Add updates up until 2024-11-15 18:59:06
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4 changed files with 2179 additions and 83 deletions
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<mods:title id="63AC8297369F38808459B06E97AE1304">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<document id="536D6438232EA4BB041EC80CA1370932" ID-DOI="10.3897/zookeys.1218.120744" ID-ZooBank="094646BE-BF9F-4890-BD59-FF26FA95A3FE" ID-publisher-id="120744" URI-arpha="6E4DF739-56EF-5B8D-AEB7-2839D486DC50" URI-zoobank="http://zoobank.org/094646BE-BF9F-4890-BD59-FF26FA95A3FE" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946914" checkinUser="pensoft" docAuthor="Murányi, Dávid & Kovács, Tibor" docDate="2024" docId="B7518924CBFD591FA01FC896BA1EA93B" docLanguage="en" docName="ZooKeys 1218: 49-79" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/120744/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Leuctra puskasi Muranyi & Kovacs 2024, sp. nov." docType="treatment" docUuid="9C2CF7D9-F7F4-469C-AF6D-B78117E2078E" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="79" masterDocId="6E4DF73956EF5B8DAEB72839D486DC50" masterDocTitle="Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)" masterLastPageNumber="79" masterPageNumber="49" pageNumber="49" updateTime="1731697136958" updateUser="admin">
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<mods:title id="D8CC7C66E325FE2407BA2F93DB0EEE6B">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:namePart id="F8C3357207642BAAB55B9BF0BE148E73">Murányi, Dávid</mods:namePart>
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<mods:nameIdentifier id="D12B4EDEE77919BDA35178A8D84CA69D" type="ORCID">0000-0002-3907-5590</mods:nameIdentifier>
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<mods:affiliation id="DC8E4650C235BD4893866161C92CEFD8">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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<mods:namePart id="A78D410C617F74A9D64869E8F72C82F2">Murányi, Dávid</mods:namePart>
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<mods:nameIdentifier id="C56A0E38D27AB11425F4615F8CA5FE0B" type="ORCID">0000-0002-3907-5590</mods:nameIdentifier>
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<mods:affiliation id="92F241CE0B0AC2CBF194754C22A511C9">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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</mods:name>
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<mods:name id="3BC7127099C9F77B5549D09425375971" type="personal">
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<mods:namePart id="9CEF06FD7D30556F8C9477F7FB272930">Kovács, Tibor</mods:namePart>
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<mods:nameIdentifier id="4E31DE22BD4E5B7950301D73A5918D09" type="ORCID">0000-0003-4577-0394</mods:nameIdentifier>
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<mods:affiliation id="337143A993AAA40582F505F238B2B8B4">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<mods:namePart id="27C91C7F22F1CA4824E8763FB907649D">Kovács, Tibor</mods:namePart>
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<mods:nameIdentifier id="9896EFB5046CA73E99B6B38BF1909254" type="ORCID">0000-0003-4577-0394</mods:nameIdentifier>
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<mods:affiliation id="2D395B05A44AE09C4CFFCF4B5B6BABC3">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<mods:typeOfResource id="B624FDFED9E5C79BCA2BFC83E9D8257A">text</mods:typeOfResource>
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<mods:title id="C068B75E8E404F7FE245716098567026">ZooKeys</mods:title>
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<mods:date id="B328BB7682A8F67B148419CE999CE741">2024</mods:date>
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<mods:classification id="EA75E42809CD3C1512E73176EBD2EB6D">journal article</mods:classification>
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<mods:identifier id="7326DE8857417CA51DEF871D632DCB93" type="ZooBank">094646BE-BF9F-4890-BD59-FF26FA95A3FE</mods:identifier>
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<treatment id="B7518924CBFD591FA01FC896BA1EA93B" ID-DOI="http://doi.org/10.5281/zenodo.14171279" ID-Zenodo-Dep="14171279" ID-arpha="B7518924-CBFD-591F-A01F-C896BA1EA93B" ID-zoobank="https://zoobank.org/9C2CF7D9-F7F4-469C-AF6D-B78117E2078E" LSID="urn:lsid:zoobank.org:act:9C2CF7D9-F7F4-469C-AF6D-B78117E2078E" LSID-ZBK="urn:lsid:zoobank.org:act:9C2CF7D9-F7F4-469C-AF6D-B78117E2078E" httpUri="http://treatment.plazi.org/id/B7518924CBFD591FA01FC896BA1EA93B">
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<subSubSection id="3001862477E418D038970AA1086F9B3F" type="nomenclature">
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@ -68,39 +68,40 @@ Murányi & Kovács
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<heading id="BA9EC575EF4E8F152AD14DC6D41BABAE" reason="title">Type material.</heading>
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</paragraph>
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<paragraph id="518CC0EEF550530E9046C2530F82CE6A">
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<materialsCitation id="2DC30931E8455274475C51977B9A5079" collectingDate="2012-03-16" collectionCode="T, HNHM" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 4185" specimenCount="1" typeStatus="holotype">
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<materialsCitation id="2DC30931E8455274475C51977B9A5079" collectingDate="2012-03-16" collectionCode="HNHM" collectorName="T. Kovacs & D. Muranyi & G. Puskas" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 4185" specimenCount="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="holotype">
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||||
<emphasis id="3EB9DA76951FA213F3081D33866E3DCF" bold="true" italics="true">
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<typeStatus id="1ADBCAF2A43C45EFC0FFE7F21EA9AC43" type="holotype">Holotype</typeStatus>
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</emphasis>
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male:
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<specimenCount id="9952F6C493B65577BEE4716FC8533F47" type="male">male</specimenCount>
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:
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<collectingCountry id="7EFD3E67E65C5EBFBA3C7118524902BF" name="Bosnia and Herzegovina">Bosnia & Herzegovina</collectingCountry>
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•
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<location id="E0A69505027825BDE8D44DB4A7B4F8E2" LSID="urn:lsid:plazi:treatment:B7518924CBFD591FA01FC896BA1EA93B:E0A69505027825BDE8D44DB4A7B4F8E2" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" latitude="45.0414" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" name="Republika Srpska">Republika Srpska</location>
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<collectingRegion id="66AE56FB7008C2534262998EA579BF18">Republika Srpska</collectingRegion>
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,
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<collectingMunicipality id="C036D70D85032AC11E27CE89729A42C3">Kozara Mts</collectingMunicipality>
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, forest brook along the
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<collectingCounty id="4E16DE0F179322C245402F6FE8C55FC1">Gornji Podgradci-Kozarac</collectingCounty>
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road,
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,
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<location id="C1E29860575FA49AB30BB80B066193E7" LSID="urn:lsid:plazi:treatment:B7518924CBFD591FA01FC896BA1EA93B:C1E29860575FA49AB30BB80B066193E7" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" latitude="45.0414" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" name="forest brook along the Gornji Podgradci-Kozarac road" stateProvince="Republika Srpska">forest brook along the Gornji Podgradci-Kozarac road</location>
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,
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<quantity id="772B40030A089BC6F74A47C5AA805DD4" metricMagnitude="2" metricUnit="m" metricValue="5.95" unit="m" value="595.0">
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<elevation id="F5EB13A793FC0EC4FAF8491114F5E4F2" metricMagnitude="2" metricUnit="m" metricValue="5.95" unit="m" value="595.0">595 m</elevation>
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</quantity>
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,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EV6AG" specific-use="{"type":"Point","coordinates":[16.904000,45.041400]}">
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<geoCoordinate id="18E24858F79CDDEA5EE7FF2C5230A8D5" degrees="45.0414" direction="north" orientation="latitude" precision="5" value="45.0414">45.0414 ° N</geoCoordinate>
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<geoCoordinate id="18E24858F79CDDEA5EE7FF2C5230A8D5" degrees="45.0414" direction="north" orientation="latitude" precision="5" value="45.0414">45.0414°N</geoCoordinate>
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,
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<geoCoordinate id="4DDC80F53122873B849A22BDD41106B2" degrees="16.9040" direction="east" orientation="longitude" precision="5" value="16.904">16.9040 ° E</geoCoordinate>
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<geoCoordinate id="4DDC80F53122873B849A22BDD41106B2" degrees="16.9040" direction="east" orientation="longitude" precision="5" value="16.904">16.9040°E</geoCoordinate>
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||||
</named-content>
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||||
,
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<date id="5E82FBD4DD9A902C0BA35E8491267D9E" value="2012-03-16">
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||||
<collectingDate id="492692405B1687431E2833D1791F4E25" value="2012-03-16">16. iii. 2012</collectingDate>
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<collectingDate id="492692405B1687431E2833D1791F4E25" value="2012-03-16">16.iii.2012</collectingDate>
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||||
</date>
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||||
(field number: loc. 6), leg.
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<collectionCode id="C2C0AD4A2F2F9D6DA41DE24E1621BFE0" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
.
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<collectorName id="CF3557FDF07EF46BF89555C330C0E954">Kovács, D</collectorName>
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||||
.
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<collectorName id="4F464BC0E6B9123FEC8555435450A9A8">Murányi, G</collectorName>
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. Puskás (
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<collectorName id="BD9FA00CC1A2A63C4462AA00B5859EB4">T. Kovács</collectorName>
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||||
,
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<collectorName id="0F3FA3BB01B964F6958E6BFE04CF3087">D. Murányi</collectorName>
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,
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<collectorName id="5F4C1EA53C8CED358FBFAB9240774660">G. Puskás</collectorName>
|
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(
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<named-content id="50D3756D66CDBC9746691BCEBF4D2C22" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Collection of Smaller Insect Orders, Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
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||||
<collectionCode id="BA1C2A6CDDDAC1042A99C03CF672257C" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
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||||
</named-content>
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@ -109,7 +110,7 @@ road,
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)
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</materialsCitation>
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||||
.
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<materialsCitation id="5030C2568A25D6ED99411158393277D2" collectingDate="2012-03-16" collectionCode="BYU" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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||||
<materialsCitation id="5030C2568A25D6ED99411158393277D2" collectingDate="2012-03-16" collectionCode="BYU" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
|
||||
<emphasis id="48B47BFD1CA335F40C08A7B289E0B813" bold="true" italics="true">
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<typeStatus id="94CC5C7768A6158C84D0EEF56F1DB841" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
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||||
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@ -123,7 +124,7 @@ road,
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)
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||||
</materialsCitation>
|
||||
•
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||||
<materialsCitation id="DA18DFC1D5883CF45CDD2ABA8C4DE8E1" collectingDate="2012-03-16" collectionCode="CGV" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<materialsCitation id="DA18DFC1D5883CF45CDD2ABA8C4DE8E1" collectingDate="2012-03-16" collectionCode="CGV" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
|
||||
<specimenCount id="6A4C0A57638671E71C373013F2DF843F" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="9F54000A6F14D6C31859E2D15F11BA09" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
|
|
@ -133,7 +134,7 @@ road,
|
|||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="6556CEE5986DFD6D3E12021A7C19DBAC" collectionCode="HNHM" country="Bosnia and Herzegovina" specimenCode="PLP 3853" specimenCount="14" specimenCount-female="4" specimenCount-male="10" typeStatus="paratype">
|
||||
<materialsCitation id="6556CEE5986DFD6D3E12021A7C19DBAC" collectingDate="2012-03-16" collectionCode="HNHM" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 3853" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Republika Srpska" typeStatus="paratype">
|
||||
<specimenCount id="FFC84BB845E61235B3F0038CA61BFF8E" count="10" type="male">10 ♂</specimenCount>
|
||||
<specimenCount id="3698B56407C9DD52C3FFDAD5D8D0A30C" count="4" type="female">4 ♀</specimenCount>
|
||||
(
|
||||
|
|
@ -145,17 +146,21 @@ road,
|
|||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="6B8B5A63AD77561F29857FADE5E08B82" collectionCode="MM" country="Bosnia and Herzegovina" specimenCount="17" specimenCount-female="5" specimenCount-male="12" typeStatus="paratype">
|
||||
<materialsCitation id="6B8B5A63AD77561F29857FADE5E08B82" collectingDate="2012-03-16" collectionCode="MM" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="2012-10, PLETYP-36" specimenCount="17" specimenCount-female="5" specimenCount-male="12" stateProvince="Republika Srpska" typeStatus="paratype">
|
||||
<specimenCount id="CB5B5E993D7B2F8FDA66D67736658085" count="12" type="male">12 ♂</specimenCount>
|
||||
<specimenCount id="14B57FDBAC4668AD07D3467C10FE6694" count="5" type="female">5 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="45AD23B29E824D51AC88FEF3324A183B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||||
<collectionCode id="56F001848ED40A23AB887F5892D637E1" country="France" name="University of Montpellier">MM</collectionCode>
|
||||
</named-content>
|
||||
: 2012-10, PLETYP- 36)
|
||||
:
|
||||
<specimenCode id="B543C76CE86B469B25AEFC1A1B9C6088">2012-10</specimenCode>
|
||||
,
|
||||
<specimenCode id="7C673B3EAD26FE246198F37385A23DDE">PLETYP-36</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="AB95F7C73A3D87FC085811582410602B" collectionCode="SMNS" country="Bosnia and Herzegovina" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<materialsCitation id="AB95F7C73A3D87FC085811582410602B" collectingDate="2012-03-16" collectionCode="SMNS" collectorName="Kovacs, D & Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
|
||||
<specimenCount id="F9F8FA8E5AC4048F4354E85EEB471194" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="34596C539C68B38D1C57B5B9B3B611C9" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
|
|
@ -207,17 +212,7 @@ Male abdomen (Figs
|
|||
<figureCitation id="99A73570751D5C5177EDC13C0E5DBFDA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Terminalia of Leuctra puskasi sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D variation of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Setation omitted with the exception of subgenital plate on G. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure8" httpUri="https://binary.pensoft.net/fig/1175459">8 A – E</figureCitation>
|
||||
,
|
||||
<figureCitation id="D8F873418A09FE79297D9F18F5952505" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 A</figureCitation>
|
||||
): Tergite I sclerotised only anteriorly and in lateral stripes, with two medial spots. Transverse row of four pigmented spots distinct on terga
|
||||
<collectionCode id="67B7C744178E0FD8D0E174EAD8436285">II</collectionCode>
|
||||
– VII. Terga
|
||||
<collectionCode id="B961764EAFE79D2D02F7966B00033755">II</collectionCode>
|
||||
and
|
||||
<collectionCode id="18940996885CCBAC062B87492E8A0D36">III</collectionCode>
|
||||
with medially divided antecosta, tergite
|
||||
<collectionCode id="894BB5FE169524AA9F887E59C62D59EB">II</collectionCode>
|
||||
with triangular membranous area anteriorly. Terga IV – VII with entire antecosta and full sclerotised, tergite VII with weakly sclerotised posterior edge and small membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends into a small triangular plate; the membranous area is ~ 1 / 3 segment width, and usually do not reach the 1 / 2 of the segment length but in some specimens it reach down to 2 / 3 above the posteromedial process, posterior margin of the membranous portion wavy; the surrounding sclerotisation of the membranous area lighter than the rest of the segment; posteromedial process not darker than the lateral areas of the segment, not erect in side view and not or only slightly overhangs the segment posterior edge, nearly as wide as the membranous area above, bilobed with rounded lobes slightly pointed laterad, the space between the lobes is concave to nearly straight; posterior margin between the posteromedial process and segment sides nearly straight, not indenting nor distinctly lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 to 1 / 2 of its length; posteromedial sclerite short but wide, sublateral areas darker, the medial portion usually narrower than the lateral portions. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep notch, not wider than 1 / 3 tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk short. Cercus simple, covered with long setae. Sterna
|
||||
<collectionCode id="1DB1A43BE5D2C2D7B810E5A53BBE8CC1">II</collectionCode>
|
||||
– VIII simple, sternite IX bears a vesicle shorter than 1 / 2 segment length, its width more than 1 / 2 its length; posteromedial 1 / 2 of the sternum is delimited by a pale and weakly sclerotised area. Paraproct with moderately wide base, abruptly narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, blunt triangular process. Specillum longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, sharp tip.
|
||||
): Tergite I sclerotised only anteriorly and in lateral stripes, with two medial spots. Transverse row of four pigmented spots distinct on terga II – VII. Terga II and III with medially divided antecosta, tergite II with triangular membranous area anteriorly. Terga IV – VII with entire antecosta and full sclerotised, tergite VII with weakly sclerotised posterior edge and small membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends into a small triangular plate; the membranous area is ~ 1 / 3 segment width, and usually do not reach the 1 / 2 of the segment length but in some specimens it reach down to 2 / 3 above the posteromedial process, posterior margin of the membranous portion wavy; the surrounding sclerotisation of the membranous area lighter than the rest of the segment; posteromedial process not darker than the lateral areas of the segment, not erect in side view and not or only slightly overhangs the segment posterior edge, nearly as wide as the membranous area above, bilobed with rounded lobes slightly pointed laterad, the space between the lobes is concave to nearly straight; posterior margin between the posteromedial process and segment sides nearly straight, not indenting nor distinctly lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 to 1 / 2 of its length; posteromedial sclerite short but wide, sublateral areas darker, the medial portion usually narrower than the lateral portions. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep notch, not wider than 1 / 3 tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk short. Cercus simple, covered with long setae. Sterna II – VIII simple, sternite IX bears a vesicle shorter than 1 / 2 segment length, its width more than 1 / 2 its length; posteromedial 1 / 2 of the sternum is delimited by a pale and weakly sclerotised area. Paraproct with moderately wide base, abruptly narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, blunt triangular process. Specillum longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, sharp tip.
|
||||
</paragraph>
|
||||
<caption id="6404BCBEEF92B2192C1250928B7726F6" ID-DOI="10.3897/zookeys.1218.120744.figure8" ID-arpha="74A8422E-EEA4-5FD7-B500-E04DB1941AC0" httpUri="https://binary.pensoft.net/fig/1175459" startId="F8">
|
||||
<paragraph id="065AD1DE22E29BB8D344F05846D9F354">
|
||||
|
|
@ -274,13 +269,7 @@ Female abdomen (Figs
|
|||
<figureCitation id="3BA23C40155AC557F5A6E77793D018C9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Terminalia of Leuctra puskasi sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D variation of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Setation omitted with the exception of subgenital plate on G. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure8" httpUri="https://binary.pensoft.net/fig/1175459">8 F, G</figureCitation>
|
||||
,
|
||||
<figureCitation id="AC8F464624BD2164516EBEBDC32BAF59" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 E</figureCitation>
|
||||
): Terga I – VIII with transverse row of four pigmented spots clearly visible; terga I – IV mostly membranous but with lateral sclerites, terga
|
||||
<collectionCode id="3102681D03D3F090D8AC27E79FF36778" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
– VIII with medial sclerite as well; medial sclerite is a longitudinal stripe on tergite
|
||||
<collectionCode id="025B1D2EBB3F8DBE1E074BD3C1CD57A7" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
, gradually widens on the following segments and as wide as 1 / 2 segment width on tergite VIII; terga IX – X fully sclerotised. Sterna I – VII simple, sterna
|
||||
<collectionCode id="CB70F36E4200B92027B319F98E11BD53">II</collectionCode>
|
||||
– VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large and wide lobes, slightly darker brown than the rest of the plate; posterior margin convex, incision between the lobes is a membranous strip, widening towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is narrow in ventral view, distinctly raised, nose shaped to rounded in lateral view, lacking distinct setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging the posterior teeth.
|
||||
): Terga I – VIII with transverse row of four pigmented spots clearly visible; terga I – IV mostly membranous but with lateral sclerites, terga V – VIII with medial sclerite as well; medial sclerite is a longitudinal stripe on tergite V, gradually widens on the following segments and as wide as 1 / 2 segment width on tergite VIII; terga IX – X fully sclerotised. Sterna I – VII simple, sterna II – VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large and wide lobes, slightly darker brown than the rest of the plate; posterior margin convex, incision between the lobes is a membranous strip, widening towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is narrow in ventral view, distinctly raised, nose shaped to rounded in lateral view, lacking distinct setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging the posterior teeth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMEBG" type="Affinities">
|
||||
|
|
@ -373,9 +362,7 @@ The species was collected at a single forest brook in the Kozara Mts of northwes
|
|||
<figureCitation id="F781F301F816534F036C58F141AF81FA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus and habitat of Leuctra puskasi sp. nov. A female habitus B, C male habitus D type locality: forest brook in the Kozara Mts. E substrate of the type locality. Photographs A – C by Gellért Puskás." figureDoi="10.3897/zookeys.1218.120744.figure9" httpUri="https://binary.pensoft.net/fig/1175460">9 D</figureCitation>
|
||||
). It has a stony substrate mixed with mud and plenty of dead wood and fallen leaves (Fig.
|
||||
<figureCitation id="9A38933F6D79A9E4FF724D8B26CEFCC9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus and habitat of Leuctra puskasi sp. nov. A female habitus B, C male habitus D type locality: forest brook in the Kozara Mts. E substrate of the type locality. Photographs A – C by Gellért Puskás." figureDoi="10.3897/zookeys.1218.120744.figure9" httpUri="https://binary.pensoft.net/fig/1175460">9 E</figureCitation>
|
||||
). This habitat
|
||||
<typeStatus id="D94A788395B2A5E4B0187E7DC807597C">type</typeStatus>
|
||||
is very different from those of the closely related
|
||||
). This habitat type is very different from those of the closely related
|
||||
<taxonomicName id="66CCCB5CE61FB8AF55C5A1225BEE1A49" authorityName="Vincon & Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||||
<emphasis id="A5C5E9475DFC830B3B5E384030C3BB74" italics="true">L. dalmoni</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
@ -391,9 +378,7 @@ species complex was found in the Kozara, despite the fact that some other habita
|
|||
<taxonomicName id="4F82D6887444F8D224621E025341FACF" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||||
<emphasis id="30D60B3AC1FB733872D61F4EC2402BE5" italics="true">L. prima</emphasis>
|
||||
</taxonomicName>
|
||||
. The
|
||||
<typeStatus id="EDDFE9D59C7B7791D734BE1B071D5213">type</typeStatus>
|
||||
specimens were the only adult stoneflies found there (in partial snow cover), but last instar larvae of
|
||||
. The type specimens were the only adult stoneflies found there (in partial snow cover), but last instar larvae of
|
||||
<taxonomicName id="F6FC02AE5EFBF81B463110794F2782C0" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||||
<emphasis id="456DF089029E01262EEA28D99B1CA975" italics="true">Leuctra nigra</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
@ -408,13 +393,9 @@ specimens were the only adult stoneflies found there (in partial snow cover), bu
|
|||
</taxonomicName>
|
||||
group member were found in the brook. We visited the locality twice more, once in May (
|
||||
<date id="D3E10F98837D924D384DBA1C4F5338AF" value="2012-05-24">24. v. 2012</date>
|
||||
, leg.
|
||||
<collectionCode id="9AF131AA3B8AE65457CB3DC4FC193A60" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
. Kovács, G. Puskás) and once in November (
|
||||
, leg. T. Kovács, G. Puskás) and once in November (
|
||||
<date id="67132132AA6DD5D2A496C6C4E77EBF0D" value="2012-11-07">7. xi. 2012</date>
|
||||
, leg.
|
||||
<collectionCode id="1A8FC093D45AD93C1927C1E98674916C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||||
. Kovács, G. Magos). In May, adults of
|
||||
, leg. T. Kovács, G. Magos). In May, adults of
|
||||
<taxonomicName id="D83F0FF46958D0A414B527E9E54C1A77" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||||
<emphasis id="D30EA5B9DA79CFF10B5E741A8E2F94BA" italics="true">Leuctra nigra</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue