,411]" italics="true" pageId="7" pageNumber="484">Indopacetus pacificus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFA94B6F1E06FA395F5DFE3EFB1EFE29" author="Longman" box="[1081,1269,385,411]" firstAuthor="Longman" pageId="7" pageNumber="484" pagination="266 - 278" refId="ref11975" refString="Longman, H. A. 1926. New records of Cetacea, with a list of Queensland species. Memoirs of the Queensland Museum 8: 266 - 278." type="journal article" year="1926">Longman, 1926</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Due to a significant anterolateral expansion of the nasals forming part of the premaxillary crests, a deep notch is visible on the dorsal margin of each nasal, a feature not discernible in
|
||||
<taxonomicName id="4C384D1D1E06FA395E06FE5DFC77FDAE" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395E06FE5DFC77FDAE" italics="true" pageId="7" pageNumber="484">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="5483883C1E06FA3958C1FDBDFBE3FDAE" box="[933,1032,514,540]" pageId="7" pageNumber="484" type="holotype">holotype</typeStatus>
|
||||
due to the bad preservation of the vertex. The inclusion of the nasals in the premaxillary crests is significant, although less for the right than for the left nasal, a condition shared with
|
||||
<taxonomicName id="4C384D1D1E06FA395F53FDDCFAE5FDCC" box="[1079,1294,611,638]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA395F53FDDCFB31FDCF" box="[1079,1242,611,637]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
rather than with
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFD3AFC5FFD2C" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[808,948,644,670]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFD3AFC5FFD2C" box="[808,948,644,670]" italics="true" pageId="7" pageNumber="484">I. saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
having a greater anterolateral expansion of the right nasal.
|
||||
</paragraph>
|
||||
<paragraph id="8B87369E1E06FA39582AFD7AFA21FCF2" blockId="7.[808,1482,158,832]" pageId="7" pageNumber="484">
|
||||
The lateral margins of the nasals are weakly convex as in
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFD5AFBEBFCB2" box="[808,1024,741,768]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFD5AFC2CFD4D" box="[808,967,741,767]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
, not anteriorly diverging as in
|
||||
<emphasis id="B94CEA8C1E06FA395E00FD5AFC49FCAD" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA395E00FD5AFC75FCAD" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">Ihlengesi saldanhae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
The nasofrontal suture is anteriorly pointed as in
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFC99FC17FCF2" box="[808,1020,806,832]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFC99FC2CFCF2" box="[808,967,806,832]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
, not convex posteriorly as in
|
||||
<emphasis id="B94CEA8C1E06FA395E5EFC98FA21FCF2" box="[1338,1482,806,832]" italics="true" pageId="7" pageNumber="484">
|
||||
<taxonomicName id="4C384D1D1E06FA395E5EFC98FA2DFCF2" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1338,1478,806,832]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">I. saldanhae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E06FA39584CFCEEFAB4FAC3" pageId="7" pageNumber="484" type="discussion">
|
||||
<paragraph id="8B87369E1E06FA39584CFCEEFAB4FAC3" blockId="7.[808,1482,849,1393]" pageId="7" pageNumber="484">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFCEEFC61FCD9" box="[808,906,849,875]" italics="true" pageId="7" pageNumber="484">Remarks</emphasis>
|
||||
.—All characters listed in the diagnosis separating
|
||||
<taxonomicName id="4C384D1D1E06FA39584CFCCEFBC8FC39" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[808,1059,881,907]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="changoensis">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFCCEFBC8FC39" box="[808,1059,881,907]" italics="true" pageId="7" pageNumber="484">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<taxonomicName id="4C384D1D1E06FA395F0EFCCEFAA6FC39" baseAuthorityName="SAM PQ" baseAuthorityYear="2792" box="[1130,1357,881,907]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E06FA395F0EFCCEFAA6FC39" box="[1130,1357,881,907]" italics="true" pageId="7" pageNumber="484">Ihlengesi saldanhae</emphasis>
|
||||
</taxonomicName>
|
||||
are shared with
|
||||
<taxonomicName id="4C384D1D1E06FA395801FC2DFBD4FC1E" box="[869,1087,914,940]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B94CEA8C1E06FA395801FC2DFBE3FC1E" box="[869,1032,914,940]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
Indeed
|
||||
<taxonomicName id="4C384D1D1E06FA395FC4FC2DFAA8FC1E" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[1184,1347,914,940]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395FC4FC2DFAA8FC1E" box="[1184,1347,914,940]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
is a hyperoodontine with a cranium roughly similar to
|
||||
<taxonomicName id="4C384D1D1E06FA395E4DFC0DFA7BFC7E" box="[1321,1424,946,972]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395E4DFC0DFA7BFC7E" box="[1321,1424,946,972]" italics="true" pageId="7" pageNumber="484">Ihlengesi</emphasis>
|
||||
</taxonomicName>
|
||||
(i.e., for the size and for the moderate elevation of the vertex). Nevertheless
|
||||
<taxonomicName id="4C384D1D1E06FA3958DBFC4CFB89FBBF" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[959,1122,1011,1037]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA3958DBFC4CFB89FBBF" box="[959,1122,1011,1037]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
does not show many of the character above listed of the genus
|
||||
<taxonomicName id="4C384D1D1E06FA395FF4FBACFB1CFB9F" box="[1168,1271,1043,1069]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395FF4FBACFB1CFB9F" box="[1168,1271,1043,1069]" italics="true" pageId="7" pageNumber="484">Ihlengesi</emphasis>
|
||||
</taxonomicName>
|
||||
(i.e., rostrum narrow in dorsal view, presence of a distinct maxillary tubercle and a wide and shallow prominental notch anteromedial to the antorbital notch; premaxillary foramen distinctly posterior to the antorbital notch, roughly flat dorsal surface of the preorbital area without distinct maxillary crest; ascending process of the premaxilla in lateral view concave with posterodorsal portion partly overhanging the bony nares) and therefore we consider reliable the appurtenance of the new Chilean species to
|
||||
<taxonomicName id="4C384D1D1E06FA395F55FA88FB73FAE3" box="[1073,1176,1335,1361]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395F55FA88FB73FAE3" box="[1073,1176,1335,1361]" italics="true" pageId="7" pageNumber="484">Ihlengesi</emphasis>
|
||||
</taxonomicName>
|
||||
rather than
|
||||
<taxonomicName id="4C384D1D1E06FA395E44FA88FA29FAE3" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[1312,1474,1335,1361]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="484" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E06FA395E44FA88FA29FAE3" box="[1312,1474,1335,1361]" italics="true" pageId="7" pageNumber="484">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
, being also supported by the phylogenetic analysis.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E06FA39584CFA3DFC55FA0F" pageId="7" pageNumber="484" type="distribution">
|
||||
<paragraph id="8B87369E1E06FA39584CFA3DFC55FA0F" blockId="7.[808,1481,1410,1469]" pageId="7" pageNumber="484">
|
||||
<emphasis id="B94CEA8C1E06FA39584CFA3DFB3DFA2E" box="[808,1238,1410,1436]" italics="true" pageId="7" pageNumber="484">Stratigraphic and geographic range</emphasis>
|
||||
.—
|
||||
<typeStatus id="5483883C1E06FA395F92FA3CFADBFA2F" box="[1270,1328,1411,1437]" pageId="7" pageNumber="484">Type</typeStatus>
|
||||
locality and horizon only.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
125
data/03/91/87/039187881E03FA3D583AF922FDE7FD12.xml
Normal file
125
data/03/91/87/039187881E03FA3D583AF922FDE7FD12.xml
Normal file
|
|
@ -0,0 +1,125 @@
|
|||
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<mods:titleInfo id="9C5BFB77DC9BDCC34AABF15A7219D0BC">
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<mods:title id="7900BC5C4CB2433C0CC82BF3F6BEB6E7">A new diminutive fossil ziphiid from the deep-sea floor off northern Chile and some remarks on the body size evolution and palaeobiogeography of the beaked whales</mods:title>
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<mods:namePart id="F5E9694AA7D01AADF39B7D68C679C901">Sielfeld, Walter</mods:namePart>
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<mods:namePart id="482718711E9F76F07F6BE78FE17ED44E">Olguin, Nicole A.</mods:namePart>
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<mods:namePart id="DBA2AF7B7C2F541361F853E209BC213A">Guzmán, Guillermo</mods:namePart>
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|
||||
<mods:identifier id="8BFDD1A7C61F436A4FF8AE5E6579C834" type="ISSN">1732-2421</mods:identifier>
|
||||
<mods:identifier id="AA5DB0E27264877E33305ACE054233AC" type="Zenodo-Dep">12198764</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="039187881E03FA3D583AF922FDE7FD12" LSID="urn:lsid:plazi:treatment:039187881E03FA3D583AF922FDE7FD12" httpUri="http://treatment.plazi.org/id/039187881E03FA3D583AF922FDE7FD12" lastPageId="3" lastPageNumber="479" pageId="2" pageNumber="479">
|
||||
<subSubSection id="C32265151E03FA3C583AF922FA1BF908" box="[862,1520,1693,1722]" pageId="2" pageNumber="479" type="nomenclature">
|
||||
<paragraph id="8B87369E1E03FA3C583AF922FA1BF908" blockId="2.[862,1520,1489,1722]" box="[862,1520,1693,1722]" pageId="2" pageNumber="479">
|
||||
Genus
|
||||
<taxonomicName id="4C384D1D1E03FA3C58D3F922FA1BF908" authority="Bianucci, Lambert, and Post, 2007" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[951,1520,1693,1722]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="479" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E03FA3C58D3F922FBC7F908" box="[951,1068,1693,1722]" italics="true" pageId="2" pageNumber="479">Ihlengesi</emphasis>
|
||||
Bianucci, Lambert, and Post, 2007
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E03FA3C583AF975FBDBF8AB" pageId="2" pageNumber="479" type="type_taxon">
|
||||
<paragraph id="8B87369E1E03FA3C583AF975FBDBF8AB" blockId="2.[862,1535,1738,1851]" pageId="2" pageNumber="479">
|
||||
<emphasis id="B94CEA8C1E03FA3C583AF975FC3DF96D" box="[862,982,1738,1759]" italics="true" pageId="2" pageNumber="479">
|
||||
<typeStatus id="5483883C1E03FA3C583AF975FC60F96D" box="[862,907,1738,1759]" pageId="2" pageNumber="479">Type</typeStatus>
|
||||
species
|
||||
</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C384D1D1E03FA3C5887F975FA10F96D" authority="Bianucci, Lambert, and Post, 2007" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[995,1531,1738,1760]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="479" phylum="Chordata" rank="species" species="saldanhae">
|
||||
<emphasis id="B94CEA8C1E03FA3C5887F975FB4EF96D" box="[995,1189,1738,1759]" italics="true" pageId="2" pageNumber="479">Ihlengesi saldanhae</emphasis>
|
||||
Bianucci, Lambert, and Post, 2007
|
||||
</taxonomicName>
|
||||
, by original designation; from the sea floor off
|
||||
<location id="8EE760451E03FA3C5E56F959FA52F94E" LSID="urn:lsid:plazi:treatment:039187881E03FA3D583AF922FDE7FD12:8EE760451E03FA3C5E56F959FA52F94E" box="[1330,1465,1766,1788]" country="South Africa" name="Saldanha Bay" pageId="2" pageNumber="479">Saldanha Bay</location>
|
||||
,
|
||||
<collectingCountry id="F32F760E1E03FA3C5EA2F959FC75F8AB" name="South Africa" pageId="2" pageNumber="479">South Africa</collectingCountry>
|
||||
; age unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E03FA3C583AF89AFA31F888" box="[862,1498,1829,1851]" pageId="2" pageNumber="479" type="taxon_list">
|
||||
<paragraph id="8B87369E1E03FA3C583AF89AFA31F888" blockId="2.[862,1535,1738,1851]" box="[862,1498,1829,1851]" pageId="2" pageNumber="479">
|
||||
<emphasis id="B94CEA8C1E03FA3C583AF89AFC14F888" box="[862,1023,1829,1850]" italics="true" pageId="2" pageNumber="479">Species included</emphasis>
|
||||
:
|
||||
<typeStatus id="5483883C1E03FA3C5F6FF89AFBD0F888" box="[1035,1083,1829,1850]" pageId="2" pageNumber="479">Type</typeStatus>
|
||||
species and
|
||||
<taxonomicName id="4C384D1D1E03FA3C5FD0F89AFA62F888" authority="Bianucci & Sielfeld & Olguin & Guzmán, 2023" authorityName="Bianucci & Sielfeld & Olguin & Guzmán" authorityYear="2023" box="[1204,1417,1829,1850]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="479" phylum="Chordata" rank="species" species="changoensis" status="sp. nov.">
|
||||
<emphasis id="B94CEA8C1E03FA3C5FD0F89AFA62F888" box="[1204,1417,1829,1850]" italics="true" pageId="2" pageNumber="479">Ihlengesi changoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27F57F71E03FA3C5EEBF89AFA31F888" box="[1423,1498,1829,1850]" pageId="2" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32265151E03FA3D583AF8F3FDE7FD12" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479" type="diagnosis">
|
||||
<paragraph id="8B87369E1E03FA3D583AF8F3FDE7FD12" blockId="2.[862,1535,1868,1960]" lastBlockId="3.[98,771,158,672]" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479">
|
||||
<emphasis id="B94CEA8C1E03FA3C583AF8F3FBDDF8D4" box="[862,1078,1868,1894]" italics="true" pageId="2" pageNumber="479">Emended diagnosis</emphasis>
|
||||
.—
|
||||
<taxonomicName id="4C384D1D1E03FA3C5F3DF8F3FB55F8D4" authorityName="Bianucci, Lambert, and Post" authorityYear="2007" box="[1113,1214,1868,1894]" class="Mammalia" family="Hyperoodontidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="479" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E03FA3C5F3DF8F3FB55F8D4" box="[1113,1214,1868,1894]" italics="true" pageId="2" pageNumber="479">Ihlengesi</emphasis>
|
||||
</taxonomicName>
|
||||
differs from all other
|
||||
<taxonomicName id="4C384D1D1E03FA3C5EDDF8F2FC08F835" authorityName="Gray" authorityYear="1866" class="Mammalia" family="Hyperoodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="479" phylum="Chordata" rank="subFamily" subFamily="Hyperoodontinae">Hyperoodontinae</taxonomicName>
|
||||
for the following unique combination of characters: narrow rostrum in dorsal view; presence of a distinct maxillary tubercle and a wide and shallow prominental notch anteromedial to the antorbital notch; dorsal infraorbital foramen close to the prominental notch at the rostrum base; premaxillary foramen distinctly posterior to the antorbital notch; roughly flat dorsal surface of the preorbital area without distinct maxillary crest; moderate elevation of the vertex less than in the other hyperoodontines but
|
||||
<taxonomicName id="4C384D1D1E02FA3D5931FEDEFD19FEC9" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[597,754,353,379]" class="Mammalia" family="Hyperoodontidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E02FA3D5931FEDEFD19FEC9" box="[597,754,353,379]" italics="true" pageId="3" pageNumber="480">Khoikhoicetus</emphasis>
|
||||
</taxonomicName>
|
||||
); ascending process of the premaxilla in lateral view concave with posterodorsal portion partly overhanging the bony nares; transverse distance between the lateral margins of the premaxillary crests less than in the other hyperoodontines; wide space between the premaxillary crests (greater than in
|
||||
<taxonomicName id="4C384D1D1E02FA3D5B06FD9BFF01FD8C" authorityName="Gervais" authorityYear="1850" box="[98,234,548,574]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E02FA3D5B06FD9BFF01FD8C" box="[98,234,548,574]" italics="true" pageId="3" pageNumber="480">Mesoplodon</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C384D1D1E02FA3D5A7BFD9BFE43FD8C" authorityName="Lacepede" authorityYear="1804" box="[287,424,548,574]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="480" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B94CEA8C1E02FA3D5A7BFD9BFE43FD8C" box="[287,424,548,574]" italics="true" pageId="3" pageNumber="480">Hyperoodon</emphasis>
|
||||
</taxonomicName>
|
||||
); inclusion of the nasal into the premaxillary crest with the at least the left nasal covering about half the length of the median margin of the crest; deep anteromedian excavation of the nasals.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue