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<mods:title id="1E7A64021D4D4D2CB6CA2261F6857522">The genus Orthomus Chaudoir, 1838 in the Iberian Peninsula and Morocco</mods:title>
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<taxonomicName id="6A9CB0FCFFB09F2CFC2CFDFF0E405691" ID-CoL="6T8QC" authority="(Dejean, 1828)" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[905,1305,634,658]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFC2CFDFF0F375691" box="[905,1134,634,658]" italics="true" pageId="4" pageNumber="434">Orthomus hispanicus</emphasis>
(Dejean, 1828)
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<paragraph id="AD23CB7FFFB09F2CFBB7FD120FC856AC" blockId="4.[905,1305,634,687]" box="[1042,1169,663,687]" pageId="4" pageNumber="434">Figs 1215.</paragraph>
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<paragraph id="AD23CB7FFFB09F2CFCF7FD4108CF5729" blockId="4.[812,1398,708,810]" pageId="4" pageNumber="434">Dorsum brown dark, almost black, body length 7.09.0 mm, males shiny with fine microreticulation, females somewhat dull with visible microreticulation. Eye convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47F8A208E25399" blockId="4.[189,1399,1831,2070]" pageId="4" pageNumber="434">
Figs 1222.
<taxonomicName id="6A9CB0FCFFB09F2CFEF4F8A20A975339" box="[337,462,1831,1850]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB09F2CFEF4F8A20AFE5339" box="[337,423,1831,1850]" italics="true" pageId="4" pageNumber="434">Orthomus</emphasis>
spp
</taxonomicName>
, specimens from
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: 1215 —
<taxonomicName id="6A9CB0FCFFB09F2CFCA7F8A208235339" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[770,890,1831,1850]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFCA7F8A208235339" box="[770,890,1831,1850]" italics="true" pageId="4" pageNumber="434">O. hispanicus</emphasis>
</taxonomicName>
; 1618 —
<taxonomicName id="6A9CB0FCFFB09F2CFC44F8A20F615339" authorityName="Martinez-Saez" authorityYear="1873" box="[993,1080,1831,1850]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB09F2CFC44F8A20F615339" box="[993,1080,1831,1850]" italics="true" pageId="4" pageNumber="434">O. perezii</emphasis>
</taxonomicName>
; 1922 —
<taxonomicName id="6A9CB0FCFFB09F2CFB3BF8A20FAB5339" authorityName="Jeanne" authorityYear="1974" box="[1182,1266,1831,1850]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB09F2CFB3BF8A20FAB5339" box="[1182,1266,1831,1850]" italics="true" pageId="4" pageNumber="434">O. aubryi</emphasis>
</taxonomicName>
; 12, 16, 19 — habitus; 13 — metepisterna; 14, 17, 2021 — median lobe of aedeagus; 15, 18, 22 — genitalia of female; 12, 14, 1617, 1920 — dorsal view; 13 — right lateral view; 21 — left lateral view; 14 — sierra de Larouco, Orense; 15 — Candelario, Salamanca; 17 — Campisábalos, Guadalajara; 18 — Valdefuentes de Sangusín, Salamanca; 1921 — Moncayo Peak, Zaragoza; 22 — Linas de Broto, Huesca; 15, 18, 22 — after
<bibRefCitation id="C90DB68EFFB09F2CFF4EF8020A325399" author="Ortuno V. M." box="[235,363,1927,1946]" pageId="4" pageNumber="434" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. Scale bars: 1.2 mm (Fig. 12); 1.3 mm (Fig. 16); 1.5 mm (Fig. 19).
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47F826088E5C15" blockId="4.[189,1399,1831,2070]" pageId="4" pageNumber="434">
Рис. 1222.
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<emphasis id="9FE8176DFFB09F2CFEE8F8260AFA53B5" box="[333,419,1955,1974]" italics="true" pageId="4" pageNumber="434">Orthomus</emphasis>
spp
</taxonomicName>
, особи иЗ
<collectingCountry id="D58B8BEFFFB09F2CFD84F826092B53B5" box="[545,626,1955,1974]" name="Spain" pageId="4" pageNumber="434">Испании</collectingCountry>
: 1215 —
<taxonomicName id="6A9CB0FCFFB09F2CFD75F826081C53B5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[720,837,1955,1974]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB09F2CFD75F826081C53B5" box="[720,837,1955,1974]" italics="true" pageId="4" pageNumber="434">O. hispanicus</emphasis>
</taxonomicName>
; 1618 —
<taxonomicName id="6A9CB0FCFFB09F2CFC01F82608A153B6" authorityName="Martinez-Saez" authorityYear="1873" box="[932,1016,1955,1974]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB09F2CFC01F82608A153B6" box="[932,1016,1955,1974]" italics="true" pageId="4" pageNumber="434">O. perezii</emphasis>
</taxonomicName>
; 1922 —
<taxonomicName id="6A9CB0FCFFB09F2CFBF3F8260FFE53B5" authorityName="Jeanne" authorityYear="1974" box="[1110,1191,1955,1974]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB09F2CFBF3F8260FFE53B5" box="[1110,1191,1955,1974]" italics="true" pageId="4" pageNumber="434">O. aubryi</emphasis>
</taxonomicName>
; 12, 16, 19 — внеШний вид; 13 — Эпистерн Заднегруди; 14, 17, 2021 — среднЯЯ долЯ Эдеагуса; 15, 18, 22 — гениталии самки; 12, 14, 1617, 1920 — сверху; 13 –сбоку справа; 21 –сбоку слева; 14 — Сьерра- де- Ларуко, Оренсе; 15 — Канделарио, Саламанка; 17 — Камписабалос, Гвадалахара; 18 — ВальдефуЭнтес- де- Сангусин, Саламанка; 1921 — пик Монкайо, Сарагоса; 22 — Линас- де- Брото, УЭска; 15, 18, 22 — по Ortuсo [1996]. МасШтаб: 1,2 мм (Рис. 12); 1,3 мм (Рис. 16); 1,5 мм (Рис. 19).
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FEB909E955A2" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Pronotum shape variable, either maximum width at apical 3/4 (more trapezoid) or at middle (more quadrate) (Fig. 12); posterior angle obtuse, slightly toothed, basis not emarginate. Foveae well marked and usually punctured.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FE2309B55624" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Elytra moderately widened backward, side narrowed toward basis, humerus toothed, striae smooth or with traces of punctures, intervals flat in females and slightly convex in males; disk with distal seta on 3rd stria but it often varies as it is found in interval 3, close to stria 2 or may be absent.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FDAE09CF56AF" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">
Prosternal process forming an obtuse ridge, rounded and emarginate on bottom, ventral segments punctured including metepisterna (Fig. 13), this as long as wide (L/
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). Males with mesotibia somewhat enlarged apically and metatibia with traces of crenulation in apical internal side.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FD3409835732" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">Median lobe of aedeagus slightly bent ventrally; apical shaft asymmetrical, more narrowed in left side and prolonged in the right side (Fig. 14); membranous sac devoid of teeth and only moderately open on dorsal view. Spermatheca dilat- ed apically, spermathecal duct relatively short (Fig. 15)</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FCB309465782" blockId="5.[189,775,315,897]" pageId="5" pageNumber="435">
Northern half of
<collectingCountry id="D58B8BEFFFB19F2DFE30FCB30AB0574F" box="[405,489,822,844]" name="Portugal" pageId="5" pageNumber="435">Portugal</collectingCountry>
, Galicia, Cantabrian Mountains, Northern Sistema Ibérico, and Sistema Central from Serra da Estrela to sierra de Gredos.
</paragraph>
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<taxonomicName id="6A9CB0FCFFB19F2DFE9EF90409D1529A" ID-CoL="74WQB" authority="Jeanne, 1974" authorityName="Jeanne" authorityYear="1974" box="[315,648,1665,1689]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFE9EF9040AAD529A" box="[315,500,1665,1689]" italics="true" pageId="5" pageNumber="435">Orthomus aubryi</emphasis>
<bibRefCitation id="C90DB68EFFB19F2DFE5EF90709D1529A" author="Jeanne C." box="[507,648,1665,1689]" pageId="5" pageNumber="435" pagination="66 - 71" refId="ref16762" refString="Jeanne C. 1974. Carabiques nouveaux (5 eme note) // Bull. Soc. ent. France. Vol. 79. P. 66 - 71." type="journal article" year="1974">Jeanne, 1974</bibRefCitation>
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<paragraph id="AD23CB7FFFB19F2DFE07F91B097B52B5" blockId="5.[189,775,1665,2073]" box="[418,546,1694,1718]" pageId="5" pageNumber="435">Figs 1922.</paragraph>
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<paragraph id="AD23CB7FFFB19F2DFF47F946094D532A" blockId="5.[189,775,1665,2073]" pageId="5" pageNumber="435">Body length 8.510.0 mm. Dorsum dark piceous, appendages reddish. Antennomeres pubescent from apical third of antennomere 3 to 11, reaching the basis of elytra. Eye convex with long oblique tempora.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F8AB0A6153E7" blockId="5.[189,775,1665,2073]" pageId="5" pageNumber="435">Pronotum with anterior angle moderately projecting forward, anterior basis almost regularly curved; maximum width before middle (Fig. 19), from here side is slightly arcuate and narrowed toward the obtuse posterior angle, that has a small protruding tooth. Basal foveae finely punctured, the internal deep, linear or curved, the external moderately impressed or almost faint.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F86C0F1E5568" blockId="5.[189,775,1665,2073]" lastBlockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
Elytra relatively long (L/
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), almost parallel, humeral tooth small or almost absent, striae smooth, intervals moderately convex, parascutellar stria somewhat short, distal dorsal pore adjoining stria 2.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FEF40E2E55D4" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">Episternae and first ventrites moderately punctured. Metepisternae as long as wide in the anterior margin (L/A 1.05). Male mesotibia slightly dilated apically, metatibia distally curved and with traces of crenulation in the distal inner side.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FE5E0F105765" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
Apical shaft of median lobe of aedeagus elongate, somewhat more narrowed to the left (Fig. 20); on lateral view the median lobe of aedeagus is almost straight, the apex slightly curved upward (Fig. 21); membranous area (ostium) prolonged toward basis. The spermatheca (Fig. 22) show a similar shape to that described for
<taxonomicName id="6A9CB0FCFFB19F2DFB82FDE40FF15675" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1063,1192,609,630]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB19F2DFB82FDE40FF15675" box="[1063,1192,609,630]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
</taxonomicName>
but the duct is longer. Differences in the length of the spermathecal duct led
<bibRefCitation id="C90DB68EFFB19F2DFA96FDFE083556AF" author="Ortuno V. M." pageId="5" pageNumber="435" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
to place
<taxonomicName id="6A9CB0FCFFB19F2DFC65FD130F4356A8" authorityName="Jeanne" authorityYear="1974" box="[960,1050,662,683]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFC65FD130F4356A8" box="[960,1050,662,683]" italics="true" pageId="5" pageNumber="435">O. aubryi</emphasis>
</taxonomicName>
near the
<taxonomicName id="6A9CB0FCFFB19F2DFBD5FD130FAE56A8" authorityName="Fairmaire" authorityYear="1872" box="[1136,1271,662,683]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="planidorsis">
<emphasis id="9FE8176DFFB19F2DFBD5FD130FAE56A8" box="[1136,1271,662,683]" italics="true" pageId="5" pageNumber="435">O. planidorsis</emphasis>
</taxonomicName>
group. As
<taxonomicName id="6A9CB0FCFFB19F2DFAFAFD13083356C5" authorityName="Jeanne" authorityYear="1974" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="9FE8176DFFB19F2DFAFAFD13083356C5" italics="true" pageId="5" pageNumber="435">O. aubryi</emphasis>
</taxonomicName>
might enter in contact with
<taxonomicName id="6A9CB0FCFFB19F2DFBC8FD340FB756C5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1133,1262,689,710]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="9FE8176DFFB19F2DFBC8FD340FB756C5" box="[1133,1262,689,710]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB19F2DFABFFD340E2F56C5" authorityName="Martinez-Saez" authorityYear="1873" box="[1306,1398,689,710]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB19F2DFABFFD340E2F56C5" box="[1306,1398,689,710]" italics="true" pageId="5" pageNumber="435">O. perezii</emphasis>
</taxonomicName>
in the west (Basque Mts., north Sistema Ibérico), and with
<taxonomicName id="6A9CB0FCFFB19F2DFAFAFD4908CE56F8" authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="planidorsis">
<emphasis id="9FE8176DFFB19F2DFAFAFD4908CE56F8" italics="true" pageId="5" pageNumber="435">O. planidorsis</emphasis>
</taxonomicName>
in the east (Central Pyrenees), the geographic distribution doesnt help in solving the affinities of the species. Molecular markers are needed to seed light into this question. Basque Mountains, Northern Sistema Ibérico (Moncayo Peak), central and western Pyrenees.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FCEE0F7B579F" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">Groups of species with metepisterna longer that wider and narrowed posteriorly</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FC240FAD57B5" blockId="5.[812,1399,315,1324]" box="[850,1268,929,951]" pageId="5" pageNumber="435">
Iberian taxa: Group of
<taxonomicName id="6A9CB0FCFFB19F2DFB90FC240FAD57B5" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1077,1268,929,950]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB19F2DFB90FC240FAD57B5" box="[1077,1268,929,950]" italics="true" pageId="5" pageNumber="435">Orthomus barbarus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FC3E088F5128" blockId="5.[812,1399,315,1324]" pageId="5" pageNumber="435">
This group is made up by
<emphasis id="9FE8176DFFB19F2DFBF6FC39088D57E8" italics="true" pageId="5" pageNumber="435">
<taxonomicName id="6A9CB0FCFFB19F2DFBF6FC390F9457D3" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1107,1229,955,977]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="barbarus">O. barbarus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB19F2DFB72FC390E2A57D3" box="[1239,1395,955,977]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="velocissimus">O. velocissimus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB19F2DFC89FC53088957E8" authorityName="Breit" authorityYear="1933" box="[812,976,982,1003]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="formenterrae">O. formenterrae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6A9CB0FCFFB19F2DFBA9FC530FC557E8" authorityName="Mateu et Colas" authorityYear="1954" box="[1036,1180,982,1003]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB19F2DFBA9FC530FC557E8" box="[1036,1180,982,1003]" italics="true" pageId="5" pageNumber="435">O. penibeticus</emphasis>
</taxonomicName>
. Males of this group show a distally dilated mesotibia and a metatibia variably crenulate in the distal half. The female genitalia show a long spermathecal duct followed by a long spermatheca [
<bibRefCitation id="C90DB68EFFB19F2DFA8CFBA308065055" author="Ortuno V. M." pageId="5" pageNumber="435" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño, 1996</bibRefCitation>
;
<bibRefCitation id="C90DB68EFFB19F2DFCCEFBC40F195055" author="Pupier R. &amp; Coulon J." box="[875,1088,1089,1111]" pageId="5" pageNumber="435" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
]. Male genitalia are rather variable as the apical shaft of the median lobe of the aedeagus is triangular in some taxa, with rounded or acute tip, whereas it is truncate, with or without a tooth at tip. Taxa of this group are found below a hypothetical line that runs from the southwest of
<collectingCountry id="D58B8BEFFFB19F2DFCD2FB4308F650DF" box="[887,943,1222,1244]" name="Georgia" pageId="5" pageNumber="435">Iberia</collectingCountry>
(Algarve and western Andalusia) to the eastern Pyrenees and south
<collectingCountry id="D58B8BEFFFB19F2DFC54FB640F6C50F5" box="[1009,1077,1249,1270]" name="France" pageId="5" pageNumber="435">France</collectingCountry>
, and also occupies the southernmost archipelago of the Balearic Islands, the Pitiusas (Ibiza and Formentera).
</paragraph>
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</treatment>
</document>

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<taxonomicName id="6A9CB0FCFFB19F2DFEB5FC1009EA57AE" ID-CoL="6SWQD" authority="Martinez-Saez, 1873" authorityName="Martinez-Saez" authorityYear="1873" box="[272,691,917,941]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="9FE8176DFFB19F2DFEB5FC100A9557AE" box="[272,460,917,941]" italics="true" pageId="5" pageNumber="435">Orthomus perezii</emphasis>
Martínez-Sáez, 1873
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<paragraph id="AD23CB7FFFB19F2DFE07FC37097B57C9" blockId="5.[189,775,917,1645]" box="[418,546,946,970]" pageId="5" pageNumber="435">Figs 1618.</paragraph>
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<subSubSection id="E58698F4FFB19F2DFF47FC5D0950526E" pageId="5" pageNumber="435" type="description">
<paragraph id="AD23CB7FFFB19F2DFF47FC5D0A9A5020" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">
Dorsum brown dark piceous, body size smaller than that of
<emphasis id="9FE8176DFFB19F2DFF73FC770A395004" box="[214,352,1010,1031]" italics="true" pageId="5" pageNumber="435">
<taxonomicName id="6A9CB0FCFFB19F2DFF73FC770A045004" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[214,349,1010,1031]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="hispanicus">O. hispanicus</taxonomicName>
,
</emphasis>
6.07.5 mm, males shiny, females mat. Eye less convex, tempora long.
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FBAD09C050A4" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Pronotum transverse, narrowed anteriad, maximum width at middle, slightly constricted backward (Fig. 16); posterior angle obtuse, no or at most, slightly toothed; basis not emarginate; foveae well marked and usually punctured, space between side and external fovea notably convex.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FB2809FB5144" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Elytra relatively short and oval, sides slightly widened backward, narrowed close to basis, basis forming at humerus a fold that doesnt protrude outside; parascutellar stria normally developed with a proximal hairy pore; intervals convex, striae smooth, second disk pore located variably: on the 2nd stria, in the interval between striae 2 and 3, close to stria 3.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47FAC8085E51B0" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Prosternal process rounded, ventral segments finely punctured and with rugose areas, metepisterna as long as wide (L/A 1.0). Male mesotibia somewhat enlarged apically, metatibia with crenulation moderately developed in apical internal side.</paragraph>
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Apical shaft of median lobe of aedeagus well developed, as wide as long, evenly rounded to tip (Fig. 17); on lateral view almost straight; membranous sac open along the entire dorsal face. Spermatheca as described in
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<emphasis id="9FE8176DFFB19F2DFDEFF98D0996521F" box="[586,719,1543,1565]" italics="true" pageId="5" pageNumber="435">O. hispanicus</emphasis>
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, with a notably dilated apical end (Fig. 1).
</paragraph>
<paragraph id="AD23CB7FFFB19F2DFF47F9B80950526E" blockId="5.[189,775,917,1645]" pageId="5" pageNumber="435">Sistema Central, Sistema Iberico (N and S), North Meseta, northern part of South Meseta.</paragraph>
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<paragraph id="AD23CB7FFFB19F2DFCF8FAC50E1F515B" blockId="5.[812,1399,1344,2072]" box="[861,1350,1344,1368]" pageId="5" pageNumber="435">
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<taxonomicName id="6A9CB0FCFFB19F2DFCF8FAC50E1F515B" ID-CoL="8LVTV" authority="(Dejean, 1828)" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[861,1350,1344,1368]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="subSpecies" species="barbarus" subSpecies="barbarus">
<emphasis id="9FE8176DFFB19F2DFCF8FAC50FC5515B" box="[861,1180,1344,1368]" italics="true" pageId="5" pageNumber="435">Orthomus barbarus barbarus</emphasis>
(Dejean, 1828)
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<paragraph id="AD23CB7FFFB19F2DFBB7FAD80FCB5176" blockId="5.[812,1399,1344,2072]" box="[1042,1170,1373,1397]" pageId="5" pageNumber="435">Figs 2327.</paragraph>
</subSubSection>
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<paragraph id="AD23CB7FFFB19F2DFCF7FA070E4E51EB" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Dorsum between black and dark brown, males shiny, females less shiny or dull with isodiametric microreticulation. Body length 8.811.5 mm. Legs black, antennae and tarsi dark red. Eye notably convex, tempora short.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7FA680E0752A1" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Pronotum shape variable, most commonly showing a trapezoid contour more narrowed forward and side entirely curved, maximum width at middle or so (Fig. 23), anterior angle not markedly protruding forward; posterior angle toothed; inner basal fovea linear, outer fovea from rugose to almost erased, punctures of moderate size and at least abundant in inner fovea, postangular field moderately convex.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7F9220E32530E" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">Elytra almost parallel, slightly narrowed toward basis; humeral tooth well developed; striae smooth or vaguely punctured, intervals almost flat; discal pores on 3rd stria; parascutellar stria normally developed with proximal setose pore.</paragraph>
<paragraph id="AD23CB7FFFB19F2DFCF7F89708D05374" blockId="5.[812,1399,1344,2072]" pageId="5" pageNumber="435">
Prosternal process rounded; ventral segments almost smooth, fine punctures are found on sides of last ventrites; metepisterna smooth, long and narrowed apicad, L/
<geoCoordinate id="C8A8ADB8FFB19F2DFA83F8C20E36535E" box="[1318,1391,1863,1885]" degrees="1.25" direction="west" orientation="longitude" pageId="5" pageNumber="435" precision="555" value="-1.25">W 1.25</geoCoordinate>
) (Fig. 24).
</paragraph>
<paragraph id="AD23CB7FFFB19F2EFCF7F8F80A195568" blockId="5.[812,1399,1344,2072]" lastBlockId="6.[189,775,315,444]" lastPageId="6" lastPageNumber="436" pageId="5" pageNumber="435">
Median lobe of aedeagus on lateral view with apical region curved upward; apical shaft on dorsal view rather variable between localities but predominating an elongate triangle evenly narrowed to tip and straight (Fig. 25), or deviated to right [see
<bibRefCitation id="C90DB68EFFB19F2DFBBDF86D0FC453FE" author="Mateu J." box="[1048,1181,2023,2045]" pageId="5" pageNumber="435" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957</bibRefCitation>
]. In lateral view the median lobe is somewhat ventrally bent (Fig. 26). The spermathecal duct is rather long, even longer than the spermatheca (Fig. 27).
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF47FEF409AC55BF" blockId="6.[189,775,315,444]" pageId="6" pageNumber="436">
The species is distributed in the Mediterranean coast from eastern Almeria to south
<collectingCountry id="D58B8BEFFFB29F2EFE0BFE090AB755A2" box="[430,494,396,417]" name="France" pageId="6" pageNumber="436">France</collectingCountry>
and also colonizes the South Meseta, Betic and Iberian chains and the River Ebro Basin.
</paragraph>
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<paragraph id="AD23CB7FFFB29F2EFF4BFE55098C55EB" blockId="6.[189,775,464,792]" box="[238,725,464,488]" pageId="6" pageNumber="436">
<heading id="F66B7C13FFB29F2EFF4BFE55098C55EB" box="[238,725,464,488]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="436" reason="8">
<taxonomicName id="6A9CB0FCFFB29F2EFF4BFE5509DA55EB" ID-CoL="8LMSF" authority="(Breit, 1933)" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[238,643,464,488]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFF4BFE550AB455EB" box="[238,493,464,488]" italics="true" pageId="6" pageNumber="436">Orthomus formenterrae</emphasis>
(Breit, 1933)
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,
<emphasis id="9FE8176DFFB29F2EFD2AFE55098C55EB" bold="true" box="[655,725,464,488]" pageId="6" pageNumber="436">stat.n.</emphasis>
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<paragraph id="AD23CB7FFFB29F2EFE07FE68097B5606" blockId="6.[189,775,464,792]" box="[418,546,493,517]" pageId="6" pageNumber="436">Figs 2831.</paragraph>
</subSubSection>
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<paragraph id="AD23CB7FFFB29F2EFF47FD97085F571B" blockId="6.[189,775,464,792]" pageId="6" pageNumber="436">
This species was described as
<taxonomicName id="6A9CB0FCFFB29F2EFDB0FD970AF05641" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="balearicus" subGenus="Orthomus" subSpecies="formenterrae">
<emphasis id="9FE8176DFFB29F2EFDB0FD970AF05641" italics="true" pageId="6" pageNumber="436">Pterostichus (Orthomus) balearicus formenterrae</emphasis>
</taxonomicName>
and lately considered a member of the
<taxonomicName id="6A9CB0FCFFB29F2EFF47FDCD0A00565F" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[226,345,583,605]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFF47FDCD0A00565F" box="[226,345,583,605]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
complex by
<bibRefCitation id="C90DB68EFFB29F2EFE79FDCD093A565E" author="Mateu J." box="[476,611,583,605]" pageId="6" pageNumber="436" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
, in which it was ranked as subspecies:
<taxonomicName id="6A9CB0FCFFB29F2EFE32FDE709CD5674" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[407,660,610,631]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="barbarus" subSpecies="formenterrae">
<emphasis id="9FE8176DFFB29F2EFE32FDE709CD5674" box="[407,660,610,631]" italics="true" pageId="6" pageNumber="436">O. barbarus formenterrae</emphasis>
</taxonomicName>
, a criterion that has been followed by authors. Due to the confusion between
<taxonomicName id="6A9CB0FCFFB29F2EFEBDFD1D0ACA56AF" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[280,403,663,685]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFEBDFD1D0ACA56AF" box="[280,403,663,685]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB29F2EFE6EFD1D093C56AF" authorityName="Chaudoir" authorityYear="1859" box="[459,613,663,685]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="trapezicollis">
<emphasis id="9FE8176DFFB29F2EFE6EFD1D093C56AF" box="[459,613,663,685]" italics="true" pageId="6" pageNumber="436">O. trapezicollis</emphasis>
</taxonomicName>
the species appeared in the catalogue of Zaballos and Jeanne [1994] as
<emphasis id="9FE8176DFFB29F2EFD4AFD370A9856E1" italics="true" pageId="6" pageNumber="436">
<taxonomicName id="6A9CB0FCFFB29F2EFD4AFD370AE456E1" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="trapezicollis" subSpecies="formenterrae">O. trapezicollis formenterrae</taxonomicName>
.
</emphasis>
In recent works, as that of Wrase et al. [2014] and
<bibRefCitation id="C90DB68EFFB29F2EFECEFD6D094B56FF" author="Bousquet Y." box="[363,530,743,765]" pageId="6" pageNumber="436" pagination="469 - 521" refId="ref16324" refString="Bousquet Y. 2017. Tribe Pterostichini // Lobl L., Lobl I. (eds.). Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Leiden: Brill. P. 469 - 521." type="book chapter" year="2017">Bousquet [2017]</bibRefCitation>
, it is again considered a subspecies of
<taxonomicName id="6A9CB0FCFFB29F2EFEEAFC870A925714" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[335,459,770,791]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFEEAFC870A925714" box="[335,459,770,791]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
. The criterion put forward by
</paragraph>
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<emphasis id="9FE8176DFFB29F2EFF5AF9740AD95307" bold="true" box="[255,384,1770,1803]" pageId="6" pageNumber="436">29 33</emphasis>
</paragraph>
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Zaballos and Jeanne was possibly based on the truncate apical shaft of the median lobe of aedeagus, that is not found in the populations of
<taxonomicName id="6A9CB0FCFFB29F2EFC5AFEF40F2E5585" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1023,1143,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC5AFEF40F2E5585" box="[1023,1143,369,390]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FE0E08A55674" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">
This taxon is herewith considered as a separate species due to the truncate penis and the isolation of Ibiza and Formentera. The truncate penis shape relates it to some populations of the
<taxonomicName id="6A9CB0FCFFB29F2EFC44FE590F2055F3" box="[993,1145,475,497]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB29F2EFC44FE590F2055F3" box="[993,1145,475,497]" italics="true" pageId="6" pageNumber="436">O. velocissimus</emphasis>
</taxonomicName>
complex, whereas a close relationship to the
<taxonomicName id="6A9CB0FCFFB29F2EFC46FE730F025608" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[995,1115,502,523]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC46FE730F025608" box="[995,1115,502,523]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
complex is not supported by this character. The relatively shorter metepisterna of
<taxonomicName id="6A9CB0FCFFB29F2EFA96FD9408D65642" baseAuthorityName="Breit" baseAuthorityYear="1933" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFA96FD9408D65642" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
, in comparison to that of
<taxonomicName id="6A9CB0FCFFB29F2EFB36FDA90E755643" box="[1171,1324,555,577]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB29F2EFB36FDA90E755643" box="[1171,1324,555,577]" italics="true" pageId="6" pageNumber="436">O. velocissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB29F2EFAFAFDA908DD5658" baseAuthorityName="Dejean" baseAuthorityYear="1828" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFAFAFDA908DD5658" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
, is other character that supports ranking this taxon as a separate species.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FDF90F8A56C4" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">
Dorsum dark brown, males shiny, females dull due to strong microreticulation. Body length
<quantity id="6A64669AFFB29F2EFB08FD130FA156A8" box="[1197,1272,662,683]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="436" unit="mm" value="10.0">10 mm</quantity>
. Appendices reddish brown. Eye convex, tempora short.
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFCF7FD490E2F5714" blockId="6.[812,1399,315,791]" pageId="6" pageNumber="436">Pronotum notably narrowed forward, almost straight backward, anterior angle not much protruding ahead, posterior angle slightly obtuse, maximum pronotum width after middle</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF47F8A409AC5397" blockId="6.[189,1399,1825,2064]" pageId="6" pageNumber="436">
Figs 2335.
<taxonomicName id="6A9CB0FCFFB29F2EFEEBF8A40A935336" box="[334,458,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFEEBF8A40AFD5337" box="[334,420,1825,1844]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
, specimens from
<collectingCountry id="D58B8BEFFFB29F2EFDC7F8A409CF5337" box="[610,662,1825,1844]" name="Spain" pageId="6" pageNumber="436">Spain</collectingCountry>
: 2327 —
<taxonomicName id="6A9CB0FCFFB29F2EFD53F8A408395337" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[758,864,1825,1844]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFD53F8A408395337" box="[758,864,1825,1844]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
; 2831 —
<taxonomicName id="6A9CB0FCFFB29F2EFC67F8A40F175337" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[962,1102,1825,1844]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFC67F8A40F175337" box="[962,1102,1825,1844]" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
; 3235 —
<taxonomicName id="6A9CB0FCFFB29F2EFB14F8A40E775337" authorityName="Mateu et Colas" authorityYear="1954" box="[1201,1326,1825,1844]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB29F2EFB14F8A40E775337" box="[1201,1326,1825,1844]" italics="true" pageId="6" pageNumber="436">O. penibeticus</emphasis>
</taxonomicName>
; 23, 28, 32 — habitus; 24 — metepisterna; 2526, 2930, 3334 — median lobe of aedeagus; 27, 31, 35 — genitalia of female; 23, 25, 2829, 32 — dorsal view; 24, 26, 30, 34 — left lateral view; 2526 — Cartagena,
<collectingRegion id="6F58059DFFB29F2EFCBBF8D408065367" box="[798,863,1873,1892]" country="Spain" name="Murcia" pageId="6" pageNumber="436">Murcia</collectingRegion>
; 27 —
<collectingRegion id="6F58059DFFB29F2EFC3AF8D408B85367" box="[927,993,1873,1892]" country="Spain" name="Madrid" pageId="6" pageNumber="436">Madrid</collectingRegion>
; 2930 — Sa-Negreta, Ibiza; 31 — Formentera Island; 32 - Almería (possibly near Adra); 3334 Lanjarón, Granada; 35 — Puerto del Suspiro del Moro, Sierra Nevada, Granada; 27, 31, 35 — after
<bibRefCitation id="C90DB68EFFB29F2EFE86F8040AFA5397" author="Ortuno V. M." box="[291,419,1921,1940]" pageId="6" pageNumber="436" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. Scale bars: 1.0 mm (Figs 23, 28, 32).
</paragraph>
<paragraph id="AD23CB7FFFB29F2EFF47F8180E5F5C13" blockId="6.[189,1399,1825,2064]" pageId="6" pageNumber="436">
Рис. 2335.
<taxonomicName id="6A9CB0FCFFB29F2EFEF2F8180A8F53B3" box="[343,470,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFEF2F8180AF453B3" box="[343,429,1949,1968]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
, особи иЗ
<collectingCountry id="D58B8BEFFFB29F2EFD9AF81809C953B3" box="[575,656,1949,1968]" name="Spain" pageId="6" pageNumber="436">Испании</collectingCountry>
:
<taxonomicName id="6A9CB0FCFFB29F2EFD3AF818084653B3" box="[671,799,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB29F2EFD3AF81809AC53B3" box="[671,757,1949,1968]" italics="true" pageId="6" pageNumber="436">Orthomus</emphasis>
spp
</taxonomicName>
: 2327 —
<taxonomicName id="6A9CB0FCFFB29F2EFC2FF81808AE53B3" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[906,1015,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB29F2EFC2FF81808AE53B3" box="[906,1015,1949,1968]" italics="true" pageId="6" pageNumber="436">O. barbarus</emphasis>
</taxonomicName>
; 2831 —
<taxonomicName id="6A9CB0FCFFB29F2EFBC1F8180FAA53B3" baseAuthorityName="Breit" baseAuthorityYear="1933" box="[1124,1267,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="formenterrae">
<emphasis id="9FE8176DFFB29F2EFBC1F8180FAA53B3" box="[1124,1267,1949,1968]" italics="true" pageId="6" pageNumber="436">O. formenterrae</emphasis>
</taxonomicName>
; 3235 —
<taxonomicName id="6A9CB0FCFFB29F2EFAC4F8180A4653CB" authorityName="Mateu et Colas" authorityYear="1954" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB29F2EFAC4F8180A4653CB" italics="true" pageId="6" pageNumber="436">O. penibeticus</emphasis>
</taxonomicName>
; 23, 28, 32 — внеШний вид; 24 — Эпистерн Заднегруди; 2526, 2930, 3334 — среднЯЯ долЯ Эдеагуса; 27, 31, 35 — гениталии самки; 23, 25, 2829, 32 — сверху; 24, 26, 30, 34 –сбоку слева; 2526 — Картахена, МурсиЯ; 27 —
<collectingRegion id="6F58059DFFB29F2EFB09F8480FAC53E3" box="[1196,1269,1997,2016]" country="Spain" name="Madrid" pageId="6" pageNumber="436">Мадрид</collectingRegion>
; 2930 — Са- Негрета, Ибица; 31 — о- в Форментера; 32 — АльмериЯ (воЗможно, недалеко от Адры); 3334 — Ланхарон, Гранада; 35 — ПуЭрто- дель- Суспиро- дель- Моро, Сьерра- Невада, Гранада; 27, 31, 35 — по
<bibRefCitation id="C90DB68EFFB29F2EFC9FF87808E25C13" author="Ortuno V. M." box="[826,955,2045,2064]" pageId="6" pageNumber="436" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
. МасШтаб: 1,0 мм (Рис. 23, 28, 32).
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF18FEBE09AB55A2" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">(Fig. 28); outer fovea moderately impressed, inner fovea deep and linear or curved; fine punctures in both foveae, postangular field moderately convex; posterior angle without tooth; basal posterior border of pronotum not emarginate.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FE23090A5624" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Elytra almost parallel, regularly narrowed apicad (L/
<geoCoordinate id="C8A8ADB8FFB39F2FFD4AFE230B8755D5" degrees="3.2" direction="west" orientation="longitude" pageId="7" pageNumber="437" precision="5555" value="-3.2">W 3.2</geoCoordinate>
), basis large and with a clear fold in the humeral region that forms a moderate tooth, disk pores on stria 3, parascutellar stria normally developed, striae smooth, intervals somewhat convex in males and flat in females.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FDAE0A6B56C4" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Prosternal process rounded on bottom surface and forming a vertical obtuse ridge; ventrites almost smooth, punctures scarce, only somewhat rugose on sides; metepisterna smooth, relatively short (L/
<geoCoordinate id="C8A8ADB8FFB39F2FFE66FDFE09555692" box="[451,524,635,657]" degrees="1.05" direction="west" orientation="longitude" pageId="7" pageNumber="437" precision="555" value="-1.05">W 1.05</geoCoordinate>
) and narrowed backward. Male mesotibia dilated in apical half, metatibia with traces of crenulation.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FD4E090A5765" blockId="7.[189,776,315,871]" pageId="7" pageNumber="437">
Median lobe of aedeagus moderately curved ventrally (Fig. 30); apical shaft forming a well-developed triangle more narrowed to tip in left side (Fig. 29), truncate at tip and slightly sinuate at middle; membranous opening long, only closed near basal bulb. Characters of female genitalia as in
<emphasis id="9FE8176DFFB39F2FFD4AFCB30A4C5765" italics="true" pageId="7" pageNumber="437">
O,
<taxonomicName id="6A9CB0FCFFB39F2FFF18FCD40A4C5765" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[189,277,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">barbarus</taxonomicName>
</emphasis>
, as duct is rather long (Fig. 31).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,183 @@
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<subSubSection id="E58698F4FFB39F2FFF6FFCFF097B57B3" pageId="7" pageNumber="437" type="nomenclature">
<paragraph id="AD23CB7FFFB39F2FFF6FFCFF09A05791" blockId="7.[189,776,890,2072]" box="[202,761,890,914]" pageId="7" pageNumber="437">
<heading id="F66B7C13FFB39F2FFF6FFCFF09A05791" box="[202,761,890,914]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="437" reason="8">
<taxonomicName id="6A9CB0FCFFB39F2FFF6FFCFF09F15791" authority="Mateu et Colas, 1954" authorityName="Mateu et Colas" authorityYear="1954" box="[202,680,890,914]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFF6FFCFF0AEF5791" box="[202,438,890,914]" italics="true" pageId="7" pageNumber="437">Orthomus penibeticus</emphasis>
Mateu et Colas, 1954
</taxonomicName>
,
<emphasis id="9FE8176DFFB39F2FFD16FCFF09A05791" bold="true" box="[691,761,890,914]" pageId="7" pageNumber="437">stat.n.</emphasis>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFE07FC1D097B57B3" blockId="7.[189,776,890,2072]" box="[418,546,920,944]" pageId="7" pageNumber="437">Figs 3235.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB39F2FFF47FC380E735765" pageId="7" pageNumber="437" type="description">
<paragraph id="AD23CB7FFFB39F2FFF47FC380A065070" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
This species has also been currently considered a subspecies of
<taxonomicName id="6A9CB0FCFFB39F2FFEA4FC5D0A2157EF" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[257,376,983,1005]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFEA4FC5D0A2157EF" box="[257,376,983,1005]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
. However, the truncate shape of penis tip and the geographic isolation of its populations suggest that these have evolved to become a separate taxon from the surrounding
<taxonomicName id="6A9CB0FCFFB39F2FFE9DFBAD0A89503F" box="[312,464,1063,1085]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB39F2FFE9DFBAD0A89503F" box="[312,464,1063,1085]" italics="true" pageId="7" pageNumber="437">O. velocissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFDA4FBAD0921503F" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[513,632,1063,1085]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFDA4FBAD0921503F" box="[513,632,1063,1085]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
populations, a hypothesis to be corroborated with further studies based on other characters.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FBFD085150C0" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">Dorsum black or dark brown, appendices brown, males somewhat mat, females mat, microreticulation visible in both sexes, body length 10.011.7 mm. Eye convex, tempora short.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FB4D09E65160" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">Pronotum regularly curved from anterior to posterior basis but more narrowed anteriorly, maximum width at posterior 2/ 3 (Fig. 32), posterior angle obtuse and without tooth, posterior basis not emarginate; external fovea shallow, separate from side by a flat space, internal deep and linear; punctures on posterior basis fine and with low to moderate density.</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FAE20AD051CE" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Elytra somewhat short, L/
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, slightly widened backwards, humerus with small tooth; striae unpunctured, intervals moderately convex, disk pores on stria 3, parascutellar stria well developed.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47FA57092B528B" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Prosternal process rounded on bottom surface and forming an acute ridge from proepisterna to bottom surface; ventral face almost devoid of punctures, metepisterna smooth and longer than wider (L/
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1.4). Male mesotibia dilat- ed apically, metatibia with moderate crenulation although
<bibRefCitation id="C90DB68EFFB39F2FFF18F9DD0A1A526E" author="Mateu J." box="[189,323,1623,1645]" pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
noted that the crenulation is strong, possibly because this character varies geographically.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F9080A8E5340" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
Median lobe of aedeagus quite particular, resembling that of
<taxonomicName id="6A9CB0FCFFB39F2FFF7CF92D0A0252BF" authorityName="Lucas" authorityYear="1846" box="[217,347,1703,1725]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFB39F2FFF7CF92D0A0252BF" box="[217,347,1703,1725]" italics="true" pageId="7" pageNumber="437">O. abacoides</emphasis>
</taxonomicName>
from Maghreb, as the apical shaft forms a long triangle asymmetrically narrowed to the left and slightly truncate at tip (Fig. 33). In lateral view the median lobe is somewhat shifted upward (Fig. 34). The female genitalia are of the
<taxonomicName id="6A9CB0FCFFB39F2FFEA7F8970A225324" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[258,379,1810,1831]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFEA7F8970A225324" box="[258,379,1810,1831]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
<typeStatus id="722775DDFFB39F2FFE23F8970AE95324" box="[390,432,1810,1831]" pageId="7" pageNumber="437">type</typeStatus>
(Fig. 35), with a very long duct, longer than the spermatheca.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F8CD0AE853AE" blockId="7.[189,776,890,2072]" pageId="7" pageNumber="437">
In addition to the geographic separation,
<taxonomicName id="6A9CB0FCFFB39F2FFDDCF8CD085F535F" authorityName="Mateu et Colas" authorityYear="1954" box="[633,774,1863,1885]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFDDCF8CD085F535F" box="[633,774,1863,1885]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFF40F8E70A3C5374" authorityName="Lucas" authorityYear="1846" box="[229,357,1890,1911]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFB39F2FFF40F8E70A3C5374" box="[229,357,1890,1911]" italics="true" pageId="7" pageNumber="437">O. abacoides</emphasis>
</taxonomicName>
differ in a number of characters, especially in the female genitalia, what suggests that similarity in male genitalia is a homoplasy.
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFF47F8370E615643" blockId="7.[189,776,890,2072]" lastBlockId="7.[812,1399,315,871]" pageId="7" pageNumber="437">
The ranking of
<taxonomicName id="6A9CB0FCFFB39F2FFEDCF837095D53C4" authorityName="Mateu et Colas" authorityYear="1954" box="[377,516,1970,1991]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFEDCF837095D53C4" box="[377,516,1970,1991]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
as a separate species from
<taxonomicName id="6A9CB0FCFFB39F2FFF18F8480A6C53E1" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[189,309,1997,2018]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFF18F8480A6C53E1" box="[189,309,1997,2018]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
is based on male genitalia and the geographic isolation of
<taxonomicName id="6A9CB0FCFFB39F2FFE96F86D0A9853FF" authorityName="Mateu et Colas" authorityYear="1954" box="[307,449,2023,2045]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFE96F86D0A9853FF" box="[307,449,2023,2045]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="C90DB68EFFB39F2FFD97F86D09EE53FE" author="Mateu J." box="[562,695,2023,2045]" pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu [1957]</bibRefCitation>
considered that the distribution of
<taxonomicName id="6A9CB0FCFFB39F2FFE60F78709635C14" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[453,570,2050,2071]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFE60F78709635C14" box="[453,570,2050,2071]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
was deeply extended across Andalusia just by occupying the middle of the province of
<collectingCountry id="D58B8BEFFFB39F2FFCD2FED30890556F" box="[887,969,342,364]" name="Grenada" pageId="7" pageNumber="437">Granada</collectingCountry>
(form
<taxonomicName id="6A9CB0FCFFB39F2FFBABFED20F3F556F" authorityName="Mateu" authorityYear="1957" box="[1038,1126,343,364]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="transiens">
<emphasis id="9FE8176DFFB39F2FFBABFED20F3F556F" box="[1038,1126,343,364]" italics="true" pageId="7" pageNumber="437">transiens</emphasis>
</taxonomicName>
) and the eastern part of the province of Malaga (form
<taxonomicName id="6A9CB0FCFFB39F2FFB99FEF40FFE5585" authorityName="Jeanne" authorityYear="1981" box="[1084,1191,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="malacensis">
<emphasis id="9FE8176DFFB39F2FFB99FEF40FFE5585" box="[1084,1191,369,390]" italics="true" pageId="7" pageNumber="437">malacensis</emphasis>
</taxonomicName>
, see map of
<bibRefCitation id="C90DB68EFFB39F2FFA96FEF408CA55A3" author="Mateu J." pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957: 113</bibRefCitation>
), this hypothesis is not supported on geographic grounds; instead, it is simpler to consider populations of the so-called
<taxonomicName id="6A9CB0FCFFB39F2FFC2EFE4408BA55D5" authorityName="Mateu" authorityYear="1957" box="[907,995,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="transiens">
<emphasis id="9FE8176DFFB39F2FFC2EFE4408BA55D5" box="[907,995,449,470]" italics="true" pageId="7" pageNumber="437">transiens</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFB39F2FFBB1FE440F2655D5" authorityName="Jeanne" authorityYear="1981" box="[1044,1151,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="malacensis">
<emphasis id="9FE8176DFFB39F2FFBB1FE440F2655D5" box="[1044,1151,449,470]" italics="true" pageId="7" pageNumber="437">malacensis</emphasis>
</taxonomicName>
as part of the subspecies
<taxonomicName id="6A9CB0FCFFB39F2FFC89FE590F7855F3" authorityName="Mateu" authorityYear="1955" box="[812,1057,475,497]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="velocissimus" subSpecies="akbensis">
<emphasis id="9FE8176DFFB39F2FFC89FE590F7855F3" box="[812,1057,475,497]" italics="true" pageId="7" pageNumber="437">O. velocissimus akbensis</emphasis>
</taxonomicName>
, that is found in almost the whole basin of the River Guadalquivir. This last hypothesis was put forward by Zaballos and Jeanne [1994] and followed by Serrano in successive catalogues [2003, 2013, 2020].
</paragraph>
<paragraph id="AD23CB7FFFB39F2FFCF7FDC30E735765" blockId="7.[812,1399,315,871]" pageId="7" pageNumber="437">
Accordingly,
<taxonomicName id="6A9CB0FCFFB39F2FFC72FDC30F385658" authorityName="Mateu et Colas" authorityYear="1954" box="[983,1121,582,603]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">
<emphasis id="9FE8176DFFB39F2FFC72FDC30F385658" box="[983,1121,582,603]" italics="true" pageId="7" pageNumber="437">O. penibeticus</emphasis>
</taxonomicName>
would be a taxon distributed from the southern high slopes of Sierra Nevada (region of Las Alpujarras) to the Mediterranean coast, a relatively small region limited to west by the river Guadalfeo (Motril) and populations of
<taxonomicName id="6A9CB0FCFFB39F2FFC64FD340F0356C5" box="[961,1114,689,710]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFB39F2FFC64FD340F0356C5" box="[961,1114,689,710]" italics="true" pageId="7" pageNumber="437">O. velocissimus</emphasis>
</taxonomicName>
, and to the east by the plain between Adra and the hills west of Almería city [
<bibRefCitation id="C90DB68EFFB39F2FFA95FD4908CF56F8" author="Mateu J." pageId="7" pageNumber="437" pagination="87 - 112" refId="ref17047" refString="Mateu J. 1957. Notas sobre los Orthomus Chaudoir (tercera nota) // Eos. Vol. 33. P. 87 - 112." type="book chapter" year="1957">Mateu, 1957: 101</bibRefCitation>
]; in this last city it is found the southernmost populations of
<taxonomicName id="6A9CB0FCFFB39F2FFC19FC840F6B5715" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[956,1074,769,790]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFB39F2FFC19FC840F6B5715" box="[956,1074,769,790]" italics="true" pageId="7" pageNumber="437">O. barbarus</emphasis>
</taxonomicName>
. This hypothesis deserves further corroboration, although at present, the study of male genitalia suffices to separate without doubt the parapatric populations of
<emphasis id="9FE8176DFFB39F2FFCE2FCD40F3C5765" box="[839,1125,849,870]" italics="true" pageId="7" pageNumber="437">
<taxonomicName id="6A9CB0FCFFB39F2FFCE2FCD408985765" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[839,961,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="barbarus">O. barbarus</taxonomicName>
,
<taxonomicName id="6A9CB0FCFFB39F2FFC69FCD40F3C5765" box="[972,1125,849,870]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="velocissimus">O. velocissimus</taxonomicName>
</emphasis>
and
<emphasis id="9FE8176DFFB39F2FFB33FCD40E735765" box="[1174,1322,849,870]" italics="true" pageId="7" pageNumber="437">
<taxonomicName id="6A9CB0FCFFB39F2FFB33FCD40E7F5765" authorityName="Mateu et Colas" authorityYear="1954" box="[1174,1318,849,870]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="437" phylum="Arthropoda" rank="species" species="penibeticus">O. penibeticus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="25357A69FFB69F2AFC2DFB170F5D5C04" ID-DOI="http://doi.org/10.5281/zenodo.13178412" ID-Zenodo-Dep="13178412" LSID="urn:lsid:plazi:treatment:25357A69FFB69F2AFC2DFB170F5D5C04" httpUri="http://treatment.plazi.org/id/25357A69FFB69F2AFC2DFB170F5D5C04" lastPageNumber="432" pageId="2" pageNumber="432">
<subSubSection id="E58698F4FFB69F2AFC2DFB17089A511F" pageId="2" pageNumber="432" type="nomenclature">
<paragraph id="AD23CB7FFFB69F2AFC2DFB170E4250A9" blockId="2.[812,1399,1170,2055]" box="[904,1307,1170,1194]" pageId="2" pageNumber="432">
<heading id="F66B7C13FFB69F2AFC2DFB170E4250A9" box="[904,1307,1170,1194]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="432" reason="8">
<taxonomicName id="6A9CB0FCFFB69F2AFC2DFB170E4250A9" ID-CoL="74X2D" authority="(Antoine, 1941)" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[904,1307,1170,1194]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="tazekensis">
<emphasis id="9FE8176DFFB69F2AFC2DFB170F3F50A9" box="[904,1126,1170,1194]" italics="true" pageId="2" pageNumber="432">Orthomus tazekensis</emphasis>
(Antoine, 1941)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FB32089A511F" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Dorsum dark brown, appendages lighter, body length 7.39.7 mm. Antennomeres pubescent from middle antennomere 4, in rare cases with some fine, short setae at apex of antennomere 3.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFB69F2AFCF7FAA70F5D5C04" pageId="2" pageNumber="432" type="description">
<paragraph id="AD23CB7FFFB69F2AFCF7FAA708A451D4" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Pronotum transverse, anterior angle moderate or acutely projecting forward; maximum width before middle, side almost linearly narrowed to posterior angle, sometimes slightly sinuate. Inner basal fovea deep and linear, outer fovea frequently shallow, posterior basis with punctures in moderatelow density. Posterior basis somewhat depressed, including the postangular field.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FA59083552C4" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">
Elytra moderately convex with fine and unpunctured fine striae, intervals slightly convex, interval 3 with 2 setigerous punctures adjoining stria 3. Humerus well-marked with a moderate tooth projecting laterally. Parascutellar stria well-developed, without scutellar setigerous puncture (present in aberration
<taxonomicName id="6A9CB0FCFFB69F2AFBACF9E40F235274" authorityName="Antoine" authorityYear="1941" box="[1033,1146,1633,1655]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="scutellaris">“scutellaris</taxonomicName>
” and
<taxonomicName id="6A9CB0FCFFB69F2AFB10F9E7081F5291" authorityName="Wrase et Jeanne" authorityYear="2005" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB69F2AFB10F9E7081F5291" italics="true" pageId="2" pageNumber="432">O. tazekensis rifensis</emphasis>
</taxonomicName>
). Elytron microsculpture indistinct, elytra moderately shiny in males, isodiametric in females, that have dull elytra.
</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7F94908315381" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">
Prosternal process rounded. Meso- and metepisterna irregularly punctured, ventrites with rugose punctures. Mesotibia of males not dilated apically, metatibia almost straight, with crenulation at internal margin starting at about the beginning of apical half and consisting of about 8 tubercles. In
<taxonomicName id="6A9CB0FCFFB69F2AFC62F8D70FF85364" authorityName="Wrase et Jeanne" authorityYear="2005" box="[967,1185,1873,1895]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB69F2AFC62F8D70FF85364" box="[967,1185,1873,1895]" italics="true" pageId="2" pageNumber="432">O. tazekensis rifensis</emphasis>
</taxonomicName>
crenulation is rather weak.
</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7F8020E7D53D1" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">Apical shaft of median lobe asymmetrical, more narrowed to tip in the left side, internal sac with some groups of middle-sized thorns, almost straight in lateral view.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7F8520F5D5C04" blockId="2.[812,1399,1170,2055]" pageId="2" pageNumber="432">This subspecies is found in forests of Middle Atlas, in the surroundings of Taza.</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="9FE8176DFFB69F2AFCDAFEBE0F2D5550" box="[895,1140,315,339]" italics="true" pageId="2" pageNumber="432">Orthomus maroccanus</emphasis>
Chaudoir, 1873
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Dorsum dark piceous, appendages yellow reddish. Body length 8.09.8 mm. Males shiny, females dull with isodiametric microsculpture strongly developed. Antennomeres pubescent from the second third of antennomere 4, sometimes with small distally hairs in antennomere
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addition to the long apical setae (Fig. 1).
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<paragraph id="AD23CB7FFFB69F2AFCF7FD9B0F5D56D0" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">Pronotum transverse with anterior angles moderately projecting forward (Fig. 1), side narrowed forward and only moderately narrowed backward, maximum width before middle, posterior angle obtuse without a small tooth. Basal foveae from almost smooth to clearly punctured, area between external impression and lateral margin (postangular field) notably convex.</paragraph>
<paragraph id="AD23CB7FFFB69F2AFCF7FD5D08985770" blockId="2.[812,1399,315,1150]" pageId="2" pageNumber="432">Elytra moderately widened toward apex, slightly narrowed toward basis; striae fine, unpunctured, intervals slightly convex, interval 3 with two setigerous punctures adjoining stria 3; humerus well-marked with a small tooth; scutellar setigerous puncture present at base of a well-developed parascutellar stria.</paragraph>
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Episterna and abdomen from distinctly punctured to almost smooth, punctures on ventrites sometimes connected by shallow longitudinal ridges. Metepisterna slightly longer than wide, L/
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(Fig. 2). Prosternal process not truncate. Mesotibia of males distinctly enlarged from middle to apex, metatibia slightly curved and weakly crenulate at internal margin of distal half.
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<paragraph id="AD23CB7FFFB79F2BFE0AFEDC094D5573" blockId="3.[189,775,315,777]" box="[431,532,344,369]" pageId="3" pageNumber="433">Figs 59.</paragraph>
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<paragraph id="AD23CB7FFFB79F2BFF47FEF80A4F56D0" blockId="3.[189,775,315,777]" pageId="3" pageNumber="433">Characters indicated above for the nominal subspecies are also found in this taxon (Fig. 5). Individuals of this subspecies differ from those of the nominal subspecies in having a scutellar pore puncture; ventrites show abundant punctures giving a rugose aspect (Fig. 6); crenulation of metatibia in males is more weakly developed (it consists of about 6 tubercles); the apical shaft of the median lobe of the aedeagus is shorter and almost evenly rounded apically (Fig. 7), and is almost straight in lateral view (Fig. 8); the open membranous sac extends toward basal bulb. Spermatheca notably dilated apically, perhaps due to full filling with spermatophora (Fig. 9). Its shape when empty couldnt be assessed.</paragraph>
<paragraph id="AD23CB7FFFB79F2BFF47FD5D0949570A" blockId="3.[189,775,315,777]" pageId="3" pageNumber="433">This subspecies is found in Rif Mountains above the timber line [Wrase, Jeanne, 2005].</paragraph>
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<paragraph id="AD23CB7FFFB79F2BFCF7FEFB0E7E561A" blockId="3.[812,1399,315,777]" pageId="3" pageNumber="433">Dorsum from light to dark reddish brown; appendages somewhat lighter than body; body length 7.89.2 mm. Eye fairly flat with long and flat tempora; antennae in both sexes with antennomere 9 reaching the pronotal base, pubescent from apical third of antennomere 3 (in addition to normal setae there are some fine short setae hardly visible).</paragraph>
<paragraph id="AD23CB7FFFB79F2BFCF7FD9B0E6156D0" blockId="3.[812,1399,315,777]" pageId="3" pageNumber="433">Pronotum transverse, widest at about anterior third, anterior margin distinctly emarginate, anterior angle moderately projecting forward; posterior angle almost rectangular, side sinuate before it; posterior basis beaded from angle to the origin of internal basal impression, base slightly concave; disk convex; basal external fovea sparsely or not punctured, internal one deeper and with some distinct punctures.</paragraph>
<paragraph id="AD23CB7FFFB79F2BFCF7FD5D094D5397" blockId="3.[812,1399,315,777]" lastBlockId="3.[189,1399,1825,2065]" pageId="3" pageNumber="433">
Elytra long-oval, moderately convex, maximum width a little behind middle; striae fine and unpunctured, intervals Figs 111.
<taxonomicName id="6A9CB0FCFFB79F2BFEE6F8A40AE65336" box="[323,447,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB79F2BFEE6F8A40AC05337" box="[323,409,1825,1844]" italics="true" pageId="3" pageNumber="433">Orthomus</emphasis>
spp
</taxonomicName>
: 14 —
<taxonomicName id="6A9CB0FCFFB79F2BFDACF8A709D55337" authorityName="Chaudoir" authorityYear="1873" box="[521,652,1826,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="9FE8176DFFB79F2BFDACF8A709D55337" box="[521,652,1826,1845]" italics="true" pageId="3" pageNumber="433">O. maroccanus</emphasis>
</taxonomicName>
; 59 —
<taxonomicName id="6A9CB0FCFFB79F2BFD76F8A708D05337" authorityName="Wrase et Jeanne" authorityYear="2005" box="[723,905,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB79F2BFD76F8A708D05337" box="[723,905,1825,1845]" italics="true" pageId="3" pageNumber="433">O. tazekensis rifensis</emphasis>
</taxonomicName>
; 1011 —
<taxonomicName id="6A9CB0FCFFB79F2BFC4EF8A70F185337" box="[1003,1089,1825,1845]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="starkei">
<emphasis id="9FE8176DFFB79F2BFC4EF8A70F185337" box="[1003,1089,1825,1845]" italics="true" pageId="3" pageNumber="433">O. starkei</emphasis>
</taxonomicName>
; 1, 5 — habitus; 2 — metepisterna; 6 — metepisterna and ventrites; 3, 78, 1011 — median lobe of aedeagus; 4, 9 — genitalia of female; 1, 5, 3, 7, 11 — dorsal view; 2, 6 — right lateral view; 8, 10 — left lateral view; 1, 59 — Tidiquin Mnt., Rif,
<collectingCountry id="D58B8BEFFFB79F2BFCE6F8D708CD5366" box="[835,916,1874,1893]" name="Morocco" pageId="3" pageNumber="433">Morocco</collectingCountry>
; 3 — San Roque, Cadiz,
<collectingCountry id="D58B8BEFFFB79F2BFBD4F8D40FFC5367" box="[1137,1189,1873,1892]" name="Spain" pageId="3" pageNumber="433">Spain</collectingCountry>
; 4 — sierra del Cabrito, Tarifa, Cádiz,
<collectingCountry id="D58B8BEFFFB79F2BFE9EF8EC0A36537F" box="[315,367,1897,1916]" name="Spain" pageId="3" pageNumber="433">Spain</collectingCountry>
; 11 — Sebt-des Beni Frassem, Rif,
<collectingCountry id="D58B8BEFFFB79F2BFD0DF8EF09A0537E" box="[680,761,1898,1917]" name="Morocco" pageId="3" pageNumber="433">Morocco</collectingCountry>
; 4, 9 — after
<bibRefCitation id="C90DB68EFFB79F2BFCD7F8EC08A8537F" author="Ortuno V. M." box="[882,1009,1897,1916]" pageId="3" pageNumber="433" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
; 1011 — after Wrase, Jeanne [2005]. Scale bars: 1.3 mm (Fig. 5); 1.4 mm (Fig. 1).
</paragraph>
<paragraph id="AD23CB7FFFB79F2BFF47F81B09D35C13" blockId="3.[189,1399,1825,2065]" pageId="3" pageNumber="433">
Рис. 111.
<taxonomicName id="6A9CB0FCFFB79F2BFEE7F8180AE453B2" box="[322,445,1949,1969]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFB79F2BFEE7F8180AC153B3" box="[322,408,1949,1968]" italics="true" pageId="3" pageNumber="433">Orthomus</emphasis>
spp
</taxonomicName>
: 14 —
<taxonomicName id="6A9CB0FCFFB79F2BFDA6F81809DC53B3" authorityName="Chaudoir" authorityYear="1873" box="[515,645,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="9FE8176DFFB79F2BFDA6F81809DC53B3" box="[515,645,1949,1968]" italics="true" pageId="3" pageNumber="433">O. maroccanus</emphasis>
</taxonomicName>
; 59 —
<taxonomicName id="6A9CB0FCFFB79F2BFD6FF818082753B3" authorityName="Wrase et Jeanne" authorityYear="2005" box="[714,894,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="tazekensis" subSpecies="rifensis">
<emphasis id="9FE8176DFFB79F2BFD6FF818082753B3" box="[714,894,1949,1968]" italics="true" pageId="3" pageNumber="433">O. tazekensis rifensis</emphasis>
</taxonomicName>
; 1011 —
<taxonomicName id="6A9CB0FCFFB79F2BFC78F8180F6853B3" box="[989,1073,1949,1968]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="starkei">
<emphasis id="9FE8176DFFB79F2BFC78F8180F6853B3" box="[989,1073,1949,1968]" italics="true" pageId="3" pageNumber="433">O. starkei</emphasis>
</taxonomicName>
; 1, 5 — внеШний вид; 2 — Эпистерн Заднегруди; 6 — Эпистерн Заднегруди и вентриты; 3, 78, 1011 — среднЯЯ долЯ Эдеагуса; 4, 9 — гениталии самки; 1, 5, 3, 7, 11 — сверху; 2, 6 –сбоку справа; 8, 10 –сбоку слева; 1, 59 — горы Тидиквин, Риф,
<collectingCountry id="D58B8BEFFFB79F2BFC2CF84B088253E2" box="[905,987,1998,2017]" name="Morocco" pageId="3" pageNumber="433">Марокко</collectingCountry>
; 3 — Сан- Роке, Кадис, ИспаниЯ; 4 — сьерра дель- Кабрито, Тарифа, Кадис, ИспаниЯ; 11 — Бени Фрассем, Риф,
<collectingCountry id="D58B8BEFFFB79F2BFC90F86308DE53FA" box="[821,903,2022,2041]" name="Morocco" pageId="3" pageNumber="433">Марокко</collectingCountry>
; 4, 9 — по Ortuсo [1996]; 1011 — по Wrase, Jeanne [2005]. МасШтаб: 1,3 мм (Рис. 5); 1,4 мм (Рис. 1).
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF18FEBE0926560F" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">
slightly convex, interval 3 with two setigerous punctures adjoining stria 3; humerus well-marked with a small indistinct tooth; basal margin anteriorly concave, forming a weak acute angle (
<typeStatus id="722775DDFFB09F2CFE99FE0E0ACD55A2" box="[316,404,395,417]" pageId="4" pageNumber="434" type="holotype">holotype</typeStatus>
) or an almost rectangular angle with lateral margin at the humerus. Setigerous pore present at base of a well-developed parascutellar stria. Microsculpture mesh pattern distinct, in males weakly transverse, in females isodiametric and slightly more stronger developed.
</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD94085F5692" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">Prosternum (laterally), episterna and abdomen (laterally) with coarse and somewhat scattered irregular punctures, sometimes connected by shallow longitudinal ridges (last ventrites). Legs slender; mesotibia of males not dilated apically, metatibia not distinctly curved and without crenulation in internal side.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD130AFA56E2" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">Male with median lobe of aedeagus strongly bent; apical shaft short and evenly rounded (Figs 1011). Internal sac without thorns or teeth.</paragraph>
<paragraph id="AD23CB7FFFB09F2CFF47FD63085E5732" blockId="4.[189,776,315,817]" pageId="4" pageNumber="434">
<materialsCitation id="1DF4C122FFB09F2CFF47FD63085E5732" country="Morocco" elevation="375" location="Known" municipality="Taza" pageId="4" pageNumber="434" specimenCount="1">
<location id="A8439DA4FFB09F2CFF47FD630A7156F8" LSID="urn:lsid:plazi:treatment:25357A69FFB79F2CFCD3FEBE085E5732:A8439DA4FFB09F2CFF47FD630A7156F8" box="[226,296,742,763]" country="Morocco" municipality="Taza" name="Known" pageId="4" pageNumber="434">Known</location>
only from three localities situated not far from each other in the north-western region of
<collectingMunicipality id="4D475105FFB09F2CFDBCFC84091E5715" box="[537,583,769,790]" pageId="4" pageNumber="434">Taza</collectingMunicipality>
(
<collectingCountry id="D58B8BEFFFB09F2CFDF0FC8409E95715" box="[597,688,769,790]" name="Morocco" pageId="4" pageNumber="434">Morocco</collectingCountry>
), in agricultural lands free of forests, at elevations about
<quantity id="6A64669AFFB09F2CFD21FC9E085A5732" box="[644,771,795,817]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="4" pageNumber="434" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">
<elevation id="26B12C4CFFB09F2CFD21FC9E085A5732" box="[644,771,795,817]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="4" pageNumber="434" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300 to 450 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,81 @@
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<paragraph id="AD23CB7FFFBD9F21FC36FD7A0E495714" blockId="9.[812,1399,767,1575]" box="[915,1296,767,791]" pageId="9" pageNumber="439">
<heading id="F66B7C13FFBD9F21FC36FD7A0E495714" box="[915,1296,767,791]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="439" reason="8">
<taxonomicName id="6A9CB0FCFFBD9F21FC36FD7A0E495714" ID-CoL="6SX2B" authority="(Lucas, 1846)" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[915,1296,767,791]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFBD9F21FC36FD7A0F2B5714" box="[915,1138,767,791]" italics="true" pageId="9" pageNumber="439">Orthomus abacoides</emphasis>
(Lucas, 1846)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD23CB7FFFBD9F21FBB7FC980FCB5737" blockId="9.[812,1399,767,1575]" box="[1042,1170,796,821]" pageId="9" pageNumber="439">Figs 5155.</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFBD9F21FCF7FCC40E115224" pageId="9" pageNumber="439" type="description">
<paragraph id="AD23CB7FFFBD9F21FCF7FCC40E2E57A4" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Body size large, 10.412.4 mm, dorsum from to black dark brown, antennae and tarsomeres red brown, femora and tibiae darker. Male shiny, female less shiny with very fine microreticulation. Eye moderately convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FC290F685044" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Anterior angle of pronotum advanced ahead, well-marked; pronotum more narrowed forward, maximum width at middle (Fig. 51), from here slightly arcuate backward; posterior basis with few and fine punctures, inner fovea linear, outer fovea shallow, basis not emarginate; posterior angle almost straight and without tooth.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FBC90FC150CF" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Elytra almost parallel with small tooth at humerus and clearly sinuate preapically when the epipleura meets dorsum; striae deep, smooth or with insinuated punctures; discal pores on stria 3, intervals almost flat; parascutellar stria normally developed, with proximal hairy pore.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FB5408D45151" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">
Ventral segments with punctures, isolated or rugose (Fig. 52). Prosternal process somewhat truncate projected backward in acute ridge. Metepisterna longer than wider (L/
<geoCoordinate id="C8A8ADB8FFBD9F21FAFAFA8208145134" degrees="1.3" direction="west" orientation="longitude" pageId="9" pageNumber="439" precision="5555" value="-1.3">W 1.3</geoCoordinate>
). Male mesotibia regularly dilated to apex, metatibia not crenulate.
</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FAD20FA951BE" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">Apical shaft of median lobe of aedeagus long, asymmetrically sinuate and deviated to the left (Fig. 54), truncate at tip; in lateral view moderately arcuate ventrally (Fig. 53). Female genitalia as described above (Fig. 55).</paragraph>
<paragraph id="AD23CB7FFFBD9F21FCF7FA440E115224" blockId="9.[812,1399,767,1575]" pageId="9" pageNumber="439">
The species is found in NW of
<collectingCountry id="D58B8BEFFFBD9F21FBC9FA440FEA51D4" box="[1132,1203,1473,1495]" name="Algeria" pageId="9" pageNumber="439">Algeria</collectingCountry>
in the region of
<collectingCountry id="D58B8BEFFFBD9F21FAE2FA470E2F51D4" box="[1351,1398,1474,1495]" name="Algeria" pageId="9" pageNumber="439">Oran</collectingCountry>
and
<collectingRegion id="6F58059DFFBD9F21FCF3FA58088A51F1" box="[854,979,1501,1522]" country="Algeria" name="Mostaganem" pageId="9" pageNumber="439">Mostaganem</collectingRegion>
, some individuals have been found in ports of south
<collectingCountry id="D58B8BEFFFBD9F21FCD8FA7208E4520F" box="[893,957,1527,1548]" name="France" pageId="9" pageNumber="439">France</collectingCountry>
(Marseille), surely imported. It might be found in
<collectingCountry id="D58B8BEFFFBD9F21FCE0F99708C45224" box="[837,925,1554,1575]" name="Morocco" pageId="9" pageNumber="439">Morocco</collectingCountry>
, in the littoral near the frontier with
<collectingCountry id="D58B8BEFFFBD9F21FB5FF9940E1D5224" box="[1274,1348,1553,1575]" name="Algeria" pageId="9" pageNumber="439">Algeria</collectingCountry>
.
</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FE07FAFC097B5192" blockId="9.[189,775,1372,2073]" box="[418,546,1401,1425]" pageId="9" pageNumber="439">Figs 4750.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FF47FA1B09295200" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Dorsum brown, with blackish to reddish hue, shiny in both sexes as microreticulation is not much developed in females; appendages piceous red. Eye of large size, convex, tempora oblique. Body length 9.011.2 mm.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FF47F98C094452DA" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Pronotum transverse with maximum width at middle (Fig. 47); from here clearly narrowed forward and almost straight backward; posterior angle slightly obtuse and not toothed; anterior angle not protruding, posterior basis not emarginate; however, lateral margin and side are clearly widened in posterior third; external fovea rugose and punctured, vaguely limited, internal one deep and linear; punctures of posterior moderately dense.</paragraph>
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Elytra long, parallel, basis forming at humerus a small fold tooth-like. Intervals almost flat, striae deep, smooth or with traces of punctures; second disk pore adjoining stria 2. Prosternal process rounded with acute ridge directed toward mesosternum; ventral segments with few punctures; metepisterna longer than wide, L/
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. Male mesotibia regularly dilated to apex, metatibia slightly curved, dilated in apical 1/6 and without crenulation.
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<paragraph id="AD23CB7FFFBD9F21FF47F836096A5C1A" blockId="9.[189,775,1372,2073]" pageId="9" pageNumber="439">Median lobe of aedeagus with short triangular apical shaft, deviated to right (Fig. 48); penis in lateral view somewhat ventrally curved and elevated at tip (Fig. 49). Female genitalia (Fig. 50) as described above.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FE07FDB8097B5656" blockId="9.[189,775,544,1352]" box="[418,546,573,597]" pageId="9" pageNumber="439">Figs 4346.</paragraph>
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<paragraph id="AD23CB7FFFBD9F21FF47FDE7085B56CB" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Dorsum black or dark brown, males not very shiny, females entirely dull, microreticulation well developed in both sexes. Large body length, 11.012.3 mm. Appendages reddish brown, femora dark. Eye large and convex, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBD9F21FF47FD48091E57D0" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Pronotum almost square in some individuals (maximum width at middle, Fig. 3), or side more curved both anteriad and apicad (maximum width on apical 2/3); the most striking features are the deep and wide lateral margin, the emarginate posterior basis and the depression of the posterior basis; both foveae deep, the inner one deeper and linear, the external one large and separate from side by a flat area; punctures of posterior basis fine and from moderate to dense, mostly found within foveae; posterior angle from obtuse to almost square, devoid of tooth or with faint traces of it.</paragraph>
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Elytra long (L/
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), slightly dilated from basis, humeral tooth absent, discal pore 2 found on the 3rd stria or on interval between striae 2 and 3; parascutellar stria normally developed with proximal setigerous pore; intervals convex, striae smooth; the lateral margin of elytra is deep and wide, excavated, as is the lateral margin of pronotum.
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Ventral segments with fine and scarce punctures; metepisterna longer than wide, L/
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. Male mesotibia regularly dilated to apex, metatibia slightly curved, dilated in apical 1/6 and with traces of crenulation.
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<paragraph id="AD23CB7FFFBD9F21FF47FB67085B514B" blockId="9.[189,775,544,1352]" pageId="9" pageNumber="439">Median lobe of aedeagus with a short triangular apical shaft (Fig. 44), more narrowed on the right side; on lateral view the penis is almost straight (Fig. 45). Female genitalia as indicated above, with a short duct and a widened spermatheca (Fig. 46).</paragraph>
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<emphasis id="9FE8176DFFBD9F21FC32F9BE0F0E5250" box="[919,1111,1595,1619]" italics="true" pageId="9" pageNumber="439">Orthomus lacouri</emphasis>
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According to
<bibRefCitation id="C90DB68EFFBD9F21FC41F9F80FAD5290" author="Pupier R. &amp; Coulon J." box="[996,1268,1661,1683]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
this species includes different geographic races that are distinguished by morphological features, and thus deserve their ranking as subspecies. Three of them are found in NE
<collectingCountry id="D58B8BEFFFBD9F21FB49F94B0E1F52E0" box="[1260,1350,1742,1763]" name="Morocco" pageId="9" pageNumber="439">Morocco</collectingCountry>
,
<taxonomicName id="6A9CB0FCFFBD9F21FAF6F94B0FEC52FE" authority="(Antoine, 1941)" baseAuthorityName="Antoine" baseAuthorityYear="1941" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="lacouri">
<emphasis id="9FE8176DFFBD9F21FAF6F94B0F5752FE" italics="true" pageId="9" pageNumber="439">Orthomus lacouri lacouri</emphasis>
(Antoine, 1941)
</taxonomicName>
;
<taxonomicName id="6A9CB0FCFFBD9F21FB64F96C0F57531B" authority="Pupier et Coulon, 2013" authorityName="Pupier et Coulon" authorityYear="2013" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="haroldi">
<emphasis id="9FE8176DFFBD9F21FB64F96C0E2F52FE" box="[1217,1398,1768,1790]" italics="true" pageId="9" pageNumber="439">O. lacouri haroldi</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FC89F8860F57531B" author="Pupier R. &amp; Coulon J." box="[812,1038,1795,1817]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier et Coulon, 2013</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="6A9CB0FCFFBD9F21FBE5F8860E2A531A" authority="Mateu, 1955" authorityName="Mateu" authorityYear="1955" box="[1088,1395,1795,1817]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
<emphasis id="9FE8176DFFBD9F21FBE5F8860FA9531B" box="[1088,1264,1795,1816]" italics="true" pageId="9" pageNumber="439">O. lacouri kocheri</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FB52F8860E2A531A" author="Mateu J." box="[1271,1395,1795,1817]" pageId="9" pageNumber="439" pagination="53 - 85" refId="ref17020" refString="Mateu J. 1955. Notas sobre los Orthomus Chaudoir (segunda nota) // Eos. Vol. 31. P. 53 - 85." type="book chapter" year="1955">Mateu, 1955</bibRefCitation>
</taxonomicName>
. The fourth subspecies,
<taxonomicName id="6A9CB0FCFFBD9F21FBB3F89B0E005330" authority="Jeanne, 1988" authorityName="Jeanne" authorityYear="1988" box="[1046,1369,1821,1843]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="pupieri">
<emphasis id="9FE8176DFFBD9F21FBB3F89B0F955330" box="[1046,1228,1821,1843]" italics="true" pageId="9" pageNumber="439">O. lacouri pupieri</emphasis>
<bibRefCitation id="C90DB68EFFBD9F21FB70F89B0E005330" author="Jeanne C." box="[1237,1369,1822,1843]" pageId="9" pageNumber="439" pagination="69 - 87" refId="ref16833" refString="Jeanne C. 1988. Carabiques noveaux ou remarquables (9 eme note) // Bull. Soc. linn. Bordeaux. Vol. 16. P. 69 - 87." type="journal article" year="1988">Jeanne, 1988</bibRefCitation>
</taxonomicName>
, is proper of NW
<collectingCountry id="D58B8BEFFFBD9F21FC67F8BD0F57534D" box="[962,1038,1848,1870]" name="Algeria" pageId="9" pageNumber="439">Algeria</collectingCountry>
. Here I follow the taxa descriptions published by
<bibRefCitation id="C90DB68EFFBD9F21FC15F8D60FF7536B" author="Pupier R. &amp; Coulon J." box="[944,1198,1875,1897]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
and
<bibRefCitation id="C90DB68EFFBD9F21FB7BF8D608365380" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="9" pageNumber="439" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
, together with personal observations of material of this species deposited in the MNCN Madrid and DZAFUM. Male genitalia are the key characters that distinguish the subspecies, which share in common most external characters. Thus, a general description follows and peculiarities of the subspecies are indicated below.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47FEB9090255BF" blockId="10.[189,776,315,657]" pageId="10" pageNumber="440">
Dorsum from black to piceous brown, appendages brown. Body length 10.511.3 mm. Eye of moderate size and convex, tempora oblique. Males shiny, females shiny or somewhat dull with visible microreticulation (
<emphasis id="9FE8176DFFBE9F22FDE8FE09099F55A3" box="[589,710,395,417]" italics="true" pageId="10" pageNumber="440">
O. l.
<taxonomicName id="6A9CB0FCFFBE9F22FD25FE0E099F55A3" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[640,710,395,416]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
); both sexes of
<emphasis id="9FE8176DFFBE9F22FEB6FE230AD455B8" box="[275,397,422,443]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
show a dull dorsum.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47FE4409B05692" blockId="10.[189,776,315,657]" pageId="10" pageNumber="440">
Pronotum shape trapezoid, anterior angle briefly protruding ahead except for
<taxonomicName id="6A9CB0FCFFBE9F22FE21FE59096855F3" authorityName="Pupier et Coulon" authorityYear="2013" box="[388,561,475,497]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="haroldi">
<emphasis id="9FE8176DFFBE9F22FE21FE59096855F3" box="[388,561,475,497]" italics="true" pageId="10" pageNumber="440">O. lacouri haroldi</emphasis>
</taxonomicName>
, anterior basis moderately concave; side arcuate, more narrowed forward, slightly narrowed backward and showing maximum width near posterior basis (Fig. 56); posterior angle slightly obtuse and without tooth; basis no or slightly (
<emphasis id="9FE8176DFFBE9F22FE6BFDC309125658" box="[462,587,582,603]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
) emarginate; inner fovea linear and marked, outer fovea poorly impressed; basis with points varying between scarce and moderately dense.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FEBE0E785584" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">Elytra parallel, humeral tooth vestigial, striae punctured with variable intensity; discal pores on stria 3; 8th interval almost vertically tumbling down to elytron margin.</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FE0E0E16560F" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
Prosternal process truncate; metepisterna long, smooth and narrowed backward (Fig. 57); ventrites almost smooth, with fine punctures except for
<taxonomicName id="6A9CB0FCFFBE9F22FBF6FE440E5A55D5" authorityName="Mateu" authorityYear="1955" box="[1107,1283,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
<emphasis id="9FE8176DFFBE9F22FBF6FE440E5A55D5" box="[1107,1283,449,470]" italics="true" pageId="10" pageNumber="440">O. lacouri kocheri</emphasis>
</taxonomicName>
, that shows well-marked punctures; male mesotibia distally dilated, metatibia with marked crenulation in inner side of distal half.
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FD940F275658" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
Female genitalia (Fig. 58) with a short spermatic conduct followed by a progressively widened spermatheca of moderate length [
<bibRefCitation id="C90DB68EFFBE9F22FC3FFDC30F2A5658" author="Pupier R. &amp; Coulon J." box="[922,1139,582,604]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
].
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FCF7FDE40E2F5692" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
According to the detailed study of
<bibRefCitation id="C90DB68EFFBE9F22FB6AFDE408295692" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
, each subspecies has a “typical” shape of the median
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FE93F9420E1952E2" blockId="10.[310,649,1735,1761]" box="[310,1344,1735,1761]" lastBlockId="10.[862,1344,1735,1761]" pageId="10" pageNumber="440">
<emphasis id="9FE8176DFFBE9F22FE93F9420E1952E2" bold="true" box="[310,1344,1735,1761]" pageId="10" pageNumber="440">57 56 62 63 64 58</emphasis>
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47F9740E03537F" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
Figs 5164.
<taxonomicName id="6A9CB0FCFFBE9F22FEEAF9740A955306" box="[335,460,1777,1797]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFBE9F22FEEAF9740AFC5307" box="[335,421,1777,1796]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
spp
</taxonomicName>
: 5155 —
<taxonomicName id="6A9CB0FCFFBE9F22FD88F97409C65307" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[557,671,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFBE9F22FD88F97409C65307" box="[557,671,1777,1796]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
</taxonomicName>
; 5664 —
<taxonomicName id="6A9CB0FCFFBE9F22FCA6F97408055307" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[771,860,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
<emphasis id="9FE8176DFFBE9F22FCA6F97408055307" box="[771,860,1777,1796]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
</taxonomicName>
(5657, 63 —
<emphasis id="9FE8176DFFBE9F22FC46F9740F095307" box="[995,1104,1777,1796]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
(56 — female); 5860, 62, 64 —
<emphasis id="9FE8176DFFBE9F22FF18F88C0A73531F" box="[189,298,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
; 61 —
<emphasis id="9FE8176DFFBE9F22FECAF88C0A82531F" box="[367,475,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
); 51, 56 — habitus; 57 — metepisterna; 52 — punctures of abdominal segments; 53, 59 — median lobe of aedeagus; 54, 6164 — apex of median lobe of aedeagus; 55, 58 — genitalia of female; 51, 54, 56, 6164 — dorsal view; 5758, 60 — right lateral view; 53, 59 — left lateral view; 52 — ventral view; 51 — reservoir El Fakia,
<collectingCountry id="D58B8BEFFFBE9F22FC7CF8BC0F44534F" box="[985,1053,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
; 55, 61 — Sidi Daho ez Zahir,
<collectingCountry id="D58B8BEFFFBE9F22FA95F8BC0E2D534F" box="[1328,1396,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
; 58 — Jebel Tikermine,
<collectingCountry id="D58B8BEFFFBE9F22FE2FF8D70A825366" box="[394,475,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 62 — Tannezatte,
<collectingCountry id="D58B8BEFFFBE9F22FD21F8D7098C5366" box="[644,725,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 63 — Farkhana, Nador,
<collectingCountry id="D58B8BEFFFBE9F22FC14F8D70F5B5366" box="[945,1026,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 64 — Sidi Bouhouria, Berkane,
<collectingCountry id="D58B8BEFFFBE9F22FA86F8D70E2D5366" box="[1315,1396,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
; 5355, 5861 — after
<bibRefCitation id="C90DB68EFFBE9F22FE23F8EC0915537F" author="Pupier R. &amp; Coulon J." box="[390,588,1897,1916]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
; 6264 — after
<bibRefCitation id="C90DB68EFFBE9F22FD7BF8EC08F3537F" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[734,938,1897,1917]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
. Scale bars: 1.5 mm (Fig. 51); 1.6 mm (Fig. 56).
</paragraph>
<paragraph id="AD23CB7FFFBE9F22FF47F8000E695C13" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
Figs 5164.
<taxonomicName id="6A9CB0FCFFBE9F22FEE8F8000A9E539B" box="[333,455,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9FE8176DFFBE9F22FEE8F8000AF8539B" box="[333,417,1925,1944]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
spp
</taxonomicName>
: 5155 —
<taxonomicName id="6A9CB0FCFFBE9F22FD82F80009CE539B" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[551,663,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
<emphasis id="9FE8176DFFBE9F22FD82F80009CE539B" box="[551,663,1925,1944]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
</taxonomicName>
; 5664 —
<taxonomicName id="6A9CB0FCFFBE9F22FD5CF8000808539B" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[761,849,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
<emphasis id="9FE8176DFFBE9F22FD5CF8000808539B" box="[761,849,1925,1944]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
</taxonomicName>
(5657, 63 —
<emphasis id="9FE8176DFFBE9F22FC70F8000F66539B" box="[981,1087,1925,1944]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
(56 — самка); 5860, 62, 64 —
<emphasis id="9FE8176DFFBE9F22FAC7F8000A5353B3" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
; 61 —
<emphasis id="9FE8176DFFBE9F22FEE3F8180AF253B3" box="[326,427,1949,1968]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
); 51, 56 — внеШний вид; 57 — Эпистерн Заднегруди; 52 — пунктировка сегментов брюШка; 53, 59 — среднЯЯ долЯ Эдеагуса; 54, 6164 — верШина средней доли Эдеагуса; 55, 58 — гениталии самки; 51, 54, 56, 6164 — сверху; 5758, 60 — сбоку справа; 53, 59 — сбоку слева; 52 — сниЗу; 51 — водохранилиЩе Эль- ФакиЯ,
<collectingCountry id="D58B8BEFFFBE9F22FC35F848089653E3" box="[912,975,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
; 55, 61 — Сиди Дахо ЭЗ Захир,
<collectingCountry id="D58B8BEFFFBE9F22FB5CF8480E6153E3" box="[1273,1336,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
; 58 — Джебель Тикермин,
<collectingCountry id="D58B8BEFFFBE9F22FEDFF8600A9553FB" box="[378,460,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 62 — ТаннеЗатт,
<collectingCountry id="D58B8BEFFFBE9F22FDD0F860099E53FB" box="[629,711,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 63 — Фархана, Надор,
<collectingCountry id="D58B8BEFFFBE9F22FC0DF86008A353FB" box="[936,1018,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 64 — Сиди БухуриЯ, Беркане,
<collectingCountry id="D58B8BEFFFBE9F22FABBF8600E2953FB" box="[1310,1392,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
; 5355, 5861 — по
<bibRefCitation id="C90DB68EFFBE9F22FED0F87809625C13" author="Pupier R. &amp; Coulon J." box="[373,571,2045,2064]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
; 6264 — по
<bibRefCitation id="C90DB68EFFBE9F22FD19F87808D15C13" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[700,904,2045,2064]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
. МасШтаб: 1,5 мм (Рис. 51); 1,6 мм (Рис. 56).
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF18FEBE0A5F55D4" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">lobe of aedeagus, but this shape may vary notably depending on the degree of population gene flow, that results in “hybrid” shapes intermediate between those of “typical” exemplars. That paper should be consulted to check the variability; here I shall refer to the so-called “typical” shapes.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FE590AC856E2" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
In lateral view the distal half of the median lobe of aedeagus is almost straight in
<emphasis id="9FE8176DFFBF9F23FE4DFE73093C5608" box="[488,613,502,523]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
, sinuate in
<emphasis id="9FE8176DFFBF9F23FD7FFE730A5D5625" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
and
<emphasis id="9FE8176DFFBF9F23FE94FD940AFF5625" box="[305,422,529,550]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FEC5FD940AFF5625" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[352,422,529,550]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
, and is ventrally bent in
<emphasis id="9FE8176DFFBF9F23FD37FD94085E5625" box="[658,775,529,550]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
[
<bibRefCitation id="C90DB68EFFBF9F23FF67FDAE0AC35643" author="Pupier R. &amp; Coulon J." box="[194,410,555,577]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
] (Fig. 59). In dorsal view, the apical shaft is truncate and slightly sinuate in
<emphasis id="9FE8176DFFBF9F23FD93FDC309F55658" box="[566,684,582,603]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FDC3FDC309F55658" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[614,684,582,603]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
(Fig. 64) and truncate with a clear tooth in
<emphasis id="9FE8176DFFBF9F23FDA2FDE409265675" box="[519,639,609,630]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
(Fig. 61); the other two subspecies show a rounded apex, sinuate to the left, being wider in
<emphasis id="9FE8176DFFBF9F23FEFEFD130A8456A8" box="[347,477,662,683]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
[
<bibRefCitation id="C90DB68EFFBF9F23FE49FD13099056A8" author="Pupier R. &amp; Coulon J." box="[492,713,662,684]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
;
<bibRefCitation id="C90DB68EFFBF9F23FD7DFD130A2D56C5" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al., 2016</bibRefCitation>
] and slender and more elongate in
<emphasis id="9FE8176DFFBF9F23FD7CFD340A5D56E3" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
(Figs 6263).
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FD630A275798" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
According to the map provided by
<bibRefCitation id="C90DB68EFFBF9F23FD88FD63085E56FF" author="Chavanon G. &amp; Mabrouki Y. &amp; Taybi F. A." box="[557,775,742,764]" pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
the subspecies
<emphasis id="9FE8176DFFBF9F23FEF4FC840A975715" box="[337,462,769,790]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
is found west of River Muluya whereas
<emphasis id="9FE8176DFFBF9F23FEB6FC990AD55733" box="[275,396,795,817]" italics="true" pageId="11" pageNumber="441">
O. l.
<taxonomicName id="6A9CB0FCFFBF9F23FEE3FC9E0AD55733" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[326,396,795,816]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
</emphasis>
is found in the surroundings of Oujda and the Algerian border, to the west near Berkane, El Aioun and Guerfane;
<emphasis id="9FE8176DFFBF9F23FEEFFCD40A995765" box="[330,448,849,870]" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
is found in-between the other two subspecies. Hybrids between these subspecies are found in many contact areas.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,111 @@
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<paragraph id="AD23CB7FFFBF9F23FF4CFADC09835172" blockId="11.[189,776,1369,2071]" box="[233,730,1369,1393]" pageId="11" pageNumber="441">
<heading id="F66B7C13FFBF9F23FF4CFADC09835172" box="[233,730,1369,1393]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="441" reason="8">
<taxonomicName id="6A9CB0FCFFBF9F23FF4CFADC09835172" ID-CoL="7592B" authority="(Reiche et Saulcy, 1855)" baseAuthorityName="Reiche et Saulcy" baseAuthorityYear="1855" box="[233,730,1369,1393]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="berytensis">
<emphasis id="9FE8176DFFBF9F23FF4CFADC0A9E5172" box="[233,455,1369,1393]" italics="true" pageId="11" pageNumber="441">Orthomus berytensis</emphasis>
(Reiche et Saulcy, 1855)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E58698F4FFBF9F23FE07FAF308D356FF" pageId="11" pageNumber="441" type="description">
<paragraph id="AD23CB7FFFBF9F23FE07FAF3097B518D" blockId="11.[189,776,1369,2071]" box="[418,546,1398,1422]" pageId="11" pageNumber="441">
<figureCitation id="35A7D7FAFFBF9F23FE07FAF30947518D" box="[418,542,1398,1422]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Figs 6570</figureCitation>
.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47FA190A54521F" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">Dorsum black to dark brown, often with reddish hue, shiny in both sexes (females show fine microreticulation), appendices with lighter coloration brown red. Body length variable 8.011.0 mm. Eye convex, protruding, tempora oblique.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F9A4095F5325" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">
Pronotum transverse almost rectangular, regularly curved, well narrowed forward and moderately backward, maximum width near middle (
<figureCitation id="35A7D7FAFFBF9F23FED9F9D30A9C526F" box="[380,453,1622,1644]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 65</figureCitation>
), anterior angle briefly projected forward, anterior basis almost regularly arcuate; posterior angle obtuse with a tiny tooth (
<figureCitation id="35A7D7FAFFBF9F23FDABF90E090552A2" box="[526,604,1675,1697]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 66</figureCitation>
), posterior basis beaded up to the origin of the inner fovea; external fovea weakly impressed, inner fovea linear; punctures usually in low density or entirely absent (in East Mediterranean populations punctures are more abundant); space between external fovea and side somewhat convex.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F8AE09D45375" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">Elytra slightly enlarged from basis, humerus with a small and non-protruding tooth, striae smooth or with traces of punctures, intervals flat, discal pores on stria 3.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F8FE0A6F53E2" blockId="11.[189,776,1369,2071]" pageId="11" pageNumber="441">
Prosternal process slightly truncate; ventral segments with shallow punctures at sides (
<figureCitation id="35A7D7FAFFBF9F23FE4DF813097453AF" box="[488,557,1942,1964]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 67</figureCitation>
), metepisterna smooth. Male mesotibia slightly dilated apically, metatibia moderately crenulate.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FF47F86308D356FF" blockId="11.[189,776,1369,2071]" lastBlockId="11.[812,1399,315,764]" pageId="11" pageNumber="441">
Apical shaft of median lobe of aedeagus wide and with trapezoid shape, only briefly narrowed to tip (
<figureCitation id="35A7D7FAFFBF9F23FD36F78409865C15" box="[659,735,2048,2071]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 68</figureCitation>
), in lateral view somewhat ventrally bent (
<figureCitation id="35A7D7FAFFBF9F23FB0CFEBE0FAA5552" box="[1193,1267,315,337]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 69</figureCitation>
). The female genitalia match in all details the characters described by
<bibRefCitation id="C90DB68EFFBF9F23FC89FEF408E75585" author="Ortuno V. M." box="[812,958,368,390]" pageId="11" pageNumber="441" pagination="261 - 274" refId="ref17074" refString="Ortuno V. M. 1996. La genitalia femenina en los Poecilini (Caraboidea, Pterostichidae) de la Peninsula Iberica I. El genero Orthomus Chaudoir, 1838 // Nouv. Rev. Ent. (nouvelle serie). Vol. 13. No. 3. P. 261 - 274." type="journal article" year="1996">Ortuño [1996]</bibRefCitation>
for the
<taxonomicName id="6A9CB0FCFFBF9F23FBBBFEF40FC35585" baseAuthorityName="Dejean" baseAuthorityYear="1828" box="[1054,1178,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="9FE8176DFFBF9F23FBBBFEF40FC35585" box="[1054,1178,369,390]" italics="true" pageId="11" pageNumber="441">O. barbarus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFBF9F23FB72FEF40E2F5585" baseAuthorityName="Waltl" baseAuthorityYear="1835" box="[1239,1398,369,390]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="velocissimus">
<emphasis id="9FE8176DFFBF9F23FB72FEF40E2F5585" box="[1239,1398,369,390]" italics="true" pageId="11" pageNumber="441">O. velocissimus</emphasis>
</taxonomicName>
groups, in that there is a long spermathecal duct (where a caecum is originated) followed by a long a slightly dilated spermatheca toward apex (
<figureCitation id="35A7D7FAFFBF9F23FB9CFE440FDC55D5" box="[1081,1157,449,471]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 70</figureCitation>
). Therefore, this pattern clearly differs from that found in the other Maghrebian taxa [
<bibRefCitation id="C90DB68EFFBF9F23FC97FE730F4D5608" author="Pupier R. &amp; Coulon J." box="[818,1044,502,524]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
]. These findings suggests that the species is more closely related to the north Mediterranean lineages than to the Maghrebian ones. However, the species occupies the whole North Africa, and to the east occupies
<collectingCountry id="D58B8BEFFFBF9F23FC89FDE4083D5674" box="[812,868,609,631]" name="Israel" pageId="11" pageNumber="441">Israel</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FCCBFDE4089C5674" box="[878,965,609,631]" name="Lebanon" pageId="11" pageNumber="441">Lebanon</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FC6AFDE40F5C5674" box="[975,1029,609,631]" name="Syria" pageId="11" pageNumber="441">Syria</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FBABFDE40F0F5674" box="[1038,1110,609,631]" name="Turkey" pageId="11" pageNumber="441">Turkey</collectingCountry>
,
<collectingCountry id="D58B8BEFFFBF9F23FBC5FDE40FFF5675" box="[1120,1190,609,630]" name="Cyprus" pageId="11" pageNumber="441">Cyprus</collectingCountry>
and
<collectingCountry id="D58B8BEFFFBF9F23FB70FDE40E455675" box="[1237,1308,609,630]" name="Greece" pageId="11" pageNumber="441">Greece</collectingCountry>
, that is, a North African geographic pattern. Indeed, it is also found in Sicily, Sardinia and
<collectingCountry id="D58B8BEFFFBF9F23FC55FD130F7256AF" box="[1008,1067,662,684]" name="Malta" pageId="11" pageNumber="441">Malta</collectingCountry>
, and to west reaches the southernmost parts of
<collectingCountry id="D58B8BEFFFBF9F23FC10FD340F5556C5" box="[949,1036,689,710]" name="Morocco" pageId="11" pageNumber="441">Morocco</collectingCountry>
and the Canary Islands. Analyses of new characters are needed to better assess its phylogenetic affinities.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="E58698F4FFBF9F23FC2BFC950E4D572B" box="[910,1300,784,808]" pageId="11" pageNumber="441" type="nomenclature">
<paragraph id="AD23CB7FFFBF9F23FC2BFC950E4D572B" blockId="11.[812,1399,784,2072]" box="[910,1300,784,808]" pageId="11" pageNumber="441">
<heading id="F66B7C13FFBF9F23FC2BFC950E4D572B" box="[910,1300,784,808]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="441" reason="8">
<taxonomicName id="6A9CB0FCFFBF9F23FC2BFC950E4D572B" ID-CoL="6SWQC" authority="Antoine, 1933" authorityName="Antoine" authorityYear="1933" box="[910,1300,784,808]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="9FE8176DFFBF9F23FC2BFC950F2B572B" box="[910,1138,784,808]" italics="true" pageId="11" pageNumber="441">Orthomus dimorphus</emphasis>
Antoine, 1933
</taxonomicName>
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<paragraph id="AD23CB7FFFBF9F23FBB7FCA80FCB5746" blockId="11.[812,1399,784,2072]" box="[1042,1170,813,837]" pageId="11" pageNumber="441">
<figureCitation id="35A7D7FAFFBF9F23FBB7FCA80FD75746" box="[1042,1166,813,837]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Figs 7176</figureCitation>
.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FCD708F257EE" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
This species has two subspecies,
<taxonomicName id="6A9CB0FCFFBF9F23FB3FFCD708F05781" authority="Mateu, 1955" authorityName="Mateu" authorityYear="1955" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="antoinei">
<emphasis id="9FE8176DFFBF9F23FB3FFCD70E2F576B" box="[1178,1398,850,872]" italics="true" pageId="11" pageNumber="441">O. dimorphus antoinei</emphasis>
<bibRefCitation id="C90DB68EFFBF9F23FC89FCE808F05781" author="Mateu J." box="[812,937,876,898]" pageId="11" pageNumber="441" pagination="53 - 85" refId="ref17020" refString="Mateu J. 1955. Notas sobre los Orthomus Chaudoir (segunda nota) // Eos. Vol. 31. P. 53 - 85." type="book chapter" year="1955">Mateu, 1955</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="6A9CB0FCFFBF9F23FC45FCE80E2A5781" authority="Antoine, 1933" authorityName="Antoine" authorityYear="1933" box="[992,1395,877,899]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="dimorphus">
<emphasis id="9FE8176DFFBF9F23FC45FCE80F815781" box="[992,1240,877,898]" italics="true" pageId="11" pageNumber="441">O. dimorphus dimorphus</emphasis>
Antoine, 1933
</taxonomicName>
. The following description is based on the first one, that is more common in Atlantic
<collectingCountry id="D58B8BEFFFBF9F23FB8CFC270FD957B4" box="[1065,1152,930,951]" name="Morocco" pageId="11" pageNumber="441">Morocco</collectingCountry>
than the nominal subspecies, that is restricted to some localities of Great Atlas over
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.a.s.l.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FC770F505070" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">Dorsum dark brown, appendices brown, male shiny, female mat, microreticulation well visible in females, weak in males. Body length variable within populations and between subspecies, 7.012.0 mm. Eyes moderately convex and protruding, tempora long.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FBFD0F7C512E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Pronotum somewhat trapezoid, maximum width after middle (
<figureCitation id="35A7D7FAFFBF9F23FC25FB17089250A4" box="[896,971,1170,1192]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 71</figureCitation>
), side curved and narrowed forward, only narrowed backward near posterior angle; this shows a small protruding tooth; posterior basis beaded up to inner linear fovea; external fovea variably impressed; punctures variable in density and size, usually fine and in moderate density; postangular field convex.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FAB708B2519B" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">Elytra slightly widened from basis; humerus with a small and non-protruding tooth, parascutellar stria well-developed with proximal seta; striae smooth but with traces of punctures; intervals flat.</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7FA180E29521E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Prosternal process somewhat rounded. Metepisterna smooth, long and narrowed backward (
<figureCitation id="35A7D7FAFFBF9F23FB62FA320E4E51CF" box="[1223,1303,1463,1485]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 72</figureCitation>
). Ventral segments laterally smooth or with fine and few punctures. Male mesotibia not distally dilated; male metatibia only slightly curved and without crenulation in distal inner side.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F9A70894528B" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Median lobe of aedeagus with an elongate apical shaft (
<figureCitation id="35A7D7FAFFBF9F23FC96F9B808225251" box="[819,891,1597,1619]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 73</figureCitation>
), almost evenly narrowed apicad, and almost straight in lateral view (
<figureCitation id="35A7D7FAFFBF9F23FC65F9D20F5E526F" box="[960,1031,1623,1645]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 74</figureCitation>
); membranous area notably prolonged toward the bulb.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F9080E45530E" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
Spermatheca notably longer than duct (
<figureCitation id="35A7D7FAFFBF9F23FB45F9080E7252A1" box="[1248,1323,1676,1699]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 75</figureCitation>
), it becomes notably thick after fertilization (
<figureCitation id="35A7D7FAFFBF9F23FB0AF9220FA352BE" box="[1199,1274,1703,1725]" captionStart="Figs 6576" captionStartId="12.[226,263,1801,1820]" captionTargetBox="[189,1399,534,1748]" captionTargetId="figure-121@12.[189,1399,526,1743]" captionTargetPageId="12" captionText="Figs 6576. Orthomus spp, specimens from Morocco: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (in part) — habitus; 66 — posterior angle of pronotum (in part); 67 — metepisterna and ventrites (in part); 72 — metepisterna; 6869, 7374 — median lobe of aedeagus; 70 — spermatheca and spermathecal duct (in part); 75 — empty spermatheca of young female; 76 - full spermatheca of mature female; 65, 68, 71, 73 — dorsal view; 66 — ventral view;69, 74 — left lateral view; 72 — right lateral view; 67, 70 — Essaouira; 6869 — Agadir; 73, 76 — Rabat (plage); 75 — Meknés. Scale bars: 1.5 mm (Fig. 71); 2.0 mm (Fig. 65). Рис. 6576. Orthomus spp, особи иЗ Марокко: 6570 — O. berytensis; 7176 — O. dimorphus antoinei; 65, 71 (частично) — внеШний вид; 66 — Задний угол переднеспинки (частично); 67 — Эпистерн Заднегруди и вентриты (частично); 72 — Эпистерн Заднегруди; 6869, 7374 — среднЯЯ долЯ Эдеагуса; 70 — сперматека и её проток (частично); 75 — пустаЯ сперматека неполовоЗрелой самки; 76 — полнаЯ сперматека половоЗрелой самки; 65, 68, 71, 73 — сверху; 66 — сниЗу; 69, 74 — сбоку слева; 72 — сбоку справа; 67, 70 — Эс-Сувейра; 6869 — Агадир; 73, 76 — Рабат (плЯж); 75 — Мекнес. МасШтаб: 15 мм (Рис. 71); 20 мм (Рис. 65)." figureDoi="http://doi.org/10.5281/zenodo.13178306" httpUri="https://zenodo.org/record/13178306/files/figure.png" pageId="11" pageNumber="441">Fig. 76</figureCitation>
). This shape difference within the same species calls attention on the variable aspect that may take the female genitalia of
<taxonomicName id="6A9CB0FCFFBF9F23FAF6F958082B530F" authorityName="Chaudoir" authorityYear="1838" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="genus">
<emphasis id="9FE8176DFFBF9F23FAF6F958082B530F" italics="true" pageId="11" pageNumber="441">Orthomus</emphasis>
</taxonomicName>
depending on matureness and fertilization.
</paragraph>
<paragraph id="AD23CB7FFFBF9F23FCF7F897082753CB" blockId="11.[812,1399,784,2072]" pageId="11" pageNumber="441">
The subspecies
<taxonomicName id="6A9CB0FCFFBF9F23FC51F8970FB65324" authorityName="Antoine" authorityYear="1933" box="[1012,1263,1810,1831]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="dimorphus">
<emphasis id="9FE8176DFFBF9F23FC51F8970FB65324" box="[1012,1263,1810,1831]" italics="true" pageId="11" pageNumber="441">O. dimorphus dimorphus</emphasis>
</taxonomicName>
occupies the Great Atlas above
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. Its body length is smaller (less than
<quantity id="6A64669AFFBF9F23FCF9F8C208FB535F" box="[860,930,1863,1884]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="441" unit="mm" value="10.0">10 mm</quantity>
), the female is mat, the pronotum basis is almost smooth, the humeral tooth is small, the striae are almost smooth, and the median lobe of aedeagus is ventrally curved upwards, with an apical shaft that forms a relatively short triangle.
</paragraph>
<paragraph id="AD23CB7FFFBF9F24FCF7F8480A1455F2" blockId="11.[812,1399,784,2072]" lastBlockId="12.[189,775,315,497]" lastPageId="12" lastPageNumber="442" pageId="11" pageNumber="441">
The subspecies
<taxonomicName id="6A9CB0FCFFBF9F23FC4BF8480F9253E1" authorityName="Mateu" authorityYear="1955" box="[1006,1227,1997,2018]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="subSpecies" species="dimorphus" subSpecies="antoinei">
<emphasis id="9FE8176DFFBF9F23FC4BF8480F9253E1" box="[1006,1227,1997,2018]" italics="true" pageId="11" pageNumber="441">O. dimorphus antoinei</emphasis>
</taxonomicName>
is of larger body length and both sexes are dorsally shiny. The pronotum basis is punctured, the external fovea is better marked, the striae of elytron are well impressed and punctured, the humeral tooth is developed. The median lobe of aedeagus is almost straight on distal half in lateral view; in dorsal view it shows a more elongate and triangular apical shaft. This subspecies is found in the Atlantic plain of
<collectingCountry id="D58B8BEFFFB89F24FE3AFE230AAF55B8" box="[415,502,422,443]" name="Morocco" pageId="12" pageNumber="442">Morocco</collectingCountry>
and doesnt enter in the Rif and the Great Atlas above
<quantity id="6A64669AFFB89F24FE6DFE44094F55D5" box="[456,534,449,470]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="442" unit="m" value="2000.0">2000 m</quantity>
. See
<bibRefCitation id="C90DB68EFFB89F24FDE9FE4409BB55D5" author="Antoine M." box="[588,738,448,471]" pageId="12" pageNumber="442" pagination="179 - 314" refId="ref16250" refString="Antoine M. 1957. Coleopteres Carabiques du Maroc (deuxieme partie) // Memoires de la Societe des Sciences Naturelles et Physiques du Maroc, Zoologie (nouvelle serie). Vol. 3. P. 179 - 314." type="journal article" year="1957">Antoine [1957]</bibRefCitation>
for further details.
</paragraph>
<paragraph id="AD23CB7FFFB89F24FE2CF95B0AFF52FB" blockId="12.[393,653,1722,1784]" box="[393,422,1758,1784]" pageId="12" pageNumber="442">
<emphasis id="9FE8176DFFB89F24FE2CF95B0AFF52FB" bold="true" box="[393,422,1758,1784]" pageId="12" pageNumber="442">71</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>