Add updates up until 2024-11-15 18:59:06
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<document id="442C4462BAE0EB8A9A1D982D43071CBC" ID-DOI="10.3897/zookeys.1218.120744" ID-ZooBank="094646BE-BF9F-4890-BD59-FF26FA95A3FE" ID-publisher-id="120744" URI-arpha="6E4DF739-56EF-5B8D-AEB7-2839D486DC50" URI-zoobank="http://zoobank.org/094646BE-BF9F-4890-BD59-FF26FA95A3FE" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946914" checkinUser="pensoft" docAuthor="Murányi, Dávid & Kovács, Tibor" docDate="2024" docId="6EAE8429DA2E58F48A813DD7F30E156A" docLanguage="en" docName="ZooKeys 1218: 49-79" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/120744/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Leuctra enigma Kovacs & Muranyi 2024, sp. nov." docType="treatment" docUuid="F0FF7DC4-7EA8-4BDE-9042-34D956A63609" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="79" masterDocId="6E4DF73956EF5B8DAEB72839D486DC50" masterDocTitle="Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)" masterLastPageNumber="79" masterPageNumber="49" pageNumber="49" updateTime="1731697136958" updateUser="admin">
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<mods:titleInfo id="37995F70DC84C922DF18BA328867E622">
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<mods:title id="F42684CED5C427132B257192BAB18C80">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:name id="5BBB9AB67A5FA3281E5F56BD1B17FACB" type="personal">
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<mods:roleTerm id="AB6CAF4B028CFCEAA0C335E151259913">Author</mods:roleTerm>
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<mods:namePart id="DC4F733C541DFDD61CA6A5EEAEBEEF64">Murányi, Dávid</mods:namePart>
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<mods:nameIdentifier id="F0ECD166624783E711D784877519C3CE" type="ORCID">0000-0002-3907-5590</mods:nameIdentifier>
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<mods:affiliation id="14661DB8B9E898998DE02B828CDF3B72">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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</mods:name>
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<mods:namePart id="CB45144D08E9A56F58EDC1EC67F30A6F">Kovács, Tibor</mods:namePart>
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<mods:affiliation id="A0110070A98BA8A76E72B4573A4307D2">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<mods:typeOfResource id="F189F8FBC2BEAF1D05E60EE144029768">text</mods:typeOfResource>
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<mods:part id="F9CF59C66ECB453E00F041FBA64D3FD9">
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<mods:date id="9764A110383DBE2FA17D01991BFEA1BA">2024</mods:date>
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<mods:number id="9A2881D767B4F22198363666B084674F">2024-11-15</mods:number>
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<mods:number id="AEE386F99FF5F6DC30E08838879EDEDE">1218</mods:number>
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<mods:classification id="631350A34C096DFEFD586B4113C9FC56">journal article</mods:classification>
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<mods:identifier id="D36BD12119C3D5FC24A00FF0366C9017" type="DOI">10.3897/zookeys.1218.120744</mods:identifier>
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<mods:identifier id="A9B24A08CF8E454C4E1AE38C97C0EC07" type="ZooBank">094646BE-BF9F-4890-BD59-FF26FA95A3FE</mods:identifier>
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</mods:mods>
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<treatment id="6EAE8429DA2E58F48A813DD7F30E156A" ID-arpha="6EAE8429-DA2E-58F4-8A81-3DD7F30E156A" ID-zoobank="https://zoobank.org/F0FF7DC4-7EA8-4BDE-9042-34D956A63609" LSID="urn:lsid:zoobank.org:act:F0FF7DC4-7EA8-4BDE-9042-34D956A63609" LSID-ZBK="urn:lsid:zoobank.org:act:F0FF7DC4-7EA8-4BDE-9042-34D956A63609" httpUri="http://treatment.plazi.org/id/6EAE8429DA2E58F48A813DD7F30E156A">
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<subSubSection id="32CF0813C499F1BD6A0B9CB12F1E03CB" type="nomenclature">
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<paragraph id="7C5BCD1E5AB54FC8A17EC8F8A99FE788">
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||||
<taxonomicName id="3FF8E843145769EDEB98A2286FF0055A" authority="Kovacs & Muranyi" authorityName="Kovacs & Muranyi" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
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||||
<emphasis id="FA2DC3D910A24176CFA3B7CA1E017EAF" italics="true">Leuctra enigma</emphasis>
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||||
Kovács & Murányi
|
||||
</taxonomicName>
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||||
<taxonomicNameLabel id="B8FB01E5CD52087A1CB7AD1E202B9B60" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="6A6C55823B55DA35F410D0696AEE6EB6" type="description">
|
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<paragraph id="2E1D148F61D7C62435AC510823309942">
|
||||
<figureCitation id="00FD09877E6DEB2739D6BD0C7135AF63" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="D5B96BB91E404DEECC7F2E16407F8D09" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="640789A2AB8EEFC8D94718E1D815EB55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Putative larva of Leuctra enigma sp. nov. A head and pronotum, dorsal view B meso- and metathorax, dorsal view C hind leg, dorsal view D terminalia, dorsal view. Scale bars: 1 m." figureDoi="10.3897/zookeys.1218.120744.figure3" httpUri="https://binary.pensoft.net/fig/1175454">3</figureCitation>
|
||||
,
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||||
<figureCitation id="E5ACE05C9B71F11AB6AF1F3104752662" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="4BB965E034B5FD1F716FE02E8FDB6097" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A – D male terminalia, dorsal view E – H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski & Zwick, 1970, Sutjeska, Bosnia & Herzegovina. Scale bars: 1 mm (D for A – D, H for E – H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 A, E</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEOAC" type="materials_examined">
|
||||
<paragraph id="0126ABC37F466E8BAEB890E0EAD32356">
|
||||
<heading id="560A4010925A5D09914B3840A1EECADC" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="ABD2D5596D695672A99626FB19745397">
|
||||
<materialsCitation id="3C5355A2E3B122303F1EDAD02825C78D" collectingDate="2012-10-10" collectionCode="MM" collectorName="P. Juhasz & T. Kovacs & D. Muranyi & G. Puskas" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2012-178, PLETYP-30" specimenCount="1" specimenCount-male="1" stateProvince="Diber county" typeStatus="holotype">
|
||||
<emphasis id="7F13AE7AC28B93A9EA10E67BC34FA02F" bold="true" italics="true">
|
||||
<typeStatus id="A741F6D46ADC696A5BB2485E08EB3DF4" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="C7C1B6D1BFEDA82260C4D4456C464848" count="1" type="male">male</specimenCount>
|
||||
:
|
||||
<collectingCountry id="DB6C208F616F41F94444EC7F7199E1C5" name="Albania">Albania</collectingCountry>
|
||||
: •
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<collectingRegion id="27330240143CAC28C1698E8FA411425B" country="Albania" name="Diber">Dibër county</collectingRegion>
|
||||
,
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<collectingMunicipality id="B672AFCC40CFCEAB27510F71E0309FD3">Bulqizë municipality</collectingMunicipality>
|
||||
,
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<collectingMunicipality id="5F8D3AD7EC20EA823AEFC5FFF871D832">Çermenikë Mts</collectingMunicipality>
|
||||
,
|
||||
<location id="D9124DA58F03C65614D2A7EADB75807A" LSID="urn:lsid:plazi:treatment:6EAE8429DA2E58F48A813DD7F30E156A:D9124DA58F03C65614D2A7EADB75807A" country="Albania" county="Bulqize" latitude="41.38665" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" name="brooks in open forest beneath Mt. Kaptine" stateProvince="Diber county">brooks in open forest beneath Mt. Kaptinë</location>
|
||||
,
|
||||
<quantity id="1E04BEDFD7C6DE0401409AABFC33C2D1" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
|
||||
<elevation id="A3A8926D39E6CC4D4FF0C882831DA717" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
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||||
</quantity>
|
||||
,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESOAC" specific-use="{"type":"Point","coordinates":[20.288967,41.386650]}">
|
||||
<geoCoordinate id="827D837D81A5F0B00432A2D4DE17C060" degrees="41" direction="north" minutes="23.199" orientation="latitude" precision="1" value="41.38665">41°23.199'N</geoCoordinate>
|
||||
,
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<geoCoordinate id="35B0D6825BE8418142353A3FC2077CAB" degrees="20" direction="east" minutes="17.338" orientation="longitude" precision="1" value="20.288967">20°17.338'E</geoCoordinate>
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</named-content>
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,
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<date id="F6B118D5C55656BC6456D9E85D064F5F" value="2012-10-10">
|
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<collectingDate id="2D4C0261269DD943B2CA6580199D38F9" value="2012-10-10">10.x.2012</collectingDate>
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</date>
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(field number: loc. 25), leg.
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<collectorName id="5905C8F6D06B9D7F03EFEBABE4E40436">P. Juhász</collectorName>
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,
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<collectorName id="E0DE69E4214469A65437F0B4AF40D493">T. Kovács</collectorName>
|
||||
,
|
||||
<collectorName id="19BD116B691D3BAE221B8E90A1D5C7CE">D. Murányi</collectorName>
|
||||
,
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<collectorName id="82F77C00D9AAD8264B9F500A3071462F">G. Puskás</collectorName>
|
||||
(
|
||||
<named-content id="BA406BC178241D40ADA535E75CC81C76" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||||
<collectionCode id="EE54FE3A1A7BDDFCB525554EF5668C4A" country="France" name="University of Montpellier">MM</collectionCode>
|
||||
</named-content>
|
||||
:
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||||
<specimenCode id="50F536263C045D9558395F656BDC2AC5">2012-178</specimenCode>
|
||||
,
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<specimenCode id="63E5A67E0C00A28CD313330F0B8E7876">PLETYP-30</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3C203646BECE370B2315EE7069790EF3" collectingDate="2012-10-10" collectionCode="BYU" collectorName="P. Juhasz & Kovacs, D & Muranyi, G" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
|
||||
<emphasis id="9E5D63E817B9D3F85DD1F27E50C14E63" bold="true" italics="true">
|
||||
<typeStatus id="D25EB46D3C22182F1353E50EFCD5735A" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: • same locality and date:
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||||
<specimenCount id="3C18C539F49C20305DD4F01E3FA27278" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="8251C2078C4F266B1D94E97A11D957F7" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="4F5E622C03253AFD5F8EBEEEB7CAA12B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/monte-l-bean-life-science-museum" xlink_title="Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, US">
|
||||
<collectionCode id="29ECEDC6A0A44EE0E95A11D810FB73E0" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" type="University or college">BYU</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="0359E18E5FAB06CE4FD5BA099E29E610" collectingDate="2012-10-10" collectionCode="CGV" collectorName="P. Juhasz & Kovacs, D & Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
|
||||
<specimenCount id="EF39CA5B7BBE1E294B787BCA65003EF2" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="CDA4FA16E24C57AAE778FD1190202F8A" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<abbrev content-type="institution" id="ABBRID0EEPAC" xlink_title="Gilles Vinçon Collection, Grenoble, France">
|
||||
<collectionCode id="18BB392891DCCA3A2FDE1052F1923BA5">CGV</collectionCode>
|
||||
</abbrev>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="84E2BA28AA8D024C193AD38B7F8DE9E3" collectingDate="2012-10-10" collectionCode="HNHM" collectorName="P. Juhasz & Kovacs, D & Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCode="PLP 4184" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Diber" typeStatus="paratype">
|
||||
<specimenCount id="91630917EEED0ACEE938599BE808050D" count="5" type="male">5 ♂</specimenCount>
|
||||
<specimenCount id="47AEB02B438FA870B706F6115E28CE35" count="6" type="female">6 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="B50D0B257AC67A3ED14D647EAF59AAD4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Collection of Smaller Insect Orders, Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
|
||||
<collectionCode id="53BB43BB699D933327052F17CE599E76" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
|
||||
</named-content>
|
||||
:
|
||||
<specimenCode id="DEA222A88988C6FECFF0D41EC4CED7A7" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">PLP 4184</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="EF2B236853CDE8A511AC46E47022A4DD" collectingDate="2012-10-10" collectionCode="MM" collectorName="P. Juhasz & Kovacs, D & Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCode="2012-178, PLETYP-31" specimenCount="15" specimenCount-female="10" specimenCount-male="5" stateProvince="Diber" typeStatus="paratype">
|
||||
<specimenCount id="1C87400453B25986E670B85A3443D38E" count="5" type="male">5 ♂</specimenCount>
|
||||
<specimenCount id="FCCA2BA04FAD2F11A7BD3D25DA2C3165" count="10" type="female">10 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="257858127369A044A0054F73A02C50EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||||
<collectionCode id="1EF113F034501570CB3AE40D5FC26152" country="France" name="University of Montpellier">MM</collectionCode>
|
||||
</named-content>
|
||||
:
|
||||
<specimenCode id="D5CE0A55919D5071644E3C2A9459393D">2012-178</specimenCode>
|
||||
,
|
||||
<specimenCode id="36B5E556FC960624EC3E849A2B65B20F">PLETYP-31</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="FDFBEC229D01B683D50070DC39A3480F" collectingDate="2012-10-10" collectionCode="SMNS" collectorName="P. Juhasz & Kovacs, D & Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
|
||||
<specimenCount id="2061A9EFC1EE8C9BED20734DE2F7FBB7" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="DC6CF87D2E038DCF4A6EDFA2EEAABFD5" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="FFB8AA8B216CC59C715C7C878FCFB2F0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-fuer-naturkund-stuttgart" xlink_title="Stuttgart State Museum of Natural History, Stuttgart, Germany">
|
||||
<collectionCode id="59EC5B8FE10781CFB2E8874212C9A851" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" type="">SMNS</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="5CEC01EA0728C2E6F03118AF8B50D91B">
|
||||
<heading id="680E8E18EA394FA39A695CCBFA32DCC1" reason="title">Other materials.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="775487E14E9A8830C1829CBDE394D659">
|
||||
•
|
||||
<materialsCitation id="2A19483FAF48F90044602AC4BB2B7A4B" collectingDate="2012-06-21" collectionCode="HNHM" collectorName="Z. Feher & T. Kovacs & D. Muranyi" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="PLP 4099" specimenCount="3" specimenCount-larva="3" stateProvince="Diber county">
|
||||
Same locality,
|
||||
<date id="AB44B9CA1A5243BBEC803F2B2793EDD1" value="2012-06-21">
|
||||
<collectingDate id="794A5F440C4004ABE58D5BD8DEBA3AB4" value="2012-06-21">21.vi.2012</collectingDate>
|
||||
</date>
|
||||
(loc. 58), leg.
|
||||
<collectorName id="1A1BA45815EE9C823D3C9CC182F9DDDF">Z. Fehér</collectorName>
|
||||
,
|
||||
<collectorName id="A778FD7C8EFA460BB6A3A548855AAF84">T. Kovács</collectorName>
|
||||
,
|
||||
<collectorName id="B5CC7B7AB1751C9FCE48202326DF3E2F">D. Murányi</collectorName>
|
||||
:
|
||||
<specimenCount id="81C64CD763AB53457E63C5BADC424CEC" count="3" type="larva">3 putative larvae</specimenCount>
|
||||
(
|
||||
<named-content id="0D98F68556253705BB32CDD70433E15B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Collection of Smaller Insect Orders, Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
|
||||
<collectionCode id="EEC0B5785C423A9B80E8BBCBE39A57CB" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
|
||||
</named-content>
|
||||
:
|
||||
<specimenCode id="25911FB961AA049AAB38E3D26DA44394">PLP 4099</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="DFD8A18159A178F3F92F04AFC446415B" collectingDate="2012-06-21" collectionCode="MM" collectorName="Z. Feher & T. Kovacs & D. Muranyi" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2012-1" specimenCount="2" specimenCount-larva="2" stateProvince="Diber county">
|
||||
<specimenCount id="7558581FBA14482ABF7DBAC85A28E6EB" count="2" type="pupa">2 putative larvae</specimenCount>
|
||||
(
|
||||
<named-content id="7F5C705FA24469DD5E0DE764705201CA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||||
<collectionCode id="F937F2AF3B3F95EE120668EFCE3FF174" country="France" name="University of Montpellier">MM</collectionCode>
|
||||
</named-content>
|
||||
:
|
||||
<specimenCode id="585AEB4FFA56BED0B3C8F7114DE1F1AC">2012-1</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="2E96475B8AA9D537569B700F954E8D52" collectingDate="2013-05-27" collectionCode="MM" collectorName="P. Juhasz & T. Kovacs & G. Magos & G. Puskas" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2013-19" specimenCount="2" specimenCount-larva="2" stateProvince="Diber county">
|
||||
same locality,
|
||||
<date id="1C9F813D5E1224A66711FA3025EA8E47" value="2013-05-27">
|
||||
<collectingDate id="66112502A9D9E6EB32CD463B75AF76CF" value="2013-05-27">27.v.2013</collectingDate>
|
||||
</date>
|
||||
(loc. 19), leg.
|
||||
<collectorName id="5E778670F8D3375B8DF94A66036BC0F4">P. Juhász</collectorName>
|
||||
,
|
||||
<collectorName id="DAD424E9366B51FC5CC04AF89887CA32">T. Kovács</collectorName>
|
||||
,
|
||||
<collectorName id="E9A80D47C57C3F0A4BAF0299D2AA72EB">G. Magos</collectorName>
|
||||
,
|
||||
<collectorName id="F3F1A53D8F2F52D5A54BA1379DCF397A">G. Puskás</collectorName>
|
||||
:
|
||||
<specimenCount id="23FC3240D9C4DFB83C0845D10B14531A" count="2" type="larva">2 putative larvae</specimenCount>
|
||||
(
|
||||
<named-content id="4B4C930BF6017CCEB3440A860DC8FDE3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||||
<collectionCode id="AF8A5184FE81265141FCDC9BA42822A8" country="France" name="University of Montpellier">MM</collectionCode>
|
||||
</named-content>
|
||||
:
|
||||
<specimenCode id="14977F826BEE3873F549ED111CF7AB0E">2013-19</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EOAAE" type="diagnosis">
|
||||
<paragraph id="F295F25D32DE87B9121FE44C46A386FE">
|
||||
<heading id="D185C4FAD07D75A16AA0DC1EFC7A2E7D" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="45A473D00F8A660D232A47FC19D73135">Macropterous in both sexes. Male tergite VII with huge membranous portion; tergite VIII with large, triangular posteromedial process, terminating in bi- or trilobed tip; tergite IX with large, lobed posteromedial sclerite supported by anterior sclerotised spots; tergite X posterior margin with deep and wide notch; sternite IX bears a vesicle shorter than 1 / 5 segment length; paraproct tip pointed, specillum slightly longer than paraproct, tip blunt. Female terga I – VIII with medial sclerite; subgenital plate with large lobes terminating in swollen and raised apex, central plate with anterolateral swellings and pale medial portion, posterior portion trapezoid in ventral view and nose-shaped in lateral view, well past by the lobes. Larva stout and with long setation, eyes small, clypeus lacks pointed corners.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ETAAE" type="description">
|
||||
<paragraph id="EDF3965B6579F5BB7CBC04A97A33168C">
|
||||
<heading id="E5FD55298CB17BD8D8998A8D09689085" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A223C02EE9D7C9037CCCB4D2EFABBAF2">
|
||||
Medium sized, robust species, both sexes macropterous. Forewing length:
|
||||
<typeStatus id="EEEC63E844C50C043ED702BCAFFF79C4" type="holotype">holotype</typeStatus>
|
||||
6.0 mm, male
|
||||
<typeStatus id="7BEAD6A3A8CE8D24D65D3E7ACC7B9ABA" type="paratype">paratypes</typeStatus>
|
||||
<quantity id="34CBFA8E80568BF60D98332C5D1F45E5" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.8–6.4 mm</quantity>
|
||||
, female
|
||||
<typeStatus id="9E507600E0EED1092595DEAA2D7173E0" type="paratype">paratypes</typeStatus>
|
||||
<quantity id="772733596F3C0897617FB5E5624D611A" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.8" metricValueMin="6.5" unit="mm" value="7.15" valueMax="7.8" valueMin="6.5">6.5–7.8 mm</quantity>
|
||||
; body length:
|
||||
<typeStatus id="BE4D0B8769CE69A1D0761D98F227196A" type="holotype">holotype</typeStatus>
|
||||
<quantity id="0DBE9FADC0DD1E73A45848E2C8B8E225" metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
|
||||
, male
|
||||
<typeStatus id="B5C94B151BCA70BB3BD75F142443EE4E" type="paratype">paratypes</typeStatus>
|
||||
6.0–
|
||||
<quantity id="3AC6A75237AA7AD10DE179D7B4FEB317" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
|
||||
, female
|
||||
<typeStatus id="4BD690CF0C472CDE709EFA0F6A8C1DC8" type="paratype">paratypes</typeStatus>
|
||||
8.0–
|
||||
<quantity id="29E59B02CC18DDF6771822A3A3E9B912" metricMagnitude="-3" metricUnit="m" metricValue="8.8" unit="mm" value="8.8">8.8 mm</quantity>
|
||||
. Setation generally short and dense. General colour dark brown. Head brown with distinct M-line and occipital rugosities, compound eyes relatively small; antennae dark brown, palpi brown. Pronotum brown to dark brown, wider than long and having rounded corners, rugosities distinct. Legs brown to dark brown, tarsi slightly darker. Wings brownish but hyaline, venation brown.
|
||||
</paragraph>
|
||||
<paragraph id="2572AE07402953658E21F98D301A8EAE">
|
||||
Male abdomen (Figs
|
||||
<figureCitation id="028B93260787AD6B44B103F050534AE6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2 A – E</figureCitation>
|
||||
,
|
||||
<figureCitation id="ED80133E17CB1D8ACF44962122766431" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A – D male terminalia, dorsal view E – H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski & Zwick, 1970, Sutjeska, Bosnia & Herzegovina. Scale bars: 1 mm (D for A – D, H for E – H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 A</figureCitation>
|
||||
): Tergite I membranous posteriorly and medially, with two medial spots; tergite II poorly sclerotised anteriorly. Transverse row of four pigmented spots distinct on terga II – VIII. Terga II – III with poorly developed antecosta, terga IV – VII with entire antecosta, terga III – VI full sclerotised. Tergite VII with huge medial membranous portion, a rounded or arrow-shaped sclerotised area extends between the medial pair of pigmented spots. Tergite VIII: antecosta nearly entire, gradually weakened in the medial 1 / 5, sometimes interrupted by a narrow medial notch; the posteromedial process is triangular, originated in the anterior 1 / 3 of the segment and reach the posterior 1 / 3 by its weakly bilobed or trilobed tip, medial portion with small membranous portion, medial pair of pigmented spots recognisable by the lateral sides of the process; the process is not darker than the lateral areas of the segment, wider than 1 / 3 segment width, slightly erect in side view; lateral edges of the medial membranous area obscure, while the anterior edge is sharply defined. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 of its length; posteromedial sclerite large, weakly bilobed anteriorly while posterior portion with long, diverging lobes, the sclerite is anteriorly supported by two sclerotised spots. Anterior margin of tergite X bilobed anteriorly but sclerotisation of lobes obscure; posterior margin with deep notch, as wide as 1 / 2 tergite width; medial portion of the notch with small protrusion at the origin of epiproct. Epiproct drop-shaped, sclerotised only at its sides, stalk gradually widening. Cercus simple, covered with long setae. Sterna II – VIII simple, sternite IX bears a very small, rounded vesicle shorter than 1 / 5 of segment length; posterolateral portions of the sternum pale coloured and weakly sclerotised. Paraproct with moderately wide base, narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a pointed tip. Base of paraproct connected to a subrectangular lateral expansion. Specillum longer than the paraproct, gently curved in lateral view and ending in a blunt tip.
|
||||
</paragraph>
|
||||
<caption id="C6F74C891334CA4FB42D9636CADFCC95" ID-DOI="10.3897/zookeys.1218.120744.figure2" ID-arpha="D29E1921-6368-5CBB-9AFB-8E4A219564F2" httpUri="https://binary.pensoft.net/fig/1175453" startId="F2">
|
||||
<paragraph id="68311BF8843F9FF8571D31596AD4D613">
|
||||
<label id="3C63737BB61DB7AFBD11DACB06E22E48">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="74E17A853095798D140081A493D14EF0">
|
||||
Terminalia of
|
||||
<taxonomicName id="6D5494B556CCAF59F6476D587153CAA5" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
|
||||
<emphasis id="D87AA987BB612E507289054A73C8881C" italics="true">Leuctra enigma</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="C7E95C8CAB3A981111C29D441C25310F" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
with setation omitted
|
||||
<emphasis id="9F4705E1230B07538BB53E95053496F4" bold="true">A</emphasis>
|
||||
male terminalia, dorsal view
|
||||
<emphasis id="D8DB4DEEE3C7384F5C4298FF43525F8C" bold="true">B</emphasis>
|
||||
same, lateral view
|
||||
<emphasis id="39BD783A513D71BAC14D89F7C51F305F" bold="true">C</emphasis>
|
||||
same, ventral view
|
||||
<emphasis id="A0841E6B3AE0329FE3300802813416DC" bold="true">D</emphasis>
|
||||
variations of male tergite 8, dorsal view
|
||||
<emphasis id="15F1692F8B358D740DBE6405038DB197" bold="true">E</emphasis>
|
||||
male paraprocts and specillae, caudal view
|
||||
<emphasis id="EF2E1BCFB445DC1D8463E49334978803" bold="true">F</emphasis>
|
||||
female terminalia, ventral view
|
||||
<emphasis id="F825201A0BBF94194A023EC16F11348C" bold="true">G</emphasis>
|
||||
same, lateral view. Scale bar: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="D13F9AF2745E4692691CC060A36E789F">
|
||||
Female abdomen (Figs
|
||||
<figureCitation id="84911C26077950CE2F2263191655CC96" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2 F – G</figureCitation>
|
||||
,
|
||||
<figureCitation id="695AEA7C28E90041CF43D3ADBF33E3FC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A – D male terminalia, dorsal view E – H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski & Zwick, 1970, Sutjeska, Bosnia & Herzegovina. Scale bars: 1 mm (D for A – D, H for E – H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 E</figureCitation>
|
||||
): Terga I – VIII with transverse row of four pigmented spots clearly visible; terga I – VIII mostly membranous but with lateral sclerites and a medial sclerite as well; medial sclerite is a longitudinal stripe, more widened on terga II and III; tergite IX mostly, tergite X fully sclerotised. Sterna I – VII simple, sterna II – VII with one subrectangular median sclerite and two small anterior sclerites that are not fused. Subgenital plate of sternite VIII large and complex, not fused with other sclerites, lacks distinct setation. It has two large and well-defined lobes, darker brown than the remainder of the plate; the lobes terminate in a swollen, rounded apex that are paler than the base of the lobes, raised in lateral view and converging in ventral view. The central plate has two anterolateral swellings, medial portion pale and lightly sclerotised; the plate terminates in a trapezoidal bulge, raised and nose-shaped in lateral view, clearly separated from the lateral lobes that extend beyond it. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus, and epiproct simple. Spermathecal sclerite thin, ring-shaped, with small anterior teeth and long posterior teeth.
|
||||
</paragraph>
|
||||
<paragraph id="7C25993C8C3BF898B493C0EFAD5DCDEE">
|
||||
Putative larva (Fig.
|
||||
<figureCitation id="A73B88BF9CB7FA0271C78C4EC17811B3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Putative larva of Leuctra enigma sp. nov. A head and pronotum, dorsal view B meso- and metathorax, dorsal view C hind leg, dorsal view D terminalia, dorsal view. Scale bars: 1 m." figureDoi="10.3897/zookeys.1218.120744.figure3" httpUri="https://binary.pensoft.net/fig/1175454">3</figureCitation>
|
||||
): Body stout, length of the penultimate or younger larvae 7.0–
|
||||
<quantity id="795125E323BED14F4B18022BD0A81C30" metricMagnitude="-3" metricUnit="m" metricValue="7.5" unit="mm" value="7.5">7.5 mm</quantity>
|
||||
. General colour pale brown, without distinctive markings, pilosity long and distinct. Head much wider than long, eye small, shorter than the widely rounded occipital region, ocelli distinct, clypeus not modified. Palpi simple, antenna longer than 1 / 2 body length. Pronotum much wider than long. Wing pads under development, posterior edge of meso- and metathorax between wing pads rounded, not like the typical
|
||||
<emphasis id="F60092C8A418DDDDB3D2CFEE36FB0C36" italics="true">prima-hippopus-inermis</emphasis>
|
||||
group shape. Legs short, tibiae not longer than femora, stretched hind leg reaches back to the seventh abdominal segment. Abdomen stout, integument matt pale brown, paraproct triangular. Cercus nearly as long as abdomen, with 25 cylindrical segments, medial segments twice longer than wide. Setation of the larva: antenna and palpi with short setae but head and clypeus with dense and erect setation, longest setae as long as the distance between hind ocelli; marginal setae continuous on pronotum, as long as the setae of the head; wing pads with similar long, erect and dense setae; femora and tibiae with hairs as long as the width of each femora, distributed on the whole surface but not forming swimming fringe; tarsi with short and scarce setation; abdominal segments evenly setose, with erect setae approximately as long as the setae of the head; cercomeres bald besides the erect apical whorl, slightly shorter than abdominal setae.
|
||||
</paragraph>
|
||||
<caption id="AAB0889B5304D2E8B98789495C3C1901" ID-DOI="10.3897/zookeys.1218.120744.figure3" ID-arpha="00379A01-9C9E-5DE2-9BF7-DD7563E6CB57" httpUri="https://binary.pensoft.net/fig/1175454" startId="F3">
|
||||
<paragraph id="8BA2825452430B0BCE8C43EF0AEAF91C">
|
||||
<label id="791AA4B18D15D7B28D5ADC69E0CED2D0">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="B9BC152434A262E1FDCB8FF01BA11A8D">
|
||||
Putative larva of
|
||||
<taxonomicName id="E5EDDFAFDEB4230B415505CE1135B1B7" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
|
||||
<emphasis id="1340B40A4FC9E1DD97FBA4CD0912581D" italics="true">Leuctra enigma</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="5B33ED51049E5866E131D6DD702697C9" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="7CD88880B28C60180EE234A4235B4011" bold="true">A</emphasis>
|
||||
head and pronotum, dorsal view
|
||||
<emphasis id="5493C952A555BCB9BA7A99C4ADA73615" bold="true">B</emphasis>
|
||||
meso- and metathorax, dorsal view
|
||||
<emphasis id="C188AC4EFD090E3F21C8CC9F46DD5D36" bold="true">C</emphasis>
|
||||
hind leg, dorsal view
|
||||
<emphasis id="358B790E117E9830EFA839DF18E493D8" bold="true">D</emphasis>
|
||||
terminalia, dorsal view. Scale bars: 1 m.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMEAE" type="Affinities">
|
||||
<paragraph id="6F5A7C7C84858BA3DD5A3768809F9784">
|
||||
<heading id="C3F04B272615CD629A67D5A995B34B35" reason="title">Affinities.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="51E6B10CE08D94ADECA0FA6008FDCEDE">
|
||||
The new species is an isolated member of the
|
||||
<taxonomicName id="A855CA7D39ADF2DE6BA8748E5A1C2052" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
|
||||
<emphasis id="A6F614163113C96DE5DEEF42F5E5E77B" italics="true">hippopus</emphasis>
|
||||
</taxonomicName>
|
||||
group. The male is rather distinctive, the only
|
||||
<taxonomicName id="AF2D8B3CBC513CD23E904FED48D3B918" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9A9EEB7E24C5594CDCB3BB76A33D631A" italics="true">Leuctra</emphasis>
|
||||
</taxonomicName>
|
||||
with comparable posteromedial process of tergite VIII is the Caucasian-Anatolian
|
||||
<taxonomicName id="88165E5618477302FADF9144C520AE9A" authority="Zhiltzova, 1960" authorityName="Zhiltzova" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="martynovi">
|
||||
<emphasis id="9659771FA458234E78E2827C428F5099" italics="true">L. martynovi</emphasis>
|
||||
<bibRefCitation id="658B9CABA39350D4E515D98437AD3A9B" author="Zhiltzova LA" issue="1" journalOrPublisher="Entomologicheskoe Obozrenie" pagination="156-171" refId="B84" refString="Zhiltzova LA (1960) K poznanyu vesnyanok (Plecoptera) Kavkaza. 4. Novye vidy Leuctridae. Entomologicheskoe Obozrenie 39 (1): 156 - 171." title="K poznanyu vesnyanok (Plecoptera) Kavkaza. 4. Novye vidy Leuctridae." volume="39" year="1960">Zhiltzova, 1960</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. However,
|
||||
<taxonomicName id="E31E2BA1B27EA1E22D84E22B9A788BEC" authorityName="Zhiltzova" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="martynovi">
|
||||
<emphasis id="4E2E017F9C3505410E345CA7D611BD72" italics="true">L. martynovi</emphasis>
|
||||
</taxonomicName>
|
||||
differs by larger posteromedial sclerite of tergite IX, shallow posterior notch of tergite X, and larger vesicle of sternite IX. The subgenital plate of the female shows similarity with the narrowly defined
|
||||
<taxonomicName id="59DC3945CB751FDBC3794C50170D14C4" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
|
||||
<emphasis id="14102CD7658965A8ED02AC9479685AFC" italics="true">hippopus</emphasis>
|
||||
</taxonomicName>
|
||||
group (
|
||||
<taxonomicName id="AFA67400EA5E7CDD4F04EF6420AB4E43" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
|
||||
<emphasis id="35CE7F348E5F816718A2D33C4EFFB3B7" italics="true">hippopus</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup, sensu, e. g.,
|
||||
<bibRefCitation id="8B45A0B49E6358704BD487B65C4F44CD" author="Ravizza C" journalOrPublisher="Lauterbornia" pagination="1-42" refId="B69" refString="Ravizza C (2002 b) Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species. Lauterbornia 44: 1–42. https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" title="Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species." url="https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" volume="44">Ravizza 2002 b</bibRefCitation>
|
||||
), having large lobes and a pale central plate. However, the combination of bulging lobes of the subgenital plate, central plate with anterolateral swellings and trapezoid, bulging posterior portion, already distinguish it from all other
|
||||
<taxonomicName id="1857BC0F2F015C821C56F6C0B09FB992" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="58C3E862010CA04C3E8F7318259ED8AA" italics="true">Leuctra</emphasis>
|
||||
</taxonomicName>
|
||||
. The putative larva is distinctive by its stout body and dense, long setation. Among the Balkanian
|
||||
<taxonomicName id="FAA3ED4E4EAC5EAEB97850F692C114E4" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="912B9F485A2F8D9AB2C9CE7C498454C5" italics="true">Leuctra</emphasis>
|
||||
</taxonomicName>
|
||||
, only
|
||||
<taxonomicName id="8D1011899E36158E0B68EC38AF1A063F" authority="(Olivier, 1811)" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||||
<emphasis id="B329A83A9C6D474BC5B6BA7D2986854E" italics="true">L. nigra</emphasis>
|
||||
(
|
||||
<bibRefCitation id="32BA3B7AE0BA7C23F8F307E3D9956721" author="Olivier GA" journalOrPublisher="Encyclopédie méthodique, ou par ordre de matiéres" pagination="185-187" refId="B57" refString="Olivier GA (1811) Nemoure. Encyclopedie methodique, ou par ordre de matieres 8: 185 - 187. https: // archive. org / details / encyclopdiemthod 08 oliv / page / 186 / mode / 2 up" title="Némoure." url="https://archive.org/details/encyclopdiemthod08oliv/page/186/mode/2up" volume="8" year="1811">Olivier, 1811</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="549EF697D19D85155A86D7540224A985" authority="Bogoesco & Tabacaru, 1960" authorityName="Bogoesco & Tabacaru" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hirsuta">
|
||||
<emphasis id="033C91CB60A60BF60590D7E88AE19CB1" italics="true">L. hirsuta</emphasis>
|
||||
<bibRefCitation id="6EC5496CE4D1D44AA0679ED29B2EC070" DOI="10.5169/seals-401378" author="Bogoesco C & Tabacaru I" issue="1-2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="91-96" refId="B8" refString="Bogoesco C, Tabacaru I (1960) Contribution a letude des Leuctra (Plecopteres) des Carpathes roumaines. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 33 (1 - 2): 91 - 96. https: // doi. org / 10.5169 / seals- 401378" title="Contribution à ľétude des Leuctra (Plécoptères) des Carpathes roumaines." volume="33" year="1960">Bogoesco & Tabacaru, 1960</bibRefCitation>
|
||||
</taxonomicName>
|
||||
are having similar long setation, however, their body is less stout, and their eyes are larger. The Alpine-Carpathian
|
||||
<taxonomicName id="0E930CB682ED633D1FCAFCC0497CA67E" authority="Kempny, 1898" authorityName="Kempny" authorityYear="1898" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="braueri">
|
||||
<emphasis id="298D06D51EC988E3E74BD513EF1AECD3" italics="true">L. braueri</emphasis>
|
||||
<bibRefCitation id="FD740C0FBDB8A551A0A65ED3244FA7DF" author="Kempny P" refId="B33" refString="Kempny P (1898) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. I. Theil. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 48: 213 - 221 + Pl. III. https: // www. biodiversitylibrary. org / part / 39234" url="https://www.biodiversitylibrary.org/part/39234" year="1898">Kempny, 1898</bibRefCitation>
|
||||
</taxonomicName>
|
||||
has similarly stout and hairy larva, but its clypeus is armed with prominent pointed corners.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EAIAE" type="distribution">
|
||||
<paragraph id="F6E67EB96A4E7181E731FF659F519C0D">
|
||||
<heading id="2D2CF29CA637C602E580E7BDCE9D725A" reason="title">Distribution and ecology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4D2A369FA6A90594C83CF126C6DA117E">
|
||||
The species was collected at a single high elevation site of the Çermenikë Mts, central
|
||||
<collectingCountry id="A29BE67EE8FE99DC3766E4BA32A3474F" name="Albania">Albania</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="49E3F9D2BD38FB03E9E6739E188550F9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">1</figureCitation>
|
||||
). It was not found at any further localities, despite of intensive collecting efforts during the last twenty years conducted in central
|
||||
<collectingCountry id="DA8E16D7F57DA965236B84B2AE33109A" name="Albania">Albania</collectingCountry>
|
||||
. The habitat is disturbed by partial deforestation, most sections of the two confluent brooks run open or in bush of young beech (Fig.
|
||||
<figureCitation id="8EB7FC15C785F8F0AD41290B6AD793FA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4 A, B</figureCitation>
|
||||
). The brooks run moderately fast,
|
||||
<quantity id="489B17EF23FDB669606D23DDFA8378A0" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 meter</quantity>
|
||||
wide and not deeper than 20 centimetres (both in June and October). The substrate is stony, mixed with small sandy patches, few patches of dead wood and partly submerged plants (Fig.
|
||||
<figureCitation id="C842B61835E58C0DCE64E63D43B6CAE1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4 B, C</figureCitation>
|
||||
). The specimens were swarming together with
|
||||
<taxonomicName id="9D43477EB2527EAD87484521703235E2" authorityName="Bogoesco & Tabacaru" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hirsuta">
|
||||
<emphasis id="5AD26731567414651A6639209B431E76" italics="true">L. hirsuta</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<date id="D1035615169C006A6D9192D598859626" value="2012-10">October 2012</date>
|
||||
. When we visited the locality in late spring and early summer, a more diverse fauna was found:
|
||||
<date id="ABC002AA39179D628260C0B528D9B5DC" value="2012-06-21">21. vi. 2012</date>
|
||||
(leg. Z. Fehér, T. Kovács, D. Murányi): adults of
|
||||
<taxonomicName id="B1CA003067590B4D194952BF43101F04" authority="Aubert, 1956 b" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
|
||||
<emphasis id="E0E937E89FBA9703678FF8201B8740FF" italics="true">Brachyptera helenica</emphasis>
|
||||
Aubert, 1956 b
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="5031065F8ABB2E085CD07C55CBA762F7" authority="Kacanski & Zwick, 1970" authorityName="Kacanski & Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
|
||||
<emphasis id="F8236301CAC40959DAF72B617E2B733E" italics="true">Leuctra hippopoides</emphasis>
|
||||
<bibRefCitation id="4AD3651C6B8D6F2F332903621721C0FB" author="Kaćanski D & Zwick P" issue="1" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="1-16" refId="B30" refString="Kacanski D, Zwick P (1970) Neue und wenig bekannte Plecopteren aus Jugoslawien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 43 (1): 1 - 16. https: // www. e-periodica. ch / cntmng? pid = seg- 001 % 3 A 1970 % 3 A 43 % 3 A % 3 A 14" title="Neue und wenig bekannte Plecopteren aus Jugoslawien." url="https://www.e-periodica.ch/cntmng?pid=seg-001%3A1970%3A43%3A%3A14" volume="43" year="1970">Kaćanski & Zwick, 1970</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C4E17D06B602AB539CE5BBEA954FF134" authority="Aubert, 1956" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
|
||||
<emphasis id="6355BE2DF1C31C2607A121C4756D56D0" italics="true">L. metsovonica</emphasis>
|
||||
Aubert, 1956
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="23189C516F836AA75794A383AD3E40DB" authority="(Klapalek, 1900)" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
|
||||
<emphasis id="08B675B9DB0B58C0222C8A66440CCEFC" italics="true">Nemurella pictetii</emphasis>
|
||||
(
|
||||
<bibRefCitation id="2F401CFA3D2FEF7E84913A91A0E9200A" author="Klapálek F" issue="20" journalOrPublisher="Slovesnost a Uměni" pagination="1-34" refId="B36" refString="Klapalek F (1900) Plekopterologicke Studie. Rozpravy Ceske Akademie Cisare Frantiska Josefa pro Vedy. Slovesnost a Umeni 9 (20): 1 - 34. [+ Pl. I - II.]" title="Plekopterologické Studie. Rozpravy České Akademie Cisaře Františka Josefa pro Vědy." volume="9" year="1900">Klapálek, 1900</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="754DC00D11D0FB59970A09A1ED071DA1" authority="(Retzius, 1783)" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
|
||||
<emphasis id="E52FF16D84E81CBB4782DB668D3AE3E3" italics="true">Nemoura cinerea cinerea</emphasis>
|
||||
(
|
||||
<bibRefCitation id="14E3730D496549FD6EF86C783AA50D9B" author="Retzius AJ" journalOrPublisher="Cruse, Lipsiae" refId="B72" refString="Retzius AJ (1783) Caroli de Geer genera et species insectorum. Cruse, Lipsiae, 220 pp. https: // www. biodiversitylibrary. org / item / 80958 # page / 68 / mode / 1 up" title="Caroli de Geer genera et species insectorum." url="https://www.biodiversitylibrary.org/item/80958#page/68/mode/1up" year="1783">Retzius, 1783</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4DE46F805528AFDC594CE2DD0AE2B7C1" authority="Pictet, 1836" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
|
||||
<emphasis id="D81EB68957DBDD309E4E7ECEF0DEC34C" italics="true">N. marginata</emphasis>
|
||||
<bibRefCitation id="4E6FD3B00914EF00037DB6B3856D7B17" author="Pictet FJ" issue="1" journalOrPublisher="Memoires de la Société de physique et d’histoire naturelle de Genève" pagination="173-190" refId="B61" refString="Pictet FJ (1836) Description de quelques nouvelles especes d'Insectes du Bassin du Leman. Memoires de la Societe de physique et d'histoire naturelle de Geneve 7 (1): 173 - 190. https: // www. biodiversitylibrary. org / item / 39755 # page / 201 / mode / 1 up" title="Description de quelques nouvelles espéces d’Insectes du Bassin du Léman." url="https://www.biodiversitylibrary.org/item/39755#page/201/mode/1up" volume="7" year="1836">Pictet, 1836</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="DB0C4198EFECC59A07F64DE60DA24D9B" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" class="Insecta" family="Perlidae" genus="Perla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pallida">
|
||||
<emphasis id="A42155A39A1DE79BA41BB4441DF07341" italics="true">Perla</emphasis>
|
||||
cf.
|
||||
<emphasis id="37E1C56F90579208955DFDC8787817F9" italics="true">pallida</emphasis>
|
||||
<bibRefCitation id="E13CD5A981C94F58B6D7153698A1B632" author="Guérin-Méneville FE" refId="B21" refString="Guerin-Meneville FE (1843) Genre Perle. In: Guerin-Meneville FE (Ed.) Iconographie de Regne Animal de G. Cuvier. Insectes. Bailliere, Paris 393 - 395. https: // www. biodiversitylibrary. org / item / 88601 # page / 399 / mode / 1 up" url="https://www.biodiversitylibrary.org/item/88601#page/399/mode/1up" year="1843">Guérin-Méneville, 1843</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B01FF173BA21C95DCFC5097275AC27A9" authority="Rauser, 1962" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="buresi">
|
||||
<emphasis id="B930A4995E921877BA0F07258ED49310" italics="true">Isoperla buresi</emphasis>
|
||||
<bibRefCitation id="C62486DB9011D81952D011D5D9EC92A3" author="Raušer J" issue="70" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="67-82" refId="B66" refString="Rauser J (1962) Plecoptera bulgarica - I. Acta Faunistica Entomologica Musei Nationalis Pragae 8 (70): 67 - 82. https: // www. aemnp. eu / data / article- 91 / 72 - 8 _ 0 _ 67. pdf" title="Plecoptera bulgarica - I." url="https://www.aemnp.eu/data/article-91/72-8_0_67.pdf" volume="8" year="1962">Raušer, 1962</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="79A22F90F72ABF12B78CB174536FC0B8" authority="(Rostock, 1881)" baseAuthorityName="Rostock" baseAuthorityYear="1881" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="neglecta">
|
||||
<emphasis id="E01DB81F32872006B7ADEF7CF2E0B35D" italics="true">Siphonoperla neglecta</emphasis>
|
||||
(
|
||||
<bibRefCitation id="D9DAA556ED76D8EC07596E01E0E86683" author="Rostock M" issue="15" journalOrPublisher="Entomologische Nachrichten" pagination="217-228" refId="B73" refString="Rostock M (1881) Verzeichniss der Neuropteren Deutschlands (1), Oesterreichs (2) und der Schweiz (3). Entomologische Nachrichten 7 (15): 217 - 228. https: // www. biodiversitylibrary. org / item / 110225 # page / 221 / mode / 1 up" title="Verzeichniss der Neuropteren Deutschlands (1), Oesterreichs (2) und der Schweiz (3)." url="https://www.biodiversitylibrary.org/item/110225#page/221/mode/1up" volume="7" year="1881">Rostock, 1881</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
;
|
||||
<date id="19E05683D69CAB6D57A8F62CE5569290" value="2013-05-27">27. v. 2013</date>
|
||||
(leg. P. Juhász, T. Kovács, G. Magos, G. Puskás): adults of
|
||||
<taxonomicName id="6A4FE3EA20CAD035AFAC5EE358E7ECD1" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
|
||||
<emphasis id="F00F0AC0B87B5F163B3FDED5AC3E2D6B" italics="true">B. helenica</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4E1D8E48C9D672A6F4524C1A80BF308E" authorityName="Kacanski & Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
|
||||
<emphasis id="B263FC181753FA31B9BCC06142AC1739" italics="true">L. hippopoides</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="44B8CE2D5488ED1168BA1A47C5CBED80" authority="Vincon & Muranyi, 2007" authorityName="Vincon & Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||||
<emphasis id="A04547F6D8DDD48A43AF5C2B0B1CBB52" italics="true">L.</emphasis>
|
||||
cf.
|
||||
<emphasis id="B83C8CAD2724BFF191A184F2946A2FA8" italics="true">dalmoni</emphasis>
|
||||
<bibRefCitation id="018B05331D9BD2D1E9379F48BF9FF837" author="Vinçon G & Murányi D" issue="3" journalOrPublisher="Nouvelle Revue d’Entomologie" pagination="237-248" refId="B82" refString="Vincon G, Muranyi D (2007) Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera). Nouvelle Revue d'Entomologie 23 (3): 237 - 248." title="Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera)." volume="23" year="2007">Vinçon & Murányi, 2007</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="80913B4B1B2D6AE12EBA48DEA419A09D" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
|
||||
<emphasis id="B8C2B79820AB8C9CDEBFE4A326862350" italics="true">Nemoura cinerea cinerea</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="0E40FC64D12715B608660D5D0389CE9F" authority="Despax, 1934" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
|
||||
<emphasis id="943AF154AB664A78EF51AF27BD097531" italics="true">N. uncinata</emphasis>
|
||||
<bibRefCitation id="B6F6E8F3123F42D70F94732666B04A66" author="Despax R" issue="2" journalOrPublisher="Bulletin de la Société d’histoire Naturelle de Toulouse" pagination="255-270" refId="B15" refString="Despax R (1934) Plecopteres pyreneens. VII. Etude et description de quelques formes de Nemoures apparentees a Nemoura marginata (Pict.). Bulletin de la Societe d'histoire Naturelle de Toulouse 66 (2): 255 - 270." title="Plécoptères pyrénéens. VII. Etude et description de quelques formes de Nemoures apparentées à Nemoura marginata (Pict.)." volume="66" year="1934">Despax, 1934</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="DA218B67B88F9990FBBF70A2CA1F4383" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
|
||||
<emphasis id="F855F3AE815534D6DD11C97897BC39CA" italics="true">N. marginata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="923121EB61F39E9C3FD54465F44B2523" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
|
||||
<emphasis id="B6E362DB5BE1A38C3A1BEF31392D234A" italics="true">Nemurella pictetii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C2C61A34212082A4F37B7A173EFA2856" baseAuthorityName="Rostock" baseAuthorityYear="1881" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="neglecta">
|
||||
<emphasis id="670F5D5021FBF8BD0F35C8050CBE5B00" italics="true">Siphonoperla neglecta</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="70B7BFA1E367F777E1F46370417C4CF3" authority="(Aubert, 1956)" baseAuthorityName="Aubert" baseAuthorityYear="1956" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="graeca">
|
||||
<emphasis id="7800E1251F991AFCDD7DE8F3E4025899" italics="true">S. graeca</emphasis>
|
||||
(Aubert, 1956)
|
||||
</taxonomicName>
|
||||
, and putative larvae of the new species were found.
|
||||
<taxonomicName id="5AC9E7ABB5840518FCC2E1143D350015" authorityName="Vincon & Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||||
<emphasis id="BE2691385E5499FA856BA96C68025012" italics="true">Leuctra</emphasis>
|
||||
cf.
|
||||
<emphasis id="6B606F0801D4B6744D7F5AE575A3D0A8" italics="true">dalmoni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="3C7A3AFFAB2EC1075BBFB772097251E5" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
|
||||
<emphasis id="8E38907E8C8112A0046CF9A851558814" italics="true">Nemoura uncinata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="178676AFA9BB2B5E9221B58A315F06D0" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
|
||||
<emphasis id="59C181CCC2DF19E1EF614F14833C383E" italics="true">N. marginata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="F90CFD8D187A303381AAB4098E59973F" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="buresi">
|
||||
<emphasis id="13C9E211C60536F4437D41BF1BD8671A" italics="true">Isoperla buresi</emphasis>
|
||||
</taxonomicName>
|
||||
are new findings for
|
||||
<collectingCountry id="7237F6113BEBBB14A4679FC610DEB376" name="Albania">Albania</collectingCountry>
|
||||
. Records of other aquatic insects collected at the type locality were already published:
|
||||
<taxonomicName id="FAF377EF32C69C189CCAE4FC8F9421B7" authority="(Kovacs and Muranyi 2013)" baseAuthorityName="Kovacs and Muranyi" baseAuthorityYear="2013" class="Insecta" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="order">
|
||||
Ephemeroptera (
|
||||
<bibRefCitation id="F3457D02FC142EB3B60554A3B7B65BDD" author="Kovacs and Muranyi" firstAuthor="Kovacs" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="15-19" refId="B38" refString="Kovács T, Murányi D (2013) New country data of some mayflies (Ephemeroptera) from Europe. Folia historico-naturalia Musei Matraensis 37: 15–19. http://www.nhmus.hu/sites/default/files/balkan/literature/Ephemeroptera/Kovacs_Muranyi_Fh-nMM_15_2013.pdf" title="New country data of some mayflies (Ephemeroptera) from Europe." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Ephemeroptera/Kovacs_Muranyi_Fh-nMM_15_2013.pdf" volume="37" year="2013">Kovács and Murányi 2013</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
:
|
||||
<taxonomicName id="18D8EF744D04B92B588112D77C4DB5DA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="muticus">
|
||||
<emphasis id="2BF44927C61715D2359780C52740A76B" italics="true">Baetis muticus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="2DD6BA9C057747F7C4D52829C273555E" authorityName="McLachlan" authorityYear="1884" class="Insecta" family="Leptophlebiidae" genus="Habrophlebia" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="lauta">
|
||||
<emphasis id="18CC2CB0AFFF8C30F2AB4BC0EE80AE2C" italics="true">Habrophlebia lauta</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="D806CF5B3C24F3904C819F3CA9EE2E99" authority="(Muranyi and Kovacs 2013)" baseAuthorityName="Muranyi and Kovacs" baseAuthorityYear="2013" class="Insecta" kingdom="Animalia" order="Odonata" phylum="Arthropoda" rank="order">
|
||||
Odonata (
|
||||
<bibRefCitation id="7A5B3284AEE9F8D3C68F160112240D77" author="Muranyi and Kovacs" firstAuthor="Muranyi" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="29-41" refId="B48" refString="Murányi D, Kovács T (2013) Contribution to the Odonata fauna of Albania and Montenegro. Folia historico-naturalia Musei Matraensis 37: 29–41. http://www.nhmus.hu/sites/default/files/balkan/literature/Odonata/Muranyi_Kovacs_2013.pdf" title="Contribution to the Odonata fauna of Albania and Montenegro." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Odonata/Muranyi_Kovacs_2013.pdf" volume="37" year="2013">Murányi and Kovács 2013</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
:
|
||||
<taxonomicName id="993C21E416868E5B411206A7C1814A9A" authorityName="Selys" authorityYear="1843" class="Insecta" family="Cordulegastridae" genus="Cordulegaster" kingdom="Animalia" order="Odonata" phylum="Arthropoda" rank="species" species="bidentata">
|
||||
<emphasis id="15D18A3A955D7E57F46D9BBE16DC1421" italics="true">Cordulegaster bidentata</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="165FA26D4636BB04F95D4D7FA8E7644C" class="Insecta" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="1B93B582B68DFC2BA5560E87825C4EA5" author="Oláh J & Kovács T" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="89-104" refId="B54" refString="Oláh J, Kovács T (2012 b) New species and records of autumnal Trichoptera form Albania. Folia historico-naturalia Musei Matraensis 36: 89–104. http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_Fh-nMM_89_2012.pdf" title="New species and records of autumnal Trichoptera form Albania." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_Fh-nMM_89_2012.pdf" volume="36">Oláh and Kovács 2012 b</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="DAA80A8EEDF8D8741F10159D84106C3F" author="Olah and Kovacs" firstAuthor="Olah" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="109-121" refId="B55" refString="Oláh J, Kovács T (2013) New species and new records of Balkan Trichoptera II. Folia historico-naturalia Musei Matraensis 37: 109–121. http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_2013.pdf" title="New species and new records of Balkan Trichoptera II." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_2013.pdf" volume="37" year="2013">2013</bibRefCitation>
|
||||
):
|
||||
<taxonomicName id="F7900F4623E58251B15B6D7EE9CD0CCF" baseAuthorityName="McLachlan" baseAuthorityYear="1879" class="Insecta" family="Glossosomatidae" genus="Agapetus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="iridipennis">
|
||||
<emphasis id="733610DD353D3F47041EAA2D534CAFDC" italics="true">Agapetus iridipennis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="41819550EA61828BDE137DC2AC45772B" authorityName="Marinkovic-Gospodnetic" authorityYear="1959" class="Insecta" family="Limnephilidae" genus="Chaetopteryx" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="bosniaca">
|
||||
<emphasis id="3A772455527FD3B8EC3CA8129C496CA5" italics="true">Chaetopteryx bosniaca</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="A6B8F1632D411E60F6011DD4200705A5" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Psychomyiidae" genus="Psychomyia" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pusilla">
|
||||
<emphasis id="05BDFA19C9CD3E321BA1226DD7C90F7C" italics="true">Psychomyia pusilla</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="92E3C1251A2E513EF8A9C1FA3F092F83" baseAuthorityName="Curtis" baseAuthorityYear="1834" class="Insecta" family="Polycentropodidae" genus="Cyrnus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="trimaculatus">
|
||||
<emphasis id="D84842B959EAFF2E83A011B1A1D05C57" italics="true">Cyrnus trimaculatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="FD8B1C968CC28F7BA0B232F5BB619EAC" baseAuthorityName="Fabricius" baseAuthorityYear="1775" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="hirtum">
|
||||
<emphasis id="9FD7B7A4A53843888415D2A766EC73E0" italics="true">Lepidostoma hirtum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="938DD4EC3E58B9F4E63F0E051FF8EC53" authorityName="Radovanovic" authorityYear="1942" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="plicatus">
|
||||
<emphasis id="188E128DC127B404EB363EAF72C1C3C9" italics="true">Drusus plicatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="3C22A32B44D4E751C21A8AF6581F6E42" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" class="Insecta" family="Leptoceridae" genus="Mystacides" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="azureus">
|
||||
<emphasis id="6384D7EB7066747EDBB799BB2F17CDEC" italics="true">Mystacides azureus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="7197664E3E06018A98E32208A27A4FCA" ID-DOI="10.3897/zookeys.1218.120744.figure4" ID-arpha="A4EE17C3-ABA5-5EA7-B6E3-CBD7B4B8796E" httpUri="https://binary.pensoft.net/fig/1175455" startId="F4">
|
||||
<paragraph id="7C6A4ADFA8BB6DF4AE28ECF491E4D57F">
|
||||
<label id="37D3644262E9DB742CAFA0F0AB03F299">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="569C3E12E4CCBCDD6E2AA4F253202DD7">
|
||||
Habitat of
|
||||
<taxonomicName id="4075DD850E8B65937F60249371526DE4" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
|
||||
<emphasis id="FBAFE96B5794780A3EDFAD0A1A60342F" italics="true">Leuctra enigma</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="48CEC4B234897FDBB9FA2A5CE584F936" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="7F286AD9D4E26DC3AC78A5FF6D23F0C1" bold="true">A</emphasis>
|
||||
type locality: partly open brook in the Çermenikë Mts.
|
||||
<emphasis id="4DCB5F2DC35BAE63C0BD06E6E0E382B2" bold="true">B</emphasis>
|
||||
section at forest edge
|
||||
<emphasis id="DF334195AF19C2B43FDA4B3DA9ED2057" bold="true">C</emphasis>
|
||||
substrate of the type locality.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EIWAE" type="etymology">
|
||||
<paragraph id="9DD90B641DE90E886F592B8A2612B990">
|
||||
<heading id="811205C22358A62CCD7A7DCE4F4A3A37" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="D034F2BC25B5BE653F581DF011CA8418">
|
||||
The name
|
||||
<emphasis id="C015550C96019F475BEACBA980E8E70A" italics="true">enigma</emphasis>
|
||||
(Latin; enigma or riddle) refers to the strange fact that such a distinctive new species was found only at a single locality. Used as a noun, gender neutral.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
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