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<document id="19AD245C25DFEB37D1EC064659DE76CE" ID-DOI="10.15298/rusentj.31.2.19" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178200" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593805114" checkinUser="felipe" docAuthor="Kustov, S. Yu. &amp; Shamshev, I. V." docDate="2022" docId="03D2404EB730FF9FA762D15862DAFADC" docLanguage="en" docName="RussEntomolJ.31.2.194-200.pdf" docOrigin="Russian Entomological Journal 31 (2)" docSource="http://dx.doi.org/10.15298/rusentj.31.2.19" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Empis (Empis) windhoekiensis Kustov et Shamshev 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="199" masterDocId="FFEB3836B734FF9AA44CD35E663BFFA0" masterDocTitle="Three new species of empidid dance flies of the genus Empis Linnaeus, 1758 (Diptera: Empididae) from Namibia" masterLastPageNumber="200" masterPageNumber="194" pageNumber="198" updateTime="1722633502547" updateUser="ExternalLinkService">
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<mods:title id="A18D77C4AADC08CE2D450C07D89D4022">Three new species of empidid dance flies of the genus Empis Linnaeus, 1758 (Diptera: Empididae) from Namibia</mods:title>
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<mods:namePart id="05CCA936C3D1503128C10EE5D8DAE189">Kustov, S. Yu.</mods:namePart>
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<mods:namePart id="3C8E0A8F76838197A9E2CE1289DBEE98">Shamshev, I. V.</mods:namePart>
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<mods:title id="F467490FFD7D31607FDD0B7FA96A63A1">Russian Entomological Journal</mods:title>
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<mods:date id="3BBA8F9B18ADBE630A2EC7C350000170">2022</mods:date>
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<mods:number id="21535C8AC4BFE45BDB93FADC1601C4FC">2022-06-30</mods:number>
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<treatment id="03D2404EB730FF9FA762D15862DAFADC" LSID="urn:lsid:plazi:treatment:03D2404EB730FF9FA762D15862DAFADC" httpUri="http://treatment.plazi.org/id/03D2404EB730FF9FA762D15862DAFADC" lastPageId="5" lastPageNumber="199" pageId="4" pageNumber="198">
<subSubSection id="C361A2D3B730FF9EA762D158634FFDBD" box="[814,1396,517,541]" pageId="4" pageNumber="198" type="nomenclature">
<paragraph id="8BC4F158B730FF9EA762D158634FFDBD" blockId="4.[812,1400,517,1291]" box="[814,1396,517,541]" pageId="4" pageNumber="198">
<heading id="D08C4634B730FF9EA762D158634FFDBD" box="[814,1396,517,541]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="198" reason="4">
<taxonomicName id="4C7B8ADBB730FF9EA762D1586303FDBD" ID-CoL="9J5TP" authority="Kustov et Shamshev" authorityName="Kustov et Shamshev" authorityYear="2022" box="[814,1336,517,541]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA762D1586556FDBD" box="[814,877,518,541]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
(
<emphasis id="B90F2D4AB730FF9EA737D1586581FDBD" box="[891,954,518,541]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA785D15B6259FDBD" box="[969,1122,517,541]" italics="true" pageId="4" pageNumber="198">windhoekiensis</emphasis>
Kustov et Shamshev
</taxonomicName>
,
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<taxonomicNameLabel id="A23C9031B730FF9EA10ED15B634FFDBD" box="[1346,1396,517,541]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA06DD17C62B9FD9A" box="[1057,1154,546,570]" pageId="4" pageNumber="198" type="description">
<paragraph id="8BC4F158B730FF9EA06DD17C62B9FD9A" blockId="4.[812,1400,517,1291]" box="[1057,1154,546,570]" pageId="4" pageNumber="198">
<figureCitation id="1340EDDDB730FF9EA06DD17C6246FD9A" box="[1057,1149,546,570]" captionStart="Figs 78" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[212,1396,1334,1928]" captionTargetId="figure-687@4.[189,1399,1334,1924]" captionTargetPageId="4" captionText="Figs 78. Empis (Empis) windhoekiensis, sp.n., ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 78. Empis (Empis) windhoekiensis, sp.n., ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="4" pageNumber="198">Figs 78</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED1166360FD2A" pageId="4" pageNumber="198" type="materials_examined">
<paragraph id="8BC4F158B730FF9EA71ED1166360FD2A" blockId="4.[812,1400,517,1291]" pageId="4" pageNumber="198">
<collectionCode id="ED6A699DB730FF9EA71ED116623FFDFA" box="[850,1028,583,603]" pageId="4" pageNumber="198">TYPE MATERIAL</collectionCode>
.
<materialsCitation id="3B13FB05B730FF9EA05DD119634FFDD2" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="4" pageNumber="198" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C04FFAB730FF9EA05DD119625AFDFA" box="[1041,1121,583,602]" pageId="4" pageNumber="198" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7D3AD1B730FF9EA027D11B6244FDFB" box="[1131,1151,581,603]" count="1" pageId="4" pageNumber="198" type="male"></specimenCount>
,
<collectingCountry id="F36CB1C8B730FF9EA0C2D11962E1FDFA" box="[1166,1242,583,602]" name="Namibia" pageId="4" pageNumber="198">Namibia</collectingCountry>
,
<location id="8EA4A783B730FF9EA0A4D1196379FDFA" LSID="urn:lsid:plazi:treatment:03D2404EB730FF9FA762D15862DAFADC:8EA4A783B730FF9EA0A4D1196379FDFA" box="[1256,1346,583,602]" country="Namibia" latitude="-22.545" longLatPrecision="75" longitude="17.255" name="Windhoek" pageId="4" pageNumber="198">Windhoek</location>
env.,
<geoCoordinate id="EE4F979FB730FF9EA760D1016546FDD2" box="[812,893,607,626]" degrees="22.545" direction="south" orientation="latitude" pageId="4" pageNumber="198" precision="55" value="-22.545">22.545°S</geoCoordinate>
<geoCoordinate id="EE4F979FB730FF9EA7C8D10165ECFDD2" box="[900,983,607,626]" degrees="17.255" direction="east" orientation="longitude" pageId="4" pageNumber="198" precision="55" value="17.255">17.255°E</geoCoordinate>
,
<quantity id="4C835CBDB730FF9EA7AFD1016212FDD3" box="[995,1065,607,627]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="198" unit="m" value="1870.0">
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</quantity>
,
<date id="FFC5D798B730FF9EA078D1016293FDD2" box="[1076,1192,607,626]" pageId="4" pageNumber="198" value="2021-01-28" valueMax="2021-01-31" valueMin="2021-01-28">
<collectingDate id="EF812E70B730FF9EA078D1016293FDD2" box="[1076,1192,607,626]" pageId="4" pageNumber="198" value="2021-01-28" valueMax="2021-01-31" valueMin="2021-01-28">2831.i.2021</collectingDate>
</date>
,
<collectorName id="268E948EB730FF9EA0F8D13E6323FDD2" box="[1204,1304,607,627]" pageId="4" pageNumber="198">N. Vikhrev</collectorName>
(
<collectionCode id="ED6A699DB730FF9EA16AD1016355FDD2" box="[1318,1390,607,626]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="198" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B13FB05B730FF9EA760D126636DFD2A" box="[812,1366,629,651]" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU, ZISP" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="4" pageNumber="198" specimenCount="9" specimenCount-male="9" typeStatus="paratype">
<typeStatus id="54C04FFAB730FF9EA760D12665B8FD2B" box="[812,899,632,651]" pageId="4" pageNumber="198" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7D3AD1B730FF9EA7C1D1296589FD2B" box="[909,946,629,651]" count="8" pageId="4" pageNumber="198" type="male">8 ♂</specimenCount>
, same data as holotype (
<collectionCode id="ED6A699DB730FF9EA0C6D12962EBFD2A" box="[1162,1232,631,650]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="198" type="Museum">ZMMU</collectionCode>
,
<specimenCount id="9D7D3AD1B730FF9EA095D12962C5FD2B" box="[1241,1278,629,651]" count="1" pageId="4" pageNumber="198" type="male">1 ♂</specimenCount>
<collectionCode id="ED6A699DB730FF9EA16CD129636BFD2A" box="[1312,1360,631,650]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="198">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED1D16563FCE5" pageId="4" pageNumber="198" type="diagnosis">
<paragraph id="8BC4F158B730FF9EA71ED1D16563FCE5" blockId="4.[812,1400,517,1291]" pageId="4" pageNumber="198">
DIAGNOSIS. A species of
<emphasis id="B90F2D4AB730FF9EA015D1CE6253FD05" box="[1113,1128,656,677]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<taxonomicName id="4C7B8ADBB730FF9EA03FD1D162C2FD05" authorityName="Smith" authorityYear="1969" box="[1139,1273,655,677]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
(
<emphasis id="B90F2D4AB730FF9EA037D1CE62B1FD05" box="[1147,1162,656,677]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.)
<emphasis id="B90F2D4AB730FF9EA0D1D1CE62C2FD05" box="[1181,1273,656,677]" italics="true" pageId="4" pageNumber="198">setitarsus</emphasis>
</taxonomicName>
-group; body
<quantity id="4C835CBDB730FF9EA760D1F46593FD1F" box="[812,936,682,704]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="198" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.54.5 mm</quantity>
; proboscis short, labrum only slightly longer than head height; laterotergite with black setae; abdomen greyish pruinose, only pale setose; wing with incomplete veins M
<subScript id="17FFF31DB730FF9EA730D05765B8FCB5" attach="left" box="[892,899,777,789]" fontSize="5" pageId="4" pageNumber="198">1</subScript>
and CuA+CuP, pterostigma pale yellowish, halter brown. Male: fore basitarsus slightly thickened; wing hyaline.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9FA71ED014622AFCC7" lastPageId="5" lastPageNumber="199" pageId="4" pageNumber="198" type="description">
<paragraph id="8BC4F158B730FF9FA71ED0146490FE27" blockId="4.[812,1400,517,1291]" lastBlockId="5.[189,776,315,2071]" lastPageId="5" lastPageNumber="199" pageId="4" pageNumber="198">
DESCRIPTION. Body length 3.54.2, wing
<quantity id="4C835CBDB730FF9EA0B1D014634FFCFF" box="[1277,1396,842,864]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="4" pageNumber="198" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.24.5 mm</quantity>
.
<emphasis id="B90F2D4AB730FF9EA760D03B655BFCDA" bold="true" box="[812,864,869,890]" pageId="4" pageNumber="198">Male</emphasis>
(
<figureCitation id="1340EDDDB730FF9EA73ED03B6589FCDA" box="[882,946,869,891]" captionStart="Figs 78" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[212,1396,1334,1928]" captionTargetId="figure-687@4.[189,1399,1334,1924]" captionTargetPageId="4" captionText="Figs 78. Empis (Empis) windhoekiensis, sp.n., ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 78. Empis (Empis) windhoekiensis, sp.n., ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="4" pageNumber="198">Fig. 7</figureCitation>
). Head with greyish, dense pruinescence on frons, face, clypeus (except shiny upper margin), ocellar triangle, occiput and postgena; black setose. Eyes holoptic, with upper ommatidia enlarged. Frons represented by very small, subtriangular space just below ocellar triangle and slightly larger space just above antennae. Face broad, almost parallel-sided, bare. Ocellar triangle with several fine setae, 2 setae somewhat longer. Occiput with fine, moderately long postoculars and several similar setae laterally; postgena with numerous hair-like setae. Antenna black; scape short, slightly longer than pedicel, both with short setulae; postpedicel elongate-conical, with lower margin slightly concave distally, nearly 2.5X as long as basal width; stylus short, nearly 2.5X shorter than postpedicel. Proboscis short, with labrum only slightly longer than head height (1.21.3X); labium lacking annulations, bearing scattered, minute setulae; labella moderately long, nearly 2X shorter than labrum, with small desclerotized space on about distal 1/3 (visible
<emphasis id="B90F2D4AB731FF9FA6EBD20964DCFECC" box="[679,743,343,364]" italics="true" pageId="5" pageNumber="199">in situ</emphasis>
as notch); palpus black, with black, scattered setulae.
</paragraph>
<caption id="DF04A1D0B730FF9EA4AED4EF625DF7B1" ID-DOI="http://doi.org/10.5281/zenodo.13178206" ID-Zenodo-Dep="13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="4" pageNumber="198" startId="4.[226,263,1969,1988]" targetBox="[212,1396,1334,1928]" targetPageId="4" targetType="figure">
<paragraph id="8BC4F158B730FF9EA4AED4EF626BF87C" blockId="4.[188,1398,1967,2065]" pageId="4" pageNumber="198">
Figs 78.
<taxonomicName id="4C7B8ADBB730FF9EA574D4EC647BF864" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[312,576,1969,1988]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA574D4EC6756F864" box="[312,365,1970,1988]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
(
<emphasis id="B90F2D4AB730FF9EA536D4EC6794F864" box="[378,431,1970,1988]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA5F1D4EF647BF864" box="[445,576,1969,1988]" italics="true" pageId="4" pageNumber="198">windhoekiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB730FF9EA607D4EF644DF864" bold="true" box="[587,630,1969,1988]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA607D4EF644DF864" box="[587,630,1969,1988]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus.
</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED4B8625DF7B1" blockId="4.[188,1398,1967,2065]" pageId="4" pageNumber="198">
Рис. 78.
<taxonomicName id="4C7B8ADBB730FF9EA57BD4B86407F858" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[311,572,2021,2040]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA57BD4B86757F858" box="[311,364,2022,2040]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
(
<emphasis id="B90F2D4AB730FF9EA53BD4B86797F858" box="[375,428,2022,2040]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA5F5D4BB6407F858" box="[441,572,2021,2040]" italics="true" pageId="4" pageNumber="198">windhoekiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB730FF9EA609D4BB644BF858" bold="true" box="[581,624,2021,2040]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA609D4BB644BF858" box="[581,624,2021,2040]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус.
</paragraph>
</caption>
<paragraph id="8BC4F158B731FF9FA4AED2D5645DFC21" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Thorax black in ground-colour, black setose; mesoscutum faintly greyish pruinose (dorsal view), prothoracic sclerites and mesopleuron somewhat denser greyish pruinose.Proepisternum with 45 rather short, fine setae on lower part and 46 similar setae on upper part. Prosternum bare. Antepronotum with 3 moderately long setae on each side. Postpronotal lobe with 1 strong, long and several fine, short setae. Mesonotal setae well-differentiated; acrostichals moderately long, thin, arranged in 2 close, irregular rows, lacking on prescutellar depression; dorsocentrals longer than acrostichals, 12-serial before suture, uniserial and longer along prescutellar depression, 2 prescutellar setae longest; 1 fine, moderately long presutural intra-alar, 1 longer presutural supra-alar, 3 notopleurals (2 setae stronger; additionally, with some setulae anteriorly), 1 postsutural supra-alar (with 34 additional short, fine setae anteriorly), 1 long and 1 minute postalars, 4 scutellars (apical setae long and cruciate, lateral setae only slightly shorter). Laterotergite with 56 long and some short fine setae. Anterior and posterior spiracles dark brown.</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED0D864B2F927" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Legs rather robust, black, subshiny (except coxae), faintly greyish pruinose, black setose; coxae densely pruinose. Fore coxa and trochanter almost bare, only with scattered minute setulae; hind trochanter with long, dense setae. Fore femur with row of short anteroventral fine setae on about basal half, almost bare posteroventrally, only some minute posteroventral setulae closer to base. Fore tibia with numerous long, fine setae posterodorsally (longest setae about 2X as long as tibia width). Fore basitarsus thickened but only slightly stouter than fore tibia at apex; with moderately long, dense, fine setae posterodorsally; tarsomeres 14 with rather long setae of subapical circlet. Mid femur with complete rows of numerous anteroventral and posteroventral setae, which are long on about middle part of femur and short near its base and apex; longest anteroventral setae nearly 1.5X as long as femur middle width. Mid tibia with several intermixed moderately long to long ad setae closer to base, more or less distinct row of moderately long posterodorsal setae and rows of moderately long anteroventral and short posteroventral setae. Mid basitarsus nearly as broad as mid tibia at apex; tarsomeres 1 and 2 with short spine-like setae ventrally, tarsomeres 13 with rather long setae of subapical circlet (longer dorsally). Hind femur with complete rows of long anteroventral (longest setae on about apical half nearly 1.5X as long as femur middle width) and short posteroventral setae. Hind tibia slightly uniformly thickened (lateral view) and somewhat curved (posterior view); with numerous moderately long to long setae dorsally; no seta in posteroapical comb. Hind basitarsus slender; with dorsal setation as on hind tibia, numerous short, spine-like setae ventrally.</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED5D567A3F8FC" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">
Wing membrane hyaline, veins mostly brownish, paler proximally; M
<subScript id="17FFF31DB731FF9FA500D5EB6768F961" attach="left" box="[332,339,1717,1729]" fontSize="5" pageId="5" pageNumber="199">1</subScript>
beyond cell dm and CuA+CuP (anal vein) very thin, M
<subScript id="17FFF31DB731FF9FA579D58E6707F97C" attach="left" box="[309,316,1744,1756]" fontSize="5" pageId="5" pageNumber="199">2</subScript>
and M
<subScript id="17FFF31DB731FF9FA5CDD58E67B3F97C" attach="left" box="[385,392,1744,1756]" fontSize="5" pageId="5" pageNumber="199">4</subScript>
weakened before wing margin; M
<subScript id="17FFF31DB731FF9FA69AD58E64E6F97C" attach="left" box="[726,733,1744,1756]" fontSize="5" pageId="5" pageNumber="199">1</subScript>
and CuA+CuP incomplete; cell dm short, with slightly elongate apex. Pterostigma pale yellowish, hardly distinguishable. Basal costal seta present, long, black. Anal lobe well-developed; axillary incision right-angled. Squama brownish, pale fringed. Halter brown.
</paragraph>
<paragraph id="8BC4F158B731FF9FA4AED43F643AF7B7" blockId="5.[189,776,315,2071]" pageId="5" pageNumber="199">Abdomen black in ground-colour, uniformly greyish pruinose, mostly covered with numerous pale hair-like setae longer on tergites laterally; sternite 7 with some fine black posteromarginal setae, segment 8 with black setae of different lengths. Segment 8 with separated sclerites; tergite 8 without projections, entire, somewhat concave posteriorly; sternite 8 desclerotized medially.</paragraph>
<paragraph id="8BC4F158B731FF9FA71ED265626CFCEC" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
Terminalia (
<figureCitation id="1340EDDDB731FF9FA787D2656233FEF0" box="[971,1032,315,337]" captionStart="Figs 78" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[212,1396,1334,1928]" captionTargetId="figure-687@4.[189,1399,1334,1924]" captionTargetPageId="4" captionText="Figs 78. Empis (Empis) windhoekiensis, sp.n., ♂ lateral view: 7 — habitus,; 8 — hypopygium. Abbreviations: cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 78. Empis (Empis) windhoekiensis, sp.n., ♂, сбоку: 7 — габитус; 8 — гипопигий. СокраЩениЯ: cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178206" httpUri="https://zenodo.org/record/13178206/files/figure.png" pageId="5" pageNumber="199">Fig. 8</figureCitation>
) moderately large, concolorous with abdomen (except noted), black setose. Epandrium divided into two lamellae (dorsal bridge absent); epandrial lamella subtriangular (lateral view), with rather deep excision closer to base dorsally, with long, fine setae along lateral margins. Hypandrium short, subtriangular (ventral view), with 2 long and 2 short setae. Hypoproct strongly sclerotised, elongate, reaching phallus, narrow on middle part and broadened distally, bare. Phallus brownish yellow, mostly hidden, short, evenly curved, thick on about basal 2/3 and slender on apical 1/3. Ejaculatory apodeme large, extended far beyond basal curvature of phallus, without lateral wings. Cerci closely attached and almost fused anteriorly, separated from epandrium; cercus almost entirely divided in two lobes by narrow notch (lateral view) posteriorly; dorsal lobe rather trapezoid (lateral view), with short, dense, strong setae anteriorly and scattered, fine setae on remaining part; ventral lobe much narrower and slightly longer than dorsal lobe (lateral view), slightly extended beyond epandrial lamella, digitiform, gently curved downwards on about apical half, with some minute setulae.
</paragraph>
<paragraph id="8BC4F158B731FF9FA71ED00F622AFCC7" blockId="5.[812,1399,315,1404]" box="[850,1041,849,871]" pageId="5" pageNumber="199">
<emphasis id="B90F2D4AB731FF9FA71ED00F6598FCC6" bold="true" box="[850,931,849,870]" pageId="5" pageNumber="199">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED0356245FA8C" pageId="5" pageNumber="199" type="discussion">
<paragraph id="8BC4F158B731FF9FA71ED0356245FA8C" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
REMARKS.
<taxonomicName id="4C7B8ADBB731FF9FA798D0326290FC20" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[980,1195,875,897]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp. nov.">
<emphasis id="B90F2D4AB731FF9FA798D0326290FC20" box="[980,1195,875,897]" italics="true" pageId="5" pageNumber="199">Empis windhoekiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB731FF9FA0FFD03562DFFC20" bold="true" box="[1203,1252,875,896]" pageId="5" pageNumber="199">
<taxonomicNameLabel id="A23C9031B731FF9FA0FFD03562DFFC20" box="[1203,1252,875,896]" pageId="5" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C7B8ADBB731FF9FA760D0D9623EFC3C" authorityName="Smith" authorityYear="1969" box="[812,1029,902,924]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
<emphasis id="B90F2D4AB731FF9FA760D0D96500FC3C" box="[812,827,903,924]" italics="true" pageId="5" pageNumber="199">E</emphasis>
. (
<emphasis id="B90F2D4AB731FF9FA71AD0D965A8FC3C" box="[854,915,903,924]" italics="true" pageId="5" pageNumber="199">Empis</emphasis>
)
<emphasis id="B90F2D4AB731FF9FA7EBD0D9623EFC3C" box="[935,1029,903,924]" italics="true" pageId="5" pageNumber="199">setitarsus</emphasis>
</taxonomicName>
-group [
<bibRefCitation id="EFEA8CA9B731FF9FA015D0D86348FC3B" author="Daugeron C. &amp; Grootaert P." box="[1113,1395,902,923]" pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
;
<bibRefCitation id="EFEA8CA9B731FF9FA760D0FF621CFC16" author="Sinclair B. J. &amp; Daugeron C." box="[812,1063,929,951]" pageId="5" pageNumber="199" pagination="1221 - 1235" refId="ref6424" refString="Sinclair B. J., Daugeron C. 2017. Empididae (empidid dance flies or balloon flies) // A. H. Kirk-Spriggs, B. J. Sinclair (eds.). Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: SANBI Graphics &amp; Editing. P. 1221 - 1235." type="book chapter" year="2017">Sinclair, Daugeron, 2017</bibRefCitation>
]. Within the key to species of this group the new species would run to the couplet 19 joining
<taxonomicName id="4C7B8ADBB731FF9FA12DD0E265D8FC4C" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="B90F2D4AB731FF9FA12DD0E2634BFC71" box="[1377,1392,956,977]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D088655CFC4B" box="[812,871,982,1003]" italics="true" pageId="5" pageNumber="199">vetula</emphasis>
<bibRefCitation id="EFEA8CA9B731FF9FA721D08865D8FC4C" author="Smith K. G. V." box="[877,995,982,1004]" pageId="5" pageNumber="199" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B8ADBB731FF9FA05ED0896567FBA6" authority="Daugeron et Grootaert, 2003" authorityName="Daugeron et Grootaert" authorityYear="2003" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="cinerarius">
<emphasis id="B90F2D4AB731FF9FA05ED089621AFC4C" box="[1042,1057,983,1004]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA061D08962ABFC4C" box="[1069,1168,983,1004]" italics="true" pageId="5" pageNumber="199">cinerarius</emphasis>
<bibRefCitation id="EFEA8CA9B731FF9FA0DBD0886567FBA6" author="Daugeron C. &amp; Grootaert P." pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron et Grootaert, 2003</bibRefCitation>
</taxonomicName>
[
<bibRefCitation id="EFEA8CA9B731FF9FA726D0AF6242FBA6" author="Daugeron C. &amp; Grootaert P." box="[874,1145,1009,1030]" pageId="5" pageNumber="199" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
]. The new species can be readily distinguished from both these species primarily by black setae on the laterotergite. In addition, in the male of
<taxonomicName id="4C7B8ADBB731FF9FA12DD779655CFBF6" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="vetula">
<emphasis id="B90F2D4AB731FF9FA12DD779634BFB9C" box="[1377,1392,1063,1084]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D71F655CFBF6" box="[812,871,1089,1110]" italics="true" pageId="5" pageNumber="199">vetula</emphasis>
</taxonomicName>
the phallus is well-exposed, longer, and uniformly slender [
<bibRefCitation id="EFEA8CA9B731FF9FA733D7056217FBD1" author="Smith K. G. V." box="[895,1068,1115,1137]" pageId="5" pageNumber="199" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969: 175</bibRefCitation>
, fig. 209], whereas in the male of the new species the phallus is short, mostly hidden, thick on about basal 2/3 and slender on apical 1/3. Besides the colour of laterotergal setae,
<taxonomicName id="4C7B8ADBB731FF9FA7B5D7F26293FB60" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[1017,1192,1195,1217]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="windhoekiensis" status="sp.n.">
<emphasis id="B90F2D4AB731FF9FA7B5D7F26233FB61" box="[1017,1032,1196,1217]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA059D7F56293FB60" box="[1045,1192,1195,1216]" italics="true" pageId="5" pageNumber="199">windhoekiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB731FF9FA0E3D7F562DBFB60" bold="true" box="[1199,1248,1195,1216]" pageId="5" pageNumber="199">
<taxonomicNameLabel id="A23C9031B731FF9FA0E3D7F562DBFB60" box="[1199,1248,1195,1216]" pageId="5" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C7B8ADBB731FF9FA12DD7F265B4FB7C" authorityName="Daugeron et Grootaert" authorityYear="2003" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="cinerarius">
<emphasis id="B90F2D4AB731FF9FA12DD7F2634BFB61" box="[1377,1392,1196,1217]" italics="true" pageId="5" pageNumber="199">E</emphasis>
.
<emphasis id="B90F2D4AB731FF9FA760D79965B4FB7C" box="[812,911,1223,1244]" italics="true" pageId="5" pageNumber="199">cinerarius</emphasis>
</taxonomicName>
by shorter proboscis (labrum only slightly longer than head height versus labrum about 2X longer than head height), shorter and thicker antennal stylus as well as by the structure of the male hypopygium.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED66F6278FAC1" pageId="5" pageNumber="199" type="etymology">
<paragraph id="8BC4F158B731FF9FA71ED66F6278FAC1" blockId="5.[812,1399,315,1404]" pageId="5" pageNumber="199">
ETYMOLOGY. The name of the new species refers to the
<typeStatus id="54C04FFAB731FF9FA71DD6126540FAC1" box="[849,891,1356,1377]" pageId="5" pageNumber="199">type</typeStatus>
locality, Windhoek.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B731FF9FA71ED63862DAFADC" box="[850,1249,1382,1404]" pageId="5" pageNumber="199" type="distribution">
<paragraph id="8BC4F158B731FF9FA71ED63862DAFADC" blockId="5.[812,1399,315,1404]" box="[850,1249,1382,1404]" pageId="5" pageNumber="199">
DISTRIBUTION. Afrotropics:
<collectingCountry id="F36CB1C8B731FF9FA0CBD63862E6FADC" box="[1159,1245,1382,1404]" name="Namibia" pageId="5" pageNumber="199">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="C361A2D3B736FF98A77ED0B8634AFC5D" box="[818,1393,997,1021]" pageId="2" pageNumber="196" type="nomenclature">
<paragraph id="8BC4F158B736FF98A77ED0B8634AFC5D" blockId="2.[812,1400,997,2064]" box="[818,1393,997,1021]" pageId="2" pageNumber="196">
<heading id="D08C4634B736FF98A77ED0B8634AFC5D" box="[818,1393,997,1021]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="196" reason="4">
<taxonomicName id="4C7B8ADBB736FF98A77ED0B86309FC5D" ID-CoL="9J6ZK" authority="Kustov et Shamshev" authorityName="Kustov et Shamshev" authorityYear="2022" box="[818,1330,997,1021]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB736FF98A77ED0B8654FFC5D" box="[818,884,998,1021]" italics="true" pageId="2" pageNumber="196">Empis</emphasis>
(
<emphasis id="B90F2D4AB736FF98A7CFD0B865FFFC5D" box="[899,964,998,1021]" italics="true" pageId="2" pageNumber="196">Empis</emphasis>
)
<emphasis id="B90F2D4AB736FF98A798D0BB626AFC5D" box="[980,1105,997,1021]" italics="true" pageId="2" pageNumber="196">namibiensis</emphasis>
Kustov et Shamshev
</taxonomicName>
,
<emphasis id="B90F2D4AB736FF98A170D0BB634AFC5D" bold="true" box="[1340,1393,997,1021]" pageId="2" pageNumber="196">
<taxonomicNameLabel id="A23C9031B736FF98A170D0BB634AFC5D" box="[1340,1393,997,1021]" pageId="2" pageNumber="196" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A053D75C62BFFBBA" box="[1055,1156,1026,1050]" pageId="2" pageNumber="196" type="description">
<paragraph id="8BC4F158B736FF98A053D75C62BFFBBA" blockId="2.[812,1400,997,2064]" box="[1055,1156,1026,1050]" pageId="2" pageNumber="196">
<figureCitation id="1340EDDDB736FF98A053D75C6244FBBA" box="[1055,1151,1026,1050]" captionStart="Figs 46" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 46. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 46. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="2" pageNumber="196">Figs 46</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A71ED776634CFBCA" pageId="2" pageNumber="196" type="materials_examined">
<paragraph id="8BC4F158B736FF98A71ED776634CFBCA" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
<collectionCode id="ED6A699DB736FF98A71ED776623FFB9A" box="[850,1028,1063,1083]" pageId="2" pageNumber="196">TYPE MATERIAL</collectionCode>
.
<materialsCitation id="3B13FB05B736FF98A05DD779634FFBF2" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="2" pageNumber="196" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C04FFAB736FF98A05DD779625AFB9A" box="[1041,1121,1063,1082]" pageId="2" pageNumber="196" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7D3AD1B736FF98A027D77B6244FB9B" box="[1131,1151,1061,1083]" count="1" pageId="2" pageNumber="196" type="male"></specimenCount>
,
<collectingCountry id="F36CB1C8B736FF98A0C2D77962E1FB9A" box="[1166,1242,1063,1082]" name="Namibia" pageId="2" pageNumber="196">Namibia</collectingCountry>
,
<location id="8EA4A783B736FF98A0A4D7796379FB9A" LSID="urn:lsid:plazi:treatment:03D2404EB736FF9EA77ED0B862DAFE51:8EA4A783B736FF98A0A4D7796379FB9A" box="[1256,1346,1063,1082]" country="Namibia" latitude="-22.545" longLatPrecision="75" longitude="17.255" name="Windhoek" pageId="2" pageNumber="196">Windhoek</location>
env.,
<geoCoordinate id="EE4F979FB736FF98A760D7616546FBF2" box="[812,893,1087,1106]" degrees="22.545" direction="south" orientation="latitude" pageId="2" pageNumber="196" precision="55" value="-22.545">22.545°S</geoCoordinate>
<geoCoordinate id="EE4F979FB736FF98A7C8D76165ECFBF2" box="[900,983,1087,1106]" degrees="17.255" direction="east" orientation="longitude" pageId="2" pageNumber="196" precision="55" value="17.255">17.255°E</geoCoordinate>
,
<quantity id="4C835CBDB736FF98A7AFD7616212FBF3" box="[995,1065,1087,1107]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="2" pageNumber="196" unit="m" value="1870.0">
<elevation id="0056166BB736FF98A7AFD7616212FBF3" box="[995,1065,1087,1107]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="2" pageNumber="196" unit="m" value="1870.0">1870 m</elevation>
</quantity>
,
<date id="FFC5D798B736FF98A078D7616293FBF2" box="[1076,1192,1087,1106]" pageId="2" pageNumber="196" value="2021-01-28" valueMax="2021-01-31" valueMin="2021-01-28">
<collectingDate id="EF812E70B736FF98A078D7616293FBF2" box="[1076,1192,1087,1106]" pageId="2" pageNumber="196" value="2021-01-28" valueMax="2021-01-31" valueMin="2021-01-28">2831.i.2021</collectingDate>
</date>
,
<collectorName id="268E948EB736FF98A0F8D71E6323FBF2" box="[1204,1304,1087,1107]" pageId="2" pageNumber="196">N. Vikhrev</collectorName>
(
<collectionCode id="ED6A699DB736FF98A16AD7616355FBF2" box="[1318,1390,1087,1106]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="196" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B13FB05B736FF98A760D7066348FBCA" box="[812,1395,1109,1131]" collectingDate="2021-01-28" collectingDateMax="2021-01-31" collectingDateMin="2021-01-28" collectionCode="ZMMU, ZISP" collectorName="N. Vikhrev" country="Namibia" elevation="1870" latitude="-22.545" location="Windhoek" longLatPrecision="75" longitude="17.255" pageId="2" pageNumber="196" specimenCount="15" specimenCount-female="3" specimenCount-male="12" typeStatus="paratype">
<typeStatus id="54C04FFAB736FF98A760D70665B8FBCB" box="[812,899,1112,1131]" pageId="2" pageNumber="196" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7D3AD1B736FF98A7C0D709658DFBCB" box="[908,950,1109,1131]" count="11" pageId="2" pageNumber="196" type="male">11♂</specimenCount>
<specimenCount id="9D7D3AD1B736FF98A7F7D70965E9FBCB" box="[955,978,1109,1131]" count="3" pageId="2" pageNumber="196" type="female">3♀</specimenCount>
, same data as holotype (
<collectionCode id="ED6A699DB736FF98A0E4D70962D5FBCA" box="[1192,1262,1111,1130]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="196" type="Museum">ZMMU</collectionCode>
,
<specimenCount id="9D7D3AD1B736FF98A0BBD7096327FBCB" box="[1271,1308,1109,1131]" count="1" pageId="2" pageNumber="196" type="male">1 ♂</specimenCount>
<collectionCode id="ED6A699DB736FF98A171D7096356FBCA" box="[1341,1389,1111,1130]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="196">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF98A71ED7316374FAAB" pageId="2" pageNumber="196" type="diagnosis">
<paragraph id="8BC4F158B736FF98A71ED7316374FAAB" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
DIAGNOSIS. A species of
<emphasis id="B90F2D4AB736FF98A015D72E6253FB25" box="[1113,1128,1136,1157]" italics="true" pageId="2" pageNumber="196">E</emphasis>
.
<taxonomicName id="4C7B8ADBB736FF98A03FD73162C2FB25" authorityName="Smith" authorityYear="1969" box="[1139,1273,1135,1157]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
(
<emphasis id="B90F2D4AB736FF98A037D72E62B1FB25" box="[1147,1162,1136,1157]" italics="true" pageId="2" pageNumber="196">E</emphasis>
.)
<emphasis id="B90F2D4AB736FF98A0D1D72E62C2FB25" box="[1181,1273,1136,1157]" italics="true" pageId="2" pageNumber="196">setitarsus</emphasis>
</taxonomicName>
-group; body about
<quantity id="4C835CBDB736FF98A72BD7D46589FB3F" box="[871,946,1162,1184]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="196" unit="mm" value="4.5">4.5 mm</quantity>
; proboscis long, labrum 1.52X as long as head height; laterotergite with pale setae; abdomen densely light grey pruinose, only pale setose; wing with complete veins M 1 and CuA+CuP. Male: wing hyaline. Female: legs with simple setae, wing somewhat broadened and slightly darkened.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B736FF9EA71ED6516488FB61" lastPageId="4" lastPageNumber="198" pageId="2" pageNumber="196" type="description">
<paragraph id="8BC4F158B736FF98A71ED651627CF89B" blockId="2.[812,1400,997,2064]" pageId="2" pageNumber="196">
DESCRIPTION. Body length 4.54.8; wing length
<quantity id="4C835CBDB736FF98A10CD651656FFA9F" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="2" pageNumber="196" unit="mm" value="4.1" valueMax="4.2" valueMin="4.0">44.2 mm</quantity>
.
<emphasis id="B90F2D4AB736FF98A710D67465ABFA9F" bold="true" box="[860,912,1322,1343]" pageId="2" pageNumber="196">Male</emphasis>
(
<figureCitation id="1340EDDDB736FF98A7D0D67465ECFA9F" box="[924,983,1322,1344]" captionStart="Figs 46" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 46. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 46. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="2" pageNumber="196">Fig. 4</figureCitation>
). Head with greyish, dense pruinescence on frons, face (except shiny lower margin), clypeus (except shiny upper margin), ocellar triangle, occiput and postgena; black setose (except noted). Eyes holoptic, with upper ommatidia enlarged. Frons represented by very small, subtriangular space just below ocellar triangle and slightly larger space just above antennae. Face broad, almost parallel-sided, bare. Ocellar triangle with several subequally short, fine setae. Occiput with fine, moderately long postoculars and several similar setae laterally; postgena with numerous long, pale, hair-like setae. Antenna black; scape short, slightly longer than pedicel, both with short setulae; postpedicel elongate-conical, with narrow base and almost straight margins, nearly 3X as long as basal width and 1.51.6X longer than stylus. Proboscis moderately long, with labrum nearly 1.5X as long as head height; labium lacking annulations, bearing scattered, minute setulae; labella moderately long, nearly 2X shorter than labrum; with small, dorsal, desclerotized space on about distal third (visible
<emphasis id="B90F2D4AB736FF98A005D45562BCF880" box="[1097,1159,1803,1824]" italics="true" pageId="2" pageNumber="196">in situ</emphasis>
as notch); palpus black, with black, scattered setulae.
</paragraph>
<paragraph id="8BC4F158B736FF99A71ED461646AFDD7" blockId="2.[812,1400,997,2064]" lastBlockId="3.[189,776,315,897]" lastPageId="3" lastPageNumber="197" pageId="2" pageNumber="196">Thorax black in ground-colour, densely greyish pruinescent; mesoscutum with 4 brownish vittae (dorsal view), lateral vittae (between dorsocentral and supra-alar setae) less distinct. Proepisternum with 67 long, fine, pale setae on lower part and 23 similar setae on upper part. Prosternum bare. Antepronotum with 45 pale setae on each side. Postpronotal lobe with 1 strong, long, black seta and several fine, pale, short setae anteriorly. Mesonotal setae well-differentiated, black; notopleuron with several short, fine, pale setae anteriorly, sometimes pale setulae present just behind postpronotal lobe: acrostichals rather long, biserial, lacking on prescutellar depression; dorsocentrals irregularly 23-serial and similar to acrostichals before suture, uniserial and sparser along prescutellar depression, 2 prescutellar setae longest; 1 fine, moderately long presutural intra-alar, 1 similar presutural supra-alar, 3 long notopleurals, 1 postsutural supra-alar (with 23 additional short, fine setae anteriorly), 1 long and 1 minute postalars, 4 scutellars (apical setae long and cruciate, lateral setae short and fine).Laterotergite with numerous pale setae of different lengths. Anterior and posterior spiracles brownish.</paragraph>
<paragraph id="8BC4F158B737FF99A4AED12262ADFC91" blockId="3.[189,776,315,897]" lastBlockId="3.[812,1399,315,897]" pageId="3" pageNumber="197">Legs long, slender, entirely dark brown, mostly faintly greyish pruinose, coxae densely pruinose, black setose (except noted). Coxae and trochanters mostly with pale fine setae, mid and hind coxae often with some additional dark setae. Fore femur with rows of very short, fine, mostly pale anteroventral and posteroventral setae. Fore tibia with slightly longer setulae posterodorsally and posteriorly; setae of subapical circlet mostly short, fine, 1 anteroventral and 1 posteroventral setae longer. Fore basitarsus slender; fore tarsomeres covered with short setae (except circlet of somewhat longer subapical setae). Mid femur with row of mostly pale posteroventral setae becoming longer toward base of femur (longest setae nearly 1.5X longer than femur basal width). Mid tibia with 45 moderately long dorsal (at most 1.5X longer than tibia width) setae and some numerous, slightly shorter setae ventrally (besides circlet of short subapicals). Mid tarsomeres slender, mostly covered with short setae; mid basitarsus with numerous, short, spine-like setae ventrally. Hind femur with mostly very short anteroventral setae (slightly longer closer to apex); long, pale, fine setae on about basal half posteroventrally and posteriorly. Hind tibia slightly almost uniformly thickened viewed laterally (except extreme base), slightly curved viewed posteriorly; with rows of numerous anterodorsal and posterodorsal setae of different lengths (longest setae at most 1.5X as long as tibia middle width), some rather short setae ventrally. Hind basitarsus slightly, uniformly thickened but not broader than apex of hind tibia; with several moderately setae dorsally and numerous short, spine-like setae ventrally.</paragraph>
<paragraph id="8BC4F158B737FF9EA71ED06867F0FE01" blockId="3.[812,1399,315,897]" lastBlockId="4.[189,776,315,1297]" lastPageId="4" lastPageNumber="198" pageId="3" pageNumber="197">Wing membrane hyaline, veins brownish; longitudinal veins complete, well sclerotised, only CuA+CuP (anal vein) somewhat weakened beyond midway toward wing margin; cell dm short, with elongate apex. Pterostigma brownish. Basal costal seta present, short, fine. Anal lobe well-developed; axillary incision almost right-angled. Squama yellow, pale fringed. Halter yellow.</paragraph>
<caption id="DF04A1D0B737FF99A4AED4EF634DF7B1" ID-DOI="http://doi.org/10.5281/zenodo.13178204" ID-Zenodo-Dep="13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="3" pageNumber="197" startId="3.[226,264,1969,1988]" targetBox="[188,1393,926,1908]" targetPageId="3" targetType="figure">
<paragraph id="8BC4F158B737FF99A4AED4EF6343F87C" blockId="3.[189,1400,1967,2065]" pageId="3" pageNumber="197">
Figs 46.
<taxonomicName id="4C7B8ADBB737FF99A572D4EC640EF864" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[318,565,1969,1988]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB737FF99A572D4EC674EF864" box="[318,373,1970,1988]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
(
<emphasis id="B90F2D4AB737FF99A5C8D4EC6780F864" box="[388,443,1970,1988]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
)
<emphasis id="B90F2D4AB737FF99A586D4EF640EF864" box="[458,565,1969,1988]" italics="true" pageId="3" pageNumber="197">namibiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB737FF99A60ED4EF6454F864" bold="true" box="[578,623,1969,1988]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A23C9031B737FF99A60ED4EF6454F864" box="[578,623,1969,1988]" pageId="3" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus.
</paragraph>
<paragraph id="8BC4F158B737FF99A4AED4B8634DF7B1" blockId="3.[189,1400,1967,2065]" pageId="3" pageNumber="197">
Рис. 46.
<taxonomicName id="4C7B8ADBB737FF99A576D4B8641DF858" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[314,550,2021,2040]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov." subGenus="Empis">
<emphasis id="B90F2D4AB737FF99A576D4B86754F858" box="[314,367,2022,2040]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
(
<emphasis id="B90F2D4AB737FF99A530D4B8678AF858" box="[380,433,2022,2040]" italics="true" pageId="3" pageNumber="197">Empis</emphasis>
)
<emphasis id="B90F2D4AB737FF99A5F3D4BB641DF858" box="[447,550,2021,2040]" italics="true" pageId="3" pageNumber="197">namibiensis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2D4AB737FF99A67DD4BB6467F858" bold="true" box="[561,604,2021,2040]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A23C9031B737FF99A67DD4BB6467F858" box="[561,604,2021,2040]" pageId="3" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус.
</paragraph>
</caption>
<paragraph id="8BC4F158B730FF9EA4AED2F8679BFDE1" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">Abdomen black in ground-colour, uniformly densely light grey pruinose, tergites with narrowly pale posterior margin; covered with numerous pale hair-like setae longer laterally. Segment 8 with separated sclerites; tergite 8 without projections, entire, somewhat concave posteriorly; sternite 8 broad, desclerotized medially.</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED11867D4FB81" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">
Terminalia (
<figureCitation id="1340EDDDB730FF9EA517D118678DFDFC" box="[347,438,582,604]" captionStart="Figs 46" captionStartId="3.[226,264,1969,1988]" captionTargetBox="[188,1393,926,1908]" captionTargetId="figure-500@3.[188,1398,926,1908]" captionTargetPageId="3" captionText="Figs 46. Empis (Empis) namibiensis, sp.n., ♂, lateral view: 4 — habitus; 5 — postabdomen; 6 — hypopygium. Abbreviations: cerc — cercus; cerc d lb — dorsal lobe of cercus; cerc v lb — ventral lobe of cercus; epand — epandrium; hypd — hypandrium; ph — phallus. Рис. 46. Empis (Empis) namibiensis, sp.n., ♂, сбоку: 4 — габитус; 5 — постабдомен; 6 — гипопигий. СокраЩениЯ: cerc — церк; cerc d lb — дорсальнаЯ лопасть церка; cerc v lb — вентральнаЯ лопасть церка; epand — Эпандрий; hypd — гипандрий; ph — фаллус." figureDoi="http://doi.org/10.5281/zenodo.13178204" httpUri="https://zenodo.org/record/13178204/files/figure.png" pageId="4" pageNumber="198">Figs 56</figureCitation>
). Epandrium densely greyish pruinose, black setose, divided into two lamellae (dorsal bridge absent); epandrial lamella subtriangular (lateral view), with long setae along upper margin and apically. Hypandrium with basal part faintly greyish pruinose, bearing several moderately long, fine, pale setae; apical part subshiny, yellowish brown; short, in ventral view rather subrectangular, with slightly concave upper margin. Hypoproct slender, digitiform (dorsal view), bare. Phallus yellowish, mostly hidden, short, evenly curved, mostly thick with subapical part narrowed. Ejaculatory apodeme large, extended far beyond basal curvature of phallus, without lateral wings. Cerci black, subshiny, black setose; separated from each other by distinct membraneous space but almost fused narrowly to anterodorsal part of epandrium; cercus somewhat concave posteriorly (lateral view); dorsal lobe subrectangular, short, with short marginal setae; ventral lobe nearly as long as dorsal lobe but very slender, with 2 long setae.
</paragraph>
<paragraph id="8BC4F158B730FF9EA4AED7786488FB61" blockId="4.[189,776,315,1297]" pageId="4" pageNumber="198">
<emphasis id="B90F2D4AB730FF9EA4AED7786708FB9B" bold="true" box="[226,307,1062,1083]" pageId="4" pageNumber="198">Female.</emphasis>
Similar to male except the following characters. Frons broad, parallel-sided; labrum somewhat longer than in male Acrostichals and dorsocentrals shorter. Legs with shorter, simple setation. Wing broader, slightly darkened. Abdomen densely light grey pruinose, covered with shorter setae. Cercus black, long, with scattered hair-like setulae.
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA4AED7986307FE01" pageId="4" pageNumber="198" type="discussion">
<paragraph id="8BC4F158B730FF9EA4AED7986307FE01" blockId="4.[189,776,315,1297]" lastBlockId="4.[812,1398,315,497]" pageId="4" pageNumber="198">
REMARKS.
<taxonomicName id="4C7B8ADBB730FF9EA529D7996427FB7B" authorityName="Kustov &amp; Shamshev" authorityYear="2022" box="[357,540,1222,1244]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="B90F2D4AB730FF9EA529D7996427FB7B" box="[357,540,1222,1244]" italics="true" pageId="4" pageNumber="198">Empis namibiensis</emphasis>
</taxonomicName>
<emphasis id="B90F2D4AB730FF9EA669D798646DFB7B" bold="true" box="[549,598,1222,1243]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A23C9031B730FF9EA669D798646DFB7B" box="[549,598,1222,1243]" pageId="4" pageNumber="198" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C7B8ADBB730FF9EA6BED7996754FB56" authorityName="Smith" authorityYear="1969" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="setitarsus" subGenus="Empis">
<emphasis id="B90F2D4AB730FF9EA6BED799653AFB7C" box="[754,769,1223,1244]" italics="true" pageId="4" pageNumber="198">E</emphasis>
. (
<emphasis id="B90F2D4AB730FF9EA489D7BF673AFB56" box="[197,257,1249,1270]" italics="true" pageId="4" pageNumber="198">Empis</emphasis>
)
<emphasis id="B90F2D4AB730FF9EA55FD7BF6754FB56" box="[275,367,1249,1270]" italics="true" pageId="4" pageNumber="198">setitarsus</emphasis>
</taxonomicName>
-group [
<bibRefCitation id="EFEA8CA9B730FF9EA5F3D7BF64EBFB56" author="Daugeron C. &amp; Grootaert P." box="[447,720,1249,1270]" pageId="4" pageNumber="198" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
;
<bibRefCitation id="EFEA8CA9B730FF9EA692D7BF67A5FAB0" author="Sinclair B. J. &amp; Daugeron C." pageId="4" pageNumber="198" pagination="1221 - 1235" refId="ref6424" refString="Sinclair B. J., Daugeron C. 2017. Empididae (empidid dance flies or balloon flies) // A. H. Kirk-Spriggs, B. J. Sinclair (eds.). Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: SANBI Graphics &amp; Editing. P. 1221 - 1235." type="book chapter" year="2017">Sinclair, Daugeron, 2017</bibRefCitation>
]. Within the key to species of this group the new species would run to
<taxonomicName id="4C7B8ADBB730FF9EA0CBD262634DFEF1" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" box="[1159,1398,315,337]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="namaqua">
<emphasis id="B90F2D4AB730FF9EA0CBD26262ADFEF1" box="[1159,1174,316,337]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<emphasis id="B90F2D4AB730FF9EA0EED26262C7FEF1" box="[1186,1276,316,337]" italics="true" pageId="4" pageNumber="198">namaqua</emphasis>
<bibRefCitation id="EFEA8CA9B730FF9EA14DD265634DFEF1" author="Smith K. G. V." box="[1281,1398,315,337]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B8ADBB730FF9EA719D2096216FECC" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" box="[853,1069,342,364]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="corana">
<emphasis id="B90F2D4AB730FF9EA719D209655FFECC" box="[853,868,343,364]" italics="true" pageId="4" pageNumber="198">E</emphasis>
.
<emphasis id="B90F2D4AB730FF9EA723D2096588FECC" box="[879,947,343,364]" italics="true" pageId="4" pageNumber="198">corana</emphasis>
<bibRefCitation id="EFEA8CA9B730FF9EA7F4D2086216FECC" author="Smith K. G. V." box="[952,1069,342,364]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
[
<bibRefCitation id="EFEA8CA9B730FF9EA074D2086379FECB" author="Daugeron C. &amp; Grootaert P." box="[1080,1346,342,363]" pageId="4" pageNumber="198" pagination="339 - 360" refId="ref6227" refString="Daugeron C., Grootaert P. 2003. Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus - group // Systematic Entomology. Vol. 28. P. 339 - 360. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00214. x." type="journal article" year="2003">Daugeron, Grootaert, 2003</bibRefCitation>
]. The new species differs from both these species primarily by pale setae on the laterotergite and abdomen [
<bibRefCitation id="EFEA8CA9B730FF9EA0FFD2D56309FE01" author="Smith K. G. V." box="[1203,1330,395,417]" pageId="4" pageNumber="198" pagination="1 - 347" refId="ref6492" refString="Smith K. G. V. 1969. The Empididae of Southern Africa (Diptera) // Annals of the Natal Museum. Vol. 19. P. 1 - 347." type="book chapter" year="1969">Smith, 1969</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED2F86220FE77" pageId="4" pageNumber="198" type="etymology">
<paragraph id="8BC4F158B730FF9EA71ED2F86220FE77" blockId="4.[812,1398,315,497]" pageId="4" pageNumber="198">ETYMOLOGY. The name of the new species refers to the country of its origin.</paragraph>
</subSubSection>
<subSubSection id="C361A2D3B730FF9EA71ED28562DAFE51" box="[850,1249,475,497]" pageId="4" pageNumber="198" type="distribution">
<paragraph id="8BC4F158B730FF9EA71ED28562DAFE51" blockId="4.[812,1398,315,497]" box="[850,1249,475,497]" pageId="4" pageNumber="198">
DISTRIBUTION. Afrotropics:
<collectingCountry id="F36CB1C8B730FF9EA0CBD28562E6FE51" box="[1159,1245,475,497]" name="Namibia" pageId="4" pageNumber="198">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,234 @@
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<subSubSection id="C36129F6BF509952FCC1FCAAFB73D59B" pageId="4" pageNumber="222" type="nomenclature">
<paragraph id="8BC47A7DBF509952FCC1FCAAFA97D579" blockId="4.[812,1399,864,1851]" box="[854,1356,864,888]" pageId="4" pageNumber="222">
<heading id="D08CCD11BF509952FCC1FCAAFA97D579" box="[854,1356,864,888]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="222" reason="8">
<taxonomicName id="4C7B01FEBF509952FCC1FCAAFA97D579" authorityName="Gorochov" authorityYear="2022" box="[854,1356,864,888]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="carinata" status="sp. n." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF509952FCC1FCAAFA97D579" box="[854,1356,864,888]" italics="true" pageId="4" pageNumber="222">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BC47A7DBF509952FC6DFCB7FB73D59B" blockId="4.[812,1399,864,1851]" box="[1018,1192,893,917]" pageId="4" pageNumber="222">
Gorochov,
<emphasis id="B90FA66FBF509952FBE4FCB7FB73D59B" bold="true" box="[1139,1192,893,917]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FBE4FCB7FB73D59B" box="[1139,1192,893,917]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509952FB85FC50FB49D5BC" box="[1042,1170,922,946]" pageId="4" pageNumber="222" type="description">
<paragraph id="8BC47A7DBF509952FB85FC50FB49D5BC" blockId="4.[812,1399,864,1851]" box="[1042,1170,922,946]" pageId="4" pageNumber="222">
<figureCitation id="134066F8BF509952FB85FC50FB56D5BC" box="[1042,1165,922,946]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 2837</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509952FCC5FC75FB92D214" pageId="4" pageNumber="222" type="materials_examined">
<paragraph id="8BC47A7DBF509952FCC5FC75FB92D214" blockId="4.[812,1399,864,1851]" pageId="4" pageNumber="222">
<materialsCitation id="3B137020BF509952FCC5FC75FB92D214" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="4" pageNumber="222" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
MATERIAL EXAMINED.
<typeStatus id="54C0C4DFBF509952FBD4FC75FB48D5DC" box="[1091,1171,959,978]" pageId="4" pageNumber="222" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7DB1F4BF509952FB24FC77FB1CD5DD" box="[1203,1223,957,979]" count="1" pageId="4" pageNumber="222" type="male"></specimenCount>
,
<emphasis id="B90FA66FBF509952FB45FC75FA98D5DC" bold="true" box="[1234,1347,959,978]" pageId="4" pageNumber="222">
<collectingCountry id="F36C3AEDBF509952FB45FC75FA98D5DC" box="[1234,1347,959,978]" name="Madagascar" pageId="4" pageNumber="222">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF509952FADAFC0AFC7DD5E5" country="Madagascar" name="Toamasina" pageId="4" pageNumber="222">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF509952FC23FC12FB87D5E4" box="[948,1116,983,1003]" pageId="4" pageNumber="222">Moramanga Distr.</collectingMunicipality>
,
<location id="8EA42CA6BF509952FBFEFC1DFAACD5E4" LSID="urn:lsid:plazi:treatment:03D2CB6BBF509950FCC1FCAAFDB6D282:8EA42CA6BF509952FBFEFC1DFAACD5E4" box="[1129,1399,983,1003]" country="Madagascar" municipality="Moramanga Distr." name="Analamazaotra Forest Station" pageId="4" pageNumber="222" stateProvince="Toamasina">Analamazaotra Forest Station</location>
near Andasibe Vill., ~
<quantity id="4C83D798BF509952FC78FC25FBF1D20D" box="[1007,1066,1007,1027]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="222" unit="m" value="900.0">
<elevation id="00569D4EBF509952FC78FC25FBF1D20D" box="[1007,1066,1007,1027]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="222" unit="m" value="900.0">900 m</elevation>
</quantity>
, on leaf of bush in forest at night,
<date id="FFC55CBDBF509952FAF6FC25FC4BD214" pageId="4" pageNumber="222" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF509952FAF6FC25FC4BD214" pageId="4" pageNumber="222" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">8 20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF509952FC0EFBCDFBD1D214" box="[921,1034,1031,1050]" pageId="4" pageNumber="222">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF509952FB81FBCDFBE5D214" box="[1046,1086,1031,1050]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="222" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF509950FCC5FBD5FCDCD232" lastPageId="6" lastPageNumber="224" pageId="4" pageNumber="222" type="description">
<paragraph id="8BC47A7DBF509952FCC5FBD5FAACD135" blockId="4.[812,1399,864,1851]" pageId="4" pageNumber="222">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF509952FBACFBD5FB4ED23B" box="[1083,1173,1055,1077]" pageId="4" pageNumber="222" type="holotype">holotype</typeStatus>
). Body coloration light greyish with following marks: eye greyish with three longitudinal darkish stripes on upper half and one yellowish stripe between two lower of these darkish stripes as well as numerous dense and arcuate darkish lines running along anterior and ventral edges of eye; epicranial dorsum also darkish (almost greyish brown) with numerous lighter and less numerous darker dots; genae with a few oblique darkish stripes; mouthparts with almost rose three distal segments of each maxillary palpus and a pair of small brown spots in dorsolateral corners of clypeus; antenna with dark ventral and lateral longitudinal stripes on scape, with one dark dot and barely darkish marks around it on dorsal surface of scape, and with rather numerous small darkish and whitish spots on flagellum (
<figureCitation id="134066F8BF509952FCA4FA5FFC75D3A4" box="[819,942,1429,1451]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 28, 31</figureCitation>
); pronotum with brown median quadrate on anterior half of disc, numerous light brown to greyish brown dots on rest parts of disc, and sparser brown dots on lateral lobes (
<figureCitation id="134066F8BF509952FCFCFA2FFC01D3F4" box="[875,986,1509,1531]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 31, 33</figureCitation>
); legs with rather numerous and small light brown to greyish brown dots and small spots on all femora and tibiae (but hind femur lighter and with these dots less distinct, except for 45 dark dots distinctly visible along median line of outer femoral surface;
<figureCitation id="134066F8BF509952FB33F985FACDD06A" box="[1188,1302,1615,1637]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 2933</figureCitation>
); tegmina yellowish grey with light brown basal area (this area having whitish reticular pattern), semitransparent membranes, whitish some areas and all crossveins in medial and humeral regions as well as in lateral field, brown to brownish rest venation and numerous small marks on almost all membranes, and brown to light brown larger areas on membranes of proximal part of lateral field (
<figureCitation id="134066F8BF509952FBF2F8C0FB0DD111" box="[1125,1238,1802,1824]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="4" pageNumber="222">Figs 3233</figureCitation>
); visible parts of hind wings barely darkened but with lighter venation (Fig.
</paragraph>
<paragraph id="8BC47A7DBF509952FF2CF89FFF12D164" blockId="4.[187,1398,1877,2064]" box="[187,201,1877,1898]" pageId="4" pageNumber="222">&lt;</paragraph>
<paragraph id="8BC47A7DBF509952FF75F8A7FD12DE1E" blockId="4.[187,1398,1877,2064]" pageId="4" pageNumber="222">
Рис. 1027. Cтруктуры брюШка, самец: 1015 —
<taxonomicName id="4C7B01FEBF509952FD3AF8A7FC7BD18E" authorityName="Gorochov" authorityYear="2022" box="[685,928,1901,1920]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="longitubulosa" status="sp. nov.">
<emphasis id="B90FA66FBF509952FD3AF8A7FC7BD18E" box="[685,928,1901,1920]" italics="true" pageId="4" pageNumber="222">Asymmetrella longitubulosa</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF509952FC31F8A7FC0AD18E" bold="true" box="[934,977,1901,1920]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FC31F8A7FC0AD18E" box="[934,977,1901,1920]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 16 —
<taxonomicName id="4C7B01FEBF509952FB81F8A7FF3CD196" authority="Gor." authorityName="Gor." class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="bella" subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FB81F8A7FAADD18E" box="[1046,1398,1901,1920]" italics="true" pageId="4" pageNumber="222">Zvenellomorpha (Zvenellomorpha) bella</emphasis>
Gor.
</taxonomicName>
; 1718 —
<taxonomicName id="4C7B01FEBF509952FED0F84CFE3AD196" authority="Gor." authorityName="Gor." box="[327,481,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="recta" subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FED0F84CFE6BD196" box="[327,432,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.) recta</emphasis>
Gor.
</taxonomicName>
; 1923 —
<taxonomicName id="4C7B01FEBF509952FDD6F84CFD00D196" authorityName="Gorochov" authorityYear="2022" box="[577,731,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="intermedia" status="sp.n." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF509952FDD6F84CFD00D196" box="[577,731,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF509952FD75F84FFCD6D196" bold="true" box="[738,781,1925,1944]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="A23C1B14BF509952FD75F84FFCD6D196" box="[738,781,1925,1944]" pageId="4" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 24 —
<emphasis id="B90FA66FBF509952FCC7F84FFCBFD196" box="[848,868,1925,1944]" italics="true" pageId="4" pageNumber="222">A.</emphasis>
sp.; 25 —
<emphasis id="B90FA66FBF509952FC52F84CFC27D196" box="[965,1020,1925,1944]" italics="true" pageId="4" pageNumber="222">Z. (Z.)</emphasis>
sp.; 26 —
<emphasis id="B90FA66FBF509952FBC9F84CFAC1D196" box="[1118,1306,1925,1944]" italics="true" pageId="4" pageNumber="222">
Z. (
<taxonomicName id="4C7B01FEBF509952FBECF84FFACFD196" authorityName="Gorochov" authorityYear="2022" box="[1147,1300,1925,1944]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Azvenellomorpha">Azvenellomorpha</taxonomicName>
)
</emphasis>
sp.; 27 —
<taxonomicName id="4C7B01FEBF509952FF2BF857FEE7D1BE" box="[188,316,1949,1968]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90FA66FBF509952FF2BF857FEC1D1BE" box="[188,282,1949,1968]" italics="true" pageId="4" pageNumber="222">Ultratrella</emphasis>
sp.
</taxonomicName>
; 10 — верШина брюШка, сверху; 1113, 1617, 1921 — гениталии (11, 20 — сверху; 12, 21 — сниЗу; 13, 1617, 19 — сбоку); 14 — сперматофор сбоку, реконструкциЯ; 15 — формула с рахисом, Эндопарамером и Эктопарамером, сниЗу; 18, 22 — формула, сниЗу; 23 — дистальнаЯ часть гениталий, сбоку; 2427 — схема сагиттального сечениЯ формулы [коричневый цвет], её склеротиЗованной передней ленты [Зелёный цвет] и ближайШего участка сперматофорной трубы [красный цвет]; 10, 23 — паратип; 1122 — голотипы [16, 17 — по:
<bibRefCitation id="EFEA078CBF509952FE6FF837FD53DE1E" author="Gorochov A. V." box="[504,648,2045,2064]" pageId="4" pageNumber="222" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
,
<bibRefCitation id="EFEA078CBF509952FD05F837FD19DE1E" author="Gorochov A. V." box="[658,706,2045,2064]" pageId="4" pageNumber="222" pagination="33 - 46" refId="ref8770" refString="Gorochov A. V. 2006. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 5: new Indo-Malayan and Madagascan Podoscirtini // Zoosystematica rossica. Vol. 15. No. 1. P. 33 - 46." type="journal article" year="2006">2006</bibRefCitation>
.]
</paragraph>
<caption id="DF042AF5BF519953FF75F8A3FE96DE1F" ID-DOI="http://doi.org/10.5281/zenodo.13178132" ID-Zenodo-Dep="13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="5" pageNumber="223" startId="5.[226,263,1897,1916]" targetBox="[195,1390,322,1834]" targetPageId="5" targetType="figure">
<paragraph id="8BC47A7DBF519953FF75F8A3FB06D1A2" blockId="5.[188,1399,1897,2065]" pageId="5" pageNumber="223">
Figs 2837.
<taxonomicName id="4C7B01FEBF519953FEDAF8A3FD07D172" authorityName="Gorochov" authorityYear="2022" box="[333,732,1897,1916]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="223" phylum="Arthropoda" rank="species" species="carinata" status="sp. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF519953FEDAF8A3FD07D172" box="[333,732,1897,1916]" italics="true" pageId="5" pageNumber="223">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF519953FD76F8A3FCD7D172" bold="true" box="[737,780,1897,1916]" pageId="5" pageNumber="223">
<taxonomicNameLabel id="A23C1B14BF519953FD76F8A3FCD7D172" box="[737,780,1897,1916]" pageId="5" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction.
</paragraph>
<paragraph id="8BC47A7DBF519953FF75F87CFE96DE1F" blockId="5.[188,1399,1897,2065]" pageId="5" pageNumber="223">
Рис. 2837.
<taxonomicName id="4C7B01FEBF519953FECDF87FFD23D1C6" authorityName="Gorochov" authorityYear="2022" box="[346,760,1973,1992]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="223" phylum="Arthropoda" rank="species" species="carinata" status="sp. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF519953FECDF87FFD23D1C6" box="[346,760,1973,1992]" italics="true" pageId="5" pageNumber="223">Zvenellomorpha (Euzvenellomorpha) carinata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF519953FC94F87FFCF5D1C6" bold="true" box="[771,814,1973,1992]" pageId="5" pageNumber="223">
<taxonomicNameLabel id="A23C1B14BF519953FC94F87FFCF5D1C6" box="[771,814,1973,1992]" pageId="5" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF529950FF2AFEF1FEE9D595" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
32); venter of body with wide longitudinal greyish brown band on abdominal sternites, a pair of barely darkish areas on proximal half of genital plate, and numerous very small darkish dots on cerci (
<figureCitation id="134066F8BF529950FE1AFE41FE08D7AE" box="[397,467,395,417]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Fig. 33</figureCitation>
). Eyes slightly higher than long; rostrum roundly angular in profile but with low and thin transverse keel at apex; maxillary palpi approximately as in
<taxonomicName id="4C7B01FEBF529950FF2AFE11FE99D7FE" authorityName="Gorochov" authorityYear="2022" box="[189,322,475,496]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FF2AFE11FE99D7FE" box="[189,322,475,496]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FEDEFE11FE4BD7FE" bold="true" box="[329,400,475,496]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FEDEFE11FE4BD7FE" box="[329,400,475,496]" pageId="6" pageNumber="224" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
; pronotum also similar to that of this genus but more strongly narrowing to head and with lateral keels slightly shorter and more sinuate in profile (
<figureCitation id="134066F8BF529950FD02FDDBFF03D44E" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 28, 31, 33</figureCitation>
); tegmina significantly protruding beyond abdominal apex and hind femora, with venation as in
<figureCitation id="134066F8BF529950FDB4FD8CFD49D455" box="[547,658,582,604]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 3233</figureCitation>
; hind wings also significantly protruding beyond tegmina (
<figureCitation id="134066F8BF529950FD14FDABFD17D478" box="[643,716,609,631]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Fig. 32</figureCitation>
); fore and middle legs rather stout, with a pair of dorsal longitudinal keels on fore tibia (
<figureCitation id="134066F8BF529950FE17FD5CFE2BD4A5" box="[384,496,662,684]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 3133</figureCitation>
) having tympana as in
<figureCitation id="134066F8BF529950FD4AFD5CFF22D4C8" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 2930</figureCitation>
(inner tympanum somewhat larger than outer one), with two keels on middle tibia (along dorsal and ventral tibial edges;
<figureCitation id="134066F8BF529950FE95FD2CFEAFD4F5" box="[258,372,742,764]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 3233</figureCitation>
); hind leg more elongate and slender but also with more or less distinct dorsal keel on femur and a pair of dorsal longitudinal keels on tibia; spines and denticles of this tibia more or less similar to those of
<taxonomicName id="4C7B01FEBF529950FDC7FCFCFD0ED545" authorityName="Gorochov" authorityYear="2022" box="[592,725,822,843]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FDC7FCFCFD0ED545" box="[592,725,822,843]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FD4BFCFCFF0BD568" bold="true" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FD4BFCFCFF0BD568" pageId="6" pageNumber="224" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
; genital plate elongately triangular and with almost spine-like but soft apical lobule; genitalia and spermatophore as in
<figureCitation id="134066F8BF529950FF2AFC4CFEF5D595" box="[189,302,902,924]" captionStart="Figs 2837" captionStartId="5.[226,263,1897,1916]" captionTargetBox="[195,1390,322,1834]" captionTargetId="figure-156@5.[185,1395,314,1847]" captionTargetPageId="5" captionText="Figs 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., male: 28 — head in front; 2930 — outer (29) and inner (30) tympana; 31 — head with pronotum and fore legs from above; 32 — wings with middle and partly hind legs from above; 33 — body from side; 34 36 — genitalia (34 — from side; 35 — from above; 36 — from below); 37 — spermatophore from side, reconstruction. Рис. 2837. Zvenellomorpha (Euzvenellomorpha) carinata sp.n., самец: 28 — голова, спереди; 2930 — наружный (29) и внутренний (30) тимпанумы; 31 — голова с переднеспинкой и передними ногами, сверху; 32 — крыльЯ со средними и частично Задними ногами, сверху; 33 — тело, сбоку; 3436 — гениталии (34 — сбоку; 35 — сверху; 36 — сниЗу); 37 — сперматофор, сбоку, реконструкциЯ." figureDoi="http://doi.org/10.5281/zenodo.13178132" httpUri="https://zenodo.org/record/13178132/files/figure.png" pageId="6" pageNumber="224">Figs 3437</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC6BFE49D5B9" blockId="6.[189,775,315,1164]" box="[226,402,929,951]" pageId="6" pageNumber="224">Female unknown.</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC71FD8AD5E5" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">Length in mm. Body 20.3; body with wings 29.5; pronotum 3.6; tegmina 21.5; hind femora 13.4.</paragraph>
<paragraph id="8BC47A7DBF529950FF75FC3BFCDCD232" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
COMPARISON. The new species differs from all the other congeners in the same characters as the subgenus
<taxonomicName id="4C7B01FEBF529950FF2AFBECFEA9D235" authorityName="Gorochov" authorityYear="2022" box="[189,370,1062,1083]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF529950FF2AFBECFEA9D235" box="[189,370,1062,1083]" italics="true" pageId="6" pageNumber="224">Euzvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FEE1FBECFE03D235" bold="true" box="[374,472,1062,1083]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FEE1FBECFE03D235" box="[374,472,1062,1083]" pageId="6" pageNumber="224" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
(see the subgeneric key above).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FB8BFDB6D282" pageId="6" pageNumber="224" type="etymology">
<paragraph id="8BC47A7DBF529950FF75FB8BFDB6D282" blockId="6.[189,775,315,1164]" pageId="6" pageNumber="224">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF529950FF2AFB91FECDD27F" authorityName="Gorochov" authorityYear="2022" box="[189,278,1115,1137]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="carinata">“carinata</taxonomicName>
” (carinate, with keels) due to the presence of characteristic keels on the lateral pronotal lobes.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,266 @@
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<mods:title id="FFCDFC92494B36B19040C16856A47B24">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<subSubSection id="C36129F6BF529950FF71FB6AFDE2D2DB" pageId="6" pageNumber="224" type="nomenclature">
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<taxonomicName id="4C7B01FEBF529950FF71FB6AFD06D2B6" authorityName="Gorochov" authorityYear="2022" box="[230,733,1184,1208]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="intermedia" status="sp. n." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF529950FF71FB6AFD06D2B6" box="[230,733,1184,1208]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC47A7DBF529950FE1DFB77FDE2D2DB" blockId="6.[189,775,1184,2056]" box="[394,569,1213,1237]" pageId="6" pageNumber="224">
Gorochov,
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<taxonomicNameLabel id="A23C1B14BF529950FD94FB77FDE2D2DB" box="[515,569,1213,1237]" pageId="6" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FEEEFB10FD90D2FC" box="[377,587,1242,1266]" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FEEEFB10FD90D2FC" blockId="6.[189,775,1184,2056]" box="[377,587,1242,1266]" pageId="6" pageNumber="224">
<figureCitation id="134066F8BF529950FEEEFB10FE2ED2FC" box="[377,501,1242,1266]" captionStart="Figs 1027" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 1027. Structures of abdomen, male: 1015 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 1718 — Z. (Z.) recta Gor.; 1923 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 1113, 1617, 1921 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 2427 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 1122 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Figs 1923</figureCitation>
,
<figureCitation id="134066F8BF529950FD96FB10FD9DD2FC" box="[513,582,1242,1266]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">3846</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FB35FE70D3AC" pageId="6" pageNumber="224" type="materials_examined">
<paragraph id="8BC47A7DBF529950FF75FB35FE70D3AC" blockId="6.[189,775,1184,2056]" pageId="6" pageNumber="224">
MATERIAL EXAMINED.
<materialsCitation id="3B137020BF529950FE44FB35FEB3D354" collectingDate="2013-02-22" collectingDateMax="2013-03-11" collectingDateMin="2013-02-22" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="1000" location="Torotorofotsy Forest Reserve" municipality="Moramanga Distr." pageId="6" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54C0C4DFBF529950FE44FB35FDF8D31C" box="[467,547,1279,1298]" pageId="6" pageNumber="224" type="holotype">Holotype</typeStatus>
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,
<emphasis id="B90FA66FBF529950FDF5FB35FD08D31C" bold="true" box="[610,723,1279,1298]" pageId="6" pageNumber="224">
<collectingCountry id="F36C3AEDBF529950FDF5FB35FD08D31C" box="[610,723,1279,1298]" name="Madagascar" pageId="6" pageNumber="224">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF529950FD49FACAFEE8D325" country="Madagascar" name="Toamasina" pageId="6" pageNumber="224">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF529950FEA9FAD2FE39D324" box="[318,482,1303,1323]" pageId="6" pageNumber="224">Moramanga Distr.</collectingMunicipality>
, ~
<locationDeviation id="9A76635FBF529950FE6FFADDFCDFD324" box="[504,772,1303,1323]" location="Andasibe Vill." pageId="6" pageNumber="224">
<quantity id="4C83D798BF529950FE6FFADDFDEAD324" box="[504,561,1303,1322]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="km" value="10.0">10 km</quantity>
NW of Andasibe Vill.
</locationDeviation>
,
<location id="8EA42CA6BF529950FF2AFAE5FE64D34D" LSID="urn:lsid:plazi:treatment:03D2CB6BBF529950FF71FB6AFBFBD102:8EA42CA6BF529950FF2AFAE5FE64D34D" box="[189,447,1327,1347]" country="Madagascar" municipality="Moramanga Distr." name="Torotorofotsy Forest Reserve" pageId="6" pageNumber="224" stateProvince="Toamasina">Torotorofotsy Forest Reserve</location>
, ~
<quantity id="4C83D798BF529950FE44FAE5FDC2D34D" box="[467,537,1327,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="m" value="1000.0">
<elevation id="00569D4EBF529950FE44FAE5FDC2D34D" box="[467,537,1327,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FDFBFAE5FCD8D34C" box="[620,771,1327,1346]" pageId="6" pageNumber="224" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">
<collectingDate id="EF81A555BF529950FDFBFAE5FCD8D34C" box="[620,771,1327,1346]" pageId="6" pageNumber="224" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">22.II11.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FF2AFA8DFEF6D354" box="[189,301,1351,1370]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FEADFA8DFEB9D354" box="[314,354,1351,1370]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B137020BF529950FEE6FA82FF2AD37C" collectingDate="2013-02-22" collectingDateMax="2013-03-11" collectingDateMin="2013-02-22" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="1000" location="Torotorofotsy Forest Reserve" municipality="Moramanga Distr." pageId="6" pageNumber="224" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="54C0C4DFBF529950FEE6FA82FE13D355" box="[369,456,1352,1371]" pageId="6" pageNumber="224" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7DB1F4BF529950FE46FA8DFE2ED355" box="[465,501,1349,1371]" count="1" pageId="6" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7DB1F4BF529950FD96FA8DFDC6D355" box="[513,541,1349,1371]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same data as for holotype (
<collectionCode id="ED6AE2B8BF529950FF54FA95FF30D37C" box="[195,235,1375,1394]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B137020BF529950FF6BFA95FE7DD3AC" collectingDate="2013-03-01" collectingDateMax="2013-03-07" collectingDateMin="2013-03-01" collectionCode="ZIN" collectorName="A. Gorochov &amp; L. Anisyutkin" country="Madagascar" elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="2" specimenCount-female="2" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="9D7DB1F4BF529950FF6BFA95FEC0D37D" box="[252,283,1373,1395]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same province and district,
<collectingMunicipality id="6BA0E007BF529950FD8AFA95FF04D384" pageId="6" pageNumber="224">Analamazaotra Forest Station</collectingMunicipality>
near
<location id="8EA42CA6BF529950FE84FABDFE49D384" LSID="urn:lsid:plazi:treatment:03D2CB6BBF529950FF71FB6AFBFBD102:8EA42CA6BF529950FE84FABDFE49D384" box="[275,402,1399,1418]" country="Madagascar" municipality="Analamazaotra Forest Station" name="Andasibe Vill" pageId="6" pageNumber="224" stateProvince="Toamasina">Andasibe Vill</location>
., ~
<quantity id="4C83D798BF529950FE3CFABDFE3CD385" box="[427,487,1399,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">
<elevation id="00569D4EBF529950FE3CFABDFE3CD385" box="[427,487,1399,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FDAAFABDFD70D384" box="[573,683,1399,1418]" pageId="6" pageNumber="224" value="2013-03-01" valueMax="2013-03-07" valueMin="2013-03-01">
<collectingDate id="EF81A555BF529950FDAAFABDFD70D384" box="[573,683,1399,1418]" pageId="6" pageNumber="224" value="2013-03-01" valueMax="2013-03-07" valueMin="2013-03-01">17.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FD21FABDFF32D3AC" pageId="6" pageNumber="224">A. Gorochov</collectorName>
,
<collectorName id="268E1FABBF529950FF65FA5AFEB0D3AC" box="[242,363,1423,1443]" pageId="6" pageNumber="224">L. Anisyutkin</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FEEFFA45FE7AD3AC" box="[376,417,1423,1442]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FF75FA6DFACCD0AF" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FF75FA6DFA93D595" blockId="6.[189,775,1184,2056]" lastBlockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF529950FE44FA6DFDF6D3B3" box="[467,557,1447,1469]" pageId="6" pageNumber="224" type="holotype">holotype</typeStatus>
). General appearance similar to that of two other species of this subgenus but somewhat intermediate between them. Body coloration yellowish with greyish tinge and following marks: upper half of epicranium, pronotum and dorsal tegminal field light brown with light greyish eyes, a pair of barely distinct yellowish interrupted stripes along both lateral edges of pronotal disc, brown M, yellowish stripe between M and Cu in humeral region of tegmen, light greyish brown diagonal vein, and almost semitransparent membranes of mirror and of some nearest cells (
<figureCitation id="134066F8BF529950FED5F978FE6FD0C9" box="[322,436,1714,1736]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 3840</figureCitation>
); antenna with brown basal part of scape as well as middle and dorsal portions of flagellum, dark brown rest of scape and proximal portion of flagellum (
<figureCitation id="134066F8BF529950FD4BF92DFF27D119" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 3839</figureCitation>
); lateral tegminal field yellowish with light brown venation and semitransparent membranes between veins and veinlets (
<figureCitation id="134066F8BF529950FE80F8FDFEBAD142" box="[279,353,1847,1869]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
). Head with oval (longitudinal) but poorly distinct lateral ocelli, small round and also poorly distinct median ocellus, and large barely elongate eyes; rostrum roundly angular in profile and rather narrow (scape approximately 1.7 times as wide as space between antennal cavities); maxillary palpi more or less similar to those of
<taxonomicName id="4C7B01FEBF529950FD15F877FCDCD1DC" authorityName="Gorochov" authorityYear="2022" box="[642,775,1981,2002]" class="Ostracoda" family="Primitiopsidae" genus="Asymmetrella" kingdom="Animalia" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FA66FBF529950FD15F877FCDCD1DC" box="[642,775,1981,2002]" italics="true" pageId="6" pageNumber="224">Asymmetrella</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF529950FF2AF81DFF21D1E2" bold="true" box="[189,250,2007,2028]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FF2AF81DFF21D1E2" box="[189,250,2007,2028]" pageId="6" pageNumber="224" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
; pronotum slightly narrowing to head, insignificantly longer than wide, and almost not depressed dorsoventrally (but lateral lobes moderately low;
<figureCitation id="134066F8BF529950FBE4FEF1FB62D75E" box="[1139,1209,315,337]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
); legs without keels on femora and tibiae, with inner tympanum oval (almost as wide as half of fore tibia width and about 1.2 times as long as latter width; as in
<figureCitation id="134066F8BF529950FC72FE41FBF4D7AE" box="[997,1071,395,417]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 42</figureCitation>
), and outer tympanum similar in shape but slightly smaller;
<figureCitation id="134066F8BF529950FBAEFE6CFB59D7B5" box="[1081,1154,422,444]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 39</figureCitation>
); tegmina distinctly protruding beyond apices of hind femora, with mirror in dorsal field almost normal (less irregular than in
<taxonomicName id="4C7B01FEBF529950FB50FE16FACBD7FE" box="[1223,1296,475,497]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FB50FE16FACBD7FE" box="[1223,1296,475,497]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
), dividing vein in this mirror more curved than in
<taxonomicName id="4C7B01FEBF529950FB33FE3DFB34D402" box="[1188,1263,503,524]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB33FE3DFB34D402" box="[1188,1263,503,524]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
(
<figureCitation id="134066F8BF529950FB6CFE3CFA98D405" box="[1275,1347,502,524]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 40</figureCitation>
), and Sc having 910 branches; hind wings strongly protruding beyond tegminal apices; genital plate triangular in proximal half but with narrow, long and soft posteromedian lobule in distal half (this lobule narrowly rounded at apex). Genitalia very similar to those of
<taxonomicName id="4C7B01FEBF529950FBA3FDB6FB5ED49E" box="[1076,1157,635,657]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FBA3FDB6FB5ED49E" box="[1076,1157,635,657]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B01FEBF529950FB56FDB6FAC8D49F" box="[1217,1299,636,657]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB56FDB6FAC8D49F" box="[1217,1299,636,657]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
but with following diferences: epiphallus somewhat lower than in these species; dorsoapical epiphallic process with slightly longer both (proximal and distal) spines than in
<taxonomicName id="4C7B01FEBF529950FA99FD06FA82D4EE" box="[1294,1369,715,737]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FA99FD06FA82D4EE" box="[1294,1369,715,737]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
as well as with distinctly deeper apical notch than in
<taxonomicName id="4C7B01FEBF529950FAB5FD2DFAABD4F2" box="[1314,1392,743,764]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FAB5FD2DFAABD4F2" box="[1314,1392,743,764]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
; ventroapical epiphallic lobule slightly shorter and less arcuate (almost S-shaped) than in
<taxonomicName id="4C7B01FEBF529950FBD1FCD6FB55D53E" box="[1094,1166,795,817]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FBD1FCD6FB55D53E" box="[1094,1166,795,817]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
as well as apically more widened and hooked than in
<taxonomicName id="4C7B01FEBF529950FBD3FCFDFB54D542" box="[1092,1167,823,844]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FBD3FCFDFB54D542" box="[1092,1167,823,844]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
; ectoparameres slightly protruding beyond epiphallus (in
<taxonomicName id="4C7B01FEBF529950FB1CFC9BFB06D568" box="[1163,1245,849,870]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B90FA66FBF529950FB1CFC9BFB06D568" box="[1163,1245,849,870]" italics="true" pageId="6" pageNumber="224">Z. recta</emphasis>
</taxonomicName>
, they strongly protruding beyond epiphallus, and in
<taxonomicName id="4C7B01FEBF529950FB04FCA6FB00D58E" box="[1171,1243,875,897]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B90FA66FBF529950FB04FCA6FB00D58E" box="[1171,1243,875,897]" italics="true" pageId="6" pageNumber="224">Z. bella</emphasis>
</taxonomicName>
, they not reaching epiphallic apices; compare
<figureCitation id="134066F8BF529950FBC9FC4CFAE7D595" box="[1118,1340,902,924]" captionStart="Figs 1027" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 1027. Structures of abdomen, male: 1015 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 1718 — Z. (Z.) recta Gor.; 1923 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 1113, 1617, 1921 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 2427 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 1122 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Figs 1618 and 1922</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FC6BFB96D258" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Variations. Second male with epicranial dorsum between eyes laterally darker (greyish brown), pronotal disc having brown median band, dividing vein of tegminal mirror clearly S-shaped (
<figureCitation id="134066F8BF529950FC02FC3BFBD2D208" box="[917,1033,1009,1031]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 4142</figureCitation>
), and genitalia having barely longer proximal spine of dorsoapical epiphallic process as well as slightly narrower (lower) distal part of ventroapical epiphallic lobule in profile (
<figureCitation id="134066F8BF529950FC6FFB8BFB9AD258" box="[1016,1089,1089,1111]" captionStart="Figs 1027" captionStartId="3.[226,263,1918,1937]" captionTargetBox="[190,1399,314,1898]" captionTargetId="figure-10@3.[189,1399,314,1902]" captionTargetPageId="3" captionText="Figs 1027. Structures of abdomen, male: 1015 — Asymmetrella longitubulosa sp.n.; 16 — Zvenellomorpha (Zvenellomorpha) bella Gor.; 1718 — Z. (Z.) recta Gor.; 1923 — Z. (Z.) intermedia sp.n.; 24 — A. sp.; 25 — Z. (Z.) sp.; 26 — Z. (Azvenellomorpha) sp.; 27 — Ultratrella sp.; 10 — abdominal apex from above; 1113, 1617, 1921 — genitalia (11, 20 — from above; 12, 21 — from below; 13, 16 17, 19 — from side); 14 — spermatophore from side, reconstruction; 15 — formula with rachis, endoparamere and ectoparamere from below; 18, 22 — formula from below; 23 — distal part of genitalia from side; 2427 — scheme of sagittal section of formula [brown], of its sclerotized anterior ribbon [green] and of nearest portion of spermatophore tube [red]; 10, 23 — paratype; 1122 — holotypes [16, 17 — after Gorochov, 2004, 2006.]" figureDoi="http://doi.org/10.5281/zenodo.13178128" httpUri="https://zenodo.org/record/13178128/files/figure.png" pageId="6" pageNumber="224">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FB91FB9FD3E8" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Female. Coloration and structure of body as in male, but scape and proximal portion of antennal flagellum somewhat lighter (more or less brown), pronotal disc and dorsal tegminal field sometimes almost yellowish with greyish tinge, posterior part of this disc sometimes completely brown (
<figureCitation id="134066F8BF529950FAD8FB0CFC9CD2F8" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 43</figureCitation>
), tegminal R brown or greyish brown, median ocellus very small and round or obliterate, tegmina with 810 straight but somewhat obliquely situated longitudinal veins and moderately sparse crossveins in dorsal field as well as with 79 branches of Sc and almost without distinct crossveins in lateral field, and genital plate barely elongate as well as distally narrowing to roundly and rather widely bilobate apex (posteromedian notch of this apex short and almost angular;
<figureCitation id="134066F8BF529950FCBBFA7CFCA8D3C5" box="[812,883,1462,1484]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Fig. 44</figureCitation>
). Ovipositor normally developed, with distinctly drilling apical part (
<figureCitation id="134066F8BF529950FC50FA1BFBE1D3E8" box="[967,1082,1488,1511]" captionStart="Figs 3846" captionStartId="7.[226,263,1897,1916]" captionTargetBox="[194,1390,320,1872]" captionTargetId="figure-154@7.[188,1398,314,1876]" captionTargetPageId="7" captionText="Figs 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female. Рис. 3846. Zvenellomorpha (Zvenellomorpha) intermedia sp.n.: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="6" pageNumber="224">Figs 4546</figureCitation>
).
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5FA21FBE9D05F" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
Length in mm. Body:
<specimenCount id="9D7DB1F4BF529950FBA4FA22FB92D00E" box="[1075,1097,1512,1536]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
1314.5,
<specimenCount id="9D7DB1F4BF529950FB27FA22FB66D00E" box="[1200,1213,1512,1536]" count="1" pageId="6" pageNumber="224" type="female"></specimenCount>
1114; body with wings:
<specimenCount id="9D7DB1F4BF529950FCE4F9C9FC52D015" box="[883,905,1539,1563]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
2022,
<specimenCount id="9D7DB1F4BF529950FC4EF9C9FC3DD015" box="[985,998,1539,1563]" count="1" pageId="6" pageNumber="224" type="female"></specimenCount>
1821; pronotum:
<specimenCount id="9D7DB1F4BF529950FB36F9C9FB6CD015" box="[1185,1207,1539,1563]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
2.12.3,
<specimenCount id="9D7DB1F4BF529950FA84F9C9FAFBD015" box="[1299,1312,1539,1563]" count="1" pageId="6" pageNumber="224" type="female"></specimenCount>
2.42.8; tegmina:
<specimenCount id="9D7DB1F4BF529950FC10F9D4FC46D038" box="[903,925,1566,1590]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
1415.5;
<specimenCount id="9D7DB1F4BF529950FC68F9D4FBD7D038" box="[1023,1036,1566,1590]" count="1" pageId="6" pageNumber="224" type="female"></specimenCount>
1315; hind femora:
<specimenCount id="9D7DB1F4BF529950FB48F9D4FB2ED038" box="[1247,1269,1566,1590]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
10.811.5,
<specimenCount id="9D7DB1F4BF529950FAFEF9D4FAADD038" box="[1385,1398,1566,1590]" count="1" pageId="6" pageNumber="224" type="female"></specimenCount>
1011.5; ovipositor 78.5.
</paragraph>
<paragraph id="8BC47A7DBF529950FCC5F99CFACCD0AF" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
COMPARISON. The new species differs from the two other species of the subgenus
<taxonomicName id="4C7B01FEBF529950FBFDF9BBFAD3D088" authorityName="Gorochov" authorityYear="2022" box="[1130,1288,1649,1670]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF529950FBFDF9BBFAD3D088" box="[1130,1288,1649,1670]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha</emphasis>
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<emphasis id="B90FA66FBF529950FA83F9BBFAADD088" bold="true" box="[1300,1398,1649,1670]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="A23C1B14BF529950FA83F9BBFAADD088" box="[1300,1398,1649,1670]" pageId="6" pageNumber="224" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
mainly in the male genital characters listed above.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF529950FCC5F96CFBFBD102" pageId="6" pageNumber="224" type="etymology">
<paragraph id="8BC47A7DBF529950FCC5F96CFBFBD102" blockId="6.[812,1399,315,1804]" pageId="6" pageNumber="224">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF529950FCBBF90BFC7AD0D9" authorityName="Gorochov" authorityYear="2022" box="[812,929,1729,1751]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="intermedia">“intermedia</taxonomicName>
” (intermediate) due to the intermediate length of ectoparameres in the new species compared to both other species of this subgenus.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,289 @@
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<mods:title id="D113DE2B23CE84BF259B7C9F7F85CF00">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<taxonomicName id="4C7B01FEBF529950FCCCF8EAFBB3D15B" authority="Gorochov" authorityName="Gorochov" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF529950FCCCF8EAFA9CD139" box="[859,1351,1824,1848]" italics="true" pageId="6" pageNumber="224">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
Gorochov
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,
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<subSubSection id="C36129F6BF529950FB85F890FB49D17C" box="[1042,1170,1882,1906]" pageId="6" pageNumber="224" type="description">
<paragraph id="8BC47A7DBF529950FB85F890FB49D17C" blockId="6.[812,1399,1824,2059]" box="[1042,1170,1882,1906]" pageId="6" pageNumber="224">
<figureCitation id="134066F8BF529950FB85F890FB55D17C" box="[1042,1166,1882,1906]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="6" pageNumber="224">Figs 4759</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF52995FFCC5F8B5FF2ED768" lastPageId="9" lastPageNumber="227" pageId="6" pageNumber="224" type="materials_examined">
<paragraph id="8BC47A7DBF529950FCC5F8B5FAADDE04" blockId="6.[812,1399,1824,2059]" pageId="6" pageNumber="224">
MATERIAL EXAMINED.
<materialsCitation id="3B137020BF529950FBD4F8B5FCB9D1D4" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" county="Moramanga Distr." elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54C0C4DFBF529950FBD4F8B5FB48D19C" box="[1091,1171,1919,1938]" pageId="6" pageNumber="224" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7DB1F4BF529950FB24F8B7FB1CD19D" box="[1203,1223,1917,1939]" count="1" pageId="6" pageNumber="224" type="male"></specimenCount>
,
<emphasis id="B90FA66FBF529950FB45F8B5FA98D19C" bold="true" box="[1234,1347,1919,1938]" pageId="6" pageNumber="224">
<collectingCountry id="F36C3AEDBF529950FB45F8B5FA98D19C" box="[1234,1347,1919,1938]" name="Madagascar" pageId="6" pageNumber="224">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF529950FADAF84AFC7DD1A5" country="Madagascar" name="Toamasina" pageId="6" pageNumber="224">Toamasina Prov.</collectingRegion>
,
<collectingCounty id="62A502F1BF529950FC23F852FB87D1A4" box="[948,1116,1943,1963]" pageId="6" pageNumber="224">Moramanga Distr.</collectingCounty>
,
<collectingMunicipality id="6BA0E007BF529950FBFEF85DFAACD1A4" box="[1129,1399,1943,1963]" pageId="6" pageNumber="224">Analamazaotra Forest Station</collectingMunicipality>
near
<location id="8EA42CA6BF529950FCCEF865FC0DD1CC" LSID="urn:lsid:plazi:treatment:03D2CB6BBF52995FFCCCF8EAFB83D53F:8EA42CA6BF529950FCCEF865FC0DD1CC" box="[857,982,1967,1986]" country="Madagascar" county="Moramanga Distr." municipality="Analamazaotra Forest Station" name="Andasibe Vill" pageId="6" pageNumber="224" stateProvince="Toamasina">Andasibe Vill</location>
. ~
<quantity id="4C83D798BF529950FC7BF865FBF3D1CD" box="[1004,1064,1967,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">
<elevation id="00569D4EBF529950FC7BF865FBF3D1CD" box="[1004,1064,1967,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="224" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF529950FBE8F865FB23D1CC" box="[1151,1272,1967,1986]" pageId="6" pageNumber="224" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF529950FBE8F865FB23D1CC" box="[1151,1272,1967,1986]" pageId="6" pageNumber="224" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">820.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FA94F865FAADD1CC" box="[1283,1398,1967,1986]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FCA4F80DFC87D1D4" box="[819,860,1991,2010]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B137020BF529950FCE7F802FB02D1FC" collectingDate="2013-02-11" collectingDateMax="2013-03-20" collectingDateMin="2013-02-11" collectionCode="ZIN" collectorName="A. Gorochov &amp; L. Anisyutkin" country="Madagascar" county="Moramanga Distr." elevation="900" location="Andasibe Vill" municipality="Analamazaotra Forest Station" pageId="6" pageNumber="224" specimenCount="16" specimenCount-female="16" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="54C0C4DFBF529950FCE7F802FC12D1D5" box="[880,969,1992,2011]" pageId="6" pageNumber="224" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7DB1F4BF529950FC41F80DFBD8D1D5" box="[982,1027,1989,2011]" count="16" pageId="6" pageNumber="224" type="female">16 ♀</specimenCount>
, same data as for holotype, but
<date id="FFC55CBDBF529950FAD6F80DFC4BD1FC" pageId="6" pageNumber="224" value="2013-02-11" valueMax="2013-03-20" valueMin="2013-02-11">
<collectingDate id="EF81A555BF529950FAD6F80DFC4BD1FC" pageId="6" pageNumber="224" value="2013-02-11" valueMax="2013-03-20" valueMin="2013-02-11">11.II 20.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF529950FC0BF815FBC8D1FC" box="[924,1043,2015,2034]" pageId="6" pageNumber="224">A. Gorochov</collectorName>
,
<collectorName id="268E1FABBF529950FB88F82AFB40D1FC" box="[1055,1179,2015,2035]" pageId="6" pageNumber="224">L. Anisyutkin</collectorName>
(
<collectionCode id="ED6AE2B8BF529950FB3DF815FB09D1FC" box="[1194,1234,2015,2034]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="224" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B137020BF529950FB71F815FAADDE04" country="Madagascar" location="Torotorofotsy" pageId="6" pageNumber="224" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="9D7DB1F4BF529950FB71F815FAD6D1FD" box="[1254,1293,2013,2035]" count="1" pageId="6" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7DB1F4BF529950FA8BF815FAE7D1FD" box="[1308,1340,2013,2035]" count="2" pageId="6" pageNumber="224" type="female">2 ♀</specimenCount>
, same province and district, ~
<locationDeviation id="9A76635FBF529950FC61F83DFB2DDE04" box="[1014,1270,2039,2059]" location="Andasibe Vill." pageId="6" pageNumber="224">
<quantity id="4C83D798BF529950FC61F83DFBF7DE04" box="[1014,1068,2039,2058]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="224" unit="km" value="10.0">10 km</quantity>
NW of Andasibe Vill.
</locationDeviation>
,
<location id="8EA42CA6BF529950FB69F83DFAADDE04" LSID="urn:lsid:plazi:treatment:03D2CB6BBF52995FFCCCF8EAFB83D53F:8EA42CA6BF529950FB69F83DFAADDE04" box="[1278,1398,2039,2058]" country="Madagascar" name="Torotorofotsy" pageId="6" pageNumber="224" stateProvince="Toamasina">Torotorofotsy</location>
</materialsCitation>
</paragraph>
<caption id="DF042AF5BF539951FF75F8A3FEF4DE1F" ID-DOI="http://doi.org/10.5281/zenodo.13178134" ID-Zenodo-Dep="13178134" httpUri="https://zenodo.org/record/13178134/files/figure.png" pageId="7" pageNumber="225" startId="7.[226,263,1897,1916]" targetBox="[194,1390,320,1872]" targetPageId="7" targetType="figure">
<paragraph id="8BC47A7DBF539951FF75F8A3FAE0D1A2" blockId="7.[189,1399,1897,2065]" pageId="7" pageNumber="225">
Figs 3846.
<taxonomicName id="4C7B01FEBF539951FEDAF8A3FD07D172" authorityName="Gorochov" authorityYear="2022" box="[333,732,1897,1916]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF539951FEDAF8A3FD07D172" box="[333,732,1897,1916]" italics="true" pageId="7" pageNumber="225">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF539951FD76F8A3FCD7D172" bold="true" box="[737,780,1897,1916]" pageId="7" pageNumber="225">
<taxonomicNameLabel id="A23C1B14BF539951FD76F8A3FCD7D172" box="[737,780,1897,1916]" pageId="7" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 3839, 4243 — head with pronotum and fore legs (38 — from above; 39 — from side, 4243 — from side/above); 4041 — tegmina in rest position from above; 44 — genital plate from below; 4546 — distal part of ovipositor (45 — from side; 46 — from above); 3842 — male (3840 — holotype; 4142 — paratype); 4346 — female.
</paragraph>
<paragraph id="8BC47A7DBF539951FF75F87CFEF4DE1F" blockId="7.[189,1399,1897,2065]" pageId="7" pageNumber="225">
Рис. 3846.
<taxonomicName id="4C7B01FEBF539951FEC4F87FFD3DD1C6" authorityName="Gorochov" authorityYear="2022" box="[339,742,1973,1992]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Zvenellomorpha">
<emphasis id="B90FA66FBF539951FEC4F87FFD3DD1C6" box="[339,742,1973,1992]" italics="true" pageId="7" pageNumber="225">Zvenellomorpha (Zvenellomorpha) intermedia</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF539951FD79F87FFCC2D1C6" bold="true" box="[750,793,1973,1992]" pageId="7" pageNumber="225">
<taxonomicNameLabel id="A23C1B14BF539951FD79F87FFCC2D1C6" box="[750,793,1973,1992]" pageId="7" pageNumber="225" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 3839, 4243 — голова с переднеспинкой и передними ногами (38 — сверху; 39 — сбоку; 4243 — сбоку/сверху); 4041 — надкрыльЯ в положении покоЯ, сверху; 44 — генитальнаЯ пластинка, сниЗу; 4546 — дистальнаЯ часть Яйцеклада (45 — сбоку; 46 — сверху); 3842 — самец (3840 — голотип; 4142 — паратип); 43 46 — самка.
</paragraph>
</caption>
<caption id="DF042AF5BF5C995EFF75F8EBFDE7DE1F" ID-DOI="http://doi.org/10.5281/zenodo.13178136" ID-Zenodo-Dep="13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="8" pageNumber="226" startId="8.[226,263,1825,1844]" targetBox="[193,1395,317,1801]" targetPageId="8" targetType="figure">
<paragraph id="8BC47A7DBF5C995EFF75F8EBFEEDD19A" blockId="8.[189,1398,1825,2065]" pageId="8" pageNumber="226">
Figs 4759.
<taxonomicName id="4C7B01FEBF5C995EFEDAF8EBFD0FD13A" authorityName="Gorochov" authorityYear="2022" box="[333,724,1825,1844]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5C995EFEDAF8EBFD0FD13A" box="[333,724,1825,1844]" italics="true" pageId="8" pageNumber="226">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5C995EFD4DF8EBFCDFD13A" box="[730,772,1825,1844]" pageId="8" pageNumber="226" rank="species">
<emphasis id="B90FA66FBF5C995EFD4DF8EBFD24D13A" bold="true" box="[730,767,1825,1844]" pageId="8" pageNumber="226">sp.n</emphasis>
.
</taxonomicNameLabel>
:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female.
</paragraph>
<paragraph id="8BC47A7DBF5C995EFF75F854FDE7DE1F" blockId="8.[189,1398,1825,2065]" pageId="8" pageNumber="226">
Рис. 4759.
<taxonomicName id="4C7B01FEBF5C995EFEC7F857FD03D1BE" authorityName="Gorochov" authorityYear="2022" box="[336,728,1949,1968]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="separata" status="sp. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5C995EFEC7F857FD03D1BE" box="[336,728,1949,1968]" italics="true" pageId="8" pageNumber="226">Zvenellomorpha (Azvenellomorpha) separata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5C995EFD48F857FCD2D1BE" box="[735,777,1949,1968]" pageId="8" pageNumber="226" rank="species">
<emphasis id="B90FA66FBF5C995EFD48F857FCDFD1BE" bold="true" box="[735,772,1949,1968]" pageId="8" pageNumber="226">sp.n</emphasis>
.
</taxonomicNameLabel>
:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF5D995FFF2AFEF6FF2ED768" blockId="9.[189,776,315,2065]" pageId="9" pageNumber="227">
Forest Reserve, ~
<quantity id="4C83D798BF5D995FFEC0FEF1FE46D741" box="[343,413,315,335]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="227" unit="m" value="1000.0">1000 m</quantity>
, at light,
<date id="FFC55CBDBF5D995FFE65FEF1FD51D740" box="[498,650,315,334]" pageId="9" pageNumber="227" value="2013-02-22" valueMax="2013-03-11" valueMin="2013-02-22">22.II11.III.2013</date>
, A. Gorochov (
<collectionCode id="ED6AE2B8BF5D995FFF54FE99FF30D768" box="[195,235,339,358]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="227" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFF75FEA1FAACD4EF" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995FFF75FEA1FE77D182" blockId="9.[189,776,315,2065]" pageId="9" pageNumber="227">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF5D995FFE44FEA1FDF6D78F" box="[467,557,363,385]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
). General appearance similar to that of all species of
<taxonomicName id="4C7B01FEBF5D995FFE67FE4CFD55D795" authorityName="Gorochov" authorityYear="2004" box="[496,654,390,411]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFE67FE4CFD55D795" box="[496,654,390,411]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFD02FE4CFD10D795" box="[661,715,390,411]" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
Body coloration yellowish with following pattern: eyes greyish; dorsum of epicranium greyish brown with a pair of yellowish longitudinal stripes running from rostral apex to yellowish lateral ocelli and from these ocelli to posterior edge of head (along medial edges of eyes and behind them), with a pair of short and thin light longitudinal stripes between previous stripes on posterior part of epicranial dorsum, with also light median line on posterior two thirds of this dorsum (
<figureCitation id="134066F8BF5D995FFD26FD91FF03D485" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 47 49</figureCitation>
); upper parts of membranes of antennal cavities and small areas on upper parts of genae behind eyes slightly darkened; scape with brown dorsal band along medial edge, darkish ventral stripe along same edge and darker (greyish brown) ventromedian spot in apical part; rest of antenna with brown to light brown spots in proximal and middle parts as well as with almost uniformly light brown distal part; pronotum with dark brown disc having a few thin lightish marks, with slightly darkish longitudinal band on each lateral lobe (this band occupying upper two thirds of this lobe but separated from dark disc by longitudinal light stripe), with a few small barely darker marks on above-mentioned darkish band, and with several dark and light spots along anterior edge of this lobe; legs with small and sparse dark and darkish spots (
<figureCitation id="134066F8BF5D995FFD4BFC1BFF25D20E" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 4749</figureCitation>
); tegmen with yellowish grey dorsal field having darker (greyish brown) venation as well as yellowish both dot near plectrum and narrow stripe along lateral edge of distal half of this field, with light greyish (semitransparent) lateral field having greyish brown longitudinal veins and whitish crossveins, and with humeral region between these fields having dark (brown) proximal half of M and whitish crossveins as well as almost semimembranous membranes (
<figureCitation id="134066F8BF5D995FFD48FB6CFF03D2D8" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 51</figureCitation>
); hind wings light greyish with somewhat darkened distal parts (in rest condition); anal plate with a pair of rather large greyish brown spots in its laterodorsal corners (
<figureCitation id="134066F8BF5D995FFD17FB3CFD1CD302" box="[640,711,1270,1292]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 50</figureCitation>
); cerci with very small and moderately numerous darkish marks. Eyes rather large, slightly higher than long (
<figureCitation id="134066F8BF5D995FFD11FAE1FD27D34E" box="[646,764,1323,1345]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 4749</figureCitation>
); rostrum practically angular in profile; scape approximately 2.3 times as wide as space between antennal cavities (
<figureCitation id="134066F8BF5D995FFD25FAABFD20D378" box="[690,763,1377,1399]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 49</figureCitation>
); pronotum as in
<figureCitation id="134066F8BF5D995FFEC2FAB1FE1FD39E" box="[341,452,1403,1425]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 4748</figureCitation>
; metanotal gland absent; fore leg with only inner tympanum which oval, almost as long as width of fore tibia, and almost as wide as half of latter width; tegmina distinctly protruding beyond apices of hind femora and with venation as in
<figureCitation id="134066F8BF5D995FFE0AFA2CFE38D3F2" box="[413,483,1510,1532]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 51</figureCitation>
; hind wings clearly protruding beyond tegminal apices; anal plate small, transverse, truncate apically, widely rounded distolaterally and with large and almost longitudinally oval membranous median area (
<figureCitation id="134066F8BF5D995FFD24F9FCFD20D042" box="[691,763,1590,1612]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 50</figureCitation>
); genital plate more or less similar to that of
<taxonomicName id="4C7B01FEBF5D995FFDC3F99BFD29D068" authorityName="Gorochov" authorityYear="2004" box="[596,754,1617,1638]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFDC3F99BFD29D068" box="[596,754,1617,1638]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFD60F99BFF07D08F" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
but with slightly wider distal lobule having almost angular apex. Genitalia (
<figureCitation id="134066F8BF5D995FFE1CF94CFDD9D092" box="[395,514,1670,1692]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 5254</figureCitation>
): epiphallus more or less similar to that of other subgenera of this genus but with dorsoapical processes having much smaller (poorly distinct) distal tubercles, and with ventroapical lobules much shorter and more hooked; ectoparameres distinctly shorter; anterior sclerotized ribbon of formula longer and narrower than in
<taxonomicName id="4C7B01FEBF5D995FFF2AF8ECFE80D135" authorityName="Gorochov" authorityYear="2004" box="[189,347,1830,1851]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90FA66FBF5D995FFF2AF8ECFE80D135" box="[189,347,1830,1851]" italics="true" pageId="9" pageNumber="227">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5D995FFEF2F8ECFE45D135" box="[357,414,1830,1851]" pageId="9" pageNumber="227" sensu="stricto">s. str.</taxonomicNameLabel>
(i.e. not plate-like), located clearly less parallel to median part of rest of this formula than in the latter subgenus and in
<taxonomicName id="4C7B01FEBF5D995FFE30F891FD87D17E" authorityName="Gorochov" authorityYear="2022" box="[423,604,1883,1904]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Euzvenellomorpha">
<emphasis id="B90FA66FBF5D995FFE30F891FD87D17E" box="[423,604,1883,1904]" italics="true" pageId="9" pageNumber="227">Euzvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5D995FFDF1F891FD13D17E" bold="true" box="[614,712,1883,1904]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFDF1F891FD13D17E" box="[614,712,1883,1904]" pageId="9" pageNumber="227" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
Spermatophore as in
<figureCitation id="134066F8BF5D995FFEF7F8BCFE73D182" box="[352,424,1910,1932]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 55</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFF75F85BFA95D75F" blockId="9.[189,776,315,2065]" lastBlockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Variations. Second male distinguished from
<typeStatus id="54C0C4DFBF5D995FFD04F85BFD3CD1A9" box="[659,743,1937,1959]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
by insignificantly less contrast coloration of head, lighter pronotal disc, yellowish brown tinge of tegmina, somewhat different venation of their dorsal field (
<figureCitation id="134066F8BF5D995FFE6CF82BFD99D1F8" box="[507,578,2016,2039]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 56</figureCitation>
), as well as genitalia with slightly smaller anterolateral (curved posteriorly) parts of epiphallus and barely lower lateral plates of formula.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FE9CFBE6D428" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Female. Coloration and structure of body practically as in males, but scape somewhat lighter, dorsal tegminal field with 910 almost straight (barely oblique) longitudinal veins as well as rather numerous and partly irregular crossveins, genital plate almost as wide as long and somewhat narrowing in distal half as well as with distinct roundly angular posteromedian notch (
<figureCitation id="134066F8BF5D995FFC5FFE3CFBCAD402" box="[968,1041,502,524]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Fig. 57</figureCitation>
); ovipositor normal and with drilling distal part as in
<figureCitation id="134066F8BF5D995FFC5EFDDBFBE2D428" box="[969,1081,528,551]" captionStart="Figs 4759" captionStartId="8.[226,263,1825,1844]" captionTargetBox="[193,1395,317,1801]" captionTargetId="figure-219@8.[189,1399,314,1813]" captionTargetPageId="8" captionText="Figs 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — head with pronotum and fore leg (47 — from side; 48 — from above); 49 — head in front; 50 — anal plate from above; 51 — right tegmen; 5254 — genitalia (52 — from side; 53 — from above; 54 — from below); 55 — spermatophore from side; 56 — wings with middle leg and antennal part in rest position from above; 57 — genital plate from below; 5859 — distal part of ovipositor (58 — from below; 59 —from side); 4756 — male (4755 — holotype; 56 — paratype); 57 59 — female. Рис. 4759. Zvenellomorpha (Azvenellomorpha) separata sp.n.:4748 — голова с переднеспинкой и передней ногой (47 — сбоку; 48 — сверху); 49 — голова, спереди; 50 — анальнаЯ пластинка, сверху; 51 — правое надкрылье; 5254 — гениталии (52 — сбоку; 53 — сверху; 54 — сниЗу); 55 — сперматофор, сбоку; 56 — крыльЯ со средней ногой и частью антенны в положении покоЯ, сверху; 57 — генитальнаЯ пластинка сниЗу; 5859 — дистальнаЯ часть Яйцеклада (58 — сниЗу; 59 — сбоку); 4756 — самец (4755 — голотип; 56 — паратип); 5759 — самка." figureDoi="http://doi.org/10.5281/zenodo.13178136" httpUri="https://zenodo.org/record/13178136/files/figure.png" pageId="9" pageNumber="227">Figs 5859</figureCitation>
.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FDE1FB9DD49F" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
Length in mm. Body:
<specimenCount id="9D7DB1F4BF5D995FFBB4FDE2FBE2D44E" box="[1059,1081,552,576]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
14.515.5,
<specimenCount id="9D7DB1F4BF5D995FFB3DFDE2FB6CD44E" box="[1194,1207,552,576]" count="1" pageId="9" pageNumber="227" type="female"></specimenCount>
1517.5; body with wings:
<specimenCount id="9D7DB1F4BF5D995FFCE2FD89FC50D455" box="[885,907,579,603]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
2425.5,
<specimenCount id="9D7DB1F4BF5D995FFC66FD89FC25D455" box="[1009,1022,579,603]" count="1" pageId="9" pageNumber="227" type="female"></specimenCount>
2629; pronotum:
<specimenCount id="9D7DB1F4BF5D995FFB57FD89FB0DD455" box="[1216,1238,579,603]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
2.72.9,
<specimenCount id="9D7DB1F4BF5D995FFAA1FD89FA98D455" box="[1334,1347,579,603]" count="1" pageId="9" pageNumber="227" type="female"></specimenCount>
3.1 3.4; tegmina:
<specimenCount id="9D7DB1F4BF5D995FFC26FD94FC1CD478" box="[945,967,606,630]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
17.518.5;
<specimenCount id="9D7DB1F4BF5D995FFBACFD94FB93D478" box="[1083,1096,606,630]" count="1" pageId="9" pageNumber="227" type="female"></specimenCount>
1820; hind femora:
<specimenCount id="9D7DB1F4BF5D995FFA8FFD94FAF5D478" box="[1304,1326,606,630]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
1314,
<specimenCount id="9D7DB1F4BF5D995FFCBBFDB2FCE2D49E" box="[812,825,632,656]" count="1" pageId="9" pageNumber="227" type="female"></specimenCount>
13.515; ovipositor 99.5.
</paragraph>
<paragraph id="8BC47A7DBF5D995FFCC5FD5CFAACD4EF" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
COMPARISON. The new species differs from all the other congeners in the same characters as the subgenus
<taxonomicName id="4C7B01FEBF5D995FFCBBFD01FC0ED4EE" authorityName="Gorochov" authorityYear="2022" box="[812,981,715,736]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Azvenellomorpha">
<emphasis id="B90FA66FBF5D995FFCBBFD01FC0ED4EE" box="[812,981,715,736]" italics="true" pageId="9" pageNumber="227">Azvenellomorpha</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5D995FFC4CFD01FBE6D4EE" bold="true" box="[987,1085,715,736]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFC4CFD01FBE6D4EE" box="[987,1085,715,736]" pageId="9" pageNumber="227" rank="subGenus">subgen.n.</taxonomicNameLabel>
</emphasis>
(see the subgeneric key above).
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFCC5FD2CFB83D53F" pageId="9" pageNumber="227" type="etymology">
<paragraph id="8BC47A7DBF5D995FFCC5FD2CFB83D53F" blockId="9.[812,1399,315,817]" pageId="9" pageNumber="227">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF5D995FFCBBFCCBFC53D518" authorityName="Gorochov" authorityYear="2022" box="[812,904,769,790]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="separata">“separata</taxonomicName>
” (separate) due to some distinct differences in the structure of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,196 @@
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<mods:title id="34B2A5DA7C0EA2A53E3157A467A362A9">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<mods:date id="2B4C0EE8242BF5969DC588D18F1B2B04">2022</mods:date>
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<treatment id="03D2CB6BBF5D995DFC19FC8FFE4CD4A5" LSID="urn:lsid:plazi:treatment:03D2CB6BBF5D995DFC19FC8FFE4CD4A5" httpUri="http://treatment.plazi.org/id/03D2CB6BBF5D995DFC19FC8FFE4CD4A5" lastPageId="11" lastPageNumber="229" pageId="9" pageNumber="227">
<subSubSection id="C36129F6BF5D995FFC19FC8FFACFD553" box="[910,1300,837,861]" pageId="9" pageNumber="227" type="nomenclature">
<paragraph id="8BC47A7DBF5D995FFC19FC8FFACFD553" blockId="9.[812,1399,837,2061]" box="[910,1300,837,861]" pageId="9" pageNumber="227">
<heading id="D08CCD11BF5D995FFC19FC8FFACFD553" box="[910,1300,837,861]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="227" reason="8">
<taxonomicName id="4C7B01FEBF5D995FFC19FC8FFB0FD553" authority="Gorochov" authorityName="Gorochov" authorityYear="2022" box="[910,1236,837,861]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5D995FFC19FC8FFB85D553" box="[910,1118,837,861]" italics="true" pageId="9" pageNumber="227">Ultratrella? ornata</emphasis>
Gorochov
</taxonomicName>
,
<emphasis id="B90FA66FBF5D995FFB48FC8FFACFD553" bold="true" box="[1247,1300,837,861]" pageId="9" pageNumber="227">
<taxonomicNameLabel id="A23C1B14BF5D995FFB48FC8FFACFD553" box="[1247,1300,837,861]" pageId="9" pageNumber="227" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFB85FCA8FB49D574" box="[1042,1170,866,890]" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995FFB85FCA8FB49D574" blockId="9.[812,1399,837,2061]" box="[1042,1170,866,890]" pageId="9" pageNumber="227">
<figureCitation id="134066F8BF5D995FFB85FCA8FB56D574" box="[1042,1165,866,890]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 6067</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995FFCC5FC4DFCBED5EC" pageId="9" pageNumber="227" type="materials_examined">
<paragraph id="8BC47A7DBF5D995FFCC5FC4DFCBED5EC" blockId="9.[812,1399,837,2061]" pageId="9" pageNumber="227">
<materialsCitation id="3B137020BF5D995FFCC5FC4DFCBED5EC" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="9" pageNumber="227" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
MATERIAL EXAMINED.
<typeStatus id="54C0C4DFBF5D995FFBD4FC4DFB48D594" box="[1091,1171,903,922]" pageId="9" pageNumber="227" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7DB1F4BF5D995FFB24FC4FFB1CD595" box="[1203,1223,901,923]" count="1" pageId="9" pageNumber="227" type="male"></specimenCount>
,
<emphasis id="B90FA66FBF5D995FFB45FC4DFA98D594" bold="true" box="[1234,1347,903,922]" pageId="9" pageNumber="227">
<collectingCountry id="F36C3AEDBF5D995FFB45FC4DFA98D594" box="[1234,1347,903,922]" name="Madagascar" pageId="9" pageNumber="227">Madagascar</collectingCountry>
</emphasis>
,
<collectingRegion id="49BFB49FBF5D995FFADAFC42FC7DD5BD" country="Madagascar" name="Toamasina" pageId="9" pageNumber="227">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF5D995FFC23FC6AFB87D5BC" box="[948,1116,927,947]" pageId="9" pageNumber="227">Moramanga Distr.</collectingMunicipality>
,
<location id="8EA42CA6BF5D995FFBFEFC55FAACD5BC" LSID="urn:lsid:plazi:treatment:03D2CB6BBF5D995DFC19FC8FFE4CD4A5:8EA42CA6BF5D995FFBFEFC55FAACD5BC" box="[1129,1399,927,947]" country="Madagascar" municipality="Moramanga Distr." name="Analamazaotra Forest Station" pageId="9" pageNumber="227" stateProvince="Toamasina">Analamazaotra Forest Station</location>
near Andasibe Vill. ~
<quantity id="4C83D798BF5D995FFC7BFC7DFBF3D5C5" box="[1004,1064,951,971]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="227" unit="m" value="900.0">
<elevation id="00569D4EBF5D995FFC7BFC7DFBF3D5C5" box="[1004,1064,951,971]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="227" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
<date id="FFC55CBDBF5D995FFBE8FC7DFB23D5C4" box="[1151,1272,951,970]" pageId="9" pageNumber="227" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">
<collectingDate id="EF81A555BF5D995FFBE8FC7DFB23D5C4" box="[1151,1272,951,970]" pageId="9" pageNumber="227" value="2013-03-08" valueMax="2013-03-20" valueMin="2013-03-08">820.III.2013</collectingDate>
</date>
,
<collectorName id="268E1FABBF5D995FFA94FC7DFAADD5C4" box="[1283,1398,951,970]" pageId="9" pageNumber="227">A. Gorochov</collectorName>
(
<collectionCode id="ED6AE2B8BF5D995FFCA4FC05FC80D5EC" box="[819,859,975,994]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="227" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5D995DFCC5FC2DFE8ED452" lastPageId="11" lastPageNumber="229" pageId="9" pageNumber="227" type="description">
<paragraph id="8BC47A7DBF5D995CFCC5FC2DFAD0D428" blockId="9.[812,1399,837,2061]" lastBlockId="10.[812,1399,315,551]" lastPageId="10" lastPageNumber="228" pageId="9" pageNumber="227">
DESCRIPTION. Male (
<typeStatus id="54C0C4DFBF5D995FFBD4FC2DFB46D5F3" box="[1091,1181,999,1021]" pageId="9" pageNumber="227" type="holotype">holotype</typeStatus>
). General appearance distinctly different from that of
<taxonomicName id="4C7B01FEBF5D995FFBF4FBC9FAA8D219" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[1123,1395,1026,1048]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5D995FFBF4FBC9FB10D219" box="[1123,1227,1026,1048]" italics="true" pageId="9" pageNumber="227">U. gracilis</emphasis>
<bibRefCitation id="EFEA078CBF5D995FFB45FBC8FAA8D219" author="Gorochov A. V." box="[1234,1395,1026,1048]" pageId="9" pageNumber="227" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
, i.e. body clearly smaller, and coloration more contrast: head and pronotum (
<figureCitation id="134066F8BF5D995FFC29FBFDFB94D243" box="[958,1103,1079,1101]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 6263</figureCitation>
) dark grey with slightly lighter (grey) stripe along posterior edge of epicranial dorsum, yellowish ocelli and dorsal transverse band between anterior parts of eyes (this band situated behind median ocellus but very near it, and lateral ocelli located on this band), almost black scape and pedicel as well as rostral area, brown antennal flagellum and light brown to yellowish spots on mouthparts as well as on pronotal disc (lateral lobes of pronotum with only sparse light dots;
<figureCitation id="134066F8BF5D995FFBA3FAC7FB79D32C" box="[1076,1186,1292,1315]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 63</figureCitation>
); fore and middle legs dark greyish brown with light brown ventral surfaces of tibiae and tarsi as well as with yellowish trochanters and coxae (each coxa with darkish lateral area); hind legs brown with light brown basal parts of femora and all spines and spurs (
<figureCitation id="134066F8BF5D995FFCA4FA58FC7ED3A6" box="[819,933,1426,1448]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 6063</figureCitation>
); tegmina light grey with yellow spot at base of each tegmen (
<figureCitation id="134066F8BF5D995FFC22FA67FBF3D3CC" box="[949,1064,1452,1475]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 61, 63</figureCitation>
), greyish brown to blackish venation of dorsal field and longitudinal veins of lateral field, whitish crossveins in lateral field, slightly darkened spots on almost all membranes and rather wide whitish humeral stripe (
<figureCitation id="134066F8BF5D995FFCA4F9DDFC72D023" box="[819,937,1559,1581]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 6061</figureCitation>
); visible distal parts of hind wings also light grey; other parts of body venter (including abdominal sternites, genital plate and cerci) yellowish to light brown. Head rather short, slightly concave dorsally (between middle parts of eyes), somewhat depressed dorsoventrally, with large eyes which slightly higher than long, with rather large and almost round ocelli, with short mouthparts and very short palpi, as well as with scape barely wider than space between antennal cavities (
<figureCitation id="134066F8BF5D995FFC14F927FBC3D10C" box="[899,1048,1772,1795]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 6263</figureCitation>
); pronotum distinctly transverse and moderately high (somewhat dorsoventrally depressed), with concave anterior edge of disc and convex posterior one (
<figureCitation id="134066F8BF5D995FFADCF8E8FCAAD15C" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 60, 63</figureCitation>
); fore and middle legs rather short, not flattened, without keels and with fore tibiae having outer and inner tympana oval and almost equal to each other in size (each of them 1.61.7 times as wide as maximal width of fore tibia and barely longer than latter width); tegmina strongly protruding beyond hind femora in rest position, with mirror clearly smaller than in
<taxonomicName id="4C7B01FEBF5D995FFC52F817FBF5D1FC" box="[965,1070,2013,2034]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="227" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5D995FFC52F817FBF5D1FC" box="[965,1070,2013,2034]" italics="true" pageId="9" pageNumber="227">U. gracilis</emphasis>
</taxonomicName>
and with rest venation as in
<figureCitation id="134066F8BF5D995FFADAF817FCB2DE03" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="9" pageNumber="227">Figs 6061</figureCitation>
(lateral field with very narrow Sc-R area, 1516 branches of Sc, rather numerous crossveins in proximal half of this field and almost without crossveins in its distal half); hind wings distinctly protruding beyond tegminal apices; genital plate practically intermediate between those of
<taxonomicName id="4C7B01FEBF5E995CFD29FE41FEC0D7B5" authorityName="Gorochov" authorityYear="2004" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B90FA66FBF5E995CFD29FE41FEC0D7B5" italics="true" pageId="10" pageNumber="228">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5E995CFEB7FE6CFE99D7B2" box="[288,322,422,444]" pageId="10" pageNumber="228" sensu="lato">s. l.</taxonomicNameLabel>
and
<taxonomicName id="4C7B01FEBF5E995CFEE6FE6DFE03D7B5" box="[369,472,422,444]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFEE6FE6DFE03D7B5" box="[369,472,422,444]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
[see
<bibRefCitation id="EFEA078CBF5E995CFD9DFE6CFD70D7B5" author="Gorochov A. V." box="[522,683,422,444]" pageId="10" pageNumber="228" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
: figs X, 2 and 7]. Genitalia (
<figureCitation id="134066F8BF5E995CFEE4FE0BFE3FD7D8" box="[371,484,448,471]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="10" pageNumber="228">Figs 6466</figureCitation>
) also more or less similar to those of
<taxonomicName id="4C7B01FEBF5E995CFE82FE16FE5BD7FE" box="[277,384,475,497]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFE82FE16FE5BD7FE" box="[277,384,475,497]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
(including similar structure of formula and rachis), but: distal part of epiphallus higher (less narrow in profile) and with dorsoapical processes triangular (not bilobate) in profile and located near convex part of epiphallus (in
<taxonomicName id="4C7B01FEBF5E995CFCDAFE9DFC6ED765" box="[845,949,342,364]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5E995CFCDAFE9DFC6ED765" box="[845,949,342,364]" italics="true" pageId="10" pageNumber="228">U. gracilis</emphasis>
</taxonomicName>
, this processes and latter part located not near each other); ventroapical epiphallic lobules higher (wider in profile); formula with shorter anterior ribbon and with lateral lobes angularly projected backwards (
<emphasis id="B90FA66FBF5E995CFB0BFE6DFB6BD7B2" box="[1180,1200,423,444]" italics="true" pageId="10" pageNumber="228">vs</emphasis>
. not projected backwards); ectoparameres clearly longer; rachis with semimembranous subapical inflation (
<emphasis id="B90FA66FBF5E995CFBAAFE16FB8AD7FF" box="[1085,1105,476,497]" italics="true" pageId="10" pageNumber="228">vs</emphasis>
. without any inflation in distal part). Spermatophore with elongately rounded ampulla (oth- er structures of spermatophore missing;
<figureCitation id="134066F8BF5E995CFB21FDDBFB24D428" box="[1206,1279,528,551]" captionStart="Figs 6067" captionStartId="10.[226,263,1921,1940]" captionTargetBox="[192,1395,599,1883]" captionTargetId="figure-155@10.[189,1399,593,1901]" captionTargetPageId="10" captionText="Figs 6067. Ultratrella ornata sp.n., male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side. Рис. 6067. Ultratrella ornata sp.n., самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="10" pageNumber="228">Fig. 67</figureCitation>
).
</paragraph>
<caption id="DF042AF5BF5E995CFF75F84BFB98DE1E" ID-DOI="http://doi.org/10.5281/zenodo.13178138" ID-Zenodo-Dep="13178138" httpUri="https://zenodo.org/record/13178138/files/figure.png" pageId="10" pageNumber="228" startId="10.[226,263,1921,1940]" targetBox="[192,1395,599,1883]" targetPageId="10" targetType="figure">
<paragraph id="8BC47A7DBF5E995CFF75F84BFBD0D1CA" blockId="10.[189,1399,1921,2064]" pageId="10" pageNumber="228">
Figs 6067.
<taxonomicName id="4C7B01FEBF5E995CFEDAF84BFE33D19A" authorityName="Gorochov" authorityYear="2022" box="[333,488,1921,1940]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5E995CFEDAF84BFE33D19A" box="[333,488,1921,1940]" italics="true" pageId="10" pageNumber="228">Ultratrella ornata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5E995CFE7AF84BFDC3D19A" bold="true" box="[493,536,1921,1940]" pageId="10" pageNumber="228">
<taxonomicNameLabel id="A23C1B14BF5E995CFE7AF84BFDC3D19A" box="[493,536,1921,1940]" pageId="10" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, male: 60 — body from side; 61 — wings, middle and hind legs as well as part of antennae and cerci in rest position from above; 62 — head with fore legs in front; 63 — head with pronotum and fore legs from above; 6466 — genitalia (64 — from above; 65 — from below; 66 — from side); 67 — ampulla of spermatophore from side.
</paragraph>
<paragraph id="8BC47A7DBF5E995CFF75F807FB98DE1E" blockId="10.[189,1399,1921,2064]" pageId="10" pageNumber="228">
Рис. 6067.
<taxonomicName id="4C7B01FEBF5E995CFED8F807FE30D1EE" authorityName="Gorochov" authorityYear="2022" box="[335,491,1997,2016]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="228" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B90FA66FBF5E995CFED8F807FE30D1EE" box="[335,491,1997,2016]" italics="true" pageId="10" pageNumber="228">Ultratrella ornata</emphasis>
</taxonomicName>
<emphasis id="B90FA66FBF5E995CFE66F807FDC7D1EE" bold="true" box="[497,540,1997,2016]" pageId="10" pageNumber="228">
<taxonomicNameLabel id="A23C1B14BF5E995CFE66F807FDC7D1EE" box="[497,540,1997,2016]" pageId="10" pageNumber="228" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, самец: 60 — тело, сбоку; 61 — крыльЯ, средние и ЗаднЯЯ ноги, а также часть усиков и церки в положении покоЯ, сверху; 62 — голова c передними ногами, спереди; 63 — голова с переднеспинкой и передними ногами, сверху; 6466 — гениталии (64 — сверху; 65 — сниЗу; 66 — сбоку); 67 — ампула сперматофора, сбоку.
</paragraph>
</caption>
<paragraph id="8BC47A7DBF5F995DFF75FEF1FE49D75F" blockId="11.[189,775,315,684]" box="[226,402,315,337]" pageId="11" pageNumber="229">Female unknown.</paragraph>
<paragraph id="8BC47A7DBF5F995DFF75FE9CFDC9D788" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">Length in mm. Body 10; body with wings 27; pronotum 2.8; tegmina 18.7; hind femora 10.</paragraph>
<paragraph id="8BC47A7DBF5F995DFF75FE41FE8ED452" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">
COMPARISON. The new species is clearly different from
<taxonomicName id="4C7B01FEBF5F995DFF65FE6DFE81D7B5" box="[242,346,422,444]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5F995DFF65FE6DFE81D7B5" box="[242,346,422,444]" italics="true" pageId="11" pageNumber="229">U. gracilis</emphasis>
</taxonomicName>
(
<typeStatus id="54C0C4DFBF5F995DFEFFFE6CFE48D7B5" box="[360,403,422,443]" pageId="11" pageNumber="229">type</typeStatus>
and only species of this genus) in the body smaller, coloration much more dark and contrast, a relatively smaller mirror in the male tegmen, and the above-mentioned characters of the male genitalia. These differences may be treated also as subgeneric or even generic ones, but this question may be clarified only after the study of some additional data.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFF75FDABFE4CD4A5" pageId="11" pageNumber="229" type="etymology">
<paragraph id="8BC47A7DBF5F995DFF75FDABFE4CD4A5" blockId="11.[189,775,315,684]" pageId="11" pageNumber="229">
ETYMOLOGY. This species name is the Latin word
<taxonomicName id="4C7B01FEBF5F995DFF2AFDB1FEDED49E" authorityName="Gorochov" authorityYear="2022" box="[189,261,635,656]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="species" species="ornata">“ornata</taxonomicName>
” (ornated, adorned) due to the characteristic (contrasting) body pattern.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,146 @@
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<mods:title id="AA475EF02056601B017E3F3F074D3013">Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 15: Asymmetrella gen. n., Zvenellomorpha and Ultratrella from Madagascar</mods:title>
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<taxonomicName id="4C7B01FEBF5F995DFE88FD0AFD78D4D6" ID-CoL="7DGMJ" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[287,675,704,728]" class="Insecta" family="Gryllidae" genus="Ultratrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FA66FBF5F995DFE88FD0AFE36D4D6" box="[287,493,704,728]" italics="true" pageId="11" pageNumber="229">Ultratrella gracilis</emphasis>
<bibRefCitation id="EFEA078CBF5F995DFE62FD0AFD78D4D6" author="Gorochov A. V." box="[501,675,704,728]" pageId="11" pageNumber="229" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFF75FD2FFD04D526" pageId="11" pageNumber="229" type="materials_examined">
<paragraph id="8BC47A7DBF5F995DFF75FD2FFD04D526" blockId="11.[189,775,704,1235]" pageId="11" pageNumber="229">
<materialsCitation id="3B137020BF5F995DFF75FD2FFD00D526" collectingDate="2013-03-08" collectingDateMax="2013-03-20" collectingDateMin="2013-03-08" collectionCode="ZIN" collectorName="A. Gorochov" country="Madagascar" elevation="900" location="Analamazaotra Forest Station" municipality="Moramanga Distr." pageId="11" pageNumber="229" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina">
MATERIAL EXAMINED.
<emphasis id="B90FA66FBF5F995DFE43FD2FFD9ED4F6" bold="true" box="[468,581,741,760]" pageId="11" pageNumber="229">
<collectingCountry id="F36C3AEDBF5F995DFE43FD2FFD9ED4F6" box="[468,581,741,760]" name="Madagascar" pageId="11" pageNumber="229">Madagascar</collectingCountry>
</emphasis>
:
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,
<specimenCount id="9D7DB1F4BF5F995DFD16FD2FFD46D4F6" box="[641,669,738,760]" count="2" pageId="11" pageNumber="229" type="female">2 ♀</specimenCount>
,
<collectingRegion id="49BFB49FBF5F995DFD3FFD2FFF35D51E" country="Madagascar" name="Toamasina" pageId="11" pageNumber="229">Toamasina Prov.</collectingRegion>
,
<collectingMunicipality id="6BA0E007BF5F995DFF61FD37FE4CD51E" box="[246,407,765,784]" pageId="11" pageNumber="229">Moramanga Distr.</collectingMunicipality>
,
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near Andasibe Vill. ~
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<elevation id="00569D4EBF5F995DFE89FCDFFE83D526" box="[286,344,789,808]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="229" unit="m" value="900.0">900 m</elevation>
</quantity>
, at light,
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</date>
,
<collectorName id="268E1FABBF5F995DFDB9FCDFFD44D526" box="[558,671,789,808]" pageId="11" pageNumber="229">A. Gorochov</collectorName>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFF75FCE7FAAAD428" pageId="11" pageNumber="229" type="description">
<paragraph id="8BC47A7DBF5F995DFF75FCE7FC2DD762" blockId="11.[189,775,704,1235]" lastBlockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
DESCRIPTION. Female (nov.). General appearance similar to that of male
<typeStatus id="54C0C4DFBF5F995DFE05FC8DFE3CD553" box="[402,487,839,861]" pageId="11" pageNumber="229" type="holotype">holotype</typeStatus>
[
<bibRefCitation id="EFEA078CBF5F995DFE6FFC8DFD7BD552" author="Gorochov A. V." box="[504,672,839,861]" pageId="11" pageNumber="229" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
] but with following differences: coloration even more uniformly light greyish brown, i.e. with less distinct lighter (yellowish white) stripes on head and pronotum, and practically without whitish humeral stripe on each tegmen; tegminal dorsal field with 1314 longitudinal veins which barely S-shaped in lateral part of proximal half, as well as with numerous and slightly irregular crossveins; tegminal lateral field with 1517 oblique branches of Sc which partly and slightly arcuate, with moderately numerous crossveins, and with area between Sc stem and R moderately narrow (in male, this area very strongly narrowed); genital plate elongate but slightly shorter than two last abdominal sternites together, gradually narrowing to roundly truncate apex having very short and roundly angular posteromedian notch; ovipositor long and with distinctly drilling apical part more or less similar to that of
<taxonomicName id="4C7B01FEBF5F995DFCBBFE9CFC11D765" authorityName="Gorochov" authorityYear="2004" box="[812,970,342,363]" class="Insecta" family="Gryllidae" genus="Zvenellomorpha" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="229" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B90FA66FBF5F995DFCBBFE9CFC11D765" box="[812,970,342,363]" italics="true" pageId="11" pageNumber="229">Zvenellomorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C1B14BF5F995DFC45FE9CFC2DD762" box="[978,1014,342,364]" pageId="11" pageNumber="229" sensu="lato">s. l.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC47A7DBF5F995DFCC5FEBBFB65D7D8" blockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
Male. Body very similar to that of
<typeStatus id="54C0C4DFBF5F995DFB47FEBBFAFED789" box="[1232,1317,369,391]" pageId="11" pageNumber="229" type="holotype">holotype</typeStatus>
of this species, but tegmina with slightly longer chords and barely smaller mirror in dorsal field; genitalia almost identical to those of this
<typeStatus id="54C0C4DFBF5F995DFC3BFE0BFBDAD7D9" box="[940,1025,449,471]" pageId="11" pageNumber="229" type="holotype">holotype</typeStatus>
[
<bibRefCitation id="EFEA078CBF5F995DFB99FE0BFB6ED7D8" author="Gorochov A. V." box="[1038,1205,449,471]" pageId="11" pageNumber="229" pagination="187 - 215" refId="ref8677" refString="Gorochov A. V. 2004. Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 3: Podoscirtini from Madagascar and nearest regions // Zoosystematica rossica. Vol. 12. No. 2 (2003). P. 187 - 215." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
].
</paragraph>
<paragraph id="8BC47A7DBF5F995DFCC5FE11FAAAD428" blockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
Length in mm. Body:
<specimenCount id="9D7DB1F4BF5F995DFBB3FE12FBE1D7FE" box="[1060,1082,472,496]" count="1" pageId="11" pageNumber="229" type="male"></specimenCount>
22,
<specimenCount id="9D7DB1F4BF5F995DFBF4FE12FBABD7FE" box="[1123,1136,472,496]" count="1" pageId="11" pageNumber="229" type="female"></specimenCount>
1921.5; body with wings:
<specimenCount id="9D7DB1F4BF5F995DFCBBFE39FC99D405" box="[812,834,499,523]" count="1" pageId="11" pageNumber="229" type="male"></specimenCount>
35,
<specimenCount id="9D7DB1F4BF5F995DFCFAFE39FCA1D405" box="[877,890,499,523]" count="1" pageId="11" pageNumber="229" type="female"></specimenCount>
3739; pronotum:
<specimenCount id="9D7DB1F4BF5F995DFBA4FE39FB92D405" box="[1075,1097,499,523]" count="1" pageId="11" pageNumber="229" type="male"></specimenCount>
4.3,
<specimenCount id="9D7DB1F4BF5F995DFBEEFE39FB5DD405" box="[1145,1158,499,523]" count="1" pageId="11" pageNumber="229" type="female"></specimenCount>
4.44.7; tegmina:
<specimenCount id="9D7DB1F4BF5F995DFAABFE39FA89D405" box="[1340,1362,499,523]" count="1" pageId="11" pageNumber="229" type="male"></specimenCount>
25,
<specimenCount id="9D7DB1F4BF5F995DFCBBFDC4FCE2D428" box="[812,825,526,550]" count="1" pageId="11" pageNumber="229" type="female"></specimenCount>
2527; hind femora:
<specimenCount id="9D7DB1F4BF5F995DFB98FDC4FBFED428" box="[1039,1061,526,550]" count="1" pageId="11" pageNumber="229" type="male"></specimenCount>
16,
<specimenCount id="9D7DB1F4BF5F995DFBC4FDC4FBBBD428" box="[1107,1120,526,550]" count="1" pageId="11" pageNumber="229" type="female"></specimenCount>
1717.5; ovipositor 1617.
</paragraph>
</subSubSection>
<subSubSection id="C36129F6BF5F995DFCC5FDE1FBDAD4EF" pageId="11" pageNumber="229" type="discussion">
<paragraph id="8BC47A7DBF5F995DFCC5FDE1FC1ED479" blockId="11.[812,1399,315,631]" pageId="11" pageNumber="229">
REMARKS. This species, originally described from “Jlôt- Prune” (
<collectingCountry id="F36C3AEDBF5F995DFCE9FD8CFC23D452" box="[894,1016,582,604]" name="Madagascar" pageId="11" pageNumber="229">Madagascar</collectingCountry>
), is here recorded from another Madagascan locality.
</paragraph>
<paragraph id="8BC47A7DBF5F995DFCC5FD5CFBDAD4EF" blockId="11.[812,1398,662,737]" pageId="11" pageNumber="229">
<emphasis id="B90FA66FBF5F995DFCC5FD5CFBC4D4A2" bold="true" box="[850,1055,662,684]" pageId="11" pageNumber="229">Acknowledgements.</emphasis>
This study was performed in the frames of the state research project No. 1021051302540-6 (
<collectingCountry id="F36C3AEDBF5F995DFCA4FD01FC2DD4EF" box="[819,1014,715,737]" name="Russia" pageId="11" pageNumber="229">Russian Federation</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>