From a5c105ef0c2d0f279748c296b05daa008a21a854 Mon Sep 17 00:00:00 2001 From: ggserver Date: Thu, 5 Sep 2024 08:49:59 +0000 Subject: [PATCH] Add updates up until 2024-09-05 08:43:54 --- .../53/038F531FFFA3FFF23028FC0BFE24FD89.xml | 1344 +++++++++++++++++ .../53/038F531FFFB3FFE233A5F97AFC48FE54.xml | 679 +++++++++ .../53/038F531FFFB6FFEE302DFAF6FD4AF981.xml | 525 +++++++ .../53/038F531FFFC9FF96300DFD47FAEDF9DD.xml | 302 ++++ .../53/038F531FFFCCFF94301CFEE1FB0AFD67.xml | 552 +++++++ .../53/038F531FFFDBFF873069FE79FAE4FCC6.xml | 342 +++++ .../53/038F531FFFE7FFBB3002FEE1FAE4FB82.xml | 303 ++++ .../53/038F531FFFECFFB43016FEE1FE5DFAEB.xml | 820 ++++++++++ 8 files changed, 4867 insertions(+) create mode 100644 data/03/8F/53/038F531FFFA3FFF23028FC0BFE24FD89.xml create mode 100644 data/03/8F/53/038F531FFFB3FFE233A5F97AFC48FE54.xml create mode 100644 data/03/8F/53/038F531FFFB6FFEE302DFAF6FD4AF981.xml create mode 100644 data/03/8F/53/038F531FFFC9FF96300DFD47FAEDF9DD.xml create mode 100644 data/03/8F/53/038F531FFFCCFF94301CFEE1FB0AFD67.xml create mode 100644 data/03/8F/53/038F531FFFDBFF873069FE79FAE4FCC6.xml create mode 100644 data/03/8F/53/038F531FFFE7FFBB3002FEE1FAE4FB82.xml create mode 100644 data/03/8F/53/038F531FFFECFFB43016FEE1FE5DFAEB.xml diff --git a/data/03/8F/53/038F531FFFA3FFF23028FC0BFE24FD89.xml b/data/03/8F/53/038F531FFFA3FFF23028FC0BFE24FD89.xml new file mode 100644 index 00000000000..b484ffca315 --- /dev/null +++ b/data/03/8F/53/038F531FFFA3FFF23028FC0BFE24FD89.xml @@ -0,0 +1,1344 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus shoshone + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +E7F437ED-E934-4E01-9FCC-3151D1C3EF6F + + + + +Figs 12O, 15F, 30 + + +Proposed common name + + +Shoshone triepeolus. + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. shoshone + +sp. nov. +apart from all other + +Triepeolus + +in the + +simplex + +species group except + +T. apache + +sp. nov. +: the T1 basal band forms an inflexed arch and is thus gradually narrowed but not completely interrupted medially, such that the discal patch has a short, narrowed (i.e., inverted V-shaped) anteromedial projection ( +Figs 15F +, +30B +), and the legs are entirely reddish orange from trochanters to tarsi ( +Fig. 30A–C +). Whereas in + +T. apache + +the T1 apical transverse band is interrupted or (less commonly) at least narrowed medially and the T2 apical transverse band is narrowed or interrupted medially, in + +T. shoshone + +T2 and usually also T1 have complete apical transverse bands ( +Figs 15F +, +30B +). Although it might not be possible to separate certain individuals morphologically, throughout most of its range + +T. shoshone + +can be separated from + +T. apache + +by geography; the former is a northern species whose known range extends from northern Arizona and northern New Mexico to Western +Canada +whereas the latter is a southern species known only from the Southwestern +United States +and adjacent +Mexico +. + + + + + +Etymology + + +This species is named for the Shoshone people, whose traditional territory spans much of its known range. + + + + +Material examined + + + + +Primary +type +material + + + + +USA +• +♀ +, +holotype +; +Utah +, +Hwy +491, +San Juan County +; +37.8350° N +, +109.1110° W +; + +27 Jul. 2007 + +; +J. Gibbs +and +C.S. Sheffield +leg.; BOLD sample ID: CMNTO_063; +PCYU +PCYU-GS-07:2170 + +. + + + +Secondary +type +material + + + + +CANADA +– + +Alberta + +• +1 ♀ +, +paratype +; + +6 km +E of Writing-on-Stone Provincial Park + +(at bridge); + +12 Aug. 1982 + +; +D.B. McCorquodale +leg.; +RAM + +• + +1 ♀ +, +paratype +; +Lethbridge +; + +9Aug. 1921 + +; +H.L. Seamans +leg.; +CNC 753877 + +• + +2 ♀♀ +, +paratypes +; +Medicine Hat +; + +17 Jul. 1917 + +; +Sladen +leg.; +CNC 753824 +, +753826 + +. – + + +British Columbia + +• +1 ♀ +, +paratype +; +Fernie +; + +3 Aug. 1963 + +; +G. Bohart +and +P. Torchio +leg.; +BBSL +BBSL522497 + +. – + + +Saskatchewan + +• +1 ♀ +, +paratype +; +Grasslands National Park +; +49.1371° N +, +107.6149° W +; + +25 Jul. 1996 + +; +A.T. Finnamore +leg.; +RAM +PMAE 17947 + +• + +1 ♀ +, +paratype +; +Grasslands National Park +; +49.1643° N +, +107.4351° W +; + +23 Jul. 1996 + +; +A.T. Finnamore +leg.; +RAM +PMAE 16026 + +• + +1 ♀ +, +paratype +; +Maple Creek +Golf Course; +49.9021° N +, +109.4654° W +; + +13 Jul.–9 Aug. 2018 + +; RSM +Bee Team +leg.; BOLD sample ID: CCDB-38770 B12; +RSKM RSKM +_ENT_E-208865 + +• + +1 ♀ +, +paratype +; +Near Avonlea +; +50.0240° N +, +104.9650° W +; + +7 Jul. 2017 + +; +G. Wihlidal +leg.; +RSKM RSKM +_ENT_E-172173 + +• + +1 ♀ +, +paratype +; near +Avonlea Badlands +; +50.0229° N +, +104.9842° W +; + +29 Jul. 2016 + +; +C.S. Sheffield +leg.; BOLD sample ID: CCDB-25139 G11; +RSKM RSKM +_ENT_E-185221 + +. + + +USA – + +California + +• + +1 ♀ +, +paratype +; +Turlock +, +Stanislaus County +; + +1 Sep. 1953 + +; +R.R. Snelling +leg.; +USNM + +• + +1 ♀ +, +paratype +; same data as for preceding; + +18 Apr. 1957 + +; +R.R. Snelling +leg.; +USNM +. – + +Colorado + + +• + +2 ♀♀ +, +paratypes +; +C.F. Baker +leg.; +USNM + +• + +1 ♀ +, +paratype +; +Costilla County +; +Jul. +??20; +S. McCampbell +leg.; +CSUC + +• + +1 ♀ +, +paratype +; +Denver +; + +19 Jun. 1917 + +; +E.C. Jackson +leg.; +USNM +. – + +Idaho + + +• + +2 ♀♀ +, +paratypes +; + +4 mi +W of Homedale + +, +Owyhee County +; + +21 Jul. 1959 + +; +W.F. Barr +leg.; +BBSL BBSL +Faunal Survey Nos +000 086 098, 000 086 099 + +• + +1 ♀ +, +paratype +; +Boise +; + +15 Jun. 1941 + +; +CAS + +• + +1 ♀ +, +paratype +; +Grand View +; + +27 Jul. 1926 + +; +R.W. Haegele +leg.; +USNM + +• + +1 ♀ +, +paratype +; +Hot Springs +, +Owyhee County +; + +5 Jul. 1956 + +; +W.F. Barr +leg.; +USNM + +• + +1 ♀ +, +paratype +; +Melba +; + +18 Jun. 1926 + +; +R.W. Haegele +leg.; +USNM + +• + +1 ♀ +, +paratype +; +Oreana +; 25 +Jun. +195?; +A.R. Gittins +leg.; +BBSL BBSL +Faunal Survey No. +000 086 097 + +• + +5 ♀♀ +, +paratypes +; +Parma +, +Canyon County +; + +12 Jul. 1966 + +; +G.E. Bohart +leg.; +BBSL BBSL +Faunal Survey Nos +000 086 100 to 000 086 104 + +• + +1 ♀ +, +paratype +; same data as for preceding; + +12 Jul. 1966 + +; +G.E. Bohart +leg.; +BBSL +BBSL582464 +. – + +Montana + + +• + +2 ♀♀ +, +paratypes +; +ANSP +. – + +Nevada + + +• + +1 ♀ +, +paratype +; + +2 mi +N of Nixon + +, +Washoe County +; + +16 Jun. 1972 + +; +J.D. Pinto +leg.; +UCR +UCRC ENT 57154 + +• + +5 ♀♀ +, +paratypes +; + +4 mi +N of Nixon + +, +Washoe County +; + +7 Jun. 1996 + +; +F.D. Parker +leg.; +BBSL +BBSL581745 +, +BBSL581746 +, +BBSL581758 +, +BBSL581762 +, +BBSL581833 + +• + +1 ♀ +, +paratype +; +Beowawe +, +Eureka County +; + +30 Jul. 1969 + +; +R.M. Bohart +leg.; +UCBME + +• + +1 ♀ +, +paratype +; +Fallon +; + +18 Jun. 1930 + +; +E.L. Bell +leg.; +AMNH + +• + +1 ♀ +, +paratype +; +Nixon +, +Washoe County +; + +20 Jun. 1927 + +; +E.P. Van Duzee +leg.; +CAS + +• + +1 ♀ +, +paratype +; same data as for preceding; + +24 Jun. 1964 + +; +D.F. Veirs +leg.; +BBSL +BBSL522486 + +• + +1 ♀ +, +paratype +; same data as for preceding; + +24 Jun. 1964 + +; +J.E. Slansky +leg.; +UCBME +M.G.R. Database No. +1209 + +• + +1 ♀ +, +paratype +; same data as for preceding; + +24 Jun. 1964 + +; +R.M. Bohart +leg.; +UCBME + +• + +1 ♀ +, +paratype +; +Orovada +, +Humboldt County +; + +14 Jul. 1962 + +; +M.E. Irwin +leg.; +UCBME + +• + +2 ♀♀ +, +paratypes +; +Pyramid Lake +, +Washoe County +; + +25 Jun. 1970 + +; +B.L. Villegas +leg.; +UCBME + +• + +1 ♀ +, +paratype +; same data as for preceding; + +25 Jun. 1970 + +; +R.M. Bohart +leg.; +UCBME + +• + +1 ♀ +, +paratype +; +Sand Pass +, +Washoe County +; + +7 Aug. 1957 + +; +R.C. Bechtel +leg.; +FSCA + +• + +1 ♀ +, +paratype +; +Winnemucca +, +Humboldt County +; + +24 Jun. 1973 + +; +P.F. Torchio +leg.; +BBSL +BBSL522487 +. – + +New Mexico + + +• + +2 ♀♀ +, +paratypes +; +Koehler +; 1 +Aug. +??14; +W.R. Walton +leg.; +USNM +. – + +Oregon + + +• + +1 ♀ +, +paratype +; + +5 mi +NW of Adrian + +, +Malheur County +; + +25 Jun. 1958 + +; +R.K. Eppley +leg.; +BBSL BBSL +Faunal Survey No. +000 086 096 + +• + +1 ♀ +, +paratype +; + +6 mi +S of Adrian + +, +Malheur County +; + +28 Jul. 1957 + +; +R.K. Eppley +leg.; +BBSL BBSL +Faunal Survey No. +000 086 095 + +• + +1 ♀ +, +paratype +; +Klamath Avenue +( +SW +of +Nyssa +), +Malheur County +; + +6 Sep. 1957 + +; +R.K. Eppley +leg.; +BBSL BBSL +Faunal Survey No. +000 086 094 + +• + +7 ♀♀ +, +paratypes +; +Ontario +; + +2 Aug. 1929 + +; +H.A. Scullen +leg.; +BBSL BBSL +Faunal Survey Nos +000 086 087 to 000 086 093. – + +South Dakota + + +• + +1 ♀ +, +paratype +; +Cave Hills +; + +22 Jul. 1928 + +; +H.C. Severin +leg.; +USNM +M.G.R. Database No. +1374. – + +Utah + + +• + +3 ♀♀ +, +paratypes +; + +12 mi +NW of Fillmore + +, +Millard County +; + +20 Jun. 1972 + +; +F. Parker +and +D. Vincent +leg.; +BBSL +BBSL522489 +to +BBSL522491 + +• + +1 ♀ +, +paratype +; +3 mi +W of +Hatton +; + +3 Jun. 1959 + +; +BBSL +BBSL522484 + +• + +1 ♀ +, +paratype +; +Gandy +; + +Apr. 1949 + +; +G.E. Bohart +leg.; +BBSL +BBSL522485 + +• + +3 ♀♀ +, +paratypes +; +Hwy +491, +San Juan +County +; +37.8350° N +, +109.1110° W +; + +27 Jul. 2007 + +; +J. Gibbs +and +C.S. +Sheffield +leg.; +PCYU +PCYU-GS-07:2171, PCYU-GS-07:2173, PCYU-GS-07:2181 + +• + +1 ♀ +, +paratype +; +Sinbad Country +( +E of Block Mountain +), +Emery County +; + +24 Jul. 1981 + +; +Parker +, +Veirs +, and +Griswold +leg.; +BBSL +BBSL224656 +. – + +Wyoming + + +• + +2 ♀♀ +, +paratypes +; +Grand Teton National Park +; +Jul. +??37; +BBSL BBSL +Faunal Survey Nos +000 086 085, 000 086 086 + +• + +1 ♀ +, +paratype +; +Sheridan Local Training Area +, +Sheridan County +; + +10 Jul. 2000 + +; +B.C. Kondratieff +, +P.M. Pineda +, and +H. Al-Dhafer +leg.; +CSUC + +. + + +DNA barcoded material + + +Available. BIN: BOLD:ADH5866. See +type +material for specimens examined and sequenced (indicated by unique BOLD sample +ID +). + + + + + +Description + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length +9.8 mm +; ITW +2.3 mm +; head length +2.6 mm +; head width +3.6 mm +; fore wing length +8.2 mm +(margins of both worn). + + +Female + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with apical third golden yellow (entirely dark brown/black in some +paratypes +). Mandible with basal two-thirds; labrum along apical and lateral margins; scape, pedicel, and F1 extensively; succeeding flagellomeres to some extent; tegula; coxae to some extent; trochanters to tarsi (excluding brown meso- and metatibial spurs) entirely; metasomal terga laterally; and pseudopygidial area with underlying integument, pygidial plate, and metasomal sterna to some extent reddish orange. Fore wing membrane subhyaline, apically dusky. Hind wing membrane dusky subhyaline to hyaline. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Clypeus, upper paraocular and frontal areas, and vertexal area mostly exposed. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with well-defined paramedian band of pale-yellow tomentum, tapering slightly toward but not attaining anterior margin; pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with off-white to pale-yellow, appressed, branched setae; upper half densely setose, except behind pronotal lobe, with setae slightly sparser on hypoepimeral area; ventrolateral half sparsely setose. Mesopleuron with sparse, pale-yellow, erect/suberect, simple setae (less than ½ MOD in length) in addition to usual appressed, branched setae. Metanotum with tomentum uninterrupted, uniformly pale yellow. Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. Metasomal terga with bands of pale-yellow tomentum. T1 with apical transverse band complete (narrowly interrupted medially in some +paratypes +), transverse bands subparallel, discal patch transversely oblong or trapezoidal with short, narrowed (i.e., inverted V-shaped) anteromedial projection. T2–T4 with complete apical transverse bands, those of T2–T3 medially somewhat removed from apical margins of terga, that of T2 with pair of basomedially convergent anterolateral extensions. T5 with large patch of pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with triangular region of posteriorly directed setae with three subregions (basal patch of dense, golden setae; darker subapical band of sparser, coppery brown setae; and apical row of dense, suberect, silvery setae) within larger trapezoidal space of posteromedially directed coppery brown setae. S2–S3 with apical transverse bands of white tomentum (ill-defined and/or reduced to posterolateral patches in +holotype +and some +paratypes +). S4 with apical transverse band of white tomentum. S5 with apical fimbria of coppery bristle-like setae. + + +SURFACE +SCULPTURE +. Labrum coarsely and densely (most i <1d) rugose-punctate. Clypeus densely punctate (most i≤ 1d) but interspaces well defined, shining; with many small punctures among larger ones. Vertexal area somewhat sparsely punctate (some i>2d), especially around ocelli. Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense (most i ≤ 1d); interspaces well defined, shining. Mesopleuron with punctures in upper half nearly contiguous and denser (i<1d) than in ventrolateral half (i≤ 2d); interspaces shining where punctures not contiguous; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i≈ 1d), and evenly distributed; interspaces shining somewhat. + + +STRUCTURE +. Labral apex with pair of small denticles, each preceded by longitudinal carina. Pronotal collar short (medial length ~ ½ MOD). Mesoscutellum weakly bigibbous. Axilla extending little if at all beyond midlength of mesoscutellum; tip distinctly pointed, but mesally unattached to mesoscutellum for less than +2/5 +medial length of axilla; lateral margin relatively straight. Fore wing with three submarginal cells. T5 with concave apical margin. Pygidial plate apically truncate. S5 straight in lateral view. + + +Male + +Unknown. + + + + +Distribution + + + +Western +United States +and adjacent +Canada +(Fig. 12O). + + + + + +Ecology + + + +Host records + +Unknown. + + +Fig. 30. + +Triepeolus shoshone + +sp. nov. +A–B +. Holotype, ♀ (BOLD sample ID: CMNTO_063; PCYU PCYU-GS-07:2170). +A +. Habitus, lateral view. +B +. Habitus, dorsal view. +C +. Paratype, ♀ (RAM PMAE 16026), habitus, lateral view. +D +. Paratype, ♀ (PCYU PCYU-GS-07:2173) pseudopygidial area, dorsal view. + + + +Floral records + + +Labels of examined voucher specimens indicate that this species has been collected from the following flowering plant species: in +Asteraceae +, + +Centromadia pungens +Greene + +and + +Tetradymia canescens +DC. + +; in +Cleomaceae +, + +Cleomella +sp. + +; in +Fabaceae +, + +Melilotus albus + +; in +Grossulariaceae DC. +, + +Ribes +sp. + +; and in +Tamaricaceae +, + +Tamarix +sp. + +or spp. and + +T. gallica + +L. + + + + + +Remarks + + + + +Triepeolus shoshone + +sp. nov. +is very similar to + +T. apache + +sp. nov. +, and the two cannot be consistently separated morphologically (see Remarks under + +T. apache + +for information pertaining to their treatment as separate species). + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFB3FFE233A5F97AFC48FE54.xml b/data/03/8F/53/038F531FFFB3FFE233A5F97AFC48FE54.xml new file mode 100644 index 00000000000..e76afbac4ce --- /dev/null +++ b/data/03/8F/53/038F531FFFB3FFE233A5F97AFC48FE54.xml @@ -0,0 +1,679 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus saturninus +Cockerell & Sandhouse, 1924 + + + +Figs 12M, 18C, 28 + + + + + + +Triepeolus saturninus +Cockerell & Sandhouse, 1924: 312 + + +( +♂ +). + + + + +Proposed common name + + + + +Saturnine triepeolus. + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. saturninus + +apart from all other + +Triepeolus + +in the + +simplex + +species group: the mesopleura have sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae ( +Figs 18C +, +28A, C +); the legs are extensively dark brown/black from coxae to femora ( +Figs 18C +, +28A–C +); and the T1 discal patch is trapezoidal ( +Fig. 28B +). + +Triepeolus saturninus + +most closely resembles + +T. hirsutus + +sp. nov. +and + +T. parahirsutus + +sp. nov. +, and in all three species the mesopleura have sparse, pale-yellow, erect/ suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae; the T1 basal band is rather abruptly interrupted medially; and the T1 discal patch is quadrangular, but in + +T. hirsutus + +and + +T. parahirsutus + +the legs are entirely reddish orange from trochanters to tarsi and the underlying integument of the pseudopygidial area of the female ranges from entirely dark brown/black to predominantly reddish orange whereas in + +T. saturninus + +it is entirely dark brown/black ( +Fig. 28D +). Additionally, + +T. saturninus + +may be confused with + +T. segregatus + +or + +T. shoshone + +sp. nov. +as the ranges of the three species overlap to some extent, but in the latter two species the legs are predominantly to entirely reddish orange from trochanters to tarsi, in + +T. segregatus + +the mesopleura laterally do not have erect/suberect, simple setae or have only sparse, short (<¼ MOD), erect/suberect, simple setae in addition to the usual appressed, branched setae, and in + +T. shoshone + +the T1 basal band forms an inflexed arch and is thus gradually narrowed but not completely interrupted medially, such that the discal patch has a short, narrowed (i.e., inverted V-shaped) anteromedial projection. + + + + + +Etymology + + + +The specific epithet was inspired by the shape of the area of black tomentum on T1 (discal patch), which in the original description was said to suggest “the planet Saturn with its rings, only the rings much broader” ( +Cockerell & Sandhouse 1924 +). + + + + + +Material examined + + + + +Primary +type +material + + + + +USA +• +♂ +, +holotype +; +California +, +Millbrae +, +San Mateo County +; + +1 Sep. 1912 + +; +E.C. Van Dyke +leg.; +CAS 1605 + +. + + +DNA barcoded material + +Unavailable. + +Non-barcoded material + + + +USA +– + +California + +• +1 ♀ +; +Etna +, +Siskiyou County +; + +6 Aug. 1978 + +; +B. Villegas +leg.; +UCBME + +• + +1 ♀ +; +Lower Klamath Lake +, +Siskiyou County +; + +14 Aug. 1963 + +; +J. Schuh +leg.; +FSCA + +• + +1 ♀ +; same data as for preceding; + +17 Aug. 1963 + +; +J. Schuh +leg.; +FSCA + +• + +1 ♀ +; +Santa Cruz Island +(wash N of research station), +Santa Barbara County +; + +22 Oct. 1990 + +; +R.W. Thorp +leg.; +UCBME + +• + +1 ♀ +; +Sunol Regional Wilderness +( + +15 mi +E of Fremont + +), +Alameda County +; + +24 May 1987 + +; +S. Geohegan +leg.; +USNM + +. – + + +Idaho + +• +1 ♀ +; + +3 mi +NW of Carey + +, +Blaine County +; + +8 Aug. 1967 + +; +D.S. Horning +, +Jr. +leg.; +UCBME + +. – + + +Oregon + +• +1 ♀ +; + +6 mi +W of Bly + +, +Klamath County +; + +18 Aug. 1955 + +; +W.P. Stephen +leg.; +OSAC + +• + +1 ♀ +; ca + +5 mi +SE of Culver + +, +Jefferson County +; + +19 Aug. 1984 + +; +Schauff +and +Grissell +leg.; +USNM + +• + +1 ♀ +; +Dairy +, +Klamath County +; + +4 Sep. 1962 + +; +J. Schuh +leg.; +FSCA + +• + +1 ♀ +; +Modoc Point +, +Klamath County +; + +14 Aug. 1963 + +; +J. Schuh +leg.; +FSCA + +• + +1 ♀ +; S of +Chiloquin +, +Klamath County +; + +13 Aug. 1973 + +; +H.V. Weems +, +Jr. +leg.; +FSCA + +. – + + +Utah + +• +1 ♀ +; +High Creek +, +Cache County +; + +15 Aug. 1974 + +; +Knowlton +and +Cazier +leg.; +BBSL BBSL +Faunal Survey No. +000 077 713 + +. – + + +Washington + +• +1 ♀ +; +Yakima River +Canyon ( +Hwy +281); + +14 Sep. 1990 + +; +E. Miliczky +leg.; +ERM + +. + + +Non-preserved material + + + +USA +• +1 ♀ +; +Washington +, +Richland +, +Benton County +; +46.3097° N +, +119.2622° W +; + +1 Sep. 2023 + +; “janeabel” obs.; iNaturalist record #185099704 + +. + + + + + +Redescription + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length +9.3 mm +; ITW +1.9 mm +; head length +2.3 mm +; head width +3.1 mm +; fore wing length +7.5 mm +. + + +Both sexes + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with apical half golden yellow and middle quarter in basal half reddish orange. Antenna (except for orange spot basally on F2) dark brown in +holotype +; scape, pedicel, and F1 extensively orange in some non-type specimens. Tegula, coxae to femora to some extent, and tibiae (excluding metatibial spurs) and tarsi entirely orange. Fore wing membrane dusky subhyaline throughout. Hind wing membrane dusky subhyaline to hyaline. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Tomentum slightly sparser on clypeus; upper paraocular and frontal areas and vertexal area mostly exposed. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with well-defined paramedian band of pale-yellow tomentum; pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with off-white to pale-yellow, appressed, branched setae; densely setose except for two sparsely setose circular patches (one beneath base of fore wing (hypoepimeral area) and slightly larger one in ventrolateral half of mesopleuron). Mesopleuron with sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to usual appressed, branched setae. Metanotum with tomentum uninterrupted, uniformly pale yellow. Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. Metasomal terga with bands of pale-yellow tomentum. T1 with basal and apical transverse bands interrupted medially (former more widely than latter) and subparallel, discal patch trapezoidal. T2–T4 with complete apical transverse bands, that of T2 somewhat narrowed medially and with pair of basomedially convergent anterolateral extensions. S2–S3 with apical transverse bands of white tomentum. + + +SURFACE +SCULPTURE +. Labrum coarsely rugose-punctate, with larger punctures than clypeus, but punctures of both equally dense (most i <1d). Clypeus with glabrous midline incomplete and very short, extending from upper margin down to <⅓ length of clypeus. Vertexal area densely punctate (most i ≤ 1d). Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense (most i≤ 1d); interspaces well defined, shining. Mesopleuron with punctures in upper half nearly contiguous but not much denser (i<1d) than in ventrolateral half (most i ≤ 1d); interspaces shining where punctures not contiguous; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i ≈1d), and evenly distributed; interspaces shining somewhat. + + +STRUCTURE +. Labrum with pair of small subapical denticles, each preceded by discrete longitudinal ridge. Pronotal collar somewhat elongate (medial length ~ +4/5 +MOD). Mesoscutellum weakly bigibbous. Axilla extending little if at all beyond midlength of mesoscutellum; tip distinctly pointed, but mesally unattached to mesoscutellum for less than +2/5 +medial length of axilla; lateral margin relatively straight. Fore wing with three submarginal cells. + + +Female + +Paramedian band tapering slightly toward but not attaining anterior margin of mesoscutum. Mesopleuron with black, appressed, branched setae in sparsely setose circular patch in ventrolateral half of mesopleuron. T5 with concave apical margin and large patch of pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with triangular region of posteriorly directed coppery to silvery setae within larger trapezoidal space of posteriorly directed dark brown setae; apical margin with row of dense, suberect, silvery setae. Pygidial plate reddish brown in part and apically truncate. S4 with apical transverse band of white tomentum. S5 straight in lateral view, with apical fimbria of coppery bristle-like setae. + +Male + +Paramedian band anteriorly joined to lateral transverse band of equally dense pale-yellow tomentum along anterior margin of mesoscutum. T5–T6 with complete apical transverse bands. Pygidial plate with small orange spot subapically, relatively flat and apically rounded. S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae, those of S4 coppery to silvery, those of S5 coppery and contrasting with apical transverse bands of white tomentum of preceding sterna. + + + + +Distribution + + + +United States +west of the Rocky Mountains (Fig. 12M). + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + + +Labels of examined voucher specimens indicate that this species has been collected from the following flowering plant species: in +Asteraceae +, + +Chrysothamnus +sp. + +or spp., + +Grindelia +sp. + +, + +Machaeranthera canescens +(Pursh) A.Gray + +, and + +Senecio flaccidus +var. +douglasii +(DC.) B.L.Turner & T.M.Barkley + +; and in +Fabaceae +, + +Melilotus +sp. + + + + + +Fig. 28. + +Triepeolus saturninus +Cockerell & Sandhouse, 1924 + +. +A +, +D +. Female (USNM). +A +. Habitus, lateral view. +B +. Female (USNM), habitus, dorsal view. +C +. Holotype, ♂ (CAS 1605), habitus, lateral view. +D +. Pseudopygidial area, dorsal view. + + + + + +Remarks + + + +In the +holotype +of this species, the discal patch has a Saturn-like shape and thus resembles that of + +T. joliae +Rightmyer, 2008 + +and, to a lesser extent, some specimens of + +T. californicus +( +Cresson, 1878 +) + +. The description of the female of + +T. saturninus + +is published here for the first time. Based on known records, adults of + +T. saturninus + +are most active from August to September. + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFB6FFEE302DFAF6FD4AF981.xml b/data/03/8F/53/038F531FFFB6FFEE302DFAF6FD4AF981.xml new file mode 100644 index 00000000000..45f62eaad7b --- /dev/null +++ b/data/03/8F/53/038F531FFFB6FFEE302DFAF6FD4AF981.xml @@ -0,0 +1,525 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus roni +Genaro, 1999 + + + + + + +Figs 5B +, 11A, 12L, 27 + + + + + + + +Triepeolus roni +Genaro, 1999: 219 + + +( +♀ +, +♂ +), figs 1c, 2b, 3c. + + + + +Proposed common name + + + + +Ron’s triepeolus. + + + + +Diagnosis + + + +Unique to + +T. roni + +within the genus are the basal arched ridge overhanging the impressed apicomedial region of the pseudopygidial area of the female, which (as in all species in the + +T. simplex + +group) is triangular, with the apical margin of T5 concave, and glabrous midline extending from it to the apical margin of the tergum, which is suggestive of the cleft pseudopygidial area of + +Rhogepeolus +Moure, 1955 + +( +Hymenoptera +: +Apidae +: +Nomadinae +) ( +Fig. 27D +). Otherwise, the following morphological features in combination tell both sexes of + +T. roni + +apart from all other species of + +Triepeolus + +: the clypeus does not have a glabrous midline (Supp. file 2: +Fig. S19C +); the axillae are large, each with the tip extending well beyond the midlength of the mesoscutellum and the free portion distinctly pointed, hooked (i.e., concave along the medial margin), and mesally unattached to the mesoscutellum for approximately half the medial length of the axilla ( +Fig. 5B +); the T1 discal patch is trapezoidal with a semicircular anteromedial projection (Figs 11A, 27B); and the T1 apical transverse band is conspicuously narrowed sublaterally, at the posterior corners of the discal patch (Figs 11A, 27B). Males of + +T. roni + +are very similar in overall appearance to those of + +T. nisibonensis +Genaro, 2001 + +, a Hispaniolan (as opposed to Bahamian and Cuban) species (outside of the + +simplex + +species group), but both sexes of + +T. roni + +can easily be separated from + +T. nisibonensis + +on the basis of the vertexal area, which in the latter is uniquely notched posteromedially, as well as by geography. + + + + + +Etymology + + + +The specific epithet (declined in the genitive case) honors the late Dr Ronald ( +Ron +) J. McGinley, a friend and colleague of the taxonomic authority, in recognition of his contributions to the knowledge of bee biodiversity, conservation, and collections management ( +Genaro 1999 +). + + + + + +Material examined + + + + +Primary +type +material + + + +CUBA +• + +♀ +, +holotype +(studied from images); +Mayabeque +, + +La Fosforita + +, +Loma de Candela +, +Güines +; + +28 Dec. 1992 + +; +J.A. Genaro +leg.; +MNHNCU + +. + + + +Secondary +type +material + + + +CUBA +– + +Mayabeque + +• + +1 ♀ +, +paratype +; same data as for holotype; +KUNHM +M.G.R. Database No. +3713 + +• + +1 ♀ +, +paratype +(studied from image); same data as for holotype; +MNHNCU +. – + +Sancti Spíritus + + +• + +1 ♀ +, +paratype +(studied from image); +Punta Caguanes +, +Yaguajay +; + +16 Jul. 1996 + +; +J.A. Genaro +leg.; +MNHNCU +. – + + +Santiago de +Cuba + + + +• + +1 ♂ +, +allotype +(studied from image), +1 ♀ +, +paratype +(studied from image); +Cuabitas +; + +Oct. 1949 + +; +P. Alayo +leg.; +MNHNCU + +. + + +DNA barcoded material + +Available. BIN: BOLD:AFI6376. Specimens examined and sequenced: + +CUBA +• + +1 ♂ +(studied from image); +Pinar del Río +, +San Diego +de los +Baños +, +Los Palacios +; + +16 Sep. 2022 + +; +D. Breto +leg.; BOLD sample ID: CCDB-14520-D12; +MHNTSN + +. + + +Non-barcoded material + + + +BAHAMAS +– + +Central Abaco + +• +1 ♀ +; +Marsh Harbour +( +Great +Abaco +Island +); + +30 Apr. 1993 + +; +T.W. Schoener +leg.; +UCBME + +. – + + +Mangrove Cay + +• +1 ♀ +; +Mangrove Cay +( +Andros +Island +); + +26 Apr. 1953 + +; +E.B. Hayden +leg.; +AMNH +M.G.R. Database No. +1307. – + +San Salvador + + +• + +1 ♂ +; +San Salvador Island +; + +6 Jun. 1978 + +; +P. Salbert +leg.; +USNM +M.G.R. Database No. +5922 + +. + + + + + +Redescription + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length +8.8 mm +; ITW +2.1 mm +; head length +2.2 mm +; head width +3.1 mm +; fore wing length +6.9 mm +. + + +Both sexes + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with apical third golden yellow. Mandible with basal two-thirds; labrum along apical and lateral margins; scape, pedicel, and F1 predominantly and succeeding flagellomeres to some extent; tegula; coxae to some extent; trochanters entirely; femora predominantly to entirely; and tibiae (excluding brown meso- and metatibial spurs) and tarsi entirely orange. Fore wing membrane dusky subhyaline throughout, slightly darker beyond venation. Hind wing membrane dusky subhyaline to hyaline. Metasomal terga laterally, pygidial plate to some extent, and metasomal sterna to some extent reddish brown. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Clypeus, upper paraocular and frontal areas, and vertexal area mostly exposed. Pronotal collar with tomentum uniformly bright yellow (off-white in some non-type specimens). Mesoscutum with well-defined paramedian band of off-white to yellow tomentum (attaining anterior margin in +holotype +, +allotype +, and +paratypes +; tapering slightly toward but not attaining anterior margin in some non-type specimens); pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with dark brown/gray, appressed, branched setae ventromedially and off-white to pale-yellow, appressed, branched setae elsewhere; densely setose except for two sparsely setose circular patches (one beneath base of fore wing (hypoepimeral area) and larger one occupying much of ventrolateral half of mesopleuron). Metanotum with tomentum uninterrupted, uniformly pale yellow (off-white in some non-type specimens). Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. Metasomal terga with bands of white/ off-white to yellow tomentum. T1 with basal and apical transverse bands subparallel and discal patch trapezoidal with semicircular anteromedial projection. T1–T4 with apical transverse bands medially somewhat removed from apical margins of terga, those of T1–T2 separated into pairs of rounded lobes medially, that of T1 narrowed sublaterally mesad lateral longitudinal band (at posterior corner of discal patch), that of T2 with pair of basomedially convergent anterolateral extensions, those of T3–T4 complete or tapering until separated medially. S2–S3 with apical transverse bands of white tomentum. + + +SURFACE +SCULPTURE +. Labrum coarsely and densely (most i <1d) rugose-punctate. Clypeus densely punctate (most i ≤ 1d) but interspaces well defined, shining; with many small punctures among larger ones. Vertexal area somewhat sparsely punctate (some i> 2d) anteriorly and around ocelli, otherwise densely (most i <1d) rugose-punctate. Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense and nearly contiguous (most i<1d). Mesopleuron with punctures in upper half nearly contiguous but not much denser (i<1d) than in ventrolateral half (most i ≤ 1d); interspaces shining where punctures not contiguous; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i≈ 1d), and evenly distributed; interspaces shining somewhat. + + + +Fig. 27. + +Triepeolus roni +Genaro, 1999 + +. +A +, +D +. Paratype, ♀ (KUNHM M.G.R. Database No. 3713). +A +. Habitus, lateral view. +B +. Female (AMNH M.G.R. Database No. 1307), habitus, dorsal view. +C +. Male (USNM M.G.R. Database No. 5922), habitus, lateral view. +D +. Pseudopygidial area, dorsal view. + + + +STRUCTURE +. Labrum with pair of small subapical denticles, each preceded by discrete longitudinal ridge. Pronotal collar short (medial length ~ ½ MOD). Mesoscutellum strongly bigibbous.Axilla extending well beyond midlength of mesoscutellum but not as far back as its posterior margin; tip distinctly pointed and hooked (i.e., concave along medial margin), mesally unattached to mesoscutellum for approximately half medial length of axilla; lateral margin somewhat arcuate. Fore wing with three submarginal cells. + + +Female + +T5 with concave apical margin and large patch of pale-yellow to white/off-white tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with underlying integument reddish brown in part, forming rounded triangle with concave sides, with brown erect/suberect setae basally and brown spinelike setae laterally, with basal arched ridge overhanging impressed apicomedial region with ill-defined glabrous midline. Pygidial plate apically truncate. S4 with apical transverse band of white tomentum. S5 straight in lateral view, with apical fimbria of brown bristle-like setae and dark brown tomentum laterally and apicomedially; S5 otherwise covered in off-white tomentum (S5 uniformly covered in brown tomentum in some non-type specimens). + +Male + +T5 with complete apical transverse band of bright to pale-yellow tomentum. T6 with small posteromedial patch of pale-yellow tomentum. Pygidial plate apically rounded and slightly downturned, with lateral margin somewhat sinuate. S4–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved, predominantly coppery brown setae, contrasting with apical transverse bands of white tomentum of preceding sterna, those of S4 coppery to silvery laterally. + + + + +Distribution + + + +The Bahamas +and +Cuba +(Fig. 12L). This is the only species in the + +T. simplex + +group known to occur in the West Indies. + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + +Unknown. + + + + +Remarks + + + +In some specimens from +the Bahamas +, the dorsum of the mesosoma and metasoma have bands of white/ off-white (as opposed to yellow) tomentum. + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFC9FF96300DFD47FAEDF9DD.xml b/data/03/8F/53/038F531FFFC9FF96300DFD47FAEDF9DD.xml new file mode 100644 index 00000000000..cf482cd0891 --- /dev/null +++ b/data/03/8F/53/038F531FFFC9FF96300DFD47FAEDF9DD.xml @@ -0,0 +1,302 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus paucipunctatus + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +9B1A5AAE-DA84-497C-8467-623DD1F80D2F + + + + +Figs 12J, 18B, 25 + + +Proposed common name + + + +Oaxaca +triepeolus. + + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. paucipunctatus + +sp. nov. +apart from all other + +Triepeolus + +in the + +simplex + +species group: the T1 discal patch is crescent-shaped ( +Fig. 25B +) and the ventrolateral halves of the mesopleura are sparsely punctate (some i>4d) and have sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae ( +Figs 18B +, +25A +). + +Triepeolus paucipunctatus + +most closely resembles specimens of + +T. alvarengai + +that have an apical transverse band on T1, + +T. eumeniformis + +sp. nov. +, and + +T. segregatus + +, but in the latter three species the mesopleura are more densely punctate (i≤ 3d) and laterally do not have erect/suberect, simple setae or have only sparse, short (<¼ MOD), erect/suberect, simple setae in addition to the usual appressed, branched setae, and in + +T. alvarengai + +and + +T. eumeniformis + +the T2 apical transverse band does not have well-defined basomedially convergent anterolateral extensions. + + + + + +Etymology + + +The specific epithet is a compound adjective that combines the Latin ‘pauci’ (meaning ‘few’) and ‘punctatus’ (meaning ‘punctate’) and refers to the sparsely punctate mesopleura. + + + + +Material examined + + + + +Primary +type +material + + + + +MEXICO +• +♀ +, +holotype +; +Oaxaca +, +3 mi +W of +El Camarón +; + +5 Aug. 1963 + +; +L.A. Stange +leg.; +UCBME +M.G.R. Database No. +1216 + +. + + +DNA barcoded material + +Unavailable. + + + + +Description + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length 8.0 mm; ITW +1.6 mm +; head length 2.0 mm; head width +2.9 mm +; fore wing length +6.4 mm +. + + +Female + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with middle quarter in basal half reddish brown. Scape to some extent, pedicel and F1 extensively, tegula, coxae to femora to some extent, and tibiae (excluding metatibial spurs) and tarsi extensively orange. F2 with orange spot basally. Fore wing membrane dusky subhyaline throughout. Hind wing membrane dusky subhyaline to hyaline. Pseudopygidial area with apical margin and pygidial plate reddish brown. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Tomentum slightly sparser on clypeus; upper paraocular and frontal areas and vertexal area mostly exposed. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with well-defined paramedian band of pale-yellow tomentum, tapering slightly toward but not attaining anterior margin; pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with pale-yellow, appressed, branched setae; upper half densely setose, with setae slightly sparser on hypoepimeral area; ventrolateral half nearly bare, except along margins. Mesopleuron with sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to usual appressed, branched setae. Metanotum with tomentum sparser medially, uniformly off-white. Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. Metasomal terga with bands of pale-yellow tomentum. T1 with basal transverse band narrowly interrupted medially, arched, and continuous with (and indistinguishable from) lateral longitudinal band; apical transverse band broadened submedially and separated into pair of rounded lobes medially; discal patch crescent-shaped. T2–T4 with complete apical transverse bands, those of T2–T3 medially somewhat narrowed and removed from apical margins of terga, that of T2 with pair of basomedially convergent anterolateral extensions. T5 with large patch of pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with triangular region of posteriorly directed coppery setae within larger trapezoidal space of posteromedially directed silvery setae; apical margin with row of dense, suberect, silvery setae. S2–S4 with apical transverse bands of white tomentum. S5 with apical fimbria of coppery bristle-like setae. + + + +Fig. 25. + +Triepeolus paucipunctatus + +sp. nov. +, holotype, ♀ (UCBME M.G.R. Database No. 1216). +A +. Habitus, lateral view. +B +. Habitus, dorsal view. +C +. Head, frontal view. +D +. Pseudopygidial area, dorsal view. + + + +SURFACE +SCULPTURE +. Labrum with punctures nearly contiguous (most i <1d). Clypeus densely punctate (most i≤ 1d) but interspaces well defined, shining; with many small punctures among larger ones. Vertexal area somewhat sparsely punctate (some i>2d), especially around ocelli. Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense (most i≤ 1d); interspaces well defined, shining. Mesopleuron with punctures in upper half much denser (most i≤ 2d) than in ventrolateral half (some i>4d); interspaces shining; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i=1–2d), and evenly distributed; interspaces shining. + + +STRUCTURE +. Labral apex with pair of very small denticles, each preceded by discrete longitudinal ridge. Pronotal collar somewhat elongate (medial length ~¾ MOD). Mesoscutellum weakly bigibbous. Axilla extending little if at all beyond midlength of mesoscutellum; tip distinctly pointed and hooked (i.e., concave along medial margin), mesally unattached to mesoscutellum for approximately half medial length of axilla; lateral margin somewhat arcuate. Left fore wing with second submarginal crossvein incomplete and thus two submarginal cells, right fore wing with three submarginal cells. T5 with concave apical margin. Pygidial plate apically truncate. S5 straight in lateral view. + + +Male + +Unknown. + + + + +Distribution + + + +Presently known only from the +type +locality in +Oaxaca +, +Mexico +(Fig. 12J). + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + +Unknown. + + + + +Remarks + + + +This species is known and described only from the +holotype +, despite extensive search efforts for additional exemplars in existing entomological collections. Although it is undesirable to describe new species from limited material, the multiple unusual morphological features exhibited by the specimen clearly set it apart from other members of the + +T. simplex + +group and thus support its treatment as a separate species. + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFCCFF94301CFEE1FB0AFD67.xml b/data/03/8F/53/038F531FFFCCFF94301CFEE1FB0AFD67.xml new file mode 100644 index 00000000000..1e174f5ec34 --- /dev/null +++ b/data/03/8F/53/038F531FFFCCFF94301CFEE1FB0AFD67.xml @@ -0,0 +1,552 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus parahirsutus + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +A06B47B8-CDD7-4E83-9B51-D3C829AA9347 + + + + + +Figs 6B +, 12I, 24 + + + +Proposed common name + + + +Texas +bristly triepeolus. + + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. parahirsutus + +sp. nov. +apart from all other + +Triepeolus + +in the + +simplex + +species group except + +T. hirsutus + +sp. nov. +: the mesopleura have sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae ( +Figs 6B +, +24A, C +); the legs are entirely reddish orange from trochanters to tarsi ( +Fig. 24A–C +); and the T1 discal patch is trapezoidal with an inverted U- or horseshoe-shaped anteromedial projection ( +Fig. 24B +). Females of + +T. parahirsutus + +and + +T. hirsutus + +are readily distinguished from each other by the pseudopygidial area, which in the former does not have distinct subregions of differentiated setae but rather setae that are golden to silvery and relatively uniform in density ( +Fig. 24D +) and in the latter has three subregions of differentiated setae—a basal patch of dense, golden setae, a darker subapical band of sparser, coppery brown setae, and an apical row of dense, suberect, silvery setae. Additionally, + +T. parahirsutus + +is generally larger (body length 8.0–12.0 mm; ITW +1.7–2.5 mm +) ( +Fig. 24A–C +) than + +T. hirsutus + +(body length 5.5–9.0 mm; ITW +1.2–1.7 mm +) and in the former the axillae commonly (as opposed to almost never) have reddish tips ( +Fig. 24B +). Although it might not be possible to separate certain males morphologically, + +T. parahirsutus + +can easily be separated from + +T. hirsutus + +by geography; the former is known only from Central and South +Texas +whereas the latter is known only from the Southwestern +United States +and adjacent +Mexico +. Additionally, + +T. parahirsutus + +may be confused with + +T. segregatus + +due to their similar size and trapezoidal discal patch with an inverted U- or horseshoe-shaped anteromedial projection and as the ranges of both species overlap to some extent, but in + +T. segregatus + +the mesopleura laterally do not have erect/suberect, simple setae or have only sparse, short (<¼ MOD), erect/suberect, simple setae in addition to the usual appressed, branched setae. + + + + + +Etymology + + + +The specific epithet refers to this species’ similarity to + +T. hirsutus + +sp. nov. +The Greek prefix ‘ +para- +’ means ‘near’. + + + + + +Material examined + + + + +Primary +type +material + + + + +USA +• +♀ +, +holotype +; +Texas +, +McAllen Botanical Gardens +(McAllen), +Hidalgo County +; + +28 Jul. 1984 + +; +C. Porter +leg.; +FSCA + +. + + + +Secondary +type +material + + + + +USA +– + +Texas + +• +1 ♀ +, +paratype +; +Texas +; +Belfrage +leg.; +USNM + +• + +1 ♀ +, +paratype +; + +2 mi +S of El Sauz + +, +Starr County +; + +28 May 1980 + +; +Webb +and +Lisowski +leg.; +INHS 7838 + +• + +1 ♀ +, +paratype +; +Dallas +; +ZMB 21261 + +• + +1 ♂ +, +allotype +; same data as for preceding; +ZMB 21261 + +• + +1 ♀ +, +paratype +; +Fedor +, +Lee County +; + +18 Sep. 1897 + +; +Birkmann +leg.; +USNM + +• + +1 ♀ +, +paratype +; same data as for preceding; + +24 Sep. 1897 + +; +Birkmann +leg.; +ZMB + +• + +1 ♀ +, +paratype +; +Laguna Atascosa National Wildlife Refuge +( +Island Fields Loop +), +Cameron County +; +26.2231° N +, +97.3525° W +; + +20 Apr. 2017 + +; +M. Buck +leg.; BOLD sample ID: CMNTO_313; +RAM +pmae00140925 + +• + +1 ♀ +, +paratype +; +Laguna Madre +; + +10 Apr. 1945 + +; +D.E. Hardy +and +V.L. Wooley +leg.; +USNM + +• + +1 ♀ +, +paratype +; +McAllen Botanical Gardens +( +McAllen +), +Hidalgo County +; + +16–30 May 1974 + +; +C. Porter +leg.; +FSCA + +• + +1 ♀ +, +paratype +; same data as for preceding; + +1 Jul. 1983 + +; +C. Porter +leg.; +FSCA + +. + + +DNA barcoded material + + +Available. BIN: BOLD:AEM8829. See +type +material for specimens examined and sequenced (indicated by unique BOLD sample ID). + + + + + +Description + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length +11.6 mm +; ITW +2.5 mm +; head length +2.7 mm +; head width +3.9 mm +; fore wing length +8.6 mm +. + + +Both sexes + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with apical half golden yellow (entirely dark brown/black in some +paratypes +). Labrum entirely (partially to entirely dark brown/ black in some +paratypes +), clypeus along lower and lateral margins (entirely dark brown/black in some +paratypes +), scape, pedicel, F1 entirely and succeeding flagellomeres to some extent, pronotal lobe (entirely dark brown/black in some +paratypes +), tegula, axilla with tip (entirely dark brown/black in some +paratypes +), coxae to some extent, trochanters to tarsi (excluding brown meso- and metatibial spurs) entirely, metasomal terga laterally, pygidial plate to some extent (difficult to see in +holotype +because T6 mostly retracted; described from +paratypes +), and metasomal sterna to some extent reddish orange. Fore wing membrane subhyaline, apically dusky. Hind wing membrane dusky subhyaline to hyaline. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Clypeus, upper paraocular and frontal areas, and vertexal area mostly exposed. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with well-defined paramedian band of pale-yellow tomentum; pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with off-white to pale-yellow, appressed, branched setae; upper half densely setose, except behind pronotal lobe, with setae slightly sparser on hypoepimeral area; ventrolateral half sparsely setose. Mesopleuron with sparse, pale-yellow, erect/ suberect, simple setae (reaching about ½ MOD in length) in addition to usual appressed, branched setae. Metanotum with tomentum uninterrupted, uniformly pale yellow. Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. Metasomal terga with bands of pale-yellow tomentum. T1 with basal and apical transverse bands interrupted medially (former more widely than latter) and subparallel, discal patch trapezoidal. T2–T4 with complete apical transverse bands, those of T2–T3 medially somewhat narrowed and removed from apical margins of terga, that of T2 with pair of basomedially convergent anterolateral extensions. S2–S3 with apical transverse bands of white tomentum. + + +SURFACE +SCULPTURE +. Labrum with punctures nearly contiguous (most i <1d). Clypeus densely punctate (most i≤ 1d) but interspaces well defined, shining; with many small punctures among larger ones. Vertexal area somewhat sparsely punctate (some i>2d), especially around ocelli. Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense (most i≤ 1d); interspaces well defined, shining. Mesopleuron with punctures in upper half denser (i ≤1d) than in ventrolateral half (i ≤3d); interspaces somewhat dull due to tessellate surface microsculpture; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i ≈1d), and evenly distributed; interspaces shining somewhat. + + +STRUCTURE +. Labrum with pair of small subapical denticles, each preceded by discrete longitudinal ridge. Pronotal collar somewhat elongate (medial length ~ ¾ MOD). Mesoscutellum moderately bigibbous. Axilla extending little if at all beyond midlength of mesoscutellum; tip distinctly pointed, but mesally unattached to mesoscutellum for less than +2/5 +medial length of axilla; lateral margin relatively straight. Fore wing with three submarginal cells. + + +Female + +Paramedian band tapering slightly toward but not attaining anterior margin of mesoscutum. T5 with concave apical margin and large patch of pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with underlying integument to some extent reddish orange, with triangular region of posteriorly directed golden to silvery setae (relatively uniform in density) within larger trapezoidal space of posteromedially directed dark brown setae. Pygidial plate apically truncate. S4 with apical transverse band of white tomentum. S5 straight in lateral view, with apical fimbria of coppery bristle-like setae. + +Male + +Paramedian band anteriorly joined to lateral transverse band of equally dense pale-yellow tomentum along anterior margin of mesoscutum. T5–T6 with complete apical transverse bands. Pygidial plate relatively flat and apically rounded, with lateral margin somewhat sinuate. S4–S5 each with apical/ subapical fringe of dense, long (> 1 MOD), curved setae, those of S4 coppery to silvery, those of S5 coppery and contrasting with apical transverse bands of white tomentum of preceding sterna. + + + + +Distribution + + + +Central and South +Texas +(Fig. 12I). + + + + +Fig. 24. + +Triepeolus parahirsutus + +sp. nov. +A–B +. Holotype, ♀ (FSCA). +A +. Habitus, lateral view. +B +. Habitus, dorsal view. +C +. Allotype, ♂ (ZMB 21261), habitus, lateral view. +D +. Paratype, ♀ (USNM) pseudopygidial area, dorsal view. + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + +Unknown. + + + + +Remarks + + + +In addition to the diagnostic morphological features that separate + +T. parahirsutus + +sp. nov. +from similar species, its status as a separate species is supported by a separate BIN and large barcode sequence divergence (5.6%) from its nearest neighbor, + +T. hirsutus + +sp. nov. +(see Supp. file 4). + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFDBFF873069FE79FAE4FCC6.xml b/data/03/8F/53/038F531FFFDBFF873069FE79FAE4FCC6.xml new file mode 100644 index 00000000000..e8c628a97f3 --- /dev/null +++ b/data/03/8F/53/038F531FFFDBFF873069FE79FAE4FCC6.xml @@ -0,0 +1,342 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus nemoralis +( +Holmberg, 1886 +) + + + + + + +Fig. 13B + + + + + + + +Doeringiella nemoralis +Holmberg, 1886: 280 + + +( +♀ +). + + + + + + +Epeolus merus +Brèthes, 1909: 250 + + +( +♂ +). + + + + + +Epeolus merus + +– + +Schrottky 1913: 265 + +(in regional checklist). + + + + + +Triepeolus nemoralis + +– + +Roig-Alsina 1989: 578 + +. — + + +Onuferko +et al +. 2024: 27 + + +, figs 1f, 3b, 11, 13b (redescription). + + + + +Proposed common name + + + + +Nemoral triepeolus. + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. nemoralis + +apart from all other species of + +Triepeolus + +: the mesoscutum has a large anteromedial ovate patch of yellow tomentum, which may be sparser medially such that the patch is suggestive of ill-defined paramedian bands ( +Fig. 13B +); the T1 basal band is arched, continuous with (and indistinguishable from) the lateral longitudinal bands, and mesally concave on each side, such that the discal patch is semicircular ( +Fig. 13B +); T1 does not have an apical transverse band ( +Fig. 13B +), but the apical impressed area may have ill-defined patches of pale tomentum; and T2–T4 have complete apical transverse bands of yellow tomentum ( +Fig. 13B +). + +Triepeolus nemoralis + +most closely resembles specimens of + +T. alvarengai + +without an apical transverse band on T1 and + +T. mexicanus + +, the latter of which is known only from North and Central America, but in + +T. alvarengai + +the mesoscutum has well-defined paramedian bands and in + +T. mexicanus + +pale tomentum on the mesoscutum is restricted to the lateral and posterior margins. Males of + +T. nemoralis + +also closely resemble those of + +T. flavipennis + +without an apical transverse band on T1 and + +T. cameroni + +, which is known only from +Mexico +and Central America, but in the latter two species, which are not in the + +simplex + +species group, the mesoscutum has well-defined paramedian bands, in + +T. flavipennis + +the pair of anterolateral patches of pale tomentum on T1 are mesally convex, such that the discal patch forms a trapezoid or triangle with concave anterolateral sides, and in + +T. cameroni + +S3 has an apical/subapical fringe of dense, long (> 1 MOD), curved, pale setae (as opposed to only short, straight setae). This species is also very similar in overall appearance to + +Epeolus luteipennis + +, and in both species the mesoscutum has a large anteromedial patch of yellow tomentum (as opposed to well-defined paramedian bands), but both sexes of + +T. nemoralis + +can easily be told apart from any similar-looking + +Epeolus + +by their simple mandibles; in + +E. luteipennis + +and most other + +Epeolus +spp. + +, the mandibles each have a distinct preapical tooth. + + + + + +Etymology + + + +See + +Onuferko +et al +. (2024) + +. + + + + + +Material examined + + + +See + +Onuferko +et al +. (2024) + +. + + +DNA barcoded material + +Unavailable. + + + + +Redescription + + + +This species was recently redescribed ( + +Onuferko +et al +. 2024 + +). + + + + + +Distribution + + + +Northern and eastern South America ( + +Onuferko +et al +. 2024 + +: fig. 1F). + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + + +See + +Onuferko +et al +. (2024) + +. + + + + + +Remarks + + + +Detailed morphological and taxonomic remarks about this species are given in + +Onuferko +et al +. (2024) + +. + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFE7FFBB3002FEE1FAE4FB82.xml b/data/03/8F/53/038F531FFFE7FFBB3002FEE1FAE4FB82.xml new file mode 100644 index 00000000000..19233648679 --- /dev/null +++ b/data/03/8F/53/038F531FFFE7FFBB3002FEE1FAE4FB82.xml @@ -0,0 +1,303 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus alvarengai +Moure, 1955 + + + + + + +Fig. 13A + + + + + + + +Triepeolus alvarengai +Moure, 1955: 126 + + +( +♀ +). + + + + + +Triepeolus alvarengai + +– + + +Onuferko +et al +. 2024: 7 + + +, figs 1a, 2, 3a (redescription of female, description of male). + + + + +Proposed common name + + + + +Alvarenga’s triepeolus. + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. alvarengai + +apart from all other + +Triepeolus + +in the + +simplex + +species group: the mesoscutum has well-defined paramedian bands ( +Fig. 13A +); the T1 basal band is arched, continuous with (and indistinguishable from) the lateral longitudinal bands, and mesally concave on each side, such that the discal patch is semicircular ( +Fig. 13A +); the T1 apical transverse band (if present) is not more narrowly interrupted medially than the basal band ( +Fig. 13A +); and T2–T4 have complete apical transverse bands of yellow tomentum that are about as broad as (if not broader than) the bands on T1 and do not have basomedially convergent anterolateral extensions ( +Fig. 13A +). Specimens in which T1 has only a basal band or pair of anterolateral patches of pale tomentum (as opposed to both basal and apical transverse bands) may be confused with + +T. mexicanus + +or + +T. nemoralis + +, but in + +T. mexicanus + +pale tomentum on the mesoscutum is restricted to the lateral and posterior margins and in + +T. nemoralis + +the mesoscutum has a large anteromedial ovate patch of yellow tomentum, which may be sparser medially such that the patch is suggestive of ill-defined paramedian bands. Specimens in which T1 has an apical transverse band may be confused with + +T. eumeniformis + +sp. nov. +, but in + +T. eumeniformis + +the T1 basal band (if present) is more widely interrupted medially than the apical transverse band. Additionally, + +T. alvarengai + +can easily be separated from + +T. eumeniformis + +and + +T. mexicanus + +by geography; the former is known only from +Brazil +whereas the latter two species are known only from North and Central America. + + + + +Fig. 13. +Mesosoma and metasoma of male, dorsal view. +A +. + +Triepeolus alvarengai +Moure, 1955 + +(ZMB ed1bbe/M.G.R. Database No. 11694) (image is copyright of the Museum für Naturkunde and reproduced here with permission). +B +. + +T. nemoralis +( +Holmberg, 1886 +) + +(CNC 1801912). + + + + + +Etymology + + + +See + +Onuferko +et al +. (2024) + +. + + + + + +Material examined + + + +See + +Onuferko +et al +. (2024) + +. + + +DNA barcoded material + +Unavailable. + + + + +Redescription + + + +This species was recently redescribed ( + +Onuferko +et al +. 2024 + +). + + + + + +Distribution + + + +Eastern +Brazil +( + +Onuferko +et al +. 2024 + +: fig. 1a). + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + +Unknown. + + + + +Remarks + + + +Detailed morphological and taxonomic remarks about this species are given in + +Onuferko +et al +. (2024) + +. + + + + \ No newline at end of file diff --git a/data/03/8F/53/038F531FFFECFFB43016FEE1FE5DFAEB.xml b/data/03/8F/53/038F531FFFECFFB43016FEE1FE5DFAEB.xml new file mode 100644 index 00000000000..528983642bb --- /dev/null +++ b/data/03/8F/53/038F531FFFECFFB43016FEE1FE5DFAEB.xml @@ -0,0 +1,820 @@ + + + +A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae) + + + +Author + +Onuferko, Thomas M. +6E4CC25A-AD82-42D3-9846-C659EDEAF541 +Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada. +thomas.onuferko@gmail.com + + + +Author + +Rightmyer, Molly G. +90A04B61-0B07-4F3F-8F9E-DDCBDC108314 +Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA. +molly_rightmyer@yahoo.com + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-02 + + +950 + + +1 + + +1 +106 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195 + +journal article +10.5852/ejt.2024.950.2643 +2118-9773 +13691805 +ACFB8240-3FDC-43B8-8200-236BFE00AD94 + + + + + + +Triepeolus eumeniformis + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +639463C4-48D5-4B50-BBF9-1A2A9BF67ED4 + + + + + +Figs 6D +, +10D +, 11C–D, 12C, 16 + + + +Proposed common name + + +Potter-wasp triepeolus. + + + + +Diagnosis + + + +The following morphological features in combination tell + +T. eumeniformis + +sp. nov. +apart from all other + +Triepeolus + +in the + +simplex + +species group: the mesoscutum has well-defined paramedian bands ( +Fig. 16B +), the T1 discal patch is semicircular (Fig. 11C) or trapezoidal (Figs 11D, 16B), the T1 basal band (if present) is more widely interrupted medially than the apical transverse band (Figs 11C–D, 16B), and T3–T4 and usually also T2 have complete apical transverse bands of yellow tomentum that are about as broad as the bands on T1 and do not have basomedially convergent anterolateral extensions ( +Figs 10D +, 11C–D, 16A–C). + +Triepeolus eumeniformis + +most closely resembles specimens of + +T. alvarengai + +that have an apical transverse band on T1 and + +T. paucipunctatus + +sp. nov. +, but in + +T. alvarengai + +the T1 apical transverse band (if present) is not more narrowly interrupted medially than the basal band and in + +T. paucipunctatus + +the T2 apical transverse band has a pair of well-defined basomedially convergent anterolateral extensions and the mesopleura are more sparsely punctate (some i>4d as opposed to ≤ 3d) and have sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae. Additionally, + +T. eumeniformis + +can easily be separated from + +T. alvarengai + +by geography; the former is known only from +Mexico +and Central America whereas the latter is known only from +Brazil +. Males of + +T. eumeniformis + +also closely resemble those of + +T. rufoclypeus +(Fox, 1891) + +(outside of the + +simplex + +species group), but in + +T. rufoclypeus + +S3 has an apical/subapical fringe of dense, long (> 1 MOD), curved, pale setae (as opposed to only short, straight setae; +Fig. 10D +). This species is also very similar in overall appearance to specimens of + +Epeolus flavofasciatus +Smith, 1879 + +( +Hymenoptera +: +Apidae +: +Nomadinae +) in which the axillae and mesoscutellum are black, but both sexes of + +T. eumeniformis + +can easily be told apart from any similar-looking + +Epeolus +Latreille, 1802 + +by their simple mandibles; in + +E. flavofasciatus + +and most other + +Epeolus +spp. + +, the mandibles each have a distinct preapical tooth. + + + + + +Etymology + + + +The specific epithet is derived from + +Eumenes +Latreille, 1802 + +, the +type +genus of the subfamily +Eumeninae +( +Hymenoptera +: +Vespidae +), and was inspired by this species’ resemblance to various kinds of potter wasps. The Latin adjectival suffix ‘-formis’ means ‘having the form of’. + + + + + +Material examined + + + + +Primary +type +material + + + + +HONDURAS +• +♀ +, +holotype +; +Cortés +, Estación Experimental Café (near Peñas Blancas rainforest); + +15 Aug. 1992 + +; +C. Porter +and +L. Stange +leg.; +FSCA + +. + + + +Secondary +type +material + + + + +HONDURAS +• +1 ♀ +, +paratype +; same data as for holotype; +FSCA +M.G.R. Database No. +1321 + +. + + + +MEXICO +– + +Chiapas + +• +1 ♀ +, +paratype +; + +18 km +S of La Trinitaria + +(small road NW of +Hwy +190); + +5 Dec. 1976 + +; +D.E. and J.A. Breedlove +leg.; +CAS + +• + +1 ♀ +, +paratype +; +Ei. Las Golondrinas +( +Acacoyagua +); +15.4340° N +, +92.6522° W +; + +29 Nov. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO- TA-E-41202 + +• + +1 ♀ +, +paratype +; +Ei. Rosario Zacatonales +( +Acacoyagua +); + +8 Dec. 2004 + +; +M. Guzmán +, +M. Rincón +, +J. Esponda +, +C. Balboa +, and +J. Mérida +leg.; +ECOSUR +ECO-TA-E-41813 + +• + +1 ♂ +, +allotype +; same data as for preceding; + +10 Nov. 2006 + +; +J. Mérida +, +M. Guzmán +, +M. Cigarroa +, +J. Toto +, and +C. Balboa +leg.; +ECOSUR +ECO-TAE-50298 + +• + +1 ♀ +, +paratype +; +Ei. Rosario Zacatonales +( +Acacoyagua +); +15.4455° N +, +92.6455° W +; + +29 Nov. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO-TA-E-41255 + +• + +3 ♂♂ +, +paratypes +; same data as for preceding; + +29 Nov. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO-TA-E-41259, ECO-TA-E-41271, ECO-TA-E-41312 + +• + +1 ♂ +, +paratype +; same data as for preceding; + +8 Dec. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO-TA-E-41944 + +• + +1 ♀ +, +paratype +; +Ei. Rosario Zacatonales +( +Acacoyagua +); +15.4480° N +, +92.6435° W +; + +30 Nov. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO-TA-E-41611 + +• + +1 ♂ +, +paratype +; same data as for preceding; + +30 Nov. 2004 + +; +M. Rincón +, +R. Ayala +, and +M. Guzmán +leg.; +ECOSUR +ECO-TA-E-41610 + +• + +1 ♀ +, +paratype +; +Ei. Unión +los +Olivos +( +Mapastepec +); + +8 Nov. 2006 + +; +J. Mérida +, +M. Guzmán +, +M. Cigarroa +, +J. Toto +, and +C. Balboa +leg.; +ECOSUR +ECO-TAE-50103 + +. – + + +Estado de México + +• +1 ♀ +, +paratype +; +Tepotzotlán +; + +12 Nov. 1983 + +; +O. Morales +leg.; +UNAM + +. – + + +Jalisco + +• +1 ♀ +, +paratype +; +Chamela +; + +24 Oct. 1986 + +; “ +A. Rodriguez +P.” leg.; +UNAM + +. – + + +Morelos + +• +1 ♂ +, +paratype +; +Campamento +YMCA, +Tepoztlán +; + +21 Aug. 1958 + +; +H.F. Howden +leg.; +CNC 1801914 + +. – + + +Nayarit + +• +1 ♀ +, +paratype +; + +8 mi +N of Tepic + +; + +25.vii.1954 + +; +M. Cazier +, +W. Gertsch +, and +Bradts +leg.; +AMNH + +. – + + +Tamaulipas + +• +1 ♀ +, +paratype +; +Cañón de Peregrina +; + +20 Oct. 1974 + +; +J.E. Gillaspy +leg.; +USNM + +. – + + +Veracruz + +• +1 ♀ +, +paratype +; + +5 mi +NE of Tinajas + +; + +18 Aug. 1963 + +; +F.D. Parker +and +L.A. Stange +leg.; +UCBME + +• + +1 ♀ +, +paratype +; +Fortín +de las +Flores +; + +17 Sep. 1954 + +; +F.X. Williams +leg.; +CAS + +. + + + +Fig. 16. + +Triepeolus eumeniformis + +sp. nov. +B +, +D +. Holotype, ♀ (FSCA). +A +. Paratype, ♀ (ECOSUR ECO-TA-E-41255), habitus, lateral view. +B +. Habitus, dorsal view. +C +. Allotype, ♂ (ECOSUR ECO- TAE-50298), habitus, lateral view. +D +. Pseudopygidial area, dorsal view. + + + +DNA barcoded material + +Unavailable. + + + + +Description + + + +MEASUREMENTS +OF + +HOLOTYPE + +. Body length +9.5 mm +; ITW +1.9 mm +; head length +2.3 mm +; head width +3.3 mm +; fore wing length +7.7 mm +. + + +Both sexes + + +INTEGUMENT +COLORATION +. Dark brown to black except as follows. Mandible with basal half (except for extreme base) and labrum with lateral margin reddish orange. F1 to some extent, tegula, coxae to femora to some extent, and tibiae (excluding brown meso- and metatibial spurs) and tarsi predominantly to entirely orange. F2 with orange spot basally (entirely dark brown in some +paratypes +). Fore wing membrane dusky subhyaline throughout. Hind wing membrane dusky subhyaline to hyaline. + + +PUBESCENCE +. Face with tomentum densest around antennal socket. Tomentum slightly sparser on clypeus; upper paraocular and frontal areas and vertexal area mostly exposed. Pronotal collar with tomentum uniformly bright yellow. Mesoscutum with well-defined paramedian band of bright yellow tomentum, tapering slightly toward but not attaining anterior margin; pale tomentum otherwise mostly restricted to lateral and posterior margins. Mesopleuron with black, appressed, branched setae just anterior to hypoepimeral area and pale-yellow, appressed, branched setae elsewhere; densely setose only just below pronotal lobe and scrobal groove and along posterior margin above base of mesocoxa (upper half otherwise sparsely setose); ventrolateral half nearly bare, except along margins. Mesopleuron with sparse, pale-yellow, erect/suberect, simple setae (less than ½ MOD in length) in addition to usual appressed, branched setae. Metanotum with tomentum dark brown/gray submedially and pale yellow laterally and medially (pale yellow laterally and black medially in some +paratypes +). Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. T1 with basal transverse band of pale-yellow tomentum widely interrupted medially and arched but distinguishable from lateral longitudinal band (continuous with (and indistinguishable from) lateral longitudinal band in +allotype +and multiple +paratypes +), apical transverse band of bright yellow tomentum separated into pair of rounded lobes medially, discal patch trapezoidal (semicircular in +allotype +and multiple +paratypes +). T2–T4 with complete apical transverse bands of bright yellow tomentum without anterolateral extensions (T2 band with pair of very short and small anterolateral extensions in some +paratypes +). S2–S3 with apical transverse bands of white tomentum. + + +SURFACE +SCULPTURE +. Labrum coarsely and densely (most i <1d) rugose-punctate. Clypeus densely punctate (most i ≤ 1d) but interspaces well defined, shining; with many small punctures among larger ones. Vertexal area densely punctate (most i ≤1d). Mesoscutum, mesoscutellum, and axilla with punctures more or less equally dense (most i ≤1d); interspaces well defined, dull due to tessellate surface microsculpture. Mesopleuron with punctures in upper half denser (most i≤ 1d) than in ventrolateral half (i ≤3d); interspaces shining; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i≈ 1d), and evenly distributed; interspaces somewhat dull due to tessellate surface microsculpture. + + +STRUCTURE +. Labral apex with pair of small denticles, each preceded by discrete longitudinal ridge. Pronotal collar rather short (medial length ~ ⅔ MOD). Mesoscutellum weakly bigibbous. Axilla not extending beyond midlength of mesoscutellum; tip visible but somewhat blunt, mesally unattached to mesoscutellum for less than ⅓ medial length of axilla; lateral margin relatively straight. Fore wing with three submarginal cells. + + +Female + +Scape and pedicel extensively orange. T5 with reddish brown and concave apical margin and large patch of pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area with triangular region of posteriorly directed setae with three subregions (basal patch of dense, golden setae; darker subapical band of sparser, coppery setae; and apical row of dense, suberect, silvery setae) within larger trapezoidal space of posteromedially directed dark brown setae. Pygidial plate reddish brown in part and apically truncate. S4 with apical transverse band of white tomentum. S5 straight in lateral view, with apical fimbria of coppery bristle-like setae. + +Male + + +Pedicel orange in part in some +paratypes +. T5–T6 with complete apical transverse bands of bright yellow tomentum. Pygidial plate with small orange spot subapically, relatively flat and apically rounded. S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae, those of S4 coppery to silvery, those of S5 coppery and contrasting with apical transverse bands of white tomentum of preceding sterna. + + + + + +Distribution + + + +Western +Mexico +to the Northern Triangle of Central America (Fig. 12C). + + + + + +Ecology + + + +Host records + +Unknown. + +Floral records + + +The label of one examined voucher specimen indicates that this species has been collected from the following flowering plant species: in +Lamiaceae Martinov +, + +Salvia +sp. + + + + + + +Remarks + + + +This species exhibits continuous variation in the shape of the T1 discal patch, which ranges from strongly trapezoidal to almost perfectly semicircular. Additionally, the T1 basal band varies in the degree to which it is interrupted medially and in some specimens appears to be absent or indistinguishable from the lateral longitudinal bands. Based on known records, adults of + +T. eumeniformis + +sp. nov. +are active from July to December. + + + + \ No newline at end of file