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<mods:title id="5DADA8B2F29BB9E80E1AC2D11CF9455A">The genus Dedalopterus Sabatinelli &amp; Pontuale, 1998 (Scarabaeidae: Melolonthinae: Leucopholini) in Laos and Vietnam, with description of a new species</mods:title>
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<taxonomicName id="D8C6C59CFFD0FFD1FF11FC20FD8D3DF4" ID-CoL="6CGVD" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[151,601,898,925]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD0FFD1FF11FC20FD8D3DF4" bold="true" box="[151,601,898,925]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF11FC20FE4E3DF5" bold="true" box="[151,410,898,925]" italics="true" pageId="6" pageNumber="147">Dedalopterus malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FE27FC20FD8D3DF4" author="Bunalski, M." box="[417,601,898,924]" pageId="6" pageNumber="147" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
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(
<figureCitation id="87FDA29AFFD0FFD1FF19FC05FECA3DA9" box="[159,286,935,961]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="6" pageNumber="147">Figs 2CD</figureCitation>
;
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;
<figureCitation id="87FDA29AFFD0FFD1FEFEFC05FE683DA9" box="[376,444,935,961]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1935]" captionTargetBox="[151,1424,181,1885]" captionTargetId="figure-20@5.[151,1436,181,1887]" captionTargetPageId="5" captionText="FIGURE 4. Lateral view of elytra (AD), right lateral view of parameres tip (EI). A, E, Dedalopterus malyszi Bunalski, 2001; B, G, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, H, D. pulchellus Sabatinelli &amp; Pontuale, 1998 (paratype of D. pulchellus); D, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611475" httpUri="https://zenodo.org/record/10611475/files/figure.png" pageId="6" pageNumber="147">4A, E</figureCitation>
;
<figureCitation id="87FDA29AFFD0FFD1FE4EFC05FDE43DA9" box="[456,560,935,961]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[161,1436,192,1626]" captionTargetId="figure-27@8.[151,1436,181,1626]" captionTargetPageId="8" captionText="FIGURE 6. Antennae, protibiae, and claws of Dedalopterus species.A, D, G, D. malyszi Bunalski, 2001; B, E, H, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611479" httpUri="https://zenodo.org/record/10611479/files/figure.png" pageId="6" pageNumber="147">6A, D, G</figureCitation>
)
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<emphasis id="2DB2620DFFD0FFD1FF11FC45FEA33D97" box="[151,375,999,1023]" italics="true" pageId="6" pageNumber="147">Dedalopterus malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FE06FC45FD823D97" author="Bunalski, M." box="[384,598,999,1023]" pageId="6" pageNumber="147" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001: 248</bibRefCitation>
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, fig. 13 (
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locality:
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: Cha-Pa [Sa Pa]);
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, fig. 2ag.
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material examined.
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None.
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<emphasis id="2DB2620DFFD0FFD1FF41FBCEFDED3AEE" bold="true" box="[199,569,1132,1158]" pageId="6" pageNumber="147">Additional material examined.</emphasis>
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,
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,&gt;
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)
</materialsCitation>
;
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<collectingRegion id="DD0270FDFFD0FFD1FE64FB16FD503AA7" box="[482,644,1204,1231]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="147">Lao Cai Prov.</collectingRegion>
,
<location id="1A19E8C4FFD0FFD1FD09FB17FCC13AA6" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFD0FFD1FD09FB17FCC13AA6" box="[655,789,1204,1231]" country="Vietnam" municipality="Thac Bac" name="Hoang Lien" pageId="6" pageNumber="147" stateProvince="Lao Cai">Hoang Lien</location>
NP.
<collectingMunicipality id="FF1D2465FFD0FFD1FCCAFB16FC6E3AA7" box="[844,954,1204,1231]" pageId="6" pageNumber="147">Thac Bac</collectingMunicipality>
,
<quantity id="D83E13FAFFD0FFD1FC40FB16FBC33AA6" box="[966,1047,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="1.788" pageId="6" pageNumber="147" unit="m" value="1788.0">
<elevation id="94EB592CFFD0FFD1FC40FB16FBC33AA6" box="[966,1047,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="1.788" pageId="6" pageNumber="147" unit="m" value="1788.0">1788m</elevation>
</quantity>
, 22.3591171˚N 103.7802435˚E, at light,
<date id="6B7898DFFFD0FFD1FF5DFB7AFE873A9A" box="[219,339,1240,1266]" pageId="6" pageNumber="147" value="2010-04-12">
<collectingDate id="7B3C6137FFD0FFD1FF5DFB7AFE873A9A" box="[219,339,1240,1266]" pageId="6" pageNumber="147" value="2010-04-12">12.iv.2010</collectingDate>
</date>
, leg.
<collectorName id="B233DBC9FFD0FFD1FE15FB7BFE3B3A9B" box="[403,495,1241,1267]" pageId="6" pageNumber="147">L. Papp</collectorName>
,
<collectorName id="B233DBC9FFD0FFD1FE7AFB7BFD413A9A" box="[508,661,1240,1267]" pageId="6" pageNumber="147">L. Peregovitz</collectorName>
&amp;
<collectorName id="B233DBC9FFD0FFD1FD3BFB7BFCE63A9A" box="[701,818,1240,1267]" pageId="6" pageNumber="147">Z. Soltesz</collectorName>
,
<collectingCountry id="67D1FE8FFFD0FFD1FCB9FB7AFCBC3A9A" box="[831,872,1240,1266]" name="Vietnam" pageId="6" pageNumber="147">VN</collectingCountry>
2010PL_9” (
<collectionCode id="79D726DAFFD0FFD1FC7FFB7AFB813A9A" box="[1017,1109,1240,1266]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="147" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFD0FFD1FBECFB7AFD193B52" collectingDate="2010-04-08" collectingDateMax="2010-04-11" collectingDateMin="2010-04-08" collectionCode="MHNG" collectorName="L. Papp &amp; L. Peregovitz &amp; Z. Soltesz" country="Vietnam" elevation="1915" location="Hoang Lien" municipality="Tram Ton" pageId="6" pageNumber="147" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<specimenCount id="09C07596FFD0FFD1FBECFB7AFB593A9A" box="[1130,1165,1240,1266]" count="1" pageId="6" pageNumber="147" type="male">1♂</specimenCount>
<collectingRegion id="DD0270FDFFD0FFD1FB27FB7AFA913A9B" box="[1185,1349,1240,1267]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="147">Lao Cai Prov.</collectingRegion>
,
<location id="1A19E8C4FFD0FFD1FAD4FB7BFF1E3B7E" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFD0FFD1FAD4FB7BFF1E3B7E" country="Vietnam" municipality="Tram Ton" name="Hoang Lien" pageId="6" pageNumber="147" stateProvince="Lao Cai">Hoang Lien</location>
NP.
<collectingMunicipality id="FF1D2465FFD0FFD1FE80FB5FFEAD3B7F" box="[262,377,1277,1303]" pageId="6" pageNumber="147">Tram Ton</collectingMunicipality>
,
<quantity id="D83E13FAFFD0FFD1FE01FB5EFE0C3B7F" box="[391,472,1276,1303]" metricMagnitude="3" metricUnit="m" metricValue="1.9149999999999998" pageId="6" pageNumber="147" unit="m" value="1915.0">
<elevation id="94EB592CFFD0FFD1FE01FB5EFE0C3B7F" box="[391,472,1276,1303]" metricMagnitude="3" metricUnit="m" metricValue="1.9149999999999998" pageId="6" pageNumber="147" unit="m" value="1915.0">1915m</elevation>
</quantity>
, 22.3493723˚N 103.7704565˚E, forest edge, at light,
<date id="6B7898DFFFD0FFD1FBC5FB5EFB033B7E" box="[1091,1239,1276,1302]" pageId="6" pageNumber="147" value="2010-04-08" valueMax="2010-04-11" valueMin="2010-04-08">
<collectingDate id="7B3C6137FFD0FFD1FBC5FB5EFB033B7E" box="[1091,1239,1276,1302]" pageId="6" pageNumber="147" value="2010-04-08" valueMax="2010-04-11" valueMin="2010-04-08">8-11.IV.2010</collectingDate>
</date>
, leg.
<collectorName id="B233DBC9FFD0FFD1FA9FFB5FFAA23B7F" box="[1305,1398,1277,1303]" pageId="6" pageNumber="147">L. Papp</collectorName>
,
<collectorName id="B233DBC9FFD0FFD1FA02FB5FFEDB3B52" pageId="6" pageNumber="147">L. Peregovitz</collectorName>
&amp;
<collectorName id="B233DBC9FFD0FFD1FEB5FA83FE723B52" box="[307,422,1312,1339]" pageId="6" pageNumber="147">Z. Soltesz</collectorName>
,
<collectingCountry id="67D1FE8FFFD0FFD1FE36FA82FE0C3B52" box="[432,472,1312,1338]" name="Vietnam" pageId="6" pageNumber="147">VN</collectingCountry>
2010PL_4” (
<collectionCode id="79D726DAFFD0FFD1FDEFFA82FD113B52" box="[617,709,1312,1338]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="147" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41FAE6FD6A3B86" pageId="6" pageNumber="147" type="diagnosis">
<paragraph id="1F79BE1FFFD0FFD1FF41FAE6FD6A3B86" blockId="6.[151,1437,1096,1627]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FAE6FE963B36" bold="true" box="[199,322,1348,1374]" pageId="6" pageNumber="147">Diagnosis.</emphasis>
Costae 23 unclearly subdivided into two stripes jointed past middle; intervals 13 and elytra suture, bearing white large dense scales; interval 4 with yellow narrow scales; epipleura bear short and dense yellow hairlake scales, one row of narrow yellow scales in male and epipleura bearing white, large scales in female. Male genitalia: long and simple; basal part longer, asymmetric paramere; left paramere usually bigger and longer than right; curved at tips and very short piled beneath.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41FA5AFCC4387A" box="[199,784,1528,1555]" pageId="6" pageNumber="147" type="distribution">
<paragraph id="1F79BE1FFFD0FFD1FF41FA5AFCC4387A" blockId="6.[151,1437,1096,1627]" box="[199,784,1528,1555]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FA5AFEB5387A" bold="true" box="[199,353,1528,1554]" pageId="6" pageNumber="147">Distribution.</emphasis>
Known only from
<collectingCountry id="67D1FE8FFFD0FFD1FDB1FA5AFD42387A" box="[567,662,1528,1554]" name="Vietnam" pageId="6" pageNumber="147">Vietnam</collectingCountry>
(
<collectingRegion id="DD0270FDFFD0FFD1FD20FA5BFCD7387A" box="[678,771,1528,1555]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="147">Lao Cai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41F9BEFF3F388B" pageId="6" pageNumber="147" type="discussion">
<paragraph id="1F79BE1FFFD0FFD1FF41F9BEFAFB3832" blockId="6.[151,1437,1096,1627]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41F9BEFEEF385E" bold="true" box="[199,315,1564,1590]" pageId="6" pageNumber="147">Remarks.</emphasis>
A male specimen collected in
<collectingRegion id="DD0270FDFFD0FFD1FD15F9BFFC8C385E" box="[659,856,1564,1591]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="147">Lao Cai Province</collectingRegion>
has epipleura with short and dense yellow hair-like scales, mixed with many large white scales. This means that the species has some considerable variation.
</paragraph>
<paragraph id="1F79BE1FFFD0FFD1FF11F901FCCC38D6" blockId="6.[151,792,1699,1763]" box="[151,792,1699,1726]" pageId="6" pageNumber="147">
<taxonomicName id="D8C6C59CFFD0FFD1FF11F901FCCC38D6" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[151,792,1699,1726]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FF11F901FE6A38D6" bold="true" box="[151,446,1699,1726]" italics="true" pageId="6" pageNumber="147">Dedalopterus pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FE43F906FCCC38D6" author="Sabatinelli, G. &amp; Pontuale, G." box="[453,792,1700,1726]" pageId="6" pageNumber="147" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1F79BE1FFFD0FFD1FF11F96AFF3F388B" blockId="6.[151,792,1699,1763]" box="[151,235,1736,1763]" pageId="6" pageNumber="147">
(
<figureCitation id="87FDA29AFFD0FFD1FF19F96AFF36388B" box="[159,226,1736,1763]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1356,190,1910]" captionTargetId="figure-20@7.[221,1365,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Variation in male parameres of Dedalopterus pulchellus Sabatinelli &amp; Pontuale, 1998 (AJ, all from Laos: Phu Phan). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10611477" httpUri="https://zenodo.org/record/10611477/files/figure.png" pageId="6" pageNumber="147">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF11F8AEFC62392B" pageId="6" pageNumber="147" type="reference_group">
<paragraph id="1F79BE1FFFD0FFD1FF11F8AEFB17394C" blockId="6.[151,1219,1804,1859]" box="[151,1219,1804,1829]" pageId="6" pageNumber="147">
<treatmentCitationGroup id="3FD69931FFD0FFD1FF11F8AEFB17394C" box="[151,1219,1804,1829]" pageId="6" pageNumber="147">
<treatmentCitation id="9E67980EFFD0FFD1FF11F8AEFD30394C" author="Sabatinelli, G. &amp; Pontuale, G." box="[151,740,1804,1829]" page="69" pageId="6" pageNumber="147" year="1998">
<taxonomicName id="D8C6C59CFFD0FFD1FF11F8AEFD30394C" ID-CoL="6CGVF" authority="Sabatinelli &amp; Pontuale, 1998: 69" authorityName="Sabatinelli &amp; Pontuale" authorityPageNumber="69" authorityYear="1998" box="[151,740,1804,1829]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FF11F8AEFE5B394C" box="[151,399,1804,1828]" italics="true" pageId="6" pageNumber="147">Dedalopterus pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FE10F8AEFD30394C" author="Sabatinelli, G. &amp; Pontuale, G." box="[406,740,1804,1829]" pageId="6" pageNumber="147" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998: 69</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="9E67980EFFD0FFD1FD69F8AEFC18394D" author="Keith, D. &amp; Jingke, L. &amp; Xueping, Z. &amp; Lirong, H." box="[751,972,1804,1829]" page="387" pageId="6" pageNumber="147" year="2012">
<bibRefCitation id="7B57C3EEFFD0FFD1FD69F8AEFC18394D" author="Keith, D. &amp; Jingke, L. &amp; Xueping, Z. &amp; Lirong, H." box="[751,972,1804,1829]" pageId="6" pageNumber="147" pagination="387 - 397" refId="ref6318" refString="Keith, D., Jingke, L., Xueping, Z. &amp; Lirong, H. (2012) Scarabaeoidea Pleurosticti recoltes sur le Mt. Gaoligongshan en Republique Populaire de Chine (Insecta: Coleoptera). Vernate, 31, 387 - 397." type="journal article" year="2012">
Keith
<emphasis id="2DB2620DFFD0FFD1FCABF8AFFCB5394C" box="[813,865,1804,1828]" italics="true" pageId="6" pageNumber="147">et al.</emphasis>
2012: 387
</bibRefCitation>
</treatmentCitation>
(new record for
<collectingCountry id="67D1FE8FFFD0FFD1FBF1F8AEFB6C394C" box="[1143,1208,1804,1828]" name="China" pageId="6" pageNumber="147">China</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="1F79BE1FFFD0FFD1FF11F889FC62392B" blockId="6.[151,1219,1804,1859]" box="[151,950,1835,1859]" pageId="6" pageNumber="147">
<treatmentCitationGroup id="3FD69931FFD0FFD1FF11F889FC62392B" box="[151,950,1835,1859]" pageId="6" pageNumber="147">
<treatmentCitation id="9E67980EFFD0FFD1FF11F889FD7E392B" author="Zidek, J. &amp; Krajcik, M." box="[151,682,1835,1859]" page="9" pageId="6" pageNumber="147" year="2007">
<taxonomicName id="D8C6C59CFFD0FFD1FF11F889FD7E392B" ID-CoL="6CH83" authority="Zidek &amp; Krajcik, 2007: 9" authorityName="Zidek &amp; Krajcik" authorityPageNumber="9" authorityYear="2007" box="[151,682,1835,1859]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="bezdekorum">
<emphasis id="2DB2620DFFD0FFD1FF11F889FE75392B" box="[151,417,1835,1859]" italics="true" pageId="6" pageNumber="147">Dedalopterus bezdekorum</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FE2EF889FD7E392B" author="Zidek, J. &amp; Krajcik, M." box="[424,682,1835,1859]" pageId="6" pageNumber="147" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007: 9</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig 3ag,
<emphasis id="2DB2620DFFD0FFD1FC90F889FC62392B" bold="true" box="[790,950,1835,1859]" pageId="6" pageNumber="147">new synonymy</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFDFFF11F8D3FE6E39A7" lastPageId="8" lastPageNumber="149" pageId="6" pageNumber="147" type="materials_examined">
<paragraph id="1F79BE1FFFD0FFD1FF11F8D3FE4B399F" blockId="6.[151,1436,1905,2039]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF11F8D3FE1539E3" bold="true" box="[151,449,1905,1931]" pageId="6" pageNumber="147">Type material examined.</emphasis>
<materialsCitation id="AFAEB442FFD0FFD1FE4FF8D0FCBA39C7" collectingDate="1916-06-16" collectionCode="MHNG" collectorName="R. Vitalis de Salvaza" country="Laos" location="Vitalis de Salvaza" municipality="Kieng Kouang" pageId="6" pageNumber="147" specimenCount="1" specimenCount-male="1" stateProvince="Xiangkhouang" typeStatus="paratype">
<typeStatus id="C07D00BDFFD0FFD1FE4FF8D0FDFF39E4" box="[457,555,1906,1932]" pageId="6" pageNumber="147" type="paratype">Paratype</typeStatus>
(
<specimenCount id="09C07596FFD0FFD1FDBBF8D3FD8739E3" box="[573,595,1905,1931]" count="1" pageId="6" pageNumber="147" type="male"></specimenCount>
) of
<taxonomicName id="D8C6C59CFFD0FFD1FD05F8D0FBBD39E3" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[643,1129,1905,1931]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FD05F8D0FCCD39E3" box="[643,793,1905,1931]" italics="true" pageId="6" pageNumber="147">D. pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FCA4F8D3FBBD39E3" author="Sabatinelli, G. &amp; Pontuale, G." box="[802,1129,1905,1931]" pageId="6" pageNumber="147" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
<collectingCountry id="67D1FE8FFFD0FFD1FBFDF8D3FB1D39E3" box="[1147,1225,1905,1931]" name="Laos" pageId="6" pageNumber="147">LAOS</collectingCountry>
,
<collectingMunicipality id="FF1D2465FFD0FFD1FB50F8D3FAAF39E4" box="[1238,1403,1905,1932]" pageId="6" pageNumber="147">Kieng Kouang</collectingMunicipality>
[=
<collectingRegion id="DD0270FDFFD0FFD1FF11F837FE9639C7" box="[151,322,1941,1967]" country="Laos" name="Xiangkhouang" pageId="6" pageNumber="147">Xiangkhouang</collectingRegion>
],
<date id="6B7898DFFFD0FFD1FED5F837FE1939C7" box="[339,461,1941,1967]" pageId="6" pageNumber="147" value="1916-06-16">
<collectingDate id="7B3C6137FFD0FFD1FED5F837FE1939C7" box="[339,461,1941,1967]" pageId="6" pageNumber="147" value="1916-06-16">16.vi.1916</collectingDate>
</date>
,
<collectorName id="B233DBC9FFD0FFD1FE5EF834FD6B39C7" box="[472,703,1941,1968]" pageId="6" pageNumber="147">
R.
<location id="1A19E8C4FFD0FFD1FE7EF837FD6B39C7" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFD0FFD1FE7EF837FD6B39C7" box="[504,703,1941,1967]" country="Laos" municipality="Kieng Kouang" name="Vitalis de Salvaza" pageId="6" pageNumber="147" stateProvince="Xiangkhouang">Vitalis de Salvaza</location>
</collectorName>
leg.” (
<collectionCode id="79D726DAFFD0FFD1FC8CF837FCB239C7" box="[778,870,1941,1967]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="147" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFD0FFD1FCFFF834FE4F399F" collectingDate="2001-04-26" collectingDateMax="2001-05-11" collectingDateMin="2001-04-26" collectionCode="MHNG" collectorName="J. Bezdek" country="Laos" elevation="1750" location="Phou Pan Mt" municipality="Ban Saluei" pageId="6" pageNumber="147" specimenCount="1" specimenCount-male="1" stateProvince="Houaphan" typeStatus="paratype">
<typeStatus id="C07D00BDFFD0FFD1FCFFF834FC0F39D8" box="[889,987,1942,1968]" pageId="6" pageNumber="147" type="paratype">Paratype</typeStatus>
(
<specimenCount id="09C07596FFD0FFD1FC6CF837FBD439C7" box="[1002,1024,1941,1967]" count="1" pageId="6" pageNumber="147" type="male"></specimenCount>
) of
<taxonomicName id="D8C6C59CFFD0FFD1FBABF834FF1B39BB" authority="Zidek &amp; Krajcik, 2007" authorityName="Zidek &amp; Krajcik" authorityYear="2007" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="bezdekorum">
<emphasis id="2DB2620DFFD0FFD1FBABF834FB0239C7" box="[1069,1238,1941,1967]" italics="true" pageId="6" pageNumber="147">D. bezdekorum</emphasis>
<bibRefCitation id="7B57C3EEFFD0FFD1FB5BF837FF1B39BB" author="Zidek, J. &amp; Krajcik, M." pageId="6" pageNumber="147" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
</taxonomicName>
<collectingCountry id="67D1FE8FFFD0FFD1FF66F81BFEF839BB" box="[224,300,1977,2003]" name="Laos" pageId="6" pageNumber="147">LAOS</collectingCountry>
NE,
<collectingRegion id="DD0270FDFFD0FFD1FEE0F818FDC239BC" box="[358,534,1977,2004]" country="Laos" name="Houaphan" pageId="6" pageNumber="147">Hua Phan prov.</collectingRegion>
,
<collectingMunicipality id="FF1D2465FFD0FFD1FDA7F818FD4A39BB" box="[545,670,1977,2004]" pageId="6" pageNumber="147">Ban Saluei</collectingMunicipality>
,
<location id="1A19E8C4FFD0FFD1FD2FF81BFCE939BC" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFD0FFD1FD2FF81BFCE939BC" box="[681,829,1977,2004]" country="Laos" municipality="Ban Saluei" name="Phou Pan Mt" pageId="6" pageNumber="147" stateProvince="Houaphan">Phou Pan Mt</location>
, 20˚15N 104˚02E,
<quantity id="D83E13FAFFD0FFD1FBA0F81BFB6939BC" box="[1062,1213,1977,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="147" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">
<elevation id="94EB592CFFD0FFD1FBA0F81BFB6939BC" box="[1062,1213,1977,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="147" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">15002000m</elevation>
</quantity>
,
<date id="6B7898DFFFD0FFD1FB4EF81BFAAA39BB" box="[1224,1406,1977,2003]" pageId="6" pageNumber="147" value="2001-04-26" valueMax="2001-05-11" valueMin="2001-04-26">
<collectingDate id="7B3C6137FFD0FFD1FB4EF81BFAAA39BB" box="[1224,1406,1977,2003]" pageId="6" pageNumber="147" value="2001-04-26" valueMax="2001-05-11" valueMin="2001-04-26">26.iv.-11.v.2001</collectingDate>
</date>
,
<collectorName id="B233DBC9FFD0FFD1FA0CF818FF3F399F" pageId="6" pageNumber="147">J. Bezděk</collectorName>
leg.” (
<collectionCode id="79D726DAFFD0FFD1FEB1F87FFE47399F" box="[311,403,2013,2039]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="147" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="4BB9EE97FFD1FFD0FF11F835FE4B39BB" ID-DOI="http://doi.org/10.5281/zenodo.10611477" ID-Zenodo-Dep="10611477" httpUri="https://zenodo.org/record/10611477/files/figure.png" pageId="7" pageNumber="148" startId="7.[151,250,1943,1967]" targetBox="[230,1356,190,1910]" targetPageId="7" targetType="figure">
<paragraph id="1F79BE1FFFD1FFD0FF11F835FE4B39BB" blockId="7.[151,1436,1943,2003]" pageId="7" pageNumber="148">
<emphasis id="2DB2620DFFD1FFD0FF11F835FEC139C7" bold="true" box="[151,277,1943,1967]" pageId="7" pageNumber="148">FIGURE 5.</emphasis>
Variation in male parameres of
<taxonomicName id="D8C6C59CFFD1FFD0FDE3F835FB4639C7" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[613,1170,1943,1967]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD1FFD0FDE3F835FC8B39C7" box="[613,863,1943,1967]" italics="true" pageId="7" pageNumber="148">Dedalopterus pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFD1FFD0FCE1F835FB4639C7" author="Sabatinelli, G. &amp; Pontuale, G." box="[871,1170,1943,1967]" pageId="7" pageNumber="148" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
(AJ, all from Laos: Phu Phan). Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="4BB9EE97FFDEFFDFFF11F9D0FED338BA" ID-DOI="http://doi.org/10.5281/zenodo.10611479" ID-Zenodo-Dep="10611479" httpUri="https://zenodo.org/record/10611479/files/figure.png" pageId="8" pageNumber="149" startId="8.[151,250,1650,1674]" targetBox="[161,1436,192,1626]" targetPageId="8" targetType="figure">
<paragraph id="1F79BE1FFFDEFFDFFF11F9D0FED338BA" blockId="8.[151,1437,1650,1746]" pageId="8" pageNumber="149">
<emphasis id="2DB2620DFFDEFFDFFF11F9D0FEC638E2" bold="true" box="[151,274,1650,1674]" pageId="8" pageNumber="149">FIGURE 6.</emphasis>
Antennae, protibiae, and claws of
<taxonomicName id="D8C6C59CFFDEFFDFFDEBF9D0FCB038E3" box="[621,868,1650,1675]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="2DB2620DFFDEFFDFFDEBF9D0FD2338E2" box="[621,759,1650,1674]" italics="true" pageId="8" pageNumber="149">Dedalopterus</emphasis>
species. A
</taxonomicName>
, D, G,
<taxonomicName id="D8C6C59CFFDEFFDFFC2DF9D1FB6238E2" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[939,1206,1650,1674]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFDEFFDFFC2DF9D1FBC138E2" box="[939,1045,1650,1674]" italics="true" pageId="8" pageNumber="149">D. malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFDEFFDFFB9CF9D0FB6238E2" author="Bunalski, M." box="[1050,1206,1650,1674]" pageId="8" pageNumber="149" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
; B, E, H,
<taxonomicName id="D8C6C59CFFDEFFDFFA90F9D1FA4838E2" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[1302,1436,1650,1674]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFDEFFDFFA90F9D1FA4838E2" box="[1302,1436,1650,1674]" italics="true" pageId="8" pageNumber="149">D. pulchellus</emphasis>
</taxonomicName>
(paratype of
<taxonomicName id="D8C6C59CFFDEFFDFFEA4F935FD1038C6" authority="Zidek &amp; Krajcik, 2007" authorityName="Zidek &amp; Krajcik" authorityYear="2007" box="[290,708,1686,1710]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="bezdekorum">
<emphasis id="2DB2620DFFDEFFDFFEA4F935FE6B38C6" box="[290,447,1686,1710]" italics="true" pageId="8" pageNumber="149">D. bezdekorum</emphasis>
<bibRefCitation id="7B57C3EEFFDEFFDFFE4DF934FD1038C6" author="Zidek, J. &amp; Krajcik, M." box="[459,708,1686,1710]" pageId="8" pageNumber="149" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
</taxonomicName>
); C, F, I,
<taxonomicName id="D8C6C59CFFDEFFDFFCB5F935FAD838C6" authority="Pham, Ahrens and Sabatinelli" authorityName="Pham, Ahrens and Sabatinelli" authorityYear="2024" box="[819,1292,1686,1710]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFDEFFDFFCB5F935FC1138C6" box="[819,965,1686,1710]" italics="true" pageId="8" pageNumber="149">D. lexuanhuei</emphasis>
Pham, Ahrens and Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFDEFFDFFA9DF934FA4938C6" bold="true" box="[1307,1437,1686,1710]" pageId="8" pageNumber="149">
<taxonomicNameLabel id="3681DF76FFDEFFDFFA9DF934FA4938C6" box="[1307,1437,1686,1710]" pageId="8" pageNumber="149" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype).
</paragraph>
</caption>
<paragraph id="1F79BE1FFFDEFFDFFF41F886FE6E39A7" blockId="8.[151,1437,1828,1999]" pageId="8" pageNumber="149">
<emphasis id="2DB2620DFFDEFFDFFF41F886FDE13956" bold="true" box="[199,565,1828,1854]" pageId="8" pageNumber="149">Additional material examined.</emphasis>
<materialsCitation id="AFAEB442FFDEFFDFFDBAF887FC13390B" collectingDate="2012-04-11" collectingDateMax="2012-05-15" collectingDateMin="2012-04-11" collectionCode="ZFMK" collectorName="C. Holzschuh" country="Laos" elevation="1600" latitude="20.2" location="Phou Pan" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" pageId="8" pageNumber="149" specimenCount="156" specimenCount-female="58" specimenCount-male="98" stateProvince="Houaphan">
<specimenCount id="09C07596FFDEFFDFFDBAF887FD513957" box="[572,645,1829,1855]" count="98" pageId="8" pageNumber="149" type="male">98♂♂</specimenCount>
,
<specimenCount id="09C07596FFDEFFDFFD17F886FD023957" box="[657,726,1828,1855]" count="58" pageId="8" pageNumber="149" type="female">58♀♀</specimenCount>
<collectingCountry id="67D1FE8FFFDEFFDFFD6EF887FCE23957" box="[744,822,1829,1855]" name="Laos" pageId="8" pageNumber="149">LAOS</collectingCountry>
,
<collectingRegion id="DD0270FDFFDEFFDFFCC4F887FC643956" box="[834,944,1828,1855]" country="Laos" name="Houaphan" pageId="8" pageNumber="149">Hua Phan</collectingRegion>
Pro.,
<collectingMunicipality id="FF1D2465FFDEFFDFFC72F887FBA53956" box="[1012,1137,1828,1855]" pageId="8" pageNumber="149">Ban Saleui</collectingMunicipality>
,
<location id="1A19E8C4FFDEFFDFFBFAF886FB333957" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFDEFFDFFBFAF886FB333957" box="[1148,1255,1828,1855]" country="Laos" latitude="20.2" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" name="Phou Pan" pageId="8" pageNumber="149" stateProvince="Houaphan">Phou Pan</location>
(Mt.),
<geoCoordinate id="7AF2D8D8FFDEFFDFFABEF887FA433957" box="[1336,1431,1829,1855]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="8" pageNumber="149" precision="925" value="20.2">
20
<superScript id="E8B31357FFDEFFDFFAD2F887FA88395B" attach="none" box="[1364,1372,1829,1843]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
12N
</geoCoordinate>
,
<geoCoordinate id="7AF2D8D8FFDEFFDFFF11F8EBFED6390B" box="[151,258,1865,1891]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="149" precision="925" value="104.01667">
104
<superScript id="E8B31357FFDEFFDFFF47F8EBFF1D393F" attach="none" box="[193,201,1865,1879]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
01E
</geoCoordinate>
,
<date id="6B7898DFFFDEFFDFFE97F8EAFE1C390B" box="[273,456,1864,1891]" pageId="8" pageNumber="149" value="2012-04-11" valueMax="2012-05-15" valueMin="2012-04-11">
<collectingDate id="7B3C6137FFDEFFDFFE97F8EAFE1C390B" box="[273,456,1864,1891]" pageId="8" pageNumber="149" value="2012-04-11" valueMax="2012-05-15" valueMin="2012-04-11">11.iv15.v.2012</collectingDate>
</date>
,
<quantity id="D83E13FAFFDEFFDFFE51F8EBFDBA390B" box="[471,622,1865,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">
<elevation id="94EB592CFFDEFFDFFE51F8EBFDBA390B" box="[471,622,1865,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">13001900m</elevation>
</quantity>
,
<collectorName id="B233DBC9FFDEFFDFFDFAF8EBFCCC390A" box="[636,792,1864,1891]" pageId="8" pageNumber="149">C. Holzschuh</collectorName>
leg.” (
<collectionCode id="79D726DAFFDEFFDFFCEDF8EBFC6B390B" box="[875,959,1865,1891]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="149" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFDEFFDFFC50F8EBFA9539EF" collectingDate="2010-04-07" collectingDateMax="2010-05-25" collectingDateMin="2010-04-07" collectionCode="ZFMK" collectorName="C. Holzschuh" country="Laos" elevation="1600" latitude="20.2" location="Phou Pan" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" pageId="8" pageNumber="149" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Houaphan">
<specimenCount id="09C07596FFDEFFDFFC50F8EBFBC4390B" box="[982,1040,1865,1891]" count="2" pageId="8" pageNumber="149" type="male">2♂♂</specimenCount>
,
<specimenCount id="09C07596FFDEFFDFFB99F8EBFB96390B" box="[1055,1090,1865,1891]" count="1" pageId="8" pageNumber="149" type="female">1♀</specimenCount>
<collectingCountry id="67D1FE8FFFDEFFDFFBD1F8EBFB71390B" box="[1111,1189,1865,1891]" name="Laos" pageId="8" pageNumber="149">LAOS</collectingCountry>
,
<collectingRegion id="DD0270FDFFDEFFDFFB32F8EBFAF1390A" box="[1204,1317,1864,1891]" country="Laos" name="Houaphan" pageId="8" pageNumber="149">Hua Phan</collectingRegion>
Pro.,
<collectingMunicipality id="FF1D2465FFDEFFDFFAE9F8EBFF3439EE" pageId="8" pageNumber="149">Ban Saleui</collectingMunicipality>
,
<location id="1A19E8C4FFDEFFDFFF69F8CEFE8939EF" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFDEFFDFFF69F8CEFE8939EF" box="[239,349,1900,1927]" country="Laos" latitude="20.2" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" name="Phou Pan" pageId="8" pageNumber="149" stateProvince="Houaphan">Phou Pan</location>
(Mt.),
<geoCoordinate id="7AF2D8D8FFDEFFDFFE35F8CFFDC739EF" box="[435,531,1901,1927]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="8" pageNumber="149" precision="925" value="20.2">
20
<superScript id="E8B31357FFDEFFDFFE49F8CFFE033913" attach="none" box="[463,471,1901,1915]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
12N
</geoCoordinate>
,
<geoCoordinate id="7AF2D8D8FFDEFFDFFDA4F8CFFD5939EF" box="[546,653,1901,1927]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="149" precision="925" value="104.01667">
104
<superScript id="E8B31357FFDEFFDFFDCAF8CFFD803913" attach="none" box="[588,596,1901,1915]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
01E
</geoCoordinate>
,
<date id="6B7898DFFFDEFFDFFD1BF8CEFC9539EF" box="[669,833,1900,1927]" pageId="8" pageNumber="149" value="2010-04-07" valueMax="2010-05-25" valueMin="2010-04-07">
<collectingDate id="7B3C6137FFDEFFDFFD1BF8CEFC9539EF" box="[669,833,1900,1927]" pageId="8" pageNumber="149" value="2010-04-07" valueMax="2010-05-25" valueMin="2010-04-07">7.iv-25.v.2010</collectingDate>
</date>
,
<quantity id="D83E13FAFFDEFFDFFCD6F8CFFC3339EF" box="[848,999,1901,1927]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">
<elevation id="94EB592CFFDEFFDFFCD6F8CFFC3339EF" box="[848,999,1901,1927]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">13001900m</elevation>
</quantity>
,
<collectorName id="B233DBC9FFDEFFDFFC70F8CFFB4639EE" box="[1014,1170,1900,1927]" pageId="8" pageNumber="149">C. Holzschuh</collectorName>
leg.” (
<collectionCode id="79D726DAFFDEFFDFFB62F8CFFAEC39EF" box="[1252,1336,1901,1927]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="149" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFDEFFDFFAC9F8CFFE6139A7" collectingDate="2011-05-01" collectingDateMax="2011-05-31" collectingDateMin="2011-05-01" collectionCode="ZFMK" collectorName="C. Holzschuh" country="Laos" elevation="1600" latitude="20.2" location="Phou Pan" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" pageId="8" pageNumber="149" specimenCount="25" specimenCount-female="2" specimenCount-male="23" stateProvince="Houaphan">
<specimenCount id="09C07596FFDEFFDFFAC9F8CFFA4C39EF" box="[1359,1432,1901,1927]" count="23" pageId="8" pageNumber="149" type="male">23♂♂</specimenCount>
,
<specimenCount id="09C07596FFDEFFDFFF11F833FF1B39C3" box="[151,207,1937,1963]" count="2" pageId="8" pageNumber="149" type="female">2♀♀</specimenCount>
<collectingCountry id="67D1FE8FFFDEFFDFFF62F833FEE639C3" box="[228,306,1937,1963]" name="Laos" pageId="8" pageNumber="149">LAOS</collectingCountry>
,
<collectingRegion id="DD0270FDFFDEFFDFFEC6F833FE6539C2" box="[320,433,1936,1963]" country="Laos" name="Houaphan" pageId="8" pageNumber="149">Hua Phan</collectingRegion>
Pro.,
<collectingMunicipality id="FF1D2465FFDEFFDFFE7CF833FDAE39C2" box="[506,634,1936,1963]" pageId="8" pageNumber="149">Ban Saleui</collectingMunicipality>
,
<location id="1A19E8C4FFDEFFDFFD0EF832FD2139C3" LSID="urn:lsid:plazi:treatment:976F0F09FFD0FFDEFF11FC20FE773C99:1A19E8C4FFDEFFDFFD0EF832FD2139C3" box="[648,757,1936,1963]" country="Laos" latitude="20.2" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" name="Phou Pan" pageId="8" pageNumber="149" stateProvince="Houaphan">Phou Pan</location>
(Mt.),
<geoCoordinate id="7AF2D8D8FFDEFFDFFCCDF833FC7E39C3" box="[843,938,1937,1963]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="8" pageNumber="149" precision="925" value="20.2">
20
<superScript id="E8B31357FFDEFFDFFCE1F833FCBB39F7" attach="none" box="[871,879,1937,1951]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
12N
</geoCoordinate>
,
<geoCoordinate id="7AF2D8D8FFDEFFDFFC3CF833FBF039C3" box="[954,1060,1937,1963]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="149" precision="925" value="104.01667">
104
<superScript id="E8B31357FFDEFFDFFC62F833FC3839F7" attach="none" box="[996,1004,1937,1951]" fontSize="6" pageId="8" pageNumber="149">o</superScript>
01E
</geoCoordinate>
,
<date id="6B7898DFFFDEFFDFFBB5F833FB1B39C3" box="[1075,1231,1937,1963]" pageId="8" pageNumber="149" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">
<collectingDate id="7B3C6137FFDEFFDFFBB5F833FB1B39C3" box="[1075,1231,1937,1963]" pageId="8" pageNumber="149" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">1.v-31.v.2011</collectingDate>
</date>
,
<quantity id="D83E13FAFFDEFFDFFB58F833FAA039C3" box="[1246,1396,1937,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">
<elevation id="94EB592CFFDEFFDFFB58F833FAA039C3" box="[1246,1396,1937,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="149" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">13001900m</elevation>
</quantity>
,
<collectorName id="B233DBC9FFDEFFDFFA05F833FEDB39A6" pageId="8" pageNumber="149">C. Holzschuh</collectorName>
leg.” (
<collectionCode id="79D726DAFFDEFFDFFEDCF817FE7E39A7" box="[346,426,1973,1999]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="149" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFDFFFDEFF41FF34FD603F0D" pageId="9" pageNumber="150" type="diagnosis">
<paragraph id="1F79BE1FFFDFFFDEFF41FF34FD603F0D" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="150">
<emphasis id="2DB2620DFFDFFFDEFF41FF34FE963ED8" bold="true" box="[199,322,150,176]" pageId="9" pageNumber="150">Diagnosis.</emphasis>
Intervals 13 contain large, oval, imbricating white scale filling also interval 2 within costae 23 and mesial part of interval 2 within costa 4; interval 4 and lateral part of interval 2 within costa 4 contain short beige setae; epipleura contains beige hairs about 3x longer than setae, most of them recumbent but some erect; and elytra suture contains narrow white to yellowish-white scales only about one-half as large as those in intervals 13. Male genitalia: parameres equal length, with abruptly constricted short distal sections curving to right and have truncate tips; left tip may or may not be slightly bulbous.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFDFFFDEFF41FECCFD3A3FB9" pageId="9" pageNumber="150" type="distribution">
<paragraph id="1F79BE1FFFDFFFDEFF41FECCFD3A3FB9" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="150">
<emphasis id="2DB2620DFFDFFFDEFF41FECCFEB53FE0" bold="true" box="[199,353,366,392]" pageId="9" pageNumber="150">Distribution.</emphasis>
Known from
<collectingCountry id="67D1FE8FFFDFFFDEFD8CFECDFD943FE1" box="[522,576,367,393]" name="Laos" pageId="9" pageNumber="150">Laos</collectingCountry>
(
<bibRefCitation id="7B57C3EEFFDFFFDEFDD4FECDFC4B3FE1" author="Sabatinelli, G. &amp; Pontuale, G." box="[594,927,367,393]" pageId="9" pageNumber="150" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale 1998</bibRefCitation>
;
<bibRefCitation id="7B57C3EEFFDFFFDEFC28FECDFB613FE1" author="Zidek, J. &amp; Krajcik, M." box="[942,1205,367,393]" pageId="9" pageNumber="150" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík 2007</bibRefCitation>
;
<bibRefCitation id="7B57C3EEFFDFFFDEFB42FECDFA453FE1" author="Keith, D. &amp; Jingke, L. &amp; Xueping, Z. &amp; Lirong, H." box="[1220,1425,367,393]" pageId="9" pageNumber="150" pagination="387 - 397" refId="ref6318" refString="Keith, D., Jingke, L., Xueping, Z. &amp; Lirong, H. (2012) Scarabaeoidea Pleurosticti recoltes sur le Mt. Gaoligongshan en Republique Populaire de Chine (Insecta: Coleoptera). Vernate, 31, 387 - 397." type="journal article" year="2012">
Keith
<emphasis id="2DB2620DFFDFFFDEFA8BFECDFA963FE1" box="[1293,1346,367,393]" italics="true" pageId="9" pageNumber="150">et al</emphasis>
. 2012
</bibRefCitation>
). The species was also reported from
<collectingCountry id="67D1FE8FFFDFFFDEFDABFE31FDA73FC5" box="[557,627,403,429]" name="China" pageId="9" pageNumber="150">China</collectingCountry>
, Mt. Gaoligongshan (
<bibRefCitation id="7B57C3EEFFDFFFDEFCEEFE31FBFE3FC5" author="Keith, D. &amp; Jingke, L. &amp; Xueping, Z. &amp; Lirong, H." box="[872,1066,403,429]" pageId="9" pageNumber="150" pagination="387 - 397" refId="ref6318" refString="Keith, D., Jingke, L., Xueping, Z. &amp; Lirong, H. (2012) Scarabaeoidea Pleurosticti recoltes sur le Mt. Gaoligongshan en Republique Populaire de Chine (Insecta: Coleoptera). Vernate, 31, 387 - 397." type="journal article" year="2012">
Keith
<emphasis id="2DB2620DFFDFFFDEFC28FE31FC343FC5" box="[942,992,403,429]" italics="true" pageId="9" pageNumber="150">et al</emphasis>
. 2012
</bibRefCitation>
), but its identification need to be confirmed considering the source of material studied.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFDFFFDEFF41FE79FE773C99" pageId="9" pageNumber="150" type="discussion">
<paragraph id="1F79BE1FFFDFFFDEFF41FE79FE773C99" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="150">
<emphasis id="2DB2620DFFDFFFDEFF41FE79FEEF3F9D" bold="true" box="[199,315,475,501]" pageId="9" pageNumber="150">Remarks.</emphasis>
The
<typeStatus id="C07D00BDFFDFFFDEFEF4FE79FE063F9D" box="[370,466,475,501]" pageId="9" pageNumber="150" type="paratype">paratype</typeStatus>
of
<taxonomicName id="D8C6C59CFFDFFFDEFE73FE79FC1A3F9D" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[501,974,475,501]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFDFFFDEFE73FE79FD5D3F9D" box="[501,649,475,501]" italics="true" pageId="9" pageNumber="150">D. pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFDFFFDEFD08FE79FC1A3F9D" author="Sabatinelli, G. &amp; Pontuale, G." box="[654,974,475,501]" pageId="9" pageNumber="150" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
and the
<typeStatus id="C07D00BDFFDFFFDEFBACFE79FB5E3F9D" box="[1066,1162,475,501]" pageId="9" pageNumber="150" type="paratype">paratype</typeStatus>
of
<taxonomicName id="D8C6C59CFFDFFFDEFB2BFE79FE9B3C71" authority="Zidek &amp; Krajcik, 2007" authorityName="Zidek &amp; Krajcik" authorityYear="2007" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="bezdekorum">
<emphasis id="2DB2620DFFDFFFDEFB2BFE79FA813F9D" box="[1197,1365,475,501]" italics="true" pageId="9" pageNumber="150">D. bezdekorum</emphasis>
<bibRefCitation id="7B57C3EEFFDFFFDEFADDFE79FE9B3C71" author="Zidek, J. &amp; Krajcik, M." pageId="9" pageNumber="150" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
</taxonomicName>
examined here are almost identical in shape of aedeagus, their type localities are not distant from each other. The species of
<taxonomicName id="D8C6C59CFFDFFFDEFE38FD81FD823C55" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[446,598,547,573]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="2DB2620DFFDFFFDEFE38FD81FD823C55" box="[446,598,547,573]" italics="true" pageId="9" pageNumber="150">Dedalopterus</emphasis>
</taxonomicName>
show in shape of the parameres some considerable variation. The apparent divergence of the tip of the parameres, particularly of the left one, is a result of the flexion of the moderately sclerotized parameres along their own longitudinal axis, which is why some parameres appear to be narrower and more sharply pointed while others are wider and more convex at apex (see
<figureCitation id="87FDA29AFFDFFFDEFC62FD2DFBFC3CC1" box="[996,1064,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1356,190,1910]" captionTargetId="figure-20@7.[221,1365,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Variation in male parameres of Dedalopterus pulchellus Sabatinelli &amp; Pontuale, 1998 (AJ, all from Laos: Phu Phan). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10611477" httpUri="https://zenodo.org/record/10611477/files/figure.png" pageId="9" pageNumber="150">Fig. 5</figureCitation>
). The record of this species from
<collectingCountry id="67D1FE8FFFDFFFDEFF11FD11FF0E3CA5" box="[151,218,691,717]" name="China" pageId="9" pageNumber="150">China</collectingCountry>
is doubtful because this record is based on the material provided by Laotian dealer Jingke Li, who sometimes mislabeled his material.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,535 @@
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<mods:title id="B736D22071014BCC040BF99157F6ACE1">The genus Dedalopterus Sabatinelli &amp; Pontuale, 1998 (Scarabaeidae: Melolonthinae: Leucopholini) in Laos and Vietnam, with description of a new species</mods:title>
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<heading id="44310973FFD7FFD6FF11FB26FC193AF6" bold="true" box="[151,973,1156,1183]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
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<taxonomicName id="D8C6C59CFFD7FFD6FF11FB26FCFA3AF6" authority="Pham, Ahrens &amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[151,814,1156,1183]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="lexuanhuei" status="sp. nov.">
<emphasis id="2DB2620DFFD7FFD6FF11FB26FE1D3AF7" bold="true" box="[151,457,1156,1183]" italics="true" pageId="1" pageNumber="142">Dedalopterus lexuanhuei</emphasis>
Pham, Ahrens &amp; Sabatinelli
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,
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<paragraph id="1F79BE1FFFD7FFD6FF11FB6EFD8D3A8F" blockId="1.[151,973,1156,1255]" box="[151,601,1228,1255]" pageId="1" pageNumber="142">
(
<figureCitation id="87FDA29AFFD7FFD6FF19FB6EFF333A8F" box="[159,231,1228,1255]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="1" pageNumber="142">Figs 1</figureCitation>
;
<figureCitation id="87FDA29AFFD7FFD6FF73FB6FFEEE3A8F" box="[245,314,1229,1255]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="1" pageNumber="142">2AB</figureCitation>
;
<figureCitation id="87FDA29AFFD7FFD6FEC0FB6FFE583A8F" box="[326,396,1229,1255]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[323,1265,358,1905]" captionTargetId="figure-83@4.[310,1278,345,1905]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species (holotype,AC); D. malyszi Bunalski, 2001 (DF). A, D, parameres in left lateral view; B, E, parameres in dorsal view. C, F, parameres in right lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611473" httpUri="https://zenodo.org/record/10611473/files/figure.png" pageId="1" pageNumber="142">3AC</figureCitation>
;
<figureCitation id="87FDA29AFFD7FFD6FE1EFB6FFE3B3A8F" box="[408,495,1229,1255]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1935]" captionTargetBox="[151,1424,181,1885]" captionTargetId="figure-20@5.[151,1436,181,1887]" captionTargetPageId="5" captionText="FIGURE 4. Lateral view of elytra (AD), right lateral view of parameres tip (EI). A, E, Dedalopterus malyszi Bunalski, 2001; B, G, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, H, D. pulchellus Sabatinelli &amp; Pontuale, 1998 (paratype of D. pulchellus); D, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611475" httpUri="https://zenodo.org/record/10611475/files/figure.png" pageId="1" pageNumber="142">4D, F, I</figureCitation>
;
<figureCitation id="87FDA29AFFD7FFD6FE7AFB6EFD843A8F" box="[508,592,1228,1255]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1356,190,1910]" captionTargetId="figure-20@7.[221,1365,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Variation in male parameres of Dedalopterus pulchellus Sabatinelli &amp; Pontuale, 1998 (AJ, all from Laos: Phu Phan). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10611477" httpUri="https://zenodo.org/record/10611477/files/figure.png" pageId="1" pageNumber="142">5C, F, I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD7FFD6FF11FAB3FC073823" pageId="1" pageNumber="142" type="materials_examined">
<paragraph id="1F79BE1FFFD7FFD6FF11FAB3FEB63BB7" blockId="1.[151,1437,1296,2043]" pageId="1" pageNumber="142">
<emphasis id="2DB2620DFFD7FFD6FF11FAB3FE903B42" bold="true" box="[151,324,1296,1323]" pageId="1" pageNumber="142">Type material.</emphasis>
<materialsCitation id="AFAEB442FFD7FFD6FECCFAB3FE4B3B27" collectingDate="2004-03-31" collectingDateMax="2004-04-06" collectingDateMin="2004-03-31" collectionCode="IEBR" collectorName="Le Xuan Hue" country="Vietnam" location="Ngoc Linh Mountain" pageId="1" pageNumber="142" specimenCount="1" specimenCount-male="1" stateProvince="Kon Tum" typeStatus="holotype">
<emphasis id="2DB2620DFFD7FFD6FECCFAB3FE693B43" bold="true" box="[330,445,1297,1323]" pageId="1" pageNumber="142">
<typeStatus id="C07D00BDFFD7FFD6FECCFAB3FE6D3B43" box="[330,441,1297,1323]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
,
</emphasis>
<specimenCount id="09C07596FFD7FFD6FE45FAB3FE0C3B43" box="[451,472,1297,1323]" count="1" pageId="1" pageNumber="142" type="male"></specimenCount>
<collectingCountry id="67D1FE8FFFD7FFD6FE6EFAB3FDBB3B43" box="[488,623,1297,1323]" name="Vietnam" pageId="1" pageNumber="142">VIETNAM</collectingCountry>
,
<collectingRegion id="DD0270FDFFD7FFD6FDFFFAB3FC9A3B42" box="[633,846,1296,1323]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="142">Kon Tum Province</collectingRegion>
,
<location id="1A19E8C4FFD7FFD6FCDFFAB3FB923B42" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FCDFFAB3FB923B42" box="[857,1094,1296,1323]" country="Vietnam" name="Ngoc Linh Mountain" pageId="1" pageNumber="142" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
,
<date id="6B7898DFFFD7FFD6FBD6FAB2FAD63B42" box="[1104,1282,1296,1322]" pageId="1" pageNumber="142" value="2004-03-31" valueMax="2004-04-06" valueMin="2004-03-31">
<collectingDate id="7B3C6137FFD7FFD6FBD6FAB2FAD63B42" box="[1104,1282,1296,1322]" pageId="1" pageNumber="142" value="2004-03-31" valueMax="2004-04-06" valueMin="2004-03-31">31.iii6.iv.2004</collectingDate>
</date>
,
<collectingMethod id="C687C608FFD7FFD6FA8BFAB2FAA03B43" box="[1293,1396,1296,1323]" pageId="1" pageNumber="142">light trap</collectingMethod>
,
<collectorName id="B233DBC9FFD7FFD6FAF9FAB3FEDD3B27" pageId="1" pageNumber="142">Le Xuan Hue</collectorName>
leg.” (
<collectionCode id="79D726DAFFD7FFD6FED4FA97FE403B27" box="[338,404,1333,1359]" pageId="1" pageNumber="142">IEBR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="AFAEB442FFD7FFD6FE2CFA97FE013B1B" collectingDate="2022-04" collectingDateMax="2022-05-31" collectingDateMin="2022-04" collectionCode="IEBR" country="Vietnam" location="Ngoc Linh Mountain" pageId="1" pageNumber="142" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Kon Tum" typeStatus="paratype">
<emphasis id="2DB2620DFFD7FFD6FE2CFA97FDFD3B27" bold="true" box="[426,553,1333,1359]" pageId="1" pageNumber="142">
<typeStatus id="C07D00BDFFD7FFD6FE2CFA97FDF73B27" box="[426,547,1333,1359]" pageId="1" pageNumber="142" type="paratype">Paratypes</typeStatus>
,
</emphasis>
<specimenCount id="09C07596FFD7FFD6FDA9FA97FDBE3B27" box="[559,618,1333,1359]" count="2" pageId="1" pageNumber="142" type="male">2♂♂</specimenCount>
,
<specimenCount id="09C07596FFD7FFD6FDF3FA97FD793B27" box="[629,685,1333,1359]" count="2" pageId="1" pageNumber="142" type="female">2♀♀</specimenCount>
<collectingCountry id="67D1FE8FFFD7FFD6FD3BFA97FC903B27" box="[701,836,1333,1359]" name="Vietnam" pageId="1" pageNumber="142">VIETNAM</collectingCountry>
,
<collectingRegion id="DD0270FDFFD7FFD6FCCBFA97FBF03B26" box="[845,1060,1332,1359]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="142">Kon Tum Province</collectingRegion>
,
<location id="1A19E8C4FFD7FFD6FBA8FA97FAC83B26" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FBA8FA97FAC83B26" box="[1070,1308,1332,1359]" country="Vietnam" name="Ngoc Linh Mountain" pageId="1" pageNumber="142" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
,
<date id="6B7898DFFFD7FFD6FAA0FA96FA4C3B26" box="[1318,1432,1332,1358]" pageId="1" pageNumber="142" value="2022-04" valueMax="2022-05-31" valueMin="2022-04-00">
<collectingDate id="7B3C6137FFD7FFD6FAA0FA96FA4C3B26" box="[1318,1432,1332,1358]" pageId="1" pageNumber="142" value="2022-04" valueMax="2022-05-31" valueMin="2022-04-00">ivv.2022</collectingDate>
</date>
, local collector leg.” (
<collectionCode id="79D726DAFFD7FFD6FE01FAFBFE193B1B" box="[391,461,1369,1395]" pageId="1" pageNumber="142">IEBR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFD7FFD6FE64FAFBFEE13BFF" collectingDate="2020-05" collectionCode="ZMPC" country="Vietnam" elevation="1500" location="Ngoc Linh Mountain" pageId="1" pageNumber="142" specimenCount="4" specimenCount-male="4" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="09C07596FFD7FFD6FE64FAFBFDCE3B1B" box="[482,538,1369,1395]" count="4" pageId="1" pageNumber="142" type="male">4♂♂</specimenCount>
<collectingRegion id="DD0270FDFFD7FFD6FDABFAFBFCDE3B1A" box="[557,778,1368,1395]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="142">Kon Tum Province</collectingRegion>
,
<location id="1A19E8C4FFD7FFD6FC9EFAFBFBDE3B1A" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FC9EFAFBFBDE3B1A" box="[792,1034,1368,1395]" country="Vietnam" name="Ngoc Linh Mountain" pageId="1" pageNumber="142" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
, alt.
<quantity id="D83E13FAFFD7FFD6FBC5FAFAFB473B1B" box="[1091,1171,1368,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="142" unit="m" value="1500.0">
<elevation id="94EB592CFFD7FFD6FBC5FAFAFB473B1B" box="[1091,1171,1368,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="142" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="6B7898DFFFD7FFD6FB26FAFBFB3A3B1B" box="[1184,1262,1369,1395]" pageId="1" pageNumber="142" value="2020-05">
<collectingDate id="7B3C6137FFD7FFD6FB26FAFBFB3A3B1B" box="[1184,1262,1369,1395]" pageId="1" pageNumber="142" value="2020-05">v.2020</collectingDate>
</date>
, local collector leg.” (
<collectionCode id="79D726DAFFD7FFD6FF5DFADFFEFA3BFF" box="[219,302,1405,1431]" pageId="1" pageNumber="142">ZMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFD7FFD6FEB9FADFFF263BD3" collectingDate="2022-05" collectionCode="MHNG, ZFMK" country="Vietnam" location="Ngoc Linh Nat." municipality="Dak Glei Distr." pageId="1" pageNumber="142" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="09C07596FFD7FFD6FEB9FADFFEAE3BFF" box="[319,378,1405,1431]" count="4" pageId="1" pageNumber="142" type="male">4♂♂</specimenCount>
,
<specimenCount id="09C07596FFD7FFD6FE02FADEFE693BFF" box="[388,445,1404,1431]" count="5" pageId="1" pageNumber="142" type="female">5♀♀</specimenCount>
<collectingRegion id="DD0270FDFFD7FFD6FE4BFADFFDA83BFF" box="[461,636,1405,1431]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="142">Kon Tum Prov.</collectingRegion>
,
<collectingMunicipality id="FF1D2465FFD7FFD6FD00FADEFCE53BFE" box="[646,817,1404,1430]" pageId="1" pageNumber="142">Dak Glei Distr.</collectingMunicipality>
,
<location id="1A19E8C4FFD7FFD6FCBDFADFFC3C3BFF" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FCBDFADFFC3C3BFF" box="[827,1000,1404,1431]" country="Vietnam" municipality="Dak Glei Distr." name="Ngoc Linh Nat." pageId="1" pageNumber="142" stateProvince="Kon Tum">Ngoc Linh Nat.</location>
res.,
<date id="6B7898DFFFD7FFD6FBA4FADFFBA43BFF" box="[1058,1136,1405,1431]" pageId="1" pageNumber="142" value="2022-05">
<collectingDate id="7B3C6137FFD7FFD6FBA4FADFFBA43BFF" box="[1058,1136,1405,1431]" pageId="1" pageNumber="142" value="2022-05">v.2022</collectingDate>
</date>
, local collectors” (
<collectionCode id="79D726DAFFD7FFD6FAB8FADFFA4C3BFF" box="[1342,1432,1405,1431]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="142" type="Museum">MHNG</collectionCode>
,
<collectionCode id="79D726DAFFD7FFD6FF11FA03FF3E3BD3" box="[151,234,1441,1467]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="142" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFAEB442FFD7FFD6FF78FA02FE893BB7" collectingDate="2022-06" collectionCode="MHNG, ZFMK" country="Vietnam" county="Tai Giang" elevation="1300" location="Axan" municipality="Tay Giang" pageId="1" pageNumber="142" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="09C07596FFD7FFD6FF78FA02FEEC3BD2" box="[254,312,1440,1466]" count="6" pageId="1" pageNumber="142" type="male">6♂♂</specimenCount>
,
<specimenCount id="09C07596FFD7FFD6FEC2FA03FEA83BD3" box="[324,380,1441,1467]" count="2" pageId="1" pageNumber="142" type="female">2♀♀</specimenCount>
<collectingRegion id="DD0270FDFFD7FFD6FE0BFA03FD883BD3" box="[397,604,1441,1467]" country="Vietnam" name="Quang Nam" pageId="1" pageNumber="142">Quang Nam prov.</collectingRegion>
,
<collectingCounty id="F618C693FFD7FFD6FDE0FA02FD033BD2" box="[614,727,1440,1466]" pageId="1" pageNumber="142">Tâi Giang</collectingCounty>
[
<collectingMunicipality id="FF1D2465FFD7FFD6FD63FA03FCB63BD2" box="[741,866,1440,1467]" pageId="1" pageNumber="142">Tây Giang</collectingMunicipality>
] distr.,
<location id="1A19E8C4FFD7FFD6FC33FA03FC203BD3" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FC33FA03FC203BD3" box="[949,1012,1441,1467]" country="Vietnam" county="Tai Giang" municipality="Tay Giang" name="Axan" pageId="1" pageNumber="142" stateProvince="Quang Nam">Axan</location>
,
<quantity id="D83E13FAFFD7FFD6FB86FA03FB843BD3" box="[1024,1104,1441,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="142" unit="m" value="1300.0">
<elevation id="94EB592CFFD7FFD6FB86FA03FB843BD3" box="[1024,1104,1441,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="142" unit="m" value="1300.0">1300m</elevation>
</quantity>
,
<date id="6B7898DFFFD7FFD6FBDAFA02FB673BD2" box="[1116,1203,1440,1466]" pageId="1" pageNumber="142" value="2022-06">
<collectingDate id="7B3C6137FFD7FFD6FBDAFA02FB673BD2" box="[1116,1203,1440,1466]" pageId="1" pageNumber="142" value="2022-06">vi.2022</collectingDate>
</date>
, local collector leg.” (
<collectionCode id="79D726DAFFD7FFD6FF19FA67FF2D3BB7" box="[159,249,1477,1503]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="142" type="Museum">MHNG</collectionCode>
,
<collectionCode id="79D726DAFFD7FFD6FE82FA67FE863BB7" box="[260,338,1477,1503]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="142" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="1F79BE1FFFD7FFD6FF41FA4AFC073823" blockId="1.[151,1437,1296,2043]" pageId="1" pageNumber="142">
<emphasis id="2DB2620DFFD7FFD6FF41FA4AFDC2386B" bold="true" box="[199,534,1512,1539]" pageId="1" pageNumber="142">Specimen excluded from the</emphasis>
<materialsCitation id="AFAEB442FFD7FFD6FD9BFA4BFF233822" collectingDate="2022-05" collectionCode="MHNG" country="Vietnam" elevation="1700" location="Mu Cang Chai" pageId="1" pageNumber="142" specimenCount="1" specimenCount-male="1" stateProvince="Yen Bai">
<emphasis id="2DB2620DFFD7FFD6FD9BFA4BFD74386A" bold="true" box="[541,672,1512,1539]" pageId="1" pageNumber="142">
<typeStatus id="C07D00BDFFD7FFD6FD9BFA4BFD84386B" box="[541,592,1513,1539]" pageId="1" pageNumber="142">type</typeStatus>
series.
</emphasis>
<specimenCount id="09C07596FFD7FFD6FD20FA4BFD1D386B" box="[678,713,1513,1539]" count="1" pageId="1" pageNumber="142" type="male">1♂</specimenCount>
“N.
<collectingCountry id="67D1FE8FFFD7FFD6FD7BFA4AFC8B386A" box="[765,863,1512,1538]" name="Vietnam" pageId="1" pageNumber="142">Vietnam</collectingCountry>
,
<collectingRegion id="DD0270FDFFD7FFD6FCEEFA4BFC15386A" box="[872,961,1512,1539]" country="Vietnam" name="Yen Bai" pageId="1" pageNumber="142">Yen Bai</collectingRegion>
Pro.,
<location id="1A19E8C4FFD7FFD6FB84FA4BFB7C386A" LSID="urn:lsid:plazi:treatment:976F0F09FFD7FFD1FF11FB26FAFD3D51:1A19E8C4FFD7FFD6FB84FA4BFB7C386A" box="[1026,1192,1512,1539]" country="Vietnam" name="Mu Cang Chai" pageId="1" pageNumber="142" stateProvince="Yen Bai">Mu Cang Chai</location>
,
<quantity id="D83E13FAFFD7FFD6FB34FA4BFAD7386B" box="[1202,1283,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="142" unit="m" value="1700.0">
<elevation id="94EB592CFFD7FFD6FB34FA4BFAD7386B" box="[1202,1283,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="142" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="6B7898DFFFD7FFD6FA8BFA4BFA8F386B" box="[1293,1371,1513,1539]" pageId="1" pageNumber="142" value="2022-05">
<collectingDate id="7B3C6137FFD7FFD6FA8BFA4BFA8F386B" box="[1293,1371,1513,1539]" pageId="1" pageNumber="142" value="2022-05">v.2022</collectingDate>
</date>
, local collectors” (
<collectionCode id="79D726DAFFD7FFD6FEA6F9AFFEA8384F" box="[288,380,1549,1575]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="142" type="Museum">MHNG</collectionCode>
). This specimen is likely erroneously labelled since the collecting locality is extremely distant from the
</materialsCitation>
<typeStatus id="C07D00BDFFD7FFD6FF78F993FEFA3823" box="[254,302,1585,1611]" pageId="1" pageNumber="142">type</typeStatus>
locality but close to the range of
<taxonomicName id="D8C6C59CFFD7FFD6FD23F993FC1B3823" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[677,975,1584,1611]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="malyzsi">
<emphasis id="2DB2620DFFD7FFD6FD23F993FCCF3822" box="[677,795,1584,1610]" italics="true" pageId="1" pageNumber="142">D. malyzsi</emphasis>
<bibRefCitation id="7B57C3EEFFD7FFD6FCA4F992FC1B3823" author="Bunalski, M." box="[802,975,1584,1611]" pageId="1" pageNumber="142" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD7FFD1FF41F9F6FBBB3F0D" lastPageId="6" lastPageNumber="147" pageId="1" pageNumber="142" type="description">
<paragraph id="1F79BE1FFFD7FFD6FF41F9F6FAAC38FB" blockId="1.[151,1437,1296,2043]" pageId="1" pageNumber="142">
<emphasis id="2DB2620DFFD7FFD6FF41F9F6FDE53806" bold="true" box="[199,561,1620,1647]" pageId="1" pageNumber="142">
Description of
<typeStatus id="C07D00BDFFD7FFD6FEF5F9F7FE0E3807" box="[371,474,1621,1647]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
(male).
</emphasis>
Body length:
<quantity id="D83E13FAFFD7FFD6FD4CF9F6FCFB3807" box="[714,815,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="142" unit="mm" value="20.5">20.5 mm</quantity>
; body width: 9.0 mm, dorsal integument and appendages yellowish-brown to dark brown, canthus dark brown and abdominal ventrites light brown to chestnut (
<figureCitation id="87FDA29AFFD7FFD6FA97F9DAFAB838FB" box="[1297,1388,1656,1683]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="1" pageNumber="142">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="1F79BE1FFFD7FFD6FF41F93FFD5239E7" blockId="1.[151,1437,1296,2043]" pageId="1" pageNumber="142">
<emphasis id="2DB2620DFFD7FFD6FF41F93FFEDA38DF" bold="true" box="[199,270,1693,1719]" pageId="1" pageNumber="142">Head.</emphasis>
Antennal club 2x longer than stalk, scape 1.43x longer than wide and distally bulbous, pedicel and segments 35 equidimensional and beadlike, and segments 67 wider than long and anteriorly pronounced into sharp teeth, tooth of segment 7 longer (
<figureCitation id="87FDA29AFFD7FFD6FDDDF946FD603897" box="[603,692,1764,1791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="1" pageNumber="142">Fig. 1H</figureCitation>
). Terminal maxillary palpomere nearly 2x longer and 1.4x wider than penultimate palpomere, both margins equally inflated and apex moderately sharp, and dorsally a callus covers proximal two thirds of surface (
<figureCitation id="87FDA29AFFD7FFD6FD8DF88EFDB7392F" box="[523,611,1836,1863]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="1" pageNumber="142">Fig. 1F</figureCitation>
). Labrum glabrous except for sparse longer setae along its lower front margin. Frontoclypeal suture complete, slightly concave-forward; deep frontal depression; vertex flat, not carinate; eyes large but not very prominent (
<figureCitation id="87FDA29AFFD7FFD6FD99F8D6FDAC39E7" box="[543,632,1908,1935]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="1" pageNumber="142">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="1F79BE1FFFD7FFD6FF41F83BFED23993" blockId="1.[151,1437,1296,2043]" pageId="1" pageNumber="142">
<emphasis id="2DB2620DFFD7FFD6FF41F83BFE9339DB" bold="true" box="[199,327,1945,1971]" pageId="1" pageNumber="142">Pronotum.</emphasis>
Pronotal vestiture consists of round white scales composing sagittal vitta, on longitudinal depression; disk covered by narrow yellowish-white scales; and short beige setae and longer beige hairs along lateral margins (
<figureCitation id="87FDA29AFFD7FFD6FF19F842FF2D3993" box="[159,249,2016,2043]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="1" pageNumber="142">Fig. 1D</figureCitation>
).
</paragraph>
<caption id="4BB9EE97FFD4FFD5FF11F83AFCA13990" ID-DOI="http://doi.org/10.5281/zenodo.10611469" ID-Zenodo-Dep="10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="2" pageNumber="143" startId="2.[151,250,1944,1968]" targetBox="[172,1416,179,1918]" targetPageId="2" targetType="figure">
<paragraph id="1F79BE1FFFD4FFD5FF11F83AFCA13990" blockId="2.[151,1437,1944,2040]" pageId="2" pageNumber="143">
<emphasis id="2DB2620DFFD4FFD5FF11F83AFEC339D8" bold="true" box="[151,279,1944,1968]" pageId="2" pageNumber="143">FIGURE 1.</emphasis>
<taxonomicName id="D8C6C59CFFD4FFD5FEA6F83AFCB239D8" authority="Pham, Ahrens and Sabatinelli" authorityName="Pham, Ahrens and Sabatinelli" authorityYear="2024" box="[288,870,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="lexuanhuei" status="sp. nov.">
<emphasis id="2DB2620DFFD4FFD5FEA6F83AFDF739D8" box="[288,547,1944,1968]" italics="true" pageId="2" pageNumber="143">Dedalopterus lexuanhuei</emphasis>
Pham, Ahrens and Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD4FFD5FCF5F83AFC2739D8" bold="true" box="[883,1011,1944,1968]" pageId="2" pageNumber="143">
<taxonomicNameLabel id="3681DF76FFD4FFD5FCF5F83AFC2739D8" box="[883,1011,1944,1968]" pageId="2" pageNumber="143" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Male (holotype, BF, H, I) and female (paratype, A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm.
</paragraph>
</caption>
<caption id="4BB9EE97FFD5FFD4FF11FA06FB943B88" ID-DOI="http://doi.org/10.5281/zenodo.10611471" ID-Zenodo-Dep="10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="3" pageNumber="144" startId="3.[151,250,1444,1468]" targetBox="[370,1260,238,1348]" targetPageId="3" targetType="figure">
<paragraph id="1F79BE1FFFD5FFD4FF11FA06FB943B88" blockId="3.[151,1436,1444,1505]" pageId="3" pageNumber="144">
<emphasis id="2DB2620DFFD5FFD4FF11FA06FEC23BD4" bold="true" box="[151,278,1444,1468]" pageId="3" pageNumber="144">FIGURE 2.</emphasis>
Habitus in dorsal view. AB,
<taxonomicName id="D8C6C59CFFD5FFD4FDDEFA07FBFC3BD4" authority="Pham, Ahrens and Sabatinelli" authorityName="Pham, Ahrens and Sabatinelli" authorityYear="2024" box="[600,1064,1444,1468]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFD5FFD4FDDEFA07FD3C3BD4" box="[600,744,1444,1468]" italics="true" pageId="3" pageNumber="144">D. lexuanhuei</emphasis>
Pham, Ahrens and Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD5FFD4FBB3FA06FB613BD4" bold="true" box="[1077,1205,1444,1468]" pageId="3" pageNumber="144">
<taxonomicNameLabel id="3681DF76FFD5FFD4FBB3FA06FB613BD4" box="[1077,1205,1444,1468]" pageId="3" pageNumber="144" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A, ♂ holotype; B, ♀ paratype; CD,
<taxonomicName id="D8C6C59CFFD5FFD4FEBEFA6BFD973B88" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[312,579,1480,1504]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD5FFD4FEBEFA6BFE773B88" box="[312,419,1480,1504]" italics="true" pageId="3" pageNumber="144">D. malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFD5FFD4FE2CFA6AFD973B88" author="Bunalski, M." box="[426,579,1480,1504]" pageId="3" pageNumber="144" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
(Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm.
</paragraph>
</caption>
<paragraph id="1F79BE1FFFD5FFD4FF41F990FD5A38D0" blockId="3.[151,1437,1586,2008]" pageId="3" pageNumber="144">
<emphasis id="2DB2620DFFD5FFD4FF41F990FECE3824" bold="true" box="[199,282,1586,1612]" pageId="3" pageNumber="144">Elytra.</emphasis>
Elytral vestiture with large, oval, imbricating white scales in intervals 13; narrow yellowish-white scales in secondary intervals within costae 23; narrow off-white to beige scales on sutural edge and in mesial part of secondary interval within costa 4; beige spines in costa; and beige setae mixed some narrow yellowish-white scales in (marginal) interval 5 (
<figureCitation id="87FDA29AFFD5FFD4FE72F93CFD9B38D0" box="[500,591,1694,1720]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="3" pageNumber="144">Figs 1C</figureCitation>
,
<figureCitation id="87FDA29AFFD5FFD4FDDDF93CFD5538D0" box="[603,641,1694,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="3" pageNumber="144">2A</figureCitation>
).
</paragraph>
<paragraph id="1F79BE1FFFD5FFD4FF41F960FB303968" blockId="3.[151,1437,1586,2008]" pageId="3" pageNumber="144">
<emphasis id="2DB2620DFFD5FFD4FF41F960FE9638B4" bold="true" box="[199,322,1730,1756]" pageId="3" pageNumber="144">Pygidium.</emphasis>
Strongly convex at basal half, disk with off-white spines, off-white scales at base, off-white setae at lateral margins and apex; pygidial apex constricted to produce a slender and sharp spine (
<figureCitation id="87FDA29AFFD5FFD4FBF9F944FB023968" box="[1151,1238,1766,1792]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="3" pageNumber="144">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="1F79BE1FFFD5FFD4FF41F8A8FCB93921" blockId="3.[151,1437,1586,2008]" pageId="3" pageNumber="144">
<emphasis id="2DB2620DFFD5FFD4FF41F8A8FE97394C" bold="true" box="[199,323,1802,1828]" pageId="3" pageNumber="144">Abdomen.</emphasis>
Ventrites 15 bear white scales and spines, posterior margin of ventrites 56 with narrow white scales; posterior margin of ventrite 6 with yellowish-white setae.
</paragraph>
<paragraph id="1F79BE1FFFD5FFD4FF41F8F0FD8F39B0" blockId="3.[151,1437,1586,2008]" pageId="3" pageNumber="144">
<emphasis id="2DB2620DFFD5FFD4FF41F8F0FED23904" bold="true" box="[199,262,1874,1900]" pageId="3" pageNumber="144">Legs.</emphasis>
Protibia tridentate, with basal tooth obsolete and situated more distantly from distal two teeth (
<figureCitation id="87FDA29AFFD5FFD4FAE9F8F0FF6C39F8" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="3" pageNumber="144">Fig. 1E</figureCitation>
). Basal protarsomere longer than protarsomere 2, basal mesotarsomere as long as mesotarsomere 2, and basal metatarsomere subequal metatarsomere 2. Ventral tooth of all tarsal claws situated nearly at half-length of claw, bases inclined slightly forward (
<figureCitation id="87FDA29AFFD5FFD4FE79F81CFD9F39B0" box="[511,587,1982,2008]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="3" pageNumber="144">Fig. 1I</figureCitation>
).
</paragraph>
<paragraph id="1F79BE1FFFD2FFD3FF41FF34FD373F75" blockId="4.[151,1436,150,285]" pageId="4" pageNumber="145">
<emphasis id="2DB2620DFFD2FFD3FF41FF34FE953ED8" bold="true" box="[199,321,150,176]" pageId="4" pageNumber="145">Aedeagus.</emphasis>
Parameres asymmetric, moderately long and slender, distal parts curved to the right; right paramere shorter and thinner than left one. In lateral views, parameres slightly curved and left termination ventrally expanded into hatchet, right termination gradually tapers to blunt upper point. Ventral setation moderately long and dense covered distal half of ventral paramere (
<figureCitation id="87FDA29AFFD2FFD3FDD1FEA1FD033F75" box="[599,727,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[323,1265,358,1905]" captionTargetId="figure-83@4.[310,1278,345,1905]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species (holotype,AC); D. malyszi Bunalski, 2001 (DF). A, D, parameres in left lateral view; B, E, parameres in dorsal view. C, F, parameres in right lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611473" httpUri="https://zenodo.org/record/10611473/files/figure.png" pageId="4" pageNumber="145">Figs 3AC</figureCitation>
).
</paragraph>
<caption id="4BB9EE97FFD2FFD3FF11F82AFE833980" ID-DOI="http://doi.org/10.5281/zenodo.10611473" ID-Zenodo-Dep="10611473" httpUri="https://zenodo.org/record/10611473/files/figure.png" pageId="4" pageNumber="145" startId="4.[151,250,1928,1952]" targetBox="[323,1265,358,1905]" targetPageId="4" targetType="figure">
<paragraph id="1F79BE1FFFD2FFD3FF11F82AFE833980" blockId="4.[151,1437,1928,2024]" pageId="4" pageNumber="145">
<emphasis id="2DB2620DFFD2FFD3FF11F82AFEC639C8" bold="true" box="[151,274,1928,1952]" pageId="4" pageNumber="145">FIGURE 3.</emphasis>
Male genitalia of
<taxonomicName id="D8C6C59CFFD2FFD3FE4EF82AFC2C39C8" authority="Pham, Ahrens and Sabatinelli" authorityName="Pham, Ahrens and Sabatinelli" authorityYear="2024" box="[456,1016,1928,1952]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="lexuanhuei" status="sp. nov.">
<emphasis id="2DB2620DFFD2FFD3FE4EF82AFD1339C8" box="[456,711,1928,1952]" italics="true" pageId="4" pageNumber="145">Dedalopterus lexuanhuei</emphasis>
Pham, Ahrens and Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD2FFD3FB87F82AFBA939C8" bold="true" box="[1025,1149,1928,1952]" pageId="4" pageNumber="145">
<taxonomicNameLabel id="3681DF76FFD2FFD3FB87F82AFBA939C8" box="[1025,1149,1928,1952]" pageId="4" pageNumber="145" rank="species">new species</taxonomicNameLabel>
</emphasis>
(holotype, AC);
<taxonomicName id="D8C6C59CFFD2FFD3FAB4F82BFEE439AC" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD2FFD3FAB4F82BFA4839C8" box="[1330,1436,1928,1952]" italics="true" pageId="4" pageNumber="145">D. malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFD2FFD3FF11F80EFEE439AC" author="Bunalski, M." box="[151,304,1964,1988]" pageId="4" pageNumber="145" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
(DF). A, D, parameres in left lateral view; B, E, parameres in dorsal view. C, F, parameres in right lateral view. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="4BB9EE97FFD3FFD2FF11F8D5FB1539BF" ID-DOI="http://doi.org/10.5281/zenodo.10611475" ID-Zenodo-Dep="10611475" httpUri="https://zenodo.org/record/10611475/files/figure.png" pageId="5" pageNumber="146" startId="5.[151,250,1911,1935]" targetBox="[151,1424,181,1885]" targetPageId="5" targetType="figure">
<paragraph id="1F79BE1FFFD3FFD2FF11F8D5FB1539BF" blockId="5.[151,1437,1911,2007]" pageId="5" pageNumber="146">
<emphasis id="2DB2620DFFD3FFD2FF11F8D5FEC639E7" bold="true" box="[151,274,1911,1935]" pageId="5" pageNumber="146">FIGURE 4.</emphasis>
Lateral view of elytra (AD), right lateral view of parameres tip (EI). A, E,
<taxonomicName id="D8C6C59CFFD3FFD2FB9DF8D5FA4D39E7" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[1051,1433,1911,1935]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD3FFD2FB9DF8D5FB2339E7" box="[1051,1271,1911,1935]" italics="true" pageId="5" pageNumber="146">Dedalopterus malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFD3FFD2FB7AF8D5FA4D39E7" author="Bunalski, M." box="[1276,1433,1911,1935]" pageId="5" pageNumber="146" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
; B, G,
<taxonomicName id="D8C6C59CFFD3FFD2FF5FF83EFEB639DB" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[217,354,1947,1971]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD3FFD2FF5FF83EFEB639DB" box="[217,354,1947,1971]" italics="true" pageId="5" pageNumber="146">D. pulchellus</emphasis>
</taxonomicName>
(paratype of
<taxonomicName id="D8C6C59CFFD3FFD2FE77F83EFC5C39DB" authority="Zidek &amp; Krajcik, 2007" authorityName="Zidek &amp; Krajcik" authorityYear="2007" box="[497,904,1947,1971]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="bezdekorum">
<emphasis id="2DB2620DFFD3FFD2FE77F83EFD5939DB" box="[497,653,1947,1971]" italics="true" pageId="5" pageNumber="146">D. bezdekorum</emphasis>
<bibRefCitation id="7B57C3EEFFD3FFD2FD10F839FC5C39DB" author="Zidek, J. &amp; Krajcik, M." box="[662,904,1947,1971]" pageId="5" pageNumber="146" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
</taxonomicName>
); C, H,
<taxonomicName id="D8C6C59CFFD3FFD2FC58F83EFA4939DB" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[990,1437,1947,1971]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD3FFD2FC58F83EFBB339DB" box="[990,1127,1947,1971]" italics="true" pageId="5" pageNumber="146">D. pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFD3FFD2FBF6F839FA4939DB" author="Sabatinelli, G. &amp; Pontuale, G." box="[1136,1437,1947,1971]" pageId="5" pageNumber="146" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
(paratype of
<taxonomicName id="D8C6C59CFFD3FFD2FE9FF862FE7439BF" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[281,416,1983,2007]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD3FFD2FE9FF862FE7439BF" box="[281,416,1983,2007]" italics="true" pageId="5" pageNumber="146">D. pulchellus</emphasis>
</taxonomicName>
); D, F, I,
<taxonomicName id="D8C6C59CFFD3FFD2FD84F862FC1139BF" authority="Pham, Ahrens and Sabatinelli" authorityName="Pham, Ahrens and Sabatinelli" authorityYear="2024" box="[514,965,1983,2007]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFD3FFD2FD84F862FD4439BF" box="[514,656,1983,2007]" italics="true" pageId="5" pageNumber="146">D. lexuanhuei</emphasis>
Pham, Ahrens and Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD3FFD2FC49F81DFB9839BF" bold="true" box="[975,1100,1983,2007]" pageId="5" pageNumber="146">
<taxonomicNameLabel id="3681DF76FFD3FFD2FC49F81DFB9839BF" box="[975,1100,1983,2007]" pageId="5" pageNumber="146" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype).
</paragraph>
</caption>
<paragraph id="1F79BE1FFFD0FFD1FF41FF34FBBB3F0D" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FF34FE993ED8" bold="true" box="[199,333,150,176]" pageId="6" pageNumber="147">Variability.</emphasis>
Male
<typeStatus id="C07D00BDFFD0FFD1FE15FF35FDD73ED9" box="[403,515,151,177]" pageId="6" pageNumber="147" type="paratype">paratypes</typeStatus>
: Body length:
<quantity id="D83E13FAFFD0FFD1FD22FF34FC9D3ED9" box="[676,841,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="6" pageNumber="147" unit="mm" value="19.5" valueMax="20.5" valueMin="18.5">18.520.5 mm</quantity>
; body width:
<quantity id="D83E13FAFFD0FFD1FC66FF35FBBD3ED9" box="[992,1129,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="8.8" metricValueMin="8.3" pageId="6" pageNumber="147" unit="mm" value="8.55" valueMax="8.8" valueMin="8.3">8.38.8 mm</quantity>
; a male
<typeStatus id="C07D00BDFFD0FFD1FB42FF35FAF03ED9" box="[1220,1316,151,177]" pageId="6" pageNumber="147" type="paratype">paratype</typeStatus>
with basal tooth of protibia almost absent. Other features are identical to the
<typeStatus id="C07D00BDFFD0FFD1FC16FF19FC223EBD" box="[912,1014,187,213]" pageId="6" pageNumber="147" type="holotype">holotype</typeStatus>
. Female
<typeStatus id="C07D00BDFFD0FFD1FBD8FF19FB1D3EBD" box="[1118,1225,187,213]" pageId="6" pageNumber="147" type="paratype">paratypes</typeStatus>
(
<figureCitation id="87FDA29AFFD0FFD1FB5CFF19FAE33EBD" box="[1242,1335,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[172,1416,179,1918]" captionTargetId="figure-27@2.[172,1416,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species. Male (holotype, BF, H, I) and female (paratype,A, G).AB, pygidium; C, elytra in lateral view; D, G, head and pronotum; E, protibia; F, terminal maxillary palpomere; H, antenna; I, metatarsomeres. Scale bar: 2.0 mm, except for F=1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611469" httpUri="https://zenodo.org/record/10611469/files/figure.png" pageId="6" pageNumber="147">Figs 1B</figureCitation>
,
<figureCitation id="87FDA29AFFD0FFD1FAC0FF19FA5A3EBD" box="[1350,1422,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="6" pageNumber="147">2A, C</figureCitation>
): Body usually bigger than males, body length:
<quantity id="D83E13FAFFD0FFD1FD2BFF7CFC813E91" box="[685,853,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.135" metricValueMax="2.2199999999999998" metricValueMin="2.05" pageId="6" pageNumber="147" unit="mm" value="21.35" valueMax="22.2" valueMin="20.5">20.522.2 mm</quantity>
, body width:
<quantity id="D83E13FAFFD0FFD1FC70FF7DFB4A3E91" box="[1014,1182,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" metricValueMax="1.05" metricValueMin="1.03" pageId="6" pageNumber="147" unit="mm" value="10.4" valueMax="10.5" valueMin="10.3">
10.3
<emphasis id="2DB2620DFFD0FFD1FBA1FF7DFBE13E91" bold="true" box="[1063,1077,223,249]" pageId="6" pageNumber="147"></emphasis>
10.5 mm
</quantity>
; antennal club shorter or subequal to stalk; pygidium slightly convex, rounded apex; (marginal) interval 5 with large, oval, imbricating yellowish-white and white scales; antennomeres 35 slightly longer than wide, antennomeres 67 anteriorly less pronounced and blunt; callus correspondingly narrower; protibia broader and tridentate.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41FECCFBFC3CED" pageId="6" pageNumber="147" type="diagnosis">
<paragraph id="1F79BE1FFFD0FFD1FF41FECCFBFC3CED" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FECCFE1B3FE0" bold="true" box="[199,463,366,392]" pageId="6" pageNumber="147">Differential diagnosis.</emphasis>
<taxonomicName id="D8C6C59CFFD0FFD1FE51FECDFBE23FE1" authority="Pham, Ahrens &amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[471,1078,367,393]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="lexuanhuei" status="sp. nov.">
<emphasis id="2DB2620DFFD0FFD1FE51FECDFD263FE1" box="[471,754,367,393]" italics="true" pageId="6" pageNumber="147">Dedalopterus lexuanhuei</emphasis>
Pham, Ahrens &amp; Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD0FFD1FBC3FECDFB1B3FE0" bold="true" box="[1093,1231,366,393]" pageId="6" pageNumber="147">
<taxonomicNameLabel id="3681DF76FFD0FFD1FBC3FECDFB1B3FE0" box="[1093,1231,366,393]" pageId="6" pageNumber="147" rank="species">new species</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="D8C6C59CFFD0FFD1FF11FE31FEF83FC5" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[151,300,403,429]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FF11FE31FEF83FC5" box="[151,300,403,429]" italics="true" pageId="6" pageNumber="147">D. pulchellus</emphasis>
</taxonomicName>
and
<taxonomicName id="D8C6C59CFFD0FFD1FEE3FE31FE0F3FC5" authorityName="Bunalski" authorityYear="2001" box="[357,475,403,429]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD0FFD1FEE3FE31FE0F3FC5" box="[357,475,403,429]" italics="true" pageId="6" pageNumber="147">D. malyszi</emphasis>
</taxonomicName>
. They can be distinguished from the each other the apical segment of maxillary palp with a large callus on its outer surface and frontal depression deep in
<taxonomicName id="D8C6C59CFFD0FFD1FC34FE15FA4C3FB9" authority="Pham, Ahrens &amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[946,1432,439,465]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFD0FFD1FC34FE15FB843FB9" box="[946,1104,439,465]" italics="true" pageId="6" pageNumber="147">D. lexuanhuei</emphasis>
Pham, Ahrens &amp; Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD0FFD1FF11FE79FEF73F9C" bold="true" box="[151,291,474,501]" pageId="6" pageNumber="147">
<taxonomicNameLabel id="3681DF76FFD0FFD1FF11FE79FEF73F9C" box="[151,291,474,501]" pageId="6" pageNumber="147" rank="species">new species</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="D8C6C59CFFD0FFD1FED8FE79FE273F9D" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[350,499,475,501]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FED8FE79FE273F9D" box="[350,499,475,501]" italics="true" pageId="6" pageNumber="147">D. pulchellus</emphasis>
</taxonomicName>
, but with a minor callus and frontal depression superficial in
<taxonomicName id="D8C6C59CFFD0FFD1FB3AFE79FAE03F9D" authorityName="Bunalski" authorityYear="2001" box="[1212,1332,475,501]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD0FFD1FB3AFE79FAE03F9D" box="[1212,1332,475,501]" italics="true" pageId="6" pageNumber="147">D. malyszi</emphasis>
</taxonomicName>
; protibia tridentate in
<taxonomicName id="D8C6C59CFFD0FFD1FEA1FE5DFCDA3C71" authority="Pham, Ahrens &amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[295,782,511,537]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFD0FFD1FEA1FE5DFE123C71" box="[295,454,511,537]" italics="true" pageId="6" pageNumber="147">D. lexuanhuei</emphasis>
Pham, Ahrens &amp; Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD0FFD1FC9CFE5DFC713C70" bold="true" box="[794,933,510,537]" pageId="6" pageNumber="147">
<taxonomicNameLabel id="3681DF76FFD0FFD1FC9CFE5DFC713C70" box="[794,933,510,537]" pageId="6" pageNumber="147" rank="species">new species</taxonomicNameLabel>
</emphasis>
, but protibia bidentate in the two remaining species (
<figureCitation id="87FDA29AFFD0FFD1FF7FFD81FEA63C55" box="[249,370,547,573]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[161,1436,192,1626]" captionTargetId="figure-27@8.[151,1436,181,1626]" captionTargetPageId="8" captionText="FIGURE 6. Antennae, protibiae, and claws of Dedalopterus species.A, D, G, D. malyszi Bunalski, 2001; B, E, H, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611479" httpUri="https://zenodo.org/record/10611479/files/figure.png" pageId="6" pageNumber="147">
Fig. 6D
<emphasis id="2DB2620DFFD0FFD1FED7FD81FE8B3C55" bold="true" box="[337,351,547,573]" pageId="6" pageNumber="147"></emphasis>
F
</figureCitation>
); right paramere much shorter than the left one in
<taxonomicName id="D8C6C59CFFD0FFD1FC34FD81FA4C3C55" authority="Pham, Ahrens &amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[946,1432,547,573]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFD0FFD1FC34FD81FB843C55" box="[946,1104,547,573]" italics="true" pageId="6" pageNumber="147">D. lexuanhuei</emphasis>
Pham, Ahrens &amp; Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFD0FFD1FF11FDE5FEF03C08" bold="true" box="[151,292,582,609]" pageId="6" pageNumber="147">
<taxonomicNameLabel id="3681DF76FFD0FFD1FF11FDE5FEF03C08" box="[151,292,582,609]" pageId="6" pageNumber="147" rank="species">new species</taxonomicNameLabel>
</emphasis>
, parameres of equal length in
<taxonomicName id="D8C6C59CFFD0FFD1FD0CFDE5FCCB3C09" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[650,799,583,609]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFD0FFD1FD0CFDE5FCCB3C09" box="[650,799,583,609]" italics="true" pageId="6" pageNumber="147">D. pulchellus</emphasis>
</taxonomicName>
, and the right paramere shorter than the left one in
<taxonomicName id="D8C6C59CFFD0FFD1FA07FDE5FF3F3CED" authorityName="Bunalski" authorityYear="2001" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFD0FFD1FA07FDE5FF3F3CED" italics="true" pageId="6" pageNumber="147">D. malyszi</emphasis>
</taxonomicName>
(
<figureCitation id="87FDA29AFFD0FFD1FF7CFDC9FEB33CED" box="[250,359,619,645]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1935]" captionTargetBox="[151,1424,181,1885]" captionTargetId="figure-20@5.[151,1436,181,1887]" captionTargetPageId="5" captionText="FIGURE 4. Lateral view of elytra (AD), right lateral view of parameres tip (EI). A, E, Dedalopterus malyszi Bunalski, 2001; B, G, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, H, D. pulchellus Sabatinelli &amp; Pontuale, 1998 (paratype of D. pulchellus); D, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611475" httpUri="https://zenodo.org/record/10611475/files/figure.png" pageId="6" pageNumber="147">
Fig. 4E
<emphasis id="2DB2620DFFD0FFD1FEC8FDC9FE883CED" bold="true" box="[334,348,619,645]" pageId="6" pageNumber="147"></emphasis>
I
</figureCitation>
), and they also differ in stripes of scales of elytra (
<figureCitation id="87FDA29AFFD0FFD1FC18FDC9FBCF3CED" box="[926,1051,619,645]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1935]" captionTargetBox="[151,1424,181,1885]" captionTargetId="figure-20@5.[151,1436,181,1887]" captionTargetPageId="5" captionText="FIGURE 4. Lateral view of elytra (AD), right lateral view of parameres tip (EI). A, E, Dedalopterus malyszi Bunalski, 2001; B, G, D. pulchellus (paratype of D. bezdekorum Zídek &amp; Krajčík, 2007); C, H, D. pulchellus Sabatinelli &amp; Pontuale, 1998 (paratype of D. pulchellus); D, F, I, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species (paratype)." figureDoi="http://doi.org/10.5281/zenodo.10611475" httpUri="https://zenodo.org/record/10611475/files/figure.png" pageId="6" pageNumber="147">
Fig. 4A
<emphasis id="2DB2620DFFD0FFD1FC73FDC9FBD73CED" bold="true" box="[1013,1027,619,645]" pageId="6" pageNumber="147"></emphasis>
D
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41FD2DFED53CA5" pageId="6" pageNumber="147" type="etymology">
<paragraph id="1F79BE1FFFD0FFD1FF41FD2DFED53CA5" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FD2DFE9A3CC1" bold="true" box="[199,334,655,681]" pageId="6" pageNumber="147">Etymology.</emphasis>
This species is named after Assoc. Prof. Le Xuan Hue, former IEBR staff, who collected the
<typeStatus id="C07D00BDFFD0FFD1FF11FD11FF293CA5" box="[151,253,691,717]" pageId="6" pageNumber="147" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="57DCED94FFD0FFD1FF41FD74FAFD3D51" pageId="6" pageNumber="147" type="distribution">
<paragraph id="1F79BE1FFFD0FFD1FF41FD74FAFD3D51" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="147">
<emphasis id="2DB2620DFFD0FFD1FF41FD74FEB53C98" bold="true" box="[199,353,726,752]" pageId="6" pageNumber="147">Distribution.</emphasis>
<collectingCountry id="67D1FE8FFFD0FFD1FEEEFD75FE133C99" box="[360,455,727,753]" name="Vietnam" pageId="6" pageNumber="147">Vietnam</collectingCountry>
(
<collectingRegion id="DD0270FDFFD0FFD1FE51FD75FD643C99" box="[471,688,727,753]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="147">Kon Tum Province</collectingRegion>
, Ngoc Linh Mountain), located in the Central Highlands region of
<collectingCountry id="67D1FE8FFFD0FFD1FF11FD59FF2D3D7D" box="[151,249,763,789]" name="Vietnam" pageId="6" pageNumber="147">Vietnam</collectingCountry>
, near the border with
<collectingCountry id="67D1FE8FFFD0FFD1FE60FD59FDC83D7D" box="[486,540,763,789]" name="Laos" pageId="6" pageNumber="147">Laos</collectingCountry>
and
<collectingCountry id="67D1FE8FFFD0FFD1FDD7FD59FD1C3D7D" box="[593,712,763,789]" name="Cambodia" pageId="6" pageNumber="147">Cambodia</collectingCountry>
. Ngọc Linh (a Sino-Vietnamese name meaning “Jade Mountain”) is a mountain of the Annamite Range that stretches between the provinces of Quảng Nam and Kon Tum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,117 @@
<document id="92BED989ECA8E3BB53D5D60DE58E62E1" ID-DOI="10.11646/zootaxa.5406.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10611467" ID-ZooBank="4CB53437-B2F9-4C82-AEA9-E2E18B13A6AF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706895428130" checkinUser="plazi" docAuthor="Pham, Phu Van, Ahrens, Dirk &amp; Sabatinelli, Guido" docDate="2024" docId="976F0F09FFD7FFD6FF11FDFDFC3B3A52" docLanguage="en" docName="zootaxa.5406.1.7.pdf" docOrigin="Zootaxa 5406 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dedalopterus Sabatinelli &amp; Pontuale 1998" docType="treatment" docVersion="2" lastPageNumber="142" masterDocId="6B567771FFD6FFD7FF86FFA2FFD43E68" masterDocTitle="The genus Dedalopterus Sabatinelli &amp; Pontuale, 1998 (Scarabaeidae: Melolonthinae: Leucopholini) in Laos and Vietnam, with description of a new species" masterLastPageNumber="152" masterPageNumber="141" pageNumber="142" updateTime="1706895922865" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2CB13E0C5AA9DFFD7F6A5A45292CA45E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FE084983DAD64178EB6A144314E37D62">
<mods:title id="11D5792388A1D3882497D1CCE5E7F079">The genus Dedalopterus Sabatinelli &amp; Pontuale, 1998 (Scarabaeidae: Melolonthinae: Leucopholini) in Laos and Vietnam, with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="A6A5DA5892B98F23C7D770F018536E39" type="personal">
<mods:role id="DF11C1B5B3039D9A4869996094F38553">
<mods:roleTerm id="714EFD99B14FD1992543B27696B9E674">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2973796CC1BCD9F2FC091C2887A39A59">Pham, Phu Van</mods:namePart>
<mods:affiliation id="BEBC32ED6D7D44CD205A2112E035328A">Institute of Ecology and Biological resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam. &amp; Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn; Germany.</mods:affiliation>
</mods:name>
<mods:name id="99C4BC39B0CDE1B25FC08E1E68A5A0A0" type="personal">
<mods:role id="F93AEC68EBAEA4AC7625CD1A3EBB8790">
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</mods:role>
<mods:namePart id="4BDF788BFD881B897093BB5C700D7D54">Ahrens, Dirk</mods:namePart>
<mods:affiliation id="5D4BEC4BF2AB5ACEE18E2E7F8F49DA8C">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn; Germany.</mods:affiliation>
</mods:name>
<mods:name id="81ED83B71C2F400D9B7510762CF99E4F" type="personal">
<mods:role id="5DFF4082578240B5CD1AE9963CB6BE3B">
<mods:roleTerm id="C7FF9B3E1A6C9627B08C927BB46DBA03">Author</mods:roleTerm>
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Genus
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<emphasis id="2DB2620DFFD7FFD6FF68FDFDFE5B3C12" bold="true" box="[238,399,607,634]" italics="true" pageId="1" pageNumber="142">Dedalopterus</emphasis>
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<emphasis id="2DB2620DFFD7FFD6FF11FD1CFEF53CBE" box="[151,289,702,726]" italics="true" pageId="1" pageNumber="142">Dedalopterus</emphasis>
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:
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species.
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<emphasis id="2DB2620DFFD7FFD6FEBFFCA2FDFD3D72" box="[313,553,768,794]" italics="true" pageId="1" pageNumber="142">Cyphochilus signatus</emphasis>
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(by original designation;
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<emphasis id="2DB2620DFFD7FFD6FF41FC86FE963D56" bold="true" box="[199,322,804,830]" pageId="1" pageNumber="142">Diagnosis.</emphasis>
Elytra with four elevated costae in addition to sutural one, usually bearing scales grouped in white longitudinal stripes or white spots; eyes large but not very prominent; labrum symmetric, crescent-shaped and either of even thickness or mesally constricted; canthus narrow, black, variably covered by tan setae, and extends about one-third of eye length; frontoclypeal suture complete, slightly concave-forward; apical (outer terminal) segment oval or pointed, usually with dorsal callus; midline of pronotum broadly depressed and forms a longitudinal groove filled with round, imbricating white scales; apex of pygidium narrowly rounded, angular or drawn out into point in males and broadly rounded in females. Distal parts of parameres curve to right, and right paramere usually shorter, thinner, with apex dorsal-ventrally less expanded; ventral surface with hairs.
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Key to the species of
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(males only)
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Based on the key of
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, modified and updated
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<keyLead id="A4371D2AFFDCFFDDFF11FC3DFA483DDC" box="[151,1436,927,949]" pageId="10" pageNumber="151">1 Antennal club less than twice as long as remaining antennomeres................................................ 2</keyLead>
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<keyLead id="A4371D2AFFDCFFDDFF11FC1FFA483DBB" box="[151,1436,957,979]" pageId="10" pageNumber="151">- Antennal club twice or more than twice as long as remaining antennomeres....................................... 6</keyLead>
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<keyLead id="A4371D2AFFDCFFDDFF11FC78FA483D87" box="[151,1436,986,1008]" pageId="10" pageNumber="151">2 Parameres of unequal length............................................................................ 3</keyLead>
</paragraph>
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<keyLead id="A4371D2AFFDCFFDDFF11FC5AFA483A64" box="[151,1436,1015,1037]" pageId="10" pageNumber="151">- Parameres of equal length.............................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FBB7FA483A0C" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FBB7FA483A42" blockId="10.[151,1437,927,1595]" box="[151,1436,1045,1067]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FBB7FA483A42" box="[151,1436,1045,1067]" pageId="10" pageNumber="151">
3 Protibia bidentate, tegument of elytra light-brown to yellow, parameres short...............
<taxonomicName id="D8C6C59CFFDCFFDDFB0EFBB7FA483A42" ID-CoL="3XLRZ" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1160,1436,1045,1067]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="2DB2620DFFDCFFDDFB0EFBB7FAD93A42" box="[1160,1293,1045,1066]" italics="true" pageId="10" pageNumber="151">D. fujianensis</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFBB7FA413A42" author="Zhang, Y. W." box="[1306,1429,1045,1067]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FB90FA483A0C" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB90FA483A0C" pageId="10" pageNumber="151">
- Protibia tridentate (basal tooth small), tegument of elytra dark brown, parameres longer....................................................................................................
<taxonomicName id="D8C6C59CFFDCFFDDFBBEFBF2FA483A0C" ID-CoL="6CH84" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[1080,1436,1103,1125]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="itohi">
<emphasis id="2DB2620DFFDCFFDDFBBEFBF2FB553A0C" box="[1080,1153,1103,1125]" italics="true" pageId="10" pageNumber="151">D. itohi</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFB01FBEDFA483A0C" author="Sabatinelli, G. &amp; Pontuale, G." box="[1159,1436,1103,1125]" pageId="10" pageNumber="151" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FBCFFA483AC8" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FBCFFA483AEA" blockId="10.[151,1437,927,1595]" box="[151,1436,1132,1155]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FBCFFA483AEA" box="[151,1436,1132,1155]" pageId="10" pageNumber="151">
4 Parameres curved ventrally by 40
<superScript id="E8B31357FFDCFFDDFDA2FBCEFDFF3A10" attach="left" box="[548,555,1132,1144]" fontSize="5" pageId="10" pageNumber="151">o</superScript>
at distal third (lateral view)..........................
<taxonomicName id="D8C6C59CFFDCFFDDFBEFFBCFFA483AEA" ID-CoL="6CGVS" authority="Zidek &amp; Krajcik, 2007" authorityName="Zidek &amp; Krajcik" authorityYear="2007" box="[1129,1436,1133,1155]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="fencli">
<emphasis id="2DB2620DFFDCFFDDFBEFFBCFFB6F3AEA" box="[1129,1211,1133,1154]" italics="true" pageId="10" pageNumber="151">D. fencli</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFB44FBCFFA483AEA" author="Zidek, J. &amp; Krajcik, M." box="[1218,1436,1133,1155]" pageId="10" pageNumber="151" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FB28FA483AC8" blockId="10.[151,1437,927,1595]" box="[151,1436,1162,1184]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB28FA483AC8" box="[151,1436,1162,1184]" pageId="10" pageNumber="151">- Parameres entirely straight (lateral view)................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FB05FA483AB2" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FB05FA483AD4" blockId="10.[151,1437,927,1595]" box="[151,1436,1191,1213]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB05FA483AD4" box="[151,1436,1191,1213]" pageId="10" pageNumber="151">
5 Right paramere rather straight and larger..........................................
<taxonomicName id="D8C6C59CFFDCFFDDFBEBFB0AFA483AD4" ID-CoL="3XLS4" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1133,1436,1191,1213]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="melanodiscus">
<emphasis id="2DB2620DFFDCFFDDFBEBFB0AFAD93AD4" box="[1133,1293,1191,1213]" italics="true" pageId="10" pageNumber="151">D. melanodiscus</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFB05FA413AD4" author="Zhang, Y. W." box="[1306,1429,1191,1213]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FB67FA483AB2" blockId="10.[151,1437,927,1595]" box="[151,1436,1221,1243]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB67FA483AB2" box="[151,1436,1221,1243]" pageId="10" pageNumber="151">
- Right paramere curved moderately dorsally at apex and slender..........................
<taxonomicName id="D8C6C59CFFDCFFDDFBFBFB67FA483AB2" ID-CoL="3XLS2" authority="(Zhang, 2002)" baseAuthorityName="Zhang" baseAuthorityYear="2002" box="[1149,1436,1221,1243]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="2DB2620DFFDCFFDDFBFBFB67FAD93AB2" box="[1149,1293,1221,1242]" italics="true" pageId="10" pageNumber="151">D. hainanensis</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFB67FA423AB2" author="Zhang, Y. W." box="[1306,1430,1221,1243]" pageId="10" pageNumber="151" pagination="327 - 336" refId="ref6696" refString="Zhang, Y. W. (2002) Coleoptera: Scarabaeoidea. In: Huang, F. S. (Ed.), Forest Insects of Hainan. Science Press, Beijing, pp. 327 - 336. [in Chinese, with English description of new species]" type="book chapter" year="2002">Zhang, 2002</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FB40FA483B7C" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FB40FA483A9F" blockId="10.[151,1437,927,1595]" box="[151,1436,1250,1272]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB40FA483A9F" box="[151,1436,1250,1272]" pageId="10" pageNumber="151">6 Pygidium apically rounded.............................................................................. 7</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FAA2FA483B7C" blockId="10.[151,1437,927,1595]" box="[151,1436,1279,1301]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FAA2FA483B7C" box="[151,1436,1279,1301]" pageId="10" pageNumber="151">- Pygidium apically triangular............................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FABFFA483BE2" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FABFFA483B27" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FABFFA483B27" pageId="10" pageNumber="151">
7 Parameres rather straight and long, right paramere width almost equal to the left one (see
<figureCitation id="87FDA29AFFDCFFDDFBEAFABFFB7D3B5A" box="[1132,1193,1309,1331]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[323,1265,358,1905]" captionTargetId="figure-83@4.[310,1278,345,1905]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species (holotype,AC); D. malyszi Bunalski, 2001 (DF). A, D, parameres in left lateral view; B, E, parameres in dorsal view. C, F, parameres in right lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611473" httpUri="https://zenodo.org/record/10611473/files/figure.png" pageId="10" pageNumber="151">Fig. 3</figureCitation>
;
<bibRefCitation id="7B57C3EEFFDCFFDDFB31FABFFEC73B27" author="Sabatinelli, G. &amp; Pontuale, G." pageId="10" pageNumber="151" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
)..........................................................................
<taxonomicName id="D8C6C59CFFDCFFDDFB27FA99FA483B27" ID-CoL="6C8Y5" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[1185,1436,1338,1360]" class="Insecta" family="Scarabaeidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="2DB2620DFFDCFFDDFB27FA99FADA3B38" box="[1185,1294,1339,1360]" italics="true" pageId="10" pageNumber="151">D. signatus</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFA98FA413B27" author="Moser, J." box="[1306,1429,1338,1359]" pageId="10" pageNumber="151" pagination="353 - 367" refId="ref6425" refString="Moser, J. (1908) Neue Melolonthiden und Cetoniden. Annales de la Societe Entomologique de Belgique, 52 (12), 353 - 367. https: // doi. org / 10.5962 / bhl. part. 5014" type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FAFAFA483BE2" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FAFAFA483BE2" pageId="10" pageNumber="151">
- Parameres rather curved and short, right paramere narrower than left one (see
<figureCitation id="87FDA29AFFDCFFDDFC56FAF5FBDE3B04" box="[976,1034,1367,1389]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. AB, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; CD, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="10" pageNumber="151">Fig. 2</figureCitation>
,
<bibRefCitation id="7B57C3EEFFDCFFDDFB92FAF5FAFE3B04" author="Sabatinelli, G. &amp; Pontuale, G." box="[1044,1322,1367,1389]" pageId="10" pageNumber="151" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
)....................................................................................
<taxonomicName id="D8C6C59CFFDCFFDDFBF8FAD7FA483BE2" ID-CoL="3XLS3" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1150,1436,1397,1419]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="2DB2620DFFDCFFDDFBF8FAD7FAD93BE2" box="[1150,1293,1397,1418]" italics="true" pageId="10" pageNumber="151">D. intermedius</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFAD7FA413BE2" author="Zhang, Y. W." box="[1306,1429,1397,1419]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FA30FA483B8A" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FA30FA483BCF" blockId="10.[151,1437,927,1595]" box="[151,1436,1426,1448]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FA30FA483BCF" box="[151,1436,1426,1448]" pageId="10" pageNumber="151">8 Frontal depression deep; apical segment of maxillary palp with a large callus on its outer surface...................... 9</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FA12FA483B8A" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FA12FA483B8A" pageId="10" pageNumber="151">
- Frontal depression superficial, apical segment of maxillary palp with a minor callus on its outer surface.................................................................................................
<taxonomicName id="D8C6C59CFFDCFFDDFB26FA6FFA483B8A" ID-CoL="6CGVD" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[1184,1436,1485,1507]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFDCFFDDFB26FA6FFAD13B8A" box="[1184,1285,1485,1506]" italics="true" pageId="10" pageNumber="151">D. malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFA8DFA6FFA483B8A" author="Bunalski, M." box="[1291,1436,1485,1507]" pageId="10" pageNumber="151" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FA48FA483852" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FA48FA493875" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FA48FA493875" pageId="10" pageNumber="151">
9 Parameres unequal in length, right paramere shorter and slender than the left one............................................................................................
<taxonomicName id="D8C6C59CFFDCFFDDFC0EF9AAFAC93875" authority="Pham, Ahrens&amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[904,1309,1543,1565]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFDCFFDDFC0EF9AAFBDA3874" box="[904,1038,1543,1565]" italics="true" pageId="10" pageNumber="151">D. lexuanhuei</emphasis>
Pham, Ahrens&amp; Sabatinelli
</taxonomicName>
,
<emphasis id="2DB2620DFFDCFFDDFAA0F9A5FA493875" bold="true" box="[1318,1437,1543,1565]" pageId="10" pageNumber="151">
<taxonomicNameLabel id="3681DF76FFDCFFDDFAA0F9A5FA493875" box="[1318,1437,1543,1565]" pageId="10" pageNumber="151" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11F987FA483852" blockId="10.[151,1437,927,1595]" box="[151,1436,1573,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11F987FA483852" box="[151,1436,1573,1595]" pageId="10" pageNumber="151">
- Parameres equal in length.............................................
<taxonomicName id="D8C6C59CFFDCFFDDFB87F987FA483852" ID-CoL="6CGVF" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[1025,1436,1573,1595]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFDCFFDDFB87F987FB553852" box="[1025,1153,1573,1594]" italics="true" pageId="10" pageNumber="151">D. pulchellus</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFB01F987FA483852" author="Sabatinelli, G. &amp; Pontuale, G." box="[1159,1436,1573,1595]" pageId="10" pageNumber="151" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
</taxonomicName>
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</subSubSection>
</treatment>
</document>

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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.745.23685</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.745.23685</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-745-101</mods:identifier>
<mods:identifier type="ZooBank">8785DC6BC2A244FD94B6243EB07C717F</mods:identifier>
<mods:identifier type="Pensoft-UUID">047DFFCAFFA5F32EA97C873F4708943F</mods:identifier>
<mods:identifier type="Zenodo-Dep">1222435</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182230093" LSID="urn:lsid:plazi:treatment:976F3370CE0052E0A8BD2CCD4E114782" httpUri="http://treatment.plazi.org/id/976F3370CE0052E0A8BD2CCD4E114782" lastPageNumber="172" pageId="71" pageNumber="172">
<subSubSection pageId="71" pageNumber="172" type="nomenclature">
<paragraph pageId="71" pageNumber="172">
<taxonomicName LSID="976F3370-CE00-52E0-A8BD-2CCD4E114782" authority="Endrodi, 1964" authorityName="Endrodi" authorityYear="1964" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala huesingi" order="Coleoptera" pageId="71" pageNumber="172" phylum="Arthropoda" rank="species" species="huesingi">
Cyclocephala huesingi
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
, 1964
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="172" type="reference_group">
<paragraph pageId="71" pageNumber="172">
<taxonomicName authorityName="Endrodi" authorityYear="1964" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala huesingi" order="Coleoptera" pageId="71" pageNumber="172" phylum="Arthropoda" rank="species" species="huesingi">Cyclocephala huesingi</taxonomicName>
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
, 1964: 436-438 [original combination].
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="172" type="types">
<paragraph pageId="71" pageNumber="172">Types.</paragraph>
<paragraph pageId="71" pageNumber="172">
Holotype ♂ at MLUH (
<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B300" refString="Endrodi, S, 1964. Eine Reihe von neuen Cyclocephala-Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica 17: 433-470." title="Eine Reihe von neuen Cyclocephala-Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica 17: 433 - 470." year="1964">
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
[1964]
</bibRefCitation>
mistakenly stated the holotype was at BMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="172" type="distribution">
<paragraph pageId="71" pageNumber="172">Distribution.</paragraph>
<paragraph pageId="71" pageNumber="172">
COLOMBIA. VENEZUELA:
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="172" type="references">
<paragraph pageId="71" pageNumber="172">References.</paragraph>
<paragraph pageId="71" pageNumber="172">
<bibRefCitation author="Pike, KS" pageId="213" pageNumber="314" refId="B899" refString="Pike, KS, River, RL, Ratcliffe, BC, Oseto, CY, Mayo, ZB, 1976. A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1-32." title="A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1 - 32." year="1976">Pike et al. 1976</bibRefCitation>
,
<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B300" refString="Endrodi, S, 1964. Eine Reihe von neuen Cyclocephala-Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica 17: 433-470." title="Eine Reihe von neuen Cyclocephala-Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica 17: 433 - 470." year="1964">
<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
1964
</bibRefCitation>
, 966,
<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
, Restrepo et al. 2003,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1649/0010-065X-70.3.645" author="Gasca-Alvarez, HJ" pageId="182" pageNumber="283" refId="B383" refString="Gasca-Alvarez, HJ, Deloya, C, 2016. Description of the female of Cyclocephala monacha Ratcliffe, 2008 (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini), with a checklist of Cyclocephala Dejean species occurring in Colombia. The Coleopterists Bulletin 70: 645-653. https://doi.org/10.1649/0010-065X-70.3.645" title="Description of the female of Cyclocephala monacha Ratcliffe, 2008 (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini), with a checklist of Cyclocephala Dejean species occurring in Colombia. The Coleopterists Bulletin 70: 645 - 653." url="https://doi.org/10.1649/0010-065X-70.3.645" year="2016">
<normalizedToken originalValue="Gasca-Álvarez">Gasca-Alvarez</normalizedToken>
and Deloya 2016
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>