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<document ID-CLB-Dataset="297105" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="c7b986d2-d221-4c24-ae0a-037b5d4d8887" ID-ISBN="0-8018-8221-4" checkinTime="1670905166440" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="970B3F0C0D5513B386A3438BB54A1D06" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Geomyidae.859-870.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Orthogeomys (Macrogeomys) cavator" docType="treatment" docVersion="3" id="B149FDE7D3909CFBF93D8FFBC8EF1D48" lastPageNumber="864" masterDocId="B149FDE7D3909CFBF93D8FFBC8EF1D48" masterDocTitle="Order Rodentia - Family Geomyidae" masterLastPageNumber="870" masterPageNumber="859" pageNumber="864" updateTime="1716775291689" updateUser="ExternalLinkService">
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<mods:title id="1AE696B1136023A3E5561B7CE0806231">Order Rodentia - Family Geomyidae</mods:title>
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<mods:namePart id="C219533E27B819A4555625D697819179">Reeder, DeeAnn</mods:namePart>
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<mods:dateIssued id="09707225F0DDA7770AB08CCE0113B98D">2005</mods:dateIssued>
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<mods:title id="34B91B7FCEFC9D858897A7CC7707C3B6">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<treatment id="970B3F0C0D5513B386A3438BB54A1D06" ID-GBIF-Taxon="231543155" LSID="urn:lsid:plazi:treatment:970B3F0C0D5513B386A3438BB54A1D06" httpUri="http://treatment.plazi.org/id/970B3F0C0D5513B386A3438BB54A1D06" lastPageNumber="864" pageNumber="864" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12800099">
<subSubSection id="7E3DBA1E79602081D7BBC2CE3695AD44" pageNumber="864" type="nomenclature">
<paragraph id="0E712D1FA1EA3B5354260440ABA4A35F" pageNumber="864">
<taxonomicName id="B9B26E5F6D9828C68728569C7A8760B2" ID-CoL="7W6MG" authority="(Bangs 1902)" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageNumber="864" phylum="Chordata" rank="species" species="cavator" subGenus="Macrogeomys">
<emphasis id="4D7EC427666DB44E930C84BDFF7ADBA4" italics="true" pageNumber="864">Orthogeomys (Macrogeomys) cavator</emphasis>
(Bangs 1902)
</taxonomicName>
</paragraph>
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<subSubSection id="1408B0B2B3345C9AD5D0E0993A537E27" pageNumber="864" type="reference_group">
<paragraph id="5BC50142D928F1FD21DEC98A3050589C" pageNumber="864">
<treatmentCitation id="ABD57001E73CD0D7FC708E82AC063271" author="null" authority="(Bangs 1902)" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" journal="Bull. Mus. Comp. Zool." kingdom="Animalia" order="Rodentia" page="42" pageNumber="864" phylum="Chordata" rank="species" species="cavator" subGenus="Macrogeomys" title="Orthogeomys (Macrogeomys) cavator" volume="39" year="-1">
<taxonomicName id="A60E958761A9CCD972C0A8FBA997E917" ID-CoL="7W6MG" authority="(Bangs 1902)" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageNumber="864" phylum="Chordata" rank="species" species="cavator" subGenus="Macrogeomys">
<emphasis id="8AE4DF72378528100582AE2FD89FD016" italics="true" pageNumber="864">Orthogeomys (Macrogeomys) cavator</emphasis>
(
<bibRefCitation id="9E40A0CC93A6C624DB04B5684F01AD32" author="Bangs, O." refId="ref19700" refString="Bangs, O. 1902. Chiriqui Mammalia. Bulletin of the Museum of Comparative Zoology at Harvard College, 39: 17 - 51." year="1902">Bangs 1902</bibRefCitation>
)
</taxonomicName>
,
<bibCitation id="2ED124A5FCE4B4C0B1DAA8D6F3EB42EA" author="null" journal="Bull. Mus. Comp. Zool." pageNumber="864" pagination="42" title="Orthogeomys (Macrogeomys) cavator" volume="39" year="-1">
Bull.
<taxonomicName id="7375F49E8C3ECD913A6D3E5E6EC400A3" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">Mus</taxonomicName>
. Comp. Zool., 39: 42
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5B9A0B317052B9E40B0BEE9BA2FC942" pageNumber="864" type="materials_examined">
<paragraph id="1FAA9908498869138D97793E9D4A387E" pageNumber="864">
<emphasis id="4A1F9B624500EB1E3AC0B939E8A1A0F1" inLineHeading="true" pageNumber="864">Type Locality:</emphasis>
<materialsCitation id="ED6E678D75CBE9E15BA5B46A1702D565" ID-GBIF-Occurrence="4867332314" country="Panama" elevation="1.463" pageNumber="864" stateProvince="Chiriqui">
<collectingCountry id="529F791D1E551BE9853B93056843EEA7" name="Panama">Panama</collectingCountry>
,
<collectingRegion id="AEA7E31AC0E1A8C86790420921D76D8A" country="Panama" name="Chiriqui">Chiriqui Prov.</collectingRegion>
, Boquete,
<quantity id="E6AF6A0A3037A8D643E339DA34C35B34" metricMagnitude="0" metricUnit="m" metricValue="1.46304" unit="ft" value="4.8">4,800 ft.</quantity>
(
<quantity id="6160FFD8DA7F6C8B77FCC669B2731F5C" metricMagnitude="0" metricUnit="m" metricValue="1.463" unit="m" value="1.463">
<elevation id="AF10B9E5F9DB2DF2405B3F64871AF497">1,463 m</elevation>
</quantity>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B18B51CD8460F80C42126E54174FF63B" pageNumber="864" type="vernacular_names">
<paragraph id="CF47F2B1CF6E08E7F1CAE59BAC96BF7A" pageNumber="864">
<emphasis id="434F2423126BF6EA5329C92A643A2A64" inLineHeading="true" pageNumber="864">Vernacular Names:</emphasis>
<vernacularName id="B0889EED5D804B281986D113733DAC74" language="eng" pageNumber="864">
<collectingRegion id="D968834BC2FD4A944729CA85B1A649D7" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
Pocket Gopher
</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="3F8B071F7DE213027639C63B29EDFED0" pageNumber="864" type="child_taxa">
<paragraph id="F166D8F0F80F06D91D80DB189370E29F" pageNumber="864">
<emphasis id="CBE3731EBF8021BDD6B4370E44FAC7C1" inLineHeading="true" pageNumber="864">Subspecies:</emphasis>
:
</paragraph>
<paragraph id="FDEF6ED8E9BA0F59CD787D3056A294E3" pageNumber="864">
Subspecies
<taxonomicName id="51A055F9B71C18FC5FF562698F00842C" authority="(Bangs 1902)" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageNumber="864" phylum="Chordata" rank="subSpecies" species="cavator" subGenus="Macrogeomys" subSpecies="cavator">
<emphasis id="D448E1595C2A9C49A1B9BE904A149726" italics="true" pageNumber="864">Orthogeomys (Macrogeomys) cavator</emphasis>
subsp.
<emphasis id="A7B1E7AE5ECF0843D96AF2530A2BD0DB" italics="true" pageNumber="864">cavator</emphasis>
(
<bibRefCitation id="C78DC99BD20FFE14A2C620E0CE06EB7C" author="Bangs, O." refId="ref19700" refString="Bangs, O. 1902. Chiriqui Mammalia. Bulletin of the Museum of Comparative Zoology at Harvard College, 39: 17 - 51." year="1902">Bangs 1902</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="D32C91DF106B58ADE717E4BD07D627FD" pageNumber="864">
Subspecies
<taxonomicName id="5C86F3D4070B0F8551840D44BF2CA123" authority="(Goodwin 1943)" baseAuthorityName="Goodwin" baseAuthorityYear="1943" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageNumber="864" phylum="Chordata" rank="subSpecies" species="cavator" subGenus="Macrogeomys" subSpecies="nigrescens">
<emphasis id="69B0F29B242CC7281C6DD16276E0CCB4" italics="true" pageNumber="864">Orthogeomys (Macrogeomys) cavator</emphasis>
subsp.
<emphasis id="C48F504B98A573BF9E12B29167CCE9DF" italics="true" pageNumber="864">nigrescens</emphasis>
(Goodwin 1943)
</taxonomicName>
</paragraph>
<paragraph id="814F1D88E4E0684CEA97C25B935D7CA7" pageNumber="864">
Subspecies
<taxonomicName id="B2A055300F278BB272BC815D60605916" authority="(Bangs 1902)" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageNumber="864" phylum="Chordata" rank="subSpecies" species="cavator" subGenus="Macrogeomys" subSpecies="pansa">
<emphasis id="BB327768F1C94FF6600368F4101198F3" italics="true" pageNumber="864">Orthogeomys (Macrogeomys) cavator</emphasis>
subsp.
<emphasis id="26E1D0B266EA654979E9D6569E9B4830" italics="true" pageNumber="864">pansa</emphasis>
(
<bibRefCitation id="708E28EBB2E2066B07768A8B04F2A7D1" author="Bangs, O." refId="ref19700" refString="Bangs, O. 1902. Chiriqui Mammalia. Bulletin of the Museum of Comparative Zoology at Harvard College, 39: 17 - 51." year="1902">Bangs 1902</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D9C443AD2DE016C958B28D5A61E16D77" pageNumber="864" type="distribution">
<paragraph id="2AF46A1DB71002B95C760EAA6D010D7F" pageNumber="864">
<emphasis id="EF854E5F42A554B73BC6B343068A382E" inLineHeading="true" pageNumber="864">Distribution:</emphasis>
NW
<collectingCountry id="85E1F33D4BD45FB80B54C717326C6AC1" name="Panama">Panama</collectingCountry>
to C
<collectingCountry id="E51A96C7A2D215FDD09CB6BD27CD5579" name="Costa Rica">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="872A25116A054603794E6EF1FD0E6C3F" pageNumber="864" type="conservation">
<paragraph id="2C2410EEB4D2119C779974C7176FF3D4" pageNumber="864">
<emphasis id="693A304D758182A928E949B43D3B0E08" inLineHeading="true" pageNumber="864">Conservation:</emphasis>
<collectionCode id="DCF3400DA0865FEDC3E749B92FF31690">IUCN</collectionCode>
– Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="60D8F6FACBF4D8F2654E886C799362EA" pageNumber="864" type="discussion">
<paragraph id="20A1C23156BCBC5A4577F937095BD065" pageNumber="864">
<emphasis id="9506A814CDE05D987CECA76F3BB91807" inLineHeading="true" pageNumber="864">Discussion:</emphasis>
Subgenus
<taxonomicName id="DAE2BE94946DA0789232220388B668B1" authorityName="Merriam" authorityYear="1895" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subGenus" subGenus="Macrogeomys">
<emphasis id="D0BFBED77C41AFA6929A4DB70A2956AE" italics="true" pageNumber="864">Macrogeomys</emphasis>
</taxonomicName>
. Reviewed by
<bibRefCitation id="D5CCDDFAEEA560BF7FF476ACB223CC8E" author="Goodwin, G. G." refId="ref125480" refString="Goodwin, G. G. 1946. Mammals of Costa Rica. Bulletin of the American Museum of Natural History, 87: 271 - 471." year="1946">Goodwin (1946)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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Tribe
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(Crantz) Fritsch
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Artbeschreibung: Sehr
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wie
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Rand der
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seidenhaarig. Der mittlere Endzahn kaum kleiner als die benachbarten
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Standort und Verbreitung in der Schweiz: Rasen, Weiden, vorwiegend auf Kalk / (montan-)subalpin-alpin / A, M in
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bis SO)
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Zeigerwerte nach
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<emphasis id="1DA14B5ECD1E4BAD14F157C75D710998" bold="true" pageNumber="258">Klimafaktoren</emphasis>
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<emphasis id="8F6C5B3E1B19F88FEB3FF8420DC8B07A" bold="true" pageNumber="258">Salztoleranz</emphasis>
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<td id="E9CB259DCFF05D47D811A1B567FB2522" pageNumber="258">Feuchtezahl F</td>
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frisch; Feuchtigkeit
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wechselnd (
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<td id="A79FFD737EFB5BDB717AF28CB5872DAD" pageNumber="258">Salzzeichen</td>
<td id="168BD6CFBB4AB0F42F99CA01A046AF9D" pageNumber="258" style="text-align:left;width:60px">--</td>
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<td id="3F775A6E6A78125A36E04ADE5E7461D9" pageNumber="258">Reaktionszahl R</td>
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unter-alpin, supra-subalpin und ober-subalpin (
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)
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<normalizedToken id="CDA0260832BEBEAA9C17F01554FF00B7" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
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<normalizedToken id="3B192A063327751BDB26C184B4A42040" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
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<td id="4DBE9AD5BC18DB541F364C9D9FCE063E" pageNumber="258" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
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Volksname Deutscher Name:
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Nom
<normalizedToken id="9BA81FF97E03AD6090B2B2FDA0F48A16" originalValue="français">francais</normalizedToken>
:
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,
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Nome italiano:
<vernacularName id="0F24E459241E2C5D06E024074A38AE9B" language="ita" pageNumber="258">Cinquefoglie di Crantz</vernacularName>
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256
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<treatment LSID="urn:lsid:plazi:treatment:970B9754C1965BA085A5B629474EDB39" httpUri="http://treatment.plazi.org/id/970B9754C1965BA085A5B629474EDB39" lastPageNumber="1" pageId="0" pageNumber="1">
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8.
<taxonomicName ID-CoL="7SZWW" LSID="970B9754-C196-5BA0-85A5-B629474EDB39" authority="Blanford, 1902" authorityName="Blanford" authorityYear="1902" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma dalyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dalyi">Rhiostoma dalyi Blanford, 1902</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">Figs 18</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">, 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Blanford" authorityYear="1902" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma dalyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dalyi">Rhiostoma dalyi</taxonomicName>
Blanford, 1902: 34, 35, fig. 1. Type locality: Juxta Phitsanulok, in sylvis humidis etr densis [Phitsanulok Province, Thailand].
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065982" author="Blanford, WT" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="274 - 284" refId="B21" refString="Blanford, WT, 1903. Notes on Mr. W. M. Daly's collections of land and fresh-water Mollusca from Siam. Proceedings of the Malacological Society of London 5 (4): 274 - 284, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065982" title="Notes on Mr. W. M. Daly's collections of land and fresh-water Mollusca from Siam." url="https://doi.org/10.1093/oxfordjournals.mollus.a065982" volume="5" year="1903">Blanford 1903</bibRefCitation>
: 281.
<bibRefCitation author="Fischer, H" editor="Pavie, A" journalOrPublisher="Ernest Leroux, Editeur, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B42" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour. In: Pavie, A, Ed., Mission Pavie, Indo-Chine 1879-1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III. Ernest Leroux, Editeur, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour." volumeTitle="Mission Pavie, Indo-Chine 1879 - 1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
: 427.
<bibRefCitation author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
: 11.
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
: 21, fig. 4i, j.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei</taxonomicName>
-
<bibRefCitation author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
: 41-46, figs 4.10-4.12 (misidentified).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
NHMUK 1902.1.24.14-16 (3 shells; Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">25A</figureCitation>
) from Pitsunaloke, Siam. This syntype lot in the Blanford ex. Daly collection consists of three specimens; the figured specimen in the original description can be recognised by its curved breathing device and is figured herein.
</paragraph>
<caption doi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
Shell of
<taxonomicName authorityName="Blanford" authorityYear="1902" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma dalyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dalyi">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma dalyi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype NHMUK 1902.1.24.14-16 from Juxta Phitsanulok
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
specimens CUMZ 4848 from Tham Erawan, Long, Prae
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Phayao" specimenCount="1" stateProvince="Phayao">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Thailand">Thailand</collectingCountry>
</emphasis>
: Tham Pha Dang, Chiangkham,
<collectingRegion country="Thailand" name="Phayao">Phayao</collectingRegion>
: CUMZ 4462, 4729. Doi Phunang, Dokkamtai,
<collectingRegion country="Thailand" name="Phayao">Phayao</collectingRegion>
: CUMZ 3938, 3942, 4313, 4319, 4420. Tarnsawan Waterfall, Dokkamtai,
<collectingRegion country="Thailand" name="Phayao">Phayao</collectingRegion>
: CUMZ 3824, 3826, 3977 (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">25D</figureCitation>
). Tham Pha Tangyai, Pong,
<collectingRegion country="Thailand" name="Phayao">Phayao</collectingRegion>
: CUMZ 3926 (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">25E, F</figureCitation>
), 4784. Tham Pha Tung, Pong,
<collectingRegion country="Thailand" name="Phayao">Phayao</collectingRegion>
: CUMZ 10062. Ban Tha Sri, Muang,
<collectingRegion country="Thailand" name="Lampang">Lampang</collectingRegion>
: CUMZ 4728. Pratupha, Ngoaw,
<collectingRegion country="Thailand" name="Lampang">Lampang</collectingRegion>
: CUMZ 4463. 3 km. before the junction to Tham Sereethai, Long, Prae: CUMZ 4450. Hill Near Tham Air Tammachard, Long, Prae: CUMZ 3979, 4451. Tham Air Thammachad, Ban Pan, Long, Prae: CUMZ 4762. Tham Erawan, Long, Prae: CUMZ 4792, 4848 (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">25B, C</figureCitation>
), 10063, 10064. Wat Tham Pha Nang Koy, Rong Kwang, Prae: CUMZ 3884, 3924, 4726
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell large, thick, and depressed. Detached whorl medium in length, and slightly descending. Breathing device tubular, curved, and attached to preceding whorl. Shell whitish with brownish zigzag patterns.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is very similar to
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
</taxonomicName>
from central Thailand. The distinguishing characteristics of
<taxonomicName lsidName="R. dalyi" pageId="0" pageNumber="1" rank="species" species="dalyi">
<emphasis italics="true" pageId="0" pageNumber="1">R. dalyi</emphasis>
</taxonomicName>
are a stout last whorl and a tubular-shaped breathing device that is short, curved and attached to the preceding whorl. In comparison,
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
</taxonomicName>
has a slender last whorl, and a long and straight tubular breathing device. In addition, this species also differs from
<taxonomicName lsidName="R. trigina" pageId="0" pageNumber="1" rank="species" species="trigina">
<emphasis italics="true" pageId="0" pageNumber="1">R. trigina</emphasis>
</taxonomicName>
sp. nov. in having a stout and short detached last whorl, a short and curved tubular breathing device, and a ventral shell surface usually without a pattern. In contrast,
<taxonomicName lsidName="R. trigina" pageId="0" pageNumber="1" rank="species" species="trigina">
<emphasis italics="true" pageId="0" pageNumber="1">R. trigina</emphasis>
</taxonomicName>
sp. nov. possesses a slender detached whorl, long and straight tubular breathing device, and usually with brown zigzag patterns on both sides.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell medium to large, cW 19.9-24.6 mm, cH 9.7-13.4 mm, thick, and discoidal shape; detached-whorl length 7.5-14.0 mm. Apex acute with dark colouration; spire slightly elevated. Whorls 5 to 6, convex and increasing regularly; suture wide and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thick or thin corneous and transparent. Shell colour whitish with brownish zigzag pattern; ventral shell surface usually without pattern; with narrow and dark brown spiral band on periphery. Detached whorl of medium length, approximately the same length as apertural width. Peristome circular and double; lip thickened, slightly expanded and multi-layered. Aperture opened sub-laterally. Breathing device tubular, curved and its tip attached to preceding whorl; outer lip forming a short and closed tube, and located just behind apertural lip; inner lip with hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, cup-shaped, and multispiral (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Shell of Rhiostoma dalyi A syntype NHMUK 1902.1.24.14 - 16 from Juxta Phitsanulok B, C specimens CUMZ 4848 from Tham Erawan, Long, Prae D specimen CUMZ 3979 from Tarnsawan Waterfall, Phayao E, F specimens CUMZ 3926 from Tham Pha Tangyai, Pong, Phayao." figureDoi="10.3897/zookeys.1142.90097.figure25" httpUri="https://binary.pensoft.net/fig/799883" pageId="0" pageNumber="1">25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is distributed in the northern part of Thailand in Phitsanuloke (type locality), Lampang, Phrae and Phayao provinces (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem (1966)</bibRefCitation>
record the specimens from 'Chieng
<normalizedToken originalValue="Dao’">Dao'</normalizedToken>
as this species. However, we examined a number of specimens from Chieng Dao, Chieng Mai Province and found clearly distinct shell morphology compared to the type specimen of
<taxonomicName lsidName="R. dalyi" pageId="0" pageNumber="1" rank="species" species="dalyi">
<emphasis italics="true" pageId="0" pageNumber="1">R. dalyi</emphasis>
</taxonomicName>
. In addition, the COI barcoding suggests that
<taxonomicName lsidName="R. dalyi" pageId="0" pageNumber="1" rank="species" species="dalyi">
<emphasis italics="true" pageId="0" pageNumber="1">R. dalyi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="R. trigina" pageId="0" pageNumber="1" rank="species" species="trigina">
<emphasis italics="true" pageId="0" pageNumber="1">R. trigina</emphasis>
</taxonomicName>
sp. nov. are separate species (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum likelihood (ML) tree showing the relationships among Rhiostoma species based on 660 bp of the mitochondrial COI gene sequence. The major nodes are indicated by numbers as the Maximum likelihood (ML) bootstrap values and Bayesian posterior probabilities of Bayesian inference (BI), respectively. Nodes with sufficient support by both ML and BI are indicated by black circles, and nodes with partial support by only ML are indicated by grey circles, and only BI by white circles. Newly described species in this study are highlighted in blue." figureDoi="10.3897/zookeys.1142.90097.figure3" httpUri="https://binary.pensoft.net/fig/799861" pageId="0" pageNumber="1">3</figureCitation>
). Therefore, the record of this species by
<bibRefCitation author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem (1966)</bibRefCitation>
should be referred to
<taxonomicName lsidName="R. trigina" pageId="0" pageNumber="1" rank="species" species="trigina">
<emphasis italics="true" pageId="0" pageNumber="1">R. trigina</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
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<treatment LSID="urn:lsid:plazi:treatment:970BF0489D895243BB0DD8AA02D45FC5" httpUri="http://treatment.plazi.org/id/970BF0489D895243BB0DD8AA02D45FC5" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="970BF048-9D89-5243-BB0D-D8AA02D45FC5" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister cinctus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cinctus">Cybister cinctus Sharp, 1882</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">Madagascar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<specimenCount type="generic">1 ex.</specimenCount>
: MAK-1A;
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="male">1 ♂</specimenCount>
: MAK-2;
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">3 ♀♀</specimenCount>
: MAK-19;
<specimenCount type="male">4 ♂♂</specimenCount>
,
<specimenCount type="generic">
1
<normalizedToken originalValue="еx">ex</normalizedToken>
.
</specimenCount>
: MAK-20;
<specimenCount type="female">1 ♀</specimenCount>
: MAK-23;
<specimenCount type="female">1 ♀</specimenCount>
: MAK-41.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar (widespread and common) (
<bibRefCitation author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="0" pageNumber="1" refId="B40" refString="Guignot, F, 1959-1961. Revision des Hydrocanthares d'Afrique (ColeopteraDytiscoidea), parts 1-3. Annales du Musee Royal du Congo Belge Tervuren (8) Sciences Zoologiques 70, 78, 90: 1-995." title="Revision des Hydrocanthares d'Afrique (ColeopteraDytiscoidea), parts 1 - 3. Annales du Musee Royal du Congo Belge Tervuren (8) Sciences Zoologiques 70, 78, 90: 1 - 995." year="1959 - 1961">Guignot 1959-1961</bibRefCitation>
;
<bibRefCitation author="Bertrand, H" journalOrPublisher="Cahiers ORSTOM, serie hydrobiologie" pageId="0" pageNumber="1" pagination="241 - 249" refId="B8" refString="Bertrand, H, Legros, C, 1971. Hydrocanthares (excl. Gyrinidae) recueillis a Madagascar (mission Bertrand, 1960). Cahiers ORSTOM, serie hydrobiologie 5 (3-4): 241 - 249" title="Hydrocanthares (excl. Gyrinidae) recueillis a Madagascar (mission Bertrand, 1960)." volume="5" year="1971">Bertrand and Legros 1971</bibRefCitation>
;
<bibRefCitation author="Bameul, F" journalOrPublisher="Nouvelle Revue d'Entomologie" pageId="0" pageNumber="1" pagination="87 - 103" refId="B2" refString="Bameul, F, 1984. Haliplidae, Noteridae, Dytiscidae et Gyrinidae captures dans les iles Mascareignes et a Madagascar par Yves Gomy (Coleoptera, Hydradephaga). Nouvelle Revue d'Entomologie 1 (1): 87 - 103" title="Haliplidae, Noteridae, Dytiscidae et Gyrinidae captures dans les iles Mascareignes et a Madagascar par Yves Gomy (Coleoptera, Hydradephaga)." volume="1" year="1984">Bameul 1984</bibRefCitation>
;
<bibRefCitation author="Rocchi, S" journalOrPublisher="Frustula Entomologica" pageId="0" pageNumber="1" pagination="71 - 89" refId="B56" refString="Rocchi, S, 1991. Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera). Frustula Entomologica 14 (27): 71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0120777" author="Bukontaite, R" journalOrPublisher="Nova Series" pageId="0" pageNumber="1" refId="B24" refString="Bukontaite, R, Ranarilalatiana, T, Randriamihaja, JH, Bergsten, J, 2015. In or out-of- Madagascar? Colonization patterns for large-bodied diving beetles (Coleoptera: Dytiscidae). PLoS ONE 10(3): e0120777. https://doi.org/10.1371/journal.pone.0120777" title="In or out-of- Madagascar? Colonization patterns for large-bodied diving beetles (Coleoptera: Dytiscidae). PLoS ONE 10 (3): e 0120777." url="https://doi.org/10.1371/journal.pone.0120777" year="2015">Bukontaite et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat in study area</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei." figureDoi="10.3897/zookeys.1127.85737.figure2" httpUri="https://binary.pensoft.net/fig/764728" pageId="0" pageNumber="1">2B, C</figureCitation>
).
</emphasis>
This species was collected only at peripheral sites, in various kinds of water bodies (lentic or slowly flowing) in open areas. It prefers habitats with at least some vegetation and is tolerant to anthropogenic pressure.
</paragraph>
</subSubSection>
</treatment>
</document>