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<document id="9CC35DE3B704F739BF2AD4C74D4EA276" ENCODING="UTF8" ID-GBIF-Dataset="fcb01a24-34d0-451a-b2fa-90b583317051" ModsDocID="6175" checkinTime="1253089969668" checkinUser="christiana" docAuthor="Collingwood, C. A." docDate="1979" docId="8DDA1C67539F5BDC475D1FA714E00D1B" docLanguage="en" docName="6175_gg1.xml" docOrigin="Fauna Entomologica Scandinavica 8" docSource="http://antbase.org/ants/publications/6175/6175.pdf" docTitle="Lasius alienus" docType="treatment" docVersion="10" lastPageNumber="98" masterDocId="BD3B0D337E1DDAA0E4761CC6B14CB110" masterDocTitle="The Formicidae (Hymenoptera) of Fennoscandia and Denmark." masterLastPageNumber="174" masterPageNumber="1" pageNumber="97" updateTime="1701312479499" updateUser="plazi">
<mods:mods id="7F15142BE1FD5318A3425869B6553AAB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3222EBEF0DC71725EDCC1769A2CAC27C">
<mods:title id="A63495AEDFC153490604FB0A1B29CCC4">The Formicidae (Hymenoptera) of Fennoscandia and Denmark.</mods:title>
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<mods:namePart id="8320F5AB41E9094D5A91783E888E09E8">Collingwood, C. A.</mods:namePart>
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<mods:typeOfResource id="4C99F70341A786940124F59483849179">text</mods:typeOfResource>
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<mods:title id="3A2B166ACE55DB1D02E5C2EDC56625EF">Fauna Entomologica Scandinavica</mods:title>
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<mods:part id="8F503AB08EA979F5BC86BD09E72E069F">
<mods:date id="D8A563BB6C1C476016F8B267B6EEC00A">1979</mods:date>
<mods:detail id="E74B00FC1D05D4386F5B53FB602FD7DE" type="volume">
<mods:number id="8E9FE6778281FBE5C830F695D8102378">8</mods:number>
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<mods:url id="92802F847CE818FCC435E9CD8DDD6E72">http://antbase.org/ants/publications/6175/6175.pdf</mods:url>
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<mods:classification id="650669F8F8E96E99DC5AF0B761D69290">journal article</mods:classification>
<mods:identifier id="BCD1EE3F531E0296FEDD0C82C0AC58B8" type="HNS-Pub">6175</mods:identifier>
</mods:mods>
<treatment id="8DDA1C67539F5BDC475D1FA714E00D1B" ID-DOI="http://doi.org/10.5281/zenodo.6283846" ID-GBIF-Taxon="100120067" ID-Zenodo-Dep="6283846" LSID="urn:lsid:plazi:treatment:8DDA1C67539F5BDC475D1FA714E00D1B" httpUri="http://treatment.plazi.org/id/8DDA1C67539F5BDC475D1FA714E00D1B" lastPageNumber="98" pageNumber="97">
<subSubSection id="FEF2E69C3BBDCEA238C5DDBD182111B7" type="nomenclature">
<paragraph id="42B69EB11611058592B669EA892AF00E" pageNumber="97">
<pageStartToken id="53883A1A5E65D3220183A420C678C1A9" pageNumber="97">
<pageStartToken id="C44893742A48BADF258D384DA2483A5D" pageNumber="97">34</pageStartToken>
</pageStartToken>
.
<taxonomicName id="E06D29EFB8CAED04C7EAD4ED7683242B" ID-CoL="3SGW5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29670" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" lsidName-HNS="Lasius alienus (Förster)" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="alienus">Lasius alienus</taxonomicName>
(Forster, 1850)
</paragraph>
</subSubSection>
<subSubSection id="2E81CA4797E1B1A3CC0469AE50E0D072" type="description">
<paragraph id="DA1DAE5734879F5503289B2AAC3A7B73" pageNumber="97">Figs. 128-130.</paragraph>
</subSubSection>
<subSubSection id="CA4FE039EA873AC8470CD8B86C1CDF31" type="reference_group">
<paragraph id="6981EE02E908DC29804BC14A0F7E729B" pageNumber="97">
<taxonomicName id="63C43B0B80BBF7577DCFC4634FB81211" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139044" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" lsidName-HNS="Formica aliena Förster" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="aliena">Formica aliena Forster</taxonomicName>
, 1850:36.
</paragraph>
</subSubSection>
<subSubSection id="E0A07BD3B0D323F6C2322C25A0A70E3D" type="description">
<paragraph id="FD46296F079C8249AC18FD6A4708DE37" lastPageNumber="98" pageNumber="97">
Worker. Greyish yellow to brownish black. Pubescence adpressed, moderately thick over whole body and appendages. Short erect hairs scattered over dorsum and round whole occipital margin of head. Back of head convex. Ocelli indistinct or invisible;
<pageStartToken id="8C46F9972BD459EE9E735AF6FA08F20D" pageNumber="98">frontal</pageStartToken>
furrow indistinct. Erect hairs absent on scape and front tibiae, sometimes present on hind tibiae. Length: 3.0-4.2 mm.
</paragraph>
<caption id="5FB0A83CA3B20F5F0DC258C34504628E">
<paragraph id="A2AD9FBDAEA4CD5797436C521E817B75" pageNumber="97">
Figs. 128-130.
<taxonomicName id="07864CAA46BB7EFC871AE72069555CE3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29670" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" lsidName-HNS="Lasius alienus (Förster)" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="alienus">Lasius alienus (Forster)</taxonomicName>
. - 128: head and alitrunk of queen in profile; 129: male in profile; 130: head of worker in dorsal view. Scale: 1 mm.
</paragraph>
</caption>
<caption id="CF2A2A169ED4F7E9726728225504312B">
<paragraph id="EF19D38DB26F6701CA537162D257B3FC" pageNumber="97">
Figs. 131, 132.
<taxonomicName id="DAD3A989BC3971238D168E9F911F07CE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29675" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" lsidName-HNS="Lasius brunneus (Latreille)" order="Hymenoptera" pageNumber="97" phylum="Arthropoda" rank="species" species="brunneus">Lasius brunneus (Latr.)</taxonomicName>
. - 131: head of worker in dorsal view; 132: head of male in dorsal view. Scale: 1 mm.
</paragraph>
</caption>
<paragraph id="B5AFD58EA85090A579E7BA163C2D16B6" pageNumber="98">
<pageStartToken id="F22CBACFD73496AA6CE96C4BE886FB7E" pageNumber="98">Queen</pageStartToken>
. Greyish black to brownish black. Wings hyaline. Head much narrower than broad alitrunk. Pubescence and pilosity as in worker, no erect hairs on scape and front tibiae. Length: 8.0-9.0 mm.
</paragraph>
<paragraph id="A3B42067C6716B4EC80603F1ECC2301C" pageNumber="98">Male. Greyish black. Wings hyaline. No erect hairs on tibiae or scapes. Length: 3.0-3.8 mm.</paragraph>
</subSubSection>
<subSubSection id="07D3A22E73FB699F0DC272E995A17E78" type="distribution">
<paragraph id="0BE03519913FF46F12FE5D466CD9F085" pageNumber="98">Distribution. Common in Denmark and South Sweden; in Norway only recorded from. O, HE and VE; in Finland recorded as far north as Ob and Ks. - Common in South Britain, local in Ireland and Southwest Scotland. - Range: Portugal to Japan also Himalayas, North Africa to Finland. Also North America (Wilson, 1955).</paragraph>
</subSubSection>
<subSubSection id="D422CC33DAE620FB6CB6577A07113CB0" type="biology_ecology">
<paragraph id="895E6B292A4F945FFEA730828E156BA6" pageNumber="98">
Biology. This wide ranging species nests in the soil on sandy lowland heaths, dry open pasture, sea cliffs and rocky outcrops in North Europe. Its habits are mainly subterranean, feeding on the exudates of root aphids but also by scavenging and predating small insects. Workers are generally unobtrusive and non aggressive compared with
<taxonomicName id="F1E2336A15B8F0E8CFF875377C3CB468" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29723" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" lsidName-HNS="Lasius niger (Linnaeus)" order="Hymenoptera" pageNumber="98" phylum="Arthropoda" rank="species" species="niger">Lasius niger</taxonomicName>
. Nests are single queened founded by solitary fertilised queens. Mating swarms occur in August.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.80492" ID-Pensoft-Pub="1860-1324-1-45" ID-Pensoft-UUID="0A5DABFBAA6551E194C35CEA6054B588" ID-ZooBank="C402CAE814FD402AAD0554646B1493E8" LSID="urn:lsid:zoobank.org:act:E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" ModsDocID="1860-1324-69-1-45" checkinTime="1649136064731" checkinUser="pensoft" docAuthor="Morishita, Shunsuke &amp; Watanabe, Kyohei" docDate="2022" docId="8DDA9D88629C59378D2AAEA99213B44E" docLanguage="en" docName="DeutEntomolZeit 69(1): 45-64" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-04-04" docSource="http://dx.doi.org/10.3897/dez.69.80492" docTitle="Woldstedtius alpicola Morishita &amp; Watanabe 2022, sp. nov." docType="treatment" docUuid="E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" docUuidSource="ZooBank" docVersion="1" id="0A5DABFBAA6551E194C35CEA6054B588" lastPageNumber="45" masterDocId="0A5DABFBAA6551E194C35CEA6054B588" masterDocTitle="Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan" masterLastPageNumber="64" masterPageNumber="45" pageId="0" pageNumber="45" updateTime="1649136064731" updateUser="pensoft">
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<mods:title>Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan</mods:title>
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<mods:namePart>Morishita, Shunsuke</mods:namePart>
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<mods:affiliation>Hatchodori 4 - 9, Toyohashi, Aichi 440 - 0806, Japan</mods:affiliation>
<mods:nameIdentifier type="email">ezogengoroumodoki4.23@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Watanabe, Kyohei</mods:namePart>
<mods:affiliation>Kanagawa Prefectural Museum of Natural History, Iryuda 499, Odawara, Kanagawa 250 - 0031, Japan</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-04-04</mods:number>
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<treatment LSID="urn:lsid:zoobank.org:act:E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" httpUri="http://treatment.plazi.org/id/8DDA9D88629C59378D2AAEA99213B44E" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/E030CAC0-73B1-4BDC-BD63-F5E6BD3BAC4B" authority="Morishita &amp; Watanabe, 2022" authorityName="Morishita &amp; Watanabe" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius alpicola" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="alpicola" status="sp. nov.">Woldstedtius alpicola</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">Figs 1A-F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">, 9E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type series">
<paragraph pageId="0" pageNumber="45">Type series.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
: F, Japan, Honshu, Mie Pref., Inabe City, Mt. Fujiwaradake, 1 Jun 2021, S. Morishita leg. (KPMNH).
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Paratypes</emphasis>
: Japan: [Honshu] 1 F, Yamanashi Pref., Koshu City, Mt. Daibosatsu, Kaminikkawa-toge, 16 Jun 2007, K. Watanabe leg. (TMNH); 1 M, Kanagawa Pref., Yamakita Town, Mt. Oomuroyama-Mt. Kanyuudousan, 15 Jun 2008, H. Kawai leg. (KPMNH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, 13-25 Jul 2015, S. Shimizu leg. (MT) (KPMNH).
</paragraph>
<caption doi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 1.</emphasis>
<taxonomicName authorityName="Morishita &amp; Watanabe" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius alpicola" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis italics="true" pageId="0" pageNumber="45">Woldstedtius alpicola</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="45">A-D.</emphasis>
female, holotype;
<emphasis bold="true" pageId="0" pageNumber="45">E, F.</emphasis>
male, paratype) -
<emphasis bold="true" pageId="0" pageNumber="45">A, E.</emphasis>
Habitus;
<emphasis bold="true" pageId="0" pageNumber="45">B, F.</emphasis>
Head, frontal view;
<emphasis bold="true" pageId="0" pageNumber="45">C.</emphasis>
Mesonotum, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="45">D.</emphasis>
Propodeum, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">Female (n = 3). Body length 8.5-9.2 (HT: 8.5) mm, polished, coriaceous and covered with silver setae.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Head</emphasis>
0.51-0.54 (HT: 0.53)
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Clypeus 2.0-2.35 (HT: 2.0)
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high, slightly convex basally in lateral view. Face 2.1-2.4 (HT: 2.4)
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high, densely punctate, convex medially in lateral view and separated from clypeus by shallow clypeal sulcus. Inner orbits almost parallel (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1B</figureCitation>
). Length of malar space 0.9-1.15 (HT: 0.9)
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width. POL 1.9-2.1 (HT: 1.9)
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. OOL 0.9-1.18 (HT: 0.9)
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. POL 1.86-1.95 (HT: 1.86)
<normalizedToken originalValue="×">x</normalizedToken>
as long as OOL. Antenna with 24-25 (HT: 24) flagellomeres. FL I 1.29-1.35 (HT: 1.35)
<normalizedToken originalValue="×">x</normalizedToken>
as long as FL II. MP IV1.42-1.53 (HT: 1.53)
<normalizedToken originalValue="×">x</normalizedToken>
as long as MP V.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Mesosoma</emphasis>
.
</emphasis>
Lateral aspect of pronotum strigose anteriorly. Mesoscutum finely and densely punctate (separated by ca.1.0
<normalizedToken originalValue="×">x</normalizedToken>
their diameter) (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1C</figureCitation>
). Scutellum finely and sparsely punctate (separated by ca.1.5-2.5
<normalizedToken originalValue="×">x</normalizedToken>
their diameter) (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1C</figureCitation>
). Anterior part and lower half of mesopleuron coarsely and densely punctate. Speculum smooth. Sternaulus weakly impressed. Propodeum finely rugulose (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1D</figureCitation>
), rounded in lateral view and pleural carina entirely present. Fore wing length 7.4-8.5 (HT: 7.4) mm. Nervellus intercepted below middle. Hind femur 4.7-5.3 (HT: 4.7)
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Hind tibia 7.5-8.0 (HT: 8.0)
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.6: 0.4: 0.2: 0.2-0.3 (HT: 0.2).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Metasoma</emphasis>
.
</emphasis>
T I rectangular in dorsal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9E</figureCitation>
), 1.4-1.5 (HT: 1.4)
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, irregular rugulose laterally, sometimes striate between latero-median carinae. Latero-median carina present basal ca. 0.5 of T I (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9E</figureCitation>
). T II 0.6-0.73 (HT: 0.73)
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as long as maximum width, striate anteriorly and strigose laterally.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1A-D</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black. Face with a large yellow spot medially. Palpi, subtegular ridge and mesepisternum yellow. Mandible yellow, except for apex and base. Lateral aspect of pronotum with a yellow spot posteriorly. Mesoscutum with yellow shoulder marks. Tegula tinged with yellow anteriorly. Scutellum with a yellow spot apically. Wings hyaline. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Legs blackish-brown to black. Fore trochanter, trochantellus, apex of hind femur and base of hind tibia tinged with white. Fore femur, tibia and tarsomeres orange to brown. Apex of hind trochantellus and base of hind femur tinged with reddish-brown.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Male (n = 1).</emphasis>
Similar to female. Body length (excluding antennae) 8.1 mm. Face 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high. Length of malar space 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width. POL 2.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. OOL 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. POL 2.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as OOL. Inner orbits weakly divergent downward (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1F</figureCitation>
). Fore wing length 6.3 mm.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Woldstedtius alpicola sp. nov. (A-D. female, holotype; E, F. male, paratype) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure1" httpUri="https://binary.pensoft.net/fig/667921" pageId="0" pageNumber="45">1E, F</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black. Clypeus, palpi, face, ventral surface of antenna, malar space, propleuron, epicnemium, mesosternum, tegula, subtegular ridge and mesepisternum yellow. Gena tinged with yellow ventrally. Mandible yellow, except for apex. Lateral aspect of pronotum tinged with yellow ventrally and posteriorly. Mesopleuron with a large yellow marking, it enlarged anteriorly. Scutellum with a yellow spot apically. T IV to T VII with a transverse yellow band posteriorly. Wings hyaline. Veins and pterostigma brown to blackish-brown except for yellow wing base. Legs yellow to yellowish-brown. Hind coxa and trochanter each with a blackish-brown stripe dorsally. Hind trochantellus and tibia tinged with blackish-brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">Japan (Honshu).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="bionomics">
<paragraph pageId="0" pageNumber="45">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="45">Host unknown. Adults were collected in broad-leaved forest at altitudes of ca. 1,000-2,000 meters.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">The species name refers that this species inhabits alpine region.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="remarks">
<paragraph pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="45">
This species resembles
<taxonomicName genus="W." lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
but can be distinguished from the latter by the following combination of character states: pleural carina of propodeum entirely present in both sexes (absent posteriorly in both sexes of
<taxonomicName genus="W." lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
); propodeum finely rugulose in both sexes (coriaceous in both sexes of
<taxonomicName genus="W." lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
); T I 1.4-1.5
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as long as maximum width in both sexes (1.1-1.25 in females, 1.14-1.26 in males of
<taxonomicName genus="W." lsidName="W. kuroashii" pageId="0" pageNumber="45" rank="species" species="kuroashii">
<emphasis italics="true" pageId="0" pageNumber="45">W. kuroashii</emphasis>
</taxonomicName>
). This species also resembles a Korean species,
<taxonomicName genus="W." lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
Balueva &amp; Lee, 2016 (male is unknown), but can be distinguished by the following combination of character states in females: a large yellow median spot of face present (absent in
<taxonomicName genus="W." lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); antenna with 24-25 flagellomeres (22-23 in
<taxonomicName genus="W." lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); scutellum finely and sparsely punctate on coriaceous background (entirely coriaceous in
<taxonomicName genus="W." lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
); propodeum finely rugulose (coriaceous in
<taxonomicName genus="W." lsidName="W. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">W. pallidus</emphasis>
</taxonomicName>
).
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55
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<paragraph id="5F5B8C200E6E62DE6CE8E0DC1328AE91" pageNumber="24">Operaria: Long. 4 mm. Castanea, abdomine fusco, abstante albido-pilosa, haud pubescens; mandibulae striolatae, apice sublaeves, prope marginem masticatorium serie punctorum; clypeus deplanatus longitudinaliter rugosus, medio laevigatus, tenuissime coriaceo-rugulosus, margine antico paulo emarginata; laminae frontales angustissimae, antice latiores; funiculi clava distincte triarticulata; frons, vertex antice et capitis latera haud dense rude et longitudinaliter rugosa, vertex postice rude reticulato-rugosus; thorax rude longitudinaliter rugosus, sutura meso-metanotali paulo impressa; pronotum postice utrimque obtuse angulatum; metanotum spinis duabus longis, parte metanoti basali fere duplo longioribus, oblique sursum et postice versus directis; petioli segmentum primum nitidum antice tenuiter petiolatum et sublaeve, postice nodo elevato transverso parte antica petiolata breviori, longitudinaliter rugoso et subtilissime coriaceo-ruguloso, segmentum secundum, subglobosum, sublaeve et nitidum.; abdomen nitidum et laevissimum; pedes pilis abstantibus, albicantibus.</paragraph>
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<paragraph id="27FC9B84225F91B52CC2CDAF60F754C2" pageNumber="24">Von der Goldkueste in Afrika.</paragraph>
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<paragraph id="A3F465978286B0796B4B5D1B9FD34AF1" pageNumber="24">
Diese Art hat habituell viele Aehnlichkeit mit
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, unterscheidet sich aber, ausser dem generischen Charakter, besonders durch den flachen Clypeus, durch die langen, duennen, nadeifoermigen Dornen des Metanotum und durch den ziemlich langen, duennen stielfoermigen vorderen Theil des ersten Stielchensegmentes.
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<taxonomicName id="C3B5502844B411F3C168A3AB4CF383A9" authorityName="Casey" authorityYear="1909" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="844" pageNumber="845" phylum="Arthropoda" rank="species" species="castanea">Pseudomorpha castanea</taxonomicName>
Casey, 1909: 278. Type locality: &quot;Stockton [Tooele County], Utah&quot; (original citation). Syntype(s) in USNM.
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<subSubSection id="C55DCEC0823704739770BE25DA1AA83F" pageId="844" pageNumber="845" type="distribution">
<paragraph id="07C96A67736AE5E4406854D478710084" pageId="844" pageNumber="845">Distribution.</paragraph>
<paragraph id="BE32FFD83B52204385A601F26B7F6A5B" pageId="844" pageNumber="845">This species is known from southeastern Oregon (Hatch 1953: 187), Sacramento County in California, and northwestern Utah (Notman 1925: 22).</paragraph>
</subSubSection>
<subSubSection id="526CA7F465C3B94FE889CD41BD13BFED" pageId="844" pageNumber="845" type="records">
<paragraph id="164CAE242F56266E276AC457E910255A" pageId="844" pageNumber="845">Records.</paragraph>
<paragraph id="9058EFEACB82EF52051031F4045EDEC6" pageId="844" pageNumber="845">
<emphasis id="544B721DBB8B67D698423F809BFAA6F0" bold="true" pageId="844" pageNumber="845">USA</emphasis>
: CA, OR, UT
</paragraph>
</subSubSection>
</treatment>
</document>