Add updates up until 2025-02-26 13:17:52

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Key to the apterous (or micropterous) subgenera of
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1 Metathoracic sternite with a pair of long digitate projections.................
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2 Hind femora unarmed.........................................
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3 Apical area of paraproctal processes bifurcate................
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<bibRefCitation id="D12C4B699C344340FB78FDC1EDF79131" author="Pang, S. Y. &amp; Lu, X. Y. &amp; Bian, X." box="[1214,1436,537,559]" pageId="1" pageNumber="374" pagination="253 - 267" refId="ref2550" refString="Pang, S. Y., Lu, X. Y. &amp; Bian, X. (2023) Study of the Chinese members of the subfamily Anabropsinae (Orthoptera: Anostostomatidae) VII: Description of two new subgenera and five new species of Anabropsis from China. Zootaxa, 5318 (2), 253-267. https://doi.org/10.11646/zootaxa.5318.2.5" type="journal article" year="2023">Pang, Lu &amp; Bian, 2023</bibRefCitation>
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- Apical area of paraproctal processes unbranched........................
<taxonomicName id="72BD4D1B9C344340FC1CFDEEEDF79154" ID-CoL="8QKGR" authority="Gorochov, 1988" authorityName="Gorochov" authorityYear="1988" box="[986,1436,566,588]" class="Insecta" family="Anostostomatidae" genus="Anabropsis" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="subGenus" subGenus="Apteranabropsis">
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(
<emphasis id="87C9EA8A9C344340FB93FDEEEC9F9154" box="[1109,1268,566,587]" italics="true" pageId="1" pageNumber="374">Apteranabropsis</emphasis>
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<bibRefCitation id="D12C4B699C344340FAC4FDEEEDF79154" author="Gorochov, A. V." box="[1282,1436,566,588]" pageId="1" pageNumber="374" pagination="353 - 366" refId="ref2244" refString="Gorochov, A. V. (1988) System and phylogeny of the recent Orthoptera of the superfamilies Hagloidea and Stenopelmatoidea with a description of new taxa. Communication 1. Zoologicheskii Zhurnal, 67 (3), 353-366." type="journal article" year="1988">Gorochov, 1988</bibRefCitation>
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<typeStatus id="6A06883A9C344340FF51FBC9E96D9734" box="[151,262,1041,1067]" pageId="1" pageNumber="374" type="holotype">Holotype</typeStatus>
.
@ -80,11 +84,9 @@
,
<collectingRegion id="7779F87A9C344340FE48FBCAEA3F9734" box="[398,596,1041,1068]" country="China" name="Yunnan" pageId="1" pageNumber="374">Yunnan Province</collectingRegion>
,
<collectingCounty id="5C634E149C344340FDA4FBC9EA869734" box="[610,749,1041,1067]" pageId="1" pageNumber="374">Dehong Dai</collectingCounty>
and
<collectingMunicipality id="5566ACE29C344340FCECFBC9ECE59734" box="[810,1166,1041,1068]" pageId="1" pageNumber="374">Jingpo Autonomous Prefecture</collectingMunicipality>
<collectingCounty id="5C634E149C344340FDA4FBC9ECE59734" box="[610,1166,1041,1068]" pageId="1" pageNumber="374">Dehong Dai and Jingpo Autonomous Prefecture</collectingCounty>
,
<location id="B06260439C344340FB5AFBC9E8A69750" LSID="urn:lsid:plazi:treatment:3D14878E9C344343FF51FC5DEA4A92EA:B06260439C344340FB5AFBC9E8A69750" country="China" county="Dehong Dai" latitude="24.8" longLatPrecision="7503" longitude="98.8" municipality="Jingpo Autonomous Prefecture" name="Gaoligongshan Nature Park" pageId="1" pageNumber="374" stateProvince="Yunnan">Gaoligongshan Nature Park</location>
<location id="B06260439C344340FB5AFBC9E8A69750" LSID="urn:lsid:plazi:treatment:3D14878E9C344343FF51FC5DEA4A92EA:B06260439C344340FB5AFBC9E8A69750" country="China" county="Dehong Dai and Jingpo Autonomous Prefecture" latitude="24.8" longLatPrecision="7503" longitude="98.8" municipality="Jingpo Autonomous Prefecture" name="Gaoligongshan Nature Park" pageId="1" pageNumber="374" stateProvince="Yunnan Province">Gaoligongshan Nature Park</location>
,
<geoCoordinate id="D089505F9C344340FF1FFBEDE9479750" box="[217,300,1077,1103]" degrees="24.8" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="5555" value="24.8">24.8°N</geoCoordinate>
,
@ -108,15 +110,14 @@ and
</materialsCitation>
</paragraph>
<paragraph id="B50236989C344340FF01FB81EAD7976C" blockId="1.[151,1437,1041,2040]" box="[199,700,1113,1140]" pageId="1" pageNumber="374">
<materialsCitation id="05D53CC59C344340FF01FB81EAD3976C" box="[199,696,1113,1140]" pageId="1" pageNumber="374" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<materialsCitation id="05D53CC59C344340FF01FB81EAD3976C" box="[199,696,1113,1140]" collectingDate="2022-05-04" collectorName="Michael Zelun Lee &amp; Yejie Lin &amp; Tongyao Jiang &amp; Zhitao Zhang" country="China" county="Dehong Dai and Jingpo Autonomous Prefecture" elevation="2000" latitude="24.8" location="Gaoligongshan Nature Park" longitude="98.8" municipality="Jingpo Autonomous Prefecture" pageId="1" pageNumber="374" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<emphasis id="87C9EA8A9C344340FF01FB81E92D976C" bold="true" box="[199,326,1113,1139]" pageId="1" pageNumber="374">
<typeStatus id="6A06883A9C344340FF01FB81E92A976C" box="[199,321,1113,1139]" pageId="1" pageNumber="374" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="A3BBFD119C344340FE8BFB81E91A976C" box="[333,369,1113,1139]" count="1" pageId="1" pageNumber="374" type="male">1♂</specimenCount>
<specimenCount id="A3BBFD119C344340FEB7FB81E9FF976C" box="[369,404,1113,1139]" count="2" pageId="1" pageNumber="374" type="female">2♀</specimenCount>
, same data as in
<typeStatus id="6A06883A9C344340FD94FB81EAD3976C" box="[594,696,1113,1139]" pageId="1" pageNumber="374" type="holotype">holotype</typeStatus>
, same data as in holotype
</materialsCitation>
.
</paragraph>
@ -149,9 +150,9 @@ Fore and mid coxae with a spine (
). Genicular lobes of all femora without spines (
<figureCitation id="2D862A1D9C344340FAD8FAA1ED04968C" box="[1310,1391,1401,1427]" captionStart="FIGURE 2" captionStartId="3.[151,250,830,854]" captionTargetBox="[155,1432,180,801]" captionTargetId="figure-15@3.[151,1435,179,804]" captionTargetPageId="3" captionText="FIGURE 2. Right tibiae of Anabropsis (Spinanabropsis) zeluni sp. nov. A, D. external side; B, C. internal side." figureDoi="http://doi.org/10.5281/zenodo.14896281" httpUri="https://zenodo.org/record/14896281/files/figure.png" pageId="1" pageNumber="374">Figs. 2</figureCitation>
,
<figureCitation id="2D862A1D9C344340FAB8FAA1EDE5968C" box="[1406,1422,1401,1427]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." pageId="1" pageNumber="374">3</figureCitation>
<figureCitation id="2D862A1D9C344340FAB8FAA1EDE5968C" box="[1406,1422,1401,1427]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionTargetId="figure-38@3.[355,1232,912,1983]" captionTargetPageId="3" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14930814" httpUri="https://zenodo.org/record/14930814/files/figure.png" pageId="1" pageNumber="374">3</figureCitation>
). External surface of hind femur with distinct chevron ridges (
<figureCitation id="2D862A1D9C344340FC86FA45EBEE96A8" box="[832,901,1437,1463]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." pageId="1" pageNumber="374">Fig. 3</figureCitation>
<figureCitation id="2D862A1D9C344340FC86FA45EBEE96A8" box="[832,901,1437,1463]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionTargetId="figure-38@3.[355,1232,912,1983]" captionTargetPageId="3" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14930814" httpUri="https://zenodo.org/record/14930814/files/figure.png" pageId="1" pageNumber="374">Fig. 3</figureCitation>
). Tympana elongated oval on both sides of fore tibiae (
<figureCitation id="2D862A1D9C344340FF23FA19E94196C4" box="[229,298,1473,1499]" captionStart="FIGURE 2" captionStartId="3.[151,250,830,854]" captionTargetBox="[155,1432,180,801]" captionTargetId="figure-15@3.[151,1435,179,804]" captionTargetPageId="3" captionText="FIGURE 2. Right tibiae of Anabropsis (Spinanabropsis) zeluni sp. nov. A, D. external side; B, C. internal side." figureDoi="http://doi.org/10.5281/zenodo.14896281" httpUri="https://zenodo.org/record/14896281/files/figure.png" pageId="1" pageNumber="374">Fig. 2</figureCitation>
AB). All femora without spinules. Fore tibia dorsally with 1 internal spine and 1 pair of apical spurs; ventrally with 4 pairs of moveable spines and 1 pair of apical spurs, with the internal spurs of both pairs being longer (
@ -159,7 +160,7 @@ AB). All femora without spinules. Fore tibia dorsally with 1 internal spine and
AB). Mid tibia dorsally with 3 internal spines, 2 external spines and 1 pair of apical spurs; ventrally with 4 pairs of spines, and 1 pair of apical spurs (
<figureCitation id="2D862A1D9C344340FDB1F9F5EAD19558" box="[631,698,1581,1607]" captionStart="FIGURE 2" captionStartId="3.[151,250,830,854]" captionTargetBox="[155,1432,180,801]" captionTargetId="figure-15@3.[151,1435,179,804]" captionTargetPageId="3" captionText="FIGURE 2. Right tibiae of Anabropsis (Spinanabropsis) zeluni sp. nov. A, D. external side; B, C. internal side." figureDoi="http://doi.org/10.5281/zenodo.14896281" httpUri="https://zenodo.org/record/14896281/files/figure.png" pageId="1" pageNumber="374">Fig. 2</figureCitation>
CD). Hind tibia dorsally with 9~10 internal, 9~10 external spines and 1 pair of elongated apical spines; ventrally with 1 pair of short spines, and 3 pairs of apical spurs gradually increasing in length and size towards tip (
<figureCitation id="2D862A1D9C344340FDACF9ADEAC59590" box="[618,686,1653,1679]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." pageId="1" pageNumber="374">Fig. 3</figureCitation>
<figureCitation id="2D862A1D9C344340FDACF9ADEAC59590" box="[618,686,1653,1679]" captionStart="FIGURE 3" captionStartId="3.[151,250,2009,2033]" captionTargetId="figure-38@3.[355,1232,912,1983]" captionTargetPageId="3" captionText="FIGURE 3. Hind legs of Anabropsis (Spinanabropsis) zeluni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14930814" httpUri="https://zenodo.org/record/14930814/files/figure.png" pageId="1" pageNumber="374">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="B50236989C344340FF01F941EBBF9478" blockId="1.[151,1437,1041,2040]" pageId="1" pageNumber="374">

View file

@ -0,0 +1,139 @@
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<heading id="8F6481EDE500935C22BBFC017514FC23" box="[175,585,976,1003]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="313" reason="2">
<taxonomicName id="13934D02E500935C22BBFC017514FC23" authority="(Boisduval, 1835)" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[175,585,976,1003]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="sulcigera">
<emphasis id="E6E7EA93E500935C22BBFC0176DDFC23" bold="true" box="[175,384,976,1003]" italics="true" pageId="8" pageNumber="313">Promethis sulcigera</emphasis>
(
<bibRefCitation id="B0024B70E500935C239BFC01751CFC23" author="BOISDUVAL J. B. A. D. DE" bookContentInfo="716 p." box="[399,577,976,1003]" journalOrPublisher="J. Tastu, Paris" pageId="8" pageNumber="313" refId="ref6675" refString="BOISDUVAL J. B. A. D. DE 1835. - Voyage de decouvertes de l' Astrolabe execute par l'ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, de M. Dumont d'Urville. Faune entomologique de l'ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Deuxieme partie. Coleopteres et autres ordres. J. Tastu, Paris, VII, 716 p." title="Voyage de decouvertes de l' Astrolabe execute par l'ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, de M. Dumont d'Urville. Faune entomologique de l'ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Deuxieme partie. Coleopteres et autres ordres" type="book" year="1835">Boisduval, 1835</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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<subSubSection id="9C89650AE500935C2265FBDE768DFBEE" box="[113,464,1038,1062]" pageId="8" pageNumber="313" type="reference_group">
<paragraph id="D42C3681E500935C2265FBDE768DFBEE" blockId="8.[113,464,1038,1062]" box="[113,464,1038,1062]" pageId="8" pageNumber="313">
<taxonomicName id="13934D02E500935C2265FBDE7691FBEE" authority="Boisduval, 1835: 256" authorityName="Boisduval" authorityPageNumber="256" authorityYear="1835" box="[113,460,1038,1062]" class="Insecta" family="Tenebrionidae" genus="Upis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="sulcigera">
<emphasis id="E6E7EA93E500935C2265FBDE77AFFBEE" box="[113,242,1038,1062]" italics="true" pageId="8" pageNumber="313">Upis sulcigera</emphasis>
<bibRefCitation id="B0024B70E500935C22ECFBDF7691FBEE" author="BOISDUVAL J. B. A. D. DE" bookContentInfo="716 p." box="[248,460,1038,1062]" journalOrPublisher="J. Tastu, Paris" pageId="8" pageNumber="313" refId="ref6675" refString="BOISDUVAL J. B. A. D. DE 1835. - Voyage de decouvertes de l' Astrolabe execute par l'ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, de M. Dumont d'Urville. Faune entomologique de l'ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Deuxieme partie. Coleopteres et autres ordres. J. Tastu, Paris, VII, 716 p." title="Voyage de decouvertes de l' Astrolabe execute par l'ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, de M. Dumont d'Urville. Faune entomologique de l'ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Deuxieme partie. Coleopteres et autres ordres" type="book" year="1835">Boisduval, 1835: 256</bibRefCitation>
</taxonomicName>
.
</paragraph>
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MATERIAL EXAMINED. —
<emphasis id="E6E7EA93E500935C2348FB9276EEFB93" bold="true" box="[348,435,1091,1115]" pageId="8" pageNumber="313">
<collectingCountry id="AC847611E500935C2348FB9276F2FB93" box="[348,431,1091,1115]" name="Vanuatu" pageId="8" pageNumber="313">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Luganville,CETRAV in Saraoutou (
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,
<geoCoordinate id="B1A75046E500935C2011FB8F7521FBBE" box="[517,636,1118,1142]" degrees="167" direction="east" minutes="12" orientation="longitude" pageId="8" pageNumber="313" precision="15" seconds="32" value="167.2089">167°1232”E</geoCoordinate>
), 1-
<date id="A02D1041E500935C2294FBA97628FB58" box="[128,373,1144,1168]" pageId="8" pageNumber="313" value="2006-10-11" valueMax="2006-10-18" valueMin="2006-10-11">11.X and 18-20.X.2006</date>
, at night with a headlamp, on the decaying trunk of an old mango tree (
<taxonomicName id="13934D02E500935C203DFB4277F7FB0D" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="313" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="E6E7EA93E500935C203DFB4277F7FB0D" italics="true" pageId="8" pageNumber="313">Mangifera indica</emphasis>
</taxonomicName>
), leg. L. Soldati,
<specimenCount id="C295FD08E500935C235CFB7C7623FB0C" box="[328,382,1197,1221]" pageId="8" pageNumber="313" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="B282AE44E500935C239EFB7C7680FB0D" box="[394,477,1197,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="313">MNHN</collectionCode>
),
<specimenCount id="C295FD08E500935C23FBFB7C757BFB0C" box="[495,550,1197,1221]" pageId="8" pageNumber="313" type="female">2 ♀♀</specimenCount>
(coll.
<collectionCode id="B282AE44E500935C204AFB7C7521FB0D" box="[606,636,1197,1221]" country="United Kingdom" name="Linnean Society of London" pageId="8" pageNumber="313">LS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE500935C2265FB32763CFA7B" pageId="8" pageNumber="313" type="distribution">
<paragraph id="D42C3681E500935C2265FB32763CFA7B" blockId="8.[111,649,1250,1459]" pageId="8" pageNumber="313">
DISTRIBUTION. — Described from “île dAmboine” (
<collectingCountry id="AC847611E500935C226EFB2D7798FADC" box="[122,197,1276,1300]" name="Indonesia" pageId="8" pageNumber="313">Ambon</collectingCountry>
,
<collectingCountry id="AC847611E500935C22D8FB2D7676FADC" box="[204,299,1276,1300]" name="Indonesia" pageId="8" pageNumber="313">Indonesia</collectingCountry>
), this widespread species is also known from
<collectingCountry id="AC847611E500935C22B5FAC677B5FAE7" box="[161,232,1303,1327]" name="Taiwan" pageId="8" pageNumber="313">Taiwan</collectingCountry>
,
<collectingCountry id="AC847611E500935C22FBFAC67606FAE7" box="[239,347,1303,1327]" name="Philippines" pageId="8" pageNumber="313">Philippines</collectingCountry>
, Moluccan Islands, New
<collectingCountry id="AC847611E500935C2055FAC675DBFAE7" box="[577,646,1303,1327]" name="Guinea-Bissau" pageId="8" pageNumber="313">Guinea</collectingCountry>
and the surrounding islands and
<collectingCountry id="AC847611E500935C23B2FAE076A6FA81" box="[422,507,1329,1353]" name="Australia" pageId="8" pageNumber="313">Australia</collectingCountry>
(
<collectingRegion id="1657F863E500935C2013FAE07523FA81" box="[519,638,1329,1353]" country="Australia" name="Queensland" pageId="8" pageNumber="313">Queensland</collectingRegion>
) (
<bibRefCitation id="B0024B70E500935C2262FA9D77A3FAAC" author="Kaszab" box="[118,254,1356,1380]" firstAuthor="Kaszab" pageId="8" pageNumber="313" pagination="67 - 170" refId="ref7513" refString="KASZAB Z. 1988 a. - Katalog und Bestimmungstabelle der Gattung Promethis Pascoe, 1869 (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae 34 (2 - 3): 67 - 170." type="journal article" year="1988" yearSuffix="a">Kaszab 1988a</bibRefCitation>
). It is also present in the
<collectingCountry id="AC847611E500935C23F5FA9D75DCFAAC" box="[481,641,1356,1380]" name="Solomon Islands" pageId="8" pageNumber="313">Solomon Islands</collectingCountry>
, the
<collectingCountry id="AC847611E500935C2281FAB7778BFAB6" box="[149,214,1382,1406]" name="Samoa" pageId="8" pageNumber="313">Samoa</collectingCountry>
Islands,
<collectingCountry id="AC847611E500935C2338FAB7763CFAB6" box="[300,353,1382,1406]" name="Palau" pageId="8" pageNumber="313">Palau</collectingCountry>
and the
<collectingCountry id="AC847611E500935C23A3FAB67554FAB6" box="[439,521,1383,1406]" name="Vanuatu" pageId="8" pageNumber="313">Vanuatu</collectingCountry>
Archipelago (Aoba, Malo, Malekula [Mallicolo] and Epi, Erronango islands) (
<bibRefCitation id="B0024B70E500935C22D2FA4A760BFA7B" author="Kaszab" box="[198,342,1435,1459]" firstAuthor="Kaszab" pageId="8" pageNumber="313" pagination="55 - 116" refId="ref7551" refString="KASZAB Z. 1988 b. - Faunistische Angaben der Gattung Promethis Pascoe, 1869 (Coleoptera, Tenebrionidae). Folia Entomologica Hungarica 49: 55 - 116." type="journal article" year="1988" yearSuffix="b">Kaszab 1988b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,430 @@
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<mods:title id="015BB469E3F2112A1D5DC9C274043107">Preliminary report on the Tenebrionidae (Insecta, Coleoptera) collected during the SANTO 2006 expedition to Vanuatu, with description of a new species of the genus Uloma Dejean, 1821</mods:title>
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<mods:namePart id="0CDA228276B1901DA69CDB51246D3AFF">Soldati, Laurent</mods:namePart>
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<mods:namePart id="535C715600269878A2CC78F90BAB9067">Kergoat, Gael J.</mods:namePart>
<mods:affiliation id="0B3814A1AFD8A049E97A9499B57ABA9B">INRA - UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier Supagro), Campus international de Baillarguet, CS 30016, F- 34988 Montferrier-sur-Lez cedex (France) soldati @ supagro. inra. fr kergoat @ supagro. inra. fr</mods:affiliation>
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</mods:role>
<mods:namePart id="E7C07840B4F1D2DCBF1E84C8A96E98A0">Condamine, Fabien L.</mods:namePart>
<mods:affiliation id="48F87BF4FFEB703E7BADBC989A2DFAAB">Mathématiques appliquées - UMR 7641 (CNRS, École polytechnique), route de Saclay, F- 91128 Palaiseau cedex (France) fabien. condamine @ gmail. com</mods:affiliation>
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<mods:title id="0E47D63E1D35FFD4CFE0B459601290A2">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="0D8E9D0CE42DCE422D968012A92BBD6A">
<mods:date id="9A27E9A33D27A5A05B36F814758A7BEB">2012</mods:date>
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<treatment id="5C3A8797E500935E22ABF99874B0FDC7" LSID="urn:lsid:plazi:treatment:5C3A8797E500935E22ABF99874B0FDC7" httpUri="http://treatment.plazi.org/id/5C3A8797E500935E22ABF99874B0FDC7" lastPageId="10" lastPageNumber="315" pageId="8" pageNumber="313">
<subSubSection id="9C89650AE500935C22ABF9987564F9AD" box="[191,569,1609,1637]" pageId="8" pageNumber="313" type="nomenclature">
<paragraph id="D42C3681E500935C22ABF9987564F9AD" blockId="8.[191,569,1609,1637]" box="[191,569,1609,1637]" pageId="8" pageNumber="313">
<heading id="8F6481EDE500935C22ABF9987564F9AD" box="[191,569,1609,1637]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="313" reason="2">
<taxonomicName id="13934D02E500935C22ABF9987564F9AD" authority="Kulzer, 1951" authorityName="Kulzer" authorityYear="1951" box="[191,569,1609,1637]" class="Insecta" family="Tenebrionidae" genus="Thesilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="mallicolensis">
<emphasis id="E6E7EA93E500935C22ABF99876FAF9AC" bold="true" box="[191,423,1609,1636]" italics="true" pageId="8" pageNumber="313">Thesilea mallicolensis</emphasis>
<bibRefCitation id="B0024B70E500935C23BAF99B7564F9AD" author="Kulzer" box="[430,569,1610,1637]" firstAuthor="Kulzer" pageId="8" pageNumber="313" pagination="116 - 171" refId="ref7581" refString="KULZER H. 1951. - Vierter Beitrag zur Kenntnis der Tenebrioniden. Entomologischen Arbeiten aus dem Museum G. Frey 2: 116 - 171." type="journal article" year="1951">Kulzer, 1951</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<caption id="80EC6609E500935C20B1F9B373E6F969" ID-DOI="http://doi.org/10.5281/zenodo.5165437" ID-Zenodo-Dep="5165437" httpUri="https://zenodo.org/record/5165437/files/figure.png" pageId="8" pageNumber="313" startId="8.[677,688,1634,1651]" targetBox="[677,1209,216,1586]" targetPageId="8">
<paragraph id="D42C3681E500935C20B1F9B373E6F969" blockId="8.[677,1211,1634,1697]" pageId="8" pageNumber="313">
FIG. 3. —
<taxonomicName id="13934D02E500935C20E1F9B374BFF9BB" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[757,994,1634,1651]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E500935C20E1F9B374D2F9BB" box="[757,911,1634,1651]" italics="true" pageId="8" pageNumber="313">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E500935C21FCF9B3734AF9BB" box="[1000,1047,1634,1651]" pageId="8" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ (MNHN EC2280):
<emphasis id="E6E7EA93E500935C20E7F9A875A2F942" bold="true" box="[755,767,1657,1674]" pageId="8" pageNumber="313">A</emphasis>
, anterior tibia (upper face);
<emphasis id="E6E7EA93E500935C21C1F9A874BFF942" bold="true" box="[981,994,1657,1674]" pageId="8" pageNumber="313">B</emphasis>
, metathorax and abdomen (ventral view).Scale bars:A,0,.5 mm;B,1 mm.Photographs:L.Soldati.
</paragraph>
</caption>
<subSubSection id="9C89650AE500935C2265F95A76AEF96B" box="[113,499,1675,1699]" pageId="8" pageNumber="313" type="reference_group">
<paragraph id="D42C3681E500935C2265F95A76AEF96B" blockId="8.[113,499,1675,1699]" box="[113,499,1675,1699]" pageId="8" pageNumber="313">
<treatmentCitationGroup id="F48311AFE500935C2265F95A76AEF96B" box="[113,499,1675,1699]" pageId="8" pageNumber="313">
<treatmentCitation id="55321090E500935C2265F95A76B2F96B" author="KULZER H." box="[113,495,1675,1699]" page="148" pageId="8" pageNumber="313" year="1951">
<taxonomicName id="13934D02E500935C2265F95A76B2F96B" authority="Kulzer, 1951: 148" authorityName="Kulzer" authorityPageNumber="148" authorityYear="1951" box="[113,495,1675,1699]" class="Insecta" family="Tenebrionidae" genus="Thesilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="mallicolensis">
<emphasis id="E6E7EA93E500935C2265F95A766BF96B" box="[113,310,1675,1699]" italics="true" pageId="8" pageNumber="313">Thesilea mallicolensis</emphasis>
<bibRefCitation id="B0024B70E500935C2328F95A76B2F96B" author="Kulzer" box="[316,495,1675,1699]" firstAuthor="Kulzer" page="148" pageId="8" pageNumber="313" pagination="116 - 171" refId="ref7581" refString="KULZER H. 1951. - Vierter Beitrag zur Kenntnis der Tenebrioniden. Entomologischen Arbeiten aus dem Museum G. Frey 2: 116 - 171." type="journal article" year="1951">Kulzer, 1951: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE501935D2264FF0976FDFE46" pageId="9" pageNumber="314" type="materials_examined">
<paragraph id="D42C3681E501935D2264FF0976FDFE46" blockId="9.[111,649,215,398]" pageId="9" pageNumber="314">
MATERIAL EXAMINED. —
<emphasis id="E6E7EA93E501935D2379FF06769AFF27" bold="true" box="[365,455,215,239]" pageId="9" pageNumber="314">
<collectingCountry id="AC847611E501935D2379FF06769EFF27" box="[365,451,215,239]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Luganville, CETRAV in Saraoutou, ecotone between dense rainforest and coffee tree plantation, beating shrubs,
<date id="A02D1041E501935D227BFEF777ACFEF6" box="[111,241,294,318]" pageId="9" pageNumber="314" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">2-10.X.2006</date>
, leg. L. Soldati,
<specimenCount id="C295FD08E501935D2383FEF6768BFEF6" box="[407,470,295,319]" pageId="9" pageNumber="314" type="male">2 ♂♂</specimenCount>
and 15 other ex. (
<collectionCode id="B282AE44E501935D226EFE907793FE91" box="[122,206,321,345]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="314">MNHN</collectionCode>
),
<specimenCount id="C295FD08E501935D22F0FE90764EFE90" box="[228,275,321,345]" pageId="9" pageNumber="314" type="generic">5 ex.</specimenCount>
(coll.
<collectionCode id="B282AE44E501935D2345FE907630FE91" box="[337,365,321,345]" country="United Kingdom" name="Linnean Society of London" pageId="9" pageNumber="314">LS</collectionCode>
) (sex unknown). — Espiritu Santo Isl., Butmas, in rainforest,
<date id="A02D1041E501935D2383FE8A76A0FEBB" box="[407,509,347,371]" pageId="9" pageNumber="314" value="2006-10-21">21.X.2006</date>
, leg. L. Soldati,
<specimenCount id="C295FD08E501935D227BFEA777C5FE45" box="[111,152,374,398]" pageId="9" pageNumber="314" type="male">1 ♂</specimenCount>
(
<collectionCode id="B282AE44E501935D22B2FEA777A7FE46" box="[166,250,374,398]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="314">MNHN</collectionCode>
),
<specimenCount id="C295FD08E501935D231BFEA77665FE45" box="[271,312,374,398]" pageId="9" pageNumber="314" type="male">1 ♂</specimenCount>
(coll.
<collectionCode id="B282AE44E501935D2362FEA776C9FE46" box="[374,404,374,398]" country="United Kingdom" name="Linnean Society of London" pageId="9" pageNumber="314">LS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE501935D2265FE777557FDA6" pageId="9" pageNumber="314" type="distribution">
<paragraph id="D42C3681E501935D2265FE777603FDC5" blockId="9.[113,648,421,525]" pageId="9" pageNumber="314">
DISTRIBUTION. — Originally described from Mallicolo (Malekula Island) in the French
<collectingCountry id="AC847611E501935D23AEFE117516FE10" box="[442,587,448,472]" name="Vanuatu" pageId="9" pageNumber="314">New Hebrides</collectingCountry>
(now
<collectingCountry id="AC847611E501935D2267FE0B7794FE3A" box="[115,201,474,498]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
), this species also occurs in the neighbouring island of Espiritu Santo.
</paragraph>
<paragraph id="D42C3681E501935D2266FD857557FDA6" blockId="9.[114,607,596,718]" box="[114,522,596,622]" pageId="9" pageNumber="314">
BIOGEOGRAPHY OF
<collectingCountry id="AC847611E501935D2390FD857557FDA6" box="[388,522,596,622]" name="Vanuatu" pageId="9" pageNumber="314">VANUATU</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE501935E2266FDA574B0FDC7" lastPageId="10" lastPageNumber="315" pageId="9" pageNumber="314" type="description">
<paragraph id="D42C3681E501935D2266FDA5769BFD65" blockId="9.[114,607,596,718]" pageId="9" pageNumber="314">
AND ASSUMPTIONS ON THE ORIGIN OF THE
<taxonomicName id="13934D02E501935D22CAFD42769BFD65" authorityName="Latreille" authorityYear="1802" box="[222,454,659,685]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">TENEBRIONIDAE</taxonomicName>
</paragraph>
<paragraph id="D42C3681E501935D2266FD6576FAFD05" blockId="9.[114,607,596,718]" box="[114,423,691,718]" pageId="9" pageNumber="314">IN THE ARCHIPELAGO.</paragraph>
<paragraph id="D42C3681E501935D2265FD23755AFBA1" blockId="9.[111,649,754,1700]" pageId="9" pageNumber="314">
The
<collectingCountry id="AC847611E501935D228BFD2277A4FCC5" box="[159,249,755,781]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
islands and that of Santo in particular are known to be recently emerged volcanic islands (about 4 million years), and can thus be qualified oceanic islands (
<bibRefCitation id="B0024B70E501935D2334FC80751AFCA4" author="Gillespie &amp; Roderick" box="[288,583,849,876]" firstAuthor="Gillespie" pageId="9" pageNumber="314" pagination="595 - 632" refId="ref7310" refString="GILLESPIE R. &amp; RODERICK G. 2002. - Arthropods on islands: colonization, speciation, and conservation. Annual Review of Entomology 47: 595 - 632." type="journal article" year="2002">Gillespie &amp; Roderick 2002</bibRefCitation>
). This geological evidence suggests that the local fauna most likely originated by dispersal and not by vicariance. After their arrival, some of the species may have then evolved to endemic species. This leads us to raise the following question: which is (are) the origin(s) of this fauna? In other words, what were the borrowed colonisation routes to reach the archipelago of the
<collectingCountry id="AC847611E501935D23B0FB9E76A2FBA1" box="[420,511,1103,1129]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
?
</paragraph>
<paragraph id="D42C3681E501935D229CFBBF76F1F9CD" blockId="9.[111,649,754,1700]" pageId="9" pageNumber="314">
With reference to the
<taxonomicName id="13934D02E501935D2395FBBF757FFB40" authorityName="Latreille" authorityYear="1802" box="[385,546,1134,1160]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
, one can say that there were nearly (because of the humanmediated introduction of species like
<taxonomicName id="13934D02E501935D200EFB7F7781FB2F" baseAuthorityName="Herbst" baseAuthorityYear="1797" class="Insecta" family="Tenebrionidae" genus="Tribolium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="castaneum">
<emphasis id="E6E7EA93E501935D200EFB7F7781FB2F" italics="true" pageId="9" pageNumber="314">Tribolium castaneum</emphasis>
</taxonomicName>
) as many colonisation events as different genera existing in
<collectingCountry id="AC847611E501935D2330FB3F7623FAC0" box="[292,382,1262,1288]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
, which are not considered as sister clades. Note that if a genus is represented by more than one species, we cannot exclude the hypothesis of multiple colonisation events as well (e.g.,
<bibRefCitation id="B0024B70E501935D22BBFABD7634FA4E" author="Balke" box="[175,361,1388,1414]" etAl="et al." firstAuthor="Balke" pageId="9" pageNumber="314" pagination="505 - 516" refId="ref6675" refString="BALKE M., PONS J., RIBERA I., SAGATA K. &amp; VOGLER A. P. 2007. - Infrequent and unidirectional colonization of megadiverse Papuadytes diving beetles in New Caledonia and New Guinea. Molecular Phylogenetics Evolution 42: 505 - 516. BOISDUVAL J. B. A. D. DE 1835. - Voyage de decouvertes de l' Astrolabe execute par l'ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, de M. Dumont d'Urville. Faune entomologique de l'ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Deuxieme partie. Coleopteres et autres ordres. J. Tastu, Paris, VII, 716 p." type="book chapter" year="2007">
Balke
<emphasis id="E6E7EA93E501935D22E6FABC767BFA4E" box="[242,294,1388,1414]" italics="true" pageId="9" pageNumber="314">et al.</emphasis>
2007
</bibRefCitation>
). If we replace
<collectingCountry id="AC847611E501935D201AFABD7534FA4E" box="[526,617,1388,1414]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
in a geographical context, one can put forth biogeographic hypotheses as for the source of these dispersions. Indeed, these events can come from several areas around the archipelago.
</paragraph>
<paragraph id="D42C3681E501935D229CF9DB743BFA6E" blockId="9.[111,649,754,1700]" lastBlockId="9.[674,1212,216,1700]" pageId="9" pageNumber="314">
First of all, several genera of
<taxonomicName id="13934D02E501935D23B8F9DB7518F9EC" authorityName="Latreille" authorityYear="1802" box="[428,581,1546,1572]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
could originate from New
<collectingCountry id="AC847611E501935D2353F9FB76C4F98C" box="[327,409,1578,1604]" name="Guinea-Bissau" pageId="9" pageNumber="314">Guinea</collectingCountry>
, which probably holds the greatest diversity of
<taxonomicName id="13934D02E501935D2379F99B7555F9AC" authorityName="Latreille" authorityYear="1802" box="[365,520,1610,1636]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
of the Australasian region (
<bibRefCitation id="B0024B70E501935D2339F9B8769EF94C" author="Gebien" box="[301,451,1641,1668]" firstAuthor="Gebien" pageId="9" pageNumber="314" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien 1920</bibRefCitation>
). Its geographical position makes it obvious that its fauna is part of the Melanesian arch. In particular, dispersals to the
<collectingCountry id="AC847611E501935D20B7FF2975A6FEDA" box="[675,763,248,274]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
could be facilitated by the
<collectingCountry id="AC847611E501935D2613FF2673E9FED9" box="[1031,1204,247,273]" name="Solomon Islands" pageId="9" pageNumber="314">Solomon Islands</collectingCountry>
, as they are themselves strongly influenced by New Guinean fauna (
<bibRefCitation id="B0024B70E501935D2140FEE67487FE99" author="Merkl" box="[852,986,311,337]" firstAuthor="Merkl" pageId="9" pageNumber="314" pagination="235 - 254" refId="ref7681" refString="MERKL O. 1989. - Melanesian representatives of Toxicum and Cryphaeus (Coleoptera, Tenebrionidae: Toxicini). Acta Zoologica Academiae Scientiarum Hungaricae 35 (3 - 4): 235 - 254." type="journal article" year="1989">Merkl 1989</bibRefCitation>
;
<bibRefCitation id="B0024B70E501935D21F1FEE67325FE99" author="Bremer" box="[997,1144,311,337]" firstAuthor="Bremer" pageId="9" pageNumber="314" pagination="509 - 528" refId="ref6921" refString="BREMER H. - J. 1993. - Neue Arten des Genus Corticeus Piller &amp; Mitterpacher, 1783, aus der papuanischmelanesischen Region, II. Mitteilung (Coleoptera, Tenebrionidae, Hypophloeini). Entomofauna 14: 509 - 528." type="journal article" year="1993">Bremer 1993</bibRefCitation>
). This assumption is corroborated by the presence of the genus
<taxonomicName id="13934D02E501935D20F8FEA67435FE59" authorityName="Dalman" authorityYear="1823" box="[748,872,375,401]" class="Insecta" family="Tenebrionidae" genus="Amarygmus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D20F8FEA67435FE59" box="[748,872,375,401]" italics="true" pageId="9" pageNumber="314">Amarygmus</emphasis>
</taxonomicName>
in the
<collectingCountry id="AC847611E501935D21A8FEA67341FE59" box="[956,1052,375,401]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
, whose centre of diversity is located in New
<collectingCountry id="AC847611E501935D21FDFE477365FE78" box="[1001,1080,406,432]" name="Guinea-Bissau" pageId="9" pageNumber="314">Guinea</collectingCountry>
and certain representatives in the
<collectingCountry id="AC847611E501935D2188FE64730BFE07" box="[924,1110,437,463]" name="Solomon Islands" pageId="9" pageNumber="314">Solomon Islands</collectingCountry>
(
<bibRefCitation id="B0024B70E501935D267DFE6475BEFE27" author="Gebien" firstAuthor="Gebien" pageId="9" pageNumber="314" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien 1920</bibRefCitation>
). Species of
<taxonomicName id="13934D02E501935D2177FE077480FE38" authorityName="Dalman" authorityYear="1823" box="[867,989,470,496]" class="Insecta" family="Tenebrionidae" genus="Amarygmus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2177FE077480FE38" box="[867,989,470,496]" italics="true" pageId="9" pageNumber="314">Amarygmus</emphasis>
</taxonomicName>
are also known from the
<collectingCountry id="AC847611E501935D20DBFE247418FDC7" box="[719,837,501,527]" name="Fiji" pageId="9" pageNumber="314">Fiji Islands</collectingCountry>
(
<bibRefCitation id="B0024B70E501935D2141FE2474B8FDD8" author="Kaszab" box="[853,997,501,528]" firstAuthor="Kaszab" pageId="9" pageNumber="314" pagination="423 - 563" refId="ref7403" refString="KASZAB Z. 1955. - Tenebrioniden der Fiji-Inseln. Proceedings of the Hawaiian Entomological Society 15: 423 - 563." type="journal article" year="1955">Kaszab 1955</bibRefCitation>
), which attests of a colonisation front going towards southeast. In the same way, if
<taxonomicName id="13934D02E501935D2125FDE57425FD86" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[817,888,564,590]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2125FDE57425FD86" box="[817,888,564,590]" italics="true" pageId="9" pageNumber="314">Uloma</emphasis>
</taxonomicName>
of the area and the presently described
<collectingCountry id="AC847611E501935D211BFD857437FDA6" box="[783,874,596,622]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
species are examined, morphological affinities show that
<taxonomicName id="13934D02E501935D21ADFDA573E7FD46" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[953,1210,628,654]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E501935D21ADFDA57313FD45" box="[953,1102,628,653]" italics="true" pageId="9" pageNumber="314">U. vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E501935D20B1FD457583FD66" box="[677,734,660,686]" pageId="9" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
is closely related to a species group of
<taxonomicName id="13934D02E501935D2666FD4273E4FD65" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[1138,1209,659,685]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2666FD4273E4FD65" box="[1138,1209,659,685]" italics="true" pageId="9" pageNumber="314">Uloma</emphasis>
</taxonomicName>
from New
<collectingCountry id="AC847611E501935D2134FD62742DFD05" box="[800,880,691,717]" name="Guinea-Bissau" pageId="9" pageNumber="314">Guinea</collectingCountry>
(i.e. the
<taxonomicName id="13934D02E501935D21CCFD657334FD05" authorityName="Kirsch" authorityYear="1875" box="[984,1129,691,717]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="bituberosa">
<emphasis id="E6E7EA93E501935D21CCFD657334FD05" box="[984,1129,691,717]" italics="true" pageId="9" pageNumber="314">U. bituberosa</emphasis>
</taxonomicName>
group;
<bibRefCitation id="B0024B70E501935D20B1FD027465FD25" author="Gebien" box="[677,824,723,749]" firstAuthor="Gebien" pageId="9" pageNumber="314" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien 1920</bibRefCitation>
).
<collectingCountry id="AC847611E501935D2158FD0274A4FD25" box="[844,1017,723,749]" name="New Caledonia" pageId="9" pageNumber="314">New Caledonia</collectingCountry>
, the nearest insular territory to
<collectingCountry id="AC847611E501935D2156FD2274C0FCC5" box="[834,925,755,781]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
(
<quantity id="136B9B64E501935D21BFFD237359FCC5" box="[939,1028,754,781]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="314" unit="km" value="400.0">400 km</quantity>
), has the greatest
<taxonomicName id="13934D02E501935D20B1FCC375B1FCE4" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[677,748,786,812]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D20B1FCC375B1FCE4" box="[677,748,786,812]" italics="true" pageId="9" pageNumber="314">Uloma</emphasis>
</taxonomicName>
diversity with respect to its surface (
<bibRefCitation id="B0024B70E501935D2678FCC375B1FC84" author="Kaszab" firstAuthor="Kaszab" pageId="9" pageNumber="314" pagination="1 - 294" refId="ref7460" refString="KASZAB Z. 1982 b. - Die Tenebrioniden Neukaledoniens und der Loyaute-Inseln (Coleoptera), Folia Entomologica Hungarica 43 (2): 1 - 294." type="journal article" year="1982" yearSuffix="b">Kaszab 1982b</bibRefCitation>
,
<bibRefCitation id="B0024B70E501935D20E3FCE3746EFC85" author="Kaszab" box="[759,819,818,845]" firstAuthor="Kaszab" pageId="9" pageNumber="314" pagination="151 - 175" refId="ref7489" refString="KASZAB Z. 1986. - Tenebrioniden (Coleoptera) aus Neukaledonien. Annales Historico-Naturales Musei Nationalis Hungarici 78: 151 - 175." type="journal article" year="1986">1986</bibRefCitation>
). However, no New Caledonian species are morphologically related to
<taxonomicName id="13934D02E501935D2634FC837452FC43" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E501935D2634FC8373E7FCA3" box="[1056,1210,850,875]" italics="true" pageId="9" pageNumber="314">U. vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E501935D210CFCA37409FC44" box="[792,852,882,908]" pageId="9" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
On the contrary, the Fijian species,
<taxonomicName id="13934D02E501935D20CEFC407361FC63" authority="Fairmaire, 1849" authorityName="Fairmaire" authorityYear="1849" box="[730,1084,913,939]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="cavicollis">
<emphasis id="E6E7EA93E501935D20CEFC4074D9FC63" box="[730,900,913,939]" italics="true" pageId="9" pageNumber="314">Uloma cavicollis</emphasis>
<bibRefCitation id="B0024B70E501935D2198FC407361FC63" author="Fairmaire" box="[908,1084,913,939]" firstAuthor="Fairmaire" pageId="9" pageNumber="314" pagination="445 - 460" refId="ref7155" refString="FAIRMAIRE L. 1849. - Essai sur les coleopteres de la Polynesie (suite). Revue et Magasin de Zoologie pure et appliquee (2) 1: 445 - 460." type="journal article" year="1849">Fairmaire, 1849</bibRefCitation>
</taxonomicName>
, shows several morphological features bringing it closer to
<taxonomicName id="13934D02E501935D20B2FC0074EAFC22" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[678,951,976,1003]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E501935D20B2FC00741FFC22" box="[678,834,977,1002]" italics="true" pageId="9" pageNumber="314">U. vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E501935D21ABFC0074A1FC23" box="[959,1020,977,1003]" pageId="9" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
As in the case of
<taxonomicName id="13934D02E501935D20B2FC21747DFBC2" authorityName="Dalman" authorityYear="1823" box="[678,800,1008,1034]" class="Insecta" family="Tenebrionidae" genus="Amarygmus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D20B2FC21747DFBC2" box="[678,800,1008,1034]" italics="true" pageId="9" pageNumber="314">Amarygmus</emphasis>
</taxonomicName>
, it seems that the presence of
<taxonomicName id="13934D02E501935D2660FC2173E1FBC2" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[1140,1212,1008,1034]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2660FC2173E1FBC2" box="[1140,1212,1008,1034]" italics="true" pageId="9" pageNumber="314">Uloma</emphasis>
</taxonomicName>
in
<collectingCountry id="AC847611E501935D20D5FBC17441FBE2" box="[705,796,1040,1066]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
is the result of a southeastern oriented dispersal using
<collectingCountry id="AC847611E501935D2156FBFE74ADFB81" box="[834,1008,1071,1097]" name="Solomon Islands" pageId="9" pageNumber="314">Solomon Islands</collectingCountry>
as stepping-stones. It is interesting to note that this dispersal has not yet reach
<collectingCountry id="AC847611E501935D20F4FBBE74DAFB40" box="[736,903,1134,1161]" name="New Caledonia" pageId="9" pageNumber="314">New Caledonia</collectingCountry>
whose
<taxonomicName id="13934D02E501935D21C7FBBF7347FB40" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[979,1050,1134,1160]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D21C7FBBF7347FB40" box="[979,1050,1134,1160]" italics="true" pageId="9" pageNumber="314">Uloma</emphasis>
</taxonomicName>
representatives seem to originate from
<collectingCountry id="AC847611E501935D21BCFB5F7355FB60" box="[936,1032,1166,1192]" name="Australia" pageId="9" pageNumber="314">Australia</collectingCountry>
(
<bibRefCitation id="B0024B70E501935D260CFB5F740EFB01" author="Matthews and Bouchard" firstAuthor="Matthews" pageId="9" pageNumber="314" refId="ref7637" refString="MATTHEWS E. G. &amp; BOUCHARD P. 2005. - Tenebrionid Beetles of Australia: Description of Tribes, Keys to Genera, Catalogue of Species. ABRS, Canberra; Advance Press Pty Ltd, western Australia; 398 p." type="book" year="2005">Matthews and Bouchard 2005</bibRefCitation>
). Consequently, it appears clearly that New
<collectingCountry id="AC847611E501935D2105FB1C743DFB2F" box="[785,864,1229,1255]" name="Guinea-Bissau" pageId="9" pageNumber="314">Guinea</collectingCountry>
played a crucial role in the species assemblage of
<taxonomicName id="13934D02E501935D217EFB3C7358FACF" authorityName="Latreille" authorityYear="1802" box="[874,1029,1261,1287]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
in
<collectingCountry id="AC847611E501935D2633FB3F73DFFAC0" box="[1063,1154,1262,1288]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
via a dispersal that followed the Melanesian arch. The genera
<taxonomicName id="13934D02E501935D20E4FAFC7437FA8F" authorityName="Perroud" authorityYear="1864" box="[752,874,1325,1351]" class="Insecta" family="Tenebrionidae" genus="Bradymerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D20E4FAFC7437FA8F" box="[752,874,1325,1351]" italics="true" pageId="9" pageNumber="314">Bradymerus</emphasis>
</taxonomicName>
(
<taxonomicName id="13934D02E501935D216DFAFC74B2FA8F" authorityName="Gebien" authorityYear="1920" box="[889,1007,1325,1351]" class="Insecta" family="Tenebrionidae" genus="Bradymerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="E6E7EA93E501935D216DFAFC74B2FA8F" box="[889,1007,1325,1351]" italics="true" pageId="9" pageNumber="314">B. lobicollis</emphasis>
</taxonomicName>
),
<taxonomicName id="13934D02E501935D2611FAFC732CFA8F" authorityName="Laporte &amp; Brulle" authorityYear="1831" box="[1029,1137,1325,1351]" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2611FAFC732CFA8F" box="[1029,1137,1325,1351]" italics="true" pageId="9" pageNumber="314">Platydema</emphasis>
</taxonomicName>
(
<taxonomicName id="13934D02E501935D2694FAFC7475FAAE" authorityName="Gebien" authorityYear="1920" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="novaeguineense">
<emphasis id="E6E7EA93E501935D2694FAFC7475FAAE" italics="true" pageId="9" pageNumber="314">P. novaeguineense</emphasis>
</taxonomicName>
),
<taxonomicName id="13934D02E501935D212AFA9D74F9FAAE" authorityName="Pascoe" authorityYear="1869" box="[830,932,1356,1382]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D212AFA9D74F9FAAE" box="[830,932,1356,1382]" italics="true" pageId="9" pageNumber="314">Promethis</emphasis>
</taxonomicName>
(
<taxonomicName id="13934D02E501935D21A7FA9C737FFAAE" authorityName="Boisduval" authorityYear="1835" box="[947,1058,1356,1382]" class="Insecta" family="Tenebrionidae" genus="Upis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="sulcigera">
<emphasis id="E6E7EA93E501935D21A7FA9C737FFAAE" box="[947,1058,1356,1382]" italics="true" pageId="9" pageNumber="314">P. sulcigera</emphasis>
</taxonomicName>
) and
<taxonomicName id="13934D02E501935D2675FA9C73E7FAAE" authorityName="Latreille" authorityYear="1802" box="[1121,1210,1357,1382]" class="Insecta" family="Tenebrionidae" genus="Toxicum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2675FA9C73E7FAAE" box="[1121,1210,1357,1382]" italics="true" pageId="9" pageNumber="314">Toxicum</emphasis>
</taxonomicName>
(
<taxonomicName id="13934D02E501935D20B9FABC741DFA4E" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[685,832,1388,1414]" class="Insecta" family="Tenebrionidae" genus="Toxicum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="quadricorne">
<emphasis id="E6E7EA93E501935D20B9FABC741DFA4E" box="[685,832,1388,1414]" italics="true" pageId="9" pageNumber="314">T. quadricorne</emphasis>
</taxonomicName>
) are most probably originating from New
<collectingCountry id="AC847611E501935D20CAFA5D746DFA6E" box="[734,816,1420,1446]" name="Guinea-Bissau" pageId="9" pageNumber="314">Guinea</collectingCountry>
, too.
</paragraph>
<paragraph id="D42C3681E501935E20AFFA7A769CFEB9" blockId="9.[674,1212,216,1700]" lastBlockId="10.[111,649,216,1700]" lastPageId="10" lastPageNumber="315" pageId="9" pageNumber="314">
Then, other genera can come from the
<collectingCountry id="AC847611E501935D2650FA7A73EBFA0D" box="[1092,1206,1451,1477]" name="Fiji" pageId="9" pageNumber="314">Fiji Islands</collectingCountry>
, a wide archipelago including approximately 80 species of
<taxonomicName id="13934D02E501935D20FEFA3A737AF9CD" authority="(Kaszab 1955)" baseAuthorityName="Kaszab" baseAuthorityYear="1955" box="[746,1063,1514,1541]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="family">
Tenebrionidae (
<bibRefCitation id="B0024B70E501935D2185FA3A7342F9CD" author="Kaszab" box="[913,1055,1514,1541]" firstAuthor="Kaszab" pageId="9" pageNumber="314" pagination="423 - 563" refId="ref7403" refString="KASZAB Z. 1955. - Tenebrioniden der Fiji-Inseln. Proceedings of the Hawaiian Entomological Society 15: 423 - 563." type="journal article" year="1955">Kaszab 1955</bibRefCitation>
)
</taxonomicName>
. Its proximity to
<collectingCountry id="AC847611E501935D20A9F9DA7448F9ED" box="[701,789,1547,1573]" name="Vanuatu" pageId="9" pageNumber="314">Vanuatu</collectingCountry>
suggests that faunal exchange could have been frequent and that resulted in several genera occurring in both archipelagoes. For example, a Melanesian connection was probably established concerning the genus
<taxonomicName id="13934D02E501935D2199F9587357F96B" authorityName="Pascoe" authorityYear="1860" box="[909,1034,1673,1699]" class="Insecta" family="Tenebrionidae" genus="Chariotheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E501935D2199F9587357F96B" box="[909,1034,1673,1699]" italics="true" pageId="9" pageNumber="314">Chariotheca</emphasis>
</taxonomicName>
with two species (
<taxonomicName id="13934D02E502935E226EFF097784FF3A" authorityName="Kaszab" authorityYear="1955" box="[122,217,216,242]" class="Insecta" family="Tenebrionidae" genus="Chariotheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="E6E7EA93E502935E226EFF097784FF3A" box="[122,217,216,242]" italics="true" pageId="10" pageNumber="315">C. striata</emphasis>
</taxonomicName>
and
<taxonomicName id="13934D02E502935E2318FF097624FF3A" authorityName="Fairmaire" authorityYear="1849" box="[268,377,216,242]" class="Insecta" family="Tenebrionidae" genus="Olisthaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="species" species="cuprina">
<emphasis id="E6E7EA93E502935E2318FF097624FF3A" box="[268,377,216,242]" italics="true" pageId="10" pageNumber="315">C. cuprina</emphasis>
</taxonomicName>
) occuring both in the
<collectingCountry id="AC847611E502935E2077FF0977E7FED9" name="Fiji" pageId="10" pageNumber="315">Fiji Islands</collectingCountry>
and in
<collectingCountry id="AC847611E502935E2304FF297632FEDA" box="[272,367,248,274]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
. These dispersals pursued westward to reach
<collectingCountry id="AC847611E502935E2321FEC97686FEF9" box="[309,475,279,306]" name="New Caledonia" pageId="10" pageNumber="315">New Caledonia</collectingCountry>
(
<bibRefCitation id="B0024B70E502935E23F3FEC675DFFEF9" author="Kaszab" box="[487,642,279,305]" firstAuthor="Kaszab" pageId="10" pageNumber="315" pagination="1 - 294" refId="ref7460" refString="KASZAB Z. 1982 b. - Die Tenebrioniden Neukaledoniens und der Loyaute-Inseln (Coleoptera), Folia Entomologica Hungarica 43 (2): 1 - 294." type="journal article" year="1982" yearSuffix="b">Kaszab 1982b</bibRefCitation>
,
<bibRefCitation id="B0024B70E502935E2264FEE677F6FE9A" author="Kaszab" box="[112,171,311,338]" firstAuthor="Kaszab" pageId="10" pageNumber="315" pagination="151 - 175" refId="ref7489" refString="KASZAB Z. 1986. - Tenebrioniden (Coleoptera) aus Neukaledonien. Annales Historico-Naturales Musei Nationalis Hungarici 78: 151 - 175." type="journal article" year="1986">1986</bibRefCitation>
), but also northward to the
<collectingCountry id="AC847611E502935E23D9FEE675DCFE99" box="[461,641,311,337]" name="Solomon Islands" pageId="10" pageNumber="315">Solomon Islands</collectingCountry>
, where the genus is also present.
</paragraph>
<paragraph id="D42C3681E502935E229CFEA7761BFA4F" blockId="10.[111,649,216,1700]" pageId="10" pageNumber="315">
Lastly,
<collectingCountry id="AC847611E502935E22DBFEA6762FFE58" box="[207,370,374,401]" name="New Caledonia" pageId="10" pageNumber="315">New Caledonia</collectingCountry>
could have also influenced the fauna of
<collectingCountry id="AC847611E502935E22E7FE47760FFE78" box="[243,338,406,432]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
. Emerged approximately 34- 37 million years ago (
<bibRefCitation id="B0024B70E502935E2348FE6476A5FE18" author="Pelletier" box="[348,504,437,464]" firstAuthor="Pelletier" pageId="10" pageNumber="315" pagination="17 - 30" refId="ref7763" refString="PELLETIER B. 2006. - Geology of the New Caledonia region and its implications for the study of the New Caledonian biodiversity, in PAYRI C. E., RICHER DE FORGES B. (eds), Compendium of Marine Species from New Caledonia Documents Scientifiques et Techniques, II 4. Institut de Recherche pour le Developpement, Noumea: 17 - 30." type="book chapter" year="2006">Pelletier 2006</bibRefCitation>
), it has some genera in common with the
<collectingCountry id="AC847611E502935E23B7FE0776A3FE38" box="[419,510,470,496]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
archipelago. Even though
<collectingCountry id="AC847611E502935E2314FE2476FAFDC7" box="[256,423,501,527]" name="New Caledonia" pageId="10" pageNumber="315">New Caledonia</collectingCountry>
and
<collectingCountry id="AC847611E502935E23CFFE24756BFDC7" box="[475,566,501,527]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
exhibit many affinities for the plants for instance (
<bibRefCitation id="B0024B70E502935E2050FDC577ECFD86" author="Pillon" firstAuthor="Pillon" pageId="10" pageNumber="315" pagination="93 - 94" refId="ref7831" refString="PILLON Y. 2011. - Geissois (Cunoniaceae), another example of the Melanesian connection, in BOUCHET P., LE GUYADER H. &amp; PASCAL O. (eds), The Natural History of Santo. Patrimoines Naturels 70. Museum national d'Histoire naturelle, Paris; IRD, Marseille; Pro-Natura International, Paris: 93 - 94." type="book chapter" year="2011">Pillon 2011</bibRefCitation>
), most tenebrionid species from the two areas are different (
<bibRefCitation id="B0024B70E502935E235CFD857686FDA7" author="Kaszab" box="[328,475,596,623]" firstAuthor="Kaszab" pageId="10" pageNumber="315" pagination="423 - 563" refId="ref7403" refString="KASZAB Z. 1955. - Tenebrioniden der Fiji-Inseln. Proceedings of the Hawaiian Entomological Society 15: 423 - 563." type="journal article" year="1955">Kaszab 1955</bibRefCitation>
,
<bibRefCitation id="B0024B70E502935E23FDFD85756EFDA6" author="Kaszab" box="[489,563,596,622]" firstAuthor="Kaszab" pageId="10" pageNumber="315" pagination="1 - 294" refId="ref7460" refString="KASZAB Z. 1982 b. - Die Tenebrioniden Neukaledoniens und der Loyaute-Inseln (Coleoptera), Folia Entomologica Hungarica 43 (2): 1 - 294." type="journal article" year="1982" yearSuffix="b">1982b</bibRefCitation>
,
<bibRefCitation id="B0024B70E502935E2054FD857520FDA7" author="Kaszab" box="[576,637,596,623]" firstAuthor="Kaszab" pageId="10" pageNumber="315" pagination="151 - 175" refId="ref7489" refString="KASZAB Z. 1986. - Tenebrioniden (Coleoptera) aus Neukaledonien. Annales Historico-Naturales Musei Nationalis Hungarici 78: 151 - 175." type="journal article" year="1986">1986</bibRefCitation>
). Moreover, several widespread Australasian genera (
<taxonomicName id="13934D02E502935E226EFD4577A9FD66" authorityName="Dalman" authorityYear="1823" box="[122,244,660,686]" class="Insecta" family="Tenebrionidae" genus="Amarygmus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E502935E226EFD4577A9FD66" box="[122,244,660,686]" italics="true" pageId="10" pageNumber="315">Amarygmus</emphasis>
</taxonomicName>
,
<taxonomicName id="13934D02E502935E2315FD42763AFD65" authorityName="Pascoe" authorityYear="1869" box="[257,359,659,685]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E502935E2315FD42763AFD65" box="[257,359,659,685]" italics="true" pageId="10" pageNumber="315">Promethis</emphasis>
</taxonomicName>
and
<taxonomicName id="13934D02E502935E2388FD4576A8FD65" authorityName="Latreille" authorityYear="1802" box="[412,501,660,685]" class="Insecta" family="Tenebrionidae" genus="Toxicum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E502935E2388FD4576A8FD65" box="[412,501,660,685]" italics="true" pageId="10" pageNumber="315">Toxicum</emphasis>
</taxonomicName>
) are found in
<collectingCountry id="AC847611E502935E2265FD657791FD06" box="[113,204,692,718]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
but not in
<collectingCountry id="AC847611E502935E2353FD6576B3FD05" box="[327,494,691,718]" name="New Caledonia" pageId="10" pageNumber="315">New Caledonia</collectingCountry>
(
<bibRefCitation id="B0024B70E502935E23E8FD627646FD25" author="Matthews &amp; Bouchard" firstAuthor="Matthews" pageId="10" pageNumber="315" refId="ref7637" refString="MATTHEWS E. G. &amp; BOUCHARD P. 2005. - Tenebrionid Beetles of Australia: Description of Tribes, Keys to Genera, Catalogue of Species. ABRS, Canberra; Advance Press Pty Ltd, western Australia; 398 p." type="book" year="2005">Matthews &amp; Bouchard 2005</bibRefCitation>
). In the same way, many New Caledonian genera do not occur in
<collectingCountry id="AC847611E502935E23D0FD22757EFCC5" box="[452,547,755,781]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
. This low level of affinity demonstrates the scarcity of the past exchanges between these two areas in spite of their proximity. This can be explained by two major hypotheses:First, one can imagine that competition or poor adaptation prevented some tenebrionid genera from colonising one or the other archipelago.
<collectingCountry id="AC847611E502935E2045FC6077BFFC22" name="New Caledonia" pageId="10" pageNumber="315">New Caledonia</collectingCountry>
, for example, is covered by ultramafic soils that have strongly constrained the diversification of plants (
<bibRefCitation id="B0024B70E502935E231EFBDE769EFBE2" author="Pillon" box="[266,451,1039,1066]" etAl="et al." firstAuthor="Pillon" pageId="10" pageNumber="315" pagination="1108 - 1116" refId="ref7896" refString="PILLON Y., MUNZINGER J., HAMIR H. &amp; LEBRUN M. 2010. - Ultramafic soils and species sorting in the flora of New Caledonia. Journal of Ecology 98: 1108 - 1116." type="journal article" year="2010">
Pillon
<emphasis id="E6E7EA93E502935E2345FBC176DEFBE1" box="[337,387,1039,1065]" italics="true" pageId="10" pageNumber="315">et al.</emphasis>
2010
</bibRefCitation>
). Second, it can be thought that dispersal opportunities between the two archipelagoes were limited for poor dispersers (wingless species), or that abiotic (e.g., marine currents) or biotic factors prevented these dispersal events. Testing these assumptions would require a more thorough knowledge of the biodiversity of the region, as well as an estimate of their phylogenetic relationships among species existing between the distinct areas through a historical biogeography approach (interspecific level) or phylogeography (intraspecific level).
</paragraph>
<paragraph id="D42C3681E502935E229CFA5D74B0FDC7" blockId="10.[111,649,216,1700]" lastBlockId="10.[674,1212,216,527]" pageId="10" pageNumber="315">
In conclusion, the tenebrionid fauna of
<collectingCountry id="AC847611E502935E2038FA5D75DAFA6E" box="[556,647,1420,1446]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
represents a reserve of species, which provides an interesting and still incompletely known biogeographic history with questions about its origin and evolution. Thanks to this first inventory, we highlight multiple and independent origins to explain the colonisation of the archipelago. However, understanding of the evolutionary and ecological processes, which have contributed to the community assemblage, remains a fascinating challenge that requires a more thorough sampling over the entire Australasian region. Even if it appears that New
<collectingCountry id="AC847611E502935E2183FEC674BBFEF9" box="[919,998,279,305]" name="Guinea-Bissau" pageId="10" pageNumber="315">Guinea</collectingCountry>
is the major source of this biodiversity, it would be very interesting to test these biogeographic hypotheses within a phylogenetic perspective including dense taxon sampling for each genus. Phylogenetic and dating analyses would then provide additional evidence for better apprehending when and how appeared the
<taxonomicName id="13934D02E502935E20D8FE24743AFDC7" baseAuthorityName="Kaszab" baseAuthorityYear="1955" box="[716,871,501,527]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
of
<collectingCountry id="AC847611E502935E219EFE2474B4FDC7" box="[906,1001,501,527]" name="Vanuatu" pageId="10" pageNumber="315">Vanuatu</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,60 +1,60 @@
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@ -66,7 +66,7 @@
)
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<bibRefCitation id="B0024B70E50A93562604FC6073F9FC03" author="Bremer" box="[1040,1188,945,971]" firstAuthor="Bremer" pageId="2" pageNumber="307" pagination="509 - 528" refId="ref6921" refString="BREMER H. - J. 1993. - Neue Arten des Genus Corticeus Piller &amp; Mitterpacher, 1783, aus der papuanischmelanesischen Region, II. Mitteilung (Coleoptera, Tenebrionidae, Hypophloeini). Entomofauna 14: 509 - 528." type="journal article" year="1993">Bremer, 1993</bibRefCitation>
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@ -77,7 +77,7 @@
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<emphasis id="E6E7EA93E50A935620B1FC3B7475FBC9" box="[677,808,1001,1025]" italics="true" pageId="2" pageNumber="307">Corticeus levis</emphasis>
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, fig. 8.
@ -134,7 +134,7 @@ Espiritu Santo Isl., Luganville, CETRAV in Saraoutou, ecotone between dense rain
DISTRIBUTION. — So far, this species is only known from Espiritu Santo (
<collectingCountry id="AC847611E50A93562128FA6774CFFA06" box="[828,914,1462,1486]" name="Vanuatu" pageId="2" pageNumber="307">Vanuatu</collectingCountry>
) and is considered as endemic of this island (
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<bibRefCitation id="B0024B70E50A9356213BFA0174ECFA20" author="Soldati" box="[815,945,1488,1512]" firstAuthor="Soldati" pageId="2" pageNumber="307" pagination="500 - 503" refId="ref8042" refString="SOLDATI L. 2011. - Endemic, native, alien or cryptogenic? The controversy of Espiritu Santo darkling beetles (Insecta: Coleoptera: Tenebrionidae), in BOUCHET P., LE GUYADER H. &amp; PASCAL O. (eds), The Natural History of Santo. Patrimoines Naturels 70. Museum national d'Histoire naturelle, Paris; IRD, Marseille; Pro-Natura International, Paris: 500 - 503." type="book chapter" year="2011">Soldati 2011</bibRefCitation>
).
</paragraph>
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@ -151,13 +151,5 @@ DISTRIBUTION. — So far, this species is only known from Espiritu Santo (
has never been mentioned since the original description until one of the authors discovered it once again during the SANTO 2006 expedition.
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Subfamily
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Tribe
<taxonomicName id="13934D02E50B93572304FF28757AFEDA" authority="Gistel, 1848" authorityName="Gistel" authorityYear="1848" box="[272,551,247,275]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="308" phylum="Arthropoda" rank="tribe" tribe="Amarygmini">AMARYGMINI Gistel, 1848</taxonomicName>
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.
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EXAMINED. —
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<collectingCountry id="AC847611E50A935621B4FD4C74ABFD7D" box="[928,1014,669,693]" name="Vanuatu" pageId="2" pageNumber="307">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Luganville, CETRAV in Saraoutou, ecotone between dense rainforest and coffee tree plantation, under bark,
<date id="A02D1041E50A935620B1FD3D7479FCCC" box="[677,804,748,772]" pageId="2" pageNumber="307" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">2-10.X.2006</date>
, leg.
<collectionCode id="B282AE44E50A93562143FD3C7435FCCC" box="[855,872,749,772]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="307" type="Herbarium">L</collectionCode>
. Soldati,
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(
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).
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DISTRIBUTION. — Oriental region, Papuan subregion, northern
<collectingCountry id="AC847611E50A93562114FCE7740BFC86" box="[768,854,822,846]" name="Australia" pageId="2" pageNumber="307">Australia</collectingCountry>
and
<collectingCountry id="AC847611E50A93562190FCE77342FC86" box="[900,1055,822,846]" name="Comoros" pageId="2" pageNumber="307">Comoro Islands</collectingCountry>
(
<bibRefCitation id="B0024B70E50A9356263EFCE773F3FC86" author="Bremer" box="[1066,1198,822,846]" firstAuthor="Bremer" pageId="2" pageNumber="307" pagination="31 - 92" refId="ref6964" refString="BREMER H. - J. 1999. - Revision der orientalischen Corticeus - Arten (Col., Tenebrionidae, Hypophloeini). II. Teil. Acta Coleopterologica 15: 31 - 92." type="journal article" year="1999">Bremer 1999</bibRefCitation>
).
<collectingCountry id="AC847611E50A935620B1FC80741BFCA1" box="[677,838,849,873]" name="Solomon Islands" pageId="2" pageNumber="307">Solomon Islands</collectingCountry>
(Vanikoro). First record for
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.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,121 +0,0 @@
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<emphasis id="E6E7EA93E50A9356228AFA3B76E0F9CD" bold="true" box="[158,445,1514,1541]" italics="true" pageId="2" pageNumber="307">Platydema novaeguineense</emphasis>
<bibRefCitation id="B0024B70E50A935623D1FA3A7504F9CD" author="GEBIEN H." box="[453,601,1515,1541]" pageId="2" pageNumber="307" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien, 1920</bibRefCitation>
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MATERIAL EXAMINED. —
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.
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Espiritu Santo Isl., Luganville, CETRAV in Saraoutou, ecotone between dense rainforest and coffee tree plantation, in fungi growing on a rotten log,
<date id="A02D1041E50A9356218AFF067340FF27" box="[926,1053,215,239]" pageId="2" pageNumber="307" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">2-10.X.2006</date>
, leg. L. Soldati,
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DISTRIBUTION. — Described from New
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(
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locality: Doré), this species is also present in the
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(
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). First record for
<collectingCountry id="AC847611E50A93562656FE8773C5FEA5" box="[1090,1176,342,365]" name="Vanuatu" pageId="2" pageNumber="307">Vanuatu</collectingCountry>
.
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Tribe
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<emphasis id="E6E7EA93E50A9356228AFA3B76E0F9CD" bold="true" box="[158,445,1514,1541]" italics="true" pageId="2" pageNumber="307">Platydema novaeguineense</emphasis>
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<emphasis id="E6E7EA93E50A93562265F9F4763AF9F5" box="[113,359,1573,1597]" italics="true" pageId="2" pageNumber="307">Platydema novaeguineense</emphasis>
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MATERIAL EXAMINED. —
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<collectingCountry id="AC847611E50A93562379F987769EF9A6" box="[365,451,1622,1646]" name="Vanuatu" pageId="2" pageNumber="307">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Luganville, CETRAV in Saraoutou, ecotone between dense rainforest and coffee tree plantation, in fungi growing on a rotten log,
<date id="A02D1041E50A9356218AFF067340FF27" box="[926,1053,215,239]" pageId="2" pageNumber="307" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">2-10.X.2006</date>
, leg. L. Soldati,
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(
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),
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(coll.
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).
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DISTRIBUTION. — Described from New
<collectingCountry id="AC847611E50A9356262CFEF073DDFEF1" box="[1080,1152,289,313]" name="Guinea-Bissau" pageId="2" pageNumber="307">Guinea</collectingCountry>
(
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locality: Doré), this species is also present in the
<collectingCountry id="AC847611E50A93562670FEEA75BBFEA6" name="Solomon Islands" pageId="2" pageNumber="307">Solomon Islands</collectingCountry>
(
<bibRefCitation id="B0024B70E50A935620E7FE8774C4FEA6" author="Schawaller" box="[755,921,342,366]" firstAuthor="Schawaller" pageId="2" pageNumber="307" pagination="413 - 429" refId="ref7935" refString="SCHAWALLER W. 2008. - The species of Platydema Laporte &amp; Brulle (Coleoptera: Tenebrionidae) from New Guinea and the Moluccan Islands, with description of 11 new species. Stuttgarter Beitrage zur Naturkunde, n. s. 1: 413 - 429." type="book chapter" year="2008">Schawaller 2008</bibRefCitation>
). First record for
<collectingCountry id="AC847611E50A93562656FE8773C5FEA5" box="[1090,1176,342,365]" name="Vanuatu" pageId="2" pageNumber="307">Vanuatu</collectingCountry>
.
</paragraph>
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View file

@ -0,0 +1,178 @@
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MATERIAL EXAMINED. —
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<collectingCountry id="AC847611E50B935721B4FDAC74ABFD5D" box="[928,1014,637,661]" name="Vanuatu" pageId="3" pageNumber="308">Vanuatu</collectingCountry>
.
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<location id="D14C605AE50B93572616FDAC73EBFD5D" LSID="urn:lsid:plazi:treatment:5C3A8797E50B935720CEFDC574EEFBCA:D14C605AE50B93572616FDAC73EBFD5D" box="[1026,1206,637,661]" country="Vanuatu" county="Saraoutou" municipality="Luganville" name="Espiritu Santo Isl." pageId="3" pageNumber="308">Espiritu Santo Isl.</location>
,
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, CETRAV in
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, ecotone between dense rainforest and coffee tree plantation, at night on dead trunks,
<date id="A02D1041E50B93572134FD1D74C0FD2C" box="[800,925,716,740]" pageId="3" pageNumber="308" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">
<collectingDate id="B069E9A9E50B93572134FD1D74C0FD2C" box="[800,925,716,740]" pageId="3" pageNumber="308" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">2-10.X.2006</collectingDate>
</date>
, leg.
<collectorName id="79665357E50B935721D8FD1C7376FD2D" box="[972,1067,717,741]" pageId="3" pageNumber="308">L. Soldati</collectorName>
,
<specimenCount id="C295FD08E50B93572626FD1C7304FD2C" box="[1074,1113,717,741]" pageId="3" pageNumber="308" type="male">1 ♂</specimenCount>
(coll.
<collectionCode id="B282AE44E50B93572685FD1C73F2FD2D" box="[1169,1199,717,741]" country="United Kingdom" name="Linnean Society of London" pageId="3" pageNumber="308">LS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="64FB3CDCE50B935720B7FD3674CEFC87" collectingDate="2006-10-01" collectingDateMax="2006-10-11" collectingDateMin="2006-10-01" collectionCode="MNHN, LS" collectorName="L. Soldati" country="Vanuatu" location="Luganville" municipality="Saraoutou" pageId="3" pageNumber="308" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<location id="D14C605AE50B935720B7FD367453FD37" LSID="urn:lsid:plazi:treatment:5C3A8797E50B935720CEFDC574EEFBCA:D14C605AE50B935720B7FD367453FD37" box="[675,782,743,767]" country="Vanuatu" municipality="Saraoutou" name="Luganville" pageId="3" pageNumber="308">Luganville</location>
, CETRAV in
<collectingMunicipality id="3448ACFBE50B935721B5FD367357FD37" box="[929,1034,743,767]" pageId="3" pageNumber="308">Saraoutou</collectingMunicipality>
,
<date id="A02D1041E50B93572602FD3673C4FD37" box="[1046,1177,743,767]" pageId="3" pageNumber="308" value="2006-10-01" valueMax="2006-10-11" valueMin="2006-10-01">
<collectingDate id="B069E9A9E50B93572602FD3673C4FD37" box="[1046,1177,743,767]" pageId="3" pageNumber="308" value="2006-10-01" valueMax="2006-10-11" valueMin="2006-10-01">1-11.X.2006</collectingDate>
</date>
, at night with a headlamp, on the decaying trunk of an old mango tree (
<taxonomicName id="13934D02E50B9357210AFCCD749DFCFC" box="[798,960,796,820]" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="308" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="E6E7EA93E50B9357210AFCCD749DFCFC" box="[798,960,796,820]" italics="true" pageId="3" pageNumber="308">Mangifera indica</emphasis>
</taxonomicName>
L.), leg.
<collectorName id="79665357E50B93572600FCCD7328FCFC" box="[1044,1141,796,820]" pageId="3" pageNumber="308">L. Soldati</collectorName>
,
<specimenCount id="C295FD08E50B9357266AFCCD73E4FCFB" box="[1150,1209,796,820]" pageId="3" pageNumber="308" type="male">2 ♂♂</specimenCount>
(
<collectionCode id="B282AE44E50B935720B8FCE6745DFC87" box="[684,768,823,847]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="308">MNHN</collectionCode>
),
<specimenCount id="C295FD08E50B93572106FCE67469FC86" box="[786,820,823,847]" pageId="3" pageNumber="308" type="female">1 ♀</specimenCount>
(coll.
<collectionCode id="B282AE44E50B9357217BFCE674D0FC87" box="[879,909,823,847]" country="United Kingdom" name="Linnean Society of London" pageId="3" pageNumber="308">LS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="64FB3CDCE50B93572188FCE67492FCA1" collectingDate="2006-10-18" collectingDateMax="2006-10-20" collectingDateMin="2006-10-18" collectionCode="MNHN, LS" collectorName="L. Soldati" country="Vanuatu" location="Luganville" municipality="Saraoutou" pageId="3" pageNumber="308" specimenCount="2" specimenCount-male="2">
same data, but
<date id="A02D1041E50B9357263EFCE773E8FC86" box="[1066,1205,822,846]" pageId="3" pageNumber="308" value="2006-10-18" valueMax="2006-10-20" valueMin="2006-10-18">
<collectingDate id="B069E9A9E50B9357263EFCE773E8FC86" box="[1066,1205,822,846]" pageId="3" pageNumber="308" value="2006-10-18" valueMax="2006-10-20" valueMin="2006-10-18">18-20.X.2006</collectingDate>
</date>
,
<specimenCount id="C295FD08E50B935720B7FC807591FCA0" box="[675,716,849,873]" pageId="3" pageNumber="308" type="male">1 ♂</specimenCount>
(
<collectionCode id="B282AE44E50B935720CDFC807470FCA1" box="[729,813,849,873]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="308">MNHN</collectionCode>
),
<specimenCount id="C295FD08E50B93572156FC807436FCA0" box="[834,875,849,873]" pageId="3" pageNumber="308" type="male">1 ♂</specimenCount>
(coll.
<collectionCode id="B282AE44E50B935721BEFC807495FCA1" box="[938,968,849,873]" country="United Kingdom" name="Linnean Society of London" pageId="3" pageNumber="308">LS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B935720B1FC5074EEFBCA" pageId="3" pageNumber="308" type="distribution">
<paragraph id="D42C3681E50B935720B1FC5074EEFBCA" blockId="3.[675,1210,896,1026]" pageId="3" pageNumber="308">
DISTRIBUTION. — Indomalayan realm: Indochina, Sunda Islands, Andaman Islands, Sulawesi;
<collectingCountry id="AC847611E50B93572610FC4A7329FC7B" box="[1028,1140,923,947]" name="Micronesia" pageId="3" pageNumber="308">Micronesia</collectingCountry>
:
<collectingCountry id="AC847611E50B93572669FC4A73E9FC7B" box="[1149,1204,923,947]" name="Palau" pageId="3" pageNumber="308">Palau</collectingCountry>
,
<collectingCountry id="AC847611E50B935720B0FC647588FC05" box="[676,725,949,973]" name="Micronesia" pageId="3" pageNumber="308">Truk</collectingCountry>
,
<collectingCountry id="AC847611E50B935720C9FC647426FC05" box="[733,891,949,973]" name="Marshall Islands" pageId="3" pageNumber="308">Marshall Islands</collectingCountry>
,
<collectingCountry id="AC847611E50B93572197FC677491FC05" box="[899,972,950,973]" name="Micronesia" pageId="3" pageNumber="308">Ponape</collectingCountry>
; Melanesia: New
<collectingCountry id="AC847611E50B93572666FC6473E4FC05" box="[1138,1209,949,973]" name="Guinea-Bissau" pageId="3" pageNumber="308">Guinea</collectingCountry>
(Irian Jaya and
<collectingCountry id="AC847611E50B93572129FC017359FC20" box="[829,1028,976,1000]" name="Papua New Guinea" pageId="3" pageNumber="308">Papua New Guinea</collectingCountry>
),
<collectingCountry id="AC847611E50B93572602FC0173E4FC20" box="[1046,1209,976,1000]" name="Solomon Islands" pageId="3" pageNumber="308">Solomon Islands</collectingCountry>
and
<collectingCountry id="AC847611E50B935720DBFC3A747CFBCA" box="[719,801,1003,1026]" name="Vanuatu" pageId="3" pageNumber="308">Vanuatu</collectingCountry>
(
<bibRefCitation id="B0024B70E50B9357213AFC3B74FAFBCA" author="Merkl" box="[814,935,1002,1026]" firstAuthor="Merkl" pageId="3" pageNumber="308" pagination="235 - 254" refId="ref7681" refString="MERKL O. 1989. - Melanesian representatives of Toxicum and Cryphaeus (Coleoptera, Tenebrionidae: Toxicini). Acta Zoologica Academiae Scientiarum Hungaricae 35 (3 - 4): 235 - 254." type="journal article" year="1989">Merkl 1989</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,111 @@
<document id="4098FD03DCFAECE502BC23885985D652" ID-DOI="10.5252/z2012n2a8" ID-ISSN="1638-9387" ID-Zenodo-Dep="5165431" IM.bibliography_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1628213050710" checkinUser="felipe" docAuthor="Soldati, Laurent, Kergoat, Gael J. &amp; Condamine, Fabien L." docDate="2012" docId="5C3A8797E50B935720F2FB7C74A8F9FA" docLanguage="en" docName="Zoosystema.34.2.305-317.pdf" docOrigin="Zoosystema 34 (2)" docSource="http://dx.doi.org/10.5252/z2012n2a8" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Tribolium castaneum" docType="treatment" docVersion="1" lastPageNumber="308" masterDocId="A003FFEFE50893542214FFD1775DFFC8" masterDocTitle="Preliminary report on the Tenebrionidae (Insecta, Coleoptera) collected during the SANTO 2006 expedition to Vanuatu, with description of a new species of the genus Uloma Dejean, 1821" masterLastPageNumber="317" masterPageNumber="305" pageNumber="308" updateTime="1740575614240" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F2C104679D3631C7A6586917B79196D8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5AE04346549A98153A154DBEFEC62BFF">Preliminary report on the Tenebrionidae (Insecta, Coleoptera) collected during the SANTO 2006 expedition to Vanuatu, with description of a new species of the genus Uloma Dejean, 1821</mods:title>
</mods:titleInfo>
<mods:name id="A065C7955FBDF42DA286A1527934DBDB" type="personal">
<mods:role id="224850F865DC3453C8DB42139E5EF4CA">
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<mods:namePart id="4AD24D3C77D11336AE409A9517440F6C">Soldati, Laurent</mods:namePart>
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<mods:namePart id="49A7CF084A451DC5FEB6A329FD866B58">Kergoat, Gael J.</mods:namePart>
<mods:affiliation id="C4BE5D5279F5B9AC8526BA7A2FAB6163">INRA - UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier Supagro), Campus international de Baillarguet, CS 30016, F- 34988 Montferrier-sur-Lez cedex (France) soldati @ supagro. inra. fr kergoat @ supagro. inra. fr</mods:affiliation>
<mods:nameIdentifier id="589DCE77D30191A9F727259AE8DB45DD" type="email">fr@supagro.inra.fr</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="7B5069843147A57E94A6EF9974D0A496">Condamine, Fabien L.</mods:namePart>
<mods:affiliation id="68D8DCF4858791C5CE8C05E2D9FB8622">Mathématiques appliquées - UMR 7641 (CNRS, École polytechnique), route de Saclay, F- 91128 Palaiseau cedex (France) fabien. condamine @ gmail. com</mods:affiliation>
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<mods:title id="6EA89A9E9EC786520A5779C84BF6B92A">Zoosystema</mods:title>
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<mods:date id="8E84286361181870D082F0FF6E7BAF42">2012</mods:date>
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<treatment id="5C3A8797E50B935720F2FB7C74A8F9FA" LSID="urn:lsid:plazi:treatment:5C3A8797E50B935720F2FB7C74A8F9FA" httpUri="http://treatment.plazi.org/id/5C3A8797E50B935720F2FB7C74A8F9FA" lastPageNumber="308" pageId="3" pageNumber="308">
<subSubSection id="9C89650AE50B935720F2FB7C7324FB00" box="[742,1145,1197,1224]" pageId="3" pageNumber="308" type="nomenclature">
<paragraph id="D42C3681E50B935720F2FB7C7324FB00" blockId="3.[742,1145,1197,1224]" box="[742,1145,1197,1224]" pageId="3" pageNumber="308">
<heading id="8F6481EDE50B935720F2FB7C7324FB00" box="[742,1145,1197,1224]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="308" reason="2">
<taxonomicName id="13934D02E50B935720F2FB7C7324FB00" authority="(Herbst, 1797)" baseAuthorityName="Herbst" baseAuthorityYear="1797" box="[742,1145,1197,1224]" class="Insecta" family="Tenebrionidae" genus="Tribolium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="308" phylum="Arthropoda" rank="species" species="castaneum">
<emphasis id="E6E7EA93E50B935720F2FB7C748DFB00" bold="true" box="[742,976,1197,1224]" italics="true" pageId="3" pageNumber="308">Tribolium castaneum</emphasis>
(
<bibRefCitation id="B0024B70E50B935721CAFB7F732CFB00" author="Herbst" box="[990,1137,1198,1224]" firstAuthor="Herbst" pageId="3" pageNumber="308" refId="ref7341" refString="HERBST J. F. W. 1797. - Natursystem aller bekannten in- und auslandischen Insekten, als eine Forsetzung der von Buffonschen Naturgeschichte. Der Kafer siebenter Theil. Geh. Commerzien-Raths Pauli, Berlin, 346 p." type="book" year="1797">Herbst, 1797</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B935720B1FB367376FB37" box="[677,1067,1255,1279]" pageId="3" pageNumber="308" type="reference_group">
<paragraph id="D42C3681E50B935720B1FB367376FB37" blockId="3.[677,1067,1255,1279]" box="[677,1067,1255,1279]" pageId="3" pageNumber="308">
<treatmentCitationGroup id="F48311AFE50B935720B1FB367376FB37" box="[677,1067,1255,1279]" pageId="3" pageNumber="308">
<treatmentCitation id="55321090E50B935720B1FB36737AFB37" author="HERBST J. F. W." box="[677,1063,1255,1279]" page="282" pageId="3" pageNumber="308" year="1797">
<taxonomicName id="13934D02E50B935720B1FB36737AFB37" authority="Herbst, 1797: 282" authorityName="Herbst" authorityPageNumber="282" authorityYear="1797" box="[677,1063,1255,1279]" class="Insecta" family="Zopheridae" genus="Colydium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="308" phylum="Arthropoda" rank="species" species="castaneum">
<emphasis id="E6E7EA93E50B935720B1FB367435FB37" box="[677,872,1255,1279]" italics="true" pageId="3" pageNumber="308">Colydium castaneum</emphasis>
<bibRefCitation id="B0024B70E50B9357217BFB36737AFB37" author="Herbst" box="[879,1063,1255,1279]" firstAuthor="Herbst" page="282" pageId="3" pageNumber="308" refId="ref7341" refString="HERBST J. F. W. 1797. - Natursystem aller bekannten in- und auslandischen Insekten, als eine Forsetzung der von Buffonschen Naturgeschichte. Der Kafer siebenter Theil. Geh. Commerzien-Raths Pauli, Berlin, 346 p." type="book" year="1797">Herbst, 1797: 282</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B935720B1FAC67351FAAC" pageId="3" pageNumber="308" type="materials_examined">
<paragraph id="D42C3681E50B935720B1FAC67351FAAC" blockId="3.[677,1210,1302,1380]" pageId="3" pageNumber="308">
MATERIAL EXAMINED. —
<emphasis id="E6E7EA93E50B935721B4FAC674A7FAE7" bold="true" box="[928,1018,1303,1327]" pageId="3" pageNumber="308">
<collectingCountry id="AC847611E50B935721B4FAC674ABFAE7" box="[928,1014,1303,1327]" name="Vanuatu" pageId="3" pageNumber="308">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Luganville, CETRAV in Saraoutou,
<date id="A02D1041E50B9357261BFAE073C1FA81" box="[1039,1180,1329,1353]" pageId="3" pageNumber="308" value="2006-10-23" valueMax="2006-10-28" valueMin="2006-10-23">23-28.X.2006</date>
, at light, leg. L. Soldati,
<specimenCount id="C295FD08E50B93572167FA9D74C1FAAB" box="[883,924,1356,1380]" pageId="3" pageNumber="308" type="male">1 ♂</specimenCount>
(
<collectionCode id="B282AE44E50B935721BEFA9D74A3FAAC" box="[938,1022,1356,1380]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="308">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B935720B1FAAD74A8F9FA" pageId="3" pageNumber="308" type="distribution">
<paragraph id="D42C3681E50B935720B1FAAD74A8F9FA" blockId="3.[675,1210,1403,1586]" pageId="3" pageNumber="308">
DISTRIBUTION. —
<taxonomicName id="13934D02E50B93572141FAAA7349FA5B" baseAuthorityName="Herbst" baseAuthorityYear="1797" box="[853,1044,1403,1427]" class="Insecta" family="Tenebrionidae" genus="Tribolium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="308" phylum="Arthropoda" rank="species" species="castaneum">
<emphasis id="E6E7EA93E50B93572141FAAA7349FA5B" box="[853,1044,1403,1427]" italics="true" pageId="3" pageNumber="308">Tribolium castaneum</emphasis>
</taxonomicName>
is a cosmopolitan species and one of the most important stored product pests around the world. It is known to infest the following commodities: barley, bran, cacao, ginger, maize, millet, manioc, nutmeg, peanut, pepper, rice, sorghum, tapioca and yam (
<bibRefCitation id="B0024B70E50B93572111FA2E7482F9DF" author="Delobel &amp; Tran" box="[773,991,1535,1559]" firstAuthor="Delobel" pageId="3" pageNumber="308" refId="ref7000" refString="DELOBEL A. &amp; TRAN M. 1993. - Les Coleopteres des denrees alimentaires entreposees dans les regions chaudes. Faune tropicale XXXII. ORSTOM / CTA Editions, Paris, 424 p." type="book" year="1993">Delobel &amp; Tran 1993</bibRefCitation>
). This species can also damage entomological collections.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,123 +0,0 @@
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, note 1.
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MATERIAL EXAMINED. —
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Espiritu Santo Isl., Butmas, Melanesian village in rainforest, UV light trap,
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, leg. L. Soldati,
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DISTRIBUTION. — Described from Lifu Island, it occurs in the Loyalty Islands and also in the
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Tribe
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<paragraph id="D42C3681E50B93572265F993753AF992" blockId="3.[113,615,1602,1626]" box="[113,615,1602,1626]" pageId="3" pageNumber="308">
<treatmentCitationGroup id="F48311AFE50B93572265F993753AF992" box="[113,615,1602,1626]" pageId="3" pageNumber="308">
<treatmentCitation id="55321090E50B93572265F9937545F992" author="FAUVEL A." box="[113,536,1602,1626]" page="188" pageId="3" pageNumber="308" year="1867">
<taxonomicName id="13934D02E50B93572265F9937545F992" authority="Fauvel, 1867: 188" authorityName="Fauvel" authorityPageNumber="188" authorityYear="1867" box="[113,536,1602,1626]" class="Insecta" family="Tenebrionidae" genus="Gonocephalum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="308" phylum="Arthropoda" rank="species" species="irroratum">
<emphasis id="E6E7EA93E50B93572265F993763DF992" box="[113,352,1602,1626]" italics="true" pageId="3" pageNumber="308">Gonocephalum irroratum</emphasis>
<bibRefCitation id="B0024B70E50B93572372F9937545F992" author="Fauvel" box="[358,536,1602,1626]" firstAuthor="Fauvel" page="188" pageId="3" pageNumber="308" pagination="172 - 214" refId="ref7189" refString="FAUVEL A. 1867. - Catalogue des coleopteres de la Nouvelle-Caledonie et dependances avec descriptions, notes et synonymies nouvelles. Bulletin de la Societe linneenne de Normandie (2) 1: 172 - 214, pl. 1, figs 13 - 17." type="journal article" year="1867">Fauvel, 1867: 188</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, note 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B93572265F9A074D3FEC1" pageId="3" pageNumber="308" type="materials_examined">
<paragraph id="D42C3681E50B93572265F9A074D3FEC1" blockId="3.[113,647,1649,1699]" lastBlockId="3.[677,1209,215,265]" pageId="3" pageNumber="308">
MATERIAL EXAMINED. —
<emphasis id="E6E7EA93E50B93572379F9A0769AF941" bold="true" box="[365,455,1649,1673]" pageId="3" pageNumber="308">
<collectingCountry id="AC847611E50B93572379F9A0769EF941" box="[365,451,1649,1673]" name="Vanuatu" pageId="3" pageNumber="308">Vanuatu</collectingCountry>
.
</emphasis>
Espiritu Santo Isl., Butmas, Melanesian village in rainforest, UV light trap,
<date id="A02D1041E50B935720B1FF06744DFF27" box="[677,784,215,239]" pageId="3" pageNumber="308" value="2006-10-21">21.X.2006</date>
, leg. L. Soldati,
<specimenCount id="C295FD08E50B935721BAFF0674A5FF26" box="[942,1016,215,239]" pageId="3" pageNumber="308" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="C295FD08E50B93572611FF067314FF26" box="[1029,1097,215,239]" pageId="3" pageNumber="308" type="female">10 ♀♀</specimenCount>
(
<collectionCode id="B282AE44E50B93572643FF0673F6FF27" box="[1111,1195,215,239]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="308">MNHN</collectionCode>
),
<specimenCount id="C295FD08E50B935720B1FF2075BFFEC0" box="[677,738,241,265]" pageId="3" pageNumber="308" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="C295FD08E50B935720FAFF20747BFEC0" box="[750,806,241,265]" pageId="3" pageNumber="308" type="female">4 ♀♀</specimenCount>
(coll.
<collectionCode id="B282AE44E50B93572170FF2074DFFEC1" box="[868,898,241,265]" country="United Kingdom" name="Linnean Society of London" pageId="3" pageNumber="308">LS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50B935720B1FEF07419FEA6" pageId="3" pageNumber="308" type="distribution">
<paragraph id="D42C3681E50B935720B1FEF07419FEA6" blockId="3.[675,1210,289,366]" pageId="3" pageNumber="308">
DISTRIBUTION. — Described from Lifu Island, it occurs in the Loyalty Islands and also in the
<collectingCountry id="AC847611E50B935721E3FEED731AFE9B" box="[1015,1095,316,339]" name="Vanuatu" pageId="3" pageNumber="308">Vanuatu</collectingCountry>
archipelago (
<bibRefCitation id="B0024B70E50B935720BDFE877465FEA6" author="Kaszab" box="[681,824,342,366]" firstAuthor="Kaszab" pageId="3" pageNumber="308" pagination="1 - 294" refId="ref7460" refString="KASZAB Z. 1982 b. - Die Tenebrioniden Neukaledoniens und der Loyaute-Inseln (Coleoptera), Folia Entomologica Hungarica 43 (2): 1 - 294." type="journal article" year="1982" yearSuffix="b">Kaszab 1982b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,342 @@
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<mods:title id="C411BA3FFA46FC047216ECB181767CD1">Preliminary report on the Tenebrionidae (Insecta, Coleoptera) collected during the SANTO 2006 expedition to Vanuatu, with description of a new species of the genus Uloma Dejean, 1821</mods:title>
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<mods:name id="A7D681BAB191EAB337C5DFA9BCD5A6F9" type="personal">
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<mods:namePart id="4AB5EBC01E6D62E159F2A6348B69F4D2">Soldati, Laurent</mods:namePart>
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<mods:namePart id="2C4A0F5981BDBA34DEF7C84EC38369ED">Kergoat, Gael J.</mods:namePart>
<mods:affiliation id="049D2D46DA0DFECA292E05F6DD8374C1">INRA - UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier Supagro), Campus international de Baillarguet, CS 30016, F- 34988 Montferrier-sur-Lez cedex (France) soldati @ supagro. inra. fr kergoat @ supagro. inra. fr</mods:affiliation>
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<mods:namePart id="3D1CAC8B77457268A1426A848B353D83">Condamine, Fabien L.</mods:namePart>
<mods:affiliation id="6E20704272EF3515E74FE7E62B53D91F">Mathématiques appliquées - UMR 7641 (CNRS, École polytechnique), route de Saclay, F- 91128 Palaiseau cedex (France) fabien. condamine @ gmail. com</mods:affiliation>
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<treatment id="5C3A8797E50C935322A8FF0674F8FEF9" LSID="urn:lsid:plazi:treatment:5C3A8797E50C935322A8FF0674F8FEF9" httpUri="http://treatment.plazi.org/id/5C3A8797E50C935322A8FF0674F8FEF9" lastPageId="7" lastPageNumber="312" pageId="4" pageNumber="309">
<subSubSection id="9C89650AE50C935022A8FF067561FF3A" box="[188,572,215,242]" pageId="4" pageNumber="309" type="nomenclature">
<paragraph id="D42C3681E50C935022A8FF067561FF3A" blockId="4.[188,572,215,274]" box="[188,572,215,242]" pageId="4" pageNumber="309">
<taxonomicName id="13934D02E50C935022A8FF0676A7FF3A" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[188,506,215,242]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E50C935022A8FF0676D6FF3A" bold="true" box="[188,395,215,242]" italics="true" pageId="4" pageNumber="309">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50C93502015FF097561FF3A" box="[513,572,216,242]" pageId="4" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50C9350235FFF2676F1FEDA" box="[331,428,247,274]" pageId="4" pageNumber="309" type="description">
<paragraph id="D42C3681E50C9350235FFF2676F1FEDA" blockId="4.[188,572,215,274]" box="[331,428,247,274]" pageId="4" pageNumber="309">
(
<figureCitation id="4CA82A04E50C93502347FF2676F9FEDA" box="[339,420,247,274]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="5.[113,124,1635,1652]" captionStartId-1="6.[113,124,1657,1674]" captionStartId-2="8.[677,688,1634,1651]" captionTargetBox-0="[120,1199,273,1593]" captionTargetBox-1="[191,1132,229,1628]" captionTargetBox-2="[677,1209,216,1586]" captionTargetId-0="figure-77@5.[112,662,773,1334]" captionTargetId-1="figure-71@6.[191,1132,214,1628]" captionTargetId-2="figure-358@8.[677,1211,216,1587]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="FIG. 1. — Uloma vanuatensis L. Soldati n. sp.holotype ♂ (MNHN EC2280):A, habitus (dorsal view);B, habitus (lateral view); C, habitus (ventral view); D, forebody (lateral view); E, aedeagus (lateral view); F, aedeagus (tergal face). Scale bars: A-C, 10 mm; D-F, 1 mm. Photographs: L. Soldati." captionText-1="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." captionText-2="FIG. 3. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280):A, anterior tibia (upper face);B, metathorax and abdomen (ventralview).Scalebars:A,0,.5 mm;B,1 mm.Photographs:L.Soldati." figureDoi-0="http://doi.org/10.5281/zenodo.5165433" figureDoi-1="http://doi.org/10.5281/zenodo.5165435" figureDoi-2="http://doi.org/10.5281/zenodo.5165437" httpUri-0="https://zenodo.org/record/5165433/files/figure.png" httpUri-1="https://zenodo.org/record/5165435/files/figure.png" httpUri-2="https://zenodo.org/record/5165437/files/figure.png" pageId="4" pageNumber="309">Figs1-3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50C93502265FEE07548FD4E" pageId="4" pageNumber="309" type="materials_examined">
<paragraph id="D42C3681E50C93502265FEE0766BFE50" blockId="4.[111,649,304,646]" pageId="4" pageNumber="309">
<materialsCitation id="64FB3CDCE50C93502265FEE0766BFE50" collectingDate="2006-10-02" collectingDateMax="2006-10-10" collectingDateMin="2006-10-02" collectionCode="MNHN" collectorName="L. Soldati" country="Vanuatu" county="Saraoutou" latitude="-15.451388" location="Santo Isl." longLatPrecision="21" longitude="167.18445" municipality="Luganville" pageId="4" pageNumber="309" specimenCode="EC2280" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="0B288823E50C93502357FEE176F4FE80" box="[323,425,304,328]" pageId="4" pageNumber="309" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="AC847611E50C935023A1FEE07551FE80" box="[437,524,305,328]" name="Vanuatu" pageId="4" pageNumber="309">Vanuatu</collectingCountry>
,
<location id="D14C605AE50C93502008FEE075DFFE80" LSID="urn:lsid:plazi:treatment:5C3A8797E50C935322A8FF0674F8FEF9:D14C605AE50C93502008FEE075DFFE80" box="[540,642,304,329]" country="Vanuatu" county="Saraoutou" latitude="-15.451388" longLatPrecision="21" longitude="167.18445" municipality="Luganville" name="Santo Isl." pageId="4" pageNumber="309">Santo Isl.</location>
,
<collectingMunicipality id="3448ACFBE50C93502267FE9A7780FEAB" box="[115,221,331,355]" pageId="4" pageNumber="309">Luganville</collectingMunicipality>
, CETRAV in
<collectingCounty id="3D4D4E0DE50C9350237AFE9A7688FEAB" box="[366,469,331,355]" pageId="4" pageNumber="309">Saraoutou</collectingCounty>
,
<date id="A02D1041E50C935023F5FE9A753EFEAB" box="[481,611,331,355]" pageId="4" pageNumber="309" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">
<collectingDate id="B069E9A9E50C935023F5FE9A753EFEAB" box="[481,611,331,355]" pageId="4" pageNumber="309" value="2006-10-02" valueMax="2006-10-10" valueMin="2006-10-02">
2-10.X.
<quantity id="136B9B64E50C9350203EFE9A753EFEAB" box="[554,611,331,355]" metricMagnitude="3" metricUnit="m" metricValue="2.0063" pageId="4" pageNumber="309" unit="m" value="2006.3">2006</quantity>
</collectingDate>
</date>
, 30 m alt.,
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,
<geoCoordinate id="B1A75046E50C93502356FEB47695FEB5" box="[322,456,357,381]" degrees="167" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="309" precision="15" seconds="04" value="167.18445">167°1104”E</geoCoordinate>
, leg.
<collectorName id="79665357E50C93502015FEB7753AFEB5" box="[513,615,357,381]" pageId="4" pageNumber="309">L. Soldati</collectorName>
,
<specimenCount id="C295FD08E50C93502060FEB875D5FEB5" box="[628,648,361,381]" pageId="4" pageNumber="309" type="male"></specimenCount>
(
<collectionCode id="B282AE44E50C9350226DFE517790FE50" box="[121,205,384,408]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="309">MNHN</collectionCode>
<specimenCode id="84359EFAE50C935022C7FE517677FE50" box="[211,298,384,408]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="309">EC2280</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D42C3681E50C93502265FE4A7548FD4E" blockId="4.[111,649,304,646]" pageId="4" pageNumber="309">
<typeStatus id="0B288823E50C93502265FE4A7790FE7B" box="[113,205,411,435]" pageId="4" pageNumber="309" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="C295FD08E50C935022C9FE4B7648FE79" box="[221,277,410,434]" pageId="4" pageNumber="309" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="C295FD08E50C93502337FE4B7602FE79" box="[291,351,410,434]" pageId="4" pageNumber="309" type="male">3 ♂♂</specimenCount>
): same data as
<typeStatus id="0B288823E50C935023EEFE4B750FFE7A" box="[506,594,410,434]" pageId="4" pageNumber="309" type="holotype">holotype</typeStatus>
,
<specimenCount id="C295FD08E50C9350204AFE4B75DBFE79" box="[606,646,410,434]" pageId="4" pageNumber="309" type="female">1 ♀</specimenCount>
(
<collectionCode id="B282AE44E50C9350226CFE647794FE05" box="[120,201,437,461]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="309">MNHN</collectionCode>
EC2281). — Santo Isl., Saraoutou, littoral forest, rotten log,
<date id="A02D1041E50C935022C3FE1E760BFE2F" box="[215,342,463,487]" pageId="4" pageNumber="309" value="2006-10-09" valueMax="2006-10-10" valueMin="2006-10-09">9-10.X.2006</date>
,
<geoCoordinate id="B1A75046E50C9350234BFE1E768DFE2F" box="[351,464,463,487]" degrees="15" direction="south" minutes="26" orientation="latitude" pageId="4" pageNumber="309" precision="15" seconds="38" value="-15.443889">15°2638”S</geoCoordinate>
,
<geoCoordinate id="B1A75046E50C935023CDFE1E7507FE2F" box="[473,602,463,487]" degrees="167" direction="east" minutes="12" orientation="longitude" pageId="4" pageNumber="309" precision="15" seconds="52" value="167.21445">167°1252”E</geoCoordinate>
, leg. L. Soldati,
<specimenCount id="C295FD08E50C935022CFFE3B764EFDC9" box="[219,275,490,514]" pageId="4" pageNumber="309" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="B282AE44E50C93502335FE3B763AFDCA" box="[289,359,490,514]" pageId="4" pageNumber="309">CBGP</collectionCode>
),
<specimenCount id="C295FD08E50C9350236DFE3B76FFFDC9" box="[377,418,490,514]" pageId="4" pageNumber="309" type="male">1 ♂</specimenCount>
and
<specimenCount id="C295FD08E50C935023C6FE3B76A5FDC9" box="[466,504,490,514]" pageId="4" pageNumber="309" type="female">1 ♀</specimenCount>
(Coll.
<collectionCode id="B282AE44E50C9350202AFE3B7501FDCA" box="[574,604,490,514]" country="United Kingdom" name="Linnean Society of London" pageId="4" pageNumber="309">LS</collectionCode>
). — Espiritu Santo Isl., Big Bay, Matantas, Vatthe Area Conservation,
<date id="A02D1041E50C93502318FDCF76C2FDFE" box="[268,415,542,566]" pageId="4" pageNumber="309" value="2006-10-12" valueMax="2006-10-17" valueMin="2006-10-12">12-17.X.2006</date>
, leg. L. Soldati,
<specimenCount id="C295FD08E50C93502058FDCE75D4FDFE" box="[588,649,543,567]" pageId="4" pageNumber="309" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="B282AE44E50C9350226DFDE87790FD99" box="[121,205,569,593]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="309">MNHN</collectionCode>
EC2282- EC2283),
<specimenCount id="C295FD08E50C9350238BFDE87694FD98" box="[415,457,569,593]" pageId="4" pageNumber="309" type="male">1 ♂</specimenCount>
(coll.
<collectionCode id="B282AE44E50C9350201DFDE87578FD99" box="[521,549,569,593]" country="United Kingdom" name="Linnean Society of London" pageId="4" pageNumber="309">LS</collectionCode>
) and
<specimenCount id="C295FD08E50C9350204BFDE875DBFD98" box="[607,646,569,593]" pageId="4" pageNumber="309" type="female">1 ♀</specimenCount>
<typeStatus id="0B288823E50C93502265FD857799FDA4" box="[113,196,596,620]" pageId="4" pageNumber="309" type="paratype">paratype</typeStatus>
(
<collectionCode id="B282AE44E50C935022C0FD857646FDA4" box="[212,283,596,620]" pageId="4" pageNumber="309">CBGP</collectionCode>
). —
<taxonomicName id="13934D02E50C93502359FD8276D3FDA3" authority="Soldati &amp; Kergoat &amp; Condamine, 2012" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[333,398,595,619]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="E6E7EA93E50C93502359FD8276D3FDA3" box="[333,398,595,619]" italics="true" pageId="4" pageNumber="309">Uloma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50C93502381FD857691FDA3" box="[405,460,596,619]" pageId="4" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
aff.
<emphasis id="E6E7EA93E50C935023EDFD857516FDA4" box="[505,587,596,620]" italics="true" pageId="4" pageNumber="309">cavifrons</emphasis>
Kasz. Santo, Zoltán Kaszab det.,
<specimenCount id="C295FD08E50C9350236EFDBF76FEFD4D" box="[378,419,622,646]" pageId="4" pageNumber="309" type="male">1 ♂</specimenCount>
(
<collectionCode id="B282AE44E50C935023A6FDBF755AFD4D" box="[434,519,622,645]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="309">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50C93502265FD4F76E4FD22" pageId="4" pageNumber="309" type="distribution">
<paragraph id="D42C3681E50C93502265FD4F76E4FD22" blockId="4.[113,646,670,747]" pageId="4" pageNumber="309">
DISTRIBUTION. — This
<taxonomicName id="13934D02E50C9350234DFD4F76C7FD7E" authorityName="Soldati &amp; Kergoat &amp; Condamine" authorityYear="2012" box="[345,410,670,694]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="E6E7EA93E50C9350234DFD4F76C7FD7E" box="[345,410,670,694]" italics="true" pageId="4" pageNumber="309">Uloma</emphasis>
</taxonomicName>
is currently known only from the island of Espiritu Santo in the
<collectingCountry id="AC847611E50C935023E2FD687515FD18" box="[502,584,697,720]" name="Vanuatu" pageId="4" pageNumber="309">Vanuatu</collectingCountry>
archipelago, where it may be endemic.
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50C93502265FCD2762DFB39" pageId="4" pageNumber="309" type="etymology">
<paragraph id="D42C3681E50C93502265FCD276BEFCFD" blockId="4.[113,646,770,821]" pageId="4" pageNumber="309">
ETYMOLOGY. — This new species is named in reference to its origin, the
<collectingCountry id="AC847611E50C93502300FCCC763BFCFC" box="[276,358,797,820]" name="Vanuatu" pageId="4" pageNumber="309">Vanuatu</collectingCountry>
archipelago.
</paragraph>
<paragraph id="D42C3681E50C93502265FC9C762DFB39" blockId="4.[111,649,844,1265]" pageId="4" pageNumber="309">
DIAGNOSE. —
<taxonomicName id="13934D02E50C935022E8FC9D7558FCAC" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[252,517,844,868]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E50C935022E8FC9D76FAFCAC" box="[252,423,844,868]" italics="true" pageId="4" pageNumber="309">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50C9350201DFC9C7566FCAC" box="[521,571,845,868]" pageId="4" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
belongs to a characteristic New Guinean species-group. It is closely related to
<taxonomicName id="13934D02E50C9350230FFC507517FC51" authority="Kirsch, 1875" authorityName="Kirsch" authorityYear="1875" box="[283,586,897,921]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="species" species="bituberosa">
<emphasis id="E6E7EA93E50C9350230FFC50769FFC51" box="[283,450,897,921]" italics="true" pageId="4" pageNumber="309">Uloma bituberosa</emphasis>
Kirsch, 1875
</taxonomicName>
. It has the same very characteristic
<typeStatus id="0B288823E50C93502397FC4D76F0FC7C" box="[387,429,924,948]" pageId="4" pageNumber="309">type</typeStatus>
of aedeagus, found in
<taxonomicName id="13934D02E50C93502265FC6677AFFC06" authorityName="Kirsch" authorityYear="1875" box="[113,242,950,974]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="species" species="bituberosa">
<emphasis id="E6E7EA93E50C93502265FC6677AFFC06" box="[113,242,950,974]" italics="true" pageId="4" pageNumber="309">U. bituberosa</emphasis>
</taxonomicName>
and its subspecies
<emphasis id="E6E7EA93E50C935023BBFC6776ABFC06" box="[431,502,950,974]" italics="true" pageId="4" pageNumber="309">hamata</emphasis>
<bibRefCitation id="B0024B70E50C935023EAFC6775DAFC06" author="Gebien" box="[510,647,950,974]" firstAuthor="Gebien" pageId="4" pageNumber="309" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien, 1920</bibRefCitation>
described from New
<collectingCountry id="AC847611E50C93502322FC017622FC20" box="[310,383,976,1000]" name="Guinea-Bissau" pageId="4" pageNumber="309">Guinea</collectingCountry>
, New Pomerania (now New Britain, the largest of the Bismarck Islands) and
<collectingCountry id="AC847611E50C93502044FC3A75D8FBCB" box="[592,645,1003,1027]" name="Palau" pageId="4" pageNumber="309">Palau</collectingCountry>
(
<bibRefCitation id="B0024B70E50C9350226DFBD477A0FBD5" author="Gebien" box="[121,253,1029,1053]" firstAuthor="Gebien" pageId="4" pageNumber="309" pagination="213 - 500" refId="ref7264" refString="GEBIEN H. 1920. - Coleoptera Tenebrionidae, in Nova Guinea, Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, vol. XIII, Zoologie, livre 3. Librairie et Imprimerie E. J. Brill, Leiden: 213 - 500." type="book chapter" year="1920">Gebien 1920</bibRefCitation>
), and
<emphasis id="E6E7EA93E50C9350232CFBD476CBFBD5" box="[312,406,1029,1053]" italics="true" pageId="4" pageNumber="309">sydneyana</emphasis>
<bibRefCitation id="B0024B70E50C93502388FBD4757CFBD5" author="KASZAB Z." box="[412,545,1029,1053]" pageId="4" pageNumber="309" pagination="233 - 291" refId="ref7425" refString="KASZAB Z. 1982 a. - Revision der australischen Uloma - Arten (Coleoptera: Tenebrionidae), Acta Zoologica Academiae Scientiarum Hungaricae 28 (3 - 4): 233 - 291." type="journal article" year="1982">Kaszab, 1982</bibRefCitation>
, described from Sydney,
<collectingCountry id="AC847611E50C93502314FBF17604FBF0" box="[256,345,1056,1080]" name="Australia" pageId="4" pageNumber="309">Australia</collectingCountry>
(
<bibRefCitation id="B0024B70E50C9350237CFBF176A1FBF0" author="Kaszab" box="[360,508,1056,1080]" firstAuthor="Kaszab" pageId="4" pageNumber="309" pagination="233 - 291" refId="ref7425" refString="KASZAB Z. 1982 a. - Revision der australischen Uloma - Arten (Coleoptera: Tenebrionidae), Acta Zoologica Academiae Scientiarum Hungaricae 28 (3 - 4): 233 - 291." type="journal article" year="1982" yearSuffix="a">Kaszab 1982a</bibRefCitation>
). However, it clearly differs from
<taxonomicName id="13934D02E50C93502320FBEA76E9FB9A" authorityName="Kirsch" authorityYear="1875" box="[308,436,1082,1106]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="309" phylum="Arthropoda" rank="species" species="bituberosa">
<emphasis id="E6E7EA93E50C93502320FBEA76E9FB9A" box="[308,436,1082,1106]" italics="true" pageId="4" pageNumber="309">U. bituberosa</emphasis>
</taxonomicName>
and its subspecies by the presence of two conical tubercles on the clypeus in males (
<figureCitation id="4CA82A04E50C935022A0FBBE77A6FB4F" box="[180,251,1135,1159]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2B</figureCitation>
) and the different shape of the pronotum (
<figureCitation id="4CA82A04E50C93502263FB5B7795FB6A" box="[119,200,1162,1186]" captionStart="FIG" captionStartId="5.[113,124,1635,1652]" captionTargetBox="[120,1199,273,1593]" captionTargetId="figure-77@5.[112,662,773,1334]" captionTargetPageId="5" captionText="FIG. 1. — Uloma vanuatensis L. Soldati n. sp.holotype ♂ (MNHN EC2280):A, habitus (dorsal view);B, habitus (lateral view); C, habitus (ventral view); D, forebody (lateral view); E, aedeagus (lateral view); F, aedeagus (tergal face). Scale bars: A-C, 10 mm; D-F, 1 mm. Photographs: L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165433" httpUri="https://zenodo.org/record/5165433/files/figure.png" pageId="4" pageNumber="309">Fig. 1D</figureCitation>
), whose bumps on the sides of the anterior impression of males are much lower: they are not projected forward up to the front edge or beyond it, and they are not hook-shaped inwards.
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50C93532265FAFC75D8F9EC" lastPageId="7" lastPageNumber="312" pageId="4" pageNumber="309" type="description">
<paragraph id="D42C3681E50C93502265FAFC765FFA8D" blockId="4.[113,649,1325,1477]" box="[113,258,1325,1351]" pageId="4" pageNumber="309">DESCRIPTION</paragraph>
<paragraph id="D42C3681E50C93502265FA9D7577FA0D" blockId="4.[113,649,1325,1477]" pageId="4" pageNumber="309">
Length 10-11.5 mm; width 3.5-
<quantity id="136B9B64E50C935023F3FA9D756CFAAF" box="[487,561,1356,1383]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="309" unit="mm" value="4.0">4 mm</quantity>
. Pitchy dark brown, sometimes slightly reddish (teneral specimens), brilliant. Antennae, mouthparts, legs and prosternal apophysis reddish-brown.
</paragraph>
<paragraph id="D42C3681E50C93502265FA3A764DF9CD" blockId="4.[111,647,1515,1700]" box="[113,272,1515,1541]" pageId="4" pageNumber="309">
<emphasis id="E6E7EA93E50C93502265FA3A764DF9CD" box="[113,272,1515,1541]" italics="true" pageId="4" pageNumber="309">
Head (
<figureCitation id="4CA82A04E50C935022ADFA3A7655F9CD" box="[185,264,1515,1541]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D42C3681E50C93502265F9DA7366FE99" blockId="4.[111,647,1515,1700]" lastBlockId="4.[674,1211,216,1319]" pageId="4" pageNumber="309">Transverse,genae narrower than eyes, first subparallel in a very short distance, then obliquely narrowed in curved line forward up to the clypeo-genal suture, which is limited laterally by two short faint grooves and disappears completely in the middle, in the area that normally separates the front from the clypeus. Clypeus transversely enlarged, swollen (standing out from frons despite absence of visible suture) and its anterior edge slightly concave.</paragraph>
<paragraph id="D42C3681E50C935020AFFE8773E7FD25" blockId="4.[674,1211,216,1319]" pageId="4" pageNumber="309">
Male:vertex sloping steeply forward and separated from front by a transverse depression as wide as space between eyes. Frons plane with a slight impression in front in the middle, in area of contact with the clypeal bead. Clypeus densely covered with tangled microgranulation and adorned with two small conical tubercles (
<figureCitation id="4CA82A04E50C9350212CFDC474D1FDE7" box="[824,908,533,559]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2B</figureCitation>
, tub), between them with a narrow arc-shaped strip (
<figureCitation id="4CA82A04E50C935021B9FDE574A3FD86" box="[941,1022,564,590]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2B</figureCitation>
, st) whose microchagreened/matte surface contrasts sharply with the shiny rest of upper head. Punctuation is very fine, sparse and superficial in the middle of frons and becomes progressively denser on genae and toward clypeus, and stronger and coarser on the temporae.
</paragraph>
<paragraph id="D42C3681E50C935020AFFD237416FC22" blockId="4.[674,1211,216,1319]" pageId="4" pageNumber="309">
Female: vertex coarsely punctate, convex and separated from temporae and frons by a transverse depression more or less pronounced, but still clearly visible. Contrary to
<specimenCount id="C295FD08E50C9350216AFC8474C9FCA3" box="[894,916,853,875]" pageId="4" pageNumber="309" type="male"></specimenCount>
, frons is also convex transversely and much more strongly punctate. Clypeus also swollen, but its surface only punctate, smooth and shiny, neither with conical tubercles nor microgranulation.
</paragraph>
<paragraph id="D42C3681E50C935020AFFC217425FAEF" blockId="4.[674,1211,216,1319]" pageId="4" pageNumber="309">
Antennae (
<figureCitation id="4CA82A04E50C93502123FC2174D3FBC2" box="[823,910,1008,1034]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2A</figureCitation>
) without notable features, antennomeres gradually becoming transverse and expanded from antennomere5 which is slightly asymmetrical. Mentum (
<figureCitation id="4CA82A04E50C93502161FB9E7498FBA1" box="[885,965,1103,1129]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[191,1132,229,1628]" captionTargetId="figure-71@6.[191,1132,214,1628]" captionTargetPageId="6" captionText="FIG. 2. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280): A, head (ventral view);B, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="4" pageNumber="309">Fig. 2A</figureCitation>
) heart-shaped, smooth and covered with coarse but shallow punctuation, with distinct midlongitudinal triangular depression, deepest anteriorly and gradually becoming shallower backward, and two short oblique lateral grooves near the base, arranged symmetrically in relation to midline.
</paragraph>
<paragraph id="D42C3681E50C935020B1FA9C7451FAAE" blockId="4.[674,1212,1357,1700]" box="[677,780,1357,1382]" pageId="4" pageNumber="309">
<emphasis id="E6E7EA93E50C935020B1FA9C7451FAAE" box="[677,780,1357,1382]" italics="true" pageId="4" pageNumber="309">Pronotum</emphasis>
</paragraph>
<paragraph id="D42C3681E50C935020B1FABD7490F96B" blockId="4.[674,1212,1357,1700]" pageId="4" pageNumber="309">1.3 times wider than long, sides weakly arcuate, almost parallel, widest at basal third. Rim on the anterior margin disappears completely in the middle; base immarginated, with exception of two small folds located at the level of the two concave curves of external margin and the posterior angles. Anterior angles smooth and slightly protruding forward, posterior ones obtuse. Whole upper surface of the pronotum very finely and regularly punctate, slightly denser on the sides.</paragraph>
<paragraph id="D42C3681E50D9351229CFF3B7469FF36" blockId="5.[136,580,234,254]" box="[136,820,234,254]" lastBlockId="5.[801,820,234,254]" pageId="5" pageNumber="310">A B C</paragraph>
<caption id="80EC6609E50D93512265F9B27660F969" ID-DOI="http://doi.org/10.5281/zenodo.5165433" ID-Zenodo-Dep="5165433" httpUri="https://zenodo.org/record/5165433/files/figure.png" pageId="5" pageNumber="310" startId="5.[113,124,1635,1652]" targetBox="[120,1199,273,1593]" targetPageId="5">
<paragraph id="D42C3681E50D93512265F9B27660F969" blockId="5.[113,1211,1635,1697]" pageId="5" pageNumber="310">
FIG. 1. —
<taxonomicName id="13934D02E50D935122ABF9B276F1F9BC" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[191,428,1635,1652]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="310" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E50D935122ABF9B27607F9BC" box="[191,346,1635,1652]" italics="true" pageId="5" pageNumber="310">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50D935123A5F9B276BDF9BC" box="[433,480,1635,1652]" pageId="5" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ (MNHN EC2280):
<emphasis id="E6E7EA93E50D935120C1F9B275BFF9BC" bold="true" box="[725,738,1635,1652]" pageId="5" pageNumber="310">A</emphasis>
, habitus (dorsal view);
<emphasis id="E6E7EA93E50D93512189F9B274F7F9BC" bold="true" box="[925,938,1635,1652]" pageId="5" pageNumber="310">B</emphasis>
, habitus (lateral view);
<emphasis id="E6E7EA93E50D93512671F9B2732EF9BC" bold="true" box="[1125,1139,1635,1652]" pageId="5" pageNumber="310">C</emphasis>
, habitus (ventral view);
<emphasis id="E6E7EA93E50D935122FEF9A877A5F942" bold="true" box="[234,248,1657,1674]" pageId="5" pageNumber="310">D</emphasis>
, forebody (lateral view);
<emphasis id="E6E7EA93E50D935123D3F9A8768EF942" bold="true" box="[455,467,1657,1674]" pageId="5" pageNumber="310">E</emphasis>
, aedeagus (lateral view);
<emphasis id="E6E7EA93E50D935120BEF9A875E8F942" bold="true" box="[682,693,1657,1674]" pageId="5" pageNumber="310">F</emphasis>
, aedeagus (tergal face). Scale bars: A-C, 10 mm; D-F, 1 mm. Photographs: L. Soldati.
</paragraph>
</caption>
<caption id="80EC6609E50E93522265F9A874F1F969" ID-DOI="http://doi.org/10.5281/zenodo.5165435" ID-Zenodo-Dep="5165435" httpUri="https://zenodo.org/record/5165435/files/figure.png" pageId="6" pageNumber="311" startId="6.[113,124,1657,1674]" targetBox="[191,1132,229,1628]" targetPageId="6">
<paragraph id="D42C3681E50E93522265F9A874F1F969" blockId="6.[113,1211,1657,1697]" pageId="6" pageNumber="311">
FIG. 2. —
<taxonomicName id="13934D02E50E935222AAF9A876F1F942" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[190,428,1657,1674]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="311" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E50E935222AAF9A87607F942" box="[190,346,1657,1674]" italics="true" pageId="6" pageNumber="311">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50E935223A4F9A87682F942" box="[432,479,1657,1674]" pageId="6" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ (MNHN EC2280):
<emphasis id="E6E7EA93E50E935220C1F9A875BFF942" bold="true" box="[725,738,1657,1674]" pageId="6" pageNumber="311">A</emphasis>
, head (ventral view);
<emphasis id="E6E7EA93E50E93522199F9A874C7F942" bold="true" box="[909,922,1657,1674]" pageId="6" pageNumber="311">B</emphasis>
, head (dorsal view).Abbreviations: tub, conical tubercle; st, arc-shaped strip. Scale bars: A, 1.5 mm; B, 1 mm. Photographs L. Soldati.
</paragraph>
</caption>
<paragraph id="D42C3681E50F9353229CFF097537FDC7" blockId="7.[111,648,216,623]" pageId="7" pageNumber="312">Male: antero-median depression of pronotum rather moderate, shallow, not reaching half of pronotal length and becoming gradually shallower toward posterior end in circular arch without any granule to delimit its posterior edge. The lateral bumps that initiate the depressions sides forward are quite low, not projected to anterior edge and simply sloping inwards, without forming lamellar hook-shaped expansions. Bottom of depression somewhat more strongly punctate than rest of pronotal surface.</paragraph>
<paragraph id="D42C3681E50F9353229CFDC475D8FDA7" blockId="7.[111,648,216,623]" pageId="7" pageNumber="312">Female: pronotum regularly convex, without antero-median depression. Prosternal apophysis in lateral view regularly curved beneath anterior coxae.</paragraph>
<paragraph id="D42C3681E50F93532265FD4277F2FD65" blockId="7.[111,648,659,939]" box="[113,175,659,685]" pageId="7" pageNumber="312">
<emphasis id="E6E7EA93E50F93532265FD4277F2FD65" box="[113,175,659,685]" italics="true" pageId="7" pageNumber="312">Elytra</emphasis>
</paragraph>
<paragraph id="D42C3681E50F93532265FD62752BFC63" blockId="7.[111,648,659,939]" pageId="7" pageNumber="312">Elytra quite flattened on disc, sides parallel, humeral angles right but rounded at the top. Lateral margin visible only in anterior half in dorsal view. Each elytron bears nine grooved striae of punctures and a scutellar striola. Strial punctures are slightly wider than grooves. Elytral intervals nearly flat on disc and becoming convex laterally and toward apex, covered with extremely fine and superficial punctuation.</paragraph>
<paragraph id="D42C3681E50F93532265FC01778EFC22" blockId="7.[111,648,976,1573]" box="[113,211,976,1002]" pageId="7" pageNumber="312">
<emphasis id="E6E7EA93E50F93532265FC01778EFC22" box="[113,211,976,1002]" italics="true" pageId="7" pageNumber="312">Abdomen</emphasis>
</paragraph>
<paragraph id="D42C3681E50F93532265FC217603FA6E" blockId="7.[111,648,976,1573]" pageId="7" pageNumber="312">
Abdominal ventrites (
<figureCitation id="4CA82A04E50F93532358FC2176CBFBC2" box="[332,406,1008,1034]" captionStart="FIG" captionStartId="8.[677,688,1634,1651]" captionTargetBox="[677,1209,216,1586]" captionTargetId="figure-358@8.[677,1211,216,1587]" captionTargetPageId="8" captionText="FIG. 3. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280):A, anterior tibia (upper face);B, metathorax and abdomen (ventralview).Scalebars:A,0,.5 mm;B,1 mm.Photographs:L.Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165437" httpUri="https://zenodo.org/record/5165437/files/figure.png" pageId="7" pageNumber="312">Fig.3B</figureCitation>
) very finely and densely punctate on a narrow mid-longitudinal strip and on the entire anal ventrite where punctuation becomes progressively bigger and sparser. On each side of this longitudinal strip, surface of ventrites 1-4 is finely microshagreened and striate longitudinally.Moreover, each ventrite is dotted with porous punctures mainly concentrated on leading edge. The terminal ventrite without lateral microshagreened and striate areas, but porous punctures present, and its outer margin very finely bordered except on both sides,in front of the base. Anterior tibiae (
<figureCitation id="4CA82A04E50F93532337FA9D762AFAAE" box="[291,375,1356,1382]" captionStart="FIG" captionStartId="8.[677,688,1634,1651]" captionTargetBox="[677,1209,216,1586]" captionTargetId="figure-358@8.[677,1211,216,1587]" captionTargetPageId="8" captionText="FIG. 3. — Uloma vanuatensis L. Soldati n. sp. holotype ♂ (MNHN EC2280):A, anterior tibia (upper face);B, metathorax and abdomen (ventralview).Scalebars:A,0,.5 mm;B,1 mm.Photographs:L.Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165437" httpUri="https://zenodo.org/record/5165437/files/figure.png" pageId="7" pageNumber="312">Fig. 3A</figureCitation>
) carinate on their upper surface up to the middle and slightly excavated at base on the inner side.
</paragraph>
<paragraph id="D42C3681E50F93532265FA7A75D8F9EC" blockId="7.[111,648,976,1573]" pageId="7" pageNumber="312">
Aedeagus:on tergal face (
<figureCitation id="4CA82A04E50F93532365FA7A76E6FA0D" box="[369,443,1451,1477]" captionStart="FIG" captionStartId="5.[113,124,1635,1652]" captionTargetBox="[120,1199,273,1593]" captionTargetId="figure-77@5.[112,662,773,1334]" captionTargetPageId="5" captionText="FIG. 1. — Uloma vanuatensis L. Soldati n. sp.holotype ♂ (MNHN EC2280):A, habitus (dorsal view);B, habitus (lateral view); C, habitus (ventral view); D, forebody (lateral view); E, aedeagus (lateral view); F, aedeagus (tergal face). Scale bars: A-C, 10 mm; D-F, 1 mm. Photographs: L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165433" httpUri="https://zenodo.org/record/5165433/files/figure.png" pageId="7" pageNumber="312">Fig. 1F</figureCitation>
), parameres parallel in basal half, then suddenly narrowed and pointed toward apex;base triangular backwards.In lateral view (
<figureCitation id="4CA82A04E50F93532263F9DB779BF9ED" box="[119,198,1546,1573]" captionStart="FIG" captionStartId="5.[113,124,1635,1652]" captionTargetBox="[120,1199,273,1593]" captionTargetId="figure-77@5.[112,662,773,1334]" captionTargetPageId="5" captionText="FIG. 1. — Uloma vanuatensis L. Soldati n. sp.holotype ♂ (MNHN EC2280):A, habitus (dorsal view);B, habitus (lateral view); C, habitus (ventral view); D, forebody (lateral view); E, aedeagus (lateral view); F, aedeagus (tergal face). Scale bars: A-C, 10 mm; D-F, 1 mm. Photographs: L. Soldati." figureDoi="http://doi.org/10.5281/zenodo.5165433" httpUri="https://zenodo.org/record/5165433/files/figure.png" pageId="7" pageNumber="312">Fig. 1E</figureCitation>
), apical half of parameres strongly curved.
</paragraph>
</subSubSection>
<subSubSection id="9C89650AE50F93532265F99B74F8FEF9" pageId="7" pageNumber="312" type="discussion">
<paragraph id="D42C3681E50F93532265F99B7788F9AA" blockId="7.[113,649,1610,1700]" box="[113,213,1610,1636]" pageId="7" pageNumber="312">REMARKS</paragraph>
<paragraph id="D42C3681E50F93532265F9B874F8FEF9" blockId="7.[113,649,1610,1700]" lastBlockId="7.[675,1210,216,306]" pageId="7" pageNumber="312">
<taxonomicName id="13934D02E50F93532265F9B876E1F94B" authority="L. Soldati" authorityName="L. Soldati" authorityYear="2012" box="[113,444,1641,1668]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="312" phylum="Arthropoda" rank="species" species="vanuatensis" status="sp. nov.">
<emphasis id="E6E7EA93E50F93532265F9B8761FF94B" box="[113,322,1641,1667]" italics="true" pageId="7" pageNumber="312">Uloma vanuatensis</emphasis>
L. Soldati
</taxonomicName>
<taxonomicNameLabel id="FDD457E8E50F935323D1F9BB755AF94C" box="[453,519,1642,1668]" pageId="7" pageNumber="312" rank="species">n. sp.</taxonomicNameLabel>
, like all its congeners, is a saproxylic insect living and developing beneath bark and in decaying wood. Unfortunately, neither the larvae nor the food preferences (
<typeStatus id="0B288823E50F93532679FF2973C0FEDA" box="[1133,1181,248,274]" pageId="7" pageNumber="312">type</typeStatus>
of tree species) are known.
</paragraph>
</subSubSection>
</treatment>
</document>