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<document id="9F4E36409958A53D2AAA0FB6F9D1A8A5" IM.bibliography_approvedBy="luisschmitz" IM.illustrations_approvedBy="luisschmitz" IM.materialsCitations_approvedBy="luisschmitz" IM.metadata_approvedBy="luisschmitz" IM.taxonomicNames_approvedBy="luisschmitz" IM.treatments_approvedBy="luisschmitz" checkinTime="1746403890787" checkinUser="luisschmitz" docAuthor="Lis, J. A." docDate="2001" docId="03C82166FFC1FFE7FE5EAA1AFDCB86E2" docLanguage="en" docName="Genus.12.4.429-436.pdf" docOrigin="Genus 12, No. 4" docTitle="Coleocydnus madagascariensis J. A. LIS" docType="treatment" docVersion="2" lastPageNumber="432" masterDocId="FFF1591EFFC3FFE4FFC6AE52FF5D847D" masterDocTitle="Review of the species of Coleocydnus J. A. Lis, with a description of C. sicardi n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae)" masterLastPageNumber="436" masterPageNumber="429" pageNumber="431" updateTime="1746404212573" updateUser="ExternalLinkService">
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<mods:title id="A0A1DF5E296BA4787A077979F2770737">Review of the species of Coleocydnus J. A. Lis, with a description of C. sicardi n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<mods:date id="A81641E3D7D378A412EF4E69331B889C">2001</mods:date>
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<treatment id="03C82166FFC1FFE7FE5EAA1AFDCB86E2" ID-DOI="http://doi.org/10.5281/zenodo.15338816" ID-Zenodo-Dep="15338816" LSID="urn:lsid:plazi:treatment:03C82166FFC1FFE7FE5EAA1AFDCB86E2" httpUri="http://treatment.plazi.org/id/03C82166FFC1FFE7FE5EAA1AFDCB86E2" lastPageId="3" lastPageNumber="432" pageId="2" pageNumber="431">
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<taxonomicName id="4C61EBF3FFC1FFE6FE5EAA1AFC308023" authority="J. A. LIS" authorityName="J. A. LIS" box="[408,877,1096,1120]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="B9154C62FFC1FFE6FE5EAA1AFC5F801D" bold="true" box="[408,770,1096,1120]" italics="true" pageId="2" pageNumber="431">Coleocydnus madagascariensis</emphasis>
J. A.
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<emphasis id="B9154C62FFC1FFE6FC9DAA1FFC308023" bold="true" box="[859,877,1101,1118]" pageId="2" pageNumber="431">LIS</emphasis>
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<paragraph id="8BDE9070FFC1FFE6FD8FAA3AFDE18006" blockId="2.[189,1097,230,1589]" box="[585,700,1128,1147]" pageId="2" pageNumber="431">(Figs 2, 6-7)</paragraph>
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<emphasis id="B9154C62FFC1FFE6FF7BAAF0FE8080C8" box="[189,477,1186,1205]" italics="true" pageId="2" pageNumber="431">Coleocydnus madagascariensis</emphasis>
J. A.
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, 1996: 347
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.
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Body (Fig. 2) black,
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in length,
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in width. Head punctate with numerous tiny, hardly visible punctures; clypeus entirely free, without a subapical pair of setigerous punctures; each paraclypeus with a row of 3-4 submarginal setigerous punctures bearing hair-like setae; eyes narrow, moderately large, ocular index 3.50-4.20; 3
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antennal segment only about 0.85-0.95 times as long as the 2
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segment; rostrum reaching mid coxae. Pronotum subquadrate in outline, its disc sparsely punctate; each lateral margin with a row of 5 submarginal setigerous punctures bearing hair-like setae. Propleuron alutaceous, with numerous wrinkles on its anterior convexity, and numerous coarse punctures in the depression. Scutellum with numerous well separated coarse punctures, larger than those of the pronotum. Mesocorium densely punctate only posteriorly (Fig. 2), punctures of different sizes, small to large; costa flattened, separated from exocorium only in its basal third, bearing two setigerous punctures; membrane small, flap-like, not reaching the tip of abdomen. Abdominal sterna coarsely punctate and transversely striated laterally. Female spermatheca with small proximal flange, and elongated pump region (Fig. 7).
</paragraph>
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<materialsCitation id="3B099A2DFFC0FFE7FF2DAF94FDD98663" collectionCode="SMNS" country="Madagascar" location="Foret de Kirindy" municipality="Foret de Kirindy" pageId="3" pageNumber="432" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectingCountry id="F376D0E0FFC0FFE7FF2DAFB4FEE98583" box="[235,436,486,510]" name="Madagascar" pageId="3" pageNumber="432" value="Madagascar">MADAGASCAR</collectingCountry>
:
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, Madagascar,
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, März 1990, leg. M.
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et al. (
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)
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;
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<specimenCount id="9D675BF9FFC0FFE7FD49AC54FDC18663" box="[655,668,518,542]" pageId="3" pageNumber="432" type="female">3</specimenCount>
<typeStatus id="54DA2ED2FFC0FFE7FD64AC54FC5E8663" box="[674,771,518,542]" pageId="3" pageNumber="432">paratype</typeStatus>
females, the same data as the holotype (
<collectionCode id="ED7008B5FFC0FFE7FEE8AC74FE208643" box="[302,381,550,574]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" pageId="3" pageNumber="432">SMNS</collectionCode>
,
<collectionCode id="ED7008B5FFC0FFE7FE4FAC74FE818643" box="[393,476,550,574]" collectionName="DBUO" pageId="3" pageNumber="432">DBUO</collectionCode>
)
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.
</paragraph>
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<paragraph id="8BDE9070FFC0FFE7FF2DACD5FDCB86E2" blockId="3.[189,1097,229,671]" box="[235,662,647,671]" pageId="3" pageNumber="432">Known only from Madagascar, so far.</paragraph>
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</treatment>
</document>

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<mods:title id="9BB7B1C72F63521D325FD7C852F75BC4">Review of the species of Coleocydnus J. A. Lis, with a description of C. sicardi n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<treatment id="03C82166FFC2FFE5FE1CAEB7FC6D86EF" ID-DOI="http://doi.org/10.5281/zenodo.15338812" ID-Zenodo-Dep="15338812" LSID="urn:lsid:plazi:treatment:03C82166FFC2FFE5FE1CAEB7FC6D86EF" httpUri="http://treatment.plazi.org/id/03C82166FFC2FFE5FE1CAEB7FC6D86EF" lastPageNumber="430" pageId="1" pageNumber="430">
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Genus
<taxonomicName id="4C61EBF3FFC2FFE5FDEBAEB7FC768486" authority="J. A. LIS" authorityName="J. A. LIS" box="[557,811,229,253]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B9154C62FFC2FFE5FDEBAEB7FD9C8480" bold="true" box="[557,705,229,253]" italics="true" pageId="1" pageNumber="430">Coleocydnus</emphasis>
J. A.
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<taxonomicName id="4C61EBF3FFC2FFE5FF7BAF77FEA58545" authority="J. A. LIS, 1994: 187" authorityName="J. A. LIS" authorityPageNumber="187" authorityYear="1994" box="[189,504,293,312]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B9154C62FFC2FFE5FF7BAF77FE6F8545" box="[189,306,293,312]" italics="true" pageId="1" pageNumber="430">Coleocydnus</emphasis>
J. A.
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species by original designation:
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<paragraph id="8BDE9070FFC2FFE5FF2DAFC8FC6D86EF" blockId="1.[189,1097,229,807]" pageId="1" pageNumber="430">Head broader than long, semicircular in outline, its margins with setigerous punctures bearing only hair-like setae, peg-like setae never present; clypeus free; ocelli absent; antennae 5-segmented. Pronotum broader than long, its anterior margin angularly concave centrally, the posterior margin almost straight. Scutellum very long, clearly longer than wide, its apex strongly elongated and narrowed. Hemelytra coleopteroid with very short membrane; clavo-corial suture absent. Meso- and metapleural evaporatoria large (Fig. 6), apex of peritreme with tooth posteriorly, ostiolar opening invisible in a ventral view.</paragraph>
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142
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J.A.
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1.
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: habitus
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Body (Fig. 1) reddish-brown to dark castaneous,
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in length,
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in width. Head impunctate; clypeus slightly concealed by paraclypei laterally, bearing no subapical pair of setigerous punctures; each paraclypeus with a row of 3-4 submarginal setigerous punctures bearing hair-like setae; eyes narrow and small, ocular index 4.40-4.80; 3
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segment; rostrum reaching mid coxae. Pronotum trapezoid in outline, its disc sparsely punctate; each lateral margin with a row of 5-6 submarginal setigerous punctures bearing hair-like setae. Propleuron alutaceous, depression with small punctures. Scutellum impunctate, except for a basal and lateral sparse rows of punctures. Mesocorium densely, evenly punctate, punctures large and deep, of equal size (Fig. 1); costa convex, separated from exocorium in its basal two thirds, without or with a single setigerous puncture; membrane reaching or slightly extending beyond the tip of abdomen. Abdominal sterna punctate laterally. Male paramere as in Fig. 4; aedeagus without median carina ventrally. Female spermatheca with large proximal flange, and dilated pump region (Fig. 5).
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<materialsCitation id="3B099A2DFFC1FFE6FF2DAD74FECD8723" collectingDate="1984-01-20" collectionCode="DBUO" country="Rwanda" pageId="2" pageNumber="431" stateProvince="Prov. Cyangugu" typeStatus="Holotype">
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:
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male,
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,
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(
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;
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female, the same data as the holotype (
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.
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:
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female, Zaire (Congo), Provinz
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, Mt. Kahuzi,
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,
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,
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, ex coll. E.
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.
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<paragraph id="8BDE9070FFC1FFE6FF2DADB5FC598782" blockId="2.[189,1097,230,1589]" box="[235,772,999,1023]" pageId="2" pageNumber="431">Hitherto recorded only from Rwanda and Zaire.</paragraph>
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</treatment>
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179
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<mods:title id="5026784191FECCE81B2D13497F4F14E5">Review of the species of Coleocydnus J. A. Lis, with a description of C. sicardi n. sp. from Madagascar (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<mods:namePart id="2E0EA46401C16F6441890C6FF3B1DFEA">Lis, J. A.</mods:namePart>
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<mods:date id="3FE1EF84C125E544D2EBF6916C8D336E">2001</mods:date>
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<heading id="D096271CFFC7FFE0FE2DAEB7FC468480" bold="true" box="[491,795,229,253]" centered="true" fontSize="10" level="1" pageId="4" pageNumber="433" reason="4">
<emphasis id="B9154C62FFC7FFE0FE2DAEB7FC468480" bold="true" box="[491,795,229,253]" pageId="4" pageNumber="433">
<taxonomicName id="4C61EBF3FFC7FFE0FE2DAEB7FD898480" authority="Lis, 2001" authorityName="Lis" authorityYear="2001" box="[491,724,229,253]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="sicardi" status="sp. nov.">
<emphasis id="B9154C62FFC7FFE0FE2DAEB7FD898480" bold="true" box="[491,724,229,253]" italics="true" pageId="4" pageNumber="433">Coleocydnus sicardi</emphasis>
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<taxonomicNameLabel id="A226F119FFC7FFE0FD1AAEB7FC468480" box="[732,795,229,253]" pageId="4" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BDE9070FFC7FFE0FD8FAF57FDE18565" blockId="4.[189,1096,229,697]" box="[585,700,261,280]" pageId="4" pageNumber="433">(Figs 3, 8-9)</paragraph>
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<paragraph id="8BDE9070FFC7FFE0FF2DAF12FD7A864A" blockId="4.[189,1096,229,697]" pageId="4" pageNumber="433">
DIAGNOSIS This new species is similar to
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<emphasis id="B9154C62FFC7FFE0FDF8AF32FC778505" box="[574,810,352,376]" italics="true" pageId="4" pageNumber="433">C. madagascariensis</emphasis>
</taxonomicName>
in its general outline and body size, but can be easily separated from the latter by the convex costa, separated from exocorium in its basal two thirds (costa of
<taxonomicName id="4C61EBF3FFC7FFE0FC82AFF2FB7285C5" box="[836,1071,416,440]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9154C62FFC7FFE0FC82AFF2FB7285C5" box="[836,1071,416,440]" italics="true" pageId="4" pageNumber="433">C. madagascariensis</emphasis>
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is flattened and separated from exocorium only in its basal third). Moreover, the corium of
<taxonomicName id="4C61EBF3FFC7FFE0FEF2AFB2FEF88585" authorityName="Lis" authorityYear="2001" box="[308,421,480,504]" class="Insecta" family="Cydnidae" genus="Coleocydnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="sicardi">
<emphasis id="B9154C62FFC7FFE0FEF2AFB2FEF88585" box="[308,421,480,504]" italics="true" pageId="4" pageNumber="433">C. sicardi</emphasis>
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bears coarse large punctures of the same size, whereas the corium of
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<emphasis id="B9154C62FFC7FFE0FEF4AC52FD438665" box="[306,542,512,536]" italics="true" pageId="4" pageNumber="433">C. madagascariensis</emphasis>
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has punctures of different sizes, from very small and shallow to large and coarse.
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Body (Fig. 3) castaneous to piceous,
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in length,
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in width.
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3.
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<emphasis id="B9154C62FFC7FFE0FE2AA813FDFA8229" box="[492,679,1601,1620]" italics="true" pageId="4" pageNumber="433">Coleocydnus sicardi</emphasis>
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: habitus
</paragraph>
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4-5.
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<emphasis id="B9154C62FFC6FFE1FF2EA85CFECA825C" box="[232,407,1550,1569]" italics="true" pageId="5" pageNumber="434">Coleocydnus ernsti</emphasis>
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, 6-7.
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<emphasis id="B9154C62FFC6FFE1FE0BA85CFDB0825C" box="[461,749,1550,1569]" italics="true" pageId="5" pageNumber="434">Coleocydnus madagascariensis</emphasis>
J. A.
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, 8-9.
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<emphasis id="B9154C62FFC6FFE1FCB0A85CFB6C825C" box="[886,1073,1550,1569]" italics="true" pageId="5" pageNumber="434">Coleocydnus sicardi</emphasis>
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<taxonomicNameLabel id="A226F119FFC6FFE1FBF1A85CFF878246" pageId="5" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
: 4, 8 – paramere, 5, 7, 9 – spermatheca (b – bulb, p – pump, df – distal flange, pf – proximal flange), 6 – evaporatoria
</paragraph>
</caption>
<paragraph id="8BDE9070FFC5FFE2FF2DAEB7FECF85C0" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">
Head impunctate or with a few tiny, hardly visible punctures; clypeus as long as paraclypei, entirely free, without a pair of submarginal setigerous punctures; each paraclypeus with a submarginal row of 3 setigerous punctures bearing hair-like setae; eyes moderately large, pale brown to reddish-brown, ocular index 3.20- 3.80; antennae yellowish-brown to dark brown, 3
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antennal segment 1.00-1.14 times longer than the 2
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segment; rostrum pale brown to dark brown, extending beyond mid coxae.
</paragraph>
<paragraph id="8BDE9070FFC5FFE2FF2DAF94FD258620" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">Pronotum subquadrate, its disc almost impunctate, bearing only a few irregularly scattered medium-sized punctures; each lateral margin with a submarginal row of 5 setigerous punctures bearing hair-like setae. Propleuron alutaceous, wrinkled on its anterior convexity, and bearing numerous coarse punctures in the depression and the posterior convexity.</paragraph>
<paragraph id="8BDE9070FFC5FFE2FF2DAC34FE6C86C0" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">Scutellum very long, strongly narrowed apically, its disc convex, impunctate except for a basal row of small punctures and lateral rows of large, deep, coarse punctures.</paragraph>
<paragraph id="8BDE9070FFC5FFE2FF2DAC94FD0F8703" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">Corium coarsely punctate with large, deep punctures, its basal and medial part sparsely punctate or impunctate (Fig. 3); clavo-corial suture absent, exocorium with puncturation denser than that of remaining parts of corium; costa convex, separated from exocorium in its basal two thirds, and bearing a single (sporadically two) setigerous puncture; membrane brown, small, triangular, reaching or almost reaching the tip of abdomen.</paragraph>
<paragraph id="8BDE9070FFC5FFE2FF2DADD5FEE487C3" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">Legs brown to dark brown, tarsi yellowish-brown to pale brown, tibial spines castaneous to piceous.</paragraph>
<paragraph id="8BDE9070FFC5FFE2FF2DAD94FCCF8062" blockId="6.[189,1097,229,1595]" pageId="6" pageNumber="435">Abdominal sterna conspicuously coarsely punctate laterally. Male paramere as in Fig. 8; aedeagus with median carina ventrally. Female spermatheca with small proximal flange, and slightly dilated pump region (Fig. 9).</paragraph>
</subSubSection>
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: [MADAGASCAR]:
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, Coll.
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, 1930, Muséum Paris (
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).
</materialsCitation>
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<typeStatus id="54DA2ED2FFC5FFE2FEBDAADAFEB080DD" box="[379,493,1160,1184]" pageId="6" pageNumber="435">Paratypes</typeStatus>
:
<specimenCount id="9D675BF9FFC5FFE2FE3BAAD5FD0A80E2" box="[509,599,1159,1183]" count="2" pageId="6" pageNumber="435" type="male">2 males</specimenCount>
<specimenCount id="9D675BF9FFC5FFE2FDA5AAD5FD8A80E2" box="[611,727,1159,1183]" count="2" pageId="6" pageNumber="435" type="female">2 females</specimenCount>
, the same data as the holotype (
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,
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)
</materialsCitation>
;
<materialsCitation id="3B099A2DFFC5FFE2FE48AAF5FC1B80A2" collectionCode="MNHN" country="Madagascar" location="Andevorante et Beforona" pageId="6" pageNumber="435" specimenCount="1" specimenCount-female="1" stateProvince="Tamatave">
<specimenCount id="9D675BF9FFC5FFE2FE48AAF5FEAD80C2" box="[398,496,1191,1215]" count="1" pageId="6" pageNumber="435" type="female">1 female</specimenCount>
,
<collectingCountry id="F376D0E0FFC5FFE2FE3DAAF5FDDF80C2" box="[507,642,1191,1215]" name="Madagascar" pageId="6" pageNumber="435" value="Madagascar">Madagascar</collectingCountry>
Est, Régions de
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,
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, Dr G.
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, 1905, Muséum Paris (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37BC3FBFFC5FFE2FF2DAB5AFC8F813D" pageId="6" pageNumber="435" type="etymology">
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The species is named after the collecter of the
<typeStatus id="54DA2ED2FFC5FFE2FD31AB7AFC04813D" box="[759,857,1320,1344]" pageId="6" pageNumber="435">holotype</typeStatus>
specimen.
</paragraph>
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</treatment>
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