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<mods:namePart id="4FA8FF69EC754140A5BE54128095B3B9">Livermore, Laurence</mods:namePart>
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<taxonomicName id="7C84E9065A91A6DB9E087329943DEB83" ID-CoL="3YZFQ" ID-ENA="311859" authority="Hoffmeyer, 1929" authorityName="Hoffmeyer" authorityYear="1929" class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus rafni" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="species" species="rafni">Megastigmus rafni Hoffmeyer, 1929</taxonomicName>
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<paragraph id="DCB14F58DC8E346199FD361EE1A3E3E4" pageId="0" pageNumber="8013">Notes</paragraph>
<paragraph id="78EC86FAC101C1A40F876391248DA918" pageId="0" pageNumber="8013">
An accidental introduction from North America, developing in the seeds of
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(
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Roques and
<normalizedToken id="0CA20166389393A273B7D06E783887AE" originalValue="Skrzypczyńska">Skrzypczynska</normalizedToken>
2003
</bibRefCitation>
).
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</treatment>
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<taxonomicName id="3C979CFCFF82FF8EFEF88949FDE6214B" ID-CoL="XHF9" authority="Kovařík &amp; Ahmed, 2007" authorityName="Kovařík &amp; Ahmed" authorityYear="2007" box="[314,616,855,882]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="afghanus" status="sp. nov.">
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(
<figureCitation id="63ACFBFAFF82FF8EFE4B8964FE7621AD" box="[393,504,890,916]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="4.[192,268,1084,1107]" captionStartId-1="5.[192,268,870,893]" captionStartId-2="5.[192,268,1674,1697]" captionTargetBox-0="[322,585,173,1039]" captionTargetBox-1="[191,1441,138,833]" captionTargetBox-2="[191,1440,915,1609]" captionTargetId-0="figure-507@4.[284,624,168,1046]" captionTargetId-1="figure-7@5.[192,1440,144,832]" captionTargetId-2="figure-31@5.[192,1439,916,1608]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figure 1: Compsobuthus afghanus sp. n., female holotype, chela of pedipalp." captionText-1="Figure 2: Compsobuthus afghanus sp. n., female holotype, dorsal aspect." captionText-2="Figure 3: Compsobuthus afghanus sp. n., female holotype, ventral aspect." figureDoi-0="http://doi.org/10.5281/zenodo.4649031" figureDoi-1="http://doi.org/10.5281/zenodo.4649033" figureDoi-2="http://doi.org/10.5281/zenodo.4649035" httpUri-0="https://zenodo.org/record/4649031/files/figure.png" httpUri-1="https://zenodo.org/record/4649033/files/figure.png" httpUri-2="https://zenodo.org/record/4649035/files/figure.png" pageId="2" pageNumber="1">Figs. 1–3</figureCitation>
,
<tableCitation id="B615D2C4FF82FF8EFDC78964FDD221AD" box="[517,604,890,916]" captionStart="Table 1" captionStartId="3.[192,257,1074,1097]" captionTargetId="graphics-7@3.[355,1276,192,1036]" captionTargetPageId="3" captionText="Table 1: Measurements (in mm) of type specimens of the new species. * Includes intersegment membranes." httpUri="http://table.plazi.org/id/AFE8B7F7FF83FF8FFF028E2CFB522670" pageId="2" pageNumber="1" tableUuid="AFE8B7F7FF83FF8FFF028E2CFB522670">Table 1</tableCitation>
)
</paragraph>
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<emphasis id="C9E33B6DFF82FF8EFF0289A5FEEE21EB" box="[192,352,955,978]" italics="true" pageId="2" pageNumber="1">Type material</emphasis>
.
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<emphasis id="C9E33B6DFF82FF8EFEB889A5FD8B21EB" bold="true" box="[378,517,955,978]" pageId="2" pageNumber="1">
<collectingCountry id="8380A7EFFF82FF8EFEB889A5FD8B21EB" box="[378,517,955,978]" name="Afghanistan" pageId="2" pageNumber="1">Afghanistan</collectingCountry>
</emphasis>
,
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env.,
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, 1962,
<specimenCount id="ED912CF6FF82FF8EFEAA89C4FE0721C8" box="[360,393,986,1009]" pageId="2" pageNumber="1" type="female">1♀</specimenCount>
(
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1 im. (
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), collector unknown. The types are deposited in the first author‘s collection (
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.
</paragraph>
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<heading id="A0605013FF82FF8EFF028E4CFD6A2650" box="[192,740,1106,1129]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="3">
<emphasis id="C9E33B6DFF82FF8EFF028E4CFEBC2650" box="[192,306,1106,1129]" italics="true" pageId="2" pageNumber="1">Etymology</emphasis>
: Named after the country of occurrence.
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</paragraph>
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<subSubSection id="B38DB4F4FF82FF8EFF028E92FDEF242A" pageId="2" pageNumber="1" type="diagnosis">
<paragraph id="FB28E77FFF82FF8EFF028E92FDEF242A" blockId="2.[192,803,1164,1555]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFF028E92FEA2269A" box="[192,300,1164,1187]" italics="true" pageId="2" pageNumber="1">Diagnosis</emphasis>
: Total length
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. Base color yellow to yellowish brown, only anterior part of fifth metasomal segment black. Movable finger of pedipalp bears 14 rows of granules, with external, very small granules present at all rows (
<emphasis id="C9E33B6DFF82FF8EFE6E8F19FE712727" box="[428,511,1287,1310]" italics="true" pageId="2" pageNumber="1">werneri</emphasis>
group). Internal granules present at all rows and larger than external granules. First metasomal segment with 10 carinae, second to fourth with 8 carinae. Intermediate carinae on second metasomal segment replaced by less than 10 minute, isolated granules situated mainly in posterior half; third segment bears only 2–3 posteriorly situated granules; fourth segment with lateral surface smooth, devoid of granules. Pectinal teeth number 29–31.
</paragraph>
</subSubSection>
<subSubSection id="B38DB4F4FF82FF8EFF028C28FBAD2446" pageId="2" pageNumber="1" type="description">
<paragraph id="FB28E77FFF82FF8EFF028C28FEFB24F1" blockId="2.[192,802,1590,1889]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFF028C28FEB02474" box="[192,318,1590,1613]" italics="true" pageId="2" pageNumber="1">Description</emphasis>
: The
<typeStatus id="242C59DDFF82FF8EFE4B8C28FE692474" box="[393,487,1590,1613]" pageId="2" pageNumber="1" type="holotype">holotype</typeStatus>
(adult female) is
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long. For habitus see
<figureCitation id="63ACFBFAFF82FF8EFE658C4BFE7F2455" box="[423,497,1621,1644]" captionStart="Figure 2" captionStartId="5.[192,268,870,893]" captionTargetBox="[191,1441,138,833]" captionTargetId="figure-7@5.[192,1440,144,832]" captionTargetPageId="5" captionText="Figure 2: Compsobuthus afghanus sp. n., female holotype, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4649033" httpUri="https://zenodo.org/record/4649033/files/figure.png" pageId="2" pageNumber="1">Figs. 2</figureCitation>
and
<figureCitation id="63ACFBFAFF82FF8EFDE48C4BFDB82455" box="[550,566,1621,1644]" captionStart="Figure 3" captionStartId="5.[192,268,1674,1697]" captionTargetBox="[191,1440,915,1609]" captionTargetId="figure-31@5.[192,1439,916,1608]" captionTargetPageId="5" captionText="Figure 3: Compsobuthus afghanus sp. n., female holotype, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4649035" httpUri="https://zenodo.org/record/4649035/files/figure.png" pageId="2" pageNumber="1">3</figureCitation>
. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps, and numbers of pectinal teeth are given in
<tableCitation id="B615D2C4FF82FF8EFEDF8CAFFEE124F1" box="[285,367,1713,1736]" captionStart="Table 1" captionStartId="3.[192,257,1074,1097]" captionTargetId="graphics-7@3.[355,1276,192,1036]" captionTargetPageId="3" captionText="Table 1: Measurements (in mm) of type specimens of the new species. * Includes intersegment membranes." httpUri="http://table.plazi.org/id/AFE8B7F7FF83FF8FFF028E2CFB522670" pageId="2" pageNumber="1" tableUuid="AFE8B7F7FF83FF8FFF028E2CFB522670">Table 1</tableCitation>
.
</paragraph>
<paragraph id="FB28E77FFF82FF8EFF028CCEFE32251D" blockId="2.[192,802,1590,1889]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFF028CCEFEBB24DE" box="[192,309,1744,1767]" italics="true" pageId="2" pageNumber="1">Coloration</emphasis>
: The base colour is yellow to yellowish brown, only anterior part of the fifth metasomal segment is black (
<figureCitation id="63ACFBFAFF82FF8EFEE38D13FEE4251D" box="[289,362,1805,1828]" captionStart="Figure 2" captionStartId="5.[192,268,870,893]" captionTargetBox="[191,1441,138,833]" captionTargetId="figure-7@5.[192,1440,144,832]" captionTargetPageId="5" captionText="Figure 2: Compsobuthus afghanus sp. n., female holotype, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4649033" httpUri="https://zenodo.org/record/4649033/files/figure.png" pageId="2" pageNumber="1">Figs. 2</figureCitation>
and
<figureCitation id="63ACFBFAFF82FF8EFE5C8D13FE25251D" box="[414,427,1805,1828]" captionStart="Figure 3" captionStartId="5.[192,268,1674,1697]" captionTargetBox="[191,1440,915,1609]" captionTargetId="figure-31@5.[192,1439,916,1608]" captionTargetPageId="5" captionText="Figure 3: Compsobuthus afghanus sp. n., female holotype, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4649035" httpUri="https://zenodo.org/record/4649035/files/figure.png" pageId="2" pageNumber="1">3</figureCitation>
).
</paragraph>
<paragraph id="FB28E77FFF82FF8EFF028D32FB5A21D0" blockId="2.[192,802,1590,1889]" lastBlockId="2.[830,1441,855,1663]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFF028D32FE5F257A" box="[192,465,1836,1859]" italics="true" pageId="2" pageNumber="1">Carapace and Mesosoma</emphasis>
: Tergites I–VI bear denticulate lateral carinae. Tergite VII is pentacarinate. Carapace and mesosoma are smooth, only sparsely granulated. The pectinal tooth count is 29–31. The seventh segment is smooth and bears four ventral crenulate carinae. The other sternites are smooth, and the sixth segment bears four smooth carinae without granules.
</paragraph>
<paragraph id="FB28E77FFF82FF8EFCFC89EFFAF7275A" blockId="2.[830,1441,855,1663]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFCFC89EFFBAA2631" box="[830,1060,1009,1032]" italics="true" pageId="2" pageNumber="1">Metasoma and telson</emphasis>
: The first segment has a total of 10 carinae, the second through fourth segments have eight carinae, and the fifth segment has five carinae. Intermediate carinae of the second segment are replaced by less than 10 minute and isolated granules situated mainly in the posterior half; the third segment bears only two or three posteriorly situated granules; and the fourth segment has the lateral surface smooth, entirely devoid of granules. The segments bear only a few bristles each, which are absent between ventral carinae. The telson is elongate, with the vesicle longer than the aculeus. Its surface is generally smooth and bears a few granules.
</paragraph>
<paragraph id="FB28E77FFF82FF8EFCFC8F75FBAD2446" blockId="2.[830,1441,855,1663]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFCFC8F75FC2727BB" box="[830,937,1387,1410]" italics="true" pageId="2" pageNumber="1">Pedipalps</emphasis>
: The femur has four granulose to crenulate carinae and the patella has seven partly crenulate carinae. The chela has smooth carinae, which are difficult to see. For the position and distribution of trichobothria on the chela see
<figureCitation id="63ACFBFAFF82FF8EFB6F8FF7FB7B2439" box="[1197,1269,1513,1536]" captionStart="Figure 1" captionStartId="4.[192,268,1084,1107]" captionTargetBox="[322,585,173,1039]" captionTargetId="figure-507@4.[284,624,168,1046]" captionTargetPageId="4" captionText="Figure 1: Compsobuthus afghanus sp. n., female holotype, chela of pedipalp." figureDoi="http://doi.org/10.5281/zenodo.4649031" httpUri="https://zenodo.org/record/4649031/files/figure.png" pageId="2" pageNumber="1">Fig. 1</figureCitation>
. The movable fingers of pedipalps bear 14 rows of granules, with external, very small granules present at all rows. Internal granules are also present at all rows and are larger than the external granules.
</paragraph>
</subSubSection>
<subSubSection id="B38DB4F4FF82FF8EFCFC8CB2FBC32558" pageId="2" pageNumber="1" type="discussion">
<paragraph id="FB28E77FFF82FF8EFCFC8CB2FBC32558" blockId="2.[830,1441,1708,1889]" pageId="2" pageNumber="1">
<emphasis id="C9E33B6DFF82FF8EFCFC8CB2FC1024FA" box="[830,926,1708,1731]" italics="true" pageId="2" pageNumber="1">Affinities</emphasis>
. The described features distinguish
<taxonomicName id="3C979CFCFF82FF8EFA4A8CB2FC2E24DA" authority="Kovařík &amp; Ahmed, 2007" authorityName="Kovařík &amp; Ahmed" authorityYear="2007" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="afghanus" status="sp. nov.">
<emphasis id="C9E33B6DFF82FF8EFA4A8CB2FC2E24DA" italics="true" pageId="2" pageNumber="1">C. afghanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D08616FF82FF8EFC6A8CD2FC6F24DA" box="[936,993,1740,1763]" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
from all other species of the genus. They are recounted in the key below. This species is characterized by numerous rows of granules on the movable finger of pedipalp (14 rows) and a high number of pectinal teeth (29–31).
</paragraph>
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</subSection>
</treatment>
</document>

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<heading id="A0605013FF83FF8FFEF28E8EFD372692" box="[304,697,1168,1195]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2" reason="3">
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(
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,
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)
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.
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env., 1962,
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(
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), collector unknown. The
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is deposited in the first author‘s collection (
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<mods:title id="5A15F7721D59863DCDB12B38F51FA1AB">Two new species of the genus Compsobuthus Vachon, 1949 from Afghanistan and Pakistan (Scorpiones: Buthidae)</mods:title>
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<subSubSection id="B38DB4F4FF86FF8AFCAF8C0CFAF62410" box="[877,1400,1553,1577]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="FB28E77FFF86FF8AFCAF8C0CFAF62410" blockId="6.[877,1400,1553,1577]" box="[877,1400,1553,1577]" pageId="6" pageNumber="5">
<heading id="A0605013FF86FF8AFCAF8C0CFAF62410" box="[877,1400,1553,1577]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="5" reason="3">
<emphasis id="C9E33B6DFF86FF8AFCAF8C0CFAF62410" bold="true" box="[877,1400,1553,1577]" pageId="6" pageNumber="5">
List of
<taxonomicName id="3C979CFCFF86FF8AFC7F8C0FFBEF2410" ID-CoL="3SVP" authorityName="Vachon" authorityYear="1949" box="[957,1121,1553,1577]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="C9E33B6DFF86FF8AFC7F8C0FFBEF2410" bold="true" box="[957,1121,1553,1577]" italics="true" pageId="6" pageNumber="5">Compsobuthus</emphasis>
</taxonomicName>
from
<collectingCountry id="8380A7EFFF86FF8AFB658C0CFABC2410" box="[1191,1330,1554,1577]" name="Afghanistan" pageId="6" pageNumber="5">Afghanistan</collectingCountry>
,
<collectingCountry id="8380A7EFFF86FF8AFAFD8C0CFAFD2410" box="[1343,1395,1554,1577]" name="Iran" pageId="6" pageNumber="5">Iran</collectingCountry>
,
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B38DB4F4FF86FF8BFC778C2FFD6C2775" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="FB28E77FFF86FF8AFC778C2FFAA72471" blockId="6.[949,1321,1585,1608]" box="[949,1321,1585,1608]" pageId="6" pageNumber="5">
<heading id="A0605013FF86FF8AFC778C2FFAA72471" box="[949,1321,1585,1608]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="5" reason="2">
<emphasis id="C9E33B6DFF86FF8AFC778C2FFAA72471" bold="true" box="[949,1321,1585,1608]" pageId="6" pageNumber="5">
<collectingCountry id="8380A7EFFF86FF8AFC778C2FFB942471" box="[949,1050,1585,1608]" name="Pakistan" pageId="6" pageNumber="5">Pakistan</collectingCountry>
, and adjacent countries
</emphasis>
</heading>
</paragraph>
<paragraph id="FB28E77FFF86FF8BFCFC8C72FEF62648" blockId="6.[830,1416,1644,1883]" lastBlockId="7.[192,802,1082,1356]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
<emphasis id="C9E33B6DFF86FF8AFCFC8C72FB1A24BC" bold="true" box="[830,1172,1644,1669]" pageId="6" pageNumber="5">
<taxonomicName id="3C979CFCFF86FF8AFCFC8C72FBDC24BD" authority="Kovařík &amp; Ahmed, 2007" authorityName="Kovařík &amp; Ahmed" authorityYear="2007" box="[830,1106,1644,1668]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="afghanus" status="sp. nov.">
<emphasis id="C9E33B6DFF86FF8AFCFC8C72FBDC24BD" bold="true" box="[830,1106,1644,1668]" italics="true" pageId="6" pageNumber="5">Compsobuthus afghanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D08616FF86FF8AFB9A8C70FB1A24BC" box="[1112,1172,1646,1669]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="8380A7EFFF86FF8AFB608C70FAA824BC" box="[1186,1318,1646,1669]" name="Afghanistan" pageId="6" pageNumber="5">Afghanistan</collectingCountry>
)
<taxonomicName id="3C979CFCFF86FF8AFCFC8C95FAB7249A" authority="Lourenco, 2004" authorityName="Lourenco" authorityYear="2004" box="[830,1337,1675,1699]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="C9E33B6DFF86FF8AFCFC8C95FBB2249A" bold="true" box="[830,1084,1675,1699]" italics="true" pageId="6" pageNumber="5">Compsobuthus andresi</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFB808C92FB64249A" author="LOURENCO" box="[1090,1258,1676,1699]" pageId="6" pageNumber="5" pagination="157 - 163" refId="ref2874" refString="LOURENCO, W. R. 2004. A new species of Compsobuthus Vachon, 1949 from India (Scorpiones: Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (169): 157 - 163." type="journal article" year="2004">Lourenço, 2004</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFB388C92FABC249A" box="[1274,1330,1676,1699]" name="India" pageId="6" pageNumber="5">India</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8CB4FAC324FB" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[830,1357,1706,1730]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="becvari">
<emphasis id="C9E33B6DFF86FF8AFCFC8CB4FBB424FB" bold="true" box="[830,1082,1706,1730]" italics="true" pageId="6" pageNumber="5">Compsobuthus becvari</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFB828CB5FB5424FB" author="KOVARIK" box="[1088,1242,1707,1730]" pageId="6" pageNumber="5" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFB288CB5FACA24FB" box="[1258,1348,1707,1730]" name="Pakistan" pageId="6" pageNumber="5">Pakistan</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8CD6FA0624D8" authority="Lourenco et Vachon, 2001" authorityName="Lourenco et Vachon" authorityYear="2001" box="[830,1416,1736,1761]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="garyi">
<emphasis id="C9E33B6DFF86FF8AFCFC8CD6FBAC24D8" bold="true" box="[830,1058,1736,1761]" italics="true" pageId="6" pageNumber="5">Compsobuthus garyi</emphasis>
Lourenço et Vachon, 2001 (
<collectingCountry id="8380A7EFFF86FF8AFA978CD4FA0C24D8" box="[1365,1410,1738,1761]" name="Iran" pageId="6" pageNumber="5">Iran</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8CF9FAFF24C6" authority="Amir et al., 2005" authorityName="Amir" authorityYear="2005" box="[830,1393,1767,1791]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="humaae">
<emphasis id="C9E33B6DFF86FF8AFCFC8CF9FBCD24C6" bold="true" box="[830,1091,1767,1791]" italics="true" pageId="6" pageNumber="5">Compsobuthus humaae</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFB8B8CF6FB7024C6" author="AMIR" box="[1097,1278,1768,1791]" pageId="6" pageNumber="5" pagination="117 - 121" refId="ref2685" refString="AMIR, R., S. KAMALUDDIN &amp; M. A. JABBAR KHAN. 2005. A new species of the genus Compsobuthus Vachon (Arachnida: Scorpionida: Buthidae) from Pakistan. Pakistan Journal of Zoology, 37 (2): 117 - 121." type="journal article" year="2005">Amir et al., 2005</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFACF8CF6FAE524C6" box="[1293,1387,1768,1791]" name="Pakistan" pageId="6" pageNumber="5">Pakistan</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8D18FA832527" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[830,1293,1798,1822]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="jakesi">
<emphasis id="C9E33B6DFF86FF8AFCFC8D18FBA42527" bold="true" box="[830,1066,1798,1822]" italics="true" pageId="6" pageNumber="5">Compsobuthus jakesi</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFBF28D19FB442527" author="KOVARIK" box="[1072,1226,1799,1822]" pageId="6" pageNumber="5" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFB188D19FA8B2527" box="[1242,1285,1799,1822]" name="Iraq" pageId="6" pageNumber="5">Iraq</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8D3AFA9D2504" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[830,1299,1828,1853]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="kafkai">
<emphasis id="C9E33B6DFF86FF8AFCFC8D3AFBBE2505" bold="true" box="[830,1072,1828,1852]" italics="true" pageId="6" pageNumber="5">Compsobuthus kafkai</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFBF48D38FB412504" author="KOVARIK" box="[1078,1231,1830,1853]" pageId="6" pageNumber="5" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFB1D8D38FA822504" box="[1247,1292,1830,1853]" name="Iran" pageId="6" pageNumber="5">Iran</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF86FF8AFCFC8D5DFA922562" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[830,1308,1859,1883]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="kaftani">
<emphasis id="C9E33B6DFF86FF8AFCFC8D5DFBB72562" bold="true" box="[830,1081,1859,1883]" italics="true" pageId="6" pageNumber="5">Compsobuthus kaftani</emphasis>
<bibRefCitation id="9F069A8EFF86FF8AFBFD8D5AFB572562" author="KOVARIK" box="[1087,1241,1860,1883]" pageId="6" pageNumber="5" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF86FF8AFB258D5AFA9A2562" box="[1255,1300,1860,1883]" name="Iran" pageId="6" pageNumber="5">Iran</collectingCountry>
)
</taxonomicName>
<taxonomicName id="3C979CFCFF87FF8BFF028E24FD2E266B" authority="(Birula, 1905)" baseAuthorityName="Birula" baseAuthorityYear="1905" box="[192,672,1082,1106]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="matthiesseni">
<emphasis id="C9E33B6DFF87FF8BFF028E24FE74266B" bold="true" box="[192,506,1082,1106]" italics="true" pageId="7" pageNumber="6">Compsobuthus matthiesseni</emphasis>
(Birula, 1905)
</taxonomicName>
(
<collectingCountry id="8380A7EFFF87FF8BFD708E25FD51266B" box="[690,735,1083,1106]" name="Iraq" pageId="7" pageNumber="6">Iraq</collectingCountry>
,
<collectingCountry id="8380A7EFFF87FF8BFD328E25FC90266B" box="[752,798,1083,1106]" name="Iran" pageId="7" pageNumber="6">Iran</collectingCountry>
,
<collectingCountry id="8380A7EFFF87FF8BFF1F8E44FE962648" box="[221,280,1114,1137]" name="Syria" pageId="7" pageNumber="6">Syria</collectingCountry>
,
<collectingCountry id="8380A7EFFF87FF8BFEE08E44FEFF2648" box="[290,369,1114,1137]" name="Turkey" pageId="7" pageNumber="6">Turkey</collectingCountry>
)
</paragraph>
<caption id="AFE8B7F7FF87FF8BFF0289E0FB8B262C" ID-DOI="http://doi.org/10.5281/zenodo.4649039" ID-Zenodo-Dep="4649039" box="[192,1029,1022,1045]" httpUri="https://zenodo.org/record/4649039/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,268,1022,1045]" targetBox="[194,1428,190,982]" targetPageId="7">
<paragraph id="FB28E77FFF87FF8BFF0289E0FB8B262C" blockId="7.[192,1029,1022,1045]" box="[192,1029,1022,1045]" pageId="7" pageNumber="6">
<emphasis id="C9E33B6DFF87FF8BFF0289E0FEA6262C" bold="true" box="[192,296,1022,1045]" pageId="7" pageNumber="6">Figure 5:</emphasis>
<taxonomicName id="3C979CFCFF87FF8BFEED8E1EFDA4262C" authority="Kovařík &amp; Ahmed, 2007" authorityName="Kovařík &amp; Ahmed" authorityYear="2007" box="[303,554,1024,1045]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pakistanus" status="sp. nov.">
<emphasis id="C9E33B6DFF87FF8BFEED8E1EFDA4262C" box="[303,554,1024,1045]" italics="true" pageId="7" pageNumber="6">Compsobuthus pakistanus</emphasis>
</taxonomicName>
<emphasis id="C9E33B6DFF87FF8BFDF28E1EFDE8262C" bold="true" box="[560,614,1024,1045]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="D2D08616FF87FF8BFDF28E1EFDE8262C" box="[560,614,1024,1045]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, immature female holotype, ventral aspect.
</paragraph>
</caption>
<paragraph id="FB28E77FFF87FF8BFF028E66FD1926A8" blockId="7.[192,802,1082,1356]" box="[192,663,1144,1169]" pageId="7" pageNumber="6">
<emphasis id="C9E33B6DFF87FF8BFF028E66FDAA26A8" bold="true" box="[192,548,1144,1169]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028E66FE6C26A9" authority="Kovařík &amp; Ahmed, 2007" authorityName="Kovařík &amp; Ahmed" authorityYear="2007" box="[192,482,1144,1168]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pakistanus" status="sp. nov.">
<emphasis id="C9E33B6DFF87FF8BFF028E66FE6C26A9" bold="true" box="[192,482,1144,1168]" italics="true" pageId="7" pageNumber="6">Compsobuthus pakistanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D08616FF87FF8BFE2B8E64FDAA26A8" box="[489,548,1146,1169]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="8380A7EFFF87FF8BFDF08E64FD1E26A8" box="[562,656,1146,1169]" name="Pakistan" pageId="7" pageNumber="6">Pakistan</collectingCountry>
)
</paragraph>
<paragraph id="FB28E77FFF87FF8BFF028E86FD2E2689" blockId="7.[192,802,1082,1356]" box="[192,672,1176,1200]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028E86FD2E2689" authority="Vignoli, 2005" authorityName="Vignoli" authorityYear="2005" box="[192,672,1176,1200]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="petriolii">
<emphasis id="C9E33B6DFF87FF8BFF028E86FE4F2689" bold="true" box="[192,449,1176,1200]" italics="true" pageId="7" pageNumber="6">Compsobuthus petriolii</emphasis>
<bibRefCitation id="9F069A8EFF87FF8BFE0A8E87FDD22689" author="VIGNOLI" box="[456,604,1177,1200]" pageId="7" pageNumber="6" pagination="79 - 86" refId="ref2983" refString="VIGNOLI, V. 2005. Description of a new species of Compsobuthus Vachon, 1949 (Scorpiones: Buthidae) from southern Iran. Zoology in the Middle East, 34: 79 - 86." type="journal article" year="2005">Vignoli, 2005</bibRefCitation>
(
<collectingCountry id="8380A7EFFF87FF8BFDA88E87FD192689" box="[618,663,1177,1200]" name="Iran" pageId="7" pageNumber="6">Iran</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="FB28E77FFF87FF8BFF028EA9FD3926F6" blockId="7.[192,802,1082,1356]" box="[192,695,1207,1231]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028EA9FD3926F6" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[192,695,1207,1231]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="plutenkoi">
<emphasis id="C9E33B6DFF87FF8BFF028EA9FE5D26F6" bold="true" box="[192,467,1207,1231]" italics="true" pageId="7" pageNumber="6">Compsobuthus plutenkoi</emphasis>
<bibRefCitation id="9F069A8EFF87FF8BFE188EA6FDFD26F6" author="KOVARIK" box="[474,627,1208,1231]" pageId="7" pageNumber="6" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF87FF8BFD418EA6FD2026F6" box="[643,686,1208,1231]" name="Iran" pageId="7" pageNumber="6">Iran</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="FB28E77FFF87FF8BFF028EC8FE962737" blockId="7.[192,802,1082,1356]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028EC8FD1D26D7" authority="(Pocock, 1900)" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[192,659,1238,1262]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="rugosulus">
<emphasis id="C9E33B6DFF87FF8BFF028EC8FE5026D7" bold="true" box="[192,478,1238,1262]" italics="true" pageId="7" pageNumber="6">Compsobuthus rugosulus</emphasis>
(Pocock, 1900)
</taxonomicName>
(
<collectingCountry id="8380A7EFFF87FF8BFD648EC9FD5026D7" box="[678,734,1239,1262]" name="India" pageId="7" pageNumber="6">India</collectingCountry>
,
<collectingCountry id="8380A7EFFF87FF8BFD338EC9FE812737" name="Pakistan" pageId="7" pageNumber="6">Pakistan</collectingCountry>
)
</paragraph>
<paragraph id="FB28E77FFF87FF8BFF028F0BFD3A2714" blockId="7.[192,802,1082,1356]" box="[192,692,1301,1325]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028F0BFD3A2714" authority="Kovarik, 2003" authorityName="Kovarik" authorityYear="2003" box="[192,692,1301,1325]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="sobotniki">
<emphasis id="C9E33B6DFF87FF8BFF028F0BFE5E2714" bold="true" box="[192,464,1301,1325]" italics="true" pageId="7" pageNumber="6">Compsobuthus sobotniki</emphasis>
<bibRefCitation id="9F069A8EFF87FF8BFE158F08FDFE2714" author="KOVARIK" box="[471,624,1302,1325]" pageId="7" pageNumber="6" pagination="87 - 112" refId="ref2798" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
(
<collectingCountry id="8380A7EFFF87FF8BFD428F08FD252714" box="[640,683,1302,1325]" name="Iran" pageId="7" pageNumber="6">Iran</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="FB28E77FFF87FF8BFF028F2AFD6C2775" blockId="7.[192,802,1082,1356]" box="[192,738,1332,1356]" pageId="7" pageNumber="6">
<taxonomicName id="3C979CFCFF87FF8BFF028F2AFD6C2775" authority="Lourenco, 2001" authorityName="Lourenco" authorityYear="2001" box="[192,738,1332,1356]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tofti">
<emphasis id="C9E33B6DFF87FF8BFF028F2AFE162775" bold="true" box="[192,408,1332,1356]" italics="true" pageId="7" pageNumber="6">Compsobuthus tofti</emphasis>
Lourenço, 2001 (
<collectingCountry id="8380A7EFFF87FF8BFD978F2BFD552775" box="[597,731,1333,1356]" name="Afghanistan" pageId="7" pageNumber="6">Afghanistan</collectingCountry>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>