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<document id="050B05CB21A12AC7206C218240259E6D" ID-DOI="10.11646/zootaxa.5569.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14704080" ID-ZooBank="0B9CB2FF-4BD1-4B9F-9A7C-979F6897E67D" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737372208581" checkinUser="plazi" docAuthor="Hallermann, Jakob &amp; Hawlitschek, Oliver" docDate="2025" docId="03D887FDFFD36D78FF180DA4FD19CC26" docLanguage="en" docName="zootaxa.5569.1.4.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boaedon subniger Hallermann &amp; Hawlitschek, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="131" masterDocId="FFE1FF85FFDA6D74FF8F0B43FFFFCE47" masterDocTitle="Two new species of Boaedon from Ethiopia and Somalia, with a review of the species of East Africa" masterLastPageNumber="137" masterPageNumber="119" pageNumber="128" updateTime="1737485471144" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="AFDD09018B658FE0A46770946CCD560F">Two new species of Boaedon from Ethiopia and Somalia, with a review of the species of East Africa</mods:title>
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<mods:namePart id="87BE8A356E841DD9D63C3EBD7E630D25">Hallermann, Jakob</mods:namePart>
<mods:affiliation id="BB111A2CFCA67727EA44C52214CCAE5B">Leibniz Institute for the Analysis of Biodiversity Change (LIB), Zoologisches Museum Hamburg, Martin-Luther-King Platz 3, D- 20146 Hamburg.</mods:affiliation>
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<mods:namePart id="FE531F8E8F2A65B0BF16F329791518A6">Hawlitschek, Oliver</mods:namePart>
<mods:affiliation id="480A89A5BA0C9E16A82E00E085CF722C">Department of Evolutionary Biology and Environmental Studies, Universität Zürich, Zürich, Switzerland</mods:affiliation>
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<paragraph id="8BCE36EBFFD36D7DFF180DA4FE26C945" blockId="9.[151,473,1767,1830]" box="[151,473,1767,1794]" pageId="9" pageNumber="128">
<heading id="D0868187FFD36D7DFF180DA4FE26C945" bold="true" box="[151,473,1767,1794]" fontSize="11" level="1" pageId="9" pageNumber="128" reason="1">
<emphasis id="B905EAF9FFD36D7DFF180DA4FE26C945" bold="true" box="[151,473,1767,1794]" pageId="9" pageNumber="128">
<taxonomicName id="4C714D68FFD36D7DFF180DA4FE8BC945" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[151,372,1767,1794]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="9" pageNumber="128" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD36D7DFF180DA4FE8BC945" bold="true" box="[151,372,1767,1794]" italics="true" pageId="9" pageNumber="128">Boaedon subniger</emphasis>
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<taxonomicNameLabel id="A2365782FFD36D7DFEF40DABFE26C945" box="[379,473,1768,1794]" pageId="9" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36B6560FFD36D7DFF180C48FEEDC962" box="[151,274,1803,1830]" pageId="9" pageNumber="128" type="description">
<paragraph id="8BCE36EBFFD36D7DFF180C48FEEDC962" blockId="9.[151,473,1767,1830]" box="[151,274,1803,1830]" pageId="9" pageNumber="128">
<emphasis id="B905EAF9FFD36D7DFF180C48FF5FC962" bold="true" box="[151,160,1803,1829]" pageId="9" pageNumber="128">(</emphasis>
<figureCitation id="134A2A6EFFD36D7DFF2E0C48FF10C961" box="[161,239,1803,1830]" captionStart="FIGURE 5" captionStartId="10.[152,255,1944,1968]" captionTargetBox="[192,1363,193,1842]" captionTargetId="figure-34@10.[192,1396,181,1895]" captionTargetPageId="10" captionText="FIGURE 5. Holotype of Boaedon subniger sp. nov. (ZMH R22350) 8.5 km south of Yuffleh 1758 m, Sanaag region, Somaliland Left: head in dorsal, ventral and lateral view, right side drawings of the head. Photo: Tomáš Mazuch." figureDoi="http://doi.org/10.5281/zenodo.14704094" httpUri="https://zenodo.org/record/14704094/files/figure.png" pageId="9" pageNumber="128">Figs. 5</figureCitation>
,
<figureCitation id="134A2A6EFFD36D7DFF740C48FEF5C962" box="[251,266,1803,1829]" captionStart="FIGURE 6" captionStartId="11.[151,250,1058,1082]" captionTargetBox="[151,1436,181,1034]" captionTargetId="figure-26@11.[151,1436,181,1034]" captionTargetPageId="11" captionText="FIGURE 6. Live photo of Boaedon subniger sp. nov. from nearby Yuffleh, Gumbi Hills, Somalia. Photo by Tomáš Mazuch." figureDoi="http://doi.org/10.5281/zenodo.14704098" httpUri="https://zenodo.org/record/14704098/files/figure.png" pageId="9" pageNumber="128">6</figureCitation>
)
</paragraph>
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<subSubSection id="C36B6560FFD36D7DFF180C13FBDEC92F" box="[151,1057,1872,1896]" pageId="9" pageNumber="128" type="reference_group">
<paragraph id="8BCE36EBFFD36D7DFF180C13FBDEC92F" blockId="9.[151,1057,1872,1896]" box="[151,1057,1872,1896]" pageId="9" pageNumber="128">
<treatmentCitationGroup id="AB6111C5FFD36D7DFF180C13FBDEC92F" box="[151,1057,1872,1896]" pageId="9" pageNumber="128">
<treatmentCitation id="0AD010FAFFD36D7DFF180C13FCF7C92F" author="Spawls, S. &amp; Mazuch, T. &amp; Mohamad, A." box="[151,776,1872,1896]" page="443" pageId="9" pageNumber="128" year="2023">
<taxonomicName id="4C714D68FFD36D7DFF180C13FCF7C92F" ID-CoL="M7VV" authority="Spawls, Mazuch &amp; Mohamad 2023: 443" authorityName="Spawls, Mazuch &amp; Mohamad" authorityPageNumber="443" authorityYear="2023" box="[151,776,1872,1896]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="9" pageNumber="128" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B905EAF9FFD36D7DFF180C13FE98C92F" box="[151,359,1872,1896]" italics="true" pageId="9" pageNumber="128">Boaedon fuliginosus</emphasis>
<bibRefCitation id="EFE04B1AFFD36D7DFEE20C13FCF7C92F" author="Spawls, S. &amp; Mazuch, T. &amp; Mohamad, A." box="[365,776,1872,1896]" pageId="9" pageNumber="128" refId="ref10919" refString="Spawls, S., Mazuch, T. &amp; Mohamad, A. (2023) Handbook of Amphibians and Reptiles of North-East Africa. London Bloomsbury Wildlife, 640 pp." type="book" year="2023">Spawls, Mazuch &amp; Mohamad 2023: 443</bibRefCitation>
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from Erigavo (Somaliland)
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</paragraph>
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<subSubSection id="C36B6560FFD36D7DFF180CD5FA64C9BF" pageId="9" pageNumber="128" type="materials_examined">
<paragraph id="8BCE36EBFFD36D7DFF180CD5FA64C9BF" blockId="9.[151,1436,1942,2041]" pageId="9" pageNumber="128">
<materialsCitation id="3B193CB6FFD36D7DFF180CD5FA64C9BF" collectingDate="2017-08-31" collectingDateMax="2024-09" collectingDateMin="2017-08-31" collectionCode="ZMH" collectorName="Tomas Mazuch " elevation="1758" latitude="10.3" location="Yuffleh" longLatPrecision="21" longitude="47.182777" pageId="9" pageNumber="128" specimenCode="R22350, TMHC 2023.01" specimenCount="1" stateProvince="Sanaag" typeStatus="holotype">
<emphasis id="B905EAF9FFD36D7DFF180CD5FEF2C9F7" bold="true" box="[151,269,1942,1968]" pageId="9" pageNumber="128">
<typeStatus id="54CA8849FFD36D7DFF180CD5FEF7C9F7" box="[151,264,1942,1968]" pageId="9" pageNumber="128" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED60AE2EFFD36D7DFE990CD4FEABC9F6" box="[278,340,1943,1969]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="9" pageNumber="128" type="Museum">ZMH</collectionCode>
<specimenCode id="DBD79E90FFD36D7DFED20CD4FE4AC9F6" box="[349,437,1942,1969]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="128">R22350</specimenCode>
(DNA voucher
<emphasis id="B905EAF9FFD36D7DFDE10CD5FC97C9F7" bold="true" box="[622,872,1942,1968]" pageId="9" pageNumber="128">
<specimenCode id="DBD79E90FFD36D7DFDE10CD5FCD5C9F7" box="[622,810,1942,1968]" collectionCode="TMHC" pageId="9" pageNumber="128">TMHC 2023.01</specimenCode>
.1018
</emphasis>
): a juvenile female from
<quantity id="4C899B0EFFD36D7DFB020CD5FB23C9F7" box="[1165,1244,1942,1969]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="128" unit="km" value="8.5">8.5 km</quantity>
south of
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(Yufle, Jooflie) (approx.
<geoCoordinate id="EE45502CFFD36D7DFE3B0CF9FDA8C992" box="[436,599,1978,2005]" degrees="10" direction="north" minutes="18" orientation="latitude" pageId="9" pageNumber="128" precision="15" seconds="00" value="10.3">10° 18 00” N</geoCoordinate>
<geoCoordinate id="EE45502CFFD36D7DFDEE0CF9FCFCC992" box="[609,771,1978,2005]" degrees="47" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="128" precision="15" seconds="58" value="47.182777">47° 10 58” E</geoCoordinate>
,
<quantity id="4C899B0EFFD36D7DFC9D0CF9FC93C993" box="[786,876,1978,2005]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="9" pageNumber="128" unit="m" value="1758.0">1758 m</quantity>
),
<collectingRegion id="49B5F809FFD36D7DFC0C0CF9FBD8C993" box="[899,1063,1978,2004]" country="Somalia" name="Sanaag" pageId="9" pageNumber="128">Sanaag region</collectingRegion>
,
<collectorName id="2684533DFFD36D7DFBBA0CF9FB4AC993" box="[1077,1205,1978,2004]" pageId="9" pageNumber="128">Somaliland</collectorName>
collected by
<collectorName id="2684533DFFD36D7DFADD0CF9FF0BC9BF" pageId="9" pageNumber="128">Tomáš Mazuch</collectorName>
,
<date id="FFCF102BFFD36D7DFF700C9DFE4BC9BF" box="[255,436,2014,2040]" pageId="9" pageNumber="128" value="2017-08-31">
<collectingDate id="EF8BE9C3FFD36D7DFF700C9DFE4BC9BF" box="[255,436,2014,2040]" pageId="9" pageNumber="128" value="2017-08-31">31 August 2017</collectingDate>
</date>
, and donated to the
<collectorName id="2684533DFFD36D7DFD1B0C9DFC89C9BE" box="[660,886,2014,2041]" pageId="9" pageNumber="128">Zoological Museum</collectorName>
Hamburg (
<collectionCode id="ED60AE2EFFD36D7DFC7A0C9DFBC8C9BF" box="[1013,1079,2014,2040]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="9" pageNumber="128" type="Museum">ZMH</collectionCode>
) in
<date id="FFCF102BFFD36D7DFBED0C9DFAE6C9BF" box="[1122,1305,2014,2040]" pageId="9" pageNumber="128" value="2024-09">
<collectingDate id="EF8BE9C3FFD36D7DFBED0C9DFAE6C9BF" box="[1122,1305,2014,2040]" pageId="9" pageNumber="128" value="2024-09">September 2024</collectingDate>
</date>
(
<figureCitation id="134A2A6EFFD36D7DFAA70C9DFA88C9BE" box="[1320,1399,2014,2041]" captionStart="FIGURE 5" captionStartId="10.[152,255,1944,1968]" captionTargetBox="[192,1363,193,1842]" captionTargetId="figure-34@10.[192,1396,181,1895]" captionTargetPageId="10" captionText="FIGURE 5. Holotype of Boaedon subniger sp. nov. (ZMH R22350) 8.5 km south of Yuffleh 1758 m, Sanaag region, Somaliland Left: head in dorsal, ventral and lateral view, right side drawings of the head. Photo: Tomáš Mazuch." figureDoi="http://doi.org/10.5281/zenodo.14704094" httpUri="https://zenodo.org/record/14704094/files/figure.png" pageId="9" pageNumber="128">Figs. 5</figureCitation>
,
<figureCitation id="134A2A6EFFD36D7DFA0C0C9DFA6DC9BF" box="[1411,1426,2014,2040]" captionStart="FIGURE 6" captionStartId="11.[151,250,1058,1082]" captionTargetBox="[151,1436,181,1034]" captionTargetId="figure-26@11.[151,1436,181,1034]" captionTargetPageId="11" captionText="FIGURE 6. Live photo of Boaedon subniger sp. nov. from nearby Yuffleh, Gumbi Hills, Somalia. Photo by Tomáš Mazuch." figureDoi="http://doi.org/10.5281/zenodo.14704098" httpUri="https://zenodo.org/record/14704098/files/figure.png" pageId="9" pageNumber="128">6</figureCitation>
)
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</paragraph>
</subSubSection>
<caption id="DF0E6663FFD06D7EFF170CDBFB6CC993" ID-DOI="http://doi.org/10.5281/zenodo.14704094" ID-Zenodo-Dep="14704094" httpUri="https://zenodo.org/record/14704094/files/figure.png" pageId="10" pageNumber="129" startId="10.[152,255,1944,1968]" targetBox="[192,1363,193,1842]" targetPageId="10" targetType="figure">
<paragraph id="8BCE36EBFFD06D7EFF170CDBFB6CC993" blockId="10.[151,1436,1944,2004]" pageId="10" pageNumber="129">
<emphasis id="B905EAF9FFD06D7EFF170CDBFEE8C9F7" bold="true" box="[152,279,1944,1968]" pageId="10" pageNumber="129">FIGURE 5.</emphasis>
Holotype of
<taxonomicName id="4C714D68FFD06D7EFE170CDBFDAFC9F7" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[408,592,1944,1968]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="10" pageNumber="129" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD06D7EFE170CDBFDAFC9F7" box="[408,592,1944,1968]" italics="true" pageId="10" pageNumber="129">Boaedon subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD06D7EFDDB0CDBFD5CC9F7" bold="true" box="[596,675,1944,1968]" pageId="10" pageNumber="129">
<taxonomicNameLabel id="A2365782FFD06D7EFDDB0CDBFD5CC9F7" box="[596,675,1944,1968]" pageId="10" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMH R22350) 8.5 km south of Yuffleh 1758 m, Sanaag region, Somaliland Left: head in dorsal, ventral and lateral view, right side drawings of the head. Photo: Tomáš Mazuch.
</paragraph>
</caption>
<caption id="DF0E6663FFD16D7FFF180F61FA77CA7D" ID-DOI="http://doi.org/10.5281/zenodo.14704098" ID-Zenodo-Dep="14704098" box="[151,1416,1058,1082]" httpUri="https://zenodo.org/record/14704098/files/figure.png" pageId="11" pageNumber="130" startId="11.[151,250,1058,1082]" targetBox="[151,1436,181,1034]" targetPageId="11" targetType="figure">
<paragraph id="8BCE36EBFFD16D7FFF180F61FA77CA7D" blockId="11.[151,1416,1058,1082]" box="[151,1416,1058,1082]" pageId="11" pageNumber="130">
<emphasis id="B905EAF9FFD16D7FFF180F61FEECCA7D" bold="true" box="[151,275,1058,1082]" pageId="11" pageNumber="130">FIGURE 6.</emphasis>
Live photo of
<taxonomicName id="4C714D68FFD16D7FFE250F61FD9CCA7D" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[426,611,1058,1082]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFE250F61FD9CCA7D" box="[426,611,1058,1082]" italics="true" pageId="11" pageNumber="130">Boaedon subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFDE60F61FD46CA7D" bold="true" box="[617,697,1058,1082]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFDE60F61FD46CA7D" box="[617,697,1058,1082]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from nearby Yuffleh, Gumbi Hills, Somalia. Photo by Tomáš Mazuch.
</paragraph>
</caption>
<subSubSection id="C36B6560FFD16D7FFF480FCFFBD1C885" pageId="11" pageNumber="130" type="diagnosis">
<paragraph id="8BCE36EBFFD16D7FFF480FCFFE28CB55" blockId="11.[151,1437,1164,2019]" pageId="11" pageNumber="130">
<emphasis id="B905EAF9FFD16D7FFF480FCFFEBDCAE1" bold="true" box="[199,322,1164,1190]" pageId="11" pageNumber="130">Diagnosis.</emphasis>
<taxonomicName id="4C714D68FFD16D7FFEC60FCFFDE8CAE1" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[329,535,1164,1190]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFEC60FCFFDE8CAE1" box="[329,535,1164,1190]" italics="true" pageId="11" pageNumber="130">Boaedon subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFD900FCFFD85CAE1" bold="true" box="[543,634,1164,1190]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFD900FCFFD85CAE1" box="[543,634,1164,1190]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from most congeners by its dark brown to blackbrown dorsal colour, without any light head stripes, two supralabials (4
<superScript id="7C049BA3FFD16D7FFC390FF3FC3CCAF9" attach="left" box="[950,963,1200,1214]" fontSize="6" pageId="11" pageNumber="130">th</superScript>
&amp; 5
<superScript id="7C049BA3FFD16D7FFC7A0FF3FBFDCAF9" attach="none" box="[1013,1026,1200,1214]" fontSize="6" pageId="11" pageNumber="130">th</superScript>
) touching on eye and one preocular, touching the frontal. PAR is longer than the distance between the frontal and the rostral scale. Venter whitish (for more characters see table 4).
</paragraph>
<paragraph id="8BCE36EBFFD16D7FFF480E5FFBD1C885" blockId="11.[151,1437,1164,2019]" pageId="11" pageNumber="130">
<taxonomicName id="4C714D68FFD16D7FFF480E5FFE6BCB71" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[199,404,1308,1334]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFF480E5FFE6BCB71" box="[199,404,1308,1334]" italics="true" pageId="11" pageNumber="130">Boaedon subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFE140E5FFE0ACB71" bold="true" box="[411,501,1308,1334]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFE140E5FFE0ACB71" box="[411,501,1308,1334]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C714D68FFD16D7FFD050E5EFCEBCB71" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[650,788,1308,1335]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFD050E5EFD64CB71" box="[650,667,1309,1334]" italics="true" pageId="11" pageNumber="130">B</emphasis>
.
<emphasis id="B905EAF9FFD16D7FFD250E5FFCEBCB71" box="[682,788,1308,1334]" italics="true" pageId="11" pageNumber="130">broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFC940E5FFC8ACB71" bold="true" box="[795,885,1308,1334]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFC940E5FFC8ACB71" box="[795,885,1308,1334]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having fewer MSR (, X 27.0, versus 2733, X 29.1, only one preocular (versus two) and its blackish brown or dark brown body colouration without light coloured head stripes (versus reddish, olive-brown or light brown body colouration with two short white head stripes and lips whitish coloured), with no dorsals uncoloured (versus first two rows uncoloured in
<taxonomicName id="4C714D68FFD16D7FFBDD0ECBFB43CBE5" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[1106,1212,1416,1442]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFBDD0ECBFB43CBE5" box="[1106,1212,1416,1442]" italics="true" pageId="11" pageNumber="130">broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFB4B0ECBFADFCBE5" bold="true" box="[1220,1312,1416,1442]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFB4B0ECBFADFCBE5" box="[1220,1312,1416,1442]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C714D68FFD16D7FFAB60ECBFEF5CB81" authorityName="Parker" authorityYear="1932" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B905EAF9FFD16D7FFAB60ECBFEF5CB81" italics="true" pageId="11" pageNumber="130">Boaedon maculatus</emphasis>
</taxonomicName>
has an additional row of scales between the supralabials and loreal between nasal and preocular scale, (versus no row of scales) and 1011 supralabials (versus 8). Extralimital species:
<taxonomicName id="4C714D68FFD16D7FFBB90E93FAF0CBAD" box="[1078,1295,1488,1514]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B905EAF9FFD16D7FFBB90E93FAF0CBAD" box="[1078,1295,1488,1514]" italics="true" pageId="11" pageNumber="130">Boaedon montanus</emphasis>
</taxonomicName>
in its whole distribution area differs in having two at least short head lines and whitish upper and lower lips (versus no headlines and no whitish lips),
<taxonomicName id="4C714D68FFD16D7FFE050D5BFD78C875" authorityName="Trape &amp; Mediannikov" authorityYear="2016" box="[394,647,1560,1586]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="perisilvestris">
<emphasis id="B905EAF9FFD16D7FFE050D5BFD78C875" box="[394,647,1560,1586]" italics="true" pageId="11" pageNumber="130">Boaedon perisilvestris</emphasis>
</taxonomicName>
with similar colouration differs in having more MSR (2931, X 29.4 (versus 27 X 27.0 in
<taxonomicName id="4C714D68FFD16D7FFE090D7EFDF6C811" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[390,521,1596,1622]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFE090D7EFDF6C811" box="[390,521,1596,1622]" italics="true" pageId="11" pageNumber="130">B. subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFD9D0D7FFD91C811" bold="true" box="[530,622,1596,1622]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFD9D0D7FFD91C811" box="[530,622,1596,1622]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and frequently three supralabial scales touching eye (versus two in
<taxonomicName id="4C714D68FFD16D7FFA0B0D7EFF06C83D" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFA0B0D7EFF06C83D" italics="true" pageId="11" pageNumber="130">B. subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFE8F0D23FEA5C83D" bold="true" box="[256,346,1632,1658]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFE8F0D23FEA5C83D" box="[256,346,1632,1658]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and a genetic interspecific distances of 0.024).
<taxonomicName id="4C714D68FFD16D7FFCF50D22FBD8C83D" authorityName="Trape &amp; Mediannikov" authorityYear="2016" box="[890,1063,1632,1658]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="paralineatus">
<emphasis id="B905EAF9FFD16D7FFCF50D22FBD8C83D" box="[890,1063,1632,1658]" italics="true" pageId="11" pageNumber="130">B. paralineatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C714D68FFD16D7FFBD20D22FB23C83D" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[1117,1244,1633,1658]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B905EAF9FFD16D7FFBD20D22FB23C83D" box="[1117,1244,1633,1658]" italics="true" pageId="11" pageNumber="130">B. capensis</emphasis>
</taxonomicName>
species complex differs in having clear head and/or body lines (versus no head and body lines in
<taxonomicName id="4C714D68FFD16D7FFB920DC6FB60C8D9" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[1053,1183,1668,1694]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFB920DC6FB60C8D9" box="[1053,1183,1668,1694]" italics="true" pageId="11" pageNumber="130">B. subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFB290DC7FAFFC8D9" bold="true" box="[1190,1280,1668,1694]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFB290DC7FAFFC8D9" box="[1190,1280,1668,1694]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and a brown or reddish colouration (versus blackish brown colouration in
<taxonomicName id="4C714D68FFD16D7FFCCC0DEAFC3AC885" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[835,965,1704,1730]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="11" pageNumber="130" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFD16D7FFCCC0DEAFC3AC885" box="[835,965,1704,1730]" italics="true" pageId="11" pageNumber="130">B. subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD16D7FFC430DEBFBDAC885" bold="true" box="[972,1061,1704,1730]" pageId="11" pageNumber="130">
<taxonomicNameLabel id="A2365782FFD16D7FFC430DEBFBDAC885" box="[972,1061,1704,1730]" pageId="11" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFD16D78FF480D8FFE33CFCE" lastPageId="12" lastPageNumber="131" pageId="11" pageNumber="130" type="description">
<paragraph id="8BCE36EBFFD16D78FF480D8FFA66CEBE" blockId="11.[151,1437,1164,2019]" lastBlockId="12.[151,1436,150,609]" lastPageId="12" lastPageNumber="131" pageId="11" pageNumber="130">
<emphasis id="B905EAF9FFD16D7FFF480D8FFE1FC8A1" bold="true" box="[199,480,1740,1766]" pageId="11" pageNumber="130">
Description of
<typeStatus id="54CA8849FFD16D7FFEF60D8FFE1FC8A1" box="[377,480,1740,1766]" pageId="11" pageNumber="130" type="holotype">holotype</typeStatus>
</emphasis>
: Juvenile female,
<quantity id="4C899B0EFFD16D7FFD3F0D8FFCF1C8A0" box="[688,782,1740,1767]" metricMagnitude="-1" metricUnit="m" metricValue="2.2199999999999998" pageId="11" pageNumber="130" unit="mm" value="222.0">222 mm</quantity>
SVL; head sub-triangular, slightly distinct from the neck, HL
<quantity id="4C899B0EFFD16D7FFF4B0DB3FED7C94C" box="[196,296,1776,1803]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="11" pageNumber="130" unit="mm" value="11.4">11.4 mm</quantity>
(5.1 % of SVL (
<quantity id="4C899B0EFFD16D7FFE690DB3FDB5C94D" box="[486,586,1776,1802]" metricMagnitude="-1" metricUnit="m" metricValue="2.2199999999999998" pageId="11" pageNumber="130" unit="mm" value="222.0">222 mm</quantity>
); interocular distance 4.0 mm, pupil elliptical, eye diameter
<quantity id="4C899B0EFFD16D7FFB790DB3FAAFC94C" box="[1270,1360,1776,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="130" unit="mm" value="2.3">2.3 mm</quantity>
; loreal rectangular, about 2.1 x as long (
<quantity id="4C899B0EFFD16D7FFD8C0C57FDA4C969" box="[515,603,1812,1839]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="130" unit="mm" value="1.5">1.5 mm</quantity>
) as high (
<quantity id="4C899B0EFFD16D7FFD470C57FCDCC969" box="[712,803,1812,1838]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="130" unit="mm" value="0.7">0.7 mm</quantity>
); body cylindrical; tail moderately short (8.5 % of SVL). Supralabials 8/8, 3th, 4th and 5th on left side of head and 4
<superScript id="7C049BA3FFD16D7FFC940C7BFCD7C901" attach="left" box="[795,808,1848,1862]" fontSize="6" pageId="11" pageNumber="130">th</superScript>
and 5
<superScript id="7C049BA3FFD16D7FFCE60C7BFC89C901" attach="left" box="[873,886,1848,1862]" fontSize="6" pageId="11" pageNumber="130">th</superScript>
on right side of head contacting orbit; infralabials 9/9, 1st on each side in contact behind mental, first three on both sides in contact with anterior chin shields and 4th on both sides in contact with posterior chin shields; one preocular on both sides touching; two postoculars in contact with the lower part of parietal scale caudally, upper postocular in contact with supraocular anterior-dorsally. Lower postocular in contact with 5th and 6th supralabial (both sides of head) ventrally; temporals 1+2+3 on both sides; two internasals; nasal divided; frontal longer (
<quantity id="4C899B0EFFD66D78FD2C0BD5FD03CEF6" box="[675,764,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="131" unit="mm" value="3.5">3.5 mm</quantity>
) than wide (
<quantity id="4C899B0EFFD66D78FC030BD4FC16CEF6" box="[908,1001,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="131" unit="mm" value="1.9">1.9 mm</quantity>
); Parietal length (
<quantity id="4C899B0EFFD66D78FB3D0BD4FAF3CEF6" box="[1202,1292,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="131" unit="mm" value="3.8">3.8 mm</quantity>
) longer than distance from frontal to rostral (
<quantity id="4C899B0EFFD66D78FD8D0BF8FDA0CE92" box="[514,607,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="131" unit="mm" value="2.3">2.3 mm</quantity>
); dorsal scales smooth, 22 SR one head-length posterior to jaw rictus, 27 MSR at midbody, 21 SR one head length anterior to vent; ventrals 225; anal plate entire; 43 subcaudals, all paired.
</paragraph>
<paragraph id="8BCE36EBFFD66D78FF480A41FB8FCF06" blockId="12.[151,1436,150,609]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF480A41FD98CF5A" bold="true" box="[199,615,258,285]" pageId="12" pageNumber="131">
Colouration of
<typeStatus id="54CA8849FFD66D78FEF40A40FE1DCF5A" box="[379,482,259,285]" pageId="12" pageNumber="131" type="holotype">holotype</typeStatus>
in ethanol:
</emphasis>
dorsal body colouration dark brown without any light head stripes. Venter whitish, no dorsal scales light coloured, upper and lower labials same colour as dorsum.
</paragraph>
<paragraph id="8BCE36EBFFD66D78FF180A09FCA3CF22" blockId="12.[151,1436,150,609]" box="[151,860,330,357]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF180A09FE30CF23" bold="true" box="[151,463,330,356]" pageId="12" pageNumber="131">
Colouration in life (
<figureCitation id="134A2A6EFFD66D78FE0E0A09FE38CF23" box="[385,455,330,356]" captionStart="FIGURE 6" captionStartId="11.[151,250,1058,1082]" captionTargetBox="[151,1436,181,1034]" captionTargetId="figure-26@11.[151,1436,181,1034]" captionTargetPageId="11" captionText="FIGURE 6. Live photo of Boaedon subniger sp. nov. from nearby Yuffleh, Gumbi Hills, Somalia. Photo by Tomáš Mazuch." figureDoi="http://doi.org/10.5281/zenodo.14704098" httpUri="https://zenodo.org/record/14704098/files/figure.png" pageId="12" pageNumber="131">Fig. 6</figureCitation>
)
</emphasis>
: as above, but colour almost black.
</paragraph>
<paragraph id="8BCE36EBFFD66D78FF480A2DFE33CFCE" blockId="12.[151,1436,150,609]" box="[199,460,366,393]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF480A2DFEBECFCF" bold="true" box="[199,321,366,392]" pageId="12" pageNumber="131">Variation:</emphasis>
see
<tableCitation id="C6F30350FFD66D78FEFD0A2CFE37CFCE" box="[370,456,367,393]" captionStart="TABLE 4" captionStartId="14.[152,244,151,175]" captionTargetPageId="14" captionText="TABLE 4. Morphometric (in mm) and meristic character comparison of Boaedon subniger sp. nov. available material, X: present, : absent or missing" pageId="12" pageNumber="131">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFD66D78FF480AD1FE28CC5E" pageId="12" pageNumber="131" type="distribution">
<paragraph id="8BCE36EBFFD66D78FF480AD1FF0DCF96" blockId="12.[151,1436,150,609]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF480AD1FE9CCFEB" bold="true" box="[199,355,402,428]" pageId="12" pageNumber="131">Distribution:</emphasis>
the species is found only in Northern
<collectingCountry id="F366767BFFD66D78FC920AD0FC86CFEA" box="[797,889,403,429]" name="Somalia" pageId="12" pageNumber="131">Somalia</collectingCountry>
(
<collectingRegion id="49B5F809FFD66D78FC050AD0FBD0CFEA" box="[906,1071,403,429]" country="Somalia" name="Sanaag" pageId="12" pageNumber="131">Sanaag region</collectingRegion>
, Somaliland (
<quantity id="4C899B0EFFD66D78FB420AD0FA94CFEA" box="[1229,1387,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.77" metricValueMax="1.78" metricValueMin="1.76" pageId="12" pageNumber="131" unit="m" value="1770.0" valueMax="1780.0" valueMin="1760.0">17601780 m</quantity>
asl) (
<figureCitation id="134A2A6EFFD66D78FF100AF4FF1ACF96" box="[159,229,439,465]" captionStart="FIGURE 2" captionStartId="5.[152,255,1846,1870]" captionTargetBox="[170,1368,516,1823]" captionTargetId="figure-160@5.[151,1436,486,1825]" captionTargetPageId="5" captionText="FIGURE 2. Confirmed records of Boaedon species in north-eastern and eastern Africa. Red dots: Boaedon broadleyi sp. nov.; Red star: Boaedon broadleyi sp. nov. type locality; green dots: Boaedon subniger sp. nov.; green star: Boaedon subniger sp. nov. type locality; red square: Boaedon montanus; red triangle: Boaedon paralineatus; blue dots, Boaedon capensis s.l.: orange dots Boaedon sp. black form; yellow dot: B. arabicus." figureDoi="http://doi.org/10.5281/zenodo.14704084" httpUri="https://zenodo.org/record/14704084/files/figure.png" pageId="12" pageNumber="131">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BCE36EBFFD66D78FF480A99FE28CC5E" blockId="12.[151,1436,150,609]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF480A99FD97CFB2" bold="true" box="[199,616,474,501]" pageId="12" pageNumber="131">Habitat and Natural History notes:</emphasis>
The specimens were found in high-altitude semiarid or arid habitats from Somaliland (
<collectingRegion id="49B5F809FFD66D78FEAA0ABCFE33CC5E" box="[293,460,511,537]" country="Somalia" name="Sanaag" pageId="12" pageNumber="131">Sanaag region</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFD66D78FF480960FD19CC26" pageId="12" pageNumber="131" type="etymology">
<paragraph id="8BCE36EBFFD66D78FF480960FD19CC26" blockId="12.[151,1436,150,609]" pageId="12" pageNumber="131">
<emphasis id="B905EAF9FFD66D78FF480960FEADCC7A" bold="true" box="[199,338,547,573]" pageId="12" pageNumber="131">Etymology:</emphasis>
The species is named after the Latin word
<taxonomicName id="4C714D68FFD66D78FCAD0960FC6DCC7A" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[802,914,547,573]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="12" pageNumber="131" phylum="Chordata" rank="species" species="subniger">“subniger</taxonomicName>
” for blackish, somewhat dark, from its blackish body coloration without any stripes on head or body.
</paragraph>
</subSubSection>
</treatment>
</document>

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Broadley Doria &amp; Wigge 2003: 236 Photo
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81 from
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<treatmentCitationGroup id="AB6111C5FFDC6D72FF180FE1FCFACAFD" box="[151,773,1186,1211]" pageId="6" pageNumber="125">
<treatmentCitation id="0AD010FAFFDC6D72FF180FE1FD55CAFD" author="Largen, M. J. &amp; Spawls, S." box="[151,682,1186,1211]" page="469" pageId="6" pageNumber="125" year="2010">
<taxonomicName id="4C714D68FFDC6D72FF180FE1FD55CAFD" ID-CoL="8J72G" authority="Largen &amp; Spawls 2010: 469" authorityName="Largen &amp; Spawls" authorityPageNumber="469" authorityYear="2010" box="[151,682,1186,1211]" class="Squamata" family="Lamprophiidae" genus="Lamprophis" kingdom="Animalia" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B905EAF9FFDC6D72FF180FE1FE79CAFD" box="[151,390,1186,1210]" italics="true" pageId="6" pageNumber="125">Lamprophis fuliginosus</emphasis>
<bibRefCitation id="EFE04B1AFFDC6D72FE020FE0FD55CAFD" author="Largen, M. J. &amp; Spawls, S." box="[397,682,1186,1211]" pageId="6" pageNumber="125" refId="ref9983" refString="Largen, M. J. &amp; Spawls, S. (2010) Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp." type="book" year="2010">Largen &amp; Spawls 2010: 469</bibRefCitation>
</taxonomicName>
</treatmentCitation>
Fig. 300
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCE36EBFFDC6D72FF180F82FCA1CA9E" blockId="6.[151,1051,1155,1241]" box="[151,862,1217,1241]" pageId="6" pageNumber="125">
<treatmentCitationGroup id="AB6111C5FFDC6D72FF180F82FCA1CA9E" box="[151,862,1217,1241]" pageId="6" pageNumber="125">
<treatmentCitation id="0AD010FAFFDC6D72FF180F82FDA1CA9E" author="Spawls, S. &amp; Mazuch, T. &amp; Mohamad, A." box="[151,606,1217,1241]" page="442" pageId="6" pageNumber="125" year="2023">
<taxonomicName id="4C714D68FFDC6D72FF180F82FDA1CA9E" ID-CoL="M7VV" authority="Spawls et al. 2023: 442" authorityName="Spawls" authorityPageNumber="442" authorityYear="2023" box="[151,606,1217,1241]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B905EAF9FFDC6D72FF180F82FE98CA9E" box="[151,359,1217,1241]" italics="true" pageId="6" pageNumber="125">Boaedon fuliginosus</emphasis>
<bibRefCitation id="EFE04B1AFFDC6D72FEE20F82FDA1CA9E" author="Spawls, S. &amp; Mazuch, T. &amp; Mohamad, A." box="[365,606,1217,1241]" pageId="6" pageNumber="125" refId="ref10919" refString="Spawls, S., Mazuch, T. &amp; Mohamad, A. (2023) Handbook of Amphibians and Reptiles of North-East Africa. London Bloomsbury Wildlife, 640 pp." type="book" year="2023">
Spawls
<emphasis id="B905EAF9FFDC6D72FE320F81FE16CA9E" box="[445,489,1217,1241]" italics="true" pageId="6" pageNumber="125">et al</emphasis>
. 2023: 442
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 443 from
<collectingRegion id="49B5F809FFDC6D72FD430F82FD00CA9E" box="[716,767,1217,1241]" country="South Sudan" name="Lakes" pageId="6" pageNumber="125">Lake</collectingRegion>
Langano
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFDC6D72FF180E44FEC6C926" pageId="6" pageNumber="125" type="materials_examined">
<paragraph id="8BCE36EBFFDC6D72FF180E44FF51CBCA" blockId="6.[151,1437,1287,2034]" pageId="6" pageNumber="125">
<materialsCitation id="3B193CB6FFDC6D72FF180E44FF51CBCA" collectingDate="2006-12" collectionCode="ZMH" collectorName="Dr. Peter Angenstein " elevation="2552" latitude="9.03" location="Addis Ababa" longLatPrecision="21" longitude="38.073334" pageId="6" pageNumber="125" specimenCode="R08485, JH 10" specimenCount="1" stateProvince="Adis Abeba" typeStatus="holotype">
<emphasis id="B905EAF9FFDC6D72FF180E44FEF2CB66" bold="true" box="[151,269,1287,1313]" pageId="6" pageNumber="125">
<typeStatus id="54CA8849FFDC6D72FF180E44FEF7CB66" box="[151,264,1287,1313]" pageId="6" pageNumber="125" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED60AE2EFFDC6D72FE9A0E4BFEACCB65" box="[277,339,1288,1314]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
<specimenCode id="DBD79E90FFDC6D72FED50E4BFE4CCB65" box="[346,435,1287,1314]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="125">R08485</specimenCode>
(DNA voucher
<specimenCode id="DBD79E90FFDC6D72FDE80E4BFD52CB66" box="[615,685,1287,1314]" collectionCode="JH" pageId="6" pageNumber="125">JH 10</specimenCode>
): an adult female from near
<collectingRegion id="49B5F809FFDC6D72FC660E44FB84CB66" box="[1001,1147,1287,1313]" country="Ethiopia" name="Adis Abeba" pageId="6" pageNumber="125">Addis Ababa</collectingRegion>
(Abada) (approx.
<geoCoordinate id="EE45502CFFDC6D72FAC40E44FF25CB01" degrees="09" direction="north" minutes="01" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="48" value="9.03">09° 01 48” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FF6E0E68FE83CB01" box="[225,380,1323,1350]" degrees="38" direction="east" minutes="04" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="24" value="38.073334">38° 04 24” E</geoCoordinate>
, 2552 m), Oromia Region, collected by
<collectorName id="2684533DFFDC6D72FCB20E6FFBD9CB02" box="[829,1062,1323,1350]" pageId="6" pageNumber="125">Dr. Peter Angenstein</collectorName>
during the end of the 1980s, kept in captivity until 2006 by
<collectorName id="2684533DFFDC6D72FE350E0CFDBFCB2E" box="[442,576,1359,1385]" pageId="6" pageNumber="125">Ralf Hörold</collectorName>
and donated to the
<collectorName id="2684533DFFDC6D72FC920E0CFBFFCB2D" box="[797,1024,1359,1386]" pageId="6" pageNumber="125">Zoological Museum</collectorName>
Hamburg (
<collectionCode id="ED60AE2EFFDC6D72FBF00E0CFB3ECB2E" box="[1151,1217,1359,1385]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
) in
<date id="FFCF102BFFDC6D72FB630E13FAA0CB2E" box="[1260,1375,1359,1386]" pageId="6" pageNumber="125" value="2006-12">
<collectingDate id="EF8BE9C3FFDC6D72FB630E13FAA0CB2E" box="[1260,1375,1359,1386]" pageId="6" pageNumber="125" value="2006-12">Dec. 2006</collectingDate>
</date>
(
<figureCitation id="134A2A6EFFDC6D72FAE00E0CFF5ACBCA" captionStart="FIGURE 3" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[210,1377,188,1920]" captionTargetId="figure-34@8.[203,1385,181,1922]" captionTargetPageId="8" captionText="FIGURE 3: Holotype of Boaedon broadleyi sp. nov. (ZMH R08485) from nearby Addis Ababa, Ethiopia. Left: head in dorsal, ventral and lateral view, right side drawings of the head." figureDoi="http://doi.org/10.5281/zenodo.14704088" httpUri="https://zenodo.org/record/14704088/files/figure.png" pageId="6" pageNumber="125">Fig. 3</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="8BCE36EBFFDC6D72FF480ED4FEC6C926" blockId="6.[151,1437,1287,2034]" pageId="6" pageNumber="125">
<materialsCitation id="3B193CB6FFDC6D72FF480ED4FA68CB92" collectionCode="MNHN" collectorName="A. M. J. Raffray" country="Ethiopia" county="Deyrolle" location="All" municipality="Henry" pageId="6" pageNumber="125" specimenCode="MNHN 4342" specimenCount="1" typeStatus="paratype">
<emphasis id="B905EAF9FFDC6D72FF480ED4FE80CBF6" bold="true" box="[199,383,1431,1457]" pageId="6" pageNumber="125">
<typeStatus id="54CA8849FFDC6D72FF480ED4FEC0CBF6" box="[199,319,1431,1457]" pageId="6" pageNumber="125" type="paratype">Paratypes</typeStatus>
(13):
</emphasis>
<location id="8EAE6030FFDC6D72FE0A0ED4FE56CBF6" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FE0A0ED4FE56CBF6" box="[389,425,1431,1457]" country="Ethiopia" name="All" pageId="6" pageNumber="125">All</location>
from
<collectingCountry id="F366767BFFDC6D72FE600ED4FDADCBF6" box="[495,594,1431,1457]" name="Ethiopia" pageId="6" pageNumber="125">Ethiopia</collectingCountry>
:
<specimenCode id="DBD79E90FFDC6D72FDD10EDBFD09CBF6" box="[606,758,1431,1458]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="125">MNHN 4342</specimenCode>
, an adult female from Ethiopia (Abyssinie) with no specific locality, received by natural produce dealer
<collectingCounty id="62AF4E67FFDC6D72FD070EF8FD15CB92" box="[648,746,1467,1493]" pageId="6" pageNumber="125">Deyrolle</collectingCounty>
(
<collectingMunicipality id="6BAAAC91FFDC6D72FD760EF8FCBECB92" box="[761,833,1467,1493]" pageId="6" pageNumber="125">Henry</collectingMunicipality>
or
<location id="8EAE6030FFDC6D72FCE70EF8FC52CB92" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FCE70EF8FC52CB92" box="[872,941,1467,1493]" country="Ethiopia" county="Deyrolle" municipality="Henry" name="Emile" pageId="6" pageNumber="125">Emile</location>
) and collected by
<collectorName id="2684533DFFDC6D72FBF20EF8FAC8CB92" box="[1149,1335,1467,1494]" pageId="6" pageNumber="125">A. M. J. Raffray</collectorName>
in 1874
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FF180EA3FCFFC859" collectingDate="1975-02-28" collectionCode="ZFMK" collectorName="H. Rupp" country="Ethiopia" elevation="1575" latitude="7.6833334" location="Sidamo Province" longLatPrecision="21" longitude="39.25" municipality="Negele" pageId="6" pageNumber="125" specimenCode="ZFMK 16231-32" specimenCount="1" stateProvince="Oromiya" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FF180EA3FEA3CBBE" box="[151,348,1503,1530]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 16231-32</specimenCode>
, adult females from
<collectingMunicipality id="6BAAAC91FFDC6D72FDC50E9CFD66CBBE" box="[586,665,1503,1529]" pageId="6" pageNumber="125">Negele</collectingMunicipality>
(approx.
<geoCoordinate id="EE45502CFFDC6D72FC860E9CFC54CBBD" box="[777,939,1503,1530]" degrees="07" direction="north" minutes="41" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="7.6833334">07° 41 00” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FC3B0E9CFBAACBBD" box="[948,1109,1503,1530]" degrees="39" direction="east" minutes="15" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="39.25">39° 15 00” E</geoCoordinate>
, 1575 m),
<location id="8EAE6030FFDC6D72FB5C0E9CFA67CBBE" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FB5C0E9CFA67CBBE" box="[1235,1432,1503,1529]" country="Ethiopia" latitude="7.6833334" longLatPrecision="21" longitude="39.25" municipality="Negele" name="Sidamo Province" pageId="6" pageNumber="125" stateProvince="Oromiya">Sidamo Province</location>
,
<collectingRegion id="49B5F809FFDC6D72FF180D40FEB5C85A" box="[151,330,1539,1565]" country="Ethiopia" name="Oromiya" pageId="6" pageNumber="125">Oromia Region</collectingRegion>
, collected by
<collectorName id="2684533DFFDC6D72FE630D47FDB0C859" box="[492,591,1540,1566]" pageId="6" pageNumber="125">H. Rupp</collectorName>
,
<date id="FFCF102BFFDC6D72FDD10D40FCFFC859" box="[606,768,1539,1566]" pageId="6" pageNumber="125" value="1975-02-28">
<collectingDate id="EF8BE9C3FFDC6D72FDD10D40FCFFC859" box="[606,768,1539,1566]" pageId="6" pageNumber="125" value="1975-02-28">28. Feb. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FC810D47FB77C806" collectingDate="1973-09" collectionCode="ZFMK" collectorName="H. Rupp" country="Ethiopia" elevation="1595" latitude="8.391944" location="Lake Koka" longLatPrecision="21" longitude="39.083332" pageId="6" pageNumber="125" specimenCode="ZFMK 16293" specimenCount="1" stateProvince="Oromiya" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FC810D47FC51C85A" box="[782,942,1539,1566]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 16293</specimenCode>
, an adult female from
<location id="8EAE6030FFDC6D72FB380D40FACAC85A" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FB380D40FACAC85A" box="[1207,1333,1539,1565]" country="Ethiopia" latitude="8.391944" longLatPrecision="21" longitude="39.083332" name="Lake Koka" pageId="6" pageNumber="125" stateProvince="Oromiya">Lake Koka</location>
(approx.
<geoCoordinate id="EE45502CFFDC6D72FF180D64FEC9C805" box="[151,310,1575,1602]" degrees="08" direction="north" minutes="23" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="31" value="8.391944">08° 23 31” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FEB10D64FE24C805" box="[318,475,1575,1602]" degrees="39" direction="east" minutes="05" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="39.083332">39° 05 00” E</geoCoordinate>
, 1595 m)
<collectingRegion id="49B5F809FFDC6D72FDC00D64FCFEC806" box="[591,769,1575,1601]" country="Ethiopia" name="Oromiya" pageId="6" pageNumber="125">Oromia Region</collectingRegion>
, collected by
<collectorName id="2684533DFFDC6D72FC120D6BFC00C805" box="[925,1023,1576,1602]" pageId="6" pageNumber="125">H. Rupp</collectorName>
,
<date id="FFCF102BFFDC6D72FB830D64FB77C806" box="[1036,1160,1575,1601]" pageId="6" pageNumber="125" value="1973-09">
<collectingDate id="EF8BE9C3FFDC6D72FB830D64FB77C806" box="[1036,1160,1575,1601]" pageId="6" pageNumber="125" value="1973-09">Sept. 1973</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FB1B0D6BFC33C8CD" collectingDate="1975-03-26" collectionCode="ZFMK" collectorName="H. Rupp" country="Ethiopia" elevation="1285" latitude="6.0333333" location="Peoples Region" longLatPrecision="21" longitude="37.55" municipality="Arba Minch" pageId="6" pageNumber="125" specimenCode="ZFMK 16294-95" specimenCount="1" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FB1B0D6BFAA7C805" box="[1172,1368,1575,1602]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 16294-95</specimenCode>
, adult female and juvenile respectively from
<collectingMunicipality id="6BAAAC91FFDC6D72FDB00D08FD39C822" box="[575,710,1611,1637]" pageId="6" pageNumber="125">Arba Minch</collectingMunicipality>
(approx.
<geoCoordinate id="EE45502CFFDC6D72FCBF0D08FC37C821" box="[816,968,1611,1638]" degrees="06" direction="north" minutes="02" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="6.0333333">06° 02 00” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FC410D08FB99C821" box="[974,1126,1611,1638]" degrees="37" direction="east" minutes="33" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="37.55">37° 33 00” E</geoCoordinate>
, 1285 m)
<collectingRegion id="49B5F809FFDC6D72FB5C0D08FA67C822" box="[1235,1432,1611,1637]" country="Ethiopia" name="Ye Debub Biheroch Bihereseboch na Hizboch" pageId="6" pageNumber="125">Southern Nations</collectingRegion>
, Nationalities, and
<location id="8EAE6030FFDC6D72FEE80D2CFDE5C8CE" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FEE80D2CFDE5C8CE" box="[359,538,1647,1673]" country="Ethiopia" latitude="6.0333333" longLatPrecision="21" longitude="37.55" municipality="Arba Minch" name="Peoples Region" pageId="6" pageNumber="125" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch">Peoples Region</location>
, collected by
<collectorName id="2684533DFFDC6D72FD360D33FCE4C8CD" box="[697,795,1648,1674]" pageId="6" pageNumber="125">H. Rupp</collectorName>
,
<date id="FFCF102BFFDC6D72FCA60D2CFC33C8CD" box="[809,972,1647,1674]" pageId="6" pageNumber="125" value="1975-03-26">
<collectingDate id="EF8BE9C3FFDC6D72FCA60D2CFC33C8CD" box="[809,972,1647,1674]" pageId="6" pageNumber="125" value="1975-03-26">26. Mar. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FC560D33FECCC895" collectingDate="1975-11-20" collectionCode="ZFMK" collectorName="G. Nikolaus" country="Ethiopia" elevation="3000" latitude="8.933333" location="Tulu Welel mountains" longLatPrecision="21" longitude="35.05" municipality="Kaki" pageId="6" pageNumber="125" specimenCode="ZFMK 16799" specimenCount="1" stateProvince="Oromiya" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FC560D33FB87C8CE" box="[985,1144,1647,1674]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 16799</specimenCode>
, an adult male from
<collectingMunicipality id="6BAAAC91FFDC6D72FAE90D2CFA63C8CE" box="[1382,1436,1647,1673]" pageId="6" pageNumber="125">Kaki</collectingMunicipality>
(approx.
<geoCoordinate id="EE45502CFFDC6D72FF720DD0FE62C8E9" box="[253,413,1683,1710]" degrees="08" direction="north" minutes="56" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="8.933333">08° 56 00” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FE2A0DD0FDBCC8E9" box="[421,579,1683,1710]" degrees="35" direction="east" minutes="03" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="35.05">35° 03 00” E</geoCoordinate>
, 3000 m)
<location id="8EAE6030FFDC6D72FD380DD0FC4DC8EA" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FD380DD0FC4DC8EA" box="[695,946,1683,1709]" country="Ethiopia" latitude="8.933333" longLatPrecision="21" longitude="35.05" municipality="Kaki" name="Tulu Welel mountains" pageId="6" pageNumber="125" stateProvince="Oromiya">Tulu Welel mountains</location>
, Oromia Region, collected by
<collectorName id="2684533DFFDC6D72FA820DD0FA67C8EA" box="[1293,1432,1683,1709]" pageId="6" pageNumber="125">G. Nikolaus</collectorName>
,
<date id="FFCF102BFFDC6D72FF180DF4FECCC895" box="[151,307,1719,1746]" pageId="6" pageNumber="125" value="1975-11-20">
<collectingDate id="EF8BE9C3FFDC6D72FF180DF4FECCC895" box="[151,307,1719,1746]" pageId="6" pageNumber="125" value="1975-11-20">20.Nov. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FECF0DFBFEB6C8B2" collectionCode="ZFMK" country="Ethiopia" elevation="825" latitude="11.603945" location="Sifra" longLatPrecision="1" longitude="40.20996" municipality="Chifra" pageId="6" pageNumber="125" specimenCode="ZFMK 19159" specimenCount="1" stateProvince="Afar" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FECF0DFBFE20C895" box="[320,479,1719,1746]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 19159</specimenCode>
, an adult female from
<collectingMunicipality id="6BAAAC91FFDC6D72FD6E0DF4FCD6C896" box="[737,809,1719,1745]" pageId="6" pageNumber="125">Chifra</collectingMunicipality>
(
<location id="8EAE6030FFDC6D72FCB70DF4FC8DC896" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FCB70DF4FC8DC896" box="[824,882,1719,1745]" country="Ethiopia" latitude="11.603945" longLatPrecision="1" longitude="40.20996" municipality="Chifra" name="Sifra" pageId="6" pageNumber="125" stateProvince="Afar">Sifra</location>
) (approx.
<geoCoordinate id="EE45502CFFDC6D72FC670DF4FB64C895" box="[1000,1179,1719,1746]" degrees="11" direction="north" minutes="36" orientation="latitude" pageId="6" pageNumber="125" precision="1" seconds="14.2" value="11.603945">11° 36 14.2” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FB2B0DF4FA9AC895" box="[1188,1381,1719,1746]" degrees="40" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="125" precision="1" seconds="35.86" value="40.20996">40° 12 35.86” E</geoCoordinate>
, 825 m)
<collectingRegion id="49B5F809FFDC6D72FF340D98FEB6C8B2" box="[187,329,1755,1781]" country="Ethiopia" name="Afar" pageId="6" pageNumber="125">Afar Region</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FEDA0D9FFA67C8B2" box="[341,1432,1755,1782]" collectionCode="ZFMK" country="Ethiopia" elevation="1634" latitude="9.666667" location="Jijiga" longLatPrecision="21" longitude="38.75" pageId="6" pageNumber="125" specimenCode="ZFMK 84994" specimenCount="1" stateProvince="Sumale" typeStatus="paratype">
<specimenCode id="DBD79E90FFDC6D72FEDA0D9FFE10C8B2" box="[341,495,1755,1782]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="125" type="Museum">ZFMK 84994</specimenCode>
a juvenile from
<location id="8EAE6030FFDC6D72FD280D98FD1BC8B2" LSID="urn:lsid:plazi:treatment:03D887FDFFDC6D7DFF180F58FE64CBCC:8EAE6030FFDC6D72FD280D98FD1BC8B2" box="[679,740,1755,1781]" country="Ethiopia" latitude="9.666667" longLatPrecision="21" longitude="38.75" name="Jijiga" pageId="6" pageNumber="125" stateProvince="Sumale">Jijiga</location>
(approx.
<geoCoordinate id="EE45502CFFDC6D72FCC00D98FC23C8B1" box="[847,988,1755,1782]" degrees="9" direction="north" minutes="40" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="9.666667">9° 40 00” N</geoCoordinate>
<geoCoordinate id="EE45502CFFDC6D72FC6C0D98FB82C8B1" box="[995,1149,1755,1782]" degrees="38" direction="east" minutes="45" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="00" value="38.75">38° 45 00” E</geoCoordinate>
, 1634 m)
<collectingRegion id="49B5F809FFDC6D72FB620D98FA67C8B2" box="[1261,1432,1755,1781]" country="Ethiopia" name="Sumale" pageId="6" pageNumber="125">Somali Region</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B193CB6FFDC6D72FF180C43FD13C95E" box="[151,748,1791,1818]" collectingDate="2006-12" collectionCode="ZMH" elevation="2552" latitude="9.03" location="Addis Ababa" longLatPrecision="21" longitude="38.073334" pageId="6" pageNumber="125" specimenCode="R08486" specimenCount="1" stateProvince="Adis Abeba" typeStatus="paratype">
<collectionCode id="ED60AE2EFFDC6D72FF180C43FF2AC95D" box="[151,213,1792,1818]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
<specimenCode id="DBD79E90FFDC6D72FF510C43FEC6C95E" box="[222,313,1791,1818]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="125">R08486</specimenCode>
, an adult male, same data as holotype
</materialsCitation>
,
<materialsCitation id="3B193CB6FFDC6D72FD760C43FEB0C97A" collectingDate="2006-12" collectionCode="ZMH" elevation="2552" latitude="9.03" location="Addis Ababa" longLatPrecision="21" longitude="38.073334" pageId="6" pageNumber="125" specimenCode="R11333, JH 59" specimenCount="1" stateProvince="Adis Abeba" typeStatus="paratype">
<collectionCode id="ED60AE2EFFDC6D72FD760C43FCC8C95D" box="[761,823,1792,1818]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
<specimenCode id="DBD79E90FFDC6D72FCB00C43FC68C95E" box="[831,919,1791,1818]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="125">R11333</specimenCode>
(DNA voucher
<specimenCode id="DBD79E90FFDC6D72FBC00C43FB6AC95D" box="[1103,1173,1791,1818]" collectionCode="JH" pageId="6" pageNumber="125">JH 59</specimenCode>
), an adult female same data as holotype
</materialsCitation>
,
<materialsCitation id="3B193CB6FFDC6D72FED50C67FECAC926" collectingDate="2006-12" collectionCode="ZMH" elevation="2552" latitude="9.03" location="Addis Ababa" longLatPrecision="21" longitude="38.073334" pageId="6" pageNumber="125" specimenCode="R11341-42" specimenCount="1" stateProvince="Adis Abeba" typeStatus="paratype">
<collectionCode id="ED60AE2EFFDC6D72FED50C67FE67C979" box="[346,408,1828,1854]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
<specimenCode id="DBD79E90FFDC6D72FE110C67FDE1C97A" box="[414,542,1827,1854]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="125">R11341-42</specimenCode>
, an adult male and female respectively, same data as holotype but donated to the
<collectionCode id="ED60AE2EFFDC6D72FF180C0BFF2AC925" box="[151,213,1864,1890]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="6" pageNumber="125" type="Museum">ZMH</collectionCode>
in 2019
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFDC6D73FF480C28FC15CA4D" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125" type="diagnosis">
<paragraph id="8BCE36EBFFDC6D73FF480C28FCF4CF5B" blockId="6.[151,1437,1287,2034]" lastBlockId="7.[151,1437,151,2036]" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125">
<emphasis id="B905EAF9FFDC6D72FF480C28FEBDC9C2" bold="true" box="[199,322,1899,1925]" pageId="6" pageNumber="125">Diagnosis.</emphasis>
<taxonomicName id="4C714D68FFDC6D72FEC60C28FDE2C9C2" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[329,541,1899,1925]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDC6D72FEC60C28FDE2C9C2" box="[329,541,1899,1925]" italics="true" pageId="6" pageNumber="125">Boaedon broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDC6D72FDAA0C28FD81C9C2" bold="true" box="[549,638,1899,1925]" pageId="6" pageNumber="125">
<taxonomicNameLabel id="A2365782FFDC6D72FDAA0C28FD81C9C2" box="[549,638,1899,1925]" pageId="6" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from most congeners by a beige to light brown to reddish ground colour, with two short white stripes on the sides of the head. The upper stripe begins just behind the eye on the upper postocular, runs through the lower part of the parietal and the upper first temporal and terminates on the second temporal; the lower stripe passes from the lower postocular through the first temporal and terminates on the last supralabial. Frequently three supralabials contact the eye and two preoculars, the upper one much larger than the lower one touching the frontal. PAR is equal to the distance between the frontal and the rostral scale. Middorsal scale rows varies between 27 and 33 (mean 29.1), ventrals are
<quantity id="4C899B0EFFDD6D73FC490B9DFBBACEBF" box="[966,1093,222,248]" metricMagnitude="0" metricUnit="m" metricValue="5.8801" metricValueMax="6.096" metricValueMin="5.664199999999999" pageId="7" pageNumber="126" unit="in" value="231.5" valueMax="240.0" valueMin="223.0">223240 in</quantity>
females and
<quantity id="4C899B0EFFDD6D73FB580B9DFAAACEBF" box="[1239,1365,222,248]" metricMagnitude="0" metricUnit="m" metricValue="5.2705" metricValueMax="5.6896" metricValueMin="4.8514" pageId="7" pageNumber="126" unit="in" value="207.5" valueMax="224.0" valueMin="191.0">191224 in</quantity>
males and subcaudals 4555 and 5963 respectively. (
<tableCitation id="C6F30350FFDD6D73FD200A41FCFECF5B" box="[687,769,258,284]" captionStart="TABLE 3" captionStartId="13.[150,242,543,569]" captionTargetPageId="13" captionText="TABLE 3. Morphometric (in mm) and meristic character comparison of Boaedon broadleyi sp. nov. available material, X: present, : absent or missing" pageId="7" pageNumber="126">Table 3</tableCitation>
)
</paragraph>
<paragraph id="8BCE36EBFFDD6D73FF480A66FC15CA4D" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="126">
Within Ethiopian
<taxonomicName id="4C714D68FFDD6D73FE020A66FE0ECF78" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[397,497,293,319]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="genus">
<emphasis id="B905EAF9FFDD6D73FE020A66FE0ECF78" box="[397,497,293,319]" italics="true" pageId="7" pageNumber="126">Boaedon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C714D68FFDD6D73FE710A66FD35CF78" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[510,714,293,319]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadley" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FE710A66FD35CF78" box="[510,714,293,319]" italics="true" pageId="7" pageNumber="126">Boaedon broadley</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FD5F0A65FCD6CF07" bold="true" box="[720,809,294,320]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FD5F0A65FCD6CF07" box="[720,809,294,320]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from Ethiopian
<emphasis id="B905EAF9FFDD6D73FBA20A65FBB9CF78" box="[1069,1094,294,319]" italics="true" pageId="7" pageNumber="126">B.</emphasis>
sp. (black form) and Somalian
<taxonomicName id="4C714D68FFDD6D73FF180A09FEE3CF24" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[151,284,329,356]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FF180A09FF57CF24" box="[151,168,330,355]" italics="true" pageId="7" pageNumber="126">B</emphasis>
.
<emphasis id="B905EAF9FFDD6D73FF350A0AFEE3CF24" box="[186,284,329,355]" italics="true" pageId="7" pageNumber="126">subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FEA80A0AFE7BCF24" bold="true" box="[295,388,329,355]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FEA80A0AFE7BCF24" box="[295,388,329,355]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the combination of its reddish, olive-brown or light brown body colouration with two short head stripes (versus blackish-brown or dark brown body colouration and head stripes missing) with the first two body scale rows uncoloured (versus no dorsals uncoloured) and a higher average of mid-dorsal scale counts of 29.1 (versus 27.0 in
<taxonomicName id="4C714D68FFDD6D73FE230AF6FDCFCF89" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[428,560,436,462]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="subniger" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FE230AF6FDCFCF89" box="[428,560,436,462]" italics="true" pageId="7" pageNumber="126">B. subniger</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FDB40AF6FD67CF88" bold="true" box="[571,664,437,463]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FDB40AF6FD67CF88" box="[571,664,437,463]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C714D68FFDD6D73FD3D0AF7FC6CCF89" authorityName="Parker" authorityYear="1932" box="[690,915,436,462]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B905EAF9FFDD6D73FD3D0AF7FC6CCF89" box="[690,915,436,462]" italics="true" pageId="7" pageNumber="126">Boaedon maculatus</emphasis>
</taxonomicName>
has an additional row of scales between the supralabials and loreal between nasal and preocular scale (versus no row of scales) and 1011 supralabials (versus 8). Extralimital species:
<taxonomicName id="4C714D68FFDD6D73FE270ABFFCA8CC51" authority="(Trape et al. 2022)" baseAuthorityName="Trape" baseAuthorityYear="2022" box="[424,855,508,535]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B905EAF9FFDD6D73FE270ABFFD7FCC51" box="[424,640,508,534]" italics="true" pageId="7" pageNumber="126">Boaedon montanus</emphasis>
(
<bibRefCitation id="EFE04B1AFFDD6D73FD1F0ABEFCB0CC51" author="Trape, J. - F. &amp; Mediannikov, O. &amp; Hinkel, H. &amp; Hinkel, H." box="[656,847,508,535]" pageId="7" pageNumber="126" pagination="1 - 12" refId="ref11237" refString="Trape, J. - F., Mediannikov, O., Hinkel, H. &amp; Hinkel, H. (2022) Un serpent nouveau du genre Boaedon Dumeril, Bibron &amp; Dumeril, 1854 (Squamata: Lamprophiidae) dans la region des Grands Lacs africains. Bulletin de la Societe Herpetologique de France, 181, 1 - 12." type="journal article" year="2022">
Trape
<emphasis id="B905EAF9FFDD6D73FD580ABEFCF0CC51" box="[727,783,508,534]" italics="true" pageId="7" pageNumber="126">et al.</emphasis>
2022
</bibRefCitation>
)
</taxonomicName>
differs genetically from
<taxonomicName id="4C714D68FFDD6D73FBE40ABEFB0BCC51" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[1131,1268,508,534]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FBE40ABEFB0BCC51" box="[1131,1268,508,534]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FB740ABFFAAACC51" bold="true" box="[1275,1365,508,534]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FB740ABFFAAACC51" box="[1275,1365,508,534]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
There is no differential diagnostic character to distinguish the species of
<taxonomicName id="4C714D68FFDD6D73FCF00963FBF4CC7E" box="[895,1035,544,569]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B905EAF9FFDD6D73FCF00963FBF4CC7E" box="[895,1035,544,569]" italics="true" pageId="7" pageNumber="126">B. montanus</emphasis>
</taxonomicName>
in its hole distribution area from
<taxonomicName id="4C714D68FFDD6D73FA0B0963FEFECC1A" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi">
<emphasis id="B905EAF9FFDD6D73FA0B0963FEFECC1A" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
because all meristic characters vary in different populations of this species. Within the
<typeStatus id="54CA8849FFDD6D73FB6B0907FAEBCC19" box="[1252,1300,580,606]" pageId="7" pageNumber="126">type</typeStatus>
series of
<taxonomicName id="4C714D68FFDD6D73FA0B0907FEFBCCC6" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B905EAF9FFDD6D73FA0B0907FEFBCCC6" italics="true" pageId="7" pageNumber="126">B. montanus</emphasis>
</taxonomicName>
one character is different: no contact of the upper preocular with the frontal (versus contact) and PAR longer than distance of frontal to rostral (versus PAR equal to distance from frontal to rostral). Specimens of the
<collectingCountry id="F366767BFFDD6D73FF1809ECFF11CC8E" box="[151,238,687,713]" name="Uganda" pageId="7" pageNumber="126">Uganda</collectingCountry>
clade (
<taxonomicName id="4C714D68FFDD6D73FECA09ECFE2CCC8F" box="[325,467,687,713]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B905EAF9FFDD6D73FECA09ECFEA9CC8F" box="[325,342,687,712]" italics="true" pageId="7" pageNumber="126">B</emphasis>
.
<emphasis id="B905EAF9FFDD6D73FEE909ECFE2CCC8F" box="[358,467,687,712]" italics="true" pageId="7" pageNumber="126">montanus</emphasis>
</taxonomicName>
) and from
<collectingCountry id="F366767BFFDD6D73FDD809ECFD18CC8E" box="[599,743,687,713]" name="South Sudan" pageId="7" pageNumber="126">South Sudan</collectingCountry>
(
<taxonomicName id="4C714D68FFDD6D73FD7609ECFC72CC8F" box="[761,909,687,713]" class="Squamata" family="Lamprophiidae" genus="Boaedon" isUncertain="true" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="montanus">
cf.
<emphasis id="B905EAF9FFDD6D73FCAF09ECFC72CC8F" box="[800,909,687,712]" italics="true" pageId="7" pageNumber="126">montanus</emphasis>
</taxonomicName>
) predominantly have a dark brown coloration (versus light brown in
<taxonomicName id="4C714D68FFDD6D73FE2D0991FDF3CCAB" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[418,524,722,748]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi">
<emphasis id="B905EAF9FFDD6D73FE2D0991FDF3CCAB" box="[418,524,722,748]" italics="true" pageId="7" pageNumber="126">broadleyi</emphasis>
</taxonomicName>
) and, variably, the preocular is separate from the frontal or in contact with it, predominantly two supralabials touching the eye (91%) (versus 55% in
<taxonomicName id="4C714D68FFDD6D73FC4E09B5FBD4CD57" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[961,1067,758,784]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FC4E09B5FBD4CD57" box="[961,1067,758,784]" italics="true" pageId="7" pageNumber="126">broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FBBC09B5FB72CD57" bold="true" box="[1075,1165,758,784]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FBBC09B5FB72CD57" box="[1075,1165,758,784]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and frequently a lower mean of MSR of 28.0
<quantity id="4C899B0EFFDD6D73FE130859FE15CD73" box="[412,490,794,820]" metricMagnitude="-1" metricUnit="m" metricValue="7.3152" pageId="7" pageNumber="126" unit="in" value="28.8">28.8 in</quantity>
all populations (versus
<quantity id="4C899B0EFFDD6D73FD760859FCC0CD73" box="[761,831,794,820]" metricMagnitude="-1" metricUnit="m" metricValue="7.3914" pageId="7" pageNumber="126" unit="in" value="29.1">29.1in</quantity>
<taxonomicName id="4C714D68FFDD6D73FCC80858FC2ECD73" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[839,977,794,820]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FCC80858FC2ECD73" box="[839,977,794,820]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FC560859FBCBCD73" bold="true" box="[985,1076,794,820]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FC560859FBCBCD73" box="[985,1076,794,820]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (table 2).
<taxonomicName id="4C714D68FFDD6D73FB3D0858FA9CCD73" authorityName="Trape &amp; Mediannikov" authorityYear="2016" box="[1202,1379,794,820]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="perisilvestris">
<emphasis id="B905EAF9FFDD6D73FB3D0858FA9CCD73" box="[1202,1379,794,820]" italics="true" pageId="7" pageNumber="126">B. perisilvestris</emphasis>
</taxonomicName>
with dark brown colouration and no clear head or body lines (versus headlines present and light brown or olive brown colour in
<taxonomicName id="4C714D68FFDD6D73FE8E0821FE76CD3C" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[257,393,865,891]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FE8E0821FE76CD3C" box="[257,393,865,891]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FE000822FE17CD3C" bold="true" box="[399,488,865,891]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FE000822FE17CD3C" box="[399,488,865,891]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) has PAR longer than distance of margin of frontal to end of snout (versus PAR equal in
<taxonomicName id="4C714D68FFDD6D73FF3B08C5FEC2CDD8" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[180,317,901,927]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FF3B08C5FEC2CDD8" box="[180,317,901,927]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FECB08C6FE61CDD8" bold="true" box="[324,414,901,927]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FECB08C6FE61CDD8" box="[324,414,901,927]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C714D68FFDD6D73FE3A08C5FD9DCDD8" authorityName="Trape &amp; Mediannikov" authorityYear="2016" box="[437,610,901,927]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="paralineatus">
<emphasis id="B905EAF9FFDD6D73FE3A08C5FD9DCDD8" box="[437,610,901,927]" italics="true" pageId="7" pageNumber="126">B. paralineatus</emphasis>
</taxonomicName>
differs in having clearly marginated light headlines anterior and posterior to eye and on anterior part of body (versus short head stripes not well contrasted and no body line in
<taxonomicName id="4C714D68FFDD6D73FA8608EAFA8CCD84" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[1289,1395,937,963]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FA8608EAFA8CCD84" box="[1289,1395,937,963]" italics="true" pageId="7" pageNumber="126">broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FAF408EAFF37CDA0" bold="true" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FAF408EAFF37CDA0" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
);
<taxonomicName id="4C714D68FFDD6D73FF6F088EFEA1CDA1" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[224,350,973,998]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B905EAF9FFDD6D73FF6F088EFEA1CDA1" box="[224,350,973,998]" italics="true" pageId="7" pageNumber="126">B. capensis</emphasis>
</taxonomicName>
species complex differs in having loreal length/high ratio mainly 2 or lower (versus 2 and higher) and a dorsal scale count mean of 27.0 (versus
<quantity id="4C899B0EFFDD6D73FD1308B3FD15CA4D" box="[668,746,1008,1034]" metricMagnitude="-1" metricUnit="m" metricValue="7.3914" pageId="7" pageNumber="126" unit="in" value="29.1">29.1 in</quantity>
<taxonomicName id="4C714D68FFDD6D73FD7F08B2FC86CA4D" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[752,889,1008,1034]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FD7F08B2FC86CA4D" box="[752,889,1008,1034]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FC0E08B3FC25CA4D" bold="true" box="[897,986,1008,1034]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FC0E08B3FC25CA4D" box="[897,986,1008,1034]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFDD6D73FF480F57FD7EC9B4" pageId="7" pageNumber="126" type="description">
<paragraph id="8BCE36EBFFDD6D73FF480F57FDBFCB9C" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="126">
<emphasis id="B905EAF9FFDD6D73FF480F57FE23CA69" bold="true" box="[199,476,1044,1070]" pageId="7" pageNumber="126">
Description of
<typeStatus id="54CA8849FFDD6D73FEFA0F57FE23CA69" box="[373,476,1044,1070]" pageId="7" pageNumber="126" type="holotype">holotype</typeStatus>
</emphasis>
: Adult female,
<quantity id="4C899B0EFFDD6D73FD070F57FD0ECA68" box="[648,753,1044,1071]" metricMagnitude="0" metricUnit="m" metricValue="1.196" pageId="7" pageNumber="126" unit="mm" value="1196.0">1196 mm</quantity>
SVL; head sub-triangular, slightly distinct from the neck, HL 3.3 % of SVL (
<quantity id="4C899B0EFFDD6D73FECA0F7BFE4FCA15" box="[325,432,1080,1106]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="7" pageNumber="126" unit="mm" value="39.6">39.6 mm</quantity>
); interocular distance
<quantity id="4C899B0EFFDD6D73FD250F7BFCEECA15" box="[682,785,1080,1106]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="7" pageNumber="126" unit="mm" value="13.7">13.7 mm</quantity>
, pupil elliptical, eye diameter
<quantity id="4C899B0EFFDD6D73FBE50F7BFB3BCA15" box="[1130,1220,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="126" unit="mm" value="3.2">3.2 mm</quantity>
; loreal rectangular, about 2.8 x as long (
<quantity id="4C899B0EFFDD6D73FEF50F18FE43CA31" box="[378,444,1115,1142]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="126" unit="mm" value="5.0">5 mm</quantity>
) as high (
<quantity id="4C899B0EFFDD6D73FDA60F1FFD7CCA31" box="[553,643,1116,1142]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="126" unit="mm" value="1.8">1.8 mm</quantity>
); body cylindrical; tail moderately short (10.3 % of SVL). Supralabials 8/8, 3rd, 4th and 5th on both sides contacting orbit; infralabials 9/10, 1st on each side in contact behind mental, first three on both sides in contact with anterior chin shields and 4th on both sides in contact with posterior chin shields; 2 preoculars on both sides, upper one more than two times larger than lower and touching frontal on the left side, on the right head side it is separated from the frontal by the contact of prefrontal with supraocular; 2 postoculars in contact with the lower part of parietal scale caudally, upper postocular in contact with supraocular anterior-dorsally. Lower postocular in contact with 5
<superScript id="7C049BA3FFDD6D73FDA60E71FDC9CB07" attach="left" box="[553,566,1330,1344]" fontSize="6" pageId="7" pageNumber="126">th</superScript>
and 6
<superScript id="7C049BA3FFDD6D73FDF00E71FD73CB07" attach="left" box="[639,652,1330,1344]" fontSize="6" pageId="7" pageNumber="126">th</superScript>
supralabial (both sides of head) ventrally; temporals 1+2+3 on both sides; two internasals; nasal divided; frontal longer (
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) than wide (
<quantity id="4C899B0EFFDD6D73FC350E15FBEACB37" box="[954,1045,1366,1392]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="126" unit="mm" value="4.4">4.4 mm</quantity>
); on anterior left border of frontal a small scale is intercalated between frontal and upper preocular; dorsal scales smooth, 22 scale rows one head-length posterior to jaw rictus, 29 scale rows at midbody, 21 scale rows one head length anterior to vent; ventrals 243; anal plate entire; 53 subcaudals, all paired.
</paragraph>
<paragraph id="8BCE36EBFFDD6D73FF480EA7FA9DC800" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="126">
<emphasis id="B905EAF9FFDD6D73FF480EA7FDDCCBB9" bold="true" box="[199,547,1508,1535]" pageId="7" pageNumber="126">
Coloration of
<typeStatus id="54CA8849FFDD6D73FEE30EA6FE2CCBB8" box="[364,467,1509,1535]" pageId="7" pageNumber="126" type="holotype">holotype</typeStatus>
in life.
</emphasis>
The coloration is light olive with a creamy venter, first dorsal scale row creamy. Two short white strips on side of head continuing only on head to the end of supralabials and temporals respectively. In
<emphasis id="B905EAF9FFDD6D73FF390D6FFEB6C801" bold="true" box="[182,329,1580,1606]" pageId="7" pageNumber="126">preservative</emphasis>
as above but body and head light brown above, first lower dorsal scale row creamy as venter.
</paragraph>
<paragraph id="8BCE36EBFFDD6D73FF480D0CFD7EC9B4" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="126">
<emphasis id="B905EAF9FFDD6D73FF480D0CFEC0C82E" bold="true" box="[199,319,1615,1641]" pageId="7" pageNumber="126">Variation.</emphasis>
Variation in measurements and scalation of the
<typeStatus id="54CA8849FFDD6D73FCE60D13FC2BC82D" box="[873,980,1616,1642]" pageId="7" pageNumber="126" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C714D68FFDD6D73FC710D13FB76C82D" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[1022,1161,1616,1642]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FC710D13FB76C82D" box="[1022,1161,1616,1642]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FB1D0D13FB12C82D" bold="true" box="[1170,1261,1616,1642]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FB1D0D13FB12C82D" box="[1170,1261,1616,1642]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is presented in
<tableCitation id="C6F30350FFDD6D73FF180D30FF11C8C9" box="[151,238,1651,1678]" captionStart="TABLE 3" captionStartId="13.[150,242,543,569]" captionTargetPageId="13" captionText="TABLE 3. Morphometric (in mm) and meristic character comparison of Boaedon broadleyi sp. nov. available material, X: present, : absent or missing" pageId="7" pageNumber="126">Table 3</tableCitation>
. The ground colour of the
<typeStatus id="54CA8849FFDD6D73FDAD0D37FD72C8C9" box="[546,653,1652,1678]" pageId="7" pageNumber="126" type="paratype">paratypes</typeStatus>
in preservative is predominantly light brown in 12 of
<specimenCount id="9D77FD62FFDD6D73FB730D37FA66C8CA" box="[1276,1433,1651,1678]" count="13" pageId="7" pageNumber="126" type="generic">13 specimens</specimenCount>
, one is greyish. Lower 13 dorsal scale rows have the same colouration as venter, creamy or whitish. All
<typeStatus id="54CA8849FFDD6D73FABE0DDBFA63C8F5" box="[1329,1436,1688,1714]" pageId="7" pageNumber="126" type="paratype">paratypes</typeStatus>
have two short white head lines, the extension of the upper head line varies a little, in some specimens it begins on prefrontals runs through preoculars, supraocular, upper postocular, lower part of parietals and terminate on second or third temporals. In others, it begins on upper postocular and runs until first or third temporals. In 5 of
<specimenCount id="9D77FD62FFDD6D73FA800C41FA63C95A" box="[1295,1436,1794,1821]" count="13" pageId="7" pageNumber="126" type="generic" typeStatus="paratype">13 paratypes</specimenCount>
there are three supralabials in contact with the eye. In others only two supralabials are in contact. In all
<typeStatus id="54CA8849FFDD6D73FABE0C64FA63C906" box="[1329,1436,1831,1857]" pageId="7" pageNumber="126" type="paratype">paratypes</typeStatus>
the upper preocular is in contact with the frontal and two postoculars are in contact with the descending part of the parietal scale. There exist some photo vouchers by Vladimír Tralin from Langano and Filtu which seems to belong to the new species. The colour in life varies from beige, light brown to almost brown. The brown coloured specimen (TMHC 2021.05.976) with two short light headlines from Yabello (DNA voucher TMHC 2021.05.976) belongs definitively to
<taxonomicName id="4C714D68FFDD6D73FEB30C99FE3AC9B4" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[316,453,2009,2035]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="7" pageNumber="126" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFDD6D73FEB30C99FE3AC9B4" box="[316,453,2009,2035]" italics="true" pageId="7" pageNumber="126">B. broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFDD6D73FE430C9AFDD9C9B4" bold="true" box="[460,550,2009,2035]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A2365782FFDD6D73FE430C9AFDD9C9B4" box="[460,550,2009,2035]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="134A2A6EFFDD6D73FDBA0C9AFD8FC9B4" box="[565,624,2009,2035]" captionStart="FIGURE 1" captionStartId="3.[152,255,1910,1934]" captionTargetBox="[390,1185,183,1880]" captionTargetId="graphics-29@3.[390,930,184,1833]" captionTargetPageId="3" captionText="FIGURE 1. Phylogenetic tree (BI/ML) based on the mitochondrial 16S dataset. Numbers at nodes are support values for Bayesian posterior probabilities (above nodes) and ML bootstraps (below node); values below PP &lt;0.5 and ML &lt;50 are not shown. High support values (PP ≥ 0.95 and ML ≥ 70) are marked in bold." figureDoi="http://doi.org/10.5281/zenodo.14704082" httpUri="https://zenodo.org/record/14704082/files/figure.png" pageId="7" pageNumber="126">Fig 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0E6663FFD26D7CFF170CD4FD2EC993" ID-DOI="http://doi.org/10.5281/zenodo.14704088" ID-Zenodo-Dep="14704088" httpUri="https://zenodo.org/record/14704088/files/figure.png" pageId="8" pageNumber="127" startId="8.[152,255,1943,1967]" targetBox="[210,1377,188,1920]" targetPageId="8" targetType="figure">
<paragraph id="8BCE36EBFFD26D7CFF170CD4FD2EC993" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="127">
<emphasis id="B905EAF9FFD26D7CFF170CD4FEEDC9E8" bold="true" box="[152,274,1943,1967]" pageId="8" pageNumber="127">FIGURE 3</emphasis>
: Holotype of
<taxonomicName id="4C714D68FFD26D7CFE110CDBFDA1C9F7" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[414,606,1944,1968]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="8" pageNumber="127" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFD26D7CFE110CDBFDA1C9F7" box="[414,606,1944,1968]" italics="true" pageId="8" pageNumber="127">Boaedon broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD26D7CFDEB0CDBFD4BC9F7" bold="true" box="[612,692,1944,1968]" pageId="8" pageNumber="127">
<taxonomicNameLabel id="A2365782FFD26D7CFDEB0CDBFD4BC9F7" box="[612,692,1944,1968]" pageId="8" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMH R08485) from nearby Addis Ababa, Ethiopia. Left: head in dorsal, ventral and lateral view, right side drawings of the head.
</paragraph>
</caption>
<caption id="DF0E6663FFD36D7DFF1808B2FAA2CA4E" ID-DOI="http://doi.org/10.5281/zenodo.14704092" ID-Zenodo-Dep="14704092" box="[151,1373,1009,1033]" httpUri="https://zenodo.org/record/14704092/files/figure.png" pageId="9" pageNumber="128" startId="9.[151,250,1009,1033]" targetBox="[189,1399,181,985]" targetPageId="9" targetType="figure">
<paragraph id="8BCE36EBFFD36D7DFF1808B2FAA2CA4E" blockId="9.[151,1373,1009,1033]" box="[151,1373,1009,1033]" pageId="9" pageNumber="128">
<emphasis id="B905EAF9FFD36D7DFF1808B2FEF2CA4E" bold="true" box="[151,269,1009,1033]" pageId="9" pageNumber="128">FIGURE 4</emphasis>
: Live photo of
<taxonomicName id="4C714D68FFD36D7DFE2508B2FD94CA4E" authorityName="Hallermann &amp; Hawlitschek" authorityYear="2025" box="[426,619,1009,1033]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="9" pageNumber="128" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis id="B905EAF9FFD36D7DFE2508B2FD94CA4E" box="[426,619,1009,1033]" italics="true" pageId="9" pageNumber="128">Boaedon broadleyi</emphasis>
</taxonomicName>
<emphasis id="B905EAF9FFD36D7DFDFE08B2FD3DCA4E" bold="true" box="[625,706,1009,1033]" pageId="9" pageNumber="128">
<taxonomicNameLabel id="A2365782FFD36D7DFDFE08B2FD3DCA4E" box="[625,706,1009,1033]" pageId="9" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from nearby Lake Oromia region, Ethiopia. Photo by Petr Nečas.
</paragraph>
</caption>
<subSubSection id="C36B6560FFD36D7DFF480F13FE2BCABC" pageId="9" pageNumber="128" type="distribution">
<paragraph id="8BCE36EBFFD36D7DFF480F13FB9ECAF4" blockId="9.[151,1437,1104,1419]" pageId="9" pageNumber="128">
<emphasis id="B905EAF9FFD36D7DFF480F13FE9ECA2D" bold="true" box="[199,353,1104,1130]" pageId="9" pageNumber="128">Distribution.</emphasis>
The species is currently known from different localities in elevations from approx.
<quantity id="4C899B0EFFD36D7DFA9C0F12FAA4CA2C" box="[1299,1371,1105,1132]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="128" unit="m" value="750.0">750 m</quantity>
up to
<quantity id="4C899B0EFFD36D7DFF180F36FF13CAC8" box="[151,236,1141,1167]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="128" unit="m" value="3000.0">3000 m</quantity>
asl in Ethiopian Regional States in Afar, Oromia, Somali and Southern Nations, Nationalities, and Peoples National Regional State and most probably in Northern
<collectingCountry id="F366767BFFD36D7DFC850FDAFC99CAF4" box="[778,870,1177,1203]" name="Somalia" pageId="9" pageNumber="128">Somalia</collectingCountry>
(Somaliland) (
<figureCitation id="134A2A6EFFD36D7DFB810FDAFBABCAF4" box="[1038,1108,1177,1203]" captionStart="FIGURE 2" captionStartId="5.[152,255,1846,1870]" captionTargetBox="[170,1368,516,1823]" captionTargetId="figure-160@5.[151,1436,486,1825]" captionTargetPageId="5" captionText="FIGURE 2. Confirmed records of Boaedon species in north-eastern and eastern Africa. Red dots: Boaedon broadleyi sp. nov.; Red star: Boaedon broadleyi sp. nov. type locality; green dots: Boaedon subniger sp. nov.; green star: Boaedon subniger sp. nov. type locality; red square: Boaedon montanus; red triangle: Boaedon paralineatus; blue dots, Boaedon capensis s.l.: orange dots Boaedon sp. black form; yellow dot: B. arabicus." figureDoi="http://doi.org/10.5281/zenodo.14704084" httpUri="https://zenodo.org/record/14704084/files/figure.png" pageId="9" pageNumber="128">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BCE36EBFFD36D7DFF480FFFFE2BCABC" blockId="9.[151,1437,1104,1419]" pageId="9" pageNumber="128">
<emphasis id="B905EAF9FFD36D7DFF480FFFFDA2CA90" bold="true" box="[199,605,1212,1239]" pageId="9" pageNumber="128">Habitat and Natural History notes</emphasis>
. The specimens were found in savannas, grasslands and woodlands, absent in arid or semi-arid habitats.
</paragraph>
</subSubSection>
<subSubSection id="C36B6560FFD36D7DFF480E46FE64CBCC" pageId="9" pageNumber="128" type="etymology">
<paragraph id="8BCE36EBFFD36D7DFF480E46FE64CBCC" blockId="9.[151,1437,1104,1419]" pageId="9" pageNumber="128">
<emphasis id="B905EAF9FFD36D7DFF480E46FEB1CB58" bold="true" box="[199,334,1285,1311]" pageId="9" pageNumber="128">Etymology.</emphasis>
The specific epithet is a patronym in the masculine genitive singular name of Donald George (Don) Broadley, a British born Herpetologist and emeritus curator of the Natural History Museum of
<collectingCountry id="F366767BFFD36D7DFA870E6AFA7FCB04" box="[1288,1408,1321,1347]" name="Zimbabwe" pageId="9" pageNumber="128">Zimbabwe</collectingCountry>
in
<collectingRegion id="49B5F809FFD36D7DFF180E0EFEF3CB20" box="[151,268,1357,1383]" country="Zimbabwe" name="Bulawayo" pageId="9" pageNumber="128">Bulawayo</collectingRegion>
. We honour his extensive work on African reptiles especially on
<taxonomicName id="4C714D68FFD36D7DFC7B0E0EFBA7CB20" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[1012,1112,1357,1383]" class="Squamata" family="Lamprophiidae" genus="Boaedon" kingdom="Animalia" pageId="9" pageNumber="128" phylum="Chordata" rank="genus">
<emphasis id="B905EAF9FFD36D7DFC7B0E0EFBA7CB20" box="[1012,1112,1357,1383]" italics="true" pageId="9" pageNumber="128">Boaedon</emphasis>
</taxonomicName>
, which he could not publish before he died in 2016.
</paragraph>
</subSubSection>
</treatment>
</document>