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<mods:titleInfo id="F0C26992AD91AE9D1476C173F2229B9F">
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<mods:title id="B282519ECE5A1EB4920C098FC46361B5">Pheidole in the New World. A dominant, hyperdiverse ant genus.</mods:title>
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<mods:namePart id="458A5FCE96009F2D0685A409F8776403">Wilson, E. O.</mods:namePart>
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<mods:typeOfResource id="EF2B5D5CF46E2335F15B7ED01AFA8E8B">text</mods:typeOfResource>
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<mods:dateIssued id="4070070EBD15B55A77E30DBD1616BDF4">2003</mods:dateIssued>
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<mods:publisher id="860F900EF6642706B1FC2DAB142A7268">Harvard University Press</mods:publisher>
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<mods:classification id="BA5D2A1AE0924D9F8C09CAA5EE8A1903">book</mods:classification>
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<mods:identifier id="0A0825D99B6787DC827E4BEAE0420021" type="HNS-Pub">20017</mods:identifier>
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<treatment id="8D051A50A45BAD13E73E93090E6B3730" ID-DOI="http://doi.org/10.5281/zenodo.3507597" ID-GBIF-Taxon="100119732" ID-Zenodo-Dep="3507597" LSID="urn:lsid:plazi:treatment:8D051A50A45BAD13E73E93090E6B3730" httpUri="http://treatment.plazi.org/id/8D051A50A45BAD13E73E93090E6B3730" lastPageNumber="512" pageNumber="512">
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<subSubSection id="47BEC9F9240E184F9F34ACFE2E13740F" type="nomenclature">
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<paragraph id="0EC11ED5074EBDC207F17D79C1E12455" pageNumber="512">
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<taxonomicName id="B537F002C20F452A33F5687D16B73676" ID-CoL="4FYM3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">Pheidole striaticeps</taxonomicName>
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Mayr
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</paragraph>
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</subSubSection>
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<subSubSection id="7973B05CEBFFC7D3D446484D50D42E14" type="reference_group">
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<paragraph id="DC025EBB2C2F2D9C4949A23A2B613CAF" pageNumber="512">
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<taxonomicName id="A0823F93A6C10594D1D7ED56F6852675" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">Pheidole striaticeps</taxonomicName>
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Mayr 1870b: 987.
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</paragraph>
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</subSubSection>
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<subSubSection id="F13D80F975A20B45F19E2C0E7CFB443D" type="nomenclature">
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<paragraph id="DE852A0BF53BC99767BE59A3BA2633A3" pageNumber="512">Types Naturhist. Mus. Wien.</paragraph>
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</subSubSection>
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<subSubSection id="B2DDF6C0EE300C654EAC6C8989B50649" type="etymology">
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<paragraph id="D4D80115E51E11AB3E63EEA9B9C10136" pageNumber="512">
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Etymology L
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||||
<taxonomicName id="2FC858A71960AB5C89C46C8C28753D65" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">striaticeps</taxonomicName>
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, lined (striate) head.
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</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection id="5C2CE9A7FBD4C2E0F0543C60D634BA6E" type="diagnosis">
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||||
<paragraph id="00E3543B26ACEF2E5D235DEE154AB452" pageNumber="512">
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||||
Diagnosis A member of the
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<taxonomicName id="261E953C1F370E8F8C127466401B4D78" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33744" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavens Roger" lsidName-HNS="Pheidole flavens Roger" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="flavens">flavens</taxonomicName>
|
||||
group close to
|
||||
<taxonomicName id="7323A15F6AB71504AE451B1E9EDBDDBC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181859" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole chloe Forel" lsidName-HNS="Pheidole chloe Forel" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="chloe">chloe</taxonomicName>
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||||
, less so to the other species listed in the heading, distinguished as follows. Major: antennal scrobes present; entire dorsal surface of head covered by longitudinal carinulae, which curve inwardly toward midline at occiput; humerus prominent and rounded in dorsal-oblique view; mesonotal convexity present, especially in dorsal-oblique view; head wider than long (Head Width/Head Length 1.03); pronotum transversely carinulate. Minor: carinulae absent from around thorax-propodeal suture.
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="8FC353B9BF3B805266E7162365C677B1" type="description">
|
||||
<paragraph id="22A250B0597922AFB9331FC4F4DE74FF" pageNumber="512">Measurements (mm) Lectotype major: HW 1.06, HL 1.02, SL 0.56, EL 0.12, PW 0.50. Paralectotype minor: HW 0.62, HL 0.62, SL 0.56, EL 0.08, PW 0.42. Color Major: concolorous light reddish brown. Minor: concolorous light reddish brown.</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection id="253CBE4CC0AB3510C0B553E9C22C1AE7" type="distribution">
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||||
<paragraph id="86AFF2E78175E2284CDD7A649786A7EA" pageNumber="512">
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||||
Range Mexico (almost certainly tropical Mexico), Honduras, Costa Rica, and Amazonian Peru. Longino (1997) does not distinguish this species from
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||||
<taxonomicName id="564A7B4FA8CFBAF9B8C9B1160CD85B9A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181859" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole chloe Forel" lsidName-HNS="Pheidole chloe Forel" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="chloe">chloe</taxonomicName>
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||||
, although the two are different. He reports "
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||||
<taxonomicName id="EEE1193907EAC6169862370167F54981" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">striaticeps</taxonomicName>
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, " which may include both species, from the Atlantic slope to 500 m and Pacific lowlands of Costa Rica.
|
||||
</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="DBDF9B2A8A271A8713AC59394AE2083F" type="biology_ecology">
|
||||
<paragraph id="7127AEB92604374B096AC2BBD9B18DBB" pageNumber="512">
|
||||
Biology Longino (1997) reports "
|
||||
<taxonomicName id="4EAE0E373218888551F8B02D40DAC983" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">striaticeps</taxonomicName>
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||||
, " which may include both true
|
||||
<taxonomicName id="48DEAABD270ECB49BB75C876C515CE49" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34028" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole striaticeps Mayr" lsidName-HNS="Pheidole striaticeps Mayr" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="striaticeps">striaticeps</taxonomicName>
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||||
and
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||||
<taxonomicName id="6EBFB39F17A2410E98279B46C5BF9B8A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181859" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole chloe Forel" lsidName-HNS="Pheidole chloe Forel" order="Hymenoptera" pageNumber="512" phylum="Arthropoda" rank="species" species="chloe">chloe</taxonomicName>
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||||
, from mature rainforest, nesting under the bark of rotting stumps and logs. At Cuzco Amazonico, near Puerto Maldonado, Stefan Cover and John E. Tobin found a colony in a gap within mature rainforest, beneath the bark of a rotten but still hard log.
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</paragraph>
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</subSubSection>
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<subSubSection id="26DE9C3E67A7E3C4F82B06795ABFD855" type="materials_examined">
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<paragraph id="9C0081438B0E0B8EE765036776EF1035" pageNumber="512">Figure Upper: lectotype, major. Lower: paralectotype, minor. MEXICO. Scale bars = 1 mm.</paragraph>
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</subSubSection>
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||||
</treatment>
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||||
</document>
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||||
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data/8D/05/A7/8D05A782189D587D80604D432D27B950.xml
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data/8D/05/A7/8D05A782189D587D80604D432D27B950.xml
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<mods:titleInfo>
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<mods:title>Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Giron, Jennifer C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, Purdue University, West Lafayette, IN 47907, USA & Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
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<mods:affiliation>Department of Ecology and Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
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<mods:nameIdentifier type="email">aezshort@ku.edu</mods:nameIdentifier>
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<mods:date>2021</mods:date>
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<mods:number>2021-02-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1019</mods:number>
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<treatment ID-GBIF-Taxon="179905044" LSID="urn:lsid:zoobank.org:act:A8163D90-1FF7-4273-854A-BDBB330DBB0E" httpUri="http://treatment.plazi.org/id/8D05A782189D587D80604D432D27B950" lastPageNumber="93" pageId="0" pageNumber="93">
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<taxonomicName LSID="http://zoobank.org/A8163D90-1FF7-4273-854A-BDBB330DBB0E" authority="Girón & Short, 2021" authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares benettii" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="benettii" status="sp. nov.">Tobochares benettii</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="93">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">Figs 4A-C</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">, 11A</figureCitation>
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Tobochares spp." figureDoi="10.3897/zookeys.1019.59881.figure13" httpUri="https://binary.pensoft.net/fig/512611" pageId="0" pageNumber="93">, 13</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Tobochares habitat in Brazil A, B type locality and habitat for T. benettii, seepage near Rio Preto de Eva (collecting event BR 17 - 0610 - 01 A) C type locality and habitat for T. goias, margin of Balneario Lejas (collecting event BR 18 - 0304 - 02 B) D habitat of T. kusad and T. sipaliwini, State of Roraima, near Usina de Jatapu reservoir (collecting Event BR 18 - 0117 - 01 A) E habitat and type locality of T. romanoae, and habitat of T. sipaliwini, State of Roraima, Serra do Tepequem, Igarape Preto Negro, Cachoeira Leje Preta (collecting event BR 18 - 0114 - 04 B) F type locality and habitat of T. fusus, State of Amapa, Calcoene (collecting event BR 18 - 0721 - 02 B)." figureDoi="10.3897/zookeys.1019.59881.figure14" httpUri="https://binary.pensoft.net/fig/512612" pageId="0" pageNumber="93">, 14A, B</figureCitation>
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</paragraph>
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="93" type="reference_group">
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<paragraph pageId="0" pageNumber="93">
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||||
<taxonomicName authority="Girón & Short, 2021" authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus" status="sp. nov.">Tobochares</taxonomicName>
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||||
sp. B:
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||||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="RAP Bulletin of Biological Assessment 63, Conservation International, Arlington" pageId="0" pageNumber="93" refId="B9" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology. https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology." url="https://doi.org/10.1111/syen.12467" year="2021">Short et al. (2021)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="type material">
|
||||
<paragraph pageId="0" pageNumber="93">Type material examined.</paragraph>
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||||
<paragraph pageId="0" pageNumber="93">
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Holotype</emphasis>
|
||||
(male)
|
||||
</emphasis>
|
||||
: "Brazil: Amazonas: Rio Preto da Eva;
|
||||
<geoCoordinate degrees="2.678466" direction="south" orientation="latitude" precision="1" value="-2.678466">-2.678466</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="59.401714" direction="west" orientation="longitude" precision="1" value="-59.401714">-59.401714</geoCoordinate>
|
||||
, 25 m; ca. 32 Km W of Rio Preto da Eva; seepage on sandstone with algae; 10.vi.2017; leg. Benetti and team; BR17-0610-01A" (INPA).
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Paratypes</emphasis>
|
||||
(8 exs.): Brazil: Amazonas
|
||||
</emphasis>
|
||||
: Same data as holotype (8, INPA, SEMC including DNA voucher SLE1264).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="differential diagnosis">
|
||||
<paragraph pageId="0" pageNumber="93">Differential diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares benettii" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="benettii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Tobochares benettii</emphasis>
|
||||
</taxonomicName>
|
||||
can be easily recognized from all other
|
||||
<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
species in the
|
||||
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares sulcatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="sulcatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Tobochares sulcatus</emphasis>
|
||||
</taxonomicName>
|
||||
species group by its elytral punctures seemingly uniformly distributed, not forming clear longitudinal rows, therefore completely lacking elytral striae (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4A</figureCitation>
|
||||
). This character makes it similar to
|
||||
<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares goias" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="goias">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Tobochares goias</emphasis>
|
||||
</taxonomicName>
|
||||
, from which it can be distinguished by its uniform orange brown coloration along pronotum and elytra, with dark brown head (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4A-C</figureCitation>
|
||||
), the posterior elevation of the mesoventrite forming a curved transverse ridge, which is medially prominent and acute and by characters of the aedeagus (Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">11A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93" start="Figure 4" startId="F4">
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">Figure 4.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
spp. in the
|
||||
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares sulcatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="sulcatus">Tobochares sulcatus</taxonomicName>
|
||||
species group
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">A-C</emphasis>
|
||||
<taxonomicName lsidName="T. benettii" pageId="0" pageNumber="93" rank="species" species="benettii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">T. benettii</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">A</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">B</emphasis>
|
||||
lateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">C</emphasis>
|
||||
ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">D-F</emphasis>
|
||||
<taxonomicName lsidName="T. goias" pageId="0" pageNumber="93" rank="species" species="goias">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">T. goias</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">D</emphasis>
|
||||
dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">E</emphasis>
|
||||
lateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">F</emphasis>
|
||||
ventral view. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="description">
|
||||
<paragraph pageId="0" pageNumber="93">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Size and form</emphasis>
|
||||
: Body length 1.6-2.0 mm. Body elongate oval, strongly convex (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4A, B</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="93">
|
||||
<emphasis italics="true" pageId="0" pageNumber="93">Color and punctation</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head dark brown, pronotum and elytra uniformly orange brown; antennae, mouthparts and legs orange to yellowish brown, with paler tarsi (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4C</figureCitation>
|
||||
). Ground punctation on head, pronotum and elytra moderately marked.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Head</emphasis>
|
||||
: Eyes in dorsal view with anterior margin oblique, posteriorly directed, and outer margins continuous with outline of head (as in Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2A</figureCitation>
|
||||
); in lateral view, eyes emarginate to about half the length of eye (as in Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2B</figureCitation>
|
||||
). Maxillary palps slender, nearly as long as the width of the head, uniformly yellow in color (as in Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2A</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Thorax</emphasis>
|
||||
: Elytra with slightly defined rows of punctures, not forming grooves (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4A</figureCitation>
|
||||
). Elevation of mesoventrite forming a somewhat transverse bulge (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4C</figureCitation>
|
||||
). Metaventrite with distinct median, broad, ovoid glabrous area extending along posterior two thirds (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Tobochares spp. in the Tobochares sulcatus species group A-C T. benettii A dorsal view B lateral view C ventral view D-F T. goias D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure4" httpUri="https://binary.pensoft.net/fig/512602" pageId="0" pageNumber="93">4C</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Abdomen</emphasis>
|
||||
: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">11A</figureCitation>
|
||||
) with basal piece 0.25
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
the length of a paramere; widest point of parameres (near base) nearly 2/3 greatest width of median lobe (near base), with outer margins slightly sinuate, and rounded apex; median lobe roughly triangular, dorsally concave, apical region nearly half as wide as base, broadly rounded at apex; gonopore situated at apex of median lobe.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="93">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">Named after Cesar J. Benetti, Brazilian specialist on aquatic beetles, in honor of his contributions to Neotropical beetle taxonomy and for all his assistance in the field.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
Only known from the type locality in Amazonas State, Brazil, situated slightly north of the Amazon River (Fig.
|
||||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Tobochares spp." figureDoi="10.3897/zookeys.1019.59881.figure13" httpUri="https://binary.pensoft.net/fig/512611" pageId="0" pageNumber="93">13</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="life history">
|
||||
<paragraph pageId="0" pageNumber="93">Life history.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
This only known series was collected on a vertical seepage on sedimentary rock (Fig.
|
||||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Tobochares habitat in Brazil A, B type locality and habitat for T. benettii, seepage near Rio Preto de Eva (collecting event BR 17 - 0610 - 01 A) C type locality and habitat for T. goias, margin of Balneario Lejas (collecting event BR 18 - 0304 - 02 B) D habitat of T. kusad and T. sipaliwini, State of Roraima, near Usina de Jatapu reservoir (collecting Event BR 18 - 0117 - 01 A) E habitat and type locality of T. romanoae, and habitat of T. sipaliwini, State of Roraima, Serra do Tepequem, Igarape Preto Negro, Cachoeira Leje Preta (collecting event BR 18 - 0114 - 04 B) F type locality and habitat of T. fusus, State of Amapa, Calcoene (collecting event BR 18 - 0721 - 02 B)." figureDoi="10.3897/zookeys.1019.59881.figure14" httpUri="https://binary.pensoft.net/fig/512612" pageId="0" pageNumber="93">14A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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