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<document id="EEA76E76883FDD7C2EF17ACA764A37A9" ID-DOI="10.15298/rusentj.31.2.20" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178175" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593715551" checkinUser="felipe" docAuthor="Gavryushin, D. I." docDate="2022" docId="0363D218FFC8FF9DFEEEFE86FB04F8B5" docLanguage="en" docName="RussEntomolJ.31.2.201-207.pdf" docOrigin="Russian Entomological Journal 31 (2)" docSource="http://dx.doi.org/10.15298/rusentj.31.2.20" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Tarnania dziedzickii" docType="treatment" docVersion="2" lastPageNumber="205" masterDocId="FF5AAA60FFCBFF99FFF8FFBAFFA9FFEE" masterDocTitle="Contribution to the knowledge of the fauna of fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) from the Republic of Dagestan" masterLastPageNumber="207" masterPageNumber="201" pageNumber="204" updateTime="1722633404845" updateUser="ExternalLinkService">
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<mods:title id="0E552E5D1A62763453C4CBBE1B964935">Contribution to the knowledge of the fauna of fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) from the Republic of Dagestan</mods:title>
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<mods:title id="4DBB81CE76BDCE6CAF1A8C5E96CA1BC7">Russian Entomological Journal</mods:title>
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<mods:date id="9F1E5D04208CA918F4161A5420424E98">2022</mods:date>
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<mods:identifier id="287AE991FAC9E8BBF4A6159B0D767730" type="DOI">10.15298/rusentj.31.2.20</mods:identifier>
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<treatment id="0363D218FFC8FF9DFEEEFE86FB04F8B5" LSID="urn:lsid:plazi:treatment:0363D218FFC8FF9DFEEEFE86FB04F8B5" httpUri="http://treatment.plazi.org/id/0363D218FFC8FF9DFEEEFE86FB04F8B5" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<subSubSection id="C3D03085FFC8FF9AFEEEFE86FD07FEBD" box="[278,686,315,339]" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="8B75630EFFC8FF9AFEEEFE86FD07FEBD" blockId="3.[189,775,315,425]" box="[278,686,315,339]" pageId="3" pageNumber="204">
<heading id="D03DD462FFC8FF9AFEEEFE86FD07FEBD" box="[278,686,315,339]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="204" reason="8">
<taxonomicName id="4CCA188DFFC8FF9AFEEEFE86FD07FEBD" ID-CoL="7BLSR" authority="(Edwards, 1941)" baseAuthorityName="Edwards" baseAuthorityYear="1941" box="[278,686,315,339]" class="Insecta" family="Mycetophilidae" genus="Tarnania" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="dziedzickii">
<emphasis id="B9BEBF1CFFC8FF9AFEEEFE86FE5BFEBD" box="[278,498,315,339]" italics="true" pageId="3" pageNumber="204">Tarnania dziedzickii</emphasis>
(Edwards, 1941)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3D03085FFC8FF9AFE5AFEE3FD8BFE9E" box="[418,546,344,369]" pageId="3" pageNumber="204" type="description">
<paragraph id="8B75630EFFC8FF9AFE5AFEE3FD8BFE9E" blockId="3.[189,775,315,425]" box="[418,546,344,369]" pageId="3" pageNumber="204">
<figureCitation id="13F17F8BFFC8FF9AFE5AFEE3FDB4FE9E" box="[418,541,344,369]" captionStart="Figs 13–21" captionStartId="4.[226,263,1921,1940]" captionTargetBox="[204,1375,338,1837]" captionTargetId="figure-11@4.[189,1399,314,1863]" captionTargetPageId="4" captionText="Figs 13–21. Male genitalia: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — gonostylus; 14–16, 20–21 — hypopygium; 18–19 — hypopygium (tergite 9 and cerci removed); 13, 17 — inner view; 14, 19–20 — dorsal view; 15, 16, 21 — ventral view; 18 — lateral view. Not to the same scale. Рис. 13–21. Гениталии самца: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — гоностиль; 14–16, 20–21 — гипопигий; 18–19 — гипопигий (тергит 9 и церки удалены); 13, 17 — иЗнутри; 14, 19–20 — сверху; 15, 16, 21 — сниЗу; 18 — сбоку. МасШтаб раЗный." figureDoi="http://doi.org/10.5281/zenodo.13178185" httpUri="https://zenodo.org/record/13178185/files/figure.png" pageId="3" pageNumber="204">Figs 14–15</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3D03085FFC8FF9AFF1AFEC4FE3CFE47" pageId="3" pageNumber="204" type="materials_examined">
<paragraph id="8B75630EFFC8FF9AFF1AFEC4FE3CFE47" blockId="3.[189,775,315,425]" pageId="3" pageNumber="204">
MATERIAL.
<specimenCount id="9DCCA887FFC8FF9AFEA3FEC4FE27FE7F" box="[347,398,379,401]" count="2" pageId="3" pageNumber="204" type="male">2♂♂</specimenCount>
, [5],
<date id="FF7445CEFFC8FF9AFE44FEC4FDB3FE7F" box="[444,538,382,401]" pageId="3" pageNumber="204" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFC8FF9AFDDAFEC4FDFCFE7F" box="[546,597,379,401]" count="2" pageId="3" pageNumber="204" type="male">2♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFC8FF9AFD5EFEC4FCAAFE7F" box="[678,771,382,401]" pageId="3" pageNumber="204" value="2021-10-18">18.X.2021</date>
;
<specimenCount id="9DCCA887FFC8FF9AFF45FE2CFF72FE47" box="[189,219,403,425]" count="1" pageId="3" pageNumber="204" type="male">1♂</specimenCount>
, ibidem,
<date id="FF7445CEFFC8FF9AFECBFE2CFE38FE47" box="[307,401,406,425]" pageId="3" pageNumber="204" value="2021-10-19">19.X.2021</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3D03085FFC8FF9DFCAAFE81FCABF8B5" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="distribution">
<paragraph id="8B75630EFFC8FF9AFCAAFE81FB00FE4F" blockId="3.[812,1399,315,417]" pageId="3" pageNumber="204">
DISTRIBUTION. Western Palaearctic:
<collectingCountry id="F3DD239EFFC8FF9AFB21FE81FA84FEBF" box="[1241,1325,315,337]" name="Portugal" pageId="3" pageNumber="204">Portugal</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFAC1FE81FADBFEBF" box="[1337,1394,315,337]" name="Spain" pageId="3" pageNumber="204">Spain</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFCD4FEECFCF8FE85" box="[812,849,342,363]" name="United Kingdom" pageId="3" pageNumber="204">UK</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFCA4FEECFC0AFE82" box="[860,931,342,364]" name="Ireland" pageId="3" pageNumber="204">Ireland</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC56FEECFC5BFE85" box="[942,1010,342,363]" name="France" pageId="3" pageNumber="204">France</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC04FEECFBFBFE82" box="[1020,1106,342,364]" name="Belgium" pageId="3" pageNumber="204">Belgium</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFBA4FEECFB11FE85" box="[1116,1208,342,363]" name="Germany" pageId="3" pageNumber="204">Germany</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFB3BFEECFA90FE82" box="[1219,1337,342,364]" name="Switzerland" pageId="3" pageNumber="204">Switzerland</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFABCFEECFADAFE82" box="[1348,1395,342,364]" name="Italy" pageId="3" pageNumber="204">Italy</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFCD4FECBFCD5FE68" box="[812,892,369,390]" name="Norway" pageId="3" pageNumber="204">Norway</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC71FECBFC71FE69" box="[905,984,369,391]" name="Sweden" pageId="3" pageNumber="204">Sweden</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC1CFECBFB92FE69" box="[996,1083,369,391]" name="Slovakia" pageId="3" pageNumber="204">Slovakia</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFBBFFECBFB2EFE69" box="[1095,1159,369,391]" name="Serbia" pageId="3" pageNumber="204">Serbia</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFB6CFECBFAA4FE68" box="[1172,1293,369,390]" name="Serbia" pageId="3" pageNumber="204">Montenegro</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFAE1FECBFADAFE69" box="[1305,1395,369,391]" name="Romania" pageId="3" pageNumber="204">Romania</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFCD4FE31FC2BFE4F" box="[812,898,395,417]" name="Bulgaria" pageId="3" pageNumber="204">Bulgaria</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC75FE31FC7DFE4E" box="[909,980,395,416]" name="Greece" pageId="3" pageNumber="204">Greece</collectingCountry>
,
<collectingCountry id="F3DD239EFFC8FF9AFC26FE31FBB7FE4F" box="[990,1054,395,417]" name="Russia" pageId="3" pageNumber="204">Russia</collectingCountry>
(first record).
</paragraph>
<paragraph id="8B75630EFFC8FF9DFE8EF8F5FCABF8B5" blockId="3.[374,403,1871,1897]" lastBlockId="4.[352,770,1857,1883]" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<emphasis id="B9BEBF1CFFC8FF9DFE8EF8F5FCABF8B5" bold="true" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">10 11 12 19 20</emphasis>
</paragraph>
</subSubSection>
<caption id="DFB53386FFC8FF9AFF1AF83BFDB1F7FF" ID-DOI="http://doi.org/10.5281/zenodo.13178183" ID-Zenodo-Dep="13178183" httpUri="https://zenodo.org/record/13178183/files/figure.png" pageId="3" pageNumber="204" startId="3.[226,263,1921,1940]" targetBox="[206,1376,467,1859]" targetPageId="3" targetType="figure">
<paragraph id="8B75630EFFC8FF9AFF1AF83BFDCEF82A" blockId="3.[189,1398,1921,2065]" pageId="3" pageNumber="204">
Figs 4–12. Male genitalia: 4–8 —
<taxonomicName id="4CCA188DFFC8FF9AFDF5F838FCA3F87A" authorityName="Vaisanen" authorityYear="1984" box="[525,778,1921,1940]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="matilei" subGenus="Mycomya">
<emphasis id="B9BEBF1CFFC8FF9AFDF5F838FDC9F87A" box="[525,608,1922,1940]" italics="true" pageId="3" pageNumber="204">Mycomya</emphasis>
(
<emphasis id="B9BEBF1CFFC8FF9AFD95F838FD69F87A" box="[621,704,1922,1940]" italics="true" pageId="3" pageNumber="204">Mycomya</emphasis>
)
<emphasis id="B9BEBF1CFFC8FF9AFD36F83BFCA3F87A" box="[718,778,1921,1940]" italics="true" pageId="3" pageNumber="204">matilei</emphasis>
</taxonomicName>
; 9–12 —
<taxonomicName id="4CCA188DFFC8FF9AFC9AF838FBB7F87A" authorityName="Matile" authorityYear="1963" box="[866,1054,1921,1940]" class="Insecta" family="Mycetophilidae" genus="Rymosia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="beaucournui">
<emphasis id="B9BEBF1CFFC8FF9AFC9AF838FBB7F87A" box="[866,1054,1921,1940]" italics="true" pageId="3" pageNumber="204">Rymosia beaucournui</emphasis>
</taxonomicName>
; 4–5 — sternal synsclerite; 6 — tergite 9 and proctiger; 7–11 — hypopygium; 12 — tergite 9, cerci and aedeagus; 4, 6, 9, 12 — dorsal view; 5, 8, 10 — ventral view; 7 — lateral view; 11 — caudal view. Not to the same scale.
</paragraph>
<paragraph id="8B75630EFFC8FF9AFF1AF874FDB1F7FF" blockId="3.[189,1398,1921,2065]" pageId="3" pageNumber="204">
Рис. 4–12. Гениталии самцов: 4–8 —
<taxonomicName id="4CCA188DFFC8FF9AFDCEF874FC98F80E" authorityName="Vaisanen" authorityYear="1984" box="[566,817,1997,2016]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="matilei" subGenus="Mycomya">
<emphasis id="B9BEBF1CFFC8FF9AFDCEF874FD20F80E" box="[566,649,1998,2016]" italics="true" pageId="3" pageNumber="204">Mycomya</emphasis>
(
<emphasis id="B9BEBF1CFFC8FF9AFD6EF874FD40F80E" box="[662,745,1998,2016]" italics="true" pageId="3" pageNumber="204">Mycomya</emphasis>
)
<emphasis id="B9BEBF1CFFC8FF9AFD0DF877FC98F80E" box="[757,817,1997,2016]" italics="true" pageId="3" pageNumber="204">matilei</emphasis>
</taxonomicName>
; 9–12 —
<taxonomicName id="4CCA188DFFC8FF9AFC7FF874FBEAF80E" authorityName="Matile" authorityYear="1963" box="[903,1091,1997,2016]" class="Insecta" family="Mycetophilidae" genus="Rymosia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="beaucournui">
<emphasis id="B9BEBF1CFFC8FF9AFC7FF874FBEAF80E" box="[903,1091,1997,2016]" italics="true" pageId="3" pageNumber="204">Rymosia beaucournui</emphasis>
</taxonomicName>
; 4–5 — стернальный синсклерит, сверху; 6 — тергит 9 и проктигер; 7–11 — гипопигий; 12 — тергит 9, церки и Эдеагус; 4, 6, 9, 12 — сверху; 5, 8, 10 — сниЗу; 7 — сбоку; 11 — сЗади. МасШтаб раЗный.
</paragraph>
</caption>
<subSubSection id="C3D03085FFCFFF9DFB68F8FBFB04F8B5" box="[1168,1197,1857,1883]" pageId="4" pageNumber="205" type="description">
<paragraph id="8B75630EFFCFFF9DFB68F8FBFB04F8B5" blockId="4.[1168,1197,1857,1883]" box="[1168,1197,1857,1883]" pageId="4" pageNumber="205">
<emphasis id="B9BEBF1CFFCFFF9DFB68F8FBFB04F8B5" bold="true" box="[1168,1197,1857,1883]" pageId="4" pageNumber="205">21</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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MATERIAL.
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;
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, [3],
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;
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[4],
<date id="FF7445CEFFC9FF9BFCAAFCDDFC06FC94" box="[850,943,871,890]" pageId="2" pageNumber="203" value="2021-10-14">14.X.2021</date>
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, [5],
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, [5],
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;
<specimenCount id="9DCCA887FFC9FF9BFAC6FCDDFAD8FC94" box="[1342,1393,868,890]" count="2" pageId="2" pageNumber="203" type="male">2♂♂</specimenCount>
,
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<paragraph id="8B75630EFFC9FF9BFCD4FCC4FBD8FC7C" blockId="2.[812,1398,809,940]" box="[812,1137,892,914]" pageId="2" pageNumber="203">
[5],
<date id="FF7445CEFFC9FF9BFCABFCC5FC19FC7C" box="[851,944,895,914]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
;
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, [6],
<date id="FF7445CEFFC9FF9BFBF7FCC5FBC5FC7C" box="[1039,1132,895,914]" pageId="2" pageNumber="203" value="2021-10-17">17.X.2021</date>
.
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Edwards, 1925
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;
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[3],
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;
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;
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, [4],
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;
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, [5],
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;
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, [5],
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;
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, [5],
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.
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<emphasis id="B9BEBF1CFFC9FF9BFC60FB4EFBE6FAE2" box="[920,1103,1268,1292]" italics="true" pageId="2" pageNumber="203">Exechia repanda</emphasis>
Johannsen, 1912
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MATERIAL.
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<emphasis id="B9BEBF1CFFC9FF9BFC68F809FBFBF825" box="[912,1106,1971,1995]" italics="true" pageId="2" pageNumber="203">Synplasta gracilis</emphasis>
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFC35F862FC45F805" box="[973,1004,2005,2027]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [5],
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<emphasis id="B9BEBF1CFFC9FF9BFC6AF92AFBD4F949" box="[914,1149,1679,1703]" italics="true" pageId="2" pageNumber="203">Rymosia beaucournui</emphasis>
Matile, 1963
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<figureCitation id="13F17F8BFFC9FF9BFBE0F916FB2FF92A" box="[1048,1158,1708,1732]" captionStart-0="Figs 4–12" captionStart-1="Figs 13–21" captionStartId-0="3.[226,263,1921,1940]" captionStartId-1="4.[226,263,1921,1940]" captionTargetBox-0="[206,1376,467,1859]" captionTargetBox-1="[204,1375,338,1837]" captionTargetId-0="figure-136@3.[188,1398,455,1873]" captionTargetId-1="figure-11@4.[189,1399,314,1863]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs 4–12. Male genitalia: 4–8 — Mycomya (Mycomya) matilei; 9–12 — Rymosia beaucournui; 4–5 — sternal synsclerite; 6 — tergite 9 and proctiger; 7–11 — hypopygium; 12 — tergite 9, cerci and aedeagus; 4, 6, 9, 12 — dorsal view; 5, 8, 10 — ventral view; 7 — lateral view; 11 — caudal view. Not to the same scale. Рис. 4–12. Гениталии самцов: 4–8 — Mycomya (Mycomya) matilei; 9–12 — Rymosia beaucournui; 4–5 — стернальный синсклерит, сверху; 6 — тергит 9 и проктигер; 7–11 — гипопигий; 12 — тергит 9, церки и Эдеагус; 4, 6, 9, 12 — сверху; 5, 8, 10 — сниЗу; 7 — сбоку; 11 — сЗади. МасШтаб раЗный." captionText-1="Figs 13–21. Male genitalia: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — gonostylus; 14–16, 20–21 — hypopygium; 18–19 — hypopygium (tergite 9 and cerci removed); 13, 17 — inner view; 14, 19–20 — dorsal view; 15, 16, 21 — ventral view; 18 — lateral view. Not to the same scale. Рис. 13–21. Гениталии самца: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — гоностиль; 14–16, 20–21 — гипопигий; 18–19 — гипопигий (тергит 9 и церки удалены); 13, 17 — иЗнутри; 14, 19–20 — сверху; 15, 16, 21 — сниЗу; 18 — сбоку. МасШтаб раЗный." figureDoi-0="http://doi.org/10.5281/zenodo.13178183" figureDoi-1="http://doi.org/10.5281/zenodo.13178185" httpUri-0="https://zenodo.org/record/13178183/files/figure.png" httpUri-1="https://zenodo.org/record/13178185/files/figure.png" pageId="2" pageNumber="203">Figs 9–13</figureCitation>
.
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MATERIAL.
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, [1],
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, [5],
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DISTRIBUTION. Western Palaearctic:
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,
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,
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,
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,
<collectingCountry id="F3DD239EFFC9FF9BFC3DF8BEFBA5F8F7" box="[965,1036,1796,1817]" name="Greece" pageId="2" pageNumber="203">Greece</collectingCountry>
,
<collectingCountry id="F3DD239EFFC9FF9BFBE1F8BEFBC8F8F7" box="[1049,1121,1796,1817]" name="Cyprus" pageId="2" pageNumber="203">Cyprus</collectingCountry>
,
<collectingCountry id="F3DD239EFFC9FF9BFB96F8BEFB4DF8F4" box="[1134,1252,1796,1818]" name="Switzerland" pageId="2" pageNumber="203">Switzerland</collectingCountry>
, North Africa, Near East,
<collectingCountry id="F3DD239EFFC9FF9BFC61F8A4FC70F8DA" box="[921,985,1822,1844]" name="Russia" pageId="2" pageNumber="203">Russia</collectingCountry>
(first record).
</paragraph>
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<taxonomicName id="4CCA188DFFC9FF9BFC98F8F6FAEBF88A" authority="(Stannius, 1831)" baseAuthorityName="Stannius" baseAuthorityYear="1831" box="[864,1346,1868,1892]" class="Insecta" family="Mycetophilidae" genus="Stigmatomeria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B9BEBF1CFFC9FF9BFC98F8F6FB21F88A" box="[864,1160,1868,1892]" italics="true" pageId="2" pageNumber="203">Stigmatomeria crassicornis</emphasis>
(Stannius, 1831)
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFC35F8CBFBA9F86B" box="[973,1024,1903,1925]" count="2" pageId="2" pageNumber="203" type="male">2♂♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFBCBF8CBFB38F86A" box="[1075,1169,1905,1924]" pageId="2" pageNumber="203" value="2021-10-18">18.X.2021</date>
;
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, ibidem,
<date id="FF7445CEFFC9FF9BFAE9F8CBFAC7F86A" box="[1297,1390,1905,1924]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
. GENERAL DISTRIBUTION. Holarctic.
</paragraph>
</subSubSection>
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81
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<emphasis id="B9BEBF1CFFC9FF9BFC79FE86FBDAFEBD" box="[897,1139,315,339]" italics="true" pageId="2" pageNumber="203">Brevicornu griseicolle</emphasis>
(Staeger, 1840)
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFC35FEDAFC45FE9A" box="[973,1004,350,372]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFBE7FEDAFBD5FE9D" box="[1055,1148,352,371]" pageId="2" pageNumber="203" value="2021-10-18">18.X.2021</date>
.
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<paragraph id="8B75630EFFC9FF9BFCAAFEC2FB5BFE60" blockId="2.[850,1313,315,398]" box="[850,1266,376,398]" pageId="2" pageNumber="203">GENERAL DISTRIBUTION. Palaearctic.</paragraph>
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<paragraph id="8B75630EFFC9FF9BFC70FE1DFB48FE17" blockId="2.[850,1359,422,505]" pageId="2" pageNumber="203">
<taxonomicName id="4CCA188DFFC9FF9BFC70FE1DFAB3FE50" authority="(Meigen, 1830)" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[904,1306,422,446]" class="Insecta" family="Mycetophilidae" genus="Brevicornu" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="sericoma">
<emphasis id="B9BEBF1CFFC9FF9BFC70FE1DFBC3FE50" box="[904,1130,423,446]" italics="true" pageId="2" pageNumber="203">Brevicornu sericoma</emphasis>
(Meigen, 1830)
</taxonomicName>
MATERIAL.
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, [5],
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;
<specimenCount id="9DCCA887FFC9FF9BFB63FE71FB13FE30" box="[1179,1210,456,478]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFB15FE71FAE2FE30" box="[1261,1355,459,478]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
. GENERAL DISTRIBUTION. Holarctic.
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Lundström, 1912
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFC35FA5DFC45FA14" box="[973,1004,1508,1530]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
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.
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. GENERAL DISTRIBUTION. Palaearctic.
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MATERIAL.
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<emphasis id="B9BEBF1CFFC9FF9BFEE6FAA4FDA2FAD8" box="[286,523,1310,1334]" italics="true" pageId="2" pageNumber="203">Sciophila quadriterga</emphasis>
Hutson, 1979
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MATERIAL.
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, [4],
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.
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Subfamily
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MATERIAL.
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, [2],
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;
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, [5],
<date id="FF7445CEFFC9FF9BFED8FA65FED5FA1C" box="[288,380,1503,1522]" pageId="2" pageNumber="203" value="2021-10-16">16.X.2021</date>
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, [5],
<date id="FF7445CEFFC9FF9BFE2FFA5AFD9BFA1D" box="[471,562,1504,1523]" pageId="2" pageNumber="203" value="2021-10-18">18.X.2021</date>
;
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, [5],
<date id="FF7445CEFFC9FF9BFD76FA5AFD40FA1D" box="[654,745,1504,1523]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
.
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<emphasis id="B9BEBF1CFFC9FF9BFEFDFB02FE44FB3E" box="[261,493,1208,1232]" italics="true" pageId="2" pageNumber="203">Polylepta guttiventris</emphasis>
(Zetterstedt, 1852)
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFEA5FB67FEDDFB1E" box="[349,372,1242,1264]" count="1" pageId="2" pageNumber="203" type="female">1♀</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFE5FFB67FDADFB1E" box="[423,516,1245,1264]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
. GENERAL DISTRIBUTION. Europe.
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<emphasis id="B9BEBF1CFFC9FF9BFF14FD58FEFDFD17" box="[236,340,738,761]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
(
<emphasis id="B9BEBF1CFFC9FF9BFE9DFD58FE65FD17" box="[357,460,738,761]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
)
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Väisänen, 1984
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<figureCitation id="13F17F8BFFC9FF9BFE57FD45FDB9FCF9" box="[431,528,767,791]" captionStart="Figs 4–12" captionStartId="3.[226,263,1921,1940]" captionTargetBox="[206,1376,467,1859]" captionTargetId="figure-136@3.[188,1398,455,1873]" captionTargetPageId="3" captionText="Figs 4–12. Male genitalia: 4–8 — Mycomya (Mycomya) matilei; 9–12 — Rymosia beaucournui; 4–5 — sternal synsclerite; 6 — tergite 9 and proctiger; 7–11 — hypopygium; 12 — tergite 9, cerci and aedeagus; 4, 6, 9, 12 — dorsal view; 5, 8, 10 — ventral view; 7 — lateral view; 11 — caudal view. Not to the same scale. Рис. 4–12. Гениталии самцов: 4–8 — Mycomya (Mycomya) matilei; 9–12 — Rymosia beaucournui; 4–5 — стернальный синсклерит, сверху; 6 — тергит 9 и проктигер; 7–11 — гипопигий; 12 — тергит 9, церки и Эдеагус; 4, 6, 9, 12 — сверху; 5, 8, 10 — сниЗу; 7 — сбоку; 11 — сЗади. МасШтаб раЗный." figureDoi="http://doi.org/10.5281/zenodo.13178183" httpUri="https://zenodo.org/record/13178183/files/figure.png" pageId="2" pageNumber="203">Figs 4–8</figureCitation>
.
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFEAFFC9EFE20FCD9" box="[343,393,801,823]" count="4" pageId="2" pageNumber="203" type="male">4♂♂</specimenCount>
, [2],
<date id="FF7445CEFFC9FF9BFE4CFC9EFDA7FCD9" box="[436,526,804,823]" pageId="2" pageNumber="203" value="2021-10-11">11.X.2021</date>
;
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, [3],
<date id="FF7445CEFFC9FF9BFDA7FC9EFD10FCD9" box="[607,697,804,823]" pageId="2" pageNumber="203" value="2021-10-12">12.X.2021</date>
;
<specimenCount id="9DCCA887FFC9FF9BFD39FC9EFD76FCD9" box="[705,735,801,823]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [4],
<date id="FF7445CEFFC9FF9BFF45FC86FEB0FCA1" box="[189,281,828,847]" pageId="2" pageNumber="203" value="2021-10-13">13.X.2021</date>
.
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DISTRIBUTION. Europe:
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,
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(first record).
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<emphasis id="B9BEBF1CFFC9FF9BFF1BFCC4FEE2FC7B" box="[227,331,894,917]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
(
<emphasis id="B9BEBF1CFFC9FF9BFEA4FCC4FE6AFC7B" box="[348,451,894,917]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
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<emphasis id="B9BEBF1CFFC9FF9BFE2CFCC7FDB7FC7B" box="[468,542,893,917]" italics="true" pageId="2" pageNumber="203">tumida</emphasis>
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MATERIAL.
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,
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, [1],
<date id="FF7445CEFFC9FF9BFE06FC19FDF3FC58" box="[510,602,931,950]" pageId="2" pageNumber="203" value="2021-10-10">10.X.2021</date>
;
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,
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, [2],
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<date id="FF7445CEFFC9FF9BFF45FC01FEB1FC20" box="[189,280,955,974]" pageId="2" pageNumber="203" value="2021-10-11">11.X.2021</date>
;
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, [2],
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;
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, [3],
<date id="FF7445CEFFC9FF9BFDA4FC01FD11FC20" box="[604,696,955,974]" pageId="2" pageNumber="203" value="2021-10-12">12.X.2021</date>
;
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,
</paragraph>
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<specimenCount id="9DCCA887FFC9FF9BFF45FC69FF76FC08" box="[189,223,976,998]" count="2" pageId="2" pageNumber="203" type="female">2♀♀</specimenCount>
, [3],
<date id="FF7445CEFFC9FF9BFEE1FC69FEDEFC08" box="[281,375,979,998]" pageId="2" pageNumber="203" value="2021-10-12">12.X.2021</date>
;
<specimenCount id="9DCCA887FFC9FF9BFE7DFC69FE0DFC08" box="[389,420,976,998]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
,
<specimenCount id="9DCCA887FFC9FF9BFE4CFC69FE62FC08" box="[436,459,976,998]" count="1" pageId="2" pageNumber="203" type="female">1♀</specimenCount>
, [4],
<date id="FF7445CEFFC9FF9BFDFDFC69FDCBFC08" box="[517,610,979,998]" pageId="2" pageNumber="203" value="2021-10-13">13.X.2021</date>
;
<specimenCount id="9DCCA887FFC9FF9BFD89FC69FD0DFC08" box="[625,676,976,998]" count="7" pageId="2" pageNumber="203" type="male">7♂♂</specimenCount>
,
<specimenCount id="9DCCA887FFC9FF9BFD4CFC69FD7EFC08" box="[692,727,976,998]" count="3" pageId="2" pageNumber="203" type="female">3♀♀</specimenCount>
, [4],
</paragraph>
<paragraph id="8B75630EFFC9FF9BFF45FC51FD2BFBF6" blockId="2.[189,774,893,1048]" pageId="2" pageNumber="203">
<date id="FF7445CEFFC9FF9BFF45FC51FEB1FC10" box="[189,280,1003,1022]" pageId="2" pageNumber="203" value="2021-10-14">14.X.2021</date>
;
<specimenCount id="9DCCA887FFC9FF9BFEDBFC51FEEBFC10" box="[291,322,1000,1022]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFE8DFC51FE78FC10" box="[373,465,1003,1022]" pageId="2" pageNumber="203" value="2021-10-18">18.X.2021</date>
. GENERAL DISTRIBUTION. Palaearctic.
</paragraph>
</subSubSection>
</treatment>
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76
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<taxonomicName id="4CCA188DFFC9FF9BFF34FDC6FD51FD7D" ID-CoL="44X36" authority="(Zetterstedt, 1852)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1852" box="[204,760,635,659]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="flavicollis" subGenus="Mycomya">
<emphasis id="B9BEBF1CFFC9FF9BFF34FDC6FE9DFD7D" box="[204,308,636,659]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
(
<emphasis id="B9BEBF1CFFC9FF9BFEBCFDC6FE02FD7D" box="[324,427,636,659]" italics="true" pageId="2" pageNumber="203">Mycomya</emphasis>
)
<emphasis id="B9BEBF1CFFC9FF9BFE45FDC1FD8EFD7D" box="[445,551,635,659]" italics="true" pageId="2" pageNumber="203">flavicollis</emphasis>
(Zetterstedt, 1852)
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MATERIAL.
<specimenCount id="9DCCA887FFC9FF9BFEA3FD1AFE24FD5D" box="[347,397,669,691]" count="3" pageId="2" pageNumber="203" type="male">3♂♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFE47FD1AFDB3FD5D" box="[447,538,672,691]" pageId="2" pageNumber="203" value="2021-10-18">18.X.2021</date>
;
<specimenCount id="9DCCA887FFC9FF9BFDDDFD1AFDEDFD5D" box="[549,580,669,691]" count="1" pageId="2" pageNumber="203" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFC9FF9BFD8DFD1AFD78FD5D" box="[629,721,672,691]" pageId="2" pageNumber="203" value="2021-10-19">19.X.2021</date>
.
<materialsCitation id="3BA26953FFC9FF9BFF1AFD02FD7CFD20" box="[226,725,696,718]" location="Western Palaearctic" pageId="2" pageNumber="203" specimenCount="1">
GENERAL DISTRIBUTION.
<location id="8E1535D5FFC9FF9BFDE8FD02FD7CFD20" LSID="urn:lsid:plazi:treatment:0363D218FFC9FF9BFF34FDC6FD70FD20:8E1535D5FFC9FF9BFDE8FD02FD7CFD20" box="[528,725,696,718]" name="Western Palaearctic" pageId="2" pageNumber="203">Western Palaearctic</location>
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.
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<emphasis id="B9BEBF1CFFCEFF9CFC4FFBA3FBCAFBDF" box="[951,1123,1049,1073]" italics="true" pageId="5" pageNumber="206">Trichonta foeda</emphasis>
Loew, 1869
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<figureCitation id="13F17F8BFFCEFF9CFBEAFB8CFB24FBA0" box="[1042,1165,1078,1102]" captionStart="Figs 13–21" captionStartId="4.[226,263,1921,1940]" captionTargetBox="[204,1375,338,1837]" captionTargetId="figure-11@4.[189,1399,314,1863]" captionTargetPageId="4" captionText="Figs 13–21. Male genitalia: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — gonostylus; 14–16, 20–21 — hypopygium; 18–19 — hypopygium (tergite 9 and cerci removed); 13, 17 — inner view; 14, 19–20 — dorsal view; 15, 16, 21 — ventral view; 18 — lateral view. Not to the same scale. Рис. 13–21. Гениталии самца: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — гоностиль; 14–16, 20–21 — гипопигий; 18–19 — гипопигий (тергит 9 и церки удалены); 13, 17 — иЗнутри; 14, 19–20 — сверху; 15, 16, 21 — сниЗу; 18 — сбоку. МасШтаб раЗный." figureDoi="http://doi.org/10.5281/zenodo.13178185" httpUri="https://zenodo.org/record/13178185/files/figure.png" pageId="5" pageNumber="206">Figs 20–21</figureCitation>
.
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<paragraph id="8B75630EFFCEFF9CFCAAFBE1FC70FB68" blockId="5.[812,1399,1049,1212]" pageId="5" pageNumber="206">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFC32FBE1FC40FB81" box="[970,1001,1113,1135]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [2],
<date id="FF7445CEFFCEFF9CFBEDFBE1FBDBFB80" box="[1045,1138,1115,1134]" pageId="5" pageNumber="206" value="2021-10-11">11.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFB82FBE1FB04FB81" box="[1146,1197,1113,1135]" count="2" pageId="5" pageNumber="206" type="male">2♂♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFB21FBE1FA9FFB80" box="[1241,1334,1115,1134]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFAC6FBE1FAD8FB81" box="[1342,1393,1113,1135]" count="2" pageId="5" pageNumber="206" type="male">2♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFC80FBC9FC7CFB68" box="[888,981,1139,1158]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3D03085FFCEFF9CFCAAFB31FAE1FB52" pageId="5" pageNumber="206" type="distribution">
<paragraph id="8B75630EFFCEFF9CFCAAFB31FAE1FB52" blockId="5.[812,1399,1049,1212]" pageId="5" pageNumber="206">
DISTRIBUTION. Europe:
<collectingCountry id="F3DD239EFFCEFF9CFBA4FB36FB28FB4F" box="[1116,1153,1164,1185]" name="United Kingdom" pageId="5" pageNumber="206">UK</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFB74FB31FB7AFB4F" box="[1164,1235,1163,1185]" name="Ireland" pageId="5" pageNumber="206">Ireland</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFB25FB31FA90FB4E" box="[1245,1337,1163,1184]" name="Germany" pageId="5" pageNumber="206">Germany</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFABDFB36FCF0FB55" name="Hungary" pageId="5" pageNumber="206">Hungary</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFC9CFB1CFC11FB52" box="[868,952,1190,1212]" name="Portugal" pageId="5" pageNumber="206">Portugal</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFC3CFB1CFBBAFB52" box="[964,1043,1190,1212]" name="Sweden" pageId="5" pageNumber="206">Sweden</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFBE5FB1CFBF4FB52" box="[1053,1117,1190,1212]" name="Russia" pageId="5" pageNumber="206">Russia</collectingCountry>
(first record); Nearctic.
</paragraph>
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MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFC37FB4FFC47FAE6" box="[975,1006,1266,1288]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [3],
<date id="FF7445CEFFCEFF9CFBDEFB4FFB2AFAE6" box="[1062,1155,1269,1288]" pageId="5" pageNumber="206" value="2021-10-12">12.X.2021</date>
;
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, [4],
<date id="FF7445CEFFCEFF9CFB1FFB4FFAEDFAE6" box="[1255,1348,1269,1288]" pageId="5" pageNumber="206" value="2021-10-13">13.X.2021</date>
;
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, ibidem,
<date id="FF7445CEFFCEFF9CFC78FAB7FC4FFACE" box="[896,998,1293,1312]" pageId="5" pageNumber="206" value="2021-10-14">14.X.2021</date>
;
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, [5],
<date id="FF7445CEFFCEFF9CFB97FAB7FB7DFACE" box="[1135,1236,1293,1312]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
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, ibidem,
<date id="FF7445CEFFCEFF9CFCD4FA9FFC20FAD6" box="[812,905,1317,1336]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
;
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, ibidem,
<date id="FF7445CEFFCEFF9CFBD9FA9FFBD7FAD6" box="[1057,1150,1317,1336]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
.
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;
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, [5],
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<subSubSection id="C3D03085FFCEFF9CFC5CFD75FB57FD08" box="[932,1278,718,742]" pageId="5" pageNumber="206" type="nomenclature">
<paragraph id="8B75630EFFCEFF9CFC5CFD75FB57FD08" blockId="5.[812,1399,718,854]" box="[932,1278,718,742]" pageId="5" pageNumber="206">
<heading id="D03DD462FFCEFF9CFC5CFD75FB57FD08" box="[932,1278,718,742]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="206" reason="8">
<taxonomicName id="4CCA188DFFCEFF9CFC5CFD75FB57FD08" ID-CoL="8DTD6" authority="Edwards, 1925" authorityName="Edwards" authorityYear="1925" box="[932,1278,718,742]" class="Insecta" family="Mycetophilidae" genus="Sceptonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="B9BEBF1CFFCEFF9CFC5CFD75FBFCFD08" box="[932,1109,719,742]" italics="true" pageId="5" pageNumber="206">Sceptonia tenuis</emphasis>
Edwards, 1925
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<paragraph id="8B75630EFFCEFF9CFBEAFD56FB38FCEA" blockId="5.[812,1399,718,854]" box="[1042,1169,748,772]" pageId="5" pageNumber="206">
<figureCitation id="13F17F8BFFCEFF9CFBEAFD56FB24FCEA" box="[1042,1165,748,772]" captionStart="Figs 13–21" captionStartId="4.[226,263,1921,1940]" captionTargetBox="[204,1375,338,1837]" captionTargetId="figure-11@4.[189,1399,314,1863]" captionTargetPageId="4" captionText="Figs 13–21. Male genitalia: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — gonostylus; 14–16, 20–21 — hypopygium; 18–19 — hypopygium (tergite 9 and cerci removed); 13, 17 — inner view; 14, 19–20 — dorsal view; 15, 16, 21 — ventral view; 18 — lateral view. Not to the same scale. Рис. 13–21. Гениталии самца: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — гоностиль; 14–16, 20–21 — гипопигий; 18–19 — гипопигий (тергит 9 и церки удалены); 13, 17 — иЗнутри; 14, 19–20 — сверху; 15, 16, 21 — сниЗу; 18 — сбоку. МасШтаб раЗный." figureDoi="http://doi.org/10.5281/zenodo.13178185" httpUri="https://zenodo.org/record/13178185/files/figure.png" pageId="5" pageNumber="206">Figs 18–19</figureCitation>
.
</paragraph>
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<paragraph id="8B75630EFFCEFF9CFCAAFCABFADEFCD2" blockId="5.[812,1399,718,854]" pageId="5" pageNumber="206">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFC37FCABFC47FCCA" box="[975,1006,782,804]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [2],
<date id="FF7445CEFFCEFF9CFBDEFCABFB2AFCCA" box="[1062,1155,785,804]" pageId="5" pageNumber="206" value="2021-10-11">11.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFB69FCABFB19FCCA" box="[1169,1200,782,804]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [3],
<date id="FF7445CEFFCEFF9CFB1FFCABFAECFCCA" box="[1255,1349,785,804]" pageId="5" pageNumber="206" value="2021-10-12">12.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFAAAFCABFAD8FCCA" box="[1362,1393,782,804]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFCA9FC93FC07FCD2" box="[849,942,809,828]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFC40FC93FC42FCD2" box="[952,1003,806,828]" count="2" pageId="5" pageNumber="206" type="male">2♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFBC5FC93FB32FCD2" box="[1085,1179,809,828]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFB5CFC93FB6AFCD2" box="[1188,1219,806,828]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFAEEFC93FADAFCD2" box="[1302,1395,809,828]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
.
</paragraph>
<paragraph id="8B75630EFFCEFF9CFCAAFCFBFB64FCB8" blockId="5.[812,1399,718,854]" box="[850,1229,833,854]" pageId="5" pageNumber="206">GENERAL DISTRIBUTION. Europe.</paragraph>
</subSubSection>
</treatment>
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64
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<emphasis id="B9BEBF1CFFCEFF9CFC62FADDFBE6FA90" box="[922,1103,1383,1406]" italics="true" pageId="5" pageNumber="206">Zygomyia notata</emphasis>
(Stannius, 1831)
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<subSubSection id="C3D03085FFCEFF9CFCAAFA31FB5BFA57" pageId="5" pageNumber="206" type="materials_examined">
<paragraph id="8B75630EFFCEFF9CFCAAFA31FB28FA70" blockId="5.[850,1289,1382,1465]" box="[850,1153,1417,1439]" pageId="5" pageNumber="206">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFC35FA31FC45FA71" box="[973,1004,1417,1439]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFBE7FA31FBD5FA70" box="[1055,1148,1419,1438]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
.
</paragraph>
<paragraph id="8B75630EFFCEFF9CFCAAFA19FB5BFA57" blockId="5.[850,1289,1382,1465]" box="[850,1266,1443,1465]" pageId="5" pageNumber="206">GENERAL DISTRIBUTION. Palaearctic.</paragraph>
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<emphasis id="B9BEBF1CFFCEFF9CFC75FDF7FBC2FD8B" box="[909,1131,589,613]" italics="true" pageId="5" pageNumber="206">Sceptonia humerella</emphasis>
Edwards, 1925
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MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFC35FDC8FC45FD6B" box="[973,1004,623,645]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFBE7FDC8FBD5FD6B" box="[1055,1148,626,645]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
.
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DISTRIBUTION. Europe:
<collectingCountry id="F3DD239EFFCEFF9CFBA1FD30FBD7FD71" box="[1113,1150,650,671]" name="United Kingdom" pageId="5" pageNumber="206">UK</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFB7FFD30FB4AFD71" box="[1159,1251,650,671]" name="Germany" pageId="5" pageNumber="206">Germany</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFB15FD30FCEFFD55" name="Czech Republic" pageId="5" pageNumber="206">Czech Republic</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFCAAFD1FFC28FD55" box="[850,897,677,699]" name="Italy" pageId="5" pageNumber="206">Italy</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFC73FD1FFC48FD55" box="[907,993,677,699]" name="Bulgaria" pageId="5" pageNumber="206">Bulgaria</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFC14FD1FFB85FD55" box="[1004,1068,677,699]" name="Russia" pageId="5" pageNumber="206">Russia</collectingCountry>
(first record).
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Lundström, 1913
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MATERIAL.
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, [2],
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;
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, [3],
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;
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, [4],
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;
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, ibidem,
<date id="FF7445CEFFCEFF9CFB9AFC1DFB69FC54" box="[1122,1216,935,954]" pageId="5" pageNumber="206" value="2021-10-14">14.X.2021</date>
;
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, [5],
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;
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, ibidem,
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;
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, ibidem,
<date id="FF7445CEFFCEFF9CFB55FC05FAA3FC3C" box="[1197,1290,959,978]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
;
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, ibidem,
<date id="FF7445CEFFCEFF9CFCA7FC6DFC15FC04" box="[863,956,983,1002]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
;
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, [6],
<date id="FF7445CEFFCEFF9CFBE2FC6DFBD1FC04" box="[1050,1144,983,1002]" pageId="5" pageNumber="206" value="2021-10-17">17.X.2021</date>
. GENERAL DISTRIBUTION. Palaearctic.
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.
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<taxonomicName id="4CCA188DFFCEFF9CFECCFA35FD26FA49" ID-CoL="4GV2H" authority="Winnertz, 1864" authorityName="Winnertz" authorityYear="1864" box="[308,655,1423,1447]" class="Insecta" family="Mycetophilidae" genus="Phronia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B9BEBF1CFFCEFF9CFECCFA35FE49FA49" box="[308,480,1423,1447]" italics="true" pageId="5" pageNumber="206">Phronia basalis</emphasis>
Winnertz, 1864
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<paragraph id="8B75630EFFCEFF9CFE5DFA16FDB4FA2A" blockId="5.[189,775,1423,1659]" box="[421,541,1452,1476]" pageId="5" pageNumber="206">
<figureCitation id="13F17F8BFFCEFF9CFE5DFA16FDB4FA2A" box="[421,541,1452,1476]" captionStart="Figs 13–21" captionStartId="4.[226,263,1921,1940]" captionTargetBox="[204,1375,338,1837]" captionTargetId="figure-11@4.[189,1399,314,1863]" captionTargetPageId="4" captionText="Figs 13–21. Male genitalia: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — gonostylus; 14–16, 20–21 — hypopygium; 18–19 — hypopygium (tergite 9 and cerci removed); 13, 17 — inner view; 14, 19–20 — dorsal view; 15, 16, 21 — ventral view; 18 — lateral view. Not to the same scale. Рис. 13–21. Гениталии самца: 13 — Rymosia beaucournui; 14— Tarnania dziedzickii; 16–17 — Phronia basalis; 18–19 — Sceptonia tenuis;20–21 — Trichonta foeda; 13, 17 — гоностиль; 14–16, 20–21 — гипопигий; 18–19 — гипопигий (тергит 9 и церки удалены); 13, 17 — иЗнутри; 14, 19–20 — сверху; 15, 16, 21 — сниЗу; 18 — сбоку. МасШтаб раЗный." figureDoi="http://doi.org/10.5281/zenodo.13178185" httpUri="https://zenodo.org/record/13178185/files/figure.png" pageId="5" pageNumber="206">Figs 16–17</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8B75630EFFCEFF9CFF1AFA74FE88F9FF" blockId="5.[189,775,1423,1659]" pageId="5" pageNumber="206">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFEA2FA74FE3EFA0F" box="[346,407,1483,1505]" count="18" pageId="5" pageNumber="206" type="male">18♂♂</specimenCount>
, [4],
<date id="FF7445CEFFCEFF9CFE3AFA74FDB6FA0F" box="[450,543,1486,1505]" pageId="5" pageNumber="206" value="2021-10-13">13.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFDDEFA74FDF0FA0F" box="[550,601,1483,1505]" count="4" pageId="5" pageNumber="206" type="male">4♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFD5EFA74FCAAFA0F" box="[678,771,1486,1505]" pageId="5" pageNumber="206" value="2021-10-14">14.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFF45FA5CFF46FA17" box="[189,239,1507,1529]" count="9" pageId="5" pageNumber="206" type="male">9♂♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFEDAFA5FFE29FA16" box="[290,384,1509,1528]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFE73FA5CFE17FA17" box="[395,446,1507,1529]" count="7" pageId="5" pageNumber="206" type="male">7♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFDEBFA5CFDD9FA17" box="[531,624,1510,1529]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFD83FA5CFD10FA17" box="[635,697,1507,1529]" count="21" pageId="5" pageNumber="206" type="male">21♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFF45FA44FEB5F9FF" box="[189,284,1534,1553]" pageId="5" pageNumber="206" value="2021-10-19">19.X.2021</date>
.
</paragraph>
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<paragraph id="8B75630EFFCEFF9CFF1AF9AFFD25F995" blockId="5.[189,775,1423,1659]" pageId="5" pageNumber="206">
DISTRIBUTION. Western Palaearctic:
<collectingCountry id="F3DD239EFFCEFF9CFD98F9ACFD2CF9C5" box="[608,645,1558,1579]" name="United Kingdom" pageId="5" pageNumber="206">UK</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFD76F9AFFD7CF9C5" box="[654,725,1557,1579]" name="Ireland" pageId="5" pageNumber="206">Ireland</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFD25F9ACFF59F9A8" name="Portugal" pageId="5" pageNumber="206">Portugal</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFF05F98AFE9FF9A8" box="[253,310,1584,1606]" name="Spain" pageId="5" pageNumber="206">Spain</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFEBCF98AFE33F9A8" box="[324,410,1584,1606]" name="Belgium" pageId="5" pageNumber="206">Belgium</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFE50F98BFE45F9A8" box="[424,492,1585,1606]" name="France" pageId="5" pageNumber="206">France</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFE02F98AFD30F9A8" box="[506,665,1584,1606]" name="Netherlands" pageId="5" pageNumber="206">the Netherlands</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFD5FF98AFCAAF9A8" box="[679,771,1584,1606]" name="Denmark" pageId="5" pageNumber="206">Denmark</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFF45F9F1FE9DF98F" box="[189,308,1611,1633]" name="Switzerland" pageId="5" pageNumber="206">Switzerland</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFEBBF9F1FE4FF98F" box="[323,486,1611,1633]" name="Czech Republic" pageId="5" pageNumber="206">Czech Republic</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFE0EF9F1FDE4F98E" box="[502,589,1611,1632]" name="Hungary" pageId="5" pageNumber="206">Hungary</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFDA6F9F1FD0DF98F" box="[606,676,1611,1633]" name="Poland" pageId="5" pageNumber="206">Poland</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFD4CF9F1FCAAF98F" box="[692,771,1611,1633]" name="Sweden" pageId="5" pageNumber="206">Sweden</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFF45F9DFFEBAF995" box="[189,275,1637,1659]" name="Bulgaria" pageId="5" pageNumber="206">Bulgaria</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFEE5F9DFFECDF994" box="[285,356,1637,1658]" name="Greece" pageId="5" pageNumber="206">Greece</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFE97F9DFFE1EF994" box="[367,439,1637,1658]" name="Cyprus" pageId="5" pageNumber="206">Cyprus</collectingCountry>
,
<collectingCountry id="F3DD239EFFCEFF9CFE3AF9DFFDABF995" box="[450,514,1637,1659]" name="Russia" pageId="5" pageNumber="206">Russia</collectingCountry>
(first record).
</paragraph>
</subSubSection>
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80
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<taxonomicName id="4CCA188DFFCEFF9CFED1F879FD33F835" ID-CoL="4GV3Y" authority="Winnertz, 1864" authorityName="Winnertz" authorityYear="1864" box="[297,666,1987,2011]" class="Insecta" family="Mycetophilidae" genus="Phronia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis id="B9BEBF1CFFCEFF9CFED1F879FE43F835" box="[297,490,1987,2011]" italics="true" pageId="5" pageNumber="206">Phronia forcipata</emphasis>
Winnertz, 1864
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<paragraph id="8B75630EFFCEFF9CFF1AF852FE67F7FD" blockId="5.[189,775,1987,2067]" pageId="5" pageNumber="206">
<materialsCitation id="3BA26953FFCEFF9CFF1AF852FE67F7FD" collectingDate="2021-06-22" collectorName="Oleg Kosterin" elevation="1300" latitude="41.95" location="Kuzhnik" longLatPrecision="4168" longitude="47.8" pageId="5" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFEA4F852FED2F812" box="[348,379,2022,2044]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
,
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,
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,
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,
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<collectingDate id="EF30BC26FFCEFF9CFF45F7BAFE89F7FD" box="[189,288,2048,2067]" pageId="5" pageNumber="206" value="2021-06-22">22.VI.2021</collectingDate>
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,
<collectorName id="263F06D8FFCEFF9CFED3F7BAFE0FF7FD" box="[299,422,2048,2067]" pageId="5" pageNumber="206">Oleg Kosterin</collectorName>
leg.
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<paragraph id="8B75630EFFCEFF9CFCAAFE81FB5BFEBF" blockId="5.[850,1266,315,337]" box="[850,1266,315,337]" pageId="5" pageNumber="206">GENERAL DISTRIBUTION. Palaearctic.</paragraph>
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<emphasis id="B9BEBF1CFFCEFF9CFEF4FB91FDA1FBAD" box="[268,520,1067,1091]" italics="true" pageId="5" pageNumber="206">Mycetophila perpallida</emphasis>
Chandler, 1993
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MATERIAL.
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, [4],
<date id="FF7445CEFFCEFF9CFE35FBEAFD83FB8D" box="[461,554,1104,1123]" pageId="5" pageNumber="206" value="2021-10-13">13.X.2021</date>
;
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, [5],
<date id="FF7445CEFFCEFF9CFD5DFBEAFCAAFB8D" box="[677,771,1104,1123]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFF45FBD2FF46FB95" box="[189,239,1125,1147]" count="2" pageId="5" pageNumber="206" type="male">2♂♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFEBEFBD2FE0AFB95" box="[326,419,1128,1147]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
;
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, [6],
<date id="FF7445CEFFCEFF9CFDEEFBD2FDDAFB95" box="[534,627,1128,1147]" pageId="5" pageNumber="206" value="2021-10-17">17.X.2021</date>
.
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<emphasis id="B9BEBF1CFFCEFF9CFEF7FE79FE5AFE35" box="[271,499,451,475]" italics="true" pageId="5" pageNumber="206">Mycetophila formosa</emphasis>
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MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFEA5FE52FED5FE15" box="[349,380,485,507]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [5],
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. GENERAL DISTRIBUTION. Palaearctic.
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<subSubSection id="C3D03085FFCEFF9CFF05FB10FD6CFB2C" box="[253,709,1194,1218]" pageId="5" pageNumber="206" type="nomenclature">
<paragraph id="8B75630EFFCEFF9CFF05FB10FD6CFB2C" blockId="5.[189,775,1194,1301]" box="[253,709,1194,1218]" pageId="5" pageNumber="206">
<heading id="D03DD462FFCEFF9CFF05FB10FD6CFB2C" box="[253,709,1194,1218]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="206" reason="8">
<taxonomicName id="4CCA188DFFCEFF9CFF05FB10FD6CFB2C" ID-CoL="44VJ5" authority="Dziedzicki, 1884" authorityName="Dziedzicki" authorityYear="1884" box="[253,709,1194,1218]" class="Insecta" family="Mycetophilidae" genus="Mycetophila" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="signatoides">
<emphasis id="B9BEBF1CFFCEFF9CFF05FB10FDACFB2C" box="[253,517,1194,1218]" italics="true" pageId="5" pageNumber="206">Mycetophila signatoides</emphasis>
Dziedzicki, 1884
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3D03085FFCEFF9CFF1AFB75FDDBFAFB" pageId="5" pageNumber="206" type="materials_examined">
<paragraph id="8B75630EFFCEFF9CFF1AFB75FDCAFB14" blockId="5.[189,775,1194,1301]" pageId="5" pageNumber="206">
MATERIAL.
<specimenCount id="9DCCA887FFCEFF9CFEA6FB75FED4FB0C" box="[350,381,1228,1250]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, [3],
<date id="FF7445CEFFCEFF9CFE49FB75FDA6FB0C" box="[433,527,1231,1250]" pageId="5" pageNumber="206" value="2021-10-12">12.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFDE2FB75FDE4FB0C" box="[538,589,1228,1250]" count="4" pageId="5" pageNumber="206" type="male">4♂♂</specimenCount>
,
<specimenCount id="9DCCA887FFCEFF9CFDA2FB75FDD8FB0C" box="[602,625,1228,1250]" count="1" pageId="5" pageNumber="206" type="female">1♀</specimenCount>
, [4],
<date id="FF7445CEFFCEFF9CFD5DFB75FCABFB0C" box="[677,770,1231,1250]" pageId="5" pageNumber="206" value="2021-10-13">13.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFF45FB5DFF46FB14" box="[189,239,1252,1274]" count="2" pageId="5" pageNumber="206" type="male">2♂♂</specimenCount>
, [5],
<date id="FF7445CEFFCEFF9CFEDBFB5DFE28FB14" box="[291,385,1255,1274]" pageId="5" pageNumber="206" value="2021-10-16">16.X.2021</date>
;
<specimenCount id="9DCCA887FFCEFF9CFE74FB5DFE02FB14" box="[396,427,1252,1274]" count="1" pageId="5" pageNumber="206" type="male">1♂</specimenCount>
, ibidem,
<date id="FF7445CEFFCEFF9CFDFAFB5DFDF6FB14" box="[514,607,1255,1274]" pageId="5" pageNumber="206" value="2021-10-18">18.X.2021</date>
.
</paragraph>
<paragraph id="8B75630EFFCEFF9CFF1AFB45FDDBFAFB" blockId="5.[189,775,1194,1301]" box="[226,626,1279,1301]" pageId="5" pageNumber="206">GENERAL DISTRIBUTION. Holarctic.</paragraph>
</subSubSection>
</treatment>
</document>

481
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<taxonomicName id="4C2A4D2542165514FC84FD6FE74DD409" ID-CoL="BM4BV" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[840,1305,744,769]" class="Insecta" family="Sesiidae" genus="Bembecia" isSensuStricto="true" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="khovdensis" status="sp. nov.">
<emphasis id="B95EEAB442165514FC84FD6FE1E5D409" box="[840,945,744,768]" italics="true" pageId="1" pageNumber="199">Bembecia</emphasis>
(s. str.)
<emphasis id="B95EEAB442165514FBC0FD6FE6D5D409" box="[1036,1153,744,768]" italics="true" pageId="1" pageNumber="199">khovdensis</emphasis>
O. Gorbunov
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,
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<taxonomicNameLabel id="A26D57CF42165514FAE9FD6FE70ED409" box="[1317,1370,744,768]" pageId="1" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
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<subSubSection id="C330652D42165514FC3FFC81E6E4D417" box="[1011,1200,774,798]" pageId="1" pageNumber="199" type="description">
<paragraph id="8B9536A642165514FC3FFC81E6E4D417" blockId="1.[812,1399,744,2067]" box="[1011,1200,774,798]" pageId="1" pageNumber="199">
<figureCitation id="13112A2342165514FC3FFC81E606D417" box="[1011,1106,774,798]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="1" pageNumber="199">Figs 1–2</figureCitation>
,
<figureCitation id="13112A2342165514FB93FC81E6DDD417" box="[1119,1161,774,798]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="1" pageNumber="199">5–9</figureCitation>
,
<figureCitation id="13112A2342165514FB59FC81E6E4D417" box="[1173,1200,774,798]" captionStart="Fig" captionStartId="6.[226,257,1969,1988]" captionTargetBox="[189,1399,1147,1951]" captionTargetId="figure-693@6.[189,1399,1147,1951]" captionTargetPageId="6" captionText="Fig. 15. The type locality of Bembecia khovdensis O. Gorbunov, sp.n.: Mongolia, Khovd Aimag, 8 km N Uyench, 1800 m, 46°07´N, 092°03´E, 22.VI.2007. Photo by O. Gorbunov. Рис. 15. Типовое местонахождение Bembecia khovdensis O. Gorbunov, sp.n.: МонголиЯ, Кобдоский аймак, 8 км С Уенча, 1800 м, 46°07´с.Ш., 092°03´в.д., 22.VI.2007. Фото О. Горбунов." pageId="1" pageNumber="199">15</figureCitation>
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<paragraph id="8B9536A642165514FC9EFCACE13DD4C7" blockId="1.[812,1399,744,2067]" pageId="1" pageNumber="199">
<materialsCitation id="3B423CFB42165514FC9EFCACE13DD4C7" collectingDate="2007-06-22" collectorName="O. Gorbunov" country="Mongolia" elevation="1800" latitude="46.116665" location="Uyench" longLatPrecision="1125" longitude="92.05" pageId="1" pageNumber="199" specimenCount="2" specimenCount-male="2" stateProvince="Hovd" typeStatus="holotype">
MATERIAL.
<emphasis id="B95EEAB442165514FC01FCACE674D437" bold="true" box="[973,1056,811,830]" pageId="1" pageNumber="199">
<typeStatus id="5491880442165514FC01FCACE674D437" box="[973,1056,811,830]" pageId="1" pageNumber="199" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D2CFD2F42165514FBE5FCAAE669D434" box="[1065,1085,813,829]" count="1" pageId="1" pageNumber="199" type="male">♂</specimenCount>
(
<figureCitation id="13112A2342165514FB86FCACE6CCD437" box="[1098,1176,811,830]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="1" pageNumber="199">Figs 1–2</figureCitation>
) with labels: “
<collectingCountry id="F33D763642165514FAD7FCACE727D437" box="[1307,1395,811,830]" name="Mongolia" pageId="1" pageNumber="199">Mongolia</collectingCountry>
,
<collectingRegion id="49EEF84442165514FCE0FCC4E1FFD45F" box="[812,939,835,854]" country="Mongolia" name="Hovd" pageId="1" pageNumber="199">Khovd Aimag</collectingRegion>
, /
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N
<location id="8EF5607D42165514FBC2FCC4E607D45F" LSID="urn:lsid:plazi:treatment:038387B042165511FC84FD6FE02AD378:8EF5607D42165514FBC2FCC4E607D45F" box="[1038,1107,835,854]" country="Mongolia" latitude="46.116665" longLatPrecision="1125" longitude="92.05" name="Uyench" pageId="1" pageNumber="199" stateProvince="Hovd">Uyench</location>
, /
<quantity id="4CD29B4342165514FBA6FCC4E6E4D45F" box="[1130,1200,835,854]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="199" unit="m" value="1800.0">
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</quantity>
,
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,
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, /
<date id="FF94106642165514FCE0FCDCE1C5D467" box="[812,913,859,878]" pageId="1" pageNumber="199" value="2007-06-22">
<collectingDate id="EFD0E98E42165514FCE0FCDCE1C5D467" box="[812,913,859,878]" pageId="1" pageNumber="199" value="2007-06-22">22.VI.2007</collectingDate>
</date>
, /
<collectorName id="26DF537042165514FC68FCDCE641D467" box="[932,1045,859,878]" pageId="1" pageNumber="199">O. Gorbunov</collectorName>
leg.”; “
<taxonomicName id="4C2A4D2542165514FB9AFCDCE6ECD467" authorityName="Boisduval" authorityYear="1828" box="[1110,1208,859,878]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="family">SESIIDAE</taxonomicName>
/ Pictures Nos / 0033- 0034–2023 / Photo by
<collectorName id="26DF537042165514FC3AFCF4E639D48F" box="[1014,1133,883,902]" pageId="1" pageNumber="199">O. Gorbunov</collectorName>
”; “Genitalia examined / by
<collectorName id="26DF537042165514FAADFCF4E1D7D497" pageId="1" pageNumber="199">O. Gorbunov</collectorName>
/ Preparation No OG–008-2023”; “
<typeStatus id="5491880442165514FB04FC0CE71ED497" box="[1224,1354,907,926]" pageId="1" pageNumber="199" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="9D2CFD2F42165514FA9FFC0AE733D494" box="[1363,1383,909,925]" count="1" pageId="1" pageNumber="199" type="male">♂</specimenCount>
/
<taxonomicName id="4C2A4D2542165514FCE0FC24E1BDD4BF" authorityName="O. Gorbunov" authorityYear="2023" box="[812,1001,931,950]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="khovdensis">
<emphasis id="B95EEAB442165514FCE0FC24E1BDD4BF" box="[812,1001,931,950]" italics="true" pageId="1" pageNumber="199">Bembecia khovdensis</emphasis>
</taxonomicName>
/ O.
<bibRefCitation id="EFBB4B5742165514FBEEFC24E6E0D4BF" author="Gorbunov O. G." box="[1058,1204,931,950]" pageId="1" pageNumber="199" pagination="68 - 75" refId="ref7645" refString="Gorbunov O. G. 2023. A new species of the genus Bembecia Hubner, 1819 [&quot; 1816 &quot;] (Lepidoptera: Sesiidae) from the Gissar Mountain Range // Russian Entomol. J. Vol. 32. No. 1. P. 68 - 75." type="journal article" year="2023">Gorbunov, 2023</bibRefCitation>
/
<collectorName id="26DF537042165514FB02FC24E710D4BF" box="[1230,1348,931,950]" pageId="1" pageNumber="199">O. Gorbunov</collectorName>
des., 2022”.
</materialsCitation>
</paragraph>
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<paragraph id="8B9536A642165514FC9EFC54E612D314" blockId="1.[812,1399,744,2067]" pageId="1" pageNumber="199">
DESCRIPTION.
<emphasis id="B95EEAB442165514FC32FC54E666D4E1" bold="true" box="[1022,1074,979,1000]" pageId="1" pageNumber="199">Male</emphasis>
(
<typeStatus id="5491880442165514FB8FFC54E6CFD4E0" box="[1091,1179,979,1001]" pageId="1" pageNumber="199" type="holotype">holotype</typeStatus>
) (
<figureCitation id="13112A2342165514FB7EFC54E759D4E1" box="[1202,1293,979,1001]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="1" pageNumber="199">Figs 1–2</figureCitation>
). Alar expanse
<quantity id="4CD29B4342165514FCA1FC6AE19CD30A" box="[877,968,1005,1027]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="1" pageNumber="199" unit="mm" value="14.9">14.9 mm</quantity>
; body length 9.0 mm; forewing length
<quantity id="4CD29B4342165514FA94FC6AE100D314" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="199" unit="mm" value="6.9">6.9 mm</quantity>
; antenna length
<quantity id="4CD29B4342165514FC38FB8FE615D314" box="[1012,1089,1032,1053]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="199" unit="mm" value="3.8">3.8 mm</quantity>
.
</paragraph>
<paragraph id="8B9536A642165514FC9EFBA4E12FD3FA" blockId="1.[812,1399,744,2067]" pageId="1" pageNumber="199">Head: antenna black with dark blue shine, scapus black exterior-dorsally and pale yellow interior-ventrally; frons pale yellow with few gray-brown scales medially; labial palpus with long hair-like scales ventrally, pale yellow and narrowly black exterior-ventrally; vertex black with dark blue shine; occipital fringe pale yellow to white; neck plate pale yellow with few gray-brown scales with blue shine distally.</paragraph>
<paragraph id="8B9536A642165514FC9EFB7FE1ABD13A" blockId="1.[812,1399,744,2067]" pageId="1" pageNumber="199">Thorax: patagia black with blue-violet shine, admixture of pale yellow scales anteriorly and small pale yellow spot laterally; tegula black with blue-violet shine, small pale yellow spot at base of forewing and few pale yellow scales both posteriorly and at inner margin anteriorly; both meso- and metathorax black with dark greenish-blue shine; besides this, tegula, meso- and metathorax densely covered with white, long, hair-like scales; thorax laterally dark gray-brown with bright blue shine and large yellow spot with golden shine anteriorly; both metepimeron and metameron posteriorly black with dark violet shine densely covered with white, long, hair-like scales.</paragraph>
<paragraph id="8B9536A642165517FC9EF9BFE387D6DE" blockId="1.[812,1399,744,2067]" lastBlockId="2.[189,775,315,1111]" lastPageId="2" lastPageNumber="200" pageId="1" pageNumber="199">Legs: fore coxa dark brown to black with blue-bronze shine, densely covered with white, long, hair-like scales and narrowly pale yellow at margins; fore femur dark brown to black with blue-bronze shine and dense admixture of pale yellow scales and white, long, hair-like scales at posterior margin; fore tibia pale yellow to yellow with few black scales dorsally; fore tarsus pale yellow to yellow with golden shine; mid coxa dark brown to black with blue-bronze shine; mid femur dark brown to black with blue-bronze shine and dense admixture of pale yellow scales and white, long, hair-like scales at posterior margin; mid tibia dark brown to black with bronze shine and large yellow with golden shine spot exterior-medially and admixture of pale yellow scales internally; spurs pale yellow with golden shine; mid tarsus completely pale yellow to yellow with golden shine; hind coxa dark brown to black with blue-bronze shine and admixture of pale yellow scales; hind femur dark brown to black with blue-bronze shine and dense admixture of pale yellow scales and white, long, hair-like scales at posterior margin; hind tibia pale yellow to yellow with golden shine and dark brown to black scales with bronze shine basally, ventrally from base to base of mid spurs and at base of apical spurs; spurs pale yellow with golden shine; hind tarsus completely pale yellow to yellow with golden shine.</paragraph>
<paragraph id="8B9536A642155517FF2EFE5CE0C6D5F5" blockId="2.[189,775,315,1111]" pageId="2" pageNumber="200">
Forewing dorsally with basal part black with bright greenish shine; costal margin brown with bronze shine and thin pale yellow line between veins Sc and R-stem; Cu-stem and anal margin yellow with golden shine and admixture of brown scales with bronze shine; discal spot trapeziform, black with dark violet shine; veins distally of discal spot yellow with admixture of brown scales distally; ventrally forewing yellow, discal spot and surface between veins R
<subScript id="17AE34E342155517FD33FD22E152D5B8" attach="left" box="[767,774,677,689]" fontSize="5" pageId="2" pageNumber="200">1</subScript>
and R
<subScript id="17AE34E342155517FF30FD47E357D5C5" attach="left" box="[252,259,704,716]" fontSize="5" pageId="2" pageNumber="200">3</subScript>
dark brown with bronze shine; transparent areas undeveloped, completely covered with yellow scales with golden hue; cilia dark brown with bronze shine.
</paragraph>
<paragraph id="8B9536A642155517FF2EFC86E04DD30E" blockId="2.[189,775,315,1111]" pageId="2" pageNumber="200">
Hindwing transparent but densely covered with translucent and yellowish scales with light golden hue dorsally and bluish hue ventrally; dorsally veins, discal spot and outer margin brown with bronze shine and admixture of individual yellow scales; ventrally veins, discal spot and outer margin yellow with admixture of brown scales with bronze shine, more dense on Cu-stem and veins M
<subScript id="17AE34E342155517FDE8FC37E07FD4B5" attach="left" box="[548,555,944,956]" fontSize="5" pageId="2" pageNumber="200">3</subScript>
and CuA
<subScript id="17AE34E342155517FD46FC37E0C5D4B5" attach="right" box="[650,657,944,956]" fontSize="5" pageId="2" pageNumber="200">1</subScript>
; discal spot cuneiform, reaching base of common stem M 3 –CuA 1; outer margin relatively broad, about 0.5 times as broad as cilia; cilia dark brown with bronze shine.
</paragraph>
<paragraph id="8B9536A642155517FF2EFB8CE715D505" blockId="2.[189,775,315,1111]" lastBlockId="2.[812,1399,315,1111]" pageId="2" pageNumber="200">Abdomen dark brown to black with bright blue shine; dorsally tergite 2 with narrow yellow stripe with golden shine distally; tergites 4 and 5 each with broad yellow stripe with golden shine distally; tergites 6 and 7 each nearly completely yellow with golden shine; ventrally sternite 1+2 with narrow yellow stripe with golden shine distally; sternite 3 with few yellow scales medially; sternite 4 completely yellow with golden shine; sternites 5–7 each with broad yellow stripe with golden shine distally; anal tuft well-developed, dorsally lateral parts black with bronze-violet shine, medial part yellow with golden shine; ventrally yellow with golden shine.</paragraph>
<paragraph id="8B9536A642155517FC9EFD96E63ED489" blockId="2.[812,1399,315,1111]" pageId="2" pageNumber="200">
<emphasis id="B95EEAB442155517FC9EFD96E1BCD52F" bold="true" box="[850,1000,528,550]" pageId="2" pageNumber="200">Male genitalia</emphasis>
(
<typeStatus id="5491880442155517FC34FD96E604D52F" box="[1016,1104,529,550]" pageId="2" pageNumber="200" type="paratype">paratype</typeStatus>
; genital preparation No OG– 008-2023) (
<figureCitation id="13112A2342155517FC6FFDACE655D548" box="[931,1025,555,577]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Figs 5–9</figureCitation>
). Tegumen-uncus complex relatively narrow; scopula androconialis well-developed, about 0.6 times as long as tegumen-uncus complex (
<figureCitation id="13112A2342155517FB24FDE6E77DD57F" box="[1256,1321,608,631]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 5</figureCitation>
); crista gnathi medialis semicircular, long, broad and poorly sclerotized in basal half; crista gnathi lateralis subcordiform, about half as long as and slightly narrower than crista gnathi medialis (
<figureCitation id="13112A2342155517FC5CFD4CE185D5E8" box="[912,977,715,737]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 5</figureCitation>
); valva (
<figureCitation id="13112A2342155517FBE3FD4CE63BD5E8" box="[1071,1135,715,737]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 6</figureCitation>
) trapeziform-ovoid, crista sacculi oblique, covered with apically pointed setae anteriorly and flat-topped ones posteriorly; caudal part of row of setae bent towards ventral margin; saccus (
<figureCitation id="13112A2342155517FB1EFC9CE759D439" box="[1234,1293,795,817]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 7</figureCitation>
) relatively narrow, long, about twice as long as vinculum; phallus (
<figureCitation id="13112A2342155517FA83FCB1E16CD46F" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 8</figureCitation>
) rather narrow, straight, about as long as valva; vesica with numerous small cornuti (
<figureCitation id="13112A2342155517FBEDFCECE60AD489" box="[1057,1118,875,897]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="2" pageNumber="200">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B9536A642155517FC9EFC01E645D495" blockId="2.[812,1399,315,1111]" box="[850,1041,902,924]" pageId="2" pageNumber="200">
<emphasis id="B95EEAB442155517FC9EFC01E1C9D492" bold="true" box="[850,925,902,923]" pageId="2" pageNumber="200">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B9536A642155517FC9EFC26E6ACD4BE" blockId="2.[812,1399,315,1111]" box="[850,1272,929,951]" pageId="2" pageNumber="200">INDIVIDUAL VARIABILITY. Unknown.</paragraph>
<paragraph id="8B9536A642155516FC9EFC3CE189D5F2" blockId="2.[812,1399,315,1111]" lastBlockId="3.[812,1399,315,764]" lastPageId="3" lastPageNumber="201" pageId="2" pageNumber="200">
DIFFERENTIAL DIAGNOSIS. Both superficially and by the structure of the male genitalia, this new species seems to be closest to
<taxonomicName id="4C2A4D2542155517FC5BFC76E6B4D30F" authority="O. Gorbunov, 1994" authorityName="O. Gorbunov" authorityYear="1994" box="[919,1248,1009,1031]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="volgensis">
<emphasis id="B95EEAB442155517FC5BFC76E65BD30F" box="[919,1039,1009,1030]" italics="true" pageId="2" pageNumber="200">B. volgensis</emphasis>
O.
<bibRefCitation id="EFBB4B5742155517FBF2FC76E6B4D30F" author="Gorbunov O. G." box="[1086,1248,1009,1031]" pageId="2" pageNumber="200" pagination="563 - 566" refId="ref7425" refString="Gorbunov O. G. 1994. A new species of the genus Bembecia Hubner, [1819] from the European part of Russia // Atalanta (Marktleuthen). Bd. 25. No. 3 / 4. S. 563 - 566, 623." type="journal article" year="1994">Gorbunov, 1994</bibRefCitation>
</taxonomicName>
(
<typeStatus id="5491880442155517FB3EFC76E748D30F" box="[1266,1308,1009,1030]" pageId="2" pageNumber="200">type</typeStatus>
locality:
<collectingCountry id="F33D763642155517FCE0FB8CE13AD328" box="[812,878,1035,1057]" name="Russia" pageId="2" pageNumber="200">Russia</collectingCountry>
,
<collectingRegion id="49EEF84442155517FCB4FB8CE672D328" box="[888,1062,1035,1057]" country="Russia" name="Ul'yanovsk" pageId="2" pageNumber="200">Ulianovsk Region</collectingRegion>
,
<quantity id="4CD29B4342155517FBFCFB8CE62DD328" box="[1072,1145,1035,1057]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="200" unit="km" value="160.0">160 km</quantity>
S of
<collectingRegion id="49EEF84442155517FB60FB8CE744D328" box="[1196,1296,1035,1057]" country="Russia" name="Ul'yanovsk" pageId="2" pageNumber="200">Ulianovsk</collectingRegion>
, Ryabina), from which it can be distinguished by the colouration of the vertex (black mixed with yellow scales in
<taxonomicName id="4C2A4D2542155517FB1BFBC6E719D35F" authorityName="O. Gorbunov" authorityYear="1994" box="[1239,1357,1089,1110]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="volgensis">
<emphasis id="B95EEAB442155517FB1BFBC6E719D35F" box="[1239,1357,1089,1110]" italics="true" pageId="2" pageNumber="200">B. volgensis</emphasis>
</taxonomicName>
,
<emphasis id="B95EEAB442155517FA91FBC6E725D35F" box="[1373,1393,1089,1110]" italics="true" pageId="2" pageNumber="200">vs</emphasis>
. completely black in the new species), patagia (black with violet shine dorsally and pale yellow laterally in
<taxonomicName id="4C2A4D2542145516FDABFED0E089D662" authorityName="O. Gorbunov" authorityYear="1994" box="[615,733,342,364]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="volgensis">
<emphasis id="B95EEAB442145516FDABFED0E089D662" box="[615,733,342,364]" italics="true" pageId="3" pageNumber="201">B. volgensis</emphasis>
</taxonomicName>
,
<emphasis id="B95EEAB442145516FD21FED0E155D665" box="[749,769,343,364]" italics="true" pageId="3" pageNumber="201">vs</emphasis>
. black with blue-violet shine, admixture of pale yellow scales anteriorly and small pale yellow spot laterally in
<taxonomicName id="4C2A4D2542145516FD49FE0BE153D6A9" authorityName="O. Gorbunov" authorityYear="2023" box="[645,775,395,417]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442145516FD49FE0BE153D6A9" box="[645,775,395,417]" italics="true" pageId="3" pageNumber="201">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442145516FF71FE21E2B8D6B2" bold="true" box="[189,236,422,443]" pageId="3" pageNumber="201">
<taxonomicNameLabel id="A26D57CF42145516FF71FE21E2B8D6B2" box="[189,236,422,443]" pageId="3" pageNumber="201" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
), fore coxa (black with violet shine, densely covered with yellow, long, hairy-like scales and broadly white at exterior margin in the species compared,
<emphasis id="B95EEAB442145516FE3BFE5BE05FD6F8" box="[503,523,476,497]" italics="true" pageId="3" pageNumber="201">vs</emphasis>
. dark brown to black with blue-bronze shine, densely covered with white, long, hair-like scales and narrowly pale yellow at margins in
<taxonomicName id="4C2A4D2542145516FD59FD96E282D548" authorityName="O. Gorbunov" authorityYear="2023" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442145516FD59FD96E282D548" italics="true" pageId="3" pageNumber="201">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442145516FF17FDACE358D549" bold="true" box="[219,268,555,576]" pageId="3" pageNumber="201">
<taxonomicNameLabel id="A26D57CF42145516FF17FDACE358D549" box="[219,268,555,576]" pageId="3" pageNumber="201" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
), abdomen (black with greenish-violet shine; dorsally tergite 2 with narrow pale yellow stripe distally; tergites 4, 6 and 7 each with broad pale yellow to yellow stripe distally; tergite 5 with admixture of pale yellow scales medially; ventrally sternites 1+2 and 3 each with few pale yellow scales; sternites 5–7 each with more numerous pale yellow scales; sternite 4 entirely pale yellow in
<taxonomicName id="4C2A4D2542145516FDF1FD4BE0E4D5E9" authorityName="O. Gorbunov" authorityYear="1994" box="[573,688,715,737]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="volgensis">
<emphasis id="B95EEAB442145516FDF1FD4BE0E4D5E9" box="[573,688,715,737]" italics="true" pageId="3" pageNumber="201">B. volgensis</emphasis>
</taxonomicName>
,
<emphasis id="B95EEAB442145516FD70FD4BE084D5E8" box="[700,720,716,737]" italics="true" pageId="3" pageNumber="201">vs</emphasis>
. dark brown to black with bright blue shine; dorsally tergite 2 with narrow yellow stripe with golden shine distally; tergites 4 and 5 each with broad yellow stripe with golden shine distally; tergites 6 and 7 each nearly completely yellow with golden shine; ventrally sternite 1+2 with narrow yellow stripe with golden shine distally; sternite 3 with few yellow scales medially; sternite 4 completely yellow with golden shine; sternites 5–7 each with broad yellow stripe with golden shine distally in
<taxonomicName id="4C2A4D2542145516FC84FE70E19BD502" authorityName="O. Gorbunov" authorityYear="2023" box="[840,975,502,524]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442145516FC84FE70E19BD502" box="[840,975,502,524]" italics="true" pageId="3" pageNumber="201">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442145516FC15FE71E65DD502" bold="true" box="[985,1033,502,523]" pageId="3" pageNumber="201">
<taxonomicNameLabel id="A26D57CF42145516FC15FE71E65DD502" box="[985,1033,502,523]" pageId="3" pageNumber="201" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
) and in the shape of the transparent areas of the forewing (well-developed but densely covered with colourless and yellowish scales in the species compared,
<emphasis id="B95EEAB442145516FCE0FDC0E114D555" box="[812,832,583,604]" italics="true" pageId="3" pageNumber="201">vs</emphasis>
. transparent areas undeveloped, completely covered with yellow scales with golden hue in the new species; cp.
<figureCitation id="13112A2342145516FAE3FDE6E16CD599" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="3" pageNumber="201">
Figs
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</figureCitation>
in this publication with pl. XVIIIb, figs
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<bibRefCitation id="EFBB4B5742145516FAC2FDFCE108D5A2" author="Gorbunov O. G." pageId="3" pageNumber="201" pagination="563 - 566" refId="ref7425" refString="Gorbunov O. G. 1994. A new species of the genus Bembecia Hubner, [1819] from the European part of Russia // Atalanta (Marktleuthen). Bd. 25. No. 3 / 4. S. 563 - 566, 623." type="journal article" year="1994">Gorbunov, 1994</bibRefCitation>
or with figs 5i–k in
<bibRefCitation id="EFBB4B5742145516FBD5FD11E751D5A2" author="Gorbunov O. G. &amp; Efetov K. A." box="[1049,1285,662,684]" pageId="3" pageNumber="201" pagination="812 - 839" refId="ref7697" refString="Gorbunov O. G., Efetov K. A. 2018. The clearwing moth genus Bembecia Hubner 1819 [&quot; 1816 &quot;] (Lepidoptera, Sesiidae) in Crimea, with the description of a new species // Zool. Zhurn. Vol. 97. No. 7. P. 812 - 839." type="journal article" year="2018">Gorbunov, Efetov, 2018</bibRefCitation>
). These two species have nearly no differences in the structure of the male genitalia (cp.
<figureCitation id="13112A2342145516FC7BFD4CE65AD5E8" box="[951,1038,715,737]" captionStart="Figs 5–9" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[189,1438,799,1955]" captionTargetId="figure-442@3.[189,1466,794,1956]" captionTargetPageId="3" captionText="Figs 5–9. Genitalia of Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9. Рис. 5–9. Гениталии Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9." pageId="3" pageNumber="201">
Figs
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in this publication with figs
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<bibRefCitation id="EFBB4B5742145516FCE0FD61E185D5F2" author="Gorbunov O. G." box="[812,977,742,764]" pageId="3" pageNumber="201" pagination="563 - 566" refId="ref7425" refString="Gorbunov O. G. 1994. A new species of the genus Bembecia Hubner, [1819] from the European part of Russia // Atalanta (Marktleuthen). Bd. 25. No. 3 / 4. S. 563 - 566, 623." type="journal article" year="1994">Gorbunov, 1994</bibRefCitation>
).
</paragraph>
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<paragraph id="8B9536A642155517FF2EF806E044D0CD" blockId="2.[189,1399,1921,2065]" pageId="2" pageNumber="200">
Figs 1–4.
<taxonomicName id="4C2A4D2542155517FEFAF806E3EAD09D" box="[310,446,1921,1941]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B95EEAB442155517FEFAF806E3DFD09D" box="[310,395,1921,1940]" italics="true" pageId="2" pageNumber="200">Bembecia</emphasis>
spp. 1
</taxonomicName>
–2 —
<taxonomicName id="4C2A4D2542155517FE39F806E170D09D" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[501,804,1921,1940]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="khovdensis" status="sp. nov.">
<emphasis id="B95EEAB442155517FE39F806E0F8D09D" box="[501,684,1921,1940]" italics="true" pageId="2" pageNumber="200">Bembecia khovdensis</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442155517FCE7F806E102D09D" bold="true" box="[811,854,1921,1940]" pageId="2" pageNumber="200">
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</emphasis>
, holotype ♂, alar expanse 14.9 mm.
<taxonomicName id="4C2A4D2542155517FB5DF806E68DD09D" authorityName="Boisduval" authorityYear="1828" box="[1169,1241,1921,1940]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
picture Nos 0033- 0034–2023; 3–4 —
<taxonomicName id="4C2A4D2542155517FEA7F81EE0C7D0A5" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[363,659,1945,1964]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov.">
<emphasis id="B95EEAB442155517FEA7F81EE043D0A5" box="[363,535,1945,1964]" italics="true" pageId="2" pageNumber="200">Bembecia yakovlevi</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442155517FD50F81EE093D0A5" bold="true" box="[668,711,1945,1964]" pageId="2" pageNumber="200">
<taxonomicNameLabel id="A26D57CF42155517FD50F81EE093D0A5" box="[668,711,1945,1964]" pageId="2" pageNumber="200" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype ♂, alar expanse 21.4 mm.
<taxonomicName id="4C2A4D2542155517FBC3F81EE603D0A5" authorityName="Boisduval" authorityYear="1828" box="[1039,1111,1945,1964]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view.
</paragraph>
<paragraph id="8B9536A642155517FF2EF849E018DF18" blockId="2.[189,1399,1921,2065]" pageId="2" pageNumber="200">
Рис. 1–4.
<taxonomicName id="4C2A4D2542155517FEF7F84AE393D0E9" box="[315,455,1997,2017]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B95EEAB442155517FEF7F84AE3C4D0E9" box="[315,400,1997,2016]" italics="true" pageId="2" pageNumber="200">Bembecia</emphasis>
spp. 1
</taxonomicName>
–2:
<taxonomicName id="4C2A4D2542155517FE27F84AE14BD0E9" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[491,799,1997,2016]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="khovdensis" status="sp. nov.">
<emphasis id="B95EEAB442155517FE27F84AE0F7D0E9" box="[491,675,1997,2016]" italics="true" pageId="2" pageNumber="200">Bembecia khovdensis</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442155517FCEBF84AE106D0E9" bold="true" box="[807,850,1997,2016]" pageId="2" pageNumber="200">
<taxonomicNameLabel id="A26D57CF42155517FCEBF84AE106D0E9" box="[807,850,1997,2016]" pageId="2" pageNumber="200" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, голотип ♂, раЗмах крыльев 14.9 мм.
<taxonomicName id="4C2A4D2542155517FB7AF84AE6AAD0E9" authorityName="Boisduval" authorityYear="1828" box="[1206,1278,1997,2016]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
снимки №№ 0033-0034–2023; 3–4 —
<taxonomicName id="4C2A4D2542155517FE55F862E0E8D0F1" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[409,700,2021,2040]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov.">
<emphasis id="B95EEAB442155517FE55F862E010D0F1" box="[409,580,2021,2040]" italics="true" pageId="2" pageNumber="200">Bembecia yakovlevi</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442155517FD08F862E0BBD0F1" bold="true" box="[708,751,2021,2040]" pageId="2" pageNumber="200">
<taxonomicNameLabel id="A26D57CF42155517FD08F862E0BBD0F1" box="[708,751,2021,2040]" pageId="2" pageNumber="200" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype ♂, раЗмах крыльев 21.4 мм.
<taxonomicName id="4C2A4D2542155517FB87F862E6C7D0F1" authorityName="Boisduval" authorityYear="1828" box="[1099,1171,2021,2040]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="200" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу.
</paragraph>
</caption>
<caption id="DF55662E42145516FF2EF836E6E2DF19" pageId="3" pageNumber="201" startId="3.[226,263,1969,1988]" targetBox="[189,1438,799,1955]" targetPageId="3" targetType="figure">
<paragraph id="8B9536A642145516FF2EF836E664D0D5" blockId="3.[189,1398,1969,2065]" pageId="3" pageNumber="201">
Figs 5–9. Genitalia of
<taxonomicName id="4C2A4D2542145516FE6CF836E084D0CD" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[416,720,1969,1988]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="khovdensis" status="sp. nov.">
<emphasis id="B95EEAB442145516FE6CF836E003D0CD" box="[416,599,1969,1988]" italics="true" pageId="3" pageNumber="201">Bembecia khovdensis</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442145516FD1BF836E156D0CD" bold="true" box="[727,770,1969,1988]" pageId="3" pageNumber="201">
<taxonomicNameLabel id="A26D57CF42145516FD1BF836E156D0CD" box="[727,770,1969,1988]" pageId="3" pageNumber="201" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype ♂. Genital preparation No OG–008-2023: 5 — tegumen-uncus complex; 6 — valva; 7 — saccus; 8 — phallus; 9 — vesica. Scale bar 0.5 mm for 5–8 and 0.2 for 9.
</paragraph>
<paragraph id="8B9536A642145516FF2EF861E6E2DF19" blockId="3.[189,1398,1969,2065]" pageId="3" pageNumber="201">
Рис. 5–9. Гениталии
<taxonomicName id="4C2A4D2542145516FE6AF862E089D0F1" authority="O. Gorbunov" authorityName="O. Gorbunov" authorityYear="2023" box="[422,733,2021,2040]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="species" species="khovdensis" status="sp. nov.">
<emphasis id="B95EEAB442145516FE6AF862E00BD0F1" box="[422,607,2021,2040]" italics="true" pageId="3" pageNumber="201">Bembecia khovdensis</emphasis>
O. Gorbunov
</taxonomicName>
,
<emphasis id="B95EEAB442145516FD2BF862E146D0F1" bold="true" box="[743,786,2021,2040]" pageId="3" pageNumber="201">
<taxonomicNameLabel id="A26D57CF42145516FD2BF862E146D0F1" box="[743,786,2021,2040]" pageId="3" pageNumber="201" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, голотип ♂. Препарат гениталий № OG–008-2023: 5 — тегумен- ункусный комплекс; 6 — вальва; 7 — саккус; 8 — фаллюс; 9 — веЗика. МасШтаб 0,5 мм длЯ 5–8 и 0,2 длЯ 9.
</paragraph>
</caption>
<paragraph id="8B9536A642135511FF2EFEBBE0BFD599" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
From
<taxonomicName id="4C2A4D2542135511FED1FEBBE073D658" authority="(Pungeler, 1905)" baseAuthorityName="Pungeler" baseAuthorityYear="1905" box="[285,551,315,337]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="tancrei">
<emphasis id="B95EEAB442135511FED1FEBBE328D658" box="[285,380,316,337]" italics="true" pageId="4" pageNumber="202">B. tancrei</emphasis>
(Püngeler, 1905)
</taxonomicName>
(
<typeStatus id="5491880442135511FDF9FEBCE034D659" box="[565,608,315,336]" pageId="4" pageNumber="202">type</typeStatus>
locality: “… aus dem Altyn-Tagh.” [Püngeler, 1905: 270] [=
<collectingCountry id="F33D763642135511FDA7FED1E0FCD665" box="[619,680,342,364]" name="China" pageId="4" pageNumber="202">China</collectingCountry>
:
<collectingRegion id="49EEF84442135511FD7EFED1E2AFD68F" country="China" name="Xinjiang Uygur" pageId="4" pageNumber="202">Xinjiang Uygur</collectingRegion>
Autonomous Region, Altyn Tagh Mts.]),
<taxonomicName id="4C2A4D2542135511FD5FFEF6E282D6A8" authorityName="O. Gorbunov" authorityYear="2023" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442135511FD5FFEF6E282D6A8" italics="true" pageId="4" pageNumber="202">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442135511FF17FE0CE358D6A9" bold="true" box="[219,268,395,416]" pageId="4" pageNumber="202">
<taxonomicNameLabel id="A26D57CF42135511FF17FE0CE358D6A9" box="[219,268,395,416]" pageId="4" pageNumber="202" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs in markedly smaller size (wing span about
<quantity id="4CD29B4342135511FD22FE0CE2B6D6B2" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="202" unit="mm" value="25.0">25 mm</quantity>
in
<taxonomicName id="4C2A4D2542135511FEC7FE20E324D6B5" baseAuthorityName="Pungeler" baseAuthorityYear="1905" box="[267,368,423,444]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="tancrei">
<emphasis id="B95EEAB442135511FEC7FE20E324D6B5" box="[267,368,423,444]" italics="true" pageId="4" pageNumber="202">B. tancrei</emphasis>
</taxonomicName>
,
<emphasis id="B95EEAB442135511FE4EFE20E3C2D6B5" box="[386,406,423,444]" italics="true" pageId="4" pageNumber="202">vs</emphasis>
.
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in the new species) and colouration of the abdomen dorsally (black with bluish shine, tergites 2, 4 and 6 each with yellow, narrow, posterior margin in
<taxonomicName id="4C2A4D2542135511FF1AFE70E361D505" baseAuthorityName="Pungeler" baseAuthorityYear="1905" box="[214,309,503,524]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="tancrei">
<emphasis id="B95EEAB442135511FF1AFE70E361D505" box="[214,309,503,524]" italics="true" pageId="4" pageNumber="202">B. tancrei</emphasis>
</taxonomicName>
,
<emphasis id="B95EEAB442135511FE8EFE70E302D505" box="[322,342,503,524]" italics="true" pageId="4" pageNumber="202">vs</emphasis>
. dark brown to black with bright blue shine; dorsally tergite 2 with narrow yellow stripe with golden shine distally; tergites 4 and 5 each with broad yellow stripe with golden shine distally; tergites 6 and 7 each nearly completely yellow with golden shine in
<taxonomicName id="4C2A4D2542135511FE01FDE6E005D57F" authorityName="O. Gorbunov" authorityYear="2023" box="[461,593,609,630]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442135511FE01FDE6E005D57F" box="[461,593,609,630]" italics="true" pageId="4" pageNumber="202">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442135511FD9BFDE6E0D3D57F" bold="true" box="[599,647,609,630]" pageId="4" pageNumber="202">
<taxonomicNameLabel id="A26D57CF42135511FD9BFDE6E0D3D57F" box="[599,647,609,630]" pageId="4" pageNumber="202" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; cp.
<figureCitation id="13112A2342135511FD7AFDE6E0BAD57F" box="[694,750,609,631]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="4" pageNumber="202">
Fig.
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</figureCitation>
in this article with pl. 25, fig.
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<bibRefCitation id="EFBB4B5742135511FDC2FDF1E0B4D599" author="Spatenka K. &amp; Gorbunov O. &amp; Lastuvka Z. &amp; Tosevski I. &amp; Arita Y." box="[526,736,630,657]" pageId="4" pageNumber="202" refId="ref8253" refString="Spatenka K., Gorbunov O., Lastuvka Z., Tosevski I., Arita Y. 1999. Sesiidae, Clearwing Moths // Naumann C. M. (ed.). Handbook of Palaearctic Macrolepidoptera. Vol. 1. Wallingford: Gem Publishing Company. 569 p., 57 pls." type="book" year="1999">
Špatenka
<emphasis id="B95EEAB442135511FDA1FDFBE0CCD599" box="[621,664,635,657]" italics="true" pageId="4" pageNumber="202">et al</emphasis>
., 1999
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9536A642135511FF2EFD11E3CFD439" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
From all other Mongolian congeners,
<taxonomicName id="4C2A4D2542135511FD81FD10E085D5A2" authorityName="O. Gorbunov" authorityYear="2023" box="[589,721,662,684]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442135511FD81FD10E085D5A2" box="[589,721,662,684]" italics="true" pageId="4" pageNumber="202">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442135511FD1AFD11E153D5A2" bold="true" box="[726,775,662,683]" pageId="4" pageNumber="202">
<taxonomicNameLabel id="A26D57CF42135511FD1AFD11E153D5A2" box="[726,775,662,683]" pageId="4" pageNumber="202" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is easily separable by the smaller size (not less than
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in all species compared,
<emphasis id="B95EEAB442135511FE57FD4BE3FBD5E8" box="[411,431,716,737]" italics="true" pageId="4" pageNumber="202">vs</emphasis>
.
<quantity id="4CD29B4342135511FE73FD4CE043D5E8" box="[447,535,715,737]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="4" pageNumber="202" unit="mm" value="14.9">14.9 mm</quantity>
in
<taxonomicName id="4C2A4D2542135511FDF1FD4BE091D5E9" authorityName="O. Gorbunov" authorityYear="2023" box="[573,709,715,737]" class="Insecta" family="Sesiidae" genus="Bembecia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="202" phylum="Arthropoda" rank="species" species="khovdensis" status="sp.n.">
<emphasis id="B95EEAB442135511FDF1FD4BE091D5E9" box="[573,709,715,737]" italics="true" pageId="4" pageNumber="202">B. khovdensis</emphasis>
</taxonomicName>
<emphasis id="B95EEAB442135511FD02FD4CE0ABD5E9" bold="true" box="[718,767,715,736]" pageId="4" pageNumber="202">
<taxonomicNameLabel id="A26D57CF42135511FD02FD4CE0ABD5E9" box="[718,767,715,736]" pageId="4" pageNumber="202" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
) colouration of various parts of the body and wings (compare
<figureCitation id="13112A2342135511FF71FC86E2AED41F" box="[189,250,769,791]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[189,1399,1154,1900]" captionTargetId="figure-647@2.[189,1399,1153,1900]" captionTargetPageId="2" captionText="Figs 1–4. Bembecia spp. 1–2 — Bembecia khovdensis O. Gorbunov, sp.n., holotype ♂, alar expanse 14.9 mm. Sesiidae picture Nos 0033- 0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, alar expanse 21.4 mm. Sesiidae picture Nos 0027-0028–2023. 1, 3 — dorsal view; 2, 4 — ventral view. Рис. 1–4. Bembecia spp. 1–2: Bembecia khovdensis O. Gorbunov, sp.n., голотип ♂, раЗмах крыльев 14.9 мм. Sesiidae снимки №№ 0033-0034–2023; 3–4 — Bembecia yakovlevi O. Gorbunov, sp.n., holotype ♂, раЗмах крыльев 21.4 мм. Sesiidae снимки №№ 0027-0028– 2023. 1, 3 — вид сверху; 2, 4 — вид сниЗу." pageId="4" pageNumber="202">
Fig.
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</figureCitation>
in this article with figs 196, 198, 218 and
<quantity id="4CD29B4342135511FD09FC86E153D41E" box="[709,775,769,791]" metricMagnitude="0" metricUnit="m" metricValue="7.2898" pageId="4" pageNumber="202" unit="in" value="287.0">287 in</quantity>
<bibRefCitation id="EFBB4B5742135511FF71FC91E3DBD439" author="Spatenka K. &amp; Gorbunov O. &amp; Lastuvka Z. &amp; Tosevski I. &amp; Arita Y." box="[189,399,790,817]" pageId="4" pageNumber="202" refId="ref8253" refString="Spatenka K., Gorbunov O., Lastuvka Z., Tosevski I., Arita Y. 1999. Sesiidae, Clearwing Moths // Naumann C. M. (ed.). Handbook of Palaearctic Macrolepidoptera. Vol. 1. Wallingford: Gem Publishing Company. 569 p., 57 pls." type="book" year="1999">
Špatenka
<emphasis id="B95EEAB442135511FED0FC9BE31CD439" box="[284,328,795,817]" italics="true" pageId="4" pageNumber="202">et al</emphasis>
., 1999
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9536A642135511FF2EFCB1E011D495" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
BIONOMICS. The host plant and larval bionomics are unknown. The
<typeStatus id="5491880442135511FE96FCD6E3FBD46E" box="[346,431,849,871]" pageId="4" pageNumber="202" type="holotype">holotype</typeStatus>
was collected using non-specific artificial sex attractants in late June. It was active slightly before noon at about 11 a.m. local time.
</paragraph>
<paragraph id="8B9536A642135511FF2EFC26E395D4D8" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
HABITAT. Rocky semi-desert or even desert with very little grass cover (
<figureCitation id="13112A2342135511FEA1FC3CE3E1D4D8" box="[365,437,955,977]" captionStart="Fig" captionStartId="6.[226,257,1969,1988]" captionTargetBox="[189,1399,1147,1951]" captionTargetId="figure-693@6.[189,1399,1147,1951]" captionTargetPageId="6" captionText="Fig. 15. The type locality of Bembecia khovdensis O. Gorbunov, sp.n.: Mongolia, Khovd Aimag, 8 km N Uyench, 1800 m, 46°07´N, 092°03´E, 22.VI.2007. Photo by O. Gorbunov. Рис. 15. Типовое местонахождение Bembecia khovdensis O. Gorbunov, sp.n.: МонголиЯ, Кобдоский аймак, 8 км С Уенча, 1800 м, 46°07´с.Ш., 092°03´в.д., 22.VI.2007. Фото О. Горбунов." pageId="4" pageNumber="202">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330652D42135511FF2EFC51E325D335" pageId="4" pageNumber="202" type="distribution">
<paragraph id="8B9536A642135511FF2EFC51E325D335" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
<materialsCitation id="3B423CFB42135511FF2EFC51E339D335" country="Mongolia" elevation="1800" location="Uyench Somon" municipality="Uyench" pageId="4" pageNumber="202" specimenCount="1" stateProvince="Hovd">
DISTRIBUTION. This species is known only from the
<typeStatus id="5491880442135511FF71FC76E2B3D30F" box="[189,231,1009,1030]" pageId="4" pageNumber="202">type</typeStatus>
locality in the vicinity of
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<collectingMunicipality id="6BF1ACDC42135511FE12FC76E07DD30E" box="[478,553,1009,1031]" pageId="4" pageNumber="202">Uyench</collectingMunicipality>
Somon
</location>
in the southern part of
<collectingRegion id="49EEF84442135511FECDFB8CE3D9D328" box="[257,397,1035,1057]" country="Mongolia" name="Hovd" pageId="4" pageNumber="202">Khovd Aimag</collectingRegion>
,
<collectingCountry id="F33D763642135511FE5AFB8CE3A2D328" box="[406,502,1035,1057]" name="Mongolia" pageId="4" pageNumber="202">Mongolia</collectingCountry>
, at an altitude of about
<elevation id="0007D19542135511FD1AFB8CE339D335" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="202" unit="m" value="1800.0">
<quantity id="4CD29B4342135511FD1AFB8CE284D332" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="202" unit="m" value="1800.0">1800 m</quantity>
above sea level
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330652D42135511FF2EFBC6E02AD378" pageId="4" pageNumber="202" type="etymology">
<paragraph id="8B9536A642135511FF2EFBC6E02AD378" blockId="4.[189,775,315,1137]" pageId="4" pageNumber="202">
ETYMOLOGY. This new species is named after the
<collectingRegion id="49EEF84442135511FF71FBDCE313D378" box="[189,327,1115,1137]" country="Mongolia" name="Hovd" pageId="4" pageNumber="202">Khovd Aimag</collectingRegion>
of Mongolia, where it inhabits.
</paragraph>
</subSubSection>
</treatment>
</document>

272
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<mods:title id="A939AEC5BD531109EDBCE994317D5246">Review of the Aphrophora salicina (Goeze, 1778) (Homoptera: Cercopoidea: Aphrophoridae) species group of Russia and adjacent territories with description of a new species</mods:title>
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<taxonomicName id="4C382B90FB62FFE2FE86FE070BB7B2F7" authority="Tishechkin" authorityName="Tishechkin" authorityYear="2023" box="[297,650,460,483]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="itiliensis" status="sp. nov.">
<emphasis id="B94C8C01FB62FFE2FE86FE070B36B2F7" box="[297,523,460,483]" italics="true" pageId="7" pageNumber="260">Aphrophora itiliensis</emphasis>
Tishechkin
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,
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<subSubSection id="C3220398FB62FFE2FE80FE220BF8B114" box="[303,709,489,512]" pageId="7" pageNumber="260" type="description">
<paragraph id="8B875013FB62FFE2FE80FE220BF8B114" blockId="7.[297,714,460,512]" box="[303,709,489,512]" pageId="7" pageNumber="260">
<figureCitation id="13034C96FB62FFE2FE80FE220892B114" box="[303,431,489,512]" captionStart="Figs 1–16" captionStartId="2.[283,320,1932,1951]" captionTargetBox="[283,1443,322,1857]" captionTargetId="figure-124@2.[269,1454,320,1882]" captionTargetPageId="2" captionText="Figs 1–16. Aphrophora spp., male calling signal oscillograms. 1–11 ― A. salicina; 12–13 ― A. willemsi; 14–16 ― A. itiliensis sp.n. Faster oscillograms of the parts of signals indicated as “17–31” are given under the same numbers, signal recording localities are given under the oscillograms. Scale mark at the bottom is the same for all oscillograms. Рис. 1–16. Aphrophora spp., осциллограммы приЗывных сигналов самцов. 1–11 ― A. salicina; 12–13 ― A. willemsi; 14–16 ― A.itiliensis sp.n. Фрагменты сигналов, помеченные цифрами “17–31”, представлены при большей скорости раЗвёртки на осциллограммах под соответствуюЩими номерами, места сбора насекомых длЯ Записи сигналов укаЗаны под осциллограммами. Отметка времени вниЗу обЩаЯ длЯ всех осциллограмм." figureDoi="http://doi.org/10.5281/zenodo.13178514" httpUri="https://zenodo.org/record/13178514/files/figure.png" pageId="7" pageNumber="260">Figs. 14–16</figureCitation>
,
<figureCitation id="13034C96FB62FFE2FE15FE2208C2B114" box="[442,511,489,512]" captionStart-0="Figs 17–31" captionStart-1="Fig" captionStartId-0="3.[245,282,1969,1988]" captionStartId-1="4.[284,315,740,759]" captionTargetBox-0="[222,1420,323,1908]" captionTargetBox-1="[245,1477,321,701]" captionTargetId-0="figure-95@3.[213,1434,320,1931]" captionTargetId-1="figure-866@4.[245,1477,320,701]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs 17–31. Aphrophora spp., male calling signal oscillograms. 17–26 ― A. salicina; 27–28 ― A. willemsi; 29–31 ― A. itiliensis sp.n. Signal recording localities are given under the oscillograms. Scale mark at the bottom is the same for all oscillograms. Рис. 17–31. Aphrophora spp., осциллограммы приЗывных сигналов самцов. 17–26 ― A. salicina; 27–28 ― A. willemsi; 29–31 ― A. itiliensis sp.n. Места сбора насекомых длЯ Записи сигналов укаЗаны под осциллограммами. Отметка времени вниЗу обЩаЯ длЯ всех осциллограмм." captionText-1="Fig. 32. Map of froghopper signal recording localities. Рис. 32. Точки сбора насекомых длЯ Записи сигналов." figureDoi-0="http://doi.org/10.5281/zenodo.13178516" figureDoi-1="http://doi.org/10.5281/zenodo.13178518" httpUri-0="https://zenodo.org/record/13178516/files/figure.png" httpUri-1="https://zenodo.org/record/13178518/files/figure.png" pageId="7" pageNumber="260">29–32</figureCitation>
, 42, 45, 51, 63–69
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<paragraph id="8B875013FB62FFE2FF5AFDEF0881B068" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
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MATERIAL EXAMINED.
<typeStatus id="5483EEB1FB62FFE2FE51FDEF0B67B12D" box="[510,602,548,569]" pageId="7" pageNumber="260" type="holotype">Holotype</typeStatus>
, male,
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,
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part of
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, environs of
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, from
<taxonomicName id="4C382B90FB62FFE2FEACFD910842B17B" box="[259,383,602,623]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="B94C8C01FB62FFE2FEACFD910842B17B" box="[259,383,602,623]" italics="true" pageId="7" pageNumber="260">Salix cinerea</emphasis>
</taxonomicName>
,
<date id="FF8676D3FB62FFE2FE25FD910B3CB17B" box="[394,513,602,623]" pageId="7" pageNumber="260" value="2004-07-12">
<collectingDate id="EFC28F3BFB62FFE2FE25FD910B3CB17B" box="[394,513,602,623]" pageId="7" pageNumber="260" value="2004-07-12">12.VII.2004</collectingDate>
</date>
,
<collectorName id="26CD35C5FB62FFE2FDA5FD910BAFB17B" box="[522,658,602,623]" pageId="7" pageNumber="260">D. Tishechkin</collectorName>
, calling signals recorded at 26–29
<superScript id="7C4DFD5BFB62FFE2FE2CFDB808A1B19E" attach="right" box="[387,412,627,650]" fontSize="5" pageId="7" pageNumber="260">oC</superScript>
;
<typeStatus id="5483EEB1FB62FFE2FE08FDBE0B3AB19E" box="[423,519,629,650]" pageId="7" pageNumber="260" type="paratype">paratypes</typeStatus>
: same locality, date, host, and collector,
<specimenCount id="9D3E9B9AFB62FFE2FE9EFD5B0857B1B1" box="[305,362,656,677]" count="3" pageId="7" pageNumber="260" type="male">3 ♂♂</specimenCount>
, calling signals recorded at 26–29
<superScript id="7C4DFD5BFB62FFE2FD63FD450BD8B1B1" attach="right" box="[716,741,654,677]" fontSize="5" pageId="7" pageNumber="260">oC</superScript>
; same locality, host, and collector,
<date id="FF8676D3FB62FFE2FE47FD600B62B1D4" box="[488,607,683,704]" pageId="7" pageNumber="260" value="2004-07-10">
<collectingDate id="EFC28F3BFB62FFE2FE47FD600B62B1D4" box="[488,607,683,704]" pageId="7" pageNumber="260" value="2004-07-10">10.VII.2004</collectingDate>
</date>
,
<specimenCount id="9D3E9B9AFB62FFE2FDC3FD600BAEB1D4" box="[620,659,683,704]" count="1" pageId="7" pageNumber="260" type="male">1 ♂</specimenCount>
; same locality and collector, from
<taxonomicName id="4C382B90FB62FFE2FE39FD0D0B11B1CF" box="[406,556,710,731]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="vinogradovii">
<emphasis id="B94C8C01FB62FFE2FE39FD0D0B11B1CF" box="[406,556,710,731]" italics="true" pageId="7" pageNumber="260">S. vinogradovii</emphasis>
</taxonomicName>
,
<date id="FF8676D3FB62FFE2FD94FD0D0B8EB1CF" box="[571,691,710,731]" pageId="7" pageNumber="260" value="2004-07-13">
<collectingDate id="EFC28F3BFB62FFE2FD94FD0D0B8EB1CF" box="[571,691,710,731]" pageId="7" pageNumber="260" value="2004-07-13">13.VII.2004</collectingDate>
</date>
,
<specimenCount id="9D3E9B9AFB62FFE2FD6FFD0D0BDAB1CF" box="[704,743,710,731]" count="1" pageId="7" pageNumber="260" type="male">1 ♂</specimenCount>
;
<collectingCountry id="F32F1083FB62FFE2FD58FD0D09D1B1E1" name="Russia" pageId="7" pageNumber="260">Russia</collectingCountry>
, northern part of
<collectingRegion id="49FC9EF1FB62FFE2FE05FD2B0B66B1E1" box="[426,603,736,757]" country="Russia" name="Volgograd" pageId="7" pageNumber="260">Volgograd Oblast</collectingRegion>
,
<location id="8EE706C8FB62FFE2FDC5FD2B0A1CB1E1" LSID="urn:lsid:plazi:treatment:0391E105FB62FFE2FE86FE070D22B096:8EE706C8FB62FFE2FDC5FD2B0A1CB1E1" box="[618,801,736,757]" country="Russia" county="Transvolga" municipality="Dyakovka village" name="Kamyshin Region" pageId="7" pageNumber="260" stateProvince="Saratov">Kamyshin Region</location>
, environs of
<location id="8EE706C8FB62FFE2FEE9FD300B1DB004" LSID="urn:lsid:plazi:treatment:0391E105FB62FFE2FE86FE070D22B096:8EE706C8FB62FFE2FEE9FD300B1DB004" box="[326,544,763,784]" country="Russia" county="Transvolga" municipality="Dyakovka village" name="Shcherbatovka village" pageId="7" pageNumber="260" stateProvince="Saratov">Shcherbatovka village</location>
,
<location id="8EE706C8FB62FFE2FD84FD3009C0B03F" LSID="urn:lsid:plazi:treatment:0391E105FB62FFE2FE86FE070D22B096:8EE706C8FB62FFE2FD84FD3009C0B03F" country="Russia" county="Transvolga" municipality="Dyakovka village" name="Shcherbakovsky National Park" pageId="7" pageNumber="260" stateProvince="Saratov">Shcherbakovsky National Park</location>
, from
<taxonomicName id="4C382B90FB62FFE2FE91FCDD08A3B03F" box="[318,414,790,811]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="B94C8C01FB62FFE2FE91FCDD08A3B03F" box="[318,414,790,811]" italics="true" pageId="7" pageNumber="260">S. cinerea</emphasis>
</taxonomicName>
,
<date id="FF8676D3FB62FFE2FE04FCDD0B1CB03F" box="[427,545,790,811]" pageId="7" pageNumber="260" value="2005-07-11">
<collectingDate id="EFC28F3BFB62FFE2FE04FCDD0B1CB03F" box="[427,545,790,811]" pageId="7" pageNumber="260" value="2005-07-11">11.VII.2005</collectingDate>
</date>
,
<collectorName id="26CD35C5FB62FFE2FD83FCDD0B8BB03F" box="[556,694,790,811]" pageId="7" pageNumber="260">D. Tishechkin</collectorName>
,
<specimenCount id="9D3E9B9AFB62FFE2FD6EFCDD0BD8B03F" box="[705,741,790,811]" count="1" pageId="7" pageNumber="260" type="female">1 ♀</specimenCount>
; same locality, host, and collector,
<date id="FF8676D3FB62FFE2FE45FCFA0B5CB052" box="[490,609,817,838]" pageId="7" pageNumber="260" value="2005-07-12">
<collectingDate id="EFC28F3BFB62FFE2FE45FCFA0B5CB052" box="[490,609,817,838]" pageId="7" pageNumber="260" value="2005-07-12">12.VII.2005</collectingDate>
</date>
,
<specimenCount id="9D3E9B9AFB62FFE2FDC1FCFA0B9AB052" box="[622,679,817,838]" count="3" pageId="7" pageNumber="260" type="male">3 ♂♂</specimenCount>
, calling signals recorded at 22–23
<superScript id="7C4DFD5BFB62FFE2FE18FC8008EDB075" attach="right" box="[439,464,843,865]" fontSize="5" pageId="7" pageNumber="260">oC</superScript>
; same locality, host, and collector,
<date id="FF8676D3FB62FFE2FF60FCAC087BB068" box="[207,326,871,892]" pageId="7" pageNumber="260" value="2005-07-13">
<collectingDate id="EFC28F3BFB62FFE2FF60FCAC087BB068" box="[207,326,871,892]" pageId="7" pageNumber="260" value="2005-07-13">13.VII.2005</collectingDate>
</date>
,
<specimenCount id="9D3E9B9AFB62FFE2FEFFFCAC0849B068" box="[336,372,871,892]" count="1" pageId="7" pageNumber="260" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3E9B9AFB62FFE2FE2FFCAC088BB068" box="[384,438,871,892]" count="3" pageId="7" pageNumber="260" type="female">3 ♀♀</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3220398FB62FFE2FF5AFC490811B6D9" pageId="7" pageNumber="260" type="description">
<paragraph id="8B875013FB62FFE2FF5AFC490851B79E" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
MALE CALLING SIGNALS. Calling signal is a phrase lasting for about 1–4 s (
<figureCitation id="13034C96FB62FFE2FE6EFC560B09B0A6" box="[449,564,925,946]" captionStart="Figs 1–16" captionStartId="2.[283,320,1932,1951]" captionTargetBox="[283,1443,322,1857]" captionTargetId="figure-124@2.[269,1454,320,1882]" captionTargetPageId="2" captionText="Figs 1–16. Aphrophora spp., male calling signal oscillograms. 1–11 ― A. salicina; 12–13 ― A. willemsi; 14–16 ― A. itiliensis sp.n. Faster oscillograms of the parts of signals indicated as “17–31” are given under the same numbers, signal recording localities are given under the oscillograms. Scale mark at the bottom is the same for all oscillograms. Рис. 1–16. Aphrophora spp., осциллограммы приЗывных сигналов самцов. 1–11 ― A. salicina; 12–13 ― A. willemsi; 14–16 ― A.itiliensis sp.n. Фрагменты сигналов, помеченные цифрами “17–31”, представлены при большей скорости раЗвёртки на осциллограммах под соответствуюЩими номерами, места сбора насекомых длЯ Записи сигналов укаЗаны под осциллограммами. Отметка времени вниЗу обЩаЯ длЯ всех осциллограмм." figureDoi="http://doi.org/10.5281/zenodo.13178514" httpUri="https://zenodo.org/record/13178514/files/figure.png" pageId="7" pageNumber="260">Figs 14–16</figureCitation>
). Usually, the male produces several phrases separated by gaps averaging from 2–3 up to 10 s. The phrase is a succession of syllables; each syllable consists of one shorter higher-amplitude pulse followed by longer lower-amplitude one (
<figureCitation id="13034C96FB62FFE2FDA4FBC20B43B70A" box="[523,638,1033,1054]" captionStart="Figs 17–31" captionStartId="3.[245,282,1969,1988]" captionTargetBox="[222,1420,323,1908]" captionTargetId="figure-95@3.[213,1434,320,1931]" captionTargetPageId="3" captionText="Figs 17–31. Aphrophora spp., male calling signal oscillograms. 17–26 ― A. salicina; 27–28 ― A. willemsi; 29–31 ― A. itiliensis sp.n. Signal recording localities are given under the oscillograms. Scale mark at the bottom is the same for all oscillograms. Рис. 17–31. Aphrophora spp., осциллограммы приЗывных сигналов самцов. 17–26 ― A. salicina; 27–28 ― A. willemsi; 29–31 ― A. itiliensis sp.n. Места сбора насекомых длЯ Записи сигналов укаЗаны под осциллограммами. Отметка времени вниЗу обЩаЯ длЯ всех осциллограмм." figureDoi="http://doi.org/10.5281/zenodo.13178516" httpUri="https://zenodo.org/record/13178516/files/figure.png" pageId="7" pageNumber="260">Figs 29–31</figureCitation>
). Syllable repetition frequency averages 19–20 syllables/s at 26–29 °C and 16 syllables/s at 22–23 °C. As a rule, at the end of the phrase there are 2–4 syllables of another shape, following each other with longer intervals.
</paragraph>
<paragraph id="8B875013FB62FFE2FF5AFB5B0BFEB7CF" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
DESCRIPTION. Similar to
<taxonomicName id="4C382B90FB62FFE2FDAEFB5B0B55B7B1" baseAuthorityName="Goeze" baseAuthorityYear="1778" box="[513,616,1168,1189]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="salicina">
<emphasis id="B94C8C01FB62FFE2FDAEFB5B0B55B7B1" box="[513,616,1168,1189]" italics="true" pageId="7" pageNumber="260">A. salicina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C382B90FB62FFE2FD39FB5B0A3DB7B1" authorityName="Lallemand" authorityYear="1946" box="[662,768,1168,1189]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="willemsi">
<emphasis id="B94C8C01FB62FFE2FD39FB5B0A3DB7B1" box="[662,768,1168,1189]" italics="true" pageId="7" pageNumber="260">A. willemsi</emphasis>
</taxonomicName>
but coloration grayish, almost without yellowish tinge even at the base of the costal margin of the forewings (Fig. 42).
</paragraph>
<paragraph id="8B875013FB62FFE2FF5AFB2A0BB8B68C" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
The hairs on the forewings only slightly exceed the diameter of punctures (Fig. 45). The end of the ovipositor does not extend or only slightly extends beyond the end of the anal tube (Fig. 51). Pygofer appendages elongated, of almost equal width throughout all their length, with widely rounded apices (Figs 63–65). Styles and penis of the same shape as in other species of the
<taxonomicName id="4C382B90FB62FFE2FEF7FA480883B68C" baseAuthorityName="Goeze" baseAuthorityYear="1778" box="[344,446,1411,1432]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="salicina">
<emphasis id="B94C8C01FB62FFE2FEF7FA480883B68C" box="[344,446,1411,1432]" italics="true" pageId="7" pageNumber="260">A. salicina</emphasis>
</taxonomicName>
group (Figs 66–69).
</paragraph>
<paragraph id="8B875013FB62FFE2FF5AFA550811B6D9" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
Body length (including tegmina):
<specimenCount id="9D3E9B9AFB62FFE2FD92FA550B72B6A7" box="[573,591,1438,1459]" count="1" pageId="7" pageNumber="260" type="male">♂</specimenCount>
, 8.5–9.4 mm;
<specimenCount id="9D3E9B9AFB62FFE2FD73FA550BD3B6A7" box="[732,750,1438,1459]" count="1" pageId="7" pageNumber="260" type="female">♀</specimenCount>
, 9.7– 10.2 mm.
</paragraph>
</subSubSection>
<subSubSection id="C3220398FB62FFE2FF5AFA180B97B540" pageId="7" pageNumber="260" type="diagnosis">
<paragraph id="8B875013FB62FFE2FF5AFA180B97B540" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
DIAGNOSIS. Distinctly differs from other species of the
<taxonomicName id="4C382B90FB62FFE2FF60FA250804B517" baseAuthorityName="Goeze" baseAuthorityYear="1778" box="[207,313,1518,1539]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="salicina">
<emphasis id="B94C8C01FB62FFE2FF60FA250804B517" box="[207,313,1518,1539]" italics="true" pageId="7" pageNumber="260">A. salicina</emphasis>
</taxonomicName>
group by another male calling signal pattern and longer pygofer appendages without expansion near the middle and with widely rounded apices (widened near the middle, with narrow tips in other members of this group).
</paragraph>
</subSubSection>
<subSubSection id="C3220398FB62FFE2FF5AF9910817B5CF" pageId="7" pageNumber="260" type="distribution">
<paragraph id="8B875013FB62FFE2FF5AF9910817B5CF" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
<materialsCitation id="3B505A4EFB62FFE2FF5AF991081BB5CF" country="Russia" location="The Lower Volga region" pageId="7" pageNumber="260" specimenCount="1" stateProvince="Saratov">
DISTRIBUTION.
<location id="8EE706C8FB62FFE2FE03F9910B9EB57B" LSID="urn:lsid:plazi:treatment:0391E105FB62FFE2FE86FE070D22B096:8EE706C8FB62FFE2FE03F9910B9EB57B" box="[428,675,1626,1647]" country="Russia" name="The Lower Volga region" pageId="7" pageNumber="260" stateProvince="Saratov">The Lower Volga region</location>
,
<collectingRegion id="49FC9EF1FB62FFE2FD00F9910BC5B57B" box="[687,760,1626,1647]" country="Russia" name="Saratov" pageId="7" pageNumber="260">Saratov</collectingRegion>
and
<collectingRegion id="49FC9EF1FB62FFE2FF60F9BE080EB59E" box="[207,307,1653,1674]" country="Russia" name="Volgograd" pageId="7" pageNumber="260">Volgograd</collectingRegion>
Oblasts. Despite repeated searches, was not found in the
<collectingRegion id="49FC9EF1FB62FFE2FEA1F95B0896B5B1" box="[270,427,1680,1701]" country="Russia" name="Moscow" pageId="7" pageNumber="260">Moscow Oblast</collectingRegion>
. On the other hand, further findings in the southern regions of European
<collectingCountry id="F32F1083FB62FFE2FD87F9600B55B5D4" box="[552,616,1707,1728]" name="Russia" pageId="7" pageNumber="260">Russia</collectingCountry>
seems to be quite probable
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3220398FB62FFE2FF5AF92A0838B453" pageId="7" pageNumber="260" type="biology_ecology">
<paragraph id="8B875013FB62FFE2FF5AF92A0838B453" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">
HOST PLANTS. Was found mainly on
<taxonomicName id="4C382B90FB62FFE2FDDEF92A0BEDB5E2" box="[625,720,1761,1782]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="B94C8C01FB62FFE2FDDEF92A0BEDB5E2" box="[625,720,1761,1782]" italics="true" pageId="7" pageNumber="260">S. cinerea</emphasis>
</taxonomicName>
, both on single-growing bushes and in mixed thickets of several willow species; only once single male was collected from
<taxonomicName id="4C382B90FB62FFE2FD16F8DC09C2B453" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="vinogradovii">
<emphasis id="B94C8C01FB62FFE2FD16F8DC09C2B453" italics="true" pageId="7" pageNumber="260">S. vinogradovii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3220398FB62FFE2FF5AF8860D22B096" pageId="7" pageNumber="260" type="etymology">
<paragraph id="8B875013FB62FFE2FF5AF88608A8B4A7" blockId="7.[207,805,548,1971]" pageId="7" pageNumber="260">ETYMOLOGY. The name of the new species derives from the name Itil, the ancient name of the Volga River in the Turkic languages. It is given because this species was collected in the Lower Volga region.</paragraph>
<paragraph id="8B875013FB62FFE2FF5AF81F0D45B10D" blockId="7.[207,804,2004,2052]" lastBlockId="7.[842,1441,321,537]" pageId="7" pageNumber="260">
Thus, in
<collectingCountry id="F32F1083FB62FFE2FEFDF81F08AFB4FD" box="[338,402,2004,2025]" name="Russia" pageId="7" pageNumber="260">Russia</collectingCountry>
and adjacent territories, the
<taxonomicName id="4C382B90FB62FFE2FD16F81F0A19B4FD" baseAuthorityName="Goeze" baseAuthorityYear="1778" box="[697,804,2004,2025]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="salicina">
<emphasis id="B94C8C01FB62FFE2FD16F81F0A19B4FD" box="[697,804,2004,2025]" italics="true" pageId="7" pageNumber="260">A. salicina</emphasis>
</taxonomicName>
group includes three taxa. Distinct differences in the calling signal temporal patterns and the fact that in some localities these taxa are sympatric and can even live on the same plant, convincingly indicate that they are good biological species. Distinguishing between these species by coloration is very difficult.
<taxonomicName id="4C382B90FB62FFE2FC0CFE7B0D36B2D1" baseAuthorityName="Goeze" baseAuthorityYear="1778" box="[931,1035,432,453]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="salicina">
<emphasis id="B94C8C01FB62FFE2FC0CFE7B0D36B2D1" box="[931,1035,432,453]" italics="true" pageId="7" pageNumber="260">A. salicina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C382B90FB62FFE2FB93FE7B0D95B2D1" authorityName="Lallemand" authorityYear="1946" box="[1084,1192,432,453]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="willemsi">
<emphasis id="B94C8C01FB62FFE2FB93FE7B0D95B2D1" box="[1084,1192,432,453]" italics="true" pageId="7" pageNumber="260">A. willemsi</emphasis>
</taxonomicName>
differ from each other in the length of the ovipositor and the length of the hairs on the forewings;
<taxonomicName id="4C382B90FB62FFE2FC17FE230D1BB2E9" authorityName="Tishechkin" authorityYear="2023" box="[952,1062,488,509]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="itiliensis" status="sp.n.">
<emphasis id="B94C8C01FB62FFE2FC17FE230D1BB2E9" box="[952,1062,488,509]" italics="true" pageId="7" pageNumber="260">A. itiliensis</emphasis>
</taxonomicName>
<emphasis id="B94C8C01FB62FFE2FB81FE230D63B2E9" bold="true" box="[1070,1118,488,509]" pageId="7" pageNumber="260">
<taxonomicNameLabel id="A27F317AFB62FFE2FB81FE230D63B2E9" box="[1070,1118,488,509]" pageId="7" pageNumber="260" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from these species in the shape of the pygofer processes.
</paragraph>
<paragraph id="8B875013FB62FFE2FCDFFDF00D22B096" blockId="7.[842,1440,571,898]" pageId="7" pageNumber="260">
<emphasis id="B94C8C01FB62FFE2FCDFFDF00D0AB144" bold="true" box="[880,1079,571,592]" pageId="7" pageNumber="260">Acknowledgements</emphasis>
. I am most grateful to Akynaly Dubanaev (Sary-Chelek Biosphere Nature Reserve,
<collectingCountry id="F32F1083FB62FFE2FA8CFD9C0CA8B178" box="[1315,1429,599,620]" name="Kyrgyzstan" pageId="7" pageNumber="260">Kyrgyzstan</collectingCountry>
), Asek Abdykulov (Kara-Balty,
<collectingCountry id="F32F1083FB62FFE2FBC0FDB80DDCB19C" box="[1135,1249,627,648]" name="Kyrgyzstan" pageId="7" pageNumber="260">Kyrgyzstan</collectingCountry>
), and Asel Lee (Kara-Oy,
<collectingCountry id="F32F1083FB62FFE2FC21FD440D3DB1B0" box="[910,1024,655,676]" name="Kazakhstan" pageId="7" pageNumber="260">Kazakhstan</collectingCountry>
) for invaluable help and hospitality during my visits to Central Asia and to Dr. Kirill Kolesnichenko and Dr. Anastasia Antonovskaya (Department of Entomology, Faculty of Biology, M.V. Lomonosov
<collectingRegion id="49FC9EF1FB62FFE2FB30FD290C17B1E3" box="[1183,1322,738,759]" country="Russia" name="Moscow" pageId="7" pageNumber="260">Moscow State</collectingRegion>
University) for the help and support during the fieldwork in the Russian Far East. The reported study was carried out as a part of the Scientific Project of the State Order of the Government of
<collectingCountry id="F32F1083FB62FFE2FCE5FC9A0D3BB072" box="[842,1030,849,870]" name="Russia" pageId="7" pageNumber="260">Russian Federation</collectingCountry>
to Lomonosov
<collectingRegion id="49FC9EF1FB62FFE2FB09FC9A0C0CB072" box="[1190,1329,849,870]" country="Russia" name="Moscow" pageId="7" pageNumber="260">Moscow State</collectingRegion>
University No. 121032300063-3.
</paragraph>
</subSubSection>
</treatment>
</document>

391
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<mods:title id="F8EBB223EC4641A845E000A83CB57B23">Dance flies (Diptera: Empididae) in A. P. Fedtschenko’s Collection from Turkestan: Empis negrobovi species group</mods:title>
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<mods:date id="AE8B117B07A8F9B3774A51C16F8616EA">2023</mods:date>
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<taxonomicName id="4C084D24FFFDFFBEFE80FA42FDD6FA06" ID-CoL="BDP5T" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[292,607,1427,1451]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="jiptykiensis" status="sp. nov.">
<emphasis id="B97CEAB5FFFDFFBEFE80FA42FE6FFA06" box="[292,486,1427,1451]" italics="true" pageId="1" pageNumber="90">Empis jiptykiensis</emphasis>
Shamshev
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,
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<paragraph id="8BB736A7FFFDFFBEFE0BFA67FD9DFA64" blockId="1.[189,775,1427,2062]" box="[431,532,1457,1481]" pageId="1" pageNumber="90">
<figureCitation id="13332A22FFFDFFBEFE0BFA67FD86FA64" box="[431,527,1457,1481]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="1" pageNumber="90">Figs 1–3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFDFFBEFF46FA00FDC1F9E4" pageId="1" pageNumber="90" type="materials_examined">
<paragraph id="8BB736A7FFFDFFBEFF46FA00FDC1F9E4" blockId="1.[189,775,1427,2062]" pageId="1" pageNumber="90">
<materialsCitation id="3B603CFAFFFDFFBEFF46FA00FDC1F9E4" collectingDate="1871-06-24" collectionCode="TYPE, MATERIAL, ZMMU" country="Kyrgyzstan" elevation="3510" latitude="39.6" location="Jiptyk" longLatPrecision="1168" longitude="70.566666" pageId="1" pageNumber="90" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="ED19AE62FFFDFFBEFF46FA00FE18FA44" box="[226,401,1494,1513]" pageId="1" pageNumber="90">TYPE MATERIAL</collectionCode>
.
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,
<specimenCount id="9D0EFD2EFFFDFFBEFE52FA05FD83FA44" box="[502,522,1491,1513]" count="1" pageId="1" pageNumber="90" type="male">♂</specimenCount>
, [
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<collectingCountry id="F31F7637FFFDFFBEFDBAFA00FD01FA44" box="[542,648,1494,1513]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
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]
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[Pamir-Alay mountain system; “
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, on mountain slopes along Shchurovsky Glacier” [
<bibRefCitation id="EF994B56FFFDFFBEFE29F9D0FDB0F9B4" author="Fedtschenko A. P." box="[397,569,1542,1561]" pageId="1" pageNumber="90" refId="ref3281" refString="Fedtschenko A. P. [1871? *] [List of localities visited by the Turkestan Scientific Expedition of the Imperial Society of Friends of Natural History, Anthropology and Ethnography, 1869 - 1871]. Moscow: University printing house (Katkoff &amp; Co.). 9 pp. [In Russian]" type="book" year="1871">Fedtschenko, 1871</bibRefCitation>
]; ~
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,
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; see additional remarks below],
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(
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, terminalia dissected, pinned in a microvial with
<typeStatus id="54B38805FFFDFFBEFE4BF9E0FDB6F9E4" box="[495,575,1590,1609]" pageId="1" pageNumber="90" type="holotype">holotype</typeStatus>
).
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</paragraph>
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DIAGNOSIS. Mid-sized (body about
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) species with brown palpus, legs and halteres; occiput behind eyes, proepisternum, hind femur and abdomen with flattened setae; phallus gently curved, broadened on apical third.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFDFFBCFF46F96FFD71FD3D" lastPageId="3" lastPageNumber="92" pageId="1" pageNumber="90" type="description">
<paragraph id="8BB736A7FFFDFFBEFF46F96FFC19FE5C" blockId="1.[189,775,1427,2062]" lastBlockId="1.[812,1399,315,2071]" pageId="1" pageNumber="90">
DESCRIPTION.
<emphasis id="B97CEAB5FFFDFFBEFE28F96FFE49F963" bold="true" box="[396,448,1721,1742]" pageId="1" pageNumber="90">Male</emphasis>
(
<figureCitation id="13332A22FFFDFFBEFE6AF96EFD81F963" box="[462,520,1720,1742]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="1" pageNumber="90">Fig. 1</figureCitation>
). Body
<quantity id="4CF09B42FFFDFFBEFDFDF96FFD2FF963" box="[601,678,1721,1742]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="90" unit="mm" value="4.8">4.8 mm</quantity>
, wing
<quantity id="4CF09B42FFFDFFBEFD4CF96EFF6FF944" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="90" unit="mm" value="4.5">4.5 mm</quantity>
. Head black, with faint, greyish pruinosity on frons, face (except shiny lower margin), ocellar triangle, postgena and occiput; clypeus shiny; setation black to brown. Eyes dichoptic, ommatidia equally small. Frons broad, below ocellar triangle slightly broader than distance between outer margins of posterior ocelli, somewhat more broadened above antennae; with curiously long marginal setulae. Face very broad, bare. Ocellar setae undifferentiated; ocellar triangle with several subequally long, fine setae. Occiput somewhat convex behind eye laterally; postoculars long, fine; similar dense setae on upper part and numerous long, flattened setae on convex part of occiput; postgena with paler hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than pedicel, with longer setulae; postpedicel 3X longer than basal width, with straight margins; stylus 1.5X shorter than postpedicel. Proboscis long, labrum 1.4X as long as eye height; palpus rather long and somewhat thickened, brown; with numerous long, black, fine setae.
</paragraph>
<paragraph id="8BB736A7FFFDFFBEFCF6FE20FC2DFB91" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="90">
Thorax brown, faintly greyish pruinose, black setose; mesoscutum (
<figureCitation id="13332A22FFFDFFBEFC1CFDC7FC71FD8B" box="[952,1016,529,551]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="1" pageNumber="90">Fig. 2</figureCitation>
) with 4 narrow, indistinct, brownish vittae (dorsal view). Prosternum bare. Proepisternum with tuft of numerous, intermixed slightly flattened and simple setae on lower part, bare on upper part in front of anterior spiracle. Antepronotum with several short, strong setae dorsally and 3–4 longer fine setae laterally on each side. Postpronotal lobe with tuft of mostly long, simple setae. Mesonotal setation (somewhat variable on right and left sides): 1 weak presutural supra-alar (with 3–4 additional finer setae), 1 postsutural supra-alar (with several additional finer setae of different lengths anteriorly and posteriorly), 4–5 notopleurals, 1 very long and 1 very short postalars, about 16 scutellars of different lengths and robustness; additional short, fine setae present on notopleuron anteriorly; acrostichals arranged in 2 close irregular rows, long, fine, lacking on prescutellar depression; presutural dorsocentrals irregularly 3–4-serial, fine, nearly as long as acrostichals, flanked with several similar intra-alar setae, postsutural dorsocentrals mostly uniserial, 3 pairs of strong prescutellars longest. Laterotergite with numerous setae. Anterior and posterior spiracles brownish yellow.
</paragraph>
<paragraph id="8BB736A7FFFDFFBEFCF6FB97FAE7F9C1" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="90">Legs entirely brown; faintly greyish pruinose, including coxae; black setose. Legs structure: robust; hind femur on middle slightly broader than fore and mid femora; hind tibia not-geniculate, slightly narrower on about apical half; fore and hind basitarsi slender. Coxae and trochanters with simple setae. Fore femur with short, stronger (especially closer to apex) setae anteroventrally; very long (longer than femur width) hair-like setae posteroventrally and posteriorly. Fore tibia with moderately long, fine setae dorsally and posterodorsally. Fore tarsomeres with somewhat longer setae dorsally and posterodorsally. Mid femur with complete rows of short, strong anteroventral and similar, longer posteroventral setae, covered with dense, spinule-like setae ventrally. Mid tibia with somewhat longer setulae dorsally, bearing erect spinule-like setulae ventrally. Hind femur with similar ventral setation to mid femur; in addition, bearing anterodorsal fringe of flattened setae (except narrow subapical portion). Hind tibia with similar dorsal setation to mid tibia; covered with erect setulae ventrally; no seta in “comb” at tip behind. Fore and mid basitarsi with some minute, spinule-like setulae ventrally; hind basitarsus only whitish pubescent ventrally.</paragraph>
<paragraph id="8BB736A7FFFDFFBEFCF6F9A7FBC4F8A1" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="90">Wing membrane almost hyaline, faintly uniformly infuscate, with brownish veins; CuA+CuP (anal vein) complete but weakened on middle portion; cell dm short, with elongate apex. Pterostigma absent. Basal costal seta absent. Anal lobe well-developed; axillary incision acute. Squama brownish, brown fringed. Halter brown.</paragraph>
<paragraph id="8BB736A7FFFDFFBEFCF6F8C7FB6BF84C" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="90">Abdomen brown, subshiny, faintly brownish grey pruinose; black setose, covered with long, dense, slightly flattened setae on tergites laterally and sternites (intermixed with some simple setae); short and sparse setae on tergites dorsally, sternite 1 bare. Pregenital segments: segment 8 with tergite and sternite separated; tergite 8 mostly membranous, represented by two weakly sclerotized, narrow lateral sclerites each bearing 5–6 setae; sternite 8 simple.</paragraph>
<paragraph id="8BB736A7FFFDFFBDFCF6F830FB67FDA1" blockId="1.[812,1399,315,2071]" lastBlockId="2.[812,1399,315,551]" lastPageId="2" lastPageNumber="91" pageId="1" pageNumber="90">
Terminalia (
<figureCitation id="13332A22FFFDFFBEFC6EF830FB8DF856" box="[970,1028,2022,2044]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="1" pageNumber="90">Fig. 3</figureCitation>
) moderately large; cercus mostly yellowish to brownish yellow, brownish near base; pubescent with microtrichia, with scattered, dark, fine setulae on outer surface; erect setulae on inner surface anteriorly; epandrial lamella brown, subshiny, covered with dense, uniformly moderately long, fine setae; phallus yellowish. Epandrium divided into two lamellae (dorsal bridge absent); epandrial lamella subtriangular, narrow, with posterior margin broadly rounded (lateral view). Hypandrium mostly membranous, rim-like sclerotized along posterior margin, bare. Cerci separated from epandrium and each other; cercus large, unilobate, subrectangular in lateral view, with rounded posterior margin and straight upper margin, posterior margin reaching nearly level of epandrial lamella apex. Hypoproct inconspicuous. Phallus hidden on about basal 2/3; thick, with broader apical 1/3 extended beyond cerci, more or less gently curved, with apex directed obliquely forward; sulcate on subapical portion. Ejaculatory apodeme extended far beyond basal curvature of phallus, with small lateral wings.
</paragraph>
<paragraph id="8BB736A7FFFEFFBDFCF6FDC7FB98FD8A" blockId="2.[812,1399,315,551]" box="[850,1041,529,551]" pageId="2" pageNumber="91">
<emphasis id="B97CEAB5FFFEFFBDFCF6FDC7FC2AFD8B" bold="true" box="[850,931,529,550]" pageId="2" pageNumber="91">Female.</emphasis>
Unknown.
</paragraph>
<caption id="DF77662FFFFEFFBDFF46F857FAFFF7BC" ID-DOI="http://doi.org/10.5281/zenodo.13178372" ID-Zenodo-Dep="13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="2" pageNumber="91" startId="2.[226,263,1921,1940]" targetBox="[192,1399,577,1906]" targetPageId="2" targetType="figure">
<paragraph id="8BB736A7FFFEFFBDFF46F857FAFEF868" blockId="2.[188,1399,1921,2065]" pageId="2" pageNumber="91">
Figs 1–4.
<taxonomicName id="4C084D24FFFEFFBDFE93F854FE4DF839" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[311,452,1921,1940]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAB5FFFEFFBDFE93F854FEE5F839" box="[311,364,1922,1940]" italics="true" pageId="2" pageNumber="91">Empis</emphasis>
Linnaeus
</taxonomicName>
: 1–3 —
<taxonomicName id="4C084D24FFFEFFBDFDA8F854FD6DF839" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[524,740,1921,1941]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="jiptykiensis" status="sp.n.">
<emphasis id="B97CEAB5FFFEFFBDFDA8F854FD90F839" box="[524,537,1922,1940]" italics="true" pageId="2" pageNumber="91">E</emphasis>
.
<emphasis id="B97CEAB5FFFEFFBDFD80F857FD0CF839" box="[548,645,1921,1940]" italics="true" pageId="2" pageNumber="91">jiptykiensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B97CEAB5FFFEFFBDFD49F857FC91F839" bold="true" box="[749,792,1921,1940]" pageId="2" pageNumber="91">
<taxonomicNameLabel id="A24F57CEFFFEFFBDFD49F857FC91F839" box="[749,792,1921,1940]" pageId="2" pageNumber="91" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 —
<taxonomicName id="4C084D24FFFEFFBDFDD9F84CFC05F801" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" box="[637,908,1945,1965]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B97CEAB5FFFEFFBDFDD9F84CFD03F801" box="[637,650,1946,1964]" italics="true" pageId="2" pageNumber="91">E</emphasis>
.
<emphasis id="B97CEAB5FFFEFFBDFD31F84FFD7DF801" box="[661,756,1945,1964]" italics="true" pageId="2" pageNumber="91">nartshukae</emphasis>
<bibRefCitation id="EF994B56FFFEFFBDFD5FF84FFC05F801" author="Shamshev I. V." box="[763,908,1945,1964]" pageId="2" pageNumber="91" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
, male, phallus, lateral view (from
<bibRefCitation id="EF994B56FFFEFFBDFB12F84FFACEF801" author="Shamshev I. V." box="[1206,1351,1945,1964]" pageId="2" pageNumber="91" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm.
</paragraph>
<paragraph id="8BB736A7FFFEFFBDFF46F818FAFFF7BC" blockId="2.[188,1399,1921,2065]" pageId="2" pageNumber="91">
Рис. 1–4.
<taxonomicName id="4C084D24FFFEFFBDFEE7F818FE5DF84D" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[323,468,1997,2016]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAB5FFFEFFBDFEE7F818FEF1F84D" box="[323,376,1998,2016]" italics="true" pageId="2" pageNumber="91">Empis</emphasis>
Linnaeus
</taxonomicName>
: 1–3 —
<taxonomicName id="4C084D24FFFEFFBDFD8FF818FC84F84D" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[555,781,1997,2017]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="jiptykiensis" status="sp.n.">
<emphasis id="B97CEAB5FFFEFFBDFD8FF818FDB1F84D" box="[555,568,1998,2016]" italics="true" pageId="2" pageNumber="91">E</emphasis>
.
<emphasis id="B97CEAB5FFFEFFBDFDECF81BFD20F84D" box="[584,681,1997,2016]" italics="true" pageId="2" pageNumber="91">jiptykiensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="B97CEAB5FFFEFFBDFCBFF81BFCCFF84D" bold="true" box="[795,838,1997,2016]" pageId="2" pageNumber="91">
<taxonomicNameLabel id="A24F57CEFFFEFFBDFCBFF81BFCCFF84D" box="[795,838,1997,2016]" pageId="2" pageNumber="91" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 —
<taxonomicName id="4C084D24FFFEFFBDFD7FF830FC69F855" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" box="[731,992,2021,2041]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B97CEAB5FFFEFFBDFD7FF830FD61F855" box="[731,744,2022,2040]" italics="true" pageId="2" pageNumber="91">E</emphasis>
.
<emphasis id="B97CEAB5FFFEFFBDFD56F833FCC6F855" box="[754,847,2021,2040]" italics="true" pageId="2" pageNumber="91">nartshukae</emphasis>
<bibRefCitation id="EF994B56FFFEFFBDFCF0F833FC69F855" author="Shamshev I. V." box="[852,992,2021,2040]" pageId="2" pageNumber="91" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
, male, phallus, lateral view (иЗ
<bibRefCitation id="EF994B56FFFEFFBDFB42F833FAFBF855" author="Shamshev I. V." box="[1254,1394,2021,2040]" pageId="2" pageNumber="91" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм.
</paragraph>
</caption>
<paragraph id="8BB736A7FFFFFFBCFF46FEEDFD71FD3D" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
DIFFERENTIAL DIAGNOSIS. Within the key to species of the
<taxonomicName id="4C084D24FFFFFFBCFE87FE81FE4EFEC6" authorityName="Shamshev" authorityYear="2001" box="[291,455,342,364]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="negrobovi">
<emphasis id="B97CEAB5FFFFFFBCFE87FE81FE4EFEC6" box="[291,455,342,364]" italics="true" pageId="3" pageNumber="92">Empis negrobovi</emphasis>
</taxonomicName>
group the new species would run to
<taxonomicName id="4C084D24FFFFFFBCFF7DFEA7FD87FE2B" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" box="[217,526,369,391]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B97CEAB5FFFFFFBCFF7DFEA7FF61FE2B" box="[217,232,369,390]" italics="true" pageId="3" pageNumber="92">E</emphasis>
.
<emphasis id="B97CEAB5FFFFFFBCFF53FEA7FEEBFE2B" box="[247,354,369,390]" italics="true" pageId="3" pageNumber="92">nartshukae</emphasis>
<bibRefCitation id="EF994B56FFFFFFBCFEC8FEA7FD87FE2B" author="Shamshev I. V." box="[364,526,369,391]" pageId="3" pageNumber="92" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
described from the East Pamir of
<collectingCountry id="F31F7637FFFFFFBCFEBEFE5DFEF2FE0C" box="[282,379,395,417]" name="Tajikistan" pageId="3" pageNumber="92">Tajikistan</collectingCountry>
[
<bibRefCitation id="EF994B56FFFFFFBCFE2CFE5DFDB8FE0D" author="Shamshev I. V." box="[392,561,395,417]" pageId="3" pageNumber="92" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
]. These species differ from all other members of the group by a combination of brown palpus, legs and halteres. The new species can be distinguished from
<taxonomicName id="4C084D24FFFFFFBCFE25FE0AFD85FE5D" authorityName="Shamshev" authorityYear="2001" box="[385,524,475,497]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B97CEAB5FFFFFFBCFE25FE0AFE19FE5C" box="[385,400,476,497]" italics="true" pageId="3" pageNumber="92">E</emphasis>
.
<emphasis id="B97CEAB5FFFFFFBCFE05FE0DFD85FE5D" box="[417,524,475,496]" italics="true" pageId="3" pageNumber="92">nartshukae</emphasis>
</taxonomicName>
by 3–4-serial presutural dorsocentral setae (versus irregularly biserial), longer posterodorsal setae on the fore tibia, brownish yellow spiracles and by absolutely different shape of the phallus (
<figureCitation id="13332A22FFFFFFBCFD22FDFDFD6BFDED" box="[646,738,555,577]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="3" pageNumber="92">Figs 3–4</figureCitation>
). It should be noted that the new species differs from all other species of the
<taxonomicName id="4C084D24FFFFFFBCFEEFFDB7FE43FDDB" authorityName="Shamshev" authorityYear="2001" box="[331,458,609,630]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="negrobovi">
<emphasis id="B97CEAB5FFFFFFBCFEEFFDB7FED3FDDB" box="[331,346,609,630]" italics="true" pageId="3" pageNumber="92">E</emphasis>
.
<emphasis id="B97CEAB5FFFFFFBCFECCFDB7FE43FDDB" box="[360,458,609,630]" italics="true" pageId="3" pageNumber="92">negrobovi</emphasis>
</taxonomicName>
group by gently curved phallus (versus S-like bend, with apex directed back, as in
<figureCitation id="13332A22FFFFFFBCFD14FDADFD64FD3D" box="[688,749,635,657]" captionStart="Figs 1–4" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[192,1399,577,1906]" captionTargetId="figure-166@2.[189,1399,576,1907]" captionTargetPageId="2" captionText="Figs 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., male, holotype: 1 — habitus, lateral view; 2 — head and mesonotum, anterior view; 3 — hypopygium, lateral view; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (from Shamshev, 2001, with modificatins). Abbreviations: cerc — cercus; epand — epandrium; hypd — hypandrium; ph — phallus; tg 8 — tergite 8. Scale bar is 1 mm. Рис. 1–4. Empis Linnaeus: 1–3 — E. jiptykiensis Shamshev, sp.n., самец, голотип: 1 — габитус, вид сбоку; 2 — голова и среднеспинка, вид спереди; 3 — гипопигий, вид сбоку; 4 — E. nartshukae Shamshev, 2001, male, phallus, lateral view (иЗ Shamshev, 2001, с иЗменениЯми). СокраЩениЯ: cerc — церк; epand — Эпандрий; hypd — гипандрий; ph — фаллус; tg 8 — 8-й тергит. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178372" httpUri="https://zenodo.org/record/13178372/files/figure.png" pageId="3" pageNumber="92">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFF46FD40FD35FD6A" pageId="3" pageNumber="92" type="etymology">
<paragraph id="8BB736A7FFFFFFBCFF46FD40FD35FD6A" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
<materialsCitation id="3B603CFAFFFFFFBCFF46FD40FD30FD6A" country="Kyrgyzstan" location="Jiptyk" pageId="3" pageNumber="92" specimenCount="1" stateProvince="Osh Shaary">
ETYMOLOGY. The epithet refers to the
<typeStatus id="54B38805FFFFFFBCFDD0FD40FD17FD06" box="[628,670,662,683]" pageId="3" pageNumber="92">type</typeStatus>
locality of the new species,
<location id="8ED7607CFFFFFFBCFEC7FD67FE17FD6A" LSID="urn:lsid:plazi:treatment:03A187B1FFFDFFBCFE80FA42FB10FB71:8ED7607CFFFFFFBCFEC7FD67FE17FD6A" box="[355,414,689,711]" country="Kyrgyzstan" name="Jiptyk" pageId="3" pageNumber="92" stateProvince="Osh Shaary">Jiptyk</location>
(
<collectingCountry id="F31F7637FFFFFFBCFE0FFD67FD97FD6B" box="[427,542,689,710]" name="Kyrgyzstan" pageId="3" pageNumber="92">Kyrgyzstan</collectingCountry>
,
<collectingRegion id="49CCF845FFFFFFBCFD8DFD67FD3BFD6A" box="[553,690,689,711]" country="Kyrgyzstan" name="Osh Shaary" pageId="3" pageNumber="92">Osh Province</collectingRegion>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFF46FD1DFE2CFC7C" pageId="3" pageNumber="92" type="discussion">
<paragraph id="8BB736A7FFFFFFBCFF46FD1DFE2CFC7C" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
REMARKS. Fedtschenko indicated “Jiptyk” (alternative spelling “Jiptik”) as a basic locality of the expedition 1871 during June 22–25 [
<bibRefCitation id="EF994B56FFFFFFBCFE26FCD7FDC9FCBB" author="Fedtschenko A. P." box="[386,576,769,791]" pageId="3" pageNumber="92" refId="ref3281" refString="Fedtschenko A. P. [1871? *] [List of localities visited by the Turkestan Scientific Expedition of the Imperial Society of Friends of Natural History, Anthropology and Ethnography, 1869 - 1871]. Moscow: University printing house (Katkoff &amp; Co.). 9 pp. [In Russian]" type="book" year="1871">Fedtschenko, 1871</bibRefCitation>
]. However, he noted in his comments a pass and a river with the same name. The new species was collected on mountain slopes along Shchurovsky Glacier, i.e., close to upper reaches of Jiptyk River (meeting point of Turkestan and Alay Ranges).
<bibRefCitation id="EF994B56FFFFFFBCFD28FCBDFEAEFC36" author="Fedtschenko A. P." pageId="3" pageNumber="92" refId="ref3339" refString="Fedtschenko A. P. 1875. [A journey to Turkestan. Vol. 1. Pt. 2. In Kokandskoe khanate]. St. Petersburg; Moscow: M. Stasyulevich's printing house. 160 pp. (Izvestia Imperatorskogo Obschestva Lyubiteley Estestvoznania, Antropologii i Etnografii. Vol. 11. Is. 7) [In Russian]" type="book" year="1875">Fedtschenko [1875: 78]</bibRefCitation>
described in details his excursion of June 24, when the new species was collected (e.g., he mentioned many insects visited flowers).
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFF46FC00FF79FBAB" pageId="3" pageNumber="92" type="distribution">
<paragraph id="8BB736A7FFFFFFBCFF46FC00FF79FBAB" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
DISTRIBUTION.
<collectingCountry id="F31F7637FFFFFFBCFE30FC00FD8AFC46" box="[404,515,982,1003]" name="Kyrgyzstan" pageId="3" pageNumber="92">Kyrgyzstan</collectingCountry>
(Pamir-Alay mountain system).
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFE84FBC2FD9DFBE5" pageId="3" pageNumber="92" type="reference_group">
<paragraph id="8BB736A7FFFFFFBCFE84FBC2FD2AFB86" blockId="3.[189,775,315,1249]" box="[288,675,1043,1067]" pageId="3" pageNumber="92">
<taxonomicName id="4C084D24FFFFFFBCFE84FBC2FD2AFB86" ID-CoL="39JNH" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" box="[288,675,1043,1067]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="rohdendorfi">
<emphasis id="B97CEAB5FFFFFFBCFE84FBC2FE63FB86" box="[288,490,1043,1067]" italics="true" pageId="3" pageNumber="92">Empis rohdendorfi</emphasis>
<bibRefCitation id="EF994B56FFFFFFBCFE56FBC5FD2AFB86" author="Shamshev I. V." box="[498,675,1043,1067]" pageId="3" pageNumber="92" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BB736A7FFFFFFBCFE0BFBE6FD9DFBE5" blockId="3.[189,775,315,1249]" box="[431,532,1072,1096]" pageId="3" pageNumber="92">
<figureCitation id="13332A22FFFFFFBCFE0BFBE6FD99FBE5" box="[431,528,1072,1096]" captionStart="Figs 5—6" captionStartId="3.[204,241,2017,2036]" captionTargetBox="[190,1391,1272,2002]" captionTargetId="figure-645@3.[189,1399,1266,2003]" captionTargetPageId="3" captionText="Figs 5—6. Empis rohdendorfi Shamshev, 2001: 5 — male, habitus, lateral view; 6 — female, habitus, lateral view. Scale bar is 1 mm. Рис. 5—6. Empis rohdendorfi Shamshev, 2001: 5 — самец, габитус, вид сбоку; 6 — самка, габитус, вид сбоку. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178374" httpUri="https://zenodo.org/record/13178374/files/figure.png" pageId="3" pageNumber="92">Figs 5–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFF46FB83FE9DFB2D" pageId="3" pageNumber="92" type="description">
<paragraph id="8BB736A7FFFFFFBCFF46FB83FE9DFB2D" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
<materialsCitation id="3B603CFAFFFFFFBCFF46FB83FE98FB2D" country="Uzbekistan" location="Tashkent" pageId="3" pageNumber="92" specimenCount="1">
<bibRefCitation id="EF994B56FFFFFFBCFF46FB83FE2AFBC5" author="Shamshev I. V." box="[226,419,1109,1128]" pageId="3" pageNumber="92" pagination="195 - 4227" refId="ref3457" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region // An International Journal of Dipterological Research. Vol. 12. P. 195 - 4227." type="book chapter" year="2001">Shamshev, 2001: 221</bibRefCitation>
, figs 23, 24.
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locality:
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,
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFF46FB53FD03FB4D" pageId="3" pageNumber="92" type="materials_examined">
<paragraph id="8BB736A7FFFFFFBCFF46FB53FD03FB4D" blockId="3.[189,775,315,1249]" pageId="3" pageNumber="92">
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<collectionCode id="ED19AE62FFFFFFBCFF46FB53FEC5FB35" box="[226,332,1157,1176]" pageId="3" pageNumber="92">MATERIAL</collectionCode>
EXAMINED.
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<collectingCountry id="F31F7637FFFFFFBCFE63FB53FDA0FB35" box="[455,553,1157,1176]" name="Uzbekistan" pageId="3" pageNumber="92">Uzbekistan</collectingCountry>
.
</emphasis>
<location id="8ED7607CFFFFFFBCFD94FB53FDF7FB35" LSID="urn:lsid:plazi:treatment:03A187B1FFFDFFBCFE80FA42FB10FB71:8ED7607CFFFFFFBCFD94FB53FDF7FB35" box="[560,638,1157,1176]" country="Uzbekistan" name="Tashkent" pageId="3" pageNumber="92">Tashkent</location>
</materialsCitation>
:
<date id="FFB61067FFFFFFBCFD20FB53FD52FB35" box="[644,731,1157,1176]" pageId="3" pageNumber="92" value="1871-04-11">11.iv.1871</date>
(
<specimenCount id="9D0EFD2EFFFFFFBCFD43FB53FD76FB34" box="[743,767,1155,1177]" count="1" pageId="3" pageNumber="92" type="female">1 ♀</specimenCount>
) [“
<quantity id="4CF09B42FFFFFFBCFF6DFB4BFE81FB1C" box="[201,264,1181,1201]" metricMagnitude="2" metricUnit="m" metricValue="4.26" pageId="3" pageNumber="92" unit="m" value="426.0">426 m</quantity>
, Chimkent road to Bossa, steppe” [
<bibRefCitation id="EF994B56FFFFFFBCFDEFFB4BFD7DFB1D" author="Fedtschenko A. P." box="[587,756,1181,1200]" pageId="3" pageNumber="92" refId="ref3281" refString="Fedtschenko A. P. [1871? *] [List of localities visited by the Turkestan Scientific Expedition of the Imperial Society of Friends of Natural History, Anthropology and Ethnography, 1869 - 1871]. Moscow: University printing house (Katkoff &amp; Co.). 9 pp. [In Russian]" type="book" year="1871">Fedtschenko, 1871</bibRefCitation>
]],
<date id="FFB61067FFFFFFBCFF19FB63FE9AFB65" box="[189,275,1205,1224]" pageId="3" pageNumber="92" value="1871-04-22">22.iv.1871</date>
(
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),
<date id="FFB61067FFFFFFBCFEF7FB63FE23FB65" box="[339,426,1205,1224]" pageId="3" pageNumber="92" value="1871-04-23">23.iv.1871</date>
(
<specimenCount id="9D0EFD2EFFFFFFBCFE12FB63FE64FB64" box="[438,493,1203,1225]" count="3" pageId="3" pageNumber="92" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0EFD2EFFFFFFBCFE53FB63FD9BFB64" box="[503,530,1203,1225]" count="4" pageId="3" pageNumber="92" type="female">4 ♀</specimenCount>
),
<materialsCitation id="3B603CFAFFFFFFBCFD87FB63FF59FB4D" collectingDate="1871-04-24" collectionCode="ZISP" country="Uzbekistan" pageId="3" pageNumber="92" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<date id="FFB61067FFFFFFBCFD87FB63FDF3FB65" box="[547,634,1205,1224]" pageId="3" pageNumber="92" value="1871-04-24">
<collectingDate id="EFF2E98FFFFFFFBCFD87FB63FDF3FB65" box="[547,634,1205,1224]" pageId="3" pageNumber="92" value="1871-04-24">24.iv.1871</collectingDate>
</date>
(
<collectionCode id="ED19AE62FFFFFFBCFD21FB63FD3BFB65" box="[645,690,1205,1224]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="92">ZISP</collectionCode>
,
<specimenCount id="9D0EFD2EFFFFFFBCFD1EFB63FD78FB64" box="[698,753,1203,1225]" count="2" pageId="3" pageNumber="92" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0EFD2EFFFFFFBCFD58FB63FF41FB4C" count="1" pageId="3" pageNumber="92" type="female">1 ♀</specimenCount>
)
</materialsCitation>
,
<date id="FFB61067FFFFFFBCFF7DFB1BFEB8FB4D" box="[217,305,1229,1248]" pageId="3" pageNumber="92" value="1871-04-25">25.iv.1871</date>
(
<specimenCount id="9D0EFD2EFFFFFFBCFE9AFB1BFED6FB4C" box="[318,351,1227,1249]" count="1" pageId="3" pageNumber="92" type="male">1 ♂</specimenCount>
), no locality label,
<date id="FFB61067FFFFFFBCFE5BFB1BFDDFFB4D" box="[511,598,1229,1248]" pageId="3" pageNumber="92" value="1871-04-22">22.iv.1871</date>
(
<specimenCount id="9D0EFD2EFFFFFFBCFDC0FB1BFDF4FB4C" box="[612,637,1227,1249]" count="1" pageId="3" pageNumber="92" type="female">1 ♀</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFCF6FEEDFA87FE5C" pageId="3" pageNumber="92" type="discussion">
<paragraph id="8BB736A7FFFFFFBCFCF6FEEDFA87FE5C" blockId="3.[812,1399,315,1031]" pageId="3" pageNumber="92">
REMARKS. This species was described after single male collected by B.B. Rohdendorf from “Tashkent”. The collection of A.P. Fedtschenko includes
<specimenCount id="9D0EFD2EFFFFFFBCFB21FEA7FA82FE2A" box="[1157,1291,369,391]" count="13" pageId="3" pageNumber="92" type="generic">13 specimens</specimenCount>
of
<taxonomicName id="4C084D24FFFFFFBCFA8BFEA7FCF7FE0D" authorityName="Shamshev" authorityYear="2001" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="rohdendorfi">
<emphasis id="B97CEAB5FFFFFFBCFA8BFEA7FAB7FE2B" box="[1327,1342,369,390]" italics="true" pageId="3" pageNumber="92">E</emphasis>
.
<emphasis id="B97CEAB5FFFFFFBCFAE9FEA7FCF7FE0D" italics="true" pageId="3" pageNumber="92">rohdendorfi</emphasis>
</taxonomicName>
taken from the same locality and
<specimenCount id="9D0EFD2EFFFFFFBCFB49FE5AFAFFFE0C" box="[1261,1398,395,417]" count="1" pageId="3" pageNumber="92" type="generic">one specimen</specimenCount>
without locality label. The female of
<taxonomicName id="4C084D24FFFFFFBCFB00FE71FABCFE16" authorityName="Shamshev" authorityYear="2001" box="[1188,1333,422,444]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="rohdendorfi">
<emphasis id="B97CEAB5FFFFFFBCFB00FE71FB3AFE11" box="[1188,1203,423,444]" italics="true" pageId="3" pageNumber="92">E</emphasis>
.
<emphasis id="B97CEAB5FFFFFFBCFB66FE70FABCFE16" box="[1218,1333,422,443]" italics="true" pageId="3" pageNumber="92">rohdendorfi</emphasis>
</taxonomicName>
is described for the first time herein. In addition, we provide a habitus photo of the male of this species (
<figureCitation id="13332A22FFFFFFBCFB61FE0DFA88FE5C" box="[1221,1281,475,497]" captionStart="Figs 5—6" captionStartId="3.[204,241,2017,2036]" captionTargetBox="[190,1391,1272,2002]" captionTargetId="figure-645@3.[189,1399,1266,2003]" captionTargetPageId="3" captionText="Figs 5—6. Empis rohdendorfi Shamshev, 2001: 5 — male, habitus, lateral view; 6 — female, habitus, lateral view. Scale bar is 1 mm. Рис. 5—6. Empis rohdendorfi Shamshev, 2001: 5 — самец, габитус, вид сбоку; 6 — самка, габитус, вид сбоку. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178374" httpUri="https://zenodo.org/record/13178374/files/figure.png" pageId="3" pageNumber="92">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFCF6FE20FC6CFC41" pageId="3" pageNumber="92" type="description">
<paragraph id="8BB736A7FFFFFFBCFCF6FE20FC6CFC41" blockId="3.[812,1399,315,1031]" pageId="3" pageNumber="92">
DESCRIPTION.
<emphasis id="B97CEAB5FFFFFFBCFC59FE20FBC1FDA6" bold="true" box="[1021,1096,502,523]" pageId="3" pageNumber="92">Female</emphasis>
(
<figureCitation id="13332A22FFFFFFBCFBF3FE20FB1DFDA1" box="[1111,1172,502,524]" captionStart="Figs 5—6" captionStartId="3.[204,241,2017,2036]" captionTargetBox="[190,1391,1272,2002]" captionTargetId="figure-645@3.[189,1399,1266,2003]" captionTargetPageId="3" captionText="Figs 5—6. Empis rohdendorfi Shamshev, 2001: 5 — male, habitus, lateral view; 6 — female, habitus, lateral view. Scale bar is 1 mm. Рис. 5—6. Empis rohdendorfi Shamshev, 2001: 5 — самец, габитус, вид сбоку; 6 — самка, габитус, вид сбоку. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178374" httpUri="https://zenodo.org/record/13178374/files/figure.png" pageId="3" pageNumber="92">Fig. 6</figureCitation>
, described for the first time). Body 4.1–4.7 (in male
<quantity id="4CF09B42FFFFFFBCFBE2FDC6FB37FD8B" box="[1094,1214,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.1" metricValueMin="4.7" pageId="3" pageNumber="92" unit="mm" value="4.9" valueMax="5.1" valueMin="4.7">4.7–5.1 mm</quantity>
); wing
<quantity id="4CF09B42FFFFFFBCFAA7FDC6FAFFFD8B" box="[1283,1398,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="3" pageNumber="92" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
(in male
<quantity id="4CF09B42FFFFFFBCFC29FDFDFC70FDED" box="[909,1017,555,577]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="3" pageNumber="92" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
). Frons broader than in male, below ocellar triangle slightly broader than distance between outer margins of posterior ocelli, parallel-sided. Occiput with sparser, simple setae laterally. Mesonotum with shorter setae. Legs generally with sparser and shorter setae; mid and hind femora not pubescent ventrally, with shorter, finer and spars- er anteroventral and posteroventral setae; mid and hind tibiae with simple setulae ventrally (in male mid tibia covered with dense, minute, erect setulae ventrally; hind tibia with scattered minute setulae ventrally), 2 short anteroventral setae on middle portion. Wing slightly uniformly infuscate. Abdomen with segments 1–7 densely whitish grey pruinose, segment 8 black, subshiny; mostly covered with scattered, minute setulae; tergites 1 and 2 with short, sparse, pale, hair-like setae laterally; sternites 6 and 7 with several, rather long, fine posteromarginal setae. Cercus dark, long, slender, with simple minute setulae.
</paragraph>
</subSubSection>
<subSubSection id="C312652CFFFFFFBCFCF6FC27FB10FB71" pageId="3" pageNumber="92" type="distribution">
<paragraph id="8BB736A7FFFFFFBCFCF6FC27FBF7FBAA" blockId="3.[812,1399,315,1031]" box="[850,1150,1009,1031]" pageId="3" pageNumber="92">
DISTRIBUTION.
<collectingCountry id="F31F7637FFFFFFBCFBAEFC27FBF3FBAA" box="[1034,1146,1009,1031]" name="Uzbekistan" pageId="3" pageNumber="92">Uzbekistan</collectingCountry>
.
</paragraph>
<paragraph id="8BB736A7FFFFFFBCFCF6FBF0FB10FB71" blockId="3.[812,1399,1062,1244]" pageId="3" pageNumber="92">
<emphasis id="B97CEAB5FFFFFFBCFCF6FBF0FB90FB91" bold="true" box="[850,1049,1062,1084]" pageId="3" pageNumber="92">Acknowledgements</emphasis>
. I am indebted to Andrey Ozerov (ZMMU) for the loan of specimens used in this study. Boris Korotyaev and Julia Astafurova kindly helped with publications of A.P. Fedtschenko. This study was performed within the frames of the Russian State Research Project no. 122031100272-3 of the Ministry of Science and Higher Education of the
<collectingCountry id="F31F7637FFFFFFBCFC72FB10FB1CFB71" box="[982,1173,1222,1244]" name="Russia" pageId="3" pageNumber="92">Russian Federation</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="4C194D30F301FFFCE49A6C3C9CFED2E7" authority="Kataev" authorityName="Kataev" authorityYear="2021" box="[925,1221,628,652]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F301FFFCE49A6C3C9C54D2E0" box="[925,1135,628,652]" italics="true" pageId="1" pageNumber="449">Coleolissus inessae</emphasis>
Kataev
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,
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<subSubSection id="C3036538F301FFFCE3186CD99CB8D2C5" box="[1055,1155,657,681]" pageId="1" pageNumber="449" type="description">
<paragraph id="8BA636B3F301FFFCE3186CD99CB8D2C5" blockId="1.[812,1399,628,2068]" box="[1055,1155,657,681]" pageId="1" pageNumber="449">
<figureCitation id="13222A36F301FFFCE3186CD99C44D2C5" box="[1055,1151,657,681]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Figs 1–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F301FFFCE4556CFE9BD8D295" pageId="1" pageNumber="449" type="materials_examined">
<paragraph id="8BA636B3F301FFFCE4556CFE9BD8D295" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
<materialsCitation id="3B713CEEF301FFFCE4556CFE9BD8D295" collectingDate="2013-11-02" collectingDateMax="2013-11-05" collectingDateMin="2013-11-02" collectionCode="ZIN" collectorName="S. Saluk" country="India" elevation="530" latitude="14.220667" location="Falls" longLatPrecision="1" longitude="74.80785" municipality="Gats Mts" pageId="1" pageNumber="449" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
MATERIAL.
<typeStatus id="54A28811F301FFFCE4CD6CFE9C25D2A5" box="[970,1054,694,713]" pageId="1" pageNumber="449" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1FFD3AF301FFFCE3236CF19C03D2A5" box="[1060,1080,697,713]" count="1" pageId="1" pageNumber="449" type="male">♂</specimenCount>
, “
<collectingCountry id="F30E7623F301FFFCE34E6CFE9CB0D2A5" box="[1097,1163,694,713]" name="India" pageId="1" pageNumber="449">INDIA</collectingCountry>
: W
<collectingRegion id="49DDF851F301FFFCE3AD6CFE9D3ED2A5" box="[1194,1285,694,713]" country="India" name="Karnataka" pageId="1" pageNumber="449">Karnataka</collectingRegion>
, W
<collectingMunicipality id="6BC2ACC9F301FFFCE2236CFE9D48D2A6" box="[1316,1395,694,714]" pageId="1" pageNumber="449">Gats Mts</collectingMunicipality>
, jog
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,
<quantity id="4CE19B56F301FFFCE4816C869BFAD28E" box="[902,961,718,738]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="449" unit="m" value="530.0">
<elevation id="0034D180F301FFFCE4816C869BFAD28E" box="[902,961,718,738]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="449" unit="m" value="530.0">530 m</elevation>
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, at light,
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<geoCoordinate id="EE2D5074F301FFFCE3886C869CC5D28D" box="[1167,1278,718,737]" degrees="74" direction="east" minutes="48.471" orientation="longitude" pageId="1" pageNumber="449" precision="1" value="74.80785">74°48.471´E</geoCoordinate>
,
<date id="FFA71073F301FFFCE20E6C869D4DD28D" box="[1289,1398,718,737]" pageId="1" pageNumber="449" value="2013-11-02" valueMax="2013-11-05" valueMin="2013-11-02">
<collectingDate id="EFE3E99BF301FFFCE20E6C869D4DD28D" box="[1289,1398,718,737]" pageId="1" pageNumber="449" value="2013-11-02" valueMax="2013-11-05" valueMin="2013-11-02">2–5.XI.2013</collectingDate>
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<collectorName id="26EC5365F301FFFCE42B6CAE9B4ED295" box="[812,885,742,761]" pageId="1" pageNumber="449">S. Saluk</collectorName>
leg.” (
<collectionCode id="ED08AE76F301FFFCE4B66CAE9BE2D295" box="[945,985,742,761]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="449" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036538F301FFFEE4556CB69988D72D" lastPageId="3" lastPageNumber="451" pageId="1" pageNumber="449" type="description">
<paragraph id="8BA636B3F301FFFCE4556CB69D19D342" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
DESCRIPTION (male). Body elongate, medium-sized, length
<quantity id="4CE19B56F301FFFCE4686D519B86D342" box="[879,957,793,814]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="449" unit="mm" value="7.8">7.8 mm</quantity>
, width 3.0 mm. Habitus as in
<figureCitation id="13222A36F301FFFCE3E46D519D26D342" box="[1251,1309,793,815]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="8BA636B3F301FFFCE4556D7B9C58D3D8" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">Body pitch black, shiny and iridescent on dorsal and ventral sides, with labrum, base of mandibles and very narrow lateral margins of pronotum reddish brown; palpi, antennae and legs yellowish brown; antennomeres 3–11 and apices of tibiae very slightly infuscate.</paragraph>
<paragraph id="8BA636B3F301FFFCE4556DF19D37D735" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
Head medium-sized (HWmax/PWmax 0.68, HWmin/ PWmax 0.53), impunctate. Eyes large, moderately convex (HWmax/HWmin 1.28), separated ventrally from buccal fissure by very short distance about as wide as antennomere 1 basally. Genae glabrous. Tempora short, oblique, gently sloping to neck. Labrum slightly convex, very shallowly concave anteriorly. Clypeus somewhat flat, slightly emarginated along anterior margin, with a pair of lateral setigerous pores and a pair of very short longitudinal wrinkles each adjacent medially to lateral pore. Fronto-clypeal suture fine, slightly impressed, almost straight. Frontal foveae like elongate short pit without prolongation on clypeus. Fronto-ocular furrows very fine, vaguely reaching supraocular furrows. Frons with angulate shallow depression and with a few very fine micropunctures at lateral margins of this depression. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by diameter of pores.
<collectingRegion id="49DDF851F301FFFCE35E6BD19C90D5C2" box="[1113,1195,1433,1454]" country="Myanmar" name="Chin" pageId="1" pageNumber="449">Mentum</collectingRegion>
separated from submentum by complete transverse suture (
<figureCitation id="13222A36F301FFFCE3B16BFB9CC8D5A5" box="[1206,1267,1459,1481]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Fig. 2</figureCitation>
), with a very short and wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite almost parallel-sided, truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, moderately wide, markedly longer than ligular sclerite and separat- ed from it apically by narrow notches. Basal labial palpomere not carinate; penultimate labial palpomere about as long as ultimate one. Mandibles very moderately elongate, evenly curved apically, with wide bases; left mandible blunted at tip. Dorsal microsculpture very fine, obliterated, consisting of indistinct, more or less isodiametric meshes. Antennae slen- der, surpassing pronotal basal edge by two apical antennomeres, pubescent from apical half of antennomere 3, with antennomeres 4–8 about two times as long as wide and basal antennomere slightly longer than antennomere 3.
</paragraph>
<paragraph id="8BA636B3F301FFFFE45569179CB7D7BB" blockId="1.[812,1399,628,2068]" lastBlockId="2.[812,1399,1799,2061]" lastPageId="2" lastPageNumber="450" pageId="1" pageNumber="449">Pronotum transverse, comparatively long for genus (PWmax/PL 1.34), widest slightly before the middle, very moderately narrowed apically and basally (PWmax/PWmin-ap 1.44, PWmax/PWmin-bas 1.20), with one lateral seta inserted at the end of the anterior third. Sides rounded along entire length; lateral bead complete, throughout very narrow. Apical margin shallowly emarginated, almost straight medially, very narrowly bordered along entire length; border slightly obliterated at the middle. Apical angles rounded, slightly protruding anteriorly. Basal margin almost straight in middle portion, slightly rounded laterally, bordered along entire length, not ciliate, slightly longer than apical margin (PWmin-bas/PWmin-ap 1.20) and almost as long as base of elytra between humeral angles. Basal angles obtuse, rounded at apex. Disc convex. Lateral depressions beginning from lateral setae, very shallow, widened basally and disappearing before the apical quarter. Basal foveae elongate oval, shallow, isolated from lateral margins and from each other by convex areas. Median line distinct, slightly impressed, isolat- ed from apical margin and touching basal margin. Anterior transverse depression wide and shallow. Surface with dense distinct non-setigerous punctures basally and along sides, with sparser and finer punctation at apical margin including anterior transverse depression. Microsculpture highly obliterated, not forming distinct meshes.</paragraph>
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<paragraph id="8BA636B3F302FFFFE7E568CF9BA5D6DE" blockId="2.[189,1399,1671,1766]" pageId="2" pageNumber="450">
Figs 1–6.
<taxonomicName id="4C194D30F302FFFFE63168CF99E6D6F5" authorityName="Kataev" authorityYear="2021" box="[310,477,1671,1690]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F302FFFFE63168CF99E6D6F5" box="[310,477,1671,1690]" italics="true" pageId="2" pageNumber="450">Coleolissus inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F302FFFFE6E668CF9A37D6F6" bold="true" box="[481,524,1671,1690]" pageId="2" pageNumber="450">
<taxonomicNameLabel id="A25E57DAF302FFFFE6E668CF9A37D6F6" box="[481,524,1671,1690]" pageId="2" pageNumber="450" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6).
</paragraph>
<paragraph id="8BA636B3F302FFFFE7E568F39C73D689" blockId="2.[189,1399,1671,1766]" pageId="2" pageNumber="450">
Рис. 1–6.
<taxonomicName id="4C194D30F302FFFFE63868F299D2D6A1" authorityName="Kataev" authorityYear="2021" box="[319,489,1722,1741]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F302FFFFE63868F299D2D6A1" box="[319,489,1722,1741]" italics="true" pageId="2" pageNumber="450">Coleolissus inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F302FFFFE6F568F29A27D6A1" bold="true" box="[498,540,1722,1741]" pageId="2" pageNumber="450">
<taxonomicNameLabel id="A25E57DAF302FFFFE6F568F29A27D6A1" box="[498,540,1722,1741]" pageId="2" pageNumber="450" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6).
</paragraph>
</caption>
<paragraph id="8BA636B3F302FFFEE45569949A20D3DA" blockId="2.[812,1399,1799,2061]" lastBlockId="3.[189,775,315,2071]" lastPageId="3" lastPageNumber="451" pageId="2" pageNumber="450">
Elytra convex, elongate, only slightly wider than pronotum (EL/EW 1.64, EL/PL 2.61, EW/PWmax 1.19), widest before the apical third; sides very slightly diverging in the apical two thirds, with weak sinuation before the middle, and roundly converging in the apical third; subapical sinuation distinct but shallow. Humeri rounded, with an indistinct denticle recognizable in caudal view. Sutural angle narrowly rounded at tip, not extended posteriorly. Basal edge slightly arched, forming a sharp, very obtuse angle with lateral margin. Striae comparatively wide, impunctate, impressed along entire length, reaching anteriorly basal elytral border, with granulate microsculpture on bottom. Intervals glabrous, almost flat on disc, gradually becoming convex and narrow at apex; intervals 8 and 9 throughout and other intervals just at apex irregularly punctate; punctation of intervals 8 and
<quantity id="4CE19B56F303FFFEE5E76C339B3CD2FD" box="[736,775,635,657]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="451" unit="in" value="9.0">9 in</quantity>
apical half very fine and sparse. Parascutellar (abbreviate) striole long, about 0.2 as long as elytron, with a large setigerous pore basally isolated from basal elytral border. Interval 3 with a series of seven very distinct discal setigerous pores adjoining to stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series without distinct gap at middle, consisting of 22–25 setigerous pores. Lateral groove narrow and flat throughout, without distinct elongate convexity along this groove apically. Microsculpture highly obliterated on intervals, without distinct meshes or lines, granulate in lateral groove.
</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56DF399F3D3BD" blockId="3.[189,775,315,2071]" box="[226,456,955,977]" pageId="3" pageNumber="451">Wings fully developed.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56D9E9A69D450" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Ventral side of thorax glabrous except for sparse very fine short setae on prosternum anteriorly, almost throughout finely punctate laterally. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56A099A10D58A" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Metacoxae each only with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side. Protibia on dorsal side without longitudinal sulcus, with two similar sized preapical spines on outer margin. Metafemur ventrally with two setae at posterior margin and without setae at anterior margin. Pro- and mesotarsi dorsally with a few very fine and short setae. Protarsomeres 1–4 and mesotarsomeres 2–4 (in male) markedly widened, short, with biseriate adhesive scales ventrally; mesotarsomere 1 not widened, elongate, only slightly shorter than mesotarsomeres 2 and 3 combined, with a pair of very small scales just at apex. Metatarsus slender, about as long as HWmax, with tarsomeres 2 and 3 weakly and tarsomeres 3 and 4 markedly widened distally; metatarsomere 1 slightly longer than metatarsomeres 2 and 3 combined. Tarsomere 5 with two pairs of ventro-lateral setae.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56BA39AF3D63D" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Abdominal sternites mostly smooth, only two basal ones with very fine non-setigerous punctures; last visible sternite (VII) (in male) very widely rounded at apex, with one pair of marginal setae; these setae not removed from margin.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E5681E9988D72D" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">
Aedeagus (
<figureCitation id="13222A36F303FFFEE64A681E9999D600" box="[333,418,1622,1644]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Figs 3–6</figureCitation>
) with comparatively large parameres (
<figureCitation id="13222A36F303FFFEE7C468399923D6EA" box="[195,280,1649,1671]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Figs 3, 4</figureCitation>
) and basal bulb. Median lobe of aedeagus in lateral view (
<figureCitation id="13222A36F303FFFEE7FA68C39900D6CD" box="[253,315,1675,1697]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Fig. 5</figureCitation>
) arcuate on dorsal side, with ventral margin almost straight in middle portion and convex in the apical quarter; apex sharply hooged ventrally; in dorsal view medi- an lobe (
<figureCitation id="13222A36F303FFFEE6126893996AD69D" box="[277,337,1755,1777]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Fig. 6</figureCitation>
) almost straight, with terminal lamella very wide and short, rounded apically. Apical orifice in dorsal position, wide apically. Internal sac with spiny formation medially, without spines.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE7E5690E9C58D107" pageId="3" pageNumber="451" type="etymology">
<paragraph id="8BA636B3F303FFFEE7E5690E9AEBD7C0" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">
ETYMOLOGY. Named after Prof. Inessa Kh. Sharova (1931–2021), the famous entomologist and soil zoologist, the teacher of many carabidologists in the former Soviet Union and
<collectingCountry id="F30E7623F303FFFEE7E269DE991DD7C0" box="[229,294,1942,1964]" name="Russia" pageId="3" pageNumber="451">Russia</collectingCountry>
, to whose memory this volume is dedicated.
</paragraph>
<paragraph id="8BA636B3F303FFFEE7E569F99C58D107" blockId="3.[189,775,315,2071]" lastBlockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">COMPARISON. This new species is markedly distinguished from all other congeners by elongate body with comparatively long pronotum which is not depressed latero-basally. In addition, unlike most of other species, its mandibles are shorter and abdominal sternite VII (in male) is with only one pair of marginal setae.</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE4556F399CA9D1CD" pageId="3" pageNumber="451" type="distribution">
<paragraph id="8BA636B3F303FFFEE4556F399CA9D1CD" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">
DISTRIBUTION. Known from the Western Ghats near Jog Falls, Western
<collectingRegion id="49DDF851F303FFFEE4EE6FC39C74D1CD" box="[1001,1103,395,417]" country="India" name="Karnataka" pageId="3" pageNumber="451">Karnataka</collectingRegion>
,
<collectingCountry id="F30E7623F303FFFEE35E6FC39CB5D1CD" box="[1113,1166,395,417]" name="India" pageId="3" pageNumber="451">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE4556FEE9B4DD87A" pageId="3" pageNumber="451" type="discussion">
<paragraph id="8BA636B3F303FFFEE4556FEE9B4DD87A" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">
REMARKS. This new species is rather dissimilar to any other described species and at first I hesitated to place it in any described genus. Such characters as the distinct fronto-ocular furrows, glabrous paraglossae, several setae on penultimate labial palpomere, elytral interval 3 with row of setigerous pores, long metatarsomere 1 and pro- and mesotarsi of male with biseriate adhesive scales are clearly indicate that it is a member of the Selenophori group. In having moderately elongate mandibles and glabrous, almost subcylindrical body without latero-basal depressions on pronotum, it looks as somewhat intermediate form between
<taxonomicName id="4C194D30F303FFFEE42B6C839BA6D28C" authorityName="Bates" authorityYear="1892" box="[812,925,715,736]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE42B6C839BA6D28C" box="[812,925,715,736]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE4CB6C849C07D28D" authorityName="Chaudoir" authorityYear="1854" box="[972,1084,716,737]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4CB6C849C07D28D" box="[972,1084,716,737]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
. Although these taxa are always regarded as separate genera, the structural differences between them are not evident [
<bibRefCitation id="EF884B42F303FFFEE3406D499CEAD37A" author="Kataev B. M." box="[1095,1233,768,790]" pageId="3" pageNumber="451" pagination="43 - 55" refId="ref6963" refString="Kataev B. M. 2006. A taxonomic review of the genus Oesyperus (Coleoptera: Carabidae: Harpalini) // Zootaxa. Vol. 1286. P. 43 - 55." type="book chapter" year="2006">Kataev, 2006</bibRefCitation>
] and assignment of some species to one or another genus is difficult since the body form is rather variable in the both genera. For example, two apparently closely related species with elongate body were described by one author [
<bibRefCitation id="EF884B42F303FFFEE3A26D249D29D3ED" author="Ito N." box="[1189,1298,875,897]" pageId="3" pageNumber="451" pagination="39 - 43" refId="ref6259" refString="Ito N. 1997 b. Two new species of the genus Oxycentrus from Vietnam (Coleoptera, Carabidae, Harpalini) // Ent. Rev. Japan. Vol. 52. No. 1. P. 39 - 43." type="journal article" year="1997">Ito, 1997b</bibRefCitation>
,
<bibRefCitation id="EF884B42F303FFFEE2186D239D69D3EC" author="Ito N." box="[1311,1362,875,896]" pageId="3" pageNumber="451" pagination="165 - 168" refId="ref6302" refString="Ito N. 1999. A new species of the genus Coleolissus from Laos (Coleoptera: Carabidae: Harpalini) // Ent. Rev. Japan. Vol. 54. No. 2. P. 165 - 168." type="journal article" year="1999">1999</bibRefCitation>
] as
<taxonomicName id="4C194D30F303FFFEE42B6DCE9CD7D3F7" authority="Ito, 1997" authorityName="Ito" authorityYear="1997" box="[812,1260,902,923]" class="Insecta" family="Carabidae" genus="Oxycentrus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="subarcuaticollis">
<emphasis id="B96DEAA1F303FFFEE42B6DCE9BA7D3F7" box="[812,924,902,923]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
(s. str.)
<emphasis id="B96DEAA1F303FFFEE4E96DCE9CB6D3F7" box="[1006,1165,902,923]" italics="true" pageId="3" pageNumber="451">subarcuaticollis</emphasis>
Ito, 1997
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE21B6DCE9C95D3DA" authority="Ito, 1999" authorityName="Ito" authorityYear="1999" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="iridipennis" subGenus="Tenuistilus">
<emphasis id="B96DEAA1F303FFFEE21B6DCE9B70D3DA" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
(
<emphasis id="B96DEAA1F303FFFEE4596DE99BF1D3DA" box="[862,970,929,950]" italics="true" pageId="3" pageNumber="451">Tenuistilus</emphasis>
)
<emphasis id="B96DEAA1F303FFFEE4DA6DE99C73D3DA" box="[989,1096,929,950]" italics="true" pageId="3" pageNumber="451">iridipennis</emphasis>
<bibRefCitation id="EF884B42F303FFFEE3566DE99C95D3DA" author="Ito N." box="[1105,1198,929,950]" pageId="3" pageNumber="451" pagination="165 - 168" refId="ref6302" refString="Ito N. 1999. A new species of the genus Coleolissus from Laos (Coleoptera: Carabidae: Harpalini) // Ent. Rev. Japan. Vol. 54. No. 2. P. 165 - 168." type="journal article" year="1999">Ito, 1999</bibRefCitation>
</taxonomicName>
without comparing them, and their generic affiliation remains unclear. They both markedly differ from
<taxonomicName id="4C194D30F303FFFEE3306D9E9CA5D380" authorityName="Kataev" authorityYear="2021" box="[1079,1182,982,1004]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3306D9E9CA5D380" box="[1079,1182,982,1004]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE3A26D9E9CEDD387" bold="true" box="[1189,1238,982,1003]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE3A26D9E9CEDD387" box="[1189,1238,982,1003]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
at least in body size larger (about 9.5–12.0 mm) and in having pronotum markedly depressed latero-basally, with more rounded sides and basal angles, and abdominal sternite VII of male with two pairs of marginal setae. The new species is included in the genus
<taxonomicName id="4C194D30F303FFFEE4966A139C39D41C" authorityName="Bates" authorityYear="1892" box="[913,1026,1115,1136]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4966A139C39D41C" box="[913,1026,1115,1136]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
on the basis of the following characters: frontal foveae without prolongation on clypeus, parascutellar striole long, interval 3 with a row of seven setigerous pores, and protibia with two preapical spines on outer margin. Each of these characters occurs in
<taxonomicName id="4C194D30F303FFFEE3D86A8E9D74D4B7" authorityName="Chaudoir" authorityYear="1854" box="[1247,1359,1222,1243]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3D86A8E9D74D4B7" box="[1247,1359,1222,1243]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
but not in such combination. As opposed to
<taxonomicName id="4C194D30F303FFFEE3C06AA99D03D49A" authorityName="Bates" authorityYear="1892" box="[1223,1336,1249,1270]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3C06AA99D03D49A" box="[1223,1336,1249,1270]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
, most species of
<taxonomicName id="4C194D30F303FFFEE4936AB49C3FD57D" authorityName="Chaudoir" authorityYear="1854" box="[916,1028,1276,1297]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4936AB49C3FD57D" box="[916,1028,1276,1297]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
have frontal foveae with more or less distinct prolongation on clypeus, parascutellar striole short or absent, interval 3 with less number of setigerous pores (in some species these pores absent), and protibia with more than two preapical spines on outer margin. The very moderately elongate mandibles of
<taxonomicName id="4C194D30F303FFFEE3476BC99C9DD5FA" authorityName="Kataev" authorityYear="2021" box="[1088,1190,1409,1430]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3476BC99C9DD5FA" box="[1088,1190,1409,1430]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE3AC6BC99CE6D5FA" bold="true" box="[1195,1245,1409,1430]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE3AC6BC99CE6D5FA" box="[1195,1245,1409,1430]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, shorter than in most
<taxonomicName id="4C194D30F303FFFEE4606BD39BE3D5DC" authorityName="Bates" authorityYear="1892" box="[871,984,1435,1456]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4606BD39BE3D5DC" box="[871,984,1435,1456]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE3136BD49CBFD5DD" authorityName="Chaudoir" authorityYear="1854" box="[1044,1156,1436,1457]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3136BD49CBFD5DD" box="[1044,1156,1436,1457]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
, and its comparatively long pronotum without latero-basal depressions, which are present probably in all congeners, suggest that this species is rather isolated within
<taxonomicName id="4C194D30F303FFFEE31B6BA39CB6D66C" authorityName="Bates" authorityYear="1892" box="[1052,1165,1515,1536]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE31B6BA39CB6D66C" box="[1052,1165,1515,1536]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
. The similarity to
<taxonomicName id="4C194D30F303FFFEE2406BA49B4FD670" authorityName="Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE2406BA49B4FD670" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
may be convergent (most species of
<taxonomicName id="4C194D30F303FFFEE3DB684E9D76D677" authorityName="Bates" authorityYear="1892" box="[1244,1357,1542,1563]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3DB684E9D76D677" box="[1244,1357,1542,1563]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE42B68699BA7D65A" authorityName="Chaudoir" authorityYear="1854" box="[812,924,1569,1590]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE42B68699BA7D65A" box="[812,924,1569,1590]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
have the plesiomorphic state of the abdominal sternite VII with two pairs of marginal setae as opposed to the apomorphic state in the new species with only one, inner, pair of setae), but it is not improbably that
<taxonomicName id="4C194D30F303FFFEE21668399D4CD6EA" authorityName="Kataev" authorityYear="2021" box="[1297,1399,1649,1670]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE21668399D4CD6EA" box="[1297,1399,1649,1670]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE42B68C39B65D6CC" bold="true" box="[812,862,1675,1696]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE42B68C39B65D6CC" box="[812,862,1675,1696]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is phylogenetically close to the branching node of two phyletic lineages sensu
<bibRefCitation id="EF884B42F303FFFEE31D68EE9CFBD6D0" author="Noonan G. R." box="[1050,1216,1702,1724]" pageId="3" pageNumber="451" pagination="1 - 33" refId="ref7311" refString="Noonan G. R. 1985 b. Reconstructed phylogeny and zoogeography of the genera and subgenera of the Selenophori group (Insecta: Coleoptera: Carabidae: Harpalini: Harpalina) // Milwaukee Public Museum, Contributions in Biology and Geology. No. 65. P. 1 - 33." type="book chapter" year="1985">Noonan [1985b]</bibRefCitation>
: the less modified lineage
<taxonomicName id="4C194D30F303FFFEE47C68899BCFD6BA" authorityName="MacLeay" authorityYear="1825" box="[891,1012,1729,1750]" class="Insecta" family="Carabidae" genus="Hyphaereon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE47C68899BCFD6BA" box="[891,1012,1729,1750]" italics="true" pageId="3" pageNumber="451">Hyphaereon</emphasis>
</taxonomicName>
and the more specialized lineage
<taxonomicName id="4C194D30F303FFFEE24068899B4FD69D" authorityName="Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE24068899B4FD69D" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
. The relationships and status of
<taxonomicName id="4C194D30F303FFFEE3D768949D01D69D" authorityName="Kataev" authorityYear="2021" box="[1232,1338,1756,1777]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3D768949D01D69D" box="[1232,1338,1756,1777]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE24268939D4DD69C" bold="true" box="[1349,1398,1755,1776]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE24268939D4DD69C" box="[1349,1398,1755,1776]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
need further study. Interestingly, the new species is somewhat similar in appearance to some species of
<taxonomicName id="4C194D30F303FFFEE22469599B8FD72D" authority="Murray, 1859" authorityName="Murray" authorityYear="1859" class="Insecta" family="Carabidae" genus="Siopelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE22469599D4DD74A" box="[1315,1398,1809,1830]" italics="true" pageId="3" pageNumber="451">Siopelus</emphasis>
Murray, 1859
</taxonomicName>
having smooth elytra, but well distinguished from them by having longer mandibles and glabrous genae and outer elytral intervals. The predominantly Ethiopian genus
<taxonomicName id="4C194D30F303FFFEE46969339BFAD7FC" authorityName="Murray" authorityYear="1859" box="[878,961,1915,1936]" class="Insecta" family="Carabidae" genus="Siopelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE46969339BFAD7FC" box="[878,961,1915,1936]" italics="true" pageId="3" pageNumber="451">Siopelus</emphasis>
</taxonomicName>
seems to occupy ancestral position in relation to the phyletic lineages
<taxonomicName id="4C194D30F303FFFEE35169DE9CF4D7C7" authorityName="MacLeay" authorityYear="1825" box="[1110,1231,1942,1963]" class="Insecta" family="Carabidae" genus="Hyphaereon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE35169DE9CF4D7C7" box="[1110,1231,1942,1963]" italics="true" pageId="3" pageNumber="451">Hyphaereon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE20169DE9C34D7AA" authority="sensu Noonan [1985 b]" authorityName="sensu Noonan" authorityYear="1985" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE20169DE9D4DD7C7" box="[1286,1398,1942,1963]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
sensu
<bibRefCitation id="EF884B42F303FFFEE46D69F99C34D7AA" author="Noonan G. R." box="[874,1039,1968,1990]" pageId="3" pageNumber="451" pagination="1 - 33" refId="ref7311" refString="Noonan G. R. 1985 b. Reconstructed phylogeny and zoogeography of the genera and subgenera of the Selenophori group (Insecta: Coleoptera: Carabidae: Harpalini: Harpalina) // Milwaukee Public Museum, Contributions in Biology and Geology. No. 65. P. 1 - 33." type="book chapter" year="1985">Noonan [1985b]</bibRefCitation>
</taxonomicName>
, and, in my opinion, does not differ morphologically from the American genus
<taxonomicName id="4C194D30F303FFFEE3F769839B8AD797" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" class="Insecta" family="Carabidae" genus="Selenophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3F769839D4DD78C" box="[1264,1398,1995,2016]" italics="true" pageId="3" pageNumber="451">Selenophorus</emphasis>
Dejean, 1829
</taxonomicName>
which is ancestral group for many American genera.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B96A2B63A840883FFEEDFB9836C6FB8B" box="[263,490,1031,1054]" italics="true" pageId="3" pageNumber="242">Gymnometriocnemus</emphasis>
(
<emphasis id="B96A2B63A840883FFE15FB993598FB8B" box="[511,692,1030,1054]" italics="true" pageId="3" pageNumber="242">Rhaphidocladius</emphasis>
)
<emphasis id="B96A2B63A840883FFF13FBB93685FBAB" box="[249,425,1062,1086]" italics="true" pageId="3" pageNumber="242">kamimegavirgus</emphasis>
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</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BA1F771A840883FFE45FBD9353FFBCB" blockId="3.[249,713,1030,1118]" box="[431,531,1094,1118]" pageId="3" pageNumber="242">
<figureCitation id="1325EBF4A840883FFE45FBD9353CFBCB" box="[431,528,1094,1118]" captionStart="Figs 2–5" captionStartId="2.[226,263,1702,1721]" captionTargetBox="[259,1268,344,1625]" captionTargetId="figure-12@2.[209,1373,314,1625]" captionTargetPageId="2" captionText="Figs 2–5. General view of hypopigium of Gymnometriocnemus (Rhaphidocladius) species: 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. Scale bar 50 µm (after Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9). Рис. 2–5. ОбЩий вид гипопигиЯ самцов Gymnometriocnemus (Rhaphidocladius): 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. МасШтабнаЯ линейка 50 мкм (по Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9)." figureDoi="http://doi.org/10.5281/zenodo.13178570" httpUri="https://zenodo.org/record/13178570/files/figure.png" pageId="3" pageNumber="242">Figs 4, 5</figureCitation>
.
</paragraph>
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<paragraph id="8BA1F771A840883FFF08FB1C3503FB75" blockId="3.[189,776,1154,1605]" pageId="3" pageNumber="242">
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MATERIAL.
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<collectingCountry id="F309B7E1A840883FFEB7FB1D36B6FB00" box="[349,410,1154,1173]" name="Russia" pageId="3" pageNumber="242">Russia</collectingCountry>
</emphasis>
:
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, village
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, left bank of
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, pine forest with admixture of deciduous species
<geoCoordinate id="EE2A91B6A840883FFED8FB2B368CFB52" box="[306,416,1204,1223]" degrees="55.46164" direction="north" orientation="latitude" pageId="3" pageNumber="242" precision="1" value="55.46164">55.46164°N</geoCoordinate>
,
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, 27.04–
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<collectingDate id="EFE42859A840883FFD98FB2B35E1FB52" box="[626,717,1204,1223]" pageId="3" pageNumber="242" value="2022-05-07">7.05.2022</collectingDate>
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,
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,
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(
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,
<specimenCount id="9D183CF8A840883FFE27FB523509FB75" box="[461,549,1229,1248]" count="7" pageId="3" pageNumber="242" type="female">7 females</specimenCount>
)
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.
</paragraph>
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<paragraph id="8BA1F771A840883FFF08FB79342BFA43" blockId="3.[189,776,1154,1605]" pageId="3" pageNumber="242">
DIAGNOSIS. Males of this species seem to be much darker in comparison with specimens of
<taxonomicName id="4C1E8CF2A840883FFDB0FA9E342CFA83" baseAuthorityName="Edwards" baseAuthorityYear="1929" box="[602,768,1281,1303]" class="Insecta" family="Chironomidae" genus="Gymnometriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="brumalis" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A840883FFDB0FA9E355EFA83" box="[602,626,1281,1302]" italics="true" pageId="3" pageNumber="242">G.</emphasis>
(
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)
<emphasis id="B96A2B63A840883FFD40FA9E342CFA83" box="[682,768,1281,1302]" italics="true" pageId="3" pageNumber="242">brumalis</emphasis>
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: thorax and abdominal tergites dark brown, almost black. Wing membrane has setae on the most apex only. The species differs from
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<emphasis id="B96A2B63A840883FFE8CFACB365BFAFC" box="[358,375,1364,1385]" italics="true" pageId="3" pageNumber="242">G</emphasis>
. (
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.)
<emphasis id="B96A2B63A840883FFE5BFACB350FFAFC" box="[433,547,1364,1385]" italics="true" pageId="3" pageNumber="242">tairaprimus</emphasis>
</taxonomicName>
by simple, not double, lower appendage of gonocoxite. The species differs from
<taxonomicName id="4C1E8CF2A840883FFD05FAF03612FA0A" baseAuthorityName="Edwards" baseAuthorityYear="1929" class="Insecta" family="Chironomidae" genus="Gymnometriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="brumalis" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A840883FFD05FAF0342BFA11" box="[751,775,1391,1412]" italics="true" pageId="3" pageNumber="242">G.</emphasis>
(
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)
<emphasis id="B96A2B63A840883FFF02FA153612FA0A" box="[232,318,1418,1439]" italics="true" pageId="3" pageNumber="242">brumalis</emphasis>
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by strongly sclerotized and well visible lamella of virga and rounded, not angulate lower appendage of gonocoxite (
<figureCitation id="1325EBF4A840883FFEDCFA5E36A1FA42" box="[310,397,1473,1495]" captionStart="Figs 2–5" captionStartId="2.[226,263,1702,1721]" captionTargetBox="[259,1268,344,1625]" captionTargetId="figure-12@2.[209,1373,314,1625]" captionTargetPageId="2" captionText="Figs 2–5. General view of hypopigium of Gymnometriocnemus (Rhaphidocladius) species: 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. Scale bar 50 µm (after Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9). Рис. 2–5. ОбЩий вид гипопигиЯ самцов Gymnometriocnemus (Rhaphidocladius): 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. МасШтабнаЯ линейка 50 мкм (по Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9)." figureDoi="http://doi.org/10.5281/zenodo.13178570" httpUri="https://zenodo.org/record/13178570/files/figure.png" pageId="3" pageNumber="242">Figs 4, 5</figureCitation>
) [
<bibRefCitation id="EF8F8A80A840883FFE48FA5E35D1FA43" author="Makarchenko E. A. &amp; Makarchenko M. A." box="[418,765,1473,1495]" pageId="3" pageNumber="242" pagination="333 - 340" refId="ref2550" refString="Makarchenko E. A., Makarchenko M. A. 2019. Review of the genus Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East // Eurasian entomological Journal. Vol. 18. No. 5. P. 333 - 340. doi: 10.15298 / euroasentj. 18.5.5." type="book chapter" year="2019">Makarchenko, Makarchenko, 2019</bibRefCitation>
].
</paragraph>
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<paragraph id="8BA1F771A840883FFF08FA4233B8FEF8" blockId="3.[189,776,1154,1605]" lastBlockId="3.[812,1398,315,365]" pageId="3" pageNumber="242">
DISTRIBUTION. The species was described from the territory of
<collectingCountry id="F309B7E1A840883FFED9FA663646F99B" box="[307,362,1529,1550]" name="Japan" pageId="3" pageNumber="242">Japan</collectingCountry>
(
<collectingRegion id="49DA3993A840883FFE91FA6736E6F998" box="[379,458,1528,1549]" country="Japan" name="Nagano" pageId="3" pageNumber="242">Nagano</collectingRegion>
, Honshu Island) [
<bibRefCitation id="EF8F8A80A840883FFD94FA67361EF9BC" author="Sasa M. &amp; Hirabayashi K." pageId="3" pageNumber="242" pagination="361 - 393" refId="ref2608" refString="Sasa M., Hirabayashi K. 1993. Studies on the additional chironomids (Diptera, Chironomidae) collected at Kamikochi and Asama-Onsen, Nagano, Japan // Japanese Journal of sanitary Zoology. Vol. 44. P. 361 - 393." type="journal article" year="1993">Sasa, Hirabayashi, 1993</bibRefCitation>
]. Registered in
<collectingCountry id="F309B7E1A840883FFE26F98B3536F9BC" box="[460,538,1556,1577]" name="Norway" pageId="3" pageNumber="242">Norway</collectingCountry>
[
<bibRefCitation id="EF8F8A80A840883FFDCDF98B35CEF9BF" author="Stur E. &amp; Ekrem T." box="[551,738,1556,1578]" pageId="3" pageNumber="242" pagination="127 - 142" refId="ref2653" refString="Stur E., Ekrem T. 2015. A review of Norwegian Gymnometriocnemus (Diptera, Chironomidae) including the description of two new species and a new name for Gymnometriocnemus volitans (Goetghebuer) sensu Brundin // ZooKeys. Vol. 508. P. 127 - 142. doi: 10.3897 / zookeys. 508.9874" type="book chapter" year="2015">Stur, Ekrem, 2015</bibRefCitation>
]. In
<collectingCountry id="F309B7E1A840883FFF57F9B037D1F9D0" box="[189,253,1583,1605]" name="Russia" pageId="3" pageNumber="242">Russia</collectingCountry>
it was found in the Far East (
<collectingRegion id="49DA3993A840883FFDCEF9B035E1F9D0" box="[548,717,1583,1605]" country="Russia" name="Magadan" pageId="3" pageNumber="242">Magadan Region</collectingRegion>
) only [
<bibRefCitation id="EF8F8A80A840883FFCD8FEA433A2FEC4" author="Makarchenko E. A. &amp; Makarchenko M. A." box="[818,1166,315,337]" pageId="3" pageNumber="242" pagination="333 - 340" refId="ref2550" refString="Makarchenko E. A., Makarchenko M. A. 2019. Review of the genus Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East // Eurasian entomological Journal. Vol. 18. No. 5. P. 333 - 340. doi: 10.15298 / euroasentj. 18.5.5." type="book chapter" year="2019">Makarchenko, Makarchenko, 2019</bibRefCitation>
]. Our registration is the first for the European part of
<collectingCountry id="F309B7E1A840883FFBA7FEC833BDFEF8" box="[1101,1169,343,365]" name="Russia" pageId="3" pageNumber="242">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8BA1F771A840883FFCB8FE1834A9FE2D" blockId="3.[812,1398,391,440]" pageId="3" pageNumber="242">
<emphasis id="B96A2B63A840883FFCB8FE18330BFE08" bold="true" box="[850,1063,391,413]" pageId="3" pageNumber="242">Competing interests.</emphasis>
The authors declare no competing interests.
</paragraph>
</subSubSection>
<subSubSection id="C304A4FAA840883FFCB8FE4E321AFD97" pageId="3" pageNumber="242" type="description">
<paragraph id="8BA1F771A840883FFCB8FE4E321AFD97" blockId="3.[812,1398,465,514]" pageId="3" pageNumber="242">
<emphasis id="B96A2B63A840883FFCB8FE4E3335FE72" bold="true" box="[850,1049,465,487]" pageId="3" pageNumber="242">Acknowledgements</emphasis>
. The investigation was supported by the Russian Science Foundation No 22-14-00363.
</paragraph>
</subSubSection>
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<mods:namePart id="F8EAB52A69C03310906DE1087D2D0BB9">Krivosheina, M. G.</mods:namePart>
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<mods:name id="C5FCFCC8AFF40A021FECC2607784FD35" type="personal">
<mods:role id="F7D8027E4F4BE58EE6733FE8784014B2">
<mods:roleTerm id="3ECC5C563A3F54CBDEEB9D9BD24AA2D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="041FA41A11A30D72434747CFFFA0EAC5">Vinogradov, D. D.</mods:namePart>
</mods:name>
<mods:name id="10AFE2C27D0714F3FC56F597421A033E" type="personal">
<mods:role id="867AA09B8A870F2A11D96838325FB727">
<mods:roleTerm id="5CE1A2A4E7194A2A35D9ACB0BE53BA36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7B640BB02D0D3DC325A9BA13E6D5059">Tsurikov, S. M.</mods:namePart>
</mods:name>
<mods:name id="A1604B897EA0036AF500D32FF7C93756" type="personal">
<mods:role id="A8BDAC5B939665B130358D8D96E83B89">
<mods:roleTerm id="A372444C7D9DE9300DD88650ED7561D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DDE0137456357B383B9D97C98379FA3B">Zuev, A. G.</mods:namePart>
</mods:name>
<mods:name id="51F35BF035BE0549AD95D8F95AD43F5A" type="personal">
<mods:role id="966B480DE438DEDFCF90CBDB99EA614D">
<mods:roleTerm id="186D95E2DC9C552E82409B24B0E1FA5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9975EE16E82BDF772D677DA8F2CEF64F">Sotnikov, I. V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5BEAD6B7215EF3FE646515048AB9E5B5">text</mods:typeOfResource>
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<mods:titleInfo id="1A8E765CACB8E9000BC50AD7CEE459B6">
<mods:title id="BC42D8CD741E138A6FE2A63B7E5BB837">Russian Entomological Journal</mods:title>
</mods:titleInfo>
<mods:part id="E6FAFCB9EC5DE7C198139E6CEE37883D">
<mods:date id="4268053A7DEE3297B033E55D9A53885A">2023</mods:date>
<mods:detail id="B86A8DC5CAC38EC5F6ABE364F16BA48E" type="pubDate">
<mods:number id="EE5CBC052929365C4F7CA0AAE2B703E5">2023-06-30</mods:number>
</mods:detail>
<mods:detail id="489ECA15345BF1C96E503B1E464AFC07" type="volume">
<mods:number id="FCF7A74886A2A7789335BFC350582773">32</mods:number>
</mods:detail>
<mods:detail id="F88428DC8B227C9CD9F7CD5E7D86DC9F" type="issue">
<mods:number id="6672475537BB4CD068575DD12CE42D3C">2</mods:number>
</mods:detail>
<mods:extent id="74A6105A699D7B8A5B85B2A56A336647" unit="page">
<mods:start id="8C124843916EC7A23188A89A4198C2BD">239</mods:start>
<mods:end id="925E31C75631D65C8FA2A53B53E6A40E">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="F31F956DD83BA0B4B2C1E8C6490E2CA6">
<mods:url id="1ECCEDAD47EBCAEC8B864DF22C758A27">http://dx.doi.org/10.15298/rusentj.32.2.15</mods:url>
</mods:location>
<mods:classification id="E61B8FEA95FD338B8389685BB52F962F">journal article</mods:classification>
<mods:identifier id="988DC5B61DA0B44D2DCD819CFA61B5C0" type="DOI">10.15298/rusentj.32.2.15</mods:identifier>
<mods:identifier id="153D0CF84AA424C1B0A02C691A5008AE" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="0D4120F568782B878B4B042CE8DC0BA3" type="Zenodo-Dep">13178566</mods:identifier>
</mods:mods>
<treatment id="03B74667A841883FFCA9F8D43510FC4D" LSID="urn:lsid:plazi:treatment:03B74667A841883FFCA9F8D43510FC4D" httpUri="http://treatment.plazi.org/id/03B74667A841883FFCA9F8D43510FC4D" lastPageId="3" lastPageNumber="242" pageId="2" pageNumber="241">
<subSubSection id="C304A4FAA841883EFCA9F8D43380F815" pageId="2" pageNumber="241" type="nomenclature">
<paragraph id="8BA1F771A841883EFCA9F8D43380F815" blockId="2.[835,1375,1867,1950]" pageId="2" pageNumber="241">
<heading id="D0E9401DA841883EFCA9F8D43380F815" centered="true" fontSize="10" level="2" pageId="2" pageNumber="241" reason="8">
<taxonomicName id="4C1E8CF2A841883EFCA9F8D43380F815" ID-CoL="8C8S4" authority="(Edwards, 1929)" baseAuthorityName="Edwards" baseAuthorityYear="1929" class="Insecta" family="Chironomidae" genus="Gymnometriocnemus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="241" phylum="Arthropoda" rank="species" species="brumalis" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A841883EFCA9F8D4330AF8F7" box="[835,1062,1867,1890]" italics="true" pageId="2" pageNumber="241">Gymnometriocnemus</emphasis>
(
<emphasis id="B96A2B63A841883EFBD3F8D433C2F8F6" box="[1081,1262,1867,1891]" italics="true" pageId="2" pageNumber="241">Rhaphidocladius</emphasis>
)
<emphasis id="B96A2B63A841883EFAEAF8D43273F8F6" box="[1280,1375,1867,1891]" italics="true" pageId="2" pageNumber="241">brumalis</emphasis>
(
<bibRefCitation id="EF8F8A80A841883EFC15F8F73388F815" author="Edwards F. W." box="[1023,1188,1896,1920]" pageId="2" pageNumber="241" pagination="279 - 430" refId="ref2345" refString="Edwards F. W. 1929. British non-biting midges (Diptera, Chironomidae) // Transactions of the Entomological Society of London. Vol. 77. P. 279 - 430. doi: 10.1111 / j. 1365 - 2311.1929. tb 00692. x" type="book chapter" year="1929">Edwards, 1929</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304A4FAA841883EFBF5F81A33AFF80B" box="[1055,1155,1925,1950]" pageId="2" pageNumber="241" type="description">
<paragraph id="8BA1F771A841883EFBF5F81A33AFF80B" blockId="2.[835,1375,1867,1950]" box="[1055,1155,1925,1950]" pageId="2" pageNumber="241">
<figureCitation id="1325EBF4A841883EFBF5F81A3353F80B" box="[1055,1151,1925,1950]" captionStart="Figs 2–5" captionStartId="2.[226,263,1702,1721]" captionTargetBox="[259,1268,344,1625]" captionTargetId="figure-12@2.[209,1373,314,1625]" captionTargetPageId="2" captionText="Figs 2–5. General view of hypopigium of Gymnometriocnemus (Rhaphidocladius) species: 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. Scale bar 50 µm (after Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9). Рис. 2–5. ОбЩий вид гипопигиЯ самцов Gymnometriocnemus (Rhaphidocladius): 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. МасШтабнаЯ линейка 50 мкм (по Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9)." figureDoi="http://doi.org/10.5281/zenodo.13178570" httpUri="https://zenodo.org/record/13178570/files/figure.png" pageId="2" pageNumber="241">Figs 2, 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304A4FAA841883EFCB8F8293366F785" pageId="2" pageNumber="241" type="materials_examined">
<paragraph id="8BA1F771A841883EFCB8F8293366F785" blockId="2.[812,1399,1973,2065]" pageId="2" pageNumber="241">
<materialsCitation id="3B76FD2CA841883EFCB8F829336AF785" collectingDate="2022-05-07" collectorName="S. M. Tsurikov &amp; A. G. Zuev" country="Russia" latitude="55.46164" location="Zhiletovka river" longLatPrecision="1" longitude="37.18284" municipality="Pervomayskoe" pageId="2" pageNumber="241" specimenCount="19" specimenCount-male="19" stateProvince="Moscow">
MATERIAL.
<emphasis id="B96A2B63A841883EFC27F82A3326F85D" bold="true" box="[973,1034,1973,1992]" pageId="2" pageNumber="241">
<collectingCountry id="F309B7E1A841883EFC27F82A3326F85D" box="[973,1034,1973,1992]" name="Russia" pageId="2" pageNumber="241">Russia</collectingCountry>
</emphasis>
:
<collectingRegion id="49DA3993A841883EFBFCF829338BF85D" box="[1046,1191,1973,1993]" country="Russia" name="Moscow" pageId="2" pageNumber="241">Moscow Region</collectingRegion>
, village
<collectingMunicipality id="6BC56D0BA841883EFB18F82A325FF85D" box="[1266,1395,1973,1992]" pageId="2" pageNumber="241">Pervomayskoe</collectingMunicipality>
, left bank of
<location id="8EC1A1AAA841883EFC72F852330BF875" LSID="urn:lsid:plazi:treatment:03B74667A841883FFCA9F8D43510FC4D:8EC1A1AAA841883EFC72F852330BF875" box="[920,1063,1997,2016]" country="Russia" latitude="55.46164" longLatPrecision="1" longitude="37.18284" municipality="Pervomayskoe" name="Zhiletovka river" pageId="2" pageNumber="241" stateProvince="Moscow">Zhiletovka river</location>
, pine forest with admixture of deciduous species
<geoCoordinate id="EE2A91B6A841883EFC48F87A3323F86D" box="[930,1039,2021,2040]" degrees="55.46164" direction="north" orientation="latitude" pageId="2" pageNumber="241" precision="1" value="55.46164">55.46164°N</geoCoordinate>
,
<geoCoordinate id="EE2A91B6A841883EFBF5F87933BAF86C" box="[1055,1174,2022,2041]" degrees="037.18284" direction="east" orientation="longitude" pageId="2" pageNumber="241" precision="1" value="37.18284">037.18284°E</geoCoordinate>
, 27.04–
<date id="FFA0D1B1A841883EFB08F87A3211F86D" box="[1250,1341,2021,2040]" pageId="2" pageNumber="241" value="2022-05-07">
<collectingDate id="EFE42859A841883EFB08F87A3211F86D" box="[1250,1341,2021,2040]" pageId="2" pageNumber="241" value="2022-05-07">7.05.2022</collectingDate>
</date>
,
<collectorName id="26EB92A7A841883EFAA1F8793450F785" pageId="2" pageNumber="241">S.M. Tsurikov</collectorName>
,
<collectorName id="26EB92A7A841883EFC69F86134CCF784" box="[899,992,2046,2065]" pageId="2" pageNumber="241">A.G. Zuev</collectorName>
(
<specimenCount id="9D183CF8A841883EFC07F861336CF785" box="[1005,1088,2045,2065]" count="19" pageId="2" pageNumber="241" type="male">19 males</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304A4FAA840883FFF08FEA337D1FDCB" pageId="3" pageNumber="242" type="diagnosis">
<paragraph id="8BA1F771A840883FFF08FEA337D1FDCB" blockId="3.[189,775,315,984]" pageId="3" pageNumber="242">
DIAGNOSIS. The species, like other members of the subgenus
<taxonomicName id="4C1E8CF2A840883FFECBFEC936E8FEFE" box="[289,452,342,363]" pageId="3" pageNumber="242" rank="subGenus" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A840883FFECBFEC936E8FEFE" box="[289,452,342,363]" italics="true" pageId="3" pageNumber="242">Rhaphidocladius</emphasis>
</taxonomicName>
, is characterized by a strongly developed virga beginning in tergite 8 and the absence of a process on tergite 9, which has only a medial fold. The species differs from
<taxonomicName id="4C1E8CF2A840883FFE6EFE37356DFE28" authorityName="Sasa et Okazawa" authorityYear="1994" box="[388,577,423,445]" class="Insecta" family="Chironomidae" genus="Gymnometriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="tairaprimus" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A840883FFE6EFE3736B0FE28" box="[388,412,424,445]" italics="true" pageId="3" pageNumber="242">G.</emphasis>
(
<emphasis id="B96A2B63A840883FFE41FE3736ECFE28" box="[427,448,424,445]" italics="true" pageId="3" pageNumber="242">R.</emphasis>
)
<emphasis id="B96A2B63A840883FFE25FE37356DFE28" box="[463,577,424,445]" italics="true" pageId="3" pageNumber="242">tairaprimus</emphasis>
</taxonomicName>
by simple, not double, lower appendage of gonocoxite. The species differs from
<taxonomicName id="4C1E8CF2A840883FFF18FE4236F7FE67" authorityName="Sasa et Hirabayashi" authorityYear="1993" box="[242,475,477,499]" class="Insecta" family="Chironomidae" genus="Gymnometriocnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="kamimegavirgus" subGenus="Rhaphidocladius">
<emphasis id="B96A2B63A840883FFF18FE423626FE67" box="[242,266,477,498]" italics="true" pageId="3" pageNumber="242">G.</emphasis>
(
<emphasis id="B96A2B63A840883FFEF2FE413602FE66" box="[280,302,478,499]" italics="true" pageId="3" pageNumber="242">R.</emphasis>
)
<emphasis id="B96A2B63A840883FFED6FE4236F7FE67" box="[316,475,477,498]" italics="true" pageId="3" pageNumber="242">kamimegavirgus</emphasis>
</taxonomicName>
by wing membrane which has setae on at least half of wing, weakly sclerotized and often hardly visible lamella of virga and angulate lower appendage of gonocoxite (
<figureCitation id="1325EBF4A840883FFE90FDB136FEFDD6" box="[378,466,558,580]" captionStart="Figs 2–5" captionStartId="2.[226,263,1702,1721]" captionTargetBox="[259,1268,344,1625]" captionTargetId="figure-12@2.[209,1373,314,1625]" captionTargetPageId="2" captionText="Figs 2–5. General view of hypopigium of Gymnometriocnemus (Rhaphidocladius) species: 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. Scale bar 50 µm (after Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9). Рис. 2–5. ОбЩий вид гипопигиЯ самцов Gymnometriocnemus (Rhaphidocladius): 2, 3 — G. (R.) brumalis; 4, 5 — G. (R.) kamumegavirgus. МасШтабнаЯ линейка 50 мкм (по Makarchenko &amp; Makarchenko, 2019, Figs 4, 5, 8, 9)." figureDoi="http://doi.org/10.5281/zenodo.13178570" httpUri="https://zenodo.org/record/13178570/files/figure.png" pageId="3" pageNumber="242">Figs 2, 3</figureCitation>
) [
<bibRefCitation id="EF8F8A80A840883FFE0DFDB137D8FDCB" author="Makarchenko E. A. &amp; Makarchenko M. A." pageId="3" pageNumber="242" pagination="333 - 340" refId="ref2550" refString="Makarchenko E. A., Makarchenko M. A. 2019. Review of the genus Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East // Eurasian entomological Journal. Vol. 18. No. 5. P. 333 - 340. doi: 10.15298 / euroasentj. 18.5.5." type="book chapter" year="2019">Makarchenko, Makarchenko, 2019</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C304A4FAA840883FFF08FDFB3510FC4D" pageId="3" pageNumber="242" type="distribution">
<paragraph id="8BA1F771A840883FFF08FDFB3510FC4D" blockId="3.[189,775,315,984]" pageId="3" pageNumber="242">
DISTRIBUTION. The species was described for
<collectingCountry id="F309B7E1A840883FFD3BFDFB362CFD00" name="United Kingdom" pageId="3" pageNumber="242">Great Britain</collectingCountry>
[
<bibRefCitation id="EF8F8A80A840883FFEE0FDE0368DFD01" author="Edwards F. W." box="[266,417,639,661]" pageId="3" pageNumber="242" pagination="279 - 430" refId="ref2345" refString="Edwards F. W. 1929. British non-biting midges (Diptera, Chironomidae) // Transactions of the Entomological Society of London. Vol. 77. P. 279 - 430. doi: 10.1111 / j. 1365 - 2311.1929. tb 00692. x" type="book chapter" year="1929">Edwards, 1929</bibRefCitation>
]. Registered in
<collectingCountry id="F309B7E1A840883FFDD9FDE035ADFD01" box="[563,641,639,660]" name="Norway" pageId="3" pageNumber="242">Norway</collectingCountry>
[
<bibRefCitation id="EF8F8A80A840883FFD61FDE037DEFD25" author="Stur E. &amp; Ekrem T." pageId="3" pageNumber="242" pagination="127 - 142" refId="ref2653" refString="Stur E., Ekrem T. 2015. A review of Norwegian Gymnometriocnemus (Diptera, Chironomidae) including the description of two new species and a new name for Gymnometriocnemus volitans (Goetghebuer) sensu Brundin // ZooKeys. Vol. 508. P. 127 - 142. doi: 10.3897 / zookeys. 508.9874" type="book chapter" year="2015">Stur, Ekrem, 2015</bibRefCitation>
]. According to the data in Fauna Europaea the species is also known from
<collectingCountry id="F309B7E1A840883FFE6FFD2A36F8FD5E" box="[389,468,693,715]" name="Sweden" pageId="3" pageNumber="242">Sweden</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFE35FD2A3507FD5E" box="[479,555,693,715]" name="Finland" pageId="3" pageNumber="242">Finland</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFDD2FD2A3553FD5E" box="[568,639,693,715]" name="Ireland" pageId="3" pageNumber="242">Ireland</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFD60FD2A35E2FD5F" box="[650,718,693,714]" name="France" pageId="3" pageNumber="242">France</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFD30FD2A37D9FD70" name="Germany" pageId="3" pageNumber="242">Germany</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFEE9FD4F3676FD70" box="[259,346,720,741]" name="Hungary" pageId="3" pageNumber="242">Hungary</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFE83FD5036F0FD70" box="[361,476,719,741]" name="Luxembourg" pageId="3" pageNumber="242">Luxemburg</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFE01FD50351DFD70" box="[491,561,719,741]" name="Poland" pageId="3" pageNumber="242">Poland</collectingCountry>
,
<collectingCountry id="F309B7E1A840883FFDD5FD503599FD70" box="[575,693,719,741]" name="Switzerland" pageId="3" pageNumber="242">Switzerland</collectingCountry>
, Nederlands and
<collectingCountry id="F309B7E1A840883FFECDFD753672FC95" box="[295,350,746,768]" name="Spain" pageId="3" pageNumber="242">Spain</collectingCountry>
[
<bibRefCitation id="EF8F8A80A840883FFE84FD743565FC95" author="De Jong Y. &amp; Verbeek M. &amp; Michelsen V." box="[366,585,746,768]" pageId="3" pageNumber="242" refId="ref2293" refString="De Jong Y., Verbeek M., Michelsen V. et al. 2014. Fauna Europaea - all European animal species on the web // Biodiversity Data Journal. Vol. 2. No. e 4034. doi: 10.3897 / BDJ. 2. e 4034." type="journal volume" year="2014">De Jong et al., 2014</bibRefCitation>
]. According to the World database of
<taxonomicName id="4C1E8CF2A840883FFE9FFC9A3691FC8E" box="[373,445,773,795]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="242" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
the species is also widely distributed in
<collectingCountry id="F309B7E1A840883FFEE3FCBF367DFCA3" box="[265,337,800,822]" name="Canada" pageId="3" pageNumber="242">Canada</collectingCountry>
and
<collectingCountry id="F309B7E1A840883FFE6FFCBF3699FCA0" box="[389,437,800,821]" name="United States of America" pageId="3" pageNumber="242">USA</collectingCountry>
[
<bibRefCitation id="EF8F8A80A840883FFE28FCBF35B3FCA0" author="Evenhuis N. L. &amp; Pape T." box="[450,671,800,822]" pageId="3" pageNumber="242" refId="ref2393" refString="Evenhuis N. L., Pape T. (editors). 2023. Systema Dipterorum, Version 4.1, last updated 15 March 2023. http: // diptera. org /, accessed on 3 May 2023." type="url" year="2023">Evenhuis, Pape, 2023</bibRefCitation>
]. The species is not rare but for a long time there was no information about its distribution in
<collectingCountry id="F309B7E1A840883FFE47FCC936DDFCF9" box="[429,497,854,876]" name="Russia" pageId="3" pageNumber="242">Russia</collectingCountry>
. It was recently registerd in
<collectingRegion id="49DA3993A840883FFF57FCEE3675FC12" box="[189,345,881,903]" country="Russia" name="Primorskiy" pageId="3" pageNumber="242">Primorsky Krai</collectingRegion>
(Lazovsky District),
<collectingRegion id="49DA3993A840883FFDD1FCEE3551FC12" box="[571,637,881,903]" country="Russia" name="Yevrey" pageId="3" pageNumber="242">Jewish</collectingRegion>
Autonomous Region and
<collectingRegion id="49DA3993A840883FFED8FC1336FBFC37" box="[306,471,908,930]" country="Russia" name="Khabarovsk" pageId="3" pageNumber="242">Khabarovsk Krai</collectingRegion>
(
<collectingRegion id="49DA3993A840883FFE0EFC133587FC37" box="[484,683,908,930]" country="Russia" name="Khabarovsk" pageId="3" pageNumber="242">Khabarovsk District</collectingRegion>
) [
<bibRefCitation id="EF8F8A80A840883FFD57FC1336F5FC29" author="Makarchenko E. A. &amp; Makarchenko M. A." pageId="3" pageNumber="242" pagination="333 - 340" refId="ref2550" refString="Makarchenko E. A., Makarchenko M. A. 2019. Review of the genus Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East // Eurasian entomological Journal. Vol. 18. No. 5. P. 333 - 340. doi: 10.15298 / euroasentj. 18.5.5." type="book chapter" year="2019">Makarchenko, Makarchenko, 2019</bibRefCitation>
]. We prove the distribution of the species in European Part of
<collectingCountry id="F309B7E1A840883FFE1FFC5D3515FC4D" box="[501,569,962,984]" name="Russia" pageId="3" pageNumber="242">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

289
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<mods:title id="076AB3AE026421038EEF593D94452EEF">Notes on the genera Aristopus, Cosmodiscus and Metabacetus (Coleoptera: Carabidae: Pterostichini: Abacetina), with description of three new species from Vietnam</mods:title>
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<subSubSection id="C371651B6538FF8F12F2FECD0D0FFE6B" pageId="1" pageNumber="414" type="nomenclature">
<paragraph id="8BD436906538FF8F12F2FECD0CA3FE4B" blockId="1.[812,1399,368,1140]" box="[968,1243,368,393]" pageId="1" pageNumber="414">
<heading id="D09C81FC6538FF8F12F2FECD0CA3FE4B" box="[968,1243,368,393]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="414" reason="8">
<taxonomicName id="4C6B4D136538FF8F12F2FECD0CA3FE4B" ID-CoL="8KXQT" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[968,1243,368,393]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA826538FF8F12F2FECD0C2BFE4A" bold="true" box="[968,1107,368,392]" italics="true" pageId="1" pageNumber="414">Metabacetus</emphasis>
<bibRefCitation id="EFFA4B616538FF8F1561FECC0CA3FE4B" author="Bates H. W." box="[1115,1243,369,393]" pageId="1" pageNumber="414" pagination="267 - 428" refId="ref16950" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regione vicine. XLIV. List of the Carabidae // Ann. Mus. Stor. Nat. Genova. Ser. 2. Vol. 12. No. 32. P. 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD436906538FF8F1268FE2B0D0FFE6B" blockId="1.[812,1399,368,1140]" box="[850,1399,406,425]" pageId="1" pageNumber="414">
<bibRefCitation id="EFFA4B616538FF8F1268FE2B0B92FE6B" author="Bates H. W." box="[850,1002,406,425]" pageId="1" pageNumber="414" pagination="267 - 428" refId="ref16950" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regione vicine. XLIV. List of the Carabidae // Ann. Mus. Stor. Nat. Genova. Ser. 2. Vol. 12. No. 32. P. 267 - 428." type="journal article" year="1892">Bates, 1892: 364</bibRefCitation>
;
<bibRefCitation id="EFFA4B616538FF8F12CEFE2B0CDFFE6B" author="Straneo S. L." box="[1012,1191,406,425]" pageId="1" pageNumber="414" pagination="151 - 158" refId="ref17503" refString="Straneo S. L. 1938 a. Le genre Metabacetus Bates [Coleoptera, Harpalidae] // Rev. Franc. Ent. Vol. 5. P. 151 - 158." type="journal article" year="1938">Straneo, 1938a: 151</bibRefCitation>
;
<bibRefCitation id="EFFA4B616538FF8F158BFE2B0D0BFE6B" author="Will K. W. &amp; Park J. K." box="[1201,1395,406,425]" pageId="1" pageNumber="414" pagination="189 - 196" refId="ref18022" refString="Will K. W., Park J. K. 2008. A new species of the oriental abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus // Coleopt. Bull. Vol. 62. P. 189 - 196. https: // doi. org / 10.1649 / 980.1." type="journal article" year="2008">Will, Park, 2008: 191</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C371651B6538FF8F1268FE130B19FE1B" pageId="1" pageNumber="414" type="materials_examined">
<paragraph id="8BD436906538FF8F1268FE130B19FE1B" blockId="1.[812,1399,368,1140]" pageId="1" pageNumber="414">
<typeStatus id="54D088326538FF8F1268FE130B06FE03" box="[850,894,430,449]" pageId="1" pageNumber="414">Type</typeStatus>
species:
<taxonomicName id="4C6B4D136538FF8F12F0FE130D62FE03" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[970,1306,430,449]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="immarginatus">
<emphasis id="B91FEA826538FF8F12F0FE130CCAFE02" box="[970,1202,430,449]" italics="true" pageId="1" pageNumber="414">Metabacetus immarginatus</emphasis>
<bibRefCitation id="EFFA4B616538FF8F158CFE130D62FE03" author="Bates H. W." box="[1206,1306,430,449]" pageId="1" pageNumber="414" pagination="267 - 428" refId="ref16950" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regione vicine. XLIV. List of the Carabidae // Ann. Mus. Stor. Nat. Genova. Ser. 2. Vol. 12. No. 32. P. 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
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<subSubSection id="C371651B6538FF8F1268FE630BE9FD6C" pageId="1" pageNumber="414" type="diagnosis">
<paragraph id="8BD436906538FF8F1268FE630BE9FD6C" blockId="1.[812,1399,368,1140]" pageId="1" pageNumber="414">
DIAGNOSIS. Macropterous species of small size (BL
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), uniform black, with antennal pedicel centrically inserted in scape, antennomeres 4–11 pubescent, pedicel glabrous but ventral seta, mentum pits well-developed, body integuments impilose, metatibia externally glabrous, pronotum impunctate, elytral interval 3 without discal setigerous pores, except for vestigial setigerous pore d2 (median) that lacks seta.
</paragraph>
</subSubSection>
<subSubSection id="C371651B6538FF8D1268FD0E0A5CF9C3" lastPageId="3" lastPageNumber="416" pageId="1" pageNumber="414" type="description">
<paragraph id="8BD436906538FF8F1268FD0E0D7CFC7B" blockId="1.[812,1399,368,1140]" pageId="1" pageNumber="414">
REDESCRIPTION. With characters of other
<taxonomicName id="4C6B4D136538FF8F142AFD0E0D0CFD0B" authorityName="Chaudoir" authorityYear="1873" box="[1296,1396,691,713]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="subTribe" subTribe="Abacetina">Abacetina</taxonomicName>
: frontal impressions sulciform, divergent towards anterior supra-ocular seta; pronotum with one, sulciform, basal fovea on each side; elytral parascutellar striole ‘missing’ (
<emphasis id="B91FEA826538FF8F1439FCB90D58FCDB" box="[1283,1312,772,793]" italics="true" pageId="1" pageNumber="414">i.e.</emphasis>
, directly extended into stria 1 that lost its primary angular base); mesoventrite with medial groove to receive subcarinate posterior inclination of prosternal process; abdominal sternites not basally sulcate; metacoxa bisetose (inner seta missing), metatrochanter with seta; supra-ocular groove often directly extended into orbital groove (that between gena and neck).
</paragraph>
<paragraph id="8BD436906538FF8F1268FC030BB7FBFC" blockId="1.[812,1399,368,1140]" pageId="1" pageNumber="414">
Body (
<figureCitation id="13502A156538FF8F12AEFC030B93FC11" box="[916,1003,958,980]" captionStart="Figs 1–3" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[193,1383,1172,1928]" captionTargetId="figure-522@1.[183,1393,1155,1936]" captionTargetPageId="1" captionText="Figs 1–3. Dorsal habitus of Metabacetus spp.: 1 — M. hermanni, ♂ from Xuan Son NP; 2 — M. laotinus, ♀ from Cao Bang Province; 3 — M. perakianus, ♀ from Song Thanh NP. Рис. 1–3. Габитус дорЗально Metabacetus spp.: 1 — M. hermanni, ♂ иЗ Национального парка Суансон; 2 — M. laotinus, ♀ иЗ провинции Каобанг; 3 — M. perakianus, ♀ иЗ Национального парка Сонгтхань." figureDoi="http://doi.org/10.5281/zenodo.13178242" httpUri="https://zenodo.org/record/13178242/files/figure.png" pageId="1" pageNumber="414">Figs 1–3</figureCitation>
) convex, uniform black, dorsum mostly iridescent and slightly sericeous due to dense and very transverse microsculpture, body appendages and extreme lateral margins of both pronotum and elytra red. Body setation nearly complete.
</paragraph>
<paragraph id="8BD436906538FF8F1268FBFE0D0EFBB6" blockId="1.[812,1399,368,1140]" pageId="1" pageNumber="414">Head rather small, without neck constriction; eyes large, almost hemispherical, nearly reaching maxillary grooves be-</paragraph>
<paragraph id="8BD436906538FF8F15AEF8C10CDBF854" blockId="1.[1172,1187,1916,1942]" box="[1172,1187,1916,1942]" pageId="1" pageNumber="414">
<emphasis id="B91FEA826538FF8F15AEF8C10CDBF854" bold="true" box="[1172,1187,1916,1942]" pageId="1" pageNumber="414">3</emphasis>
</paragraph>
<caption id="DF1466186538FF8F11D8F80C0BE4F7D3" ID-DOI="http://doi.org/10.5281/zenodo.13178242" ID-Zenodo-Dep="13178242" httpUri="https://zenodo.org/record/13178242/files/figure.png" pageId="1" pageNumber="414" startId="1.[226,263,1969,1988]" targetBox="[193,1383,1172,1928]" targetPageId="1" targetType="figure">
<paragraph id="8BD436906538FF8F11D8F80C0A30F81F" blockId="1.[189,1399,1967,2065]" pageId="1" pageNumber="414">
Figs 1–3. Dorsal habitus of
<taxonomicName id="4C6B4D136538FF8F10E9F80C0A0CF807" box="[467,628,1969,1989]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA826538FF8F10E9F80C0A39F806" box="[467,577,1969,1988]" italics="true" pageId="1" pageNumber="414">Metabacetus</emphasis>
spp.
</taxonomicName>
: 1 —
<taxonomicName id="4C6B4D136538FF8F1392F80F0B6EF806" authorityName="Will et Park" authorityYear="2008" box="[680,790,1969,1988]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA826538FF8F1392F80F0B6EF806" box="[680,790,1969,1988]" italics="true" pageId="1" pageNumber="414">M. hermanni</emphasis>
</taxonomicName>
, ♂ from Xuan Son NP; 2 —
<taxonomicName id="4C6B4D136538FF8F1523F80F0C03F806" authorityName="Straneo" authorityYear="1938" box="[1049,1147,1969,1988]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA826538FF8F1523F80F0C03F806" box="[1049,1147,1969,1988]" italics="true" pageId="1" pageNumber="414">M. laotinus</emphasis>
</taxonomicName>
, ♀ from Cao Bang Province; 3 —
<taxonomicName id="4C6B4D136538FF8F11D0F877091FF81E" authorityName="Straneo" authorityYear="1938" box="[234,359,1993,2012]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA826538FF8F11D0F877091FF81E" box="[234,359,1993,2012]" italics="true" pageId="1" pageNumber="414">M. perakianus</emphasis>
</taxonomicName>
, ♀ from Song Thanh NP.
</paragraph>
<paragraph id="8BD436906538FF8F11D8F85B0BE4F7D3" blockId="1.[189,1399,1967,2065]" pageId="1" pageNumber="414">
Рис. 1–3. Габитус дорЗально
<taxonomicName id="4C6B4D136538FF8F10C9F8580AEFF83B" box="[499,663,2021,2041]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA826538FF8F10C9F8580A19F83A" box="[499,609,2021,2040]" italics="true" pageId="1" pageNumber="414">Metabacetus</emphasis>
spp.
</taxonomicName>
: 1 —
<taxonomicName id="4C6B4D136538FF8F13F5F85B0B47F83A" authorityName="Will et Park" authorityYear="2008" box="[719,831,2021,2040]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA826538FF8F13F5F85B0B47F83A" box="[719,831,2021,2040]" italics="true" pageId="1" pageNumber="414">M. hermanni</emphasis>
</taxonomicName>
, ♂ иЗ Национального парка Суансон; 2 —
<taxonomicName id="4C6B4D136538FF8F15E4F85B0D39F83A" authorityName="Straneo" authorityYear="1938" box="[1246,1345,2021,2040]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA826538FF8F15E4F85B0D39F83A" box="[1246,1345,2021,2040]" italics="true" pageId="1" pageNumber="414">M. laotinus</emphasis>
</taxonomicName>
, ♀ иЗ провинции Каобанг; 3 —
<taxonomicName id="4C6B4D136538FF8F1094F8430A55F7D2" authorityName="Straneo" authorityYear="1938" box="[430,557,2045,2064]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA826538FF8F1094F8430A55F7D2" box="[430,557,2045,2064]" italics="true" pageId="1" pageNumber="414">M. perakianus</emphasis>
</taxonomicName>
, ♀ иЗ Национального парка Сонгтхань.
</paragraph>
</caption>
<paragraph id="8BD43690653BFF8C1187FE860AA8FD9E" blockId="2.[189,776,315,764]" pageId="2" pageNumber="415">
neath; genae fairly long, smooth, subtransverse, meeting neck at sharp obtuse angle, anteroventrally without antennal grooves; frontal sulci reaching level of anterior supra-ocular seta, posteriorly diverging or slightly S-shaped at the seta. Labrum mostly subsinuate at apical margin. Mandibles rather long, narrow and slightly incurved apically, with ventral scrobal ridge straight or subconvex. Apical maxillary and labial palpomere fusiform.
<collectingRegion id="49AFF872653BFF8C10F8FE4B0A6CFDC9" box="[450,532,502,523]" country="Myanmar" name="Chin" pageId="2" pageNumber="415">Mentum</collectingRegion>
tooth triangular, narrowly rounded, faintly longitudinally grooved. Setation: two supraocular setae and one clypeal seta on each side; labrum sexsetose; submentum quadrisetose, mentum bisetose.
</paragraph>
<paragraph id="8BD43690653BFF8C11D8FDDC0B93FE45" blockId="2.[189,776,315,764]" lastBlockId="2.[812,1399,315,764]" pageId="2" pageNumber="415">Pronotum quadrisetose, subquadrate. Sides rounded, finely beaded, subsinuate or straight in front of obtuse yet sharp basal angles, these with minute, almost imperceptible, blunt tooth; lateral margin distinctly explanate and reflexed or not. Base laterally oblique, without bead; apical bead obliterate in middle 1/5–1/3; basal transverse impression missing, apical one vague. Basal sulci narrow, deep, smooth to crenulate at bottom, slightly converging apicad, reaching base, and running on basal third.</paragraph>
<paragraph id="8BD43690653BFF8C1268FE360D0DFD6E" blockId="2.[812,1399,315,764]" pageId="2" pageNumber="415">
Elytra parallel-sided, truncate basally, with distinct humeri, a nearly imperceptible humeral tooth and a slight preapical sinuation. Basal ridge entire, humeral angle obtuse or very so. Striae entire, deep, more or less distinctly punctate, stria 7 separately reaching apex, stria 8 adjoining humerus or lateral margin just outside. Intervals 8 and 9 subequally wide except basally, interval 9 with umbilical seta series (USS) consisting of 6–1–7 umbilical setae/setigerous pores (
<collectingCountry id="F37C7600653BFF8C1589FDFB0CACFD99" box="[1203,1236,582,603]" name="United States of America" pageId="2" pageNumber="415">US</collectingCountry>
), i.e., six
<collectingCountry id="F37C7600653BFF8C1401FDFB0D21FD99" box="[1339,1369,582,603]" name="United States of America" pageId="2" pageNumber="415">US</collectingCountry>
in anterior group, seven (3+2+2) in posterior group, and one intermediate seta. Parascutellar seta at base of stria 2. Stria 7 with two setae before apex, apical one being smaller than preapical.
</paragraph>
<paragraph id="8BD43690653BFF8C1268FD0C0D0EFD3E" blockId="2.[812,1399,315,764]" pageId="2" pageNumber="415">Ventral side: prosternum with median groove distinct or not, prosternal process not bordered. Metepisternum longer than wide. Metaventrite laterally and abdominal sternite II rather</paragraph>
<paragraph id="8BD43690653BFF8C11D8F83C0B21F807" blockId="2.[189,1399,1921,2065]" pageId="2" pageNumber="415">
Figs 4–18. Aedeagus of
<taxonomicName id="4C6B4D13653BFF8C1080F83C0A25F857" box="[442,605,1921,1941]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA82653BFF8C1080F83C0A50F856" box="[442,552,1921,1940]" italics="true" pageId="2" pageNumber="415">Metabacetus</emphasis>
spp.
</taxonomicName>
: 4–5, 11, 15–16 —
<taxonomicName id="4C6B4D13653BFF8C1234F83F0B06F856" authorityName="Will et Park" authorityYear="2008" box="[782,894,1921,1940]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653BFF8C1234F83F0B06F856" box="[782,894,1921,1940]" italics="true" pageId="2" pageNumber="415">M. hermanni</emphasis>
</taxonomicName>
; 6–7, 12, 17–18 —
<taxonomicName id="4C6B4D13653BFF8C1509F83F0CEEF856" authorityName="Straneo" authorityYear="1938" box="[1075,1174,1921,1940]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA82653BFF8C1509F83F0CEEF856" box="[1075,1174,1921,1940]" italics="true" pageId="2" pageNumber="415">M. laotinus</emphasis>
</taxonomicName>
; 8–14 —
<taxonomicName id="4C6B4D13653BFF8C15C8F83F0D08F856" authorityName="Straneo" authorityYear="1938" box="[1266,1392,1921,1940]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA82653BFF8C15C8F83F0D08F856" box="[1266,1392,1921,1940]" italics="true" pageId="2" pageNumber="415">M. perakianus</emphasis>
</taxonomicName>
; 4–12 — median lobe; 13, 15, 17 — right paramere; 14, 16, 18 — left paramere; 4, 6, 8, 13, 15, 17 — left aspect; 5, 7, 9, 14, 16, 18 — right aspect; 10–12 — ventral aspect;
<emphasis id="B91FEA82653BFF8C10A3F80F09D4F807" bold="true" box="[409,428,1970,1989]" italics="true" pageId="2" pageNumber="415">vc</emphasis>
— ventral longitudinal carina. Scale bar:
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.
</paragraph>
<paragraph id="8BD43690653BFF8C11D8F8730BF8F7D3" blockId="2.[189,1399,1921,2065]" pageId="2" pageNumber="415">
Рис. 4–18. Эдеагус
<taxonomicName id="4C6B4D13653BFF8C10AFF8700A4FF823" box="[405,567,1997,2017]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA82653BFF8C10AFF8700A7BF822" box="[405,515,1997,2016]" italics="true" pageId="2" pageNumber="415">Metabacetus</emphasis>
spp.
</taxonomicName>
: 4–5, 11, 15–16 —
<taxonomicName id="4C6B4D13653BFF8C13DEF8730B2CF822" authorityName="Will et Park" authorityYear="2008" box="[740,852,1997,2016]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653BFF8C13DEF8730B2CF822" box="[740,852,1997,2016]" italics="true" pageId="2" pageNumber="415">M. hermanni</emphasis>
</taxonomicName>
; 6–7, 12, 17–18 —
<taxonomicName id="4C6B4D13653BFF8C153FF8730C10F822" authorityName="Straneo" authorityYear="1938" box="[1029,1128,1997,2016]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA82653BFF8C153FF8730C10F822" box="[1029,1128,1997,2016]" italics="true" pageId="2" pageNumber="415">M. laotinus</emphasis>
</taxonomicName>
; 8–14 —
<taxonomicName id="4C6B4D13653BFF8C15F8F8730D47F822" authorityName="Straneo" authorityYear="1938" box="[1218,1343,1997,2016]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA82653BFF8C15F8F8730D47F822" box="[1218,1343,1997,2016]" italics="true" pageId="2" pageNumber="415">M. perakianus</emphasis>
</taxonomicName>
; 4–12 — среднЯЯ долЯ; 13, 15, 17 — праваЯ парамера; 14, 16, 18 — леваЯ парамера; 4, 6, 8, 13, 15, 17 — слева; 5, 7, 9, 14, 16, 18 — справа; 10–12 — вентрально;
<emphasis id="B91FEA82653BFF8C10B0F84009E5F7D2" bold="true" box="[394,413,2045,2064]" italics="true" pageId="2" pageNumber="415">vc</emphasis>
— продольный вентральный киль. Масштаб: 1 мм.
</paragraph>
<paragraph id="8BD43690653AFF8D1187FE860A9DFE63" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">densely punctate; mesepisterna basally, metepisterna, and sides of abdominal sternites III–V less distinctly so. Sternites IV–VI with one pair of obligatory paramedian setae. Sternite VII bisetose in male, quadrisetose in female, all setae marginal.</paragraph>
<paragraph id="8BD43690653AFF8D11D8FE1B0A81FC43" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">Legs slender, protibia not apically dilated, with two close lateroapical setae (spinules), apical seta being stronger than preapical. Setation otherwise as follows: Profemur without anterior (anteroventral) setae, with three posterior setae, basal, medioventral, and apical. Mesotibia with lateral armature consisting of moderately strong spiniform setae, 3–4 anterolateral and one posterolateral near distal anterolateral seta (not counting apical setae); inner setal brush (at posteromesal margin) continuous and nearly unspecialized, consisting of ca. 16 setae. Metatibia externally glabrous. Meso- and metatarsi with tarsomere 1 externally (anteriorly) sulcate and slightly shorter than tarsomeres 2 to 4 combined, without distinct lateral setae. Tarsomere 5 ventrally glabrous, dorsal (dorsolateral) seta almost median in position; tarsomeres 1 and 2 with dorsoapical setae (DAS), these being much shorter in metatarsi; pro- and mesotarsomere 4 with lamellate ventroapical setae. Basal three protarsomeres dilated and biserially squamose in male, basal two protarsomeres toothed at latero-apical angles in female.</paragraph>
<paragraph id="8BD43690653AFF8D11D8FC3B09B3FBC5" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">Aedeagus (Figs 4–18): median lobe geniculate, with a well-developed longitudinal median carina in ventral curve; apex short, narrow, more or less triangular; internal sac unarmed. Parameres subequal in length, right one apically rounded and spoon-shaped.</paragraph>
<paragraph id="8BD43690653AFF8D11D8FBB60911FAE9" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">
Tergite VIII and sternite VIII in female as in Figs 26–26. Female genitalia (urite IX) and reproductive tract (examined in
<taxonomicName id="4C6B4D13653AFF8D11EDFBFC092DFB94" authorityName="Will et Park" authorityYear="2008" box="[215,341,1089,1110]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653AFF8D11EDFBFC092DFB94" box="[215,341,1089,1110]" italics="true" pageId="3" pageNumber="416">M. hermanni</emphasis>
</taxonomicName>
): laterotergite ventrally with a long process which is directed apicad and bears a brush of longer setae at apex (Fig. 34); gonosubcoxite at apex with 2–3 very short, dense, apically blunt, setae; gonocoxite slender, crescent, as long as gonosubcoxite, with a small ensiform, apically blunt, ventral (outer) seta a third from base and double preapical nematiform seta, no dorsal setae;
<emphasis id="B91FEA82653AFF8D133DFB5C0AD0FB34" box="[519,680,1249,1270]" italics="true" pageId="3" pageNumber="416">bursa copulatrix</emphasis>
membranous, spermatheca convoluted and very long, consisting of ca. 60 tight turns.
</paragraph>
<paragraph id="8BD43690653AFF8D11D8FA8C0940FAA3" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">
GEOGRAPHIC DISTRIBUTION. Throughout the
<collectingRegion id="49AFF872653AFF8D13E4FA8C094CFAA3" country="Democratic Republic of the Congo" name="Orientale" pageId="3" pageNumber="416">Oriental region</collectingRegion>
.
</paragraph>
<paragraph id="8BD43690653AFF8D11D8FADB0A5CF9C3" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">
<materialsCitation id="3B033CCD653AFF8D11D8FADB0A67F9C3" country="Vietnam" elevation="1" location="Vietnam" pageId="3" pageNumber="416" specimenCount="1">
HABITATS AND HABITS. Many adults of the three species reviewed below were collected at light at night, at the edges of broad-leaved monsoon forests at 600–
<quantity id="4C939B75653AFF8D13B0FA260AA1FA73" box="[650,729,1435,1457]" metricMagnitude="0" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="416" unit="m" value="1.45">
<elevation id="0046D1A3653AFF8D13B0FA260AA1FA73" box="[650,729,1435,1457]" metricMagnitude="0" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="416" unit="m" value="1.45">1.450 m</elevation>
</quantity>
altitudes. Those of the most common species
<taxonomicName id="4C6B4D13653AFF8D136AFA0A0AB3FA09" authorityName="Will et Park" authorityYear="2008" box="[592,715,1462,1484]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653AFF8D136AFA0A0AB3FA09" box="[592,715,1462,1484]" italics="true" pageId="3" pageNumber="416">M. hermanni</emphasis>
</taxonomicName>
occur in montane regions of
<collectingCountry id="F37C7600653AFF8D1090FA6C0985FA25" box="[426,509,1489,1511]" name="Vietnam" pageId="3" pageNumber="416">Vietnam</collectingCountry>
at 400–
<quantity id="4C939B75653AFF8D136CFA6C0AD2FA24" box="[598,682,1489,1510]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="416" unit="m" value="1.7">
<elevation id="0046D1A3653AFF8D136CFA6C0AD2FA24" box="[598,682,1489,1510]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="416" unit="m" value="1.7">1.700 m</elevation>
</quantity>
altitudes where they dwell in forest leaf-litter
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C371651B653AFF8D11D8F9BB0B7EF883" pageId="3" pageNumber="416" type="discussion">
<paragraph id="8BD43690653AFF8D11D8F9BB0AA4F979" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">
COMMENTS.
<bibRefCitation id="EFFA4B61653AFF8D1048F9BB0A43F9D9" author="Will K. W. &amp; Park J. K." box="[370,571,1542,1564]" pageId="3" pageNumber="416" pagination="189 - 196" refId="ref18022" refString="Will K. W., Park J. K. 2008. A new species of the oriental abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus // Coleopt. Bull. Vol. 62. P. 189 - 196. https: // doi. org / 10.1649 / 980.1." type="journal article" year="2008">Will and Park [2008]</bibRefCitation>
stated that most characters of
<taxonomicName id="4C6B4D13653AFF8D1029F99C09F1F9F4" authorityName="Bates" authorityYear="1892" box="[275,393,1569,1590]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA82653AFF8D1029F99C09F1F9F4" box="[275,393,1569,1590]" italics="true" pageId="3" pageNumber="416">Metabacetus</emphasis>
</taxonomicName>
are symplesiotypic while they described well-developed mentum pits, apparently apotypic character, for at least
<taxonomicName id="4C6B4D13653AFF8D103DF9EA0989F9A9" authority="Will, 2008" authorityName="Will et Park" authorityYear="2008" box="[263,497,1622,1644]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653AFF8D103DF9EA09F9F9A9" box="[263,385,1622,1644]" italics="true" pageId="3" pageNumber="416">M. hermanni</emphasis>
<bibRefCitation id="EFFA4B61653AFF8D10B2F9EB0989F9A9" author="Will K. W. &amp; Park J. K." box="[392,497,1622,1644]" pageId="3" pageNumber="416" pagination="189 - 196" refId="ref18022" refString="Will K. W., Park J. K. 2008. A new species of the oriental abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus // Coleopt. Bull. Vol. 62. P. 189 - 196. https: // doi. org / 10.1649 / 980.1." type="journal article" year="2008">Will, 2008</bibRefCitation>
</taxonomicName>
. The fact that these pits have been reported [
<bibRefCitation id="EFFA4B61653AFF8D1071F9CC0A7DF944" author="Gueourguiev B. V." box="[331,517,1649,1671]" pageId="3" pageNumber="416" pagination="35 - 50" refId="ref17046" refString="Gueourguiev B. V. 2013. On five species of the tribes Abacetini and Pterostichini (Coleoptera, Carabidae) // ZooKeys. Vol. 352. P. 35 - 50. doi: 10.3897 / zookeys. 352.6294." type="book chapter" year="2013">Guéourguiev, 2013</bibRefCitation>
] for
<taxonomicName id="4C6B4D13653AFF8D130FF9CC0896F962" authority="Gueourguiev, 2013" authorityName="Gueourguiev" authorityYear="2013" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="willi">
<emphasis id="B91FEA82653AFF8D130FF9CC0AF8F944" box="[565,640,1649,1670]" italics="true" pageId="3" pageNumber="416">M. willi</emphasis>
<bibRefCitation id="EFFA4B61653AFF8D13B2F9CC0896F962" author="Gueourguiev B. V." pageId="3" pageNumber="416" pagination="35 - 50" refId="ref17046" refString="Gueourguiev B. V. 2013. On five species of the tribes Abacetini and Pterostichini (Coleoptera, Carabidae) // ZooKeys. Vol. 352. P. 35 - 50. doi: 10.3897 / zookeys. 352.6294." type="book chapter" year="2013">Guéourguiev, 2013</bibRefCitation>
</taxonomicName>
, and found in
<taxonomicName id="4C6B4D13653AFF8D1056F9310A20F962" authority="Straneo, 1938" authorityName="Straneo" authorityYear="1938" box="[364,600,1675,1697]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA82653AFF8D1056F93109ACF962" box="[364,468,1675,1697]" italics="true" pageId="3" pageNumber="416">M. laotinus</emphasis>
Straneo, 1938
</taxonomicName>
and
<taxonomicName id="4C6B4D13653AFF8D13B8F9310938F979" authority="Straneo, 1938" authorityName="Straneo" authorityYear="1938" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA82653AFF8D13B8F9310B7FF962" box="[642,775,1675,1697]" italics="true" pageId="3" pageNumber="416">M. perakianus</emphasis>
Straneo, 1938
</taxonomicName>
, suggests monophyletic origin of the genus.
</paragraph>
<paragraph id="8BD43690653AFF8D11D8F97C0B7EF883" blockId="3.[189,776,315,1857]" pageId="3" pageNumber="416">
Another apotypic feature of
<taxonomicName id="4C6B4D13653AFF8D10D5F97C0A12F914" authorityName="Bates" authorityYear="1892" box="[495,618,1729,1750]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA82653AFF8D10D5F97C0A12F914" box="[495,618,1729,1750]" italics="true" pageId="3" pageNumber="416">Metabacetus</emphasis>
</taxonomicName>
, the elytra without discal setae, should be corrected because vestigial d2 has been found in all of the three species examined by me. It is very minute and thence hard to detect in
<taxonomicName id="4C6B4D13653AFF8D1360F8AC0AA0F8E4" authorityName="Will et Park" authorityYear="2008" box="[602,728,1809,1830]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="hermanni">
<emphasis id="B91FEA82653AFF8D1360F8AC0AA0F8E4" box="[602,728,1809,1830]" italics="true" pageId="3" pageNumber="416">M. hermanni</emphasis>
</taxonomicName>
, but more or less well traceable in
<taxonomicName id="4C6B4D13653AFF8D10E0F8910A30F882" authorityName="Straneo" authorityYear="1938" box="[474,584,1835,1857]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="laotinus">
<emphasis id="B91FEA82653AFF8D10E0F8910A30F882" box="[474,584,1835,1857]" italics="true" pageId="3" pageNumber="416">M. laotinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D13653AFF8D134CF8910B78F882" authorityName="Straneo" authorityYear="1938" box="[630,768,1835,1857]" class="Insecta" family="Carabidae" genus="Metabacetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="perakianus">
<emphasis id="B91FEA82653AFF8D134CF8910B78F882" box="[630,768,1835,1857]" italics="true" pageId="3" pageNumber="416">M. perakianus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

59
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@ -1,42 +1,43 @@
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</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED1D16563FCE5" pageId="4" pageNumber="198" type="diagnosis">
<paragraph id="8BC4F158B730FF9EA71ED1D16563FCE5" blockId="4.[812,1400,517,1291]" pageId="4" pageNumber="198">
DIAGNOSIS. A species of
<emphasis id="B90F2D4AB730FF9EA015D1CE6253FD05" box="[1113,1128,656,677]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<taxonomicName id="4C7B8ADBB730FF9EA03FD1D162C2FD05" authorityName="Smith" authorityYear="1969" box="[1139,1273,655,677]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
(
<emphasis id="B90F2D4AB730FF9EA037D1CE62B1FD05" box="[1147,1162,656,677]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.)
<emphasis id="B90F2D4AB730FF9EA0D1D1CE62C2FD05" box="[1181,1273,656,677]" italics="true" pageId="4" pageNumber="198">setitarsus</emphasis>
</taxonomicName>
-group; body
<quantity id="4C835CBDB730FF9EA760D1F46593FD1F" box="[812,936,682,704]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="198" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
; proboscis short, labrum only slightly longer than head height; laterotergite with black setae; abdomen greyish pruinose, only pale setose; wing with incomplete veins M
<subScript id="17FFF31DB730FF9EA730D05765B8FCB5" attach="left" box="[892,899,777,789]" fontSize="5" pageId="4" pageNumber="198">1</subScript>
and CuA+CuP, pterostigma pale yellowish, halter brown. Male: fore basitarsus slightly thickened; wing hyaline.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9FA71ED014622AFCC7" lastPageId="5" lastPageNumber="199" pageId="4" pageNumber="198" type="description">
<paragraph id="8BC4F158B730FF9FA71ED0146490FE27" blockId="4.[812,1400,517,1291]" lastBlockId="5.[189,776,315,2071]" lastPageId="5" lastPageNumber="199" pageId="4" pageNumber="198">
DESCRIPTION. Body length 3.5–4.2, wing
<quantity id="4C835CBDB730FF9EA0B1D014634FFCFF" box="[1277,1396,842,864]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="4" pageNumber="198" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
.
<emphasis id="B90F2D4AB730FF9EA760D03B655BFCDA" bold="true" box="[812,864,869,890]" pageId="4" pageNumber="198">Male</emphasis>
(
<figureCitation id="1340EDDDB730FF9EA73ED03B6589FCDA" box="[882,946,869,891]" captionStart="Figs 7–8" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[212,1396,1334,1928]" captionTargetId="figure-687@4.[189,1399,1334,1924]" captionTargetPageId="4" captionText="Figs 7–8. Empis (Empis) windhoekiensis, sp.n., ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 7–8. Empis (Empis) windhoekiensis, sp.n., ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="4" pageNumber="198">Fig. 7</figureCitation>
). Head with greyish, dense pruinescence on frons, face, clypeus (except shiny upper margin), ocellar triangle, occiput and postgena; black setose. Eyes holoptic, with upper ommatidia enlarged. Frons represented by very small, subtriangular space just below ocellar triangle and slightly larger space just above antennae. Face broad, almost parallel-sided, bare. Ocellar triangle with several fine setae, 2 setae somewhat longer. Occiput with fine, moderately long postoculars and several similar setae laterally; postgena with numerous hair-like setae. Antenna black; scape short, slightly longer than pedicel, both with short setulae; postpedicel elongate-conical, with lower margin slightly concave distally, nearly 2.5X as long as basal width; stylus short, nearly 2.5X shorter than postpedicel. Proboscis short, with labrum only slightly longer than head height (1.2–1.3X); labium lacking annulations, bearing scattered, minute setulae; labella moderately long, nearly 2X shorter than labrum, with small desclerotized space on about distal 1/3 (visible
<emphasis id="B90F2D4AB731FF9FA6EBD20964DCFECC" box="[679,743,343,364]" italics="true" pageId="5" pageNumber="199">in situ</emphasis>
as notch); palpus black, with black, scattered setulae.
</paragraph>
<caption id="DF04A1D0B730FF9EA4AED4EF625DF7B1" ID-DOI="http://doi.org/10.5281/zenodo.13178206" ID-Zenodo-Dep="13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="4" pageNumber="198" startId="4.[226,263,1969,1988]" targetBox="[212,1396,1334,1928]" targetPageId="4" targetType="figure">
<paragraph id="8BC4F158B730FF9EA4AED4EF626BF87C" blockId="4.[188,1398,1967,2065]" pageId="4" pageNumber="198">
Figs 7–8.
<taxonomicName id="4C7B8ADBB730FF9EA574D4EC647BF864" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[312,576,1969,1988]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA574D4EC6756F864" box="[312,365,1970,1988]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
(
<emphasis id="B90F2D4AB730FF9EA536D4EC6794F864" box="[378,431,1970,1988]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA5F1D4EF647BF864" box="[445,576,1969,1988]" italics="true" pageId="4" pageNumber="198">windhoekiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB730FF9EA607D4EF644DF864" bold="true" box="[587,630,1969,1988]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA607D4EF644DF864" box="[587,630,1969,1988]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus.
</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED4B8625DF7B1" blockId="4.[188,1398,1967,2065]" pageId="4" pageNumber="198">
Рис. 7–8.
<taxonomicName id="4C7B8ADBB730FF9EA57BD4B86407F858" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[311,572,2021,2040]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA57BD4B86757F858" box="[311,364,2022,2040]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
(
<emphasis id="B90F2D4AB730FF9EA53BD4B86797F858" box="[375,428,2022,2040]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA5F5D4BB6407F858" box="[441,572,2021,2040]" italics="true" pageId="4" pageNumber="198">windhoekiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB730FF9EA609D4BB644BF858" bold="true" box="[581,624,2021,2040]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA609D4BB644BF858" box="[581,624,2021,2040]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус.
</paragraph>
</caption>
<paragraph id="8BC4F158B731FF9FA4AED2D5645DFC21" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Thorax black in ground-colour, black setose; mesoscutum faintly greyish pruinose (dorsal view), prothoracic sclerites and mesopleuron somewhat denser greyish pruinose.Proepisternum with 4–5 rather short, fine setae on lower part and 4–6 similar setae on upper part. Prosternum bare. Antepronotum with 3 moderately long setae on each side. Postpronotal lobe with 1 strong, long and several fine, short setae. Mesonotal setae well-differentiated; acrostichals moderately long, thin, arranged in 2 close, irregular rows, lacking on prescutellar depression; dorsocentrals longer than acrostichals, 1–2-serial before suture, uniserial and longer along prescutellar depression, 2 prescutellar setae longest; 1 fine, moderately long presutural intra-alar, 1 longer presutural supra-alar, 3 notopleurals (2 setae stronger; additionally, with some setulae anteriorly), 1 postsutural supra-alar (with 3–4 additional short, fine setae anteriorly), 1 long and 1 minute postalars, 4 scutellars (apical setae long and cruciate, lateral setae only slightly shorter). Laterotergite with 5–6 long and some short fine setae. Anterior and posterior spiracles dark brown.</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED0D864B2F927" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Legs rather robust, black, subshiny (except coxae), faintly greyish pruinose, black setose; coxae densely pruinose. Fore coxa and trochanter almost bare, only with scattered minute setulae; hind trochanter with long, dense setae. Fore femur with row of short anteroventral fine setae on about basal half, almost bare posteroventrally, only some minute posteroventral setulae closer to base. Fore tibia with numerous long, fine setae posterodorsally (longest setae about 2X as long as tibia width). Fore basitarsus thickened but only slightly stouter than fore tibia at apex; with moderately long, dense, fine setae posterodorsally; tarsomeres 1–4 with rather long setae of subapical circlet. Mid femur with complete rows of numerous anteroventral and posteroventral setae, which are long on about middle part of femur and short near its base and apex; longest anteroventral setae nearly 1.5X as long as femur middle width. Mid tibia with several intermixed moderately long to long ad setae closer to base, more or less distinct row of moderately long posterodorsal setae and rows of moderately long anteroventral and short posteroventral setae. Mid basitarsus nearly as broad as mid tibia at apex; tarsomeres 1 and 2 with short spine-like setae ventrally, tarsomeres 1–3 with rather long setae of subapical circlet (longer dorsally). Hind femur with complete rows of long anteroventral (longest setae on about apical half nearly 1.5X as long as femur middle width) and short posteroventral setae. Hind tibia slightly uniformly thickened (lateral view) and somewhat curved (posterior view); with numerous moderately long to long setae dorsally; no seta in posteroapical comb. Hind basitarsus slender; with dorsal setation as on hind tibia, numerous short, spine-like setae ventrally.</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED5D567A3F8FC" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">
Wing membrane hyaline, veins mostly brownish, paler proximally; M
<subScript id="17FFF31DB731FF9FA500D5EB6768F961" attach="left" box="[332,339,1717,1729]" fontSize="5" pageId="5" pageNumber="199">1</subScript>
beyond cell dm and CuA+CuP (anal vein) very thin, M
<subScript id="17FFF31DB731FF9FA579D58E6707F97C" attach="left" box="[309,316,1744,1756]" fontSize="5" pageId="5" pageNumber="199">2</subScript>
and M
<subScript id="17FFF31DB731FF9FA5CDD58E67B3F97C" attach="left" box="[385,392,1744,1756]" fontSize="5" pageId="5" pageNumber="199">4</subScript>
weakened before wing margin; M
<subScript id="17FFF31DB731FF9FA69AD58E64E6F97C" attach="left" box="[726,733,1744,1756]" fontSize="5" pageId="5" pageNumber="199">1</subScript>
and CuA+CuP incomplete; cell dm short, with slightly elongate apex. Pterostigma pale yellowish, hardly distinguishable. Basal costal seta present, long, black. Anal lobe well-developed; axillary incision right-angled. Squama brownish, pale fringed. Halter brown.
</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED43F643AF7B7" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Abdomen black in ground-colour, uniformly greyish pruinose, mostly covered with numerous pale hair-like setae longer on tergites laterally; sternite 7 with some fine black posteromarginal setae, segment 8 with black setae of different lengths. Segment 8 with separated sclerites; tergite 8 without projections, entire, somewhat concave posteriorly; sternite 8 desclerotized medially.</paragraph>
<paragraph id="8BC4F158B731FF9FA71ED265626CFCEC" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
Terminalia (
<figureCitation id="1340EDDDB731FF9FA787D2656233FEF0" box="[971,1032,315,337]" captionStart="Figs 7–8" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[212,1396,1334,1928]" captionTargetId="figure-687@4.[189,1399,1334,1924]" captionTargetPageId="4" captionText="Figs 7–8. Empis (Empis) windhoekiensis, sp.n., ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 7–8. Empis (Empis) windhoekiensis, sp.n., ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="5" pageNumber="199">Fig. 8</figureCitation>
) moderately large, concolorous with abdomen (except noted), black setose. Epandrium divided into two lamellae (dorsal bridge absent); epandrial lamella subtriangular (lateral view), with rather deep excision closer to base dorsally, with long, fine setae along lateral margins. Hypandrium short, subtriangular (ventral view), with 2 long and 2 short setae. Hypoproct strongly sclerotised, elongate, reaching phallus, narrow on middle part and broadened distally, bare. Phallus brownish yellow, mostly hidden, short, evenly curved, thick on about basal 2/3 and slender on apical 1/3. Ejaculatory apodeme large, extended far beyond basal curvature of phallus, without lateral wings. Cerci closely attached and almost fused anteriorly, separated from epandrium; cercus almost entirely divided in two lobes by narrow notch (lateral view) posteriorly; dorsal lobe rather trapezoid (lateral view), with short, dense, strong setae anteriorly and scattered, fine setae on remaining part; ventral lobe much narrower and slightly longer than dorsal lobe (lateral view), slightly extended beyond epandrial lamella, digitiform, gently curved downwards on about apical half, with some minute setulae.
</paragraph>
<paragraph id="8BC4F158B731FF9FA71ED00F622AFCC7" blockId="5.[812,1399,315,1404]" box="[850,1041,849,871]" pageId="5" pageNumber="199">
<emphasis id="B90F2D4AB731FF9FA71ED00F6598FCC6" bold="true" box="[850,931,849,870]" pageId="5" pageNumber="199">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED0356245FA8C" pageId="5" pageNumber="199" type="discussion">
<paragraph id="8BC4F158B731FF9FA71ED0356245FA8C" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
REMARKS.
<taxonomicName id="4C7B8ADBB731FF9FA798D0326290FC20" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[980,1195,875,897]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov.">
<emphasis id="B90F2D4AB731FF9FA798D0326290FC20" box="[980,1195,875,897]" italics="true" pageId="5" pageNumber="199">Empis windhoekiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB731FF9FA0FFD03562DFFC20" bold="true" box="[1203,1252,875,896]" pageId="5" pageNumber="199">
<taxonomicNameLabel id="A23C9031B731FF9FA0FFD03562DFFC20" box="[1203,1252,875,896]" pageId="5" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C7B8ADBB731FF9FA760D0D9623EFC3C" authorityName="Smith" authorityYear="1969" box="[812,1029,902,924]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
<emphasis id="B90F2D4AB731FF9FA760D0D96500FC3C" box="[812,827,903,924]" italics="true" pageId="5" pageNumber="199">E</emphasis>
. (
<emphasis id="B90F2D4AB731FF9FA71AD0D965A8FC3C" box="[854,915,903,924]" italics="true" pageId="5" pageNumber="199">Empis</emphasis>
)
<emphasis id="B90F2D4AB731FF9FA7EBD0D9623EFC3C" box="[935,1029,903,924]" italics="true" pageId="5" pageNumber="199">setitarsus</emphasis>
</taxonomicName>
-group [
<bibRefCitation id="EFEA8CA9B731FF9FA015D0D86348FC3B" author="Daugeron C. &amp; Grootaert P." box="[1113,1395,902,923]" pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
;
<bibRefCitation id="EFEA8CA9B731FF9FA760D0FF621CFC16" author="Sinclair B. J. &amp; Daugeron C." box="[812,1063,929,951]" pageId="5" pageNumber="199" pagination="1221 - 1235" refId="ref6424" refString="Sinclair B. J., Daugeron C. 2017. Empididae (empidid dance flies or balloon flies) // A. H. Kirk-Spriggs, B. J. Sinclair (eds.). Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: SANBI Graphics &amp; Editing. P. 1221 - 1235." type="book chapter" year="2017">Sinclair, Daugeron, 2017</bibRefCitation>
]. Within the key to species of this group the new species would run to the couplet 19 joining
<taxonomicName id="4C7B8ADBB731FF9FA12DD0E265D8FC4C" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="B90F2D4AB731FF9FA12DD0E2634BFC71" box="[1377,1392,956,977]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D088655CFC4B" box="[812,871,982,1003]" italics="true" pageId="5" pageNumber="199">vetula</emphasis>
<bibRefCitation id="EFEA8CA9B731FF9FA721D08865D8FC4C" author="Smith K. G. V." box="[877,995,982,1004]" pageId="5" pageNumber="199" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B8ADBB731FF9FA05ED0896567FBA6" authority="Daugeron et Grootaert, 2003" authorityName="Daugeron et Grootaert" authorityYear="2003" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="cinerarius">
<emphasis id="B90F2D4AB731FF9FA05ED089621AFC4C" box="[1042,1057,983,1004]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA061D08962ABFC4C" box="[1069,1168,983,1004]" italics="true" pageId="5" pageNumber="199">cinerarius</emphasis>
<bibRefCitation id="EFEA8CA9B731FF9FA0DBD0886567FBA6" author="Daugeron C. &amp; Grootaert P." pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron et Grootaert, 2003</bibRefCitation>
</taxonomicName>
[
<bibRefCitation id="EFEA8CA9B731FF9FA726D0AF6242FBA6" author="Daugeron C. &amp; Grootaert P." box="[874,1145,1009,1030]" pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
]. The new species can be readily distinguished from both these species primarily by black setae on the laterotergite. In addition, in the male of
<taxonomicName id="4C7B8ADBB731FF9FA12DD779655CFBF6" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="B90F2D4AB731FF9FA12DD779634BFB9C" box="[1377,1392,1063,1084]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D71F655CFBF6" box="[812,871,1089,1110]" italics="true" pageId="5" pageNumber="199">vetula</emphasis>
</taxonomicName>
the phallus is well-exposed, longer, and uniformly slender [
<bibRefCitation id="EFEA8CA9B731FF9FA733D7056217FBD1" author="Smith K. G. V." box="[895,1068,1115,1137]" pageId="5" pageNumber="199" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969: 175</bibRefCitation>
, fig. 209], whereas in the male of the new species the phallus is short, mostly hidden, thick on about basal 2/3 and slender on apical 1/3. Besides the colour of laterotergal setae,
<taxonomicName id="4C7B8ADBB731FF9FA7B5D7F26293FB60" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[1017,1192,1195,1217]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp.n.">
<emphasis id="B90F2D4AB731FF9FA7B5D7F26233FB61" box="[1017,1032,1196,1217]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA059D7F56293FB60" box="[1045,1192,1195,1216]" italics="true" pageId="5" pageNumber="199">windhoekiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB731FF9FA0E3D7F562DBFB60" bold="true" box="[1199,1248,1195,1216]" pageId="5" pageNumber="199">
<taxonomicNameLabel id="A23C9031B731FF9FA0E3D7F562DBFB60" box="[1199,1248,1195,1216]" pageId="5" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C7B8ADBB731FF9FA12DD7F265B4FB7C" authorityName="Daugeron et Grootaert" authorityYear="2003" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="cinerarius">
<emphasis id="B90F2D4AB731FF9FA12DD7F2634BFB61" box="[1377,1392,1196,1217]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D79965B4FB7C" box="[812,911,1223,1244]" italics="true" pageId="5" pageNumber="199">cinerarius</emphasis>
</taxonomicName>
by shorter proboscis (labrum only slightly longer than head height versus labrum about 2X longer than head height), shorter and thicker antennal stylus as well as by the structure of the male hypopygium.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED66F6278FAC1" pageId="5" pageNumber="199" type="etymology">
<paragraph id="8BC4F158B731FF9FA71ED66F6278FAC1" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
ETYMOLOGY. The name of the new species refers to the
<typeStatus id="54C04FFAB731FF9FA71DD6126540FAC1" box="[849,891,1356,1377]" pageId="5" pageNumber="199">type</typeStatus>
locality, Windhoek.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED63862DAFADC" box="[850,1249,1382,1404]" pageId="5" pageNumber="199" type="distribution">
<paragraph id="8BC4F158B731FF9FA71ED63862DAFADC" blockId="5.[812,1399,315,1404]" box="[850,1249,1382,1404]" pageId="5" pageNumber="199">
DISTRIBUTION. Afrotropics:
<collectingCountry id="F36CB1C8B731FF9FA0CBD63862E6FADC" box="[1159,1245,1382,1404]" name="Namibia" pageId="5" pageNumber="199">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

245
data/03/D2/40/03D2404EB736FF9EA77ED0B862DAFE51.xml Normal file
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@ -0,0 +1,245 @@
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<mods:title id="F3050417BEC40B6B1FCC7D1C2E40AF93">Three new species of empidid dance flies of the genus Empis Linnaeus, 1758 (Diptera: Empididae) from Namibia</mods:title>
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<paragraph id="8BC4F158B736FF98A77ED0B8634AFC5D" blockId="2.[812,1400,997,2064]" box="[818,1393,997,1021]" pageId="2" pageNumber="196">
<heading id="D08C4634B736FF98A77ED0B8634AFC5D" box="[818,1393,997,1021]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="196" reason="4">
<taxonomicName id="4C7B8ADBB736FF98A77ED0B86309FC5D" ID-CoL="9J6ZK" authority="Kustov et Shamshev" authorityName="Kustov et Shamshev" authorityYear="2022" box="[818,1330,997,1021]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB736FF98A77ED0B8654FFC5D" box="[818,884,998,1021]" italics="true" pageId="2" pageNumber="196">Empis</emphasis>
(
<emphasis id="B90F2D4AB736FF98A7CFD0B865FFFC5D" box="[899,964,998,1021]" italics="true" pageId="2" pageNumber="196">Empis</emphasis>
)
<emphasis id="B90F2D4AB736FF98A798D0BB626AFC5D" box="[980,1105,997,1021]" italics="true" pageId="2" pageNumber="196">namibiensis</emphasis>
Kustov et Shamshev
</taxonomicName>
,
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<taxonomicNameLabel id="A23C9031B736FF98A170D0BB634AFC5D" box="[1340,1393,997,1021]" pageId="2" pageNumber="196" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A053D75C62BFFBBA" box="[1055,1156,1026,1050]" pageId="2" pageNumber="196" type="description">
<paragraph id="8BC4F158B736FF98A053D75C62BFFBBA" blockId="2.[812,1400,997,2064]" box="[1055,1156,1026,1050]" pageId="2" pageNumber="196">
<figureCitation id="1340EDDDB736FF98A053D75C6244FBBA" box="[1055,1151,1026,1050]" captionStart="Figs 4–6" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 4–6. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 4–6. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="2" pageNumber="196">Figs 4–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A71ED776634CFBCA" pageId="2" pageNumber="196" type="materials_examined">
<paragraph id="8BC4F158B736FF98A71ED776634CFBCA" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
<collectionCode id="ED6A699DB736FF98A71ED776623FFB9A" box="[850,1028,1063,1083]" pageId="2" pageNumber="196">TYPE MATERIAL</collectionCode>
.
<materialsCitation id="3B13FB05B736FF98A05DD779634FFBF2" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="2" pageNumber="196" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C04FFAB736FF98A05DD779625AFB9A" box="[1041,1121,1063,1082]" pageId="2" pageNumber="196" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7D3AD1B736FF98A027D77B6244FB9B" box="[1131,1151,1061,1083]" count="1" pageId="2" pageNumber="196" type="male">♂</specimenCount>
,
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,
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env.,
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,
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,
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,
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(
<collectionCode id="ED6A699DB736FF98A16AD7616355FBF2" box="[1318,1390,1087,1106]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="196" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B13FB05B736FF98A760D7066348FBCA" box="[812,1395,1109,1131]" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU, ZISP" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="2" pageNumber="196" specimenCount="15" specimenCount-female="3" specimenCount-male="12" typeStatus="paratype">
<typeStatus id="54C04FFAB736FF98A760D70665B8FBCB" box="[812,899,1112,1131]" pageId="2" pageNumber="196" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7D3AD1B736FF98A7C0D709658DFBCB" box="[908,950,1109,1131]" count="11" pageId="2" pageNumber="196" type="male">11♂</specimenCount>
<specimenCount id="9D7D3AD1B736FF98A7F7D70965E9FBCB" box="[955,978,1109,1131]" count="3" pageId="2" pageNumber="196" type="female">3♀</specimenCount>
, same data as holotype (
<collectionCode id="ED6A699DB736FF98A0E4D70962D5FBCA" box="[1192,1262,1111,1130]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="196" type="Museum">ZMMU</collectionCode>
,
<specimenCount id="9D7D3AD1B736FF98A0BBD7096327FBCB" box="[1271,1308,1109,1131]" count="1" pageId="2" pageNumber="196" type="male">1 ♂</specimenCount>
—
<collectionCode id="ED6A699DB736FF98A171D7096356FBCA" box="[1341,1389,1111,1130]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="196">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A71ED7316374FAAB" pageId="2" pageNumber="196" type="diagnosis">
<paragraph id="8BC4F158B736FF98A71ED7316374FAAB" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
DIAGNOSIS. A species of
<emphasis id="B90F2D4AB736FF98A015D72E6253FB25" box="[1113,1128,1136,1157]" italics="true" pageId="2" pageNumber="196">E</emphasis>
.
<taxonomicName id="4C7B8ADBB736FF98A03FD73162C2FB25" authorityName="Smith" authorityYear="1969" box="[1139,1273,1135,1157]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
(
<emphasis id="B90F2D4AB736FF98A037D72E62B1FB25" box="[1147,1162,1136,1157]" italics="true" pageId="2" pageNumber="196">E</emphasis>
.)
<emphasis id="B90F2D4AB736FF98A0D1D72E62C2FB25" box="[1181,1273,1136,1157]" italics="true" pageId="2" pageNumber="196">setitarsus</emphasis>
</taxonomicName>
-group; body about
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; proboscis long, labrum 1.5–2X as long as head height; laterotergite with pale setae; abdomen densely light grey pruinose, only pale setose; wing with complete veins M 1 and CuA+CuP. Male: wing hyaline. Female: legs with simple setae, wing somewhat broadened and slightly darkened.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF9EA71ED6516488FB61" lastPageId="4" lastPageNumber="198" pageId="2" pageNumber="196" type="description">
<paragraph id="8BC4F158B736FF98A71ED651627CF89B" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
DESCRIPTION. Body length 4.5–4.8; wing length
<quantity id="4C835CBDB736FF98A10CD651656FFA9F" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="2" pageNumber="196" unit="mm" value="4.1" valueMax="4.2" valueMin="4.0">4–4.2 mm</quantity>
.
<emphasis id="B90F2D4AB736FF98A710D67465ABFA9F" bold="true" box="[860,912,1322,1343]" pageId="2" pageNumber="196">Male</emphasis>
(
<figureCitation id="1340EDDDB736FF98A7D0D67465ECFA9F" box="[924,983,1322,1344]" captionStart="Figs 4–6" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 4–6. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 4–6. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="2" pageNumber="196">Fig. 4</figureCitation>
). Head with greyish, dense pruinescence on frons, face (except shiny lower margin), clypeus (except shiny upper margin), ocellar triangle, occiput and postgena; black setose (except noted). Eyes holoptic, with upper ommatidia enlarged. Frons represented by very small, subtriangular space just below ocellar triangle and slightly larger space just above antennae. Face broad, almost parallel-sided, bare. Ocellar triangle with several subequally short, fine setae. Occiput with fine, moderately long postoculars and several similar setae laterally; postgena with numerous long, pale, hair-like setae. Antenna black; scape short, slightly longer than pedicel, both with short setulae; postpedicel elongate-conical, with narrow base and almost straight margins, nearly 3X as long as basal width and 1.5–1.6X longer than stylus. Proboscis moderately long, with labrum nearly 1.5X as long as head height; labium lacking annulations, bearing scattered, minute setulae; labella moderately long, nearly 2X shorter than labrum; with small, dorsal, desclerotized space on about distal third (visible
<emphasis id="B90F2D4AB736FF98A005D45562BCF880" box="[1097,1159,1803,1824]" italics="true" pageId="2" pageNumber="196">in situ</emphasis>
as notch); palpus black, with black, scattered setulae.
</paragraph>
<paragraph id="8BC4F158B736FF99A71ED461646AFDD7" blockId="2.[812,1400,997,2064]" lastBlockId="3.[189,776,315,897]" lastPageId="3" lastPageNumber="197" pageId="2" pageNumber="196">Thorax black in ground-colour, densely greyish pruinescent; mesoscutum with 4 brownish vittae (dorsal view), lateral vittae (between dorsocentral and supra-alar setae) less distinct. Proepisternum with 6–7 long, fine, pale setae on lower part and 2–3 similar setae on upper part. Prosternum bare. Antepronotum with 4–5 pale setae on each side. Postpronotal lobe with 1 strong, long, black seta and several fine, pale, short setae anteriorly. Mesonotal setae well-differentiated, black; notopleuron with several short, fine, pale setae anteriorly, sometimes pale setulae present just behind postpronotal lobe: acrostichals rather long, biserial, lacking on prescutellar depression; dorsocentrals irregularly 2–3-serial and similar to acrostichals before suture, uniserial and sparser along prescutellar depression, 2 prescutellar setae longest; 1 fine, moderately long presutural intra-alar, 1 similar presutural supra-alar, 3 long notopleurals, 1 postsutural supra-alar (with 2–3 additional short, fine setae anteriorly), 1 long and 1 minute postalars, 4 scutellars (apical setae long and cruciate, lateral setae short and fine).Laterotergite with numerous pale setae of different lengths. Anterior and posterior spiracles brownish.</paragraph>
<paragraph id="8BC4F158B737FF99A4AED12262ADFC91" blockId="3.[189,776,315,897]" lastBlockId="3.[812,1399,315,897]" pageId="3" pageNumber="197">Legs long, slender, entirely dark brown, mostly faintly greyish pruinose, coxae densely pruinose, black setose (except noted). Coxae and trochanters mostly with pale fine setae, mid and hind coxae often with some additional dark setae. Fore femur with rows of very short, fine, mostly pale anteroventral and posteroventral setae. Fore tibia with slightly longer setulae posterodorsally and posteriorly; setae of subapical circlet mostly short, fine, 1 anteroventral and 1 posteroventral setae longer. Fore basitarsus slender; fore tarsomeres covered with short setae (except circlet of somewhat longer subapical setae). Mid femur with row of mostly pale posteroventral setae becoming longer toward base of femur (longest setae nearly 1.5X longer than femur basal width). Mid tibia with 4–5 moderately long dorsal (at most 1.5X longer than tibia width) setae and some numerous, slightly shorter setae ventrally (besides circlet of short subapicals). Mid tarsomeres slender, mostly covered with short setae; mid basitarsus with numerous, short, spine-like setae ventrally. Hind femur with mostly very short anteroventral setae (slightly longer closer to apex); long, pale, fine setae on about basal half posteroventrally and posteriorly. Hind tibia slightly almost uniformly thickened viewed laterally (except extreme base), slightly curved viewed posteriorly; with rows of numerous anterodorsal and posterodorsal setae of different lengths (longest setae at most 1.5X as long as tibia middle width), some rather short setae ventrally. Hind basitarsus slightly, uniformly thickened but not broader than apex of hind tibia; with several moderately setae dorsally and numerous short, spine-like setae ventrally.</paragraph>
<paragraph id="8BC4F158B737FF9EA71ED06867F0FE01" blockId="3.[812,1399,315,897]" lastBlockId="4.[189,776,315,1297]" lastPageId="4" lastPageNumber="198" pageId="3" pageNumber="197">Wing membrane hyaline, veins brownish; longitudinal veins complete, well sclerotised, only CuA+CuP (anal vein) somewhat weakened beyond midway toward wing margin; cell dm short, with elongate apex. Pterostigma brownish. Basal costal seta present, short, fine. Anal lobe well-developed; axillary incision almost right-angled. Squama yellow, pale fringed. Halter yellow.</paragraph>
<caption id="DF04A1D0B737FF99A4AED4EF634DF7B1" ID-DOI="http://doi.org/10.5281/zenodo.13178204" ID-Zenodo-Dep="13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="3" pageNumber="197" startId="3.[226,264,1969,1988]" targetBox="[188,1393,926,1908]" targetPageId="3" targetType="figure">
<paragraph id="8BC4F158B737FF99A4AED4EF6343F87C" blockId="3.[189,1400,1967,2065]" pageId="3" pageNumber="197">
Figs 4–6.
<taxonomicName id="4C7B8ADBB737FF99A572D4EC640EF864" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[318,565,1969,1988]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB737FF99A572D4EC674EF864" box="[318,373,1970,1988]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
(
<emphasis id="B90F2D4AB737FF99A5C8D4EC6780F864" box="[388,443,1970,1988]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
)
<emphasis id="B90F2D4AB737FF99A586D4EF640EF864" box="[458,565,1969,1988]" italics="true" pageId="3" pageNumber="197">namibiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB737FF99A60ED4EF6454F864" bold="true" box="[578,623,1969,1988]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A23C9031B737FF99A60ED4EF6454F864" box="[578,623,1969,1988]" pageId="3" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus.
</paragraph>
<paragraph id="8BC4F158B737FF99A4AED4B8634DF7B1" blockId="3.[189,1400,1967,2065]" pageId="3" pageNumber="197">
Рис. 4–6.
<taxonomicName id="4C7B8ADBB737FF99A576D4B8641DF858" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[314,550,2021,2040]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB737FF99A576D4B86754F858" box="[314,367,2022,2040]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
(
<emphasis id="B90F2D4AB737FF99A530D4B8678AF858" box="[380,433,2022,2040]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
)
<emphasis id="B90F2D4AB737FF99A5F3D4BB641DF858" box="[447,550,2021,2040]" italics="true" pageId="3" pageNumber="197">namibiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB737FF99A67DD4BB6467F858" bold="true" box="[561,604,2021,2040]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A23C9031B737FF99A67DD4BB6467F858" box="[561,604,2021,2040]" pageId="3" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус.
</paragraph>
</caption>
<paragraph id="8BC4F158B730FF9EA4AED2F8679BFDE1" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">Abdomen black in ground-colour, uniformly densely light grey pruinose, tergites with narrowly pale posterior margin; covered with numerous pale hair-like setae longer laterally. Segment 8 with separated sclerites; tergite 8 without projections, entire, somewhat concave posteriorly; sternite 8 broad, desclerotized medially.</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED11867D4FB81" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">
Terminalia (
<figureCitation id="1340EDDDB730FF9EA517D118678DFDFC" box="[347,438,582,604]" captionStart="Figs 4–6" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 4–6. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 4–6. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="4" pageNumber="198">Figs 5–6</figureCitation>
). Epandrium densely greyish pruinose, black setose, divided into two lamellae (dorsal bridge absent); epandrial lamella subtriangular (lateral view), with long setae along upper margin and apically. Hypandrium with basal part faintly greyish pruinose, bearing several moderately long, fine, pale setae; apical part subshiny, yellowish brown; short, in ventral view rather subrectangular, with slightly concave upper margin. Hypoproct slender, digitiform (dorsal view), bare. Phallus yellowish, mostly hidden, short, evenly curved, mostly thick with subapical part narrowed. Ejaculatory apodeme large, extended far beyond basal curvature of phallus, without lateral wings. Cerci black, subshiny, black setose; separated from each other by distinct membraneous space but almost fused narrowly to anterodorsal part of epandrium; cercus somewhat concave posteriorly (lateral view); dorsal lobe subrectangular, short, with short marginal setae; ventral lobe nearly as long as dorsal lobe but very slender, with 2 long setae.
</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED7786488FB61" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">
<emphasis id="B90F2D4AB730FF9EA4AED7786708FB9B" bold="true" box="[226,307,1062,1083]" pageId="4" pageNumber="198">Female.</emphasis>
Similar to male except the following characters. Frons broad, parallel-sided; labrum somewhat longer than in male Acrostichals and dorsocentrals shorter. Legs with shorter, simple setation. Wing broader, slightly darkened. Abdomen densely light grey pruinose, covered with shorter setae. Cercus black, long, with scattered hair-like setulae.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA4AED7986307FE01" pageId="4" pageNumber="198" type="discussion">
<paragraph id="8BC4F158B730FF9EA4AED7986307FE01" blockId="4.[189,776,315,1297]" lastBlockId="4.[812,1398,315,497]" pageId="4" pageNumber="198">
REMARKS.
<taxonomicName id="4C7B8ADBB730FF9EA529D7996427FB7B" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[357,540,1222,1244]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="B90F2D4AB730FF9EA529D7996427FB7B" box="[357,540,1222,1244]" italics="true" pageId="4" pageNumber="198">Empis namibiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB730FF9EA669D798646DFB7B" bold="true" box="[549,598,1222,1243]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA669D798646DFB7B" box="[549,598,1222,1243]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C7B8ADBB730FF9EA6BED7996754FB56" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA6BED799653AFB7C" box="[754,769,1223,1244]" italics="true" pageId="4" pageNumber="198">E</emphasis>
. (
<emphasis id="B90F2D4AB730FF9EA489D7BF673AFB56" box="[197,257,1249,1270]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA55FD7BF6754FB56" box="[275,367,1249,1270]" italics="true" pageId="4" pageNumber="198">setitarsus</emphasis>
</taxonomicName>
-group [
<bibRefCitation id="EFEA8CA9B730FF9EA5F3D7BF64EBFB56" author="Daugeron C. &amp; Grootaert P." box="[447,720,1249,1270]" pageId="4" pageNumber="198" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
;
<bibRefCitation id="EFEA8CA9B730FF9EA692D7BF67A5FAB0" author="Sinclair B. J. &amp; Daugeron C." pageId="4" pageNumber="198" pagination="1221 - 1235" refId="ref6424" refString="Sinclair B. J., Daugeron C. 2017. Empididae (empidid dance flies or balloon flies) // A. H. Kirk-Spriggs, B. J. Sinclair (eds.). Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: SANBI Graphics &amp; Editing. P. 1221 - 1235." type="book chapter" year="2017">Sinclair, Daugeron, 2017</bibRefCitation>
]. Within the key to species of this group the new species would run to
<taxonomicName id="4C7B8ADBB730FF9EA0CBD262634DFEF1" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" box="[1159,1398,315,337]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="namaqua">
<emphasis id="B90F2D4AB730FF9EA0CBD26262ADFEF1" box="[1159,1174,316,337]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<emphasis id="B90F2D4AB730FF9EA0EED26262C7FEF1" box="[1186,1276,316,337]" italics="true" pageId="4" pageNumber="198">namaqua</emphasis>
<bibRefCitation id="EFEA8CA9B730FF9EA14DD265634DFEF1" author="Smith K. G. V." box="[1281,1398,315,337]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B8ADBB730FF9EA719D2096216FECC" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" box="[853,1069,342,364]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="corana">
<emphasis id="B90F2D4AB730FF9EA719D209655FFECC" box="[853,868,343,364]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<emphasis id="B90F2D4AB730FF9EA723D2096588FECC" box="[879,947,343,364]" italics="true" pageId="4" pageNumber="198">corana</emphasis>
<bibRefCitation id="EFEA8CA9B730FF9EA7F4D2086216FECC" author="Smith K. G. V." box="[952,1069,342,364]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
[
<bibRefCitation id="EFEA8CA9B730FF9EA074D2086379FECB" author="Daugeron C. &amp; Grootaert P." box="[1080,1346,342,363]" pageId="4" pageNumber="198" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
]. The new species differs from both these species primarily by pale setae on the laterotergite and abdomen [
<bibRefCitation id="EFEA8CA9B730FF9EA0FFD2D56309FE01" author="Smith K. G. V." box="[1203,1330,395,417]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED2F86220FE77" pageId="4" pageNumber="198" type="etymology">
<paragraph id="8BC4F158B730FF9EA71ED2F86220FE77" blockId="4.[812,1398,315,497]" pageId="4" pageNumber="198">ETYMOLOGY. The name of the new species refers to the country of its origin.</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED28562DAFE51" box="[850,1249,475,497]" pageId="4" pageNumber="198" type="distribution">
<paragraph id="8BC4F158B730FF9EA71ED28562DAFE51" blockId="4.[812,1398,315,497]" box="[850,1249,475,497]" pageId="4" pageNumber="198">
DISTRIBUTION. Afrotropics:
<collectingCountry id="F36CB1C8B730FF9EA0CBD28562E6FE51" box="[1159,1245,475,497]" name="Namibia" pageId="4" pageNumber="198">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:location id="CB0DD59BEE2A7ACE578860CDE133F152">
<mods:url id="77D51562254BC853F66EB66E1040F18C">http://dx.doi.org/10.15298/rusentj.31.3.02</mods:url>
</mods:location>
<mods:classification id="C6625F51748EC4C41F297721BB364F60">journal article</mods:classification>
<mods:identifier id="EA22EA8B8F83FB4719A34395904D5EEB" type="DOI">10.15298/rusentj.31.3.02</mods:identifier>
<mods:identifier id="FB0DC63D4C631207284021ECA6675938" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="3911B67682939BCA4FA57E25A416B397" type="Zenodo-Dep">13178120</mods:identifier>
</mods:mods>
<treatment id="03D2CB6BBF509950FCC1FCAAFDB6D282" LSID="urn:lsid:plazi:treatment:03D2CB6BBF509950FCC1FCAAFDB6D282" httpUri="http://treatment.plazi.org/id/03D2CB6BBF509950FCC1FCAAFDB6D282" lastPageId="6" lastPageNumber="224" pageId="4" pageNumber="222">
<subSubSection id="C36129F6BF509952FCC1FCAAFB73D59B" pageId="4" pageNumber="222" type="nomenclature">
<paragraph id="8BC47A7DBF509952FCC1FCAAFA97D579" blockId="4.[812,1399,864,1851]" box="[854,1356,864,888]" pageId="4" pageNumber="222">
<heading id="D08CCD11BF509952FCC1FCAAFA97D579" box="[854,1356,864,888]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="222" reason="8">
<taxonomicName id="4C7B01FEBF509952FCC1FCAAFA97D579" authorityName="Gorochov" authorityYear="2022" box="[854,1356,864,888]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="carinata" status="sp. n." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF509952FCC1FCAAFA97D579" box="[854,1356,864,888]" italics="true" pageId="4" pageNumber="222">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC47A7DBF509952FC6DFCB7FB73D59B" blockId="4.[812,1399,864,1851]" box="[1018,1192,893,917]" pageId="4" pageNumber="222">
Gorochov,
<emphasis id="B90FA66FBF509952FBE4FCB7FB73D59B" bold="true" box="[1139,1192,893,917]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FBE4FCB7FB73D59B" box="[1139,1192,893,917]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509952FB85FC50FB49D5BC" box="[1042,1170,922,946]" pageId="4" pageNumber="222" type="description">
<paragraph id="8BC47A7DBF509952FB85FC50FB49D5BC" blockId="4.[812,1399,864,1851]" box="[1042,1170,922,946]" pageId="4" pageNumber="222">
<figureCitation id="134066F8BF509952FB85FC50FB56D5BC" box="[1042,1165,922,946]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 28–37</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509952FCC5FC75FB92D214" pageId="4" pageNumber="222" type="materials_examined">
<paragraph id="8BC47A7DBF509952FCC5FC75FB92D214" blockId="4.[812,1399,864,1851]" pageId="4" pageNumber="222">
<materialsCitation id="3B137020BF509952FCC5FC75FB92D214" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="4" pageNumber="222" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
MATERIAL EXAMINED.
<typeStatus id="54C0C4DFBF509952FBD4FC75FB48D5DC" box="[1091,1171,959,978]" pageId="4" pageNumber="222" type="holotype">Holotype</typeStatus>
—
<specimenCount id="9D7DB1F4BF509952FB24FC77FB1CD5DD" box="[1203,1223,957,979]" count="1" pageId="4" pageNumber="222" type="male">♂</specimenCount>
,
<emphasis id="B90FA66FBF509952FB45FC75FA98D5DC" bold="true" box="[1234,1347,959,978]" pageId="4" pageNumber="222">
<collectingCountry id="F36C3AEDBF509952FB45FC75FA98D5DC" box="[1234,1347,959,978]" name="Madagascar" pageId="4" pageNumber="222">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF509952FADAFC0AFC7DD5E5" country="Madagascar" name="Toamasina" pageId="4" pageNumber="222">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF509952FC23FC12FB87D5E4" box="[948,1116,983,1003]" pageId="4" pageNumber="222">Moramanga Distr.</collectingMunicipality>
,
<location id="8EA42CA6BF509952FBFEFC1DFAACD5E4" LSID="urn:lsid:plazi:treatment:03D2CB6BBF509950FCC1FCAAFDB6D282:8EA42CA6BF509952FBFEFC1DFAACD5E4" box="[1129,1399,983,1003]" country="Madagascar" municipality="Moramanga Distr." name="Analamazaotra Forest Station" pageId="4" pageNumber="222" stateProvince="Toamasina">Analamazaotra Forest Station</location>
near Andasibe Vill., ~
<quantity id="4C83D798BF509952FC78FC25FBF1D20D" box="[1007,1066,1007,1027]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="222" unit="m" value="900.0">
<elevation id="00569D4EBF509952FC78FC25FBF1D20D" box="[1007,1066,1007,1027]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="222" unit="m" value="900.0">900 m</elevation>
</quantity>
, on leaf of bush in forest at night,
<date id="FFC55CBDBF509952FAF6FC25FC4BD214" pageId="4" pageNumber="222" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF509952FAF6FC25FC4BD214" pageId="4" pageNumber="222" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">8– 20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF509952FC0EFBCDFBD1D214" box="[921,1034,1031,1050]" pageId="4" pageNumber="222">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF509952FB81FBCDFBE5D214" box="[1046,1086,1031,1050]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="222" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509950FCC5FBD5FCDCD232" lastPageId="6" lastPageNumber="224" pageId="4" pageNumber="222" type="description">
<paragraph id="8BC47A7DBF509952FCC5FBD5FAACD135" blockId="4.[812,1399,864,1851]" pageId="4" pageNumber="222">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF509952FBACFBD5FB4ED23B" box="[1083,1173,1055,1077]" pageId="4" pageNumber="222" type="holotype">holotype</typeStatus>
). Body coloration light greyish with following marks: eye greyish with three longitudinal darkish stripes on upper half and one yellowish stripe between two lower of these darkish stripes as well as numerous dense and arcuate darkish lines running along anterior and ventral edges of eye; epicranial dorsum also darkish (almost greyish brown) with numerous lighter and less numerous darker dots; genae with a few oblique darkish stripes; mouthparts with almost rose three distal segments of each maxillary palpus and a pair of small brown spots in dorsolateral corners of clypeus; antenna with dark ventral and lateral longitudinal stripes on scape, with one dark dot and barely darkish marks around it on dorsal surface of scape, and with rather numerous small darkish and whitish spots on flagellum (
<figureCitation id="134066F8BF509952FCA4FA5FFC75D3A4" box="[819,942,1429,1451]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 28, 31</figureCitation>
); pronotum with brown median quadrate on anterior half of disc, numerous light brown to greyish brown dots on rest parts of disc, and sparser brown dots on lateral lobes (
<figureCitation id="134066F8BF509952FCFCFA2FFC01D3F4" box="[875,986,1509,1531]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 31, 33</figureCitation>
); legs with rather numerous and small light brown to greyish brown dots and small spots on all femora and tibiae (but hind femur lighter and with these dots less distinct, except for 4–5 dark dots distinctly visible along median line of outer femoral surface;
<figureCitation id="134066F8BF509952FB33F985FACDD06A" box="[1188,1302,1615,1637]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 29–33</figureCitation>
); tegmina yellowish grey with light brown basal area (this area having whitish reticular pattern), semitransparent membranes, whitish some areas and all crossveins in medial and humeral regions as well as in lateral field, brown to brownish rest venation and numerous small marks on almost all membranes, and brown to light brown larger areas on membranes of proximal part of lateral field (
<figureCitation id="134066F8BF509952FBF2F8C0FB0DD111" box="[1125,1238,1802,1824]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 32–33</figureCitation>
); visible parts of hind wings barely darkened but with lighter venation (Fig.
</paragraph>
<paragraph id="8BC47A7DBF509952FF2CF89FFF12D164" blockId="4.[187,1398,1877,2064]" box="[187,201,1877,1898]" pageId="4" pageNumber="222">&lt;</paragraph>
<paragraph id="8BC47A7DBF509952FF75F8A7FD12DE1E" blockId="4.[187,1398,1877,2064]" pageId="4" pageNumber="222">
Рис. 10–27. Cтруктуры брюШка, самец: 10–15 —
<taxonomicName id="4C7B01FEBF509952FD3AF8A7FC7BD18E" authorityName="Gorochov" authorityYear="2022" box="[685,928,1901,1920]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="longitubulosa" status="sp. nov.">
<emphasis id="B90FA66FBF509952FD3AF8A7FC7BD18E" box="[685,928,1901,1920]" italics="true" pageId="4" pageNumber="222">Asymmetrella longitubulosa</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF509952FC31F8A7FC0AD18E" bold="true" box="[934,977,1901,1920]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FC31F8A7FC0AD18E" box="[934,977,1901,1920]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 16 —
<taxonomicName id="4C7B01FEBF509952FB81F8A7FF3CD196" authority="Gor." authorityName="Gor." class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="bella" subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FB81F8A7FAADD18E" box="[1046,1398,1901,1920]" italics="true" pageId="4" pageNumber="222">Zvenellomorpha (Zvenellomorpha) bella</emphasis>
Gor.
</taxonomicName>
; 17–18 —
<taxonomicName id="4C7B01FEBF509952FED0F84CFE3AD196" authority="Gor." authorityName="Gor." box="[327,481,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="recta" subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FED0F84CFE6BD196" box="[327,432,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.) recta</emphasis>
Gor.
</taxonomicName>
; 19–23 —
<taxonomicName id="4C7B01FEBF509952FDD6F84CFD00D196" authorityName="Gorochov" authorityYear="2022" box="[577,731,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="intermedia" status="sp.n." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FDD6F84CFD00D196" box="[577,731,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF509952FD75F84FFCD6D196" bold="true" box="[738,781,1925,1944]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FD75F84FFCD6D196" box="[738,781,1925,1944]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 24 —
<emphasis id="B90FA66FBF509952FCC7F84FFCBFD196" box="[848,868,1925,1944]" italics="true" pageId="4" pageNumber="222">A.</emphasis>
sp.; 25 —
<emphasis id="B90FA66FBF509952FC52F84CFC27D196" box="[965,1020,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.)</emphasis>
sp.; 26 —
<emphasis id="B90FA66FBF509952FBC9F84CFAC1D196" box="[1118,1306,1925,1944]" italics="true" pageId="4" pageNumber="222">
Z. (
<taxonomicName id="4C7B01FEBF509952FBECF84FFACFD196" authorityName="Gorochov" authorityYear="2022" box="[1147,1300,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Azvenellomorpha">Azvenellomorpha</taxonomicName>
)
</emphasis>
sp.; 27 —
<taxonomicName id="4C7B01FEBF509952FF2BF857FEE7D1BE" box="[188,316,1949,1968]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90FA66FBF509952FF2BF857FEC1D1BE" box="[188,282,1949,1968]" italics="true" pageId="4" pageNumber="222">Ultratrella</emphasis>
sp.
</taxonomicName>
; 10 — верШина брюШка, сверху; 11–13, 16–17, 19–21 — гениталии (11, 20 — сверху; 12, 21 — сниЗу; 13, 16–17, 19 — сбоку); 14 — сперматофор сбоку, реконструкциЯ; 15 — формула с рахисом, Эндопарамером и Эктопарамером, сниЗу; 18, 22 — формула, сниЗу; 23 — дистальнаЯ часть гениталий, сбоку; 24–27 — схема сагиттального сечениЯ формулы [коричневый цвет], её склеротиЗованной передней ленты [Зелёный цвет] и ближайШего участка сперматофорной трубы [красный цвет]; 10, 23 — паратип; 11–22 — голотипы [16, 17 — по:
<bibRefCitation id="EFEA078CBF509952FE6FF837FD53DE1E" author="Gorochov A. V." box="[504,648,2045,2064]" pageId="4" pageNumber="222" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
,
<bibRefCitation id="EFEA078CBF509952FD05F837FD19DE1E" author="Gorochov A. V." box="[658,706,2045,2064]" pageId="4" pageNumber="222" pagination="33 - 46" refId="ref8770" refString="Gorochov A. V. 2006. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 5: new Indo-Malayan and Madagascan Podoscirtini // Zoosystematica rossica. Vol. 15. No. 1. P. 33 - 46." type="journal article" year="2006">2006</bibRefCitation>
.]
</paragraph>
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<paragraph id="8BC47A7DBF519953FF75F8A3FB06D1A2" blockId="5.[188,1399,1897,2065]" pageId="5" pageNumber="223">
Figs 28–37.
<taxonomicName id="4C7B01FEBF519953FEDAF8A3FD07D172" authorityName="Gorochov" authorityYear="2022" box="[333,732,1897,1916]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="223" phylum="Arthropoda" rank="species" species="carinata" status="sp. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF519953FEDAF8A3FD07D172" box="[333,732,1897,1916]" italics="true" pageId="5" pageNumber="223">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF519953FD76F8A3FCD7D172" bold="true" box="[737,780,1897,1916]" pageId="5" pageNumber="223">
<taxonomicNameLabel id="A23C1B14BF519953FD76F8A3FCD7D172" box="[737,780,1897,1916]" pageId="5" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction.
</paragraph>
<paragraph id="8BC47A7DBF519953FF75F87CFE96DE1F" blockId="5.[188,1399,1897,2065]" pageId="5" pageNumber="223">
Рис. 28–37.
<taxonomicName id="4C7B01FEBF519953FECDF87FFD23D1C6" authorityName="Gorochov" authorityYear="2022" box="[346,760,1973,1992]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="223" phylum="Arthropoda" rank="species" species="carinata" status="sp. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF519953FECDF87FFD23D1C6" box="[346,760,1973,1992]" italics="true" pageId="5" pageNumber="223">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF519953FC94F87FFCF5D1C6" bold="true" box="[771,814,1973,1992]" pageId="5" pageNumber="223">
<taxonomicNameLabel id="A23C1B14BF519953FC94F87FFCF5D1C6" box="[771,814,1973,1992]" pageId="5" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF529950FF2AFEF1FEE9D595" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
32); venter of body with wide longitudinal greyish brown band on abdominal sternites, a pair of barely darkish areas on proximal half of genital plate, and numerous very small darkish dots on cerci (
<figureCitation id="134066F8BF529950FE1AFE41FE08D7AE" box="[397,467,395,417]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Fig. 33</figureCitation>
). Eyes slightly higher than long; rostrum roundly angular in profile but with low and thin transverse keel at apex; maxillary palpi approximately as in
<taxonomicName id="4C7B01FEBF529950FF2AFE11FE99D7FE" authorityName="Gorochov" authorityYear="2022" box="[189,322,475,496]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FF2AFE11FE99D7FE" box="[189,322,475,496]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FEDEFE11FE4BD7FE" bold="true" box="[329,400,475,496]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FEDEFE11FE4BD7FE" box="[329,400,475,496]" pageId="6" pageNumber="224" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
; pronotum also similar to that of this genus but more strongly narrowing to head and with lateral keels slightly shorter and more sinuate in profile (
<figureCitation id="134066F8BF529950FD02FDDBFF03D44E" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 28, 31, 33</figureCitation>
); tegmina significantly protruding beyond abdominal apex and hind femora, with venation as in
<figureCitation id="134066F8BF529950FDB4FD8CFD49D455" box="[547,658,582,604]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 32–33</figureCitation>
; hind wings also significantly protruding beyond tegmina (
<figureCitation id="134066F8BF529950FD14FDABFD17D478" box="[643,716,609,631]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Fig. 32</figureCitation>
); fore and middle legs rather stout, with a pair of dorsal longitudinal keels on fore tibia (
<figureCitation id="134066F8BF529950FE17FD5CFE2BD4A5" box="[384,496,662,684]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 31–33</figureCitation>
) having tympana as in
<figureCitation id="134066F8BF529950FD4AFD5CFF22D4C8" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 29–30</figureCitation>
(inner tympanum somewhat larger than outer one), with two keels on middle tibia (along dorsal and ventral tibial edges;
<figureCitation id="134066F8BF529950FE95FD2CFEAFD4F5" box="[258,372,742,764]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 32–33</figureCitation>
); hind leg more elongate and slender but also with more or less distinct dorsal keel on femur and a pair of dorsal longitudinal keels on tibia; spines and denticles of this tibia more or less similar to those of
<taxonomicName id="4C7B01FEBF529950FDC7FCFCFD0ED545" authorityName="Gorochov" authorityYear="2022" box="[592,725,822,843]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FDC7FCFCFD0ED545" box="[592,725,822,843]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FD4BFCFCFF0BD568" bold="true" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FD4BFCFCFF0BD568" pageId="6" pageNumber="224" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
; genital plate elongately triangular and with almost spine-like but soft apical lobule; genitalia and spermatophore as in
<figureCitation id="134066F8BF529950FF2AFC4CFEF5D595" box="[189,302,902,924]" captionStart="Figs 28–37" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 29–30 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34– 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 28–37. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 29–30 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 34–36 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 34–37</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC6BFE49D5B9" blockId="6.[189,775,315,1164]" box="[226,402,929,951]" pageId="6" pageNumber="224">Female unknown.</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC71FD8AD5E5" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">Length in mm. Body 20.3; body with wings 29.5; pronotum 3.6; tegmina 21.5; hind femora 13.4.</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC3BFCDCD232" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
COMPARISON. The new species differs from all the other congeners in the same characters as the subgenus
<taxonomicName id="4C7B01FEBF529950FF2AFBECFEA9D235" authorityName="Gorochov" authorityYear="2022" box="[189,370,1062,1083]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF529950FF2AFBECFEA9D235" box="[189,370,1062,1083]" italics="true" pageId="6" pageNumber="224">Euzvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FEE1FBECFE03D235" bold="true" box="[374,472,1062,1083]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FEE1FBECFE03D235" box="[374,472,1062,1083]" pageId="6" pageNumber="224" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
(see the subgeneric key above).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FB8BFDB6D282" pageId="6" pageNumber="224" type="etymology">
<paragraph id="8BC47A7DBF529950FF75FB8BFDB6D282" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF529950FF2AFB91FECDD27F" authorityName="Gorochov" authorityYear="2022" box="[189,278,1115,1137]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="carinata">“carinata</taxonomicName>
” (carinate, with keels) due to the presence of characteristic keels on the lateral pronotal lobes.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:start id="EA4264EEBC357B83B6E648439E5B31CF">218</mods:start>
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<mods:location id="591C4801B8CDE6E3BE453EBA56E46F93">
<mods:url id="FE99E4CF88D6C1D94CE043612F4EF88D">http://dx.doi.org/10.15298/rusentj.31.3.02</mods:url>
</mods:location>
<mods:classification id="A93F4C426245D56D55300202C6AD8D78">journal article</mods:classification>
<mods:identifier id="82DAA66D3C365C013980A626BEC42912" type="DOI">10.15298/rusentj.31.3.02</mods:identifier>
<mods:identifier id="AB8FB7DB345DE3D9EAF402D73660D6BF" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="15FB87ABDE22AE06966754A3C79D3CC6" type="Zenodo-Dep">13178120</mods:identifier>
</mods:mods>
<treatment id="03D2CB6BBF529950FF71FB6AFBFBD102" ID-DOI="http://doi.org/10.5281/zenodo.13178264" ID-Zenodo-Dep="13178264" LSID="urn:lsid:plazi:treatment:03D2CB6BBF529950FF71FB6AFBFBD102" httpUri="http://treatment.plazi.org/id/03D2CB6BBF529950FF71FB6AFBFBD102" lastPageNumber="224" pageId="6" pageNumber="224">
<subSubSection id="C36129F6BF529950FF71FB6AFDE2D2DB" pageId="6" pageNumber="224" type="nomenclature">
<paragraph id="8BC47A7DBF529950FF71FB6AFD06D2B6" blockId="6.[189,775,1184,2056]" box="[230,733,1184,1208]" pageId="6" pageNumber="224">
<heading id="D08CCD11BF529950FF71FB6AFD06D2B6" box="[230,733,1184,1208]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="224" reason="8">
<taxonomicName id="4C7B01FEBF529950FF71FB6AFD06D2B6" authorityName="Gorochov" authorityYear="2022" box="[230,733,1184,1208]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="intermedia" status="sp. n." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF529950FF71FB6AFD06D2B6" box="[230,733,1184,1208]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC47A7DBF529950FE1DFB77FDE2D2DB" blockId="6.[189,775,1184,2056]" box="[394,569,1213,1237]" pageId="6" pageNumber="224">
Gorochov,
<emphasis id="B90FA66FBF529950FD94FB77FDE2D2DB" bold="true" box="[515,569,1213,1237]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FD94FB77FDE2D2DB" box="[515,569,1213,1237]" pageId="6" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FEEEFB10FD90D2FC" box="[377,587,1242,1266]" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FEEEFB10FD90D2FC" blockId="6.[189,775,1184,2056]" box="[377,587,1242,1266]" pageId="6" pageNumber="224">
<figureCitation id="134066F8BF529950FEEEFB10FE2ED2FC" box="[377,501,1242,1266]" captionStart="Figs 10–27" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 10–27. Structures of abdomen, male: 10–15 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 17–18 — Z. (Z.) recta Gor.; 19–23 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 11–13, 16–17, 19–21 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16– 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 24–27 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 11–22 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Figs 19–23</figureCitation>
,
<figureCitation id="134066F8BF529950FD96FB10FD9DD2FC" box="[513,582,1242,1266]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">38–46</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FB35FE70D3AC" pageId="6" pageNumber="224" type="materials_examined">
<paragraph id="8BC47A7DBF529950FF75FB35FE70D3AC" blockId="6.[189,775,1184,2056]" pageId="6" pageNumber="224">
MATERIAL EXAMINED.
<materialsCitation id="3B137020BF529950FE44FB35FEB3D354" collectingDate="2013-02-22" collectingDateMax="2013-03-11" collectingDateMin="2013-02-22" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="1000" location="Torotorofotsy Forest Reserve" municipality="Moramanga Distr." pageId="6" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54C0C4DFBF529950FE44FB35FDF8D31C" box="[467,547,1279,1298]" pageId="6" pageNumber="224" type="holotype">Holotype</typeStatus>
—
<specimenCount id="9D7DB1F4BF529950FDD4FB37FD8CD31D" box="[579,599,1277,1299]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
,
<emphasis id="B90FA66FBF529950FDF5FB35FD08D31C" bold="true" box="[610,723,1279,1298]" pageId="6" pageNumber="224">
<collectingCountry id="F36C3AEDBF529950FDF5FB35FD08D31C" box="[610,723,1279,1298]" name="Madagascar" pageId="6" pageNumber="224">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF529950FD49FACAFEE8D325" country="Madagascar" name="Toamasina" pageId="6" pageNumber="224">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF529950FEA9FAD2FE39D324" box="[318,482,1303,1323]" pageId="6" pageNumber="224">Moramanga Distr.</collectingMunicipality>
, ~
<locationDeviation id="9A76635FBF529950FE6FFADDFCDFD324" box="[504,772,1303,1323]" location="Andasibe Vill." pageId="6" pageNumber="224">
<quantity id="4C83D798BF529950FE6FFADDFDEAD324" box="[504,561,1303,1322]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="km" value="10.0">10 km</quantity>
NW of Andasibe Vill.
</locationDeviation>
,
<location id="8EA42CA6BF529950FF2AFAE5FE64D34D" LSID="urn:lsid:plazi:treatment:03D2CB6BBF529950FF71FB6AFBFBD102:8EA42CA6BF529950FF2AFAE5FE64D34D" box="[189,447,1327,1347]" country="Madagascar" municipality="Moramanga Distr." name="Torotorofotsy Forest Reserve" pageId="6" pageNumber="224" stateProvince="Toamasina">Torotorofotsy Forest Reserve</location>
, ~
<quantity id="4C83D798BF529950FE44FAE5FDC2D34D" box="[467,537,1327,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="m" value="1000.0">
<elevation id="00569D4EBF529950FE44FAE5FDC2D34D" box="[467,537,1327,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FDFBFAE5FCD8D34C" box="[620,771,1327,1346]" pageId="6" pageNumber="224" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">
<collectingDate id="EF81A555BF529950FDFBFAE5FCD8D34C" box="[620,771,1327,1346]" pageId="6" pageNumber="224" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">22.II–11.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FF2AFA8DFEF6D354" box="[189,301,1351,1370]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FEADFA8DFEB9D354" box="[314,354,1351,1370]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B137020BF529950FEE6FA82FF2AD37C" collectingDate="2013-02-22" collectingDateMax="2013-03-11" collectingDateMin="2013-02-22" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="1000" location="Torotorofotsy Forest Reserve" municipality="Moramanga Distr." pageId="6" pageNumber="224" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="54C0C4DFBF529950FEE6FA82FE13D355" box="[369,456,1352,1371]" pageId="6" pageNumber="224" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7DB1F4BF529950FE46FA8DFE2ED355" box="[465,501,1349,1371]" count="1" pageId="6" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7DB1F4BF529950FD96FA8DFDC6D355" box="[513,541,1349,1371]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same data as for holotype (
<collectionCode id="ED6AE2B8BF529950FF54FA95FF30D37C" box="[195,235,1375,1394]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B137020BF529950FF6BFA95FE7DD3AC" collectingDate="2013-03-01" collectingDateMax="2013-03-07" collectingDateMin="2013-03-01" collectionCode="ZIN" collectorName="A. Gorochov &amp; L. Anisyutkin" country="Madagascar" elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="2" specimenCount-female="2" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="9D7DB1F4BF529950FF6BFA95FEC0D37D" box="[252,283,1373,1395]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same province and district,
<collectingMunicipality id="6BA0E007BF529950FD8AFA95FF04D384" pageId="6" pageNumber="224">Analamazaotra Forest Station</collectingMunicipality>
near
<location id="8EA42CA6BF529950FE84FABDFE49D384" LSID="urn:lsid:plazi:treatment:03D2CB6BBF529950FF71FB6AFBFBD102:8EA42CA6BF529950FE84FABDFE49D384" box="[275,402,1399,1418]" country="Madagascar" municipality="Analamazaotra Forest Station" name="Andasibe Vill" pageId="6" pageNumber="224" stateProvince="Toamasina">Andasibe Vill</location>
., ~
<quantity id="4C83D798BF529950FE3CFABDFE3CD385" box="[427,487,1399,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">
<elevation id="00569D4EBF529950FE3CFABDFE3CD385" box="[427,487,1399,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FDAAFABDFD70D384" box="[573,683,1399,1418]" pageId="6" pageNumber="224" value="2013-03-01" valueMax="2013-03-07" valueMin="2013-03-01">
<collectingDate id="EF81A555BF529950FDAAFABDFD70D384" box="[573,683,1399,1418]" pageId="6" pageNumber="224" value="2013-03-01" valueMax="2013-03-07" valueMin="2013-03-01">1–7.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FD21FABDFF32D3AC" pageId="6" pageNumber="224">A. Gorochov</collectorName>
,
<collectorName id="268E1FABBF529950FF65FA5AFEB0D3AC" box="[242,363,1423,1443]" pageId="6" pageNumber="224">L. Anisyutkin</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FEEFFA45FE7AD3AC" box="[376,417,1423,1442]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FA6DFACCD0AF" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FF75FA6DFA93D595" blockId="6.[189,775,1184,2056]" lastBlockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF529950FE44FA6DFDF6D3B3" box="[467,557,1447,1469]" pageId="6" pageNumber="224" type="holotype">holotype</typeStatus>
). General appearance similar to that of two other species of this subgenus but somewhat intermediate between them. Body coloration yellowish with greyish tinge and following marks: upper half of epicranium, pronotum and dorsal tegminal field light brown with light greyish eyes, a pair of barely distinct yellowish interrupted stripes along both lateral edges of pronotal disc, brown M, yellowish stripe between M and Cu in humeral region of tegmen, light greyish brown diagonal vein, and almost semitransparent membranes of mirror and of some nearest cells (
<figureCitation id="134066F8BF529950FED5F978FE6FD0C9" box="[322,436,1714,1736]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 38–40</figureCitation>
); antenna with brown basal part of scape as well as middle and dorsal portions of flagellum, dark brown rest of scape and proximal portion of flagellum (
<figureCitation id="134066F8BF529950FD4BF92DFF27D119" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 38–39</figureCitation>
); lateral tegminal field yellowish with light brown venation and semitransparent membranes between veins and veinlets (
<figureCitation id="134066F8BF529950FE80F8FDFEBAD142" box="[279,353,1847,1869]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
). Head with oval (longitudinal) but poorly distinct lateral ocelli, small round and also poorly distinct median ocellus, and large barely elongate eyes; rostrum roundly angular in profile and rather narrow (scape approximately 1.7 times as wide as space between antennal cavities); maxillary palpi more or less similar to those of
<taxonomicName id="4C7B01FEBF529950FD15F877FCDCD1DC" authorityName="Gorochov" authorityYear="2022" box="[642,775,1981,2002]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FD15F877FCDCD1DC" box="[642,775,1981,2002]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FF2AF81DFF21D1E2" bold="true" box="[189,250,2007,2028]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FF2AF81DFF21D1E2" box="[189,250,2007,2028]" pageId="6" pageNumber="224" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
; pronotum slightly narrowing to head, insignificantly longer than wide, and almost not depressed dorsoventrally (but lateral lobes moderately low;
<figureCitation id="134066F8BF529950FBE4FEF1FB62D75E" box="[1139,1209,315,337]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
); legs without keels on femora and tibiae, with inner tympanum oval (almost as wide as half of fore tibia width and about 1.2 times as long as latter width; as in
<figureCitation id="134066F8BF529950FC72FE41FBF4D7AE" box="[997,1071,395,417]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 42</figureCitation>
), and outer tympanum similar in shape but slightly smaller;
<figureCitation id="134066F8BF529950FBAEFE6CFB59D7B5" box="[1081,1154,422,444]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
); tegmina distinctly protruding beyond apices of hind femora, with mirror in dorsal field almost normal (less irregular than in
<taxonomicName id="4C7B01FEBF529950FB50FE16FACBD7FE" box="[1223,1296,475,497]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FB50FE16FACBD7FE" box="[1223,1296,475,497]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
), dividing vein in this mirror more curved than in
<taxonomicName id="4C7B01FEBF529950FB33FE3DFB34D402" box="[1188,1263,503,524]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB33FE3DFB34D402" box="[1188,1263,503,524]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
(
<figureCitation id="134066F8BF529950FB6CFE3CFA98D405" box="[1275,1347,502,524]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 40</figureCitation>
), and Sc having 9–10 branches; hind wings strongly protruding beyond tegminal apices; genital plate triangular in proximal half but with narrow, long and soft posteromedian lobule in distal half (this lobule narrowly rounded at apex). Genitalia very similar to those of
<taxonomicName id="4C7B01FEBF529950FBA3FDB6FB5ED49E" box="[1076,1157,635,657]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FBA3FDB6FB5ED49E" box="[1076,1157,635,657]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B01FEBF529950FB56FDB6FAC8D49F" box="[1217,1299,636,657]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB56FDB6FAC8D49F" box="[1217,1299,636,657]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
but with following diferences: epiphallus somewhat lower than in these species; dorsoapical epiphallic process with slightly longer both (proximal and distal) spines than in
<taxonomicName id="4C7B01FEBF529950FA99FD06FA82D4EE" box="[1294,1369,715,737]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FA99FD06FA82D4EE" box="[1294,1369,715,737]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
as well as with distinctly deeper apical notch than in
<taxonomicName id="4C7B01FEBF529950FAB5FD2DFAABD4F2" box="[1314,1392,743,764]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FAB5FD2DFAABD4F2" box="[1314,1392,743,764]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
; ventroapical epiphallic lobule slightly shorter and less arcuate (almost S-shaped) than in
<taxonomicName id="4C7B01FEBF529950FBD1FCD6FB55D53E" box="[1094,1166,795,817]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FBD1FCD6FB55D53E" box="[1094,1166,795,817]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
as well as apically more widened and hooked than in
<taxonomicName id="4C7B01FEBF529950FBD3FCFDFB54D542" box="[1092,1167,823,844]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FBD3FCFDFB54D542" box="[1092,1167,823,844]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
; ectoparameres slightly protruding beyond epiphallus (in
<taxonomicName id="4C7B01FEBF529950FB1CFC9BFB06D568" box="[1163,1245,849,870]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB1CFC9BFB06D568" box="[1163,1245,849,870]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
, they strongly protruding beyond epiphallus, and in
<taxonomicName id="4C7B01FEBF529950FB04FCA6FB00D58E" box="[1171,1243,875,897]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FB04FCA6FB00D58E" box="[1171,1243,875,897]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
, they not reaching epiphallic apices; compare
<figureCitation id="134066F8BF529950FBC9FC4CFAE7D595" box="[1118,1340,902,924]" captionStart="Figs 10–27" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 10–27. Structures of abdomen, male: 10–15 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 17–18 — Z. (Z.) recta Gor.; 19–23 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 11–13, 16–17, 19–21 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16– 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 24–27 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 11–22 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Figs 16–18 and 19–22</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FC6BFB96D258" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Variations. Second male with epicranial dorsum between eyes laterally darker (greyish brown), pronotal disc having brown median band, dividing vein of tegminal mirror clearly S-shaped (
<figureCitation id="134066F8BF529950FC02FC3BFBD2D208" box="[917,1033,1009,1031]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 41–42</figureCitation>
), and genitalia having barely longer proximal spine of dorsoapical epiphallic process as well as slightly narrower (lower) distal part of ventroapical epiphallic lobule in profile (
<figureCitation id="134066F8BF529950FC6FFB8BFB9AD258" box="[1016,1089,1089,1111]" captionStart="Figs 10–27" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 10–27. Structures of abdomen, male: 10–15 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 17–18 — Z. (Z.) recta Gor.; 19–23 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 11–13, 16–17, 19–21 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16– 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 24–27 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 11–22 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FB91FB9FD3E8" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Female. Coloration and structure of body as in male, but scape and proximal portion of antennal flagellum somewhat lighter (more or less brown), pronotal disc and dorsal tegminal field sometimes almost yellowish with greyish tinge, posterior part of this disc sometimes completely brown (
<figureCitation id="134066F8BF529950FAD8FB0CFC9CD2F8" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 43</figureCitation>
), tegminal R brown or greyish brown, median ocellus very small and round or obliterate, tegmina with 8–10 straight but somewhat obliquely situated longitudinal veins and moderately sparse crossveins in dorsal field as well as with 7–9 branches of Sc and almost without distinct crossveins in lateral field, and genital plate barely elongate as well as distally narrowing to roundly and rather widely bilobate apex (posteromedian notch of this apex short and almost angular;
<figureCitation id="134066F8BF529950FCBBFA7CFCA8D3C5" box="[812,883,1462,1484]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 44</figureCitation>
). Ovipositor normally developed, with distinctly drilling apical part (
<figureCitation id="134066F8BF529950FC50FA1BFBE1D3E8" box="[967,1082,1488,1511]" captionStart="Figs 38–46" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female. Рис. 38–46. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 45–46</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FA21FBE9D05F" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Length in mm. Body:
<specimenCount id="9D7DB1F4BF529950FBA4FA22FB92D00E" box="[1075,1097,1512,1536]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
13–14.5,
<specimenCount id="9D7DB1F4BF529950FB27FA22FB66D00E" box="[1200,1213,1512,1536]" count="1" pageId="6" pageNumber="224" type="female">♀</specimenCount>
11–14; body with wings:
<specimenCount id="9D7DB1F4BF529950FCE4F9C9FC52D015" box="[883,905,1539,1563]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
20–22,
<specimenCount id="9D7DB1F4BF529950FC4EF9C9FC3DD015" box="[985,998,1539,1563]" count="1" pageId="6" pageNumber="224" type="female">♀</specimenCount>
18–21; pronotum:
<specimenCount id="9D7DB1F4BF529950FB36F9C9FB6CD015" box="[1185,1207,1539,1563]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
2.1–2.3,
<specimenCount id="9D7DB1F4BF529950FA84F9C9FAFBD015" box="[1299,1312,1539,1563]" count="1" pageId="6" pageNumber="224" type="female">♀</specimenCount>
2.4–2.8; tegmina:
<specimenCount id="9D7DB1F4BF529950FC10F9D4FC46D038" box="[903,925,1566,1590]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
14–15.5;
<specimenCount id="9D7DB1F4BF529950FC68F9D4FBD7D038" box="[1023,1036,1566,1590]" count="1" pageId="6" pageNumber="224" type="female">♀</specimenCount>
13–15; hind femora:
<specimenCount id="9D7DB1F4BF529950FB48F9D4FB2ED038" box="[1247,1269,1566,1590]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
10.8–11.5,
<specimenCount id="9D7DB1F4BF529950FAFEF9D4FAADD038" box="[1385,1398,1566,1590]" count="1" pageId="6" pageNumber="224" type="female">♀</specimenCount>
10–11.5; ovipositor 7–8.5.
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5F99CFACCD0AF" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
COMPARISON. The new species differs from the two other species of the subgenus
<taxonomicName id="4C7B01FEBF529950FBFDF9BBFAD3D088" authorityName="Gorochov" authorityYear="2022" box="[1130,1288,1649,1670]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF529950FBFDF9BBFAD3D088" box="[1130,1288,1649,1670]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FA83F9BBFAADD088" bold="true" box="[1300,1398,1649,1670]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FA83F9BBFAADD088" box="[1300,1398,1649,1670]" pageId="6" pageNumber="224" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
mainly in the male genital characters listed above.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FCC5F96CFBFBD102" pageId="6" pageNumber="224" type="etymology">
<paragraph id="8BC47A7DBF529950FCC5F96CFBFBD102" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF529950FCBBF90BFC7AD0D9" authorityName="Gorochov" authorityYear="2022" box="[812,929,1729,1751]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="intermedia">“intermedia</taxonomicName>
” (intermediate) due to the intermediate length of ectoparameres in the new species compared to both other species of this subgenus.
</paragraph>
</subSubSection>
</treatment>
</document>

289
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<mods:title id="D113DE2B23CE84BF259B7C9F7F85CF00">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<paragraph id="8BC47A7DBF529950FCCCF8EAFB73D15B" blockId="6.[812,1399,1824,2059]" pageId="6" pageNumber="224">
<heading id="D08CCD11BF529950FCCCF8EAFB73D15B" centered="true" fontSize="10" level="2" pageId="6" pageNumber="224" reason="8">
<taxonomicName id="4C7B01FEBF529950FCCCF8EAFBB3D15B" authority="Gorochov" authorityName="Gorochov" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF529950FCCCF8EAFA9CD139" box="[859,1351,1824,1848]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
Gorochov
</taxonomicName>
,
<emphasis id="B90FA66FBF529950FBE4F8F7FB73D15B" bold="true" box="[1139,1192,1853,1877]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FBE4F8F7FB73D15B" box="[1139,1192,1853,1877]" pageId="6" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FB85F890FB49D17C" box="[1042,1170,1882,1906]" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FB85F890FB49D17C" blockId="6.[812,1399,1824,2059]" box="[1042,1170,1882,1906]" pageId="6" pageNumber="224">
<figureCitation id="134066F8BF529950FB85F890FB55D17C" box="[1042,1166,1882,1906]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="6" pageNumber="224">Figs 47–59</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF52995FFCC5F8B5FF2ED768" lastPageId="9" lastPageNumber="227" pageId="6" pageNumber="224" type="materials_examined">
<paragraph id="8BC47A7DBF529950FCC5F8B5FAADDE04" blockId="6.[812,1399,1824,2059]" pageId="6" pageNumber="224">
MATERIAL EXAMINED.
<materialsCitation id="3B137020BF529950FBD4F8B5FCB9D1D4" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" county="Moramanga Distr." elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54C0C4DFBF529950FBD4F8B5FB48D19C" box="[1091,1171,1919,1938]" pageId="6" pageNumber="224" type="holotype">Holotype</typeStatus>
—
<specimenCount id="9D7DB1F4BF529950FB24F8B7FB1CD19D" box="[1203,1223,1917,1939]" count="1" pageId="6" pageNumber="224" type="male">♂</specimenCount>
,
<emphasis id="B90FA66FBF529950FB45F8B5FA98D19C" bold="true" box="[1234,1347,1919,1938]" pageId="6" pageNumber="224">
<collectingCountry id="F36C3AEDBF529950FB45F8B5FA98D19C" box="[1234,1347,1919,1938]" name="Madagascar" pageId="6" pageNumber="224">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF529950FADAF84AFC7DD1A5" country="Madagascar" name="Toamasina" pageId="6" pageNumber="224">Toamasina Prov.</collectingRegion>
,
<collectingCounty id="62A502F1BF529950FC23F852FB87D1A4" box="[948,1116,1943,1963]" pageId="6" pageNumber="224">Moramanga Distr.</collectingCounty>
,
<collectingMunicipality id="6BA0E007BF529950FBFEF85DFAACD1A4" box="[1129,1399,1943,1963]" pageId="6" pageNumber="224">Analamazaotra Forest Station</collectingMunicipality>
near
<location id="8EA42CA6BF529950FCCEF865FC0DD1CC" LSID="urn:lsid:plazi:treatment:03D2CB6BBF52995FFCCCF8EAFB83D53F:8EA42CA6BF529950FCCEF865FC0DD1CC" box="[857,982,1967,1986]" country="Madagascar" county="Moramanga Distr." municipality="Analamazaotra Forest Station" name="Andasibe Vill" pageId="6" pageNumber="224" stateProvince="Toamasina">Andasibe Vill</location>
. ~
<quantity id="4C83D798BF529950FC7BF865FBF3D1CD" box="[1004,1064,1967,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">
<elevation id="00569D4EBF529950FC7BF865FBF3D1CD" box="[1004,1064,1967,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FBE8F865FB23D1CC" box="[1151,1272,1967,1986]" pageId="6" pageNumber="224" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF529950FBE8F865FB23D1CC" box="[1151,1272,1967,1986]" pageId="6" pageNumber="224" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">8–20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FA94F865FAADD1CC" box="[1283,1398,1967,1986]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FCA4F80DFC87D1D4" box="[819,860,1991,2010]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B137020BF529950FCE7F802FB02D1FC" collectingDate="2013-02-11" collectingDateMax="2013-03-20" collectingDateMin="2013-02-11" collectionCode="ZIN" collectorName="A. Gorochov &amp; L. Anisyutkin" country="Madagascar" county="Moramanga Distr." elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="16" specimenCount-female="16" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="54C0C4DFBF529950FCE7F802FC12D1D5" box="[880,969,1992,2011]" pageId="6" pageNumber="224" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7DB1F4BF529950FC41F80DFBD8D1D5" box="[982,1027,1989,2011]" count="16" pageId="6" pageNumber="224" type="female">16 ♀</specimenCount>
, same data as for holotype, but
<date id="FFC55CBDBF529950FAD6F80DFC4BD1FC" pageId="6" pageNumber="224" value="2013-02-11" valueMax="2013-03-20" valueMin="2013-02-11">
<collectingDate id="EF81A555BF529950FAD6F80DFC4BD1FC" pageId="6" pageNumber="224" value="2013-02-11" valueMax="2013-03-20" valueMin="2013-02-11">11.II– 20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FC0BF815FBC8D1FC" box="[924,1043,2015,2034]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
,
<collectorName id="268E1FABBF529950FB88F82AFB40D1FC" box="[1055,1179,2015,2035]" pageId="6" pageNumber="224">L. Anisyutkin</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FB3DF815FB09D1FC" box="[1194,1234,2015,2034]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B137020BF529950FB71F815FAADDE04" country="Madagascar" location="Torotorofotsy" pageId="6" pageNumber="224" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="9D7DB1F4BF529950FB71F815FAD6D1FD" box="[1254,1293,2013,2035]" count="1" pageId="6" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7DB1F4BF529950FA8BF815FAE7D1FD" box="[1308,1340,2013,2035]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same province and district, ~
<locationDeviation id="9A76635FBF529950FC61F83DFB2DDE04" box="[1014,1270,2039,2059]" location="Andasibe Vill." pageId="6" pageNumber="224">
<quantity id="4C83D798BF529950FC61F83DFBF7DE04" box="[1014,1068,2039,2058]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="km" value="10.0">10 km</quantity>
NW of Andasibe Vill.
</locationDeviation>
,
<location id="8EA42CA6BF529950FB69F83DFAADDE04" LSID="urn:lsid:plazi:treatment:03D2CB6BBF52995FFCCCF8EAFB83D53F:8EA42CA6BF529950FB69F83DFAADDE04" box="[1278,1398,2039,2058]" country="Madagascar" name="Torotorofotsy" pageId="6" pageNumber="224" stateProvince="Toamasina">Torotorofotsy</location>
</materialsCitation>
</paragraph>
<caption id="DF042AF5BF539951FF75F8A3FEF4DE1F" ID-DOI="http://doi.org/10.5281/zenodo.13178134" ID-Zenodo-Dep="13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="7" pageNumber="225" startId="7.[226,263,1897,1916]" targetBox="[194,1390,320,1872]" targetPageId="7" targetType="figure">
<paragraph id="8BC47A7DBF539951FF75F8A3FAE0D1A2" blockId="7.[189,1399,1897,2065]" pageId="7" pageNumber="225">
Figs 38–46.
<taxonomicName id="4C7B01FEBF539951FEDAF8A3FD07D172" authorityName="Gorochov" authorityYear="2022" box="[333,732,1897,1916]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF539951FEDAF8A3FD07D172" box="[333,732,1897,1916]" italics="true" pageId="7" pageNumber="225">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF539951FD76F8A3FCD7D172" bold="true" box="[737,780,1897,1916]" pageId="7" pageNumber="225">
<taxonomicNameLabel id="A23C1B14BF539951FD76F8A3FCD7D172" box="[737,780,1897,1916]" pageId="7" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 38–39, 42–43 — head with pronotum and fore legs (38 — from above; 39 — from side, 42–43 — from side/above); 40–41 — tegmina in rest position from above; 44 — genital plate from below; 45–46 — distal part of ovipositor (45 — from side; 46 — from above); 38–42 — male (38–40 — holotype; 41–42 — paratype); 43–46 — female.
</paragraph>
<paragraph id="8BC47A7DBF539951FF75F87CFEF4DE1F" blockId="7.[189,1399,1897,2065]" pageId="7" pageNumber="225">
Рис. 38–46.
<taxonomicName id="4C7B01FEBF539951FEC4F87FFD3DD1C6" authorityName="Gorochov" authorityYear="2022" box="[339,742,1973,1992]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF539951FEC4F87FFD3DD1C6" box="[339,742,1973,1992]" italics="true" pageId="7" pageNumber="225">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF539951FD79F87FFCC2D1C6" bold="true" box="[750,793,1973,1992]" pageId="7" pageNumber="225">
<taxonomicNameLabel id="A23C1B14BF539951FD79F87FFCC2D1C6" box="[750,793,1973,1992]" pageId="7" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 38–39, 42–43 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 42–43 — сбоку/сверху); 40–41 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 45–46 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 38–42 — самец (38–40 — голотип; 41–42 — паратип); 43– 46 — самка.
</paragraph>
</caption>
<caption id="DF042AF5BF5C995EFF75F8EBFDE7DE1F" ID-DOI="http://doi.org/10.5281/zenodo.13178136" ID-Zenodo-Dep="13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="8" pageNumber="226" startId="8.[226,263,1825,1844]" targetBox="[193,1395,317,1801]" targetPageId="8" targetType="figure">
<paragraph id="8BC47A7DBF5C995EFF75F8EBFEEDD19A" blockId="8.[189,1398,1825,2065]" pageId="8" pageNumber="226">
Figs 47–59.
<taxonomicName id="4C7B01FEBF5C995EFEDAF8EBFD0FD13A" authorityName="Gorochov" authorityYear="2022" box="[333,724,1825,1844]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5C995EFEDAF8EBFD0FD13A" box="[333,724,1825,1844]" italics="true" pageId="8" pageNumber="226">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5C995EFD4DF8EBFCDFD13A" box="[730,772,1825,1844]" pageId="8" pageNumber="226" rank="species">
<emphasis id="B90FA66FBF5C995EFD4DF8EBFD24D13A" bold="true" box="[730,767,1825,1844]" pageId="8" pageNumber="226">sp.n</emphasis>
.
</taxonomicNameLabel>
:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female.
</paragraph>
<paragraph id="8BC47A7DBF5C995EFF75F854FDE7DE1F" blockId="8.[189,1398,1825,2065]" pageId="8" pageNumber="226">
Рис. 47–59.
<taxonomicName id="4C7B01FEBF5C995EFEC7F857FD03D1BE" authorityName="Gorochov" authorityYear="2022" box="[336,728,1949,1968]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5C995EFEC7F857FD03D1BE" box="[336,728,1949,1968]" italics="true" pageId="8" pageNumber="226">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5C995EFD48F857FCD2D1BE" box="[735,777,1949,1968]" pageId="8" pageNumber="226" rank="species">
<emphasis id="B90FA66FBF5C995EFD48F857FCDFD1BE" bold="true" box="[735,772,1949,1968]" pageId="8" pageNumber="226">sp.n</emphasis>
.
</taxonomicNameLabel>
:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF5D995FFF2AFEF6FF2ED768" blockId="9.[189,776,315,2065]" pageId="9" pageNumber="227">
Forest Reserve, ~
<quantity id="4C83D798BF5D995FFEC0FEF1FE46D741" box="[343,413,315,335]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="227" unit="m" value="1000.0">1000 m</quantity>
, at light,
<date id="FFC55CBDBF5D995FFE65FEF1FD51D740" box="[498,650,315,334]" pageId="9" pageNumber="227" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">22.II–11.III.2013</date>
, A. Gorochov (
<collectionCode id="ED6AE2B8BF5D995FFF54FE99FF30D768" box="[195,235,339,358]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="227" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFF75FEA1FAACD4EF" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995FFF75FEA1FE77D182" blockId="9.[189,776,315,2065]" pageId="9" pageNumber="227">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF5D995FFE44FEA1FDF6D78F" box="[467,557,363,385]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
). General appearance similar to that of all species of
<taxonomicName id="4C7B01FEBF5D995FFE67FE4CFD55D795" authorityName="Gorochov" authorityYear="2004" box="[496,654,390,411]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFE67FE4CFD55D795" box="[496,654,390,411]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFD02FE4CFD10D795" box="[661,715,390,411]" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
Body coloration yellowish with following pattern: eyes greyish; dorsum of epicranium greyish brown with a pair of yellowish longitudinal stripes running from rostral apex to yellowish lateral ocelli and from these ocelli to posterior edge of head (along medial edges of eyes and behind them), with a pair of short and thin light longitudinal stripes between previous stripes on posterior part of epicranial dorsum, with also light median line on posterior two thirds of this dorsum (
<figureCitation id="134066F8BF5D995FFD26FD91FF03D485" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 47– 49</figureCitation>
); upper parts of membranes of antennal cavities and small areas on upper parts of genae behind eyes slightly darkened; scape with brown dorsal band along medial edge, darkish ventral stripe along same edge and darker (greyish brown) ventromedian spot in apical part; rest of antenna with brown to light brown spots in proximal and middle parts as well as with almost uniformly light brown distal part; pronotum with dark brown disc having a few thin lightish marks, with slightly darkish longitudinal band on each lateral lobe (this band occupying upper two thirds of this lobe but separated from dark disc by longitudinal light stripe), with a few small barely darker marks on above-mentioned darkish band, and with several dark and light spots along anterior edge of this lobe; legs with small and sparse dark and darkish spots (
<figureCitation id="134066F8BF5D995FFD4BFC1BFF25D20E" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 47–49</figureCitation>
); tegmen with yellowish grey dorsal field having darker (greyish brown) venation as well as yellowish both dot near plectrum and narrow stripe along lateral edge of distal half of this field, with light greyish (semitransparent) lateral field having greyish brown longitudinal veins and whitish crossveins, and with humeral region between these fields having dark (brown) proximal half of M and whitish crossveins as well as almost semimembranous membranes (
<figureCitation id="134066F8BF5D995FFD48FB6CFF03D2D8" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 51</figureCitation>
); hind wings light greyish with somewhat darkened distal parts (in rest condition); anal plate with a pair of rather large greyish brown spots in its laterodorsal corners (
<figureCitation id="134066F8BF5D995FFD17FB3CFD1CD302" box="[640,711,1270,1292]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 50</figureCitation>
); cerci with very small and moderately numerous darkish marks. Eyes rather large, slightly higher than long (
<figureCitation id="134066F8BF5D995FFD11FAE1FD27D34E" box="[646,764,1323,1345]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 47–49</figureCitation>
); rostrum practically angular in profile; scape approximately 2.3 times as wide as space between antennal cavities (
<figureCitation id="134066F8BF5D995FFD25FAABFD20D378" box="[690,763,1377,1399]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 49</figureCitation>
); pronotum as in
<figureCitation id="134066F8BF5D995FFEC2FAB1FE1FD39E" box="[341,452,1403,1425]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 47–48</figureCitation>
; metanotal gland absent; fore leg with only inner tympanum which oval, almost as long as width of fore tibia, and almost as wide as half of latter width; tegmina distinctly protruding beyond apices of hind femora and with venation as in
<figureCitation id="134066F8BF5D995FFE0AFA2CFE38D3F2" box="[413,483,1510,1532]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 51</figureCitation>
; hind wings clearly protruding beyond tegminal apices; anal plate small, transverse, truncate apically, widely rounded distolaterally and with large and almost longitudinally oval membranous median area (
<figureCitation id="134066F8BF5D995FFD24F9FCFD20D042" box="[691,763,1590,1612]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 50</figureCitation>
); genital plate more or less similar to that of
<taxonomicName id="4C7B01FEBF5D995FFDC3F99BFD29D068" authorityName="Gorochov" authorityYear="2004" box="[596,754,1617,1638]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFDC3F99BFD29D068" box="[596,754,1617,1638]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFD60F99BFF07D08F" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
but with slightly wider distal lobule having almost angular apex. Genitalia (
<figureCitation id="134066F8BF5D995FFE1CF94CFDD9D092" box="[395,514,1670,1692]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 52–54</figureCitation>
): epiphallus more or less similar to that of other subgenera of this genus but with dorsoapical processes having much smaller (poorly distinct) distal tubercles, and with ventroapical lobules much shorter and more hooked; ectoparameres distinctly shorter; anterior sclerotized ribbon of formula longer and narrower than in
<taxonomicName id="4C7B01FEBF5D995FFF2AF8ECFE80D135" authorityName="Gorochov" authorityYear="2004" box="[189,347,1830,1851]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFF2AF8ECFE80D135" box="[189,347,1830,1851]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFEF2F8ECFE45D135" box="[357,414,1830,1851]" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
(i.e. not plate-like), located clearly less parallel to median part of rest of this formula than in the latter subgenus and in
<taxonomicName id="4C7B01FEBF5D995FFE30F891FD87D17E" authorityName="Gorochov" authorityYear="2022" box="[423,604,1883,1904]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF5D995FFE30F891FD87D17E" box="[423,604,1883,1904]" italics="true" pageId="9" pageNumber="227">Euzvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5D995FFDF1F891FD13D17E" bold="true" box="[614,712,1883,1904]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFDF1F891FD13D17E" box="[614,712,1883,1904]" pageId="9" pageNumber="227" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
Spermatophore as in
<figureCitation id="134066F8BF5D995FFEF7F8BCFE73D182" box="[352,424,1910,1932]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 55</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFF75F85BFA95D75F" blockId="9.[189,776,315,2065]" lastBlockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Variations. Second male distinguished from
<typeStatus id="54C0C4DFBF5D995FFD04F85BFD3CD1A9" box="[659,743,1937,1959]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
by insignificantly less contrast coloration of head, lighter pronotal disc, yellowish brown tinge of tegmina, somewhat different venation of their dorsal field (
<figureCitation id="134066F8BF5D995FFE6CF82BFD99D1F8" box="[507,578,2016,2039]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 56</figureCitation>
), as well as genitalia with slightly smaller anterolateral (curved posteriorly) parts of epiphallus and barely lower lateral plates of formula.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FE9CFBE6D428" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Female. Coloration and structure of body practically as in males, but scape somewhat lighter, dorsal tegminal field with 9–10 almost straight (barely oblique) longitudinal veins as well as rather numerous and partly irregular crossveins, genital plate almost as wide as long and somewhat narrowing in distal half as well as with distinct roundly angular posteromedian notch (
<figureCitation id="134066F8BF5D995FFC5FFE3CFBCAD402" box="[968,1041,502,524]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 57</figureCitation>
); ovipositor normal and with drilling distal part as in
<figureCitation id="134066F8BF5D995FFC5EFDDBFBE2D428" box="[969,1081,528,551]" captionStart="Figs 47–59" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 52–54 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 58–59 — distal part of ovipositor (58 — from below; 59 —from side); 47–56 — male (47–55 — holotype; 56 — paratype); 57– 59 — female. Рис. 47–59. Zvenellomorpha (Azvenellomorpha) separata sp.n.:47–48 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 52–54 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 58–59 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 47–56 — самец (47–55 — голотип; 56 — паратип); 57–59 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 58–59</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FDE1FB9DD49F" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Length in mm. Body:
<specimenCount id="9D7DB1F4BF5D995FFBB4FDE2FBE2D44E" box="[1059,1081,552,576]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
14.5–15.5,
<specimenCount id="9D7DB1F4BF5D995FFB3DFDE2FB6CD44E" box="[1194,1207,552,576]" count="1" pageId="9" pageNumber="227" type="female">♀</specimenCount>
15–17.5; body with wings:
<specimenCount id="9D7DB1F4BF5D995FFCE2FD89FC50D455" box="[885,907,579,603]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
24–25.5,
<specimenCount id="9D7DB1F4BF5D995FFC66FD89FC25D455" box="[1009,1022,579,603]" count="1" pageId="9" pageNumber="227" type="female">♀</specimenCount>
26–29; pronotum:
<specimenCount id="9D7DB1F4BF5D995FFB57FD89FB0DD455" box="[1216,1238,579,603]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
2.7–2.9,
<specimenCount id="9D7DB1F4BF5D995FFAA1FD89FA98D455" box="[1334,1347,579,603]" count="1" pageId="9" pageNumber="227" type="female">♀</specimenCount>
3.1– 3.4; tegmina:
<specimenCount id="9D7DB1F4BF5D995FFC26FD94FC1CD478" box="[945,967,606,630]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
17.5–18.5;
<specimenCount id="9D7DB1F4BF5D995FFBACFD94FB93D478" box="[1083,1096,606,630]" count="1" pageId="9" pageNumber="227" type="female">♀</specimenCount>
18–20; hind femora:
<specimenCount id="9D7DB1F4BF5D995FFA8FFD94FAF5D478" box="[1304,1326,606,630]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
13–14,
<specimenCount id="9D7DB1F4BF5D995FFCBBFDB2FCE2D49E" box="[812,825,632,656]" count="1" pageId="9" pageNumber="227" type="female">♀</specimenCount>
13.5–15; ovipositor 9–9.5.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FD5CFAACD4EF" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
COMPARISON. The new species differs from all the other congeners in the same characters as the subgenus
<taxonomicName id="4C7B01FEBF5D995FFCBBFD01FC0ED4EE" authorityName="Gorochov" authorityYear="2022" box="[812,981,715,736]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5D995FFCBBFD01FC0ED4EE" box="[812,981,715,736]" italics="true" pageId="9" pageNumber="227">Azvenellomorpha</emphasis>
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<emphasis id="B90FA66FBF5D995FFC4CFD01FBE6D4EE" bold="true" box="[987,1085,715,736]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFC4CFD01FBE6D4EE" box="[987,1085,715,736]" pageId="9" pageNumber="227" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
(see the subgeneric key above).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFCC5FD2CFB83D53F" pageId="9" pageNumber="227" type="etymology">
<paragraph id="8BC47A7DBF5D995FFCC5FD2CFB83D53F" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF5D995FFCBBFCCBFC53D518" authorityName="Gorochov" authorityYear="2022" box="[812,904,769,790]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="separata">“separata</taxonomicName>
” (separate) due to some distinct differences in the structure of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>

196
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<mods:title id="34B2A5DA7C0EA2A53E3157A467A362A9">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<paragraph id="8BC47A7DBF5D995FFC19FC8FFACFD553" blockId="9.[812,1399,837,2061]" box="[910,1300,837,861]" pageId="9" pageNumber="227">
<heading id="D08CCD11BF5D995FFC19FC8FFACFD553" box="[910,1300,837,861]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="227" reason="8">
<taxonomicName id="4C7B01FEBF5D995FFC19FC8FFB0FD553" authority="Gorochov" authorityName="Gorochov" authorityYear="2022" box="[910,1236,837,861]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5D995FFC19FC8FFB85D553" box="[910,1118,837,861]" italics="true" pageId="9" pageNumber="227">Ultratrella? ornata</emphasis>
Gorochov
</taxonomicName>
,
<emphasis id="B90FA66FBF5D995FFB48FC8FFACFD553" bold="true" box="[1247,1300,837,861]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFB48FC8FFACFD553" box="[1247,1300,837,861]" pageId="9" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFB85FCA8FB49D574" box="[1042,1170,866,890]" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995FFB85FCA8FB49D574" blockId="9.[812,1399,837,2061]" box="[1042,1170,866,890]" pageId="9" pageNumber="227">
<figureCitation id="134066F8BF5D995FFB85FCA8FB56D574" box="[1042,1165,866,890]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60–67</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFCC5FC4DFCBED5EC" pageId="9" pageNumber="227" type="materials_examined">
<paragraph id="8BC47A7DBF5D995FFCC5FC4DFCBED5EC" blockId="9.[812,1399,837,2061]" pageId="9" pageNumber="227">
<materialsCitation id="3B137020BF5D995FFCC5FC4DFCBED5EC" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="9" pageNumber="227" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
MATERIAL EXAMINED.
<typeStatus id="54C0C4DFBF5D995FFBD4FC4DFB48D594" box="[1091,1171,903,922]" pageId="9" pageNumber="227" type="holotype">Holotype</typeStatus>
—
<specimenCount id="9D7DB1F4BF5D995FFB24FC4FFB1CD595" box="[1203,1223,901,923]" count="1" pageId="9" pageNumber="227" type="male">♂</specimenCount>
,
<emphasis id="B90FA66FBF5D995FFB45FC4DFA98D594" bold="true" box="[1234,1347,903,922]" pageId="9" pageNumber="227">
<collectingCountry id="F36C3AEDBF5D995FFB45FC4DFA98D594" box="[1234,1347,903,922]" name="Madagascar" pageId="9" pageNumber="227">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF5D995FFADAFC42FC7DD5BD" country="Madagascar" name="Toamasina" pageId="9" pageNumber="227">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF5D995FFC23FC6AFB87D5BC" box="[948,1116,927,947]" pageId="9" pageNumber="227">Moramanga Distr.</collectingMunicipality>
,
<location id="8EA42CA6BF5D995FFBFEFC55FAACD5BC" LSID="urn:lsid:plazi:treatment:03D2CB6BBF5D995DFC19FC8FFE4CD4A5:8EA42CA6BF5D995FFBFEFC55FAACD5BC" box="[1129,1399,927,947]" country="Madagascar" municipality="Moramanga Distr." name="Analamazaotra Forest Station" pageId="9" pageNumber="227" stateProvince="Toamasina">Analamazaotra Forest Station</location>
near Andasibe Vill. ~
<quantity id="4C83D798BF5D995FFC7BFC7DFBF3D5C5" box="[1004,1064,951,971]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="227" unit="m" value="900.0">
<elevation id="00569D4EBF5D995FFC7BFC7DFBF3D5C5" box="[1004,1064,951,971]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="227" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF5D995FFBE8FC7DFB23D5C4" box="[1151,1272,951,970]" pageId="9" pageNumber="227" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF5D995FFBE8FC7DFB23D5C4" box="[1151,1272,951,970]" pageId="9" pageNumber="227" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">8–20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF5D995FFA94FC7DFAADD5C4" box="[1283,1398,951,970]" pageId="9" pageNumber="227">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF5D995FFCA4FC05FC80D5EC" box="[819,859,975,994]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="227" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995DFCC5FC2DFE8ED452" lastPageId="11" lastPageNumber="229" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995CFCC5FC2DFAD0D428" blockId="9.[812,1399,837,2061]" lastBlockId="10.[812,1399,315,551]" lastPageId="10" lastPageNumber="228" pageId="9" pageNumber="227">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF5D995FFBD4FC2DFB46D5F3" box="[1091,1181,999,1021]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
). General appearance distinctly different from that of
<taxonomicName id="4C7B01FEBF5D995FFBF4FBC9FAA8D219" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[1123,1395,1026,1048]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5D995FFBF4FBC9FB10D219" box="[1123,1227,1026,1048]" italics="true" pageId="9" pageNumber="227">U. gracilis</emphasis>
<bibRefCitation id="EFEA078CBF5D995FFB45FBC8FAA8D219" author="Gorochov A. V." box="[1234,1395,1026,1048]" pageId="9" pageNumber="227" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
, i.e. body clearly smaller, and coloration more contrast: head and pronotum (
<figureCitation id="134066F8BF5D995FFC29FBFDFB94D243" box="[958,1103,1079,1101]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 62–63</figureCitation>
) dark grey with slightly lighter (grey) stripe along posterior edge of epicranial dorsum, yellowish ocelli and dorsal transverse band between anterior parts of eyes (this band situated behind median ocellus but very near it, and lateral ocelli located on this band), almost black scape and pedicel as well as rostral area, brown antennal flagellum and light brown to yellowish spots on mouthparts as well as on pronotal disc (lateral lobes of pronotum with only sparse light dots;
<figureCitation id="134066F8BF5D995FFBA3FAC7FB79D32C" box="[1076,1186,1292,1315]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 63</figureCitation>
); fore and middle legs dark greyish brown with light brown ventral surfaces of tibiae and tarsi as well as with yellowish trochanters and coxae (each coxa with darkish lateral area); hind legs brown with light brown basal parts of femora and all spines and spurs (
<figureCitation id="134066F8BF5D995FFCA4FA58FC7ED3A6" box="[819,933,1426,1448]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60–63</figureCitation>
); tegmina light grey with yellow spot at base of each tegmen (
<figureCitation id="134066F8BF5D995FFC22FA67FBF3D3CC" box="[949,1064,1452,1475]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 61, 63</figureCitation>
), greyish brown to blackish venation of dorsal field and longitudinal veins of lateral field, whitish crossveins in lateral field, slightly darkened spots on almost all membranes and rather wide whitish humeral stripe (
<figureCitation id="134066F8BF5D995FFCA4F9DDFC72D023" box="[819,937,1559,1581]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60–61</figureCitation>
); visible distal parts of hind wings also light grey; other parts of body venter (including abdominal sternites, genital plate and cerci) yellowish to light brown. Head rather short, slightly concave dorsally (between middle parts of eyes), somewhat depressed dorsoventrally, with large eyes which slightly higher than long, with rather large and almost round ocelli, with short mouthparts and very short palpi, as well as with scape barely wider than space between antennal cavities (
<figureCitation id="134066F8BF5D995FFC14F927FBC3D10C" box="[899,1048,1772,1795]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 62–63</figureCitation>
); pronotum distinctly transverse and moderately high (somewhat dorsoventrally depressed), with concave anterior edge of disc and convex posterior one (
<figureCitation id="134066F8BF5D995FFADCF8E8FCAAD15C" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 63</figureCitation>
); fore and middle legs rather short, not flattened, without keels and with fore tibiae having outer and inner tympana oval and almost equal to each other in size (each of them 1.6–1.7 times as wide as maximal width of fore tibia and barely longer than latter width); tegmina strongly protruding beyond hind femora in rest position, with mirror clearly smaller than in
<taxonomicName id="4C7B01FEBF5D995FFC52F817FBF5D1FC" box="[965,1070,2013,2034]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5D995FFC52F817FBF5D1FC" box="[965,1070,2013,2034]" italics="true" pageId="9" pageNumber="227">U. gracilis</emphasis>
</taxonomicName>
and with rest venation as in
<figureCitation id="134066F8BF5D995FFADAF817FCB2DE03" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60–61</figureCitation>
(lateral field with very narrow Sc-R area, 15–16 branches of Sc, rather numerous crossveins in proximal half of this field and almost without crossveins in its distal half); hind wings distinctly protruding beyond tegminal apices; genital plate practically intermediate between those of
<taxonomicName id="4C7B01FEBF5E995CFD29FE41FEC0D7B5" authorityName="Gorochov" authorityYear="2004" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B90FA66FBF5E995CFD29FE41FEC0D7B5" italics="true" pageId="10" pageNumber="228">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5E995CFEB7FE6CFE99D7B2" box="[288,322,422,444]" pageId="10" pageNumber="228" sensu="lato">s. l.</taxonomicNameLabel>
and
<taxonomicName id="4C7B01FEBF5E995CFEE6FE6DFE03D7B5" box="[369,472,422,444]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFEE6FE6DFE03D7B5" box="[369,472,422,444]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
[see
<bibRefCitation id="EFEA078CBF5E995CFD9DFE6CFD70D7B5" author="Gorochov A. V." box="[522,683,422,444]" pageId="10" pageNumber="228" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
: figs X, 2 and 7]. Genitalia (
<figureCitation id="134066F8BF5E995CFEE4FE0BFE3FD7D8" box="[371,484,448,471]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="10" pageNumber="228">Figs 64–66</figureCitation>
) also more or less similar to those of
<taxonomicName id="4C7B01FEBF5E995CFE82FE16FE5BD7FE" box="[277,384,475,497]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFE82FE16FE5BD7FE" box="[277,384,475,497]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
(including similar structure of formula and rachis), but: distal part of epiphallus higher (less narrow in profile) and with dorsoapical processes triangular (not bilobate) in profile and located near convex part of epiphallus (in
<taxonomicName id="4C7B01FEBF5E995CFCDAFE9DFC6ED765" box="[845,949,342,364]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFCDAFE9DFC6ED765" box="[845,949,342,364]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
, this processes and latter part located not near each other); ventroapical epiphallic lobules higher (wider in profile); formula with shorter anterior ribbon and with lateral lobes angularly projected backwards (
<emphasis id="B90FA66FBF5E995CFB0BFE6DFB6BD7B2" box="[1180,1200,423,444]" italics="true" pageId="10" pageNumber="228">vs</emphasis>
. not projected backwards); ectoparameres clearly longer; rachis with semimembranous subapical inflation (
<emphasis id="B90FA66FBF5E995CFBAAFE16FB8AD7FF" box="[1085,1105,476,497]" italics="true" pageId="10" pageNumber="228">vs</emphasis>
. without any inflation in distal part). Spermatophore with elongately rounded ampulla (oth- er structures of spermatophore missing;
<figureCitation id="134066F8BF5E995CFB21FDDBFB24D428" box="[1206,1279,528,551]" captionStart="Figs 60–67" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 60–67. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 60–67. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="10" pageNumber="228">Fig. 67</figureCitation>
).
</paragraph>
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<paragraph id="8BC47A7DBF5E995CFF75F84BFBD0D1CA" blockId="10.[189,1399,1921,2064]" pageId="10" pageNumber="228">
Figs 60–67.
<taxonomicName id="4C7B01FEBF5E995CFEDAF84BFE33D19A" authorityName="Gorochov" authorityYear="2022" box="[333,488,1921,1940]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5E995CFEDAF84BFE33D19A" box="[333,488,1921,1940]" italics="true" pageId="10" pageNumber="228">Ultratrella ornata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5E995CFE7AF84BFDC3D19A" bold="true" box="[493,536,1921,1940]" pageId="10" pageNumber="228">
<taxonomicNameLabel id="A23C1B14BF5E995CFE7AF84BFDC3D19A" box="[493,536,1921,1940]" pageId="10" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 64–66 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side.
</paragraph>
<paragraph id="8BC47A7DBF5E995CFF75F807FB98DE1E" blockId="10.[189,1399,1921,2064]" pageId="10" pageNumber="228">
Рис. 60–67.
<taxonomicName id="4C7B01FEBF5E995CFED8F807FE30D1EE" authorityName="Gorochov" authorityYear="2022" box="[335,491,1997,2016]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5E995CFED8F807FE30D1EE" box="[335,491,1997,2016]" italics="true" pageId="10" pageNumber="228">Ultratrella ornata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5E995CFE66F807FDC7D1EE" bold="true" box="[497,540,1997,2016]" pageId="10" pageNumber="228">
<taxonomicNameLabel id="A23C1B14BF5E995CFE66F807FDC7D1EE" box="[497,540,1997,2016]" pageId="10" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 64–66 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF5F995DFF75FEF1FE49D75F" blockId="11.[189,775,315,684]" box="[226,402,315,337]" pageId="11" pageNumber="229">Female unknown.</paragraph>
<paragraph id="8BC47A7DBF5F995DFF75FE9CFDC9D788" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">Length in mm. Body 10; body with wings 27; pronotum 2.8; tegmina 18.7; hind femora 10.</paragraph>
<paragraph id="8BC47A7DBF5F995DFF75FE41FE8ED452" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">
COMPARISON. The new species is clearly different from
<taxonomicName id="4C7B01FEBF5F995DFF65FE6DFE81D7B5" box="[242,346,422,444]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5F995DFF65FE6DFE81D7B5" box="[242,346,422,444]" italics="true" pageId="11" pageNumber="229">U. gracilis</emphasis>
</taxonomicName>
(
<typeStatus id="54C0C4DFBF5F995DFEFFFE6CFE48D7B5" box="[360,403,422,443]" pageId="11" pageNumber="229">type</typeStatus>
and only species of this genus) in the body smaller, coloration much more dark and contrast, a relatively smaller mirror in the male tegmen, and the above-mentioned characters of the male genitalia. These differences may be treated also as subgeneric or even generic ones, but this question may be clarified only after the study of some additional data.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFF75FDABFE4CD4A5" pageId="11" pageNumber="229" type="etymology">
<paragraph id="8BC47A7DBF5F995DFF75FDABFE4CD4A5" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF5F995DFF2AFDB1FEDED49E" authorityName="Gorochov" authorityYear="2022" box="[189,261,635,656]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="species" species="ornata">“ornata</taxonomicName>
” (ornated, adorned) due to the characteristic (contrasting) body pattern.
</paragraph>
</subSubSection>
</treatment>
</document>

146
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<emphasis id="B90FA66FBF5F995DFE88FD0AFE36D4D6" box="[287,493,704,728]" italics="true" pageId="11" pageNumber="229">Ultratrella gracilis</emphasis>
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<materialsCitation id="3B137020BF5F995DFF75FD2FFD00D526" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="11" pageNumber="229" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina">
MATERIAL EXAMINED.
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<collectingCountry id="F36C3AEDBF5F995DFE43FD2FFD9ED4F6" box="[468,581,741,760]" name="Madagascar" pageId="11" pageNumber="229">Madagascar</collectingCountry>
</emphasis>
:
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,
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,
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,
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,
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near Andasibe Vill. ~
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, at light,
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,
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(
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.
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DESCRIPTION. Female (nov.). General appearance similar to that of male
<typeStatus id="54C0C4DFBF5F995DFE05FC8DFE3CD553" box="[402,487,839,861]" pageId="11" pageNumber="229" type="holotype">holotype</typeStatus>
[
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] but with following differences: coloration even more uniformly light greyish brown, i.e. with less distinct lighter (yellowish white) stripes on head and pronotum, and practically without whitish humeral stripe on each tegmen; tegminal dorsal field with 13–14 longitudinal veins which barely S-shaped in lateral part of proximal half, as well as with numerous and slightly irregular crossveins; tegminal lateral field with 15–17 oblique branches of Sc which partly and slightly arcuate, with moderately numerous crossveins, and with area between Sc stem and R moderately narrow (in male, this area very strongly narrowed); genital plate elongate but slightly shorter than two last abdominal sternites together, gradually narrowing to roundly truncate apex having very short and roundly angular posteromedian notch; ovipositor long and with distinctly drilling apical part more or less similar to that of
<taxonomicName id="4C7B01FEBF5F995DFCBBFE9CFC11D765" authorityName="Gorochov" authorityYear="2004" box="[812,970,342,363]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B90FA66FBF5F995DFCBBFE9CFC11D765" box="[812,970,342,363]" italics="true" pageId="11" pageNumber="229">Zvenellomorpha</emphasis>
</taxonomicName>
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</paragraph>
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Male. Body very similar to that of
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of this species, but tegmina with slightly longer chords and barely smaller mirror in dorsal field; genitalia almost identical to those of this
<typeStatus id="54C0C4DFBF5F995DFC3BFE0BFBDAD7D9" box="[940,1025,449,471]" pageId="11" pageNumber="229" type="holotype">holotype</typeStatus>
[
<bibRefCitation id="EFEA078CBF5F995DFB99FE0BFB6ED7D8" author="Gorochov A. V." box="[1038,1205,449,471]" pageId="11" pageNumber="229" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
].
</paragraph>
<paragraph id="8BC47A7DBF5F995DFCC5FE11FAAAD428" blockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
Length in mm. Body:
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22,
<specimenCount id="9D7DB1F4BF5F995DFBF4FE12FBABD7FE" box="[1123,1136,472,496]" count="1" pageId="11" pageNumber="229" type="female">♀</specimenCount>
19–21.5; body with wings:
<specimenCount id="9D7DB1F4BF5F995DFCBBFE39FC99D405" box="[812,834,499,523]" count="1" pageId="11" pageNumber="229" type="male">♂</specimenCount>
35,
<specimenCount id="9D7DB1F4BF5F995DFCFAFE39FCA1D405" box="[877,890,499,523]" count="1" pageId="11" pageNumber="229" type="female">♀</specimenCount>
37–39; pronotum:
<specimenCount id="9D7DB1F4BF5F995DFBA4FE39FB92D405" box="[1075,1097,499,523]" count="1" pageId="11" pageNumber="229" type="male">♂</specimenCount>
4.3,
<specimenCount id="9D7DB1F4BF5F995DFBEEFE39FB5DD405" box="[1145,1158,499,523]" count="1" pageId="11" pageNumber="229" type="female">♀</specimenCount>
4.4–4.7; tegmina:
<specimenCount id="9D7DB1F4BF5F995DFAABFE39FA89D405" box="[1340,1362,499,523]" count="1" pageId="11" pageNumber="229" type="male">♂</specimenCount>
25,
<specimenCount id="9D7DB1F4BF5F995DFCBBFDC4FCE2D428" box="[812,825,526,550]" count="1" pageId="11" pageNumber="229" type="female">♀</specimenCount>
25–27; hind femora:
<specimenCount id="9D7DB1F4BF5F995DFB98FDC4FBFED428" box="[1039,1061,526,550]" count="1" pageId="11" pageNumber="229" type="male">♂</specimenCount>
16,
<specimenCount id="9D7DB1F4BF5F995DFBC4FDC4FBBBD428" box="[1107,1120,526,550]" count="1" pageId="11" pageNumber="229" type="female">♀</specimenCount>
17–17.5; ovipositor 16–17.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFCC5FDE1FBDAD4EF" pageId="11" pageNumber="229" type="discussion">
<paragraph id="8BC47A7DBF5F995DFCC5FDE1FC1ED479" blockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
REMARKS. This species, originally described from “Jlôt- Prune” (
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), is here recorded from another Madagascan locality.
</paragraph>
<paragraph id="8BC47A7DBF5F995DFCC5FD5CFBDAD4EF" blockId="11.[812,1398,662,737]" pageId="11" pageNumber="229">
<emphasis id="B90FA66FBF5F995DFCC5FD5CFBC4D4A2" bold="true" box="[850,1055,662,684]" pageId="11" pageNumber="229">Acknowledgements.</emphasis>
This study was performed in the frames of the state research project No. 1021051302540-6 (
<collectingCountry id="F36C3AEDBF5F995DFCA4FD01FC2DD4EF" box="[819,1014,715,737]" name="Russia" pageId="11" pageNumber="229">Russian Federation</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

214
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<emphasis id="B93610627378BB455FDB552CFE2DFBD4" box="[195,455,1054,1078]" italics="true" pageId="17" pageNumber="218">Scathophaga incompleta</emphasis>
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<paragraph id="8BFDCC707378BB455E6A5509FDBBFBB1" blockId="17.[195,769,1054,1107]" box="[370,593,1083,1107]" pageId="17" pageNumber="218">
<figureCitation id="1379D0F57378BB455E6A5509FE2FFBB1" box="[370,453,1083,1107]" captionStart="26 27 28 29" captionStartId="9.[189,250,1754,1812]" captionText="26 27 28 29 Figs 14–29. Male sternite 4 of Scathophaga spp.: 14 — S. apicalis (Curtis); 15 — S. buryatica Ozerov et Krivosheina; 16 — S. calida (Haliday); 17 — S. cordylurina (Holmgren); 18 — S. crinita (Coquillett); 19 — S. dasythrix (Becker); 20 — S. decipiens (Haliday); 21 — S. exalata Ozerov; 22 — S. furcata (Say); 23 — S. incola (Becker); 24 — S. incompleta Ozerov et Krivosheina; 25 — S. inquinata (Meigen); 26 — S. intermedia (Walker); 27 — S. karelica Ozerov; 28 — S. lapponica (Ringdahl); 29 — S. litorea (Fallén). 15 — after Ozerov &amp; Krivosheina, 2019, fig. 1; 20, 24 — after Ozerov &amp; Krivosheina, 2020, figs 3, 9; 27 — after Ozerov, 2013, fig.13. Рис. 14–29. Стернит 4 самца Scathophaga spp.: 14 — S. apicalis (Curtis); 15 — S. buryatica Ozerov et Krivosheina; 16 — S. calida (Haliday); 17 — S. cordylurina (Holmgren); 18 — S. crinita (Coquillett); 19 — S. dasythrix (Becker); 20 — S. decipiens (Haliday); 21 — S. exalata Ozerov; 22 — S. furcata (Say); 23 — S. incola (Becker); 24 — S. incompleta Ozerov et Krivosheina; 25 — S. inquinata (Meigen); 26 — S. intermedia (Walker); 27 — S. karelica Ozerov; 28 — S. lapponica (Ringdahl); 29 — S. litorea (Fallén). 15 — по Ozerov &amp; Krivosheina, 2019, fig. 1; 20, 24 — по Ozerov &amp; Krivosheina, 2020, figs 3, 9; 27 — по Ozerov, 2013, fig.13." pageId="17" pageNumber="218">Figs 24</figureCitation>
,
<figureCitation id="1379D0F57378BB455EC95509FDFCFBB1" box="[465,534,1083,1107]" captionStart="76 77 78" captionStartId="20.[400,461,1797,1855]" captionText="76 77 78 Figs 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) and S. inquinata (Meigen) (76–78): 70, 73, 76 — male sternite 5; 71, 74, 77 — epandrium, cercal plate and surstyli, lateral view; 72, 75, 78 — same, dorsal view. 73–75 — after Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13. Рис. 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) и S. inquinata (Meigen) (76–78): 70, 73, 76 — стернит 5 самца; 71, 74, 77 — Эпандрий, церки и сурстили, сбоку; 72, 75, 78 — то же, сверху. 73–75 — по Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13." pageId="17" pageNumber="218">73–75</figureCitation>
,
<figureCitation id="1379D0F57378BB455D3A5509FDA7FBB1" box="[546,589,1083,1107]" captionStart="Figs 130–137" captionStartId="41.[226,263,1885,1904]" captionTargetBox="[229,1357,318,1859]" captionTargetId="figure-7@41.[209,1380,303,1879]" captionTargetPageId="41" captionText="Figs 130–137. Aedeagus of Scathophaga spp., lateral view: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — after Ozerov &amp; Krivosheina, 2019, fig. 2; 131, 132 — after Ozerov &amp; Krivosheina, 2020, figs 5, 11. Рис. 130–137. Эдеагус Scathophaga spp., сбоку: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — по Ozerov &amp; Krivosheina, 2019, fig. 2; 131, 132 — по Ozerov &amp; Krivosheina, 2020, figs 5, 11." figureDoi="http://doi.org/10.5281/zenodo.13178494" httpUri="https://zenodo.org/record/13178494/files/figure.png" pageId="17" pageNumber="218">132</figureCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="B93610627378BB455FFA554BFEA9FB6E" box="[226,323,1145,1164]" italics="true" pageId="17" pageNumber="218">incompleta</emphasis>
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. Type-locality: Priyutnoe env. (
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) (
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:
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REMARK. This species was recorded in
<collectingCountry id="F3558CE07378BB455D91559BFD25FB23" box="[649,719,1193,1217]" name="Russia" pageId="17" pageNumber="218">Russia</collectingCountry>
from
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, Crimea and
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[
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].
</paragraph>
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MATERIAL EXAMINED.
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<collectingRegion id="498602927378BB455EC55433FDD8FAF6" box="[477,562,1281,1300]" country="Russia" name="Dagestan" pageId="17" pageNumber="218">Dagestan</collectingRegion>
</emphasis>
:
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(
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),
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<collectingDate id="EFB813587378BB455E3A542BFE95FACE" box="[290,383,1305,1324]" pageId="17" pageNumber="218" value="1921-10-12">12.X.1921</collectingDate>
</date>
, Ryabov (
<specimenCount id="9D4407F97378BB455EF9542BFDEDFACE" box="[481,519,1302,1324]" count="1" pageId="17" pageNumber="218" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED5354B57378BB455D0E542BFDACFACE" box="[534,582,1305,1324]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="218">ZISP</collectionCode>
). See also
<bibRefCitation id="EFD3B1817378BB455DB5542BFE72FAA6" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="17" pageNumber="218" pagination="227 - 231" refId="ref51989" refString="Ozerov A. L., Krivosheina M. G. 2020. To taxonomy of the genus Scathophaga Meigen, 1803 (Diptera: Scathophagidae) with description of a new species // Russian Entomological Journal. Vol. 29. No. 2. P. 227 - 231." type="journal article" year="2020">Ozerov &amp; Krivosheina [2020: 228]</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFDCC707378BB455FFA547BFED6FA6E" blockId="17.[189,776,1145,2065]" pageId="17" pageNumber="218">
<materialsCitation id="3B2AC62D7378BB455FFA547BFED3FA6E" collectingDate="1932-11-02" collectionCode="ZISP" country="Turkmenistan" latitude="37.658" location="Etrek" longLatPrecision="70" longitude="54.763" pageId="17" pageNumber="218" specimenCount="1" specimenCount-male="1">
ADDITIONAL MATERIAL EXAMINED.
<emphasis id="B93610627378BB455D61547AFCEAFAB9" bold="true" box="[633,768,1352,1371]" pageId="17" pageNumber="218">
<collectingCountry id="F3558CE07378BB455D61547AFCEAFAB9" box="[633,768,1352,1371]" name="Turkmenistan" pageId="17" pageNumber="218">Turkmenistan</collectingCountry>
</emphasis>
:
<location id="8E9D9AAB7378BB455FA55453FF06FA96" LSID="urn:lsid:plazi:treatment:03EB7D667378BB455FDB552CFC63FC3B:8E9D9AAB7378BB455FA55453FF06FA96" box="[189,236,1377,1396]" country="Turkmenistan" latitude="37.658" longLatPrecision="70" longitude="54.763" name="Etrek" pageId="17" pageNumber="218">Etrek</location>
(
<geoCoordinate id="EE76AAB77378BB455FE55453FEBAFA96" box="[253,336,1377,1396]" degrees="37.658" direction="north" orientation="latitude" pageId="17" pageNumber="218" precision="55" value="37.658">37.658ºN</geoCoordinate>
<geoCoordinate id="EE76AAB77378BB455E445453FE45FA96" box="[348,431,1377,1396]" degrees="54.763" direction="east" orientation="longitude" pageId="17" pageNumber="218" precision="55" value="54.763">54.763ºE</geoCoordinate>
),
<date id="FFFCEAB07378BB455EDC5453FDF5FA96" box="[452,543,1377,1396]" pageId="17" pageNumber="218" value="1932-11-02">
<collectingDate id="EFB813587378BB455EDC5453FDF5FA96" box="[452,543,1377,1396]" pageId="17" pageNumber="218" value="1932-11-02">2.XI.1932</collectingDate>
</date>
, Ushinskiy (
<specimenCount id="9D4407F97378BB455D815453FD28FA96" box="[665,706,1374,1396]" count="1" pageId="17" pageNumber="218" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED5354B57378BB455DCA5453FCEAFA96" box="[722,768,1377,1396]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="218">ZISP</collectionCode>
) (
<emphasis id="B93610627378BB455FDC544AFEC5FA6E" bold="true" box="[196,303,1400,1420]" pageId="17" pageNumber="218">first record</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3589FFB7378BB455FFA54A3FAA3FC63" pageId="17" pageNumber="218" type="description">
<paragraph id="8BFDCC707378BB455FFA54A3FEFFFA27" blockId="17.[189,776,1145,2065]" pageId="17" pageNumber="218">
DESCRIPTION. Male. Female. Body-length
<quantity id="4CBA61957378BB455DC054A2FEFBFA27" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.6" metricValueMin="5.8" pageId="17" pageNumber="218" unit="mm" value="6.199999999999999" valueMax="6.6" valueMin="5.8">5.8– 6.6 mm</quantity>
.
</paragraph>
<paragraph id="8BFDCC707378BB455FFA54F9FEB1F971" blockId="17.[189,776,1145,2065]" pageId="17" pageNumber="218">
<emphasis id="B93610627378BB455FFA54F9FEF6FA01" box="[226,284,1483,1507]" italics="true" pageId="17" pageNumber="218">Head</emphasis>
. Frontal vitta yellow, with delicate whitish reflection; fronto-orbital plate blackish, densely pale grey dusted, ocellar triangle blackish. Face and gena yellow, with whitish reflection. Postcranium blackish. 3 orbital and 2–4 frontal setae present. Antenna black; postpedicel about twice as long as wide. Arista bare. Palpus yellow.
</paragraph>
<paragraph id="8BFDCC707378BB455FFA57ABFD25F89F" blockId="17.[189,776,1145,2065]" pageId="17" pageNumber="218">
<emphasis id="B93610627378BB455FFA57ABFEC4F953" box="[226,302,1689,1713]" italics="true" pageId="17" pageNumber="218">Thorax</emphasis>
black, densely pale grey dusted; scutum with a double brownish line down the middle, and an obscure one on each side. Acrostichals setulose in two rows, dorsocentrals 2+3, intra-alars 1+(1–2) (posterior postsutural intra-alar seta small, about 0.5 times as long as anterior one or absent), supra-alars 1+2, postpronotals 2. Scutellum greyish dusted, with a pair of strong basal scutellar and a pair of strong apical scutellar setae.
</paragraph>
<paragraph id="8BFDCC707378BB455FFA56B6FC14FDB9" blockId="17.[189,776,1145,2065]" lastBlockId="17.[812,1399,315,985]" pageId="17" pageNumber="218">
<emphasis id="B93610627378BB455FFA56B6FEFEF879" box="[226,276,1924,1947]" italics="true" pageId="17" pageNumber="218">Legs</emphasis>
. All coxae black, greyish dusted. All femora greyish dusted, black, except yellow apex. All tibiae and tarsi yellow. Fore femur with whitish hairs, longer ventrally, with 4–5 dorsal setulae in apical third. Fore tibia with 2–3 dorsal, 3–4 posterodorsal, 1–2 posterior, 1 preapical posterodorsal, 1 apical posteroventral, 1 apical posterior setae. Mid femur with a row of anterodorsal setae, with 1 preapical posterodorsal and 1 preapical posterior setae. Mid tibia with 1–2 anterodorsal, 1–2 posterodorsal, 2 posterior (thin), 1 ventral (strong in female) setae, also with a ring of apicals. Hind femur with a row of dorsal/anterodorsal setae. Hind tibia with 2–3 posterodorsal, 2–3 anterodorsal, 1 preapical dorsal, 1 preapical anterodorsal and 1 apical anteroventral setae.
</paragraph>
<paragraph id="8BFDCC707378BB455C4A5353FC8CFD74" blockId="17.[812,1399,315,985]" pageId="17" pageNumber="218">
<emphasis id="B93610627378BB455C4A5353FC60FD9A" box="[850,906,609,632]" italics="true" pageId="17" pageNumber="218">Wing</emphasis>
clear; crossveins r-m and dm-cu not darkened.
</paragraph>
<paragraph id="8BFDCC707378BB455C4A53A9FAA3FC63" blockId="17.[812,1399,315,985]" pageId="17" pageNumber="218">
<emphasis id="B93610627378BB455C4A53A9FC5FFD51" box="[850,949,667,691]" italics="true" pageId="17" pageNumber="218">Abdomen</emphasis>
black, densely pale grey dusted, covered with whitish hairs. Tergites
<quantity id="4CBA61957378BB455B49538AFB7EFD33" box="[1105,1172,696,721]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="12.7" metricValueMin="5.08" pageId="17" pageNumber="218" unit="in" value="3.5" valueMax="5.0" valueMin="2.0">2–5 in</quantity>
male and tergiters
<quantity id="4CBA61957378BB455A44538BFCB3FD0C" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="17" pageNumber="218" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">2– 6 in</quantity>
female each with a row of marginal setae. Male sternite 4 almost twice as long as wide (
<figureCitation id="1379D0F57378BB455BC453C1FAC6FCE9" box="[1244,1324,755,779]" captionStart="26 27 28 29" captionStartId="9.[189,250,1754,1812]" captionText="26 27 28 29 Figs 14–29. Male sternite 4 of Scathophaga spp.: 14 — S. apicalis (Curtis); 15 — S. buryatica Ozerov et Krivosheina; 16 — S. calida (Haliday); 17 — S. cordylurina (Holmgren); 18 — S. crinita (Coquillett); 19 — S. dasythrix (Becker); 20 — S. decipiens (Haliday); 21 — S. exalata Ozerov; 22 — S. furcata (Say); 23 — S. incola (Becker); 24 — S. incompleta Ozerov et Krivosheina; 25 — S. inquinata (Meigen); 26 — S. intermedia (Walker); 27 — S. karelica Ozerov; 28 — S. lapponica (Ringdahl); 29 — S. litorea (Fallén). 15 — after Ozerov &amp; Krivosheina, 2019, fig. 1; 20, 24 — after Ozerov &amp; Krivosheina, 2020, figs 3, 9; 27 — after Ozerov, 2013, fig.13. Рис. 14–29. Стернит 4 самца Scathophaga spp.: 14 — S. apicalis (Curtis); 15 — S. buryatica Ozerov et Krivosheina; 16 — S. calida (Haliday); 17 — S. cordylurina (Holmgren); 18 — S. crinita (Coquillett); 19 — S. dasythrix (Becker); 20 — S. decipiens (Haliday); 21 — S. exalata Ozerov; 22 — S. furcata (Say); 23 — S. incola (Becker); 24 — S. incompleta Ozerov et Krivosheina; 25 — S. inquinata (Meigen); 26 — S. intermedia (Walker); 27 — S. karelica Ozerov; 28 — S. lapponica (Ringdahl); 29 — S. litorea (Fallén). 15 — по Ozerov &amp; Krivosheina, 2019, fig. 1; 20, 24 — по Ozerov &amp; Krivosheina, 2020, figs 3, 9; 27 — по Ozerov, 2013, fig.13." pageId="17" pageNumber="218">Fig. 24</figureCitation>
). Male sternite 5 with moderately long and narrow lobes, with a small projection medially between lobes (
<figureCitation id="1379D0F57378BB455BFD521CFADFFCA4" box="[1253,1333,814,838]" captionStart="76 77 78" captionStartId="20.[400,461,1797,1855]" captionText="76 77 78 Figs 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) and S. inquinata (Meigen) (76–78): 70, 73, 76 — male sternite 5; 71, 74, 77 — epandrium, cercal plate and surstyli, lateral view; 72, 75, 78 — same, dorsal view. 73–75 — after Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13. Рис. 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) и S. inquinata (Meigen) (76–78): 70, 73, 76 — стернит 5 самца; 71, 74, 77 — Эпандрий, церки и сурстили, сбоку; 72, 75, 78 — то же, сверху. 73–75 — по Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13." pageId="17" pageNumber="218">Fig. 73</figureCitation>
). Cercal plate longer than surstyli and narrowing to a slender incurved tip (
<figureCitation id="1379D0F57378BB455CA0525BFBD9FC62" box="[952,1075,872,897]" captionStart="76 77 78" captionStartId="20.[400,461,1797,1855]" captionText="76 77 78 Figs 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) and S. inquinata (Meigen) (76–78): 70, 73, 76 — male sternite 5; 71, 74, 77 — epandrium, cercal plate and surstyli, lateral view; 72, 75, 78 — same, dorsal view. 73–75 — after Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13. Рис. 70–78. Scathophaga incola (Becker) (70–72), S. incompleta Ozerov et Krivosheina (73–75) и S. inquinata (Meigen) (76–78): 70, 73, 76 — стернит 5 самца; 71, 74, 77 — Эпандрий, церки и сурстили, сбоку; 72, 75, 78 — то же, сверху. 73–75 — по Ozerov &amp; Krivosheina, 2020, figs 10, 12, 13." pageId="17" pageNumber="218">Figs 74, 75</figureCitation>
). Aedeagus as in
<figureCitation id="1379D0F57378BB455BF3525BFAACFC63" box="[1259,1350,873,897]" captionStart="Figs 130–137" captionStartId="41.[226,263,1885,1904]" captionTargetBox="[229,1357,318,1859]" captionTargetId="figure-7@41.[209,1380,303,1879]" captionTargetPageId="41" captionText="Figs 130–137. Aedeagus of Scathophaga spp., lateral view: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — after Ozerov &amp; Krivosheina, 2019, fig. 2; 131, 132 — after Ozerov &amp; Krivosheina, 2020, figs 5, 11. Рис. 130–137. Эдеагус Scathophaga spp., сбоку: 130 — S. buryatica Ozerov et Krivosheina; 131 — S. decipiens (Haliday); 132 — S. incompleta Ozerov et Krivosheina; 133 — S. lapponica (Ringdahl); 134 — S. mollis (Becker); 135 — S. nigripalpis (Becker); 136 — S. obscura (Fallén); 137 — S. tinctinervis (Becker). 130 — по Ozerov &amp; Krivosheina, 2019, fig. 2; 131, 132 — по Ozerov &amp; Krivosheina, 2020, figs 5, 11." figureDoi="http://doi.org/10.5281/zenodo.13178494" httpUri="https://zenodo.org/record/13178494/files/figure.png" pageId="17" pageNumber="218">Fig. 132</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3589FFB7378BB455C4A52B4FC63FC3B" pageId="17" pageNumber="218" type="distribution">
<paragraph id="8BFDCC707378BB455C4A52B4FC63FC3B" blockId="17.[812,1399,315,985]" pageId="17" pageNumber="218">
DISTRIBUTION.
<collectingCountry id="F3558CE07378BB455B3B52B4FB85FC7C" box="[1059,1135,902,926]" name="Russia" pageId="17" pageNumber="218">Russia</collectingCountry>
:
<collectingRegion id="498602927378BB455B9952B4FB32FC7C" box="[1153,1240,902,926]" country="Russia" name="Adygeya" pageId="17" pageNumber="218">Adygea</collectingRegion>
, Crimea,
<collectingRegion id="498602927378BB455A5652B4FC99FC5E" country="Russia" name="Dagestan" pageId="17" pageNumber="218">Dagestan</collectingRegion>
,
<collectingRegion id="498602927378BB455C985291FC07FC59" box="[896,1005,931,955]" country="Russia" name="Kalmykiya" pageId="17" pageNumber="218">Kalmykia</collectingRegion>
. —
<collectingCountry id="F3558CE07378BB455B045291FB7EFC59" box="[1052,1172,931,955]" name="Azerbaijan" pageId="17" pageNumber="218">Azerbaijan</collectingCountry>
,
<collectingCountry id="F3558CE07378BB455BB95291FADCFC59" box="[1185,1334,931,955]" name="Turkmenistan" pageId="17" pageNumber="218">Turkmenistan</collectingCountry>
(
<emphasis id="B93610627378BB455A5F5291FC92FC3B" bold="true" pageId="17" pageNumber="218">first record</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>

117
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<subSubSection id="E58698F4FFB09F2CFC2CFDFF0FC856AC" pageId="4" pageNumber="434" type="nomenclature">
<paragraph id="AD23CB7FFFB09F2CFC2CFDFF0E405691" blockId="4.[905,1305,634,687]" box="[905,1305,634,658]" pageId="4" pageNumber="434">
<heading id="F66B7C13FFB09F2CFC2CFDFF0E405691" box="[905,1305,634,658]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="434" reason="8">
<taxonomicName id="6A9CB0FCFFB09F2CFC2CFDFF0E405691" ID-CoL="6T8QC" authority="(Dejean, 1828)" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[905,1305,634,658]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFC2CFDFF0F375691" box="[905,1134,634,658]" italics="true" pageId="4" pageNumber="434">Orthomus hispanicus</emphasis>
(Dejean, 1828)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFBB7FD120FC856AC" blockId="4.[905,1305,634,687]" box="[1042,1169,663,687]" pageId="4" pageNumber="434">Figs 12–15.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB09F2DFCF7FD4109465782" lastPageId="5" lastPageNumber="435" pageId="4" pageNumber="434" type="description">
<paragraph id="AD23CB7FFFB09F2CFCF7FD4108CF5729" blockId="4.[812,1398,708,810]" pageId="4" pageNumber="434">Dorsum brown dark, almost black, body length 7.0–9.0 mm, males shiny with fine microreticulation, females somewhat dull with visible microreticulation. Eye convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47F8A208E25399" blockId="4.[189,1399,1831,2070]" pageId="4" pageNumber="434">
Figs 12–22.
<taxonomicName id="6A9CB0FCFFB09F2CFEF4F8A20A975339" box="[337,462,1831,1850]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB09F2CFEF4F8A20AFE5339" box="[337,423,1831,1850]" italics="true" pageId="4" pageNumber="434">Orthomus</emphasis>
spp
</taxonomicName>
, specimens from
<collectingCountry id="D58B8BEFFFB09F2CFDCFF8A209C75339" box="[618,670,1831,1850]" name="Spain" pageId="4" pageNumber="434">Spain</collectingCountry>
: 12–15 —
<taxonomicName id="6A9CB0FCFFB09F2CFCA7F8A208235339" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[770,890,1831,1850]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFCA7F8A208235339" box="[770,890,1831,1850]" italics="true" pageId="4" pageNumber="434">O. hispanicus</emphasis>
</taxonomicName>
; 16–18 —
<taxonomicName id="6A9CB0FCFFB09F2CFC44F8A20F615339" authorityName="Martinez-Saez" authorityYear="1873" box="[993,1080,1831,1850]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB09F2CFC44F8A20F615339" box="[993,1080,1831,1850]" italics="true" pageId="4" pageNumber="434">O. perezii</emphasis>
</taxonomicName>
; 19–22 —
<taxonomicName id="6A9CB0FCFFB09F2CFB3BF8A20FAB5339" authorityName="Jeanne" authorityYear="1974" box="[1182,1266,1831,1850]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB09F2CFB3BF8A20FAB5339" box="[1182,1266,1831,1850]" italics="true" pageId="4" pageNumber="434">O. aubryi</emphasis>
</taxonomicName>
; 12, 16, 19 — habitus; 13 — metepisterna; 14, 17, 20–21 — median lobe of aedeagus; 15, 18, 22 — genitalia of female; 12, 14, 16–17, 19–20 — dorsal view; 13 — right lateral view; 21 — left lateral view; 14 — sierra de Larouco, Orense; 15 — Candelario, Salamanca; 17 — Campisábalos, Guadalajara; 18 — Valdefuentes de Sangusín, Salamanca; 19–21 — Moncayo Peak, Zaragoza; 22 — Linas de Broto, Huesca; 15, 18, 22 — after
<bibRefCitation id="C90DB68EFFB09F2CFF4EF8020A325399" author="Ortuno V. M." box="[235,363,1927,1946]" pageId="4" pageNumber="434" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. Scale bars: 1.2 mm (Fig. 12); 1.3 mm (Fig. 16); 1.5 mm (Fig. 19).
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47F826088E5C15" blockId="4.[189,1399,1831,2070]" pageId="4" pageNumber="434">
Рис. 12–22.
<taxonomicName id="6A9CB0FCFFB09F2CFEE8F8260A9E53B5" box="[333,455,1955,1974]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB09F2CFEE8F8260AFA53B5" box="[333,419,1955,1974]" italics="true" pageId="4" pageNumber="434">Orthomus</emphasis>
spp
</taxonomicName>
, особи иЗ
<collectingCountry id="D58B8BEFFFB09F2CFD84F826092B53B5" box="[545,626,1955,1974]" name="Spain" pageId="4" pageNumber="434">Испании</collectingCountry>
: 12–15 —
<taxonomicName id="6A9CB0FCFFB09F2CFD75F826081C53B5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[720,837,1955,1974]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFD75F826081C53B5" box="[720,837,1955,1974]" italics="true" pageId="4" pageNumber="434">O. hispanicus</emphasis>
</taxonomicName>
; 16–18 —
<taxonomicName id="6A9CB0FCFFB09F2CFC01F82608A153B6" authorityName="Martinez-Saez" authorityYear="1873" box="[932,1016,1955,1974]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB09F2CFC01F82608A153B6" box="[932,1016,1955,1974]" italics="true" pageId="4" pageNumber="434">O. perezii</emphasis>
</taxonomicName>
; 19–22 —
<taxonomicName id="6A9CB0FCFFB09F2CFBF3F8260FFE53B5" authorityName="Jeanne" authorityYear="1974" box="[1110,1191,1955,1974]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB09F2CFBF3F8260FFE53B5" box="[1110,1191,1955,1974]" italics="true" pageId="4" pageNumber="434">O. aubryi</emphasis>
</taxonomicName>
; 12, 16, 19 — внеШний вид; 13 — Эпистерн Заднегруди; 14, 17, 20–21 — среднЯЯ долЯ Эдеагуса; 15, 18, 22 — гениталии самки; 12, 14, 16–17, 19–20 — сверху; 13 –сбоку справа; 21 –сбоку слева; 14 — Сьерра- де- Ларуко, Оренсе; 15 — Канделарио, Саламанка; 17 — Камписабалос, Гвадалахара; 18 — ВальдефуЭнтес- де- Сангусин, Саламанка; 19–21 — пик Монкайо, Сарагоса; 22 — Линас- де- Брото, УЭска; 15, 18, 22 — по Ortuсo [1996]. МасШтаб: 1,2 мм (Рис. 12); 1,3 мм (Рис. 16); 1,5 мм (Рис. 19).
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FEB909E955A2" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Pronotum shape variable, either maximum width at apical 3/4 (more trapezoid) or at middle (more quadrate) (Fig. 12); posterior angle obtuse, slightly toothed, basis not emarginate. Foveae well marked and usually punctured.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FE2309B55624" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Elytra moderately widened backward, side narrowed toward basis, humerus toothed, striae smooth or with traces of punctures, intervals flat in females and slightly convex in males; disk with distal seta on 3rd stria but it often varies as it is found in interval 3, close to stria 2 or may be absent.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FDAE09CF56AF" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">
Prosternal process forming an obtuse ridge, rounded and emarginate on bottom, ventral segments punctured including metepisterna (Fig. 13), this as long as wide (L/
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). Males with mesotibia somewhat enlarged apically and metatibia with traces of crenulation in apical internal side.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FD3409835732" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Median lobe of aedeagus slightly bent ventrally; apical shaft asymmetrical, more narrowed in left side and prolonged in the right side (Fig. 14); membranous sac devoid of teeth and only moderately open on dorsal view. Spermatheca dilat- ed apically, spermathecal duct relatively short (Fig. 15)</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FCB309465782" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">
Northern half of
<collectingCountry id="D58B8BEFFFB19F2DFE30FCB30AB0574F" box="[405,489,822,844]" name="Portugal" pageId="5" pageNumber="435">Portugal</collectingCountry>
, Galicia, Cantabrian Mountains, Northern Sistema Ibérico, and Sistema Central from Serra da Estrela to sierra de Gredos.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="E58698F4FFB19F2DFE9EF904097B52B5" pageId="5" pageNumber="435" type="nomenclature">
<paragraph id="AD23CB7FFFB19F2DFE9EF90409D1529A" blockId="5.[189,775,1665,2073]" box="[315,648,1665,1689]" pageId="5" pageNumber="435">
<heading id="F66B7C13FFB19F2DFE9EF90409D1529A" box="[315,648,1665,1689]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="435" reason="8">
<taxonomicName id="6A9CB0FCFFB19F2DFE9EF90409D1529A" ID-CoL="74WQB" authority="Jeanne, 1974" authorityName="Jeanne" authorityYear="1974" box="[315,648,1665,1689]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFE9EF9040AAD529A" box="[315,500,1665,1689]" italics="true" pageId="5" pageNumber="435">Orthomus aubryi</emphasis>
<bibRefCitation id="C90DB68EFFB19F2DFE5EF90709D1529A" author="Jeanne C." box="[507,648,1665,1689]" pageId="5" pageNumber="435" pagination="66 - 71" refId="ref16762" refString="Jeanne C. 1974. Carabiques nouveaux (5 eme note) // Bull. Soc. ent. France. Vol. 79. P. 66 - 71." type="journal article" year="1974">Jeanne, 1974</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFE07F91B097B52B5" blockId="5.[189,775,1665,2073]" box="[418,546,1694,1718]" pageId="5" pageNumber="435">Figs 19–22.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB19F2DFF47F946088F5128" pageId="5" pageNumber="435" type="description">
<paragraph id="AD23CB7FFFB19F2DFF47F946094D532A" blockId="5.[189,775,1665,2073]" pageId="5" pageNumber="435">Body length 8.5–10.0 mm. Dorsum dark piceous, appendages reddish. Antennomeres pubescent from apical third of antennomere 3 to 11, reaching the basis of elytra. Eye convex with long oblique tempora.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F8AB0A6153E7" blockId="5.[189,775,1665,2073]" pageId="5" pageNumber="435">Pronotum with anterior angle moderately projecting forward, anterior basis almost regularly curved; maximum width before middle (Fig. 19), from here side is slightly arcuate and narrowed toward the obtuse posterior angle, that has a small protruding tooth. Basal foveae finely punctured, the internal deep, linear or curved, the external moderately impressed or almost faint.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F86C0F1E5568" blockId="5.[189,775,1665,2073]" lastBlockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
Elytra relatively long (L/
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), almost parallel, humeral tooth small or almost absent, striae smooth, intervals moderately convex, parascutellar stria somewhat short, distal dorsal pore adjoining stria 2.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FEF40E2E55D4" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">Episternae and first ventrites moderately punctured. Metepisternae as long as wide in the anterior margin (L/A 1.05). Male mesotibia slightly dilated apically, metatibia distally curved and with traces of crenulation in the distal inner side.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FE5E0F105765" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
Apical shaft of median lobe of aedeagus elongate, somewhat more narrowed to the left (Fig. 20); on lateral view the median lobe of aedeagus is almost straight, the apex slightly curved upward (Fig. 21); membranous area (ostium) prolonged toward basis. The spermatheca (Fig. 22) show a similar shape to that described for
<taxonomicName id="6A9CB0FCFFB19F2DFB82FDE40FF15675" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1063,1192,609,630]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB19F2DFB82FDE40FF15675" box="[1063,1192,609,630]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
</taxonomicName>
but the duct is longer. Differences in the length of the spermathecal duct led
<bibRefCitation id="C90DB68EFFB19F2DFA96FDFE083556AF" author="Ortuno V. M." pageId="5" pageNumber="435" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
to place
<taxonomicName id="6A9CB0FCFFB19F2DFC65FD130F4356A8" authorityName="Jeanne" authorityYear="1974" box="[960,1050,662,683]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFC65FD130F4356A8" box="[960,1050,662,683]" italics="true" pageId="5" pageNumber="435">O. aubryi</emphasis>
</taxonomicName>
near the
<taxonomicName id="6A9CB0FCFFB19F2DFBD5FD130FAE56A8" authorityName="Fairmaire" authorityYear="1872" box="[1136,1271,662,683]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="planidorsis">
<emphasis id="9FE8176DFFB19F2DFBD5FD130FAE56A8" box="[1136,1271,662,683]" italics="true" pageId="5" pageNumber="435">O. planidorsis</emphasis>
</taxonomicName>
group. As
<taxonomicName id="6A9CB0FCFFB19F2DFAFAFD13083356C5" authorityName="Jeanne" authorityYear="1974" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFAFAFD13083356C5" italics="true" pageId="5" pageNumber="435">O. aubryi</emphasis>
</taxonomicName>
might enter in contact with
<taxonomicName id="6A9CB0FCFFB19F2DFBC8FD340FB756C5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1133,1262,689,710]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB19F2DFBC8FD340FB756C5" box="[1133,1262,689,710]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB19F2DFABFFD340E2F56C5" authorityName="Martinez-Saez" authorityYear="1873" box="[1306,1398,689,710]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB19F2DFABFFD340E2F56C5" box="[1306,1398,689,710]" italics="true" pageId="5" pageNumber="435">O. perezii</emphasis>
</taxonomicName>
in the west (Basque Mts., north Sistema Ibérico), and with
<taxonomicName id="6A9CB0FCFFB19F2DFAFAFD4908CE56F8" authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="planidorsis">
<emphasis id="9FE8176DFFB19F2DFAFAFD4908CE56F8" italics="true" pageId="5" pageNumber="435">O. planidorsis</emphasis>
</taxonomicName>
in the east (Central Pyrenees), the geographic distribution doesn’t help in solving the affinities of the species. Molecular markers are needed to seed light into this question. Basque Mountains, Northern Sistema Ibérico (Moncayo Peak), central and western Pyrenees.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FCEE0F7B579F" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">Groups of species with metepisterna longer that wider and narrowed posteriorly</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FC240FAD57B5" blockId="5.[812,1399,315,1324]" box="[850,1268,929,951]" pageId="5" pageNumber="435">
Iberian taxa: Group of
<taxonomicName id="6A9CB0FCFFB19F2DFB90FC240FAD57B5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1077,1268,929,950]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB19F2DFB90FC240FAD57B5" box="[1077,1268,929,950]" italics="true" pageId="5" pageNumber="435">Orthomus barbarus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FC3E088F5128" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
This group is made up by
<emphasis id="9FE8176DFFB19F2DFBF6FC39088D57E8" italics="true" pageId="5" pageNumber="435">
<taxonomicName id="6A9CB0FCFFB19F2DFBF6FC390F9457D3" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1107,1229,955,977]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="barbarus">O. barbarus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB19F2DFB72FC390E2A57D3" box="[1239,1395,955,977]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="velocissimus">O. velocissimus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB19F2DFC89FC53088957E8" authorityName="Breit" authorityYear="1933" box="[812,976,982,1003]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="formenterrae">O. formenterrae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6A9CB0FCFFB19F2DFBA9FC530FC557E8" authorityName="Mateu et Colas" authorityYear="1954" box="[1036,1180,982,1003]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB19F2DFBA9FC530FC557E8" box="[1036,1180,982,1003]" italics="true" pageId="5" pageNumber="435">O. penibeticus</emphasis>
</taxonomicName>
. Males of this group show a distally dilated mesotibia and a metatibia variably crenulate in the distal half. The female genitalia show a long spermathecal duct followed by a long spermatheca [
<bibRefCitation id="C90DB68EFFB19F2DFA8CFBA308065055" author="Ortuno V. M." pageId="5" pageNumber="435" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño, 1996</bibRefCitation>
;
<bibRefCitation id="C90DB68EFFB19F2DFCCEFBC40F195055" author="Pupier R. &amp; Coulon J." box="[875,1088,1089,1111]" pageId="5" pageNumber="435" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
]. Male genitalia are rather variable as the apical shaft of the median lobe of the aedeagus is triangular in some taxa, with rounded or acute tip, whereas it is truncate, with or without a tooth at tip. Taxa of this group are found below a hypothetical line that runs from the southwest of
<collectingCountry id="D58B8BEFFFB19F2DFCD2FB4308F650DF" box="[887,943,1222,1244]" name="Georgia" pageId="5" pageNumber="435">Iberia</collectingCountry>
(Algarve and western Andalusia) to the eastern Pyrenees and south
<collectingCountry id="D58B8BEFFFB19F2DFC54FB640F6C50F5" box="[1009,1077,1249,1270]" name="France" pageId="5" pageNumber="435">France</collectingCountry>
, and also occupies the southernmost archipelago of the Balearic Islands, the Pitiusas (Ibiza and Formentera).
</paragraph>
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<taxonomicName id="6A9CB0FCFFB19F2DFEB5FC1009EA57AE" ID-CoL="6SWQD" authority="Martinez-Saez, 1873" authorityName="Martinez-Saez" authorityYear="1873" box="[272,691,917,941]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB19F2DFEB5FC100A9557AE" box="[272,460,917,941]" italics="true" pageId="5" pageNumber="435">Orthomus perezii</emphasis>
Martínez-Sáez, 1873
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<paragraph id="AD23CB7FFFB19F2DFE07FC37097B57C9" blockId="5.[189,775,917,1645]" box="[418,546,946,970]" pageId="5" pageNumber="435">Figs 16–18.</paragraph>
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<paragraph id="AD23CB7FFFB19F2DFF47FC5D0A9A5020" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">
Dorsum brown dark piceous, body size smaller than that of
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<taxonomicName id="6A9CB0FCFFB19F2DFF73FC770A045004" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[214,349,1010,1031]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">O. hispanicus</taxonomicName>
,
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6.0–7.5 mm, males shiny, females mat. Eye less convex, tempora long.
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<paragraph id="AD23CB7FFFB19F2DFF47FBAD09C050A4" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Pronotum transverse, narrowed anteriad, maximum width at middle, slightly constricted backward (Fig. 16); posterior angle obtuse, no or at most, slightly toothed; basis not emarginate; foveae well marked and usually punctured, space between side and external fovea notably convex.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FB2809FB5144" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Elytra relatively short and oval, sides slightly widened backward, narrowed close to basis, basis forming at humerus a fold that doesn’t protrude outside; parascutellar stria normally developed with a proximal hairy pore; intervals convex, striae smooth, second disk pore located variably: on the 2nd stria, in the interval between striae 2 and 3, close to stria 3.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FAC8085E51B0" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Prosternal process rounded, ventral segments finely punctured and with rugose areas, metepisterna as long as wide (L/A 1.0). Male mesotibia somewhat enlarged apically, metatibia with crenulation moderately developed in apical internal side.</paragraph>
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Apical shaft of median lobe of aedeagus well developed, as wide as long, evenly rounded to tip (Fig. 17); on lateral view almost straight; membranous sac open along the entire dorsal face. Spermatheca as described in
<taxonomicName id="6A9CB0FCFFB19F2DFDEFF98D0996521F" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[586,719,1543,1565]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB19F2DFDEFF98D0996521F" box="[586,719,1543,1565]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
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, with a notably dilated apical end (Fig. 1).
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F9B80950526E" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Sistema Central, Sistema Iberico (N and S), North Meseta, northern part of South Meseta.</paragraph>
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<emphasis id="9FE8176DFFB19F2DFCF8FAC50FC5515B" box="[861,1180,1344,1368]" italics="true" pageId="5" pageNumber="435">Orthomus barbarus barbarus</emphasis>
(Dejean, 1828)
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<paragraph id="AD23CB7FFFB19F2DFCF7FA070E4E51EB" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Dorsum between black and dark brown, males shiny, females less shiny or dull with isodiametric microreticulation. Body length 8.8–11.5 mm. Legs black, antennae and tarsi dark red. Eye notably convex, tempora short.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FA680E0752A1" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Pronotum shape variable, most commonly showing a trapezoid contour more narrowed forward and side entirely curved, maximum width at middle or so (Fig. 23), anterior angle not markedly protruding forward; posterior angle toothed; inner basal fovea linear, outer fovea from rugose to almost erased, punctures of moderate size and at least abundant in inner fovea, postangular field moderately convex.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7F9220E32530E" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Elytra almost parallel, slightly narrowed toward basis; humeral tooth well developed; striae smooth or vaguely punctured, intervals almost flat; discal pores on 3rd stria; parascutellar stria normally developed with proximal setose pore.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7F89708D05374" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">
Prosternal process rounded; ventral segments almost smooth, fine punctures are found on sides of last ventrites; metepisterna smooth, long and narrowed apicad, L/
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) (Fig. 24).
</paragraph>
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Median lobe of aedeagus on lateral view with apical region curved upward; apical shaft on dorsal view rather variable between localities but predominating an elongate triangle evenly narrowed to tip and straight (Fig. 25), or deviated to right [see
<bibRefCitation id="C90DB68EFFB19F2DFBBDF86D0FC453FE" author="Mateu J." box="[1048,1181,2023,2045]" pageId="5" pageNumber="435" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957</bibRefCitation>
]. In lateral view the median lobe is somewhat ventrally bent (Fig. 26). The spermathecal duct is rather long, even longer than the spermatheca (Fig. 27).
</paragraph>
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The species is distributed in the Mediterranean coast from eastern Almeria to south
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and also colonizes the South Meseta, Betic and Iberian chains and the River Ebro Basin.
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<mods:title id="45B99FEE5B98611F7775BAC16EFBB549">The genus Orthomus Chaudoir, 1838 in the Iberian Peninsula and Morocco</mods:title>
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<subSubSection id="E58698F4FFB29F2EFF4BFE55097B5606" pageId="6" pageNumber="436" type="nomenclature">
<paragraph id="AD23CB7FFFB29F2EFF4BFE55098C55EB" blockId="6.[189,775,464,792]" box="[238,725,464,488]" pageId="6" pageNumber="436">
<heading id="F66B7C13FFB29F2EFF4BFE55098C55EB" box="[238,725,464,488]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="436" reason="8">
<taxonomicName id="6A9CB0FCFFB29F2EFF4BFE5509DA55EB" ID-CoL="8LMSF" authority="(Breit, 1933)" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[238,643,464,488]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFF4BFE550AB455EB" box="[238,493,464,488]" italics="true" pageId="6" pageNumber="436">Orthomus formenterrae</emphasis>
(Breit, 1933)
</taxonomicName>
,
<emphasis id="9FE8176DFFB29F2EFD2AFE55098C55EB" bold="true" box="[655,725,464,488]" pageId="6" pageNumber="436">stat.n.</emphasis>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFE07FE68097B5606" blockId="6.[189,775,464,792]" box="[418,546,493,517]" pageId="6" pageNumber="436">Figs 28–31.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB29F2FFF47FD97090A5765" lastPageId="7" lastPageNumber="437" pageId="6" pageNumber="436" type="description">
<paragraph id="AD23CB7FFFB29F2EFF47FD97085F571B" blockId="6.[189,775,464,792]" pageId="6" pageNumber="436">
This species was described as
<taxonomicName id="6A9CB0FCFFB29F2EFDB0FD970AF05641" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="balearicus" subGenus="Orthomus" subSpecies="formenterrae">
<emphasis id="9FE8176DFFB29F2EFDB0FD970AF05641" italics="true" pageId="6" pageNumber="436">Pterostichus (Orthomus) balearicus formenterrae</emphasis>
</taxonomicName>
and lately considered a member of the
<taxonomicName id="6A9CB0FCFFB29F2EFF47FDCD0A00565F" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[226,345,583,605]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFF47FDCD0A00565F" box="[226,345,583,605]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
complex by
<bibRefCitation id="C90DB68EFFB29F2EFE79FDCD093A565E" author="Mateu J." box="[476,611,583,605]" pageId="6" pageNumber="436" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
, in which it was ranked as subspecies:
<taxonomicName id="6A9CB0FCFFB29F2EFE32FDE709CD5674" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[407,660,610,631]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="barbarus" subSpecies="formenterrae">
<emphasis id="9FE8176DFFB29F2EFE32FDE709CD5674" box="[407,660,610,631]" italics="true" pageId="6" pageNumber="436">O. barbarus formenterrae</emphasis>
</taxonomicName>
, a criterion that has been followed by authors. Due to the confusion between
<taxonomicName id="6A9CB0FCFFB29F2EFEBDFD1D0ACA56AF" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[280,403,663,685]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFEBDFD1D0ACA56AF" box="[280,403,663,685]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB29F2EFE6EFD1D093C56AF" authorityName="Chaudoir" authorityYear="1859" box="[459,613,663,685]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="trapezicollis">
<emphasis id="9FE8176DFFB29F2EFE6EFD1D093C56AF" box="[459,613,663,685]" italics="true" pageId="6" pageNumber="436">O. trapezicollis</emphasis>
</taxonomicName>
the species appeared in the catalogue of Zaballos and Jeanne [1994] as
<emphasis id="9FE8176DFFB29F2EFD4AFD370A9856E1" italics="true" pageId="6" pageNumber="436">
<taxonomicName id="6A9CB0FCFFB29F2EFD4AFD370AE456E1" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="trapezicollis" subSpecies="formenterrae">O. trapezicollis formenterrae</taxonomicName>
.
</emphasis>
In recent works, as that of Wrase et al. [2014] and
<bibRefCitation id="C90DB68EFFB29F2EFECEFD6D094B56FF" author="Bousquet Y." box="[363,530,743,765]" pageId="6" pageNumber="436" pagination="469 - 521" refId="ref16324" refString="Bousquet Y. 2017. Tribe Pterostichini // Lobl L., Lobl I. (eds.). Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Leiden: Brill. P. 469 - 521." type="book chapter" year="2017">Bousquet [2017]</bibRefCitation>
, it is again considered a subspecies of
<taxonomicName id="6A9CB0FCFFB29F2EFEEAFC870A925714" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[335,459,770,791]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFEEAFC870A925714" box="[335,459,770,791]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
. The criterion put forward by
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF5AF9740AD95307" blockId="6.[255,384,1770,1803]" box="[255,384,1770,1803]" pageId="6" pageNumber="436">
<emphasis id="9FE8176DFFB29F2EFF5AF9740AD95307" bold="true" box="[255,384,1770,1803]" pageId="6" pageNumber="436">29 33</emphasis>
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFC89FEBE0F255585" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">
Zaballos and Jeanne was possibly based on the truncate apical shaft of the median lobe of aedeagus, that is not found in the populations of
<taxonomicName id="6A9CB0FCFFB29F2EFC5AFEF40F2E5585" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1023,1143,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC5AFEF40F2E5585" box="[1023,1143,369,390]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FE0E08A55674" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">
This taxon is herewith considered as a separate species due to the truncate penis and the isolation of Ibiza and Formentera. The truncate penis shape relates it to some populations of the
<taxonomicName id="6A9CB0FCFFB29F2EFC44FE590F2055F3" box="[993,1145,475,497]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB29F2EFC44FE590F2055F3" box="[993,1145,475,497]" italics="true" pageId="6" pageNumber="436">O. velocissimus</emphasis>
</taxonomicName>
complex, whereas a close relationship to the
<taxonomicName id="6A9CB0FCFFB29F2EFC46FE730F025608" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[995,1115,502,523]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC46FE730F025608" box="[995,1115,502,523]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
complex is not supported by this character. The relatively shorter metepisterna of
<taxonomicName id="6A9CB0FCFFB29F2EFA96FD9408D65642" baseAuthorityName="Breit" baseAuthorityYear="1933" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFA96FD9408D65642" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
, in comparison to that of
<taxonomicName id="6A9CB0FCFFB29F2EFB36FDA90E755643" box="[1171,1324,555,577]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB29F2EFB36FDA90E755643" box="[1171,1324,555,577]" italics="true" pageId="6" pageNumber="436">O. velocissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB29F2EFAFAFDA908DD5658" baseAuthorityName="Dejean" baseAuthorityYear="1828" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFAFAFDA908DD5658" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
, is other character that supports ranking this taxon as a separate species.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FDF90F8A56C4" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">
Dorsum dark brown, males shiny, females dull due to strong microreticulation. Body length
<quantity id="6A64669AFFB29F2EFB08FD130FA156A8" box="[1197,1272,662,683]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="436" unit="mm" value="10.0">10 mm</quantity>
. Appendices reddish brown. Eye convex, tempora short.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FD490E2F5714" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">Pronotum notably narrowed forward, almost straight backward, anterior angle not much protruding ahead, posterior angle slightly obtuse, maximum pronotum width after middle</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF47F8A409AC5397" blockId="6.[189,1399,1825,2064]" pageId="6" pageNumber="436">
Figs 23–35.
<taxonomicName id="6A9CB0FCFFB29F2EFEEBF8A40A935336" box="[334,458,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFEEBF8A40AFD5337" box="[334,420,1825,1844]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
, specimens from
<collectingCountry id="D58B8BEFFFB29F2EFDC7F8A409CF5337" box="[610,662,1825,1844]" name="Spain" pageId="6" pageNumber="436">Spain</collectingCountry>
: 23–27 —
<taxonomicName id="6A9CB0FCFFB29F2EFD53F8A408395337" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[758,864,1825,1844]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFD53F8A408395337" box="[758,864,1825,1844]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
; 28–31 —
<taxonomicName id="6A9CB0FCFFB29F2EFC67F8A40F175337" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[962,1102,1825,1844]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFC67F8A40F175337" box="[962,1102,1825,1844]" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
; 32–35 —
<taxonomicName id="6A9CB0FCFFB29F2EFB14F8A40E775337" authorityName="Mateu et Colas" authorityYear="1954" box="[1201,1326,1825,1844]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB29F2EFB14F8A40E775337" box="[1201,1326,1825,1844]" italics="true" pageId="6" pageNumber="436">O. penibeticus</emphasis>
</taxonomicName>
; 23, 28, 32 — habitus; 24 — metepisterna; 25–26, 29–30, 33–34 — median lobe of aedeagus; 27, 31, 35 — genitalia of female; 23, 25, 28–29, 32 — dorsal view; 24, 26, 30, 34 — left lateral view; 25–26 — Cartagena,
<collectingRegion id="6F58059DFFB29F2EFCBBF8D408065367" box="[798,863,1873,1892]" country="Spain" name="Murcia" pageId="6" pageNumber="436">Murcia</collectingRegion>
; 27 —
<collectingRegion id="6F58059DFFB29F2EFC3AF8D408B85367" box="[927,993,1873,1892]" country="Spain" name="Madrid" pageId="6" pageNumber="436">Madrid</collectingRegion>
; 29–30 — Sa-Negreta, Ibiza; 31 — Formentera Island; 32 - Almería (possibly near Adra); 33–34 –Lanjarón, Granada; 35 — Puerto del Suspiro del Moro, Sierra Nevada, Granada; 27, 31, 35 — after
<bibRefCitation id="C90DB68EFFB29F2EFE86F8040AFA5397" author="Ortuno V. M." box="[291,419,1921,1940]" pageId="6" pageNumber="436" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. Scale bars: 1.0 mm (Figs 23, 28, 32).
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF47F8180E5F5C13" blockId="6.[189,1399,1825,2064]" pageId="6" pageNumber="436">
Рис. 23–35.
<taxonomicName id="6A9CB0FCFFB29F2EFEF2F8180A8F53B3" box="[343,470,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFEF2F8180AF453B3" box="[343,429,1949,1968]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
, особи иЗ
<collectingCountry id="D58B8BEFFFB29F2EFD9AF81809C953B3" box="[575,656,1949,1968]" name="Spain" pageId="6" pageNumber="436">Испании</collectingCountry>
:
<taxonomicName id="6A9CB0FCFFB29F2EFD3AF818084653B3" box="[671,799,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFD3AF81809AC53B3" box="[671,757,1949,1968]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
: 23–27 —
<taxonomicName id="6A9CB0FCFFB29F2EFC2FF81808AE53B3" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[906,1015,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC2FF81808AE53B3" box="[906,1015,1949,1968]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
; 28–31 —
<taxonomicName id="6A9CB0FCFFB29F2EFBC1F8180FAA53B3" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[1124,1267,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFBC1F8180FAA53B3" box="[1124,1267,1949,1968]" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
; 32–35 —
<taxonomicName id="6A9CB0FCFFB29F2EFAC4F8180A4653CB" authorityName="Mateu et Colas" authorityYear="1954" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB29F2EFAC4F8180A4653CB" italics="true" pageId="6" pageNumber="436">O. penibeticus</emphasis>
</taxonomicName>
; 23, 28, 32 — внеШний вид; 24 — Эпистерн Заднегруди; 25–26, 29–30, 33–34 — среднЯЯ долЯ Эдеагуса; 27, 31, 35 — гениталии самки; 23, 25, 28–29, 32 — сверху; 24, 26, 30, 34 –сбоку слева; 25–26 — Картахена, МурсиЯ; 27 —
<collectingRegion id="6F58059DFFB29F2EFB09F8480FAC53E3" box="[1196,1269,1997,2016]" country="Spain" name="Madrid" pageId="6" pageNumber="436">Мадрид</collectingRegion>
; 29–30 — Са- Негрета, Ибица; 31 — о- в Форментера; 32 — АльмериЯ (воЗможно, недалеко от Адры); 33–34 — Ланхарон, Гранада; 35 — ПуЭрто- дель- Суспиро- дель- Моро, Сьерра- Невада, Гранада; 27, 31, 35 — по
<bibRefCitation id="C90DB68EFFB29F2EFC9FF87808E25C13" author="Ortuno V. M." box="[826,955,2045,2064]" pageId="6" pageNumber="436" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. МасШтаб: 1,0 мм (Рис. 23, 28, 32).
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF18FEBE09AB55A2" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">(Fig. 28); outer fovea moderately impressed, inner fovea deep and linear or curved; fine punctures in both foveae, postangular field moderately convex; posterior angle without tooth; basal posterior border of pronotum not emarginate.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FE23090A5624" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Elytra almost parallel, regularly narrowed apicad (L/
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), basis large and with a clear fold in the humeral region that forms a moderate tooth, disk pores on stria 3, parascutellar stria normally developed, striae smooth, intervals somewhat convex in males and flat in females.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FDAE0A6B56C4" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Prosternal process rounded on bottom surface and forming a vertical obtuse ridge; ventrites almost smooth, punctures scarce, only somewhat rugose on sides; metepisterna smooth, relatively short (L/
<geoCoordinate id="C8A8ADB8FFB39F2FFE66FDFE09555692" box="[451,524,635,657]" degrees="1.05" direction="west" orientation="longitude" pageId="7" pageNumber="437" precision="555" value="-1.05">W 1.05</geoCoordinate>
) and narrowed backward. Male mesotibia dilated in apical half, metatibia with traces of crenulation.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FD4E090A5765" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Median lobe of aedeagus moderately curved ventrally (Fig. 30); apical shaft forming a well-developed triangle more narrowed to tip in left side (Fig. 29), truncate at tip and slightly sinuate at middle; membranous opening long, only closed near basal bulb. Characters of female genitalia as in
<emphasis id="9FE8176DFFB39F2FFD4AFCB30A4C5765" italics="true" pageId="7" pageNumber="437">
O,
<taxonomicName id="6A9CB0FCFFB39F2FFF18FCD40A4C5765" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[189,277,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">barbarus</taxonomicName>
</emphasis>
, as duct is rather long (Fig. 31).
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="AD23CB7FFFB39F2FFF6FFCFF09A05791" blockId="7.[189,776,890,2072]" box="[202,761,890,914]" pageId="7" pageNumber="437">
<heading id="F66B7C13FFB39F2FFF6FFCFF09A05791" box="[202,761,890,914]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="437" reason="8">
<taxonomicName id="6A9CB0FCFFB39F2FFF6FFCFF09F15791" authority="Mateu et Colas, 1954" authorityName="Mateu et Colas" authorityYear="1954" box="[202,680,890,914]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFF6FFCFF0AEF5791" box="[202,438,890,914]" italics="true" pageId="7" pageNumber="437">Orthomus penibeticus</emphasis>
Mateu et Colas, 1954
</taxonomicName>
,
<emphasis id="9FE8176DFFB39F2FFD16FCFF09A05791" bold="true" box="[691,761,890,914]" pageId="7" pageNumber="437">stat.n.</emphasis>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFE07FC1D097B57B3" blockId="7.[189,776,890,2072]" box="[418,546,920,944]" pageId="7" pageNumber="437">Figs 32–35.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB39F2FFF47FC380E735765" pageId="7" pageNumber="437" type="description">
<paragraph id="AD23CB7FFFB39F2FFF47FC380A065070" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
This species has also been currently considered a subspecies of
<taxonomicName id="6A9CB0FCFFB39F2FFEA4FC5D0A2157EF" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[257,376,983,1005]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFEA4FC5D0A2157EF" box="[257,376,983,1005]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
. However, the truncate shape of penis tip and the geographic isolation of its populations suggest that these have evolved to become a separate taxon from the surrounding
<taxonomicName id="6A9CB0FCFFB39F2FFE9DFBAD0A89503F" box="[312,464,1063,1085]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB39F2FFE9DFBAD0A89503F" box="[312,464,1063,1085]" italics="true" pageId="7" pageNumber="437">O. velocissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFDA4FBAD0921503F" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[513,632,1063,1085]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFDA4FBAD0921503F" box="[513,632,1063,1085]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
populations, a hypothesis to be corroborated with further studies based on other characters.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FBFD085150C0" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">Dorsum black or dark brown, appendices brown, males somewhat mat, females mat, microreticulation visible in both sexes, body length 10.0–11.7 mm. Eye convex, tempora short.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FB4D09E65160" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">Pronotum regularly curved from anterior to posterior basis but more narrowed anteriorly, maximum width at posterior 2/ 3 (Fig. 32), posterior angle obtuse and without tooth, posterior basis not emarginate; external fovea shallow, separate from side by a flat space, internal deep and linear; punctures on posterior basis fine and with low to moderate density.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FAE20AD051CE" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Elytra somewhat short, L/
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, slightly widened backwards, humerus with small tooth; striae unpunctured, intervals moderately convex, disk pores on stria 3, parascutellar stria well developed.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FA57092B528B" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Prosternal process rounded on bottom surface and forming an acute ridge from proepisterna to bottom surface; ventral face almost devoid of punctures, metepisterna smooth and longer than wider (L/
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–1.4). Male mesotibia dilat- ed apically, metatibia with moderate crenulation although
<bibRefCitation id="C90DB68EFFB39F2FFF18F9DD0A1A526E" author="Mateu J." box="[189,323,1623,1645]" pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
noted that the crenulation is strong, possibly because this character varies geographically.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F9080A8E5340" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Median lobe of aedeagus quite particular, resembling that of
<taxonomicName id="6A9CB0FCFFB39F2FFF7CF92D0A0252BF" authorityName="Lucas" authorityYear="1846" box="[217,347,1703,1725]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFB39F2FFF7CF92D0A0252BF" box="[217,347,1703,1725]" italics="true" pageId="7" pageNumber="437">O. abacoides</emphasis>
</taxonomicName>
from Maghreb, as the apical shaft forms a long triangle asymmetrically narrowed to the left and slightly truncate at tip (Fig. 33). In lateral view the median lobe is somewhat shifted upward (Fig. 34). The female genitalia are of the
<taxonomicName id="6A9CB0FCFFB39F2FFEA7F8970A225324" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[258,379,1810,1831]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFEA7F8970A225324" box="[258,379,1810,1831]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
<typeStatus id="722775DDFFB39F2FFE23F8970AE95324" box="[390,432,1810,1831]" pageId="7" pageNumber="437">type</typeStatus>
(Fig. 35), with a very long duct, longer than the spermatheca.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F8CD0AE853AE" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
In addition to the geographic separation,
<taxonomicName id="6A9CB0FCFFB39F2FFDDCF8CD085F535F" authorityName="Mateu et Colas" authorityYear="1954" box="[633,774,1863,1885]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFDDCF8CD085F535F" box="[633,774,1863,1885]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFF40F8E70A3C5374" authorityName="Lucas" authorityYear="1846" box="[229,357,1890,1911]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFB39F2FFF40F8E70A3C5374" box="[229,357,1890,1911]" italics="true" pageId="7" pageNumber="437">O. abacoides</emphasis>
</taxonomicName>
differ in a number of characters, especially in the female genitalia, what suggests that similarity in male genitalia is a homoplasy.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F8370E615643" blockId="7.[189,776,890,2072]" lastBlockId="7.[812,1399,315,871]" pageId="7" pageNumber="437">
The ranking of
<taxonomicName id="6A9CB0FCFFB39F2FFEDCF837095D53C4" authorityName="Mateu et Colas" authorityYear="1954" box="[377,516,1970,1991]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFEDCF837095D53C4" box="[377,516,1970,1991]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
as a separate species from
<taxonomicName id="6A9CB0FCFFB39F2FFF18F8480A6C53E1" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[189,309,1997,2018]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFF18F8480A6C53E1" box="[189,309,1997,2018]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
is based on male genitalia and the geographic isolation of
<taxonomicName id="6A9CB0FCFFB39F2FFE96F86D0A9853FF" authorityName="Mateu et Colas" authorityYear="1954" box="[307,449,2023,2045]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFE96F86D0A9853FF" box="[307,449,2023,2045]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="C90DB68EFFB39F2FFD97F86D09EE53FE" author="Mateu J." box="[562,695,2023,2045]" pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
considered that the distribution of
<taxonomicName id="6A9CB0FCFFB39F2FFE60F78709635C14" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[453,570,2050,2071]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFE60F78709635C14" box="[453,570,2050,2071]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
was deeply extended across Andalusia just by occupying the middle of the province of
<collectingCountry id="D58B8BEFFFB39F2FFCD2FED30890556F" box="[887,969,342,364]" name="Grenada" pageId="7" pageNumber="437">Granada</collectingCountry>
(form
<taxonomicName id="6A9CB0FCFFB39F2FFBABFED20F3F556F" authorityName="Mateu" authorityYear="1957" box="[1038,1126,343,364]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="transiens">
<emphasis id="9FE8176DFFB39F2FFBABFED20F3F556F" box="[1038,1126,343,364]" italics="true" pageId="7" pageNumber="437">transiens</emphasis>
</taxonomicName>
) and the eastern part of the province of Malaga (form
<taxonomicName id="6A9CB0FCFFB39F2FFB99FEF40FFE5585" authorityName="Jeanne" authorityYear="1981" box="[1084,1191,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="malacensis">
<emphasis id="9FE8176DFFB39F2FFB99FEF40FFE5585" box="[1084,1191,369,390]" italics="true" pageId="7" pageNumber="437">malacensis</emphasis>
</taxonomicName>
, see map of
<bibRefCitation id="C90DB68EFFB39F2FFA96FEF408CA55A3" author="Mateu J." pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957: 113</bibRefCitation>
), this hypothesis is not supported on geographic grounds; instead, it is simpler to consider populations of the so-called
<taxonomicName id="6A9CB0FCFFB39F2FFC2EFE4408BA55D5" authorityName="Mateu" authorityYear="1957" box="[907,995,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="transiens">
<emphasis id="9FE8176DFFB39F2FFC2EFE4408BA55D5" box="[907,995,449,470]" italics="true" pageId="7" pageNumber="437">transiens</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFBB1FE440F2655D5" authorityName="Jeanne" authorityYear="1981" box="[1044,1151,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="malacensis">
<emphasis id="9FE8176DFFB39F2FFBB1FE440F2655D5" box="[1044,1151,449,470]" italics="true" pageId="7" pageNumber="437">malacensis</emphasis>
</taxonomicName>
as part of the subspecies
<taxonomicName id="6A9CB0FCFFB39F2FFC89FE590F7855F3" authorityName="Mateu" authorityYear="1955" box="[812,1057,475,497]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="velocissimus" subSpecies="akbensis">
<emphasis id="9FE8176DFFB39F2FFC89FE590F7855F3" box="[812,1057,475,497]" italics="true" pageId="7" pageNumber="437">O. velocissimus akbensis</emphasis>
</taxonomicName>
, that is found in almost the whole basin of the River Guadalquivir. This last hypothesis was put forward by Zaballos and Jeanne [1994] and followed by Serrano in successive catalogues [2003, 2013, 2020].
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFCF7FDC30E735765" blockId="7.[812,1399,315,871]" pageId="7" pageNumber="437">
Accordingly,
<taxonomicName id="6A9CB0FCFFB39F2FFC72FDC30F385658" authorityName="Mateu et Colas" authorityYear="1954" box="[983,1121,582,603]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFC72FDC30F385658" box="[983,1121,582,603]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
would be a taxon distributed from the southern high slopes of Sierra Nevada (region of Las Alpujarras) to the Mediterranean coast, a relatively small region limited to west by the river Guadalfeo (Motril) and populations of
<taxonomicName id="6A9CB0FCFFB39F2FFC64FD340F0356C5" box="[961,1114,689,710]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB39F2FFC64FD340F0356C5" box="[961,1114,689,710]" italics="true" pageId="7" pageNumber="437">O. velocissimus</emphasis>
</taxonomicName>
, and to the east by the plain between Adra and the hills west of Almería city [
<bibRefCitation id="C90DB68EFFB39F2FFA95FD4908CF56F8" author="Mateu J." pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957: 101</bibRefCitation>
]; in this last city it is found the southernmost populations of
<taxonomicName id="6A9CB0FCFFB39F2FFC19FC840F6B5715" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[956,1074,769,790]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFC19FC840F6B5715" box="[956,1074,769,790]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
. This hypothesis deserves further corroboration, although at present, the study of male genitalia suffices to separate without doubt the parapatric populations of
<emphasis id="9FE8176DFFB39F2FFCE2FCD40F3C5765" box="[839,1125,849,870]" italics="true" pageId="7" pageNumber="437">
<taxonomicName id="6A9CB0FCFFB39F2FFCE2FCD408985765" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[839,961,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">O. barbarus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB39F2FFC69FCD40F3C5765" box="[972,1125,849,870]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">O. velocissimus</taxonomicName>
</emphasis>
and
<emphasis id="9FE8176DFFB39F2FFB33FCD40E735765" box="[1174,1322,849,870]" italics="true" pageId="7" pageNumber="437">
<taxonomicName id="6A9CB0FCFFB39F2FFB33FCD40E7F5765" authorityName="Mateu et Colas" authorityYear="1954" box="[1174,1318,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">O. penibeticus</taxonomicName>
.
</emphasis>
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<emphasis id="9FE8176DFFB69F2AFC2DFB170F3F50A9" box="[904,1126,1170,1194]" italics="true" pageId="2" pageNumber="432">Orthomus tazekensis</emphasis>
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<paragraph id="AD23CB7FFFB69F2AFCF7FB32089A511F" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Dorsum dark brown, appendages lighter, body length 7.3–9.7 mm. Antennomeres pubescent from middle antennomere 4, in rare cases with some fine, short setae at apex of antennomere 3.</paragraph>
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<paragraph id="AD23CB7FFFB69F2AFCF7FAA708A451D4" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Pronotum transverse, anterior angle moderate or acutely projecting forward; maximum width before middle, side almost linearly narrowed to posterior angle, sometimes slightly sinuate. Inner basal fovea deep and linear, outer fovea frequently shallow, posterior basis with punctures in moderatelow density. Posterior basis somewhat depressed, including the postangular field.</paragraph>
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Elytra moderately convex with fine and unpunctured fine striae, intervals slightly convex, interval 3 with 2 setigerous punctures adjoining stria 3. Humerus well-marked with a moderate tooth projecting laterally. Parascutellar stria well-developed, without scutellar setigerous puncture (present in aberration
<taxonomicName id="6A9CB0FCFFB69F2AFBACF9E40F235274" authorityName="Antoine" authorityYear="1941" box="[1033,1146,1633,1655]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="scutellaris">“scutellaris</taxonomicName>
” and
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). Elytron microsculpture indistinct, elytra moderately shiny in males, isodiametric in females, that have dull elytra.
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Prosternal process rounded. Meso- and metepisterna irregularly punctured, ventrites with rugose punctures. Mesotibia of males not dilated apically, metatibia almost straight, with crenulation at internal margin starting at about the beginning of apical half and consisting of about 8 tubercles. In
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crenulation is rather weak.
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<paragraph id="AD23CB7FFFB69F2AFCF7F8020E7D53D1" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Apical shaft of median lobe asymmetrical, more narrowed to tip in the left side, internal sac with some groups of middle-sized thorns, almost straight in lateral view.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7F8520F5D5C04" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">This subspecies is found in forests of Middle Atlas, in the surroundings of Taza.</paragraph>
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Chaudoir, 1873
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<paragraph id="AD23CB7FFFB69F2AFBBAFEDC0FDD5572" blockId="2.[812,1399,315,1150]" box="[1055,1156,345,369]" pageId="2" pageNumber="432">Figs 1–4.</paragraph>
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Dorsum dark piceous, appendages yellow reddish. Body length 8.0–9.8 mm. Males shiny, females dull with isodiametric microsculpture strongly developed. Antennomeres pubescent from the second third of antennomere 4, sometimes with small distally hairs in antennomere
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addition to the long apical setae (Fig. 1).
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<paragraph id="AD23CB7FFFB69F2AFCF7FD9B0F5D56D0" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">Pronotum transverse with anterior angles moderately projecting forward (Fig. 1), side narrowed forward and only moderately narrowed backward, maximum width before middle, posterior angle obtuse without a small tooth. Basal foveae from almost smooth to clearly punctured, area between external impression and lateral margin (postangular field) notably convex.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FD5D08985770" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">Elytra moderately widened toward apex, slightly narrowed toward basis; striae fine, unpunctured, intervals slightly convex, interval 3 with two setigerous punctures adjoining stria 3; humerus well-marked with a small tooth; scutellar setigerous puncture present at base of a well-developed parascutellar stria.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FCFD08A3502D" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">
Episterna and abdomen from distinctly punctured to almost smooth, punctures on ventrites sometimes connected by shallow longitudinal ridges. Metepisterna slightly longer than wide, L/
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(Fig. 2). Prosternal process not truncate. Mesotibia of males distinctly enlarged from middle to apex, metatibia slightly curved and weakly crenulate at internal margin of distal half.
</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FBB60F04507D" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">Apical shaft of median lobe of the aedeagus large and equally narrowed to tip (Fig. 3), internal sac membranous and with longitudinal macrotrichia.</paragraph>
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<taxonomicName id="6A9CB0FCFFB79F2BFF6FFEBE09A05550" ID-CoL="8MBWQ" authority="Wrase et Jeanne, 2005" authorityName="Wrase et Jeanne" authorityYear="2005" box="[202,761,315,339]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB79F2BFF6FFEBE0AA65550" box="[202,511,315,339]" italics="true" pageId="3" pageNumber="433">Orthomus tazekensis rifensis</emphasis>
Wrase et Jeanne, 2005
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<paragraph id="AD23CB7FFFB79F2BFE0AFEDC094D5573" blockId="3.[189,775,315,777]" box="[431,532,344,369]" pageId="3" pageNumber="433">Figs 5–9.</paragraph>
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<paragraph id="AD23CB7FFFB79F2BFF47FEF80A4F56D0" blockId="3.[189,775,315,777]" pageId="3" pageNumber="433">Characters indicated above for the nominal subspecies are also found in this taxon (Fig. 5). Individuals of this subspecies differ from those of the nominal subspecies in having a scutellar pore puncture; ventrites show abundant punctures giving a rugose aspect (Fig. 6); crenulation of metatibia in males is more weakly developed (it consists of about 6 tubercles); the apical shaft of the median lobe of the aedeagus is shorter and almost evenly rounded apically (Fig. 7), and is almost straight in lateral view (Fig. 8); the open membranous sac extends toward basal bulb. Spermatheca notably dilated apically, perhaps due to full filling with spermatophora (Fig. 9). Its shape when empty couldn’t be assessed.</paragraph>
<paragraph id="AD23CB7FFFB79F2BFF47FD5D0949570A" blockId="3.[189,775,315,777]" pageId="3" pageNumber="433">This subspecies is found in Rif Mountains above the timber line [Wrase, Jeanne, 2005].</paragraph>
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<taxonomicName id="6A9CB0FCFFB79F2BFCD3FEBE0E755550" ID-CoL="74WQD" authority="Wrase et Jeanne, 2005" authorityName="Wrase et Jeanne" authorityYear="2005" box="[886,1324,315,339]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="starkei">
<emphasis id="9FE8176DFFB79F2BFCD3FEBE0F6B5550" box="[886,1074,315,339]" italics="true" pageId="3" pageNumber="433">Orthomus starkei</emphasis>
Wrase et Jeanne, 2005
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<paragraph id="AD23CB7FFFB79F2BFBB7FEDC0FC85572" blockId="3.[812,1399,315,777]" box="[1042,1169,345,369]" pageId="3" pageNumber="433">Figs 10–11.</paragraph>
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<paragraph id="AD23CB7FFFB79F2BFCF7FEFB0E7E561A" blockId="3.[812,1399,315,777]" pageId="3" pageNumber="433">Dorsum from light to dark reddish brown; appendages somewhat lighter than body; body length 7.8–9.2 mm. Eye fairly flat with long and flat tempora; antennae in both sexes with antennomere 9 reaching the pronotal base, pubescent from apical third of antennomere 3 (in addition to normal setae there are some fine short setae hardly visible).</paragraph>
<paragraph id="AD23CB7FFFB79F2BFCF7FD9B0E6156D0" blockId="3.[812,1399,315,777]" pageId="3" pageNumber="433">Pronotum transverse, widest at about anterior third, anterior margin distinctly emarginate, anterior angle moderately projecting forward; posterior angle almost rectangular, side sinuate before it; posterior basis beaded from angle to the origin of internal basal impression, base slightly concave; disk convex; basal external fovea sparsely or not punctured, internal one deeper and with some distinct punctures.</paragraph>
<paragraph id="AD23CB7FFFB79F2BFCF7FD5D094D5397" blockId="3.[812,1399,315,777]" lastBlockId="3.[189,1399,1825,2065]" pageId="3" pageNumber="433">
Elytra long-oval, moderately convex, maximum width a little behind middle; striae fine and unpunctured, intervals Figs 1–11.
<taxonomicName id="6A9CB0FCFFB79F2BFEE6F8A40AE65336" box="[323,447,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB79F2BFEE6F8A40AC05337" box="[323,409,1825,1844]" italics="true" pageId="3" pageNumber="433">Orthomus</emphasis>
spp
</taxonomicName>
: 1–4 —
<taxonomicName id="6A9CB0FCFFB79F2BFDACF8A709D55337" authorityName="Chaudoir" authorityYear="1873" box="[521,652,1826,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="9FE8176DFFB79F2BFDACF8A709D55337" box="[521,652,1826,1845]" italics="true" pageId="3" pageNumber="433">O. maroccanus</emphasis>
</taxonomicName>
; 5–9 —
<taxonomicName id="6A9CB0FCFFB79F2BFD76F8A708D05337" authorityName="Wrase et Jeanne" authorityYear="2005" box="[723,905,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB79F2BFD76F8A708D05337" box="[723,905,1825,1845]" italics="true" pageId="3" pageNumber="433">O. tazekensis rifensis</emphasis>
</taxonomicName>
; 10–11 —
<taxonomicName id="6A9CB0FCFFB79F2BFC4EF8A70F185337" box="[1003,1089,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="starkei">
<emphasis id="9FE8176DFFB79F2BFC4EF8A70F185337" box="[1003,1089,1825,1845]" italics="true" pageId="3" pageNumber="433">O. starkei</emphasis>
</taxonomicName>
; 1, 5 — habitus; 2 — metepisterna; 6 — metepisterna and ventrites; 3, 7–8, 10–11 — median lobe of aedeagus; 4, 9 — genitalia of female; 1, 5, 3, 7, 11 — dorsal view; 2, 6 — right lateral view; 8, 10 — left lateral view; 1, 5–9 — Tidiquin Mnt., Rif,
<collectingCountry id="D58B8BEFFFB79F2BFCE6F8D708CD5366" box="[835,916,1874,1893]" name="Morocco" pageId="3" pageNumber="433">Morocco</collectingCountry>
; 3 — San Roque, Cadiz,
<collectingCountry id="D58B8BEFFFB79F2BFBD4F8D40FFC5367" box="[1137,1189,1873,1892]" name="Spain" pageId="3" pageNumber="433">Spain</collectingCountry>
; 4 — sierra del Cabrito, Tarifa, Cádiz,
<collectingCountry id="D58B8BEFFFB79F2BFE9EF8EC0A36537F" box="[315,367,1897,1916]" name="Spain" pageId="3" pageNumber="433">Spain</collectingCountry>
; 11 — Sebt-des Beni Frassem, Rif,
<collectingCountry id="D58B8BEFFFB79F2BFD0DF8EF09A0537E" box="[680,761,1898,1917]" name="Morocco" pageId="3" pageNumber="433">Morocco</collectingCountry>
; 4, 9 — after
<bibRefCitation id="C90DB68EFFB79F2BFCD7F8EC08A8537F" author="Ortuno V. M." box="[882,1009,1897,1916]" pageId="3" pageNumber="433" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
; 10–11 — after Wrase, Jeanne [2005]. Scale bars: 1.3 mm (Fig. 5); 1.4 mm (Fig. 1).
</paragraph>
<paragraph id="AD23CB7FFFB79F2BFF47F81B09D35C13" blockId="3.[189,1399,1825,2065]" pageId="3" pageNumber="433">
Рис. 1–11.
<taxonomicName id="6A9CB0FCFFB79F2BFEE7F8180AE453B2" box="[322,445,1949,1969]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB79F2BFEE7F8180AC153B3" box="[322,408,1949,1968]" italics="true" pageId="3" pageNumber="433">Orthomus</emphasis>
spp
</taxonomicName>
: 1–4 —
<taxonomicName id="6A9CB0FCFFB79F2BFDA6F81809DC53B3" authorityName="Chaudoir" authorityYear="1873" box="[515,645,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="9FE8176DFFB79F2BFDA6F81809DC53B3" box="[515,645,1949,1968]" italics="true" pageId="3" pageNumber="433">O. maroccanus</emphasis>
</taxonomicName>
; 5–9 —
<taxonomicName id="6A9CB0FCFFB79F2BFD6FF818082753B3" authorityName="Wrase et Jeanne" authorityYear="2005" box="[714,894,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB79F2BFD6FF818082753B3" box="[714,894,1949,1968]" italics="true" pageId="3" pageNumber="433">O. tazekensis rifensis</emphasis>
</taxonomicName>
; 10–11 —
<taxonomicName id="6A9CB0FCFFB79F2BFC78F8180F6853B3" box="[989,1073,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="starkei">
<emphasis id="9FE8176DFFB79F2BFC78F8180F6853B3" box="[989,1073,1949,1968]" italics="true" pageId="3" pageNumber="433">O. starkei</emphasis>
</taxonomicName>
; 1, 5 — внеШний вид; 2 — Эпистерн Заднегруди; 6 — Эпистерн Заднегруди и вентриты; 3, 7–8, 10–11 — среднЯЯ долЯ Эдеагуса; 4, 9 — гениталии самки; 1, 5, 3, 7, 11 — сверху; 2, 6 –сбоку справа; 8, 10 –сбоку слева; 1, 5–9 — горы Тидиквин, Риф,
<collectingCountry id="D58B8BEFFFB79F2BFC2CF84B088253E2" box="[905,987,1998,2017]" name="Morocco" pageId="3" pageNumber="433">Марокко</collectingCountry>
; 3 — Сан- Роке, Кадис, ИспаниЯ; 4 — сьерра дель- Кабрито, Тарифа, Кадис, ИспаниЯ; 11 — Бени Фрассем, Риф,
<collectingCountry id="D58B8BEFFFB79F2BFC90F86308DE53FA" box="[821,903,2022,2041]" name="Morocco" pageId="3" pageNumber="433">Марокко</collectingCountry>
; 4, 9 — по Ortuсo [1996]; 10–11 — по Wrase, Jeanne [2005]. МасШтаб: 1,3 мм (Рис. 5); 1,4 мм (Рис. 1).
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF18FEBE0926560F" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">
slightly convex, interval 3 with two setigerous punctures adjoining stria 3; humerus well-marked with a small indistinct tooth; basal margin anteriorly concave, forming a weak acute angle (
<typeStatus id="722775DDFFB09F2CFE99FE0E0ACD55A2" box="[316,404,395,417]" pageId="4" pageNumber="434" type="holotype">holotype</typeStatus>
) or an almost rectangular angle with lateral margin at the humerus. Setigerous pore present at base of a well-developed parascutellar stria. Microsculpture mesh pattern distinct, in males weakly transverse, in females isodiametric and slightly more stronger developed.
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD94085F5692" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">Prosternum (laterally), episterna and abdomen (laterally) with coarse and somewhat scattered irregular punctures, sometimes connected by shallow longitudinal ridges (last ventrites). Legs slender; mesotibia of males not dilated apically, metatibia not distinctly curved and without crenulation in internal side.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD130AFA56E2" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">Male with median lobe of aedeagus strongly bent; apical shaft short and evenly rounded (Figs 10–11). Internal sac without thorns or teeth.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD63085E5732" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">
<materialsCitation id="1DF4C122FFB09F2CFF47FD63085E5732" country="Morocco" elevation="375" location="Known" municipality="Taza" pageId="4" pageNumber="434" specimenCount="1">
<location id="A8439DA4FFB09F2CFF47FD630A7156F8" LSID="urn:lsid:plazi:treatment:25357A69FFB79F2CFCD3FEBE085E5732:A8439DA4FFB09F2CFF47FD630A7156F8" box="[226,296,742,763]" country="Morocco" municipality="Taza" name="Known" pageId="4" pageNumber="434">Known</location>
only from three localities situated not far from each other in the north-western region of
<collectingMunicipality id="4D475105FFB09F2CFDBCFC84091E5715" box="[537,583,769,790]" pageId="4" pageNumber="434">Taza</collectingMunicipality>
(
<collectingCountry id="D58B8BEFFFB09F2CFDF0FC8409E95715" box="[597,688,769,790]" name="Morocco" pageId="4" pageNumber="434">Morocco</collectingCountry>
), in agricultural lands free of forests, at elevations about
<quantity id="6A64669AFFB09F2CFD21FC9E085A5732" box="[644,771,795,817]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="4" pageNumber="434" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">
<elevation id="26B12C4CFFB09F2CFD21FC9E085A5732" box="[644,771,795,817]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="4" pageNumber="434" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300 to 450 m</elevation>
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.
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</paragraph>
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</treatment>
</document>

81
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<paragraph id="AD23CB7FFFBD9F21FBB7FC980FCB5737" blockId="9.[812,1399,767,1575]" box="[1042,1170,796,821]" pageId="9" pageNumber="439">Figs 51–55.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FCF7FCC40E2E57A4" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Body size large, 10.4–12.4 mm, dorsum from to black dark brown, antennae and tarsomeres red brown, femora and tibiae darker. Male shiny, female less shiny with very fine microreticulation. Eye moderately convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FC290F685044" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Anterior angle of pronotum advanced ahead, well-marked; pronotum more narrowed forward, maximum width at middle (Fig. 51), from here slightly arcuate backward; posterior basis with few and fine punctures, inner fovea linear, outer fovea shallow, basis not emarginate; posterior angle almost straight and without tooth.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FBC90FC150CF" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Elytra almost parallel with small tooth at humerus and clearly sinuate preapically when the epipleura meets dorsum; striae deep, smooth or with insinuated punctures; discal pores on stria 3, intervals almost flat; parascutellar stria normally developed, with proximal hairy pore.</paragraph>
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Ventral segments with punctures, isolated or rugose (Fig. 52). Prosternal process somewhat truncate projected backward in acute ridge. Metepisterna longer than wider (L/
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). Male mesotibia regularly dilated to apex, metatibia not crenulate.
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<paragraph id="AD23CB7FFFBD9F21FCF7FAD20FA951BE" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Apical shaft of median lobe of aedeagus long, asymmetrically sinuate and deviated to the left (Fig. 54), truncate at tip; in lateral view moderately arcuate ventrally (Fig. 53). Female genitalia as described above (Fig. 55).</paragraph>
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The species is found in NW of
<collectingCountry id="D58B8BEFFFBD9F21FBC9FA440FEA51D4" box="[1132,1203,1473,1495]" name="Algeria" pageId="9" pageNumber="439">Algeria</collectingCountry>
in the region of
<collectingCountry id="D58B8BEFFFBD9F21FAE2FA470E2F51D4" box="[1351,1398,1474,1495]" name="Algeria" pageId="9" pageNumber="439">Oran</collectingCountry>
and
<collectingRegion id="6F58059DFFBD9F21FCF3FA58088A51F1" box="[854,979,1501,1522]" country="Algeria" name="Mostaganem" pageId="9" pageNumber="439">Mostaganem</collectingRegion>
, some individuals have been found in ports of south
<collectingCountry id="D58B8BEFFFBD9F21FCD8FA7208E4520F" box="[893,957,1527,1548]" name="France" pageId="9" pageNumber="439">France</collectingCountry>
(Marseille), surely imported. It might be found in
<collectingCountry id="D58B8BEFFFBD9F21FCE0F99708C45224" box="[837,925,1554,1575]" name="Morocco" pageId="9" pageNumber="439">Morocco</collectingCountry>
, in the littoral near the frontier with
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.
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<paragraph id="AD23CB7FFFBD9F21FE07FAFC097B5192" blockId="9.[189,775,1372,2073]" box="[418,546,1401,1425]" pageId="9" pageNumber="439">Figs 47–50.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FF47FA1B09295200" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Dorsum brown, with blackish to reddish hue, shiny in both sexes as microreticulation is not much developed in females; appendages piceous red. Eye of large size, convex, tempora oblique. Body length 9.0–11.2 mm.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FF47F98C094452DA" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Pronotum transverse with maximum width at middle (Fig. 47); from here clearly narrowed forward and almost straight backward; posterior angle slightly obtuse and not toothed; anterior angle not protruding, posterior basis not emarginate; however, lateral margin and side are clearly widened in posterior third; external fovea rugose and punctured, vaguely limited, internal one deep and linear; punctures of posterior moderately dense.</paragraph>
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Elytra long, parallel, basis forming at humerus a small fold tooth-like. Intervals almost flat, striae deep, smooth or with traces of punctures; second disk pore adjoining stria 2. Prosternal process rounded with acute ridge directed toward mesosternum; ventral segments with few punctures; metepisterna longer than wide, L/
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. Male mesotibia regularly dilated to apex, metatibia slightly curved, dilated in apical 1/6 and without crenulation.
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<paragraph id="AD23CB7FFFBD9F21FF47F836096A5C1A" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Median lobe of aedeagus with short triangular apical shaft, deviated to right (Fig. 48); penis in lateral view somewhat ventrally curved and elevated at tip (Fig. 49). Female genitalia (Fig. 50) as described above.</paragraph>
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<emphasis id="9FE8176DFFBD9F21FF61FDA50AFE563B" box="[196,423,544,568]" italics="true" pageId="9" pageNumber="439">Orthomus balearicus</emphasis>
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<paragraph id="AD23CB7FFFBD9F21FE07FDB8097B5656" blockId="9.[189,775,544,1352]" box="[418,546,573,597]" pageId="9" pageNumber="439">Figs 43–46.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FF47FDE7085B56CB" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Dorsum black or dark brown, males not very shiny, females entirely dull, microreticulation well developed in both sexes. Large body length, 11.0–12.3 mm. Appendages reddish brown, femora dark. Eye large and convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FF47FD48091E57D0" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Pronotum almost square in some individuals (maximum width at middle, Fig. 3), or side more curved both anteriad and apicad (maximum width on apical 2/3); the most striking features are the deep and wide lateral margin, the emarginate posterior basis and the depression of the posterior basis; both foveae deep, the inner one deeper and linear, the external one large and separate from side by a flat area; punctures of posterior basis fine and from moderate to dense, mostly found within foveae; posterior angle from obtuse to almost square, devoid of tooth or with faint traces of it.</paragraph>
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Elytra long (L/
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), slightly dilated from basis, humeral tooth absent, discal pore 2 found on the 3rd stria or on interval between striae 2 and 3; parascutellar stria normally developed with proximal setigerous pore; intervals convex, striae smooth; the lateral margin of elytra is deep and wide, excavated, as is the lateral margin of pronotum.
</paragraph>
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Ventral segments with fine and scarce punctures; metepisterna longer than wide, L/
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. Male mesotibia regularly dilated to apex, metatibia slightly curved, dilated in apical 1/6 and with traces of crenulation.
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<paragraph id="AD23CB7FFFBD9F21FF47FB67085B514B" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Median lobe of aedeagus with a short triangular apical shaft (Fig. 44), more narrowed on the right side; on lateral view the penis is almost straight (Fig. 45). Female genitalia as indicated above, with a short duct and a widened spermatheca (Fig. 46).</paragraph>
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<emphasis id="9FE8176DFFBD9F21FC32F9BE0F0E5250" box="[919,1111,1595,1619]" italics="true" pageId="9" pageNumber="439">Orthomus lacouri</emphasis>
(Antoine, 1941)
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According to
<bibRefCitation id="C90DB68EFFBD9F21FC41F9F80FAD5290" author="Pupier R. &amp; Coulon J." box="[996,1268,1661,1683]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
this species includes different geographic races that are distinguished by morphological features, and thus deserve their ranking as subspecies. Three of them are found in NE
<collectingCountry id="D58B8BEFFFBD9F21FB49F94B0E1F52E0" box="[1260,1350,1742,1763]" name="Morocco" pageId="9" pageNumber="439">Morocco</collectingCountry>
,
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<emphasis id="9FE8176DFFBD9F21FAF6F94B0F5752FE" italics="true" pageId="9" pageNumber="439">Orthomus lacouri lacouri</emphasis>
(Antoine, 1941)
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;
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<emphasis id="9FE8176DFFBD9F21FB64F96C0E2F52FE" box="[1217,1398,1768,1790]" italics="true" pageId="9" pageNumber="439">O. lacouri haroldi</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FC89F8860F57531B" author="Pupier R. &amp; Coulon J." box="[812,1038,1795,1817]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier et Coulon, 2013</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="6A9CB0FCFFBD9F21FBE5F8860E2A531A" authority="Mateu, 1955" authorityName="Mateu" authorityYear="1955" box="[1088,1395,1795,1817]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
<emphasis id="9FE8176DFFBD9F21FBE5F8860FA9531B" box="[1088,1264,1795,1816]" italics="true" pageId="9" pageNumber="439">O. lacouri kocheri</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FB52F8860E2A531A" author="Mateu J." box="[1271,1395,1795,1817]" pageId="9" pageNumber="439" pagination="53 - 85" refId="ref17020" refString="Mateu J. 1955. Notas sobre los Orthomus Chaudoir (segunda nota) // Eos. Vol. 31. P. 53 - 85." type="book chapter" year="1955">Mateu, 1955</bibRefCitation>
</taxonomicName>
. The fourth subspecies,
<taxonomicName id="6A9CB0FCFFBD9F21FBB3F89B0E005330" authority="Jeanne, 1988" authorityName="Jeanne" authorityYear="1988" box="[1046,1369,1821,1843]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="pupieri">
<emphasis id="9FE8176DFFBD9F21FBB3F89B0F955330" box="[1046,1228,1821,1843]" italics="true" pageId="9" pageNumber="439">O. lacouri pupieri</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FB70F89B0E005330" author="Jeanne C." box="[1237,1369,1822,1843]" pageId="9" pageNumber="439" pagination="69 - 87" refId="ref16833" refString="Jeanne C. 1988. Carabiques noveaux ou remarquables (9 eme note) // Bull. Soc. linn. Bordeaux. Vol. 16. P. 69 - 87." type="journal article" year="1988">Jeanne, 1988</bibRefCitation>
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, is proper of NW
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. Here I follow the taxa descriptions published by
<bibRefCitation id="C90DB68EFFBD9F21FC15F8D60FF7536B" author="Pupier R. &amp; Coulon J." box="[944,1198,1875,1897]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
and
<bibRefCitation id="C90DB68EFFBD9F21FB7BF8D608365380" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="9" pageNumber="439" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
, together with personal observations of material of this species deposited in the MNCN Madrid and DZAFUM. Male genitalia are the key characters that distinguish the subspecies, which share in common most external characters. Thus, a general description follows and peculiarities of the subspecies are indicated below.
</paragraph>
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Dorsum from black to piceous brown, appendages brown. Body length 10.5–11.3 mm. Eye of moderate size and convex, tempora oblique. Males shiny, females shiny or somewhat dull with visible microreticulation (
<emphasis id="9FE8176DFFBE9F22FDE8FE09099F55A3" box="[589,710,395,417]" italics="true" pageId="10" pageNumber="440">
O. l.
<taxonomicName id="6A9CB0FCFFBE9F22FD25FE0E099F55A3" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[640,710,395,416]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
); both sexes of
<emphasis id="9FE8176DFFBE9F22FEB6FE230AD455B8" box="[275,397,422,443]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
show a dull dorsum.
</paragraph>
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Pronotum shape trapezoid, anterior angle briefly protruding ahead except for
<taxonomicName id="6A9CB0FCFFBE9F22FE21FE59096855F3" authorityName="Pupier et Coulon" authorityYear="2013" box="[388,561,475,497]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="haroldi">
<emphasis id="9FE8176DFFBE9F22FE21FE59096855F3" box="[388,561,475,497]" italics="true" pageId="10" pageNumber="440">O. lacouri haroldi</emphasis>
</taxonomicName>
, anterior basis moderately concave; side arcuate, more narrowed forward, slightly narrowed backward and showing maximum width near posterior basis (Fig. 56); posterior angle slightly obtuse and without tooth; basis no or slightly (
<emphasis id="9FE8176DFFBE9F22FE6BFDC309125658" box="[462,587,582,603]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
) emarginate; inner fovea linear and marked, outer fovea poorly impressed; basis with points varying between scarce and moderately dense.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FEBE0E785584" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">Elytra parallel, humeral tooth vestigial, striae punctured with variable intensity; discal pores on stria 3; 8th interval almost vertically tumbling down to elytron margin.</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FE0E0E16560F" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
Prosternal process truncate; metepisterna long, smooth and narrowed backward (Fig. 57); ventrites almost smooth, with fine punctures except for
<taxonomicName id="6A9CB0FCFFBE9F22FBF6FE440E5A55D5" authorityName="Mateu" authorityYear="1955" box="[1107,1283,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
<emphasis id="9FE8176DFFBE9F22FBF6FE440E5A55D5" box="[1107,1283,449,470]" italics="true" pageId="10" pageNumber="440">O. lacouri kocheri</emphasis>
</taxonomicName>
, that shows well-marked punctures; male mesotibia distally dilated, metatibia with marked crenulation in inner side of distal half.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FD940F275658" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
Female genitalia (Fig. 58) with a short spermatic conduct followed by a progressively widened spermatheca of moderate length [
<bibRefCitation id="C90DB68EFFBE9F22FC3FFDC30F2A5658" author="Pupier R. &amp; Coulon J." box="[922,1139,582,604]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
].
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FDE40E2F5692" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
According to the detailed study of
<bibRefCitation id="C90DB68EFFBE9F22FB6AFDE408295692" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
, each subspecies has a “typical” shape of the median
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FE93F9420E1952E2" blockId="10.[310,649,1735,1761]" box="[310,1344,1735,1761]" lastBlockId="10.[862,1344,1735,1761]" pageId="10" pageNumber="440">
<emphasis id="9FE8176DFFBE9F22FE93F9420E1952E2" bold="true" box="[310,1344,1735,1761]" pageId="10" pageNumber="440">57 56 62 63 64 58</emphasis>
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47F9740E03537F" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
Figs 51–64.
<taxonomicName id="6A9CB0FCFFBE9F22FEEAF9740A955306" box="[335,460,1777,1797]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFBE9F22FEEAF9740AFC5307" box="[335,421,1777,1796]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
spp
</taxonomicName>
: 51–55 —
<taxonomicName id="6A9CB0FCFFBE9F22FD88F97409C65307" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[557,671,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFBE9F22FD88F97409C65307" box="[557,671,1777,1796]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
</taxonomicName>
; 56–64 —
<taxonomicName id="6A9CB0FCFFBE9F22FCA6F97408055307" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[771,860,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
<emphasis id="9FE8176DFFBE9F22FCA6F97408055307" box="[771,860,1777,1796]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
</taxonomicName>
(56–57, 63 —
<emphasis id="9FE8176DFFBE9F22FC46F9740F095307" box="[995,1104,1777,1796]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
(56 — female); 58–60, 62, 64 —
<emphasis id="9FE8176DFFBE9F22FF18F88C0A73531F" box="[189,298,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
; 61 —
<emphasis id="9FE8176DFFBE9F22FECAF88C0A82531F" box="[367,475,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
); 51, 56 — habitus; 57 — metepisterna; 52 — punctures of abdominal segments; 53, 59 — median lobe of aedeagus; 54, 61–64 — apex of median lobe of aedeagus; 55, 58 — genitalia of female; 51, 54, 56, 61–64 — dorsal view; 57–58, 60 — right lateral view; 53, 59 — left lateral view; 52 — ventral view; 51 — reservoir El Fakia,
<collectingCountry id="D58B8BEFFFBE9F22FC7CF8BC0F44534F" box="[985,1053,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
; 55, 61 — Sidi Daho ez Zahir,
<collectingCountry id="D58B8BEFFFBE9F22FA95F8BC0E2D534F" box="[1328,1396,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
; 58 — Jebel Tikermine,
<collectingCountry id="D58B8BEFFFBE9F22FE2FF8D70A825366" box="[394,475,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 62 — Tannezatte,
<collectingCountry id="D58B8BEFFFBE9F22FD21F8D7098C5366" box="[644,725,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 63 — Farkhana, Nador,
<collectingCountry id="D58B8BEFFFBE9F22FC14F8D70F5B5366" box="[945,1026,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 64 — Sidi Bouhouria, Berkane,
<collectingCountry id="D58B8BEFFFBE9F22FA86F8D70E2D5366" box="[1315,1396,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 53–55, 58–61 — after
<bibRefCitation id="C90DB68EFFBE9F22FE23F8EC0915537F" author="Pupier R. &amp; Coulon J." box="[390,588,1897,1916]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
; 62–64 — after
<bibRefCitation id="C90DB68EFFBE9F22FD7BF8EC08F3537F" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[734,938,1897,1917]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
. Scale bars: 1.5 mm (Fig. 51); 1.6 mm (Fig. 56).
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47F8000E695C13" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
Figs 51–64.
<taxonomicName id="6A9CB0FCFFBE9F22FEE8F8000A9E539B" box="[333,455,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFBE9F22FEE8F8000AF8539B" box="[333,417,1925,1944]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
spp
</taxonomicName>
: 51–55 —
<taxonomicName id="6A9CB0FCFFBE9F22FD82F80009CE539B" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[551,663,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFBE9F22FD82F80009CE539B" box="[551,663,1925,1944]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
</taxonomicName>
; 56–64 —
<taxonomicName id="6A9CB0FCFFBE9F22FD5CF8000808539B" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[761,849,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
<emphasis id="9FE8176DFFBE9F22FD5CF8000808539B" box="[761,849,1925,1944]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
</taxonomicName>
(56–57, 63 —
<emphasis id="9FE8176DFFBE9F22FC70F8000F66539B" box="[981,1087,1925,1944]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
(56 — самка); 58–60, 62, 64 —
<emphasis id="9FE8176DFFBE9F22FAC7F8000A5353B3" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
; 61 —
<emphasis id="9FE8176DFFBE9F22FEE3F8180AF253B3" box="[326,427,1949,1968]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
); 51, 56 — внеШний вид; 57 — Эпистерн Заднегруди; 52 — пунктировка сегментов брюШка; 53, 59 — среднЯЯ долЯ Эдеагуса; 54, 61–64 — верШина средней доли Эдеагуса; 55, 58 — гениталии самки; 51, 54, 56, 61–64 — сверху; 57–58, 60 — сбоку справа; 53, 59 — сбоку слева; 52 — сниЗу; 51 — водохранилиЩе Эль- ФакиЯ,
<collectingCountry id="D58B8BEFFFBE9F22FC35F848089653E3" box="[912,975,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
; 55, 61 — Сиди Дахо ЭЗ Захир,
<collectingCountry id="D58B8BEFFFBE9F22FB5CF8480E6153E3" box="[1273,1336,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
; 58 — Джебель Тикермин,
<collectingCountry id="D58B8BEFFFBE9F22FEDFF8600A9553FB" box="[378,460,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 62 — ТаннеЗатт,
<collectingCountry id="D58B8BEFFFBE9F22FDD0F860099E53FB" box="[629,711,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 63 — Фархана, Надор,
<collectingCountry id="D58B8BEFFFBE9F22FC0DF86008A353FB" box="[936,1018,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 64 — Сиди БухуриЯ, Беркане,
<collectingCountry id="D58B8BEFFFBE9F22FABBF8600E2953FB" box="[1310,1392,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 53–55, 58–61 — по
<bibRefCitation id="C90DB68EFFBE9F22FED0F87809625C13" author="Pupier R. &amp; Coulon J." box="[373,571,2045,2064]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
; 62–64 — по
<bibRefCitation id="C90DB68EFFBE9F22FD19F87808D15C13" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[700,904,2045,2064]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
. МасШтаб: 1,5 мм (Рис. 51); 1,6 мм (Рис. 56).
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF18FEBE0A5F55D4" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">lobe of aedeagus, but this shape may vary notably depending on the degree of population gene flow, that results in “hybrid” shapes intermediate between those of “typical” exemplars. That paper should be consulted to check the variability; here I shall refer to the so-called “typical” shapes.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FE590AC856E2" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
In lateral view the distal half of the median lobe of aedeagus is almost straight in
<emphasis id="9FE8176DFFBF9F23FE4DFE73093C5608" box="[488,613,502,523]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
, sinuate in
<emphasis id="9FE8176DFFBF9F23FD7FFE730A5D5625" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
and
<emphasis id="9FE8176DFFBF9F23FE94FD940AFF5625" box="[305,422,529,550]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FEC5FD940AFF5625" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[352,422,529,550]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
, and is ventrally bent in
<emphasis id="9FE8176DFFBF9F23FD37FD94085E5625" box="[658,775,529,550]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
[
<bibRefCitation id="C90DB68EFFBF9F23FF67FDAE0AC35643" author="Pupier R. &amp; Coulon J." box="[194,410,555,577]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
] (Fig. 59). In dorsal view, the apical shaft is truncate and slightly sinuate in
<emphasis id="9FE8176DFFBF9F23FD93FDC309F55658" box="[566,684,582,603]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FDC3FDC309F55658" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[614,684,582,603]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
(Fig. 64) and truncate with a clear tooth in
<emphasis id="9FE8176DFFBF9F23FDA2FDE409265675" box="[519,639,609,630]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
(Fig. 61); the other two subspecies show a rounded apex, sinuate to the left, being wider in
<emphasis id="9FE8176DFFBF9F23FEFEFD130A8456A8" box="[347,477,662,683]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
[
<bibRefCitation id="C90DB68EFFBF9F23FE49FD13099056A8" author="Pupier R. &amp; Coulon J." box="[492,713,662,684]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
;
<bibRefCitation id="C90DB68EFFBF9F23FD7DFD130A2D56C5" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al., 2016</bibRefCitation>
] and slender and more elongate in
<emphasis id="9FE8176DFFBF9F23FD7CFD340A5D56E3" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
(Figs 62–63).
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FD630A275798" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
According to the map provided by
<bibRefCitation id="C90DB68EFFBF9F23FD88FD63085E56FF" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[557,775,742,764]" pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
the subspecies
<emphasis id="9FE8176DFFBF9F23FEF4FC840A975715" box="[337,462,769,790]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
is found west of River Muluya whereas
<emphasis id="9FE8176DFFBF9F23FEB6FC990AD55733" box="[275,396,795,817]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FEE3FC9E0AD55733" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[326,396,795,816]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
is found in the surroundings of Oujda and the Algerian border, to the west near Berkane, El Aioun and Guerfane;
<emphasis id="9FE8176DFFBF9F23FEEFFCD40A995765" box="[330,448,849,870]" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
is found in-between the other two subspecies. Hybrids between these subspecies are found in many contact areas.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="25357A69FFBF9F23FF4CFADC08D356FF" ID-DOI="http://doi.org/10.5281/zenodo.13178454" ID-Zenodo-Dep="13178454" LSID="urn:lsid:plazi:treatment:25357A69FFBF9F23FF4CFADC08D356FF" httpUri="http://treatment.plazi.org/id/25357A69FFBF9F23FF4CFADC08D356FF" lastPageNumber="441" pageId="11" pageNumber="441">
<subSubSection id="E58698F4FFBF9F23FF4CFADC09835172" box="[233,730,1369,1393]" pageId="11" pageNumber="441" type="nomenclature">
<paragraph id="AD23CB7FFFBF9F23FF4CFADC09835172" blockId="11.[189,776,1369,2071]" box="[233,730,1369,1393]" pageId="11" pageNumber="441">
<heading id="F66B7C13FFBF9F23FF4CFADC09835172" box="[233,730,1369,1393]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="441" reason="8">
<taxonomicName id="6A9CB0FCFFBF9F23FF4CFADC09835172" ID-CoL="7592B" authority="(Reiche et Saulcy, 1855)" baseAuthorityName="Reiche et Saulcy" baseAuthorityYear="1855" box="[233,730,1369,1393]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="berytensis">
<emphasis id="9FE8176DFFBF9F23FF4CFADC0A9E5172" box="[233,455,1369,1393]" italics="true" pageId="11" pageNumber="441">Orthomus berytensis</emphasis>
(Reiche et Saulcy, 1855)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFBF9F23FE07FAF308D356FF" pageId="11" pageNumber="441" type="description">
<paragraph id="AD23CB7FFFBF9F23FE07FAF3097B518D" blockId="11.[189,776,1369,2071]" box="[418,546,1398,1422]" pageId="11" pageNumber="441">
<figureCitation id="35A7D7FAFFBF9F23FE07FAF30947518D" box="[418,542,1398,1422]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Figs 65–70</figureCitation>
.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FA190A54521F" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">Dorsum black to dark brown, often with reddish hue, shiny in both sexes (females show fine microreticulation), appendices with lighter coloration brown red. Body length variable 8.0–11.0 mm. Eye convex, protruding, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F9A4095F5325" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">
Pronotum transverse almost rectangular, regularly curved, well narrowed forward and moderately backward, maximum width near middle (
<figureCitation id="35A7D7FAFFBF9F23FED9F9D30A9C526F" box="[380,453,1622,1644]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 65</figureCitation>
), anterior angle briefly projected forward, anterior basis almost regularly arcuate; posterior angle obtuse with a tiny tooth (
<figureCitation id="35A7D7FAFFBF9F23FDABF90E090552A2" box="[526,604,1675,1697]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 66</figureCitation>
), posterior basis beaded up to the origin of the inner fovea; external fovea weakly impressed, inner fovea linear; punctures usually in low density or entirely absent (in East Mediterranean populations punctures are more abundant); space between external fovea and side somewhat convex.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F8AE09D45375" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">Elytra slightly enlarged from basis, humerus with a small and non-protruding tooth, striae smooth or with traces of punctures, intervals flat, discal pores on stria 3.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F8FE0A6F53E2" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">
Prosternal process slightly truncate; ventral segments with shallow punctures at sides (
<figureCitation id="35A7D7FAFFBF9F23FE4DF813097453AF" box="[488,557,1942,1964]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 67</figureCitation>
), metepisterna smooth. Male mesotibia slightly dilated apically, metatibia moderately crenulate.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F86308D356FF" blockId="11.[189,776,1369,2071]" lastBlockId="11.[812,1399,315,764]" pageId="11" pageNumber="441">
Apical shaft of median lobe of aedeagus wide and with trapezoid shape, only briefly narrowed to tip (
<figureCitation id="35A7D7FAFFBF9F23FD36F78409865C15" box="[659,735,2048,2071]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 68</figureCitation>
), in lateral view somewhat ventrally bent (
<figureCitation id="35A7D7FAFFBF9F23FB0CFEBE0FAA5552" box="[1193,1267,315,337]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 69</figureCitation>
). The female genitalia match in all details the characters described by
<bibRefCitation id="C90DB68EFFBF9F23FC89FEF408E75585" author="Ortuno V. M." box="[812,958,368,390]" pageId="11" pageNumber="441" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
for the
<taxonomicName id="6A9CB0FCFFBF9F23FBBBFEF40FC35585" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1054,1178,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFBF9F23FBBBFEF40FC35585" box="[1054,1178,369,390]" italics="true" pageId="11" pageNumber="441">O. barbarus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFBF9F23FB72FEF40E2F5585" baseAuthorityName="Waltl" baseAuthorityYear="1835" box="[1239,1398,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFBF9F23FB72FEF40E2F5585" box="[1239,1398,369,390]" italics="true" pageId="11" pageNumber="441">O. velocissimus</emphasis>
</taxonomicName>
groups, in that there is a long spermathecal duct (where a caecum is originated) followed by a long a slightly dilated spermatheca toward apex (
<figureCitation id="35A7D7FAFFBF9F23FB9CFE440FDC55D5" box="[1081,1157,449,471]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 70</figureCitation>
). Therefore, this pattern clearly differs from that found in the other Maghrebian taxa [
<bibRefCitation id="C90DB68EFFBF9F23FC97FE730F4D5608" author="Pupier R. &amp; Coulon J." box="[818,1044,502,524]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
]. These findings suggests that the species is more closely related to the north Mediterranean lineages than to the Maghrebian ones. However, the species occupies the whole North Africa, and to the east occupies
<collectingCountry id="D58B8BEFFFBF9F23FC89FDE4083D5674" box="[812,868,609,631]" name="Israel" pageId="11" pageNumber="441">Israel</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FCCBFDE4089C5674" box="[878,965,609,631]" name="Lebanon" pageId="11" pageNumber="441">Lebanon</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FC6AFDE40F5C5674" box="[975,1029,609,631]" name="Syria" pageId="11" pageNumber="441">Syria</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FBABFDE40F0F5674" box="[1038,1110,609,631]" name="Turkey" pageId="11" pageNumber="441">Turkey</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FBC5FDE40FFF5675" box="[1120,1190,609,630]" name="Cyprus" pageId="11" pageNumber="441">Cyprus</collectingCountry>
and
<collectingCountry id="D58B8BEFFFBF9F23FB70FDE40E455675" box="[1237,1308,609,630]" name="Greece" pageId="11" pageNumber="441">Greece</collectingCountry>
, that is, a North African geographic pattern. Indeed, it is also found in Sicily, Sardinia and
<collectingCountry id="D58B8BEFFFBF9F23FC55FD130F7256AF" box="[1008,1067,662,684]" name="Malta" pageId="11" pageNumber="441">Malta</collectingCountry>
, and to west reaches the southernmost parts of
<collectingCountry id="D58B8BEFFFBF9F23FC10FD340F5556C5" box="[949,1036,689,710]" name="Morocco" pageId="11" pageNumber="441">Morocco</collectingCountry>
and the Canary Islands. Analyses of new characters are needed to better assess its phylogenetic affinities.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4D91ED5C8ADF99F18F9E3960149DA32A">The genus Orthomus Chaudoir, 1838 in the Iberian Peninsula and Morocco</mods:title>
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<subSubSection id="E58698F4FFBF9F23FC2BFC950E4D572B" box="[910,1300,784,808]" pageId="11" pageNumber="441" type="nomenclature">
<paragraph id="AD23CB7FFFBF9F23FC2BFC950E4D572B" blockId="11.[812,1399,784,2072]" box="[910,1300,784,808]" pageId="11" pageNumber="441">
<heading id="F66B7C13FFBF9F23FC2BFC950E4D572B" box="[910,1300,784,808]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="441" reason="8">
<taxonomicName id="6A9CB0FCFFBF9F23FC2BFC950E4D572B" ID-CoL="6SWQC" authority="Antoine, 1933" authorityName="Antoine" authorityYear="1933" box="[910,1300,784,808]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="9FE8176DFFBF9F23FC2BFC950F2B572B" box="[910,1138,784,808]" italics="true" pageId="11" pageNumber="441">Orthomus dimorphus</emphasis>
Antoine, 1933
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFBF9F24FBB7FCA80AFF52FB" lastPageId="12" lastPageNumber="442" pageId="11" pageNumber="441" type="description">
<paragraph id="AD23CB7FFFBF9F23FBB7FCA80FCB5746" blockId="11.[812,1399,784,2072]" box="[1042,1170,813,837]" pageId="11" pageNumber="441">
<figureCitation id="35A7D7FAFFBF9F23FBB7FCA80FD75746" box="[1042,1166,813,837]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Figs 71–76</figureCitation>
.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FCD708F257EE" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
This species has two subspecies,
<taxonomicName id="6A9CB0FCFFBF9F23FB3FFCD708F05781" authority="Mateu, 1955" authorityName="Mateu" authorityYear="1955" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="antoinei">
<emphasis id="9FE8176DFFBF9F23FB3FFCD70E2F576B" box="[1178,1398,850,872]" italics="true" pageId="11" pageNumber="441">O. dimorphus antoinei</emphasis>
<bibRefCitation id="C90DB68EFFBF9F23FC89FCE808F05781" author="Mateu J." box="[812,937,876,898]" pageId="11" pageNumber="441" pagination="53 - 85" refId="ref17020" refString="Mateu J. 1955. Notas sobre los Orthomus Chaudoir (segunda nota) // Eos. Vol. 31. P. 53 - 85." type="book chapter" year="1955">Mateu, 1955</bibRefCitation>
</taxonomicName>
and
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<emphasis id="9FE8176DFFBF9F23FC45FCE80F815781" box="[992,1240,877,898]" italics="true" pageId="11" pageNumber="441">O. dimorphus dimorphus</emphasis>
Antoine, 1933
</taxonomicName>
. The following description is based on the first one, that is more common in Atlantic
<collectingCountry id="D58B8BEFFFBF9F23FB8CFC270FD957B4" box="[1065,1152,930,951]" name="Morocco" pageId="11" pageNumber="441">Morocco</collectingCountry>
than the nominal subspecies, that is restricted to some localities of Great Atlas over
<quantity id="6A64669AFFBF9F23FC89FC52082757EE" box="[812,894,983,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="441" unit="m" value="2000.0">2000 m</quantity>
.a.s.l.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FC770F505070" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">Dorsum dark brown, appendices brown, male shiny, female mat, microreticulation well visible in females, weak in males. Body length variable within populations and between subspecies, 7.0–12.0 mm. Eyes moderately convex and protruding, tempora long.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FBFD0F7C512E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Pronotum somewhat trapezoid, maximum width after middle (
<figureCitation id="35A7D7FAFFBF9F23FC25FB17089250A4" box="[896,971,1170,1192]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 71</figureCitation>
), side curved and narrowed forward, only narrowed backward near posterior angle; this shows a small protruding tooth; posterior basis beaded up to inner linear fovea; external fovea variably impressed; punctures variable in density and size, usually fine and in moderate density; postangular field convex.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FAB708B2519B" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">Elytra slightly widened from basis; humerus with a small and non-protruding tooth, parascutellar stria well-developed with proximal seta; striae smooth but with traces of punctures; intervals flat.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FA180E29521E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Prosternal process somewhat rounded. Metepisterna smooth, long and narrowed backward (
<figureCitation id="35A7D7FAFFBF9F23FB62FA320E4E51CF" box="[1223,1303,1463,1485]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 72</figureCitation>
). Ventral segments laterally smooth or with fine and few punctures. Male mesotibia not distally dilated; male metatibia only slightly curved and without crenulation in distal inner side.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F9A70894528B" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Median lobe of aedeagus with an elongate apical shaft (
<figureCitation id="35A7D7FAFFBF9F23FC96F9B808225251" box="[819,891,1597,1619]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 73</figureCitation>
), almost evenly narrowed apicad, and almost straight in lateral view (
<figureCitation id="35A7D7FAFFBF9F23FC65F9D20F5E526F" box="[960,1031,1623,1645]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 74</figureCitation>
); membranous area notably prolonged toward the bulb.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F9080E45530E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Spermatheca notably longer than duct (
<figureCitation id="35A7D7FAFFBF9F23FB45F9080E7252A1" box="[1248,1323,1676,1699]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 75</figureCitation>
), it becomes notably thick after fertilization (
<figureCitation id="35A7D7FAFFBF9F23FB0AF9220FA352BE" box="[1199,1274,1703,1725]" captionStart="Figs 65–76" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 65–76. Orthomus spp, specimens from Morocco: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 68–69, 73–74 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 68–69 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 65–76. Orthomus spp, особи иЗ Марокко: 65–70 — O. berytensis; 71–76 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 68–69, 73–74 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 68–69 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 76</figureCitation>
). This shape difference within the same species calls attention on the variable aspect that may take the female genitalia of
<taxonomicName id="6A9CB0FCFFBF9F23FAF6F958082B530F" authorityName="Chaudoir" authorityYear="1838" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="genus">
<emphasis id="9FE8176DFFBF9F23FAF6F958082B530F" italics="true" pageId="11" pageNumber="441">Orthomus</emphasis>
</taxonomicName>
depending on matureness and fertilization.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F897082753CB" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
The subspecies
<taxonomicName id="6A9CB0FCFFBF9F23FC51F8970FB65324" authorityName="Antoine" authorityYear="1933" box="[1012,1263,1810,1831]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="dimorphus">
<emphasis id="9FE8176DFFBF9F23FC51F8970FB65324" box="[1012,1263,1810,1831]" italics="true" pageId="11" pageNumber="441">O. dimorphus dimorphus</emphasis>
</taxonomicName>
occupies the Great Atlas above
<quantity id="6A64669AFFBF9F23FC4DF8A80F6E5341" box="[1000,1079,1837,1858]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="441" unit="m" value="2000.0">2000 m</quantity>
. Its body length is smaller (less than
<quantity id="6A64669AFFBF9F23FCF9F8C208FB535F" box="[860,930,1863,1884]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="441" unit="mm" value="10.0">10 mm</quantity>
), the female is mat, the pronotum basis is almost smooth, the humeral tooth is small, the striae are almost smooth, and the median lobe of aedeagus is ventrally curved upwards, with an apical shaft that forms a relatively short triangle.
</paragraph>
<paragraph id="AD23CB7FFFBF9F24FCF7F8480A1455F2" blockId="11.[812,1399,784,2072]" lastBlockId="12.[189,775,315,497]" lastPageId="12" lastPageNumber="442" pageId="11" pageNumber="441">
The subspecies
<taxonomicName id="6A9CB0FCFFBF9F23FC4BF8480F9253E1" authorityName="Mateu" authorityYear="1955" box="[1006,1227,1997,2018]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="antoinei">
<emphasis id="9FE8176DFFBF9F23FC4BF8480F9253E1" box="[1006,1227,1997,2018]" italics="true" pageId="11" pageNumber="441">O. dimorphus antoinei</emphasis>
</taxonomicName>
is of larger body length and both sexes are dorsally shiny. The pronotum basis is punctured, the external fovea is better marked, the striae of elytron are well impressed and punctured, the humeral tooth is developed. The median lobe of aedeagus is almost straight on distal half in lateral view; in dorsal view it shows a more elongate and triangular apical shaft. This subspecies is found in the Atlantic plain of
<collectingCountry id="D58B8BEFFFB89F24FE3AFE230AAF55B8" box="[415,502,422,443]" name="Morocco" pageId="12" pageNumber="442">Morocco</collectingCountry>
and doesn’t enter in the Rif and the Great Atlas above
<quantity id="6A64669AFFB89F24FE6DFE44094F55D5" box="[456,534,449,470]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="442" unit="m" value="2000.0">2000 m</quantity>
. See
<bibRefCitation id="C90DB68EFFB89F24FDE9FE4409BB55D5" author="Antoine M." box="[588,738,448,471]" pageId="12" pageNumber="442" pagination="179 - 314" refId="ref16250" refString="Antoine M. 1957. Coleopteres Carabiques du Maroc (deuxieme partie) // Memoires de la Societe des Sciences Naturelles et Physiques du Maroc, Zoologie (nouvelle serie). Vol. 3. P. 179 - 314." type="journal article" year="1957">Antoine [1957]</bibRefCitation>
for further details.
</paragraph>
<paragraph id="AD23CB7FFFB89F24FE2CF95B0AFF52FB" blockId="12.[393,653,1722,1784]" box="[393,422,1758,1784]" pageId="12" pageNumber="442">
<emphasis id="9FE8176DFFB89F24FE2CF95B0AFF52FB" bold="true" box="[393,422,1758,1784]" pageId="12" pageNumber="442">71</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

79
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@ -1,48 +1,49 @@
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@ -60,7 +61,7 @@ Ozerov et Krivosheina
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<figureCitation id="58F89E820243FFD43774FA61CE94F9D8" box="[1075,1170,1522,1545]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="0" pageNumber="136">Figs 1–5</figureCitation> <figureCitation id="58F89E820243FFD43774FA61CE94F9D8" box="[1075,1170,1522,1545]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="0" pageNumber="136">Figs 1–5</figureCitation>
. .
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@ -102,7 +103,7 @@ DESCRIPTION. Male (
<emphasis id="F2B75E150243FFD43018F8E6C9A2F85B" box="[863,932,1909,1930]" italics="true" pageId="0" pageNumber="136">Thorax</emphasis> <emphasis id="F2B75E150243FFD43018F8E6C9A2F85B" box="[863,932,1909,1930]" italics="true" pageId="0" pageNumber="136">Thorax</emphasis>
completely black, grey dusted, with dark stripes and fuzzy spots on scutum.Acrostichals not differentiated from the other hairs on scutum, 2 postpronotals, 2 notopleurals, 1+2 supra-alars, 0+2 intra-alars, 2 postalars, and 2+3 dorsocentrals. Proepisternum centrally covered with whitish hairs, without strong setae ventrally. Proepimeron covered with whitish hairs anteriorly. Anepisternum covered with hairs completely and with 2 strong setae along posterior margin. Katepisternum covered with hairs completely, with one strong seta in posterodorsal corner. Postmetacoxal bridge present. Scutellum greyish dusted, with a pair of strong basal scutellar and a pair of strong apical scutellar setae. completely black, grey dusted, with dark stripes and fuzzy spots on scutum.Acrostichals not differentiated from the other hairs on scutum, 2 postpronotals, 2 notopleurals, 1+2 supra-alars, 0+2 intra-alars, 2 postalars, and 2+3 dorsocentrals. Proepisternum centrally covered with whitish hairs, without strong setae ventrally. Proepimeron covered with whitish hairs anteriorly. Anepisternum covered with hairs completely and with 2 strong setae along posterior margin. Katepisternum covered with hairs completely, with one strong seta in posterodorsal corner. Postmetacoxal bridge present. Scutellum greyish dusted, with a pair of strong basal scutellar and a pair of strong apical scutellar setae.
</paragraph> </paragraph>
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<paragraph id="C07C82070242FFD533FAF82CC939F83B" blockId="1.[189,1421,1983,2074]" pageId="1" pageNumber="137"> <paragraph id="C07C82070242FFD533FAF82CC939F83B" blockId="1.[189,1421,1983,2074]" pageId="1" pageNumber="137">
<emphasis id="F2B75E150242FFD533FAF82CCB08F803" bold="true" box="[189,270,1983,2002]" pageId="1" pageNumber="137">Figs 1–5.</emphasis> <emphasis id="F2B75E150242FFD533FAF82CCB08F803" bold="true" box="[189,270,1983,2002]" pageId="1" pageNumber="137">Figs 1–5.</emphasis>
<taxonomicName id="07C3F9840242FFD53253F82CCBCCF803" authorityName="Ozerov &amp; Krivosheina" authorityYear="2024" box="[276,458,1983,2002]" class="Insecta" family="Scathophagidae" genus="Scatomyza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="gobaensis" status="sp. nov."> <taxonomicName id="07C3F9840242FFD53253F82CCBCCF803" authorityName="Ozerov &amp; Krivosheina" authorityYear="2024" box="[276,458,1983,2002]" class="Insecta" family="Scathophagidae" genus="Scatomyza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="gobaensis" status="sp. nov.">
@ -137,15 +138,15 @@ tinged with slightly brownish, veins brownish. Calypters, including margins, yel
<paragraph id="C07C82070241FFD633A4FC59CB12FBAE" blockId="2.[189,787,303,1337]" pageId="2" pageNumber="138"> <paragraph id="C07C82070241FFD633A4FC59CB12FBAE" blockId="2.[189,787,303,1337]" pageId="2" pageNumber="138">
<emphasis id="F2B75E150241FFD633A4FC59CB38FC0E" box="[227,318,970,991]" italics="true" pageId="2" pageNumber="138">Abdomen</emphasis> <emphasis id="F2B75E150241FFD633A4FC59CB38FC0E" box="[227,318,970,991]" italics="true" pageId="2" pageNumber="138">Abdomen</emphasis>
black, greyish dusted, tergites 2–6 each with a lateral marginal seta. Male sternite 4 oval, about 1.5 times as long as wide ( black, greyish dusted, tergites 2–6 each with a lateral marginal seta. Male sternite 4 oval, about 1.5 times as long as wide (
<figureCitation id="58F89E820241FFD6320BFC6CCB81FBC5" box="[332,391,1023,1044]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="2" pageNumber="138">Fig. 1</figureCitation> <figureCitation id="58F89E820241FFD6320BFC6CCB81FBC5" box="[332,391,1023,1044]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="2" pageNumber="138">Fig. 1</figureCitation>
). Male sternite 5 with moderately long lateral lobes, curved like propeller in apical half and with short median lobes ( ). Male sternite 5 with moderately long lateral lobes, curved like propeller in apical half and with short median lobes (
<figureCitation id="58F89E820241FFD63208FBA7CB8FFB98" box="[335,393,1076,1097]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="2" pageNumber="138">Fig. 1</figureCitation> <figureCitation id="58F89E820241FFD63208FBA7CB8FFB98" box="[335,393,1076,1097]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="2" pageNumber="138">Fig. 1</figureCitation>
). Cercal plate as in ). Cercal plate as in
<figureCitation id="58F89E820241FFD63110FBA7C8A9FB98" box="[599,687,1076,1097]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="2" pageNumber="138">Figs 3–5</figureCitation> <figureCitation id="58F89E820241FFD63110FBA7C8A9FB98" box="[599,687,1076,1097]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="2" pageNumber="138">Figs 3–5</figureCitation>
; surstylus with a long process near base ( ; surstylus with a long process near base (
<figureCitation id="58F89E820241FFD632B6FBDCC82AFBB5" box="[497,556,1103,1124]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="2" pageNumber="138">Fig. 3</figureCitation> <figureCitation id="58F89E820241FFD632B6FBDCC82AFBB5" box="[497,556,1103,1124]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="2" pageNumber="138">Fig. 3</figureCitation>
). Aedeagus laterally as in ). Aedeagus laterally as in
<figureCitation id="58F89E820241FFD63391FBF9CB16FBAE" box="[214,272,1130,1151]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." pageId="2" pageNumber="138">Fig. 2</figureCitation> <figureCitation id="58F89E820241FFD63391FBF9CB16FBAE" box="[214,272,1130,1151]" captionStart="Figs 1–5" captionStartId="1.[189,227,1983,2002]" captionTargetBox="[251,1356,385,1869]" captionTargetId="figure-7@1.[189,1420,332,1914]" captionTargetPageId="1" captionText="Figs 1–5. Scatomyza gobaensis, sp.n., male: 1 — abdominal sternites 4 (lower) and 5 (upper); 2 — aedeagus; 3 — epandrium, cercal plate and surstyli, dorsolateral view; 4 — same, dorsal view; 5 — same, lateral view. Рис. 1–5. Scatomyza gobaensis, sp.n., самец: 1 — стерниты брюшка 4 (вниЗу) и 5 (вверху); 2 — Эдеагус; 3 — Эпандрий, церкальнаЯ пластинка и сурстили, дорсолатерально; 4 — то же, сверху; 5 — то же, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178047" httpUri="https://zenodo.org/record/13178047/files/figure.png" pageId="2" pageNumber="138">Fig. 2</figureCitation>
. .
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<paragraph id="C07C82070241FFD633A4FB17CB4AFB65" blockId="2.[189,787,303,1337]" pageId="2" pageNumber="138"> <paragraph id="C07C82070241FFD633A4FB17CB4AFB65" blockId="2.[189,787,303,1337]" pageId="2" pageNumber="138">

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)
<emphasis id="E8A1EAD9BB01FFD003CF1B6963186852" box="[455,606,1831,1855]" italics="true" pageId="7" pageNumber="228">kazakhstanica</emphasis>
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(
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), fig. 15.
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locality:
<collectingCountry id="A2C2765BBB01FFD000B51B1A61A46812" name="Kazakhstan" pageId="7" pageNumber="228">Kazakhstan</collectingCountry>
,
<location id="DF0A6010BB01FFD002E41B22600E6812" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFD002C41B6665BA6E83:DF0A6010BB01FFD002E41B22600E6812" box="[236,328,1900,1919]" country="Kazakhstan" county="Aksu-Dzhabagly" municipality="Talasskiy Alatau" name="Tian Shan" pageId="7" pageNumber="228">Tian Shan</location>
,
<collectingMunicipality id="3A0EACB1BB01FFD0035A1B2260AE6812" box="[338,488,1900,1919]" pageId="7" pageNumber="228">Talasskiy Alatau</collectingMunicipality>
,
<collectingCounty id="330B4E47BB01FFD003FA1B2263C06812" box="[498,646,1900,1919]" pageId="7" pageNumber="228">Aksu-Dzhabagly</collectingCounty>
,
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<elevation id="51F8D1F8BB01FFD000991B2263896812" box="[657,719,1900,1919]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="228" unit="m" value="1200.0">1200m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD002EA1BCA601A68AA" pageId="7" pageNumber="228" type="materials_examined">
<paragraph id="DA6A36CBBB01FFD002EA1BCA601A68AA" blockId="7.[189,775,1873,2071]" pageId="7" pageNumber="228">
<materialsCitation id="6ABD3C96BB01FFD002EA1BCA601E68AA" collectingDate="1871-03-24" collectingDateMax="1871-04-01" collectingDateMin="1871-03-24" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Tashkent" pageId="7" pageNumber="228" specimenCount="13" specimenCount-female="3" specimenCount-male="10">
<collectionCode id="BCC4AE0EBB01FFD002EA1BCA601768FA" box="[226,337,1924,1943]" pageId="7" pageNumber="228">MATERIAL</collectionCode>
EXAMINED.
<emphasis id="E8A1EAD9BB01FFD003D01BCA630268FA" bold="true" box="[472,580,1924,1943]" pageId="7" pageNumber="228">
<collectingCountry id="A2C2765BBB01FFD003D01BCA630668FA" box="[472,576,1924,1943]" name="Uzbekistan" pageId="7" pageNumber="228">Uzbekistan</collectingCountry>
.
</emphasis>
<location id="DF0A6010BB01FFD000441BCA63D968FA" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFD002C41B6665BA6E83:DF0A6010BB01FFD000441BCA63D968FA" box="[588,671,1924,1943]" country="Uzbekistan" name="Tashkent" pageId="7" pageNumber="228">Tashkent</location>
:
<date id="AE6B100BBB01FFD000A11BCA624168FA" box="[681,775,1924,1943]" pageId="7" pageNumber="228" value="1871-03-24">
<collectingDate id="BE2FE9E3BB01FFD000A11BCA624168FA" box="[681,775,1924,1943]" pageId="7" pageNumber="228" value="1871-03-24">24.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD002CB1BD261BA68C2" box="[195,252,1945,1967]" count="2" pageId="7" pageNumber="228" type="male">2 ♂♂</specimenCount>
),
<date id="AE6B100BBB01FFD003061BD2602A68C2" box="[270,364,1948,1967]" pageId="7" pageNumber="228" value="1871-03-26">
<collectingDate id="BE2FE9E3BB01FFD003061BD2602A68C2" box="[270,364,1948,1967]" pageId="7" pageNumber="228" value="1871-03-26">26.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD003711BD260E768C2" box="[377,417,1945,1967]" count="3" pageId="7" pageNumber="228" type="female">3 ♀♀</specimenCount>
),
<date id="AE6B100BBB01FFD003BB1BD2635768C2" box="[435,529,1948,1967]" pageId="7" pageNumber="228" value="1871-03-27">
<collectingDate id="BE2FE9E3BB01FFD003BB1BD2635768C2" box="[435,529,1948,1967]" pageId="7" pageNumber="228" value="1871-03-27">27.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD000151BD2631068C2" box="[541,598,1945,1967]" count="4" pageId="7" pageNumber="228" type="male">4 ♂♂</specimenCount>
),
<date id="AE6B100BBB01FFD000611BD263FC68C2" box="[617,698,1948,1967]" pageId="7" pageNumber="228" value="1871-04-01">
<collectingDate id="BE2FE9E3BB01FFD000611BD263FC68C2" box="[617,698,1948,1967]" pageId="7" pageNumber="228" value="1871-04-01">1.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD000CE1BD263B968C2" box="[710,767,1945,1967]" count="4" pageId="7" pageNumber="228" type="male">4 ♂♂</specimenCount>
) [
<collectorName id="7720531DBB01FFD002CA1BFA601568AA" box="[194,339,1972,1991]" pageId="7" pageNumber="228">A. Fedtschenko</collectorName>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD002EA1B8263DE6896" pageId="7" pageNumber="228" type="discussion">
<paragraph id="DA6A36CBBB01FFD002EA1B8263DE6896" blockId="7.[189,775,1873,2071]" pageId="7" pageNumber="228">
REMARKS. This species was known only by a male and a female taken from
<collectingCountry id="A2C2765BBB01FFD003801BA860BE6891" box="[392,504,2022,2044]" name="Kazakhstan" pageId="7" pageNumber="228">Kazakhstan</collectingCountry>
[
<bibRefCitation id="BE444B3ABB01FFD0000D1BA863C86896" author="Chvala M." box="[517,654,2022,2044]" pageId="7" pageNumber="228" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD002EA144F63CA677A" box="[226,652,2049,2071]" pageId="7" pageNumber="228" type="distribution">
<paragraph id="DA6A36CBBB01FFD002EA144F63CA677A" blockId="7.[189,775,1873,2071]" box="[226,652,2049,2071]" pageId="7" pageNumber="228">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB01FFD00391144F634A677A" box="[409,524,2049,2071]" name="Kazakhstan" pageId="7" pageNumber="228">Kazakhstan</collectingCountry>
,
<collectingCountry id="A2C2765BBB01FFD00010144F63CE677A" box="[536,648,2049,2071]" name="Uzbekistan" pageId="7" pageNumber="228">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD0014C1D7264316EE6" pageId="7" pageNumber="228" type="reference_group">
<paragraph id="DA6A36CBBB01FFD0014C1D7264316EE6" blockId="7.[812,1400,315,494]" pageId="7" pageNumber="228">
<treatmentCitationGroup id="FAC511E5BB01FFD0014C1D7264316EE6" pageId="7" pageNumber="228">
<treatmentCitation id="5B7410DABB01FFD0014C1D7262BB6E1E" author="Chvala M." page="188" pageId="7" pageNumber="228" year="1999">
<taxonomicName id="1DD54D48BB01FFD0014C1D7262BB6E1E" ID-CoL="8BZPB" authority="Chvala, 1999 Chvala, 1999: 188" authorityName="Chvala, 1999 Chvala" authorityYear="1999" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="mediasiatica" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB01FFD0014C1D7262C16E3E" box="[836,903,316,339]" italics="true" pageId="7" pageNumber="228">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB01FFD0019F1D75656B6E3E" box="[919,1069,315,339]" italics="true" pageId="7" pageNumber="228">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB01FFD006361D7565806E3E" box="[1086,1222,315,339]" italics="true" pageId="7" pageNumber="228">mediasiatica</emphasis>
<materialsCitation id="6ABD3C96BB01FFD006C71D7562BB6E1E" country="Kazakhstan" county="Aksu-Dzhabagly" elevation="1200" location="Tian Shan" municipality="Talasskiy Alatau" pageId="7" pageNumber="228" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="BE444B3ABB01FFD006C71D7562BB6E1E" author="Chvala M." pageId="7" pageNumber="228" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999 Chvála, 1999: 188</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
((
<specimenCount id="CCD3FD42BB01FFD0061F1D10656D6E19" box="[1047,1067,350,372]" count="1" pageId="7" pageNumber="228" type="male">♂</specimenCount>
and
<specimenCount id="CCD3FD42BB01FFD006681D10652A6E19" box="[1120,1132,350,372]" count="1" pageId="7" pageNumber="228" type="female">♀</specimenCount>
), figs 23–24.
<typeStatus id="056E8869BB01FFD006FF1D2F64626E19" box="[1271,1316,353,372]" pageId="7" pageNumber="228">Type</typeStatus>
locality:
<collectingCountry id="A2C2765BBB01FFD001241D3662D56EE6" box="[812,915,376,395]" name="Kazakhstan" pageId="7" pageNumber="228">Kazakhstan</collectingCountry>
,
<location id="DF0A6010BB01FFD001931D3662B36EE6" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFD002C41B6665BA6E83:DF0A6010BB01FFD001931D3662B36EE6" box="[923,1013,376,395]" country="Kazakhstan" county="Aksu-Dzhabagly" municipality="Talasskiy Alatau" name="Tian Shan" pageId="7" pageNumber="228">Tian Shan</location>
,
<collectingMunicipality id="3A0EACB1BB01FFD001F51D3665D76EE6" box="[1021,1169,376,395]" pageId="7" pageNumber="228">Talasskiy Alatau</collectingMunicipality>
,
<collectingCounty id="330B4E47BB01FFD006911D36646B6EE6" box="[1177,1325,376,395]" pageId="7" pageNumber="228">Aksu-Dzhabagly</collectingCounty>
,
<quantity id="1D2D9B2EBB01FFD0073D1D3664356EE6" box="[1333,1395,376,395]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="228" unit="m" value="1200.0">
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.
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</paragraph>
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<paragraph id="DA6A36CBBB01FFD0015A1DDE62DB6EBE" blockId="7.[812,1400,315,494]" pageId="7" pageNumber="228">
<materialsCitation id="6ABD3C96BB01FFD0015A1DDE62DF6EBE" collectingDate="1871-04-08" collectingDateMax="1871-04-11" collectingDateMin="1871-04-08" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Tashkent" pageId="7" pageNumber="228" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<collectionCode id="BCC4AE0EBB01FFD0015A1DDE62876ECE" box="[850,961,400,419]" pageId="7" pageNumber="228">MATERIAL</collectionCode>
EXAMINED.
<emphasis id="E8A1EAD9BB01FFD006401DDE65F06ECE" bold="true" box="[1096,1206,400,419]" pageId="7" pageNumber="228">
<collectingCountry id="A2C2765BBB01FFD006401DDE65F46ECE" box="[1096,1202,400,419]" name="Uzbekistan" pageId="7" pageNumber="228">Uzbekistan</collectingCountry>
:
</emphasis>
<location id="DF0A6010BB01FFD006B61DDE64496ECE" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFD002C41B6665BA6E83:DF0A6010BB01FFD006B61DDE64496ECE" box="[1214,1295,400,419]" country="Uzbekistan" name="Tashkent" pageId="7" pageNumber="228">Tashkent</location>
,
<date id="AE6B100BBB01FFD007121DDE64306ECE" box="[1306,1398,400,419]" pageId="7" pageNumber="228" value="1871-04-11">
<collectingDate id="BE2FE9E3BB01FFD007121DDE64306ECE" box="[1306,1398,400,419]" pageId="7" pageNumber="228" value="1871-04-11">11.iv.1871</collectingDate>
</date>
[
<collectorName id="7720531DBB01FFD0013A1DE662FD6ED6" box="[818,955,424,443]" pageId="7" pageNumber="228">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB01FFD001C31DE662AA6ED1" box="[971,1004,422,444]" count="1" pageId="7" pageNumber="228" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB01FFD001FD1DE6655D6ED1" box="[1013,1051,422,444]" count="2" pageId="7" pageNumber="228" type="female">2 ♀♀</specimenCount>
); same locality,
<date id="AE6B100BBB01FFD006AC1DE665B36ED6" box="[1188,1269,424,443]" pageId="7" pageNumber="228" value="1871-04-08">
<collectingDate id="BE2FE9E3BB01FFD006AC1DE665B36ED6" box="[1188,1269,424,443]" pageId="7" pageNumber="228" value="1871-04-08">8.iv.1871</collectingDate>
</date>
[
<collectorName id="7720531DBB01FFD006F61DE662026EBE" pageId="7" pageNumber="228">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB01FFD0015E1D8E62D66EB9" box="[854,912,446,468]" count="2" pageId="7" pageNumber="228" type="male">2 ♂♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD0015A1D9665BA6E83" box="[850,1276,472,494]" pageId="7" pageNumber="228" type="distribution">
<paragraph id="DA6A36CBBB01FFD0015A1D9665BA6E83" blockId="7.[812,1400,315,494]" box="[850,1276,472,494]" pageId="7" pageNumber="228">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB01FFD006011D96653A6E83" box="[1033,1148,472,494]" name="Kazakhstan" pageId="7" pageNumber="228">Kazakhstan</collectingCountry>
,
<collectingCountry id="A2C2765BBB01FFD006801D9665BE6E83" box="[1160,1272,472,494]" name="Uzbekistan" pageId="7" pageNumber="228">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

319
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<heading id="812281A7BB01FFD001481E4D64246D77" box="[832,1378,514,538]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="228" reason="8">
<taxonomicName id="1DD54D48BB01FFD001481E4D64646D77" ID-CoL="C2PSL" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[832,1314,514,538]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="sogdiensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB01FFD001481E4D62C56D77" box="[832,899,515,538]" italics="true" pageId="7" pageNumber="228">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB01FFD0019B1E4C656F6D77" box="[915,1065,514,538]" italics="true" pageId="7" pageNumber="228">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB01FFD006321E4C65EF6D77" box="[1082,1193,514,538]" italics="true" pageId="7" pageNumber="228">sogdiensis</emphasis>
Shamshev
</taxonomicName>
,
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<paragraph id="DA6A36CBBB01FFD0061A1E5165D76D5A" blockId="7.[812,1399,514,2063]" box="[1042,1169,543,567]" pageId="7" pageNumber="228">Figs 13–17.</paragraph>
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<subSubSection id="92CF6540BB01FFD0015A1E0B62E46C32" pageId="7" pageNumber="228" type="materials_examined">
<paragraph id="DA6A36CBBB01FFD0015A1E0B64226DF2" blockId="7.[812,1399,514,2063]" pageId="7" pageNumber="228">
<materialsCitation id="6ABD3C96BB01FFD0015A1E0B64226DF2" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Tajikistan" latitude="39.483334" location="Ziddy" longLatPrecision="1169" longitude="67.95" municipality="Stackelberg" pageId="7" pageNumber="228" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="BCC4AE0EBB01FFD0015A1E0B62B96D3A" box="[850,1023,580,600]" pageId="7" pageNumber="228">TYPE MATERIAL</collectionCode>
.
<typeStatus id="056E8869BB01FFD006001E0A651C6D3A" box="[1032,1114,580,599]" pageId="7" pageNumber="228" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CCD3FD42BB01FFD0066B1E0C65316D35" box="[1123,1143,578,600]" count="1" pageId="7" pageNumber="228" type="male">♂</specimenCount>
, labels as in Fig. 17, [printed in Cyrillic, Russian]
<location id="DF0A6010BB01FFD001FA1E1265606D02" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFDE01481E4D65AA6B1C:DF0A6010BB01FFD001FA1E1265606D02" box="[1010,1062,604,623]" country="Tajikistan" latitude="39.483334" longLatPrecision="1169" longitude="67.95" municipality="Stackelberg" name="Ziddy" pageId="7" pageNumber="228">Ziddy</location>
[
<geoCoordinate id="BFE1500CBB01FFD0063E1E1265C46D02" box="[1078,1154,604,623]" degrees="39" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="228" precision="925" value="39.483334">39°29´N</geoCoordinate>
<geoCoordinate id="BFE1500CBB01FFD006861E12659F6D02" box="[1166,1241,604,623]" degrees="67" direction="east" minutes="57" orientation="longitude" pageId="7" pageNumber="228" precision="925" value="67.95">67°57´E</geoCoordinate>
], S skl. [=sklon, slope] Gissarsk. khr. [=Gissarskiy khrebet, Gissar Range], Taj. [=
<emphasis id="E8A1EAD9BB01FFD001481EC262D96DF2" bold="true" box="[832,927,652,671]" pageId="7" pageNumber="228">
<collectingCountry id="A2C2765BBB01FFD001481EC262D96DF2" box="[832,927,652,671]" name="Tajikistan" pageId="7" pageNumber="228">Tajikistan</collectingCountry>
</emphasis>
] 14.vi.44,
<collectingMunicipality id="3A0EACB1BB01FFD006091EC265216DF2" box="[1025,1127,652,671]" pageId="7" pageNumber="228">Stackelberg</collectingMunicipality>
(
<collectionCode id="BCC4AE0EBB01FFD0067D1EC265E36DF2" box="[1141,1189,652,671]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
,
<collectionCode id="BCC4AE0EBB01FFD006B81EC265946DF2" box="[1200,1234,652,671]" pageId="7" pageNumber="228">INS</collectionCode>
_DIP_0000623).
</materialsCitation>
</paragraph>
<paragraph id="DA6A36CBBB01FFD0015A1EEB62E46C32" blockId="7.[812,1399,514,2063]" pageId="7" pageNumber="228">
<materialsCitation id="6ABD3C96BB01FFD0015A1EEB62246DA2" collectionCode="ZISP" country="Tajikistan" latitude="39.483334" location="Ziddy" longLatPrecision="1169" longitude="67.95" municipality="Stackelberg" pageId="7" pageNumber="228" specimenCount="8" specimenCount-female="1" specimenCount-male="7" typeStatus="paratype">
<typeStatus id="056E8869BB01FFD0015A1EEB62EF6DD5" box="[850,937,677,696]" pageId="7" pageNumber="228" type="paratype">Paratypes</typeStatus>
.
<emphasis id="E8A1EAD9BB01FFD001BE1EEA655A6DDA" bold="true" box="[950,1052,676,695]" pageId="7" pageNumber="228">
<collectingCountry id="A2C2765BBB01FFD001BE1EEA655E6DDA" box="[950,1048,676,695]" name="Tajikistan" pageId="7" pageNumber="228">Tajikistan</collectingCountry>
:
</emphasis>
same data as holotype (
<specimenCount id="CCD3FD42BB01FFD0070B1EEA64066DD5" box="[1283,1344,674,696]" count="7" pageId="7" pageNumber="228" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB01FFD007581EEA64376DD5" box="[1360,1393,674,696]" count="1" pageId="7" pageNumber="228" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB01FFD001241EF2621A6DA2" box="[812,860,700,719]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB01FFD001621EF362246D8A" collectingDate="1944-06-15" collectionCode="ZISP" country="Tajikistan" latitude="39.483334" location="Ziddy" longLatPrecision="1169" longitude="67.95" municipality="Stackelberg" pageId="7" pageNumber="228" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
same locality and collector as
<typeStatus id="056E8869BB01FFD006631EF265FF6DA2" box="[1131,1209,700,719]" pageId="7" pageNumber="228" type="holotype">holotype</typeStatus>
,
<date id="AE6B100BBB01FFD006C81EF2645A6DA2" box="[1216,1308,700,719]" pageId="7" pageNumber="228" value="1944-06-15">
<collectingDate id="BE2FE9E3BB01FFD006C81EF2645A6DA2" box="[1216,1308,700,719]" pageId="7" pageNumber="228" value="1944-06-15">15.vi.1944</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD007221EF2640A6DBD" box="[1322,1356,698,720]" count="1" pageId="7" pageNumber="228" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB01FFD0075E1EF264376DBD" box="[1366,1393,698,720]" count="1" pageId="7" pageNumber="228" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB01FFD001241E9A621A6D8A" box="[812,860,724,743]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB01FFD001621E9B62246D92" collectingDate="1944-06-17" collectionCode="ZISP" country="Tajikistan" latitude="39.483334" location="Ziddy" longLatPrecision="1169" longitude="67.95" municipality="Stackelberg" pageId="7" pageNumber="228" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
same locality and collector as
<typeStatus id="056E8869BB01FFD006631E9A65FF6D8A" box="[1131,1209,724,743]" pageId="7" pageNumber="228" type="holotype">holotype</typeStatus>
,
<date id="AE6B100BBB01FFD006C81E9A645A6D8A" box="[1216,1308,724,743]" pageId="7" pageNumber="228" value="1944-06-17">
<collectingDate id="BE2FE9E3BB01FFD006C81E9A645A6D8A" box="[1216,1308,724,743]" pageId="7" pageNumber="228" value="1944-06-17">17.vi.1944</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB01FFD007221E9A640A6D85" box="[1322,1356,722,744]" count="1" pageId="7" pageNumber="228" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB01FFD0075E1E9A64376D85" box="[1366,1393,722,744]" count="1" pageId="7" pageNumber="228" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB01FFD001241EA2621A6D92" box="[812,860,748,767]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB01FFD001631EA265056C7A" collectingDate="1940-07-11" collectionCode="ZISP" collectorName="Gussakovskij" country="Tajikistan" location="Gofilabad" municipality="Lyucheb River" pageId="7" pageNumber="228" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB01FFD001631EA262826D92" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFDE01481E4D65AA6B1C:DF0A6010BB01FFD001631EA262826D92" box="[875,964,748,767]" country="Tajikistan" municipality="Lyucheb River" name="Gofilabad" pageId="7" pageNumber="228">Gofilabad</location>
, upper flow of
<collectingMunicipality id="3A0EACB1BB01FFD0064C1EA265806D92" box="[1092,1222,748,767]" pageId="7" pageNumber="228">Lyucheb River</collectingMunicipality>
,
<date id="AE6B100BBB01FFD006C61EA264746D92" box="[1230,1330,748,767]" pageId="7" pageNumber="228" value="1940-07-11">
<collectingDate id="BE2FE9E3BB01FFD006C61EA264746D92" box="[1230,1330,748,767]" pageId="7" pageNumber="228" value="1940-07-11">11.vii.1940</collectingDate>
</date>
,
<collectorName id="7720531DBB01FFD007321EA2622A6C7A" pageId="7" pageNumber="228">Gussakovskij</collectorName>
(
<specimenCount id="CCD3FD42BB01FFD001751F4A62E46C75" box="[893,930,770,792]" count="1" pageId="7" pageNumber="228" type="male">1 ♂</specimenCount>
, dissected,
<collectionCode id="BCC4AE0EBB01FFD006041F4A657A6C7A" box="[1036,1084,772,791]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ABD3C96BB01FFD006461F4A64086C42" collectingDate="1869-05-25" collectionCode="ZMMU" collectorName="A. Fedtschenko" country="Uzbekistan" location="Sangy Dzhuman" pageId="7" pageNumber="228" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="E8A1EAD9BB01FFD006461F4A65FA6C7A" bold="true" box="[1102,1212,772,791]" pageId="7" pageNumber="228">
<collectingCountry id="A2C2765BBB01FFD006461F4A65FE6C7A" box="[1102,1208,772,791]" name="Uzbekistan" pageId="7" pageNumber="228">Uzbekistan</collectingCountry>
:
</emphasis>
<location id="DF0A6010BB01FFD006CC1F4A64136C7A" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFDE01481E4D65AA6B1C:DF0A6010BB01FFD006CC1F4A64136C7A" box="[1220,1365,772,791]" country="Uzbekistan" name="Sangy Dzhuman" pageId="7" pageNumber="228">Sangy Dzhuman</location>
[or Sangy-juman Pass],
<date id="AE6B100BBB01FFD001E81F5265736C42" box="[992,1077,796,815]" pageId="7" pageNumber="228" value="1869-05-25">
<collectingDate id="BE2FE9E3BB01FFD001E81F5265736C42" box="[992,1077,796,815]" pageId="7" pageNumber="228" value="1869-05-25">25.v.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB01FFD0064A1F5265886C42" box="[1090,1230,796,815]" pageId="7" pageNumber="228">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB01FFD006EA1F5465B06C5D" box="[1250,1270,794,816]" count="1" pageId="7" pageNumber="228" type="male">♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB01FFD0070A1F52640F6C42" box="[1282,1353,796,815]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="7" pageNumber="228" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB01FFD007501F5362D86C32" collectingDate="1984-06-03" collectionCode="ZISP" collectorName="Tanasijchuk" country="Uzbekistan" location="Takhta-Karacha Pass" pageId="7" pageNumber="228" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<location id="DF0A6010BB01FFD007501F5362896C25" LSID="urn:lsid:plazi:treatment:527C87DDBB01FFDE01481E4D65AA6B1C:DF0A6010BB01FFD007501F5362896C25" country="Uzbekistan" name="Takhta-Karacha Pass" pageId="7" pageNumber="228">Takhta-Karacha Pass</location>
,
<locationDeviation id="CBD82FE9BB01FFD001D11F7A65E36C2A" box="[985,1189,820,839]" location="Samarkand" pageId="7" pageNumber="228">
<quantity id="1D2D9B2EBB01FFD001D11F7A65566C2A" box="[985,1040,820,839]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="228" unit="km" value="70.0">70 km</quantity>
S of Samarkand
</locationDeviation>
,
<date id="AE6B100BBB01FFD006A71F7A64456C2A" box="[1199,1283,820,839]" pageId="7" pageNumber="228" value="1984-06-03">
<collectingDate id="BE2FE9E3BB01FFD006A71F7A64456C2A" box="[1199,1283,820,839]" pageId="7" pageNumber="228" value="1984-06-03">3.vi.1984</collectingDate>
</date>
,
<collectorName id="7720531DBB01FFD007051F7A64306C2A" box="[1293,1398,820,839]" pageId="7" pageNumber="228">Tanasijchuk</collectorName>
(
<specimenCount id="CCD3FD42BB01FFD0013B1F02621A6C0D" box="[819,860,842,864]" count="3" pageId="7" pageNumber="228" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB01FFD0016F1F0262D16C32" box="[871,919,844,863]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="228">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD0015A1F2A642B6B59" pageId="7" pageNumber="228" type="diagnosis">
<paragraph id="DA6A36CBBB01FFD0015A1F2A642B6B59" blockId="7.[812,1399,514,2063]" pageId="7" pageNumber="228">
DIAGNOSIS. Rather large species (body about
<quantity id="1D2D9B2EBB01FFD007161F2A642D6C14" box="[1310,1387,868,890]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="228" unit="mm" value="7.5">7.5 mm</quantity>
); male eyes dichoptic; labrum nearly 2X longer than eye height; palpus brownish to yellowish brown, usually with yellow tip; mesoscutum indistinctly vittate, acrostichal setae biserial, dorsocentral setae uniserial, scutellum with 2 setae, laterotergite with brown setae; legs with coxae and femora yellow; abdomen brown, tergites extensively shiny, only tergites 1–2 with posteromarginal setae; female similar to male, legs with simple setae.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFDE015A187462836B6B" lastPageId="9" lastPageNumber="230" pageId="7" pageNumber="228" type="description">
<paragraph id="DA6A36CBBB01FFD0015A187462E06927" blockId="7.[812,1399,514,2063]" pageId="7" pageNumber="228">
DESCRIPTION. Body length about
<quantity id="1D2D9B2EBB01FFD006AD187765A86B22" box="[1189,1262,1081,1103]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="228" unit="mm" value="7.5">7.5 mm</quantity>
, wing 6.9–7.1.
<emphasis id="E8A1EAD9BB01FFD00124181A62186B04" bold="true" box="[812,862,1108,1129]" pageId="7" pageNumber="228">Male</emphasis>
(Fig. 13). Head capsule regions mostly densely greyish pruinose; face on lower margin and clypeus shiny; head setation mostly black. Dichoptic, ommatidia equally small. Frons on middle part nearly as broad as distance between inner margins of posterior ocelli, slightly divergent just below ocellar triangle and above antennae; with marginal setulae. Face broad, bare. Ocellar setae long, strong, lateroclinate; two pairs of posterior setulae. Occiput with row of postocular setae, including 3–5 long setae on upper portion and short, slender setae on lower portion; transverse row of long, strong setae; postgena with pale hair-like setae. Antenna with scape and pedicel light brown, postpedicel and stylus black; scape nearly 2X longer than subglobular pedicel, both with short setulae; postpedicel 3.8–4X as long as basal width, with straight margins; stylus nearly 2X shorter than postpedicel basal width. Proboscis with labrum mostly yellowish, usually brownish yellow near base; nearly 2X longer than eye height; palpus short, brownish to yellowish brown, usually with yellow tip; bearing scattered pale fine setae, 1 subapical seta usually dark.
</paragraph>
<paragraph id="DA6A36CBBB01FFDF015A1A0160AE6ECC" blockId="7.[812,1399,514,2063]" lastBlockId="8.[189,775,315,417]" lastPageId="8" lastPageNumber="229" pageId="7" pageNumber="228">Thorax almost entirely brown in ground-colour, postpronotal lobe and postalar tubercle somewhat yellowish brown translucent; mostly densely greyish pruinose; mesoscutum in dorsal view mostly densely brownish grey pruinose, with some traces of darker vittae between acrostichal and dorsocentral setae only near anterior margin; thoracic setation black. Prosternum bare. Proepisternum with 5–6 brownish to pale setae on lower part, bare on upper part. Antepronotum with 7–8 short, spine-like setae dorsally. Postpronotal lobe bearing 1 long and 2–3 shorter setae (with a few additional setulae). Mesonotal setae: acrostichals irregularly biserial, very short, scattered (6–7 per row), absent on prescutellar depression; dorsocentral setae uniserial, presutural dorsocentrals similar to but slightly longer than acrostichals, postsutural dorsocentral setae becoming longer towards scutellum, 2 pairs of prescutellars longest and strongest; 1 long, strong presutural supra-alar, 4 notopleurals, 1 postsutural supra-alar, 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair very short, fine). Laterotergite with 4–5 strong brownish setae and, usually, 2–3 fine, paler setae. Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="DA6A36CBBB0EFFDF015A1D7264306ECC" blockId="8.[812,1398,315,417]" pageId="8" pageNumber="229">Legs with coxae faintly greyish pruinose, remaining podomeres subshiny; black setose. Legs colour: coxae and trochanters yellow (sometimes fore coxa slightly darkened near base anteriorly); femora yellow to brownish yellow</paragraph>
<paragraph id="DA6A36CBBB0EFFDF03F31B19635E681C" blockId="8.[189,1399,1879,2061]" box="[507,536,1879,1905]" pageId="8" pageNumber="229">
<emphasis id="E8A1EAD9BB0EFFDF03F31B19635E681C" bold="true" box="[507,536,1879,1905]" pageId="8" pageNumber="229">14</emphasis>
</paragraph>
<paragraph id="DA6A36CBBB0EFFDF02EA1B30656F68AC" blockId="8.[189,1399,1879,2061]" pageId="8" pageNumber="229">
Figs 13–17.
<taxonomicName id="1DD54D48BB0EFFDF03471B30639468FC" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[335,722,1918,1937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="sogdiensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0EFFDF03471B3060C268FD" box="[335,388,1918,1936]" italics="true" pageId="8" pageNumber="229">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0EFFDF039A1B30634D68FC" box="[402,523,1918,1937]" italics="true" pageId="8" pageNumber="229">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0EFFDF00101B30633768FC" box="[536,625,1918,1937]" italics="true" pageId="8" pageNumber="229">sogdiensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB0EFFDF00D41B30624168FC" bold="true" box="[732,775,1918,1937]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="F39257A2BB0EFFDF00D41B30624168FC" box="[732,775,1918,1937]" pageId="8" pageNumber="229" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 13 — habitus,
<typeStatus id="056E8869BB0EFFDF01C11B30655168FC" box="[969,1047,1918,1937]" pageId="8" pageNumber="229" type="holotype">holotype</typeStatus>
; 14 — hypopygium; 15 — hypandrium; 16 — cerci; 17 —
<typeStatus id="056E8869BB0EFFDF03681BD860ED68C4" box="[352,427,1942,1961]" pageId="8" pageNumber="229" type="holotype">holotype</typeStatus>
labels; 13–14 — lateral view; 15 — ventral view; 16 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is
<quantity id="1D2D9B2EBB0EFFDF01F81BE0656368AC" box="[1008,1061,1966,1985]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="229" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="DA6A36CBBB0EFFDF02EA1B8462DE6760" blockId="8.[189,1399,1879,2061]" pageId="8" pageNumber="229">
Рис. 13–17.
<taxonomicName id="1DD54D48BB0EFFDF03581B84639568B1" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[336,723,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="sogdiensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0EFFDF03581B8460C368B1" box="[336,389,1994,2012]" italics="true" pageId="8" pageNumber="229">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0EFFDF039B1B87634A68B1" box="[403,524,1993,2012]" italics="true" pageId="8" pageNumber="229">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0EFFDF00111B87633468B1" box="[537,626,1993,2012]" italics="true" pageId="8" pageNumber="229">sogdiensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB0EFFDF00D41B87624168B1" bold="true" box="[732,775,1993,2012]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="F39257A2BB0EFFDF00D41B87624168B1" box="[732,775,1993,2012]" pageId="8" pageNumber="229" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 13 — габитус, голотип; 14 — гипопигий; 15 — гипандрий; 16 — церки; 17 — Этикетки голотипа; 13–14 — сбоку; 15 — сниЗу; 16 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм.
</paragraph>
<paragraph id="DA6A36CBBB0FFFDE02B51D7563BA6C0A" blockId="9.[189,776,315,2071]" pageId="9" pageNumber="230">(somewhat darker closer to apex dorsally); tibiae yellow near base, becoming gradually darker towards apex, brownish on apex; tarsi brown. Coxae and trochanters with simple setae. Fore femur bare ventrally. Fore tibia covered with short setulae; bearing 2–3 short anterodorsal and 2–3 similar posterodorsal setae (besides circlet of subapical setae; number, position and robustness variable). Fore basitarsus slender, with short, strong setae ventrally. Mid femur whitish pubescent ventrally; with minute anteroventral and posteroventral setulae; 1 long strong seta near apex anteriorly. Mid tibia with 2–3 anterodorsal, 1 dorsal near base and 2–3 posterodorsal setae; dense, fine setulae ventrally. Mid tarsomeres 1–3 with short spine-like setae ventrally (longer on basitarsus). Hind femur slender, only slightly broader than mid femur; whitish pubescent ventrally; with rows of short strong anteroventral and longer posteroventral setae; dense spinule-like setulae over entire ventral face; 1 long strong seta near apex anteriorly. Hind tibia slender; with 2–3 anterodorsal and 3–4 posterodorsal setae; erect fine setulae ventrally; very short seta in posteroapical comb. Hind basitarsus slender; tarsomeres 1–3 with short spine-like setae ventrally (longer on basitarsus).</paragraph>
<paragraph id="DA6A36CBBB0FFFDE02EA1F25632D6B1C" blockId="9.[189,776,315,2071]" pageId="9" pageNumber="230">
Wing membrane faintly infuscate; entirely covered with microtrichia; veins yellowish brown, well-sclerotized. Veins R
<subScript id="4651348EBB0FFFDE02C51FFE61926CD1" attach="left" box="[205,212,944,956]" fontSize="5" pageId="9" pageNumber="230">5</subScript>
and M
<subScript id="4651348EBB0FFFDE031E1FFE605B6CD1" attach="left" box="[278,285,944,956]" fontSize="5" pageId="9" pageNumber="230">1</subScript>
strongly divergent toward wing margin; radial fork acute; R
<subScript id="4651348EBB0FFFDE03071F8460506CBB" attach="left" box="[271,278,970,982]" fontSize="5" pageId="9" pageNumber="230">5</subScript>
meeting costa before wing apex; CuA+CuP (anal vein) complete (slightly weakened closer to wing margin). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 2X longer than its middle portion. Pterostigma pale brownish, very narrow. Basal costal seta absent. Anal lobe well-developed; axillary incision 90°. Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<paragraph id="DA6A36CBBB0FFFDE02EA18386302699C" blockId="9.[189,776,315,2071]" pageId="9" pageNumber="230">Abdomen brown in ground colour, segments 1–3 slightly paler, tergites 6–7 somewhat yellow translucent posteriorly; tergites extensively shiny, tergite 1 entirely, tergites 2–6 narrowly anteriorly very faintly greyish pruinose (sometimes pruinosity inconspicuous on tergites 3–6), tergites 6–7 with golden brown iridescent spots; sternites mostly faintly greyish pruinose, sternites 2–6 with some shiny space becoming gradually smaller from sternite 2 to sternite 6. Abdominal setation greatly reduced; tergites covered with very short, hair-like, pale setae, only tergites 1 and 2 with long, strong, black posteromarginal setae; sternite 1 bare, sternites 2–6 mostly with very short, pale, hair-like setae, sternites 7–8 with black setae including long strong posteromarginals. Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; small isolated sclerite just beyond posterior margin of pleuron 7; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, somewhat constricted anteriorly, with 2 lateral projections on each side anteriorly (anterior small projection close to upper margin; posterior projection larger, slightly concave apically), deep rounded concavity between projections.</paragraph>
<paragraph id="DA6A36CBBB0FFFDE02EA1AB862A46D2C" blockId="9.[189,776,315,2071]" lastBlockId="9.[812,1399,315,1137]" pageId="9" pageNumber="230">Hypopygium (Fig. 14) moderately large; epandrial lamella mostly brown, narrowly yellowish brown along lower margin, subshiny; hypandrium brown, bare; cercus brown; phallus yellowish brown. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella rather subtriangular (lateral view), with rounded apex; bearing black setae more numerous and longer along lower margin. Hypandrium (Fig. 15) separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin; with two small notches on truncate apex (ventral view); bare; gonocoxal apodeme small. Cerci (Fig. 16) separated with each other and from epandrium; cercus elongate ovate in lateral view, rather subtriangular in dorsal view; shorter than epandrial lamella, with hardly distinguishable additional lobe dorsally, smoothed on inner margin; with black setulae. Phallus almost entirely hidden; zigzag bent, strongly constricted beyond middle; basal portion thick, gently curved and becoming narrower distad; apical portion semicircular, nearly of uniform thickness, with its tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.</paragraph>
<paragraph id="DA6A36CBBB0FFFDE015A1E0864316C7A" blockId="9.[812,1399,315,1137]" pageId="9" pageNumber="230">
<emphasis id="E8A1EAD9BB0FFFDE015A1E0862E56D36" bold="true" box="[850,931,582,603]" pageId="9" pageNumber="230">Female.</emphasis>
Similar to male except as follows. Frons slightly broader, almost parallel-sided. Mid tibia with 1 anteroventral seta. Hind tibia with 1 anteroventral seta on apical half; with simple setulae ventrally. Mid femur with 2–3 short, strong anterodorsal and row of 3–6 short anteroventral setae; no posteroventral setae and ventral spinules. Abdominal tergites shiny, except faintly pruinose tergite 1 and anterior margin of tergite 2. Cercus brownish, very short, with scattered setulae.
</paragraph>
<paragraph id="DA6A36CBBB0FFFDE015A1F5562836B6B" blockId="9.[812,1399,315,1137]" pageId="9" pageNumber="230">
DIFFERENTIAL DIAGNOSIS. The new species is most similar to
<taxonomicName id="1DD54D48BB0FFFDE019B1F7965866C26" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[915,1216,822,844]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="uzbekistanica">
<emphasis id="E8A1EAD9BB0FFFDE019B1F7962E46C21" box="[915,930,823,844]" italics="true" pageId="9" pageNumber="230">E</emphasis>
.
<emphasis id="E8A1EAD9BB0FFFDE01B91F7865726C26" box="[945,1076,822,843]" italics="true" pageId="9" pageNumber="230">uzbekistanica</emphasis>
<bibRefCitation id="BE444B3ABB0FFFDE06351F7865866C26" author="Chvala M." box="[1085,1216,822,844]" pageId="9" pageNumber="230" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
(known from
<collectingCountry id="A2C2765BBB0FFFDE075A1F7862386C0A" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
) sharing dichoptic eyes of the male and extensively yellow legs (including coxae).
<taxonomicName id="1DD54D48BB0FFFDE065D1F2265BA6CED" authorityName="Shamshev" authorityYear="2023" box="[1109,1276,875,897]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="sogdiensis" status="sp. nov.">
<emphasis id="E8A1EAD9BB0FFFDE065D1F2265BA6CED" box="[1109,1276,875,897]" italics="true" pageId="9" pageNumber="230">Empis sogdiensis</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB0FFFDE07091F2564746CED" bold="true" box="[1281,1330,875,896]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="F39257A2BB0FFFDE07091F2564746CED" box="[1281,1330,875,896]" pageId="9" pageNumber="230" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="1DD54D48BB0FFFDE01681FC962BB6CF6" authorityName="Chvala" authorityYear="1999" box="[864,1021,902,924]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="uzbekistanica">
<emphasis id="E8A1EAD9BB0FFFDE01681FC962296CF1" box="[864,879,903,924]" italics="true" pageId="9" pageNumber="230">E</emphasis>
.
<emphasis id="E8A1EAD9BB0FFFDE01721FC862BB6CF6" box="[890,1021,902,923]" italics="true" pageId="9" pageNumber="230">uzbekistanica</emphasis>
</taxonomicName>
primarily by brown abdomen, 4 scutellar setae and black posteromarginal setae on tergites 1 and 2 only. In
<taxonomicName id="1DD54D48BB0FFFDE01881FF265596CBD" authorityName="Chvala" authorityYear="1999" box="[896,1055,955,977]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="uzbekistanica">
<emphasis id="E8A1EAD9BB0FFFDE01881FF262C96CBC" box="[896,911,956,977]" italics="true" pageId="9" pageNumber="230">E</emphasis>
.
<emphasis id="E8A1EAD9BB0FFFDE01941FF565596CBD" box="[924,1055,955,976]" italics="true" pageId="9" pageNumber="230">uzbekistanica</emphasis>
</taxonomicName>
the abdomen is yellow, the scutellum bears only 2 setae and posteromarginal setae are present on tergites 1–5.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDE015A184565426B3A" pageId="9" pageNumber="230" type="etymology">
<paragraph id="DA6A36CBBB0FFFDE015A184565426B3A" blockId="9.[812,1399,315,1137]" pageId="9" pageNumber="230">
ETYMOLOGY. The epithet refers to the name of an ancient civilization in present-day
<collectingCountry id="A2C2765BBB0FFFDE067E186865916B51" box="[1142,1239,1062,1084]" name="Tajikistan" pageId="9" pageNumber="230">Tajikistan</collectingCountry>
and
<collectingCountry id="A2C2765BBB0FFFDE070B186864346B51" box="[1283,1394,1062,1084]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
, Sogdia (or Sogdiana).
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDE015A181565AA6B1C" box="[850,1260,1115,1137]" pageId="9" pageNumber="230" type="distribution">
<paragraph id="DA6A36CBBB0FFFDE015A181565AA6B1C" blockId="9.[812,1399,315,1137]" box="[850,1260,1115,1137]" pageId="9" pageNumber="230">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB0FFFDE06011815652A6B1C" box="[1033,1132,1115,1137]" name="Tajikistan" pageId="9" pageNumber="230">Tajikistan</collectingCountry>
,
<collectingCountry id="A2C2765BBB0FFFDE0670181565AE6B1C" box="[1144,1256,1115,1137]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

228
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<subSubSection id="92CF6540BB03FFD2013F183D642D6BE7" box="[823,1387,1138,1162]" pageId="5" pageNumber="226" type="nomenclature">
<paragraph id="DA6A36CBBB03FFD2013F183D642D6BE7" blockId="5.[812,1399,1138,2067]" box="[823,1387,1138,1162]" pageId="5" pageNumber="226">
<heading id="812281A7BB03FFD2013F183D642D6BE7" box="[823,1387,1138,1162]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="226" reason="8">
<taxonomicName id="1DD54D48BB03FFD2013F183D646D6BE7" ID-CoL="C2R36" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[823,1323,1138,1162]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="hirsutitarsis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB03FFD2013F183D623C6BE7" box="[823,890,1139,1162]" italics="true" pageId="5" pageNumber="226">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB03FFD20183183C65676BE7" box="[907,1057,1138,1162]" italics="true" pageId="5" pageNumber="226">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB03FFD20639183C65F46BE7" box="[1073,1202,1138,1162]" italics="true" pageId="5" pageNumber="226">hirsutitarsis</emphasis>
Shamshev
</taxonomicName>
,
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<taxonomicNameLabel id="F39257A2BB03FFD2073E183C642D6BE7" box="[1334,1387,1138,1162]" pageId="5" pageNumber="226" rank="species">sp.n.</taxonomicNameLabel>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD2061A18C165D76BCA" box="[1042,1169,1167,1191]" pageId="5" pageNumber="226" type="description">
<paragraph id="DA6A36CBBB03FFD2061A18C165D76BCA" blockId="5.[812,1399,1138,2067]" box="[1042,1169,1167,1191]" pageId="5" pageNumber="226">
<figureCitation id="42EE2A4EBB03FFD2061A18C165CB6BCA" box="[1042,1165,1167,1191]" captionStart="Figs 10–12" captionStartId="6.[226,263,1965,1984]" captionTargetBox="[197,1386,1185,1939]" captionTargetId="figure-614@6.[188,1398,1177,1947]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., male, holotype: 10–11 — habitus; 10 — lateral view; 11 — dorsal view; 12 — labels. Scale bar is 1 mm. Рис. 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., самец, голотип: 10–11 — габитус; 10 — сбоку; 11 — сверху; 12 — Этикетки. Шкала = 1 mm." pageId="5" pageNumber="226">Figs 10–12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD2015A18F664746A2E" pageId="5" pageNumber="226" type="materials_examined">
<paragraph id="DA6A36CBBB03FFD2015A18F665A16A46" blockId="5.[812,1399,1138,2067]" pageId="5" pageNumber="226">
<materialsCitation id="6ABD3C96BB03FFD2015A18F665A16A46" collectingDate="1871-06-25" collectionCode="TYPE, MATERIAL, ZMMU" collectorName="A. Fedtschenko &amp; Cyrillic &amp; Russian &amp; Jiptyk &amp; Jiptik" country="Kyrgyzstan" latitude="39.6" location="Pamir-Alay mountain" longLatPrecision="1168" longitude="70.566666" pageId="5" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="BCC4AE0EBB03FFD2015A18F665476BA6" box="[850,1025,1208,1227]" pageId="5" pageNumber="226">TYPE MATERIAL</collectionCode>
.
<typeStatus id="056E8869BB03FFD2060318F6651B6BA6" box="[1035,1117,1208,1227]" pageId="5" pageNumber="226" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CCD3FD42BB03FFD2066118FB653B6BA6" box="[1129,1149,1205,1227]" count="1" pageId="5" pageNumber="226" type="male">♂</specimenCount>
, labels as in
<figureCitation id="42EE2A4EBB03FFD206FE18F664716BA6" box="[1270,1335,1208,1227]" captionStart="Figs 10–12" captionStartId="6.[226,263,1965,1984]" captionTargetBox="[197,1386,1185,1939]" captionTargetId="figure-614@6.[188,1398,1177,1947]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., male, holotype: 10–11 — habitus; 10 — lateral view; 11 — dorsal view; 12 — labels. Scale bar is 1 mm. Рис. 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., самец, голотип: 10–11 — габитус; 10 — сбоку; 11 — сверху; 12 — Этикетки. Шкала = 1 mm." pageId="5" pageNumber="226">Fig. 12</figureCitation>
, [
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<collectingCountry id="A2C2765BBB03FFD2074218F662376B8E" name="Kyrgyzstan" pageId="5" pageNumber="226">Kyrgyzstan</collectingCountry>
</emphasis>
] small bluish green label with red line and number 25. [=
<date id="AE6B100BBB03FFD2012418A662CC6B96" box="[812,906,1256,1275]" pageId="5" pageNumber="226" value="1871-06-25">
<collectingDate id="BE2FE9E3BB03FFD2012418A662CC6B96" box="[812,906,1256,1275]" pageId="5" pageNumber="226" value="1871-06-25">25.vi.1871</collectingDate>
</date>
] [
<collectorName id="7720531DBB03FFD2019618A6656A6B96" box="[926,1068,1256,1275]" pageId="5" pageNumber="226">A. Fedtschenko</collectorName>
] // [printed in
<collectorName id="7720531DBB03FFD206BA18A665B06B96" box="[1202,1270,1256,1275]" pageId="5" pageNumber="226">Cyrillic</collectorName>
,
<collectorName id="7720531DBB03FFD2070A18A6640F6B96" box="[1282,1353,1256,1275]" pageId="5" pageNumber="226">Russian</collectorName>
, pre- 1918 orthography]
<collectorName id="7720531DBB03FFD201D3194E65566A7E" box="[987,1040,1280,1299]" pageId="5" pageNumber="226">Jiptyk</collectorName>
[alternative spelling
<collectorName id="7720531DBB03FFD206DB194E64436A7E" box="[1235,1285,1280,1299]" pageId="5" pageNumber="226">Jiptik</collectorName>
, ~
<geoCoordinate id="BFE1500CBB03FFD2072E194E64346A7E" box="[1318,1394,1280,1299]" degrees="39" direction="north" minutes="36" orientation="latitude" pageId="5" pageNumber="226" precision="925" value="39.6">39°36´N</geoCoordinate>
,
<geoCoordinate id="BFE1500CBB03FFD20124195662306A46" box="[812,886,1304,1323]" degrees="70" direction="east" minutes="34" orientation="longitude" pageId="5" pageNumber="226" precision="925" value="70.566666">70°34´E</geoCoordinate>
,
<location id="DF0A6010BB03FFD20189195665786A46" LSID="urn:lsid:plazi:treatment:527C87DDBB03FFD0013F183D63496866:DF0A6010BB03FFD20189195665786A46" box="[897,1086,1304,1323]" country="Kyrgyzstan" latitude="39.6" longLatPrecision="1168" longitude="70.566666" name="Pamir-Alay mountain" pageId="5" pageNumber="226">Pamir-Alay mountain</location>
system] (
<collectionCode id="BCC4AE0EBB03FFD2069E195665986A46" box="[1174,1246,1304,1323]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="226" type="Museum">ZMMU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DA6A36CBBB03FFD2015A197E64746A2E" blockId="5.[812,1399,1138,2067]" box="[850,1330,1325,1347]" pageId="5" pageNumber="226">
<materialsCitation id="6ABD3C96BB03FFD2015A197E646B6A2E" box="[850,1325,1325,1347]" collectingDate="1871-06-25" collectionCode="ZISP" collectorName="A. Fedtschenko &amp; Cyrillic &amp; Russian &amp; Jiptyk &amp; Jiptik" country="Kyrgyzstan" latitude="39.6" location="Pamir-Alay mountain" longLatPrecision="1168" longitude="70.566666" pageId="5" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="056E8869BB03FFD2015A197E62E66A2E" box="[850,928,1328,1347]" pageId="5" pageNumber="226" type="paratype">Paratype</typeStatus>
. Same data as holotype (
<specimenCount id="CCD3FD42BB03FFD20674196365D66A2E" box="[1148,1168,1325,1347]" count="1" pageId="5" pageNumber="226" type="male">♂</specimenCount>
, dissected,
<collectionCode id="BCC4AE0EBB03FFD206FF197E64616A2E" box="[1271,1319,1328,1347]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="226">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD2015A1906645D6974" pageId="5" pageNumber="226" type="diagnosis">
<paragraph id="DA6A36CBBB03FFD2015A1906645D6974" blockId="5.[812,1399,1138,2067]" pageId="5" pageNumber="226">
DIAGNOSIS. Mid-sized (body length about
<quantity id="1D2D9B2EBB03FFD20717190664366A30" box="[1311,1392,1352,1374]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="226" unit="mm" value="5.5">5.5 mm</quantity>
) species; male eyes holoptic; postpedicel 4X as long as basal width; labrum 1.5X longer than eye height, palpus brownish on apical 1/3; mesoscutum indistinctly vittate, acrostichal setae biserial, dorsocentrals uniserial, laterotergite with pale setae; femora extensively brownish to brownish yellow, basitarsus of all legs with long setae dorsally; abdomen pale setose, abdominal tergites 2–5 shiny, cerci yellow.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD0015A1A53637069D1" lastPageId="7" lastPageNumber="228" pageId="5" pageNumber="226" type="description">
<paragraph id="DA6A36CBBB03FFD2015A1A5365B7677F" blockId="5.[812,1399,1138,2067]" pageId="5" pageNumber="226">
DESCRIPTION. Body length
<quantity id="1D2D9B2EBB03FFD206811A53659D695E" box="[1161,1243,1565,1587]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="226" unit="mm" value="5.4">5.4 mm</quantity>
, wing
<quantity id="1D2D9B2EBB03FFD2072F1A536430695E" box="[1319,1398,1565,1587]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="226" unit="mm" value="6.1">6.1 mm</quantity>
(
<typeStatus id="056E8869BB03FFD2013B1A7662C86923" box="[819,910,1592,1614]" pageId="5" pageNumber="226" type="holotype">holotype</typeStatus>
).
<emphasis id="E8A1EAD9BB03FFD201AC1A76629E6920" bold="true" box="[932,984,1592,1613]" pageId="5" pageNumber="226">Male</emphasis>
(
<figureCitation id="42EE2A4EBB03FFD201E41A76657D6920" box="[1004,1083,1592,1614]" captionStart="Figs 10–12" captionStartId="6.[226,263,1965,1984]" captionTargetBox="[197,1386,1185,1939]" captionTargetId="figure-614@6.[188,1398,1177,1947]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., male, holotype: 10–11 — habitus; 10 — lateral view; 11 — dorsal view; 12 — labels. Scale bar is 1 mm. Рис. 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., самец, голотип: 10–11 — габитус; 10 — сбоку; 11 — сверху; 12 — Этикетки. Шкала = 1 mm." pageId="5" pageNumber="226">Fig. 10</figureCitation>
). Head capsule regions black, mostly densely greyish pruinose, face along lower margin and clypeus shiny.Holoptic, upper ommatidia enlarged.Frons represented by very small subtriangular space just below ocellar triangle and somewhat larger space above antennae, bare. Face broad, bare. Ocellar setae black, long, fine. Postocular setae black, long and strong on upper portion and short on lower portion; occiput with almost regular transverse row of long, strong mostly black setae (some lateral setae brownish yellow to pale); postgena with pale, mostly fine setae. Antenna with scape and pedicel brownish, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with short setulae; postpedicel 4X as long as basal width; stylus nearly 2X longer than postpedicel basal width. Proboscis with labrum brownish, paler apically; nearly 1.5X longer than eye height; palpus short, yellowish brown on about basal 2/3, brown on apical 1/3; bearing sparse, long, mostly pale, fine setae (1 dark subapical seta).
</paragraph>
<paragraph id="DA6A36CBBB00FFD102EA1D7563C36C7A" blockId="6.[189,775,315,1137]" pageId="6" pageNumber="227">
Thorax almost entirely brown in ground-colour, postalar tubercle somewhat yellowish brown translucent; mostly densely greyish pruinose and black setose (except noted); mesoscutum with 4 brownish, very indistinct vittae (
<figureCitation id="42EE2A4EBB00FFD100361DC563C06ECD" box="[574,646,395,417]" captionStart="Figs 10–12" captionStartId="6.[226,263,1965,1984]" captionTargetBox="[197,1386,1185,1939]" captionTargetId="figure-614@6.[188,1398,1177,1947]" captionTargetPageId="6" captionText="Figs 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., male, holotype: 10–11 — habitus; 10 — lateral view; 11 — dorsal view; 12 — labels. Scale bar is 1 mm. Рис. 10–12. Empis (Polyblepharis) hirsutitarsis Shamshev, sp.n., самец, голотип: 10–11 — габитус; 10 — сбоку; 11 — сверху; 12 — Этикетки. Шкала = 1 mm." pageId="6" pageNumber="227">Fig. 11</figureCitation>
). Prosternum bare. Proepisternum with tuft of pale, fine setae on lower part, bare on upper part. Antepronotum with 2–3 black, short, strong and 2–3 pale finer setae. Postpronotal lobe with 1 long, black seta and several pale and dark setulae anteriorly. Mesonotal setae: acrostichals irregularly biserial, short, fine, absent on prescutellar depression; dorsocentral setae uniserial, presuturals slightly longer than acrostichals, postsuturals becoming longer towards scutellum, 2 pairs of long, strong prescutellars; 1 long presutural supra-alar; 3 notopleurals, 1 postsutural supra-alar (with 1 short, fine seta anteriorly), 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair slightly shorter); in addition, some pale setulae present on anterior half of notopleuron. Laterotergite with numerous pale to pale yellow setae. Anterior and posterior spiracles pale.
</paragraph>
<paragraph id="DA6A36CBBB00FFD102EA1F5265C96C81" blockId="6.[189,775,315,1137]" lastBlockId="6.[812,1399,315,1137]" pageId="6" pageNumber="227">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose, coxae with fine pale and stronger black setae. Legs colour: coxae and trochanters brownish; fore femur entirely brownish, mid femur mostly brownish (narrowly brownish yellow near base), hind femur yellowish on about basal 1/3 and rather brownish yellow on apical 2/3; fore and mid tibiae yellowish near base and becoming gradually darker towards brownish apex, hind tibia mostly yellowish (brownish at apex); tarsi brown. Coxae and trochanters with simple setae. All femora whitish pubescent ventrally (more distinct on mid and hind femora). Fore femur with rows of minute, fine anteroventral and similar, somewhat longer posteroventral setae. Fore tibia with rows of anterodorsal and posterodorsal setae, which are short on about basal 1/3 and moderately long on about apical 2/3 (longest setae lightly longer than tibia width). Fore basitarsus slender; with moderately long setae dorsally (as fore tibia), scattered anteroventral and posteroventral spine-like, short setae; tarsomere 2 with slightly longer setulae dorsally, bearing anteroventral and posteroventral spinules. Mid femur with rows of minute anteroventral and posteroventral setulae, only 3–4 short posteroventral setae near extreme apex, row of short anterodorsal setae (more distinct closer to apex). Mid tibia with numerous long, strong anterodorsal and slightly finer posterodorsal setae (at least 1.5X longer than tibia width); simple setulae ventrally. Mid tarsus as fore tarsus but basitarsus with more numerous and somewhat longer anteroventral and posteroventral spine-like setae. Hind femur slightly broader on about apical 2/3 (anterior view), on middle slightly broader than mid femur (nearly 1.3X); with row of 5–6 anterodorsal setae on about apical 1/3; anteroventral setae stronger than posteroventrals, of different lengths, short closer to base and apex, moderately long on middle part (longest setae shorter than femur maximal width); spinules on ventral face; long fine setae posteroventrally (except subapical portion). Hind tibia simple; bearing numerous long anterodorsal and posterodorsal setae; simple setulae ventrally; no seta in posteroapical comb. Hind tarsus with similar setation to mid tarsus; basitarsus slightly thickened.</paragraph>
<paragraph id="DA6A36CBBB00FFD0015A1FBF624E6ED1" blockId="6.[812,1399,315,1137]" lastBlockId="7.[189,776,315,1803]" lastPageId="7" lastPageNumber="228" pageId="6" pageNumber="227">
Wing membrane hyaline; entirely covered with microtrichia; veins mostly brownish, well-sclerotized (except not- ed). Veins R
<subScript id="4651348EBB00FFD101AF187B62E86B2C" attach="left" box="[935,942,1077,1089]" fontSize="5" pageId="6" pageNumber="227">5</subScript>
and M
<subScript id="4651348EBB00FFD101FD187B62BA6B2C" attach="left" box="[1013,1020,1077,1089]" fontSize="5" pageId="6" pageNumber="227">1</subScript>
divergent towards wing margin; radial fork acute; R
<subScript id="4651348EBB00FFD101AC181E62ED6B31" attach="left" box="[932,939,1104,1116]" fontSize="5" pageId="6" pageNumber="227">5</subScript>
meeting costa before wing apex; CuA+CuP (anal vein) complete (slightly weakened beyond middle). Cell dm moderately large, with elongate apex; apical portion of M 4 nearly 2.5X longer than its middle portion. Pterostigma brownish yellow, very narrow. Basal costal seta present, moderately long. Anal lobe well-developed; axillary incision acute but closer to 90°. Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<caption id="8EAA6643BB00FFD102EA1BE360806760" pageId="6" pageNumber="227" startId="6.[226,263,1965,1984]" targetBox="[197,1386,1185,1939]" targetPageId="6" targetType="figure">
<paragraph id="DA6A36CBBB00FFD102EA1BE3609168B4" blockId="6.[189,1399,1965,2061]" pageId="6" pageNumber="227">
Figs 10–12.
<taxonomicName id="1DD54D48BB00FFD103431BE0639168AD" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[331,727,1965,1985]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="227" phylum="Arthropoda" rank="species" species="hirsutitarsis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB00FFD103431BE060C668AD" box="[331,384,1966,1984]" italics="true" pageId="6" pageNumber="227">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB00FFD103841BE3634368AD" box="[396,517,1965,1984]" italics="true" pageId="6" pageNumber="227">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB00FFD100191BE3633F68AD" box="[529,633,1965,1984]" italics="true" pageId="6" pageNumber="227">hirsutitarsis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB00FFD100D71BE0624C68AC" bold="true" box="[735,778,1966,1985]" pageId="6" pageNumber="227">
<taxonomicNameLabel id="F39257A2BB00FFD100D71BE0624C68AC" box="[735,778,1966,1985]" pageId="6" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male, holotype: 10–11 — habitus; 10 — lateral view; 11 — dorsal view; 12 — labels. Scale bar is 1 mm.
</paragraph>
<paragraph id="DA6A36CBBB00FFD102EA1BAC60806760" blockId="6.[189,1399,1965,2061]" pageId="6" pageNumber="227">
Рис. 10–12.
<taxonomicName id="1DD54D48BB00FFD103591BAC63A56899" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[337,739,2017,2036]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="227" phylum="Arthropoda" rank="species" species="hirsutitarsis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB00FFD103591BAC60C06899" box="[337,390,2018,2036]" italics="true" pageId="6" pageNumber="227">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB00FFD1039C1BAF634B6899" box="[404,525,2017,2036]" italics="true" pageId="6" pageNumber="227">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB00FFD100131BAF63C56899" box="[539,643,2017,2036]" italics="true" pageId="6" pageNumber="227">hirsutitarsis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB00FFD100E51BAF625E6899" bold="true" box="[749,792,2017,2036]" pageId="6" pageNumber="227">
<taxonomicNameLabel id="F39257A2BB00FFD100E51BAF625E6899" box="[749,792,2017,2036]" pageId="6" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец, голотип: 10–11 — габитус; 10 — сбоку; 11 — сверху; 12 — Этикетки. Шкала = 1 mm.
</paragraph>
</caption>
<paragraph id="DA6A36CBBB01FFD002EA1D8F638A6CBC" blockId="7.[189,776,315,1803]" pageId="7" pageNumber="228">Abdomen long cylindrical, mostly brown, covered with pale to yellow setae; sternites 1–6 somewhat paler; sternite 8 rather yellowish brown, paler closer to posterior margin. Tergite 1 entirely faintly greyish pruinose; tergites 2–5 mostly shiny, narrowly faintly greyish pruinose anterodorsally; tergites 6–7 with brownish iridescent spot dorsally, shiny literally; tergites covered with mostly fine setae longer laterally, tergites 1–3 with hardly distinguishable, slightly stronger posteromarginal lateral setae. Sternites faintly greyish pruinose; sternite 1 bare, remaining sternites with long, fine setae. Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; large, isolated, convex sclerite near posterior margin of pleuron 7 (modified posterior part of pleuron?); segment 8 with separated tergite and sternite; tergite 8 represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, somewhat constricted anteriorly, with 2 projections on each side (anterior projection close to upper margin, small; posterior projection large, subconical).</paragraph>
<paragraph id="DA6A36CBBB01FFD002EA1F98633D6AA1" blockId="7.[189,776,315,1803]" pageId="7" pageNumber="228">Hypopygium moderately large; epandrial lamella mostly brownish yellow, yellow along margins; cerci yellow; hypandrium brownish yellow; phallus with basal portion brownish yellow, curved apical portion yellow. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella subtriangular (lateral view), with broadly rounded apex; bearing brownish yellow to yellow fine setae more numerous along lower margin. Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, with rounded apex; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus shorter than epandrial lamella, flattened dorsally forming additional lobe, serrate on inner margin in subapical portion; with dark setulae. Phallus almost entirely hidden; zigzag bent; very thick and gently curved on basal portion, strongly constricted beyond middle; apical flexible portion semicircularly bend, thickened proximally, with its tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.</paragraph>
<paragraph id="DA6A36CBBB01FFD002EA199F60E76A8A" blockId="7.[189,776,315,1803]" box="[226,417,1489,1511]" pageId="7" pageNumber="228">
<emphasis id="E8A1EAD9BB01FFD002EA199F60756A8B" bold="true" box="[226,307,1489,1510]" pageId="7" pageNumber="228">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="DA6A36CBBB01FFD002EA19A5637069D1" blockId="7.[189,776,315,1803]" pageId="7" pageNumber="228">
DIFFERENTIAL DIAGNOSIS. The new species is most similar to
<taxonomicName id="1DD54D48BB01FFD003231A49637E6976" authority="Shamshev, 2003" authorityName="Shamshev" authorityYear="2003" box="[299,568,1542,1564]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="zimini">
<emphasis id="E8A1EAD9BB01FFD003231A49607C6971" box="[299,314,1543,1564]" italics="true" pageId="7" pageNumber="228">E</emphasis>
.
<emphasis id="E8A1EAD9BB01FFD003441A4960C16971" box="[332,391,1543,1564]" italics="true" pageId="7" pageNumber="228">zimini</emphasis>
<bibRefCitation id="BE444B3ABB01FFD0039B1A48637E6976" author="Shamshev I. V." box="[403,568,1542,1564]" pageId="7" pageNumber="228" pagination="19 - 27" refId="ref12875" refString="Shamshev I. V. 2003. Descriptions of three new species of the Empis subgenus Polyblepharis (Diptera: Empididae) from the Ukraine and Middle Asia // An international Journal of dipterological Research. Vol. 14. P. 19 - 27." type="book chapter" year="2003">Shamshev, 2003</bibRefCitation>
</taxonomicName>
sharing extensively brownish legs, pale setae of the laterotergite and abdomen as well as shiny abdominal tergites 2–5 and yellow cerci.
<taxonomicName id="1DD54D48BB01FFD000C21A7260746906" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="hirsutitarsis" status="sp. nov.">
<emphasis id="E8A1EAD9BB01FFD000C21A7260746906" italics="true" pageId="7" pageNumber="228">Empis hirsutitarsis</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB01FFD0033F1A1860216906" bold="true" box="[311,359,1622,1643]" pageId="7" pageNumber="228">
<taxonomicNameLabel id="F39257A2BB01FFD0033F1A1860216906" box="[311,359,1622,1643]" pageId="7" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
can be readily distinguished from
<taxonomicName id="1DD54D48BB01FFD000B91A1962416901" authorityName="Shamshev" authorityYear="2003" box="[689,775,1622,1644]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="zimini">
<emphasis id="E8A1EAD9BB01FFD000B91A1963866901" box="[689,704,1623,1644]" italics="true" pageId="7" pageNumber="228">E</emphasis>
.
<emphasis id="E8A1EAD9BB01FFD000C41A1962416901" box="[716,775,1623,1644]" italics="true" pageId="7" pageNumber="228">zimini</emphasis>
</taxonomicName>
primarily by brownish palpi (versus pale yellow), indistinctly vittate mesoscutum (four more or less distinct dark vittae) and long setose fore and mid basitarsi.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD002EA1A8F6010699C" pageId="7" pageNumber="228" type="etymology">
<paragraph id="DA6A36CBBB01FFD002EA1A8F6010699C" blockId="7.[189,776,315,1803]" pageId="7" pageNumber="228">ETYMOLOGY. The epithet refers to long setose tarsi in the new species</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB01FFD002EA1AB863496866" box="[226,527,1782,1803]" pageId="7" pageNumber="228" type="distribution">
<paragraph id="DA6A36CBBB01FFD002EA1AB863496866" blockId="7.[189,776,315,1803]" box="[226,527,1782,1803]" pageId="7" pageNumber="228">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB01FFD003911AB8634A6866" box="[409,524,1782,1803]" name="Kyrgyzstan" pageId="7" pageNumber="228">Kyrgyzstan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

469
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@ -0,0 +1,469 @@
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<mods:title id="C1D44BD3B823A92D9D851BAF9FE30FA9">Dance flies (Diptera: Empididae) in A. P. Fedtschenko’s Collection from Turkestan: Empis subgenus Polyblepharis</mods:title>
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<mods:role id="79FCD065689C279E95BDA728E8F17994">
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</mods:role>
<mods:namePart id="84075CA8662737F470165E3FA5DFF05D">Shamshev, I. V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="16AF49A6BAC216B7DC135DF851251F36">text</mods:typeOfResource>
<mods:relatedItem id="A74CAEB8DCEAF356915F390401455E90" type="host">
<mods:titleInfo id="D7AEE477D85932F7CDAA42F465329BB4">
<mods:title id="33A31657F048EEDEEEA287CD143747C7">Russian Entomological Journal</mods:title>
</mods:titleInfo>
<mods:part id="9FBE2D6170167407CCB986D64CC108D3">
<mods:date id="F80B45CAA54561DB3805365276107F3F">2023</mods:date>
<mods:detail id="240FDF55A56432F29F9C54B24F7D6059" type="pubDate">
<mods:number id="7D9D4BA274D4A02BC83653018E5EDFB5">2023-06-30</mods:number>
</mods:detail>
<mods:detail id="C84F58E01D845B8C9AEA139D09D30295" type="volume">
<mods:number id="1FC2DA7E4704159E81564F8179ED01A0">32</mods:number>
</mods:detail>
<mods:detail id="32DC7CCEEB949D4E7888B3AD3DFB4E94" type="issue">
<mods:number id="DAAC43C4D8132F7A006FAF7294022CBA">2</mods:number>
</mods:detail>
<mods:extent id="9BC33034BEECB8BA51BB7BB029990C41" unit="page">
<mods:start id="E90DCA45C866E108D0641A55F4B001E1">221</mods:start>
<mods:end id="0F7DBAF4B4F036588541EC93D108C547">233</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="8A2031DFD863B03F856E05B475539A47">
<mods:url id="1E6F8FB66F371444926A8F7CF14A69CD">http://dx.doi.org/10.15298/rusentj.32.2.13</mods:url>
</mods:location>
<mods:classification id="88E5B10A11327777EA1B6C730C3118FD">journal article</mods:classification>
<mods:identifier id="B23640534F1C3756323BC39D8997DE0D" type="DOI">10.15298/rusentj.32.2.13</mods:identifier>
<mods:identifier id="5DB2A962DB5E493DA6CA48619040EBC8" type="ISSN">0132-8069</mods:identifier>
</mods:mods>
<treatment id="527C87DDBB05FFD202CD195D65AA6B3A" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A" httpUri="http://treatment.plazi.org/id/527C87DDBB05FFD202CD195D65AA6B3A" lastPageId="5" lastPageNumber="226" pageId="3" pageNumber="224">
<subSubSection id="92CF6540BB05FFD402CD195D63B86A47" box="[197,766,1298,1322]" pageId="3" pageNumber="224" type="nomenclature">
<paragraph id="DA6A36CBBB05FFD402CD195D63B86A47" blockId="3.[197,766,1298,1351]" box="[197,766,1298,1322]" pageId="3" pageNumber="224">
<heading id="812281A7BB05FFD402CD195D63B86A47" box="[197,766,1298,1322]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="224" reason="8">
<taxonomicName id="1DD54D48BB05FFD402CD195D63FB6A47" ID-CoL="C2T2B" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[197,701,1298,1322]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB05FFD402CD195D604E6A47" box="[197,264,1299,1322]" italics="true" pageId="3" pageNumber="224">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB05FFD40310195C60E86A47" box="[280,430,1298,1322]" italics="true" pageId="3" pageNumber="224">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB05FFD403B7195C63026A47" box="[447,580,1298,1322]" italics="true" pageId="3" pageNumber="224">fedtschenkoi</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB05FFD400C0195C63B86A47" bold="true" box="[712,766,1298,1322]" pageId="3" pageNumber="224">
<taxonomicNameLabel id="F39257A2BB05FFD400C0195C63B86A47" box="[712,766,1298,1322]" pageId="3" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD403A7196163526A2A" box="[431,532,1327,1351]" pageId="3" pageNumber="224" type="description">
<paragraph id="DA6A36CBBB05FFD403A7196163526A2A" blockId="3.[197,766,1298,1351]" box="[431,532,1327,1351]" pageId="3" pageNumber="224">
<figureCitation id="42EE2A4EBB05FFD403A7196163576A2A" box="[431,529,1327,1351]" captionStart="Figs 6–9" captionStartId="4.[226,263,1917,1936]" captionTargetBox="[189,1390,607,1892]" captionTargetId="figure-311@4.[188,1398,587,1900]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="3" pageNumber="224">Figs 6–9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD402EA191263016982" pageId="3" pageNumber="224" type="materials_examined">
<paragraph id="DA6A36CBBB05FFD402EA191263646ADA" blockId="3.[189,776,1369,2068]" pageId="3" pageNumber="224">
<materialsCitation id="6ABD3C96BB05FFD402EA191263646ADA" collectingDate="1869-06-19" collectionCode="TYPE, MATERIAL, ZMMU" collectorName="A. Fedtschenko &amp; Cyrillic &amp; Russian &amp; Fan" country="Tajikistan" location="Fan Darya River" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="BCC4AE0EBB05FFD402EA191260D66A02" box="[226,400,1372,1391]" pageId="3" pageNumber="224">TYPE MATERIAL</collectionCode>
.
<typeStatus id="056E8869BB05FFD40392191260AA6A02" box="[410,492,1372,1391]" pageId="3" pageNumber="224" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CCD3FD42BB05FFD403FE1917634C6A02" box="[502,522,1369,1391]" count="1" pageId="3" pageNumber="224" type="male">♂</specimenCount>
, labels as in
<figureCitation id="42EE2A4EBB05FFD40088191263F36A02" box="[640,693,1372,1391]" captionStart="Figs 6–9" captionStartId="4.[226,263,1917,1936]" captionTargetBox="[189,1390,607,1892]" captionTargetId="figure-311@4.[188,1398,587,1900]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="3" pageNumber="224">Fig. 9</figureCitation>
, [
<emphasis id="E8A1EAD9BB05FFD400CF191261A56AEA" bold="true" pageId="3" pageNumber="224">
<collectingCountry id="A2C2765BBB05FFD400CF191261A56AEA" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
</emphasis>
] small bluish green label only with number 19. [=
<date id="AE6B100BBB05FFD400AB193A62476AEA" box="[675,769,1396,1415]" pageId="3" pageNumber="224" value="1869-06-19">
<collectingDate id="BE2FE9E3BB05FFD400AB193A62476AEA" box="[675,769,1396,1415]" pageId="3" pageNumber="224" value="1869-06-19">19.vi.1869</collectingDate>
</date>
] [
<collectorName id="7720531DBB05FFD402CA19C260086AF2" box="[194,334,1420,1439]" pageId="3" pageNumber="224">A. Fedtschenko</collectorName>
] // [printed in
<collectorName id="7720531DBB05FFD403C019C2634A6AF2" box="[456,524,1420,1439]" pageId="3" pageNumber="224">Cyrillic</collectorName>
,
<collectorName id="7720531DBB05FFD4001C19C2631D6AF2" box="[532,603,1420,1439]" pageId="3" pageNumber="224">Russian</collectorName>
, pre-1918 orthography]
<collectorName id="7720531DBB05FFD402E219EA604C6ADA" box="[234,266,1444,1463]" pageId="3" pageNumber="224">Fan</collectorName>
[=
<location id="DF0A6010BB05FFD4032319EA60F86ADA" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD4032319EA60F86ADA" box="[299,446,1444,1463]" country="Tajikistan" name="Fan Darya River" pageId="3" pageNumber="224">Fan Darya River</location>
] (
<collectionCode id="BCC4AE0EBB05FFD403D819EA635E6ADA" box="[464,536,1444,1463]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="224" type="Museum">ZMMU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DA6A36CBBB05FFD402EA19F263016982" blockId="3.[189,776,1369,2068]" pageId="3" pageNumber="224">
<materialsCitation id="6ABD3C96BB05FFD402EA19F260976A8A" collectingDate="1869-06-19" collectionCode="ZISP" collectorName="A. Fedtschenko" country="Tajikistan" location="Fan Darya River" pageId="3" pageNumber="224" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="056E8869BB05FFD402EA19F2607F6AA2" box="[226,313,1468,1487]" pageId="3" pageNumber="224" type="paratype">Paratypes</typeStatus>
.
<emphasis id="E8A1EAD9BB05FFD4034919F260E16AA2" bold="true" box="[321,423,1468,1487]" pageId="3" pageNumber="224">
<collectingCountry id="A2C2765BBB05FFD4034919F260E56AA2" box="[321,419,1468,1487]" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
:
</emphasis>
same data as holotype [
<collectorName id="7720531DBB05FFD4007F19F262476AA2" box="[631,769,1468,1487]" pageId="3" pageNumber="224">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB05FFD402CD199A61A16A8A" box="[197,231,1489,1511]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB05FFD402F9199A604A6A8A" box="[241,268,1489,1511]" count="1" pageId="3" pageNumber="224" type="female">1 ♀</specimenCount>
;
<specimenCount id="CCD3FD42BB05FFD4031E199A607F6A8A" box="[278,313,1489,1511]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
, dissected,
<collectionCode id="BCC4AE0EBB05FFD40393199A608D6A8A" box="[411,459,1492,1511]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="224">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403D1199A60276A92" collectingDate="1869-06-21" collectionCode="ZMMU" collectorName="A. Fedtschenko" country="Tajikistan" location="Yagnob" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB05FFD403D1199A63596A8A" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD403D1199A63596A8A" box="[473,543,1492,1511]" country="Tajikistan" name="Yagnob" pageId="3" pageNumber="224">Yagnob</location>
,
<date id="AE6B100BBB05FFD4002F199A63C56A8A" box="[551,643,1492,1511]" pageId="3" pageNumber="224" value="1869-06-21">
<collectingDate id="BE2FE9E3BB05FFD4002F199A63C56A8A" box="[551,643,1492,1511]" pageId="3" pageNumber="224" value="1869-06-21">21.vi.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB05FFD40085199A61936A92" pageId="3" pageNumber="224">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB05FFD402EF19A2604C6A92" box="[231,266,1513,1535]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD4031C19A2601D6A92" box="[276,347,1516,1535]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="224" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD4036219A26017697A" collectingDate="1869-06-02" collectionCode="ZMMU" collectorName="A. Fedtschenko" country="Tajikistan" location="Turkestanskiy" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB05FFD4036219A260A26A92" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD4036219A260A26A92" box="[362,484,1516,1535]" country="Tajikistan" name="Turkestanskiy" pageId="3" pageNumber="224">Turkestanskiy</location>
kray,
<date id="AE6B100BBB05FFD4001219A2632A6A92" box="[538,620,1516,1535]" pageId="3" pageNumber="224" value="1869-06-02">
<collectingDate id="BE2FE9E3BB05FFD4001219A2632A6A92" box="[538,620,1516,1535]" pageId="3" pageNumber="224" value="1869-06-02">2.vi.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB05FFD4007F19A262446A92" box="[631,770,1516,1535]" pageId="3" pageNumber="224">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB05FFD402CD1A4A61A9697A" box="[197,239,1537,1559]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD4030A1A4A600D697A" box="[258,331,1540,1559]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="224" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403691A4A609A6942" collectingDate="1986-06-07" collectingDateMax="1986-06-08" collectingDateMin="1986-06-07" collectionCode="ZISP" collectorName="M. Dolgovskaya" country="Tajikistan" location="Novabad" municipality="Komarou Gorge" pageId="3" pageNumber="224" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<location id="DF0A6010BB05FFD403691A4A60F3697A" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD403691A4A60F3697A" box="[353,437,1540,1559]" country="Tajikistan" municipality="Komarou Gorge" name="Novabad" pageId="3" pageNumber="224">Novabad</location>
,
<collectingMunicipality id="3A0EACB1BB05FFD403CC1A4A6319697A" box="[452,607,1540,1559]" pageId="3" pageNumber="224">Komarou Gorge</collectingMunicipality>
,
<date id="AE6B100BBB05FFD400671A4A6398697A" box="[623,734,1540,1559]" pageId="3" pageNumber="224" value="1986-06-07" valueMax="1986-06-08" valueMin="1986-06-07">
<collectingDate id="BE2FE9E3BB05FFD400671A4A6398697A" box="[623,734,1540,1559]" pageId="3" pageNumber="224" value="1986-06-07" valueMax="1986-06-08" valueMin="1986-06-07">7–8.vi.1986</collectingDate>
</date>
,
<collectorName id="7720531DBB05FFD400E51A4A60696942" pageId="3" pageNumber="224">M. Dolgovskaya</collectorName>
(
<specimenCount id="CCD3FD42BB05FFD403321A5260356942" box="[314,371,1561,1583]" count="2" pageId="3" pageNumber="224" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB05FFD403771A5260DD6942" box="[383,411,1561,1583]" count="1" pageId="3" pageNumber="224" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD403AE1A5260906942" box="[422,470,1564,1583]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="224">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403ED1A5260AF6932" collectingDate="2016-05-24" collectionCode="SZMN, ZISP" collectorName="A. Barkalov" country="Tajikistan" elevation="1900" latitude="38.65" location="Tavildara District" longLatPrecision="704" longitude="70.52" municipality="Kolon" pageId="3" pageNumber="224" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<location id="DF0A6010BB05FFD403ED1A5263C76942" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD403ED1A5263C76942" box="[485,641,1564,1583]" country="Tajikistan" latitude="38.65" longLatPrecision="704" longitude="70.52" municipality="Kolon" name="Tavildara District" pageId="3" pageNumber="224">Tavildara District</location>
, village Dekhi-
<collectingMunicipality id="3A0EACB1BB05FFD402B51A7A61B3692A" box="[189,245,1588,1607]" pageId="3" pageNumber="224">Kolon</collectingMunicipality>
,
<geoCoordinate id="BFE1500CBB05FFD402F41A7A6003692A" box="[252,325,1588,1607]" degrees="38,65" direction="north" orientation="latitude" pageId="3" pageNumber="224" precision="555" value="38.65">38,65°N</geoCoordinate>
<geoCoordinate id="BFE1500CBB05FFD403411A7A60D4692A" box="[329,402,1588,1607]" degrees="70,52" direction="east" orientation="longitude" pageId="3" pageNumber="224" precision="555" value="70.52">70,52°E</geoCoordinate>
,
<quantity id="1D2D9B2EBB05FFD403911A7A634A692A" box="[409,524,1588,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="3" pageNumber="224" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<elevation id="51F8D1F8BB05FFD403911A7A634A692A" box="[409,524,1588,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="3" pageNumber="224" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000m</elevation>
</quantity>
,
<date id="AE6B100BBB05FFD4001C1A7A632A692A" box="[532,620,1588,1607]" pageId="3" pageNumber="224" value="2016-05-24">
<collectingDate id="BE2FE9E3BB05FFD4001C1A7A632A692A" box="[532,620,1588,1607]" pageId="3" pageNumber="224" value="2016-05-24">24.v.2016</collectingDate>
</date>
, coll.
<collectorName id="7720531DBB05FFD400961A7A6240692A" box="[670,774,1588,1607]" pageId="3" pageNumber="224">A. Barkalov</collectorName>
(
<specimenCount id="CCD3FD42BB05FFD402CC1A0261B86932" box="[196,254,1609,1631]" count="3" pageId="3" pageNumber="224" type="male">3 ♂♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD403031A02600F6932" box="[267,329,1612,1631]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="3" pageNumber="224">SZMN</collectionCode>
;
<specimenCount id="CCD3FD42BB05FFD4035C1A02603C6932" box="[340,378,1609,1631]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB05FFD4038F1A0260E06932" box="[391,422,1609,1631]" count="1" pageId="3" pageNumber="224" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD403BB1A0260A56932" box="[435,483,1612,1631]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="224">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403FC1A026044691A" collectingDate="2016-05-26" collectionCode="SZMN" collectorName="A. Barkalov" country="Tajikistan" elevation="1900" latitude="38.65" location="Tavildara District" longLatPrecision="704" longitude="70.52" municipality="Kolon" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality,
<date id="AE6B100BBB05FFD4007E1A02638A6932" box="[630,716,1612,1631]" pageId="3" pageNumber="224" value="2016-05-26">
<collectingDate id="BE2FE9E3BB05FFD4007E1A02638A6932" box="[630,716,1612,1631]" pageId="3" pageNumber="224" value="2016-05-26">26.v.2016</collectingDate>
</date>
(
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,
<collectionCode id="BCC4AE0EBB05FFD402B51A2A61BB691A" box="[189,253,1636,1655]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="3" pageNumber="224">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403041A2A60A3691A" box="[268,485,1633,1655]" collectingDate="2016-05-15" collectionCode="SZMN" collectorName="A. Barkalov" country="Tajikistan" elevation="1900" isTailEnumeration="true" latitude="38.65" location="Tavildara District" longLatPrecision="704" longitude="70.52" municipality="Kolon" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="AE6B100BBB05FFD403041A2A6024691A" box="[268,354,1636,1655]" pageId="3" pageNumber="224" value="2016-05-15">
<collectingDate id="BE2FE9E3BB05FFD403041A2A6024691A" box="[268,354,1636,1655]" pageId="3" pageNumber="224" value="2016-05-15">15.v.2016</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB05FFD403781A2A60D2691A" box="[368,404,1633,1655]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD403971A2A6099691A" box="[415,479,1636,1655]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="3" pageNumber="224">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD403E71A2A638E691A" box="[495,712,1633,1655]" collectingDate="2016-05-17" collectionCode="SZMN" country="Tajikistan" pageId="3" pageNumber="224" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB05FFD403E71A2A6302691A" box="[495,580,1636,1655]" pageId="3" pageNumber="224" value="2016-05-17">
<collectingDate id="BE2FE9E3BB05FFD403E71A2A6302691A" box="[495,580,1636,1655]" pageId="3" pageNumber="224" value="2016-05-17">17.v.2016</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB05FFD4005A1A2A6331691A" box="[594,631,1633,1655]" count="2" pageId="3" pageNumber="224" type="male">2 ♂</specimenCount>
,
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)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD400DA1A2A637669CA" collectingDate="2016-05-25" collectionCode="SZMN" collectorName="A. Barkalov" country="Tajikistan" elevation="1886" latitude="38.66" location="Tavildara District" longLatPrecision="704" longitude="70.49" municipality="Sary-Dzhangal" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB05FFD400DA1A2A606E69E2" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD400DA1A2A606E69E2" country="Tajikistan" latitude="38.66" longLatPrecision="704" longitude="70.49" municipality="Sary-Dzhangal" name="Tavildara District" pageId="3" pageNumber="224">Tavildara District</location>
,village
<collectingMunicipality id="3A0EACB1BB05FFD403641A3260A969E2" box="[364,495,1660,1679]" pageId="3" pageNumber="224">Sary-Dzhangal</collectingMunicipality>
,
<geoCoordinate id="BFE1500CBB05FFD403FD1A32637869E2" box="[501,574,1660,1679]" degrees="38,66" direction="north" orientation="latitude" pageId="3" pageNumber="224" precision="555" value="38.66">38,66°N</geoCoordinate>
<geoCoordinate id="BFE1500CBB05FFD400491A3263CF69E2" box="[577,649,1660,1679]" degrees="70,49" direction="east" orientation="longitude" pageId="3" pageNumber="224" precision="555" value="70.49">70,49°E</geoCoordinate>
,
<quantity id="1D2D9B2EBB05FFD400981A32624269E2" box="[656,772,1660,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.8855" metricValueMax="1.9449999999999998" metricValueMin="1.8259999999999998" pageId="3" pageNumber="224" unit="m" value="1885.5" valueMax="1945.0" valueMin="1826.0">
<elevation id="51F8D1F8BB05FFD400981A32624269E2" box="[656,772,1660,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.8855" metricValueMax="1.9449999999999998" metricValueMin="1.8259999999999998" pageId="3" pageNumber="224" unit="m" value="1885.5" valueMax="1945.0" valueMin="1826.0">1826–1945m</elevation>
</quantity>
,
<date id="AE6B100BBB05FFD402B51ADA605369CA" box="[189,277,1684,1703]" pageId="3" pageNumber="224" value="2016-05-25">
<collectingDate id="BE2FE9E3BB05FFD402B51ADA605369CA" box="[189,277,1684,1703]" pageId="3" pageNumber="224" value="2016-05-25">25.v.2016</collectingDate>
</date>
, coll.
<collectorName id="7720531DBB05FFD403401ADA60F769CA" box="[328,433,1684,1703]" pageId="3" pageNumber="224">A. Barkalov</collectorName>
(
<specimenCount id="CCD3FD42BB05FFD403B61ADA60A669CA" box="[446,480,1681,1703]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD403E21ADA636C69CA" box="[490,554,1684,1703]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="3" pageNumber="224">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB05FFD400301ADA60CF69BA" collectingDate="2017-06-30" collectionCode="SZMN" collectorName="A. Barkalov" country="Tajikistan" elevation="2440" latitude="39.06" location="Varzab Gorge" longLatPrecision="703" longitude="68.87" municipality="Kalon" pageId="3" pageNumber="224" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="DF0A6010BB05FFD400301ADA63F469CA" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD400301ADA63F469CA" box="[568,690,1684,1703]" country="Tajikistan" latitude="39.06" longLatPrecision="703" longitude="68.87" municipality="Kalon" name="Varzab Gorge" pageId="3" pageNumber="224">Varzab Gorge</location>
,
<quantity id="1D2D9B2EBB05FFD400B21ADA63A369CA" box="[698,741,1684,1703]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="224" unit="km" value="3.0">3 km</quantity>
NE of village
<collectingMunicipality id="3A0EACB1BB05FFD403161AE2601369D2" box="[286,341,1708,1727]" pageId="3" pageNumber="224">Kalon</collectingMunicipality>
,
<quantity id="1D2D9B2EBB05FFD4036A1AE260D969D2" box="[354,415,1708,1727]" metricMagnitude="3" metricUnit="m" metricValue="2.44" pageId="3" pageNumber="224" unit="m" value="2440.0">
<elevation id="51F8D1F8BB05FFD4036A1AE260D969D2" box="[354,415,1708,1727]" metricMagnitude="3" metricUnit="m" metricValue="2.44" pageId="3" pageNumber="224" unit="m" value="2440.0">2440m</elevation>
</quantity>
,
<geoCoordinate id="BFE1500CBB05FFD403A51AE260B069D2" box="[429,502,1708,1727]" degrees="39,06" direction="north" orientation="latitude" pageId="3" pageNumber="224" precision="555" value="39.06">39,06°N</geoCoordinate>
<geoCoordinate id="BFE1500CBB05FFD400081AE2630E69D2" box="[512,584,1708,1727]" degrees="68,87" direction="east" orientation="longitude" pageId="3" pageNumber="224" precision="555" value="68.87">68,87°E</geoCoordinate>
,
<date id="AE6B100BBB05FFD4005E1AE263F269D2" box="[598,692,1708,1727]" pageId="3" pageNumber="224" value="2017-06-30">
<collectingDate id="BE2FE9E3BB05FFD4005E1AE263F269D2" box="[598,692,1708,1727]" pageId="3" pageNumber="224" value="2017-06-30">30.vi.2017</collectingDate>
</date>
, coll.
<collectorName id="7720531DBB05FFD400FA1AE2604A69BA" pageId="3" pageNumber="224">A. Barkalov</collectorName>
(
<specimenCount id="CCD3FD42BB05FFD403131A8A607169BA" box="[283,311,1729,1751]" count="1" pageId="3" pageNumber="224" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD4034B1A8A60C569BA" box="[323,387,1732,1751]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="3" pageNumber="224">SZMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ABD3C96BB05FFD4039B1A8A63056982" collectingDate="1897-05-31" collectionCode="ZISP" collectorName="Kaznakov" country="Uzbekistan" location="Salbar" municipality="Ak-su" pageId="3" pageNumber="224" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="E8A1EAD9BB05FFD4039B1A8A634769BA" bold="true" box="[403,513,1732,1751]" pageId="3" pageNumber="224">
<collectingCountry id="A2C2765BBB05FFD4039B1A8A60BB69BA" box="[403,509,1732,1751]" name="Uzbekistan" pageId="3" pageNumber="224">Uzbekistan</collectingCountry>
:
</emphasis>
<location id="DF0A6010BB05FFD4000F1A8A630669BA" LSID="urn:lsid:plazi:treatment:527C87DDBB05FFD202CD195D65AA6B3A:DF0A6010BB05FFD4000F1A8A630669BA" box="[519,576,1732,1751]" country="Uzbekistan" municipality="Ak-su" name="Salbar" pageId="3" pageNumber="224">Salbar</location>
, valley of
<collectingMunicipality id="3A0EACB1BB05FFD400971A8A639069BA" box="[671,726,1732,1751]" pageId="3" pageNumber="224">Ak-su</collectingMunicipality>
,
<locationDeviation id="CBD82FE9BB05FFD400E91A8A604E6982" location="Bukhara" pageId="3" pageNumber="224">E of Bukhara</locationDeviation>
,
<date id="AE6B100BBB05FFD4031A1A92602C6982" box="[274,362,1756,1775]" pageId="3" pageNumber="224" value="1897-05-31">
<collectingDate id="BE2FE9E3BB05FFD4031A1A92602C6982" box="[274,362,1756,1775]" pageId="3" pageNumber="224" value="1897-05-31">31.v.1897</collectingDate>
</date>
,
<collectorName id="7720531DBB05FFD4037D1A92608A6982" box="[373,460,1756,1775]" pageId="3" pageNumber="224">Kaznakov</collectorName>
(
<specimenCount id="CCD3FD42BB05FFD403D31A9263466982" box="[475,512,1753,1775]" count="1" pageId="3" pageNumber="224" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB05FFD400041A92637A6982" box="[524,572,1756,1775]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="224">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD402EA1ABA63EB68B3" pageId="3" pageNumber="224" type="diagnosis">
<paragraph id="DA6A36CBBB05FFD402EA1ABA63EB68B3" blockId="3.[189,776,1369,2068]" pageId="3" pageNumber="224">
DIAGNOSIS. Mid-sized species (body length
<quantity id="1D2D9B2EBB05FFD400901ABA63BA6864" box="[664,764,1780,1802]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="3" pageNumber="224" unit="mm" value="6.25" valueMax="6.5" valueMin="6.0">6–6.5 mm</quantity>
); male eyes holoptic; labrum nearly 1.5X longer than eye height, palpus yellow to brownish yellow on apical half; prosternum setose; mesoscutum with 4 distinct vittae, acrostichal and dorsocentral setae multiserial, laterotergite with pale setae; legs entirely brown, fore and mid basitarsi long setose; abdomen mostly pale setose, tergites 2–5 shiny dorsally; female hind femur flattened, somewhat convex dorsally and straight ventrally, with slightly flattened setae.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD202EA1BAA62B96C86" lastPageId="5" lastPageNumber="226" pageId="3" pageNumber="224" type="description">
<paragraph id="DA6A36CBBB05FFD402EA1BAA64796D8C" blockId="3.[189,776,1369,2068]" lastBlockId="3.[812,1399,315,2071]" pageId="3" pageNumber="224">
DESCRIPTION. Body length about
<quantity id="1D2D9B2EBB05FFD400581BAA638F6894" box="[592,713,2020,2042]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="3" pageNumber="224" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.8–6.4 mm</quantity>
, wing
<quantity id="1D2D9B2EBB05FFD402B51BB060716779" box="[189,311,2046,2068]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.5" metricValueMin="5.9" pageId="3" pageNumber="224" unit="mm" value="6.2" valueMax="6.5" valueMin="5.9">5.9–6.5 mm</quantity>
.
<emphasis id="E8A1EAD9BB05FFD4034E1BB0603F677E" bold="true" box="[326,377,2046,2067]" pageId="3" pageNumber="224">Male</emphasis>
(
<figureCitation id="42EE2A4EBB05FFD403841BB0608A6779" box="[396,460,2046,2068]" captionStart="Figs 6–9" captionStartId="4.[226,263,1917,1936]" captionTargetBox="[189,1390,607,1892]" captionTargetId="figure-311@4.[188,1398,587,1900]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="3" pageNumber="224">Fig. 6</figureCitation>
). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Holoptic, upper ommatidia enlarged. Frons represented by very small subtriangular space just below ocellar triangle and somewhat larger space above antennae; bare. Face broad, bare. Ocellar setae undifferentiated; ocellar triangle with several subequally short, hair-like setae. Occiput covered with long, dense, fine setae (including postoculars); postgena with pale hair-like setae. Antenna with scape and pedicel light brown, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with moderately long setulae; postpedicel 3X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow; nearly 1.5X longer than eye height; palpus short, yellowish to brownish yellow on about apical half, brownish towards base; bearing numerous long, black, fine setae.
</paragraph>
<paragraph id="DA6A36CBBB05FFD4015A1EA8653C6A8A" blockId="3.[812,1399,315,2071]" pageId="3" pageNumber="224">Thorax almost entirely black in ground-colour, postalar tubercle somewhat yellowish brown translucent;mostly densely greyish pruinose; mesoscutum with some bluish green tinge, 4 distinct brown vittae (lateral vittae slightly broader). Prosternum with 3–4 long pale setae near apex on each side.Proepisternum with tuft of numerous, pale, fine setae on lower part and several similar setae on upper part. Antepronotum with several short to moderately long, strong, black setae dorsally and long, fine, pale setae laterally. Postpronotal lobe covered with pale, dense, long, fine setae; postpronotal seta undifferentiated, at most 1–4 black setae of the same robustness and length as surrounding pale setae. Mesonotal setae: acrostichals arranged in 4–5 irregular rows, long, fine, dense, usually intermixed black and pale anteriorly, absent on prescutellar depression, separated from dorsocentrals by narrow space; presutural dorsocentral setae multiserial and not separated from setae of presutural supra-alar space, similar to acrostichals, pale anteriorly at least near postpronotal lobe, postsutural dorsocentrals arranged in 2–3 irregular rows, sparser, becoming longer and stronger towards scutellum, 1–2 very long strong prescutellars; presutural supra-alar seta undifferentiated, at most 1–2 black setae of the same robustness and length as surrounding pale setae; 4 notopleurals, 1 postsutural supra-alar, 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair slightly shorter); in addition, supra-alar space entirely covered with dense fine setae (similar to dorsocentrals and mostly pale before suture), numerous long, fine, pale setae on anterior half of notopleuron. Laterotergite with numerous paler setae. Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="DA6A36CBBB05FFD3015A19A265DA6D2C" blockId="3.[812,1399,315,2071]" lastBlockId="4.[812,1399,315,577]" lastPageId="4" lastPageNumber="225" pageId="3" pageNumber="224">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose. Legs colour: entirely dark brown to black. Coxae and trochanters mostly with pale hair-like setae, a few black stronger setae (fore coxa sometimes only with pale setae). Fore femur bare along ventral face; with moderately long, fine setae anteroventrally (pale on about basal half); row of 5–7 strong antero-anteroventral setae; numerous fine, long (longer than femur width), mostly pale setae posteroventrally and posteriorly. Fore tibia with row of short, rather strong, anterodorsal setae; covered with long, dense, mostly fine setae posterodorsally and posteriorly (longest setae nearly 1.5X longer than tibia width, somewhat stronger near apex). Fore basitarsus slightly thickened towards base but not broader than fore tibia at apex, with 3–4 short strong anterodorsal setae; dense, intermixed fine and strong, long setae posterodorsally and posteriorly (as on fore tibia apically); anteroventral and posteroventral spine-like setae; long, strong setae of subapical circlet; tarsomere 2 with spinule-like anteroventral and posteroventral setulae. Mid femur with fine, moderately long setae ventrally (pale on about basal half), similar slightly longer setae near apex posteriorly; whitish pubescent ventrally. Mid tibia with rows of long anterodorsal and posterodorsal strong setae (about 7 setae per row, longest setae nearly 1.5X longer than tibia width); 2–3 rather short anteroventral and row of similar posteroventral setae. Mid basitarsus slender; with several long, strong setae dorsally; anteroventral and posteroventral spine-like setae; tarsomere 2 with spinule-like anteroventral and posteroventral setulae. Hind femur slender; with 4–5 moderately long, anterodorsal setae near apex; fine setulae along ventral face; anteroventral and posteroventral setae short, mostly fine, partly pale on basal part, slightly longer, strong anteroventral setae on about apical half (number and robustness variable); no whitish ventral pubescence. Hind tibia simple, slightly narrower near base; with rows of moderately long anterodorsal and posterodorsal setae; simple setulae ventrally; very short seta in posteroapical comb. Hind basitarsus slightly thickened towards base but not broader than hind tibia at apex; several long setae dorsally; tarsomeres 1–3 with short spine-like anteroventral and posteroventral setae ventrally (longer on basitarsus).</paragraph>
<caption id="8EAA6643BB02FFD302EA1B3362DE6760" pageId="4" pageNumber="225" startId="4.[226,263,1917,1936]" targetBox="[189,1390,607,1892]" targetPageId="4" targetType="figure">
<paragraph id="DA6A36CBBB02FFD302EA1B33628568AC" blockId="4.[189,1398,1917,2061]" pageId="4" pageNumber="225">
Figs 6–9.
<taxonomicName id="1DD54D48BB02FFD303321B30638968FD" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[314,719,1917,1937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB02FFD303321B30602968FD" box="[314,367,1918,1936]" italics="true" pageId="4" pageNumber="225">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB02FFD303751B3360B068FD" box="[381,502,1917,1936]" italics="true" pageId="4" pageNumber="225">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB02FFD3000B1B33632968FD" box="[515,623,1917,1936]" italics="true" pageId="4" pageNumber="225">fedtschenkoi</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB02FFD300D11B30624268FC" bold="true" box="[729,772,1918,1937]" pageId="4" pageNumber="225">
<taxonomicNameLabel id="F39257A2BB02FFD300D11B30624268FC" box="[729,772,1918,1937]" pageId="4" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm.
</paragraph>
<paragraph id="DA6A36CBBB02FFD302EA1B8462DE6760" blockId="4.[189,1398,1917,2061]" pageId="4" pageNumber="225">
Рис. 6–9.
<taxonomicName id="1DD54D48BB02FFD303341B84639568B1" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[316,723,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB02FFD303341B84603768B1" box="[316,369,1994,2012]" italics="true" pageId="4" pageNumber="225">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB02FFD303881B8760BF68B1" box="[384,505,1993,2012]" italics="true" pageId="4" pageNumber="225">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB02FFD3000F1B87633568B1" box="[519,627,1993,2012]" italics="true" pageId="4" pageNumber="225">fedtschenkoi</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB02FFD300D61B87624E68B1" bold="true" box="[734,776,1993,2012]" pageId="4" pageNumber="225">
<taxonomicNameLabel id="F39257A2BB02FFD300D61B87624E68B1" box="[734,776,1993,2012]" pageId="4" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм.
</paragraph>
</caption>
<paragraph id="DA6A36CBBB03FFD202EA1D7560316D31" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="226">
Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins mostly brownish, brownish yellow near base, well-sclerotized. Veins R
<subScript id="4651348EBB03FFD200451DCE63126EE1" attach="left" box="[589,596,384,396]" fontSize="5" pageId="5" pageNumber="226">5</subScript>
and M
<subScript id="4651348EBB03FFD200941DCE63E56EE1" attach="left" box="[668,675,384,396]" fontSize="5" pageId="5" pageNumber="226">1</subScript>
divergent towards wing margin; radial fork acute; R
<subScript id="4651348EBB03FFD200631DD463346ECB" attach="left" box="[619,626,410,422]" fontSize="5" pageId="5" pageNumber="226">5</subScript>
meeting costa before wing apex; CuA+CuP (anal vein) complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 2.5X longer than its middle portion. Pterostigma pale brownish, very narrow. Basal costal seta absent. Anal lobe well-developed; axillary incision very acute.Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<paragraph id="DA6A36CBBB03FFD202EA1E2F63236B3A" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="226">Abdomen dark brown in ground colour; tergite 1 entirely densely greyish pruinose, tergites 2–5 shiny dorsally and greyish pruinose laterally (in addition, tergite 2 narrowly pruinose anteriorly); tergites 6 and 7 with spots of brown iridescent pruinosity dorsally, greyish pruinose laterally; sternites entirely greyish pruinose, slightly finer on sternite 8. Abdomen pale setose, only tergites 2–5 with strong yellow and sternite 8 with black posteromarginal setae; tergites 1–5 laterally and sternites (including sternite 1) covered with dense, long, fine setae. Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; pleuron 7 with strongly sclerotised posterior margin covered with minute setulae; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, somewhat constricted anteriorly, with 2 subequal lateral projections on each side anteriorly (anterior projection close to upper margin).</paragraph>
<paragraph id="DA6A36CBBB03FFD202EA1815634B695A" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="226">
Hypopygium (
<figureCitation id="42EE2A4EBB03FFD20364181560E26B1D" box="[364,420,1115,1137]" captionStart="Figs 6–9" captionStartId="4.[226,263,1917,1936]" captionTargetBox="[189,1390,607,1892]" captionTargetId="figure-311@4.[188,1398,587,1900]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="5" pageNumber="226">Fig. 7</figureCitation>
) moderately large; epandrium brown, subshiny; cerci brownish yellow (darker anteriorly); phallus yellowish brown. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella subtriangular (lateral view); with long, fine, pale setae more numerous closer to lower margin. Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, with rounded apex; bare; gonocoxal apodeme small. Cerci (
<figureCitation id="42EE2A4EBB03FFD202F7197F607F6A2B" box="[255,313,1329,1351]" captionStart="Figs 6–9" captionStartId="4.[226,263,1917,1936]" captionTargetBox="[189,1390,607,1892]" captionTargetId="figure-311@4.[188,1398,587,1900]" captionTargetPageId="4" captionText="Figs 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., male: 6 — holotype, habitus; 7 — hypopygium, setation of epandrium and cerci omitted; 8 — cerci; 9 — holotype labels; 6–7 — lateral view; 8 — dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 6–9. Empis (Polyblepharis) fedtschenkoi Shamshev, sp.n., самец: 6 — голотип, габитус; 7 — гипопигий, Щетинки ЭпандриЯ и церок не укаЗаны; 8 — церки; 9 — Этикетки голотипа; 6–7 — сбоку; 8 — сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="5" pageNumber="226">Fig. 8</figureCitation>
) separated with each other and from hypandrium; cercus elongate ovate (lateral view), shorter than epandrial lamella, without additional lobe dorsally, serrate on inner margin; with dark and pale setulae. Phallus almost entirely hidden; zigzag bent, strongly constricted beyond middle; basal portion thick, gently curved and becoming narrower distad; apical bend portion semicircular, nearly of uniform thickness, with its tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="DA6A36CBBB03FFD202EA1A7565C06D2C" blockId="5.[189,776,315,2071]" lastBlockId="5.[812,1399,315,1111]" pageId="5" pageNumber="226">
<emphasis id="E8A1EAD9BB03FFD202EA1A756072693D" bold="true" box="[226,308,1595,1616]" pageId="5" pageNumber="226">Female.</emphasis>
Similar to male except as follows. Eyes dichoptic, ommatidia equally small. Frons broad, parallel-sided. Ocellar triangle with 2 long, stronger and several short, fine setae. Occiput with somewhat sparser and stronger setae. Palpus mostly yellow, brownish near base. Postpronotal lobe and mesonotum with slightly shorter fine setae; postpronotal and presutural supra-alar setae usually more or less differentiated but only slightly stronger than surrounding fine setae. Legs generally shorter setose. Fore tibia with short fine setae posterodorsally and posteriorly. Fore basitarsus slender; only with simple setulae dorsally and posteriorly. Mid tibia with slightly shorter anterodorsal and posterodorsal setae; almost complete rows of similar anteroventral and posteroventral setae. Mid basitarsus without long setae dorsally. Hind femur flattened, somewhat convex dorsally and straight ventrally; with fringe of short, slightly flattened, short, dense setae almost over entire length dorsally and ventrally (setae finer near base), row of anteroanteroventral moderately long setae. Hind tibia slightly flattened; with denser setulae dorsally and ventrally; rows of short anterodorsal and posterodorsal setae, 2–3 similar anteroventral setae on apical half. Hind basitarsus slender; without long setae dorsally. Abdomen with tergites 2–8 extensively shiny; tergite 1 entirely and tergites 2–6 narrowly laterally densely greyish pruinose; tergites covered with mostly short fine setae (longer on tergites 1–3 laterally), no stronger posteromarginal setae. Cercus brown, short, covered with dense, pale, erect setulae.
</paragraph>
<paragraph id="DA6A36CBBB03FFD2015A1E0864306C0B" blockId="5.[812,1399,315,1111]" pageId="5" pageNumber="226">
DIFFERENTIAL DIAGNOSIS. Within the key to the Palearctic species of
<taxonomicName id="1DD54D48BB03FFD206091E2F65CE6D1B" box="[1025,1160,609,630]" pageId="5" pageNumber="233" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB03FFD206091E2F65CE6D1B" box="[1025,1160,609,630]" italics="true" pageId="5" pageNumber="226">Polyblepharis</emphasis>
</taxonomicName>
the new species would run to
<taxonomicName id="1DD54D48BB03FFD2017E1E3265E86DFD" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[886,1198,635,657]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="freidbergi">
<emphasis id="E8A1EAD9BB03FFD2017E1E3265586DFD" box="[886,1054,635,657]" italics="true" pageId="5" pageNumber="226">Empis freidbergi</emphasis>
<bibRefCitation id="BE444B3ABB03FFD206211E3565E86DFD" author="Chvala M." box="[1065,1198,635,657]" pageId="5" pageNumber="226" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
known from
<collectingCountry id="A2C2765BBB03FFD207491E3564316DFC" box="[1345,1399,635,657]" name="Israel" pageId="5" pageNumber="226">Israel</collectingCountry>
[
<bibRefCitation id="BE444B3ABB03FFD2013A1ED862FF6DC6" author="Chvala M." box="[818,953,662,684]" pageId="5" pageNumber="226" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
]. Both species share a combination of holoptic eyes of the male, entirely black to brown legs, setose prosternum, pale setose laterotergite and undifferentiated postpronotal and presutural supra-alar setae.
<taxonomicName id="1DD54D48BB03FFD206BD1EA964306D96" authorityName="Shamshev" authorityYear="2023" box="[1205,1398,742,764]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp. nov.">
<emphasis id="E8A1EAD9BB03FFD206BD1EA964306D96" box="[1205,1398,742,764]" italics="true" pageId="5" pageNumber="226">Empis fedtschenkoi</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB03FFD201241F4F621B6C7B" bold="true" box="[812,861,769,790]" pageId="5" pageNumber="226">
<taxonomicNameLabel id="F39257A2BB03FFD201241F4F621B6C7B" box="[812,861,769,790]" pageId="5" pageNumber="226" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
can be readily distinguished from
<taxonomicName id="1DD54D48BB03FFD206A01F4F64646C7B" authorityName="Chvala" authorityYear="1999" box="[1192,1314,769,790]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="freidbergi">
<emphasis id="E8A1EAD9BB03FFD206A01F4F65F16C7B" box="[1192,1207,769,790]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8A1EAD9BB03FFD206CA1F4F64646C7B" box="[1218,1314,769,790]" italics="true" pageId="5" pageNumber="226">freidbergi</emphasis>
</taxonomicName>
primarily by four mesoscutal vittae (versus three). In addition, the male of
<taxonomicName id="1DD54D48BB03FFD201741F7965576C26" authorityName="Shamshev" authorityYear="2023" box="[892,1041,822,844]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp.n.">
<emphasis id="E8A1EAD9BB03FFD201741F7962CD6C21" box="[892,907,823,844]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8A1EAD9BB03FFD201901F7865576C26" box="[920,1041,822,843]" italics="true" pageId="5" pageNumber="226">fedtschenkoi</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB03FFD2061F1F78650E6C26" bold="true" box="[1047,1096,822,843]" pageId="5" pageNumber="226">
<taxonomicNameLabel id="F39257A2BB03FFD2061F1F78650E6C26" box="[1047,1096,822,843]" pageId="5" pageNumber="226" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
has mostly yellowish to brownish yellow palpus (entirely black in the male of
<taxonomicName id="1DD54D48BB03FFD206F91F1F642E6C0B" authorityName="Chvala" authorityYear="1999" box="[1265,1384,849,870]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="freidbergi">
<emphasis id="E8A1EAD9BB03FFD206F91F1F64466C0B" box="[1265,1280,849,870]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8A1EAD9BB03FFD207021F1F642E6C0B" box="[1290,1384,849,870]" italics="true" pageId="5" pageNumber="226">freidbergi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="DA6A36CBBB03FFD2015A1F2562B96C86" blockId="5.[812,1399,315,1111]" pageId="5" pageNumber="226">
Amongst species of
<taxonomicName id="1DD54D48BB03FFD206281F2565E16CED" box="[1056,1191,875,896]" pageId="5" pageNumber="233" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB03FFD206281F2565E16CED" box="[1056,1191,875,896]" italics="true" pageId="5" pageNumber="226">Polyblepharis</emphasis>
</taxonomicName>
of the Middle Asia, setose prosternum was described only in
<taxonomicName id="1DD54D48BB03FFD206CC1FC962F46CDB" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="alatauensis">
<emphasis id="E8A1EAD9BB03FFD206CC1FC964306CF6" box="[1220,1398,902,924]" italics="true" pageId="5" pageNumber="226">Empis alatauensis</emphasis>
<bibRefCitation id="BE444B3ABB03FFD201241FEF62F46CDB" author="Chvala M." box="[812,946,929,951]" pageId="5" pageNumber="226" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="1DD54D48BB03FFD2062C1FEF65E96CDB" authorityName="Chvala" authorityYear="1999" box="[1060,1199,929,950]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="alatauensis">
<emphasis id="E8A1EAD9BB03FFD2062C1FEF65756CDB" box="[1060,1075,929,950]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8A1EAD9BB03FFD206491FEF65E96CDB" box="[1089,1199,929,950]" italics="true" pageId="5" pageNumber="226">alatauensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="1DD54D48BB03FFD2073C1FEF62C16CBD" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="fedtschenkoi" status="sp.n.">
<emphasis id="E8A1EAD9BB03FFD2073C1FEF64056CDB" box="[1332,1347,929,950]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8A1EAD9BB03FFD207591FEF62C16CBD" italics="true" pageId="5" pageNumber="226">fedtschenkoi</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB03FFD201861FF562F96CBD" bold="true" box="[910,959,955,976]" pageId="5" pageNumber="226">
<taxonomicNameLabel id="F39257A2BB03FFD201861FF562F96CBD" box="[910,959,955,976]" pageId="5" pageNumber="226" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
primarily by distinct postpronotal and presutural supra-alar setae.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD2015A1FBF653E6B56" pageId="5" pageNumber="226" type="etymology">
<paragraph id="DA6A36CBBB03FFD2015A1FBF653E6B56" blockId="5.[812,1399,315,1111]" pageId="5" pageNumber="226">ETYMOLOGY. The specific name is a patronym in honour of the Russian naturalist and explorer Aleksey Pavlovich Fedtschenko (1844–1873).</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB03FFD2015A180F65AA6B3A" box="[850,1260,1089,1111]" pageId="5" pageNumber="226" type="distribution">
<paragraph id="DA6A36CBBB03FFD2015A180F65AA6B3A" blockId="5.[812,1399,315,1111]" box="[850,1260,1089,1111]" pageId="5" pageNumber="226">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB03FFD20601180F652A6B3A" box="[1033,1132,1089,1111]" name="Tajikistan" pageId="5" pageNumber="226">Tajikistan</collectingCountry>
,
<collectingCountry id="A2C2765BBB03FFD20670180F65AE6B3A" box="[1144,1256,1089,1111]" name="Uzbekistan" pageId="5" pageNumber="226">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="A8D73A3DF75FB8FAE1FEF3C27D14DC87" type="DOI">10.15298/rusentj.32.2.13</mods:identifier>
<mods:identifier id="F7A71909884D99C1EECEB8991A543E66" type="ISSN">0132-8069</mods:identifier>
</mods:mods>
<treatment id="527C87DDBB07FFD402CE184663446B9A" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A" httpUri="http://treatment.plazi.org/id/527C87DDBB07FFD402CE184663446B9A" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222">
<subSubSection id="92CF6540BB07FFD602CE184663BB6B72" box="[198,765,1031,1055]" pageId="1" pageNumber="222" type="nomenclature">
<paragraph id="DA6A36CBBB07FFD602CE184663BB6B72" blockId="1.[189,776,1031,2063]" box="[198,765,1031,1055]" pageId="1" pageNumber="222">
<heading id="812281A7BB07FFD602CE184663BB6B72" box="[198,765,1031,1055]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="222" reason="8">
<taxonomicName id="1DD54D48BB07FFD602CE184663FA6B72" ID-CoL="C2Q7T" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[198,700,1031,1055]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="222" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB07FFD602CE1846604F6B72" box="[198,265,1032,1055]" italics="true" pageId="1" pageNumber="222">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB07FFD60311184960E96B72" box="[281,431,1031,1055]" italics="true" pageId="1" pageNumber="222">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB07FFD603C8184963026B72" box="[448,580,1031,1055]" italics="true" pageId="1" pageNumber="222">fandariensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB07FFD600C0184963BB6B72" bold="true" box="[712,765,1031,1055]" pageId="1" pageNumber="222">
<taxonomicNameLabel id="F39257A2BB07FFD600C0184963BB6B72" box="[712,765,1031,1055]" pageId="1" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB07FFD603A7186A63526B51" box="[431,532,1060,1084]" pageId="1" pageNumber="222" type="description">
<paragraph id="DA6A36CBBB07FFD603A7186A63526B51" blockId="1.[189,776,1031,2063]" box="[431,532,1060,1084]" pageId="1" pageNumber="222">
<figureCitation id="42EE2A4EBB07FFD603A7186A63566B51" box="[431,528,1060,1084]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="1" pageNumber="222">Figs 1–5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB07FFD602EA180763036BB9" pageId="1" pageNumber="222" type="materials_examined">
<paragraph id="DA6A36CBBB07FFD602EA180763D86BC9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
<materialsCitation id="6ABD3C96BB07FFD602EA180763D86BC9" collectingDate="1869-06-19" collectionCode="MATERIAL, ZMMU" collectorName="A. Fedtschenko &amp; Cyrillic &amp; Russian &amp; Fan" country="Tajikistan" location="Fan Darya River" pageId="1" pageNumber="222" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="BCC4AE0EBB07FFD602EA180760176B31" box="[226,337,1097,1116]" pageId="1" pageNumber="222">MATERIAL</collectionCode>
EXAMINED.
<typeStatus id="056E8869BB07FFD603D21807636B6B31" box="[474,557,1097,1116]" pageId="1" pageNumber="222" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CCD3FD42BB07FFD600311809630B6B30" box="[569,589,1095,1117]" count="1" pageId="1" pageNumber="222" type="male">♂</specimenCount>
, labels as in
<figureCitation id="42EE2A4EBB07FFD600C3180762446B31" box="[715,770,1097,1116]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="1" pageNumber="222">Fig. 4</figureCitation>
, [
<emphasis id="E8A1EAD9BB07FFD602CC182F60656B19" bold="true" box="[196,291,1121,1140]" pageId="1" pageNumber="222">
<collectingCountry id="A2C2765BBB07FFD602CC182F60656B19" box="[196,291,1121,1140]" name="Tajikistan" pageId="1" pageNumber="222">Tajikistan</collectingCountry>
</emphasis>
] small bluish green label only with number 19. [=
<date id="AE6B100BBB07FFD602B51837605D6BE1" box="[189,283,1145,1164]" pageId="1" pageNumber="222" value="1869-06-19">
<collectingDate id="BE2FE9E3BB07FFD602B51837605D6BE1" box="[189,283,1145,1164]" pageId="1" pageNumber="222" value="1869-06-19">19.vi.1869</collectingDate>
</date>
] [
<collectorName id="7720531DBB07FFD60326183760FA6BE1" box="[302,444,1145,1164]" pageId="1" pageNumber="222">A. Fedtschenko</collectorName>
] // [printed in
<collectorName id="7720531DBB07FFD6004A183763C06BE1" box="[578,646,1145,1164]" pageId="1" pageNumber="222">Cyrillic</collectorName>
,
<collectorName id="7720531DBB07FFD6009A1837639F6BE1" box="[658,729,1145,1164]" pageId="1" pageNumber="222">Russian</collectorName>
, pre- 1918 orthography]
<collectorName id="7720531DBB07FFD6036E18DC60C06BC8" box="[358,390,1170,1189]" pageId="1" pageNumber="222">Fan</collectorName>
[=
<location id="DF0A6010BB07FFD603AE18DC637D6BC9" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A:DF0A6010BB07FFD603AE18DC637D6BC9" box="[422,571,1169,1189]" country="Tajikistan" name="Fan Darya River" pageId="1" pageNumber="222">Fan Darya River</location>
] (
<collectionCode id="BCC4AE0EBB07FFD6004418DF63D26BC9" box="[588,660,1169,1188]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="222" type="Museum">ZMMU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DA6A36CBBB07FFD602EA18E463036BB9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
<materialsCitation id="6ABD3C96BB07FFD602EA18E463076BB9" collectingDate="1869-06-19" collectionCode="ZMMU, ZISP" collectorName="A. Fedtschenko &amp; Cyrillic &amp; Russian &amp; Fan" country="Tajikistan" location="Fan Darya River" pageId="1" pageNumber="222" specimenCount="7" specimenCount-female="3" specimenCount-male="4" typeStatus="paratype">
<typeStatus id="056E8869BB07FFD602EA18E4607F6BD0" box="[226,313,1194,1213]" pageId="1" pageNumber="222" type="paratype">Paratypes</typeStatus>
.
<emphasis id="E8A1EAD9BB07FFD6034D18E760ED6BD1" bold="true" box="[325,427,1193,1212]" pageId="1" pageNumber="222">
<collectingCountry id="A2C2765BBB07FFD6034D18E760E16BD1" box="[325,423,1193,1212]" name="Tajikistan" pageId="1" pageNumber="222">Tajikistan</collectingCountry>
:
</emphasis>
same data as holotype (
<specimenCount id="CCD3FD42BB07FFD6008318E763816BD0" box="[651,711,1191,1213]" count="2" pageId="1" pageNumber="222" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB07FFD600DE18E762476BD0" box="[726,769,1191,1213]" count="2" pageId="1" pageNumber="222" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB07FFD602B5188C60446BB8" box="[189,258,1218,1237]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="222" type="Museum">ZMMU</collectionCode>
;
<specimenCount id="CCD3FD42BB07FFD60304188F60036BB8" box="[268,325,1215,1237]" count="2" pageId="1" pageNumber="222" type="male">2 ♂♂</specimenCount>
(both dissected),
<specimenCount id="CCD3FD42BB07FFD603EA188F60B96BB8" box="[482,511,1215,1237]" count="1" pageId="1" pageNumber="222" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB07FFD60003188F637D6BB9" box="[523,571,1217,1236]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="222">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB07FFD602EA189760C76AA9" pageId="1" pageNumber="222" type="diagnosis">
<paragraph id="DA6A36CBBB07FFD602EA189760C76AA9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
DIAGNOSIS. Mid-sized species (body about
<quantity id="1D2D9B2EBB07FFD600B2189763BA6B83" box="[698,764,1241,1263]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="222" unit="mm" value="5.0">5 mm</quantity>
); male eyes holoptic; proboscis with labrum nearly 2X longer than eye height; palpus mostly brownish in male, yellow in female; mesoscutum with 3 brownish vittae along rows of acrostichal and dorsocentral setae; acrostichal setae irregularly 2–3-serial, presutural dorsocentrals irregularly 3–4- serial, laterotergite with pale setae; abdominal tergites 2–6 extensively shiny (lateral view); female mid and hind femora with flattened setae.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB07FFD402EA198763F86BCA" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222" type="description">
<paragraph id="DA6A36CBBB07FFD602EA198760986802" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
DESCRIPTION. Body length
<quantity id="1D2D9B2EBB07FFD60000198763386AB2" box="[520,638,1481,1503]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.1" metricValueMin="4.8" pageId="1" pageNumber="222" unit="mm" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8–5.1 mm</quantity>
, wing
<quantity id="1D2D9B2EBB07FFD600B6198761A36A94" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="1" pageNumber="222" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
.
<emphasis id="E8A1EAD9BB07FFD602FC19AA606E6A94" bold="true" box="[244,296,1508,1529]" pageId="1" pageNumber="222">Male</emphasis>
(
<figureCitation id="42EE2A4EBB07FFD6033319AA603C6A94" box="[315,378,1508,1530]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="1" pageNumber="222">Fig. 1</figureCitation>
). Head capsule regions black, mostly densely greyish pruinose, face along lower margin and clypeus shiny. Eyes holoptic, upper ommatidia enlarged. Frons represented by small subtriangular space just below ocellar triangle and larger space above antennae, bare. Face broad, bare. Ocellar setae black, long, fine. Occiput covered with numerous, long, fine, black setae (including postoculars); postgena with pale, long, hair-like setae. Antenna with scape and pedicel brownish, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with short setulae; postpedicel nearly 3X as long as basal width; stylus nearly as long as postpedicel basal width. Proboscis with labrum brownish, paler apically; nearly 2X longer than eye height; palpus short, mostly brown, rather yellowish brown at tip; bearing long, black setae.
</paragraph>
<paragraph id="DA6A36CBBB07FFD602EA1B3A641B6DC1" blockId="1.[189,776,1031,2063]" lastBlockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">Thorax black in ground-colour, mostly densely greyish pruinose; mesoscutum with 3 brownish, indistinct vittae along rows of acrostichal and dorsocentral setae. Prosternum bare. Proepisternum with tuft of numerous pale, fine setae on lower part, bare on upper part. Antepronotum with 4–5 black, short setae dorsally and numerous pale, long, fine setae laterally. Postpronotal lobe with 1 long, strong, black and numerous short, fine, black and pale setae. Mesonotal setae black (except noted): acrostichals arranged in 2–3 irregular rows (often entirely biserial), rather long, fine, absent on prescutellar depression; presutural dorsocentral setae arranged in 3–4 irregular rows, similar to acrostichals; postsutural dorsocentrals sparser,becoming uniserial,longer and stronger towards scutellum, 1 pair of prescutellars longest; 1 rather fine presutural supra-alar; 3 notopleurals, 2 postsutural supra-alars (anterior seta shorter and finer), 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair slightly shorter); in addition, supra-alar face with numerous black and pale setulae, notopleuron with similar pale setulae anteriorly. Laterotergite with numerous pale setae. Anterior and posterior spiracles pale.</paragraph>
<paragraph id="DA6A36CBBB07FFD6015A1EFF62CC6901" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">Legs with coxae densely greyish pruinose, remaining podomeres subshiny. Legs colour: almost entirely brown, usually only hind femur at apex and hind tibia near base narrowly brownish yellow (sometimes hind femur entirely brownish). Legs mostly black setose. Fore coxa covered with pale, dense, hair-like setae (2–3 stronger, black, subapical setae); hind trochanter with numerous pale, short, fine setae, 4–5 brownish to black, long, strong, subapical setae ventrally and 1–2 similar setae dorsally. Fore and mid femora whitish pubescent ventrally; hind femur with indistinct ventral pubescence only closer to apex. Fore femur with rows of fine anteroventral and posteroventral setae, which are pale on about basal half, short closer to apex and becoming longer towards base (longest setae nearly as long as femur width); dense, long, fine setae posteriorly (pale closer to base). Fore tibia with 4–6 short anterodorsal setae and subequal to them in length, fine setae posterodorsally (2–3 setae on apical half usually stronger). Fore basitarsus slender; with 2 short anterodorsal setae; anteroventral and posteroventral spine-like setae. Mid femur with rows of subequally long anteroventral and posteroventral setae; additional fine setae near extreme base and spinule-like ventral setulae on basal half; 1 anterior subapical seta. Mid tibia with 4–5 anterodorsal and 4–5 posterodorsal long setae; stronger setulae closer to apex ventrally. Mid tarsus with anteroventral and posteroventral, spine-like setae. Hind femur only slightly broader than mid femur; with 2 anterodorsal setae near apex, 1 anterior subapical seta; complete row of moderately long strong anteroventral setae (longest setae at most half as long as femur width); row of short posteroventral setae intermixed with additional fine setae; dense ventral spinule-like setulae. Hind tibia slightly, gently thickened towards apex; bearing rows of 4–5 moderately long anterodorsal and posterodorsal setae; simple setulae ventrally; no seta in posteroapical comb. Hind tarsomeres rather slightly thickened; basitarsus with 2–3 short anterodorsal and posterodorsal setae; spine-like setae ventrally.</paragraph>
<paragraph id="DA6A36CBBB07FFD6015A1A3F65FE681A" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">
Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins mostly brownish, well-sclerotized (except noted). Veins R
<subScript id="4651348EBB07FFD606491AFB650E69AC" attach="left" box="[1089,1096,1717,1729]" fontSize="5" pageId="1" pageNumber="222">5</subScript>
and M
<subScript id="4651348EBB07FFD606821AFB65D769AC" attach="left" box="[1162,1169,1717,1729]" fontSize="5" pageId="1" pageNumber="222">1</subScript>
divergent towards wing margin; radial fork acute; R
<subScript id="4651348EBB07FFD6063D1A9E657A69B1" attach="left" box="[1077,1084,1744,1756]" fontSize="5" pageId="1" pageNumber="222">5</subScript>
meeting costa before wing apex; CuA+CuP (anal vein) complete but weakened beyond middle. Cell dm rather short, with elongate apex; apical portion of M
<subScript id="4651348EBB07FFD601531B6E62246841" attach="left" box="[859,866,1824,1836]" fontSize="5" pageId="1" pageNumber="222">4</subScript>
nearly 3.5X longer than its middle portion. Pterostigma absent. Basal costal seta present, short. Anal lobe well-developed; axillary incision acute but closer to 90°. Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<paragraph id="DA6A36CBBB07FFD5015A1B3560B96DFC" blockId="1.[812,1399,315,2071]" lastBlockId="2.[189,775,315,791]" lastPageId="2" lastPageNumber="223" pageId="1" pageNumber="222">Abdomen entirely dark-brown. Tergite 1 entirely greyish pruinose; tergites 2–6 mostly shiny, narrowly brown pruinose antero-dorsally (dorsal view); tergite 7 with brownish, slightly iridescent pruinosity dorsally and greyish pruinose laterally; tergite 8 invisible; tergite 1 with pale setae, tergites 2–6 with black setulae dorsally and numerous long pale setae laterally, slightly stronger lateral posteromarginal setae pale yellow; tergite 7 with scattered pale setulae laterally. Sternites 1–7 densely greyish pruinose, with pale setae, sternite 1 bare; sternite 8 contrastingly subshiny, mostly with black setae. Pregenital segments: segment 6 unmodified; tergite 7 with straight posterior margin, slightly humped mid-dorsally (lateral view); sternite 7 simple; pleuron 7 unmodified; segment 8 with separated tergite and sternite; tergite 8 represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, unmodified anteriorly, with 2 digitiform, short, lateral projections on each side closer to upper margin anteriorly (posterior projection slightly longer).</paragraph>
<paragraph id="DA6A36CBBB04FFD502EA1ED864316DAA" blockId="2.[189,775,315,791]" lastBlockId="2.[812,1399,315,791]" pageId="2" pageNumber="223">
Hypopygium (
<figureCitation id="42EE2A4EBB04FFD503661ED860E06DC6" box="[366,422,662,684]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="2" pageNumber="223">Fig. 2</figureCitation>
) moderately large; epandrium black; cerci pale brown to yellowish brown; hypandrium invisible; phallus brownish yellow. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella subtriangular (lateral view), with broadly rounded apex; bearing dark fine setae more numerous along lower margin and dense at apex. Hypandrium separated from epandrium; mostly membranous, narrowly sclerotized along margin, with truncate apex, narrowly separated on apex; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus egg-shaped, with posterior margin extending beyond apex of epandrial lamella, undivided, with smoothed inner margin; covered with dark setulae. Phallus (
<figureCitation id="42EE2A4EBB04FFD506E01DB864636D66" box="[1256,1317,502,524]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="2" pageNumber="223">Fig. 3</figureCitation>
) almost entirely hidden, only medial curvature and, often, short apical portion visible; zigzag bent; its basal portion almost horizontal, mostly thick, with mid-dorsal tubercle; apical portion long, almost vertical and straight, slightly curved on apex (with its tip pointing almost to rear), becoming gently slenderer towards apex. Ejaculatory apodeme very large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="DA6A36CBBB04FFD4015A1E8563406C5C" blockId="2.[812,1399,315,791]" lastBlockId="3.[189,776,315,1271]" lastPageId="3" lastPageNumber="224" pageId="2" pageNumber="223">
<emphasis id="E8A1EAD9BB04FFD5015A1E8562DB6D8D" bold="true" box="[850,925,715,736]" pageId="2" pageNumber="223">Female</emphasis>
(
<figureCitation id="42EE2A4EBB04FFD501B91E8562B56D8C" box="[945,1011,715,737]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="2" pageNumber="223">Fig. 5</figureCitation>
). Similar to male except as follows. Dichoptic, ommatidia equally small.Frons very broad, broader than distance between outer margins of posterior ocelli; with marginal setulae. Occiput with sparser and stronger setae. Palpus yellow, with scattered short setae. Mesoscutum with shorter acrostichal and dorsocentral setae; presutural supra-alar space only with black fine setae. Legs colour: mid and hind femora narrowly yellow at apex; mid and hind tibiae rather brownish yellow (the former generally slightly darker). Hind trochanter only with pale fine setae ventrally. Fore leg with simple setae (similar pattern but somewhat shorter than in male). Mid femur with short, fine anteroventral setae longer closer to base; moderately long, flattened setae on about apical 2/3 posteroventrally. Mid tibia with 2–3 anteroventral and 1–2 posteroventral setae. Hind femur with fridge of short, dense, slightly flattened setae posteroventrally (except extreme base). Hind tibia with 1–2 anteroventral setae. Abdomen with shorter setae, only pale setose; tergites mostly shiny, narrowly greyish pruinose along lateral margin (in addition, tergite 2 narrowly pollinose anteriorly; tergite 1 entirely pruinose); sternite 8 subshiny. Cercus very short, covered with dense, erect setulae.
</paragraph>
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<paragraph id="DA6A36CBBB04FFD502EA1B30625468AC" blockId="2.[189,1398,1917,2061]" pageId="2" pageNumber="223">
Figs 1–5.
<taxonomicName id="1DD54D48BB04FFD503351B30639068FC" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[317,726,1918,1937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB04FFD503351B30603468FD" box="[317,370,1918,1936]" italics="true" pageId="2" pageNumber="223">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB04FFD5038A1B3060BD68FC" box="[386,507,1918,1937]" italics="true" pageId="2" pageNumber="223">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB04FFD500021B30633268FC" box="[522,628,1918,1937]" italics="true" pageId="2" pageNumber="223">fandariensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB04FFD500EA1B30624B68FC" bold="true" box="[738,781,1918,1937]" pageId="2" pageNumber="223">
<taxonomicNameLabel id="F39257A2BB04FFD500EA1B30624B68FC" box="[738,781,1918,1937]" pageId="2" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm.
</paragraph>
<paragraph id="DA6A36CBBB04FFD502EA1B8463AC6760" blockId="2.[189,1398,1917,2061]" pageId="2" pageNumber="223">
Рис. 1–5.
<taxonomicName id="1DD54D48BB04FFD503351B84639368B1" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[317,725,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB04FFD503351B84603468B1" box="[317,370,1994,2012]" italics="true" pageId="2" pageNumber="223">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB04FFD503891B8760BC68B1" box="[385,506,1993,2012]" italics="true" pageId="2" pageNumber="223">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB04FFD500011B87633568B1" box="[521,627,1993,2012]" italics="true" pageId="2" pageNumber="223">fandariensis</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB04FFD500D71B87624C68B1" bold="true" box="[735,778,1993,2012]" pageId="2" pageNumber="223">
<taxonomicNameLabel id="F39257A2BB04FFD500D71B87624C68B1" box="[735,778,1993,2012]" pageId="2" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм.
</paragraph>
</caption>
<paragraph id="DA6A36CBBB05FFD402EA1F7863F86BCA" blockId="3.[189,776,315,1271]" pageId="3" pageNumber="224">
DIFFERENTIAL DIAGNOSIS. The new species is very similar to
<taxonomicName id="1DD54D48BB05FFD403141F1F631E6C0B" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[284,600,849,871]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
<emphasis id="E8A1EAD9BB05FFD403141F1F60976C0B" box="[284,465,849,870]" italics="true" pageId="3" pageNumber="224">Empis tajikistanica</emphasis>
<bibRefCitation id="BE444B3ABB05FFD403DF1F1F631E6C0B" author="Chvala M." box="[471,600,849,871]" pageId="3" pageNumber="224" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="1DD54D48BB05FFD4006A1F1F61906CEC" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov.">
<emphasis id="E8A1EAD9BB05FFD4006A1F1F61906CEC" italics="true" pageId="3" pageNumber="224">Empis fandariensis</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB05FFD402E91F2560566CED" bold="true" box="[225,272,875,896]" pageId="3" pageNumber="224">
<taxonomicNameLabel id="F39257A2BB05FFD402E91F2560566CED" box="[225,272,875,896]" pageId="3" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="1DD54D48BB05FFD403951F2263766CED" authorityName="Chvala" authorityYear="1999" box="[413,560,875,897]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
<emphasis id="E8A1EAD9BB05FFD403951F2260EA6CEC" box="[413,428,876,897]" italics="true" pageId="3" pageNumber="224">E</emphasis>
.
<emphasis id="E8A1EAD9BB05FFD403B41F2563766CED" box="[444,560,875,896]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
</taxonomicName>
by shorter proboscis (labrum nearly 2X longer than head height) and extensively shiny abdominal tergites 2–6 (lateral view). In addition, the male of the new species has sparser and finer subapical, ventral setae on hind trochanter and rounded cerci. In
<taxonomicName id="1DD54D48BB05FFD4007F1F9962406C86" authorityName="Chvala" authorityYear="1999" box="[631,774,982,1004]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
<emphasis id="E8A1EAD9BB05FFD4007F1F9963C36C81" box="[631,645,983,1004]" italics="true" pageId="3" pageNumber="224">E</emphasis>
.
<emphasis id="E8A1EAD9BB05FFD4009A1F9862406C86" box="[658,774,982,1003]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
</taxonomicName>
the labrum of proboscis is nearly 3X longer than head height and abdominal tergites 2–6 extensively greyish pruinose (lateral view). The male of
<taxonomicName id="1DD54D48BB05FFD403AE1869637E6B56" authorityName="Chvala" authorityYear="1999" box="[422,568,1062,1084]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
<emphasis id="E8A1EAD9BB05FFD403AE186960F36B51" box="[422,437,1063,1084]" italics="true" pageId="3" pageNumber="224">E</emphasis>
.
<emphasis id="E8A1EAD9BB05FFD403CC1868637E6B56" box="[452,568,1062,1083]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
</taxonomicName>
has a tuft of strong, black, ventral setae on the hind trochanter and somewhat pointed cerci [
<bibRefCitation id="BE444B3ABB05FFD4034F181560B96B1D" author="Chvala M." box="[327,511,1115,1137]" pageId="3" pageNumber="224" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999: 210</bibRefCitation>
, fig. 28]. The female of
<taxonomicName id="1DD54D48BB05FFD400FA181260746BE6" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="fandariensis" status="sp.n.">
<emphasis id="E8A1EAD9BB05FFD400FA181262466B1C" box="[754,768,1116,1137]" italics="true" pageId="3" pageNumber="224">E</emphasis>
.
<emphasis id="E8A1EAD9BB05FFD402B5183860746BE6" box="[189,306,1142,1163]" italics="true" pageId="3" pageNumber="224">fandariensis</emphasis>
</taxonomicName>
<emphasis id="E8A1EAD9BB05FFD403311838602F6BE6" bold="true" box="[313,361,1142,1163]" pageId="3" pageNumber="224">
<taxonomicNameLabel id="F39257A2BB05FFD403311838602F6BE6" box="[313,361,1142,1163]" pageId="3" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from the female of
<taxonomicName id="1DD54D48BB05FFD4007D183962406BE6" authorityName="Chvala" authorityYear="1999" box="[629,774,1142,1164]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
<emphasis id="E8A1EAD9BB05FFD4007D183963C26BE1" box="[629,644,1143,1164]" italics="true" pageId="3" pageNumber="224">E</emphasis>
.
<emphasis id="E8A1EAD9BB05FFD4009A183862406BE6" box="[658,774,1142,1163]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
</taxonomicName>
primarily by somewhat pennate mid and hind femora.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD402EA18E562466BB1" pageId="3" pageNumber="224" type="etymology">
<paragraph id="DA6A36CBBB05FFD402EA18E562466BB1" blockId="3.[189,776,315,1271]" pageId="3" pageNumber="224">
<materialsCitation id="6ABD3C96BB05FFD402EA18E563BB6BB1" country="Tajikistan" location="Fan Darya River" pageId="3" pageNumber="224" specimenCount="1">
ETYMOLOGY. The epithet refers to the
<typeStatus id="056E8869BB05FFD4007C18E263D86BAC" box="[628,670,1196,1217]" pageId="3" pageNumber="224">type</typeStatus>
locality of the new species, an area near
<location id="DF0A6010BB05FFD403D7188863C46BB1" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A:DF0A6010BB05FFD403D7188863C46BB1" box="[479,642,1222,1244]" country="Tajikistan" name="Fan Darya River" pageId="3" pageNumber="224">Fan Darya River</location>
(
<collectingCountry id="A2C2765BBB05FFD40087188863B06BB1" box="[655,758,1222,1244]" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB05FFD402EA18AF63446B9A" box="[226,514,1249,1271]" pageId="3" pageNumber="224" type="distribution">
<paragraph id="DA6A36CBBB05FFD402EA18AF63446B9A" blockId="3.[189,776,315,1271]" box="[226,514,1249,1271]" pageId="3" pageNumber="224">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB05FFD4039218AF60B86B9A" box="[410,510,1249,1271]" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="DA6A36CBBB0AFFDB02EF1EB8639D6C60" blockId="12.[189,776,757,1037]" box="[231,731,757,781]" pageId="12" pageNumber="233">
<heading id="812281A7BB0AFFDB02EF1EB8639D6C60" box="[231,731,757,781]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="233" reason="8">
<taxonomicName id="1DD54D48BB0AFFDB02EF1EB8639D6C60" ID-CoL="8BZQP" authority="Shamshev, 2003" authorityName="Shamshev" authorityYear="2003" box="[231,731,757,781]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="233" phylum="Arthropoda" rank="species" species="zimini" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0AFFDB02EF1EB8606C6C60" box="[231,298,758,781]" italics="true" pageId="12" pageNumber="233">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0AFFDB03331EBB60976C60" box="[315,465,757,781]" italics="true" pageId="12" pageNumber="233">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0AFFDB03E91EB863646C60" box="[481,546,758,781]" italics="true" pageId="12" pageNumber="233">zimini</emphasis>
<bibRefCitation id="BE444B3ABB0AFFDB00221EBB639D6C60" author="Shamshev I. V." box="[554,731,757,781]" pageId="12" pageNumber="233" pagination="19 - 27" refId="ref12875" refString="Shamshev I. V. 2003. Descriptions of three new species of the Empis subgenus Polyblepharis (Diptera: Empididae) from the Ukraine and Middle Asia // An international Journal of dipterological Research. Vol. 14. P. 19 - 27." type="book chapter" year="2003">Shamshev, 2003</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DA6A36CBBB0AFFDB02EA1F5463846C28" blockId="12.[189,776,757,1037]" pageId="12" pageNumber="233">
<materialsCitation id="6ABD3C96BB0AFFDB02EA1F5463F86C28" collectorName="NW Bukhara" country="Uzbekistan" location="Uzbekistan" pageId="12" pageNumber="233" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="BE444B3ABB0AFFDB02EA1F5460DA6C40" author="Shamshev I. V." box="[226,412,794,813]" pageId="12" pageNumber="233" pagination="19 - 27" refId="ref12875" refString="Shamshev I. V. 2003. Descriptions of three new species of the Empis subgenus Polyblepharis (Diptera: Empididae) from the Ukraine and Middle Asia // An international Journal of dipterological Research. Vol. 14. P. 19 - 27." type="book chapter" year="2003">Shamshev, 2003: 24</bibRefCitation>
(
<specimenCount id="CCD3FD42BB0AFFDB03A71F5960856C40" box="[431,451,791,813]" count="1" pageId="12" pageNumber="233" type="male">♂</specimenCount>
and
<specimenCount id="CCD3FD42BB0AFFDB03F01F5963426C40" box="[504,516,791,813]" count="1" pageId="12" pageNumber="233" type="female">♀</specimenCount>
), figs. 9–12.
<typeStatus id="056E8869BB0AFFDB00801F5463F26C40" box="[648,692,794,813]" pageId="12" pageNumber="233">Type</typeStatus>
locality:
<collectingCountry id="A2C2765BBB0AFFDB02B51F7C60666C28" box="[189,288,818,837]" name="Uzbekistan" pageId="12" pageNumber="233">Uzbekistan</collectingCountry>
,
<collectorName id="7720531DBB0AFFDB03221F7C60E66C28" box="[298,416,818,837]" pageId="12" pageNumber="233">NW Bukhara</collectorName>
, Yargak, tributary of Chatyrchy
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0AFFDB02EA1F04636B6CD0" pageId="12" pageNumber="233" type="materials_examined">
<paragraph id="DA6A36CBBB0AFFDB02EA1F04636B6CD0" blockId="12.[189,776,757,1037]" pageId="12" pageNumber="233">
<materialsCitation id="6ABD3C96BB0AFFDB02EA1F0463596C18" collectingDate="1869-04-03" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Zeravshan Valley" pageId="12" pageNumber="233" specimenCount="5" specimenCount-female="4" specimenCount-male="1">
<collectionCode id="BCC4AE0EBB0AFFDB02EA1F0460146C30" box="[226,338,842,861]" pageId="12" pageNumber="233">MATERIAL</collectionCode>
EXAMINED.
<emphasis id="E8A1EAD9BB0AFFDB03EB1F0463126C30" bold="true" box="[483,596,842,861]" pageId="12" pageNumber="233">
<collectingCountry id="A2C2765BBB0AFFDB03EB1F0463166C30" box="[483,592,842,861]" name="Uzbekistan" pageId="12" pageNumber="233">Uzbekistan</collectingCountry>
:
</emphasis>
<location id="DF0A6010BB0AFFDB00571F0462456C30" LSID="urn:lsid:plazi:treatment:527C87DDBB0AFFDB02EF1EB863486B60:DF0A6010BB0AFFDB00571F0462456C30" box="[607,771,842,861]" country="Uzbekistan" name="Zeravshan Valley" pageId="12" pageNumber="233">Zeravshan Valley</location>
,
<date id="AE6B100BBB0AFFDB02B51F2C60486C18" box="[189,270,866,885]" pageId="12" pageNumber="233" value="1869-04-03">
<collectingDate id="BE2FE9E3BB0AFFDB02B51F2C60486C18" box="[189,270,866,885]" pageId="12" pageNumber="233" value="1869-04-03">3.iv.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB0AFFDB03131F2C60EE6C18" box="[283,424,866,885]" pageId="12" pageNumber="233">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB0AFFDB03B41F2C60A66C18" box="[444,480,863,885]" count="1" pageId="12" pageNumber="233" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0AFFDB03E41F2C63506C18" box="[492,534,863,885]" count="4" pageId="12" pageNumber="233" type="female">4 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0AFFDB00221F2C60C26CE0" collectingDate="1869-04-25" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Zeravshan Valley" pageId="12" pageNumber="233" specimenCount="1" specimenCount-female="1">
same locality,
<date id="AE6B100BBB0AFFDB00A21F2C62406C18" box="[682,774,866,885]" pageId="12" pageNumber="233" value="1869-04-25">
<collectingDate id="BE2FE9E3BB0AFFDB00A21F2C62406C18" box="[682,774,866,885]" pageId="12" pageNumber="233" value="1869-04-25">25.iv.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB0AFFDB02CA1F3460086CE0" box="[194,334,890,909]" pageId="12" pageNumber="233">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB0AFFDB03681F34603D6CE0" box="[352,379,887,909]" count="1" pageId="12" pageNumber="233" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0AFFDB03851F34606D6CC8" collectingDate="1869-04-11" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Samarkand" pageId="12" pageNumber="233" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<location id="DF0A6010BB0AFFDB03851F3460B66CE0" LSID="urn:lsid:plazi:treatment:527C87DDBB0AFFDB02EF1EB863486B60:DF0A6010BB0AFFDB03851F3460B66CE0" box="[397,496,890,909]" country="Uzbekistan" name="Samarkand" pageId="12" pageNumber="233">Samarkand</location>
,
<date id="AE6B100BBB0AFFDB03F11F3463136CE0" box="[505,597,890,909]" pageId="12" pageNumber="233" value="1869-04-11">
<collectingDate id="BE2FE9E3BB0AFFDB03F11F3463136CE0" box="[505,597,890,909]" pageId="12" pageNumber="233" value="1869-04-11">11.iv.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB0AFFDB00681F3463AC6CE0" box="[608,746,890,909]" pageId="12" pageNumber="233">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB0AFFDB00F41F3461A36CC8" count="3" pageId="12" pageNumber="233" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0AFFDB02FD1FDC60646CC8" box="[245,290,911,933]" count="2" pageId="12" pageNumber="233" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0AFFDB03321FDC62446CC8" box="[314,770,911,933]" collectingDate="1869-04-17" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Samarkand" pageId="12" pageNumber="233" specimenCount="1" specimenCount-female="1">
same locality,
<date id="AE6B100BBB0AFFDB03CB1FDC63596CC8" box="[451,543,914,933]" pageId="12" pageNumber="233" value="1869-04-17">
<collectingDate id="BE2FE9E3BB0AFFDB03CB1FDC63596CC8" box="[451,543,914,933]" pageId="12" pageNumber="233" value="1869-04-17">17.iv.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB0AFFDB00381FDC63876CC8" box="[560,705,914,933]" pageId="12" pageNumber="233">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB0AFFDB00D11FDC63BF6CC8" box="[729,761,911,933]" count="1" pageId="12" pageNumber="233" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0AFFDB02B51FE4636F6CD0" box="[189,553,935,957]" collectingDate="1871-04-22" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Keles" pageId="12" pageNumber="233" specimenCount="1" specimenCount-female="1">
<location id="DF0A6010BB0AFFDB02B51FE461B66CD0" LSID="urn:lsid:plazi:treatment:527C87DDBB0AFFDB02EF1EB863486B60:DF0A6010BB0AFFDB02B51FE461B66CD0" box="[189,240,938,957]" country="Uzbekistan" name="Keles" pageId="12" pageNumber="233">Keles</location>
,
<date id="AE6B100BBB0AFFDB02F21FE460116CD0" box="[250,343,938,957]" pageId="12" pageNumber="233" value="1871-04-22">
<collectingDate id="BE2FE9E3BB0AFFDB02F21FE460116CD0" box="[250,343,938,957]" pageId="12" pageNumber="233" value="1871-04-22">22.iv.1871</collectingDate>
</date>
[
<collectorName id="7720531DBB0AFFDB036B1FE460B66CD0" box="[355,496,938,957]" pageId="12" pageNumber="233">A. Fedtschenko</collectorName>
] (
<specimenCount id="CCD3FD42BB0AFFDB000C1FE463666CD0" box="[516,544,935,957]" count="1" pageId="12" pageNumber="233" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0AFFDB02EA1F8C606D6C9E" pageId="12" pageNumber="233" type="discussion">
<paragraph id="DA6A36CBBB0AFFDB02EA1F8C606D6C9E" blockId="12.[189,776,757,1037]" pageId="12" pageNumber="233">
REMARKS. The species has been known only from the
<typeStatus id="056E8869BB0AFFDB02B51F9361A16C9F" box="[189,231,989,1010]" pageId="12" pageNumber="233">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0AFFDB02EA1FB963486B60" box="[226,526,1015,1037]" pageId="12" pageNumber="233" type="distribution">
<paragraph id="DA6A36CBBB0AFFDB02EA1FB963486B60" blockId="12.[189,776,757,1037]" box="[226,526,1015,1037]" pageId="12" pageNumber="233">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB0AFFDB03921FB9634C6B60" box="[410,522,1015,1037]" name="Uzbekistan" pageId="12" pageNumber="233">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

889
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@ -0,0 +1,889 @@
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<treatment id="527C87DDBB0FFFDB013E1924624E6D8C" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C" httpUri="http://treatment.plazi.org/id/527C87DDBB0FFFDB013E1924624E6D8C" lastPageId="12" lastPageNumber="233" pageId="9" pageNumber="230">
<subSubSection id="92CF6540BB0FFFDE013E1924642B6AEC" box="[822,1389,1385,1409]" pageId="9" pageNumber="230" type="nomenclature">
<paragraph id="DA6A36CBBB0FFFDE013E1924642B6AEC" blockId="9.[812,1399,1385,2071]" box="[822,1389,1385,1409]" pageId="9" pageNumber="230">
<heading id="812281A7BB0FFFDE013E1924642B6AEC" box="[822,1389,1385,1409]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="230" reason="8">
<taxonomicName id="1DD54D48BB0FFFDE013E1924646A6AEC" ID-CoL="C2V7L" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[822,1324,1385,1409]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="turkestanica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0FFFDE013E1924623F6AEC" box="[822,889,1386,1409]" italics="true" pageId="9" pageNumber="230">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0FFFDE0181192765596AEC" box="[905,1055,1385,1409]" italics="true" pageId="9" pageNumber="230">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0FFFDE0627192765F56AEC" box="[1071,1203,1385,1409]" italics="true" pageId="9" pageNumber="230">turkestanica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB0FFFDE073F1927642B6AEC" bold="true" box="[1335,1389,1385,1409]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="F39257A2BB0FFFDE073F1927642B6AEC" box="[1335,1389,1385,1409]" pageId="9" pageNumber="230" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDE061A19C865D76AF3" box="[1042,1169,1414,1438]" pageId="9" pageNumber="230" type="description">
<paragraph id="DA6A36CBBB0FFFDE061A19C865D76AF3" blockId="9.[812,1399,1385,2071]" box="[1042,1169,1414,1438]" pageId="9" pageNumber="230">
<figureCitation id="42EE2A4EBB0FFFDE061A19C865CB6AF3" box="[1042,1165,1414,1438]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="9" pageNumber="230">Figs 18–23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDD015A19E263316DEB" lastPageId="10" lastPageNumber="231" pageId="9" pageNumber="230" type="materials_examined">
<paragraph id="DA6A36CBBB0FFFDE015A19E264506A83" blockId="9.[812,1399,1385,2071]" pageId="9" pageNumber="230">
<materialsCitation id="6ABD3C96BB0FFFDE015A19E264506A83" collectionCode="TYPE, MATERIAL, ZISP, INS" country="Uzbekistan" county="Bos-su" location="Tashkent" municipality="Dobzhansky" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="BCC4AE0EBB0FFFDE015A19E262B86AD3" box="[850,1022,1451,1471]" pageId="9" pageNumber="230">TYPE MATERIAL</collectionCode>
.
<typeStatus id="056E8869BB0FFFDE060D19E565116AD3" box="[1029,1111,1451,1470]" pageId="9" pageNumber="230" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CCD3FD42BB0FFFDE066819E765326AD2" box="[1120,1140,1449,1471]" count="1" pageId="9" pageNumber="230" type="male">♂</specimenCount>
, labels as in
<figureCitation id="42EE2A4EBB0FFFDE06E919E564596AD3" box="[1249,1311,1451,1470]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="9" pageNumber="230">Fig. 23</figureCitation>
, [
<emphasis id="E8A1EAD9BB0FFFDE072719E562156ABB" bold="true" pageId="9" pageNumber="230">
<collectingCountry id="A2C2765BBB0FFFDE072719E562156ABB" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
</emphasis>
] [printed in Cyrillic, Russian]
<location id="DF0A6010BB0FFFDE068F198D659F6ABB" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE068F198D659F6ABB" box="[1159,1241,1475,1494]" country="Uzbekistan" county="Bos-su" municipality="Dobzhansky" name="Tashkent" pageId="9" pageNumber="230">Tashkent</location>
/ 25.iv.[1]925 /
<collectingMunicipality id="3A0EACB1BB0FFFDE0124199562DE6A83" box="[812,920,1499,1518]" pageId="9" pageNumber="230">Dobzhansky</collectingMunicipality>
// aryk
<collectingCounty id="330B4E47BB0FFFDE01D71992655D6A82" box="[991,1051,1500,1519]" pageId="9" pageNumber="230">Bos-su</collectingCounty>
(
<collectionCode id="BCC4AE0EBB0FFFDE0620199565116A83" box="[1064,1111,1499,1518]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
,
<collectionCode id="BCC4AE0EBB0FFFDE0669199565C56A83" box="[1121,1155,1499,1518]" pageId="9" pageNumber="230">INS</collectionCode>
_DIP_0000624).
</materialsCitation>
</paragraph>
<paragraph id="DA6A36CBBB0FFFDD015A19BA60836D4B" blockId="9.[812,1399,1385,2071]" lastBlockId="10.[189,776,313,2060]" lastPageId="10" lastPageNumber="231" pageId="9" pageNumber="230">
<materialsCitation id="6ABD3C96BB0FFFDE015A19BA62246973" collectionCode="ZISP" country="Uzbekistan" county="Bos-su" location="Tashkent" municipality="Dobzhansky" pageId="9" pageNumber="230" specimenCount="19" specimenCount-female="15" specimenCount-male="4" typeStatus="paratype">
<typeStatus id="056E8869BB0FFFDE015A19BA62EF696A" box="[850,937,1524,1543]" pageId="9" pageNumber="230" type="paratype">Paratypes</typeStatus>
.
<emphasis id="E8A1EAD9BB0FFFDE01BA19BD6566696B" bold="true" box="[946,1056,1523,1542]" pageId="9" pageNumber="230">
<collectingCountry id="A2C2765BBB0FFFDE01BA19BD655A696B" box="[946,1052,1523,1542]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
:
</emphasis>
same data as holotype (
<specimenCount id="CCD3FD42BB0FFFDE06FE19BD6476696A" box="[1270,1328,1521,1543]" count="4" pageId="9" pageNumber="230" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE073419BD6437696A" box="[1340,1393,1521,1543]" count="15" pageId="9" pageNumber="230" type="female">15 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01241A45621A6973" box="[812,860,1547,1566]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01671A4564356973" box="[879,1395,1545,1567]" collectingDate="1925-04-22" collectionCode="ZISP" collectorName="Dobzhansky" country="Uzbekistan" location="Tashkent" pageId="9" pageNumber="230" specimenCount="21" specimenCount-female="15" specimenCount-male="6" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE01671A4562F86973" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01671A4562F86973" box="[879,958,1547,1566]" country="Uzbekistan" name="Tashkent" pageId="9" pageNumber="230">Tashkent</location>
<date id="AE6B100BBB0FFFDE01C01A4565626973" box="[968,1060,1547,1566]" pageId="9" pageNumber="230" value="1925-04-22">
<collectingDate id="BE2FE9E3BB0FFFDE01C01A4565626973" box="[968,1060,1547,1566]" pageId="9" pageNumber="230" value="1925-04-22">22.iv.1925</collectingDate>
</date>
<collectorName id="7720531DBB0FFFDE06261A4565DC6973" box="[1070,1178,1547,1566]" pageId="9" pageNumber="230">Dobzhansky</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE06A21A4565A16972" box="[1194,1255,1545,1567]" count="6" pageId="9" pageNumber="230" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE06FE1A45646B6972" box="[1270,1325,1545,1567]" count="15" pageId="9" pageNumber="230" type="female">15 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE07351A45642B6973" box="[1341,1389,1547,1566]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01241A6A6435695B" box="[812,1395,1569,1591]" collectingDate="1871-03-25" collectionCode="ZMMU" collectorName="A. Fedtschenko" country="Uzbekistan" location="Tashkent" pageId="9" pageNumber="230" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="paratype">
same locality, [
<collectorName id="7720531DBB0FFFDE01A71A6D657B695B" box="[943,1085,1571,1590]" pageId="9" pageNumber="230">A. Fedtschenko</collectorName>
]:
<date id="AE6B100BBB0FFFDE06411A6D65E1695B" box="[1097,1191,1571,1590]" pageId="9" pageNumber="230" value="1871-03-25">
<collectingDate id="BE2FE9E3BB0FFFDE06411A6D65E1695B" box="[1097,1191,1571,1590]" pageId="9" pageNumber="230" value="1871-03-25">25.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE06BA1A6D65AD695A" box="[1202,1259,1569,1591]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE06FD1A6D645B695A" box="[1269,1309,1569,1591]" count="4" pageId="9" pageNumber="230" type="female">4 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE072F1A6D642B695B" box="[1319,1389,1571,1590]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE01241A7565306923" box="[812,1142,1593,1615]" collectingDate="1871-03-26" collectionCode="ZMMU" collectorName="A. Fedtschenko" country="Uzbekistan" isTailEnumeration="true" location="Tashkent" pageId="9" pageNumber="230" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE01241A7562CD6923" box="[812,907,1595,1614]" pageId="9" pageNumber="230" value="1871-03-26">
<collectingDate id="BE2FE9E3BB0FFFDE01241A7562CD6923" box="[812,907,1595,1614]" pageId="9" pageNumber="230" value="1871-03-26">26.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE01941A75629D6922" box="[924,987,1593,1615]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE01E31A75655F6922" box="[1003,1049,1593,1615]" count="2" pageId="9" pageNumber="230" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06211A7565366923" box="[1065,1136,1595,1614]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE068C1A75623F690B" collectingDate="1871-03-27" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE068C1A7565A46923" box="[1156,1250,1595,1614]" pageId="9" pageNumber="230" value="1871-03-27">
<collectingDate id="BE2FE9E3BB0FFFDE068C1A7565A46923" box="[1156,1250,1595,1614]" pageId="9" pageNumber="230" value="1871-03-27">27.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE06FC1A7564746922" box="[1268,1330,1593,1615]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE074B1A7564376922" box="[1347,1393,1593,1615]" count="5" pageId="9" pageNumber="230" type="female">5 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01241A1D6234690B" box="[812,882,1619,1638]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE018B1A1D65FC690B" box="[899,1210,1617,1639]" collectingDate="1871-03-28" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE018B1A1D62A7690B" box="[899,993,1619,1638]" pageId="9" pageNumber="230" value="1871-03-28">
<collectingDate id="BE2FE9E3BB0FFFDE018B1A1D62A7690B" box="[899,993,1619,1638]" pageId="9" pageNumber="230" value="1871-03-28">28.iii.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE01E71A1D656C690A" box="[1007,1066,1617,1639]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE063E1A1D6527690A" box="[1078,1121,1617,1639]" count="2" pageId="9" pageNumber="230" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06651A1D65F5690B" box="[1133,1203,1619,1638]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE06CC1A1D62DA6913" collectingDate="1871-04-01" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="13" specimenCount-female="9" specimenCount-male="4" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE06CC1A1D6450690B" box="[1220,1302,1619,1638]" pageId="9" pageNumber="230" value="1871-04-01">
<collectingDate id="BE2FE9E3BB0FFFDE06CC1A1D6450690B" box="[1220,1302,1619,1638]" pageId="9" pageNumber="230" value="1871-04-01">1.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE072C1A1D6419690A" box="[1316,1375,1617,1639]" count="4" pageId="9" pageNumber="230" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE07631A1D62026912" count="9" pageId="9" pageNumber="230" type="female">9 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01581A2562D06913" box="[848,918,1643,1662]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE01AE1A25653C6913" box="[934,1146,1641,1663]" collectingDate="1871-04-02" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE01AE1A2562B16913" box="[934,1015,1643,1662]" pageId="9" pageNumber="230" value="1871-04-02">
<collectingDate id="BE2FE9E3BB0FFFDE01AE1A2562B16913" box="[934,1015,1643,1662]" pageId="9" pageNumber="230" value="1871-04-02">2.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE060E1A2565646912" box="[1030,1058,1641,1663]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06251A2565356913" box="[1069,1139,1643,1662]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE068B1A2564356913" box="[1155,1395,1641,1663]" collectingDate="1871-04-03" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE068B1A2565936913" box="[1155,1237,1643,1662]" pageId="9" pageNumber="230" value="1871-04-03">
<collectingDate id="BE2FE9E3BB0FFFDE068B1A2565936913" box="[1155,1237,1643,1662]" pageId="9" pageNumber="230" value="1871-04-03">3.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE06EA1A25645D6912" box="[1250,1307,1641,1663]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE072F1A25642B6913" box="[1319,1389,1643,1662]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE01241ACD655569FB" box="[812,1043,1665,1687]" collectingDate="1871-04-10" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE01241ACD62CF69FB" box="[812,905,1667,1686]" pageId="9" pageNumber="230" value="1871-04-10">
<collectingDate id="BE2FE9E3BB0FFFDE01241ACD62CF69FB" box="[812,905,1667,1686]" pageId="9" pageNumber="230" value="1871-04-10">10.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE019C1ACD62FB69FA" box="[916,957,1665,1687]" count="2" pageId="9" pageNumber="230" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01CF1ACD654B69FB" box="[967,1037,1667,1686]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE06131ACD65B169FB" box="[1051,1271,1665,1687]" collectingDate="1871-04-11" collectionCode="ZMMU" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE06131ACD653E69FB" box="[1051,1144,1667,1686]" pageId="9" pageNumber="230" value="1871-04-11">
<collectingDate id="BE2FE9E3BB0FFFDE06131ACD653E69FB" box="[1051,1144,1667,1686]" pageId="9" pageNumber="230" value="1871-04-11">11.iv.1871</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE068D1ACD65E669FA" box="[1157,1184,1665,1687]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06A21ACD65B769FB" box="[1194,1265,1667,1686]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE06F71ACA653969C3" collectingDate="1923-04-11" collectionCode="ZMMU" collectorName="E. Smirnov" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality,
<date id="AE6B100BBB0FFFDE01241AD562CC69C3" box="[812,906,1691,1710]" pageId="9" pageNumber="230" value="1923-04-11">
<collectingDate id="BE2FE9E3BB0FFFDE01241AD562CC69C3" box="[812,906,1691,1710]" pageId="9" pageNumber="230" value="1923-04-11">11.iv.1923</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE019C1AD262B069C3" box="[916,1014,1691,1711]" pageId="9" pageNumber="230">E. Smirnov</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE060C1AD5656169C2" box="[1028,1063,1689,1711]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE063B1AD5653C69C3" box="[1075,1146,1691,1710]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE06801AD2654169AB" collectingDate="1929-04-04" collectionCode="ZISP" collectorName="B. Rohdendorf" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality,
<date id="AE6B100BBB0FFFDE070E1AD5641F69C3" box="[1286,1369,1691,1710]" pageId="9" pageNumber="230" value="1929-04-04">
<collectingDate id="BE2FE9E3BB0FFFDE070E1AD5641F69C3" box="[1286,1369,1691,1710]" pageId="9" pageNumber="230" value="1929-04-04">4.iv.1929</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE076A1AD262D569AB" pageId="9" pageNumber="230">B. Rohdendorf</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE01AA1AFD628069AA" box="[930,966,1713,1735]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01D91AFD654769AB" box="[977,1025,1715,1734]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE06191AFA622469B3" collectingDate="1929-04-17" collectionCode="ZISP" collectorName="L. Zimin" country="Uzbekistan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
same locality,
<date id="AE6B100BBB0FFFDE06981AFD65A969AB" box="[1168,1263,1715,1734]" pageId="9" pageNumber="230" value="1929-04-17">
<collectingDate id="BE2FE9E3BB0FFFDE06981AFD65A969AB" box="[1168,1263,1715,1734]" pageId="9" pageNumber="230" value="1929-04-17">17.iv.1929</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE06F11AFA640069AB" box="[1273,1350,1715,1735]" pageId="9" pageNumber="230">L. Zimin</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE075D1AFD643769AA" box="[1365,1393,1713,1735]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01241A85621A69B3" box="[812,860,1739,1758]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01651A8562DF699B" collectingDate="1922-06-07" collectionCode="ZISP" collectorName="N. Kuznetsov" country="Uzbekistan" location="Tashkent" municipality="Min-Bulak" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE01651A8562F869B3" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01651A8562F869B3" box="[877,958,1739,1758]" country="Uzbekistan" municipality="Min-Bulak" name="Tashkent" pageId="9" pageNumber="230">Tashkent</location>
, environs of
<collectingMunicipality id="3A0EACB1BB0FFFDE063C1A8565D369B3" box="[1076,1173,1739,1758]" pageId="9" pageNumber="230">Min-Bulak</collectingMunicipality>
,
<date id="AE6B100BBB0FFFDE06971A8565B569B3" box="[1183,1267,1739,1758]" pageId="9" pageNumber="230" value="1922-06-07">
<collectingDate id="BE2FE9E3BB0FFFDE06971A8565B569B3" box="[1183,1267,1739,1758]" pageId="9" pageNumber="230" value="1922-06-07">7.vi.1922</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE06F61A82643069B2" box="[1278,1398,1740,1759]" pageId="9" pageNumber="230">N. Kuznetsov</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE013D1AAD621E699A" box="[821,856,1761,1783]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE016B1AAD62D5699B" box="[867,915,1763,1782]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01AB1AAA6435699B" box="[931,1395,1761,1783]" collectingDate="1922-06-15" collectionCode="ZISP" collectorName="N. Kuznetsov" country="Uzbekistan" location="Tashkent" municipality="Min-Bulak" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality,
<date id="AE6B100BBB0FFFDE06291AAD6539699B" box="[1057,1151,1763,1782]" pageId="9" pageNumber="230" value="1922-06-15">
<collectingDate id="BE2FE9E3BB0FFFDE06291AAD6539699B" box="[1057,1151,1763,1782]" pageId="9" pageNumber="230" value="1922-06-15">15.vi.1922</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE06811AAA65B9699A" box="[1161,1279,1764,1783]" pageId="9" pageNumber="230">N. Kuznetsov</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE07061AAD6477699A" box="[1294,1329,1761,1783]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE07351AAD642B699B" box="[1341,1389,1763,1782]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01241AB26545684B" collectingDate="1958-04-18" collectionCode="ZISP" collectorName="Sychevskaya" country="Uzbekistan" elevation="1100" location="Kuram Ridge" municipality="Angren" pageId="9" pageNumber="230" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE01241AB262E16863" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01241AB262E16863" box="[812,935,1787,1807]" country="Uzbekistan" municipality="Angren" name="Kuram Ridge" pageId="9" pageNumber="230">Kuram Ridge</location>
, environs of
<collectingMunicipality id="3A0EACB1BB0FFFDE062A1AB565236863" box="[1058,1125,1787,1806]" pageId="9" pageNumber="230">Angren</collectingMunicipality>
,
<quantity id="1D2D9B2EBB0FFFDE06791AB565FE6862" box="[1137,1208,1787,1807]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="230" unit="m" value="1100.0">
<elevation id="51F8D1F8BB0FFFDE06791AB565FE6862" box="[1137,1208,1787,1807]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="230" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="AE6B100BBB0FFFDE06CD1AB564656863" box="[1221,1315,1787,1806]" pageId="9" pageNumber="230" value="1958-04-18">
<collectingDate id="BE2FE9E3BB0FFFDE06CD1AB564656863" box="[1221,1315,1787,1806]" pageId="9" pageNumber="230" value="1958-04-18">18.iv.1958</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE07381AB5621B684B" pageId="9" pageNumber="230">Sychevskaya</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE01641B5D62C9684A" box="[876,911,1809,1831]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE01921B5D6284684A" box="[922,962,1809,1831]" count="2" pageId="9" pageNumber="230" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01C51B5D62BB684B" box="[973,1021,1811,1830]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE06041B5A65DC6853" collectingDate="1963-05-16" collectingDateMax="1963-05-17" collectingDateMin="1963-05-16" collectionCode="ZISP" collectorName="Sugonyaev" country="Uzbekistan" elevation="1200" location="Ugam Ridge" pageId="9" pageNumber="230" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE06041B5A653A684B" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE06041B5A653A684B" box="[1036,1148,1811,1831]" country="Uzbekistan" name="Ugam Ridge" pageId="9" pageNumber="230">Ugam Ridge</location>
,
<locationDeviation id="CBD82FE9BB0FFFDE068D1B5D6461684B" box="[1157,1319,1811,1830]" location="Khumsan" pageId="9" pageNumber="230">
<quantity id="1D2D9B2EBB0FFFDE068D1B5D65FD684B" box="[1157,1211,1811,1830]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="230" unit="km" value="20.0">20 km</quantity>
S Khumsan
</locationDeviation>
,
<quantity id="1D2D9B2EBB0FFFDE07381B5D6435684A" box="[1328,1395,1811,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="230" unit="m" value="1200.0">
<elevation id="51F8D1F8BB0FFFDE07381B5D6435684A" box="[1328,1395,1811,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="230" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="AE6B100BBB0FFFDE01241B6562E76853" box="[812,929,1835,1854]" pageId="9" pageNumber="230" value="1963-05-16" valueMax="1963-05-17" valueMin="1963-05-16">
<collectingDate id="BE2FE9E3BB0FFFDE01241B6562E76853" box="[812,929,1835,1854]" pageId="9" pageNumber="230" value="1963-05-16" valueMax="1963-05-17" valueMin="1963-05-16">16-17.v.1963</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE01A51B65654B6853" box="[941,1037,1835,1854]" pageId="9" pageNumber="230">Sugonyaev</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE06141B6565116852" box="[1052,1111,1833,1855]" count="3" pageId="9" pageNumber="230" type="male">3 ♂♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE066C1B6565D26853" box="[1124,1172,1835,1854]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE06AE1B65645A683B" collectingDate="1963-05-23" collectingDateMax="1963-05-25" collectingDateMin="1963-05-23" collectionCode="ZISP" collectorName="Gurjeva" country="Uzbekistan" elevation="3000" location="Pskem Ridge" municipality="Nanay" pageId="9" pageNumber="230" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE06AE1B6564586853" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE06AE1B6564586853" box="[1190,1310,1835,1854]" country="Uzbekistan" municipality="Nanay" name="Pskem Ridge" pageId="9" pageNumber="230">Pskem Ridge</location>
, near
<collectingMunicipality id="3A0EACB1BB0FFFDE075E1B62620B683A" pageId="9" pageNumber="230">Nanay</collectingMunicipality>
,
<quantity id="1D2D9B2EBB0FFFDE015F1B0D62DD683A" box="[855,923,1859,1879]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="230" unit="m" value="3000.0">
<elevation id="51F8D1F8BB0FFFDE015F1B0D62DD683A" box="[855,923,1859,1879]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="230" unit="m" value="3000.0">3000 m</elevation>
</quantity>
,
<date id="AE6B100BBB0FFFDE01AC1B0D655B683B" box="[932,1053,1859,1878]" pageId="9" pageNumber="230" value="1963-05-23" valueMax="1963-05-25" valueMin="1963-05-23">
<collectingDate id="BE2FE9E3BB0FFFDE01AC1B0D655B683B" box="[932,1053,1859,1878]" pageId="9" pageNumber="230" value="1963-05-23" valueMax="1963-05-25" valueMin="1963-05-23">23–25.v.1963</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE062F1B0D652A683B" box="[1063,1132,1859,1878]" pageId="9" pageNumber="230">Gurjeva</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE06711B0D65F4683A" box="[1145,1202,1857,1879]" count="4" pageId="9" pageNumber="230" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE06B61B0D659D683A" box="[1214,1243,1857,1879]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06EE1B0D6450683B" box="[1254,1302,1859,1878]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE072E1B0D622468EB" collectingDate="1963-05-12" collectionCode="ZISP" collectorName="Sugonyaev" country="Uzbekistan" location="Tau Ridge" pageId="9" pageNumber="230" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
Karzhan-
<location id="DF0A6010BB0FFFDE01241B1262CF6803" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01241B1262CF6803" box="[812,905,1883,1903]" country="Uzbekistan" name="Tau Ridge" pageId="9" pageNumber="230">Tau Ridge</location>
,
<locationDeviation id="CBD82FE9BB0FFFDE019A1B1565026803" box="[914,1092,1883,1903]" location="Khumsan" pageId="9" pageNumber="230">
<quantity id="1D2D9B2EBB0FFFDE019A1B1562FB6803" box="[914,957,1883,1902]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="230" unit="km" value="5.0">5 km</quantity>
N of Khumsan
</locationDeviation>
,
<date id="AE6B100BBB0FFFDE06441B1565E36803" box="[1100,1189,1883,1902]" pageId="9" pageNumber="230" value="1963-05-12">
<collectingDate id="BE2FE9E3BB0FFFDE06441B1565E36803" box="[1100,1189,1883,1902]" pageId="9" pageNumber="230" value="1963-05-12">12.v.1963</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE06A51B15644B6803" box="[1197,1293,1883,1902]" pageId="9" pageNumber="230">Sugonyaev</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE07131B1564786802" box="[1307,1342,1881,1903]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE07411B1564376802" box="[1353,1393,1881,1903]" count="3" pageId="9" pageNumber="230" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01241B3D621A68EB" box="[812,860,1907,1926]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ABD3C96BB0FFFDE01641B3D643568F3" collectingDate="1907-05-06" collectionCode="ZISP" collectorName="A. Jacobson" country="Kazakhstan" elevation="960" location="Chak-pak" municipality="Vysokoe" pageId="9" pageNumber="230" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<emphasis id="E8A1EAD9BB0FFFDE01641B3D62A668EB" bold="true" box="[876,992,1907,1926]" pageId="9" pageNumber="230">
<collectingCountry id="A2C2765BBB0FFFDE01641B3D629A68EB" box="[876,988,1907,1926]" name="Kazakhstan" pageId="9" pageNumber="230">Kazakhstan</collectingCountry>
.
</emphasis>
<location id="DF0A6010BB0FFFDE01EF1B3D657B68EB" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01EF1B3D657B68EB" box="[999,1085,1907,1926]" country="Kazakhstan" municipality="Vysokoe" name="Chak-pak" pageId="9" pageNumber="230">Chak-pak</location>
, village
<collectingMunicipality id="3A0EACB1BB0FFFDE06801B3D659168EB" box="[1160,1239,1907,1926]" pageId="9" pageNumber="230">Vysokoe</collectingMunicipality>
,
<quantity id="1D2D9B2EBB0FFFDE06E91B3D645D68EA" box="[1249,1307,1907,1927]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="230" unit="m" value="960.0">
<elevation id="51F8D1F8BB0FFFDE06E91B3D645D68EA" box="[1249,1307,1907,1927]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="230" unit="m" value="960.0">960 m</elevation>
</quantity>
,
<date id="AE6B100BBB0FFFDE072D1B3D643568EB" box="[1317,1395,1907,1926]" pageId="9" pageNumber="230" value="1907-05-06">
<collectingDate id="BE2FE9E3BB0FFFDE072D1B3D643568EB" box="[1317,1395,1907,1926]" pageId="9" pageNumber="230" value="1907-05-06">6.v.1907</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE01241BC562D068F3" box="[812,918,1931,1950]" pageId="9" pageNumber="230">A. Jacobson</collectorName>
// coll. A. Semenov-Tian-Shansky (
<specimenCount id="CCD3FD42BB0FFFDE06D81BC5644C68F2" box="[1232,1290,1929,1951]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE071D1BC5647468F2" box="[1301,1330,1929,1951]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE07351BC5642B68F3" box="[1341,1389,1931,1950]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="230">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ABD3C96BB0FFFDE01241BED628B68A3" collectingDate="1965-05-24" collectionCode="ZMMU" collectorName="L. Zimina" country="Kazakhstan" location="Aksu-Zhabagly Nature Reserve" pageId="9" pageNumber="230" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB0FFFDE01241BED650768DA" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0FFFDE01241BED650768DA" box="[812,1089,1955,1975]" country="Kazakhstan" name="Aksu-Zhabagly Nature Reserve" pageId="9" pageNumber="230">Aksu-Zhabagly Nature Reserve</location>
,
<date id="AE6B100BBB0FFFDE06411BED65E468DB" box="[1097,1186,1955,1974]" pageId="9" pageNumber="230" value="1965-05-24">
<collectingDate id="BE2FE9E3BB0FFFDE06411BED65E468DB" box="[1097,1186,1955,1974]" pageId="9" pageNumber="230" value="1965-05-24">24.v.1965</collectingDate>
</date>
,
<collectorName id="7720531DBB0FFFDE06A31BEA644768DB" box="[1195,1281,1955,1975]" pageId="9" pageNumber="230">L. Zimina</collectorName>
(
<specimenCount id="CCD3FD42BB0FFFDE07181BED647568DA" box="[1296,1331,1953,1975]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE07361BED641D68DA" box="[1342,1371,1953,1975]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
on one pin,
<collectionCode id="BCC4AE0EBB0FFFDE01891BF5628E68A3" box="[897,968,1979,1998]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDE01D31BF2623F688B" collectingDate="1965-05-14" collectionCode="ZMMU" collectorName="L. Zimina" country="Kazakhstan" location="Aksu-Zhabagly Nature Reserve" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality and collector,
<date id="AE6B100BBB0FFFDE06E81BF5647168A3" box="[1248,1335,1979,1998]" pageId="9" pageNumber="230" value="1965-05-14">
<collectingDate id="BE2FE9E3BB0FFFDE06E81BF5647168A3" box="[1248,1335,1979,1998]" pageId="9" pageNumber="230" value="1965-05-14">14.v.1965</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE07411BF5643768A2" box="[1353,1393,1977,1999]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01241B9D6234688B" box="[812,882,2003,2022]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE018E1B9D653E688B" box="[902,1144,2001,2023]" collectingDate="1965-05-14" collectionCode="ZMMU" collectorName="L. Zimina" country="Kazakhstan" isTailEnumeration="true" location="Aksu-Zhabagly Nature Reserve" pageId="9" pageNumber="230" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE018E1B9D629B688B" box="[902,989,2003,2022]" pageId="9" pageNumber="230" value="1965-05-14">
<collectingDate id="BE2FE9E3BB0FFFDE018E1B9D629B688B" box="[902,989,2003,2022]" pageId="9" pageNumber="230" value="1965-05-14">14.v.1965</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE01E61B9D655A688A" box="[1006,1052,2001,2023]" count="2" pageId="9" pageNumber="230" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06231B9D6537688B" box="[1067,1137,2003,2022]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE068E1B9D6435688B" box="[1158,1395,2001,2023]" collectingDate="1965-05-21" collectionCode="ZMMU" country="Kazakhstan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE068E1B9D659A688B" box="[1158,1244,2003,2022]" pageId="9" pageNumber="230" value="1965-05-21">
<collectingDate id="BE2FE9E3BB0FFFDE068E1B9D659A688B" box="[1158,1244,2003,2022]" pageId="9" pageNumber="230" value="1965-05-21">21.v.1965</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE06E71B9D6451688A" box="[1263,1303,2001,2023]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE072F1B9D642B688B" box="[1319,1389,2003,2022]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE01241BA565026893" box="[812,1092,2025,2047]" collectingDate="1965-05-23" collectionCode="ZMMU" country="Kazakhstan" pageId="9" pageNumber="230" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE01241BA562C46893" box="[812,898,2027,2046]" pageId="9" pageNumber="230" value="1965-05-23">
<collectingDate id="BE2FE9E3BB0FFFDE01241BA562C46893" box="[812,898,2027,2046]" pageId="9" pageNumber="230" value="1965-05-23">23.v.1965</collectingDate>
</date>
(
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,
<specimenCount id="CCD3FD42BB0FFFDE01CA1BA562AA6892" box="[962,1004,2025,2047]" count="3" pageId="9" pageNumber="230" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01F01BA565786893" box="[1016,1086,2027,2046]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE06461BA564356893" box="[1102,1395,2025,2047]" collectingDate="1966-06-11" collectionCode="ZMMU" country="Kazakhstan" pageId="9" pageNumber="230" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE06461BA565EC6893" box="[1102,1194,2027,2046]" pageId="9" pageNumber="230" value="1966-06-11">
<collectingDate id="BE2FE9E3BB0FFFDE06461BA565EC6893" box="[1102,1194,2027,2046]" pageId="9" pageNumber="230" value="1966-06-11">11.vi.1966</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE06BF1BA565B76892" box="[1207,1265,2025,2047]" count="2" pageId="9" pageNumber="230" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0FFFDE06F51BA5645D6892" box="[1277,1307,2025,2047]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE072F1BA5642B6893" box="[1319,1389,2027,2046]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE0124144D6541677B" box="[812,1031,2049,2071]" collectingDate="1966-05-26" collectionCode="ZMMU" country="Kazakhstan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE0124144D62C5677B" box="[812,899,2051,2070]" pageId="9" pageNumber="230" value="1966-05-26">
<collectingDate id="BE2FE9E3BB0FFFDE0124144D62C5677B" box="[812,899,2051,2070]" pageId="9" pageNumber="230" value="1966-05-26">26.v.1966</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE019A144D62E8677A" box="[914,942,2049,2071]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE01B2144D6546677B" box="[954,1024,2051,2070]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="6ABD3C96BB0FFFDE0619144D65B5677B" box="[1041,1267,2049,2071]" collectingDate="1966-05-30" collectionCode="ZMMU" country="Kazakhstan" pageId="9" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="AE6B100BBB0FFFDE0619144D6521677B" box="[1041,1127,2051,2070]" pageId="9" pageNumber="230" value="1966-05-30">
<collectingDate id="BE2FE9E3BB0FFFDE0619144D6521677B" box="[1041,1127,2051,2070]" pageId="9" pageNumber="230" value="1966-05-30">30.v.1966</collectingDate>
</date>
(
<specimenCount id="CCD3FD42BB0FFFDE067E144D65DC677A" box="[1142,1178,2049,2071]" count="1" pageId="9" pageNumber="230" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0FFFDE06AE144D65AB677B" box="[1190,1261,2051,2070]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="230" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0FFFDD06F5144A63CF6E23" collectingDate="1977-06-30" collectionCode="ZMMU" collectorName="A. Shatalkin" country="Kazakhstan" lastPageId="10" lastPageNumber="231" pageId="9" pageNumber="230" specimenCount="11" specimenCount-female="3" specimenCount-male="8" typeStatus="paratype">
same locality,
<date id="AE6B100BBB0CFFDD02B51D75605D6E23" box="[189,283,315,334]" pageId="10" pageNumber="231" value="1977-06-30">
<collectingDate id="BE2FE9E3BB0CFFDD02B51D75605D6E23" box="[189,283,315,334]" pageId="10" pageNumber="231" value="1977-06-30">30.vi.1977</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD03221D7560DA6E23" box="[298,412,315,334]" pageId="10" pageNumber="231">A. Shatalkin</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD03A61D7560AA6E22" box="[430,492,313,335]" count="8" pageId="10" pageNumber="231" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0CFFDD03F51D75636D6E22" box="[509,555,313,335]" count="3" pageId="10" pageNumber="231" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD00331D7563C46E23" box="[571,642,315,334]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="231" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ABD3C96BB0CFFDD009E1D7563916E0B" collectingDate="1925-05-27" collectionCode="ZISP" collectorName="Dobzhansky" country="Kyrgyzstan" location="Kyrk-Kichiktau Mts." pageId="10" pageNumber="231" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="E8A1EAD9BB0CFFDD009E1D7562416E23" bold="true" box="[662,775,315,334]" pageId="10" pageNumber="231">
<collectingCountry id="A2C2765BBB0CFFDD009E1D7562446E23" box="[662,770,315,334]" name="Kyrgyzstan" pageId="10" pageNumber="231">Kyrgyzstan</collectingCountry>
.
</emphasis>
<location id="DF0A6010BB0CFFDD02B51D1D603F6E0A" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD02B51D1D603F6E0A" box="[189,377,339,359]" country="Kyrgyzstan" name="Kyrk-Kichiktau Mts." pageId="10" pageNumber="231">Kyrk-Kichiktau Mts.</location>
,
<date id="AE6B100BBB0CFFDD038E1D1D60986E0B" box="[390,478,339,358]" pageId="10" pageNumber="231" value="1925-05-27">
<collectingDate id="BE2FE9E3BB0CFFDD038E1D1D60986E0B" box="[390,478,339,358]" pageId="10" pageNumber="231" value="1925-05-27">27.v.1925</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD03E41D1D631E6E0B" box="[492,600,339,358]" pageId="10" pageNumber="231">Dobzhansky</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD00621D1D63D46E0A" box="[618,658,337,359]" count="1" pageId="10" pageNumber="231" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD00A91D1D63976E0B" box="[673,721,339,358]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD00ED1D1D63C56E13" collectingDate="1925-04-30" collectionCode="ZISP" collectorName="Dobzhansky" country="Kyrgyzstan" location="Aktash" pageId="10" pageNumber="231" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD00ED1D1D61A46E13" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD00ED1D1D61A46E13" country="Kyrgyzstan" name="Aktash" pageId="10" pageNumber="231">Aktash</location>
,
<date id="AE6B100BBB0CFFDD02E41D25600C6E13" box="[236,330,363,382]" pageId="10" pageNumber="231" value="1925-04-30">
<collectingDate id="BE2FE9E3BB0CFFDD02E41D25600C6E13" box="[236,330,363,382]" pageId="10" pageNumber="231" value="1925-04-30">30.iv.1925</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD035D1D2560876E13" box="[341,449,363,382]" pageId="10" pageNumber="231">Dobzhansky</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD03C71D25634C6E12" box="[463,522,361,383]" count="2" pageId="10" pageNumber="231" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0CFFDD001E1D2563066E12" box="[534,576,361,383]" count="3" pageId="10" pageNumber="231" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD00451D25633B6E13" box="[589,637,363,382]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD00861D2563886EFB" collectingDate="1925-05-25" collectionCode="ZISP" collectorName="Dobzhansky" country="Kyrgyzstan" location="Mikhaylovka" municipality="Kugart River" pageId="10" pageNumber="231" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD00861D2562456E13" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD00861D2562456E13" box="[654,771,363,382]" country="Kyrgyzstan" municipality="Kugart River" name="Mikhaylovka" pageId="10" pageNumber="231">Mikhaylovka</location>
, valley of
<collectingMunicipality id="3A0EACB1BB0CFFDD03071DCA60C56EFB" box="[271,387,387,407]" pageId="10" pageNumber="231">Kugart River</collectingMunicipality>
,
<date id="AE6B100BBB0CFFDD03851DCD60A36EFB" box="[397,485,387,406]" pageId="10" pageNumber="231" value="1925-05-25">
<collectingDate id="BE2FE9E3BB0CFFDD03851DCD60A36EFB" box="[397,485,387,406]" pageId="10" pageNumber="231" value="1925-05-25">25.v.1925</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD03E71DCD631D6EFB" box="[495,603,387,406]" pageId="10" pageNumber="231">Dobzhansky</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD00611DCD63CA6EFA" box="[617,652,385,407]" count="1" pageId="10" pageNumber="231" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD00901DCD638E6EFB" box="[664,712,387,406]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD00D01DCD63F06EC3" collectingDate="1925-05-08" collectionCode="ZISP" collectorName="Dobzhansky" country="Kyrgyzstan" location="Kugart-su River" pageId="10" pageNumber="231" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
shore of
<location id="DF0A6010BB0CFFDD02D01DD2602D6EC3" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD02D01DD2602D6EC3" box="[216,363,411,431]" country="Kyrgyzstan" name="Kugart-su River" pageId="10" pageNumber="231">Kugart-su River</location>
,
<date id="AE6B100BBB0CFFDD03701DD560836EC3" box="[376,453,411,430]" pageId="10" pageNumber="231" value="1925-05-08">
<collectingDate id="BE2FE9E3BB0CFFDD03701DD560836EC3" box="[376,453,411,430]" pageId="10" pageNumber="231" value="1925-05-08">8.v.1925</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD03DB1DD563796EC3" box="[467,575,411,430]" pageId="10" pageNumber="231">Dobzhansky</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD00591DD563366EC2" box="[593,624,409,431]" count="1" pageId="10" pageNumber="231" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD00881DD563F66EC3" box="[640,688,411,430]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD00CB1DD5638D6EAB" collectingDate="1963-06-09" collectionCode="ZISP" collectorName="Gurjeva" country="Kyrgyzstan" elevation="2000" location="Chatkal Nature Reserve" pageId="10" pageNumber="231" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD00CB1DD560006EAA" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD00CB1DD560006EAA" country="Kyrgyzstan" name="Chatkal Nature Reserve" pageId="10" pageNumber="231">Chatkal Nature Reserve</location>
,
<quantity id="1D2D9B2EBB0CFFDD03581DFD60D36EAA" box="[336,405,435,455]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="231" unit="m" value="2000.0">
<elevation id="51F8D1F8BB0CFFDD03581DFD60D36EAA" box="[336,405,435,455]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="231" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="AE6B100BBB0CFFDD03961DFD60B46EAB" box="[414,498,435,454]" pageId="10" pageNumber="231" value="1963-06-09">
<collectingDate id="BE2FE9E3BB0CFFDD03961DFD60B46EAB" box="[414,498,435,454]" pageId="10" pageNumber="231" value="1963-06-09">9.vi.1963</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD03F41DFD63076EAB" box="[508,577,435,454]" pageId="10" pageNumber="231">Gurjeva</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD00471DFD63CF6EAA" box="[591,649,433,455]" count="2" pageId="10" pageNumber="231" type="male">2 ♂♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD009D1DFD63836EAB" box="[661,709,435,454]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD00DE1DFA61B56E9B" collectingDate="1966-06-18" collectionCode="V, ZISP" collectorName="Zaitsev" country="Kyrgyzstan" elevation="2300" location="Kamchik Pass" municipality="Kuram Ridge" pageId="10" pageNumber="231" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD00DE1DFA60556EB2" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD00DE1DFA60556EB2" country="Kyrgyzstan" municipality="Kuram Ridge" name="Kamchik Pass" pageId="10" pageNumber="231">Kamchik Pass</location>
,
<quantity id="1D2D9B2EBB0CFFDD03291D85602F6EB2" box="[289,361,459,479]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="231" unit="m" value="2300.0">
<elevation id="51F8D1F8BB0CFFDD03291D85602F6EB2" box="[289,361,459,479]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="231" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<collectingMunicipality id="3A0EACB1BB0CFFDD037E1D8260B76EB3" box="[374,497,459,479]" pageId="10" pageNumber="231">Kuram Ridge</collectingMunicipality>
,
<date id="AE6B100BBB0CFFDD03F61D85631A6EB3" box="[510,604,459,478]" pageId="10" pageNumber="231" value="1966-06-18">
<collectingDate id="BE2FE9E3BB0CFFDD03F61D85631A6EB3" box="[510,604,459,478]" pageId="10" pageNumber="231" value="1966-06-18">18.vi.1966</collectingDate>
</date>
,
<collectionCode id="BCC4AE0EBB0CFFDD00611D82633D6EB2" box="[617,635,460,479]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="231" type="Museum">V</collectionCode>
.
<collectorName id="7720531DBB0CFFDD00801D8563816EB3" box="[648,711,459,478]" pageId="10" pageNumber="231">Zaitsev</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD00D21D8562476EB2" box="[730,769,457,479]" count="1" pageId="10" pageNumber="231" type="male">1 ♂</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD02B51DAD61AB6E9B" box="[189,237,483,502]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="231">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD02F41DAD608B6D63" collectingDate="1952-05-29" collectionCode="ZMMU" collectorName="Zhelokhovtsev" country="Kyrgyzstan" elevation="2000" location="Chatkal Ridge" municipality="Sary-Chilek" pageId="10" pageNumber="231" specimenCount="9" specimenCount-female="4" specimenCount-male="5" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD02F41DAD603A6E9B" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD02F41DAD603A6E9B" box="[252,380,483,502]" country="Kyrgyzstan" municipality="Sary-Chilek" name="Chatkal Ridge" pageId="10" pageNumber="231">Chatkal Ridge</location>
,
<collectingMunicipality id="3A0EACB1BB0CFFDD038D1DAD60B66E9B" box="[389,496,483,502]" pageId="10" pageNumber="231">Sary-Chilek</collectingMunicipality>
,
<quantity id="1D2D9B2EBB0CFFDD03F11DAD63706E9B" box="[505,566,483,502]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="231" unit="m" value="2000.0">
<elevation id="51F8D1F8BB0CFFDD03F11DAD63706E9B" box="[505,566,483,502]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="231" unit="m" value="2000.0">2000m</elevation>
</quantity>
,
<date id="AE6B100BBB0CFFDD00481DAD63DE6E9B" box="[576,664,483,502]" pageId="10" pageNumber="231" value="1952-05-29">
<collectingDate id="BE2FE9E3BB0CFFDD00481DAD63DE6E9B" box="[576,664,483,502]" pageId="10" pageNumber="231" value="1952-05-29">29.v.1952</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD00AA1DAD61A66D62" pageId="10" pageNumber="231">Zhelokhovtsev</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD02FA1DB560766D62" box="[242,304,505,527]" count="5" pageId="10" pageNumber="231" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="CCD3FD42BB0CFFDD03491DB560296D62" box="[321,367,505,527]" count="4" pageId="10" pageNumber="231" type="female">4 ♀♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD03881DB560816D63" box="[384,455,507,526]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="231" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6ABD3C96BB0CFFDD03D41DB560876D4B" collectingDate="1952-06-06" collectionCode="ZMMU" collectorName="Zhelokhovtsev" country="Kyrgyzstan" location="Chatkal Ridge" municipality="Arkit" pageId="10" pageNumber="231" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="DF0A6010BB0CFFDD03D41DB563276D63" LSID="urn:lsid:plazi:treatment:527C87DDBB0FFFDB013E1924624E6D8C:DF0A6010BB0CFFDD03D41DB563276D63" box="[476,609,507,526]" country="Kyrgyzstan" municipality="Arkit" name="Chatkal Ridge" pageId="10" pageNumber="231">Chatkal Ridge</location>
,
<collectingMunicipality id="3A0EACB1BB0CFFDD00781DB563E66D63" box="[624,672,507,526]" pageId="10" pageNumber="231">Arkit</collectingMunicipality>
,
<date id="AE6B100BBB0CFFDD00A61DB562456D63" box="[686,771,507,526]" pageId="10" pageNumber="231" value="1952-06-06">
<collectingDate id="BE2FE9E3BB0CFFDD00A61DB562456D63" box="[686,771,507,526]" pageId="10" pageNumber="231" value="1952-06-06">6.vi.1952</collectingDate>
</date>
,
<collectorName id="7720531DBB0CFFDD02B51E5D607B6D4B" box="[189,317,531,550]" pageId="10" pageNumber="231">Zhelokhovtsev</collectorName>
(
<specimenCount id="CCD3FD42BB0CFFDD03441E5D602E6D4A" box="[332,360,529,551]" count="1" pageId="10" pageNumber="231" type="female">1 ♀</specimenCount>
,
<collectionCode id="BCC4AE0EBB0CFFDD037C1E5D60FD6D4B" box="[372,443,531,550]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="231" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DA6A36CBBB0CFFDD02EA1E6563316DEB" blockId="10.[189,776,313,2060]" pageId="10" pageNumber="231">
<materialsCitation id="6ABD3C96BB0CFFDD02EA1E6563356DEB" collectingDate="1871-03-25" collectingDateMax="1871-04-10" collectingDateMin="1871-03-25" collectionCode="MATERIAL" collectorName="A. Fedtschenko" country="Uzbekistan" location="Uzbekistan" pageId="10" pageNumber="231" specimenCount="9" specimenCount-female="8" specimenCount-male="1">
ADDITIONAL
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EXAMINED.
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<collectingCountry id="A2C2765BBB0CFFDD006A1E65638C6D53" box="[610,714,555,574]" name="Uzbekistan" pageId="10" pageNumber="231">Uzbekistan</collectingCountry>
.
</emphasis>
Tashkent, [
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]:
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<collectingDate id="BE2FE9E3BB0CFFDD03A31E0D63486D3B" box="[427,526,579,598]" pageId="10" pageNumber="231" value="1871-03-25">25.iii.1871</collectingDate>
</date>
(
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),
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</date>
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),
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<collectingDate id="BE2FE9E3BB0CFFDD02B51E15605C6D03" box="[189,282,603,622]" pageId="10" pageNumber="231" value="1871-03-28">28.iii.1871</collectingDate>
</date>
(
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),
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<collectingDate id="BE2FE9E3BB0CFFDD03521E1560ED6D03" box="[346,427,603,622]" pageId="10" pageNumber="231" value="1871-04-05">5.iv.1871</collectingDate>
</date>
(
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),
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<collectingDate id="BE2FE9E3BB0CFFDD03FC1E1563166D03" box="[500,592,603,622]" pageId="10" pageNumber="231" value="1871-04-10">10.iv.1871</collectingDate>
</date>
(
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),
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(
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),
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</date>
(
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) (mostly eaten by dermestids)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0CFFDD02EA1EC5607E6C2C" pageId="10" pageNumber="231" type="diagnosis">
<paragraph id="DA6A36CBBB0CFFDD02EA1EC5607E6C2C" blockId="10.[189,776,313,2060]" pageId="10" pageNumber="231">
DIAGNOSIS. Mid-sized species (body length
<quantity id="1D2D9B2EBB0CFFDD00901EC563BA6DCD" box="[664,764,651,673]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="10" pageNumber="231" unit="mm" value="6.25" valueMax="6.5" valueMin="6.0">6–6.5 mm</quantity>
); eyes dichoptic, palpus black; postpronotal lobe shiny on outer side; acrostichal and presutural dorsocentral setae very short, the former biserial, the latter uniserial, presutural supra-alar seta present; femora entirely black; hind femur strongly thickened, hind tibia geniculate; abdominal tergites extensively shiny.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0CFFDB02EA1F08639F6D36" lastPageId="12" lastPageNumber="233" pageId="10" pageNumber="231" type="description">
<paragraph id="DA6A36CBBB0CFFDD02EA1F0863BF6A56" blockId="10.[189,776,313,2060]" pageId="10" pageNumber="231">
DESCRIPTION. Body length 5.9–6.6– mm, wing 6.0–
<quantity id="1D2D9B2EBB0CFFDD02B51F2E604A6C1B" box="[189,268,864,886]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="231" unit="mm" value="6.4">6.4 mm</quantity>
.
<emphasis id="E8A1EAD9BB0CFFDD03101F2F600A6C1B" bold="true" box="[280,332,865,886]" pageId="10" pageNumber="231">Male</emphasis>
(
<figureCitation id="42EE2A4EBB0CFFDD03551F2F60EE6C1B" box="[349,424,865,887]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="10" pageNumber="231">Fig. 18</figureCitation>
). Head with occiput, postgena and gena densely greyish pruinose; ocellar triangle subshiny, clypeus shiny, frons and face (except shiny lower margin) densely brownish pruinose; head setation mostly black. Dichoptic, ommatidia equally small. Frons broad, nearly as broad as distance between outer margins of posterior ocelli, almost parallel-sided, with marginal setulae. Face broader than frons, bare. Ocellar setae long, strong. Postocular setae long, strong on upper portion and very short on lower portion; occiput with almost regular transverse row of long, strong setae (more numerous laterally); postgena with yellowish, mostly fine setae. Antenna with scape and pedicel brownish, postpedicel and stylus black; scape nearly 2X longer than subglobular pedicel, both with short setulae; postpedicel 3–3.5X as long as basal width; stylus long, only slightly shorter than postpedicel. Proboscis with labrum brownish, paler apically; nearly 2X longer than eye height; palpus short, black; bearing sparse, long, black, fine setae.
</paragraph>
<paragraph id="DA6A36CBBB0CFFDD02EA190F63AB68BA" blockId="10.[189,776,313,2060]" pageId="10" pageNumber="231">
Thorax almost entirely black in ground-colour, postalar tubercle somewhat yellowish brown translucent; mostly densely greyish pruinose and black setose (except noted); postpronotal lobe shiny on outer side; mesoscutum (
<figureCitation id="42EE2A4EBB0CFFDD007B19DF63FC6ACB" box="[627,698,1425,1447]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="10" pageNumber="231">Fig. 19</figureCitation>
) with 4 brown vittae (median vittae usually more distinct). Prosternum bare. Proepisternum with tuft of 4–5 strong setae and usually a few pale fine setae on lower part, bare on upper part (rarely 1 black fine seta present). Antepronotum with 5–6 short, spine-like setae. Postpronotal lobe with 7–10 strong, moderately long setae anteriorly (number, length, robustness and position variable); postpronotal seta usually nearly as long as longest anterior setae (sometimes slightly longer). Mesonotal setae: acrostichals irregularly biserial, mostly very short and fine (2 anteriormost setae spine-like and somewhat longer), absent on prescutellar depression; dorsocentral setae uniserial, presuturals similar to acrostichals, postsuturals becoming longer towards scutellum, 2 pairs of long, strong prescutellars; 1 fine presutural intra-alar (sometimes absent), 1 moderately long presutural supra-alar; 3 notopleurals, 1 postsutural supra-alar, 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair slightly shorter); in addition, a few setulae present on anterior half of notopleuron. Laterotergite with 3–5 strong, long and a few short, fine setae (at least 3 strong setae always black, fine setae varying from black to pale). Anterior and posterior spiracles brownish.
</paragraph>
<paragraph id="DA6A36CBBB0CFFDD02EA1B92658B6B4C" blockId="10.[189,776,313,2060]" lastBlockId="10.[812,1399,315,2071]" pageId="10" pageNumber="231">Legs with coxae densely greyish pruinose, remaining podomeres subshiny; black setose. Legs colour: coxae, trochanters, femora and tarsi entirely dark brown; fore tibia mostly brown, brownish yellow near base; mid and hind tibiae extensively yellow, narrowly brownish on apex. Coxae and trochanters with simple setae. Fore femur with row of anteroventral setae, which are moderately long, spine-like on apical 1/2 to 2/3 and short, fine closer to base; row of short, fine posteroventral setae; short, strong setae closer to apex anteriorly. Fore tibia with row of 5–6 short, strong anterodorsal and 2–3 similar posterodorsal setae closer to apex; ventral setulae somewhat longer on about basal half. Fore basitarsus slender, mostly covered with simple setulae; with short, strong, anteroventral and posteroventral setae. Mid femur without whitish ventral pubescence; with row of short setae on about apical half anteriorly; rows of anteroventral and posteroventral, short, mostly strong setae (shorter and finer near apex); covered with spinule-like setulae on ventral surface; 1 short, strong anterior subapical seta. Mid tibia with reduced setulae (denser ventrally); 3–4 short, spine-like anterodorsal, 1 dorsal near base and 1–2 posterodorsal setae on basal half. Mid tarsus as fore tarsus. Hind femur strongly thickened, nearly 2X broader than mid femur; with moderately long, spine-like anteroventral and posteroventral setae; dense spines on ventral face; 1 short, strong anterior subapical seta. Hind tibia geniculate, slightly arcuate (anterior view); with reduced setulae; bearing 4–5 short setae of different robustness dorsally; sparse erect setulae ventrally; very short, fine seta in posteroapical comb. Hind basitarsus slightly thickened; setation as in fore basitarsus.</paragraph>
<paragraph id="DA6A36CBBB0CFFDD015A1868640A6A41" blockId="10.[812,1399,315,2071]" pageId="10" pageNumber="231">
Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins mostly brownish, brownish yellow near base, well-sclerotized (except noted). Veins R
<subScript id="4651348EBB0CFFDD0741182464166B1B" attach="left" box="[1353,1360,1130,1142]" fontSize="5" pageId="10" pageNumber="231">5</subScript>
and M
<subScript id="4651348EBB0CFFDD014918CB620E6BFC" attach="left" box="[833,840,1157,1169]" fontSize="5" pageId="10" pageNumber="231">1</subScript>
strongly divergent towards wing margin; radial fork acute; R
<subScript id="4651348EBB0CFFDD013418EE62056BC1" attach="left" box="[828,835,1184,1196]" fontSize="5" pageId="10" pageNumber="231">5</subScript>
meeting costa before wing apex; CuA+CuP (anal vein) complete but slightly weakened beyond middle. Cell dm moderately large, with elongate apex; apical portion of M
<subScript id="4651348EBB0CFFDD0726189B64736B8C" attach="left" box="[1326,1333,1237,1249]" fontSize="5" pageId="10" pageNumber="231">4</subScript>
nearly 3X longer than its middle portion. Pterostigma absent. Basal costal seta absent. Anal lobe well-developed; axillary incision obtuse. Squama yellow, pale fringed. Halter pale yellow.
</paragraph>
<paragraph id="DA6A36CBBB0CFFDD015A197F646A6831" blockId="10.[812,1399,315,2071]" pageId="10" pageNumber="231">Abdomen black to dark brown in ground colour. Tergite 1 shiny along posterior margin; tergites 2–5 extensively shiny, narrowly densely grey pruinose along anterior margin and somewhat broader laterally; tergites 6 and 7 with brownish grey iridescent spot dorsally, shiny laterally; tergite 8 hidden by tergite 7; tergites with greatly reduced setation, mostly covered with sparse, fine, black setulae; tergites 1–5 with long, strong, black posteromarginal setae laterally. Sternites 1–6 densely grey pruinose, sternites 7–8 subshiny; sternite 1 bare, sternites 2–6 with sparse black and pale setulae; sternites 7–8 black setose, with moderately long posteromarginal setae. Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; small, isolated sclerite just beyond posterior margin of pleuron 7; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, somewhat constricted anteriorly, with 3 lateral projections and concavity between them (anterior projections small; posterior projection large, subglobular).</paragraph>
<paragraph id="DA6A36CBBB0CFFDC015A1B2F647F6E3C" blockId="10.[812,1399,315,2071]" lastBlockId="11.[812,1399,315,604]" lastPageId="11" lastPageNumber="232" pageId="10" pageNumber="231">
Hypopygium (
<figureCitation id="42EE2A4EBB0CFFDD01D51B2F6564681B" box="[989,1058,1889,1911]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="10" pageNumber="231">Fig. 20</figureCitation>
) moderately large; epandrial lamella and cerci dark brown; hypandrium pale brown; phallus mostly yellow, curved slender apical portion dark brown. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella rather subrectangular (lateral view); with short, fine, black setae, more numerous along lower margin.Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, with rounded apex; bare; gonocoxal apodeme small. Cerci (
<figureCitation id="42EE2A4EBB0DFFDC00D71D1861906EEB" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="11" pageNumber="232">Fig. 21</figureCitation>
) separated with each other and from epandrium; cercus elongate ovate (lateral view), slightly shorter than epandrial lamella, with additional lobe dorsally, smoothed on inner margin; with dense black setulae. Phallus (
<figureCitation id="42EE2A4EBB0DFFDC00651D8F63F06EBB" box="[621,694,449,471]" captionStart="Figs 18–23" captionStartId="11.[226,263,1918,1937]" captionTargetBox="[194,1399,642,1897]" captionTargetId="figure-316@11.[189,1399,633,1901]" captionTargetPageId="11" captionText="Figs 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 18–23. Empis (Polyblepharis) turkestanica Shamshev, sp.n., самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." pageId="11" pageNumber="232">Fig. 22</figureCitation>
) almost entirely hidden; zigzag bent, strongly constricted beyond middle; basal portion thick, gently curved and becoming narrower distad (lateral view); apical, bend portion broader proximally, its subapical part slender, with tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.
</paragraph>
<paragraph id="DA6A36CBBB0DFFDC015A1D18651F6D31" blockId="11.[812,1399,315,604]" pageId="11" pageNumber="232">
<emphasis id="E8A1EAD9BB0DFFDC015A1D1862E56E06" bold="true" box="[850,931,342,363]" pageId="11" pageNumber="232">Female.</emphasis>
Similar to male except as follows. Fore tibia with 6 anterodorsal, 2 posterodorsal, 4–5 finer anteroventral and 4–5 posteroventral setae. Mid femur with minute anteroventral and posteroventral setae. Mid tibia with 4 anterodorsal, 1 dorsal near base, 2 posterodorsal, 2–3 anteroventral and 2 posteroventral setae. Abdominal tergites 2–5 somewhat broader pruinose laterally; only tergites 1–3 with strong posteromarginal setae; sternite 6 faintly pruinose, tergite 6 entirely shiny, segments 7–10 shiny; cercus black, long, slender, with scattered setulae.
</paragraph>
<caption id="8EAA6643BB0DFFDC02EA1B3062DE6760" pageId="11" pageNumber="232" startId="11.[226,263,1918,1937]" targetBox="[194,1399,642,1897]" targetPageId="11" targetType="figure">
<paragraph id="DA6A36CBBB0DFFDC02EA1B30628468AC" blockId="11.[189,1399,1918,2061]" pageId="11" pageNumber="232">
Figs 18–23.
<taxonomicName id="1DD54D48BB0DFFDC03581B3063A168FC" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[336,743,1918,1937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="232" phylum="Arthropoda" rank="species" species="turkestanica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0DFFDC03581B3060C368FD" box="[336,389,1918,1936]" italics="true" pageId="11" pageNumber="232">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0DFFDC039C1B30634B68FC" box="[404,525,1918,1937]" italics="true" pageId="11" pageNumber="232">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0DFFDC00131B3063C368FC" box="[539,645,1918,1937]" italics="true" pageId="11" pageNumber="232">turkestanica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB0DFFDC00F91B30625A68FC" bold="true" box="[753,796,1918,1937]" pageId="11" pageNumber="232">
<taxonomicNameLabel id="F39257A2BB0DFFDC00F91B30625A68FC" box="[753,796,1918,1937]" pageId="11" pageNumber="232" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 18–19 — habitus, holotype; 20 — hypopygium; 21 — cerci; 22 — phallus; 23 — holotype labels; 18, 20, 22 — lateral view; 19, 21— dorsal view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm.
</paragraph>
<paragraph id="DA6A36CBBB0DFFDC02EA1B8462DE6760" blockId="11.[189,1399,1918,2061]" pageId="11" pageNumber="232">
Рис. 18–23.
<taxonomicName id="1DD54D48BB0DFFDC03471B8463A468B1" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[335,738,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="232" phylum="Arthropoda" rank="species" species="turkestanica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0DFFDC03471B8460C268B1" box="[335,388,1994,2012]" italics="true" pageId="11" pageNumber="232">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0DFFDC039A1B87634D68B1" box="[402,523,1993,2012]" italics="true" pageId="11" pageNumber="232">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0DFFDC00101B8763C468B1" box="[536,642,1993,2012]" italics="true" pageId="11" pageNumber="232">turkestanica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="E8A1EAD9BB0DFFDC00E31B87625068B1" bold="true" box="[747,790,1993,2012]" pageId="11" pageNumber="232">
<taxonomicNameLabel id="F39257A2BB0DFFDC00E31B87625068B1" box="[747,790,1993,2012]" pageId="11" pageNumber="232" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 18–19 — габитус, голотип; 20 — гипопигий; 21 — церки; 22 — фаллус; 23 — Этикетки голотипа; 18, 20, 22 — сбоку; 19, 21— сверху. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм.
</paragraph>
</caption>
<paragraph id="DA6A36CBBB0AFFDB02EA1D75639F6D36" blockId="12.[189,776,315,737]" pageId="12" pageNumber="233">
DIFFERENTIAL DIAGNOSIS. The new species resembles very much the species of the
<taxonomicName id="1DD54D48BB0AFFDB001F1D1963C16E06" authorityName="Meigen" authorityYear="1822" box="[535,647,342,364]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="233" phylum="Arthropoda" rank="species" species="albicans">
<emphasis id="E8A1EAD9BB0AFFDB001F1D1963606E01" box="[535,550,343,364]" italics="true" pageId="12" pageNumber="233">E</emphasis>
.
<emphasis id="E8A1EAD9BB0AFFDB003D1D1863C16E06" box="[565,647,342,363]" italics="true" pageId="12" pageNumber="233">albicans</emphasis>
</taxonomicName>
group sensu
<bibRefCitation id="BE444B3ABB0AFFDB02B51D3F60006EEB" author="Chvala M." box="[189,326,369,391]" pageId="12" pageNumber="233" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála [1999]</bibRefCitation>
occurring in stepper regions of Central Europe and the European part of
<collectingCountry id="A2C2765BBB0AFFDB03CC1DC5634E6ECC" box="[452,520,395,417]" name="Russia" pageId="12" pageNumber="233">Russia</collectingCountry>
. The new species can be readily distinguished from all species of this group primarily by black palpus. Amongst species of
<taxonomicName id="1DD54D48BB0AFFDB003D1D8F63FA6EBB" box="[565,700,449,470]" pageId="12" pageNumber="233" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0AFFDB003D1D8F63FA6EBB" box="[565,700,449,470]" italics="true" pageId="12" pageNumber="233">Polyblepharis</emphasis>
</taxonomicName>
known from Middle Asia, the new species is most similar to
<taxonomicName id="1DD54D48BB0AFFDB00FA1D9260386D61" authority="Engel, 1946" authorityName="Engel" authorityYear="1946" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="233" phylum="Arthropoda" rank="species" species="saltans">
<emphasis id="E8A1EAD9BB0AFFDB00FA1D9262476E9C" box="[754,769,476,497]" italics="true" pageId="12" pageNumber="233">E</emphasis>
.
<emphasis id="E8A1EAD9BB0AFFDB02B51DB860476D66" box="[189,257,502,523]" italics="true" pageId="12" pageNumber="233">saltans</emphasis>
Engel, 1946
</taxonomicName>
. The new species differs from
<taxonomicName id="1DD54D48BB0AFFDB00AF1DB962406D66" authorityName="Engel" authorityYear="1946" box="[679,774,502,524]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="233" phylum="Arthropoda" rank="species" species="saltans">
<emphasis id="E8A1EAD9BB0AFFDB00AF1DB963F06D61" box="[679,694,503,524]" italics="true" pageId="12" pageNumber="233">E</emphasis>
.
<emphasis id="E8A1EAD9BB0AFFDB00CA1DB862406D66" box="[706,774,502,523]" italics="true" pageId="12" pageNumber="233">saltans</emphasis>
</taxonomicName>
primarily by partly shiny postpronotal lobe, presence of a presutural supra-alar seta and by entirely dark brown femora (in
<taxonomicName id="1DD54D48BB0AFFDB02D61E0960786D36" authorityName="Engel" authorityYear="1946" box="[222,318,582,604]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="233" phylum="Arthropoda" rank="species" species="saltans">
<emphasis id="E8A1EAD9BB0AFFDB02D61E0961AB6D31" box="[222,237,583,604]" italics="true" pageId="12" pageNumber="233">E</emphasis>
.
<emphasis id="E8A1EAD9BB0AFFDB02F21E0860786D36" box="[250,318,582,603]" italics="true" pageId="12" pageNumber="233">saltans</emphasis>
</taxonomicName>
all femora are brownish yellow on apex).
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0AFFDB02EA1E2F607B6DAA" pageId="12" pageNumber="233" type="etymology">
<paragraph id="DA6A36CBBB0AFFDB02EA1E2F607B6DAA" blockId="12.[189,776,315,737]" pageId="12" pageNumber="233">ETYMOLOGY. The epithet refers to the name of a historical region, where the new species was collected, partly coinciding with an area now commonly called Central Asia or Middle Asia.</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0AFFDB02EA1E85624E6D8C" box="[226,776,715,737]" pageId="12" pageNumber="233" type="distribution">
<paragraph id="DA6A36CBBB0AFFDB02EA1E85624E6D8C" blockId="12.[189,776,315,737]" box="[226,776,715,737]" pageId="12" pageNumber="233">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB0AFFDB03911E85634A6D8C" box="[409,524,715,737]" name="Kazakhstan" pageId="12" pageNumber="233">Kazakhstan</collectingCountry>
,
<collectingCountry id="A2C2765BBB0AFFDB001F1E8263CF6D8C" box="[535,649,716,737]" name="Kyrgyzstan" pageId="12" pageNumber="233">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="A2C2765BBB0AFFDB009C1E8562426D8C" box="[660,772,715,737]" name="Uzbekistan" pageId="12" pageNumber="233">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

129
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<taxonomicName id="1DD54D48BB0FFFDE014E18C8641A6BF0" ID-CoL="8BZQ5" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[838,1372,1157,1181]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="tajikistanica" subGenus="Polyblepharis">
<emphasis id="E8A1EAD9BB0FFFDE014E18C862CF6BF0" box="[838,905,1158,1181]" italics="true" pageId="9" pageNumber="230">Empis</emphasis>
(
<emphasis id="E8A1EAD9BB0FFFDE019218CB65766BF0" box="[922,1072,1157,1181]" italics="true" pageId="9" pageNumber="230">Polyblepharis</emphasis>
)
<emphasis id="E8A1EAD9BB0FFFDE064818CB65826BF0" box="[1088,1220,1157,1181]" italics="true" pageId="9" pageNumber="230">tajikistanica</emphasis>
<bibRefCitation id="BE444B3ABB0FFFDE06C518CB641A6BF0" author="Chvala M." box="[1229,1372,1157,1181]" pageId="9" pageNumber="230" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
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<bibRefCitation id="BE444B3ABB0FFFDE015A18E462BD6BD0" author="Chvala M." box="[850,1019,1194,1213]" pageId="9" pageNumber="230" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999: 209</bibRefCitation>
(
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and
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), figs. 28, 29.
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locality:
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,
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,
<collectingMunicipality id="3A0EACB1BB0FFFDE061A188C65CC6BB8" box="[1042,1162,1218,1237]" pageId="9" pageNumber="230">Varzob River</collectingMunicipality>
,
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.
</paragraph>
</subSubSection>
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<paragraph id="DA6A36CBBB0FFFDE015A189464796A68" blockId="9.[812,1400,1157,1365]" pageId="9" pageNumber="230">
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<collectionCode id="BCC4AE0EBB0FFFDE015A189462876B80" box="[850,961,1242,1261]" pageId="9" pageNumber="230">MATERIAL</collectionCode>
EXAMINED.
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<collectingCountry id="A2C2765BBB0FFFDE0643189465F36B80" box="[1099,1205,1242,1261]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
:
</emphasis>
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[or Sangy-juman Pass],
<date id="AE6B100BBB0FFFDE01E818BC65706A68" box="[992,1078,1266,1285]" pageId="9" pageNumber="230" value="1869-05-25">
<collectingDate id="BE2FE9E3BB0FFFDE01E818BC65706A68" box="[992,1078,1266,1285]" pageId="9" pageNumber="230" value="1869-05-25">25.v.1869</collectingDate>
</date>
[
<collectorName id="7720531DBB0FFFDE064B18BC65896A68" box="[1091,1231,1266,1285]" pageId="9" pageNumber="230">A. Fedtschenko</collectorName>
] (
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,
<specimenCount id="CCD3FD42BB0FFFDE071D18BC64746A68" box="[1301,1330,1263,1285]" count="1" pageId="9" pageNumber="230" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDE015A194464316A57" pageId="9" pageNumber="230" type="discussion">
<paragraph id="DA6A36CBBB0FFFDE015A194464316A57" blockId="9.[812,1400,1157,1365]" pageId="9" pageNumber="230">
<materialsCitation id="6ABD3C96BB0FFFDE015A194464356A57" country="Tajikistan" location="Tajikistan" pageId="9" pageNumber="230" specimenCount="1">
REMARKS. This species was known only by the
<typeStatus id="056E8869BB0FFFDE0744194464306A72" box="[1356,1398,1290,1311]" pageId="9" pageNumber="230">type</typeStatus>
material taken from
<collectingCountry id="A2C2765BBB0FFFDE01E1196B650F6A56" box="[1001,1097,1317,1339]" name="Tajikistan" pageId="9" pageNumber="230">Tajikistan</collectingCountry>
and
<collectingCountry id="A2C2765BBB0FFFDE067B196B65996A56" box="[1139,1247,1317,1339]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
[
<bibRefCitation id="BE444B3ABB0FFFDE06E0196B642B6A57" author="Chvala M." box="[1256,1389,1317,1339]" pageId="9" pageNumber="230" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92CF6540BB0FFFDE015A197165AA6A38" box="[850,1260,1343,1365]" pageId="9" pageNumber="230" type="distribution">
<paragraph id="DA6A36CBBB0FFFDE015A197165AA6A38" blockId="9.[812,1400,1157,1365]" box="[850,1260,1343,1365]" pageId="9" pageNumber="230">
DISTRIBUTION.
<collectingCountry id="A2C2765BBB0FFFDE06011971652A6A38" box="[1033,1132,1343,1365]" name="Tajikistan" pageId="9" pageNumber="230">Tajikistan</collectingCountry>
,
<collectingCountry id="A2C2765BBB0FFFDE0670197165AE6A38" box="[1144,1256,1343,1365]" name="Uzbekistan" pageId="9" pageNumber="230">Uzbekistan</collectingCountry>
.
</paragraph>
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Genus
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<emphasis id="CB940B7CAE211547DECDF99EAF0DF94A" box="[962,1035,1658,1682]" italics="true" pageId="6" pageNumber="101">Delina</emphasis>
Robineau-Desvoidy, 1830
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<figureCitation id="61DBCBEBAE211547D927F97CAF70F968" box="[1064,1142,1688,1712]" captionStart="Figs 13–18" captionStartId="5.[227,262,1920,1939]" captionTargetBox="[269,1372,318,1903]" captionTargetPageId="5" captionText="Figs 13–18. Head of Scathophagidae, lateral view: 13 — Delina nigrita (Fallén); 14 — Hexamitocera loxocerata (Fallén); 15 — Leptopa filiformis (Zetterstedt); 16 — Megaphthalma pallida (Fallén); 17 — Micropselapha filiformis (Zetterstedt); 18 — Mixocordylura parva Ozerov. 14 — after Ozerov &amp; Krivosheina, 2015, fig. 3; 17 — after Ozerov, 2014, fig. 7; 18 — after Ozerov &amp; Krivosheina, 2012a, fig. 10. Рис. 13–18. Голова Scathophagidae, сбоку: 13 — Delina nigrita (Fallén); 14 — Hexamitocera loxocerata (Fallén); 15 — Leptopa filiformis (Zetterstedt); 16 — Megaphthalma pallida (Fallén); 17 — Micropselapha filiformis (Zetterstedt); 18 — Mixocordylura parva Ozerov. 14 — по Ozerov &amp; Krivosheina, 2015, fig. 3; 17 — по Ozerov, 2014, fig. 7; 18 — по Ozerov &amp; Krivosheina, 2012a, fig. 10." figureDoi="http://doi.org/10.5281/zenodo.13178430" httpUri="https://zenodo.org/record/13178430/files/figure.png" pageId="6" pageNumber="101">Fig. 13</figureCitation>
.
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REMARKS. One species of this genus has been recorded in
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—
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(
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)
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. Diagnosis of the genus and description of
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is given by
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Genus
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<emphasis id="CB940B7CAE211547DEFDF882AF63F8A5" box="[1010,1125,1894,1917]" italics="true" pageId="6" pageNumber="101">Ernoneura</emphasis>
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<figureCitation id="61DBCBEBAE211547DEB6F866AF0AF842" box="[953,1036,1922,1946]" captionStart="Figs 28–32" captionStartId="8.[226,263,1870,1889]" captionTargetBox="[505,1061,1498,1793]" captionTargetId="figure-23@8.[486,1073,1516,1793]" captionTargetPageId="8" captionText="Figs 28–32. Fragments of thorax of Scathophagidae: 28 — Scathophaga furcata (Say), anepisternum and katepisternum; 29 — Ernoneura argus (Zetterstedt), anepisternum and katepisternum; 30 — Scathophaga stercoraria (Linnaeus), postmetacoxal area; 31 — Scatomyza scybalaria (Linnaeus), postmetacoxal area; 32 — Gimnomera hirta (Hendel), anterior part of scutum, dorsal view. 30, 31 — after Ozerov &amp; Krivosheina, 2011b, figs 1, 2; 32 — after Ozerov, 2019a, fig. 2. Рис. 28–32. Фрагменты груди Scathophagidae: 28 — Scathophaga furcata (Say), анЭпистерн и катЭпистерн; 29 — Ernoneura argus (Zetterstedt), анЭпистерн и катЭпистерн; 30 — Scathophaga stercoraria (Linnaeus), постметакоксальнаЯ область; 31 — Scatomyza scybalaria (Linnaeus), постметакоксальнаЯ область; 11 — Gimnomera hirta (Hendel), переднЯЯ часть среднеспинки, сверху. 30, 31 — по Ozerov &amp; Krivosheina, 2011b, figs 1, 2; 32 — по Ozerov, 2019a, fig. 2." figureDoi="http://doi.org/10.5281/zenodo.13178440" httpUri="https://zenodo.org/record/13178440/files/figure.png" pageId="6" pageNumber="101">Figs 29</figureCitation>
,
<figureCitation id="61DBCBEBAE211547D918F866AF32F842" box="[1047,1076,1922,1946]" captionStart="Figs 50–58" captionStartId="12.[228,266,1822,1841]" captionTargetBox="[200,768,1098,1764]" captionTargetId="figure-21@12.[200,778,1360,1753]" captionTargetPageId="12" captionText="Figs 50–58. Lower and upper calypters of wing (50) and wings (51–58) of Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — after Ozerov, 2016, figs 7, 8; 55 — after Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — after Ozerov, 2017, fig. 9. Рис. 50–58. Закрыловые чеШуйки крыла (50) и крыльЯ (51–58) Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — по Ozerov, 2016, figs 7, 8; 55 — по Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — по Ozerov, 2017, fig. 9." figureDoi="http://doi.org/10.5281/zenodo.13178448" httpUri="https://zenodo.org/record/13178448/files/figure.png" pageId="6" pageNumber="101">53</figureCitation>
,
<figureCitation id="61DBCBEBAE211547D94FF866AF5BF842" box="[1088,1117,1922,1946]" captionStart="Figs 61–67" captionStartId="14.[733,770,1821,1840]" captionTargetBox="[211,1399,894,2060]" captionTargetId="figure-18@14.[619,1003,952,1545]" captionTargetPageId="14" captionText="Figs 61–67. Male sternites 4 (lower) and 5 (upper) of Scathophagidae: 61 — Ernoneura argus (Zetterstedt); 62 — Hydromyza livens (Fabricius); 63 — Leptopa filiformis Zetterstedt; 64 — Megaphthalma pallida (Fallén, 1819); 65 — Megaphthalma unilineata (Zetterstedt); 66 — Staegeria kunzei (Zetterstedt); 67 — Scatomyza magnipennis (Portschinsky). Рис. 61–67. Стерниты 4 (вниЗу) и 5 (вверху) самцов Scathophagidae: 61 — Ernoneura argus (Zetterstedt); 62 — Hydromyza livens (Fabricius); 63 — Leptopa filiformis Zetterstedt; 64 — Megaphthalma pallida (Fallén); 65 — Megaphthalma unilineata (Zetterstedt); 66 — Staegeria kunzei (Zetterstedt); 67 — Scatomyza magnipennis (Portschinsky)." figureDoi="http://doi.org/10.5281/zenodo.13178452" httpUri="https://zenodo.org/record/13178452/files/figure.png" pageId="6" pageNumber="101">61</figureCitation>
,
<figureCitation id="61DBCBEBAE211547D966F866AFA9F842" box="[1129,1199,1922,1946]" captionStart="89 90 91 92" captionStartId="16.[231,294,1805,1862]" captionText="89 90 91 92 Figs 77–92. Epandrium, cerci and surstyli of Scathophagidae, lateral view (77, 79, 81, 83, 85, 87, 89, 91) and dorsal view (78, 80, 82, 84, 86, 88, 90, 92): 77, 78 — Cosmetopus dentimanus (Zetterstedt); 79, 80 — Cosmetopus longus (Walker); 81, 82 — Ernoneura argus (Zetterstedt); 83, 84 — Hydromyza livens (Fabricius); 85, 86 — Leptopa filiformis Zetterstedt; 87, 88 — Megaphthalma pallida (Fallén); 89, 90 — Megaphthalma unilineata (Zetterstedt); 91, 92 — Staegeria kunzei (Zetterstedt). Рис. 1–7. Эпандрий, церки и сурстили Scathophagidae, сбоку (77, 79, 81, 83, 85, 87, 89, 91) и сверху (78, 80, 82, 84, 86, 88, 90, 92): 77, 78 — Cosmetopus dentimanus (Zetterstedt); 79, 80 — Cosmetopus longus (Walker); 81, 82 — Ernoneura argus (Zetterstedt); 83, 84 — Hydromyza livens (Fabricius); 85, 86 — Leptopa filiformis Zetterstedt; 87, 88 — Megaphthalma pallida (Fallén); 89, 90 — Megaphthalma unilineata (Zetterstedt); 91, 92 — Staegeria kunzei (Zetterstedt)." pageId="6" pageNumber="101">81, 82</figureCitation>
,
<figureCitation id="61DBCBEBAE211547D9B4F866AFE0F842" box="[1211,1254,1922,1946]" captionStart="Figs 112–117" captionStartId="21.[229,266,1918,1937]" captionTargetBox="[195,1404,315,1901]" captionTargetPageId="21" captionText="Figs 112–117. Habitus of Scathophagidae: 112 — Neochirosia veratri (Hendel); 113 — Ernoneura argus (Zetterstedt); 114 — Gimnomera tarsea (Fallén); 115 — Hydromyza livens (Fabricius); 116 — Leptopa filiformis Zetterstedt; 117 — Megaphthalma pallida (Fallén). 114 — after Ozerov, 2019a, fig.1. Рис. 112–117. ОбЩий вид Scathophagidae: 112 — Neochirosia veratri (Hendel); 113 — Ernoneura argus (Zetterstedt); 114 — Gimnomera tarsea (Fallén); 115 — Hydromyza livens (Fabricius); 116 — Leptopa filiformis Zetterstedt; 117 — Megaphthalma pallida (Fallén). 114 — по Ozerov, 2019a, fig.1." figureDoi="http://doi.org/10.5281/zenodo.13178460" httpUri="https://zenodo.org/record/13178460/files/figure.png" pageId="6" pageNumber="101">113</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE211548DE5DF859A9DAF8E6" lastPageId="9" lastPageNumber="104" pageId="6" pageNumber="101" type="diagnosis">
<paragraph id="F95FD76EAE211549DE5DF859AFBEFA3D" blockId="6.[812,1399,1981,2064]" lastBlockId="8.[735,1208,1473,1509]" lastPageId="8" lastPageNumber="103" pageId="6" pageNumber="101">
DIAGNOSIS. Medium-sized flies (
<quantity id="3E187A8BAE211547D9DCF859AE29F80D" box="[1235,1327,1981,2005]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="101" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
long). Proepisternum is covered with hairs at middle or in anterior part. Anepisternum covered with hairs comscapular setae notopleural spinules
</paragraph>
<caption id="AD9F87E6AE201546DDEDF909A89AF7D4" ID-DOI="http://doi.org/10.5281/zenodo.13178438" ID-Zenodo-Dep="13178438" httpUri="https://zenodo.org/record/13178438/files/figure.png" pageId="7" pageNumber="102" startId="7.[226,263,1773,1792]" targetBox="[190,1399,317,1732]" targetPageId="7" targetType="figure">
<paragraph id="F95FD76EAE201546DDEDF909A84FF8A1" blockId="7.[188,1398,1773,2061]" pageId="7" pageNumber="102">
Figs 22–27. Fragments of thorax of
<taxonomicName id="3EE0ACEDAE201546DF11F909A9ACF8D8" box="[542,682,1773,1792]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 22 —
<taxonomicName id="3EE0ACEDAE201546DFE5F909A8EBF8D8" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[746,1005,1773,1792]" class="Insecta" family="Scathophagidae" genus="Hydromyza" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="livens">
<emphasis id="CB940B7CAE201546DFE5F909A883F8D8" box="[746,901,1773,1792]" italics="true" pageId="7" pageNumber="102">Hydromyza livens</emphasis>
(Fabricius)
</taxonomicName>
, anepisternum; 23 —
<taxonomicName id="3EE0ACEDAE201546D9A1F909AA16F8C0" authority="(Meigen)" baseAuthorityName="Meigen" baseAuthorityYear="1826" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="CB940B7CAE201546D9A1F909AE70F8D9" box="[1198,1398,1773,1793]" italics="true" pageId="7" pageNumber="102">Trichopalpus fraternus</emphasis>
(Meigen)
</taxonomicName>
, anepisternum; 24 —
<taxonomicName id="3EE0ACEDAE201546DCE8F8E1A9F7F8C0" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[487,753,1797,1816]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="litorea">
<emphasis id="CB940B7CAE201546DCE8F8E1A998F8C0" box="[487,670,1797,1816]" italics="true" pageId="7" pageNumber="102">Scathophaga litorea</emphasis>
(Fallén)
</taxonomicName>
, anepisternum; 25 —
<taxonomicName id="3EE0ACEDAE201546DEC6F8E1AFC1F8C0" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[969,1223,1797,1816]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="CB940B7CAE201546DEC6F8E1AF72F8C0" box="[969,1140,1797,1816]" italics="true" pageId="7" pageNumber="102">Cleigastra flavipes</emphasis>
(Fallén)
</taxonomicName>
, anepisternum and katepisternum; 26 —
<taxonomicName id="3EE0ACEDAE201546DC8EF8F9A97EF8E8" authority="(Becker)" baseAuthorityName="Becker" baseAuthorityYear="1907" box="[385,632,1821,1840]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="arctica">
<emphasis id="CB940B7CAE201546DC8EF8F9A927F8E8" box="[385,545,1821,1840]" italics="true" pageId="7" pageNumber="102">Cleigastra arctica</emphasis>
(Becker)
</taxonomicName>
, anepisternum and katepisternum; 27 —
<taxonomicName id="3EE0ACEDAE201546DEE3F8F9AE13F8E8" authority="(Meigen)" authorityName="" baseAuthorityName="Meigen" baseAuthorityYear="1826" box="[1004,1301,1821,1840]" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="punctipes">
<emphasis id="CB940B7CAE201546DEE3F8F9AFBDF8E8" box="[1004,1211,1821,1840]" italics="true" pageId="7" pageNumber="102">Trichopalpus punctipes</emphasis>
(Meigen)
</taxonomicName>
, anepisternum and katepisternum. Abbreviations: a kepst s — anterior katepisternal seta; anepst — anepisternum; kepst — katepisternum; lp kepst s — lower posterior katepisternal seta; prepst — proepisternum; up kepst s — upper posterior katepisternal seta. 24 — after
<bibRefCitation id="9D71AA9FAE201546D811F8AAAA59F8A1" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="7" pageNumber="102" pagination="201 - 246" refId="ref16309" refString="Ozerov A. L., Krivosheina M. G. 2021. A review of the genus Scathophaga Meigen, 1803 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 30. No. 2. P. 201 - 246." type="journal article" year="2021">Ozerov &amp; Krivosheina, 2021</bibRefCitation>
, fig. 6; 25 — after
<bibRefCitation id="9D71AA9FAE201546DF04F882A80CF8A0" author="Ozerov A. L. &amp; Krivosheina M. G." box="[523,778,1893,1913]" pageId="7" pageNumber="102" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 1.
</paragraph>
<paragraph id="F95FD76EAE201546DDEDF866A89AF7D4" blockId="7.[188,1398,1773,2061]" pageId="7" pageNumber="102">
Рис. 22–27. Фрагменты груди
<taxonomicName id="3EE0ACEDAE201546DF06F865A995F84C" box="[521,659,1921,1940]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 22 —
<taxonomicName id="3EE0ACEDAE201546DFD0F865A8E1F84C" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[735,999,1921,1940]" class="Insecta" family="Scathophagidae" genus="Hydromyza" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="livens">
<emphasis id="CB940B7CAE201546DFD0F865A87BF84C" box="[735,893,1921,1940]" italics="true" pageId="7" pageNumber="102">Hydromyza livens</emphasis>
(Fabricius)
</taxonomicName>
, анЭпистерн; 23 —
<taxonomicName id="3EE0ACEDAE201546D9A2F865AA08F874" authority="(Meigen)" baseAuthorityName="Meigen" baseAuthorityYear="1826" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="CB940B7CAE201546D9A2F865AE70F84C" box="[1197,1398,1921,1940]" italics="true" pageId="7" pageNumber="102">Trichopalpus fraternus</emphasis>
(Meigen)
</taxonomicName>
, анЭпистерн; 24 —
<taxonomicName id="3EE0ACEDAE201546DCC8F87DA9C0F874" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[455,710,1945,1964]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="litorea">
<emphasis id="CB940B7CAE201546DCC8F87DA97FF874" box="[455,633,1945,1964]" italics="true" pageId="7" pageNumber="102">Scathophaga litorea</emphasis>
(Fallén)
</taxonomicName>
, анЭпистерн; 25 —
<taxonomicName id="3EE0ACEDAE201546DE8FF87DAF74F874" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[896,1138,1945,1964]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="CB940B7CAE201546DE8FF87DAF23F874" box="[896,1061,1945,1964]" italics="true" pageId="7" pageNumber="102">Cleigastra flavipes</emphasis>
(Fallén)
</taxonomicName>
, анЭпистерн и катЭпистерн; 26 —
<taxonomicName id="3EE0ACEDAE201546DDF3F855AAF4F81C" authority="(Becker)" baseAuthorityName="Becker" baseAuthorityYear="1907" box="[252,498,1969,1988]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="arctica">
<emphasis id="CB940B7CAE201546DDF3F855AA9AF81C" box="[252,412,1969,1988]" italics="true" pageId="7" pageNumber="102">Cleigastra arctica</emphasis>
(Becker)
</taxonomicName>
, анЭпистерн и катЭпистерн; 27 —
<taxonomicName id="3EE0ACEDAE201546DE49F855AF6BF81C" authority="(Meigen)" authorityName="" baseAuthorityName="Meigen" baseAuthorityYear="1826" box="[838,1133,1969,1988]" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="102" phylum="Arthropoda" rank="species" species="punctipes">
<emphasis id="CB940B7CAE201546DE49F855AF14F81C" box="[838,1042,1969,1988]" italics="true" pageId="7" pageNumber="102">Trichopalpus punctipes</emphasis>
(Meigen)
</taxonomicName>
, анЭпистерн и катЭпистерн. ОбоЗначениЯ: a kepst s — переднЯЯ катЭпистернальнаЯ Щетинка; anepst — анЭпистернум; kepst — катЭпистернум; lp kepst s — нижнЯЯ ЗаднЯЯ катЭпистернальнаЯ Щетинка; prepst — проЭпистернум; up kepst s — верхнЯЯ ЗаднЯЯ катЭпистернальнаЯ Щетинка. 24 — по
<bibRefCitation id="9D71AA9FAE201546DDD7F81DAAD4F7D4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[216,466,2041,2060]" pageId="7" pageNumber="102" pagination="201 - 246" refId="ref16309" refString="Ozerov A. L., Krivosheina M. G. 2021. A review of the genus Scathophaga Meigen, 1803 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 30. No. 2. P. 201 - 246." type="journal article" year="2021">Ozerov &amp; Krivosheina, 2021</bibRefCitation>
, fig. 6; 25 — по
<bibRefCitation id="9D71AA9FAE201546DF68F81DA867F7D4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[615,865,2041,2060]" pageId="7" pageNumber="102" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 1.
</paragraph>
</caption>
<paragraph id="F95FD76EAE2F1549DCA5F935AAEFF8D2" blockId="8.[426,489,1745,1802]" box="[426,489,1745,1802]" pageId="8" pageNumber="103">32</paragraph>
<caption id="AD9F87E6AE2F1549DDEDF8AAA8FFF7D4" ID-DOI="http://doi.org/10.5281/zenodo.13178440" ID-Zenodo-Dep="13178440" httpUri="https://zenodo.org/record/13178440/files/figure.png" pageId="8" pageNumber="103" startId="8.[226,263,1870,1889]" targetBox="[505,1061,1498,1793]" targetPageId="8" targetType="figure">
<paragraph id="F95FD76EAE2F1549DDEDF8AAA87CF871" blockId="8.[189,1398,1870,2061]" pageId="8" pageNumber="103">
Figs 28–32. Fragments of thorax of
<taxonomicName id="3EE0ACEDAE2F1549DF39F8AAA9C4F8B9" box="[566,706,1870,1889]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 28 —
<taxonomicName id="3EE0ACEDAE2F1549DE01F8AAAF07F8B9" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[782,1025,1870,1889]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="furcata">
<emphasis id="CB940B7CAE2F1549DE01F8AAA8C0F8B9" box="[782,966,1870,1889]" italics="true" pageId="8" pageNumber="103">Scathophaga furcata</emphasis>
(Say)
</taxonomicName>
, anepisternum and katepisternum; 29 —
<taxonomicName id="3EE0ACEDAE2F1549DDB2F882AACFF8A1" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[189,457,1894,1913]" class="Insecta" family="Scathophagidae" genus="Ernoneura" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="CB940B7CAE2F1549DDB2F882AA55F8A0" box="[189,339,1894,1912]" italics="true" pageId="8" pageNumber="103">Ernoneura argus</emphasis>
(Zetterstedt)
</taxonomicName>
, anepisternum and katepisternum; 30 —
<taxonomicName id="3EE0ACEDAE2F1549DE31F882AF79F8A1" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[830,1151,1894,1913]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="stercoraria">
<emphasis id="CB940B7CAE2F1549DE31F882AF11F8A0" box="[830,1047,1894,1913]" italics="true" pageId="8" pageNumber="103">Scathophaga stercoraria</emphasis>
(Linnaeus)
</taxonomicName>
, postmetacoxal area; 31 —
<taxonomicName id="3EE0ACEDAE2F1549DDB2F89AAAE4F849" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,482,1918,1937]" class="Insecta" family="Scathophagidae" genus="Scatomyza" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="scybalaria">
<emphasis id="CB940B7CAE2F1549DDB2F89AAA7DF849" box="[189,379,1918,1937]" italics="true" pageId="8" pageNumber="103">Scatomyza scybalaria</emphasis>
(Linnaeus)
</taxonomicName>
, postmetacoxal area; 32 —
<taxonomicName id="3EE0ACEDAE2F1549DFD3F89AA8CDF849" authority="(Hendel)" baseAuthorityName="Hendel" box="[732,971,1918,1937]" class="Insecta" family="Scathophagidae" genus="Gimnomera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="hirta">
<emphasis id="CB940B7CAE2F1549DFD3F89AA872F849" box="[732,884,1918,1937]" italics="true" pageId="8" pageNumber="103">Gimnomera hirta</emphasis>
(Hendel)
</taxonomicName>
, anterior part of scutum, dorsal view. 30, 31 — after
<bibRefCitation id="9D71AA9FAE2F1549DDE4F872AAF3F871" author="Ozerov A. L. &amp; Krivosheina M. G." box="[235,501,1942,1961]" pageId="8" pageNumber="103" pagination="1 - 10" refId="ref15738" refString="Ozerov A. L., Krivosheina M. G. 2011 b. To the knowledge of dung flies (Diptera, Scathophagidae) of Vietnam, with the description of a new species // Far Eastern Entomologist. No. 228. P. 1 - 10." type="book chapter" year="2011">Ozerov &amp; Krivosheina, 2011b</bibRefCitation>
, figs 1, 2; 32 — after
<bibRefCitation id="9D71AA9FAE2F1549DFB5F872A83DF871" author="Ozerov A. L." box="[698,827,1942,1961]" pageId="8" pageNumber="103" pagination="195 - 218" refId="ref15523" refString="Ozerov A. L. 2019 a. A review of the genus Gimnomera Rondani, 1866 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 28. No. 2. P. 195 - 218." type="journal article" year="2019">Ozerov, 2019a</bibRefCitation>
, fig. 2.
</paragraph>
<paragraph id="F95FD76EAE2F1549DDEDF856A8FFF7D4" blockId="8.[189,1398,1870,2061]" pageId="8" pageNumber="103">
Рис. 28–32. Фрагменты груди
<taxonomicName id="3EE0ACEDAE2F1549DF0AF855A997F81C" box="[517,657,1969,1988]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 28 —
<taxonomicName id="3EE0ACEDAE2F1549DFD6F855A8CDF81C" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[729,971,1969,1988]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="furcata">
<emphasis id="CB940B7CAE2F1549DFD6F855A896F81C" box="[729,912,1969,1988]" italics="true" pageId="8" pageNumber="103">Scathophaga furcata</emphasis>
(Say)
</taxonomicName>
, анЭпистерн и катЭпистерн; 29 —
<taxonomicName id="3EE0ACEDAE2F1549D816F856AA6EF804" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" class="Insecta" family="Scathophagidae" genus="Ernoneura" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="CB940B7CAE2F1549D816F856ABE8F804" italics="true" pageId="8" pageNumber="103">Ernoneura argus</emphasis>
(Zetterstedt)
</taxonomicName>
, анЭпистерн и катЭпистерн; 30 —
<taxonomicName id="3EE0ACEDAE2F1549DFC8F82DAF17F804" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[711,1041,1993,2012]" class="Insecta" family="Scathophagidae" genus="Scathophaga" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="stercoraria">
<emphasis id="CB940B7CAE2F1549DFC8F82DA8A3F804" box="[711,933,1993,2012]" italics="true" pageId="8" pageNumber="103">Scathophaga stercoraria</emphasis>
(Linnaeus)
</taxonomicName>
, постметакоксальнаЯ область; 31 —
<taxonomicName id="3EE0ACEDAE2F1549DDB2F806AAEFF82C" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,489,2017,2037]" class="Insecta" family="Scathophagidae" genus="Scatomyza" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="scybalaria">
<emphasis id="CB940B7CAE2F1549DDB2F806AA78F82C" box="[189,382,2017,2037]" italics="true" pageId="8" pageNumber="103">Scatomyza scybalaria</emphasis>
(Linnaeus)
</taxonomicName>
, постметакоксальнаЯ область; 11 —
<taxonomicName id="3EE0ACEDAE2F1549DE5CF806AF4EF82C" authority="(Hendel)" baseAuthorityName="Hendel" box="[851,1096,2017,2037]" class="Insecta" family="Scathophagidae" genus="Gimnomera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="103" phylum="Arthropoda" rank="species" species="hirta">
<emphasis id="CB940B7CAE2F1549DE5CF806A8E9F82C" box="[851,1007,2017,2037]" italics="true" pageId="8" pageNumber="103">Gimnomera hirta</emphasis>
(Hendel)
</taxonomicName>
, переднЯЯ часть среднеспинки, сверху. 30, 31 — по
<bibRefCitation id="9D71AA9FAE2F1549DC73F81DA980F7D4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[380,646,2041,2060]" pageId="8" pageNumber="103" pagination="1 - 10" refId="ref15738" refString="Ozerov A. L., Krivosheina M. G. 2011 b. To the knowledge of dung flies (Diptera, Scathophagidae) of Vietnam, with the description of a new species // Far Eastern Entomologist. No. 228. P. 1 - 10." type="book chapter" year="2011">Ozerov &amp; Krivosheina, 2011b</bibRefCitation>
, figs 1, 2; 32 — по
<bibRefCitation id="9D71AA9FAE2F1549DE36F81DA8BCF7D4" author="Ozerov A. L." box="[825,954,2041,2060]" pageId="8" pageNumber="103" pagination="195 - 218" refId="ref15523" refString="Ozerov A. L. 2019 a. A review of the genus Gimnomera Rondani, 1866 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 28. No. 2. P. 195 - 218." type="journal article" year="2019">Ozerov, 2019a</bibRefCitation>
, fig. 2.
</paragraph>
</caption>
<paragraph id="F95FD76EAE2E1548D927FA97AF61FA74" blockId="9.[1064,1127,1395,1452]" box="[1064,1127,1395,1452]" pageId="9" pageNumber="104">37</paragraph>
<caption id="AD9F87E6AE2E1548DDEBFA0EAFABF9A0" ID-DOI="http://doi.org/10.5281/zenodo.13178444" ID-Zenodo-Dep="13178444" httpUri="https://zenodo.org/record/13178444/files/figure.png" pageId="9" pageNumber="104" startId="9.[228,265,1514,1533]" targetBox="[202,1400,314,1422]" targetPageId="9" targetType="figure">
<paragraph id="F95FD76EAE2E1548DDEBFA0EAE7EF9F5" blockId="9.[190,1400,1513,1657]" pageId="9" pageNumber="104">
Figs 33–37. Scutellum of
<taxonomicName id="3EE0ACEDAE2E1548DCC2FA0EA95EFA25" box="[461,600,1514,1533]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, dorsal view: 33 —
<taxonomicName id="3EE0ACEDAE2E1548DE1CFA0EA8F8FA25" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[787,1022,1514,1533]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="CB940B7CAE2E1548DE1CFA0EA8A9FA25" box="[787,943,1514,1533]" italics="true" pageId="9" pageNumber="104">Cordilura albipes</emphasis>
(Fallén)
</taxonomicName>
; 34 —
<taxonomicName id="3EE0ACEDAE2E1548D94BFA0EAE3FFA25" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[1092,1337,1514,1533]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="CB940B7CAE2E1548D94BFA0EAFECFA25" box="[1092,1258,1514,1533]" italics="true" pageId="9" pageNumber="104">Cleigastra flavipes</emphasis>
(Fallén)
</taxonomicName>
; 35 —
<taxonomicName id="3EE0ACEDAE2E1548DDB1F9E6AAB1F9CD" authority="(Loew)" baseAuthorityName="Loew" box="[190,439,1538,1557]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="CB940B7CAE2E1548DDB1F9E6AA6FF9CC" box="[190,361,1538,1557]" italics="true" pageId="9" pageNumber="104">Cleigastra cornuta</emphasis>
(Loew)
</taxonomicName>
; 36 —
<taxonomicName id="3EE0ACEDAE2E1548DF08F9E6A83DF9CD" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[519,827,1538,1557]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="spinimanum">
<emphasis id="CB940B7CAE2E1548DF08F9E6A9E1F9CC" box="[519,743,1538,1557]" italics="true" pageId="9" pageNumber="104">Norellisoma spinimanum</emphasis>
(Fallén)
</taxonomicName>
; 37 —
<taxonomicName id="3EE0ACEDAE2E1548DE84F9E6AF9FF9CD" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[907,1177,1538,1557]" class="Insecta" family="Scathophagidae" genus="Norellia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="tipularia">
<emphasis id="CB940B7CAE2E1548DE84F9E6AF2DF9CD" box="[907,1067,1538,1557]" italics="true" pageId="9" pageNumber="104">Norellia tipularia</emphasis>
(Fabricius)
</taxonomicName>
. 33 — after
<bibRefCitation id="9D71AA9FAE2E1548D813F9E6AA6DF9F5" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="9" pageNumber="104" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, fig. 12; 34, 35 — after
<bibRefCitation id="9D71AA9FAE2E1548DF32F9FEA83CF9F4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[573,826,1561,1581]" pageId="9" pageNumber="104" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, figs 7, 8; 36, 37 — after
<bibRefCitation id="9D71AA9FAE2E1548D914F9FEAE25F9F5" author="Ozerov A. L. &amp; Krivosheina M. G." box="[1051,1315,1562,1581]" pageId="9" pageNumber="104" pagination="101 - 103" refId="ref15684" refString="Ozerov A. L., Krivosheina M. G. 2011 a. The first record of Norellia tipularia (Fabricius, 1794) (Diptera, Scathophagidae) for the fauna of Russia // Russian Entomological Journal. Vol. 20. No. 1. P. 101 - 103." type="book chapter" year="2011">Ozerov &amp; Krivosheina, 2011a</bibRefCitation>
, figs 4, 3.
</paragraph>
<paragraph id="F95FD76EAE2E1548DDEBF9D2AFABF9A0" blockId="9.[190,1400,1513,1657]" pageId="9" pageNumber="104">
Рис. 33–37. Щиток
<taxonomicName id="3EE0ACEDAE2E1548DC94F9D1A920F990" box="[411,550,1589,1608]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, сверху: 33 —
<taxonomicName id="3EE0ACEDAE2E1548DFBAF9D1A8A6F990" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[693,928,1589,1608]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="CB940B7CAE2E1548DFBAF9D1A857F990" box="[693,849,1589,1608]" italics="true" pageId="9" pageNumber="104">Cordilura albipes</emphasis>
(Fallén)
</taxonomicName>
; 34 —
<taxonomicName id="3EE0ACEDAE2E1548DEEBF9D1AFDEF990" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[996,1240,1589,1608]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="CB940B7CAE2E1548DEEBF9D1AF8CF990" box="[996,1162,1589,1608]" italics="true" pageId="9" pageNumber="104">Cleigastra flavipes</emphasis>
(Fallén)
</taxonomicName>
; 35 —
<taxonomicName id="3EE0ACEDAE2E1548D812F9D1AA4CF9B8" authority="(Loew)" baseAuthorityName="Loew" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="CB940B7CAE2E1548D812F9D1AA07F9B8" italics="true" pageId="9" pageNumber="104">Cleigastra cornuta</emphasis>
(Loew)
</taxonomicName>
; 36 —
<taxonomicName id="3EE0ACEDAE2E1548DC9FF9A9A9BCF9B8" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[400,698,1613,1632]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="spinimanum">
<emphasis id="CB940B7CAE2E1548DC9FF9A9A96DF9B8" box="[400,619,1613,1632]" italics="true" pageId="9" pageNumber="104">Norellisoma spinimanum</emphasis>
(Fallén)
</taxonomicName>
; 37 —
<taxonomicName id="3EE0ACEDAE2E1548DFF0F9A9AF02F9B8" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[767,1028,1613,1632]" class="Insecta" family="Scathophagidae" genus="Norellia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="tipularia">
<emphasis id="CB940B7CAE2E1548DFF0F9A9A89CF9B8" box="[767,922,1613,1632]" italics="true" pageId="9" pageNumber="104">Norellia tipularia</emphasis>
(Fabricius)
</taxonomicName>
. 33 — по
<bibRefCitation id="9D71AA9FAE2E1548D969F9A9AE73F9B8" author="Ozerov A. L. &amp; Krivosheina M. G." box="[1126,1397,1613,1632]" pageId="9" pageNumber="104" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, fig. 12; 34, 35 — по
<bibRefCitation id="9D71AA9FAE2E1548DC76F981A97FF9A0" author="Ozerov A. L. &amp; Krivosheina M. G." box="[377,633,1637,1656]" pageId="9" pageNumber="104" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, figs 7, 8; 36, 37 — по
<bibRefCitation id="9D71AA9FAE2E1548DE42F981AF50F9A0" author="Ozerov A. L. &amp; Krivosheina M. G." box="[845,1110,1637,1656]" pageId="9" pageNumber="104" pagination="101 - 103" refId="ref15684" refString="Ozerov A. L., Krivosheina M. G. 2011 a. The first record of Norellia tipularia (Fabricius, 1794) (Diptera, Scathophagidae) for the fauna of Russia // Russian Entomological Journal. Vol. 20. No. 1. P. 101 - 103." type="book chapter" year="2011">Ozerov &amp; Krivosheina, 2011a</bibRefCitation>
, figs 4, 3.
</paragraph>
</caption>
<paragraph id="F95FD76EAE2E1548DDB2F94CA9DAF8E6" blockId="9.[189,776,1704,2062]" pageId="9" pageNumber="104">
pletely or almost completely, with hairs posterior to anterior spiracle. Katepisternum without strong setae (
<figureCitation id="61DBCBEBAE2E1548DDCBF903AA1CF927" box="[196,282,1767,1791]" captionStart="Figs 28–32" captionStartId="8.[226,263,1870,1889]" captionTargetBox="[505,1061,1498,1793]" captionTargetId="figure-23@8.[486,1073,1516,1793]" captionTargetPageId="8" captionText="Figs 28–32. Fragments of thorax of Scathophagidae: 28 — Scathophaga furcata (Say), anepisternum and katepisternum; 29 — Ernoneura argus (Zetterstedt), anepisternum and katepisternum; 30 — Scathophaga stercoraria (Linnaeus), postmetacoxal area; 31 — Scatomyza scybalaria (Linnaeus), postmetacoxal area; 32 — Gimnomera hirta (Hendel), anterior part of scutum, dorsal view. 30, 31 — after Ozerov &amp; Krivosheina, 2011b, figs 1, 2; 32 — after Ozerov, 2019a, fig. 2. Рис. 28–32. Фрагменты груди Scathophagidae: 28 — Scathophaga furcata (Say), анЭпистерн и катЭпистерн; 29 — Ernoneura argus (Zetterstedt), анЭпистерн и катЭпистерн; 30 — Scathophaga stercoraria (Linnaeus), постметакоксальнаЯ область; 31 — Scatomyza scybalaria (Linnaeus), постметакоксальнаЯ область; 11 — Gimnomera hirta (Hendel), переднЯЯ часть среднеспинки, сверху. 30, 31 — по Ozerov &amp; Krivosheina, 2011b, figs 1, 2; 32 — по Ozerov, 2019a, fig. 2." figureDoi="http://doi.org/10.5281/zenodo.13178440" httpUri="https://zenodo.org/record/13178440/files/figure.png" pageId="9" pageNumber="104">Fig. 29</figureCitation>
). Postmetacoxal bridge absent. Wing membrane with dark spots (
<figureCitation id="61DBCBEBAE2E1548DCBEF8E2A907F8C6" box="[433,513,1798,1822]" captionStart="Figs 50–58" captionStartId="12.[228,266,1822,1841]" captionTargetBox="[200,768,1098,1764]" captionTargetId="figure-21@12.[200,778,1360,1753]" captionTargetPageId="12" captionText="Figs 50–58. Lower and upper calypters of wing (50) and wings (51–58) of Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — after Ozerov, 2016, figs 7, 8; 55 — after Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — after Ozerov, 2017, fig. 9. Рис. 50–58. Закрыловые чеШуйки крыла (50) и крыльЯ (51–58) Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — по Ozerov, 2016, figs 7, 8; 55 — по Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — по Ozerov, 2017, fig. 9." figureDoi="http://doi.org/10.5281/zenodo.13178448" httpUri="https://zenodo.org/record/13178448/files/figure.png" pageId="9" pageNumber="104">Fig. 53</figureCitation>
). Male sternites 4 and 5, epandrium, cerci and surstyli as in
<figureCitation id="61DBCBEBAE2E1548DF3BF8C1A980F8E5" box="[564,646,1829,1853]" captionStart="Figs 61–67" captionStartId="14.[733,770,1821,1840]" captionTargetBox="[211,1399,894,2060]" captionTargetId="figure-18@14.[619,1003,952,1545]" captionTargetPageId="14" captionText="Figs 61–67. Male sternites 4 (lower) and 5 (upper) of Scathophagidae: 61 — Ernoneura argus (Zetterstedt); 62 — Hydromyza livens (Fabricius); 63 — Leptopa filiformis Zetterstedt; 64 — Megaphthalma pallida (Fallén, 1819); 65 — Megaphthalma unilineata (Zetterstedt); 66 — Staegeria kunzei (Zetterstedt); 67 — Scatomyza magnipennis (Portschinsky). Рис. 61–67. Стерниты 4 (вниЗу) и 5 (вверху) самцов Scathophagidae: 61 — Ernoneura argus (Zetterstedt); 62 — Hydromyza livens (Fabricius); 63 — Leptopa filiformis Zetterstedt; 64 — Megaphthalma pallida (Fallén); 65 — Megaphthalma unilineata (Zetterstedt); 66 — Staegeria kunzei (Zetterstedt); 67 — Scatomyza magnipennis (Portschinsky)." figureDoi="http://doi.org/10.5281/zenodo.13178452" httpUri="https://zenodo.org/record/13178452/files/figure.png" pageId="9" pageNumber="104">Figs 61</figureCitation>
,
<figureCitation id="61DBCBEBAE2E1548DF9DF8C2A9DEF8E6" box="[658,728,1830,1854]" captionStart="89 90 91 92" captionStartId="16.[231,294,1805,1862]" captionText="89 90 91 92 Figs 77–92. Epandrium, cerci and surstyli of Scathophagidae, lateral view (77, 79, 81, 83, 85, 87, 89, 91) and dorsal view (78, 80, 82, 84, 86, 88, 90, 92): 77, 78 — Cosmetopus dentimanus (Zetterstedt); 79, 80 — Cosmetopus longus (Walker); 81, 82 — Ernoneura argus (Zetterstedt); 83, 84 — Hydromyza livens (Fabricius); 85, 86 — Leptopa filiformis Zetterstedt; 87, 88 — Megaphthalma pallida (Fallén); 89, 90 — Megaphthalma unilineata (Zetterstedt); 91, 92 — Staegeria kunzei (Zetterstedt). Рис. 1–7. Эпандрий, церки и сурстили Scathophagidae, сбоку (77, 79, 81, 83, 85, 87, 89, 91) и сверху (78, 80, 82, 84, 86, 88, 90, 92): 77, 78 — Cosmetopus dentimanus (Zetterstedt); 79, 80 — Cosmetopus longus (Walker); 81, 82 — Ernoneura argus (Zetterstedt); 83, 84 — Hydromyza livens (Fabricius); 85, 86 — Leptopa filiformis Zetterstedt; 87, 88 — Megaphthalma pallida (Fallén); 89, 90 — Megaphthalma unilineata (Zetterstedt); 91, 92 — Staegeria kunzei (Zetterstedt)." pageId="9" pageNumber="104">81, 82</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE2E1548DDEDF8A2A90CF865" pageId="9" pageNumber="104" type="discussion">
<paragraph id="F95FD76EAE2E1548DDEDF8A2A90CF865" blockId="9.[189,776,1704,2062]" pageId="9" pageNumber="104">
REMARKS. Monotypic Holarctic genus. The species was recorded by
<bibRefCitation id="9D71AA9FAE2E1548DCAEF881A952F8A6" author="Gorodkov K. B." box="[417,596,1893,1918]" pageId="9" pageNumber="104" pagination="440 - 458" refId="ref14639" refString="Gorodkov K. B. 1970. [Family Scathophagidae (Cordyluridae, Scatomyzidae, Scopeumatidae)] // Bei-Bienko G. Ya. (ed.). Opredelitel' nasekomykh Evropeiskoi chasti SSSR. Vol. 5. Diptera, Siphonaptera. Part 2. P. 440 - 458 [in Russian]." type="book chapter" year="1970">Gorodkov [1970</bibRefCitation>
,
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] in
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from the European part,
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and Far East but without indicating specific locality.
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MATERIAL in
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.
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Genus
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Macquart, 1835
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REMARKS. Description of the genus, descriptions of most species noted in
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and a key for their identification are given by
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[
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]. Currently, 27 species have been recorded in
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(see checklist below).
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Genus
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<figureCitation id="61DBCBEBAE241542DC94FA03AAE8FA27" box="[411,494,1511,1535]" captionStart="Figs 38–43" captionStartId="10.[501,538,1321,1340]" captionTargetBox="[835,1357,1059,1287]" captionTargetId="figure-12@10.[815,1374,1035,1299]" captionTargetPageId="10" captionText="Figs 38–43. Legs of Scathophagidae: 38 — Bostrichopyga crassipes (Zetterstedt), fore tibia, an- teroventral view; 39 — Acanthocnema sternalis Suwa, fore tibia, posterior view; 40 — Norellia tipularia (Fabricius), fore tibia, ventral view; 41 — Norellisoma spinimanum (Fallén), fore tibia, ventral view; 42, 43 — Staegeria kunzei (Zetterstedt), mid tibia, anterodorsal view (42) and mid femur, anterior view (43). 38 — after Ozerov &amp; Krivosheina, 2014a, fig. 2; 39 — after Ozerov &amp; Krivosheina, 2014b, fig. 1 (as A. vikhrevi); 40, 41 — after Ozerov &amp; Krivosheina, 2011a, figs 1, 2. Рис. 38–43. Ноги Scathophagidae: 38 — Bostrichopyga crassipes (Zetterstedt), голень передней ноги, антеровентрально ; 39 — Acanthocnema sternalis Suwa, голень передней ноги, сЗади; 40 — Norellia tipularia (Fabricius ), голень передней ноги, сниЗу; 41 — Norellisoma spinimanum (Fallén), 42 голень передней ноги, сниЗу; 42, 43 — Staegeria kunzei (Zetterstedt), голень средней ноги, антеро- дорсально (42) и бедро средней ноги, спереди (43). 38 — по Ozerov &amp; Krivosheina, 2014a, fig. 2; 39 — по Ozerov &amp; Krivosheina , 2014b, fig. 1 (as A. vikhrevi); 40, 41 — по Ozerov &amp; Krivosheina, 2011a, figs 1, 2." figureDoi="http://doi.org/10.5281/zenodo.13178446" httpUri="https://zenodo.org/record/13178446/files/figure.png" pageId="3" pageNumber="98">Figs 38</figureCitation>
,
<figureCitation id="61DBCBEBAE241542DCF6FA03A922FA27" box="[505,548,1511,1535]" captionStart="Figs 100–105" captionStartId="19.[226,263,1870,1889]" captionTargetBox="[216,1405,315,1820]" captionTargetPageId="19" captionText="Figs 100–105. Habitus of Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — after Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — after Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — after Ozerov, 2015, fig. 1; 104 — after Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — after Ozerov, 2013, fig. 19. Рис. 100–105. ОбЩий вид Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — по Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — по Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — по Ozerov, 2015, fig. 1; 104 — по Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — по Ozerov, 2013, fig. 19." figureDoi="http://doi.org/10.5281/zenodo.13178458" httpUri="https://zenodo.org/record/13178458/files/figure.png" pageId="3" pageNumber="98">104</figureCitation>
.
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REMARKS. Monotypic genus. Diagnosis of the genus, changes in nomenclature, description of male, later female of
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<emphasis id="CB940B7CAE241542DC7BF9B9AAFAF9AC" box="[372,508,1629,1652]" italics="true" pageId="3" pageNumber="98">B. crassipes</emphasis>
(Zetterstedt, 1838)
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are given by
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and by
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. In
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,
<taxonomicName id="3EE0ACEDAE241542DCC2F973A950F976" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[461,598,1687,1710]" class="Insecta" family="Scathophagidae" genus="Bostrichopyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="CB940B7CAE241542DCC2F973A950F976" box="[461,598,1687,1710]" italics="true" pageId="3" pageNumber="98">B. crassipes</emphasis>
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is known from materials taken from
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and Yakutia [
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].
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Genus
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REMARKS.
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is the largest genus within the family
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. Diagnosis and description of the genus, descriptions of all species noted in
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and a key for their identification are given by
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. There are currently 38 species in
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<subSubSection id="B1FA84E5AE241542DE91F8ADAE28F8A7" pageId="3" pageNumber="98" type="nomenclature">
<paragraph id="F95FD76EAE241542DE91F8ADAE02F8B9" blockId="3.[885,1326,1865,1919]" box="[926,1284,1865,1889]" pageId="3" pageNumber="98">
<heading id="A2176002AE241542DE91F8ADAE02F8B9" box="[926,1284,1865,1889]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="98" reason="2">
Genus
<taxonomicName id="3EE0ACEDAE241542DEE5F8AEAE02F8B9" ID-CoL="62NP2" authority="Becker, 1894" authorityName="Becker" authorityYear="1894" box="[1002,1284,1865,1889]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE241542DEE5F8AEAF6AF8B9" box="[1002,1132,1866,1889]" italics="true" pageId="3" pageNumber="98">Cosmetopus</emphasis>
<bibRefCitation id="9D71AA9FAE241542D97BF8ADAE02F8B9" author="Becker T." box="[1140,1284,1865,1889]" pageId="3" pageNumber="98" pagination="77 - 196" refId="ref14329" refString="Becker T. 1894. Dipterologische Studien. I. Scatomyzidae // Berliner Entomologische Zeitschrift. Bd. 39. H. 1. S. 77 - 196." type="journal article" year="1894">Becker, 1894</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F95FD76EAE241542DE7AF883AE28F8A7" blockId="3.[885,1326,1865,1919]" box="[885,1326,1894,1919]" pageId="3" pageNumber="98">Figs 5, 6, 44, 68, 69, 72–75, 77–80, 111.</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE241547DE5DF845AA87FA39" lastPageId="6" lastPageNumber="101" pageId="3" pageNumber="98" type="diagnosis">
<paragraph id="F95FD76EAE241542DE5DF845AE70F7C9" blockId="3.[812,1399,1953,2066]" pageId="3" pageNumber="98">
DIAGNOSIS. Medium-sized flies (
<quantity id="3E187A8BAE241542D9DCF845AE29F861" box="[1235,1327,1953,1977]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="98" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long). Proepisternum is covered with hairs at middle or in anterior part. Anepisternum covered with hairs in posterior half. Katepisternum with one strong seta in upper
</paragraph>
<caption id="AD9F87E6AE231545DDEAF8AAAF8DF7D5" ID-DOI="http://doi.org/10.5281/zenodo.13178428" ID-Zenodo-Dep="13178428" httpUri="https://zenodo.org/record/13178428/files/figure.png" pageId="4" pageNumber="99" startId="4.[229,265,1870,1889]" targetBox="[298,1313,338,1851]" targetPageId="4" targetType="figure">
<paragraph id="F95FD76EAE231545DDEAF8AAAF91F871" blockId="4.[191,1402,1870,2061]" pageId="4" pageNumber="99">
Figs 7–12. Head of
<taxonomicName id="3EE0ACEDAE231545DC90F8AAA921F8B9" box="[415,551,1870,1889]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, lateral view: 7 —
<taxonomicName id="3EE0ACEDAE231545DFD0F8AAAF7EF8B9" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2018" box="[735,1144,1870,1889]" class="Insecta" family="Scathophagidae" genus="Acanthocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="CB940B7CAE231545DFD0F8AAA8A4F8B9" box="[735,930,1870,1889]" italics="true" pageId="4" pageNumber="99">Acanthocnema altaica</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 8 —
<taxonomicName id="3EE0ACEDAE231545D9B7F8AAAA2DF8A1" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="breviseta">
<emphasis id="CB940B7CAE231545D9B7F8AAAE7FF8B9" box="[1208,1401,1870,1889]" italics="true" pageId="4" pageNumber="99">Acerocnema breviseta</emphasis>
(Zetterstedt)
</taxonomicName>
; 9 —
<taxonomicName id="3EE0ACEDAE231545DC50F882A9E9F8A1" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2018" box="[351,751,1894,1913]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="gorodkovi">
<emphasis id="CB940B7CAE231545DC50F882A922F8A1" box="[351,548,1894,1913]" italics="true" pageId="4" pageNumber="99">Acerocnema gorodkovi</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 10 —
<taxonomicName id="3EE0ACEDAE231545DE21F882AF35F8A1" authority="(Becker)" baseAuthorityName="Becker" baseAuthorityYear="1894" box="[814,1075,1894,1913]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="articulata">
<emphasis id="CB940B7CAE231545DE21F882A8E6F8A1" box="[814,992,1894,1913]" italics="true" pageId="4" pageNumber="99">Cleigastra articulata</emphasis>
(Becker)
</taxonomicName>
; 11 —
<taxonomicName id="3EE0ACEDAE231545D97DF882AA2FF849" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2016" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="maritima">
<emphasis id="CB940B7CAE231545D97DF882AE18F8A0" box="[1138,1310,1894,1913]" italics="true" pageId="4" pageNumber="99">Cleigastra maritima</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 12 —
<taxonomicName id="3EE0ACEDAE231545DC62F89AA984F849" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1805" box="[365,642,1918,1937]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="albilabris">
<emphasis id="CB940B7CAE231545DC62F89AA91DF849" box="[365,539,1918,1937]" italics="true" pageId="4" pageNumber="99">Cordilura albilabris</emphasis>
(Fabricius)
</taxonomicName>
. 7 — after
<bibRefCitation id="9D71AA9FAE231545DFE5F89AA8F4F849" author="Ozerov A. L. &amp; Krivosheina M. G." box="[746,1010,1918,1937]" pageId="4" pageNumber="99" pagination="93 - 100" refId="ref16109" refString="Ozerov A. L., Krivosheina M. G. 2018 a. A review of the genus Acanthocnema Becker, 1894 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 27. No. 1. P. 93 - 100." type="journal article" year="2018">Ozerov &amp; Krivosheina, 2018a</bibRefCitation>
, fig. 3; 8, 9 — after
<bibRefCitation id="9D71AA9FAE231545D9A1F89AABF0F871" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="4" pageNumber="99" pagination="210 - 214" refId="ref16158" refString="Ozerov A. L., Krivosheina M. G. 2018 b. Two new species of the genus Acerocnema Becker, 1894 (Diptera: Scathophagidae) from Far East of Russia // Russian Entomological Journal. Vol. 27. No. 2. P. 210 - 214." type="journal article" year="2018">Ozerov &amp; Krivosheina, 2018b</bibRefCitation>
, figs 2, 1; 11 — after
<bibRefCitation id="9D71AA9FAE231545DCBAF872A9A9F871" author="Ozerov A. L. &amp; Krivosheina M. G." box="[437,687,1942,1961]" pageId="4" pageNumber="99" pagination="97 - 102" refId="ref16062" refString="Ozerov A. L., Krivosheina M. G. 2016. To taxonomy of the genus Cleigastra Macquart, 1835 with description of two new species // Russian Entomological Journal. Vol. 25. No. 1. P. 97 - 102." type="journal article" year="2016">Ozerov &amp; Krivosheina, 2016</bibRefCitation>
, fig. 5; 12 — after
<bibRefCitation id="9D71AA9FAE231545DE59F872AF5DF871" author="Ozerov A. L. &amp; Krivosheina M. G." box="[854,1115,1942,1961]" pageId="4" pageNumber="99" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, fig. 7.
</paragraph>
<paragraph id="F95FD76EAE231545DDEAF856AF8DF7D5" blockId="4.[191,1402,1870,2061]" pageId="4" pageNumber="99">
Рис. 7–12. Голова
<taxonomicName id="3EE0ACEDAE231545DC9BF855A919F81C" box="[404,543,1969,1988]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, сбоку: 7 —
<taxonomicName id="3EE0ACEDAE231545DF94F855AF34F81C" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2018" box="[667,1074,1969,1988]" class="Insecta" family="Scathophagidae" genus="Acanthocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="CB940B7CAE231545DF94F855A859F81C" box="[667,863,1969,1988]" italics="true" pageId="4" pageNumber="99">Acanthocnema altaica</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 8 —
<taxonomicName id="3EE0ACEDAE231545D962F856ABF4F804" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="breviseta">
<emphasis id="CB940B7CAE231545D962F856AE29F81C" box="[1133,1327,1969,1989]" italics="true" pageId="4" pageNumber="99">Acerocnema breviseta</emphasis>
(Zetterstedt)
</taxonomicName>
; 9 —
<taxonomicName id="3EE0ACEDAE231545DC22F82EA9CAF804" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2018" box="[301,716,1993,2013]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="gorodkovi">
<emphasis id="CB940B7CAE231545DC22F82EAAFEF804" box="[301,504,1993,2013]" italics="true" pageId="4" pageNumber="99">Acerocnema gorodkovi</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 10 —
<taxonomicName id="3EE0ACEDAE231545DE1DF82DAF27F804" authority="(Becker)" baseAuthorityName="Becker" baseAuthorityYear="1894" box="[786,1057,1993,2012]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="articulata">
<emphasis id="CB940B7CAE231545DE1DF82DA8CCF804" box="[786,970,1993,2012]" italics="true" pageId="4" pageNumber="99">Cleigastra articulata</emphasis>
(Becker)
</taxonomicName>
; 11 —
<taxonomicName id="3EE0ACEDAE231545D968F82DAA2DF82C" authority="Ozerov &amp; Krivosheina" authorityName="Ozerov &amp; Krivosheina" authorityYear="2016" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="maritima">
<emphasis id="CB940B7CAE231545D968F82DAE1FF804" box="[1127,1305,1993,2012]" italics="true" pageId="4" pageNumber="99">Cleigastra maritima</emphasis>
Ozerov &amp; Krivosheina
</taxonomicName>
; 12 —
<taxonomicName id="3EE0ACEDAE231545DC7DF805A988F82C" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1805" box="[370,654,2017,2036]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="albilabris">
<emphasis id="CB940B7CAE231545DC7DF805A923F82C" box="[370,549,2017,2036]" italics="true" pageId="4" pageNumber="99">Cordilura albilabris</emphasis>
(Fabricius)
</taxonomicName>
. 7 — по
<bibRefCitation id="9D71AA9FAE231545DFE6F805A8FEF82C" author="Ozerov A. L. &amp; Krivosheina M. G." box="[745,1016,2017,2036]" pageId="4" pageNumber="99" pagination="93 - 100" refId="ref16109" refString="Ozerov A. L., Krivosheina M. G. 2018 a. A review of the genus Acanthocnema Becker, 1894 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 27. No. 1. P. 93 - 100." type="journal article" year="2018">Ozerov &amp; Krivosheina, 2018a</bibRefCitation>
, fig. 3; 8, 9 — по
<bibRefCitation id="9D71AA9FAE231545D9A6F805ABF1F7D4" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="4" pageNumber="99" pagination="210 - 214" refId="ref16158" refString="Ozerov A. L., Krivosheina M. G. 2018 b. Two new species of the genus Acerocnema Becker, 1894 (Diptera: Scathophagidae) from Far East of Russia // Russian Entomological Journal. Vol. 27. No. 2. P. 210 - 214." type="journal article" year="2018">Ozerov &amp; Krivosheina, 2018b</bibRefCitation>
, figs 2, 1; 11 — по
<bibRefCitation id="9D71AA9FAE231545DCA5F81DA9AFF7D4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[426,681,2041,2060]" pageId="4" pageNumber="99" pagination="97 - 102" refId="ref16062" refString="Ozerov A. L., Krivosheina M. G. 2016. To taxonomy of the genus Cleigastra Macquart, 1835 with description of two new species // Russian Entomological Journal. Vol. 25. No. 1. P. 97 - 102." type="journal article" year="2016">Ozerov &amp; Krivosheina, 2016</bibRefCitation>
, fig. 5; 12 — по
<bibRefCitation id="9D71AA9FAE231545DE4CF81DAF4BF7D4" author="Ozerov A. L. &amp; Krivosheina M. G." box="[835,1101,2041,2060]" pageId="4" pageNumber="99" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, fig. 7.
</paragraph>
</caption>
<caption id="AD9F87E6AE221544DDECF864AFDDF7D7" ID-DOI="http://doi.org/10.5281/zenodo.13178430" ID-Zenodo-Dep="13178430" httpUri="https://zenodo.org/record/13178430/files/figure.png" pageId="5" pageNumber="100" startId="5.[227,262,1920,1939]" targetBox="[269,1372,318,1903]" targetPageId="5" targetType="figure">
<paragraph id="F95FD76EAE221544DDECF864AF7DF81C" blockId="5.[189,1400,1920,2064]" pageId="5" pageNumber="100">
Figs 13–18. Head of
<taxonomicName id="3EE0ACEDAE221544DC83F864A90DF84B" box="[396,523,1920,1939]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, lateral view: 13 —
<taxonomicName id="3EE0ACEDAE221544DFA2F864A862F84B" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[685,868,1920,1939]" class="Insecta" family="Scathophagidae" genus="Delina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="CB940B7CAE221544DFA2F864A819F84B" box="[685,799,1920,1939]" italics="true" pageId="5" pageNumber="100">Delina nigrita</emphasis>
(Fallén)
</taxonomicName>
; 14 —
<taxonomicName id="3EE0ACEDAE221544DE90F865AFA8F84B" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1826" box="[927,1198,1920,1939]" class="Insecta" family="Scathophagidae" genus="Hexamitocera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="loxocerata">
<emphasis id="CB940B7CAE221544DE90F865AF6EF84B" box="[927,1128,1920,1939]" italics="true" pageId="5" pageNumber="100">Hexamitocera loxocerata</emphasis>
(Fallén)
</taxonomicName>
; 15 —
<taxonomicName id="3EE0ACEDAE221544D9E6F865AA26F873" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE221544D9E6F865AE70F84B" box="[1257,1398,1920,1939]" italics="true" pageId="5" pageNumber="100">Leptopa filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 16 —
<taxonomicName id="3EE0ACEDAE221544DC6BF87CA963F873" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[356,613,1944,1963]" class="Insecta" family="Scathophagidae" genus="Megaphthalma" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="CB940B7CAE221544DC6BF87CA91AF873" box="[356,540,1944,1963]" italics="true" pageId="5" pageNumber="100">Megaphthalma pallida</emphasis>
(Fallén)
</taxonomicName>
; 17 —
<taxonomicName id="3EE0ACEDAE221544DFA7F87CA8DCF873" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" box="[680,986,1944,1963]" class="Insecta" family="Scathophagidae" genus="Micropselapha" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE221544DFA7F87CA876F873" box="[680,880,1944,1963]" italics="true" pageId="5" pageNumber="100">Micropselapha filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 18 —
<taxonomicName id="3EE0ACEDAE221544D911F87CAE12F874" authority="Ozerov." authorityName="Ozerov et Krivosheina" authorityYear="2012" box="[1054,1300,1944,1964]" class="Insecta" family="Scathophagidae" genus="Mixocordylura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="CB940B7CAE221544D911F87CAFCDF873" box="[1054,1227,1944,1963]" italics="true" pageId="5" pageNumber="100">Mixocordylura parva</emphasis>
Ozerov.
</taxonomicName>
14 — after
<bibRefCitation id="9D71AA9FAE221544DDB2F855AAA1F81B" author="Ozerov A. L. &amp; Krivosheina M. G." box="[189,423,1968,1988]" pageId="5" pageNumber="100" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 3; 17 — after Ozerov, 2014, fig. 7; 18 — after
<bibRefCitation id="9D71AA9FAE221544DE48F855AF3FF81C" author="Ozerov A. L. &amp; Krivosheina M. G." box="[839,1081,1968,1988]" pageId="5" pageNumber="100" pagination="431 - 435" refId="ref15788" refString="Ozerov A. L., Krivosheina M. G. 2012 a. Revision of dung flies of the genus Mixocordylura Hendel (Diptera, Scathophagidae), with the description of new species from Altay // Russian Entomological Journal. Vol. 20. No. 4 (for 2011). P. 431 - 435." type="journal article" year="2012">Ozerov &amp; Krivosheina, 2012a</bibRefCitation>
, fig. 10.
</paragraph>
<paragraph id="F95FD76EAE221544DDECF829AFDDF7D7" blockId="5.[189,1400,1920,2064]" pageId="5" pageNumber="100">
Рис. 13–18. Голова
<taxonomicName id="3EE0ACEDAE221544DC91F828A925F807" box="[414,547,1996,2015]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, сбоку: 13 —
<taxonomicName id="3EE0ACEDAE221544DFBFF828A87EF807" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[688,888,1996,2015]" class="Insecta" family="Scathophagidae" genus="Delina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="CB940B7CAE221544DFBFF828A82CF807" box="[688,810,1996,2015]" italics="true" pageId="5" pageNumber="100">Delina nigrita</emphasis>
(Fallén)
</taxonomicName>
; 14 —
<taxonomicName id="3EE0ACEDAE221544DECCF829AFEEF807" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1826" box="[963,1256,1996,2015]" class="Insecta" family="Scathophagidae" genus="Hexamitocera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="loxocerata">
<emphasis id="CB940B7CAE221544DECCF829AF9CF807" box="[963,1178,1996,2015]" italics="true" pageId="5" pageNumber="100">Hexamitocera loxocerata</emphasis>
(Fallén)
</taxonomicName>
; 15 —
<taxonomicName id="3EE0ACEDAE221544D83CF829AA7EF82F" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE221544D83CF829AA0EF82F" italics="true" pageId="5" pageNumber="100">Leptopa filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 16 —
<taxonomicName id="3EE0ACEDAE221544DCB8F800A9B9F82F" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[439,703,2020,2039]" class="Insecta" family="Scathophagidae" genus="Megaphthalma" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="CB940B7CAE221544DCB8F800A970F82F" box="[439,630,2020,2039]" italics="true" pageId="5" pageNumber="100">Megaphthalma pallida</emphasis>
(Fallén)
</taxonomicName>
; 17 —
<taxonomicName id="3EE0ACEDAE221544DFF0F800AF3BF82F" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" box="[767,1085,2020,2039]" class="Insecta" family="Scathophagidae" genus="Micropselapha" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE221544DFF0F800A8C8F82F" box="[767,974,2020,2039]" italics="true" pageId="5" pageNumber="100">Micropselapha filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 18 —
<taxonomicName id="3EE0ACEDAE221544D973F800AE7EF82F" authority="Ozerov." authorityName="Ozerov et Krivosheina" authorityYear="2012" box="[1148,1400,2020,2039]" class="Insecta" family="Scathophagidae" genus="Mixocordylura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="CB940B7CAE221544D973F800AE28F82F" box="[1148,1326,2020,2039]" italics="true" pageId="5" pageNumber="100">Mixocordylura parva</emphasis>
Ozerov.
</taxonomicName>
14 — по
<bibRefCitation id="9D71AA9FAE221544DC01F818A905F7D7" author="Ozerov A. L. &amp; Krivosheina M. G." box="[270,515,2044,2063]" pageId="5" pageNumber="100" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 3; 17 — по Ozerov, 2014, fig. 7; 18 — по
<bibRefCitation id="9D71AA9FAE221544DE97F818AF90F7D7" author="Ozerov A. L. &amp; Krivosheina M. G." box="[920,1174,2044,2063]" pageId="5" pageNumber="100" pagination="431 - 435" refId="ref15788" refString="Ozerov A. L., Krivosheina M. G. 2012 a. Revision of dung flies of the genus Mixocordylura Hendel (Diptera, Scathophagidae), with the description of new species from Altay // Russian Entomological Journal. Vol. 20. No. 4 (for 2011). P. 431 - 435." type="journal article" year="2012">Ozerov &amp; Krivosheina, 2012a</bibRefCitation>
, fig. 10.
</paragraph>
</caption>
<paragraph id="F95FD76EAE211547DC8BFCD4AAC5FCB1" blockId="6.[388,451,816,873]" box="[388,451,816,873]" pageId="6" pageNumber="101">19</paragraph>
<caption id="AD9F87E6AE211547DE5DFBC5A8CBFB38" ID-DOI="http://doi.org/10.5281/zenodo.13178434" ID-Zenodo-Dep="13178434" httpUri="https://zenodo.org/record/13178434/files/figure.png" pageId="6" pageNumber="101" startId="6.[850,887,1057,1076]" targetBox="[900,1388,355,882]" targetPageId="6" targetType="figure">
<paragraph id="F95FD76EAE211547DE5DFBC5A8CBFBA4" blockId="6.[812,1398,1057,1248]" pageId="6" pageNumber="101">
Figs 19–21. Head of
<taxonomicName id="3EE0ACEDAE211547D904FBC5AF90FBEC" box="[1035,1174,1057,1076]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, lateral view (19, 20) and dorsolateral view (21): 19 —
<taxonomicName id="3EE0ACEDAE211547D94DFBDDAE74FB94" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[1090,1394,1081,1100]" class="Insecta" family="Scathophagidae" genus="Spaziphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="hydromyzina">
<emphasis id="CB940B7CAE211547D94DFBDDAE26FB94" box="[1090,1312,1081,1100]" italics="true" pageId="6" pageNumber="101">Spaziphora hydromyzina</emphasis>
(Fallén)
</taxonomicName>
; 20 —
<taxonomicName id="3EE0ACEDAE211547DE62FBB5AFC4FBBC" authority="(Vockeroth)" baseAuthorityName="Vockeroth" baseAuthorityYear="1995" box="[877,1218,1105,1124]" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="nearcticus">
<emphasis id="CB940B7CAE211547DE62FBB5AF4EFBBC" box="[877,1096,1105,1124]" italics="true" pageId="6" pageNumber="101">Trichopalpus nearcticus</emphasis>
(Vockeroth)
</taxonomicName>
; 21 —
<taxonomicName id="3EE0ACEDAE211547D81DFBB5A8CFFBA4" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" class="Insecta" family="Scathophagidae" genus="Hydromyza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="livens">
<emphasis id="CB940B7CAE211547D81DFBB5A858FBA4" italics="true" pageId="6" pageNumber="101">Hydromyza livens</emphasis>
(Fabricius)
</taxonomicName>
.
</paragraph>
<paragraph id="F95FD76EAE211547DE5DFB62A8CBFB38" blockId="6.[812,1398,1057,1248]" pageId="6" pageNumber="101">
Рис. 19–21. Голова
<taxonomicName id="3EE0ACEDAE211547D92BFB61AFBCFB40" box="[1060,1210,1157,1176]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
, сбоку (19, 20) и дорсолатерально (21): 19 —
<taxonomicName id="3EE0ACEDAE211547D949FB79AE74FB68" authority="(Fallen)" baseAuthorityName="Fallen" baseAuthorityYear="1819" box="[1094,1394,1181,1200]" class="Insecta" family="Scathophagidae" genus="Spaziphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="hydromyzina">
<emphasis id="CB940B7CAE211547D949FB79AE27FB68" box="[1094,1313,1181,1200]" italics="true" pageId="6" pageNumber="101">Spaziphora hydromyzina</emphasis>
(Fallén)
</taxonomicName>
; 20 —
<taxonomicName id="3EE0ACEDAE211547DE62FB51AFC4FB10" authority="(Vockeroth)" baseAuthorityName="Vockeroth" baseAuthorityYear="1995" box="[877,1218,1205,1224]" class="Insecta" family="Scathophagidae" genus="Trichopalpus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="nearcticus">
<emphasis id="CB940B7CAE211547DE62FB51AF4EFB10" box="[877,1096,1205,1224]" italics="true" pageId="6" pageNumber="101">Trichopalpus nearcticus</emphasis>
(Vockeroth)
</taxonomicName>
; 21 —
<taxonomicName id="3EE0ACEDAE211547D81DFB51A8CFFB38" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1794" class="Insecta" family="Scathophagidae" genus="Hydromyza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="livens">
<emphasis id="CB940B7CAE211547D81DFB51A858FB38" italics="true" pageId="6" pageNumber="101">Hydromyza livens</emphasis>
(Fabricius)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F95FD76EAE211547DDB2FAC8AA87FA39" blockId="6.[189,775,1324,2065]" pageId="6" pageNumber="101">
posterior corner. Postmetacoxal bridge absent. Fore tibia ventrally with short black spinules. Male fore femur ventrally near midlength with bifurcate tubercle, matching with ventral emargination of fore tibia (
<figureCitation id="61DBCBEBAE211547DFD4FA6EABDAFA1A" captionStart="48 49" captionStartId="11.[292,355,1751,1808]" captionText="48 49 Figs 44–49. Legs of Scathophagidae: 44 — Cosmetopus longus (Walker), male fore leg, posterior view; 45 — Microprosopa pallidicauda (Zetterstedt), male fore leg, anterior view; 46 — Norellisoma spinimanum (Fallén), fore leg, posterior view; 47 — Paracosmetopus helleni Hackman, female fore leg, anterior view; 48 — Pogonota stackelbergi (Gorodkov), male fore leg, posterior view; 49 — Pogonota dasyprocta (Loew), female fore leg, anterior view. 45 — after Ozerov, 2017, fig. 4; 47 — after Ozerov &amp; Krivosheina, 2014a, fig. 12; 48, 49 — after Ozerov, 2016, figs 2, 4. Рис. 44–49. Ноги Scathophagidae: 44 — Cosmetopus longus (Walker), переднЯЯ нога самца, сЗади; 45 — Microprosopa pallidicauda (Zetterstedt), переднЯЯ нога самца, спереди; 46 — Norellisoma spinimanum (Fallén), переднЯЯ нога, сЗади; 47 — Paracosmetopus helleni Hackman, переднЯЯ нога самки, спереди; 48 — Pogonota stackelbergi (Gorodkov), переднЯЯ нога самца, сЗади; 49 — Pogonota dasyprocta (Loew), переднЯЯ нога самки, спереди. 45 — по Ozerov, 2017, fig. 4; 47 — по Ozerov &amp; Krivosheina, 2014a, fig. 12; 48, 49 — по Ozerov, 2016, figs 2, 4." pageId="6" pageNumber="101">Fig. 44</figureCitation>
). Female fore femur simple: without numerous short spinules ventrally.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE211547DDEDFA0DA986F946" pageId="6" pageNumber="101" type="discussion">
<paragraph id="F95FD76EAE211547DDEDFA0DA986F946" blockId="6.[189,775,1324,2065]" pageId="6" pageNumber="101">
REMARKS. There are 2 species in
<collectingCountry id="81F797FEAE211547DF6DFA0DA9AAF9D9" box="[610,684,1513,1537]" name="Russia" pageId="6" pageNumber="101">Russia</collectingCountry>
,
<taxonomicName id="3EE0ACEDAE211547DFB8FA0DAAEBF9F8" authority="(Zetterstedt, 1838)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="dentimanus">
<emphasis id="CB940B7CAE211547DFB8FA0DAA17F9F8" italics="true" pageId="6" pageNumber="101">C. dentimanus</emphasis>
(Zetterstedt, 1838)
</taxonomicName>
and
<taxonomicName id="3EE0ACEDAE211547DF21F9EDABF8F998" authority="(Walker, 1849)" baseAuthorityName="Walker" baseAuthorityYear="1849" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="CB940B7CAE211547DF21F9EDA99DF9F8" box="[558,667,1544,1568]" italics="true" pageId="6" pageNumber="101">C. longus</emphasis>
(Walker, 1849)
</taxonomicName>
, of which only
<taxonomicName id="3EE0ACEDAE211547DCABF9CCA90CF998" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[420,522,1576,1600]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="CB940B7CAE211547DCABF9CCA90CF998" box="[420,522,1576,1600]" italics="true" pageId="6" pageNumber="101">C. longus</emphasis>
</taxonomicName>
was previously record- ed in
<collectingCountry id="81F797FEAE211547DDF3F9A3AA45F987" box="[252,323,1607,1631]" name="Russia" pageId="6" pageNumber="101">Russia</collectingCountry>
from south of European part and
<taxonomicName id="3EE0ACEDAE211547DFB4F9A3A801F987" authorityName="Blackwelder" authorityYear="1952" box="[699,775,1607,1631]" class="Insecta" family="Staphylinidae" genus="Siberia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="112" phylum="Arthropoda" rank="genus">Siberia</taxonomicName>
by
<bibRefCitation id="9D71AA9FAE211547DDD1F983AA96F9A7" author="Gorodkov K. B." box="[222,400,1639,1663]" pageId="6" pageNumber="101" pagination="440 - 458" refId="ref14639" refString="Gorodkov K. B. 1970. [Family Scathophagidae (Cordyluridae, Scatomyzidae, Scopeumatidae)] // Bei-Bienko G. Ya. (ed.). Opredelitel' nasekomykh Evropeiskoi chasti SSSR. Vol. 5. Diptera, Siphonaptera. Part 2. P. 440 - 458 [in Russian]." type="book chapter" year="1970">Gorodkov [1970</bibRefCitation>
(as
<taxonomicName id="3EE0ACEDAE211547DCB2F983A806F9A7" authority="(Zetterstedt), 1986" authorityName="" authorityYear="1986" baseAuthorityName="Zetterstedt" baseAuthorityYear="1986" box="[445,768,1639,1663]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="CB940B7CAE211547DCB2F983A929F9A7" box="[445,559,1639,1663]" italics="true" pageId="6" pageNumber="101">C. fulvipes</emphasis>
(Zetterstedt), 1986
</taxonomicName>
] without specifying the place of collecting.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE211547DDEDF942AA9BF7C8" pageId="6" pageNumber="101" type="materials_examined">
<paragraph id="F95FD76EAE211547DDEDF942AA9BF7C8" blockId="6.[189,775,1324,2065]" pageId="6" pageNumber="101">
MATERIAL in
<collectionCode id="9FF14FABAE211547DC61F942AAB5F961" box="[366,435,1702,1721]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="6" pageNumber="101">ZMUM</collectionCode>
.
<taxonomicName id="3EE0ACEDAE211547DCB3F942A938F961" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[444,574,1702,1721]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="dentimanus">
<emphasis id="CB940B7CAE211547DCB3F942A938F961" bold="true" box="[444,574,1702,1721]" italics="true" pageId="6" pageNumber="101">C. dentimanus</emphasis>
</taxonomicName>
.
<taxonomicName id="3EE0ACEDAE211547DF46F942A981F961" authorityName="Blackwelder" authorityYear="1952" box="[585,647,1702,1721]" class="Insecta" family="Staphylinidae" genus="Siberia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE211547DF46F942A981F961" box="[585,647,1702,1721]" italics="true" pageId="6" pageNumber="101">Siberia</emphasis>
</taxonomicName>
:
<materialsCitation id="4988DD33AE211547DF9BF942AA47F8D0" collectingDate="2008-07-08" collectorName="A. Barkalov. &amp; European" country="Russia" county="Altai" elevation="2450" latitude="49.3" location="Ukok" longLatPrecision="5567" longitude="87.65" municipality="Kosh-Agach" pageId="6" pageNumber="101" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="EFE61CE7AE211547DF9BF942A9CBF961" box="[660,717,1699,1721]" count="2" pageId="6" pageNumber="101" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="EFE61CE7AE211547DFD7F942A807F961" box="[728,769,1699,1721]" count="2" pageId="6" pageNumber="101" type="female">2 ♀♀</specimenCount>
,
<collectingCounty id="103EAFE2AE211547DDB2F924ABEAF90B" box="[189,236,1728,1747]" pageId="6" pageNumber="101">Altai</collectingCounty>
,
<collectingMunicipality id="193B4D14AE211547DDF4F924AA68F90B" box="[251,366,1728,1747]" pageId="6" pageNumber="101">Kosh-Agach</collectingMunicipality>
, plato
<location id="FC3F81B5AE211547DCB9F924AAECF90B" LSID="urn:lsid:plazi:treatment:71496678AE241547DE91F8ADAA9BF7C8:FC3F81B5AE211547DCB9F924AAECF90B" box="[438,490,1728,1747]" country="Russia" county="Altai" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Kosh-Agach" name="Ukok" pageId="6" pageNumber="101">Ukok</location>
,
<quantity id="3E187A8BAE211547DCF7F924A945F90C" box="[504,579,1728,1748]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="101" unit="m" value="2450.0">
<elevation id="72CD305DAE211547DCF7F924A945F90C" box="[504,579,1728,1748]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="101" unit="m" value="2450.0">2450 m</elevation>
</quantity>
,
<location id="FC3F81B5AE211547DF5CF924A801F90B" LSID="urn:lsid:plazi:treatment:71496678AE241547DE91F8ADAA9BF7C8:FC3F81B5AE211547DF5CF924A801F90B" box="[595,775,1728,1747]" country="Russia" county="Altai" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Kosh-Agach" name="Lake Muzdy-Bulak" pageId="6" pageNumber="101">Lake Muzdy-Bulak</location>
(
<geoCoordinate id="9CD4B1A9AE211547DDCCF93FAA04F936" box="[195,258,1755,1774]" degrees="49.3" direction="north" orientation="latitude" pageId="6" pageNumber="101" precision="5555" value="49.3">49.3ºN</geoCoordinate>
,
<geoCoordinate id="9CD4B1A9AE211547DC02F93FAA53F936" box="[269,341,1755,1774]" degrees="87.65" direction="east" orientation="longitude" pageId="6" pageNumber="101" precision="555" value="87.65">87.65ºE</geoCoordinate>
),
<date id="8D5EF1AEAE211547DC68F93FAAC1F936" box="[359,455,1755,1774]" pageId="6" pageNumber="101" value="2008-07-08">
<collectingDate id="9D1A0846AE211547DC68F93FAAC1F936" box="[359,455,1755,1774]" pageId="6" pageNumber="101" value="2008-07-08">8.VII.2008</collectingDate>
</date>
,
<collectorName id="5415B2B8AE211547DCDCF93FA940F936" box="[467,582,1755,1774]" pageId="6" pageNumber="101">A. Barkalov.</collectorName>
<taxonomicName id="3EE0ACEDAE211547DF42F93FA9A2F935" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[589,676,1754,1774]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="101" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="CB940B7CAE211547DF42F93FA9A2F935" bold="true" box="[589,676,1754,1774]" italics="true" pageId="6" pageNumber="101">C. longus</emphasis>
</taxonomicName>
.
<emphasis id="CB940B7CAE211547DFBDF93FAA47F8D0" italics="true" pageId="6" pageNumber="101">
<collectorName id="5415B2B8AE211547DFBDF93FA801F935" box="[690,775,1755,1773]" pageId="6" pageNumber="101">European</collectorName>
part of
<collectingCountry id="81F797FEAE211547DC08F912AA47F8D0" box="[263,321,1782,1800]" name="Russia" pageId="6" pageNumber="101">Russia</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="4988DD33AE211547DC5CF911AABCF8FA" collectingDate="2010-08-04" collectorName="D. Gavryushin" country="Russia" latitude="64.546" location="Arkhangelsk" longLatPrecision="60" longitude="40.567" pageId="6" pageNumber="101" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Arkhangel'sk">
<specimenCount id="EFE61CE7AE211547DC5CF911AA7AF8D1" box="[339,380,1779,1801]" count="1" pageId="6" pageNumber="101" type="male">1 ♂</specimenCount>
,
<specimenCount id="EFE61CE7AE211547DC82F911AABDF8D1" box="[397,443,1779,1801]" count="2" pageId="6" pageNumber="101" type="female">2 ♀♀</specimenCount>
,
<collectingRegion id="3B24198CAE211547DCC4F911A93CF8D0" box="[459,570,1781,1800]" country="Russia" name="Arkhangel'sk" pageId="6" pageNumber="101">Arkhangelsk</collectingRegion>
(
<geoCoordinate id="9CD4B1A9AE211547DF44F911A999F8D0" box="[587,671,1781,1800]" degrees="64.546" direction="north" orientation="latitude" pageId="6" pageNumber="101" precision="55" value="64.546">64.546ºN</geoCoordinate>
<geoCoordinate id="9CD4B1A9AE211547DFA5F911A9FBF8D0" box="[682,765,1781,1800]" degrees="40.567" direction="east" orientation="longitude" pageId="6" pageNumber="101" precision="55" value="40.567">40.567ºE</geoCoordinate>
),
<date id="8D5EF1AEAE211547DDB2F8F4AA22F8FB" box="[189,292,1808,1827]" pageId="6" pageNumber="101" value="2010-08-04">
<collectingDate id="9D1A0846AE211547DDB2F8F4AA22F8FB" box="[189,292,1808,1827]" pageId="6" pageNumber="101" value="2010-08-04">4.VIII.2010</collectingDate>
</date>
,
<collectorName id="5415B2B8AE211547DC3CF8F4AABCF8FA" box="[307,442,1807,1827]" pageId="6" pageNumber="101">D. Gavryushin</collectorName>
</materialsCitation>
;
<materialsCitation id="4988DD33AE211547DCC7F8F4AAF7F8E5" collectingDate="2008-07-07" collectorName="A. L. Ozerov" country="Russia" latitude="67.631" location="Nar'yan-Mar" longLatPrecision="59" longitude="52.985" pageId="6" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Arkhangel'sk">
<specimenCount id="EFE61CE7AE211547DCC7F8F4AAF7F8FB" box="[456,497,1805,1827]" count="1" pageId="6" pageNumber="101" type="male">1 ♂</specimenCount>
,
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<subSubSection id="B1FA84E5AE251543DECFF8AAAFE5F8BE" box="[960,1251,1870,1894]" pageId="2" pageNumber="97" type="description">
<paragraph id="F95FD76EAE251543DECFF8AAAFE5F8BE" blockId="2.[925,1285,1839,1894]" box="[960,1251,1870,1894]" pageId="2" pageNumber="97">
<figureCitation id="61DBCBEBAE251543DECFF8AAAF26F8BE" box="[960,1056,1870,1894]" captionStart="Figs 7–12" captionStartId="4.[229,265,1870,1889]" captionTargetBox="[298,1313,338,1851]" captionTargetPageId="4" captionText="Figs 7–12. Head of Scathophagidae, lateral view: 7 — Acanthocnema altaica Ozerov &amp; Krivosheina; 8 — Acerocnema breviseta (Zetterstedt); 9 — Acerocnema gorodkovi Ozerov &amp; Krivosheina; 10 — Cleigastra articulata (Becker); 11 — Cleigastra maritima Ozerov &amp; Krivosheina; 12 — Cordilura albilabris (Fabricius). 7 — after Ozerov &amp; Krivosheina, 2018a, fig. 3; 8, 9 — after Ozerov &amp; Krivosheina, 2018b, figs 2, 1; 11 — after Ozerov &amp; Krivosheina, 2016, fig. 5; 12 — after Ozerov &amp; Krivosheina, 2020b, fig. 7. Рис. 7–12. Голова Scathophagidae, сбоку: 7 — Acanthocnema altaica Ozerov &amp; Krivosheina; 8 — Acerocnema breviseta (Zetterstedt); 9 — Acerocnema gorodkovi Ozerov &amp; Krivosheina; 10 — Cleigastra articulata (Becker); 11 — Cleigastra maritima Ozerov &amp; Krivosheina; 12 — Cordilura albilabris (Fabricius). 7 — по Ozerov &amp; Krivosheina, 2018a, fig. 3; 8, 9 — по Ozerov &amp; Krivosheina, 2018b, figs 2, 1; 11 — по Ozerov &amp; Krivosheina, 2016, fig. 5; 12 — по Ozerov &amp; Krivosheina, 2020b, fig. 7." figureDoi="http://doi.org/10.5281/zenodo.13178428" httpUri="https://zenodo.org/record/13178428/files/figure.png" pageId="2" pageNumber="97">Figs 8, 9</figureCitation>
,
<figureCitation id="61DBCBEBAE251543D923F8AAAF74F8BE" box="[1068,1138,1870,1894]" captionStart="Figs 50–58" captionStartId="12.[228,266,1822,1841]" captionTargetBox="[200,768,1098,1764]" captionTargetId="figure-21@12.[200,778,1360,1753]" captionTargetPageId="12" captionText="Figs 50–58. Lower and upper calypters of wing (50) and wings (51–58) of Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — after Ozerov, 2016, figs 7, 8; 55 — after Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — after Ozerov, 2017, fig. 9. Рис. 50–58. Закрыловые чеШуйки крыла (50) и крыльЯ (51–58) Scathophagidae: 50 — Scathophaga stercoraria (Linnaeus); 51 — Pogonota barbata (Zetterstedt); 52 — Acerocnema arctica Ozerov; 53 — Ernoneura argus (Zetterstedt); 54 — Pogonota immunda (Zetterstedt); 55 — Pleurochaetella barkalovi Ozerov; 56 — Microprosopa pallidicauda (Zetterstedt); 57 — Cleigastra apicalis (Meigen); 58 — Acerocnema lobanovi Ozerov, fragment. 51, 54 — по Ozerov, 2016, figs 7, 8; 55 — по Ozerov &amp; Krivosheina, 2012b, fig. 6; 56 — по Ozerov, 2017, fig. 9." figureDoi="http://doi.org/10.5281/zenodo.13178448" httpUri="https://zenodo.org/record/13178448/files/figure.png" pageId="2" pageNumber="97">52, 58</figureCitation>
,
<figureCitation id="61DBCBEBAE251543D971F8AAAFD9F8BE" box="[1150,1247,1870,1894]" captionStart="Figs 100–105" captionStartId="19.[226,263,1870,1889]" captionTargetBox="[216,1405,315,1820]" captionTargetPageId="19" captionText="Figs 100–105. Habitus of Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — after Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — after Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — after Ozerov, 2015, fig. 1; 104 — after Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — after Ozerov, 2013, fig. 19. Рис. 100–105. ОбЩий вид Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — по Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — по Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — по Ozerov, 2015, fig. 1; 104 — по Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — по Ozerov, 2013, fig. 19." figureDoi="http://doi.org/10.5281/zenodo.13178458" httpUri="https://zenodo.org/record/13178458/files/figure.png" pageId="2" pageNumber="97">101–103</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE251542DE5DF86EAF7BFA57" lastPageId="3" lastPageNumber="98" pageId="2" pageNumber="97" type="discussion">
<paragraph id="F95FD76EAE251543DE5DF86EAF62F823" blockId="2.[812,1398,1929,2043]" pageId="2" pageNumber="97">
REMARKS. Diagnosis of the genus and a key for identification the species noted in
<collectingCountry id="81F797FEAE251543D993F843AFE5F867" box="[1180,1251,1959,1983]" name="Russia" pageId="2" pageNumber="97">Russia</collectingCountry>
are given by
<bibRefCitation id="9D71AA9FAE251543DE23F821AF86F805" author="Ozerov A. L. &amp; Krivosheina M. G." box="[812,1152,1989,2013]" pageId="2" pageNumber="97" pagination="210 - 214" refId="ref16158" refString="Ozerov A. L., Krivosheina M. G. 2018 b. Two new species of the genus Acerocnema Becker, 1894 (Diptera: Scathophagidae) from Far East of Russia // Russian Entomological Journal. Vol. 27. No. 2. P. 210 - 214." type="journal article" year="2018">Ozerov &amp; Krivosheina [2018b]</bibRefCitation>
. There are 9 species in
<collectingCountry id="81F797FEAE251543DE23F807A875F823" box="[812,883,2019,2043]" name="Russia" pageId="2" pageNumber="97">Russia</collectingCountry>
(see checklist below).
</paragraph>
<paragraph id="F95FD76EAE241542DDF7FB59AE35FB36" blockId="3.[191,1401,1204,1424]" box="[248,1331,1204,1271]" pageId="3" pageNumber="98">5 6</paragraph>
<paragraph id="F95FD76EAE241542DDEAFAE4AE52FA9B" blockId="3.[191,1401,1204,1424]" pageId="3" pageNumber="98">
Figs 1–6. Antenna, lateral view (1, 2) and palpus (3–6) of
<taxonomicName id="3EE0ACEDAE241542DE07FAE4A892FACB" box="[776,916,1280,1299]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 1 —
<taxonomicName id="3EE0ACEDAE241542DEDCFAE4AFE5FACB" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[979,1251,1280,1299]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="CB940B7CAE241542DEDCFAE4AF6DFACB" box="[979,1131,1280,1299]" italics="true" pageId="3" pageNumber="98">Cleigastra pilosa</emphasis>
(Zetterstedt)
</taxonomicName>
; 2 —
<taxonomicName id="3EE0ACEDAE241542D82CFAE4AA4BFAF3" authority="(Collin)" baseAuthorityName="Collin" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="aemula">
<emphasis id="CB940B7CAE241542D82CFAE4ABF8FAF3" italics="true" pageId="3" pageNumber="98">Cordilura aemula</emphasis>
(Collin)
</taxonomicName>
; 3 —
<taxonomicName id="3EE0ACEDAE241542DC86FAFDA99BFAF3" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" box="[393,669,1304,1323]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE241542DC86FAFDA921FAF3" box="[393,551,1304,1323]" italics="true" pageId="3" pageNumber="98">Leptopa filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 4 —
<taxonomicName id="3EE0ACEDAE241542DFD7FAFCA8C6FAF3" authority="(Elberg)" baseAuthorityName="Elberg" baseAuthorityYear="1972" box="[728,960,1304,1323]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="remmi">
<emphasis id="CB940B7CAE241542DFD7FAFCA868FAF3" box="[728,878,1304,1323]" italics="true" pageId="3" pageNumber="98">Cordilura remmi</emphasis>
(Elberg)
</taxonomicName>
; 5 —
<taxonomicName id="3EE0ACEDAE241542DEF3FAFDAE40FAF3" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[1020,1350,1304,1324]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="dentimanus">
<emphasis id="CB940B7CAE241542DEF3FAFDAFD6FAF3" box="[1020,1232,1304,1324]" italics="true" pageId="3" pageNumber="98">Cosmetopus dentimanus</emphasis>
(Zetterstedt)
</taxonomicName>
; 6 —
<taxonomicName id="3EE0ACEDAE241542DDB0FAD5AAB9FA9B" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[191,447,1328,1348]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="CB940B7CAE241542DDB0FAD5AA61FA9B" box="[191,359,1328,1348]" italics="true" pageId="3" pageNumber="98">Cosmetopus longus</emphasis>
(Walker)
</taxonomicName>
. 1 — after
<bibRefCitation id="9D71AA9FAE241542DF2BFAD4A822FA9B" author="Ozerov A. L. &amp; Krivosheina M. G." box="[548,804,1328,1347]" pageId="3" pageNumber="98" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 177; 2, 4 — after
<bibRefCitation id="9D71AA9FAE241542DEFEFAD4AFFAFA9B" author="Ozerov A. L. &amp; Krivosheina M. G." box="[1009,1276,1328,1347]" pageId="3" pageNumber="98" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, figs 4, 8.
</paragraph>
<paragraph id="F95FD76EAE241542DDEAFAA9AF7BFA57" blockId="3.[191,1401,1204,1424]" pageId="3" pageNumber="98">
Рис. 1–6. Усик, сбоку (1, 2) и Щупик (3–6)
<taxonomicName id="3EE0ACEDAE241542DF75FAA8A80FFA87" box="[634,777,1356,1375]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="family">Scathophagidae</taxonomicName>
: 1 —
<taxonomicName id="3EE0ACEDAE241542DE4FFAA8AF56FA87" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[832,1104,1356,1375]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="CB940B7CAE241542DE4FFAA8A8DFFA87" box="[832,985,1356,1375]" italics="true" pageId="3" pageNumber="98">Cleigastra pilosa</emphasis>
(Zetterstedt)
</taxonomicName>
; 2 —
<taxonomicName id="3EE0ACEDAE241542D987FAA8AE73FA87" authority="(Collin)" baseAuthorityName="Collin" box="[1160,1397,1356,1375]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="aemula">
<emphasis id="CB940B7CAE241542D987FAA8AE20FA87" box="[1160,1318,1356,1375]" italics="true" pageId="3" pageNumber="98">Cordilura aemula</emphasis>
(Collin)
</taxonomicName>
; 3 —
<taxonomicName id="3EE0ACEDAE241542DDE2FA81AAF8FAAF" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1846" box="[237,510,1380,1399]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE241542DDE2FA81AA8FFAAF" box="[237,393,1380,1399]" italics="true" pageId="3" pageNumber="98">Leptopa filiformis</emphasis>
(Zetterstedt)
</taxonomicName>
; 4 —
<taxonomicName id="3EE0ACEDAE241542DF3AFA80A81CFAAF" authority="(Elberg)" baseAuthorityName="Elberg" baseAuthorityYear="1972" box="[565,794,1380,1399]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="remmi">
<emphasis id="CB940B7CAE241542DF3AFA80A9CFFAAF" box="[565,713,1380,1399]" italics="true" pageId="3" pageNumber="98">Cordilura remmi</emphasis>
(Elberg)
</taxonomicName>
; 5 —
<taxonomicName id="3EE0ACEDAE241542DE5EFA80AF9FFAAF" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" box="[849,1177,1380,1399]" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="dentimanus">
<emphasis id="CB940B7CAE241542DE5EFA80AF22FAAF" box="[849,1060,1380,1399]" italics="true" pageId="3" pageNumber="98">Cosmetopus dentimanus</emphasis>
(Zetterstedt)
</taxonomicName>
; 6 —
<taxonomicName id="3EE0ACEDAE241542D9DFFA80AA16FA57" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1849" class="Insecta" family="Scathophagidae" genus="Cosmetopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="98" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="CB940B7CAE241542D9DFFA80AE7FFAAF" box="[1232,1401,1380,1399]" italics="true" pageId="3" pageNumber="98">Cosmetopus longus</emphasis>
(Walker)
</taxonomicName>
. 1 — по
<bibRefCitation id="9D71AA9FAE241542DC6DFA98A967FA57" author="Ozerov A. L. &amp; Krivosheina M. G." box="[354,609,1404,1423]" pageId="3" pageNumber="98" pagination="201 - 258" refId="ref16000" refString="Ozerov A. L., Krivosheina M. G. 2015. A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia // Zootaxa. Vol. 4012. No. 2. P. 201 - 258." type="journal article" year="2015">Ozerov &amp; Krivosheina, 2015</bibRefCitation>
, fig. 177; 2, 4 — по
<bibRefCitation id="9D71AA9FAE241542DE13FA98AF20FA57" author="Ozerov A. L. &amp; Krivosheina M. G." box="[796,1062,1404,1423]" pageId="3" pageNumber="98" pagination="439 - 480" refId="ref16260" refString="Ozerov A. L., Krivosheina M. G. 2020 b. A review of the genus Cordilura Fallen, 1810 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 29. No. 4. P. 439 - 480." type="journal article" year="2020">Ozerov &amp; Krivosheina, 2020b</bibRefCitation>
, figs 4, 8.
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<subSubSection id="B1FA84E5AE251543DE9FF9FBAE14F9EF" box="[912,1298,1567,1591]" pageId="2" pageNumber="97" type="nomenclature">
<paragraph id="F95FD76EAE251543DE9FF9FBAE14F9EF" blockId="2.[912,1298,1567,1622]" box="[912,1298,1567,1591]" pageId="2" pageNumber="97">
<heading id="A2176002AE251543DE9FF9FBAE14F9EF" box="[912,1298,1567,1591]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="97" reason="2">
Genus
<taxonomicName id="3EE0ACEDAE251543DED3F9FBAE14F9EF" ID-CoL="M5C" authority="Becker, 1894" authorityName="Becker" authorityYear="1894" box="[988,1298,1567,1591]" class="Insecta" family="Scathophagidae" genus="Acanthocnema" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE251543DED3F9FBAF7CF9EF" box="[988,1146,1567,1591]" italics="true" pageId="2" pageNumber="97">Acanthocnema</emphasis>
<bibRefCitation id="9D71AA9FAE251543D98CF9FBAE14F9EF" author="Becker T." box="[1155,1298,1567,1591]" pageId="2" pageNumber="97" pagination="77 - 196" refId="ref14329" refString="Becker T. 1894. Dipterologische Studien. I. Scatomyzidae // Berliner Entomologische Zeitschrift. Bd. 39. H. 1. S. 77 - 196." type="journal article" year="1894">Becker, 1894</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE251543DEF2F9DAAFA0F98E" box="[1021,1190,1598,1622]" pageId="2" pageNumber="97" type="description">
<paragraph id="F95FD76EAE251543DEF2F9DAAFA0F98E" blockId="2.[912,1298,1567,1622]" box="[1021,1190,1598,1622]" pageId="2" pageNumber="97">
<figureCitation id="61DBCBEBAE251543DEF2F9DAAF44F98D" box="[1021,1090,1598,1622]" captionStart="Figs 7–12" captionStartId="4.[229,265,1870,1889]" captionTargetBox="[298,1313,338,1851]" captionTargetPageId="4" captionText="Figs 7–12. Head of Scathophagidae, lateral view: 7 — Acanthocnema altaica Ozerov &amp; Krivosheina; 8 — Acerocnema breviseta (Zetterstedt); 9 — Acerocnema gorodkovi Ozerov &amp; Krivosheina; 10 — Cleigastra articulata (Becker); 11 — Cleigastra maritima Ozerov &amp; Krivosheina; 12 — Cordilura albilabris (Fabricius). 7 — after Ozerov &amp; Krivosheina, 2018a, fig. 3; 8, 9 — after Ozerov &amp; Krivosheina, 2018b, figs 2, 1; 11 — after Ozerov &amp; Krivosheina, 2016, fig. 5; 12 — after Ozerov &amp; Krivosheina, 2020b, fig. 7. Рис. 7–12. Голова Scathophagidae, сбоку: 7 — Acanthocnema altaica Ozerov &amp; Krivosheina; 8 — Acerocnema breviseta (Zetterstedt); 9 — Acerocnema gorodkovi Ozerov &amp; Krivosheina; 10 — Cleigastra articulata (Becker); 11 — Cleigastra maritima Ozerov &amp; Krivosheina; 12 — Cordilura albilabris (Fabricius). 7 — по Ozerov &amp; Krivosheina, 2018a, fig. 3; 8, 9 — по Ozerov &amp; Krivosheina, 2018b, figs 2, 1; 11 — по Ozerov &amp; Krivosheina, 2016, fig. 5; 12 — по Ozerov &amp; Krivosheina, 2020b, fig. 7." figureDoi="http://doi.org/10.5281/zenodo.13178428" httpUri="https://zenodo.org/record/13178428/files/figure.png" pageId="2" pageNumber="97">Figs 7</figureCitation>
,
<figureCitation id="61DBCBEBAE251543D941F9DAAF6DF98E" box="[1102,1131,1598,1622]" captionStart="Figs 38–43" captionStartId="10.[501,538,1321,1340]" captionTargetBox="[835,1357,1059,1287]" captionTargetId="figure-12@10.[815,1374,1035,1299]" captionTargetPageId="10" captionText="Figs 38–43. Legs of Scathophagidae: 38 — Bostrichopyga crassipes (Zetterstedt), fore tibia, an- teroventral view; 39 — Acanthocnema sternalis Suwa, fore tibia, posterior view; 40 — Norellia tipularia (Fabricius), fore tibia, ventral view; 41 — Norellisoma spinimanum (Fallén), fore tibia, ventral view; 42, 43 — Staegeria kunzei (Zetterstedt), mid tibia, anterodorsal view (42) and mid femur, anterior view (43). 38 — after Ozerov &amp; Krivosheina, 2014a, fig. 2; 39 — after Ozerov &amp; Krivosheina, 2014b, fig. 1 (as A. vikhrevi); 40, 41 — after Ozerov &amp; Krivosheina, 2011a, figs 1, 2. Рис. 38–43. Ноги Scathophagidae: 38 — Bostrichopyga crassipes (Zetterstedt), голень передней ноги, антеровентрально ; 39 — Acanthocnema sternalis Suwa, голень передней ноги, сЗади; 40 — Norellia tipularia (Fabricius ), голень передней ноги, сниЗу; 41 — Norellisoma spinimanum (Fallén), 42 голень передней ноги, сниЗу; 42, 43 — Staegeria kunzei (Zetterstedt), голень средней ноги, антеро- дорсально (42) и бедро средней ноги, спереди (43). 38 — по Ozerov &amp; Krivosheina, 2014a, fig. 2; 39 — по Ozerov &amp; Krivosheina , 2014b, fig. 1 (as A. vikhrevi); 40, 41 — по Ozerov &amp; Krivosheina, 2011a, figs 1, 2." figureDoi="http://doi.org/10.5281/zenodo.13178446" httpUri="https://zenodo.org/record/13178446/files/figure.png" pageId="2" pageNumber="97">39</figureCitation>
,
<figureCitation id="61DBCBEBAE251543D978F9DAAFA4F98E" box="[1143,1186,1598,1622]" captionStart="Figs 100–105" captionStartId="19.[226,263,1870,1889]" captionTargetBox="[216,1405,315,1820]" captionTargetPageId="19" captionText="Figs 100–105. Habitus of Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — after Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — after Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — after Ozerov, 2015, fig. 1; 104 — after Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — after Ozerov, 2013, fig. 19. Рис. 100–105. ОбЩий вид Scathophagidae: 100 — Acanthocnema altaica Ozerov &amp; Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov &amp; Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — по Ozerov &amp; Krivosheina, 2018a, fig. 1; 102 — по Ozerov &amp; Krivosheina, 2018b, fig. 6; 103 — по Ozerov, 2015, fig. 1; 104 — по Ozerov &amp; Krivosheina, 2014a, fig. 1; 105 — по Ozerov, 2013, fig. 19." figureDoi="http://doi.org/10.5281/zenodo.13178458" httpUri="https://zenodo.org/record/13178458/files/figure.png" pageId="2" pageNumber="97">100</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE251543DE5DF99EAE69F8D1" pageId="2" pageNumber="97" type="discussion">
<paragraph id="F95FD76EAE251543DE5DF99EAE69F8D1" blockId="2.[812,1399,1657,1801]" pageId="2" pageNumber="97">
REMARKS. Diagnosis of the genus, descriptions of species noted in
<collectingCountry id="81F797FEAE251543DEF7F973AF44F977" box="[1016,1090,1687,1711]" name="Russia" pageId="2" pageNumber="97">Russia</collectingCountry>
, a key for their identification and changes in nomenclature are given by
<bibRefCitation id="9D71AA9FAE251543D806F951AF0FF933" author="Ozerov A. L. &amp; Krivosheina M. G." pageId="2" pageNumber="97" pagination="93 - 100" refId="ref16109" refString="Ozerov A. L., Krivosheina M. G. 2018 a. A review of the genus Acanthocnema Becker, 1894 (Diptera: Scathophagidae) of Russia // Russian Entomological Journal. Vol. 27. No. 1. P. 93 - 100." type="journal article" year="2018">Ozerov &amp; Krivosheina [2018a]</bibRefCitation>
and by
<bibRefCitation id="9D71AA9FAE251543D96DF937AE17F933" author="Ozerov A. L." box="[1122,1297,1747,1771]" pageId="2" pageNumber="97" pagination="328 - 330" refId="ref15566" refString="Ozerov A. L. 2019 b. Nomenclatural and morphological notes on some Scathophagidae (Diptera) // Russian Entomological Journal. Vol. 28. No. 3. P. 328 - 330." type="book chapter" year="2019">Ozerov [2019b]</bibRefCitation>
. Currently, there are 4 species in
<collectingCountry id="81F797FEAE251543D938F915AF78F8D1" box="[1079,1150,1777,1801]" name="Russia" pageId="2" pageNumber="97">Russia</collectingCountry>
(see checklist below).
</paragraph>
</subSubSection>
</treatment>
</document>

165
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<heading id="A2176002AE2A154CDC3EFC26A994FC02" box="[305,658,961,986]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="108" reason="2">
Genus
<taxonomicName id="3EE0ACEDAE2A154CDC72FC26A994FC02" ID-CoL="5D87" authority="Zetterstedt, 1838" authorityName="Zetterstedt" authorityYear="1838" box="[381,658,961,986]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDC72FC26AAD2FC01" box="[381,468,962,985]" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
Zetterstedt, 1838
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F95FD76EAE2A154CDC5FFC3BA975FC2F" blockId="13.[305,658,961,1015]" box="[336,627,991,1015]" pageId="13" pageNumber="108">Figs 3, 15, 63, 85, 86, 116.</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE2A154CDDEDFBFEA801FAE4" pageId="13" pageNumber="108" type="diagnosis">
<paragraph id="F95FD76EAE2A154CDDEDFBFEA801FAE4" blockId="13.[189,775,1050,2063]" pageId="13" pageNumber="108">
DIAGNOSIS. Medium-sized flies (
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long). Palpus short, noticeably shorter than proboscis, with strong apical seta about as long as or longer than palpus (Fig. 3). Proepisternum bare without hairs at middle or in anterior part. Anepisternum covered with setulae usually along dorsal margin and in posterior part only, without setulae posterior to anterior spiracle. Katepisternum with one strong seta in upper posterior corner. Postmetacoxal bridge absent. Fore tibia ventrally covered with hairs, without spinules forming rows.
</paragraph>
</subSubSection>
<subSubSection id="B1FA84E5AE2A154CDDEDFAA6AA3DF7D7" pageId="13" pageNumber="108" type="discussion">
<paragraph id="F95FD76EAE2A154CDDEDFAA6A9D4FA6C" blockId="13.[189,775,1050,2063]" pageId="13" pageNumber="108">
REMARKS. One species of
<taxonomicName id="3EE0ACEDAE2A154CDF2DFAA7A97FFA82" authorityName="Zetterstedt" authorityYear="1838" box="[546,633,1347,1370]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDF2DFAA7A97FFA82" box="[546,633,1347,1370]" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
</taxonomicName>
is known in the Palearctic —
<taxonomicName id="3EE0ACEDAE2A154CDC73FA85A805FAA0" authority="Zetterstedt, 1838" authorityName="Zetterstedt" authorityYear="1838" box="[380,771,1376,1400]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE2A154CDC73FA85A939FAA0" box="[380,575,1376,1400]" italics="true" pageId="13" pageNumber="108">Leptopa filiformis</emphasis>
Zetterstedt, 1838
</taxonomicName>
. In this paper, we also consider species of the genus
<taxonomicName id="3EE0ACEDAE2A154CDDB2FA78AAE7FA6D" authority="Strobl, 1894" authorityName="Strobl" authorityYear="1894" box="[189,481,1436,1461]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDDB2FA78AA50FA6C" box="[189,342,1436,1460]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
Strobl,
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</taxonomicName>
in the genus
<taxonomicName id="3EE0ACEDAE2A154CDF7CFA79A9CCFA6C" authorityName="Zetterstedt" authorityYear="1838" box="[627,714,1437,1460]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDF7CFA79A9CCFA6C" box="[627,714,1437,1460]" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F95FD76EAE2A154CDDEDFA5FAA3DF7D7" blockId="13.[189,775,1050,2063]" pageId="13" pageNumber="108">
The description of the genus
<taxonomicName id="3EE0ACEDAE2A154CDF37FA5FA9D7FA0B" authorityName="Strobl" authorityYear="1894" box="[568,721,1467,1491]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDF37FA5FA9D7FA0B" box="[568,721,1467,1491]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
</taxonomicName>
was given by
<bibRefCitation id="9D71AA9FAE2A154CDC25FA3DAACAFA29" author="Becker T." box="[298,460,1497,1521]" pageId="13" pageNumber="108" pagination="77 - 196" refId="ref14329" refString="Becker T. 1894. Dipterologische Studien. I. Scatomyzidae // Berliner Entomologische Zeitschrift. Bd. 39. H. 1. S. 77 - 196." type="journal article" year="1894">Becker [1894]</bibRefCitation>
. In the same paper, Becker cites the differences between
<taxonomicName id="3EE0ACEDAE2A154CDF0BFA13A99BF9D7" authorityName="Strobl" authorityYear="1894" box="[516,669,1527,1551]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDF0BFA13A99BF9D7" box="[516,669,1527,1551]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
</taxonomicName>
and
<taxonomicName id="3EE0ACEDAE2A154CDFD9FA1CABEAF9F5" authorityName="Zetterstedt" authorityYear="1838" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDFD9FA1CABEAF9F5" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
</taxonomicName>
. However, Becker mistakenly believed that
<taxonomicName id="3EE0ACEDAE2A154CDFFEF9F2AA19F993" authorityName="Zetterstedt" authorityYear="1838" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE2A154CDFFEF9F2AA19F993" italics="true" pageId="13" pageNumber="108">L. filiformis</emphasis>
</taxonomicName>
does not have a long seta on the palpus, while it is present in this species. Therefore, if we use the key proposed by Becker [
<bibRefCitation id="9D71AA9FAE2A154CDCA6F98BA96BF95F" author="Becker T." box="[425,621,1647,1672]" pageId="13" pageNumber="108" pagination="77 - 196" refId="ref14329" refString="Becker T. 1894. Dipterologische Studien. I. Scatomyzidae // Berliner Entomologische Zeitschrift. Bd. 39. H. 1. S. 77 - 196." type="journal article" year="1894">Becker, 1894: 85</bibRefCitation>
, 86] to determine genera, given the fact that
<taxonomicName id="3EE0ACEDAE2A154CDF00F96AA98BF97D" authorityName="Zetterstedt" authorityYear="1838" box="[527,653,1677,1701]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE2A154CDF00F96AA98BF97D" box="[527,653,1677,1701]" italics="true" pageId="13" pageNumber="108">L. filiformis</emphasis>
</taxonomicName>
has seta on the palpus, then
<taxonomicName id="3EE0ACEDAE2A154CDC63F948AAEDF91C" authorityName="Zetterstedt" authorityYear="1838" box="[364,491,1708,1732]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE2A154CDC63F948AAEDF91C" box="[364,491,1708,1732]" italics="true" pageId="13" pageNumber="108">L. filiformis</emphasis>
</taxonomicName>
enters into the genus
<taxonomicName id="3EE0ACEDAE2A154CDFD9F948AA28F93A" authorityName="Strobl" authorityYear="1894" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDFD9F948AA28F93A" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
</taxonomicName>
. All other characters indicated by Becker for the genus
<taxonomicName id="3EE0ACEDAE2A154CDC54F90CAAF2F8D8" authorityName="Strobl" authorityYear="1894" box="[347,500,1768,1792]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDC54F90CAAF2F8D8" box="[347,500,1768,1792]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
</taxonomicName>
also correspond to
<taxonomicName id="3EE0ACEDAE2A154CDFD9F90DABEAF8C6" authorityName="Zetterstedt" authorityYear="1838" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDFD9F90DABEAF8C6" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
</taxonomicName>
.
<taxonomicName id="3EE0ACEDAE2A154CDDF3F8E3AA7BF8C6" authorityName="Zetterstedt" authorityYear="1838" box="[252,381,1798,1822]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="CB940B7CAE2A154CDDF3F8E3AA7BF8C6" box="[252,381,1798,1822]" italics="true" pageId="13" pageNumber="108">L. filiformis</emphasis>
</taxonomicName>
has enlarged vibrissal angle with a row of centrally directed setae and postgena with strong setae (
<figureCitation id="61DBCBEBAE2A154CDC0CF8A6AA53F882" box="[259,341,1858,1882]" captionStart="Figs 13–18" captionStartId="5.[227,262,1920,1939]" captionTargetBox="[269,1372,318,1903]" captionTargetPageId="5" captionText="Figs 13–18. Head of Scathophagidae, lateral view: 13 — Delina nigrita (Fallén); 14 — Hexamitocera loxocerata (Fallén); 15 — Leptopa filiformis (Zetterstedt); 16 — Megaphthalma pallida (Fallén); 17 — Micropselapha filiformis (Zetterstedt); 18 — Mixocordylura parva Ozerov. 14 — after Ozerov &amp; Krivosheina, 2015, fig. 3; 17 — after Ozerov, 2014, fig. 7; 18 — after Ozerov &amp; Krivosheina, 2012a, fig. 10. Рис. 13–18. Голова Scathophagidae, сбоку: 13 — Delina nigrita (Fallén); 14 — Hexamitocera loxocerata (Fallén); 15 — Leptopa filiformis (Zetterstedt); 16 — Megaphthalma pallida (Fallén); 17 — Micropselapha filiformis (Zetterstedt); 18 — Mixocordylura parva Ozerov. 14 — по Ozerov &amp; Krivosheina, 2015, fig. 3; 17 — по Ozerov, 2014, fig. 7; 18 — по Ozerov &amp; Krivosheina, 2012a, fig. 10." figureDoi="http://doi.org/10.5281/zenodo.13178430" httpUri="https://zenodo.org/record/13178430/files/figure.png" pageId="13" pageNumber="108">Fig. 15</figureCitation>
), which distinguishes this species from
<taxonomicName id="3EE0ACEDAE2A154CDDB2F884AAB6F8A0" box="[189,432,1888,1912]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="CB940B7CAE2A154CDDB2F884AA50F8A0" box="[189,342,1888,1912]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
species.
</taxonomicName>
We attribute these characters to the characters of the species rank and therefore propose to include the species
<taxonomicName id="3EE0ACEDAE2A154CDCEBF879A97BF86D" authorityName="Strobl" authorityYear="1894" box="[484,637,1949,1973]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDCEBF879A97BF86D" box="[484,637,1949,1973]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="3EE0ACEDAE2A154CDDB2F858AA12F80B" authorityName="Zetterstedt" authorityYear="1838" box="[189,276,1980,2003]" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDDB2F858AA12F80B" box="[189,276,1980,2003]" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
</taxonomicName>
, and the name
<taxonomicName id="3EE0ACEDAE2A154CDCCDF85FA9EDF80B" authority="Strobl, 1894" authorityName="Strobl" authorityYear="1894" box="[450,747,1979,2003]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDCCDF85FA95DF80B" box="[450,603,1979,2003]" italics="true" pageId="13" pageNumber="108">Parallelomma</emphasis>
Strobl, 1894
</taxonomicName>
is considered a new junior synonym of
<taxonomicName id="3EE0ACEDAE2A154CDF53F83EAA31F7D7" authority="Zetterstedt, 1838" authorityName="Zetterstedt" authorityYear="1838" class="Insecta" family="Scathophagidae" genus="Leptopa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="CB940B7CAE2A154CDF53F83EA9B5F829" box="[604,691,2010,2033]" italics="true" pageId="13" pageNumber="108">Leptopa</emphasis>
Zetterstedt, 1838
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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