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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="8D8A109AA8895DA1BA59C34BA7990A55" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Metasarcus limachii Benedetti &amp; Pinto-da-Rocha 2022, sp. nov." docType="treatment" docUuid="DD6B9FC9-7490-4502-8EA9-F4CC949918C6" docUuidSource="ZooBank" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>309</mods:start>
<mods:end>388</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:DD6B9FC9-7490-4502-8EA9-F4CC949918C6" httpUri="http://treatment.plazi.org/id/8D8A109AA8895DA1BA59C34BA7990A55" lastPageNumber="309" pageId="0" pageNumber="309">
<subSubSection pageId="0" pageNumber="309" type="nomenclature">
<paragraph pageId="0" pageNumber="309">
3.46.
<taxonomicName LSID="http://zoobank.org/DD6B9FC9-7490-4502-8EA9-F4CC949918C6" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii" status="sp. nov.">Metasarcus limachii</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5G</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10K, L</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">, 14G, H</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">, 26A, B</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">Description.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 6.1; DSL: 6.8; CL: 1.4. FIVL: 14.2. ChL: 2.4.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">26A</figureCitation>
) Yellow background with black spots covering virtually the entire DS; scutal area III, pedipalpus, chelicerae and legs I-III brown. Leg IV black.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5G</figureCitation>
) Gamma-P-type DSS, with strongly concave posterior margin of DS. Median elevation of anterior margin of DS with some scattered granules. Ocularium with median depression well marked with large and small granules around the eyes, and in lower concentrations in depression. Carapace with sparse granules. Areas I-IV with small-scattered granules. Area I undivided; with a pair of median tubercles. Areas II unarmed. Area III with a median pair of spines. Area IV with a pair of median tubercles. Lateral margins of DS with a denser concentration of granules. Posterior margin of DS and free tergites I-III with a row of granules. Free tergite III with a long spiniform apophysis, curved laterally and with a ventral short and sharp projection (This curvature is probably an individual anomaly. Furthermore, the first author has already had the opportunity to examine a photograph taken by Dr. Arthur Anker of a live male specimen whose apophysis was straight).
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5G</figureCitation>
) Not swollen. Segment I smooth. Segment II with granules concentrated near the fixed finger; finger with three teeth. Segment III with four teeth.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Trochanter with a ventroapical setiferous tubercle. Femur with two small ventrobasal spines; a ventral row of large spines and a proapical spine. Patella with a proapical spine. Tibia retrolateral iiiIi, prolateral IiIi. Tarsus: retrolateral iIiI, prolateral IiIi/iIiIi.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a median row of 5-6 small setiferous tubercles near the apex. Coxae II-IV with sparse granules. Coxa IV with the lower granules. Genital area with few granules. Free sternites with rows of larger granules. Anal operculum with granules, concentrated in the apical part.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5G</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10K, L</figureCitation>
) Coxa I-II each one with a retrolateral and a prolateral apophysis. Coxa III with dorsal small tubercles. Coxa IV with sparse granules; with 4-5 dorsoapical tubercles; a retrolateral short apophysis, with a tubercle at the base of apophysis, and a retolateral tubercle located more ventrally than apophysis. Trochanters I-III with sparse granules. Trochanter IV with a small probasal spiniform apophysis; a retrobasal and large retroapical spiniform apophyses. Femora I-III unarmed, granular. Femur IV slightly sigmoid, with a large dorsoapical tubercle; granular. Patellae I-IV unarmed. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) 8, 16, 10, 11.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14G, H</figureCitation>
) VP rectangular, with distal margin straight; straight in lateral view. MS C1-C3 apical long and straight; MS A1 median long and slightly curved; MS B1 sub basal long and slightly curved. Lateral sacs short, apically tapered, with short T3-like microsetae. Stylus with inflated apex, with an elongated ventral projection and tiny projections in the apical portion. Dorsal process long. Promontory convex. -
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) DSW: 5.6-5.7; DSL: 5.3-5.7; CL: 1.1-1.2. FIVL: 11.2-14.0. ChL: 2.0-2.3. (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">26B</figureCitation>
) Areas I-II with three pairs of small tubercles. Area III with a paramedian pair of small spines. Trochanter IV unarmed. Apophysis of free tergite III shorter and without ventral projection. Coloration with less brown spots, highlighting the most yellow body, resulting in a spotty pattern. Area IV with brown pigmentation more obvious. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) 7-8, 13-15, 9-11, 7-10.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
It differs from other species of the genus because by having area I undivided; long spiniform apophysis on free tergite III (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5G</figureCitation>
); trochanter IV with a probasal, a retrobasal and a retroapical apophyses (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10K, L</figureCitation>
); DS with yellow background with black spots (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">26A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="etymology">
<paragraph pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="309">
The specific epithet of masculine gender, in the genitive form, dedicated Miguel Limachi, from
<normalizedToken originalValue="Coleccíon">Coleccion</normalizedToken>
Boliviana de Fauna (CBF), who provided invaluable help with all formalities for collecting in Bolivia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
) BOLIVIA. La Paz. La Paz-Coroico; Yanacachi.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation collectingDate="2010-12-12" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, A. Saravia" country="BOLIVIA" latitude="-16.398472" location="Yanacachi" longLatPrecision="1" longitude="-67.73655" specimenCount="♂" stateProvince="La Paz" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
, '
<collectingCountry name="Bolivia">BOLIVIA</collectingCountry>
,
<collectingRegion country="Bolivia" name="La Paz">La Paz</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:8D8A109AA8895DA1BA59C34BA7990A55:71005DDF6BF24A37C64A822FE1AB12F5" country="BOLIVIA" latitude="-16.398472" longLatPrecision="1" longitude="-67.73655" name="Yanacachi" stateProvince="La Paz">Yanacachi</location>
,
<geoCoordinate degrees="16" direction="south" minutes="23" orientation="latitude" precision="1" seconds="54.5" value="-16.398472">16°23′54.5″S</geoCoordinate>
<geoCoordinate degrees="67" direction="west" minutes="44" orientation="longitude" precision="1" seconds="11.6" value="-67.73655">67°44′11.6″W</geoCoordinate>
,
<collectingDate value="2010-12-12">12/XII/2010</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
,
<collectorName>A. Benedetti</collectorName>
,
<collectorName>J. Ochoa</collectorName>
&amp;
<collectorName>A. Saravia</collectorName>
leg. (CBF)
</materialsCitation>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
,
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36991).
<materialsCitation collectingDate="2010-11-30" collectorName="La Paz-Coroico, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, A. Saravia" country="BOLIVIA" latitude="-16.229168" location="La Paz-Coroico" longLatPrecision="15" longitude="-67.82142" specimenCount="♀" stateProvince="La Paz" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
, '
<collectingCountry name="Bolivia">BOLIVIA</collectingCountry>
,
<collectingRegion country="Bolivia" name="La Paz">La Paz</collectingRegion>
, road
<collectorName>
<location LSID="urn:lsid:plazi:treatment:8D8A109AA8895DA1BA59C34BA7990A55:9E06BAE13B747AAD3C6BBA7EB3DD29F6" country="BOLIVIA" latitude="-16.229168" longLatPrecision="15" longitude="-67.82142" name="La Paz-Coroico" stateProvince="La Paz">La Paz-Coroico</location>
</collectorName>
,
<geoCoordinate degrees="16" direction="south" minutes="13" orientation="latitude" precision="15" seconds="45" value="-16.229168">16°13′45″S</geoCoordinate>
<geoCoordinate degrees="67" direction="west" minutes="49" orientation="longitude" precision="1" seconds="17.1" value="-67.82142">67°49′17.1″W</geoCoordinate>
,
<collectingDate value="2010-11-30">30/XI/2010</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
,
<collectorName>A. Benedetti</collectorName>
,
<collectorName>J. Ochoa</collectorName>
&amp;
<collectorName>A. Saravia</collectorName>
leg. (MZSP 36992)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.413.7172" ID-GBIF-Dataset="f45c1ebe-039d-4180-bfd0-a9d772f72281" ID-PMC="PMC4086027" ID-Pensoft-Pub="1313-2970-413-1" ID-PubMed="25009414" ID-ZBK="5791CE9C1CC0472095838A585DA79446" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-413-1" ModsDocOrigin="ZooKeys 413" ModsDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" checkinTime="1451245833965" checkinUser="pensoft" docAuthor="Hita Garcia, Francisco &amp; Fisher, Brian L." docDate="2014" docId="8D8A1AE994B25BFF296400E74544F3CB" docLanguage="en" docName="ZooKeys 413: 1-170" docOrigin="ZooKeys 413" docSource="http://dx.doi.org/10.3897/zookeys.413.7172" docTitle="Tetramorium myrmidon Hita Garcia &amp; Fisher, sp. n." docType="treatment" docUuid="11BD4CF2-149D-4A6A-9B9C-C59C0030B466" docUuidSource="ZooBank" docVersion="4" lastPageNumber="66" masterDocId="FFFAFFE7FFB3C93AB161C804401EFFC2" masterDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" masterLastPageNumber="170" masterPageNumber="1" pageNumber="64" updateTime="1668158573061" updateUser="ExternalLinkService">
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<mods:title>The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hita Garcia, Francisco</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>413</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.413.7172</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.413.7172</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-413-1</mods:identifier>
<mods:identifier type="ZBK">5791CE9C1CC0472095838A585DA79446</mods:identifier>
<mods:identifier type="ZooBank">5791CE9C1CC0472095838A585DA79446</mods:identifier>
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<treatment ID-GBIF-Taxon="152053299" LSID="urn:lsid:zoobank.org:act:11BD4CF2-149D-4A6A-9B9C-C59C0030B466" httpUri="http://treatment.plazi.org/id/8D8A1AE994B25BFF296400E74544F3CB" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
<subSubSection pageId="63" pageNumber="64" type="nomenclature">
<paragraph pageId="63" pageNumber="64">
<taxonomicName LSID="http://zoobank.org/11BD4CF2-149D-4A6A-9B9C-C59C0030B466" authority="Hita Garcia &amp; Fisher" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon Hita Garcia &amp; Fisher</taxonomicName>
<taxonomicNameLabel pageId="63" pageNumber="64">sp. n.</taxonomicNameLabel>
Figs 23C, 25B, 26C, 36, 64
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="type material">
<paragraph pageId="63" pageNumber="64">Type material.</paragraph>
<paragraph pageId="63" pageNumber="64">
Holotype, pinned worker, MADAGASCAR, Toliara,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken originalValue="d’Ambohijanahary">d'Ambohijanahary</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
, 35.2 km 312° NW Ambaravaranala,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.26667">18.26667°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.40667">45.40667°E</geoCoordinate>
, 1050 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF07020, 13.-17.I.2003 (B.L. Fisher et al.) (CAS: CASENT0028635). Paratypes, two workers with same data as holotype (CAS: CASENT0028642; CASENT0028643); and three workers MADAGASCAR, Toliara,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken originalValue="d’Ambohijanahary">d'Ambohijanahary</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
, 34.6 km 314° NW Ambaravaranala,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-18.26">18.26°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.41833">45.41833°E</geoCoordinate>
, 1100 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF07086, 16.I.2003 (B.L. Fisher et al.) (CAS: CASENT0029810; CASENT0029821; CASENT0030099).
</paragraph>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 36.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
holotype worker (CASENT0028635). A Body in profile B Body in dorsal view C Head in full-face view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="diagnosis">
<paragraph pageId="64" pageNumber="65">
<pageBreakToken pageId="64" pageNumber="65" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="64" pageNumber="65">
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
can be easily distinguished from the other members of the group by the following combination of characters: eyes relatively large (OI 24-25); antennal scapes short (SI 74-76); frontal carinae well developed, noticeably raised, ending shortly before posterior head margin; petiolar node high rounded nodiform, in profile around 1.7 times higher than long (LPeI 58-60), in dorsal view around 1.2 to 1.3 times wider than long (DPeI 121-129); in dorsal view postpetiole around 1.3 to 1.5 times broader than petiolar node (PPI 133-141); dorsum of mesosoma with two pairs of long, standing hairs on pronotum and mesonotum.
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="worker measurements">
<paragraph pageId="64" pageNumber="65">Worker measurements</paragraph>
<paragraph pageId="64" pageNumber="65">(N=6). HL 0.70-0.76 (0.73); HW 0.62-0.69 (0.65); SL 0.47-0.52 (0.49); EL 0.16-0.18 (0.16); PH 0.33-0.37 (0.35); PW 0.45-0.50 (0.47); WL 0.85-0.94 (0.89); PSL 0.13-0.16 (0.14); PTL 0.14-0.16 (0.15); PTH 0.24-0.27 (0.25); PTW 0.18-0.20 (0.19); PPL 0.18-0.21 (0.19); PPH 0.24-0.28 (0.26); PPW 0.24-0.28 (0.26); CI 88-91 (90); SI 74-76 (75); OI 24-25 (25); DMI 52-54 (53); LMI 38-40 (39); PSLI 18-20 (19); PeNI 39-40 (39); LPeI 58-60 (59); DPeI 121-129 (125); PpNI 52-55 (54); LPpI 73-77 (76); DPpI 129-136 (133); PPI 133-141 (138).</paragraph>
</subSubSection>
<subSubSection lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="worker description">
<paragraph pageId="64" pageNumber="65">Worker description.</paragraph>
<paragraph lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
Head much longer than wide (CI 88-91); in full-face view posterior head margin weakly concave. Anterior clypeal margin with distinct median impression. Frontal carinae well developed, noticeably raised, diverging posteriorly, merging with surrounding sculpture shortly before posterior head margin. Antennal scrobes very weakly developed, almost absent, shallow and without clear and distinct posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 74-76). Eyes relatively large (OI 24-25). Mesosomal outline in profile flat to weakly convex, moderately low and long (LMI 38-40), moderately marginate from lateral to dorsal mesosoma; promesonotal suture absent; metanotal groove present, but weakly developed. Propodeal spines short to moderate, elongate-triangular, and acute (PSLI 18-20), propodeal lobes triangular and short, always much shorter than propodeal spines. Petiolar node in profile high nodiform with well rounded margins, around 1.7 times higher than long (LPeI 58-60), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, anterodorsal margin slightly more angulate than posterodorsal, more rounded margin, petiolar dorsum relatively flat to weakly convex; node in dorsal view 1.2 to 1.3 times wider than long (DPeI 121-129), in dorsal view pronotum between 2.5 to 2.6 times wider than petiolar node (PeNI 39-40). Postpetiole in profile globular, between 1.3 to 1.4 times higher than long (LPpI 73-77); in dorsal view around 1.3 to 1.4 times wider than long (DPpI 129-136), pronotum around 1.8 to 1.9 times wider than postpetiole (PpNI 52-55). Postpetiole in profile appearing more voluminous than petiolar node, postpetiole in dorsal view around 1.3 to 1.4 times wider than petiolar node (PPI 133-141). Mandibles completely unsculptured, smooth, and shiny; clypeus irregularly longitudinally rugulose with three to seven rugulae, median ruga usually fully developed, one to three mostly unbroken lateral rugulae present on each side; cephalic dorsum between frontal carinae longitudinally rugulose with seven to nine rugulae, rugulae usually running from posterior clypeal margin to posterior head margin, often interrupted or with cross-meshes, especially posteriorly; scrobal area mostly unsculptured
<pageBreakToken pageId="65" pageNumber="66" start="start">with</pageBreakToken>
ground sculpture only, lateral head reticulate-rugose to longitudinally rugulose. Ground sculpture on head well developed and distinct, mostly reticulate-punctate. Dorsum and sides of mesosoma irregularly longitudinally rugulose to reticulate-rugulose. Forecoxae dorsally weakly rugulose, but mostly unsculptured, smooth, and shining. Ground sculpture on mesosoma weakly to moderately punctate. Both waist segments and gaster fully unsculptured, smooth, and shining. Dorsum of head with several pairs of standing, long, fine hairs; dorsum of mesosoma with two pairs only, one on anterior pronotum and one on anterior mesonotum; propodeum, waist segments, and first gastral tergite without any standing hairs; first gastral tergite with very short, scarce, appressed pubescence. Anterior edges of antennal scapes and dorsal (outer) surfaces of hind tibiae with appressed hairs. Head, mesosoma, waist segments, and gaster uniformly brown to dark brown, appendages yellowish to light brown.
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="66" type="etymology">
<paragraph pageId="65" pageNumber="66">Etymology.</paragraph>
<paragraph pageId="65" pageNumber="66">
The name of the new species is inspired by the ancient Greek
<normalizedToken originalValue="“myrmidons”">&quot;myrmidons&quot;</normalizedToken>
, which were skilled warriors and known as the legendary
<normalizedToken originalValue="“ant-people”">&quot;ant-people&quot;</normalizedToken>
who inhabited the Greek island of Aegina. The species epithet is a noun in apposition, and thus invariant.
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="66" type="distribution and biology">
<paragraph pageId="65" pageNumber="66">Distribution and biology.</paragraph>
<paragraph pageId="65" pageNumber="66">
The new species is so far only known from Ambohijanahary, which is an isolated montane rainforest located in the midwest of Madagascar (Fig. 64), where it was collected only twice. With only six known specimens it represents an extremely rare species.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
was collected from leaf litter at elevations of 1050 to 1100 m.
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="66" type="discussion">
<paragraph pageId="65" pageNumber="66">Discussion.</paragraph>
<paragraph pageId="65" pageNumber="66">
Based on its larger body size and very well developed frontal carinae,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
differs strongly from the smaller species
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium aspis" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="aspis">Tetramorium aspis</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium camelliae" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="camelliae">Tetramorium camelliae</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium cognatum" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="cognatum">Tetramorium cognatum</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium karthala" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="karthala">Tetramorium karthala</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium rumo" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rumo">Tetramorium rumo</taxonomicName>
, all with more weakly developed frontal carinae, while being presumably morphologically closer to
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium gladius" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="gladius">Tetramorium gladius</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
. Of these,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium gladius" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="gladius">Tetramorium gladius</taxonomicName>
possesses very small eyes (OI 19-20), while the eyes of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
are much larger (OI 24-25).
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
has shorter antennal scapes (SI 66-70) and a thinner petiolar node, in profile 1.8 to 2.2 times higher than long (LPeI 45-54), in contrast to
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
with its longer scapes (SI 74-76) and lower and thicker petiolar node, which is in profile around 1.7 times higher than long (LPeI 58-60). The widespread
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
, however, appears to be the species morphologically closest to
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
and most of their morphometric ranges and characters overlap. Nonetheless, we consider both sufficiently demarcated from each other since they are found to co-occur in sympatry in Ambohijanahary. The specimens of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
from this locality differ from
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
by having five to six pairs of long, standing hairs on the mesosoma and a generally thinner and higher petiolar node. Finally,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
is unlikely to be confused with the last species of the complex,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium freya" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="freya">Tetramorium freya</taxonomicName>
, since the latter lacks long, standing pilosity on the mesosomal dorsum (present in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
).
</paragraph>
<paragraph pageId="65" pageNumber="66">The small number of specimens from just two collection events in the same locality does not permit proper assessment of intraspecific variation.</paragraph>
</subSubSection>
</treatment>
</document>