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<document ID-CLB-Dataset="297097" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="f9373c90-73c5-4ee4-b0f8-4c371f527d91" ID-ISBN="0-8018-8221-4" checkinTime="1670905209323" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="A31A098593A6284D753C904FBCA94B7B" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Muridae.1189-1531.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Rattus adustus Sody 1940" docType="treatment" docVersion="4" id="A2267A57FF8A65DA07E1D2E970F9422A" lastPageNumber="1463" masterDocId="A2267A57FF8A65DA07E1D2E970F9422A" masterDocTitle="Order Rodentia - Family Muridae" masterLastPageNumber="1531" masterPageNumber="1189" pageNumber="1463" updateTime="1716781534040" updateUser="ExternalLinkService">
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<mods:title id="E7C3F037FD36B1A7F47B0B0557A23A97">Order Rodentia - Family Muridae</mods:title>
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<mods:title id="94BC77EF77CFD616CA6FE857EA26CD54">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<paragraph id="C4E7EF926010030CC9167E2E429C0A13" pageNumber="1463">
<taxonomicName id="9C95108EFB778350CD556DCA3D2E1FBB" ID-CoL="4RM3Q" authority="Sody 1940" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1463" phylum="Chordata" rank="species" species="adustus">
<emphasis id="B69D9E8557D8B16FE90C8A2C2835F51D" italics="true" pageNumber="1463">Rattus adustus</emphasis>
Sody 1940
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<paragraph id="6949F3ED7F6337C132554F9DB43B8F2E" pageNumber="1463">
<treatmentCitation id="FE5506FAFF6FB8F2A8E7AC3843D0B5E3" author="Sody" authority="Sody 1940" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" journal="Treubia" kingdom="Animalia" order="Rodentia" page="397" pageNumber="1463" phylum="Chordata" rank="species" species="adustus" title="Rattus adustus" volume="17" year="1940">
<taxonomicName id="7D7221BA36A0446F69435B292879BFF4" ID-CoL="4RM3Q" authority="Sody 1940" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1463" phylum="Chordata" rank="species" species="adustus">
<emphasis id="D78D2DCF563313D3E151BBB18260F0AE" italics="true" pageNumber="1463">Rattus adustus</emphasis>
<bibRefCitation id="7429ACB9DB45D44B5291829F001CB9C6" author="Sody, H. J. V." refId="ref332575" refString="Sody, H. J. V. 1940. On the mammals of Enggano. Treubia, 17: 391 - 405." year="1940">Sody 1940</bibRefCitation>
</taxonomicName>
,
<bibCitation id="0AB679445307830B8B891F751379FD67" author="Sody" journal="Treubia" pageNumber="1463" pagination="397" title="Rattus adustus" volume="17" year="1940">Treubia, 17: 397</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6D4ACAD7F4AEF7D2DDD09819F0D4973A" pageNumber="1463" type="materials_examined">
<paragraph id="17E9644299EFC645AD31412A24FB15D1" pageNumber="1463">
<emphasis id="91D4DD3CA40DD68EB27834E3B4B957FA" inLineHeading="true" pageNumber="1463">Type Locality:</emphasis>
<materialsCitation id="EDD4F6B91E27D8CA2335999D0A719275" ID-GBIF-Occurrence="4867324804" country="Indonesia" pageNumber="1463" stateProvince="Sumatera Utara">
<collectingCountry id="9A751BE833D1A038F5B5EC6C68A16C61" name="Indonesia">Indonesia</collectingCountry>
, Pulau Enggano, off the coast of W
<collectingRegion id="2EB4FBF9479AEAA33151CA3277AB2080" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
(and off the continental shelf), Kiojoh, sea level.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="456EF7A8EB34B2E66A8182A84FECECDD" pageNumber="1463" type="vernacular_names">
<paragraph id="0C71B8DB3FF03F17CAC83AD930387512" pageNumber="1463">
<emphasis id="272ED0FAB868CEEDE5F74E4B693DE2AB" inLineHeading="true" pageNumber="1463">Vernacular Names:</emphasis>
<vernacularName id="E5F3C4AEEB0BAC40A5D610350B207D57" language="eng" pageNumber="1463">Burnished Enggano Rat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="0895BA78F54CFE6EC26A68C541B13A42" pageNumber="1463" type="distribution">
<paragraph id="B51DF4CF20540F9D30A06E5F0A5DCDDC" pageNumber="1463">
<emphasis id="DCED4075664EEB0A17ACACB647720096" inLineHeading="true" pageNumber="1463">Distribution:</emphasis>
Pulau Enggano.
</paragraph>
</subSubSection>
<subSubSection id="E3E69834533BB29087D203A137AB0E77" pageNumber="1463" type="conservation">
<paragraph id="7D6D7A627672D6EA1188B69C77583A96" pageNumber="1463">
<emphasis id="9B5BCFDB5C23936488229D55568A08E4" inLineHeading="true" pageNumber="1463">Conservation:</emphasis>
<collectionCode id="A208CBA00E5ED1133FCBE85E6C1937AA">IUCN</collectionCode>
Vulnerable.
</paragraph>
</subSubSection>
<subSubSection id="FB23CF5FD087AAFFAF8164C47E20C1D4" pageNumber="1463" type="discussion">
<paragraph id="55CDFB7200A36A33506F779393A2FCBE" pageNumber="1463">
<emphasis id="77953438E6B65DDB339A73F1CA3BEF14" inLineHeading="true" pageNumber="1463">Discussion:</emphasis>
<taxonomicName id="2378BF7403F646357BFB5F588412FA55" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="rattus">
<emphasis id="8C35D9EB52B404F0CF6A0C0A154B27CF" italics="true" pageNumber="1463">Rattus rattus</emphasis>
</taxonomicName>
species group. Still known only by the
<typeStatus id="764367C7EDFCA1E23E4B510F38B327BC">holotype</typeStatus>
. Although
<bibRefCitation id="61E05BA59728FF32F52CBFBB5B2CC92A" author="Sody, H. J. V." refId="ref332575" refString="Sody, H. J. V. 1940. On the mammals of Enggano. Treubia, 17: 391 - 405." year="1940">Sody (1940)</bibRefCitation>
described
<taxonomicName id="D9237B50A1D6919E4B8500661E7971DC" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="adustus">
<emphasis id="BD3B27D8C98100200919CDB2FF502F9B" italics="true" pageNumber="1463">adustus</emphasis>
</taxonomicName>
as a species, he later listed it as a subspecies of
<taxonomicName id="3879FADC5F8031E88D05D532A316AC81" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="rattus">
<emphasis id="D7EE430F67B4E77445BF9515FE3B3FB1" italics="true" pageNumber="1463">R. rattus</emphasis>
</taxonomicName>
in a section also containing
<taxonomicName id="CF97AEDD65927870A054FC30EB008B0E" authorityName="Miller" authorityYear="1903" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="lugens">
<emphasis id="10A8F331EF1CFB7A2C545D6D7C55E263" italics="true" pageNumber="1463">lugens</emphasis>
</taxonomicName>
and
<taxonomicName id="F035E059BF80F127E2542E7229AD8455" authorityName="Chasen and Kloss" authorityYear="1928" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="mentawai">
<emphasis id="21A19EFD28585CB1E5C0BA05B355D9C8" italics="true" pageNumber="1463">mentawai</emphasis>
</taxonomicName>
, populations endemic to the Mentawai Isls (
<bibRefCitation id="9CB216781DB659608335F837E97DB43E" author="Sody, H. J. V." refId="ref332599" refString="Sody, H. J. V. 1941. On a collection of rats from the Indo- Malayan and Indo- Australian regions (with descriptions of 43 new genera, species and subspecies). Treubia, 18: 255 - 325." year="1941">Sody, 1941</bibRefCitation>
). In morphology and geographic proximity,
<taxonomicName id="BE8D1E815B2317C52D795EC7BB08997D" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="adustus">
<emphasis id="F388F197CC3051385C838123E5521A17" italics="true" pageNumber="1463">R. adustus</emphasis>
</taxonomicName>
is related to
<taxonomicName id="90FB6EDD8660A795E0DB38DAF745C45F" authorityName="Miller" authorityYear="1903" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="lugens">
<emphasis id="B683809E3A8365EAF9991993759DFE8A" italics="true" pageNumber="1463">R. lugens</emphasis>
</taxonomicName>
, the two are allies of
<taxonomicName id="DC66AE87C4EC42291EE2AEE2A7CE25B8" authorityName="Miller" authorityYear="1903" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="simalurensis">
<emphasis id="D255E6905EFFE572A0906AE3E1C820EB" italics="true" pageNumber="1463">R. simalurensis</emphasis>
</taxonomicName>
from the Simalur Arch., and all three share close kinship with Sundaic
<taxonomicName id="AC4975B679B90D111B7950FDA08AF30D" authorityName="Miller" authorityYear="1900" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="tiomanicus">
<emphasis id="5CD0081145F635043904A798A14F651E" italics="true" pageNumber="1463">R. tiomanicus</emphasis>
</taxonomicName>
(Musser and Heaney, 1985; Musser, ms.).
<taxonomicName id="ECDAD6BCC630E292095E56E4C2FF7467" authorityName="Sody" authorityYear="1940" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="adustus">
<emphasis id="D5D8A6160DF023C2E7EF07EF80BADB43" italics="true" pageNumber="1463">Rattus adustus</emphasis>
</taxonomicName>
,
<taxonomicName id="B524A9E1E49A066D0581DA881DBCED3B" authorityName="Miller" authorityYear="1906" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="enganus">
<emphasis id="C08CF20614F01EF1378412E5231D1451" italics="true" pageNumber="1463">R. enganus</emphasis>
</taxonomicName>
(see that account), and
<taxonomicName id="F9ACE92E5E0460C5F592A3F3EDFD5493" authorityName="Miller" authorityYear="1900" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="tiomanicus">
<emphasis id="516C94507A768A99D07E429DF35C6633" italics="true" pageNumber="1463">R. tiomanicus</emphasis>
</taxonomicName>
are the only native
<taxonomicName id="3C1F3C38DC2C4E2F87CB91A99D21DD78" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="D187791151AD33261422A864F48E65DA" italics="true" pageNumber="1463">Rattus</emphasis>
</taxonomicName>
(and the sole native murines) on Pulau Enggano.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="18F47772E8BFE5F8351653EE9E4DA324" ID-DOI="10.5281/zenodo.5356276" ID-ISSN="0253-116X" ID-Zenodo-Dep="5356276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630216915060" checkinUser="marcus" docAuthor="W, David &amp; M, Boris" docDate="2016" docId="A31A87A70D09FFF552C5EFBFFD683E3B" docLanguage="en" docName="LinzerbiolBeitr.2016.48.2.1783-1806.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Acinopus (Acinopus) laevigatus MENETRIES 1832" docType="treatment" docVersion="3" lastPageNumber="1804" masterDocId="5F23FFDF0D1DFFE05277EA3CFF833D7C" masterDocTitle="Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)" masterLastPageNumber="1806" masterPageNumber="1783" pageNumber="1803" updateTime="1699190026385" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="134D9FE39EA32EB172927EC6F141F967">Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)</mods:title>
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<taxonomicName id="ECB34D320D09FFF452C5EFBFFD4238E4" ID-CoL="9BVN" authority="MENETRIES, 1832" authorityName="MENETRIES" authorityYear="1832" box="[178,705,1410,1433]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="laevigatus" subGenus="Acinopus">
<emphasis id="19C7EAA30D09FFF452C5EFBFFD0238E4" bold="true" box="[178,641,1410,1433]" pageId="20" pageNumber="1803">
<emphasis id="19C7EAA30D09FFF452C5EFBFFE9738E4" bold="true" box="[178,276,1411,1432]" italics="true" pageId="20" pageNumber="1803">Acinopus</emphasis>
(
<emphasis id="19C7EAA30D09FFF45354EFBFFE0638E4" bold="true" box="[291,389,1411,1432]" italics="true" pageId="20" pageNumber="1803">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D09FFF453E2EFBFFE7C38E5" bold="true" box="[405,511,1411,1433]" italics="true" pageId="20" pageNumber="1803">laevigatus</emphasis>
MÉNÉTRIÉS
</emphasis>
,
<emphasis id="19C7EAA30D09FFF450F9EFBFFD4238E4" bold="true" box="[654,705,1411,1432]" pageId="20" pageNumber="1803">1832</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D09FFF552C5EF95FD3F3C8D" lastPageId="21" lastPageNumber="1804" pageId="20" pageNumber="1803" type="reference_group">
<paragraph id="2B0C36B10D09FFF452C5EF95FE2338AA" blockId="20.[178,1085,1449,1495]" pageId="20" pageNumber="1803">
<treatmentCitationGroup id="0BA3119F0D09FFF452C5EF95FE2338AA" pageId="20" pageNumber="1803">
<treatmentCitation id="AA1210A00D09FFF452C5EF95FC8F38C2" author="SCHAUBERGER E" box="[178,780,1449,1470]" page="7" pageId="20" pageNumber="1803" year="1927">
<taxonomicName id="ECB34D320D09FFF452C5EF95FC8F38C2" ID-CoL="9BWP" authority="SCHAUBERGER, 1927: 7" authorityName="SCHAUBERGER" authorityPageNumber="7" authorityYear="1927" box="[178,780,1449,1470]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="subSpecies" species="laevigatus" subSpecies="kashmirensis">
<emphasis id="19C7EAA30D09FFF452C5EF95FEED38C2" box="[178,366,1449,1470]" italics="true" pageId="20" pageNumber="1803">Acinopus laevigatus</emphasis>
ssp.
<emphasis id="19C7EAA30D09FFF453D1EF95FD9F38C2" box="[422,540,1449,1470]" italics="true" pageId="20" pageNumber="1803">kashmirensis</emphasis>
<bibRefCitation id="4F224B400D09FFF45050EF95FC8F38C2" author="SCHAUBERGER E" box="[551,780,1449,1470]" pageId="20" pageNumber="1803" pagination="1927 - 1928" refId="ref14927" refString="SCHAUBERGER E. (1927): Beitrag zur Kenntnis der palaarktischen Harpalinen, III. - Coleopterologisches Centralblatt 2 [1927 - 1928]: 6 - 19." type="journal article" year="1927">SCHAUBERGER, 1927: 7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="F40888130D09FFF4516AEF95FCC538C2" box="[797,838,1449,1470]" pageId="20" pageNumber="1803">type</typeStatus>
loc.: &quot;Poo im westlichen Himalaja&quot;)
<emphasis id="19C7EAA30D09FFF45326EFFDFE2338AA" bold="true" box="[337,416,1473,1494]" pageId="20" pageNumber="1803">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2B0C36B10D09FFF552C5EFD9FD3F3C8D" blockId="20.[178,1085,1509,1588]" lastBlockId="21.[178,1086,218,497]" lastPageId="21" lastPageNumber="1804" pageId="20" pageNumber="1803">
<bibRefCitation id="4F224B400D09FFF452C5EFD9FE2E3886" author="SCHAUBERGER E" box="[178,429,1509,1530]" pageId="20" pageNumber="1803" pagination="1927 - 1928" refId="ref14927" refString="SCHAUBERGER E. (1927): Beitrag zur Kenntnis der palaarktischen Harpalinen, III. - Coleopterologisches Centralblatt 2 [1927 - 1928]: 6 - 19." type="journal article" year="1927">SCHAUBERGER (1927: 7)</bibRefCitation>
described the subspecies
<emphasis id="19C7EAA30D09FFF450C1EFD9FCE83886" box="[694,875,1509,1530]" italics="true" pageId="20" pageNumber="1803">A. l. kashmirensis</emphasis>
based on several (at least six: see
<bibRefCitation id="4F224B400D09FFF45335EC3DFDD33B6B" author="GUSENLEITNER F" box="[322,592,1537,1559]" pageId="20" pageNumber="1803" pagination="725 - 785" refId="ref14373" refString="GUSENLEITNER F. (1990): Die Typen der Carabidae-Sammlung Dr. Erwin Schauberger am Oberosterreichischen Landesmuseum in Linz, Austria (Insecta: Coleoptera: Carabidae). - Linzer biologische Beitrage 22 (2): 725 - 785." type="journal article" year="1990">GUSENLEITNER 1990: 763</bibRefCitation>
) specimens which differ from specimens from South
<collectingCountry id="53A476210D09FFF45283EC22FEBB3B48" box="[244,312,1566,1588]" name="Russia" pageId="20" pageNumber="1803">Russia</collectingCountry>
and West Asia by the excision of the right mandible longer and more shal- low and by the pronotum stronger and more evenly rounded. Though not having examined the
<typeStatus id="F40888130D08FFF55324EACBFE083C70" box="[339,395,247,268]" pageId="21" pageNumber="1804">types</typeStatus>
, we investigated several hundreds of specimens of
<taxonomicName id="ECB34D320D08FFF551A8EACBFE993C54" ID-CoL="9BVN" authorityName="MENETRIES" authorityYear="1832" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1804" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D08FFF551A8EACBFE993C54" italics="true" pageId="21" pageNumber="1804">Acinopus laevigatus</emphasis>
</taxonomicName>
from all areas of its western distribution and compared them with specimens from the western Himalaya (including specimens coming from the area of the
<typeStatus id="F40888130D08FFF551A0EB0CFB803C39" box="[983,1027,304,325]" pageId="21" pageNumber="1804">type</typeStatus>
locality) not only in external characters, but also concerning the construction of the median lobe of the aedeagus, stating that the characters Schauberger mentioned are very variable and can be met also in specimens coming from western areas outside the West Himalaya, hence, a variability in geographical aspect cannot be stated. Therefore we propose to treat
<emphasis id="19C7EAA30D08FFF55290EB83FE1D3CAF" box="[231,414,446,468]" italics="true" pageId="21" pageNumber="1804">A. l. kashmirensis</emphasis>
as a junior subjective synonym of
<taxonomicName id="ECB34D320D08FFF5517DEB83FC0C3CAF" ID-CoL="9BVN" authorityName="MENETRIES" authorityYear="1832" box="[778,911,446,468]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1804" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D08FFF5517DEB83FC9A3CA8" box="[778,793,447,468]" italics="true" pageId="21" pageNumber="1804">A</emphasis>
.
<emphasis id="19C7EAA30D08FFF55150EB82FC0C3CAF" box="[807,911,446,467]" italics="true" pageId="21" pageNumber="1804">laevigatus</emphasis>
</taxonomicName>
. For other synonyms of
<taxonomicName id="ECB34D320D08FFF5537CEBE7FE0C3C8C" ID-CoL="9BVN" authorityName="MENETRIES" authorityYear="1832" box="[267,399,475,496]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1804" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D08FFF5537CEBE7FE0C3C8C" box="[267,399,475,496]" italics="true" pageId="21" pageNumber="1804">A. laevigatus</emphasis>
</taxonomicName>
see
<bibRefCitation id="4F224B400D08FFF553CBEBE7FD3B3C8D" author="KATAEV B &amp; WRASE D" box="[444,696,475,497]" pageId="21" pageNumber="1804" pagination="367 - 397" refId="ref14649" refString="KATAEV B. M., WRASE D. W. &amp; N. ITO (2003): Harpalina, pp. 367 - 397. - In: LOBL I. &amp; A. SMETANA (eds), Catalogue of Palaearctic Coleoptera, Vol. 1 - Myxophaga - Adephaga. Stenstrup: Apollo Books, 819 pp." type="book chapter" year="2003">KATAEV et al. 2003: 368</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D08FFF552C5E83CFD683E3B" pageId="21" pageNumber="1804" type="description">
<paragraph id="2B0C36B10D08FFF552C5E83CFB973F6A" blockId="21.[178,1087,512,839]" box="[178,1044,512,534]" pageId="21" pageNumber="1804">
M a t e r i a l f r o m w e s t e r n H i m a l a y a e x a m i n e d (
<specimenCount id="3DB5FD380D08FFF551F7E83CFB8B3F6A" box="[896,1032,512,534]" pageId="21" pageNumber="1804" type="generic">32specimens</specimenCount>
):
</paragraph>
<paragraph id="2B0C36B10D08FFF552C5E81DFDA93FA2" blockId="21.[178,1087,512,839]" pageId="21" pageNumber="1804">
<materialsCitation id="9BDB3CEC0D08FFF552C5E81DFB8A3F32" collectionCode="MFNB" county="Lahaul" location="Keylong" municipality="Spiti district" pageId="21" pageNumber="1804" specimenCount="2" specimenCount-female="2" stateProvince="Himachal Pradesh">
I n d i a:
<collectingRegion id="E977F8530D08FFF5536DE81DFE473F4B" box="[282,452,545,567]" country="India" name="Himachal Pradesh" pageId="21" pageNumber="1804">Himachal Pradesh</collectingRegion>
:
<collectingCounty id="C26D4E3D0D08FFF553A5E81DFD8C3F4B" box="[466,527,545,567]" pageId="21" pageNumber="1804">Lahaul</collectingCounty>
and
<collectingMunicipality id="CB68ACCB0D08FFF55030E81DFD383F4B" box="[583,699,545,567]" pageId="21" pageNumber="1804">Spiti district</collectingMunicipality>
:
<location id="2E6C606A0D08FFF550BDE81DFC953F4B" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF550BDE81DFC953F4B" box="[714,790,545,567]" county="Lahaul" municipality="Spiti district" name="Keylong" pageId="21" pageNumber="1804" stateProvince="Himachal Pradesh">Keylong</location>
(&quot;
<location id="2E6C606A0D08FFF55158E81DFCFC3F4B" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF55158E81DFCFC3F4B" box="[815,895,545,567]" county="Lahaul" municipality="Spiti district" name="Kyelang" pageId="21" pageNumber="1804" stateProvince="Himachal Pradesh">Kyelang</location>
&quot;, 1 spcm.,
<collectionCode id="4DA2AE740D08FFF55185E81DFBB63F4A" box="[1010,1077,545,566]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="1804">MFNB</collectionCode>
).
<location id="2E6C606A0D08FFF552A8E805FEEF3F33" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF552A8E805FEEF3F33" box="[223,364,569,591]" county="Lahaul" municipality="Spiti district" name="Kinnaur district" pageId="21" pageNumber="1804" stateProvince="Himachal Pradesh">Kinnaur district</location>
:
<location id="2E6C606A0D08FFF55304E805FE153F32" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF55304E805FE153F32" box="[371,406,569,590]" county="Lahaul" municipality="Spiti district" name="Poo" pageId="21" pageNumber="1804" stateProvince="Himachal Pradesh">Poo</location>
(&quot;
<location id="2E6C606A0D08FFF553DFE805FDC13F32" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF553DFE805FDC13F32" box="[424,578,569,591]" county="Lahaul" municipality="Spiti district" name="Bashar State Poo" pageId="21" pageNumber="1804" stateProvince="Himachal Pradesh">Bashar State Poo</location>
&quot;, 1066/
<specimenCount id="3DB5FD380D08FFF550E4E802FD293F2F" box="[659,682,574,595]" pageId="21" pageNumber="1804" type="female">♀♀</specimenCount>
; &quot;Kaschmir Poo&quot;, 566 /
<specimenCount id="3DB5FD380D08FFF551E6E802FC2B3F2F" box="[913,936,574,595]" pageId="21" pageNumber="1804" type="female">♀♀</specimenCount>
,
<collectionCode id="4DA2AE740D08FFF551B6E805FB803F32" box="[961,1027,569,590]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="1804">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9BDB3CEC0D08FFF55665E805FDA53FA2" collectingDate="1914-08" collectingDateMax="2003-06-27" collectingDateMin="1914-08" collectionCode="BMNH, MFNB, MNHN, ZIN" collectorName="R. Novak &amp; G. Babault &amp; Gy. Topal" county="Shimla District" elevation="3400" location="Simla" municipality="Shimla" pageId="21" pageNumber="1804" specimenCount="3" specimenCount-female="3" stateProvince="Jammu and Kashmir">
&quot;Poo Kashmir&quot; (
<specimenCount id="3DB5FD380D08FFF55335E86DFEE63F17" box="[322,357,593,619]" pageId="21" pageNumber="1804" type="female">2♀♀</specimenCount>
,
<collectionCode id="4DA2AE740D08FFF553F6E86DFE443F1A" box="[385,455,593,614]" pageId="21" pageNumber="1804">BMNH</collectionCode>
).
<collectingCounty id="C26D4E3D0D08FFF553AFE86DFDE13F1B" box="[472,610,593,615]" pageId="21" pageNumber="1804">Shimla District</collectingCounty>
:
<collectingMunicipality id="CB68ACCB0D08FFF5501AE86DFD283F1B" box="[621,683,593,615]" pageId="21" pageNumber="1804">Shimla</collectingMunicipality>
(&quot;
<location id="2E6C606A0D08FFF550B6E86DFD7A3F1B" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF550B6E86DFD7A3F1B" box="[705,761,593,615]" county="Shimla District" municipality="Shimla" name="Simla" pageId="21" pageNumber="1804" stateProvince="Jammu and Kashmir">Simla</location>
&quot;, 2 spcms., MFNB, one with label:
<taxonomicName id="ECB34D320D08FFF552A8E855FDFA3F03" authority="Men. H. W. Andrewes" box="[223,633,617,639]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1804" phylum="Arthropoda" rank="species" species="laevigatus">Acinopus laevigatus Mén. H.W. Andrewes</taxonomicName>
det.). -
<collectingRegion id="E977F8530D08FFF550A2E855FC1C3F03" box="[725,927,617,639]" country="India" name="Jammu and Kashmir" pageId="21" pageNumber="1804">Jammu and Kashmir</collectingRegion>
:
<location id="2E6C606A0D08FFF551D8E855FBB93F03" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF551D8E855FBB93F03" box="[943,1082,617,639]" county="Shimla District" municipality="Shimla" name="Ladakh District" pageId="21" pageNumber="1804" stateProvince="Jammu and Kashmir">Ladakh District</location>
: &quot;
<location id="2E6C606A0D08FFF55291E8BDFEB73FEB" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF55291E8BDFEB73FEB" box="[230,308,641,663]" county="Shimla District" municipality="Shimla" name="Khalatse" pageId="21" pageNumber="1804" stateProvince="Jammu and Kashmir">Khalatse</location>
15000.&quot; (966/
<specimenCount id="3DB5FD380D08FFF553A0E8BAFE6D3FE7" box="[471,494,646,667]" pageId="21" pageNumber="1804" type="female">♀♀</specimenCount>
,
<collectionCode id="4DA2AE740D08FFF55061E8BDFDDB3FEA" box="[534,600,641,662]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="1804">MFNB</collectionCode>
). Ganderbal District: Sonamarg,
<quantity id="EC4B9B540D08FFF551C4E8BDFBB93FEA" box="[947,1082,641,662]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="4.0" metricValueMin="2.8" pageId="21" pageNumber="1804" unit="m" value="3400.0" valueMax="4000.0" valueMin="2800.0">
<elevation id="A09ED1820D08FFF551C4E8BDFBB93FEA" box="[947,1082,641,662]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="4.0" metricValueMin="2.8" pageId="21" pageNumber="1804" unit="m" value="3400.0" valueMax="4000.0" valueMin="2800.0">2800-4000 m</elevation>
</quantity>
, 23./
<date id="5F0D10710D08FFF55377E8A5FEE53FD2" box="[256,358,665,686]" pageId="21" pageNumber="1804" value="2003-06-27">
<collectingDate id="4F49E9990D08FFF55377E8A5FEE53FD2" box="[256,358,665,686]" pageId="21" pageNumber="1804" value="2003-06-27">27.VI.2003</collectingDate>
</date>
,
<collectorName id="864653670D08FFF55305E8A5FE483FD3" box="[370,459,665,687]" pageId="21" pageNumber="1804">R. Novák</collectorName>
leg. (16, cWR); &quot;Sonemarg (Cachemire)&quot;, &quot;
<collectorName id="864653670D08FFF5511BE8A5FC523FD3" box="[876,977,665,687]" pageId="21" pageNumber="1804">G. Babault</collectorName>
,
<date id="5F0D10710D08FFF551A9E8A5FBB53FD2" box="[990,1078,665,686]" pageId="21" pageNumber="1804" value="1914-08">
<collectingDate id="4F49E9990D08FFF551A9E8A5FBB53FD2" box="[990,1078,665,686]" pageId="21" pageNumber="1804" value="1914-08">VIII.1914</collectingDate>
</date>
&quot; (16,
<collectionCode id="4DA2AE740D08FFF55378E88DFED43FBA" box="[271,343,689,710]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="1804">MNHN</collectionCode>
).
<location id="2E6C606A0D08FFF55311E88DFE783FBB" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF55311E88DFE783FBB" box="[358,507,689,711]" county="Shimla District" municipality="Shimla" name="Srinagar District" pageId="21" pageNumber="1804" stateProvince="Jammu and Kashmir">Srinagar District</location>
: Srinagar, Zabarwan Range, Pari Mahal, from under stones/dung,
<date id="5F0D10710D08FFF552A8E8F5FEBE3FA3" box="[223,317,713,735]" pageId="21" pageNumber="1804" value="1967-05-25">
<collectingDate id="4F49E9990D08FFF552A8E8F5FEBE3FA3" box="[223,317,713,735]" pageId="21" pageNumber="1804" value="1967-05-25">25.V.1967</collectingDate>
</date>
,
<collectorName id="864653670D08FFF55331E8F5FE1D3FA3" box="[326,414,713,735]" pageId="21" pageNumber="1804">Gy. Topál</collectorName>
leg. (16,
<collectionCode id="4DA2AE740D08FFF55380E8F5FD9C3FA2" box="[503,543,713,734]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="1804" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2B0C36B10D08FFF552C5E8D5FD683E3B" blockId="21.[178,1087,512,839]" pageId="21" pageNumber="1804">
<materialsCitation id="9BDB3CEC0D08FFF552C5E8D5FBB93F82" box="[178,1082,745,767]" collectingDate="2008-06-07" location="Swat Valley" pageId="21" pageNumber="1804" specimenCount="1" stateProvince="Khyber Pakhtunkhwa">
P a k i s t a n:
<collectingRegion id="E977F8530D08FFF5533BE8D5FDE33F83" box="[332,608,745,767]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="21" pageNumber="1804">Khyber Pakhtunkhwa Province</collectingRegion>
:
<location id="2E6C606A0D08FFF55005E8D5FD623F83" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF55005E8D5FD623F83" box="[626,737,745,767]" name="Swat Valley" pageId="21" pageNumber="1804" stateProvince="Khyber Pakhtunkhwa">Swat Valley</location>
,
<date id="5F0D10710D08FFF5509CE8D5FCC63F82" box="[747,837,745,766]" pageId="21" pageNumber="1804" value="2008-06-07">
<collectingDate id="4F49E9990D08FFF5509CE8D5FCC63F82" box="[747,837,745,766]" pageId="21" pageNumber="1804" value="2008-06-07">7.VI.2008</collectingDate>
</date>
, local collector (16, cWR)
</materialsCitation>
;
<materialsCitation id="9BDB3CEC0D08FFF55297E93DFD643E3B" collectingDate="1971-06-19" collectingDateMax="2003-07-10" collectingDateMin="1971-06-19" collectionCode="NME" collectorName="C. Holzschuh &amp; Azad Jammu &amp; Kashmir &amp; Naran &amp; V. Gurko &amp; S. Ovchinnikov" country="Pakistan" elevation="1400" location="Madyan" municipality="Swat Prov." pageId="21" pageNumber="1804" specimenCount="1">
NW
<collectingCountry id="53A476210D08FFF5537BE93DFEDB3E6B" box="[268,344,769,791]" name="Pakistan" pageId="21" pageNumber="1804">Pakistan</collectingCountry>
,
<collectingMunicipality id="CB68ACCB0D08FFF55316E93DFE463E6A" box="[353,453,769,790]" pageId="21" pageNumber="1804">Swat Prov.</collectingMunicipality>
,
<location id="2E6C606A0D08FFF553BAE93DFD943E6B" LSID="urn:lsid:plazi:treatment:A31A87A70D09FFF552C5EFBFFD683E3B:2E6C606A0D08FFF553BAE93DFD943E6B" box="[461,535,769,791]" country="Pakistan" municipality="Swat Prov." name="Madyan" pageId="21" pageNumber="1804">Madyan</location>
, 35°70' N 71°90' E,
<quantity id="EC4B9B540D08FFF550BBE93DFC923E6A" box="[716,785,769,790]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="1804" unit="m" value="1400.0">
<elevation id="A09ED1820D08FFF550BBE93DFC923E6A" box="[716,785,769,790]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="1804" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="5F0D10710D08FFF5516CE93DFC3F3E6A" box="[795,956,769,790]" pageId="21" pageNumber="1804" value="1971-06-19" valueMax="1971-07-04" valueMin="1971-06-19">
<collectingDate id="4F49E9990D08FFF5516CE93DFC3F3E6A" box="[795,956,769,790]" pageId="21" pageNumber="1804" value="1971-06-19" valueMax="1971-07-04" valueMin="1971-06-19">19.VI.-4.VII.1971</collectingDate>
</date>
,
<collectorName id="864653670D08FFF551B2E93DFBBE3E6B" box="[965,1085,769,791]" pageId="21" pageNumber="1804">C. Holzschuh</collectorName>
leg. (16,
<collectionCode id="4DA2AE740D08FFF55349E925FEF03E52" box="[318,371,793,814]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="21" pageNumber="1804">NME</collectionCode>
). -
<collectorName id="864653670D08FFF553E0E925FD933E52" box="[407,528,793,815]" pageId="21" pageNumber="1804">Azad Jammu</collectorName>
and
<collectorName id="864653670D08FFF55033E925FD103E53" box="[580,659,793,815]" pageId="21" pageNumber="1804">Kashmir</collectorName>
: env.
<collectorName id="864653670D08FFF550B9E925FC873E52" box="[718,772,793,814]" pageId="21" pageNumber="1804">Naran</collectorName>
&quot;NW from Junkar&quot;,
<quantity id="EC4B9B540D08FFF551B9E925FB9B3E52" box="[974,1048,793,814]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="1804" unit="m" value="3000.0">
<elevation id="A09ED1820D08FFF551B9E925FB9B3E52" box="[974,1048,793,814]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="1804" unit="m" value="3000.0">3000 m</elevation>
</quantity>
,
<date id="5F0D10710D08FFF55652E925FECF3E3A" pageId="21" pageNumber="1804" value="2003-07-01" valueMax="2003-07-10" valueMin="2003-07-01">
<collectingDate id="4F49E9990D08FFF55652E925FECF3E3A" pageId="21" pageNumber="1804" value="2003-07-01" valueMax="2003-07-10" valueMin="2003-07-01">1.- 10.VII.2003</collectingDate>
</date>
,
<collectorName id="864653670D08FFF55323E90DFE253E3B" box="[340,422,817,839]" pageId="21" pageNumber="1804">V. Gurko</collectorName>
&amp;
<collectorName id="864653670D08FFF553B5E90DFDC83E3B" box="[450,587,817,839]" pageId="21" pageNumber="1804">S. Ovchinnikov</collectorName>
leg. (16, cGRK)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,293 @@
<document id="3F0C129B8079E7E77B2CFEC766CC2FEC" ID-DOI="10.5281/zenodo.5356276" ID-ISSN="0253-116X" ID-Zenodo-Dep="5356276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630216915060" checkinUser="marcus" docAuthor="W, David &amp; M, Boris" docDate="2016" docId="A31A87A70D0CFFF452C5EF4BFC4E3824" docLanguage="en" docName="LinzerbiolBeitr.2016.48.2.1783-1806.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Acinopus (Acinopus) arabicus W &amp; M 2016, nov.sp." docType="treatment" docVersion="4" lastPageNumber="1803" masterDocId="5F23FFDF0D1DFFE05277EA3CFF833D7C" masterDocTitle="Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)" masterLastPageNumber="1806" masterPageNumber="1783" pageNumber="1800" updateTime="1699190026385" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="83D9439ECBFC216B3F48459295B1BFFC">Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)</mods:title>
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<mods:namePart id="3DC7BC6CE953B682B3E4B2DE6D4E9750">W, David</mods:namePart>
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<mods:namePart id="05DFAB5A5470680B0EF8ACE60601BB5C">M, Boris</mods:namePart>
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<mods:date id="3F982915E0CE33704D1F1DEEC7F67293">2016</mods:date>
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<mods:number id="F615490C6DB1E3D9A54E6A407823DA52">2016-12-19</mods:number>
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<mods:start id="293A7B8CA05B52400575400DC1C20765">1783</mods:start>
<mods:end id="14A5085C76FD0EA8CE74DA2CE8BEF1FA">1806</mods:end>
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<paragraph id="2B0C36B10D0CFFF152C5EF4BFC8938F0" blockId="17.[178,778,1398,1420]" box="[178,778,1398,1420]" pageId="17" pageNumber="1800">
<heading id="704481DD0D0CFFF152C5EF4BFC8938F0" box="[178,778,1398,1420]" fontSize="8" level="2" pageId="17" pageNumber="1800" reason="2">
<emphasis id="19C7EAA30D0CFFF152C5EF4BFC8938F0" bold="true" box="[178,778,1398,1420]" pageId="17" pageNumber="1800">
<taxonomicName id="ECB34D320D0CFFF152C5EF4BFE6C38F0" ID-CoL="64KDY" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[178,495,1398,1420]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1800" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov." subGenus="Acinopus">
<emphasis id="19C7EAA30D0CFFF152C5EF4BFE9738F0" bold="true" box="[178,276,1399,1420]" italics="true" pageId="17" pageNumber="1800">Acinopus</emphasis>
(
<emphasis id="19C7EAA30D0CFFF15354EF4BFE0638F0" bold="true" box="[291,389,1399,1420]" italics="true" pageId="17" pageNumber="1800">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D0CFFF153E2EF4AFE6C38F0" bold="true" box="[405,495,1398,1420]" italics="true" pageId="17" pageNumber="1800">arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D0CFFF15382EF4BFDC238F0" box="[501,577,1399,1420]" pageId="17" pageNumber="1800" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="B3882A340D0CFFF15039EF4AFD1238F0" box="[590,657,1398,1420]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="17" pageNumber="1800">Figs 4</figureCitation>
,
<figureCitation id="B3882A340D0CFFF150EAEF4BFD5E38F0" box="[669,733,1399,1420]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="17" pageNumber="1800">11, 12</figureCitation>
,
<figureCitation id="B3882A340D0CFFF1509EEF4AFC8038F0" box="[745,771,1398,1420]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="17" pageNumber="1800">16</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="2B0C36B10D0CFFF152C5EFA1FDF4389E" blockId="17.[178,1087,1436,1511]" pageId="17" pageNumber="1800">
T y p e m a t e r i a l:
<typeStatus id="F40888130D0CFFF153DEEFA0FE7838CE" box="[425,507,1436,1458]" pageId="17" pageNumber="1800" type="holotype">Holotype</typeStatus>
6, labelled: &quot;
<collectingCountry id="53A476210D0CFFF1500CEFA1FC9E38CE" box="[635,797,1437,1458]" name="Saudi Arabia" pageId="17" pageNumber="1800">SAUDI ARABIA</collectingCountry>
/
<collectingRegion id="E977F8530D0CFFF15145EFA0FC1238CE" box="[818,913,1436,1458]" country="Saudi Arabia" name="'Asir" pageId="17" pageNumber="1800">Asir Prov.</collectingRegion>
,
<quantity id="EC4B9B540D0CFFF151EDEFA0FB9038CE" box="[922,1043,1436,1458]" metricMagnitude="3" metricUnit="m" metricValue="2.925" metricValueMax="3.05" metricValueMin="2.8" pageId="17" pageNumber="1800" unit="m" value="2925.0" valueMax="3050.0" valueMin="2800.0">2800-3050 m</quantity>
/ Al Soudah, ca.
<quantity id="EC4B9B540D0CFFF1533DEF88FE0038B6" box="[330,387,1460,1482]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="1800" unit="km" value="50.0">50 km</quantity>
NW Abha /
<date id="5F0D10710D0CFFF15383EF89FDC238B6" box="[500,577,1461,1482]" pageId="17" pageNumber="1800" value="1994-01-04">4.1.1994</date>
/ H.J. Bremer leg.&quot; (black print on white label), (MFNB).
<typeStatus id="F40888130D0CFFF152A8EFF1FEB4389E" box="[223,311,1485,1506]" pageId="17" pageNumber="1800" type="paratype">Paratypes</typeStatus>
: 16,
<specimenCount id="3DB5FD380D0CFFF15310EFF1FEFD389B" box="[359,382,1485,1511]" pageId="17" pageNumber="1800" type="female">1♀</specimenCount>
: same data (MFNB, cWR).
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D0FFFF452C5EAE7FC4E3824" lastPageId="20" lastPageNumber="1803" pageId="18" pageNumber="1801" type="description">
<paragraph id="2B0C36B10D0FFFF252C5EAE7FDC13C71" blockId="18.[178,1085,218,269]" pageId="18" pageNumber="1801">E t y m o l o g y: The specific epithet is a latinizid adjective, based on the name of the region in which this species was found.</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5EB20FE983F4D" blockId="18.[178,1085,283,562]" pageId="18" pageNumber="1801">
D i a g n o s i s: A micropterous species of small to medium size for
<taxonomicName id="ECB34D320D0FFFF251D6EB27FC7C3C4C" authorityName="DEJEAN" authorityYear="1821" box="[929,1023,283,304]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF251D6EB27FC7C3C4C" box="[929,1023,283,304]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
, with excision of dorsal edge of right mandible (subgenus
<taxonomicName id="ECB34D320D0FFFF250A8EB04FCBA3C31" authorityName="DEJEAN" authorityYear="1821" box="[735,825,312,333]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF250A8EB04FCBA3C31" box="[735,825,312,333]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
), black, tarsomeres and antennae dark reddish-brown, labrum moderately incised at anterior margin, clypeus with one setiferous pore puncture at anterior angles, pronotum laterally almost rectilinearly or very weakly convexly narrowed toward the obtuse-angled posterior angles and weakly convexly narrowed apicad, anterior angles somewhat protruding, narrowly rounded at tip, posterior angles widely rounded, elytra short-oval, moderately convex on disc, metepisternum very short, pro- and mesotarsi in males moderately widened, apical lamella of median lobe short, about somewhat elongate-triangular (dorsal view). Habitus see
<figureCitation id="B3882A340D0FFFF252AEE820FE983F4D" box="[217,283,540,562]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4.</figureCitation>
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E87DFEE73FF3" blockId="18.[178,1085,577,655]" pageId="18" pageNumber="1801">
D e s c r i p t i o n: Body length in males
<quantity id="EC4B9B540D0FFFF2500FE87DFC8F3F2A" box="[632,780,577,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.455" metricValueMax="1.47" metricValueMin="1.44" pageId="18" pageNumber="1801" unit="mm" value="14.55" valueMax="14.7" valueMin="14.4">14.4-14.7 mm</quantity>
, in
<typeStatus id="F40888130D0FFFF25142E87DFC0D3F2B" box="[821,910,577,599]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D0FFFF251E1E87DFC773F2A" box="[918,1012,577,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="18" pageNumber="1801" unit="mm" value="14.7">14.7 mm</quantity>
, in the female
<typeStatus id="F40888130D0FFFF2528AE862FED73F0F" box="[253,340,606,627]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
<quantity id="EC4B9B540D0FFFF2532CE861FE3A3F0F" box="[347,441,605,627]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="1801" unit="mm" value="12.9">12.9 mm</quantity>
; width in males
<quantity id="EC4B9B540D0FFFF25017E861FD5B3F0F" box="[608,728,605,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="5.5" metricValueMin="5.4" pageId="18" pageNumber="1801" unit="mm" value="5.45" valueMax="5.5" valueMin="5.4">5.4-5.5 mm</quantity>
, in
<typeStatus id="F40888130D0FFFF2508AE861FCD53F0F" box="[765,854,605,627]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D0FFFF2512AE861FC2E3F0F" box="[861,941,605,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="1801" unit="mm" value="5.5">5.5 mm</quantity>
; in the female
<typeStatus id="F40888130D0FFFF252C5E846FE8A3FF3" box="[178,265,634,655]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
<quantity id="EC4B9B540D0FFFF25367E845FEE23FF3" box="[272,353,633,655]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="18" pageNumber="1801" unit="mm" value="5.1">5.1 mm</quantity>
.
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E8A2FD6B3FC8" blockId="18.[178,744,670,692]" box="[178,744,670,692]" pageId="18" pageNumber="1801">Colour: Black, tarsomeres and antennae reddish-brown.</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E8FFFBBD3EA5" blockId="18.[178,1086,707,985]" pageId="18" pageNumber="1801">
Head: Wide (as normal in
<taxonomicName id="ECB34D320D0FFFF253B2E8FFFDA03FA4" authorityName="DEJEAN" authorityYear="1821" box="[453,547,707,728]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF253B2E8FFFDA03FA4" box="[453,547,707,728]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
), only somewhat narrower than pronotum (PW/HW in males 1.18-1.21, in
<typeStatus id="F40888130D0FFFF253EBE8E3FE763F89" box="[412,501,735,757]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.18; in the female
<typeStatus id="F40888130D0FFFF250B0E8DCFC9D3F89" box="[711,798,736,757]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.24). Eyes small and relatively flat, tempora oblique, about three fourth as long as diameter of eye or somewhat shorter in female
<typeStatus id="F40888130D0FFFF25310E925FE3D3E52" box="[359,446,793,814]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
(dorsally seen), rectilinearly converging to neck. Disc smooth, laterad with small, weakly impressed and widely spaced punctures. Labrum strongly and somewhat angulately incised at anterior margin, clypeus weakly semicircularly incised, with anterior angles with one setiferous pore puncture each. Mentum tooth small, submentum with one long seta laterally each side. Excision of dorsal lateral edge of right mandible well developed, almost semicircular, relatively close to base of mandible, basal angle of excision situated distinctly anterior to level of anterior margin of clypeus (with mandibles closed).
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E9D4FC1D380D" blockId="18.[178,1085,1000,1393]" pageId="18" pageNumber="1801">
Pronotum (
<figureCitation id="B3882A340D0FFFF2535EE9D4FEED3E82" box="[297,366,1000,1022]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4</figureCitation>
): Transverse (PW/PL in males 1.52-1.58, in
<typeStatus id="F40888130D0FFFF2512DE9D4FC303E82" box="[858,947,1000,1022]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.52; in the female
<typeStatus id="F40888130D0FFFF25374EE39FED93966" box="[259,346,1029,1050]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.50), widest at about end of anterior third, where the lateral seta inserted. Anterior margin a little wider than posterior margin, only weakly excavate, anterior angles moderately projecting forward, relatively narrowly rounded at tip. Sides moderately curved apicad, basad from widest point almost rectilinear at a short distance, and anterior to posterior angles gently convexly narrowed. Posterior angles somewhat obtuse-angled, widely rounded at tip. Basal edge almost rectlinear between the basal foveae, posterior angles very weakly shifted backwards. Basal foveae short, elongate, distinctly converging basad, shallowly or more strongly impressed, some punctures of unequal size in and around fovae, more or less widely spaced and continuing to the area of posterior angles. Disc convex, area of posterior angles more or less strongly flattened, anterior and posterior transverse impressions very shallow or only indicated, median line weakly impressed, terminated anterior to apical margin and hardly reaching basal margin. Lateral gutter narrow, somewhat widened toward anterior angles.
</paragraph>
<paragraph id="2B0C36B10D0FFFF352C5EFBCFC6C3C1E" blockId="18.[178,1086,1408,1601]" lastBlockId="19.[178,1085,218,354]" lastPageId="19" lastPageNumber="1802" pageId="18" pageNumber="1801">
Elytra (
<figureCitation id="B3882A340D0FFFF25288EFBCFEBC38EA" box="[255,319,1408,1430]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4</figureCitation>
): short-oval (EL/EW in males 1.39-1.43, in
<typeStatus id="F40888130D0FFFF25171EFBCFCDC38EA" box="[774,863,1408,1430]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.39; in female
<typeStatus id="F40888130D0FFFF2567EEFBCFF5D38CE" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.42), somewhat wider than pronotum (EW/PW in males 1.08-1.11, in
<typeStatus id="F40888130D0FFFF25193EFA0FBBE38CE" box="[996,1085,1436,1458]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.11; in female
<typeStatus id="F40888130D0FFFF25324EF85FE2938B2" box="[339,426,1465,1486]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.12). On disc moderately convex, humeri distinctly developed, rounded at tip, without humeral tooth, not fused at suture; toward behind somewhat convexely enlarged, widest about in middle. Basal bead weakly sinuate, arcuately curving inside humerus and weakly angled or round towards lateral margin. Scutellar pore puncture present, scutellar stria long. Striae fine, smooth, intervals flat on disc, becoming convex toward elytral declivity, interval 3 without pore puncture (
<specimenCount id="3DB5FD380D0EFFF351AFEAE6FF683C70" pageId="19" pageNumber="1802" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
),
<typeStatus id="F40888130D0EFFF35377EACBFEDB3C71" box="[256,344,247,269]" pageId="19" pageNumber="1802" type="holotype">holotype</typeStatus>
with pore puncture on right elytron only at about beginning of apical fourth close to stria 2. Interval 5 with one to tree pore punctures apically, interval 7 with two to four pore punctures apically (
<typeStatus id="F40888130D0EFFF35051EB0CFD073C3A" box="[550,644,304,326]" pageId="19" pageNumber="1802" type="holotype">holotype</typeStatus>
), or such punctures lacking (remaining
<specimenCount id="3DB5FD380D0EFFF35660EB0CFE9B3C1E" pageId="19" pageNumber="1802" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
). Preapical sinuation weak in males, only suggested in female
<typeStatus id="F40888130D0EFFF351E6EB71FC6F3C1E" box="[913,1004,333,354]" pageId="19" pageNumber="1802" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EB4DFCAF3CFB" blockId="19.[178,812,369,391]" box="[178,812,369,391]" pageId="19" pageNumber="1802">Hindwings: Reduced, about one third as long as elytral length.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EBA9FDC83F2A" blockId="19.[178,1085,405,599]" pageId="19" pageNumber="1802">Ventral surface: Prosternum with short setae, apex of prosternal process with long setae, proepisternum and anterior part of mesepisternum with sparse, fine setae, metepisternum and metasternum (laterally) with sparse, coarse seta-bearing punctures. Metepisternum very short, ratio of anterior margin/internal margin (visible parts) about 0.91, moderately narrowed behind. Abdominal sternites III-V with scattered setae of unequal length. Last sternite, beside some fine scattered setae, with two pore punctures bearing a long seta on each side at apical margin in both sexes.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E85AFB8D3E75" blockId="19.[178,1086,613,778]" pageId="19" pageNumber="1802">
Legs: Normal for
<taxonomicName id="ECB34D320D0EFFF35303E859FDAE3F07" box="[372,557,613,635]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D0EFFF35303E859FE513F06" box="[372,466,613,634]" italics="true" pageId="19" pageNumber="1802">Acinopus</emphasis>
species.
</taxonomicName>
Pro- and mesotarsi moderately widened in males (less wider than in
<taxonomicName id="ECB34D320D0EFFF353F6E8BEFD8B3FEB" authorityName="MENETRIES" authorityYear="1832" box="[385,520,642,663]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D0EFFF353F6E8BEFD8B3FEB" box="[385,520,642,663]" italics="true" pageId="19" pageNumber="1802">A. laevigatus</emphasis>
</taxonomicName>
), pro- and mesotarsomere 1 with two small adhesive scales apically, pro- and mesotarsomere 2-4 with biseriate adhesive vestiture. Ventroapical tubercle of protibia with three to five spines, arranged in a transverse row. Outer distal margin of fore tibia with five to eight spines. Eight to nine spines on lower surface of protibia arranged in one row. Tarsi smooth on superior surface except obligatory setae.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E924FCC13EF8" blockId="19.[178,1085,792,900]" pageId="19" pageNumber="1802">Microsculpture of surface: Head and pronotum in males with transverse to almost isodiametric meshes weakly engraved and somewhat irregular, elytra with about the same kind of meshes somewhat more strongly engraved as on the forebody, surface very shiny, in females meshes somewhat stronger engraved, surface less shiny.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E9AEFC2C3E82" blockId="19.[178,1085,914,1022]" pageId="19" pageNumber="1802">
Median lobe of aedeagus (
<figureCitation id="B3882A340D0EFFF353BBE9AEFDC93ED5" box="[460,586,914,937]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="19" pageNumber="1802">Figs 11, 12</figureCitation>
): Of normal construction for
<taxonomicName id="ECB34D320D0EFFF351F2E9AFFBB93ED4" box="[901,1082,914,936]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D0EFFF351F2E9AFFC603ED4" box="[901,995,915,936]" italics="true" pageId="19" pageNumber="1802">Acinopus</emphasis>
species
</taxonomicName>
, middle and apical part somewhat bent to the left, terminal lamella wide and flat, somewhat elongate-triangular (dorsal view), and somewhat reflexed apically (lateral view), internal sac with a small, elongate group of about ten medium-sized spines.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EE31FD343904" blockId="19.[178,1085,1037,1144]" pageId="19" pageNumber="1802">
Female genitalia (
<figureCitation id="B3882A340D0EFFF3531CEE31FE3B395F" box="[363,440,1037,1059]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="19" pageNumber="1802">Fig. 16</figureCitation>
): Hemisternite asymmetrical, with numerous scattered setae in its internal, somewhat hyaline part. Gonocoxite 1 elongate, apically with numerous setae, gonocoxite 2 scoop-shaped, apically somewhat blunt, with a double-sensilla in a furrow, and with numerous fine long setae at concave side.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EEBBFCAC3872" blockId="19.[178,1085,1159,1295]" pageId="19" pageNumber="1802">
I n t r a s p e c i f i c v a r i a b i l i t y: The material is too small to make an extended statement on this item. Concerning the
<specimenCount id="3DB5FD380D0EFFF350B7EE9FFCE539C5" box="[704,870,1187,1209]" pageId="19" pageNumber="1802" type="generic">three specimens</specimenCount>
examined it can be noted that the body size varies somewhat and that the elytra can have or have not one pore puncture in interval 3, with the possibility that only one elytron has this puncture, also the number of pore punctures in interval 5 and 7 can vary.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EF22FCD33B48" blockId="19.[178,1086,1309,1588]" pageId="19" pageNumber="1802">
C o m p a r i s o n s: The species is closest to
<taxonomicName id="ECB34D320D0EFFF350DCEF22FC94384E" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[683,791,1309,1331]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="brittoni" status="sp. nov.">
<emphasis id="19C7EAA30D0EFFF350DCEF22FC94384E" box="[683,791,1309,1331]" italics="true" pageId="19" pageNumber="1802">A. brittoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D0EFFF35156EF22FCE8384F" box="[801,875,1310,1331]" pageId="19" pageNumber="1802" rank="species">nov.sp.</taxonomicNameLabel>
in having the same body colour, a somewhat flattened body, a clypeus with anterior angles with one setiferous pore puncture each, and the same position of the excision of the dorsal lateral edge of the right mandible, but differs in smaller body size, a wider head (compared with pronotum), a longer pronotum with a somewhat narrower lateral gutter and with less protruding anterior angles, and with a less strongly flattened area of the posterior angles. In addition, the elytra of the new species are somewhat longer and a little less oval, the median lobe of the aedeagus is somewhat shorter and stouter, the apical lamella a little longer and almost evenly narrowed, apically more acute (dorsal view), somewhat reflexed apically (lateral view), the spines of the internal sac are lower in number (compare
<figureCitation id="B3882A340D0EFFF35011EC22FD5A3B4F" box="[614,729,1566,1588]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="19" pageNumber="1802">Figs 11, 12</figureCitation>
, and 9, 10).
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5EAE7FDC43CE7" blockId="20.[178,1085,218,411]" pageId="20" pageNumber="1803">
H a b i t a t: Nothing is known about biotope and circumstances of collecting. Al Soudah is situated west of Abah City on the Sirwat Mountains, a massif with heights of more than
<quantity id="EC4B9B540D09FFF45352EB2FFEF73C55" box="[293,372,275,297]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="1803" unit="m" value="3000.0">3000 m</quantity>
a.s.l. After a source in the internet (http://www.the-saudi.net/saudiarabia/abha/abha_city.htm) the Abha region has the highest level of rainfall of any part of
<collectingCountry id="53A476210D09FFF452A3EB70FEE13C1E" box="[212,354,332,354]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, the mountains are covered with stands of Araar trees,
<taxonomicName id="ECB34D320D09FFF451B8EB70FE613C02" authority="(VAHL) MASTERS" authorityName="MASTERS" baseAuthorityName="VAHL" class="Pinopsida" family="Cupressaceae" genus="Tetraclinis" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="1803" phylum="Tracheophyta" rank="species" species="articulata">
<emphasis id="19C7EAA30D09FFF451B8EB70FE953C02" italics="true" pageId="20" pageNumber="1803">Tetraclinis articulata</emphasis>
(VAHL) MASTERS
</taxonomicName>
, an evergreen coniferous tree of the cypress (family
<taxonomicName id="ECB34D320D09FFF452C5EBB9FEBD3CE7" box="[178,318,389,411]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="1803" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
) and other natural forests.
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5EB96FCD73C85" blockId="20.[178,1085,426,505]" pageId="20" pageNumber="1803">
D i s t r i b u t i o n: Up to present only known from one locality in the Sirwat Mountains (
<collectingCountry id="53A476210D09FFF45283EBFAFE073CA0" box="[244,388,454,476]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
), situated in the Sarat '
<collectingRegion id="E977F8530D09FFF450F7EBFAFD793CA0" box="[640,762,454,476]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir region</collectingRegion>
in the middle of the Sarawat Mountain range, but surely more widely distributed in that massif.
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5E834FC4E3824" blockId="20.[178,1086,519,1369]" pageId="20" pageNumber="1803">
R e m a r k s: The Sarawat mountain range, a massif running parallel to the western coast of the Arabian Peninsula and is among the Peninsula's most prominent geographical features, starts from the border of
<collectingCountry id="53A476210D09FFF45038E87CFD113F2A" box="[591,658,576,598]" name="Jordan" pageId="20" pageNumber="1803">Jordan</collectingCountry>
in the north to the Gulf of Aden in the south, running through
<collectingCountry id="53A476210D09FFF453DEE861FDB13F0F" box="[425,562,605,627]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
and
<collectingCountry id="53A476210D09FFF4501EE861FD363F0E" box="[617,693,605,626]" name="Yemen" pageId="20" pageNumber="1803">Yemen</collectingCountry>
. The range's northern half, known as Sarat al-Hejaz rarely rises about 2100 meters, while the middle and southern portions (Sarat '
<collectingRegion id="E977F8530D09FFF4528DE8AAFEAB3FD0" box="[250,296,662,684]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir</collectingRegion>
and Sarat al-Yemen, respectively) can reach heights of over 3,300 meters. The discovery of two members of the subgenus
<taxonomicName id="ECB34D320D09FFF450D3E88FFC813FB4" authorityName="DEJEAN" authorityYear="1821" box="[676,770,691,712]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D09FFF450D3E88FFC813FB4" box="[676,770,691,712]" italics="true" pageId="20" pageNumber="1803">Acinopus</emphasis>
</taxonomicName>
in western
<collectingCountry id="53A476210D09FFF4510CE88FFB873FB5" box="[891,1028,691,713]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, both similar to the North African
<taxonomicName id="ECB34D320D09FFF453ACE8F3FDDF3F98" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[475,604,719,740]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF453ACE8F3FDDF3F98" box="[475,604,719,740]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
, is very interesting in a zoogeographical point of view. There is a wide gap of about thousand kilometres between eastern populations of
<taxonomicName id="ECB34D320D09FFF452A5E935FED53E61" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[210,342,776,798]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF452A5E935FED53E61" box="[210,342,776,798]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
from
<collectingCountry id="53A476210D09FFF453EBE934FE543E62" box="[412,471,776,798]" name="Libya" pageId="20" pageNumber="1803">Libya</collectingCountry>
and the populations of two new species in
<collectingCountry id="53A476210D09FFF451C3E934FBBE3E62" box="[948,1085,776,798]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
(from
<collectingCountry id="53A476210D09FFF45284E919FEB13E46" box="[243,306,805,826]" name="Egypt" pageId="20" pageNumber="1803">Egypt</collectingCountry>
, in fact from the
<collectingRegion id="E977F8530D09FFF4539FE919FD0A3E47" box="[488,649,805,827]" country="Egypt" name="Shamal Sina'" pageId="20" pageNumber="1803">Sinai Peninsula</collectingRegion>
, only one rather dissimilar and apparently unrelated species,
<taxonomicName id="ECB34D320D09FFF45318E97EFE6E3E2B" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[367,493,834,855]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D09FFF45318E97EFE6E3E2B" box="[367,493,834,855]" italics="true" pageId="20" pageNumber="1803">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D09FFF45383E97EFDC33E2B" box="[500,576,834,855]" pageId="20" pageNumber="1803" rank="species">nov.sp.</taxonomicNameLabel>
, is known). It can probably be hypothetizied, due to the relative similarity of
<taxonomicName id="ECB34D320D09FFF453B8E962FDD33E0F" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[463,592,862,883]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF453B8E962FDD33E0F" box="[463,592,862,883]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
and the two species from
<collectingCountry id="53A476210D09FFF45113E962FC6D3E08" box="[868,1006,862,884]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, that
<taxonomicName id="ECB34D320D09FFF45650E962FE953EF3" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF45650E962FE953EF3" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
on the one hand and the two Arabian species on the other hand are adelphotaxa with an ancestor widely distributed over North Africa and the Arabian Peninsula in the Neogene, most likely not earlier than the middle Miocene, because in the Paleogene and the early Miocene, North Africa was dominated by tropical forests; by the middle Miocene grass-dominated ecosystems begin to spread and vegetation changed into a more open grassland vegetation towards the end of the Miocene (e.g.,
<bibRefCitation id="4F224B400D09FFF451EFEE35FE9F3947" author="MICHEELS A &amp; ERONEN J" pageId="20" pageNumber="1803" pagination="193 - 204" refId="ref14796" refString="MICHEELS A., ERONEN J. &amp; V. MOSBRUGGER (2009): The Late Miocene climate response to a modern Sahara desert. - Global and Planetary Change 67: 193 - 204." type="journal article" year="2009">MICHEELS et al. 2009: 193</bibRefCitation>
;
<bibRefCitation id="4F224B400D09FFF45350EE1AFD913947" author="JACOBS B &amp; PAN A" box="[295,530,1061,1083]" pageId="20" pageNumber="1803" pagination="57 - 72" refId="ref14487" refString="JACOBS B. F., PAN A. D. &amp; C. R. SCOTESE (2010): A review of the Cenozoic vegetation history of Africa. Pp. 57 - 72. - In: WERDELIN L. &amp; W. SANDERS (eds), Cenozoic Mammals of Africa. Berkeley: University of California Press, 986 pp." type="book chapter" year="2010">JACOBS et al. 2010: 69</bibRefCitation>
). The continuous distribution of the common ancestor of the species discussed was probably divided into the African and Arabian parts by lowland desert areas appeared for the first time in North Africa at around the Miocene/Pliocene boundary (e.g.,
<bibRefCitation id="4F224B400D09FFF45064EE40FD5B39ED" author="LE HOUEROU H" box="[531,728,1147,1169]" pageId="20" pageNumber="1803" pagination="619 - 647" refId="ref14711" refString="LE HOUEROU H. N. (1997): Climate, flora and fauna changes in the Sahara over the past 500 million years. - Journal of Arid Environments 37: 619 - 647." type="journal article" year="1997">LE HOUEROU 1997</bibRefCitation>
). This separation was aided by the Red Sea which formation began in the Eocene and became a sea in the Miocene, causing the isolation of the mountains of South-West Arabia from those of Africa (
<bibRefCitation id="4F224B400D09FFF45195EE88FE8D399B" author="BRITTON E" pageId="20" pageNumber="1803" pagination="126 - 131" refId="ref14287" refString="BRITTON E. B. (1948): Coleoptera: Cicindelidae and Carabidae. Pp. 87 - 125, pl. 7 - 9. - In: Expedition to South-west Arabia 1937 - 8, Vol. 1. London: British Museum (Natural History), pp. 67 - 178, pl. 4 - 9 [with an appendix by P. BASILEWSKY: Trois nouveaux Carabiques du Sud-Ouest de l'Arabie, pp. 126 - 131]." type="book chapter" year="1948">BRITTON 1948: 89</bibRefCitation>
). The result of alternating dry and damp climatic periods in a region today drier than in the past, caused a further splitting in two species with a relict-like occurrence, which need for surviving plants and herewith connected moisture, in the high mountains of western Arabia, in one species in the Sarat al-Hejaz in the northern portion of the Sarawat mountain range, and a second one in the Sarat '
<collectingRegion id="E977F8530D09FFF45091EF7FFC973825" box="[742,788,1347,1369]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir</collectingRegion>
in its middle part.
</paragraph>
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@ -0,0 +1,335 @@
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<mods:title id="D44246D532327555F7540A48EF7555E0">Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)</mods:title>
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<taxonomicName id="ECB34D320D16FFEB52C5E96DFE7A3E1A" ID-CoL="9BVX" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[178,505,848,870]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov." subGenus="Acinopus">
<emphasis id="19C7EAA30D16FFEB52C5E96DFE973E1A" bold="true" box="[178,276,849,870]" italics="true" pageId="11" pageNumber="1794">Acinopus</emphasis>
(
<emphasis id="19C7EAA30D16FFEB5354E96DFE063E1A" bold="true" box="[291,389,849,870]" italics="true" pageId="11" pageNumber="1794">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D16FFEB53E2E96DFE7A3E1A" bold="true" box="[405,505,849,870]" italics="true" pageId="11" pageNumber="1794">sinaiticus</emphasis>
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(
<figureCitation id="B3882A340D16FFEB502FE96CFD183E19" box="[600,667,848,870]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="11" pageNumber="1794">Figs 2</figureCitation>
,
<figureCitation id="B3882A340D16FFEB50D0E96DFD4E3E19" box="[679,717,848,870]" captionStart="Figs 5-8" captionStartId="6.[178,218,1273,1295]" captionTargetBox="[198,1071,235,1238]" captionTargetId="figure-0@6.[178,1085,204,1274]" captionTargetPageId="6" captionText="Figs 5-8: Acinopus DEJEAN, 1821. Median lobe of male. (5, 6) A. orszuliki nov.sp. (5) Lateral view (holotype). (6) dorsal view (paratype, type locality). (7, 8) A. sinaiticus nov.sp. (7) Lateral view (holotype). (8) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="11" pageNumber="1794">7, 8</figureCitation>
,
<figureCitation id="B3882A340D16FFEB50AEE96CFD703E1A" box="[729,755,848,870]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="11" pageNumber="1794">14</figureCitation>
)
</emphasis>
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</paragraph>
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<paragraph id="2B0C36B10D16FFEB52C5E94BFD363EF0" blockId="11.[178,704,886,987]" box="[178,693,886,908]" pageId="11" pageNumber="1794">
<treatmentCitationGroup id="0BA3119F0D16FFEB52C5E94BFD363EF0" box="[178,693,886,908]" pageId="11" pageNumber="1794">
<taxonomicName id="ECB34D320D16FFEB52C5E94BFE583EF6" ID-CoL="9BVN" authority="MENETRIES" authorityName="MENETRIES" authorityYear="1832" box="[178,475,886,908]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D16FFEB52C5E94BFEEA3EF7" box="[178,361,886,908]" italics="true" pageId="11" pageNumber="1794">Acinopus laevigatus</emphasis>
MÉNÉTRIÉS
</taxonomicName>
:
<treatmentCitation id="AA1210A00D16FFEB5390E94BFD363EF0" author="PEYERIMHOFF P" box="[487,693,886,908]" page="6" pageId="11" pageNumber="1794" year="1907">
<bibRefCitation id="4F224B400D16FFEB5390E94BFD363EF0" author="PEYERIMHOFF P" box="[487,693,886,908]" pageId="11" pageNumber="1794" pagination="1907 - 1923" refId="ref14833" refString="PEYERIMHOFF P. de (1907): Liste des coleopteres du Sinai. - L`Abeille. Journal d'Entomologie 31 [1907 - 1923]: 1 - 55." type="journal article" year="1907">PEYERIMHOFF 1907: 6</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5E9ADFD433EDA" blockId="11.[178,704,886,987]" box="[178,704,912,934]" pageId="11" pageNumber="1794">
<treatmentCitationGroup id="0BA3119F0D16FFEB52C5E9ADFD433EDA" box="[178,704,912,934]" pageId="11" pageNumber="1794">
<taxonomicName id="ECB34D320D16FFEB52C5E9ADFE583ED9" ID-CoL="9BVN" authority="MENETRIES" authorityName="MENETRIES" authorityYear="1832" box="[178,475,912,934]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D16FFEB52C5E9ADFEEA3ED9" box="[178,361,912,934]" italics="true" pageId="11" pageNumber="1794">Acinopus laevigatus</emphasis>
MÉNÉTRIÉS
</taxonomicName>
:
<treatmentCitation id="AA1210A00D16FFEB5390E9ADFD433EDA" author="SCHATZMAYR A" box="[487,704,912,934]" page="76" pageId="11" pageNumber="1794" year="1936">
<bibRefCitation id="4F224B400D16FFEB5390E9ADFD433EDA" author="SCHATZMAYR A" box="[487,704,912,934]" pageId="11" pageNumber="1794" pagination="1 - 114" refId="ref14863" refString="SCHATZMAYR A. (1936): Risultati scientifici della spedizione entomologica di S. A. S. il Principe Alessandro della Torre e Tasso in Egitto e nella penisola del Sinai. XII. Catalogo ragionato dei Carabidi finora noti d'Egitto e del Sinai. - Publicazioni del Museo Entomologico &quot; Pietro Rossi &quot;, Duino 1: 1 - 114." type="journal article" year="1936">SCHATZMAYR 1936: 76</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5E997FD0B3EBC" blockId="11.[178,704,886,987]" box="[178,648,939,961]" pageId="11" pageNumber="1794">
<treatmentCitationGroup id="0BA3119F0D16FFEB52C5E997FD0B3EBC" box="[178,648,939,961]" pageId="11" pageNumber="1794">
<taxonomicName id="ECB34D320D16FFEB52C5E997FE583EC3" ID-CoL="9BVN" authority="MENETRIES" authorityName="MENETRIES" authorityYear="1832" box="[178,475,939,960]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D16FFEB52C5E997FEEA3EBC" box="[178,361,939,960]" italics="true" pageId="11" pageNumber="1794">Acinopus laevigatus</emphasis>
MÉNÉTRIÉS
</taxonomicName>
:
<treatmentCitation id="AA1210A00D16FFEB5390E997FD0B3EBC" author="ALFIERI A" box="[487,648,939,961]" page="21" pageId="11" pageNumber="1794" year="1976">
<bibRefCitation id="4F224B400D16FFEB5390E997FD0B3EBC" author="ALFIERI A" box="[487,648,939,961]" pageId="11" pageNumber="1794" pagination="1 - 361" refId="ref14125" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xiii, 1 - 361." type="book chapter" year="1976">ALFIERI 1976: 21</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5E9F9FD353EA7" blockId="11.[178,704,886,987]" box="[178,694,965,987]" pageId="11" pageNumber="1794">
<treatmentCitationGroup id="0BA3119F0D16FFEB52C5E9F9FD353EA7" box="[178,694,965,987]" pageId="11" pageNumber="1794">
<taxonomicName id="ECB34D320D16FFEB52C5E9F9FE583EA5" ID-CoL="9BVN" authority="MENETRIES" authorityName="MENETRIES" authorityYear="1832" box="[178,475,965,986]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D16FFEB52C5E9F9FEEA3EA6" box="[178,361,965,986]" italics="true" pageId="11" pageNumber="1794">Acinopus laevigatus</emphasis>
MÉNÉTRIÉS
</taxonomicName>
:
<treatmentCitation id="AA1210A00D16FFEB5390E9F9FD353EA7" author="HOSNI M &amp; SHARKAWY A &amp; OSHAIBAH A &amp; HASSAN M" box="[487,694,965,987]" page="561" pageId="11" pageNumber="1794" year="2003">
<bibRefCitation id="4F224B400D16FFEB5390E9F9FD353EA7" author="HOSNI M &amp; SHARKAWY A &amp; OSHAIBAH A &amp; HASSAN M" box="[487,694,965,987]" pageId="11" pageNumber="1794" pagination="549 - 573" refId="ref14416" refString="HOSNI M. T., AL- SHARKAWY A. Z., OSHAIBAH A. A., HASSAN M. I. &amp; A. A. EL- GHARBAWY (2003): Classification of subfamily Harpalinae (family: Carabidae - order: Coleoptera) in Egypt. II - tribes Harpalini, Masoreini and Amarini. - Egyptian Journal of Entomology 40: 549 - 573." type="journal article" year="2003">HOSNI et al. 2003: 561</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5E9D0FCB33921" blockId="11.[178,1086,1003,1117]" pageId="11" pageNumber="1794">
T y p e m a t e r i a l:
<typeStatus id="F40888130D16FFEB53D0E9D0FE7F397E" box="[423,508,1004,1026]" pageId="11" pageNumber="1794" type="holotype">Holotype</typeStatus>
6, labelled: &quot;Sinai /
<collectingCountry id="53A476210D16FFEB50CAE9D0FC8E397D" box="[701,781,1004,1025]" name="Israel" pageId="11" pageNumber="1794">ISRAEL</collectingCountry>
/ W[adi El]. Arbain /
<date id="5F0D10710D16FFEB51A2E9D0FBB3397D" box="[981,1072,1004,1025]" pageId="11" pageNumber="1794" value="1974-04-04">4.IV.1974</date>
/ D.Furth&quot; (TAU).
<typeStatus id="F40888130D16FFEB530AEE3FFE5A3964" box="[381,473,1027,1048]" pageId="11" pageNumber="1794" type="paratype">Paratypes</typeStatus>
: 266,
<specimenCount id="3DB5FD380D16FFEB5054EE3FFDB93961" box="[547,570,1027,1053]" pageId="11" pageNumber="1794" type="female">1♀</specimenCount>
, labelled: &quot;Sinai
<quantity id="EC4B9B540D16FFEB5090EE3EFCAC3964" box="[743,815,1026,1048]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="1794" unit="m" value="2500.0">2500 m</quantity>
/
<collectingCountry id="53A476210D16FFEB5131EE3FFC163964" box="[838,917,1027,1048]" name="Israel" pageId="11" pageNumber="1794">ISRAEL</collectingCountry>
/ Mt. Katherine /
<date id="5F0D10710D16FFEB52AFEE25FEB73952" box="[216,308,1049,1070]" pageId="11" pageNumber="1794" value="1974-04-08">8.IV.1974</date>
/ D.Furth&quot; (TAU, ZIN, cWR).
<specimenCount id="3DB5FD380D16FFEB501CEE25FD00394F" box="[619,643,1049,1075]" pageId="11" pageNumber="1794" type="female">1♀</specimenCount>
, labelled: &quot;
<taxonomicName id="ECB34D320D16FFEB508DEE25FC443953" ID-CoL="9BVN" authority="Men" authorityName="MENETRIES" authorityYear="1832" box="[762,967,1049,1071]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="laevigatus">Acin. laevigatus Mén</taxonomicName>
/ Sinai: Dj. Safsafeh /
<date id="5F0D10710D16FFEB5343EE0CFE133939" box="[308,400,1072,1093]" pageId="11" pageNumber="1794" value="1902-02-22">22.2.1902</date>
&quot; (handwritten by Peyerimhoff, species name in black, locality and date in red), &quot;Bedel / vidit (handwritten in red by Peyerimhoff) (MNHN).
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D16FFEE52C5EE57FD703CDC" lastPageId="14" lastPageNumber="1797" pageId="11" pageNumber="1794" type="description">
<paragraph id="2B0C36B10D16FFEB52C5EE57FDC139E2" blockId="11.[178,1085,1130,1182]" pageId="11" pageNumber="1794">E t y m o l o g y: The specific epithet is a latinizid adjective, based on the name of the region in which this species was found.</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5EE92FBA438B6" blockId="11.[178,1085,1197,1483]" pageId="11" pageNumber="1794">
D i a g n o s i s: A macropterous species of small to medium size for
<taxonomicName id="ECB34D320D16FFEB51DFEE91FB8539BE" authorityName="DEJEAN" authorityYear="1821" box="[936,1030,1197,1218]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D16FFEB51DFEE91FB8539BE" box="[936,1030,1197,1218]" italics="true" pageId="11" pageNumber="1794">Acinopus</emphasis>
</taxonomicName>
with excision of dorsal edge of right mandible (subgenus
<taxonomicName id="ECB34D320D16FFEB50C5EEF7FC8F399C" authorityName="DEJEAN" authorityYear="1821" box="[690,780,1227,1248]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D16FFEB50C5EEF7FC8F399C" box="[690,780,1227,1248]" italics="true" pageId="11" pageNumber="1794">Acinopus</emphasis>
</taxonomicName>
), dark rufopiceous (in mature condition), with appendages somewhat lighter, labrum strongly incised at anterior margin, clypeus with one setiferous pore puncture at anterior angles, pronotum almost cordate with base rectilinear throughout, toward the posterior angles rectilinearly and anterior to posterior angles weakly sinuately narrowed, anterior angles acutely or more or less roundly protruding, posterior angles obtuse-angled, relatively narrowly rounded at tip, elytra cylindrical, somewhat flattened on disc, metepisternum slightly longer than wide. Pro- and mesotarsi in males distinctly widened. Apical lamella of median lobe moderately long, about somewhat elongate-triangular (dorsal view). Habitus see
<figureCitation id="B3882A340D16FFEB5191EF89FBA438B6" box="[998,1063,1461,1483]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="11" pageNumber="1794">Fig. 2.</figureCitation>
</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5EFE7FDDE3B57" blockId="11.[178,1085,1498,1579]" pageId="11" pageNumber="1794">
D e s c r i p t i o n: Body length in males
<quantity id="EC4B9B540D16FFEB500FEFE6FC8F388C" box="[632,780,1498,1520]" metricMagnitude="-2" metricUnit="m" metricValue="1.345" metricValueMax="1.53" metricValueMin="1.16" pageId="11" pageNumber="1794" unit="mm" value="13.45" valueMax="15.3" valueMin="11.6">11.6-15.3 mm</quantity>
, in
<typeStatus id="F40888130D16FFEB5142EFE6FC0D388C" box="[821,910,1498,1520]" pageId="11" pageNumber="1794" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D16FFEB51E1EFE6FC77388C" box="[918,1012,1498,1520]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="11" pageNumber="1794" unit="mm" value="14.6">14.6 mm</quantity>
, in the female
<typeStatus id="F40888130D16FFEB528AEFC4FEDD3B71" box="[253,350,1528,1549]" pageId="11" pageNumber="1794" type="paratype">paratypes</typeStatus>
13.2 and
<quantity id="EC4B9B540D16FFEB53B4EFC4FDA23B71" box="[451,545,1528,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="11" pageNumber="1794" unit="mm" value="15.1">15.1 mm</quantity>
; width in males
<quantity id="EC4B9B540D16FFEB50B0EFC4FCBC3B71" box="[711,831,1528,1550]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.3" metricValueMin="4.3" pageId="11" pageNumber="1794" unit="mm" value="4.8" valueMax="5.3" valueMin="4.3">4.3-5.3 mm</quantity>
, in
<typeStatus id="F40888130D16FFEB5114EFC4FC3F3B72" box="[867,956,1528,1550]" pageId="11" pageNumber="1794" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D16FFEB51B5EFC4FB9C3B71" box="[962,1055,1528,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="11" pageNumber="1794" unit="mm" value="15.1">15.1 mm</quantity>
, in the female
<typeStatus id="F40888130D16FFEB5355EC2AFE003B57" box="[290,387,1558,1579]" pageId="11" pageNumber="1794" type="paratype">paratypes</typeStatus>
<quantity id="EC4B9B540D16FFEB53FDEC29FE543B57" box="[394,471,1557,1579]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="1794" unit="mm" value="5.6">5.6 mm</quantity>
and
<quantity id="EC4B9B540D16FFEB507EEC29FDD93B57" box="[521,602,1557,1579]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="11" pageNumber="1794" unit="mm" value="5.9">5.9 mm</quantity>
.
</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5EAE7FF7F3C57" blockId="12.[178,1085,219,299]" pageId="12" pageNumber="1795">Colour: Lighter to darker rufo-piceous or piceous also in mature condition (never completely black, if dark always with a piceous ground colour), appendages somewhat lighter.</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5EB07FC273F08" blockId="12.[178,1086,315,628]" pageId="12" pageNumber="1795">
Head: wide (as normal in
<taxonomicName id="ECB34D320D11FFEC53B5EB07FDA33C2C" authorityName="DEJEAN" authorityYear="1821" box="[450,544,315,336]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1795" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D11FFEC53B5EB07FDA33C2C" box="[450,544,315,336]" italics="true" pageId="12" pageNumber="1795">Acinopus</emphasis>
</taxonomicName>
), only somewhat narrower than pronotum (PW/HW in males 1.13-1.20, average 1.17, in
<typeStatus id="F40888130D11FFEC504AEB64FD153C12" box="[573,662,344,366]" pageId="12" pageNumber="1795" type="holotype">holotype</typeStatus>
1.13; in the female
<typeStatus id="F40888130D11FFEC5102EB64FC553C11" box="[885,982,344,365]" pageId="12" pageNumber="1795" type="paratype">paratypes</typeStatus>
1.16 and 1.20). Eyes small and relatively flat, tempora oblique, about three fourth as long as diameter of eye or somewhat shorter (dorsally seen), rectilinearly converging to neck. Frontal fovae mostly small and shallow, sometimes elongately prolonged backwards. Disc smooth. Excision of dorsal lateral edge of right mandible relatively small, almost semicircular, relatively close to base of mandible, proximal (basal) angle of excision situated somewhat anterior to level of anterior margin of clypeus (with mandibles closed). Labrum strongly and angulately incised at anterior margin, clypeus weakly, somewhat angulately incised, with anterior angles with one setiferous pore puncture each. Mentum tooth small, submentum with one long and one to three short, fine setae laterally each side.
</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5E8B8FC143E85" blockId="12.[178,1085,643,1017]" pageId="12" pageNumber="1795">
Pronotum (
<figureCitation id="B3882A340D11FFEC5353E8BFFEE03FE5" box="[292,355,643,665]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="12" pageNumber="1795">Fig. 2</figureCitation>
): Transverse, almost cordate (PW/PL in males 1.47-1.57, average 1.53, in
<typeStatus id="F40888130D11FFEC52BAE89CFEA53FCA" box="[205,294,672,694]" pageId="12" pageNumber="1795" type="holotype">holotype</typeStatus>
1.55; in the female
<typeStatus id="F40888130D11FFEC5380E89DFDDB3FCA" box="[503,600,673,694]" pageId="12" pageNumber="1795" type="paratype">paratypes</typeStatus>
1.54 and 1.61), widest at about end of anterior third, there lateral seta inserted. Disc convex, area of posterior angles more or less strongly flattened, anterior and posterior transverse impressions very shallow or only indicated, median line indistinctly impressed, terminated anterior to transverse impressions and hardly reaching basal margin. Basal foveae short, elongate, distinctly converging basad, shallowly or more strongly impressed. Anterior margin a little wider than posterior margin, only weakly, somewhat angulately excavate, anterior angles moderately projecting forward, relatively widely rounded at tip. Lateral gutter narrow, somewhat widened toward anterior angles. Sides moderately curved apicad, basad from widest point almost rectlinearly, and anterior to posterior angles weakly sinuately narrowed. Posterior angles distinct, obtuse-angled, relatively narrowly rounded at tip. Base rectilinear throughout or between the basal foveae very weakly excavate.
</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5EE34FCD93802" blockId="12.[178,1085,1032,1406]" pageId="12" pageNumber="1795">
Elytra (
<figureCitation id="B3882A340D11FFEC5377EE34FEC33963" box="[256,320,1032,1055]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="12" pageNumber="1795">Fig. 2</figureCitation>
): Longer or shorter cylindrical (EL/EW in males 1.48-1.58, average 1.52, in
<typeStatus id="F40888130D11FFEC52A7EE1AFEAA3940" box="[208,297,1062,1084]" pageId="12" pageNumber="1795" type="holotype">holotype</typeStatus>
1.52; in female
<typeStatus id="F40888130D11FFEC53ABEE1AFDBE3947" box="[476,573,1062,1083]" pageId="12" pageNumber="1795" type="paratype">paratypes</typeStatus>
1.41 and 1.55), hardly or only somewhat wider than pronotum (EW/PW in males 1.06-1.15, average 1.10, in
<typeStatus id="F40888130D11FFEC514BEE7FFC163925" box="[828,917,1091,1113]" pageId="12" pageNumber="1795" type="holotype">holotype</typeStatus>
1.10; in female
<typeStatus id="F40888130D11FFEC52C5EE5DFE90390A" box="[178,275,1121,1142]" pageId="12" pageNumber="1795" type="paratype">paratypes</typeStatus>
1.09 and 1.17). On disc somewhat flattened, humeri distinctly developed, widely rounded at tip, or with a suggested angle meeting the lateral margin, without humeral tooth; toward behind somewhat convexely or almost parallely enlarged, widest somewhat behind middle. Basal bead weakly sinuate, arcuately curving inside humerus and weakly angled or round towards lateral margin. Scutellar pore puncture present, scutellar stria long. Striae fine, smooth, intervals flat on disc, becoming convex toward elytral declivity, interval 3 without pore puncture (
<typeStatus id="F40888130D11FFEC50B2EF2CFC9C385A" box="[709,799,1296,1318]" pageId="12" pageNumber="1795" type="holotype">holotype</typeStatus>
and
<specimenCount id="3DB5FD380D11FFEC5122EF2DFC5D385A" box="[853,990,1297,1318]" pageId="12" pageNumber="1795" type="generic" typeStatus="paratype">one paratype</specimenCount>
), or with pore puncture (
<specimenCount id="3DB5FD380D11FFEC533BEF12FE68383F" box="[332,491,1326,1348]" pageId="12" pageNumber="1795" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
) at about beginning of apical fourth close to stria 2. Interval 5 with one to two pore punctures apically (in
<specimenCount id="3DB5FD380D11FFEC50DFEF70FC9B381D" box="[680,792,1355,1377]" pageId="12" pageNumber="1795" type="female">one female</specimenCount>
<typeStatus id="F40888130D11FFEC5169EF70FCF6381D" box="[798,885,1356,1377]" pageId="12" pageNumber="1795" type="paratype">paratype</typeStatus>
lacking), interval 7 with two to four pore punctures apically. Preapical sinuation weak.
</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5EFB2FC9B38DF" blockId="12.[178,792,1422,1444]" box="[178,792,1422,1444]" pageId="12" pageNumber="1795">Hindwings: Completely developed (folded anterior to apex).</paragraph>
<paragraph id="2B0C36B10D11FFEC52C5EF8FFBBD3B42" blockId="12.[178,1086,1459,1598]" pageId="12" pageNumber="1795">Ventral surface: Prosternum and apex of prosternal process with long setae, proepisternum and anterior part of mesepisternum with sparse, fine setae, metepisternum and metasternum (laterally) with sparse, coarse seta-bearing punctures. Metepisternum relatively long, ratio of anterior margin/internal margin (visible parts) about 0.71, moderately narrowed behind. Abdominal sternites III-V with scattered setae of unequal length.</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EAE7FD1A3C71" blockId="13.[178,1085,219,270]" pageId="13" pageNumber="1796">Last sternite, beside some fine scattered setae with two pore punctures bearing a long seta on each side at apical margin in both sexes.</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EB22FE503CB9" blockId="13.[178,1086,285,454]" pageId="13" pageNumber="1796">
Legs: Normal for
<taxonomicName id="ECB34D320D10FFED530DEB22FDB73C4F" box="[378,564,285,307]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D10FFED530DEB22FE5B3C4F" box="[378,472,286,307]" italics="true" pageId="13" pageNumber="1796">Acinopus</emphasis>
species.
</taxonomicName>
Pro- and mesotarsi distinctly widened in males (about as strong as in
<taxonomicName id="ECB34D320D10FFED53D3EB07FDAF3C2C" authorityName="MENETRIES" authorityYear="1832" box="[420,556,315,336]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D10FFED53D3EB07FDAF3C2C" box="[420,556,315,336]" italics="true" pageId="13" pageNumber="1796">A. laevigatus</emphasis>
</taxonomicName>
), with pro- and mesotarsomere 2-4 with biseriate adhesive vestiture. Ventroapical tubercle of protibia with about four to five spines, arranged in a transverse row. Eight to ten spines on lower surface of protibia arranged in one row. Outer distal margin of fore tibia with six to eight spines. Tarsi smooth on superior surface except obligatory setae.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EBE9FCC13F3F" blockId="13.[178,1085,469,579]" pageId="13" pageNumber="1796">Microsculpture of surface: Head with weakly engraved somewhat irregular isodiametric meshes, pronotum and elytra with somewhat irregular slightly transverse meshes, somewhat more strongly engraved and more confused as on head, surface very shiny, in females meshes somewhat stronger engraved, surface less shiny.</paragraph>
<paragraph id="2B0C36B10D10FFED52C5E86FFCD83FBD" blockId="13.[178,1085,595,705]" pageId="13" pageNumber="1796">
Median lobe of aedeagus (
<figureCitation id="B3882A340D10FFED53B5E86FFD9D3F15" box="[450,542,595,617]" captionStart="Figs 5-8" captionStartId="6.[178,218,1273,1295]" captionTargetBox="[198,1071,235,1238]" captionTargetId="figure-0@6.[178,1085,204,1274]" captionTargetPageId="6" captionText="Figs 5-8: Acinopus DEJEAN, 1821. Median lobe of male. (5, 6) A. orszuliki nov.sp. (5) Lateral view (holotype). (6) dorsal view (paratype, type locality). (7, 8) A. sinaiticus nov.sp. (7) Lateral view (holotype). (8) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="13" pageNumber="1796">Figs 7, 8</figureCitation>
): Of normal construction for
<taxonomicName id="ECB34D320D10FFED5139E86FFB873F15" box="[846,1028,595,617]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D10FFED5139E86FFC2F3F14" box="[846,940,595,616]" italics="true" pageId="13" pageNumber="1796">Acinopus</emphasis>
species.
</taxonomicName>
Middle and apical part somewhat bent to the left, terminal lamella wide and flat, somewhat elongate-triangular (dorsal view), and somewhat reflexed apically (lateral view), internal sac with a small, elongate group of about ten medium-sized spines.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5E8ECFD343E42" blockId="13.[178,1085,720,830]" pageId="13" pageNumber="1796">
Female genitalia (
<figureCitation id="B3882A340D10FFED531CE8ECFE3B3F9A" box="[363,440,720,742]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="13" pageNumber="1796">Fig. 14</figureCitation>
): Hemisternite asymmetrical, with numerous scattered setae in its internal, somewhat hyaline part. Gonocoxite 1 elongate, apically with numerous setae, gonocoxite 2 scoop-shaped, apically somewhat blunt, with a double-sensilla in a furrow, and with numerous fine long setae at concave side.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5E972FCAC396F" blockId="13.[178,1085,845,1043]" pageId="13" pageNumber="1796">
I n t r a s p e c i f i c v a r i a b i l i t y Though the material is small, at least a statement can be made concerning the body size (varies from
<quantity id="EC4B9B540D10FFED5149E957FC573EFC" box="[830,980,875,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.345" metricValueMax="1.53" metricValueMin="1.16" pageId="13" pageNumber="1796" unit="mm" value="13.45" valueMax="15.3" valueMin="11.6">11.6-15.3 mm</quantity>
), the construction of the pronotal base (rectilinear throughout or between the basal foveae very weakly excavate), and of basal foveae (shallowly or more distinctly impressed). The elytra are longer or shorter cylindrical with interval 3 lacking pore puncture or with pore puncture at about beginning of apical fourth close to stria 2, and with interval 5 with one to two pore punctures apically or these pore punctures lacking.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EE1FFD5539B7" blockId="13.[178,1085,1059,1227]" pageId="13" pageNumber="1796">
C o m p a r i s o n s: Similar in its colour to
<taxonomicName id="ECB34D320D10FFED50FEEE1FFD7F3944" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[649,764,1059,1080]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D10FFED50FEEE1FFD7F3944" box="[649,764,1059,1080]" italics="true" pageId="13" pageNumber="1796">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D10FFED5174EE1FFCCC3944" box="[771,847,1059,1080]" pageId="13" pageNumber="1796" rank="species">nov.sp.</taxonomicNameLabel>
, but different in having pro- and mesotarsomeres distinctly enlarged, the labrum distinctly incised apically, the anterior pronotal angles not as strong and less acutely projecting forward, the pronotal base more or less rectilinear, the pronotum anterior to the posterior angles laterally somewhat sinuate, the elytra somewhat flattened, and the hind wings fully developed (folded anterior to apex), and the metepisterna longer.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EEE7FD14381A" blockId="13.[178,1085,1243,1382]" pageId="13" pageNumber="1796">
The species is distinguishable from
<taxonomicName id="ECB34D320D10FFED5051EEE7FD0F398C" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[550,652,1243,1264]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D10FFED5051EEE7FD0F398C" box="[550,652,1243,1264]" italics="true" pageId="13" pageNumber="1796">A. picipes</emphasis>
</taxonomicName>
and
<taxonomicName id="ECB34D320D10FFED50B7EEE7FCC5398C" authorityName="MENETRIES" authorityYear="1832" box="[704,838,1243,1264]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D10FFED50B7EEE7FCC5398C" box="[704,838,1243,1264]" italics="true" pageId="13" pageNumber="1796">A. laevigatus</emphasis>
</taxonomicName>
by its dark rufopiceous colour, the pronotum anterior to the posterior angles somewhat sinuate and with posterior angles distinctly angled, the elytra somewhat flattened, from
<taxonomicName id="ECB34D320D10FFED5145EF2AFC1A3857" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[818,921,1302,1323]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D10FFED5145EF2AFC1A3857" box="[818,921,1302,1323]" italics="true" pageId="13" pageNumber="1796">A. picipes</emphasis>
</taxonomicName>
additionally by having the clypeal anterior angles with only one setiferous pore puncture each (in
<taxonomicName id="ECB34D320D10FFED51AEEF0FFBBE3834" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[985,1085,1331,1352]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D10FFED51AEEF0FFBBE3834" box="[985,1085,1331,1352]" italics="true" pageId="13" pageNumber="1796">A. picipes</emphasis>
</taxonomicName>
as a rule with two setiferous pore punctures each).
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EF4AFEA038BA" blockId="13.[178,1085,1397,1478]" pageId="13" pageNumber="1796">H a b i t a t: Nothing is known about biotope and circumstances of collecting, but it can be supposed that the members of this species live under stones on moist soil with vegetation.</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EFEAFD053B75" blockId="13.[178,1085,1493,1545]" pageId="13" pageNumber="1796">
D i s t r i b u t i o n: Up to present only known from
<collectingCountry id="53A476210D10FFED5090EFEAFCA73897" box="[743,804,1494,1515]" name="Egypt" pageId="13" pageNumber="1796">Egypt</collectingCountry>
from some localities in the Mount
<collectingRegion id="E977F8530D10FFED528BEFCFFEB33B75" box="[252,304,1523,1545]" country="Egypt" name="Shamal Sina'" pageId="13" pageNumber="1796">Sinai</collectingRegion>
near the town of Saint Catherine.
</paragraph>
<paragraph id="2B0C36B10D10FFED52C5EC24FBBF3B52" blockId="13.[178,1084,1560,1582]" box="[178,1084,1560,1582]" pageId="13" pageNumber="1796">
R e m a r k s: There is only one species of
<taxonomicName id="ECB34D320D10FFED500EEC24FD543B51" authorityName="DEJEAN" authorityYear="1821" box="[633,727,1560,1581]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1796" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D10FFED500EEC24FD543B51" box="[633,727,1560,1581]" italics="true" pageId="13" pageNumber="1796">Acinopus</emphasis>
</taxonomicName>
recorded from
<collectingCountry id="53A476210D10FFED5100EC24FC373B51" box="[887,948,1560,1581]" name="Egypt" pageId="13" pageNumber="1796">Egypt</collectingCountry>
(in fact from
</paragraph>
<paragraph id="2B0C36B10D13FFEE52C5EAE7FD703CDC" blockId="14.[178,1086,217,417]" pageId="14" pageNumber="1797">
the
<collectingRegion id="E977F8530D13FFEE52AFEAE7FEF43D8D" box="[216,375,219,241]" country="Egypt" name="Shamal Sina'" pageId="14" pageNumber="1797">Sinai Peninsula</collectingRegion>
) with the name
<taxonomicName id="ECB34D320D13FFEE506AEAE7FCDC3D8C" authority="MENETRIES, 1832" authorityName="MENETRIES" authorityYear="1832" box="[541,863,217,240]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1797" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D13FFEE506AEAE7FD213D8C" box="[541,674,219,240]" italics="true" pageId="14" pageNumber="1797">A. laevigatus</emphasis>
MÉNÉTRIÉS, 1832
</taxonomicName>
(
<bibRefCitation id="4F224B400D13FFEE511AEAE7FF433C72" author="PEYERIMHOFF P" pageId="14" pageNumber="1797" pagination="1907 - 1923" refId="ref14833" refString="PEYERIMHOFF P. de (1907): Liste des coleopteres du Sinai. - L`Abeille. Journal d'Entomologie 31 [1907 - 1923]: 1 - 55." type="journal article" year="1907">PEYERIMHOFF 1907: 6</bibRefCitation>
, mentioned the locality &quot;Djebel Safsafeh, vers. 1800&quot;, based on his own finding, obviously only
<specimenCount id="3DB5FD380D13FFEE5351EB2AFE373C57" box="[294,436,277,299]" pageId="14" pageNumber="1797" type="generic">one specimen</specimenCount>
, which we could examine, see &quot;
<typeStatus id="F40888130D13FFEE5177EB2AFCB53C57" box="[768,822,278,299]" pageId="14" pageNumber="1797">Type</typeStatus>
material&quot;).
<bibRefCitation id="4F224B400D13FFEE51C7EB2AFE9D3C35" author="SCHATZMAYR A" pageId="14" pageNumber="1797" pagination="1 - 114" refId="ref14863" refString="SCHATZMAYR A. (1936): Risultati scientifici della spedizione entomologica di S. A. S. il Principe Alessandro della Torre e Tasso in Egitto e nella penisola del Sinai. XII. Catalogo ragionato dei Carabidi finora noti d'Egitto e del Sinai. - Publicazioni del Museo Entomologico &quot; Pietro Rossi &quot;, Duino 1: 1 - 114." type="journal article" year="1936">SCHATZMAYR (1936: 76)</bibRefCitation>
and
<bibRefCitation id="4F224B400D13FFEE5325EB0FFDCD3C35" author="HOSNI M &amp; SHARKAWY A &amp; OSHAIBAH A &amp; HASSAN M" box="[338,590,307,329]" pageId="14" pageNumber="1797" pagination="549 - 573" refId="ref14416" refString="HOSNI M. T., AL- SHARKAWY A. Z., OSHAIBAH A. A., HASSAN M. I. &amp; A. A. EL- GHARBAWY (2003): Classification of subfamily Harpalinae (family: Carabidae - order: Coleoptera) in Egypt. II - tribes Harpalini, Masoreini and Amarini. - Egyptian Journal of Entomology 40: 549 - 573." type="journal article" year="2003">HOSNI et al. (2003: 561)</bibRefCitation>
refered on PEYERIMHOFF (l. c.).
<bibRefCitation id="4F224B400D13FFEE51D3EB0FFF563C1A" author="ALFIERI A" pageId="14" pageNumber="1797" pagination="1 - 361" refId="ref14125" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xiii, 1 - 361." type="book chapter" year="1976">ALFIERI (1976: 21)</bibRefCitation>
, beside the reference on PEYERIMHOFF (l. c.) added a second locality: Wadi El Lega (which is identic with Wadi el Arbain). Without doubt, the species mentioned as
<taxonomicName id="ECB34D320D13FFEE5650EB52FE993CDC" authorityName="MENETRIES" authorityYear="1832" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1797" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D13FFEE5650EB52FE993CDC" italics="true" pageId="14" pageNumber="1797">A. laevigatus</emphasis>
</taxonomicName>
by ALFIERI also refers on
<taxonomicName id="ECB34D320D13FFEE5051EBB7FD203CDC" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[550,675,395,416]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1797" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D13FFEE5051EBB7FD203CDC" box="[550,675,395,416]" italics="true" pageId="14" pageNumber="1797">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D13FFEE50DEEBB7FD703CDC" box="[681,755,395,416]" pageId="14" pageNumber="1797" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,746 @@
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<mods:title id="48432EC8FB2F94360280F885743A2C03">Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)</mods:title>
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<paragraph id="2B0C36B10D1EFFE352C5EB95FC943CC2" blockId="3.[178,791,424,446]" box="[178,791,424,446]" pageId="3" pageNumber="1786">
<heading id="704481DD0D1EFFE352C5EB95FC943CC2" box="[178,791,424,446]" fontSize="8" level="2" pageId="3" pageNumber="1786" reason="2">
<emphasis id="19C7EAA30D1EFFE352C5EB95FC943CC2" bold="true" box="[178,791,424,446]" pageId="3" pageNumber="1786">
<taxonomicName id="ECB34D320D1EFFE352C5EB95FE6C3CC2" ID-CoL="9BVS" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[178,495,424,446]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1786" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov." subGenus="Acinopus">
<emphasis id="19C7EAA30D1EFFE352C5EB95FE973CC2" bold="true" box="[178,276,425,446]" italics="true" pageId="3" pageNumber="1786">Acinopus</emphasis>
(
<emphasis id="19C7EAA30D1EFFE35354EB95FE063CC2" bold="true" box="[291,389,425,446]" italics="true" pageId="3" pageNumber="1786">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D1EFFE353E2EB94FE6C3CC2" bold="true" box="[405,495,424,446]" italics="true" pageId="3" pageNumber="1786">orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D1EFFE35382EB95FDC23CC2" box="[501,577,425,446]" pageId="3" pageNumber="1786" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="B3882A340D1EFFE35039EB94FD133CC2" box="[590,656,424,446]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="3" pageNumber="1786">Figs 1</figureCitation>
,
<figureCitation id="B3882A340D1EFFE350EAEB95FD403CC2" box="[669,707,425,446]" captionStart="Figs 5-8" captionStartId="6.[178,218,1273,1295]" captionTargetBox="[198,1071,235,1238]" captionTargetId="figure-0@6.[178,1085,204,1274]" captionTargetPageId="6" captionText="Figs 5-8: Acinopus DEJEAN, 1821. Median lobe of male. (5, 6) A. orszuliki nov.sp. (5) Lateral view (holotype). (6) dorsal view (paratype, type locality). (7, 8) A. sinaiticus nov.sp. (7) Lateral view (holotype). (8) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="3" pageNumber="1786">5, 6</figureCitation>
,
<figureCitation id="B3882A340D1EFFE350B8EB95FD6A3CC2" box="[719,745,425,446]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="3" pageNumber="1786">13</figureCitation>
,
<figureCitation id="B3882A340D1EFFE35082EB94FC8C3CC2" box="[757,783,424,446]" captionStart="Fig" captionStartId="10.[178,211,901,923]" captionTargetBox="[178,1085,330,892]" captionTargetId="figure-0@10.[178,1085,212,892]" captionTargetPageId="10" captionText="Fig. 17: One of the localities where A. orszuliki nov.sp. was found (Mahan, SE Kerman, 2500 m, Artemisia-high-altitude steppe)." figureDoi="http://doi.org/10.5281/zenodo.5356286" httpUri="https://zenodo.org/record/5356286/files/figure.png" pageId="3" pageNumber="1786">17</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="2B0C36B10D1EFFE352C5EBF2FD9C386C" blockId="3.[178,1086,461,1296]" pageId="3" pageNumber="1786">
T y p e m a t e r i a l:
<typeStatus id="F40888130D1EFFE353D0EBF2FE733C98" box="[423,496,462,484]" pageId="3" pageNumber="1786" type="holotype">Holotype</typeStatus>
6: &quot;
<collectingCountry id="53A476210D1EFFE35055EBF2FDD93C9F" box="[546,602,462,483]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
prov.
<collectingRegion id="E977F8530D1EFFE350E5EBF2FD5A3C9F" box="[658,729,462,483]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
/ Sarbizhan
<date id="5F0D10710D1EFFE35132EBF1FC113C9F" box="[837,914,461,483]" pageId="3" pageNumber="1786" value="2010-05-05">5.5.2010</date>
/
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S
<collectingRegion id="E977F8530D1EFFE35181EBF2FBBE3C9F" box="[1014,1085,462,483]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
/ lgt. Orszulik
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&quot; (cWR).
<typeStatus id="F40888130D1EFFE3539AEBD9FDC63C86" box="[493,581,485,506]" pageId="3" pageNumber="1786" type="paratype">Paratypes</typeStatus>
: 466,
<specimenCount id="3DB5FD380D1EFFE350F0EBD9FD2A3C83" box="[647,681,485,511]" pageId="3" pageNumber="1786" type="female">5♀♀</specimenCount>
: with the same data (cORSZ, cWR). 1266,
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: with the same data but &quot;
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&quot; and &quot;
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&quot; (cORSZ, cWR, MFNB, ZIN). 266,
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: &quot;
<collectingCountry id="53A476210D1EFFE35291E82FFE883F54" box="[230,267,531,552]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
(
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) / Babidan vill., W Radiabar / (Kuh-i- Lalazar)
<quantity id="EC4B9B540D1EFFE35171E82FFCC93F55" box="[774,842,531,553]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="1786" unit="m" value="2800.0">2800 m</quantity>
/ ca
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NW Dgirouft) /
<date id="5F0D10710D1EFFE352A8E816FEC23F3C" box="[223,321,554,576]" pageId="3" pageNumber="1786" value="2006-03-28">28.III.2006</date>
W. Heinz&quot; (cWR). 466: &quot;
<collectingCountry id="53A476210D1EFFE35024E817FDFB3F3C" box="[595,632,555,576]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
(
<collectingRegion id="E977F8530D1EFFE350FDE817FD573F3D" box="[650,724,555,577]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
) / Babidan-vill.
<quantity id="EC4B9B540D1EFFE35102E817FC3C3F3C" box="[885,959,555,576]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="1786" unit="m" value="2800.0">2800 m</quantity>
/ Heinz leg.
<date id="5F0D10710D1EFFE352A8E87EFEC63F24" box="[223,325,578,600]" pageId="3" pageNumber="1786" value="2006-03-28">28.III.2006</date>
&quot;, &quot;Kuh-i- Lalazar / W. Radiabar / ≈
<quantity id="EC4B9B540D1EFFE350F7E87EFD343F24" box="[640,695,578,600]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="1786" unit="km" value="60.0">60 km</quantity>
NW Dgirouft&quot; (cHZ, cWR).
<specimenCount id="3DB5FD380D1EFFE351CEE87EFC563F20" box="[953,981,578,604]" pageId="3" pageNumber="1786" type="female">1 ♀</specimenCount>
: &quot;S
<collectingCountry id="53A476210D1EFFE35675E87EFBB93F2B" box="[1026,1082,578,599]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
, prov.
<collectingRegion id="E977F8530D1EFFE35364E865FED83F12" box="[275,347,601,622]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kermān</collectingRegion>
,
<quantity id="EC4B9B540D1EFFE35313E865FE243F12" box="[356,423,601,622]" metricMagnitude="3" metricUnit="m" metricValue="2.213" pageId="3" pageNumber="1786" unit="m" value="2213.0">2213 m</quantity>
/
<quantity id="EC4B9B540D1EFFE353CEE865FE683F13" box="[441,491,601,623]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="1786" unit="km" value="45.0">45km</quantity>
NNE Sābsvārān (Jiroft) /
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,
<date id="5F0D10710D1EFFE3510FE865FC603F12" box="[888,995,601,622]" pageId="3" pageNumber="1786" value="2004-07-19">19.VII.2004</date>
/ leg. Petr Kabátek&quot; (OUMNH). 266,
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: &quot;
<collectingCountry id="53A476210D1EFFE35036E84CFDEB3FF9" box="[577,616,624,645]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
,
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, Mahan, Dareh / Bar,
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/
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,
<quantity id="EC4B9B540D1EFFE35303E8BBFE3A3FE1" box="[372,441,647,669]" metricMagnitude="3" metricUnit="m" metricValue="2.503" pageId="3" pageNumber="1786" unit="m" value="2503.0">2503 m</quantity>
18-03- / 2007 leg. R.F.F.L. Felix&quot; (cFEL). 566,
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: &quot;
<collectingCountry id="53A476210D1EFFE351B7E8BBFC643FE0" box="[960,999,647,668]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
,
<collectingRegion id="E977F8530D1EFFE35186E8B4FBB93FE1" box="[1009,1082,648,669]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
, Qal ehAskar, halverwege dal,
<geoCoordinate id="4E8750760D1EFFE3507FE8A2FD0B3FC8" box="[520,648,670,692]" degrees="29" direction="north" minutes="36" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="04.5" value="29.60125">N29°3604.5</geoCoordinate>
/
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,
<quantity id="EC4B9B540D1EFFE35148E8A2FC083FC8" box="[831,907,670,692]" metricMagnitude="3" metricUnit="m" metricValue="2.502" pageId="3" pageNumber="1786" unit="m" value="2502.0">2502 m</quantity>
. /
<date id="5F0D10710D1EFFE351DCE8A3FB923FC8" box="[939,1041,671,692]" pageId="3" pageNumber="1786" value="2007-03-22">22-03-2007</date>
leg. R.F.F.L. Felix&quot; (cFEL, cWR, MFNB).
<specimenCount id="3DB5FD380D1EFFE35033E88AFDE53FAC" box="[580,614,694,720]" pageId="3" pageNumber="1786" type="female">2♀♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE350FDE88AFD333FB7" box="[650,688,694,715]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
,
<collectingRegion id="E977F8530D1EFFE350CBE88AFC863FB7" box="[700,773,694,715]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
, Kuh e Lalehzan, /
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<geoCoordinate id="4E8750760D1EFFE352A8E8F1FEE93F9F" box="[223,362,717,739]" degrees="056" direction="east" minutes="47" orientation="longitude" pageId="3" pageNumber="1786" precision="1" seconds="33.6" value="56.792667">E056°4733.6</geoCoordinate>
, /
<quantity id="EC4B9B540D1EFFE353F7E8F1FE443F9E" box="[384,455,717,738]" metricMagnitude="3" metricUnit="m" metricValue="2.83" pageId="3" pageNumber="1786" unit="m" value="2830.0">2830 m</quantity>
.
<date id="5F0D10710D1EFFE353A7E8F1FDB53F9E" box="[464,566,717,738]" pageId="3" pageNumber="1786" value="2007-03-19">19-03-2007</date>
/ leg. R.F.F.L. Felix&quot; (cFEL).
<specimenCount id="3DB5FD380D1EFFE3513FE8F1FCE93F9B" box="[840,874,717,743]" pageId="3" pageNumber="1786" type="female">2♀♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE351FBE8F1FC313F9E" box="[908,946,717,738]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
,
<collectingRegion id="E977F8530D1EFFE351CBE8F1FB863F9E" box="[956,1029,717,738]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
,
<quantity id="EC4B9B540D1EFFE35678E8F1FBBE3F9F" box="[1039,1085,717,739]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="1786" unit="km" value="2.0">2 km</quantity>
W Sirch, /
<geoCoordinate id="4E8750760D1EFFE35337E8D8FE313F85" box="[320,434,740,761]" degrees="30" direction="north" minutes="13" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="09.1" value="30.219194">N30°1309.1</geoCoordinate>
<geoCoordinate id="4E8750760D1EFFE353CFE8D8FDC13F86" box="[440,578,740,762]" degrees="057" direction="east" minutes="27" orientation="longitude" pageId="3" pageNumber="1786" precision="1" seconds="21.7" value="57.456028">E057°2721.7</geoCoordinate>
, /
<quantity id="EC4B9B540D1EFFE35020E8D8FD1E3F85" box="[599,669,740,762]" metricMagnitude="3" metricUnit="m" metricValue="2.246" pageId="3" pageNumber="1786" unit="m" value="2246.0">2246 m</quantity>
.
<date id="5F0D10710D1EFFE350D1E8D8FC8F3F85" box="[678,780,740,761]" pageId="3" pageNumber="1786" value="2007-03-17">17-03-2007</date>
/ leg. R.F.F.L. Felix&quot; (cFEL). 16,
<specimenCount id="3DB5FD380D1EFFE352A8E8C7FE823E69" box="[223,257,763,789]" pageId="3" pageNumber="1786" type="female">4♀♀</specimenCount>
: &quot;IR
<collectingRegion id="E977F8530D1EFFE35348E8C0FE053E6D" box="[319,390,764,785]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
/ Qaleh Askar /
<date id="5F0D10710D1EFFE35069E8C7FD013E6C" box="[542,642,763,784]" pageId="3" pageNumber="1786" value="2007-03-22">22-III 2007</date>
/ Muilwijk leg&quot; (cMUIL, cWR). 366,
<specimenCount id="3DB5FD380D1EFFE35195E8C7FB863E69" box="[994,1029,763,789]" pageId="3" pageNumber="1786" type="female">2♀♀</specimenCount>
: &quot;IR
<collectingRegion id="E977F8530D1EFFE352A8E92FFEA53E54" box="[223,294,787,808]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
/ Sirch env. /
<date id="5F0D10710D1EFFE353D9E92FFD973E54" box="[430,532,787,808]" pageId="3" pageNumber="1786" value="2007-03-17">17-III 2007</date>
/ Muilwijk leg&quot; (cMUIL, cWR). 266,
<specimenCount id="3DB5FD380D1EFFE3510AE92FFC1F3E51" box="[893,924,787,813]" pageId="3" pageNumber="1786" type="female">1 ♀</specimenCount>
:
<collectingCountry id="53A476210D1EFFE351C6E92FFC643E54" box="[945,999,787,808]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
(
<collectingRegion id="E977F8530D1EFFE35182E92FFE923E43" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman Prov.</collectingRegion>
, / Jiroft County) / Darb-e Behesht area / Sinabad, ca
<quantity id="EC4B9B540D1EFFE35154E915FCEF3E43" box="[803,876,809,831]" metricMagnitude="3" metricUnit="m" metricValue="2.6149999999999998" pageId="3" pageNumber="1786" unit="m" value="2615.0">2615 m</quantity>
/ (
<quantity id="EC4B9B540D1EFFE351E7E915FC4C3E3C" box="[912,975,809,832]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="1786" unit="km" value="75.0">75 km</quantity>
N Jiroft) / 29°1400.99N / 57°2025.51E / (orchard, pit fall traps) /
<date id="5F0D10710D1EFFE35177E97DFC2D3E2A" box="[768,942,833,854]" pageId="3" pageNumber="1786" value="2013-04-18" valueMax="2013-08-08" valueMin="2013-04-18">18.IV.- 8.VIII.2013</date>
local collector&quot; (cMUIL, cWR).
<specimenCount id="3DB5FD380D1EFFE35304E964FE093E0E" box="[371,394,856,882]" pageId="3" pageNumber="1786" type="female">1♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE353D3E964FE5F3E11" box="[420,476,856,877]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
30 Km Z v / NEGAR 10-03-07 / VELDKAMP&quot; (cVELD). 16,
<specimenCount id="3DB5FD380D1EFFE3566DE964FBB33E0E" box="[1050,1072,856,882]" pageId="3" pageNumber="1786" type="female">1♀</specimenCount>
: &quot;IR CAH MAHALI / BAKHTIYARI 31- / 3-07 VELDKAMP&quot; (cVELD). 466,
<specimenCount id="3DB5FD380D1EFFE351A9E953FB8B3EF5" box="[990,1032,879,905]" pageId="3" pageNumber="1786" type="female">2 ♀♀</specimenCount>
: IR SIRCH /
<date id="5F0D10710D1EFFE35346E9BBFE0F3EE0" box="[305,396,903,924]" pageId="3" pageNumber="1786" value="2007-03-17">17-3-2007</date>
/ VELDKAMP&quot; (cVELD, cWR). 16,
<specimenCount id="3DB5FD380D1EFFE35093E9BBFD783EDD" box="[740,763,903,929]" pageId="3" pageNumber="1786" type="female">1♀</specimenCount>
: &quot;IR HEMMAD ABAD / 40 Km O
<collectingRegion id="E977F8530D1EFFE352A8E9A2FEBC3ECF" box="[223,319,926,947]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">KERMAN</collectingRegion>
/ 18-3-07 VELDKAMP&quot; (cVELD, cWR). 16,
<specimenCount id="3DB5FD380D1EFFE35098E9A2FC853EC4" box="[751,774,926,952]" pageId="3" pageNumber="1786" type="female">1♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE35156E9A2FCDA3ECF" box="[801,857,926,947]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
(
<collectingRegion id="E977F8530D1EFFE35111E9A2FC653ECF" box="[870,998,926,947]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman Prov.</collectingRegion>
) / Mahan SE
<collectingRegion id="E977F8530D1EFFE35373E989FEC83EB6" box="[260,331,949,970]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
<quantity id="EC4B9B540D1EFFE35322E989FE1D3EB6" box="[341,414,949,971]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="1786" unit="m" value="2500.0">2500 m</quantity>
/
<geoCoordinate id="4E8750760D1EFFE353CEE989FDBD3EB7" box="[441,574,949,971]" degrees="29" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="34.1" value="29.892805">N 29°53'34.1</geoCoordinate>
/
<geoCoordinate id="4E8750760D1EFFE35032E989FD4F3EB7" box="[581,716,949,971]" degrees="57" direction="east" minutes="20" orientation="longitude" pageId="3" pageNumber="1786" precision="1" seconds="43.4" value="57.345387">E 57°20'43.4</geoCoordinate>
/ (
<emphasis id="19C7EAA30D1EFFE35099E989FCC03EB6" box="[750,835,949,970]" italics="true" pageId="3" pageNumber="1786">Artemisia</emphasis>
-high-altitude steppe, under stones) /
<date id="5F0D10710D1EFFE35342E9F0FE0D3E9D" box="[309,398,972,993]" pageId="3" pageNumber="1786" value="2014-06-01">1.VI.2014</date>
Wrase &amp; Laser [06]&quot; (MFNB, cWR, *
<superScript id="DCC69BF90D1EFFE3508AE9FBFC873EA9" attach="none" box="[765,772,967,981]" fontSize="6" pageId="3" pageNumber="1786">1</superScript>
). &quot;
<collectingCountry id="53A476210D1EFFE35156E9F0FCDA3E9D" box="[801,857,972,993]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
(
<collectingRegion id="E977F8530D1EFFE35110E9F0FC6A3E9D" box="[871,1001,972,993]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman Prov.</collectingRegion>
) / Kūh-e Lālehzār / nr. Lālehzār vill.
<quantity id="EC4B9B540D1EFFE353A0E9DFFDCD3E85" box="[471,590,995,1017]" metricMagnitude="3" metricUnit="m" metricValue="3.2049999999999996" metricValueMax="3.3" metricValueMin="3.11" pageId="3" pageNumber="1786" unit="m" value="3205.0" valueMax="3300.0" valueMin="3110.0">3110-3300 m</quantity>
/
<geoCoordinate id="4E8750760D1EFFE35017E9D8FD533E85" box="[608,720,995,1017]" degrees="29" direction="north" minutes="28" orientation="latitude" pageId="3" pageNumber="1786" precision="15" seconds="35" value="29.476389">N 29°28'35</geoCoordinate>
/
<geoCoordinate id="4E8750760D1EFFE350A0E9DFFCCA3E85" box="[727,841,995,1017]" degrees="56" direction="east" minutes="49" orientation="longitude" pageId="3" pageNumber="1786" precision="15" seconds="15" value="56.82083">E 56°49'15</geoCoordinate>
/ (mountain steppe, / under stones) /
<date id="5F0D10710D1EFFE35347E9C7FE0B396C" box="[304,392,1019,1040]" pageId="3" pageNumber="1786" value="2014-06-03">3.VI.2014</date>
Wrase &amp; Laser [09B]&quot; (cWR, *). 466,
<specimenCount id="3DB5FD380D1EFFE35086E9C7FC9F3969" box="[753,796,1019,1045]" pageId="3" pageNumber="1786" type="female">7 ♀♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE35149E9C7FCE6396C" box="[830,869,1019,1040]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
, Prov.
<collectingRegion id="E977F8530D1EFFE351D4E9C7FC6F396C" box="[931,1004,1019,1040]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
,
<collectingRegion id="E977F8530D1EFFE35181E9C7FBBE396C" box="[1014,1085,1019,1040]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
S. / Mahan,
<emphasis id="19C7EAA30D1EFFE3533CEE2EFE1C395B" box="[331,415,1042,1063]" italics="true" pageId="3" pageNumber="1786">Artemisia</emphasis>
-Hochsteppe /
<geoCoordinate id="4E8750760D1EFFE35068EE2DFD1C395B" box="[543,671,1041,1063]" degrees="29" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="34.1" value="29.892805">N29°5334.1</geoCoordinate>
<geoCoordinate id="4E8750760D1EFFE350D2EE2DFCA0395B" box="[677,803,1041,1063]" degrees="57" direction="east" minutes="20" orientation="longitude" pageId="3" pageNumber="1786" precision="1" seconds="43.4" value="57.345387">E57°2043.4</geoCoordinate>
/
<date id="5F0D10710D1EFFE35158EE2EFC12395B" box="[815,913,1042,1063]" pageId="3" pageNumber="1786" value="2014-06-01">01.VI.2014</date>
<quantity id="EC4B9B540D1EFFE351E0EE2DFC58395B" box="[919,987,1041,1063]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="1786" unit="m" value="2500.0">2500 m</quantity>
üNN / leg. A. Weigel IR 06 HF&quot; (NME, cSCHN, cWEIG, cWR, *).
<specimenCount id="3DB5FD380D1EFFE35173EE15FCA5393F" box="[772,806,1065,1091]" pageId="3" pageNumber="1786" type="female">1 ♀</specimenCount>
: &quot;
<collectingCountry id="53A476210D1EFFE35133EE15FCE93942" box="[836,874,1065,1086]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
, Prov.
<collectingRegion id="E977F8530D1EFFE351C7EE15FC7A3942" box="[944,1017,1065,1086]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
, Kuh-e Lalehzar, / Lalehzar vill. Bachtal,
<geoCoordinate id="4E8750760D1EFFE3507AEE7CFD00392A" box="[525,643,1088,1110]" degrees="29" direction="north" minutes="57" orientation="latitude" pageId="3" pageNumber="1786" precision="15" seconds="59" value="29.966389">N 29°5759</geoCoordinate>
E 56°4574 /
<date id="5F0D10710D1EFFE35166EE7CFCF73929" box="[785,884,1088,1109]" pageId="3" pageNumber="1786" value="2014-06-03">03.VI.2014</date>
3.400 müNN / leg. A. Weigel IR 09a HF&quot; (cWEIG). 266: &quot;
<collectingCountry id="53A476210D1EFFE3505BEE6BFDE73910" box="[556,612,1111,1132]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
(
<collectingRegion id="E977F8530D1EFFE35018EE6BFD6E3911" box="[623,749,1111,1133]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman Prov.</collectingRegion>
) / nr. Sirch vill. SE
<collectingRegion id="E977F8530D1EFFE351EFEE64FC5C3911" box="[920,991,1112,1133]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
/
<quantity id="EC4B9B540D1EFFE35198EE6BFBB13911" box="[1007,1074,1111,1133]" metricMagnitude="3" metricUnit="m" metricValue="2.56" pageId="3" pageNumber="1786" unit="m" value="2560.0">2560 m</quantity>
/
<geoCoordinate id="4E8750760D1EFFE352A8EE53FED939F8" box="[223,346,1134,1156]" degrees="30" direction="north" minutes="10" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="56.4" value="30.182333">N 30°10'56.4&quot;</geoCoordinate>
/
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/ (partly irrigated orchard, under stones, partly running at darkness) /
<date id="5F0D10710D1EFFE352A8EEBAFEB839E7" box="[223,315,1158,1179]" pageId="3" pageNumber="1786" value="2014-05-30">30.V.2014</date>
Saeed Azadbakhsh [04]&quot; (ZMAF, cAZAD, *). 16: &quot;
<collectingCountry id="53A476210D1EFFE35135EEBAFCFF39E7" box="[834,892,1158,1179]" name="Iran" pageId="3" pageNumber="1786">IRAN</collectingCountry>
,
<collectingRegion id="E977F8530D1EFFE351FCEEBAFBA839E0" box="[907,1067,1158,1180]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman Province</collectingRegion>
/ Bardsir-Baft road: / Qaleh Askar,
<quantity id="EC4B9B540D1EFFE35066EEA1FDD239CF" box="[529,593,1181,1203]" metricMagnitude="3" metricUnit="m" metricValue="2.76" pageId="3" pageNumber="1786" unit="m" value="2760.0">2760m</quantity>
,
<geoCoordinate id="4E8750760D1EFFE3502DEEA1FD4C39CE" box="[602,719,1181,1202]" degrees="29" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="1786" precision="15" seconds="18" value="29.505">N 29°3018</geoCoordinate>
<geoCoordinate id="4E8750760D1EFFE350A2EEA1FCD739CF" box="[725,852,1181,1203]" degrees="056" direction="east" minutes="37" orientation="longitude" pageId="3" pageNumber="1786" precision="15" seconds="47" value="56.62972">E 056°3747</geoCoordinate>
/
<date id="5F0D10710D1EFFE35111EEA1FC4939CF" box="[870,970,1181,1203]" pageId="3" pageNumber="1786" value="2007-06-05">06.05.2007</date>
, lg. Frisch &amp; Serri&quot; (MFNB).
<specimenCount id="3DB5FD380D1EFFE35304EE88FE0939B2" box="[371,394,1204,1230]" pageId="3" pageNumber="1786" type="female">1♀</specimenCount>
: &quot;
<geoCoordinate id="4E8750760D1EFFE353D2EE88FDA839B6" box="[421,555,1204,1226]" degrees="29" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="1786" precision="1" seconds="34.1" value="29.892805">N 29°5334.1</geoCoordinate>
/
<geoCoordinate id="4E8750760D1EFFE35044EE88FD3A39B6" box="[563,697,1204,1226]" degrees="57" direction="east" minutes="20" orientation="longitude" pageId="3" pageNumber="1786" precision="1" seconds="43.4" value="57.345387">E 57°2043.4</geoCoordinate>
/
<collectingCountry id="53A476210D1EFFE350B9EE88FD7039B5" box="[718,755,1204,1225]" name="Iran" pageId="3" pageNumber="1786">Iran</collectingCountry>
, Prov.
<collectingRegion id="E977F8530D1EFFE35144EE88FCFF39B5" box="[819,892,1204,1225]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
,
<collectingRegion id="E977F8530D1EFFE351F1EE88FC4E39B5" box="[902,973,1204,1225]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman</collectingRegion>
S. / Mahan,
<emphasis id="19C7EAA30D1EFFE352A8EEF7FEB0399C" box="[223,307,1227,1248]" italics="true" pageId="3" pageNumber="1786">Artemisia</emphasis>
-Hochsteppe /
<date id="5F0D10710D1EFFE353CFEEF7FD99399D" box="[440,538,1227,1249]" pageId="3" pageNumber="1786" value="2014-01-06">01.06.2014</date>
<quantity id="EC4B9B540D1EFFE35055EEF7FDDC399D" box="[546,607,1227,1249]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="1786" unit="m" value="2500.0">2500m</quantity>
üNN / leg.: Schnitter IR 06 HF&quot; (cSCHN, *).
<specimenCount id="3DB5FD380D1EFFE3567BEEF7FBAC3999" box="[1036,1071,1227,1253]" pageId="3" pageNumber="1786" type="female">2♀♀</specimenCount>
&quot;dat.
<date id="5F0D10710D1EFFE3537AEEDEFEE63984" box="[269,357,1250,1272]" pageId="3" pageNumber="1786" value="2014-05-29">29.5.2014</date>
<collectingCountry id="53A476210D1EFFE3531DEEDEFE0E3984" box="[362,397,1250,1272]" name="Iran" pageId="3" pageNumber="1786">Irán</collectingCountry>
IR /
<collectingRegion id="E977F8530D1EFFE353C3EEDFFDAC3984" box="[436,559,1251,1272]" country="Iran" name="Kerman" pageId="3" pageNumber="1786">Kerman prov.</collectingRegion>
(Jabel Baref / Deh Bakri (Jass) /
<geoCoordinate id="4E8750760D1EFFE35138EEDEFC3C3984" box="[847,959,1250,1272]" degrees="28" direction="north" minutes="59" orientation="latitude" pageId="3" pageNumber="1786" precision="15" seconds="53" value="28.998056">N28°5953</geoCoordinate>
<geoCoordinate id="4E8750760D1EFFE351B3EEDEFBB13984" box="[964,1074,1250,1272]" degrees="57" direction="east" minutes="54" orientation="longitude" pageId="3" pageNumber="1786" precision="15" seconds="16" value="57.904446">E57°5416</geoCoordinate>
/ lgt. Černý Fr.
<quantity id="EC4B9B540D1EFFE3532EEEC6FE1D3873" box="[345,414,1274,1295]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="1786" unit="m" value="2300.0">2300 m</quantity>
.n.m&quot; (cVON).
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D1EFFEB52C5EF22FCFE3E5A" lastPageId="11" lastPageNumber="1794" pageId="3" pageNumber="1786" type="description">
<paragraph id="2B0C36B10D1EFFE352C5EF22FCB23812" blockId="3.[178,1084,1310,1390]" pageId="3" pageNumber="1786">
E t y m o l o g y: Latinized patronym based on the surname of Kamil Orszulik (Frýdec- Místek,
<collectingCountry id="53A476210D1EFFE35371EF07FE2D382D" box="[262,430,1339,1361]" name="Czech Republic" pageId="3" pageNumber="1786">Czech Republic</collectingCountry>
), whose indefatigable efforts in the field contributed substantial numbers of Carabid beetles, very interesting or new to science.
</paragraph>
<paragraph id="2B0C36B10D1EFFE352C5EF42FBB13897" blockId="3.[178,1085,1406,1516]" pageId="3" pageNumber="1786">
D i a g n o s i s: A micropterous species of small to medium size for
<taxonomicName id="ECB34D320D1EFFE351D6EF42FC7C38EF" authorityName="DEJEAN" authorityYear="1821" box="[929,1023,1406,1427]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1786" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D1EFFE351D6EF42FC7C38EF" box="[929,1023,1406,1427]" italics="true" pageId="3" pageNumber="1786">Acinopus</emphasis>
</taxonomicName>
, with excision of dorsal edge of right mandible (subgenus
<taxonomicName id="ECB34D320D1EFFE350C5EFA0FC8F38CD" authorityName="DEJEAN" authorityYear="1821" box="[690,780,1436,1457]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1786" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D1EFFE350C5EFA0FC8F38CD" box="[690,780,1436,1457]" italics="true" pageId="3" pageNumber="1786">Acinopus</emphasis>
</taxonomicName>
), dark rufopiceous or piceous (in mature condition), with appendages somewhat lighter, clypeus in general with one setiferous pore puncture at anterior angles, pronotum with anterior angles acutely pro-
</paragraph>
<paragraph id="2B0C36B10D1EFFE352C5EC17FF383B44" box="[178,187,1579,1592]" pageId="3" pageNumber="1786">
<emphasis id="19C7EAA30D1EFFE352C5EC17FF383B44" bold="true" box="[178,187,1579,1592]" pageId="3" pageNumber="1786">1</emphasis>
</paragraph>
<paragraph id="2B0C36B10D1EFFE552B6EC0EFBBB3C23" blockId="3.[193,813,1586,1608]" lastBlockId="5.[178,1085,217,352]" lastPageId="5" lastPageNumber="1788" pageId="3" pageNumber="1786">
An asterix (*) indicates syntopic occurrence with
<taxonomicName id="ECB34D320D1EFFE350F3EC0EFCAA3B34" authority="(OLIV.)" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[644,809,1586,1608]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1786" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D1EFFE350F3EC0EFD5E3B34" box="[644,733,1586,1608]" italics="true" pageId="3" pageNumber="1786">A. picipes</emphasis>
(OLIV.)
</taxonomicName>
. truding, and with posterior angles obtuse-angled, moderately rounded at tip, somewhat directed backwards with base rectlinear between basal fovae, elytra longer or shorter cylindrical, fused at suture, hind wings hardly half as long as elytra, metepisterna relatively short, pro- and mesotarsi in males only indistinctly widened, apical lamella of median lobe short, about somewhat elongate-triangular (dorsal view). Habitus see
<figureCitation id="B3882A340D18FFE55181EB76FBBB3C23" box="[1014,1080,330,352]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="5" pageNumber="1788">Fig. 1.</figureCitation>
</paragraph>
<caption id="7FCC66390D19FFE452C5EFD6FC9E3B52" ID-DOI="http://doi.org/10.5281/zenodo.5356278" ID-Zenodo-Dep="5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="4" pageNumber="1787" startId="4.[178,218,1514,1536]" targetBox="[277,985,216,1490]" targetPageId="4">
<paragraph id="2B0C36B10D19FFE452C5EFD6FC9E3B52" blockId="4.[178,1086,1514,1582]" pageId="4" pageNumber="1787">
<emphasis id="19C7EAA30D19FFE452C5EFD6FE833883" bold="true" box="[178,256,1514,1536]" pageId="4" pageNumber="1787">Figs 1-4</emphasis>
:
<taxonomicName id="ECB34D320D19FFE4537AEFD7FE6C3B7C" authority="DEJEAN, 1821" authorityName="DEJEAN" authorityYear="1821" box="[269,495,1515,1536]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1787" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D19FFE4537AEFD7FEE13B7C" box="[269,354,1515,1536]" italics="true" pageId="4" pageNumber="1787">Acinopus</emphasis>
DEJEAN, 1821
</taxonomicName>
. Habitus. (
<emphasis id="19C7EAA30D19FFE45022EFD6FDE33883" bold="true" box="[597,608,1514,1535]" pageId="4" pageNumber="1787">1</emphasis>
)
<taxonomicName id="ECB34D320D19FFE45018EFD7FD543B7C" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[623,727,1515,1536]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1787" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D19FFE45018EFD7FD543B7C" box="[623,727,1515,1536]" italics="true" pageId="4" pageNumber="1787">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D19FFE450A9EFD7FCA23B7C" box="[734,801,1515,1536]" pageId="4" pageNumber="1787" rank="species">nov.sp.</taxonomicNameLabel>
(male paratype, type locality). (
<emphasis id="19C7EAA30D19FFE452CDEC3DFF463B6A" bold="true" box="[186,197,1537,1558]" pageId="4" pageNumber="1787">2</emphasis>
)
<taxonomicName id="ECB34D320D19FFE452A2EC3EFEC43B6B" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[213,327,1538,1559]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1787" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D19FFE452A2EC3EFEC43B6B" box="[213,327,1538,1559]" italics="true" pageId="4" pageNumber="1787">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D19FFE45338EC3EFE113B6B" box="[335,402,1538,1559]" pageId="4" pageNumber="1787" rank="species">nov.sp.</taxonomicNameLabel>
(male paratype, Mt. Katherine). (
<emphasis id="19C7EAA30D19FFE450B8EC3DFD593B6A" bold="true" box="[719,730,1537,1558]" pageId="4" pageNumber="1787">3</emphasis>
)
<taxonomicName id="ECB34D320D19FFE4509DEC3EFCC93B6A" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[746,842,1537,1559]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1787" phylum="Arthropoda" rank="species" species="brittoni" status="sp. nov.">
<emphasis id="19C7EAA30D19FFE4509DEC3EFCC93B6A" box="[746,842,1537,1559]" italics="true" pageId="4" pageNumber="1787">A. brittoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D19FFE45125EC3EFC163B6B" box="[850,917,1538,1559]" pageId="4" pageNumber="1787" rank="species">nov.sp.</taxonomicNameLabel>
(female paratype, type locality). (
<emphasis id="19C7EAA30D19FFE4534AEC24FECB3B51" bold="true" box="[317,328,1560,1581]" pageId="4" pageNumber="1787">4</emphasis>
)
<taxonomicName id="ECB34D320D19FFE45321EC24FE3D3B51" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[342,446,1560,1581]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1787" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="19C7EAA30D19FFE45321EC24FE3D3B51" box="[342,446,1560,1581]" italics="true" pageId="4" pageNumber="1787">A. arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D19FFE453B3EC24FD843B51" box="[452,519,1560,1581]" pageId="4" pageNumber="1787" rank="species">nov.sp.</taxonomicNameLabel>
(male paratype, type locality).
</paragraph>
</caption>
<paragraph id="2B0C36B10D18FFE552C5EB53FC753CDE" blockId="5.[178,1085,367,418]" pageId="5" pageNumber="1788">
D e s c r i p t i o n: Body length 12.0-
<quantity id="EC4B9B540D18FFE55031EB53FD223CF8" box="[582,673,367,388]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="5" pageNumber="1788" unit="mm" value="14.8">14.8 mm</quantity>
in males (
<typeStatus id="F40888130D18FFE5517CEB53FCE63CF9" box="[779,869,367,389]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D18FFE5511BEB53FC493CF8" box="[876,970,367,388]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="5" pageNumber="1788" unit="mm" value="14.4">14.4 mm</quantity>
),
<quantity id="EC4B9B540D18FFE551ABEB53FF5A3CDE" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.55" metricValueMin="1.19" pageId="5" pageNumber="1788" unit="mm" value="13.7" valueMax="15.5" valueMin="11.9">11.9-15.5 mm</quantity>
in females; width 4.0-
<quantity id="EC4B9B540D18FFE553C9EBB0FE7B3CDE" box="[446,504,396,418]" metricMagnitude="-1" metricUnit="m" metricValue="1.3462" pageId="5" pageNumber="1788" unit="in" value="5.3">5.3 in</quantity>
males (
<typeStatus id="F40888130D18FFE55031EBB0FD233CDE" box="[582,672,396,418]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D18FFE550D0EBB0FD7B3CDE" box="[679,760,396,418]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="1788" unit="mm" value="5.1">5.1 mm</quantity>
),
<quantity id="EC4B9B540D18FFE5517DEBB0FCFC3CDE" box="[778,895,396,418]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.5" metricValueMin="4.3" pageId="5" pageNumber="1788" unit="mm" value="4.9" valueMax="5.5" valueMin="4.3">4.3-5.5 mm</quantity>
in females.
</paragraph>
<paragraph id="2B0C36B10D18FFE552C5EB8DFB823C99" blockId="5.[178,1086,433,485]" pageId="5" pageNumber="1788">Colour: Lighter to darker rufo-piceous or piceous also in mature condition (never completely black, if dark always with a piceous ground colour), appendages somewhat lighter.</paragraph>
<paragraph id="2B0C36B10D18FFE552C5EBC8FCCC3E16" blockId="5.[178,1086,500,874]" pageId="5" pageNumber="1788">
Head: wide (as normal in
<taxonomicName id="ECB34D320D18FFE553B5EBC8FDA33F75" authorityName="DEJEAN" authorityYear="1821" box="[450,544,500,521]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1788" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D18FFE553B5EBC8FDA33F75" box="[450,544,500,521]" italics="true" pageId="5" pageNumber="1788">Acinopus</emphasis>
</taxonomicName>
), only somewhat narrower than pronotum (PW/HW in males 1.16-1.28, average 1.22, in
<typeStatus id="F40888130D18FFE5505AE82DFD053F5B" box="[557,646,529,551]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
1.26;
<quantity id="EC4B9B540D18FFE550BEE82EFCC53F5B" box="[713,838,529,551]" metricMagnitude="-2" metricUnit="m" metricValue="3.2004" metricValueMax="3.2766" metricValueMin="3.1242" pageId="5" pageNumber="1788" unit="in" value="1.26" valueMax="1.29" valueMin="1.23">1.23-1.29 in</quantity>
females, average 1.25), Eyes small and relatively flat, tempora oblique, as long as diameter of eye or somewhat shorter (dorsally seen), rectlinearly or weakly convexly converging to neck. Frontal fovae mostly small and shallow, sometimes elongately prolonged backwards. Disc with sparse, fine, and shallow puncturation, hardly visible, sometimes between frontal fovae more or less strongly wrinkled. Excision of dorsal lateral edge of right mandible somewhat variable in size and strength, moderately strong to shallow, basal angle of excision as a rule situated somewhat anterior to or somewhat behind level of anterior margin of clypeus (with mandibles closed). Labrum weakly incised at anterior margin, clypeus weakly and semicircularly incised, anterior angles with one setiferous pore puncture each (in
<specimenCount id="3DB5FD380D18FFE55101E926FB813E53" box="[886,1026,793,815]" pageId="5" pageNumber="1788" type="generic">one specimen</specimenCount>
from Sarbizhan as well as the strong, long seta with two additional short, fine setae at both sides). Mentum with a small tooth, submentum with one pair of long setae.
</paragraph>
<paragraph id="2B0C36B10D18FFE552C5E946FDCB3993" blockId="5.[178,1086,889,1263]" pageId="5" pageNumber="1788">
Pronotum (
<figureCitation id="B3882A340D18FFE5535FE945FEEB3EEC" box="[296,360,889,912]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="5" pageNumber="1788">Fig. 1</figureCitation>
): Transverse (PW/PL
<quantity id="EC4B9B540D18FFE5502DE946FD5A3EF3" box="[602,729,889,911]" metricMagnitude="-2" metricUnit="m" metricValue="3.4798" metricValueMax="3.6576" metricValueMin="3.302" pageId="5" pageNumber="1788" unit="in" value="1.37" valueMax="1.44" valueMin="1.3">1.30-1.44 in</quantity>
males, average 1.36, in
<typeStatus id="F40888130D18FFE55193E945FBBE3EF3" box="[996,1085,889,911]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
1.44; in females 1.32-1.41, average 1.37), widest at about end of anterior third, there lateral seta inserted. Disc convex, area of posterior angles more or less strongly flattened in most specimens, anterior and posterior transverse impressions very shallow or only indicated, median line weakly impressed, mostly terminated anterior to transverse impressions. Basal foveae round or elongate, shallowly impressed, in some specimens only indicated. Anterior margin somewhat wider than posterior margin, only weakly excavate, anterior angles distinctly, acutely projecting forward, only narrowly rounded at tip in most specimens. Lateral gutter narrow, somewhat widened toward anterior angles. Sides moderately curved apicad, from widest point weakly convex, in some specimens almost rectlinearly narrowed toward posterior angles. Posterior angles obtuse-angled, moderately rounded at tip, more or less distinctly projecting backwards, base rectilinear between basal fovae in most specimens.
</paragraph>
<paragraph id="2B0C36B10D18FFE552C5EEC1FEFD3B38" blockId="5.[178,1085,1277,1604]" pageId="5" pageNumber="1788">
Elytra (
<figureCitation id="B3882A340D18FFE55377EEC1FEBE3868" box="[256,317,1277,1300]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="5" pageNumber="1788">Fig. 1</figureCitation>
): Longer or shorter cylindrical (EL/EW in males 1.48-1.61, average 1.55, in
<typeStatus id="F40888130D18FFE552B8EF25FEAB3853" box="[207,296,1305,1327]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
1.52; in females 1.54-1.68, average 1.58), hardly wider or only somewhat wider than pronotum (EW/PW in males 1.01-1.11, average 1.08, in
<typeStatus id="F40888130D18FFE551FEEF09FC613837" box="[905,994,1333,1355]" pageId="5" pageNumber="1788" type="holotype">holotype</typeStatus>
1.10; in females 1.07-1.21, average 1.14), fused at suture. On disc strongly convex, humeri distinctly developed, widely rounded at tip, without humeral tooth; toward behind somewhat convexely or almost parallely enlarged, widest somewhat behind middle. Basal bead weakly sinuate, arcuately curving inside humerus and weakly angled or round towards lateral margin. Scutellar pore puncture present, scutellar stria long. Striae fine, smooth or only weakly punctured, intervals flat or weakly convex, with a pore puncture in interval 3 at about beginning of apical fourth at or close to stria 2, sometimes lacking on one or both elytra. Interval 5 and 7 without pore punctures apically. Preapical sinuation only suggested.
</paragraph>
<caption id="7FCC66390D1BFFE652C5EEC5FCC93840" ID-DOI="http://doi.org/10.5281/zenodo.5356280" ID-Zenodo-Dep="5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="6" pageNumber="1789" startId="6.[178,218,1273,1295]" targetBox="[198,1071,235,1238]" targetPageId="6">
<paragraph id="2B0C36B10D1BFFE652C5EEC5FCC93840" blockId="6.[178,1085,1273,1341]" pageId="6" pageNumber="1789">
<emphasis id="19C7EAA30D1BFFE652C5EEC5FF7C3872" bold="true" box="[178,255,1273,1295]" pageId="6" pageNumber="1789">Figs 5-8</emphasis>
:
<taxonomicName id="ECB34D320D1BFFE6537CEEC6FE683873" authority="DEJEAN, 1821" authorityName="DEJEAN" authorityYear="1821" box="[267,491,1274,1295]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1789" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D1BFFE6537CEEC6FEE33873" box="[267,352,1274,1295]" italics="true" pageId="6" pageNumber="1789">Acinopus</emphasis>
DEJEAN, 1821
</taxonomicName>
. Median lobe of male. (
<emphasis id="19C7EAA30D1BFFE650B2EEC6FD643872" bold="true" box="[709,743,1273,1295]" pageId="6" pageNumber="1789">5, 6</emphasis>
)
<taxonomicName id="ECB34D320D1BFFE65082EEC6FCDF3872" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[757,860,1273,1295]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1789" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D1BFFE65082EEC6FCDF3872" box="[757,860,1273,1295]" italics="true" pageId="6" pageNumber="1789">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D1BFFE65115EEC6FC263873" box="[866,933,1274,1295]" pageId="6" pageNumber="1789" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="19C7EAA30D1BFFE651C5EEC6FC3E3873" bold="true" box="[946,957,1274,1295]" pageId="6" pageNumber="1789">5</emphasis>
) Lateral view (holotype). (
<emphasis id="19C7EAA30D1BFFE65351EF2CFEB23859" bold="true" box="[294,305,1296,1317]" pageId="6" pageNumber="1789">6</emphasis>
) dorsal view (paratype, type locality). (
<emphasis id="19C7EAA30D1BFFE650DCEF2CFD4C3859" bold="true" box="[683,719,1296,1317]" pageId="6" pageNumber="1789">7, 8</emphasis>
)
<taxonomicName id="ECB34D320D1BFFE65097EF2CFCD0385A" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[736,851,1296,1318]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1789" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D1BFFE65097EF2CFCD0385A" box="[736,851,1296,1318]" italics="true" pageId="6" pageNumber="1789">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D1BFFE6512CEF2CFC1D3859" box="[859,926,1296,1317]" pageId="6" pageNumber="1789" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="19C7EAA30D1BFFE651DAEF2CFC3B3859" bold="true" box="[941,952,1296,1317]" pageId="6" pageNumber="1789">7</emphasis>
) Lateral view (holotype). (
<emphasis id="19C7EAA30D1BFFE65354EF1BFEAD3840" bold="true" box="[291,302,1319,1340]" pageId="6" pageNumber="1789">8</emphasis>
) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm.
</paragraph>
</caption>
<paragraph id="2B0C36B10D1BFFE652C5EF67FD38380D" blockId="6.[178,699,1371,1393]" box="[178,699,1371,1393]" pageId="6" pageNumber="1789">Hind wings: Reduced, hardly half as long as elytra.</paragraph>
<paragraph id="2B0C36B10D1BFFE652C5EFBCFD1A3B3A" blockId="6.[178,1086,1408,1606]" pageId="6" pageNumber="1789">Ventral surface: Prosternum, proepisternum, anterior part of mesepisternum, and metepisternum with sparse, fine setae, prosternal process with long setae apically. Metepisternum relatively short, ratio of anterior margin/internal margin (visible parts) about 0.87, moderately narrowed behind. Abdominal sternites III-V with scattered setae of unequal length, some of them arranged in suggested transverse rows about at middle of sternites. Last sternite, beside some fine scattered setae with two pore punctures bearing a long seta on each side at apical margin in both sexes.</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5EAE7FD3F3CE7" blockId="7.[178,1086,218,411]" pageId="7" pageNumber="1790">
Legs: Normal for
<taxonomicName id="ECB34D320D1AFFE7531EEAE6FD9D3D8C" box="[361,542,218,240]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D1AFFE7531EEAE6FE443D93" box="[361,455,218,239]" italics="true" pageId="7" pageNumber="1790">Acinopus</emphasis>
species.
</taxonomicName>
Pro- and mesotarsi indistinctly dilated in males (only somewhat wider than in females), with pro- and mesotarsomere 2 to 4 only in apical half with biseriate adhesive vestiture. Ventroapical tubercle of protibia with about four to five spines, arranged in a transverse row. Spines of lower surface of fore tibia arranged in one row of about eight spines, often a second, internal row, somewhat irregular and consisting of spines lower in number. Outer distal margin of protibia with about seven to nine spines. Tarsi smooth on superior surface except obligatory setae.
</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5EB96FD6E3F69" blockId="7.[178,1086,426,533]" pageId="7" pageNumber="1790">Microsculpture of surface: Head and pronotum in males with weakly engraved somewhat irregular slightly transverse meshes, elytra with the same kind of meshes somewhat less strongly engraved and more confused as on the forebody, surface very shiny, in females meshes somewhat stronger engraved, surface less shiny.</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5E818FE093FEC" blockId="7.[178,1085,548,656]" pageId="7" pageNumber="1790">
Median lobe of aedeagus (
<figureCitation id="B3882A340D1AFFE753B1E818FDA53F46" box="[454,550,548,570]" captionStart="Figs 5-8" captionStartId="6.[178,218,1273,1295]" captionTargetBox="[198,1071,235,1238]" captionTargetId="figure-0@6.[178,1085,204,1274]" captionTargetPageId="6" captionText="Figs 5-8: Acinopus DEJEAN, 1821. Median lobe of male. (5, 6) A. orszuliki nov.sp. (5) Lateral view (holotype). (6) dorsal view (paratype, type locality). (7, 8) A. sinaiticus nov.sp. (7) Lateral view (holotype). (8) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="7" pageNumber="1790">Figs 5, 6</figureCitation>
): Of normal construction for
<taxonomicName id="ECB34D320D1AFFE7512DE818FB8D3F46" box="[858,1038,548,570]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D1AFFE7512DE818FC3B3F45" box="[858,952,548,569]" italics="true" pageId="7" pageNumber="1790">Acinopus</emphasis>
species
</taxonomicName>
: terminal lamella somewhat elongate-triangular (dorsal view), and somewhat reflexed apically (lateral view), internal sac with a large, elongate group of about 40-50 mediumsized to small spines.
</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5E8A2FD343E74" blockId="7.[178,1085,670,776]" pageId="7" pageNumber="1790">
Female genitalia (
<figureCitation id="B3882A340D1AFFE7531CE8A2FE343FC8" box="[363,439,670,692]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="7" pageNumber="1790">Fig. 13</figureCitation>
): Hemisternite asymmetrical, with numerous scattered setae in its internal, somewhat hyaline part. Gonocoxite 1 elongate, apically with numerous setae, gonocoxite 2 scoop-shaped, apically somewhat blunt, with a double-sensilla in a furrow, and with numerous fine long setae at concave side.
</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5E92BFCC4391A" blockId="7.[178,1086,790,1126]" pageId="7" pageNumber="1790">
I n t r a s p e c i f i c v a r i a b i l i t y: All external characters have a great variability (as normal in
<taxonomicName id="ECB34D320D1AFFE753F7E90FFE5D3E34" authorityName="DEJEAN" authorityYear="1821" box="[384,478,819,840]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D1AFFE753F7E90FFE5D3E34" box="[384,478,819,840]" italics="true" pageId="7" pageNumber="1790">Acinopus</emphasis>
</taxonomicName>
), this concerns mainly the excision of the dorsal edge of the right mandible and its position (somewhat variable in size and strength, moderately strong to shallow, proximal (basal) angle of excision as a rule situated somewhat anterior to somewhat behind level of clypeus), the form of the pronotum, especially the construction of the anterior angles (distinctly, acutely projecting forward, only narrowly rounded at tip in most specimens, sometimes less distinctly projecting forward), the excavation of the base (distinctly excavate in most specimens, with posterior angles obtuse-angled, moderately rounded at tip, distinctly projecting backwards, sometimes less distinctly excavate with posterior angles less distinctly projecting backwards), and the form of the elytra (longer or shorter cylindrical), also the elytral pore puncture in interval 3 can be absent on one or both elytra. A variability in geographical aspect cannot be stated.
</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5EE48FCCB3986" blockId="7.[178,1085,1140,1274]" pageId="7" pageNumber="1790">C o m p a r i s o n s: It's the combination of colour, the male pro- and mesotarsomeres only weakly enlarged, the labrum only weakly incised apically, the anterior pronotal angles distinctly, acutely projecting forward, the pronotal base excavate with posterior angles shifted backward, and by the hind wings reduced (and connected with that, the metepisterna short) make this species distinct from its congeners.</paragraph>
<paragraph id="2B0C36B10D1AFFE752C5EF35FC2B3B63" blockId="7.[178,1085,1288,1567]" pageId="7" pageNumber="1790">
Easily distinguished from
<taxonomicName id="ECB34D320D1AFFE753BFEF35FD673862" authority="(OLIVIER, 1795)" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[456,740,1288,1310]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D1AFFE753BFEF35FDB33862" box="[456,560,1289,1310]" italics="true" pageId="7" pageNumber="1790">A. picipes</emphasis>
(OLIVIER, 1795)
</taxonomicName>
, a species having been recorded syntopic with the new species, and
<taxonomicName id="ECB34D320D1AFFE75045EF19FCFC3846" authority="MENETRIES, 1832" authorityName="MENETRIES" authorityYear="1832" box="[562,895,1316,1338]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D1AFFE75045EF19FD383846" box="[562,699,1317,1338]" italics="true" pageId="7" pageNumber="1790">A. laevigatus</emphasis>
MÉNÉTRIÉS, 1832
</taxonomicName>
, otherwise widely distributed in the
<collectingCountry id="53A476210D1AFFE7531AEF7EFE15382B" box="[365,406,1346,1367]" name="Iran" pageId="7" pageNumber="1790">Iran</collectingCountry>
but not yet known from the
<collectingRegion id="E977F8530D1AFFE750BBEF7EFC07382B" box="[716,900,1345,1367]" country="Iran" name="Kerman" pageId="7" pageNumber="1790">Kerman Province</collectingRegion>
, by its dark rufopiceous colour and the only indistinctly widened male pro- and mesotarsomes, by its pronotum with base distinctly excavate, with posterior angles less rounded at apex and somewhat directed backwards, and with anterior angles acutely protruding, and by its hind wings reduced. The latter two species are blackish in mature condition, have distinctly widened male pro- and mesotarsomes, a pronotum with base rectilinear, with posterior angles widely rounded, and with anterior angles less acutely protruding. Additionally,
<taxonomicName id="ECB34D320D1AFFE7537BEC36FEEC3B63" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[268,367,1546,1567]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1790" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D1AFFE7537BEC36FEEC3B63" box="[268,367,1546,1567]" italics="true" pageId="7" pageNumber="1790">A. picipes</emphasis>
</taxonomicName>
has the excision of the right mandible close to the base.
</paragraph>
<caption id="7FCC66390D15FFE852C5EFD5FD1F3B50" ID-DOI="http://doi.org/10.5281/zenodo.5356282" ID-Zenodo-Dep="5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="8" pageNumber="1791" startId="8.[178,218,1513,1535]" targetBox="[209,1036,218,1469]" targetPageId="8">
<paragraph id="2B0C36B10D15FFE852C5EFD5FD1F3B50" blockId="8.[178,1085,1513,1581]" pageId="8" pageNumber="1791">
<emphasis id="19C7EAA30D15FFE852C5EFD5FE8E3882" bold="true" box="[178,269,1513,1535]" pageId="8" pageNumber="1791">Figs 9-12</emphasis>
:
<taxonomicName id="ECB34D320D15FFE8536CEFD6FD833882" authority="DEJEAN, 1821" authorityName="DEJEAN" authorityYear="1821" box="[283,512,1513,1535]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1791" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D15FFE8536CEFD6FEEC3883" box="[283,367,1514,1535]" italics="true" pageId="8" pageNumber="1791">Acinopus</emphasis>
DEJEAN, 1821
</taxonomicName>
. Median lobe of male. (
<emphasis id="19C7EAA30D15FFE85091EFD5FC943882" bold="true" box="[742,791,1513,1534]" pageId="8" pageNumber="1791">9, 10</emphasis>
)
<taxonomicName id="ECB34D320D15FFE85150EFD6FC043882" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[807,903,1513,1535]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1791" phylum="Arthropoda" rank="species" species="brittoni" status="sp. nov.">
<emphasis id="19C7EAA30D15FFE85150EFD6FC043882" box="[807,903,1513,1535]" italics="true" pageId="8" pageNumber="1791">A. brittoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D15FFE851E7EFD6FC513883" box="[912,978,1514,1535]" pageId="8" pageNumber="1791" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="19C7EAA30D15FFE85196EFD5FC6F3882" bold="true" box="[993,1004,1513,1534]" pageId="8" pageNumber="1791">9</emphasis>
) Lateral view (holotype). (
<emphasis id="19C7EAA30D15FFE85320EC3CFEED3B69" bold="true" box="[343,366,1536,1557]" pageId="8" pageNumber="1791">10</emphasis>
) dorsal view (paratype). (
<emphasis id="19C7EAA30D15FFE85028EC3CFD1A3B69" bold="true" box="[607,665,1536,1557]" pageId="8" pageNumber="1791">11, 12</emphasis>
)
<taxonomicName id="ECB34D320D15FFE850D0EC3CFC913B69" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[679,786,1536,1557]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1791" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="19C7EAA30D15FFE850D0EC3CFC913B69" box="[679,786,1536,1557]" italics="true" pageId="8" pageNumber="1791">A. arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D15FFE8516FEC3CFCD83B69" box="[792,859,1536,1557]" pageId="8" pageNumber="1791" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="19C7EAA30D15FFE8511FEC3CFCFC3B69" bold="true" box="[872,895,1536,1557]" pageId="8" pageNumber="1791">11</emphasis>
) Lateral view (paratype). (
<emphasis id="19C7EAA30D15FFE85283EC2AFE883B57" bold="true" box="[244,267,1558,1579]" pageId="8" pageNumber="1791">12</emphasis>
) dorsal view (holotype). Scale bar: 1.6 mm.
</paragraph>
</caption>
<caption id="7FCC66390D14FFE952C5EFD9FE623B43" ID-DOI="http://doi.org/10.5281/zenodo.5356284" ID-Zenodo-Dep="5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="9" pageNumber="1792" startId="9.[178,218,1509,1531]" targetBox="[257,975,222,1478]" targetPageId="9">
<paragraph id="2B0C36B10D14FFE952C5EFD9FE623B43" blockId="9.[178,1086,1509,1599]" pageId="9" pageNumber="1792">
<emphasis id="19C7EAA30D14FFE952C5EFD9FE953886" bold="true" box="[178,278,1509,1531]" pageId="9" pageNumber="1792">Figs 13-16</emphasis>
:
<taxonomicName id="ECB34D320D14FFE95353EFDAFD863886" authority="DEJEAN, 1821" authorityName="DEJEAN" authorityYear="1821" box="[292,517,1509,1531]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1792" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D14FFE95353EFDAFEFB3887" box="[292,376,1510,1531]" italics="true" pageId="9" pageNumber="1792">Acinopus</emphasis>
DEJEAN, 1821
</taxonomicName>
. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (
<emphasis id="19C7EAA30D14FFE9528CEFC0FE913B6D" bold="true" box="[251,274,1532,1553]" pageId="9" pageNumber="1792">13</emphasis>
)
<taxonomicName id="ECB34D320D14FFE95356EFC0FE083B6D" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[289,395,1532,1553]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1792" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D14FFE95356EFC0FE083B6D" box="[289,395,1532,1553]" italics="true" pageId="9" pageNumber="1792">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D14FFE953E5EFC0FE563B6D" box="[402,469,1532,1553]" pageId="9" pageNumber="1792" rank="species">nov.sp.</taxonomicNameLabel>
(paratype, type locality). (
<emphasis id="19C7EAA30D14FFE950B8EFC0FD653B6D" bold="true" box="[719,742,1532,1553]" pageId="9" pageNumber="1792">14</emphasis>
)
<taxonomicName id="ECB34D320D14FFE95081EFC0FCEB3B6E" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[758,872,1532,1554]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1792" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D14FFE95081EFC0FCEB3B6E" box="[758,872,1532,1554]" italics="true" pageId="9" pageNumber="1792">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D14FFE95107EFC0FC303B6D" box="[880,947,1532,1553]" pageId="9" pageNumber="1792" rank="species">nov.sp.</taxonomicNameLabel>
(paratype, Mt. Katherine). (
<emphasis id="19C7EAA30D14FFE9535EEC2EFEC33B5B" bold="true" box="[297,320,1554,1575]" pageId="9" pageNumber="1792">15</emphasis>
)
<taxonomicName id="ECB34D320D14FFE95327EC2FFE323B54" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[336,433,1555,1576]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1792" phylum="Arthropoda" rank="species" species="brittoni" status="sp. nov.">
<emphasis id="19C7EAA30D14FFE95327EC2FFE323B54" box="[336,433,1555,1576]" italics="true" pageId="9" pageNumber="1792">A. brittoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D14FFE953CDEC2FFE7E3B54" box="[442,509,1555,1576]" pageId="9" pageNumber="1792" rank="species">nov.sp.</taxonomicNameLabel>
(paratype, type locality). (
<emphasis id="19C7EAA30D14FFE9508CEC2EFC913B5B" bold="true" box="[763,786,1554,1575]" pageId="9" pageNumber="1792">16</emphasis>
)
<taxonomicName id="ECB34D320D14FFE95155EC2FFC0D3B54" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[802,910,1555,1576]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1792" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="19C7EAA30D14FFE95155EC2FFC0D3B54" box="[802,910,1555,1576]" italics="true" pageId="9" pageNumber="1792">A. arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D14FFE951E0EC2FFC593B54" box="[919,986,1555,1576]" pageId="9" pageNumber="1792" rank="species">nov.sp.</taxonomicNameLabel>
(paratype, type locality). Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="7FCC66390D17FFEA52C5E9B9FE503ECE" ID-DOI="http://doi.org/10.5281/zenodo.5356286" ID-Zenodo-Dep="5356286" httpUri="https://zenodo.org/record/5356286/files/figure.png" pageId="10" pageNumber="1793" startId="10.[178,211,901,923]" targetBox="[178,1085,330,892]" targetPageId="10">
<paragraph id="2B0C36B10D17FFEA52C5E9B9FE503ECE" blockId="10.[178,1085,901,946]" pageId="10" pageNumber="1793">
<emphasis id="19C7EAA30D17FFEA52C5E9B9FF753EE6" bold="true" box="[178,246,901,923]" pageId="10" pageNumber="1793">Fig. 17</emphasis>
: One of the localities where
<taxonomicName id="ECB34D320D17FFEA5072E9BAFDED3EE7" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[517,622,902,923]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D17FFEA5072E9BAFDED3EE7" box="[517,622,902,923]" italics="true" pageId="10" pageNumber="1793">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D17FFEA5001E9BAFD3B3EE7" box="[630,696,902,923]" pageId="10" pageNumber="1793" rank="species">nov.sp.</taxonomicNameLabel>
was found (Mahan, SE Kerman, 2500 m,
<emphasis id="19C7EAA30D17FFEA52C5E9A0FE893ECD" box="[178,266,924,945]" italics="true" pageId="10" pageNumber="1793">Artemisia</emphasis>
-high-altitude steppe).
</paragraph>
</caption>
<paragraph id="2B0C36B10D17FFEA52C5E9FCFDB23915" blockId="10.[178,1085,960,1129]" pageId="10" pageNumber="1793">
Similar to
<taxonomicName id="ECB34D320D17FFEA5357E9FDFE1C3EAA" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[288,415,961,982]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D17FFEA5357E9FDFE1C3EAA" box="[288,415,961,982]" italics="true" pageId="10" pageNumber="1793">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D17FFEA53D0E9FDFE723EAA" box="[423,497,961,982]" pageId="10" pageNumber="1793" rank="species">nov.sp.</taxonomicNameLabel>
in its body colour but this species has the male pro- and mesotarsomeres more strongly enlarged, the labrum distinctly incised apically, the anterior pronotal angles not as strong and less acutely projecting forward, the pronotum laterally anterior to the posterior angles somewhat sinuate and with its base more or less rectilinear, the elytra somewhat flattened, the hind wings fully developed (folded anterior to apex), and the metepisterna longer.
</paragraph>
<paragraph id="2B0C36B10D17FFEA52C5EE44FD6338AD" blockId="10.[178,1085,1144,1489]" pageId="10" pageNumber="1793">
The new species differs distinctly from
<taxonomicName id="ECB34D320D17FFEA501AEE45FEC039D0" authority="AZADBAKHSH &amp; WRASE, 2016" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis" subGenus="Acinopus">
<emphasis id="19C7EAA30D17FFEA501AEE45FD0039F2" box="[621,643,1145,1166]" italics="true" pageId="10" pageNumber="1793">A.</emphasis>
(
<emphasis id="19C7EAA30D17FFEA50EEEE45FD7439F2" box="[665,759,1145,1166]" italics="true" pageId="10" pageNumber="1793">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D17FFEA517BEE45FC0139F2" box="[780,898,1145,1166]" italics="true" pageId="10" pageNumber="1793">zagrosensis</emphasis>
AZADBAKHSH &amp; WRASE, 2016
</taxonomicName>
, recently described from the Zagros Mts. in western
<collectingCountry id="53A476210D17FFEA510AEEAAFC2B39D7" box="[893,936,1174,1195]" name="Iran" pageId="10" pageNumber="1793">Iran</collectingCountry>
, by its colour (lighter to darker rufo-piceous or piceous, in
<taxonomicName id="ECB34D320D17FFEA50F7EE8FFC9039B5" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" box="[640,787,1203,1225]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="19C7EAA30D17FFEA50F7EE8FFC9039B5" box="[640,787,1203,1225]" italics="true" pageId="10" pageNumber="1793">A. zagrosensis</emphasis>
</taxonomicName>
black), by the pronotal anterior angles distinctly, acutely projecting forward (in
<taxonomicName id="ECB34D320D17FFEA50A4EEEDFCE4399A" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" box="[723,871,1233,1254]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="19C7EAA30D17FFEA50A4EEEDFCE4399A" box="[723,871,1233,1254]" italics="true" pageId="10" pageNumber="1793">A. zagrosensis</emphasis>
</taxonomicName>
anterior angles only moderately and somewhat less acutely projecting forward), by the posterior angles obtuse-angled, only moderately rounded at tip (in
<taxonomicName id="ECB34D320D17FFEA50A5EF37FCE9385D" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" box="[722,874,1291,1313]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="19C7EAA30D17FFEA50A5EF37FCE9385D" box="[722,874,1291,1313]" italics="true" pageId="10" pageNumber="1793">A. zagrosensis</emphasis>
</taxonomicName>
widedy rounded at apex). While the new species has the hind wings only reduced on about half of the elytral lengths,
<taxonomicName id="ECB34D320D17FFEA537BEF7AFE203827" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" box="[268,419,1350,1371]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="19C7EAA30D17FFEA537BEF7AFE203827" box="[268,419,1350,1371]" italics="true" pageId="10" pageNumber="1793">A. zagrosensis</emphasis>
</taxonomicName>
has the hindwings reduced to only small scales. The external construction of the median lobe is similar in both species, but in
<taxonomicName id="ECB34D320D17FFEA5126EF5FFC463804" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[849,965,1379,1400]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D17FFEA5126EF5FFC463804" box="[849,965,1379,1400]" italics="true" pageId="10" pageNumber="1793">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D17FFEA51BAEF58FB943805" box="[973,1047,1380,1401]" pageId="10" pageNumber="1793" rank="species">nov.sp.</taxonomicNameLabel>
the internal sac has a larger group with spines more numerous (about 40-50), while in
<taxonomicName id="ECB34D320D17FFEA5650EFBDFEAB38CF" authorityName="AZADBAKHSH &amp; WRASE" authorityYear="2016" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="19C7EAA30D17FFEA5650EFBDFEAB38CF" italics="true" pageId="10" pageNumber="1793">A. zagrosensis</emphasis>
</taxonomicName>
the group is smaller by spines less numerous (about 10 to 18, compare
<figureCitation id="B3882A340D17FFEA5666EFA2FF5B38AD" captionStart="Figs 5-8" captionStartId="6.[178,218,1273,1295]" captionTargetBox="[198,1071,235,1238]" captionTargetId="figure-0@6.[178,1085,204,1274]" captionTargetPageId="6" captionText="Figs 5-8: Acinopus DEJEAN, 1821. Median lobe of male. (5, 6) A. orszuliki nov.sp. (5) Lateral view (holotype). (6) dorsal view (paratype, type locality). (7, 8) A. sinaiticus nov.sp. (7) Lateral view (holotype). (8) dorsal view (paratype, Mt. Katherine). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356280" httpUri="https://zenodo.org/record/5356280/files/figure.png" pageId="10" pageNumber="1793">Figs 5, 6</figureCitation>
with
<figureCitation id="B3882A340D17FFEA5365EF87FECE38AC" box="[274,333,1467,1489]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="10" pageNumber="1793">
Fig.
<quantity id="EC4B9B540D17FFEA5336EF87FECE38AC" box="[321,333,1467,1488]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="1793" unit="in" value="2.0">2</quantity>
</figureCitation>
in AZADBAKHSH &amp; WRASE 2016: 585).
</paragraph>
<paragraph id="2B0C36B10D17FFEA52C5EFDDFBA13B4D" blockId="10.[178,1086,1504,1585]" pageId="10" pageNumber="1793">
D i s t r i b u t i o n: Up to now only known from several localities in the mountainous regions of the Iranian Province of
<collectingRegion id="E977F8530D17FFEA538BEFC2FDCE3B6F" box="[508,589,1534,1555]" country="Iran" name="Kerman" pageId="10" pageNumber="1793">Kerman</collectingRegion>
, in altitudinal ranges of about
<quantity id="EC4B9B540D17FFEA5107EFC2FB8F3B6F" box="[880,1036,1534,1555]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="10" pageNumber="1793" unit="m" value="2800.0" valueMax="3400.0" valueMin="2200.0">2200 to 3400 m</quantity>
. Due to its inability to fly most likely a wider occurrence of
<taxonomicName id="ECB34D320D17FFEA50CAEC27FCA93B4C" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[701,810,1563,1584]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1793" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D17FFEA50CAEC27FCA93B4C" box="[701,810,1563,1584]" italics="true" pageId="10" pageNumber="1793">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D17FFEA5158EC20FCF53B4D" box="[815,886,1564,1585]" pageId="10" pageNumber="1793" rank="species">nov.sp.</taxonomicNameLabel>
can be excluded.
</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5EAE7FE4F3C85" blockId="11.[178,1085,219,505]" pageId="11" pageNumber="1794">
H a b i t a t a n d b i o n o m i c n o t e s The specimens, collected by R. Felix and J. Muilwijk in the Kūh-e Lālehzār massif were found in a green, grassy valley partly covered by snow, with low shrubs, along a little stream and in a more open stony and grassy hillside also along this stream, the specimens from Mahan on poor, dry fields and field edges, while the specimens from Sirch and Qal ehAskar were collected on grassland (R. Felix, pers. comm.). The specimens from Sinabad were found in an orchard by pit fall traps. Finally, the specimens, recently collected by S. Azadbakhsh, B. Laser, P. Schnitter, A. Weigel, and D.W. Wrase (see above under
<typeStatus id="F40888130D16FFEB5082EB94FCAD3CC1" box="[757,814,424,445]" pageId="11" pageNumber="1794">types</typeStatus>
), were taken in daytime in
<emphasis id="19C7EAA30D16FFEB52C5EBFAFE973CA7" box="[178,276,454,475]" italics="true" pageId="11" pageNumber="1794">Artemisia</emphasis>
-high-altitude steppes (
<figureCitation id="B3882A340D16FFEB5389EBF9FDCB3CA0" box="[510,584,453,476]" captionStart="Fig" captionStartId="10.[178,211,901,923]" captionTargetBox="[178,1085,330,892]" captionTargetId="figure-0@10.[178,1085,212,892]" captionTargetPageId="10" captionText="Fig. 17: One of the localities where A. orszuliki nov.sp. was found (Mahan, SE Kerman, 2500 m, Artemisia-high-altitude steppe)." figureDoi="http://doi.org/10.5281/zenodo.5356286" httpUri="https://zenodo.org/record/5356286/files/figure.png" pageId="11" pageNumber="1794">Fig. 17</figureCitation>
) and in an orchard, mainly from under stones, or running around at darkness.
</paragraph>
<paragraph id="2B0C36B10D16FFEB52C5E834FCFE3E5A" blockId="11.[178,1086,520,806]" pageId="11" pageNumber="1794">
Interestingly, the series of
<taxonomicName id="ECB34D320D16FFEB53A8E834FD163F62" box="[479,661,520,542]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D16FFEB53A8E834FDBE3F61" box="[479,573,520,541]" italics="true" pageId="11" pageNumber="1794">Acinopus</emphasis>
species
</taxonomicName>
from the
<collectingRegion id="E977F8530D16FFEB5167E834FC493F62" box="[784,970,520,542]" country="Iran" name="Kerman" pageId="11" pageNumber="1794">Kerman Province</collectingRegion>
we could examine, collected in March to middle of May, contained only
<specimenCount id="3DB5FD380D16FFEB5116E81AFC783F47" box="[865,1019,549,571]" pageId="11" pageNumber="1794" type="generic">two specimens</specimenCount>
of
<taxonomicName id="ECB34D320D16FFEB5650E81AFF793F24" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D16FFEB5650E81AFF793F24" italics="true" pageId="11" pageNumber="1794">A. picipes</emphasis>
</taxonomicName>
(Golmalek nr. Dehbakry, ca
<quantity id="EC4B9B540D16FFEB505CE87FFDEF3F25" box="[555,620,579,601]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="1794" unit="km" value="55.0">55 km</quantity>
N Jiroft,
<date id="5F0D10710D16FFEB50A5E87FFCC43F25" box="[722,839,579,601]" pageId="11" pageNumber="1794" value="2006-03-26">26.III.2006</date>
, W. Heinz leg.,
<specimenCount id="3DB5FD380D16FFEB5186E87FFBB93F25" box="[1009,1082,579,601]" pageId="11" pageNumber="1794" type="male">1 male</specimenCount>
, cHZ; Jiroft, Anbar-Abad,
<date id="5F0D10710D16FFEB53B3E85CFDCE3F0A" box="[452,589,608,630]" pageId="11" pageNumber="1794" value="1956-05-01" valueMax="1956-05-18" valueMin="1956-05-01">1.-18.V.1956</date>
, W. Richter leg.,
<specimenCount id="3DB5FD380D16FFEB517AE85CFCE93F0A" box="[781,874,608,630]" pageId="11" pageNumber="1794" type="female">1 female</specimenCount>
, cWR), contrary, in findings from May 30 to June 7 the series comprised mainly
<taxonomicName id="ECB34D320D16FFEB5154E842FC0B3FEF" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[803,904,638,659]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D16FFEB5154E842FC0B3FEF" box="[803,904,638,659]" italics="true" pageId="11" pageNumber="1794">A. picipes</emphasis>
</taxonomicName>
(often in more or less immature condition) and only a few specimens of the new species, the latter often with strongly worn tibiae and tarsi, also with lacking parts of the appendages,
<specimenCount id="3DB5FD380D16FFEB51A2E884FF5D3F97" pageId="11" pageNumber="1794" type="generic">one specimen</specimenCount>
was found dead, so seemingly both species which could be registered syntopic in three localities, share the same localities, with
<taxonomicName id="ECB34D320D16FFEB50D9E8CFFCA53E74" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[686,806,755,776]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="19C7EAA30D16FFEB50D9E8CFFCA53E74" box="[686,806,755,776]" italics="true" pageId="11" pageNumber="1794">A. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D16FFEB5145E8CFFC1D3E74" box="[818,926,755,776]" pageId="11" pageNumber="1794" rank="species">nov. spec.</taxonomicNameLabel>
as the species earlier appearing, and disappearing when
<taxonomicName id="ECB34D320D16FFEB502EE92CFD3D3E5A" baseAuthorityName="OLIVIER" baseAuthorityYear="1795" box="[601,702,784,806]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1794" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="19C7EAA30D16FFEB502EE92CFD3D3E5A" box="[601,702,784,806]" italics="true" pageId="11" pageNumber="1794">A. picipes</emphasis>
</taxonomicName>
emerges as adults.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

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<taxonomicName id="5DCDAC2551C68E167FA159E2B790988E" ID-CoL="XTKP" authority="Cameron, 1881" authorityName="Cameron" authorityYear="1881" class="Insecta" family="Megaspilidae" genus="Conostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Conostigmus mullensis" order="Hymenoptera" pageId="0" pageNumber="1167" phylum="Arthropoda" rank="species" species="mullensis">Conostigmus mullensis (Cameron, 1881)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1122E5364AB5DEC1F70FD981BAF051D4" pageId="0" pageNumber="1167" type="reference_group">
<paragraph id="08F514A937E4EBC9153DBBD55D9FC626" pageId="0" pageNumber="1167">
<taxonomicName id="9FE82DF133CE340652C4D2F3974BECFA" class="Insecta" family="Megaspilidae" genus="Megaspilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaspilus mullensis" order="Hymenoptera" pageId="0" pageNumber="1167" phylum="Arthropoda" rank="species" species="mullensis">Megaspilus mullensis</taxonomicName>
Cameron, 1881
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</subSubSection>
<subSubSection id="1DE53793FD9B55D45FC41C556F082BE7" pageId="0" pageNumber="1167" type="distribution">
<paragraph id="8F5C0311D0DB39FF5385ECFF667F348D" pageId="0" pageNumber="1167">Distribution</paragraph>
<paragraph id="C6AC4BBCD374674D26A4CCC552C97009" pageId="0" pageNumber="1167">Scotland</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Tournier, 1877" authorityName="Tournier" authorityYear="1877" class="Insecta" family="Gasteruptiidae" genus="Gasteruption" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gasteruption laticeps" order="Hymenoptera" pageId="0" pageNumber="1116" phylum="Arthropoda" rank="species" species="laticeps">Gasteruption laticeps (Tournier, 1877)</taxonomicName>
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<taxonomicName class="Insecta" family="Gasteruptiidae" genus="Foenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Foenus laticeps" order="Hymenoptera" pageId="0" pageNumber="1116" phylum="Arthropoda" rank="species" species="laticeps">Foenus laticeps</taxonomicName>
Tournier, 1877
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<taxonomicName class="Insecta" family="Gasteruptiidae" genus="Gasteruption" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gasteruption foveolatum" order="Hymenoptera" pageId="0" pageNumber="1116" phylum="Arthropoda" rank="species" species="foveolatum">Gasteruption foveolatum</taxonomicName>
Schletterer, 1889
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<taxonomicName class="Insecta" family="Gasteruptiidae" genus="Gasteruption" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gasteruption foveolum" order="Hymenoptera" pageId="0" pageNumber="1116" phylum="Arthropoda" rank="species" species="foveolum">Gasteruption foveolum</taxonomicName>
<normalizedToken originalValue="Szépligeti">Szepligeti</normalizedToken>
, 1903
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