Add updates up until 2024-08-02 21:35:03

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<mods:title id="E2D7BD54F32643ACC04181FEC05B318C">Three new species of the genus Trechus Clairville, 1806 from Sichuan (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:date id="57344E28A8F4B20CF0015340B55221C6">2021</mods:date>
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<figureCitation id="ACF12A3EFFD0D624BA6E29BD5793FA6C" box="[1048,1158,1505,1529]" captionStart-0="Fig" captionStart-1="Figs 1013" captionStartId-0="7.[226,257,2017,2036]" captionStartId-1="8.[226,263,2017,2036]" captionTargetBox-1="[191,1394,1015,1994]" captionTargetId-1="figure-472@8.[191,1398,1015,1994]" captionTargetPageId-1="8" captionText-0="Fig. 9. Habitus of Trechus sharovae sp. n., ♂, paratype. Рис. 9. ОбЩий вид Trechus sharovae sp. n., ♂, паратип." captionText-1="Figs 1013. Male genitalia of Trechus sharovae sp. n., paratypes: 10, 12 — lateral view; 11, 13 — dorsal view. Рис. 1013. Гениталии самцов Trechus sharovae sp. n., паратипы: 10, 12 — сбоку; 11, 13 — сверху." figureDoi-1="http://doi.org/10.5281/zenodo.13178260" httpUri-1="https://zenodo.org/record/13178260/files/figure.png" pageId="6" pageNumber="407">Figs 913</figureCitation>
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MATERIAL.
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,
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<collectingDate id="5030E993FFD0D624BD382AAA50A0F89C" box="[846,949,1782,1801]" pageId="6" pageNumber="407" value="2002-08-22">22.08.2002</collectingDate>
</date>
,
<collectorName id="993F536DFFD0D624BDB22AAA5724F89C" box="[964,1073,1782,1801]" pageId="6" pageNumber="407">Belousov I.</collectorName>
&amp;
<collectorName id="993F536DFFD0D624BA2F2AAA57B9F89C" box="[1113,1196,1782,1801]" pageId="6" pageNumber="407">Kabak I.</collectorName>
leg.” [
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/
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, corrected height =
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] (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD0D628BD242B7A5730FD04" lastPageId="10" lastPageNumber="411" pageId="6" pageNumber="407" type="description">
<paragraph id="347536BBFFD0D624BD242B7A50FBF785" blockId="6.[812,1399,1476,2065]" pageId="6" pageNumber="407">
DESCRIPTION. Medium-sized species, body length usually slightly less than
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, habitus robust and subparallel sided, dorsum moderately convex (
<figureCitation id="ACF12A3EFFD0D624BA0C2B0757A1F8E5" box="[1146,1204,1883,1905]" captionStart="Fig" captionStartId="7.[226,257,2017,2036]" captionText="Fig. 9. Habitus of Trechus sharovae sp. n., ♂, paratype. Рис. 9. ОбЩий вид Trechus sharovae sp. n., ♂, паратип." pageId="6" pageNumber="407">Fig. 9</figureCitation>
). Legs and antennae rather slender; middle antennomeres much longer than wide. Color of upper side rather concolorous, reddish brown, usually with darker hind part of head and disk of elytra; the suture and margins of the latter normally slightly paler, amber reddish. Legs pale amber reddish, antennae vaguely obscured from segments 34.
</paragraph>
<caption id="60B56633FFD1D625BE942BBD51F3F784" pageId="7" pageNumber="408" startId="7.[226,257,2017,2036]" targetType="figure">
<paragraph id="347536BBFFD1D625BE942BBD51C3F860" blockId="7.[226,742,2015,2065]" box="[226,726,2015,2037]" pageId="7" pageNumber="408">
Fig. 9. Habitus of
<taxonomicName id="F3CA4D38FFD1D625BFF32BBD5135F861" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[389,544,2017,2036]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="sharovae" status="sp. nov.">
<emphasis id="06BEEAA9FFD1D625BFF32BBD5135F861" box="[389,544,2017,2036]" italics="true" pageId="7" pageNumber="408">Trechus sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD1D625BC502BBD514DF861" bold="true" box="[550,600,2017,2036]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="1D8D57D2FFD1D625BC502BBD514DF861" box="[550,600,2017,2036]" pageId="7" pageNumber="408" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, paratype.
</paragraph>
<paragraph id="347536BBFFD1D625BE942BA251F3F784" blockId="7.[226,742,2015,2065]" box="[226,742,2043,2065]" pageId="7" pageNumber="408">
Рис. 9. ОбЩий вид
<taxonomicName id="F3CA4D38FFD1D625BFE12BA15127F785" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[407,562,2045,2064]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="sharovae" status="sp. nov.">
<emphasis id="06BEEAA9FFD1D625BFE12BA15127F785" box="[407,562,2045,2064]" italics="true" pageId="7" pageNumber="408">Trechus sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD1D625BC4E2BA1517FF785" bold="true" box="[568,618,2045,2064]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="1D8D57D2FFD1D625BC4E2BA1517FF785" box="[568,618,2045,2064]" pageId="7" pageNumber="408" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, паратип.
</paragraph>
</caption>
<paragraph id="347536BBFFDED62ABE942D6752BBFE29" blockId="8.[189,775,315,977]" pageId="8" pageNumber="409">Microsculpture well distinguished, consisting of isodiametric meshes on head, very transverse meshes on pronotum, and transverse lines on elytra. Pronotum with slight, elytra with distinct iridescence. Micropunctures well developed on the whole upper surface.</paragraph>
<paragraph id="347536BBFFDED62ABE942D9D524BFD74" blockId="8.[189,775,315,977]" pageId="8" pageNumber="409">Head of medium size. Frontal furrows well impressed, angularly curved in their anterior third. Parietal impression indistinct. Supraorbital setiferous pores located in lines subparallel to body axis; posterior pore more deeply foveolate, located at level clearly behind posterior edge of eye. Eyes medium-sized, evenly convex; tempora rather long, slightly convex, glabrous. Mandibles stout, evenly curved on their exterior margin. Tooth on right mandible tridentate, its shape rather variable but distal denticle always most prominent and more isolated from other denticles. Labrum with anterior margin concave.</paragraph>
<paragraph id="347536BBFFDED62ABE942EBA574BFD99" blockId="8.[189,775,315,977]" lastBlockId="8.[812,1399,315,977]" pageId="8" pageNumber="409">Pronotum moderately transverse, subconvex on disk, with maximum width clearly before its mid-length, marginally wid- er at base than at apex. Sides slightly to moderately sinuate before hind angles.Latter medium-sized, subrectangular, point- ed apically. Basal margin nearly straight, barely emarginate on sides. Anterior margin rectilinear to slightly concave; front angles rounded and barely salient. Prebasal transverse impression moderately impressed, located far from and subparallel to basal margin, sharply curved in basal foveae. The latter large and deep, often with a triangular impression in bottom. Basal surface longitudinally rugose. Preapical transverse impression vague and shallow, more distinct laterally, often with a few coarse punctures or impressions medially. Lateral groove rath- er wide, narrowed to front angles and dilated to pronotal base. Median line distinct, more deeply impressed toward base, shortened anteriorly, reaching basal margin of pronotum. Discal foveae shallow or indistinct.</paragraph>
<paragraph id="347536BBFFDED62BBD242E4D51E0FD04" blockId="8.[812,1399,315,977]" lastBlockId="9.[189,776,315,843]" lastPageId="9" lastPageNumber="410" pageId="8" pageNumber="409">Elytra ovate, subconvex, with maximum width near their mid-length, relatively narrow (as compared to other members of the group), elytra commonly rounded at apex. Humeri rounded. Preapical sinuation distinct. Two discal setiferous pores in stria 3. Preapical pore located in the apical cross, much closer to stria 2 than to stria 3, at level markedly behind anterior termination of apical striole. Apical triangle slightly elongate, its inner side normally slightly divergent posteriad or parallel to elytral suture. Angulo-apical pore approximately in middle between exterior pore and elytral suture. Umbilicate pores well aggregated, umbilicate pores 7 and 8 most spaced (compared to those within other groups). Elytral striae 15 well impressed and distinct for most of their length, though becoming increasingly shallower toward sides; all striae coarsely punctured. Stria 6 very shallow, even in its median part, stria 7 only partially discernible. Inner interspaces slightly convex, others flat. Striation on apical slope rather shallow: striae 3 and 4 joining each other at level between anterior termination of apical striole and preapical pore, striae 5 and 6 either independently directed to apical striole or disappearing posteriorly without clear connection. Striation on apical slope, although rather shallow, especially striae 5 and 6, could be summarized as follows: (5xAS|5x6xAS)(3x4)(2x3). Parascutellar pore present, parascutellar striole well developed, rather long. Apical striole sharply engraved; rather straight, directed to the site of stria 5 anteriorly and abruptly interrupted there. Lateral groove narrower than average for the group, but wider than in most other groups of the genus, margins slightly and evenly reflexed throughout. Lateral border with a distinct hook anteriorly.</paragraph>
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<paragraph id="347536BBFFDED62ABE942BBD57A9F861" blockId="8.[226,1212,2017,2065]" box="[226,1212,2017,2037]" pageId="8" pageNumber="409">
Figs 1013. Male genitalia of
<taxonomicName id="F3CA4D38FFDED62ABF9A2BBD5192F861" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[492,647,2017,2036]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="sharovae" status="sp. nov.">
<emphasis id="06BEEAA9FFDED62ABF9A2BBD5192F861" box="[492,647,2017,2036]" italics="true" pageId="8" pageNumber="409">Trechus sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFDED62ABCFB2BBD51ABF861" bold="true" box="[653,702,2017,2036]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="1D8D57D2FFDED62ABCFB2BBD51ABF861" box="[653,702,2017,2036]" pageId="8" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratypes: 10, 12 — lateral view; 11, 13 — dorsal view.
</paragraph>
<paragraph id="347536BBFFDED62ABE942BA2576AF784" blockId="8.[226,1212,2017,2065]" box="[226,1151,2045,2065]" pageId="8" pageNumber="409">
Рис. 1013. Гениталии самцов
<taxonomicName id="F3CA4D38FFDED62ABC742BA15188F785" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[514,669,2045,2064]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="sharovae" status="sp. nov.">
<emphasis id="06BEEAA9FFDED62ABC742BA15188F785" box="[514,669,2045,2064]" italics="true" pageId="8" pageNumber="409">Trechus sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFDED62ABCD52BA151C0F785" bold="true" box="[675,725,2045,2064]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="1D8D57D2FFDED62ABCD52BA151C0F785" box="[675,725,2045,2064]" pageId="8" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, паратипы: 10, 12 — сбоку; 11, 13 — сверху.
</paragraph>
</caption>
<paragraph id="347536BBFFDFD62BBE942ECA5232FCDE" blockId="9.[189,776,315,843]" pageId="9" pageNumber="410">Protibiae thick, especially in apical half, barely grooved on exterior face in distal third, their anterior surface glabrous except for its innermost apex bearing a few rudimentary hairs. Metatibiae slender, nearly straight, moderately thickened distally. Male protarsi with two basal segments markedly dilated and inwardly dentate; both segments 1 and 2 transverse.</paragraph>
<paragraph id="347536BBFFDFD62BBD242D6750F6FD04" blockId="9.[812,1399,315,844]" pageId="9" pageNumber="410">
Aedeagus relatively large (compared to body size) and thick, markedly curved at basal portion, rather bulky in middle portion (
<figureCitation id="ACF12A3EFFDFD62BBDA12D2D575AFE13" box="[983,1103,369,391]" captionStart="Figs 1013" captionStartId="8.[226,263,2017,2036]" captionTargetBox="[191,1394,1015,1994]" captionTargetId="figure-472@8.[191,1398,1015,1994]" captionTargetPageId="8" captionText="Figs 1013. Male genitalia of Trechus sharovae sp. n., paratypes: 10, 12 — lateral view; 11, 13 — dorsal view. Рис. 1013. Гениталии самцов Trechus sharovae sp. n., паратипы: 10, 12 — сбоку; 11, 13 — сверху." figureDoi="http://doi.org/10.5281/zenodo.13178260" httpUri="https://zenodo.org/record/13178260/files/figure.png" pageId="9" pageNumber="410">Figs 1013</figureCitation>
). Apical portion attenuating downward, ending with a well-developed and oblique button, adjacent portion of ventral margin markedly sinuate. Sagittal aileron missing; basal orifice slightly concave. Endophallus armature consisting of two large and heavily sclerotized sclerites typical for members of the group but their shape and position deviate compared to other species: distal margin of longer sclerite semicircular, its proximal portion markedly concave. Parameres of medium length, each bearing 45 apical setae; left paramere longer, with a ventral apophysis.
</paragraph>
<paragraph id="347536BBFFDFD62BBD242ECA57DAFC83" blockId="9.[812,1399,315,844]" pageId="9" pageNumber="410">
SEXUAL DIMORPHISM. Males differ in larger body size (on average,
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vs.
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in females, p≤0.01) and proportionally longer elytra (EL/EW, on average, 1.35 vs.
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females, p≤0.05) and longer antennae (EL/AL, on average, 1.18 vs.
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females, p≤0.05).
</paragraph>
<paragraph id="347536BBFFDFD62BBD242F475663FCD9" blockId="9.[812,1399,315,844]" pageId="9" pageNumber="410">COMPARATIVE NOTES. Externally, the species is similar to other members of the group, especially to smaller</paragraph>
<paragraph id="347536BBFFDFD62BBF372FDF5662FC0C" blockId="9.[190,1399,899,948]" box="[321,1399,899,921]" pageId="9" pageNumber="410">
Table 1. Morphometric characters of
<taxonomicName id="F3CA4D38FFDFD62BBCD92FDF5044FC0C" box="[687,849,899,921]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="06BEEAA9FFDFD62BBCD92FDF51E8FC0D" box="[687,765,899,920]" italics="true" pageId="9" pageNumber="410">Trechus</emphasis>
species.
</taxonomicName>
Abbeviations are as indicated in Material and Methods.
</paragraph>
<paragraph id="347536BBFFDFD628BEC82FC15730FD04" blockId="9.[190,1399,899,948]" lastBlockId="10.[812,1399,315,791]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
Таблица 1. Морфометрические приЗнаки видов рода
<taxonomicName id="F3CA4D38FFDFD62BBC902FC25021FC26" authorityName="Clairville" authorityYear="1806" box="[742,820,926,947]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="06BEEAA9FFDFD62BBC902FC25021FC26" box="[742,820,926,947]" italics="true" pageId="9" pageNumber="410">Trechus</emphasis>
</taxonomicName>
. СокраЩениЯ укаЗаны в раЗделе «Материал и методы». species like
<taxonomicName id="F3CA4D38FFDCD628BF472D6051BFFEC4" authority="Sciaky et Pavesi, 1995" authorityName="Sciaky et Pavesi" authorityYear="1995" box="[305,682,315,337]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="trachypachys">
<emphasis id="06BEEAA9FFDCD628BF472D6052DDFEC5" box="[305,456,315,337]" italics="true" pageId="10" pageNumber="411">T. trachypachys</emphasis>
<bibRefCitation id="505B4B4AFFDCD628BFB92D6751BFFEC4" author="Sciaky R. &amp; Pavesi M." box="[463,682,315,337]" pageId="10" pageNumber="411" pagination="23 - 32" refId="ref8125" refString="Sciaky R., Pavesi M. 1995. Nuove specie di Trechus del Sichuan (Cina) (Coleoptera, Carabidae) // Acta coleopterologica. Vol. 11. P. 23 - 32." type="book chapter" year="1995">Sciaky et Pavesi, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F3CA4D38FFDCD628BCC52D605296FEFE" authority="Deuve, 2004" authorityName="Deuve" authorityYear="2004" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="altitudinum">
<emphasis id="06BEEAA9FFDCD628BCC52D6053EDFEFE" italics="true" pageId="10" pageNumber="411">T. altitudinum</emphasis>
<bibRefCitation id="505B4B4AFFDCD628BF742D0A5296FEFE" author="Deuve T." box="[258,387,342,363]" pageId="10" pageNumber="411" pagination="215 - 234" refId="ref7956" refString="Deuve T. 2004. Nouveaux Trechidae (Trechinae et Bembidiinae) d'Afrique et d'Asie (Coleoptera, Caraboidea) // Coleopteres. Vol. 10. No. 19. P. 215 - 234." type="book chapter" year="2004">Deuve, 2004</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="F3CA4D38FFDCD628BFC92D0B53F9FE13" authority="Belousov et Kabak, 2000" authorityName="Belousov et Kabak" authorityYear="2000" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="kurbatovi">
<emphasis id="06BEEAA9FFDCD628BFC92D0B5122FEFE" box="[447,567,342,364]" italics="true" pageId="10" pageNumber="411">T. kurbatovi</emphasis>
<bibRefCitation id="505B4B4AFFDCD628BC342D0A53F9FE13" author="Belousov I. A. &amp; Kabak I. I." pageId="10" pageNumber="411" pagination="49 - 78" refId="ref7536" refString="Belousov I. A., Kabak I. I. 2000. [Contribution to the knowledge of species of the genus Trechus Clairv. (Coleoptera, Carabidae) from China] // Entomologicheskoe Obozrenie. Vol. 79. No. 1. P. 49 - 78 [in Russian, with English summary]." type="book chapter" year="2000">Belousov et Kabak, 2000</bibRefCitation>
</taxonomicName>
. From the former species which is known only from female specimens, it differs mostly in paler coloration and shape of pronotum with basal margin less oblique at sides and lateral margins less sinuate before hind angles. On the other hand, the membership of
<taxonomicName id="F3CA4D38FFDCD628BF3D2D8052CBFE65" authorityName="Sciaky et Pavesi" authorityYear="1995" box="[331,478,475,497]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="trachypachys">
<emphasis id="06BEEAA9FFDCD628BF3D2D8052CBFE65" box="[331,478,475,497]" italics="true" pageId="10" pageNumber="411">T. trachypachys</emphasis>
</taxonomicName>
in the
<taxonomicName id="F3CA4D38FFDCD628BC6F2D87516AFE65" authorityName="Sciaky et Pavesi" authorityYear="1995" box="[537,639,475,496]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="validicollis">
<emphasis id="06BEEAA9FFDCD628BC6F2D87516AFE65" box="[537,639,475,496]" italics="true" pageId="10" pageNumber="411">validicollis</emphasis>
</taxonomicName>
species group, as it was noted earlier, needs to be confirmed, see also [
<bibRefCitation id="505B4B4AFFDCD628BCCA2DAA524EFDB3" author="Belousov I. A. &amp; Kabak I. I." pageId="10" pageNumber="411" pagination="49 - 78" refId="ref7536" refString="Belousov I. A., Kabak I. I. 2000. [Contribution to the knowledge of species of the genus Trechus Clairv. (Coleoptera, Carabidae) from China] // Entomologicheskoe Obozrenie. Vol. 79. No. 1. P. 49 - 78 [in Russian, with English summary]." type="book chapter" year="2000">Belousov, Kabak, 2000</bibRefCitation>
]. Among species with known males,
<taxonomicName id="F3CA4D38FFDCD628BCC32E4D53E2FDD5" authorityName="Deuve" authorityYear="2004" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="altitudinum">
<emphasis id="06BEEAA9FFDCD628BCC32E4D53E2FDD5" italics="true" pageId="10" pageNumber="411">T. altitudinum</emphasis>
</taxonomicName>
seems to be most similar to
<taxonomicName id="F3CA4D38FFDCD628BC682E705185FDD5" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[542,656,555,577]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="sharovae" status="sp.n.">
<emphasis id="06BEEAA9FFDCD628BC682E705185FDD5" box="[542,656,555,577]" italics="true" pageId="10" pageNumber="411">T. sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFDCD628BCEC2E7751DDFDD5" bold="true" box="[666,712,555,576]" pageId="10" pageNumber="411">
<taxonomicNameLabel id="1D8D57D2FFDCD628BCEC2E7751DDFDD5" box="[666,712,555,576]" pageId="10" pageNumber="411" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Their closeness is also consistent with their neighboring distribution (approximately
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between the
<typeStatus id="EB718819FFDCD628BC432E3D514BFDE3" box="[565,606,609,630]" pageId="10" pageNumber="411">type</typeStatus>
localities). Both species share roughly the same coloration, habitus and very similar structure of the male genitalia: median lobe rather short and stout, with short apex and large endophallus armature filling up nearly all the lumen of the median lobe. However,
<taxonomicName id="F3CA4D38FFDCD628BC852E905201FD6E" authorityName="Belousov &amp; Kabak" authorityYear="2021" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="sharovae" status="sp.n.">
<emphasis id="06BEEAA9FFDCD628BC852E905201FD6E" italics="true" pageId="10" pageNumber="411">T. sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFDCD628BF6E2EBA525DFD6E" bold="true" box="[280,328,742,763]" pageId="10" pageNumber="411">
<taxonomicNameLabel id="1D8D57D2FFDCD628BF6E2EBA525DFD6E" box="[280,328,742,763]" pageId="10" pageNumber="411" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs in having the median lobe with slenderer apical portion, deeper sinuation of ventral margin before apex and more developed button-like apex; endophallus armature readily differs in the main copulatory piece with distal portion much narrowly rounded vs. broad and subangular in
<taxonomicName id="F3CA4D38FFDCD628BB552D2D507DFE35" authorityName="Deuve" authorityYear="2004" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="altitudinum">
<emphasis id="06BEEAA9FFDCD628BB552D2D507DFE35" italics="true" pageId="10" pageNumber="411">T. altitudinum</emphasis>
</taxonomicName>
.
<taxonomicName id="F3CA4D38FFDCD628BD0C2DD050E4FE35" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[890,1009,395,417]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="sharovae" status="sp.n.">
<emphasis id="06BEEAA9FFDCD628BD0C2DD050E4FE35" box="[890,1009,395,417]" italics="true" pageId="10" pageNumber="411">T. sharovae</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFDCD628BD8B2DD7573BFE35" bold="true" box="[1021,1070,395,416]" pageId="10" pageNumber="411">
<taxonomicNameLabel id="1D8D57D2FFDCD628BD8B2DD7573BFE35" box="[1021,1070,395,416]" pageId="10" pageNumber="411" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is also similar to
<emphasis id="06BEEAA9FFDCD628BA832DD05662FE35" box="[1269,1399,395,417]" italics="true" pageId="10" pageNumber="411">
<taxonomicName id="F3CA4D38FFDCD628BA832DD05666FE35" authorityName="Belousov et Kabak" authorityYear="2000" box="[1269,1395,395,417]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="kurbatovi">T. kurbatovi</taxonomicName>
.
</emphasis>
Externally, it differs from the latter species mainly in the structure of pronotum with hind angles which are smaller and less acute. The structure of the male genitalia is significantly different: median lobe more robust, ventral margin of apical portion much deeper emarginate, and the main copulatory piece more developed and complexly twisted, less elongate, with apical portion much more broadly arcuate (vs. falcate in
<taxonomicName id="F3CA4D38FFDCD628BD5A2E3D5093FDE3" authorityName="Belousov et Kabak" authorityYear="2000" box="[812,902,609,630]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="kurbatovi">
<emphasis id="06BEEAA9FFDCD628BD5A2E3D5093FDE3" box="[812,902,609,630]" italics="true" pageId="10" pageNumber="411">kurbatovi</emphasis>
</taxonomicName>
) and apical lamella more acute (broadly rounded in
<taxonomicName id="F3CA4D38FFDCD628BD5A2E275093FD05" authorityName="Belousov et Kabak" authorityYear="2000" box="[812,902,635,656]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="kurbatovi">
<emphasis id="06BEEAA9FFDCD628BD5A2E275093FD05" box="[812,902,635,656]" italics="true" pageId="10" pageNumber="411">kurbatovi</emphasis>
</taxonomicName>
) in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFDCD629BD242ECA51E4FDE2" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" type="distribution">
<paragraph id="347536BBFFDCD628BD242ECA57C4FD74" blockId="10.[812,1399,315,791]" pageId="10" pageNumber="411">
DISTRIBUTION. The species is known only from the upper valley of the Sanchagou River located north of the township of Tonghua, Li County,
<collectingRegion id="F60EF859FFDCD628BA0D2E9757D9FD74" box="[1147,1228,715,737]" country="China" name="Sichuan" pageId="10" pageNumber="411">Sichuan</collectingRegion>
.
</paragraph>
<paragraph id="347536BBFFDCD628BD242EBA5790FC83" blockId="10.[812,1399,315,791]" pageId="10" pageNumber="411">
BIONOMICS. The species was found in the forest zone at elevations from
<quantity id="F3329B5EFFDCD628BD952F5D5794FC83" box="[995,1153,769,790]" metricMagnitude="3" metricUnit="m" metricValue="3.6" metricValueMax="3.9" metricValueMin="3.3" pageId="10" pageNumber="411" unit="m" value="3600.0" valueMax="3900.0" valueMin="3300.0">3300 to 3900 m</quantity>
.
</paragraph>
<paragraph id="347536BBFFDCD629BEBB2F6052B1FE34" blockId="10.[205,1399,828,930]" lastBlockId="11.[189,775,315,417]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
Table 2. Statistically significant differences between
<taxonomicName id="F3CA4D38FFDCD628BCA02F605064FCC7" box="[726,881,828,850]" pageId="10" pageNumber="411">
<emphasis id="06BEEAA9FFDCD628BCA02F605031FCC4" box="[726,804,828,849]" italics="true" pageId="10" pageNumber="411">Trechus</emphasis>
species
</taxonomicName>
(Mann-Whitney criterion, both sexes). Abbeviations are as indicated in Material and Methods. Таблица 2. Статистически Значимые раЗличиЯ между видами рода
<taxonomicName id="F3CA4D38FFDCD628BDE32F2E50F6FC12" authorityName="Clairville" authorityYear="1806" box="[917,995,882,903]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="06BEEAA9FFDCD628BDE32F2E50F6FC12" box="[917,995,882,903]" italics="true" pageId="10" pageNumber="411">Trechus</emphasis>
</taxonomicName>
(U-критерий Манна- Уитни, оба пола). СокраЩениЯ укаЗаны в раЗделе «Материал и методы» DERIVATIO NOMINIS. We are honored to name this species in the memory of the late professor Inessa Khristianovna Sharova for her invaluable contribution to our knowledge of carabid beetles.
</paragraph>
<paragraph id="347536BBFFDDD629BE942D9C51E4FDE2" blockId="11.[189,775,448,631]" pageId="11" pageNumber="412">
<emphasis id="06BEEAA9FFDDD629BE942D9C52B0FE43" bold="true" box="[226,421,448,470]" pageId="11" pageNumber="412">Acknowledgments.</emphasis>
We are very grateful to Drs. S.-I. Uéno (Tokyo), Yu. Imura (Yokohama), A. Konstantinov (Washington), B. Březina (Prague) and A. Gitzen (Neuhofen) for their help in our research projects on Chinese
<taxonomicName id="F3CA4D38FFDDD629BECB2E77521AFDD4" authorityName="Bonelli" authorityYear="1810" box="[189,271,555,577]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
during different periods of time. We also thank Dr. A.E. Korolev (Saint-Petersburg) for his help in mounting specimens of the new
<taxonomicName id="F3CA4D38FFDDD629BFE12E3D5124FDE2" box="[407,561,609,631]" pageId="11" pageNumber="412">
<emphasis id="06BEEAA9FFDDD629BFE12E3D52F0FDE3" box="[407,485,609,630]" italics="true" pageId="11" pageNumber="412">Trechus</emphasis>
species
</taxonomicName>
collected in
<collectingCountry id="4CDD762BFFDDD629BCC62E3D51F8FDE2" box="[688,749,609,631]" name="China" pageId="11" pageNumber="412">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFDDD629BE942ECA5200FD52" pageId="11" pageNumber="412" type="description">
<paragraph id="347536BBFFDDD629BE942ECA5200FD52" blockId="11.[189,775,662,711]" pageId="11" pageNumber="412">
<emphasis id="06BEEAA9FFDDD629BE942ECA52A4FD39" bold="true" box="[226,433,662,684]" pageId="11" pageNumber="412">Competing interests</emphasis>
. The authors declare no competing interests.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,326 @@
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<emphasis id="06BEEAA9FFD2D626BF1C2D675101FEC6" box="[362,532,315,339]" italics="true" pageId="4" pageNumber="405">Trechus comma</emphasis>
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<figureCitation id="ACF12A3EFFD2D626BFD92D055105FEE5" box="[431,528,344,369]" captionStart-0="Figs 27" captionStart-1="Fig" captionStartId-0="4.[226,263,1969,1988]" captionStartId-1="5.[226,257,2017,2036]" captionTargetBox-0="[194,1396,1013,1943]" captionTargetId-0="figure-526@4.[194,1397,1013,1945]" captionTargetPageId-0="4" captionText-0="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." captionText-1="Fig. 8. Habitus of Trechus comma sp. n., ♂, paratype. Рис. 8. ОбЩий вид Trechus comma sp. n., ♂, паратип." figureDoi-0="http://doi.org/10.5281/zenodo.13178258" httpUri-0="https://zenodo.org/record/13178258/files/figure.png" pageId="4" pageNumber="405">Figs 58</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD2D626BE942DD95188FDCD" pageId="4" pageNumber="405" type="materials_examined">
<paragraph id="347536BBFFD2D626BE942DD95188FDCD" blockId="4.[189,775,387,974]" pageId="4" pageNumber="405">
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MATERIAL.
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<collectingCountry id="4CDD762BFFD2D626BFA12DD9511AFE0D" box="[471,527,389,408]" name="China" pageId="4" pageNumber="405">China</collectingCountry>
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, Belousov I. &amp;
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leg.” [
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: 12(8)
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, Belousov I. &amp;
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leg.” [
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.
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<subSubSection id="7CD06530FFD2D624BE942E0151C1F783" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405" type="description">
<paragraph id="347536BBFFD2D626BE942E015118FC86" blockId="4.[189,775,387,974]" pageId="4" pageNumber="405">
DESCRIPTION. Body ovate, markedly constricted at pronotal base, convex (
<figureCitation id="ACF12A3EFFD2D626BFD12E2452F0FD18" box="[423,485,632,654]" captionStart="Fig" captionStartId="5.[226,257,2017,2036]" captionText="Fig. 8. Habitus of Trechus comma sp. n., ♂, paratype. Рис. 8. ОбЩий вид Trechus comma sp. n., ♂, паратип." pageId="4" pageNumber="405">Fig. 8</figureCitation>
). Appendages not long, legs stout, especially femora in males. Upper side reddish brown, often with darker blackish disc of elytra; the suture, margins of elytra and occasionally head and disc of pronotum paler, reddish. Legs and antennae uniformly yellowish. Head before clypeal suture reddish brown.
</paragraph>
<paragraph id="347536BBFFD2D626BE942F4453E2FC5B" blockId="4.[189,775,387,974]" pageId="4" pageNumber="405">Microsculpture well developed on head and pronotum, though becoming shallower on disc of the latter, consisting of isodiametric meshes on median portion of head, more or less uniform transverse meshes on pronotum, and transverse shallow anastomosing lines on elytra, surface of elytra with iridescent luster. All upper side distinctly micropunctured.</paragraph>
<paragraph id="347536BBFFD2D626BD242D675645FDCE" blockId="4.[812,1399,315,977]" pageId="4" pageNumber="405">Head of medium size, eyes rather small and flat, tempora long and convex, scarcely pubescent. Frontal furrows regularly impressed, subangulate, approaching one another in middle; frons convex. Supraorbital setae located in lines, slightly convergent anteriad, posterior seta far behind posterior margin of eye. Pore of anterior supraorbital seta markedly foveolate that of posterior seta slightly so. Mandibles stout, evenly curved. Tooth on right mandible tridentate, its shape rather variable, distal and proximal denticles always most prominent, with deepest notch between median and distal denticles. Labrum with anterior margin deeply concave.</paragraph>
<paragraph id="347536BBFFD2D624BD242E3D501DFEF9" blockId="4.[812,1399,315,977]" lastBlockId="6.[189,776,315,2071]" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405">Pronotum transverse, subconvex, with maximum width clearly before middle, markedly contracted at base. Its sides evenly and broadly arcuate, barely sinuate before hind angles, latter rather small but distinct, subrectangular. Posterior margin straight medially, obliquely truncate laterally; anteri- or margin straight. Anterior angles marked but not salient. Lateral margins narrowly bordered and slightly reflexed, lateral groove of average width, slightly expanded posteriorly. Prebasal transverse impression shallow, vaguely delimit- ed, angularly curved in basal foveae, the latter rather small and shallow. Apical transverse impression irregular, occasionally interrupted medially.Discal foveae missing or slightly impressed. Anterior marginal seta placed in about anterior third of pronotum, posterior seta in hind angle. Base of pronotum distinctly rugose. Median line distinct, slightly impressed, deepest at level of prebasal transverse impression.</paragraph>
<caption id="60B56633FFD2D626BE942BED5710F784" ID-DOI="http://doi.org/10.5281/zenodo.13178258" ID-Zenodo-Dep="13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="4" pageNumber="405" startId="4.[226,263,1969,1988]" targetBox="[194,1396,1013,1943]" targetPageId="4" targetType="figure">
<paragraph id="347536BBFFD2D626BE942BED50BDF848" blockId="4.[189,1399,1969,2065]" pageId="4" pageNumber="405">
Figs 27. Male genitalia of
<taxonomicName id="F3CA4D38FFD2D626BFB92BED512AF851" box="[463,575,1969,1988]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="06BEEAA9FFD2D626BFB92BED5101F851" box="[463,532,1969,1988]" italics="true" pageId="4" pageNumber="405">Trechus</emphasis>
<emphasis id="06BEEAA9FFD2D626BC6C2BED512AF851" bold="true" box="[538,575,1969,1988]" pageId="4" pageNumber="405">spp.</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD2D626BC332BED5143F851" bold="true" box="[581,598,1969,1988]" pageId="4" pageNumber="405">n.</emphasis>
: 24 —
<taxonomicName id="F3CA4D38FFD2D626BCD72BEE5019F851" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[673,780,1970,1988]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="xiongmao" status="sp. n.">
<emphasis id="06BEEAA9FFD2D626BCD72BEE5019F851" box="[673,780,1970,1988]" italics="true" pageId="4" pageNumber="405">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD2D626BD672BED5054F851" bold="true" box="[785,833,1969,1988]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="1D8D57D2FFD2D626BD672BED5054F851" box="[785,833,1969,1988]" pageId="4" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype; 57 —
<taxonomicName id="F3CA4D38FFD2D626BD942BEE5723F851" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[994,1078,1970,1988]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="comma" status="sp. n.">
<emphasis id="06BEEAA9FFD2D626BD942BEE5723F851" box="[994,1078,1970,1988]" italics="true" pageId="4" pageNumber="405">T. comma</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD2D626BA4D2BED577EF851" bold="true" box="[1083,1131,1969,1988]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="1D8D57D2FFD2D626BA4D2BED577EF851" box="[1083,1131,1969,1988]" pageId="4" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex.
</paragraph>
<paragraph id="347536BBFFD2D626BE942BBA5710F784" blockId="4.[189,1399,1969,2065]" pageId="4" pageNumber="405">
Рис. 27. Гениталии самцов новых видов рода
<taxonomicName id="F3CA4D38FFD2D626BCEE2BB951C8F86D" authorityName="Clairville" authorityYear="1806" box="[664,733,2021,2040]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="06BEEAA9FFD2D626BCEE2BB951C8F86D" box="[664,733,2021,2040]" italics="true" pageId="4" pageNumber="405">Trechus</emphasis>
</taxonomicName>
: 24 —
<taxonomicName id="F3CA4D38FFD2D626BD5A2BBA508CF86D" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[812,921,2022,2040]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="xiongmao" status="sp. n.">
<emphasis id="06BEEAA9FFD2D626BD5A2BBA508CF86D" box="[812,921,2022,2040]" italics="true" pageId="4" pageNumber="405">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD2D626BDE92BB950C5F86D" bold="true" box="[927,976,2021,2040]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="1D8D57D2FFD2D626BDE92BB950C5F86D" box="[927,976,2021,2040]" pageId="4" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, паратип; 57 —
<taxonomicName id="F3CA4D38FFD2D626BA002BBA57D9F86D" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[1142,1228,2022,2040]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="comma" status="sp. n.">
<emphasis id="06BEEAA9FFD2D626BA002BBA57D9F86D" box="[1142,1228,2022,2040]" italics="true" pageId="4" pageNumber="405">T. comma</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD2D626BAA42BB95616F86D" bold="true" box="[1234,1283,2021,2040]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="1D8D57D2FFD2D626BAA42BB95616F86D" box="[1234,1283,2021,2040]" pageId="4" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры.
</paragraph>
</caption>
<caption id="60B56633FFD3D627BE942BBD51C1F784" pageId="5" pageNumber="406" startId="5.[226,257,2017,2036]" targetType="figure">
<paragraph id="347536BBFFD3D627BE942BBD51D1F860" blockId="5.[226,724,2015,2065]" box="[226,708,2015,2037]" pageId="5" pageNumber="406">
Fig. 8. Habitus of
<taxonomicName id="F3CA4D38FFD3D627BFF22BBD511BF861" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[388,526,2017,2036]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="comma" status="sp. nov.">
<emphasis id="06BEEAA9FFD3D627BFF22BBD511BF861" box="[388,526,2017,2036]" italics="true" pageId="5" pageNumber="406">Trechus comma</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD3D627BC632BBD5153F861" bold="true" box="[533,582,2017,2036]" pageId="5" pageNumber="406">
<taxonomicNameLabel id="1D8D57D2FFD3D627BC632BBD5153F861" box="[533,582,2017,2036]" pageId="5" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, paratype.
</paragraph>
<paragraph id="347536BBFFD3D627BE942BA251C1F784" blockId="5.[226,724,2015,2065]" box="[226,724,2043,2065]" pageId="5" pageNumber="406">
Рис. 8. ОбЩий вид
<taxonomicName id="F3CA4D38FFD3D627BFE02BA15135F785" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[406,544,2045,2064]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="comma" status="sp. nov.">
<emphasis id="06BEEAA9FFD3D627BFE02BA15135F785" box="[406,544,2045,2064]" italics="true" pageId="5" pageNumber="406">Trechus comma</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD3D627BC512BA1514DF785" bold="true" box="[551,600,2045,2064]" pageId="5" pageNumber="406">
<taxonomicNameLabel id="1D8D57D2FFD3D627BC512BA1514DF785" box="[551,600,2045,2064]" pageId="5" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, паратип.
</paragraph>
</caption>
<paragraph id="347536BBFFD0D624BE942D2D5203FBB4" blockId="6.[189,776,315,2071]" pageId="6" pageNumber="407">
Elytra oval, with evenly arcuate sides, widest just after their mid-length. Humeri broadly rounded, prehumeral margin evenly arcuate, posthumeral sinuation indistinct. Elytral apex broadly rounded, nearly truncate, with a distinct preapical sinuation. Elytral striae rather deep and regular, even stria 7 partially visible. Intervals 34 subconvex, others flat, interval 2 distinctly wider than interval
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apical portion of elytra. All elytral striae finely punctured and slightly undulate. Stria 2 surpassing level of preapical pore, with or without distinct connection with stria 3, latter joining stria 4 at level clearly behind anterior termination of apical striole. Striae 5 and 6 joining each other slightly behind median group of umbilicate series or near that level; stria 7 without clear connection with any of other striae. Both parascutellar striole and parascutellar seta present. Apical recurrent striole of average length, deep and straight anteriorly, usually surpassing level of mid-distance between umbilicate pores 7 and 8 anteriad, joining fused stria (5x6) anteriorly. Striation on apical slope as follows: ((5x6)xAS)(3x4)(2x3). Apical triangle usually subequilateral, inner sides of apical triangles of both elytra subparallel to body axis. Preapical pore placed at level of umbilicate pore 8 or slightly behind. Angulo-apical pore usually set closer to exterior pore than to elytral suture. Lateral margins bordered and narrowly reflexed in posterior part while moderately reflexed in anterior part.
</paragraph>
<paragraph id="347536BBFFD0D624BE94287A513CFBE4" blockId="6.[189,776,315,2071]" pageId="6" pageNumber="407">Protibiae depressed and slightly grooved on exterior surface in their median part; two basal segments of male protarsi dilated, markedly transverse.</paragraph>
<paragraph id="347536BBFFD0D624BE94282A5184FA02" blockId="6.[189,776,315,2071]" pageId="6" pageNumber="407">
Median lobe of aedeagus (
<figureCitation id="ACF12A3EFFD0D624BF85282A5144FB1E" box="[499,593,1142,1164]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="6" pageNumber="407">Figs 5, 7</figureCitation>
) of medium size, rather thick, angularly curved at basal third, with ventral side nearly straight for most of its length, apical portion distinctly attenuates downwards and gradually narrowing toward small and button-like apex in lateral view. Apical lamella subparallel sided, with apical margin broadly rounded in dorsal view. Sagittal aileron small. Parameres of average size, their apex variable: rounded, oblique or truncate (
<figureCitation id="ACF12A3EFFD0D624BC4E296D5184FAD3" box="[568,657,1328,1351]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="6" pageNumber="407">Figs 56</figureCitation>
). Endophallus armature S-shaped in dorsal view and somewhat triangular shaped in lateral view; one poorly defined scaly area located near left dorsal wall of the median lobe.
</paragraph>
<paragraph id="347536BBFFD0D624BE9429C75112F9C4" blockId="6.[189,776,315,2071]" pageId="6" pageNumber="407">
SEXUAL DIMORPHISM. Males differ in larger body size (on average,
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vs.
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in females, p≤0.01), proportionally longer elytra (EL/EW, on average, 1.34 vs.
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females, p≤0.001; EL/PL, on average, 2.77 vs.
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females, p≤0.05; EL/TiL, on average, 2.39 vs.
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females, p≤0.05) and narrower pronotum (PW/PL, on average, 1.38 vs.
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females, p≤0.05).
</paragraph>
<paragraph id="347536BBFFD0D624BE942A0A51C1F783" blockId="6.[189,776,315,2071]" pageId="6" pageNumber="407">
COMPARATIVE NOTES.
<taxonomicName id="F3CA4D38FFD0D624BF9B2A0B515EF9F9" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[493,587,1623,1644]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="comma" status="sp.n.">
<emphasis id="06BEEAA9FFD0D624BF9B2A0B515EF9F9" box="[493,587,1623,1644]" italics="true" pageId="6" pageNumber="407">T. comma</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD0D624BC262A0A5194F9FE" bold="true" box="[592,641,1622,1643]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="1D8D57D2FFD0D624BC262A0A5194F9FE" box="[592,641,1622,1643]" pageId="6" pageNumber="407" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is most closely related to neighboring
<taxonomicName id="F3CA4D38FFD0D624BFB72A2D5128F913" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[449,573,1649,1670]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="xiongmao" status="sp.n.">
<emphasis id="06BEEAA9FFD0D624BFB72A2D5128F913" box="[449,573,1649,1670]" italics="true" pageId="6" pageNumber="407">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD0D624BC312A2D516DF913" bold="true" box="[583,632,1649,1670]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="1D8D57D2FFD0D624BC312A2D516DF913" box="[583,632,1649,1670]" pageId="6" pageNumber="407" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
but differs in having larger size, flatter and longer eyes, less transverse pronotum with sides more deeply sinuate before larger hind angles, which are more produced, basal margin narrower, basal surface more coarsely rugose; elytra more elongate; apical recurrent striole longer, surpassing level of mid-distance between umbilicate pores 7 and 8; lateral margins of both pronotum and elytra less markedly reflexed (the differences in morphometric characters are shown in Table 2). The male genitalia of the new species differ from those of allied taxa in their rather thick shape (
<figureCitation id="ACF12A3EFFD0D624BF912B27510AF804" box="[487,543,1915,1937]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="6" pageNumber="407">Fig. 5</figureCitation>
) and transverse triangular contour of the main copulatory piece in the endophallus armature (in lateral view). Nonetheless, the principal structure of the endophallus armature is similar to that of related species like
<taxonomicName id="F3CA4D38FFD0D624BF472BBB52BCF869" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[305,425,2023,2044]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="xiongmao" status="sp.n.">
<emphasis id="06BEEAA9FFD0D624BF472BBB52BCF869" box="[305,425,2023,2044]" italics="true" pageId="6" pageNumber="407">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD0D624BFD82BBA52CAF86E" bold="true" box="[430,479,2022,2043]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="1D8D57D2FFD0D624BFD82BBA52CAF86E" box="[430,479,2022,2043]" pageId="6" pageNumber="407" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="ACF12A3EFFD0D624BF9D2BBA5131F869" box="[491,548,2022,2044]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="6" pageNumber="407">Fig. 5</figureCitation>
vs.
<figureCitation id="ACF12A3EFFD0D624BC3C2BBA5191F86E" box="[586,644,2022,2044]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="6" pageNumber="407">Fig. 2</figureCitation>
),
<taxonomicName id="F3CA4D38FFD0D624BCE32BBB52CDF783" authority="Deuve et Queinnec, 1993" authorityName="Deuve et Queinnec" authorityYear="1993" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="06BEEAA9FFD0D624BCE32BBB53C3F783" italics="true" pageId="6" pageNumber="407">T. gansuensis</emphasis>
<bibRefCitation id="505B4B4AFFD0D624BEAB245D52CDF783" author="Deuve T. &amp; Queinnec E." box="[221,472,2049,2071]" pageId="6" pageNumber="407" pagination="1 - 9" refId="ref8082" refString="Deuve T., Queinnec E. 1993. Nouveaux Trechus du Qinghai, du Sichuan et du Gansu, Chine (Coleoptera, Caraboidea: Trechidae) // Opuscula Zoologica Fluminensia. Vol. 104. P. 1 - 9." type="journal article" year="1993">Deuve et Quéinnec, 1993</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="F3CA4D38FFD0D624BC7B245D51C5F783" authority="Deuve, 1992" authorityName="Deuve" authorityYear="1992" box="[525,720,2049,2070]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="xiei">
<emphasis id="06BEEAA9FFD0D624BC7B245D515FF783" box="[525,586,2049,2070]" italics="true" pageId="6" pageNumber="407">T. xiei</emphasis>
<bibRefCitation id="505B4B4AFFD0D624BC27245D51C5F783" author="Deuve T." box="[593,720,2049,2070]" pageId="6" pageNumber="407" pagination="171 - 184" refId="ref7922" refString="Deuve T. 1992. Contribution a la connaissance des Trechidae asiatiques (Coleoptera) // Bulleten de la Societe entomologique de France. Vol. 97. P. 171 - 184." type="book chapter" year="1992">Deuve, 1992</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD0D624BD242D675741FE64" pageId="6" pageNumber="407" type="distribution">
<paragraph id="347536BBFFD0D624BD242D675636FE12" blockId="6.[812,1398,315,577]" pageId="6" pageNumber="407">
DISTRIBUTION. The species is known only from the upper valley of the Sanchagou River located north of the township of Tonghua, Wenchuan County,
<collectingRegion id="F60EF859FFD0D624BAB82D2D560AFE12" box="[1230,1311,369,391]" country="China" name="Sichuan" pageId="6" pageNumber="407">Sichuan</collectingRegion>
.
</paragraph>
<paragraph id="347536BBFFD0D624BD242DD75741FE64" blockId="6.[812,1398,315,577]" pageId="6" pageNumber="407">
BIONOMICS. The species occurs in the upper forest and subalpine zones, at elevations of
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. It was found mostly in the forest litter and along banks of mountain streams in the subalpine zone.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD0D624BD242DAA508AFDD4" pageId="6" pageNumber="407" type="description">
<paragraph id="347536BBFFD0D624BD242DAA508AFDD4" blockId="6.[812,1398,315,577]" pageId="6" pageNumber="407">DERIVATIO NOMINIS. The species epithet derives from the particular shape of the copulatory piece resembling the comma.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,363 @@
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<mods:title id="775023B7F4B8D76B9347816B6A3F32CB">Three new species of the genus Trechus Clairville, 1806 from Sichuan (Coleoptera: Carabidae: Trechini)</mods:title>
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<taxonomicName id="F3CA4D38FFD7D623BDB92A9C5783F94D" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[975,1174,1728,1752]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="xiongmao" status="sp. nov.">
<emphasis id="06BEEAA9FFD7D623BDB92A9C5783F94D" box="[975,1174,1728,1752]" italics="true" pageId="1" pageNumber="402">Trechus xiongmao</emphasis>
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.
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MATERIAL.
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, “
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,
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,
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of
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<collectingDate id="5030E993FFD7D623BD5A2B6E5085F8D0" box="[812,912,1842,1861]" pageId="1" pageNumber="402" value="2002-07-23">23.07.2002</collectingDate>
</date>
,
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&amp;
<collectorName id="993F536DFFD7D623BA582B6E576AF8D3" box="[1070,1151,1842,1862]" pageId="1" pageNumber="402">Kabak I.</collectorName>
leg.” [
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].
<emphasis id="06BEEAA9FFD7D623BDF92B1650FEF8C8" bold="true" box="[911,1003,1866,1885]" pageId="1" pageNumber="402">
<typeStatus id="EB718819FFD7D623BDF92B1650FEF8C8" box="[911,1003,1866,1885]" pageId="1" pageNumber="402" type="paratype">Paratypes</typeStatus>
</emphasis>
: 14(7)
</materialsCitation>
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<specimenCount id="22CCFD32FFD7D623BA5D2B145746F8CB" box="[1067,1107,1864,1886]" count="1" pageId="1" pageNumber="402" type="male">♂♂</specimenCount>
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, collected with holotype (
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,
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,
<collectionCode id="52DBAE7EFFD7D623BDB42B3F50E1F8E3" box="[962,1012,1891,1910]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="1" pageNumber="402">NME</collectionCode>
,
<collectionCode id="52DBAE7EFFD7D623BA772B3F5727F8E3" box="[1025,1074,1891,1910]" country="France" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="1" pageNumber="402" type="Herbarium">MPU</collectionCode>
,
<collectionCode id="52DBAE7EFFD7D623BA492B3E5769F8E0" box="[1087,1148,1890,1909]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="402">SMNS</collectionCode>
,
<collectionCode id="52DBAE7EFFD7D623BAFE2B3E57A3F8E0" box="[1160,1206,1890,1909]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="402" type="Museum">ZSM</collectionCode>
,
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,
<collectionCode id="52DBAE7EFFD7D623BB762B3E563AF8E0" box="[1280,1327,1890,1909]" pageId="1" pageNumber="402">CAK</collectionCode>
,
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); 1f
</materialsCitation>
<materialsCitation id="84A23CE6FFD7D623BD482B245779F828" collectingDate="2002-07-22" collectionCode="NW, CBK" collectorName="Belousov I. &amp; Kabak I." country="China" elevation="3350" latitude="31.702223" location="Shangmeng" longLatPrecision="20" longitude="103.039444" pageId="1" pageNumber="402" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
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, “China,
<collectionCode id="52DBAE7EFFD7D623BDD02B2750DFF81B" box="[934,970,1915,1934]" pageId="1" pageNumber="402">NW</collectionCode>
<collectingRegion id="F60EF859FFD7D623BDA62B26570DF818" box="[976,1048,1914,1933]" country="China" name="Sichuan" pageId="1" pageNumber="402">Sichuan</collectingRegion>
,
<locationDeviation id="25C72F99FFD7D623BA572B275788F818" box="[1057,1181,1914,1934]" location="Lixian" pageId="1" pageNumber="402">
<collectionCode id="52DBAE7EFFD7D623BA572B275750F81B" box="[1057,1093,1915,1934]" pageId="1" pageNumber="402">NW</collectionCode>
of Lixian
</locationDeviation>
,
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WNW of
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, forest
<quantity id="F3329B5EFFD7D623BDD02BCE50F8F833" box="[934,1005,1938,1958]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="1" pageNumber="402" unit="m" value="3350.0">
<elevation id="BFE7D188FFD7D623BDD02BCE50F8F833" box="[934,1005,1938,1958]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="1" pageNumber="402" unit="m" value="3350.0">3350 m</elevation>
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,
<date id="4074107BFFD7D623BD8C2BCE5748F830" box="[1018,1117,1938,1957]" pageId="1" pageNumber="402" value="2002-07-22">
<collectingDate id="5030E993FFD7D623BD8C2BCE5748F830" box="[1018,1117,1938,1957]" pageId="1" pageNumber="402" value="2002-07-22">22.07.2002</collectingDate>
</date>
,
<collectorName id="993F536DFFD7D623BA1C2BCE57C7F833" box="[1130,1234,1938,1958]" pageId="1" pageNumber="402">Belousov I.</collectorName>
&amp;
<collectorName id="993F536DFFD7D623BA832BCE5651F833" box="[1269,1348,1938,1958]" pageId="1" pageNumber="402">Kabak I.</collectorName>
leg.” [
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/
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] (
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)
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;
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<specimenCount id="22CCFD32FFD7D623BA032BF65799F82B" box="[1141,1164,1960,1982]" count="1" pageId="1" pageNumber="402" type="female">1♀</specimenCount>
, “China
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,
<locationDeviation id="25C72F99FFD7D623BB242BF75094F840" location="Lixian" pageId="1" pageNumber="402">
<collectionCode id="52DBAE7EFFD7D623BB242BF75663F82B" box="[1362,1398,1963,1982]" pageId="1" pageNumber="402">NW</collectionCode>
of Lixian
</locationDeviation>
,
<quantity id="F3329B5EFFD7D623BDF82B9E50D2F840" box="[910,967,1986,2005]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="402" unit="km" value="10.0">10 km</quantity>
WSW of
<location id="31156060FFD7D623BA5E2B9E5785F840" LSID="urn:lsid:plazi:treatment:BC6387ADFFD7D621BDB92A9C5662F783:31156060FFD7D623BA5E2B9E5785F840" box="[1064,1168,1986,2005]" country="China" latitude="31.66111" longLatPrecision="20" longitude="103.030556" name="Shangmeng" pageId="1" pageNumber="402" stateProvince="Sichuan">Shangmeng</location>
, alp., scree
<quantity id="F3329B5EFFD7D623BA892B9E5666F840" box="[1279,1395,1986,2005]" metricMagnitude="3" metricUnit="m" metricValue="3.925" metricValueMax="4.0" metricValueMin="3.85" pageId="1" pageNumber="402" unit="m" value="3925.0" valueMax="4000.0" valueMin="3850.0">
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,
<date id="4074107BFFD7D623BD5A2B86508CF878" box="[812,921,2010,2029]" pageId="1" pageNumber="402" value="2002-07-25">
<collectingDate id="5030E993FFD7D623BD5A2B86508CF878" box="[812,921,2010,2029]" pageId="1" pageNumber="402" value="2002-07-25">25.07.2002</collectingDate>
</date>
,
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&amp;
<collectorName id="993F536DFFD7D623BA312B86578BF87B" box="[1095,1182,2010,2030]" pageId="1" pageNumber="402">Kabak I.</collectorName>
leg.” [
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/
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] (
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)
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.
</paragraph>
</subSubSection>
<caption id="60B56633FFD4D620BE942BBD51F9F785" pageId="2" pageNumber="403" startId="2.[226,257,2017,2036]" targetType="figure">
<paragraph id="347536BBFFD4D620BE942BBD51C9F860" blockId="2.[226,748,2015,2064]" box="[226,732,2015,2037]" pageId="2" pageNumber="403">
Fig. 1. Habitus of
<taxonomicName id="F3CA4D38FFD4D620BFF32BBD5133F861" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[389,550,2017,2036]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="xiongmao" status="sp. nov.">
<emphasis id="06BEEAA9FFD4D620BFF32BBD5133F861" box="[389,550,2017,2036]" italics="true" pageId="2" pageNumber="403">Trechus xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD4D620BC5A2BBD514BF861" bold="true" box="[556,606,2017,2036]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="1D8D57D2FFD4D620BC5A2BBD514BF861" box="[556,606,2017,2036]" pageId="2" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, paratype.
</paragraph>
<paragraph id="347536BBFFD4D620BE942BA151F9F785" blockId="2.[226,748,2015,2064]" box="[226,748,2042,2064]" pageId="2" pageNumber="403">
Рис. 1. ОбЩий вид
<taxonomicName id="F3CA4D38FFD4D620BFE12BA1512DF785" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[407,568,2045,2064]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="xiongmao" status="sp. nov.">
<emphasis id="06BEEAA9FFD4D620BFE12BA1512DF785" box="[407,568,2045,2064]" italics="true" pageId="2" pageNumber="403">Trechus xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD4D620BC492BA15165F785" bold="true" box="[575,624,2045,2064]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="1D8D57D2FFD4D620BC492BA15165F785" box="[575,624,2045,2064]" pageId="2" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂, паратип.
</paragraph>
</caption>
<subSubSection id="7CD06530FFD5D621BE942D6757E4F942" pageId="3" pageNumber="404" type="description">
<paragraph id="347536BBFFD5D621BE942D6751B2FD99" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="404">
DESCRIPTION. Small-sized species with body length usually less than
<quantity id="F3329B5EFFD5D621BF192D0A52BAFEFE" box="[367,431,342,363]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="404" unit="mm" value="3.0">3 mm</quantity>
. Body broad ovate, constricted at pronotal base, very convex. Appendages not long, legs stout (
<figureCitation id="ACF12A3EFFD5D621BEB22DD753EEFE35" box="[196,251,395,417]" captionStart="Fig" captionStartId="2.[226,257,2017,2036]" captionText="Fig. 1. Habitus of Trechus xiongmao sp. n., ♂, paratype. Рис. 1. ОбЩий вид Trechus xiongmao sp. n., ♂, паратип." pageId="3" pageNumber="404">Fig. 1</figureCitation>
). Upper side usually dark brownish, suture, margins of elytra, and occasionally disc of pronotum pale reddish; more rarely whole dorsum amber brownish. Head before clypeal suture much paler than behind, boundary not sharp. Legs yellowish. Antennae concolorous, reddish yellow.
</paragraph>
<paragraph id="347536BBFFD5D621BE942E4D52E5FD74" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="404">Microsculpture well developed over entire surface of both head and pronotum, though becoming shallower on disc of the latter, consisting of isodiametric meshes on disc of head, slightly transverse meshes on anterior portion of pronotum, and markedly transverse meshes on its posterior portion; microsculpture of elytra shallow, consisting of transverse anastomosing lines, surface of elytra with iridescent luster. All upper side micropunctured.</paragraph>
<paragraph id="347536BBFFD5D621BE942EBA51ADFC7E" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="404">Head of medium size, eyes rather small and flat, tempora long and convex, with a few very short hairs. Frontal furrows regularly impressed, subangulate and approaching one another before middle; frons convex. Supraorbital setae located in lines, distinctly convergent anteriad, posterior seta far behind posterior margin of eye. Pores of both anterior and posterior setae foveolate, though posterior pore less deeply impressed. Mandibles stout, evenly curved. Tooth on right mandible tridentate, its shape rather variable but distal denticle always most prominent. Anterior margin of labrum concave.</paragraph>
<paragraph id="347536BBFFD5D621BE942FAD51BAFA72" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="404">Pronotum markedly transverse, convex, with maximum width in apical third, clearly contracted to base, approximately as wide at base as at apex. Its sides broadly arcuate in anterior part, then nearly straight and convergent, with very shallow or no sinuation before hind angles, latter obtuse, blunt, or sharp at apex, always distinct. Posterior margin straight or slightly convex medially, obliquely truncate laterally; anterior margin straight. Anterior angles marked but not salient. Lateral margins bordered and reflexed, lateral groove of average width, clearly expanded posteriorly. Prebasal transverse impression shallow, vaguely delimited, angularly curved in basal foveae, the latter are of average size. Apical transverse impression continuous, well engraved laterally, becoming shallower or even interrupted medially. Discal foveae missing or slightly impressed. Anterior marginal seta placed in about anterior third of pronotum, posterior seta — in hind angle. Basal surface of pronotum distinctly rugose; wrinkles becoming sharper near basal margin. Median line regular, slightly impressed, becoming deeper near prebasal impression.</paragraph>
<paragraph id="347536BBFFD5D621BE9429B7571BFE64" blockId="3.[189,776,315,2071]" lastBlockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
Elytra broad oval, with arcuate sides, widest just after their middle. Humeri broadly rounded, prehumeral margin evenly arcuate, posthumeral sinuation indistinct. Elytral apex broadly rounded, nearly truncate, with a distinct preapical sinuation. Elytral striae rather deep and regular, nearly entire, even stria 7 visible in its middle portion. Intervals 34 rather convex, interval 2 about 1.5 times as wide as interval
<quantity id="F3329B5EFFD5D621BCA92AD75012F934" box="[735,775,1675,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="404" unit="in" value="1.0">1 in</quantity>
their apical third. All elytral striae finely punctured and slightly undulate. Stria 2 surpassing level of preapical pore, occasionally redoubled behind it, often without clear connection with stria 3, latter joining stria 4 at level slightly behind anterior end of apical striole. Striae 5 and 6 joining each other slightly behind median group of umbilicate series or near that level; stria 7 without clear connection with any of other striae. Parascutellar striole and parascutellar seta present. Apical recurrent striole of average length, rather deep and straight, usually not reaching level of mid-distance between umbilicate pores 7 and 8, joining merged stria (5x6) anteriorly. The dominant pattern of striation on apical slope of elytra may be described as follows: ((5x6)xAS)(3x4)(2x3) where each next conjunction x is located closer to the elytral apex. Apical triangle (consisting of three pores on the apical slope of each elytron, including the preapical pore) usually slightly elongate, its inner side convergent with suture anteriorly. Preapical pore set at level clearly behind anterior termination of apical striole. Angulo-apical pore usually placed closer to exterior pore than to elytral suture. Lateral margins bordered and markedly reflexed.
</paragraph>
<paragraph id="347536BBFFD5D621BD242DAA5662FDB3" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">Protibiae flattened and grooved on exterior surface; two basal segments of male protarsi dilated, markedly transverse.</paragraph>
<paragraph id="347536BBFFD5D621BD242E77564AFCF2" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
Median lobe of aedeagus of average size, subcylindrical, sharply curved at basal third, with straight ventral margin and small apical dilation, apical lamella rather large, its sides gradually narrowed toward apex in lateral view (
<figureCitation id="ACF12A3EFFD5D621BB792E27567CFD05" box="[1295,1385,635,657]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="3" pageNumber="404">Figs 2, 4</figureCitation>
); and nearly parallel sided in dorsal view (
<figureCitation id="ACF12A3EFFD5D621BAA02ECA5601FD3E" box="[1238,1300,662,684]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
). Sagittal aileron of medium size. Parameres of average length, straight in apical half, left one clearly longer, with a well-developed ventral apophysis; each paramere bearing 4 apical setae (
<figureCitation id="ACF12A3EFFD5D621BB392EBA502FFC83" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="3" pageNumber="404">Fig. 2</figureCitation>
). Endophallus armature is similar to that of
<taxonomicName id="F3CA4D38FFD5D621BA872F5D5727FCA5" authority="Deuve et Queinnec, 1993" authorityName="Deuve et Queinnec" authorityYear="1993" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="06BEEAA9FFD5D621BA872F5D5663FC83" box="[1265,1398,769,790]" italics="true" pageId="3" pageNumber="404">T. gansuensis</emphasis>
<bibRefCitation id="505B4B4AFFD5D621BD5A2F405727FCA5" author="Deuve T. &amp; Queinnec E." box="[812,1074,795,817]" pageId="3" pageNumber="404" pagination="1 - 9" refId="ref8082" refString="Deuve T., Queinnec E. 1993. Nouveaux Trechus du Qinghai, du Sichuan et du Gansu, Chine (Coleoptera, Caraboidea: Trechidae) // Opuscula Zoologica Fluminensia. Vol. 104. P. 1 - 9." type="journal article" year="1993">Deuve et Quéinnec, 1993</bibRefCitation>
</taxonomicName>
and related taxa, consisting of poorly defined sclerite, S-shaped in dorsal view, and one small scaly area located near left wall of the median lobe.
</paragraph>
<paragraph id="347536BBFFD5D621BD242F3757DAFC79" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
SEXUAL DIMORPHISM. Males differ in smaller head (PW/HW, on average,
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vs.
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in females, p≤0.01), larger eyes (YL/TL, on average, 1.34 vs.
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females, p≤0.05), and less transverse pronotum (PW/PL, on average, 1.41 vs.
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females, p≤0.05).
</paragraph>
<paragraph id="347536BBFFD5D621BD242FAD5756FB4E" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
COMPARATIVE NOTES.
<taxonomicName id="F3CA4D38FFD5D621BA142FAD57C9FB93" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[1122,1244,1009,1030]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="xiongmao" status="sp.n.">
<emphasis id="06BEEAA9FFD5D621BA142FAD57C9FB93" box="[1122,1244,1009,1030]" italics="true" pageId="3" pageNumber="404">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD5D621BA922FAD5600FB93" bold="true" box="[1252,1301,1009,1030]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="1D8D57D2FFD5D621BA922FAD5600FB93" box="[1252,1301,1009,1030]" pageId="3" pageNumber="404" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="F3CA4D38FFD5D621BD302850571DFBB5" authority="Deuve, 1992" authorityName="Deuve" authorityYear="1992" box="[838,1032,1035,1057]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="xiei">
<emphasis id="06BEEAA9FFD5D621BD3028505096FBB4" box="[838,899,1036,1057]" italics="true" pageId="3" pageNumber="404">T. xiei</emphasis>
<bibRefCitation id="505B4B4AFFD5D621BDFC2850571DFBB5" author="Deuve T." box="[906,1032,1035,1057]" pageId="3" pageNumber="404" pagination="171 - 184" refId="ref7922" refString="Deuve T. 1992. Contribution a la connaissance des Trechidae asiatiques (Coleoptera) // Bulleten de la Societe entomologique de France. Vol. 97. P. 171 - 184." type="book chapter" year="1992">Deuve, 1992</bibRefCitation>
</taxonomicName>
. However, it differs from this species in having larger body size (2.723.15, on average, 2.93 vs. 2.443.04, on average,
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<taxonomicName id="F3CA4D38FFD5D621BA2F281D5780FBC3" authorityName="Deuve" authorityYear="1992" box="[1113,1173,1089,1110]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="xiei">
<emphasis id="06BEEAA9FFD5D621BA2F281D5780FBC3" box="[1113,1173,1089,1110]" italics="true" pageId="3" pageNumber="404">T. xiei</emphasis>
</taxonomicName>
), darker color of upper side, broader base of pronotum and longer antennae (see Table 2) and particularly it may be distinguished by the shape of the median lobe, which is much bigger, with endophallus armature more heavily sclerotized, and its distal portion more markedly curved (
<figureCitation id="ACF12A3EFFD5D621BDA8289A572DFB4E" box="[990,1080,1222,1244]" captionStart="Figs 27" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[194,1396,1013,1943]" captionTargetId="figure-526@4.[194,1397,1013,1945]" captionTargetPageId="4" captionText="Figs 27. Male genitalia of Trechus spp. n.: 24 — T. xiongmao sp. n., paratype; 57 — T. comma sp. n., paratype; 2, 5 — lateral view; 3, 7 — dorsal view; 4 — distal portion, right lateral view; 6 — right paramere, apex. Рис. 27. Гениталии самцов новых видов рода Trechus: 24 — T. xiongmao sp. n., паратип; 57 — T. comma sp. n., паратип; 2, 5 — сбоку; 3, 7 — сверху; 4 — дистальнаЯ часть, справа; 6 — верШина правой парамеры." figureDoi="http://doi.org/10.5281/zenodo.13178258" httpUri="https://zenodo.org/record/13178258/files/figure.png" pageId="3" pageNumber="404">Figs 24</figureCitation>
).
</paragraph>
<paragraph id="347536BBFFD5D621BD2428BD57E4F942" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
On the other hand,
<taxonomicName id="F3CA4D38FFD5D621BA6B28BD578CFB63" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[1053,1177,1249,1270]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="xiongmao" status="sp.n.">
<emphasis id="06BEEAA9FFD5D621BA6B28BD578CFB63" box="[1053,1177,1249,1270]" italics="true" pageId="3" pageNumber="404">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD5D621BAD228BD57C1FB63" bold="true" box="[1188,1236,1249,1270]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="1D8D57D2FFD5D621BAD228BD57C1FB63" box="[1188,1236,1249,1270]" pageId="3" pageNumber="404" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="F3CA4D38FFD5D621BB1528BD5786FA85" authority="Deuve &amp; Queinnec, 1993" authorityName="Deuve &amp; Queinnec" authorityYear="1993" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="06BEEAA9FFD5D621BB1528BD5083FA84" italics="true" pageId="3" pageNumber="404">T. gansuensis</emphasis>
<bibRefCitation id="505B4B4AFFD5D621BDEA28A05786FA85" author="Deuve T. &amp; Queinnec E." box="[924,1171,1275,1297]" pageId="3" pageNumber="404" pagination="1 - 9" refId="ref8082" refString="Deuve T., Queinnec E. 1993. Nouveaux Trechus du Qinghai, du Sichuan et du Gansu, Chine (Coleoptera, Caraboidea: Trechidae) // Opuscula Zoologica Fluminensia. Vol. 104. P. 1 - 9." type="journal article" year="1993">Deuve &amp; Quéinnec, 1993</bibRefCitation>
</taxonomicName>
described from
<collectingRegion id="F60EF859FFD5D621BB4428A75666FA85" box="[1330,1395,1275,1296]" country="China" name="Gansu" pageId="3" pageNumber="404">Gansu</collectingRegion>
: “
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au sud-ouest de Lanzhou, Ponggartang” [
<bibRefCitation id="505B4B4AFFD5D621BB59294A50DFFAD3" author="Deuve T. &amp; Queinnec E." pageId="3" pageNumber="404" pagination="1 - 9" refId="ref8082" refString="Deuve T., Queinnec E. 1993. Nouveaux Trechus du Qinghai, du Sichuan et du Gansu, Chine (Coleoptera, Caraboidea: Trechidae) // Opuscula Zoologica Fluminensia. Vol. 104. P. 1 - 9." type="journal article" year="1993">Deuve, Quéinnec, 1993</bibRefCitation>
] but differs in having more broadly ovate habitus, smaller eyes (their diameter only slightly exceeding length of both tempora and antennomere 3), longer antennae, more transverse pronotum (PW/PL 1.43 vs. 1.36, PW/HW 1.30 vs.
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<taxonomicName id="F3CA4D38FFD5D621BDB229C05753FA24" authorityName="Deuve &amp; Queinnec" authorityYear="1993" box="[964,1094,1436,1457]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="06BEEAA9FFD5D621BDB229C05753FA24" box="[964,1094,1436,1457]" italics="true" pageId="3" pageNumber="404">T. gansuensis</emphasis>
</taxonomicName>
), wider and more convex elytra (EL/BH 2.24 vs. 2.37) and larger pronotum (EL/PL 2.75 vs. 2.80); the preapical pore removed farther from the apex of elytra, umbilicate pores 14 removed from umbilicate pore 5 and placed closer to the base of elytra (the significance of differences in the above morphometric characters was based on the
<emphasis id="06BEEAA9FFD5D621BD192A60506BF9C4" box="[879,894,1596,1617]" italics="true" pageId="3" pageNumber="404">t-</emphasis>
criterion, species comparison on the Mann-Whitney criterion is shown in Table 2). Male genitalia of
<taxonomicName id="F3CA4D38FFD5D621BA882A0B5663F9F9" authorityName="Belousov &amp; Kabak" authorityYear="2021" box="[1278,1398,1623,1644]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="xiongmao" status="sp.n.">
<emphasis id="06BEEAA9FFD5D621BA882A0B5663F9F9" box="[1278,1398,1623,1644]" italics="true" pageId="3" pageNumber="404">T. xiongmao</emphasis>
</taxonomicName>
<emphasis id="06BEEAA9FFD5D621BD5A2A2D5048F913" bold="true" box="[812,861,1649,1670]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="1D8D57D2FFD5D621BD5A2A2D5048F913" box="[812,861,1649,1670]" pageId="3" pageNumber="404" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="F3CA4D38FFD5D621BA282A2D57FDF913" authorityName="Deuve &amp; Queinnec" authorityYear="1993" box="[1118,1256,1649,1670]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="06BEEAA9FFD5D621BA282A2D57FDF913" box="[1118,1256,1649,1670]" italics="true" pageId="3" pageNumber="404">T. gansuensis</emphasis>
</taxonomicName>
, but the new species can be readily distinguished by the endophallus armature more developed and by the apex of the median lobe distinctly attenuated downward in lateral view
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD5D621BD242A875711F874" pageId="3" pageNumber="404" type="distribution">
<paragraph id="347536BBFFD5D621BD242A875097F8C9" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
<materialsCitation id="84A23CE6FFD5D621BD242A87506BF8C9" collectionCode="NW" county="Li County" location="Mengdonggou River" municipality="Shangmeng" pageId="3" pageNumber="404" specimenCount="1" stateProvince="Sichuan">
DISTRIBUTION. The species is known only from the
<typeStatus id="EB718819FFD5D621BD5A2AAA5043F89E" box="[812,854,1782,1803]" pageId="3" pageNumber="404">type</typeStatus>
locality, situated in the upper valley of a small river located west of the village of
<collectingMunicipality id="D411ACC1FFD5D621BA392B4D57D4F8B2" box="[1103,1217,1809,1831]" pageId="3" pageNumber="404">Shangmeng</collectingMunicipality>
in the upper basin of the
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,
<collectionCode id="52DBAE7EFFD5D621BA4B2B705770F8D4" box="[1085,1125,1836,1857]" pageId="3" pageNumber="404">NW</collectionCode>
of
<location id="31156060FFD5D621BAF32B775610F8D4" LSID="urn:lsid:plazi:treatment:BC6387ADFFD7D621BDB92A9C5662F783:31156060FFD5D621BAF32B775610F8D4" box="[1157,1285,1835,1857]" county="Li County" municipality="Shangmeng" name="Lixian Town" pageId="3" pageNumber="404" stateProvince="Sichuan">Lixian Town</location>
,
<collectingCounty id="DD144E37FFD5D621BB782B775666F8D5" box="[1294,1395,1835,1857]" pageId="3" pageNumber="404">Li County</collectingCounty>
,
<collectingRegion id="F60EF859FFD5D621BD5A2B1A506BF8C9" box="[812,894,1862,1884]" country="China" name="Sichuan" pageId="3" pageNumber="404">Sichuan</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="347536BBFFD5D621BD242B3D5711F874" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">
BIONOMICS. The species occurs in a wide belt of altitudes in both the alpine and upper forest zones (though much more common in the latter) at elevations from
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. Most specimens were sifted from the forest litter in rather humid habitats.
</paragraph>
</subSubSection>
<subSubSection id="7CD06530FFD5D621BD242BBA5662F783" pageId="3" pageNumber="404" type="description">
<paragraph id="347536BBFFD5D621BD242BBA5662F783" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="404">DERIVATIO NOMINIS. The species epithet refers to the Chinese name of giant panda occurring in the same area.</paragraph>
</subSubSection>
</treatment>
</document>