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</mods:role>
<mods:namePart>Bousquet, Yves</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bousquety1@yahoo.com</mods:nameIdentifier>
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<treatment ID-GBIF-Taxon="127867070" LSID="urn:lsid:plazi:treatment:F4993469ACF69AEE094888A1B90532A6" httpUri="http://treatment.plazi.org/id/F4993469ACF69AEE094888A1B90532A6" lastPageNumber="423" pageId="422" pageNumber="423">
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<taxonomicName LSID="F4993469-ACF6-9AEE-0948-88A1B90532A6" authority="Lindroth, 1969" authorityName="Lindroth" authorityYear="1969" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bembidion manningense" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="manningense">Bembidion manningense Lindroth, 1969</taxonomicName>
</paragraph>
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<subSubSection pageId="422" pageNumber="423" type="reference_group">
<paragraph pageId="422" pageNumber="423">
<taxonomicName authorityName="Lindroth" authorityYear="1963" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="ochropus">Bembidion ochropus</taxonomicName>
Lindroth, 1963b: 278 [primary homonym of
<taxonomicName authorityName="Lindroth" authorityYear="1963" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="ochropus">
<emphasis italics="true" pageId="422" pageNumber="423">Bembidion ochropus</emphasis>
</taxonomicName>
Andrewes, 1935]. Type locality: &quot;Pasayten R[iver] E[ast] Manning Park, B[ritish] C[olumbia]&quot; (original citation). Holotype (♂) in CNC [# 8393].
</paragraph>
<paragraph pageId="422" pageNumber="423">
<taxonomicName authorityName="Lindroth" authorityYear="1969" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="manningense">Bembidion manningense</taxonomicName>
Lindroth, 1969a: 1114. Replacement name for
<taxonomicName authorityName="Lindroth" authorityYear="1963" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="ochropus">
<emphasis italics="true" pageId="422" pageNumber="423">Bembidion ochropus</emphasis>
</taxonomicName>
Lindroth, 1963.
</paragraph>
</subSubSection>
<subSubSection pageId="422" pageNumber="423" type="distribution">
<paragraph pageId="422" pageNumber="423">Distribution.</paragraph>
<paragraph pageId="422" pageNumber="423">
This species ranges from the Yukon River in western Alaska to the Mackenzie River delta in northwestern Northwest Territories (Lindroth 1963b: 278-279, as
<taxonomicName authorityName="Lindroth" authorityYear="1963" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="ochropus">
<emphasis italics="true" pageId="422" pageNumber="423">Bembidion ochropus</emphasis>
</taxonomicName>
), south through the Rocky Mountains to western Montana (Russell 1968: 51) and mountains in southern British Columbia (Lindroth 1963b: 279, as
<taxonomicName authorityName="Lindroth" authorityYear="1963" class="Hexapoda" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="422" pageNumber="423" phylum="Arthropoda" rank="species" species="ochropus">
<emphasis italics="true" pageId="422" pageNumber="423">Bembidion ochropus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="422" pageNumber="423" type="records">
<paragraph pageId="422" pageNumber="423">Records.</paragraph>
<paragraph pageId="422" pageNumber="423">
<emphasis bold="true" pageId="422" pageNumber="423">CAN</emphasis>
: AB, BC, NT, YT
<emphasis bold="true" pageId="422" pageNumber="423">USA</emphasis>
: AK, MT
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:date id="D5BF039F77163DBC749D7D0F000E50DB">2018</mods:date>
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<subSubSection id="A863701ABA695225F3ABB4615BD73B06" pageId="0" pageNumber="129" type="nomenclature">
<paragraph id="CD5B1C2D0ED9C89DD8BE594756C69937" pageId="0" pageNumber="129">
<taxonomicName id="63C99993154F69ABB52FF1A4DE002DA7" ID-CoL="8RLYL" authority="Dai, Zhang &amp; Webb, 2010" authorityName="Dai, Zhang &amp; Webb" authorityYear="2010" class="Insecta" family="Cicadellidae" genus="Mimotettix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimotettix robustistylus" order="Hemiptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robustistylus">Mimotettix robustistylus Dai, Zhang &amp; Webb, 2010</taxonomicName>
</paragraph>
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<paragraph id="B790234C2A8F93EF542564A01704CFD1" pageId="0" pageNumber="129">
<taxonomicName id="AE03F4C146C262693CF39B301C27B4FC" class="Insecta" family="Cicadellidae" genus="Mimotettix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimotettix robustistylus" order="Hemiptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robustistylus">Mimotettix robustistylus</taxonomicName>
Dai, Zhang &amp; Webb, 2010: 9, figs 13A-F.
</paragraph>
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<paragraph id="A3E57F4EBA0370F626AB0B723073ED2E" pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph id="DB94AF5F96A270C78AB6F586B7439674" pageId="0" pageNumber="129">China (Yunnan).</paragraph>
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<mods:title>Chapter 7: Linnaean Plant Names and their Types (part P)</mods:title>
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<mods:identifier type="Zenodo-Dep">291971</mods:identifier>
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<subSubSection pageNumber="733" type="reference_group">
<paragraph pageNumber="733">
<bibCitation author="Linnaeus" pagination="1181" title="Peziza cochleata" volumeTitle="Species Plantarum 2" year="1753">
<treatmentCitation author="Linnaeus" pageNumber="1181" year="1753">
<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Ascomycetes" family="Pezizaceae" genus="Peziza" higherTaxonomySource="GBIF" kingdom="Fungi" order="Pezizales" phylum="Ascomycota" rank="species" species="cochleata">
<emphasis bold="true" pageNumber="733">Peziza cochleata</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle>
<emphasis italics="true" pageNumber="733">Species Plantarum</emphasis>
2
</volumeTitle>
: 1181. 1753
</treatmentCitation>
</bibCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageNumber="733" type="distribution">
<paragraph pageNumber="733">&quot;Habitat in umbrosis.&quot; RCN: 8501.</paragraph>
</subSubSection>
<subSubSection pageNumber="733" type="materials_examined">
<paragraph pageNumber="733">Type not designated.</paragraph>
</subSubSection>
<subSubSection pageNumber="733" type="materials_examined">
<paragraph pageNumber="733">
Original material: [icon] in Vaillant, Bot.
<taxonomicName class="Liliopsida" family="Melanthiaceae" genus="Paris" kingdom="Plantae" order="Liliales" phylum="Tracheophyta" rank="genus">Paris</taxonomicName>
.: 57, t. 11, f. 8. 1727.
</paragraph>
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<subSubSection pageNumber="733" type="nomenclature">
<paragraph pageNumber="733">
Current name:
<taxonomicName authority="(Huds.: Fr.) Fuckel" authorityName="Fuckel" authorityYear="1870" baseAuthorityName="Huds." class="Ascomycetes" family="Pyronemataceae" genus="Otidea" higherTaxonomySource="GBIF" kingdom="Fungi" order="Pezizales" phylum="Ascomycota" rank="species" species="cochleata">
<emphasis bold="true" italics="true" pageNumber="733">Otidea cochleata</emphasis>
(Huds.: Fr.) Fuckel
</taxonomicName>
(
<taxonomicName authorityName="Corda" authorityYear="1842" rank="family">Pyronemataceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageNumber="733" type="discussion">
<paragraph pageNumber="733">
<emphasis italics="true" pageNumber="733">Note:</emphasis>
See comments by Nannfeldt (in
<emphasis italics="true" pageNumber="733">Ann. Bot. Fenn.</emphasis>
3: 312. 1966).
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="F49943DCCDD0375493526912F3ED6C87" ID-GBIF-Taxon="147660752" LSID="urn:lsid:plazi:treatment:F49943DCCDD0375493526912F3ED6C87" httpUri="http://treatment.plazi.org/id/F49943DCCDD0375493526912F3ED6C87" lastPageId="7" lastPageNumber="26" pageId="4" pageNumber="23">
<subSubSection id="5DE75EF45E3D9456D1141AE1B4DC2C1B" pageId="4" pageNumber="23" type="nomenclature">
<paragraph id="E4F2B37518DAED52E25D97F15946E755" pageId="4" pageNumber="23">
<taxonomicName id="59FE8854C3272BDA71B1085850469529" ID-CoL="DTSB" authority="Cope, 1869" authorityName="Cope" authorityYear="1869" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius Cope, 1869</taxonomicName>
Figs 1, 2, 3, 4, 6, 7, 8, 9, Tables 1, 2
</paragraph>
</subSubSection>
<subSubSection id="05A64A244FAEC5311DFC9CAAA2DDC9C2" pageId="4" pageNumber="23" type="reference_group">
<paragraph id="5EEC375E493F8CFA024F3DE3D09E2338" pageId="4" pageNumber="23">
<taxonomicName id="FF9FC175EB16EA335A3030E25178FBAC" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius</taxonomicName>
Cope, 1869: 182;
<bibRefCitation id="EA72075D6206765B3EF6ED6B4C1615CF" author="Gardner, MR" journalOrPublisher="Memoirs of the Pacific Coast Entomological Society" pageId="17" pageNumber="36" pagination="1 - 61" title="Revision of the millipede family Andrognathidae in the Nearctic region." volume="5" year="1975">Gardner 1975</bibRefCitation>
: 15, figs 1, 5-7;
<bibRefCitation id="AADCFE7E86F9B91F8DD4B831E9D5AE5E" author="Shear, WA" journalOrPublisher="Festschrift in honor of Richard Hoffman, Memoirs of the Virginia Museum of Natural History" pageId="19" pageNumber="38" pagination="149 - 158" title="Andrognathushoffmani, n. sp., a second species in the genus and the first species of Andrognathidae from Mexico (Diplopoda, Platydesmida, Andrognathidae)." url="https://www.fieldmuseum.org/sites/default/files/shear%26marek_festschrift_2009.pdf" volume="16" year="2009">Shear and Marek 2009</bibRefCitation>
: 157, figs 1-10. (There are no synonyms for
<taxonomicName id="E080F174328ECC365DF94ACAF62F4A75" family="Andrognathidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. corticarius" order="Platydesmida" pageId="4" pageNumber="23" rank="species" species="corticarius">A. corticarius</taxonomicName>
.)
</paragraph>
</subSubSection>
<subSubSection id="87D2A3CA04F0ADA6B4F5B2B66378EFE6" pageId="4" pageNumber="23" type="type species">
<paragraph id="EEC66FAC853477FFF682F589B77E5CDE" pageId="4" pageNumber="23">Type species.</paragraph>
<paragraph id="AE87D91C0288D51AE2119AD5314B6107" pageId="4" pageNumber="23">Original type material lost. Collected by Cope &quot;from Montgomery County, Virginia&quot;.</paragraph>
</subSubSection>
<subSubSection id="DA653220E8237FA0DCDD61FFD467BD69" pageId="4" pageNumber="23" type="materials_examined">
<paragraph id="D4B51FC15A48D9DC3A90D5EDA9B4150E" pageId="4" pageNumber="23">Material examined.</paragraph>
<paragraph id="C30FB6BECE1D74847CFAA2C32058F37B" pageId="4" pageNumber="23">Neotype: Male neotype (VTEC, MPE01942); 1 male from Virginia, Montgomery County, Christiansburg, Yellow Sulfur Springs Spa (37.1796, -80.3979, Elev. 607m), 30 June 2016 (Colls: P. Shorter, J. Means, V. Wong). Head and posterior body rings preserved as voucher. Collected on a hardwood log on a footpath between the old hotel gardens and an abandoned bowling alley.</paragraph>
<paragraph id="A67A8CAB77F055D2882E1BF0B1DD105D" pageId="4" pageNumber="23">Other material examined: 3 males, 3 females, details as for neotype; 109 males, 113 females, and 52 juveniles were examined from Virginia and nine other states (West Virginia, Tennessee, North Carolina, Florida, South Carolina, Alabama, Georgia, Ohio, and Kentucky), details in Suppl. material 1.</paragraph>
<paragraph id="F61456586162053BE4A875C9641BF76F" pageId="4" pageNumber="23">
We here designate a neotype for
<taxonomicName id="C9B6BE86104013382096D2CACEF157F7" lsidName="A. corticarius" pageId="4" pageNumber="23" rank="species" species="corticarius">A. corticarius</taxonomicName>
because the holotype or syntypes have not been located and are presumed lost (see Material and methods section), and because our morphological studies indicate that &quot;
<taxonomicName id="0821B4583EEFB3E14D88BFF6C3B9A4AE" lsidName="A. corticarius" pageId="4" pageNumber="23" rank="species" species="corticarius">A. corticarius</taxonomicName>
&quot; may represent more than one species. To provide a basis for taxonomy of the group we have selected a specimen from the type locality which agrees in all details with the description given by
<bibRefCitation id="4FF193AFBDE6927A5E9253509A502D5A" author="Cope, ED" journalOrPublisher="Proceedings of the American Philosophical Society" pageId="17" pageNumber="36" pagination="147 - 192" title="Synopsis of the extinct Mammalia of the cave formations in the United States, with observations on some Myriapoda found in and near the same, and on some extinct mammals of the caves of Anguilla, WI, and of other localities." url="http://biodiversitylibrary.org/page/31205032" volume="11" year="1869">Cope (1869)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AFEB829907376DD29D02371990554DBF" pageId="4" pageNumber="23" type="diagnosis">
<paragraph id="167671149D7D068F4C4F2C7EAD5989DB" pageId="4" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="6F28E7703D945151B7A57B1C141BDABA" pageId="4" pageNumber="23">
Adult males of
<taxonomicName id="B896EDB8A85B41FE7F4094DC9EADF084" lsidName="A. corticarius" pageId="4" pageNumber="23" rank="species" species="corticarius">A. corticarius</taxonomicName>
are distinct from other
<taxonomicName id="1DA7ED308CF5AB57E96C2E1A6B5E198D" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus" order="Platydesmida" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">Andrognathus</taxonomicName>
species based on the following combination of characters:
<taxonomicName id="A8D2931CF670A29F6A33033A90CC6323" genus="Exoskeleton" lsidName="Exoskeleton" pageId="4" pageNumber="23" rank="genus">Exoskeleton</taxonomicName>
. Ring five with pleuroterga distinctly bilobed, papilioform (Figure 2A); lobe of anterior corner flared anteriorly, lobe of posterior corner bulging laterally, contrasting with the reduced lobes of
<taxonomicName id="B60F8DF089D870D093BFC0C3D8F97296" lsidName="A. hoffmani" pageId="4" pageNumber="23" rank="species" species="hoffmani">A. hoffmani</taxonomicName>
and
<taxonomicName id="A455EF6C8DA8269463BC6FF24062A9CF" lsidName="A. grubbsi" pageId="4" pageNumber="23" rank="species" species="grubbsi">A. grubbsi</taxonomicName>
sp. n. Ring VI with metaterga singly lobed, with angulate paranota bearing a posterolaterally oriented ozopore, separating
<taxonomicName id="8A381B81D03F8528D8DC9E5421DBC4BC" lsidName="A. corticarius" pageId="4" pageNumber="23" rank="species" species="corticarius">A. corticarius</taxonomicName>
from
<taxonomicName id="95C027C2D7F89882F11671E5864FA499" lsidName="A. hoffmani" pageId="4" pageNumber="23" rank="species" species="hoffmani">A. hoffmani</taxonomicName>
and
<taxonomicName id="D411FDAC3C229738BF238281BDAF47DD" lsidName="A. grubbsi" pageId="4" pageNumber="23" rank="species" species="grubbsi">A. grubbsi</taxonomicName>
sp. n. in which the paranota are rounded and bear a laterally oriented ozopore (Figure 2). Paranota becoming progressively more directed posteriorly along subsequent body rings. Posterior gonopod with P6 terminating in a distally flared calyx, tip with many small serrations (Figs 6-8, Figure 9A), not spatulate as in
<taxonomicName id="76B2A95CEEF9F2F5F51DA191351B2597" lsidName="A. hoffmani" pageId="4" pageNumber="23" rank="species" species="hoffmani">A. hoffmani</taxonomicName>
(Figure 9B) and
<taxonomicName id="B8F1187494EF286B7DA4646F30DA42B6" lsidName="A. grubbsi" pageId="4" pageNumber="23" rank="species" species="grubbsi">A. grubbsi</taxonomicName>
sp. n. (Figure 9C). Claw (ungulum) of P6 present, thorn-shaped, varying in size; claw not perpendicular to the shaft of the gonopod as in
<taxonomicName id="AC516836A0535DB54AA37084012BF04B" lsidName="A. grubbsi" pageId="4" pageNumber="23" rank="species" species="grubbsi">A. grubbsi</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection id="731121D828421E2AC0A0C81288E861BE" pageId="5" pageNumber="24" type="neotype details">
<paragraph id="B587FF594F83A634E91A51E2AE90E6FB" pageId="5" pageNumber="24">
<pageBreakToken id="ABDB3E79284E18F55529304B7EAFDCE2" pageId="5" pageNumber="24" start="start">Neotype</pageBreakToken>
details.
</paragraph>
<paragraph id="AFC50400B1EFC2AE65AA96B82CFC89F7" pageId="5" pageNumber="24">Body length (BL) = 17.9 mm, head width (HW) = 0.46 mm, collum width (CW) = 0.61 mm, metazonite width at 1/4 length of body (W1) = 0.81 mm, number of podous tergites (p) = 57, number of legs (l) = 210. NCBI accession # MH282831.</paragraph>
</subSubSection>
<subSubSection id="130014D73A1B593F173CDFA0FD74DA74" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24" type="variation">
<paragraph id="52B1EA197E6A88E1BE205EABBBF141AC" pageId="5" pageNumber="24">Variation.</paragraph>
<paragraph id="3106388263445569BACC3CA483AFEE79" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24">
<taxonomicName id="A9FD682D85489D25BC357A44BBD03FC4" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius</taxonomicName>
is known from the panhandle of Florida, north into southern Indiana and Pennsylvania (Figure 3). Variation in somatic characters of males and females is given in Table 1, and variation in gonopodal characters in Table 2. The morphology of the A6 and P6 podomeres and the claw of the P6 of the male gonopods of
<taxonomicName id="B2AAC21509714FED6F6FECD67503B9BC" lsidName="A. corticarius" pageId="5" pageNumber="24" rank="species" species="corticarius">A. corticarius</taxonomicName>
differed most across the distribution of the species (Table 2).
<pageBreakToken id="0FFC1E03D5BFF0EFC1D37C2980EA5FDE" pageId="6" pageNumber="25" start="start">Four</pageBreakToken>
individuals from the subset of geographically widespread samples differed from the neotype in two of these three characters of the male gonopods (Figs 6B, 7A, 8A, 8D); a male from Chesterfield County, South Carolina, deviates from the neotype in all three of these characters (Figure 8B).
</paragraph>
<caption id="D9BA87B8F2F4F711D5B4F375530C67F2" ID-Table-UUID="D9BA87B8F2F4F711D5B4F375530C67F2" httpUri="http://table.plazi.org/id/D9BA87B8F2F4F711D5B4F375530C67F2" pageId="6" pageNumber="25">
<paragraph id="01CED258C38274C2AEBC4CB634BAFCBE" pageId="6" pageNumber="25">
Table 2. Variable gonopodal characters in
<taxonomicName id="67F85A3B00CC4C747EDB5F65FC427C4E" lsidName="A. corticarius." pageId="6" pageNumber="25" rank="species" species="corticarius.">A. corticarius.</taxonomicName>
Neotype (*).
</paragraph>
</caption>
<paragraph id="410FC5133BB323B7013A084DF9DCE0BE" pageId="6" pageNumber="25">
<table id="3AB325C729C5AC303C8F75B987CB31E0" pageId="6" pageNumber="25">
<tr id="2B8B66669E63A06A821CFA3043AE8148" pageId="6" pageNumber="25">
<th id="D98CB09D1DCF8A22BFB6A015D77D41D2" colspan="1" pageId="6" pageNumber="25" rowspan="1">A6 Ribbed</th>
<th id="86A4E928000BEDA346178C0504626F04" colspan="1" pageId="6" pageNumber="25" rowspan="1">A6 Spatulate smooth curve</th>
<th id="3E112446645BA2A589EF2E9842AB443F" colspan="1" pageId="6" pageNumber="25" rowspan="1">A6 Ribbed, scalloped edges</th>
<th id="BACF951848B0C77C3F654093DAF06BB7" colspan="1" pageId="6" pageNumber="25" rowspan="1">A6 Medial bifurcate hook</th>
<th id="767055E9C6170D8EE86B9841DFA0DE8C" colspan="1" pageId="6" pageNumber="25" rowspan="1">P6 claw Toothlike hook</th>
<th id="6C3621D990A7384DE9CD469EE47A35F2" colspan="1" pageId="6" pageNumber="25" rowspan="1">P6 claw Bulbous, Rounded</th>
<th id="0552FCD07FD99BCA3BF0995CD90FDA70" colspan="1" pageId="6" pageNumber="25" rowspan="1">P6 claw Pinched</th>
<th id="E36506C72455A7E00C44DEB2AB1FBE31" colspan="1" pageId="6" pageNumber="25" rowspan="1">P6 claw Bulbous, Rounded tooth lacking</th>
<th id="7E29F76AB03BCCF1E473080895FF8F68" colspan="1" pageId="6" pageNumber="25" rowspan="1">P6 claw Bifurcate</th>
<th id="C955FF09B6D5472E2C34887443E78FD0" colspan="1" pageId="6" pageNumber="25" rowspan="1">Apex of P6 Singular fringed crown</th>
<th id="57B409E188C49D861AE50C0A26367E31" colspan="1" pageId="6" pageNumber="25" rowspan="1">Apex of P6 Bifurcate</th>
<th id="B072A2B231A2E1146DC8E502A9AF3F0A" colspan="1" pageId="6" pageNumber="25" rowspan="1">Apex of P6 Bifurcate branched</th>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="CB92F1B55C10FA3077FF4BAF2BA33141" pageId="7" pageNumber="26" type="genetic analysis">
<paragraph id="EAAA94D7A144900F6135CE4CD5125BF0" pageId="7" pageNumber="26">
<pageBreakToken id="42C44E5608FC57F4A8048DA0C0C6E242" pageId="7" pageNumber="26" start="start">Genetic</pageBreakToken>
analysis.
</paragraph>
<paragraph id="2C77C6A9BEF6D8F93D05236E11536A64" pageId="7" pageNumber="26">
Genetic analysis using the COI region showed relatively low variation within eastern
<taxonomicName id="CF39DEBF42CEF16CCB974FC2DC1D648C" lsidName="A. corticarius" pageId="7" pageNumber="26" rank="species" species="corticarius">A. corticarius</taxonomicName>
(mean pairwise distance = 0.16%). The aligned COI sequences were length invariant, trivial to align, and resulted in a matrix of 605 DNA base-pairs for 14 individuals. Individuals from Boone County, West Virginia and Scioto County, Ohio had a maximum pairwise distance of 0.99% (605 bp). The maximum pairwise distances between individuals from the same locality in Montgomery Co., Virginia, was 0.17% (605 bp).
</paragraph>
</subSubSection>
<subSubSection id="F2955A4CCE07C3172056649D2F38216E" pageId="7" pageNumber="26" type="distribution">
<paragraph id="F099F9200C509036B55B1C568ACD8C9E" pageId="7" pageNumber="26">Distribution and ecology.</paragraph>
<paragraph id="BDE3A6FCFE61F84AD72B3DE496E192A5" pageId="7" pageNumber="26">
<taxonomicName id="F1E86F29C81913BCDFA66A55ABCADDC5" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius</taxonomicName>
is now known from Indiana, Ohio, Pennsylvania, Kentucky, West Virginia, Virginia, Tennessee, North Carolina, South Carolina, Georgia, Alabama, and Florida (Figure 3). The map assembled here extended the known range of
<taxonomicName id="EE1711899EEF8A4D303717B1F4A4B403" lsidName="A. corticarius" pageId="7" pageNumber="26" rank="species" species="corticarius">A. corticarius</taxonomicName>
northward into southwestern Ohio and southeastern Pennsylvania and synthesized and expanded distributions from the literature (Figure 3). Based on the collection dates of museum specimens, the activity period of the species appears to span March - November, with the greatest number of specimens encountered mid-summer. Individuals are often found clinging to the underside of logs in mesic deciduous forests in aggregations of millipedes, including juveniles (Figure 1B).
<taxonomicName id="049BD6EACFB98D8A7F5CBC8C26B9D23A" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius</taxonomicName>
is found from elevations of 51 m to 1160 m, and infrequently as lone adults in leaf litter and in drier habitats (Figure 10).
</paragraph>
<caption id="746F670BE6CAD703BBC9B048BDA5A482" pageId="7" pageNumber="26">
<paragraph id="84202FE761C6BFC4830C8F4628EAFDFD" pageId="7" pageNumber="26">
Figure 10.
<taxonomicName id="91B466CAFE35602EBDCEF1EC6DE1A4A9" class="Diplopoda" family="Andrognathidae" genus="Andrognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrognathus corticarius" order="Platydesmida" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="corticarius">Andrognathus corticarius</taxonomicName>
habitat. A Boone County, West Virginia B Morgan County, Tennessee C Carter County, Kentucky D Montgomery County, Virginia.
</paragraph>
</caption>
<paragraph id="BDA467E43163AF6DB64FFF374DB7F97C" pageId="7" pageNumber="26">
After some specimens of
<taxonomicName id="0DAB451A2D3FDB15509B1D4083DD4B8A" lsidName="A. corticarius" pageId="7" pageNumber="26" rank="species" species="corticarius">A. corticarius</taxonomicName>
were collected in Pulaski Co., Virginia in 2016, a male and female were observed to be coiled together at the bottom of their container (Figure 1D). Upon closer inspection, the female was coiled around a cluster of eggs, with the male on top of her. The millipedes stayed in this configuration even when removed from their container and examined for approximately 20 minutes. This is a previously unobserved behavior in the genus, which was not known to demonstrate parental care.
</paragraph>
</subSubSection>
</treatment>
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<mods:namePart>Ahrens, Dirk</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>837</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>155</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.837.32057</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.837.32057</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-837-1</mods:identifier>
<mods:identifier type="ZBK">4A18822935804DB7B1229F131F6A0AC8</mods:identifier>
<mods:identifier type="ZooBank">4A18822935804DB7B1229F131F6A0AC8</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156200636" LSID="urn:lsid:zoobank.org:act:D76D8280-422B-45F9-B26D-3E59E5051088" httpUri="http://treatment.plazi.org/id/F4998957832CF8DE8E90BA4074190B0C" lastPageNumber="39" pageId="38" pageNumber="39">
<subSubSection pageId="38" pageNumber="39" type="nomenclature">
<paragraph pageId="38" pageNumber="39">
<taxonomicName LSID="http://zoobank.org/D76D8280-422B-45F9-B26D-3E59E5051088" class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica smetsi" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="smetsi">Tetraserica smetsi</taxonomicName>
<taxonomicNameLabel pageId="38" pageNumber="39">sp. n.</taxonomicNameLabel>
Figures 14, 51
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="type material">
<paragraph pageId="38" pageNumber="39">Type material examined.</paragraph>
<paragraph pageId="38" pageNumber="39">Holotype: ♂ &quot;Coll. I. R. Sc. N. B., Cambodia, Pursat prov., Phnom Samkos W.S. forest edge, light trap 14.iv.2005 Leg. K. Smets &amp; I. Var&quot; (ISNB). Paratypes: 2 ♂♂ &quot;Coll. I. R. Sc. N. B. Cambodia (Pursat Prov.) Phnom Samkos wild life Sanctuary, light trap Forest Edge &amp; Primary 14-IV-2005 Leg. K. Smets &amp; I. Var&quot; (ISNB), 7 ♂♂ &quot;Coll. I. R. Sc. N. B. Cambodia, Pursat prov. Phnom Samkos W.S. Pramaoy Forest Edge 15-IV-2005, Light Trap Leg. K. Smets &amp; I. Var&quot; (ISNB, ZFMK).</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="description">
<paragraph pageId="38" pageNumber="39">Description.</paragraph>
<paragraph pageId="38" pageNumber="39">Length of body: 9.4 mm; length of elytra: 6.9 mm; maximum width: 5.8 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large, ratio of diameter/interocular width: 0.65. Ratio of length of metepisternum/metacoxa: 1/1.79. Metatibia short and wide, ratio width/length: 1/2.78; basal group of dorsal spines of metatibia at first third of metatibial length.</paragraph>
<paragraph pageId="38" pageNumber="39">
Aedeagus: Fig. 14
<normalizedToken originalValue="A–C">A-C</normalizedToken>
. Habitus: Fig. 14D.
</paragraph>
<caption pageId="38" pageNumber="39">
<paragraph pageId="38" pageNumber="39">
Figure 14.
<normalizedToken originalValue="A–D">A-D</normalizedToken>
<taxonomicName class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica smetsi" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="smetsi">Tetraserica smetsi</taxonomicName>
sp. n. (holotype)
<normalizedToken originalValue="E–H">E-H</normalizedToken>
<taxonomicName lsidName="T. shangsiensis" pageId="38" pageNumber="39" rank="species" species="shangsiensis">T. shangsiensis</taxonomicName>
Liu et al., 2014 (Vietnam: Tay Yen Tu Nat. Res.)
<normalizedToken originalValue="I–L">I-L</normalizedToken>
<taxonomicName lsidName="T. namnaoensis" pageId="38" pageNumber="39" rank="species" species="namnaoensis">T. namnaoensis</taxonomicName>
sp. n. (holotype) A, E, I aedeagus, left side lateral view C, G, K aedeagus, right side lateral view B, F, J parameres, dorsal view D, H, L habitus. Scale bars: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph pageId="38" pageNumber="39">Female unknown.</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="variation">
<paragraph pageId="38" pageNumber="39">Variation.</paragraph>
<paragraph pageId="38" pageNumber="39">Length of body: 9.3-9.4 mm; length of elytra: 6.6-6.9 mm; maximum width: 5.7-5.8 mm.</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="diagnosis">
<paragraph pageId="38" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="38" pageNumber="39">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Tetraserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraserica smetsi" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="smetsi">Tetraserica smetsi</taxonomicName>
sp. n. is very similar to
<taxonomicName lsidName="T. shangsiensis" pageId="38" pageNumber="39" rank="species" species="shangsiensis">T. shangsiensis</taxonomicName>
Liu et al., 2014 in shape of the aedeagus. The new species differs from the latter by the median lamina of phallobase being approximately as long as left paramere and slightly bifurcate at apex, as well as the preapical tooth of the left paramere being dorsally positioned and duplicate. In
<taxonomicName lsidName="T. shangsiensis" pageId="38" pageNumber="39" rank="species" species="shangsiensis">T. shangsiensis</taxonomicName>
the median lamina of phallobase is distinctly shorter than left paramere, simply pointed, and slightly rounded at apex.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="etymology">
<paragraph pageId="38" pageNumber="39">Etymology.</paragraph>
<paragraph pageId="38" pageNumber="39">The new species is named after one of its collectors, K Smets (noun in genitive singular).</paragraph>
</subSubSection>
</treatment>
</document>