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<mods:title id="8BE8F125B945640DA42F12F320219EFD">Growing giants: ontogeny and life history of the temnospondyl Mastodonsaurus giganteus (Stereospondyli) from the Middle Triassic of Germany</mods:title>
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<mods:namePart id="D1EF7244BF06DF15A17FC54E698458E3">Witzmann, Florian</mods:namePart>
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<mods:namePart id="D6361D2D742BC8D3C507D30C779B2ED6">Moreno, Raphael</mods:namePart>
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<mods:namePart id="BE9A916855EBB2119420F046D1CA99CF">Mujal, Eudald</mods:namePart>
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<mods:affiliation id="8CFBBCE4980F19E124E20F5DFD059EE7">Staatliches Museum fu ̈ r Naturkunde, Rosenstein 1, D- 70191 Stuttgart, Germany &amp; Institut Català de Paleontologia Miquel Crusafont (ICP-CERCA), Universitat Autònoma de Barcelona, Edifici ICTA-ICP, c / Columnes s / n, Campus de la UAB, E- 08193 Cerdanyola del Vallès, Catalonia, Spain</mods:affiliation>
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<mods:date id="8E541441D90435CDE3F8FF3F8214F214">2024</mods:date>
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<paragraph id="56FC2D563BA7B2AF0D61CFF357F5B4F4">
<heading id="B20DDD111D317688682B7670C8DD561C" reason="label">Genus</heading>
<taxonomicName id="4468D26D0006DB60C6CBFEA483C1797C" authority="Jaeger, 1828" authorityName="Jaeger" authorityYear="1828" class="Amphibia" family="Mastodonsauridae" genus="Mastodonsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Temnospondyli" phylum="Chordata" rank="genus">
<emphasis id="9680E61052CBEBF5D9512F5714B420AF" italics="true">Mastodonsaurus</emphasis>
<bibRefCitation id="E28DE159C0B03F84DD9E522A8B3DAB9D" DOI="10.5962/bhl.title.4689" author="Jaeger GF" journalOrPublisher="Metzler, Stuttgart" refId="B10" refString="Jaeger GF (1828) Ueber die fossile [sic] Reptilien, welche in Wurttemberg aufgefunden worden sind. Metzler, Stuttgart, 48 pp. https: // doi. org / 10.5962 / bhl. title. 4689" title="Ueber die fossile [sic] Reptilien, welche in Württemberg aufgefunden worden sind." year="1828">Jaeger, 1828</bibRefCitation>
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</paragraph>
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<paragraph id="11A072A871CCD9E4F2940F9A7866531F">
<heading id="AE9C26F49D2A84BDE7841CFD2DF5D4C6" reason="title">
<typeStatus id="5A6869F0F2A30E02D6B0FD2F94FF4362">Type</typeStatus>
species.
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</paragraph>
<paragraph id="9D0DF215BF1B0072752F934748D004B4">
<taxonomicName id="9596ADB067C874DF5BB3028B4C5B6843" authority="Jaeger, 1828" authorityName="Jaeger" authorityYear="1828" class="Amphibia" family="Mastodonsauridae" genus="Mastodonsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="F0B79D6F38C7A668CB6FA5CD332BD04C" italics="true">Mastodonsaurus giganteus</emphasis>
<bibRefCitation id="D339DD6ED1FD15C09191297BD2F28187" DOI="10.5962/bhl.title.4689" author="Jaeger GF" journalOrPublisher="Metzler, Stuttgart" refId="B10" refString="Jaeger GF (1828) Ueber die fossile [sic] Reptilien, welche in Wurttemberg aufgefunden worden sind. Metzler, Stuttgart, 48 pp. https: // doi. org / 10.5962 / bhl. title. 4689" title="Ueber die fossile [sic] Reptilien, welche in Württemberg aufgefunden worden sind." year="1828">Jaeger, 1828</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="SECID0E5DAE" type="diagnosis">
<paragraph id="98101CE5D28A5A4CFD6FC812DD696C75">
<heading id="D5FE5596C5C3C47F05A3DC969B3B6785" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="668139FEF825113DF633F70F8322D88F">
Autapomorphies (Figs
<figureCitation id="E039A4694F6BDF1102C5DA8C32EABEFE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mastodonsaurus giganteus Jaeger, 1828. Frequency and size distribution of specimens in six different localities and horizons of the Lower Keuper in northern Baden-Württemberg (Germany). Black squares: skulls or semi-articulated skeletons; Grey squares: single bones." figureDoi="10.3897/fr.27.125379.figure2" httpUri="https://binary.pensoft.net/fig/1206164">2</figureCitation>
<figureCitation id="40192761FDE7B7A8D86E259A34D12CCC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mandibles and postcranial material of Lower Keuper temnospondyls. A H. Symphyses. A. M. giganteus, SMNS 97035 (Kupferzell, K 3); B. M. giganteus, SMNS 97036 (Kupferzell, K 3); C. M. giganteus, MHI 1992 / 32 (Vellberg, E 6); D. M. giganteus, SMNS 81310 (Kupferzell, K 3); E. Tatrasuchus wildi, MHI - Ku 1992 / 42 (Vellberg, E 6); F. Tatrasuchus wildi, SMNS 54670 (Kupferzell, K 3); G. Callistomordax kugleri, SMNS 90520; H. Trematolestes hagdorni, SMNS 97034; I. M. giganteus, adult interclavicle in dorsal view, SMNS 97132 (Kupferzell, K 3); J. M. giganteus, small juvenile clavicle, SMNS 97044 (Vellberg, E 6); K. M. giganteus, tiny interclavicle in dorsal view, SMNS 97131 (Vellberg, E 6); L. M. giganteus, adult neural arch, SMNS 80913; M. M. giganteus, giant neural arch, SMNS 81310; N-Q. M. giganteus, intercentra in anterior view; N. MHI 1046; O. MHI 1079; P, Q. SMNS 80913." figureDoi="10.3897/fr.27.125379.figure6" httpUri="https://binary.pensoft.net/fig/1206168">6</figureCitation>
): (1) Premaxilla with openings for symphyseal tusks, set well anterior to naris; (2) orbits large, reaching one-fifth of skull length; (3) jugal slender lateral to orbit; (4) parietal elongated anterior to pineal foramen; (5) lateral line sulci very wide; (6) epipterygoid massive and complex, with six distinct processes (footplate, pr. anterior, pr. dorsalis, pr. sphenethmoidalis, pr. basalis, pr. posterior). Characters shared with other capitosaurs: (a) interclavicle with elongated anterior process; (b) elongate postglenoid area; (c) tall hamate process.
</paragraph>
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<subSubSection id="SECID0EMEAE" type="comments">
<paragraph id="688946DED9026223EFC4C6A0A0DA373C">
<heading id="C0DF35E14A90F38E636F2A25BCF927C4" reason="title">Comment.</heading>
</paragraph>
<paragraph id="5B35A4E6AD7AAD1469B137D87DF39A75">
Character (1) is shared with
<taxonomicName id="FFB9EFE7BD80194CC037C0A6A6DB1FBC" rank="species" species="naraserluki">
<emphasis id="A8A6FBA8FCE4A2413540D18E18B3819F" italics="true">C. naraserluki</emphasis>
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and
<taxonomicName id="B6CCF7DE28EFA89F08C0B30BA4A47EE5" rank="species" species="mordax">
<emphasis id="A247CEF2DCD5E8B336AAE9E64734B077" italics="true">C. mordax</emphasis>
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(Fraas 1913;
<bibRefCitation id="C0487B9E375E9E9D7691EA192C309154" DOI="10.1080/02724634.2017.1303501" author="Marzola" etAl="et al." firstAuthor="Marzola" refId="B14" refString="Marzola M, Mateus O, Shubin NH, Clemmensen LB (2017) Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology 73: e 1303501. https://doi.org/10.1080/02724634.2017.1303501" year="2017">Marzola et al. 2017</bibRefCitation>
) and also occurs in
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of
<taxonomicName id="3D439990EA88C9F5AB6D96512ED3D898" authority="(Schoch and Moreno 2024)" baseAuthorityName="Schoch and Moreno" baseAuthorityYear="2024" rank="species" species="robustus">
<emphasis id="DC6A38D70C439AEBD3FF3536255277A9" italics="true">C. robustus</emphasis>
(
<bibRefCitation id="D3179F0AD7B8A034BE86BB5510071701" DOI="10.18476/pale.v17.a2" author="Schoch and Moreno" firstAuthor="Schoch" journalOrPublisher="Paleodiversity" pagination="9-48" refId="B39" refString="Schoch RR, Moreno R (2024) Synopsis on the temnospondyls from the German Triassic. Paleodiversity 17: 948. https://doi.org/10.18476/pale.v17.a2" title="Synopsis on the temnospondyls from the German Triassic." volume="17" year="2024">Schoch and Moreno 2024</bibRefCitation>
)
</taxonomicName>
. However,
<taxonomicName id="18D081AC8AEA14B18231F52FDB2D7E64" authorityName="Jaeger" authorityYear="1828" class="Amphibia" family="Mastodonsauridae" genus="Mastodonsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" phylum="Chordata" rank="genus">
<emphasis id="89746842D5E1E44E26EA07CFBB3AD673" italics="true">Mastodonsaurus</emphasis>
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differs from all these in having a more elongated premaxilla anterior to the naris and a more lateral and posterior emplacement of the opening with respect to the anterior snout margin (
<bibRefCitation id="343102AF6FF6A9AD631388BF8EF64956" DOI="10.18476/pale.v17.a2" author="Schoch and Moreno" firstAuthor="Schoch" journalOrPublisher="Paleodiversity" pagination="9-48" refId="B39" refString="Schoch RR, Moreno R (2024) Synopsis on the temnospondyls from the German Triassic. Paleodiversity 17: 948. https://doi.org/10.18476/pale.v17.a2" title="Synopsis on the temnospondyls from the German Triassic." volume="17" year="2024">Schoch and Moreno 2024</bibRefCitation>
). Character (4) is shared with metoposaurids, but no other taxon with Capitosauroidea.
</paragraph>
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</treatment>
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