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<document ID-CLB-Dataset="297092" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="29b06347-2a16-47e8-8deb-c8084f3522bd" ID-ISBN="0-8018-8221-4" checkinTime="1670905252754" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="137D534DD05843C71C3177FB5B6D06F9" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Sciuridae.754-818.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Aeromys thomasi" docType="treatment" docVersion="4" id="91C4B717759630DB830679AF52629BE0" lastPageNumber="766" masterDocId="91C4B717759630DB830679AF52629BE0" masterDocTitle="Order Rodentia - Family Sciuridae" masterLastPageNumber="818" masterPageNumber="754" pageNumber="766" updateTime="1716778896931" updateUser="ExternalLinkService">
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<mods:title id="0CA85E607763A5FD44AF56229CADD1FD">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<emphasis id="22211EEF032B2946B4CDC83C626D3221" italics="true" pageNumber="766">Aeromys thomasi</emphasis>
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<emphasis id="EAA507CBBF7B46EB467C19ADC54B6B49" italics="true" pageNumber="766">[Aeromys] thomasi</emphasis>
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.
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<emphasis id="1EF6044C0FD79974DC33ABB6B61FE1D4" inLineHeading="true" pageNumber="766">Type Locality:</emphasis>
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&quot;Silat River, about
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&quot; [Baram,
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,
<collectingCountry id="128E963DD5F08AC319C49B6CF2FC51E2" name="Malaysia">Malaysia</collectingCountry>
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<emphasis id="98A4CAF87807D8F3E7CEE6D5FDD3AE72" inLineHeading="true" pageNumber="766">Vernacular Names:</emphasis>
<vernacularName id="0DBEB719ACE1CC9D410B2D1C39501392" language="eng" pageNumber="766">Thomas's Flying Squirrel</vernacularName>
.
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Borneo, except SE.
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<emphasis id="5832FCF135660377329D6C463A8CDD54" inLineHeading="true" pageNumber="766">Conservation:</emphasis>
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Lower Risk (lc).
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<emphasis id="A9A01B34A321DB36FF56E82DFE7BF8F9" bold="true" box="[151,411,1793,1819]" italics="true" pageId="10" pageNumber="46">Spirostreptus unciger</emphasis>
(
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<figureCitation id="03EFDBA3A321DB36FF56E808FF01F8DC" box="[151,225,1828,1854]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="10" pageNumber="46">Figs 5</figureCitation>
,
<figureCitation id="03EFDBA3A321DB36FF2FE808FF1CF8DC" box="[238,252,1828,1854]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="10" pageNumber="46">7</figureCitation>
d
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<taxonomicName id="5CD4BCA5A321DB36FF56E85EFDC8F86A" ID-CoL="5857D" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,552,1905,1929]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A321DB36FF56E85EFE75F86B" box="[151,405,1906,1929]" italics="true" pageId="10" pageNumber="46">Triaenostreptus unciger</emphasis>
<bibRefCitation id="FF45BAD7A321DB36FE5DE85DFDC8F86A" author="Attems" box="[412,552,1905,1928]" pageId="10" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
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, p. 365, figs 318320;
<bibRefCitation id="FF45BAD7A321DB36FCDDE85DFC5BF86A" author="Hoffman" box="[796,955,1905,1928]" pageId="10" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A321DB36FC07E85DFBB3F86A" author="Krabbe" box="[966,1107,1905,1928]" pageId="10" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A321DB36FB9FE85DFB03F86A" author="Hamer" box="[1118,1251,1905,1928]" pageId="10" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
<taxonomicName id="5CD4BCA5A321DB36FF56E8BEFDC4F84A" ID-CoL="4Z8Y5" authority="Hoffman 2008" authorityName="Hoffman" authorityYear="2008" box="[151,548,1937,1961]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A321DB36FF56E8BEFE9AF84B" box="[151,378,1938,1961]" italics="true" pageId="10" pageNumber="46">Spirostreptus unciger</emphasis>
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<bibRefCitation id="FF45BAD7A321DB36FE48E8BDFDC4F84A" author="Hoffman" box="[393,548,1937,1960]" pageId="10" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
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<subSubSection id="D3CE94ADA321DB37FF56E8F4FC2BFEBA" lastPageId="11" lastPageNumber="47" pageId="10" pageNumber="46" type="materials_examined">
<paragraph id="9B6BC726A321DB36FF56E8F4FB2BF810" blockId="10.[151,1227,2008,2034]" box="[151,1227,2008,2034]" pageId="10" pageNumber="46">
<emphasis id="A9A01B34A321DB36FF56E8F4FEA6F810" bold="true" box="[151,326,2008,2034]" pageId="10" pageNumber="46">
<typeStatus id="446F7984A321DB36FF56E8F4FF32F810" box="[151,210,2008,2034]" pageId="10" pageNumber="46">Type</typeStatus>
material
</emphasis>
(not examined):
<collectingCountry id="E3C387B6A321DB36FDCFE8F4FD07F810" box="[526,743,2008,2034]" name="South Africa" pageId="10" pageNumber="46">SOUTH AFRICA</collectingCountry>
:
<emphasis id="A9A01B34A321DB36FD33E8F4FBE1F810" bold="true" box="[754,1025,2008,2034]" pageId="10" pageNumber="46">North West Province:</emphasis>
Vryburg (
<collectionCode id="FDC55FE3A321DB36FBB8E8F4FB5EF810" box="[1145,1214,2008,2034]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="10" pageNumber="46">SAM</collectionCode>
).
</paragraph>
<paragraph id="9B6BC726A320DB37FF04EFB6FC2BFEBA" blockId="11.[151,1436,152,344]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04EFB6FDA5FF56" bold="true" box="[197,581,154,180]" pageId="11" pageNumber="47">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A320DB37FD8DEFB6FCC5FF56" box="[588,805,154,180]" name="South Africa" pageId="11" pageNumber="47">SOUTH AFRICA</collectingCountry>
:
<emphasis id="A9A01B34A320DB37FCF1EFB6FBFBFF56" bold="true" box="[816,1051,154,180]" pageId="11" pageNumber="47">Gauteng Province:</emphasis>
1 3, Pretoria [25
<geoCoordinate id="FEE0A1E1A320DB37FB29EFB4FAC5FF56" box="[1256,1317,152,180]" direction="south" orientation="latitude" pageId="11" pageNumber="47" precision="925" value="-0.73333335">0 44'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A320DB37FA8EEFB4FA6EFF56" box="[1359,1422,152,180]" direction="east" orientation="longitude" pageId="11" pageNumber="47" precision="925" value="0.18333334">0 11'E</geoCoordinate>
],
<date id="EF6AE1E6A320DB37FF56EFE8FF0DFF3C" box="[151,237,196,222]" pageId="11" pageNumber="47" value="1961-10">x.1961</date>
, H. Fiedler (
<collectionCode id="FDC55FE3A320DB37FE46EFE8FE00FF3C" LSID="urn:lsid:biocol.org:col:34985" box="[391,480,196,222]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="47">NMSA</collectionCode>
8896); 1 3, Pretoria [25
<geoCoordinate id="FEE0A1E1A320DB37FCD1EFEDFCB0FF3C" box="[784,848,193,222]" direction="south" orientation="latitude" pageId="11" pageNumber="47" precision="925" value="-0.73333335">0 44'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A320DB37FCBDEFEDFC5DFF3C" box="[892,957,193,222]" direction="east" orientation="longitude" pageId="11" pageNumber="47" precision="925" value="0.18333334">0 11'E</geoCoordinate>
], 2004, (
<collectionCode id="FDC55FE3A320DB37FBE8EFE8FB62FF3C" LSID="urn:lsid:biocol.org:col:34985" box="[1065,1154,196,222]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="47">NMSA</collectionCode>
21939).
<emphasis id="A9A01B34A320DB37FB31EFE8FEF2FEE5" bold="true" pageId="11" pageNumber="47">Mpumalanga Province:</emphasis>
1 3, 1 Ƥ, Barberton, Mpumalanga [25
<geoCoordinate id="FEE0A1E1A320DB37FD33EFC7FCD2FEEA" box="[754,818,235,264]" direction="south" orientation="latitude" pageId="11" pageNumber="47" precision="925" value="-0.78333336">0 47'S</geoCoordinate>
, 31
<geoCoordinate id="FEE0A1E1A320DB37FC9CEFC7FC41FEEA" box="[861,929,235,264]" direction="east" orientation="longitude" pageId="11" pageNumber="47" precision="925" value="0.05">0 03'E</geoCoordinate>
],
<date id="EF6AE1E6A320DB37FC76EFC2FBFFFEEA" box="[951,1055,238,264]" pageId="11" pageNumber="47" value="1909-12">xii.1909</date>
, Miss de
<collectingCountry id="E3C387B6A320DB37FB55EFC2FB2DFEEA" box="[1172,1229,238,264]" name="Senegal" pageId="11" pageNumber="47">Beer</collectingCountry>
(
<collectionCode id="FDC55FE3A320DB37FB1FEFC2FAD7FEEA" LSID="urn:lsid:biocol.org:col:34985" box="[1246,1335,238,264]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="47">NMSA</collectionCode>
18990).
<emphasis id="A9A01B34A320DB37FF56EE3BFE56FED3" bold="true" box="[151,438,279,305]" pageId="11" pageNumber="47">North West Province:</emphasis>
2 3, Barakolalo [25
<geoCoordinate id="FEE0A1E1A320DB37FD7AEE38FD1CFED3" box="[699,764,276,305]" direction="south" orientation="latitude" pageId="11" pageNumber="47" precision="925" value="-0.26666668">0 16'S</geoCoordinate>
, 25
<geoCoordinate id="FEE0A1E1A320DB37FCE8EE38FC8FFED3" box="[809,879,276,305]" direction="east" orientation="longitude" pageId="11" pageNumber="47" precision="925" value="0.9">0 54'E</geoCoordinate>
],
<date id="EF6AE1E6A320DB37FC44EE3BFC1EFED3" box="[901,1022,279,305]" pageId="11" pageNumber="47" value="2000-01-14">14.i.2000</date>
, M. Hamer &amp; R. Slotow (
<collectionCode id="FDC55FE3A320DB37FA83EE3BFA7CFED3" LSID="urn:lsid:biocol.org:col:34985" box="[1346,1436,279,305]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="47">NMSA</collectionCode>
21975).
<emphasis id="A9A01B34A320DB37FF39EE12FE3EFEBA" bold="true" box="[248,478,318,344]" pageId="11" pageNumber="47">Unknown locality:</emphasis>
1 3,
<date id="EF6AE1E6A320DB37FDE0EE12FD9AFEBA" box="[545,634,318,344]" pageId="11" pageNumber="47" value="1966-04">iv.1966</date>
, G. Lombard (
<collectionCode id="FDC55FE3A320DB37FCE0EE12FC98FEBA" LSID="urn:lsid:biocol.org:col:34985" box="[801,888,318,344]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="47">NMSA</collectionCode>
9609).
</paragraph>
</subSubSection>
<caption id="CFAB97AEA320DB37FF56EAA7FBB8F9E9" httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47" targetBox="[220,1366,390,1395]" targetPageId="11">
<paragraph id="9B6BC726A320DB37FF56EAA7FBB8F9E9" blockId="11.[151,1436,1419,1547]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF56EAA7FEF9FA41" bold="true" box="[151,281,1419,1443]" pageId="11" pageNumber="47">FIGURE 5.</emphasis>
<taxonomicName id="5CD4BCA5A320DB37FEDEEAA0FDE2FA41" box="[287,514,1420,1443]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A320DB37FEDEEAA0FDE2FA41" box="[287,514,1420,1443]" italics="true" pageId="11" pageNumber="47">Spirostreptus unciger</emphasis>
</taxonomicName>
(NMSA 8896).
<emphasis id="A9A01B34A320DB37FD71EAA7FD24FA41" bold="true" box="[688,708,1419,1443]" pageId="11" pageNumber="47">a,</emphasis>
oral view of gonopods;
<emphasis id="A9A01B34A320DB37FC04EAA7FC3BFA41" bold="true" box="[965,987,1419,1443]" pageId="11" pageNumber="47">b,</emphasis>
aboral view of gonopods;
<emphasis id="A9A01B34A320DB37FB34EAA7FAE8FA41" bold="true" box="[1269,1288,1419,1443]" pageId="11" pageNumber="47">c,</emphasis>
telopodite;
<emphasis id="A9A01B34A320DB37FA47EAA7FA7CFA41" bold="true" box="[1414,1436,1419,1443]" pageId="11" pageNumber="47">d,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="A9A01B34A320DB37FD19EA82FD0BFA24" bold="true" box="[728,747,1454,1478]" pageId="11" pageNumber="47">e,</emphasis>
lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe.
</paragraph>
</caption>
<subSubSection id="D3CE94ADA320DB37FF04E915FDC6F90C" pageId="11" pageNumber="47" type="diagnosis">
<paragraph id="9B6BC726A320DB37FF04E915FC8BF998" blockId="11.[151,1437,1593,2009]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E915FEADF9B1" bold="true" box="[197,333,1593,1619]" pageId="11" pageNumber="47">Diagnosis:</emphasis>
Lateral lobe very wide proximally, distal part lying tightly against lateral edge of apical proplica (
<figureCitation id="03EFDBA3A320DB37FEC6E94CFEADF998" box="[263,333,1632,1658]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47">Fig. 5</figureCitation>
a); telopodite with distal side branch (
<figureCitation id="03EFDBA3A320DB37FCC6E94CFCAEF998" box="[775,846,1632,1658]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47">Fig. 5</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A320DB37FF04E9ABFDEDF925" blockId="11.[151,1437,1593,2009]" pageId="11" pageNumber="47">
Description:
<emphasis id="A9A01B34A320DB37FEA3E9ABFE72F942" box="[354,402,1671,1696]" italics="true" pageId="11" pageNumber="47">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A320DB37FD81E9ABFD03F943" box="[576,739,1671,1697]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="11" pageNumber="47" unit="mm" value="240.0" valueMax="250.0" valueMin="230.0">230250 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A320DB37FB72E9ABFAF6F943" box="[1203,1302,1671,1697]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="47" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
and
<quantity id="5C2C6AC3A320DB37FA92E9ABFF28F925" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="11" pageNumber="47" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">1115 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A320DB37FE46E981FDE9F925" box="[391,521,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="11" pageNumber="47" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A320DB37FF04E9F8FDC6F90C" blockId="11.[151,1437,1593,2009]" box="[197,550,1748,1774]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E9F8FE25F90F" box="[197,453,1748,1773]" italics="true" pageId="11" pageNumber="47">Number of body rings</emphasis>
: 6773.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA320DB30FF04E9D7FD3CFD60" lastPageId="12" lastPageNumber="48" pageId="11" pageNumber="47" type="description">
<paragraph id="9B6BC726A320DB37FF04E9D7FBEBF8F7" blockId="11.[151,1437,1593,2009]" box="[197,1035,1787,1813]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E9D7FEF8F8F6" box="[197,280,1787,1812]" italics="true" pageId="11" pageNumber="47">Colour</emphasis>
: Body black or brownish; legs brown; antennae black or brown.
</paragraph>
<paragraph id="9B6BC726A320DB37FF04E80EFEDEF880" blockId="11.[151,1437,1593,2009]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E80EFEFDF8D9" box="[197,285,1826,1851]" italics="true" pageId="11" pageNumber="47">Collum</emphasis>
: Anterior produced laterally into a narrow, rounded lobe, with 3 complete and 12 incomplete folds (
<figureCitation id="03EFDBA3A320DB37FF20E864FEC6F880" box="[225,294,1864,1890]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47">Fig. 5</figureCitation>
e)
</paragraph>
<paragraph id="9B6BC726A320DB37FF04E85EFE3EF850" blockId="11.[151,1437,1593,2009]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E85EFD03F869" box="[197,739,1903,1931]" italics="true" pageId="11" pageNumber="47">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally U-shaped with short, apical nipple slightly deflected laterally (
<figureCitation id="03EFDBA3A320DB37FEBBE8B4FE5FF850" box="[378,447,1944,1970]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47">Fig. 5</figureCitation>
d).
</paragraph>
<paragraph id="9B6BC726A320DB37FF04E893FAC6F83B" blockId="11.[151,1437,1593,2009]" box="[197,1318,1983,2009]" pageId="11" pageNumber="47">
<emphasis id="A9A01B34A320DB37FF04E893FED2F83A" box="[197,306,1983,2008]" italics="true" pageId="11" pageNumber="47">Gonopod</emphasis>
: (
<figureCitation id="03EFDBA3A320DB37FE88E893FE74F83B" box="[329,404,1983,2009]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="11" pageNumber="47">Figs 5</figureCitation>
a, 5b)
<quantity id="5C2C6AC3A320DB37FE1CE893FDDFF83B" box="[477,575,1983,2009]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="47" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long. Sternite apex extends distally to level of paracoxite apex.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EFB4FDE8FF1D" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">Proplica tapering apically, wider at the level of telopodite spiral. Proplica not overlapping the distal lobe, resulting in much of proximal lateral lobe being visible orally; lateral edge of proximal half of proplica raised more sharply than medial edge.</paragraph>
<paragraph id="9B6BC726A327DB30FF04EE20FDC5FE91" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">Distally, metaplica with a rounded hump and nipple-shaped apical extension proximal to distal lobe apex; metaplica width approximately constant along the length except at the level of lateral lobe, a deep depression present at the level of lateral lobe.</paragraph>
<paragraph id="9B6BC726A327DB30FF04EEACFD4FFE05" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">
Free lateral lobe short and finger-like distally. Proximally lateral lobe broad and vertical. Lateral lobe lying tightly against proplica, does not extend beyond lateral edges of proplica or distal lobe (
<figureCitation id="03EFDBA3A327DB30FB17EE8AFAFBFE22" box="[1238,1307,422,448]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="12" pageNumber="48">Fig. 5</figureCitation>
a). Distally lateral lobe widely separated from distal lobe.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EED8FD24FDD6" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">Telocoxites parallel, slightly wider apart distal to sternite apex, medial edge straight, distal lobe not extending beyond lateral edge of proplica apex.</paragraph>
<paragraph id="9B6BC726A327DB30FF04ED6DFD3CFD60" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">
Post knee telopodite with a side branch distal to coil (
<figureCitation id="03EFDBA3A327DB30FC8DED6DFC75FDB9" box="[844,917,577,603]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="12" pageNumber="48">Fig. 5</figureCitation>
c); antetorsal process of telopodite straight, extending past the origin of side branch (
<figureCitation id="03EFDBA3A327DB30FDB6ED44FD5DFD60" box="[631,701,616,642]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="12" pageNumber="48">Fig. 7</figureCitation>
d).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA327DB30FF04EDA2FBAAFD4A" box="[197,1098,654,680]" pageId="12" pageNumber="48" type="distribution">
<paragraph id="9B6BC726A327DB30FF04EDA2FBAAFD4A" blockId="12.[151,1437,152,951]" box="[197,1098,654,680]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EDA2FE87FD4A" bold="true" box="[197,359,654,680]" pageId="12" pageNumber="48">Distribution:</emphasis>
Known only from north-western and north-eastern
<collectingCountry id="E3C387B6A327DB30FC76EDA2FBAAFD4A" box="[951,1098,654,680]" name="South Africa" pageId="12" pageNumber="48">South Africa</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA327DB30FF04ED99FDB0FC55" pageId="12" pageNumber="48" type="discussion">
<paragraph id="9B6BC726A327DB30FF04ED99FD09FC88" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04ED99FEA1FD2D" bold="true" box="[197,321,693,719]" pageId="12" pageNumber="48">Remarks:</emphasis>
The broad and vertical proximal part of the lateral lobe and the side branch on the telopodite distinguish
<taxonomicName id="5CD4BCA5A327DB30FEE6EDF0FE65FD17" box="[295,389,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A327DB30FEE6EDF0FE65FD17" box="[295,389,732,757]" italics="true" pageId="12" pageNumber="48">unciger</emphasis>
</taxonomicName>
from congeners (
<figureCitation id="03EFDBA3A327DB30FDA1EDF0FD50FD14" box="[608,688,732,758]" captionStart="FIGURE 5" captionStartId="11.[151,255,1419,1443]" captionTargetBox="[220,1366,390,1395]" captionTargetId="figure@11.[216,1373,375,1412]" captionTargetPageId="11" captionText="FIGURE 5. Spirostreptus unciger (NMSA 8896). a, oral view of gonopods; b, aboral view of gonopods; c, telopodite; d, oral view of prefemoral lobe of first pair of male legs; e, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mn, metaplica nipple; dl, distal lobe; d, depression; tp, telopodite; tpb, telopodite branch; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189863/files/figure.png" pageId="12" pageNumber="48">Figs 5</figureCitation>
c, 7d). The distal lobes of the gonopods are as in congeners, however, the shape and surface texture suggest closer affinities with
<taxonomicName id="5CD4BCA5A327DB30FC24EC2EFBC7FCF9" box="[997,1063,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A327DB30FC24EC2EFBC7FCF9" box="[997,1063,770,795]" italics="true" pageId="12" pageNumber="48">heros</emphasis>
</taxonomicName>
,
<taxonomicName id="5CD4BCA5A327DB30FBF9EC2EFB58FCF9" box="[1080,1208,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="batokensis">
<emphasis id="A9A01B34A327DB30FBF9EC2EFB58FCF9" box="[1080,1208,770,795]" italics="true" pageId="12" pageNumber="48">batokensis</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A327DB30FB37EC2EFABDFCF9" box="[1270,1373,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A327DB30FB37EC2EFABDFCF9" box="[1270,1373,770,795]" italics="true" pageId="12" pageNumber="48">kruegeri</emphasis>
</taxonomicName>
. The position of the lateral lobe in relation to the proplica and distal lobe, and the wide separation of the apical proplica from the distal lobe are unique to
<taxonomicName id="5CD4BCA5A327DB30FD4BEC7CFD02FC8B" box="[650,738,848,873]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A327DB30FD4BEC7CFD02FC8B" box="[650,738,848,873]" italics="true" pageId="12" pageNumber="48">unciger</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9B6BC726A327DB30FF07EC5AFDB0FC55" blockId="12.[151,1437,152,951]" pageId="12" pageNumber="48">
The
<typeStatus id="446F7984A327DB30FF3AEC5AFECDFC72" box="[251,301,886,912]" pageId="12" pageNumber="48">type</typeStatus>
material of
<taxonomicName id="5CD4BCA5A327DB30FE7DEC5BFDD2FC72" box="[444,562,887,912]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A327DB30FE7DEC5BFDD2FC72" box="[444,562,887,912]" italics="true" pageId="12" pageNumber="48">S. unciger</emphasis>
</taxonomicName>
is supposed to be in the SAM but is missing from this collection and from other institutions in southern African.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,252 @@
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<mods:title id="84CE61E6A913DBF4CE235D594BC82105">A revision of the taxonomy and distribution of Spirostreptus Brandt 1833 (Diplopoda, Spirostreptida, Spirostreptidae) with descriptions of a new species and a new genus of spirostreptid millipede</mods:title>
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<mods:namePart id="9C262EEA21E28029A9478A46F0834E45">Barraclough, David</mods:namePart>
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<treatment id="137D7630A322DB36FF56EC63FBBAFE78" ID-DOI="http://doi.org/10.5281/zenodo.6220854" ID-GBIF-Taxon="119623023" ID-Zenodo-Dep="6220854" LSID="urn:lsid:plazi:treatment:137D7630A322DB36FF56EC63FBBAFE78" httpUri="http://treatment.plazi.org/id/137D7630A322DB36FF56EC63FBBAFE78" lastPageId="10" lastPageNumber="46" pageId="9" pageNumber="45">
<subSubSection id="D3CE94ADA322DB35FF56EC63FEE6FC6E" pageId="9" pageNumber="45" type="nomenclature">
<paragraph id="9B6BC726A322DB35FF56EC63FD21FC8B" blockId="9.[151,705,847,908]" box="[151,705,847,873]" pageId="9" pageNumber="45">
<heading id="C023704AA322DB35FF56EC63FD21FC8B" bold="true" box="[151,705,847,873]" fontSize="11" level="1" pageId="9" pageNumber="45" reason="1">
<taxonomicName id="5CD4BCA5A322DB35FF56EC63FD21FC8B" ID-CoL="4Z8XX" authority="Cook &amp; Collins 1893" authorityName="Cook &amp; Collins" authorityYear="1893" box="[151,705,847,873]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FF56EC63FD21FC8B" bold="true" box="[151,705,847,873]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF56EC63FE56FC8B" bold="true" box="[151,438,847,873]" italics="true" pageId="9" pageNumber="45">Spirostreptus tripartitus</emphasis>
<bibRefCitation id="FF45BAD7A322DB35FE7CEC63FD21FC8B" author="Cook" box="[445,705,847,873]" pageId="9" pageNumber="55" refString="Cook, O. F. &amp; Collins, G. N. (1893) The Myriapoda collected by the U. S. Eclipse Expedition to West Africa, 1889 and 1890. Annals of the New York Academy of Science, 8, 24 - 40." type="journal article" year="1893">Cook &amp; Collins 1893</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B6BC726A322DB35FF56EC5EFEE6FC6E" blockId="9.[151,705,847,908]" box="[151,262,882,908]" pageId="9" pageNumber="45">
<figureCitation id="03EFDBA3A322DB35FF56EC5EFF02FC6E" box="[151,226,882,908]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Figs 4</figureCitation>
,
<figureCitation id="03EFDBA3A322DB35FF2FEC5EFF1BFC6E" box="[238,251,882,908]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="9" pageNumber="45">7</figureCitation>
f
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF56ECECFC69FC15" pageId="9" pageNumber="45" type="reference_group">
<paragraph id="9B6BC726A322DB35FF56ECECFC69FC15" blockId="9.[151,1004,960,1015]" pageId="9" pageNumber="45">
<taxonomicName id="5CD4BCA5A322DB35FF56ECECFD64FC35" ID-CoL="4Z8XX" authority="Cook &amp; Collins 1893" authorityName="Cook &amp; Collins" authorityYear="1893" box="[151,644,960,983]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FF56ECECFE76FC35" box="[151,406,960,983]" italics="true" pageId="9" pageNumber="45">Spirostreptus tripartitus</emphasis>
<bibRefCitation id="FF45BAD7A322DB35FE5DECECFD64FC35" author="Cook" box="[412,644,960,983]" pageId="9" pageNumber="55" refString="Cook, O. F. &amp; Collins, G. N. (1893) The Myriapoda collected by the U. S. Eclipse Expedition to West Africa, 1889 and 1890. Annals of the New York Academy of Science, 8, 24 - 40." type="journal article" year="1893">Cook &amp; Collins 1893</bibRefCitation>
</taxonomicName>
, p. 31, figs 1722;
<bibRefCitation id="FF45BAD7A322DB35FC90ECECFC0CFC35" author="Hoffman" box="[849,1004,960,983]" pageId="9" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
<taxonomicName id="5CD4BCA5A322DB35FF56ECCCFDAAFC15" ID-CoL="5857C" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,586,992,1015]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FF56ECCCFE50FC15" box="[151,432,992,1015]" italics="true" pageId="9" pageNumber="45">Triaenostreptus tripartitus</emphasis>
:
<bibRefCitation id="FF45BAD7A322DB35FE7FECCCFDAAFC15" author="Attems" box="[446,586,992,1015]" pageId="9" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="FF45BAD7A322DB35FD94ECCCFD13FC15" author="Hoffman" box="[597,755,992,1015]" pageId="9" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A322DB35FD3EECCCFC69FC15" author="Krabbe" box="[767,905,992,1015]" pageId="9" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF56EB0AFEA5FB73" pageId="9" pageNumber="45" type="materials_examined">
<paragraph id="9B6BC726A322DB35FF56EB0AFCCEFBA2" blockId="9.[151,1437,1062,2022]" box="[151,814,1062,1088]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF56EB0AFEADFBA2" bold="true" box="[151,333,1062,1088]" pageId="9" pageNumber="45">
<typeStatus id="446F7984A322DB35FF56EB0AFF32FBA2" box="[151,210,1062,1088]" pageId="9" pageNumber="45">Type</typeStatus>
material:
</emphasis>
<typeStatus id="446F7984A322DB35FE97EB0AFE26FBA2" box="[342,454,1062,1088]" pageId="9" pageNumber="45" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="E3C387B6A322DB35FE10EB0AFDDEFBA2" box="[465,574,1062,1088]" name="Republic of the Congo" pageId="9" pageNumber="45">CONGO</collectingCountry>
: 2 3 (
<collectionCode id="FDC55FE3A322DB35FD45EB0AFD3BFBA2" LSID="urn:lsid:biocol.org:col:34871" box="[644,731,1062,1088]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="45">USNM</collectionCode>
2384).
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EB7CFEA5FB73" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EB7CFDA6FB88" bold="true" box="[197,582,1104,1130]" pageId="9" pageNumber="45">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A322DB35FD8FEB7CFD2FFB88" box="[590,719,1104,1130]" name="Angola" pageId="9" pageNumber="45">ANGOLA</collectingCountry>
: 1 3,
<collectingCountry id="E3C387B6A322DB35FCD7EB7CFCB5FB88" box="[790,853,1104,1130]" name="Egypt" pageId="9" pageNumber="45">Egito</collectingCountry>
near Benguela [12
<geoCoordinate id="FEE0A1E1A322DB35FBF7EB61FB93FB88" box="[1078,1139,1101,1130]" direction="south" orientation="latitude" pageId="9" pageNumber="45" precision="925" value="-0.55">0 33'S</geoCoordinate>
, 13
<geoCoordinate id="FEE0A1E1A322DB35FB5FEB61FB3FFB88" box="[1182,1247,1101,1130]" direction="east" orientation="longitude" pageId="9" pageNumber="45" precision="925" value="0.4">0 24'E</geoCoordinate>
], 1975, (
<collectionCode id="FDC55FE3A322DB35FA84EB7CFA7CFB88" box="[1349,1436,1104,1130]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="45">ZMUC</collectionCode>
uncatalogued).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF04EBB1FD04FB3C" pageId="9" pageNumber="45" type="diagnosis">
<paragraph id="9B6BC726A322DB35FF04EBB1FD04FB3C" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EBB1FEA7FB55" bold="true" box="[197,327,1181,1207]" pageId="9" pageNumber="45">Diagnosis:</emphasis>
Gonopods with angular distal lobe, and with oral surface with folds and a prominent diagonal groove; distal lobe length equal to width (
<figureCitation id="03EFDBA3A322DB35FD41EBE8FD28FB3C" box="[640,712,1220,1246]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Fig. 4</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF04EBC6FA89F87B" pageId="9" pageNumber="45" type="description">
<paragraph id="9B6BC726A322DB35FF04EBC6FA68FAE6" blockId="9.[151,1437,1062,2022]" box="[197,1416,1258,1284]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EBC6FEBFFAE6" bold="true" box="[197,351,1258,1284]" pageId="9" pageNumber="45">Description:</emphasis>
<emphasis id="A9A01B34A322DB35FEAFEBC7FE7DFAE6" box="[366,413,1259,1284]" italics="true" pageId="9" pageNumber="45">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A322DB35FD85EBC6FD48FAE6" box="[580,680,1258,1284]" metricMagnitude="-1" metricUnit="m" metricValue="1.76" pageId="9" pageNumber="45" unit="mm" value="176.0">176 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A322DB35FBA8EBC6FB2BFAE6" box="[1129,1227,1258,1284]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="9" pageNumber="45" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
and
<quantity id="5C2C6AC3A322DB35FAC2EBC6FA64FAE6" box="[1283,1412,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="9" pageNumber="45" unit="mm" value="12.5" valueMax="14.0" valueMin="11.0">1114 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EA3DFDC6FAC9" blockId="9.[151,1437,1062,2022]" box="[197,550,1297,1323]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EA3DFE25FAC8" box="[197,453,1297,1322]" italics="true" pageId="9" pageNumber="45">Number of body rings</emphasis>
: 6672.
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EA14FC68FAB0" blockId="9.[151,1437,1062,2022]" box="[197,904,1336,1362]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EA14FEF8FAB3" box="[197,280,1336,1361]" italics="true" pageId="9" pageNumber="45">Colour</emphasis>
: Body black or brown; legs and antennae red-brown.
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EA73FE25FA7D" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EA73FEC2FA9A" box="[197,290,1375,1400]" italics="true" pageId="9" pageNumber="45">Collum</emphasis>
: With narrow anterior lobe slightly produced ventro-laterally, 34 complete folds and an incomplete fold (
<figureCitation id="03EFDBA3A322DB35FE9EEAA9FE46FA7D" box="[351,422,1413,1439]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Fig. 4</figureCitation>
d).
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EA83FEB3FA12" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EA83FD2AFA2A" box="[197,714,1452,1480]" italics="true" pageId="9" pageNumber="45">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally rounded, with a long slightly laterally deflected nipple (
<figureCitation id="03EFDBA3A322DB35FF31EAFAFED8FA12" box="[240,312,1494,1520]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Fig. 4</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EAD0FB1BF9F4" blockId="9.[151,1437,1062,2022]" box="[197,1275,1532,1558]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04EAD0FED2F9F7" box="[197,306,1532,1557]" italics="true" pageId="9" pageNumber="45">Gonopod</emphasis>
: (
<figureCitation id="03EFDBA3A322DB35FE88EAD0FE75F9F4" box="[329,405,1532,1558]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Figs 4</figureCitation>
a, 4b)
<quantity id="5C2C6AC3A322DB35FE1CEAD0FDC1F9F4" box="[477,545,1532,1558]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="45" unit="mm" value="6.0">6 mm</quantity>
long. Sternite short, not reaching the level of paracoxite apex.
</paragraph>
<paragraph id="9B6BC726A322DB35FF04E90FFAAAF9DF" blockId="9.[151,1437,1062,2022]" box="[197,1354,1571,1597]" pageId="9" pageNumber="45">Telocoxites parallel, not touching, wider apart distal to sternite apex (at the level of proplica apex).</paragraph>
<paragraph id="9B6BC726A322DB35FF04E966FD18F986" blockId="9.[151,1437,1062,2022]" box="[197,760,1610,1636]" pageId="9" pageNumber="45">
Proplica wide and angular subapically (
<figureCitation id="03EFDBA3A322DB35FD55E966FD3CF986" box="[660,732,1610,1636]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Fig. 4</figureCitation>
a).
</paragraph>
<paragraph id="9B6BC726A322DB35FF04E95CFC2CF953" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">Metaplica wider proximal to telopodite knee, distally metaplica with small horizontal hump, raised medial edge and a small central, conical apex giving a two prong appearance.</paragraph>
<paragraph id="9B6BC726A322DB35FF04E992FBA4F91C" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">Lateral lobe tightly fused to distal lobe and apical proplica, folded and rounded distally. Lateral lobe extending slightly beyond the lateral edges of subapical proplica and distal lobe.</paragraph>
<paragraph id="9B6BC726A322DB35FF04E827FA89F8AE" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
Distal lobe extends to or beyond the lateral edge of proplica apex, lobe as long as it is wide. Orally, distal lobe surface with folds and a deep diagonal groove (
<figureCitation id="03EFDBA3A322DB35FD3DE81EFCA4F8AE" box="[764,836,1842,1868]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Fig. 4</figureCitation>
a). Distal lobe overlaps lateral lobe proximally.
</paragraph>
<paragraph id="9B6BC726A322DB35FF04E874FA89F87B" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
Telopodite with crescent-shaped antetorsal process which does not extend to or past paracoxite apex, strongly bent medially but not crossing metaplica (
<figureCitation id="03EFDBA3A322DB35FD28E853FCD5F87B" box="[745,821,1919,1945]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="9" pageNumber="45">Figs 4</figureCitation>
a, 4b, 7f); post knee telopodite with loose spiral.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF04E889FF0BF804" pageId="9" pageNumber="45" type="distribution">
<paragraph id="9B6BC726A322DB35FF04E889FF0BF804" blockId="9.[151,1437,1062,2022]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04E889FE8CF85D" bold="true" box="[197,364,1957,1983]" pageId="9" pageNumber="45">Distribution:</emphasis>
Benguela, west coast of
<collectingCountry id="E3C387B6A322DB35FD54E88AFD0CF822" box="[661,748,1958,1984]" name="Angola" pageId="9" pageNumber="45">Angola</collectingCountry>
and from an unknown locality in
<collectingCountry id="E3C387B6A322DB35FB41E88AFF07F804" name="Democratic Republic of the Congo" pageId="9" pageNumber="45">Democratic Republic of Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA321DB36FF04EFBBFBBAFE78" pageId="10" pageNumber="46" type="discussion">
<paragraph id="9B6BC726A321DB36FF04EFBBFBEFFEC4" blockId="10.[151,1437,151,410]" pageId="10" pageNumber="46">
<emphasis id="A9A01B34A321DB36FF04EFBBFEA2FF53" bold="true" box="[197,322,151,177]" pageId="10" pageNumber="46">Remarks:</emphasis>
Gonopod morphology suggests that
<taxonomicName id="5CD4BCA5A321DB36FD3BEFB4FC93FF53" box="[762,883,152,177]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A321DB36FD3BEFB4FC93FF53" box="[762,883,152,177]" italics="true" pageId="10" pageNumber="46">tripartitus</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="5CD4BCA5A321DB36FBCEEFB4FBB2FF53" box="[1039,1106,152,177]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A321DB36FBCEEFB4FBB2FF53" box="[1039,1106,152,177]" italics="true" pageId="10" pageNumber="46">sebae</emphasis>
</taxonomicName>
. Like
<taxonomicName id="5CD4BCA5A321DB36FB60EFB4FB04FF53" box="[1185,1252,152,177]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A321DB36FB60EFB4FB04FF53" box="[1185,1252,152,177]" italics="true" pageId="10" pageNumber="46">sebae</emphasis>
</taxonomicName>
, the distal lobe has surface folds, however,
<taxonomicName id="5CD4BCA5A321DB36FE1EEF93FDB6FF3A" box="[479,598,191,216]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A321DB36FE1EEF93FDB6FF3A" box="[479,598,191,216]" italics="true" pageId="10" pageNumber="46">tripartitus</emphasis>
</taxonomicName>
is distinguished by a prominent diagonal groove (
<figureCitation id="03EFDBA3A321DB36FB66EF92FB0DFF3A" box="[1191,1261,190,216]" captionStart="FIGURE 4" captionStartId="10.[151,255,1585,1609]" captionTargetBox="[158,1425,483,1546]" captionTargetId="figure@10.[151,1436,467,1561]" captionTargetPageId="10" captionText="FIGURE 4. Spirostreptus tripartitus (USNM 2384). a, oral view of gonopods (without left telopodite); b, aboral view of gonopods (without left telopodite); c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; mp, metaplica prong; dl, distal lobe; dlg, distal lobe groove; tp, telopodite; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189862/files/figure.png" pageId="10" pageNumber="46">Fig. 4</figureCitation>
a) on the distal lobe surface and the smaller size of the distal lobe. In
<taxonomicName id="5CD4BCA5A321DB36FCDEEFC9FC82FF1C" box="[799,866,229,254]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A321DB36FCDEEFC9FC82FF1C" box="[799,866,229,254]" italics="true" pageId="10" pageNumber="46">sebae</emphasis>
</taxonomicName>
the distal lobe is wider (lateral width) than it is long and the apex is horizontal, unlike in
<taxonomicName id="5CD4BCA5A321DB36FDBDEE20FD13FEC7" box="[636,755,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A321DB36FDBDEE20FD13FEC7" box="[636,755,268,293]" italics="true" pageId="10" pageNumber="46">tripartitus</emphasis>
</taxonomicName>
(see remarks on
<taxonomicName id="5CD4BCA5A321DB36FC7DEE20FC1FFEC7" box="[956,1023,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A321DB36FC7DEE20FC1FFEC7" box="[956,1023,268,293]" italics="true" pageId="10" pageNumber="46">sebae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9B6BC726A321DB36FF04EE1EFBBAFE78" blockId="10.[151,1437,151,410]" pageId="10" pageNumber="46">
According to the literature (
<bibRefCitation id="FF45BAD7A321DB36FDD5EE1EFD53FEAE" author="Krabbe" box="[532,691,306,332]" pageId="10" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
) the
<typeStatus id="446F7984A321DB36FD30EE1EFCB9FEAE" box="[753,857,306,332]" pageId="10" pageNumber="46" type="holotype">holotype</typeStatus>
of
<taxonomicName id="5CD4BCA5A321DB36FC42EE1EFBFCFEA9" box="[899,1052,306,331]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A321DB36FC42EE1EFBFCFEA9" box="[899,1052,306,331]" italics="true" pageId="10" pageNumber="46">S. tripartitus</emphasis>
</taxonomicName>
is in the
<collectingCountry id="E3C387B6A321DB36FB4DEE1EFACDFEAE" box="[1164,1325,306,332]" name="United States of America" pageId="10" pageNumber="46">United States</collectingCountry>
National Museum, Washington,
<collectingCountry id="E3C387B6A321DB36FE6EEE75FE0FFE91" box="[431,495,345,371]" name="United States of America" pageId="10" pageNumber="46">USA</collectingCountry>
. However, only a
<typeStatus id="446F7984A321DB36FD0DEE75FCCAFE91" box="[716,810,345,371]" pageId="10" pageNumber="46" type="syntype">syntype</typeStatus>
was made available for examination. Therefore the
<typeStatus id="446F7984A321DB36FF56EEACFF15FE78" box="[151,245,384,410]" pageId="10" pageNumber="46" type="syntype">syntype</typeStatus>
, material from
<collectingCountry id="E3C387B6A321DB36FE69EEACFE1EFE78" box="[424,510,384,410]" name="Angola" pageId="10" pageNumber="46">Angola</collectingCountry>
and the original description were used in the study.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,244 @@
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FBA1B8D78D93BFD4F224BCA6C57EF14D">journal article</mods:classification>
<mods:identifier id="5559088F11D136A42667EC8CEDB3A0B0" type="DOI">10.5281/zenodo.189858</mods:identifier>
<mods:identifier id="9D5DC3D3B36977520863D8E106E63251" type="GBIF-Dataset">048c2675-8e1e-4546-8f82-c1392523dbab</mods:identifier>
<mods:identifier id="7E6B1163A887F7CFEF694A0A1BD3AA23" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B95A76FEBB72DA717C45BB476A27F689" type="Zenodo-Dep">189858</mods:identifier>
</mods:mods>
<treatment id="137D7630A325DB2CFF56E94EFCEFFCA1" ID-DOI="http://doi.org/10.5281/zenodo.6220866" ID-GBIF-Taxon="119623019" ID-Zenodo-Dep="6220866" LSID="urn:lsid:plazi:treatment:137D7630A325DB2CFF56E94EFCEFFCA1" httpUri="http://treatment.plazi.org/id/137D7630A325DB2CFF56E94EFCEFFCA1" lastPageId="16" lastPageNumber="52" pageId="14" pageNumber="50">
<subSubSection id="D3CE94ADA325DB32FF56E94EFF3BF97D" pageId="14" pageNumber="50" type="nomenclature">
<paragraph id="9B6BC726A325DB32FF56E94EFD2EF99E" blockId="14.[151,718,1634,1695]" box="[151,718,1634,1660]" pageId="14" pageNumber="50">
<heading id="C023704AA325DB32FF56E94EFD2EF99E" bold="true" box="[151,718,1634,1660]" fontSize="11" level="1" pageId="14" pageNumber="50" reason="1">
<taxonomicName id="5CD4BCA5A325DB32FF56E94EFD2EF99E" ID-CoL="45LJS" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[151,718,1634,1660]" class="Diplopoda" family="Spirostreptidae" genus="Namibostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="50" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A325DB32FF56E94EFD2EF99E" bold="true" box="[151,718,1634,1660]" pageId="14" pageNumber="50">
<emphasis id="A9A01B34A325DB32FF56E94EFDF7F99E" bold="true" box="[151,535,1634,1660]" italics="true" pageId="14" pageNumber="50">Namibostreptus kymatorhabdus</emphasis>
(
<bibRefCitation id="FF45BAD7A325DB32FDE6E94EFD26F99E" author="Attems" box="[551,710,1634,1660]" pageId="14" pageNumber="55" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25 (65 / 66), 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B6BC726A325DB32FF56E9A9FF3BF97D" blockId="14.[151,718,1634,1695]" box="[151,219,1669,1695]" pageId="14" pageNumber="50">
<figureCitation id="03EFDBA3A325DB32FF56E9A9FF3BF97D" box="[151,219,1669,1695]" captionStart="FIGURE 8" captionStartId="15.[151,255,1222,1246]" captionTargetBox="[158,1426,221,1181]" captionTargetId="figure@15.[151,1436,194,1198]" captionTargetPageId="15" captionText="FIGURE 8. Namibostreptus kymatorhabdus (ZMB 2043). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; clb, convex lateral lobe; dlp, distal lateral process; ep, end process; tp, telopodite; at, antetorsal process; pc, paracoxite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189866/files/figure.png" pageId="14" pageNumber="50">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA325DB32FF56E9FEFBB3F8EB" pageId="14" pageNumber="50" type="reference_group">
<paragraph id="9B6BC726A325DB32FF56E9FEFC10F90B" blockId="14.[151,1107,1746,1801]" box="[151,1008,1746,1769]" pageId="14" pageNumber="50">
<taxonomicName id="5CD4BCA5A325DB32FF56E9FEFD80F90B" ID-CoL="6ZB43" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[151,608,1746,1769]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="50" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A325DB32FF56E9FEFE2EF90B" box="[151,462,1746,1769]" italics="true" pageId="14" pageNumber="50">Spirostreptus kymatorhabdus</emphasis>
<bibRefCitation id="FF45BAD7A325DB32FE15E9FEFD80F90B" author="Attems" box="[468,608,1746,1769]" pageId="14" pageNumber="55" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25 (65 / 66), 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
</taxonomicName>
, p. 151, figs 147149;
<bibRefCitation id="FF45BAD7A325DB32FC95E9FEFC10F90B" author="Hoffman" box="[852,1008,1746,1769]" pageId="14" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
</paragraph>
<paragraph id="9B6BC726A325DB32FF56E9DEFBB3F8EB" blockId="14.[151,1107,1746,1801]" box="[151,1107,1778,1801]" pageId="14" pageNumber="50">
<taxonomicName id="5CD4BCA5A325DB32FF56E9DEFD61F8EB" ID-CoL="58576" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,641,1778,1801]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="50" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A325DB32FF56E9DEFE08F8EB" box="[151,488,1778,1801]" italics="true" pageId="14" pageNumber="50">Triaenostreptus kymatorhabdus</emphasis>
:
<bibRefCitation id="FF45BAD7A325DB32FE37E9DEFD61F8EB" author="Attems" box="[502,641,1778,1801]" pageId="14" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="FF45BAD7A325DB32FD4CE9DEFCCBF8EB" author="Hoffman" box="[653,811,1778,1801]" pageId="14" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A325DB32FCF7E9DEFC24F8EB" author="Krabbe" box="[822,964,1778,1801]" pageId="14" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A325DB32FC0EE9DEFBB3F8EB" author="Hamer" box="[975,1107,1778,1801]" pageId="14" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA325DB32FF56E817FDB6F811" pageId="14" pageNumber="50" type="materials_examined">
<paragraph id="9B6BC726A325DB32FF56E817FF03F89E" blockId="14.[151,1436,1849,2035]" pageId="14" pageNumber="50">
<emphasis id="A9A01B34A325DB32FF56E817FEB7F8B7" bold="true" box="[151,343,1851,1877]" pageId="14" pageNumber="50">
<typeStatus id="446F7984A325DB32FF56E817FF36F8B7" box="[151,214,1851,1877]" pageId="14" pageNumber="50">Type</typeStatus>
material:
</emphasis>
<typeStatus id="446F7984A325DB32FEA1E810FE36F8B4" box="[352,470,1852,1878]" pageId="14" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="E3C387B6A325DB32FE23E810FD8BF8B4" box="[482,619,1852,1878]" name="Namibia" pageId="14" pageNumber="50">NAMIBIA</collectingCountry>
: 1 3, Walfishbay [22
<geoCoordinate id="FEE0A1E1A325DB32FCB0E815FC51F8B4" box="[881,945,1849,1878]" direction="south" orientation="latitude" pageId="14" pageNumber="50" precision="925" value="-0.6333333">0 38'S</geoCoordinate>
, 14
<geoCoordinate id="FEE0A1E1A325DB32FC1CE815FBC2F8B4" box="[989,1058,1849,1878]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="925" value="0.65">0 39'E</geoCoordinate>
], Colonoal-Gesellschaft (
<collectionCode id="FDC55FE3A325DB32FA98E810FA7CF8B4" box="[1369,1436,1852,1878]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="14" pageNumber="50">ZMB</collectionCode>
2043).
</paragraph>
<paragraph id="9B6BC726A325DB32FF04E8A4FDB6F811" blockId="14.[151,1436,1849,2035]" pageId="14" pageNumber="50">
<emphasis id="A9A01B34A325DB32FF04E8A4FDA4F840" bold="true" box="[197,580,1928,1954]" pageId="14" pageNumber="50">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A325DB32FD8AE8A5FD30F841" box="[587,720,1929,1955]" name="Namibia" pageId="14" pageNumber="50">NAMIBIA</collectingCountry>
: 1 3, Lubbert (
<collectionCode id="FDC55FE3A325DB32FCBEE8A5FC20F841" box="[895,960,1929,1955]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="14" pageNumber="50">ZMB</collectionCode>
5197); 1 3, Daan Viljoen Game Park
<quantity id="5C2C6AC3A325DB32FABEE8A5FF5DF82E" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="50" unit="km" value="30.0">30 km</quantity>
north of Windhoek [22
<geoCoordinate id="FEE0A1E1A325DB32FE0EE89CFDECF82E" box="[463,524,1968,1996]" direction="south" orientation="latitude" pageId="14" pageNumber="50" precision="925" value="-0.53333336">0 32'S</geoCoordinate>
, 17
<geoCoordinate id="FEE0A1E1A325DB32FDF7E89CFD96F82E" box="[566,630,1968,1996]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="925" value="0.68333334">0 41'E</geoCoordinate>
],
<date id="EF6AE1E6A325DB32FD4AE89EFCEBF82E" box="[651,779,1970,1996]" pageId="14" pageNumber="50" value="1978-02-24">24.ii. 1978</date>
,
<collectingCountry id="E3C387B6A325DB32FCD6E89EFCCFF82E" box="[791,815,1970,1996]" name="Iceland" pageId="14" pageNumber="50">O</collectingCountry>
. Lomholdt &amp; E. Wederkinch (
<collectionCode id="FDC55FE3A325DB32FB51E89EFB07F82E" box="[1168,1255,1970,1996]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="50">ZMUC</collectionCode>
uncatalogued).
<emphasis id="A9A01B34A325DB32FF07E8F5FEA8F811" bold="true" box="[198,328,2009,2035]" pageId="14" pageNumber="50">Diagnosis:</emphasis>
See generic diagnosis.
</paragraph>
</subSubSection>
<caption id="CFAB97AEA324DB33FF56EBEAFD52FAA7" httpUri="https://zenodo.org/record/189866/files/figure.png" pageId="15" pageNumber="51" targetBox="[158,1426,221,1181]" targetPageId="15">
<paragraph id="9B6BC726A324DB33FF56EBEAFD52FAA7" blockId="15.[151,1436,1222,1349]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF56EBEAFEFAFB3C" bold="true" box="[151,282,1222,1246]" pageId="15" pageNumber="51">FIGURE 8.</emphasis>
<taxonomicName id="5CD4BCA5A324DB33FEE1EBEBFD92FB3C" box="[288,626,1223,1246]" class="Diplopoda" family="Spirostreptidae" genus="Namibostreptus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="51" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A324DB33FEE1EBEBFD92FB3C" box="[288,626,1223,1246]" italics="true" pageId="15" pageNumber="51">Namibostreptus kymatorhabdus</emphasis>
</taxonomicName>
(ZMB 2043).
<emphasis id="A9A01B34A324DB33FCCFEBEAFCC3FB3C" bold="true" box="[782,803,1222,1246]" pageId="15" pageNumber="51">a,</emphasis>
oral view of gonopods;
<emphasis id="A9A01B34A324DB33FBE7EBEAFBDCFB3C" bold="true" box="[1062,1084,1222,1246]" pageId="15" pageNumber="51">b,</emphasis>
aboral view of gonopods;
<emphasis id="A9A01B34A324DB33FA98EBEAFA8CFB3C" bold="true" box="[1369,1388,1222,1246]" pageId="15" pageNumber="51">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="A9A01B34A324DB33FD69EBC5FD5EFAE3" bold="true" box="[680,702,1257,1281]" pageId="15" pageNumber="51">d,</emphasis>
lateral view of collum. Abbreviations: p, proplica; m, metaplica; clb, convex lateral lobe; dlp, distal lateral process; ep, end process; tp, telopodite; at, antetorsal process; pc, paracoxite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe.
</paragraph>
</caption>
<subSubSection id="D3CE94ADA324DB2CFF04EA5FFAD1FDEC" lastPageId="16" lastPageNumber="52" pageId="15" pageNumber="51" type="description">
<paragraph id="9B6BC726A324DB33FF04EA5FFE21FA56" blockId="15.[151,1437,1395,2005]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04EA5FFEBFFA6F" bold="true" box="[197,351,1395,1421]" pageId="15" pageNumber="51">Description:</emphasis>
<emphasis id="A9A01B34A324DB33FEA9EA58FE77FA6F" box="[360,407,1396,1421]" italics="true" pageId="15" pageNumber="51">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A324DB33FD81EA58FD45FA6C" box="[576,677,1396,1422]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="51" unit="mm" value="210.0">210 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A324DB33FBADEA58FB3BFA6C" box="[1132,1243,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="15" pageNumber="51" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
and
<quantity id="5C2C6AC3A324DB33FAD7EA58FA77FA6C" box="[1302,1431,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="15" pageNumber="51" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A324DB33FE8BEAB6FE5DFA56" box="[330,445,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="15" pageNumber="51" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04EAEDFDC6FA39" blockId="15.[151,1437,1395,2005]" box="[197,550,1473,1499]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04EAEDFE25FA38" box="[197,453,1473,1498]" italics="true" pageId="15" pageNumber="51">Number of body rings</emphasis>
: 6274.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04EAC4FC75F9E0" blockId="15.[151,1437,1395,2005]" box="[197,917,1512,1538]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04EAC4FEF8F9E3" box="[197,280,1512,1537]" italics="true" pageId="15" pageNumber="51">Colour</emphasis>
: Body black or dark brown; legs and antennae brown.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E923FD74F9AD" blockId="15.[151,1437,1395,2005]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04E923FEEEF9CA" box="[197,270,1551,1576]" italics="true" pageId="15" pageNumber="51">Head:</emphasis>
Regularly spaced striations present behind and between the lateral edges of eyes; gnathochilarium with a triangular median prebasilar sclerite.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E970FE83F97E" blockId="15.[151,1437,1395,2005]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04E970FEC2F997" box="[197,290,1628,1653]" italics="true" pageId="15" pageNumber="51">Collum</emphasis>
: Anteriorly produced into a short rounded lateral lobe, with 34 complete and 2 or more incomplete folds.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E980FDDAF924" blockId="15.[151,1437,1395,2005]" box="[197,570,1705,1734]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04E980FE8DF927" box="[197,365,1708,1733]" italics="true" pageId="15" pageNumber="51">First ozopore:</emphasis>
On 6th body ring.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E9FFFAD4F90F" blockId="15.[151,1437,1395,2005]" box="[197,1332,1747,1773]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04E9FFFE5DF90E" box="[197,445,1747,1772]" italics="true" pageId="15" pageNumber="51">Pre-femoral process:</emphasis>
Proximally rounded with a short laterally deflected medial nipple (
<figureCitation id="03EFDBA3A324DB33FB10E9FFFAF6F90F" box="[1233,1302,1747,1773]" captionStart="FIGURE 8" captionStartId="15.[151,255,1222,1246]" captionTargetBox="[158,1426,221,1181]" captionTargetId="figure@15.[151,1436,194,1198]" captionTargetPageId="15" captionText="FIGURE 8. Namibostreptus kymatorhabdus (ZMB 2043). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; clb, convex lateral lobe; dlp, distal lateral process; ep, end process; tp, telopodite; at, antetorsal process; pc, paracoxite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189866/files/figure.png" pageId="15" pageNumber="51">Fig. 8</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E9D6FCF2F883" blockId="15.[151,1437,1395,2005]" pageId="15" pageNumber="51">
<emphasis id="A9A01B34A324DB33FF04E9D6FED9F8F1" box="[197,313,1786,1811]" italics="true" pageId="15" pageNumber="51">Gonopod</emphasis>
s: (
<figureCitation id="03EFDBA3A324DB33FEA3E9D6FE53F8F6" box="[354,435,1786,1812]" captionStart="FIGURE 8" captionStartId="15.[151,255,1222,1246]" captionTargetBox="[158,1426,221,1181]" captionTargetId="figure@15.[151,1436,194,1198]" captionTargetPageId="15" captionText="FIGURE 8. Namibostreptus kymatorhabdus (ZMB 2043). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; clb, convex lateral lobe; dlp, distal lateral process; ep, end process; tp, telopodite; at, antetorsal process; pc, paracoxite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189866/files/figure.png" pageId="15" pageNumber="51">Figs 8</figureCitation>
a, 8b)
<quantity id="5C2C6AC3A324DB33FDC4E9D6FD8DF8F6" box="[517,621,1786,1812]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="15" pageNumber="51" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
long. Gonopod surface smooth, without ridges or humps; distal telocoxite with an end process. Orally medial edges of telocoxite parallel, raised and folded laterally, fold becomes broader distally, ending in a flat end process.
</paragraph>
<paragraph id="9B6BC726A324DB33FF04E841FE41F84C" blockId="15.[151,1437,1395,2005]" pageId="15" pageNumber="51">Sternite triangular, reaching the level of the paracoxite apex; paracoxite apically horizontal and tightly fused to the metaplica.</paragraph>
<paragraph id="9B6BC726A324DB33FF04E897FA7CF837" blockId="15.[151,1437,1395,2005]" box="[197,1436,1979,2005]" pageId="15" pageNumber="51">Proplica about half the length of gonopod; apical proplica broad with a concavity and acute medial tip.</paragraph>
<paragraph id="9B6BC726A33BDB2CFF56EFB4FE44FF1D" blockId="16.[151,1437,152,835]" pageId="16" pageNumber="52">Lateral and medial edges of proplica pronounced subapically, with shallow central groove, lateral edge more pronounced than medial edge. Orally distal depression separates proplica from the larger convex lateral lobe of the distal telocoxite.</paragraph>
<paragraph id="9B6BC726A33BDB2CFF04EE20FB3CFE78" blockId="16.[151,1437,152,835]" pageId="16" pageNumber="52">Metaplica with a broad, convex-shaped lateral lobe (wider than it is long and distal to proplica apex) which extends beyond lateral edge of proplica; a smaller distal lateral process parallel to paracoxite apex present proximal to end process. Metaplica with a window through which part of telopodite is visible. Wide depression present adjacent to the convex lobe, distal to the point of emergence of telopodite.</paragraph>
<paragraph id="9B6BC726A33BDB2CFF04EE8AFAD1FDEC" blockId="16.[151,1437,152,835]" pageId="16" pageNumber="52">
Telopodite long, with a trifurcate ending. Antetorsal process crescent-shaped and tapering (
<figureCitation id="03EFDBA3A33BDB2CFAC5EE8AFAB0FE22" box="[1284,1360,422,448]" captionStart="FIGURE 8" captionStartId="15.[151,255,1222,1246]" captionTargetBox="[158,1426,221,1181]" captionTargetId="figure@15.[151,1436,194,1198]" captionTargetPageId="15" captionText="FIGURE 8. Namibostreptus kymatorhabdus (ZMB 2043). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; clb, convex lateral lobe; dlp, distal lateral process; ep, end process; tp, telopodite; at, antetorsal process; pc, paracoxite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189866/files/figure.png" pageId="16" pageNumber="52">Figs 8</figureCitation>
a, 8b), extending past paracoxite apex. Distally, antetorsal process with a terminal spine, bending medially past lateral edge of proplica; post knee telopodite lamellate with a loose coil preceding abrupt narrowing.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA33BDB2CFF04ED36FB37FDD6" box="[197,1239,538,564]" pageId="16" pageNumber="52" type="distribution">
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<emphasis id="A9A01B34A33BDB2CFF04ED36FE87FDD6" bold="true" box="[197,359,538,564]" pageId="16" pageNumber="52">Distribution:</emphasis>
Known only from the west coast of
<collectingCountry id="E3C387B6A33BDB2CFCC8ED36FC8CFDD6" box="[777,876,538,564]" name="Namibia" pageId="16" pageNumber="52">Namibia</collectingCountry>
and near Windhoek (
<collectingCountry id="E3C387B6A33BDB2CFBA0ED36FB2BFDD6" box="[1121,1227,538,564]" name="Namibia" pageId="16" pageNumber="52">Namibia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA33BDB2CFF04ED6CFCEFFCA1" pageId="16" pageNumber="52" type="discussion">
<paragraph id="9B6BC726A33BDB2CFF04ED6CFCEFFCA1" blockId="16.[151,1437,152,835]" pageId="16" pageNumber="52">
<emphasis id="A9A01B34A33BDB2CFF04ED6CFEA2FDB8" bold="true" box="[197,322,576,602]" pageId="16" pageNumber="52">Remarks:</emphasis>
The lack of a lateral lobe and the lack of a laterally rounded distal lobe, the shorter proplica, and the abrupt narrowing of the telopodite (distal to the telopodite coil) are character states that do not support placement in
<taxonomicName id="5CD4BCA5A33BDB2CFEF8EDA3FE36FD4A" box="[313,470,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="A9A01B34A33BDB2CFEF8EDA3FE36FD4A" box="[313,470,655,680]" italics="true" pageId="16" pageNumber="52">Spirostreptus</emphasis>
</taxonomicName>
. Based on Hoffmans (2008) diagnosis of the Spirostreptini,
<taxonomicName id="5CD4BCA5A33BDB2CFB75EDA3FA8BFD4A" box="[1204,1387,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Namibostreptus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="52" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A33BDB2CFB75EDA3FA8BFD4A" box="[1204,1387,655,680]" italics="true" pageId="16" pageNumber="52">kymatorhabdus</emphasis>
</taxonomicName>
is a member of the tribe, the telopodite structure and the prostatic groove resemble member genera. However, the shape and size of the distal coxite of the gonopod of
<taxonomicName id="5CD4BCA5A33BDB2CFD3CEDF0FC51FD17" box="[765,945,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Namibostreptus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="52" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A33BDB2CFD3CEDF0FC51FD17" box="[765,945,732,757]" italics="true" pageId="16" pageNumber="52">kymatorhabdus</emphasis>
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differ from other Spirostreptini genera by having a unique convex lobe. Therefore, we erect
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<emphasis id="A9A01B34A33BDB2CFCC7EC2EFC27FCF9" box="[774,967,770,795]" italics="true" pageId="16" pageNumber="52">Namibostreptus</emphasis>
</taxonomicName>
to accommodate
<taxonomicName id="5CD4BCA5A33BDB2CFB62EC2EFABFFCF9" box="[1187,1375,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Namibostreptus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="52" phylum="Arthropoda" rank="species" species="kymatorhabdus">
<emphasis id="A9A01B34A33BDB2CFB62EC2EFABFFCF9" box="[1187,1375,770,795]" italics="true" pageId="16" pageNumber="52">kymatorhabdus</emphasis>
</taxonomicName>
, this bringing the number of genera in Spirostreptini to six.
</paragraph>
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</document>

View file

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<mods:title id="9675ADB640D988509929AECB9E3B64FB">A revision of the taxonomy and distribution of Spirostreptus Brandt 1833 (Diplopoda, Spirostreptida, Spirostreptidae) with descriptions of a new species and a new genus of spirostreptid millipede</mods:title>
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species:
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<emphasis id="A9A01B34A326DB31FF56E81BFEC8F8B3" bold="true" box="[151,296,1847,1873]" pageId="13" pageNumber="49">Etymology:</emphasis>
Named for
<collectingCountry id="E3C387B6A326DB31FE74E814FDFDF8B0" box="[437,541,1848,1874]" name="Namibia" pageId="13" pageNumber="49">Namibia</collectingCountry>
, the country of provenance of the
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Proplica length approximately half that of gonopod (
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a). Apical proplica with an acute medial tip; proplica wide, with a concavity apically; distal metaplica with large proximal convex lateral lobe, short lateral process and short laterally deflected end process. Convex lobe extending beyond the lateral edge of proplica.
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process of distal metaplica at approximately right angle with the short lateral process.
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<treatment id="137D7630A327DB31FF56EB3DFB27FEAE" ID-DOI="http://doi.org/10.5281/zenodo.6220860" ID-GBIF-Taxon="119623022" ID-Zenodo-Dep="6220860" LSID="urn:lsid:plazi:treatment:137D7630A327DB31FF56EB3DFB27FEAE" httpUri="http://treatment.plazi.org/id/137D7630A327DB31FF56EB3DFB27FEAE" lastPageId="13" lastPageNumber="49" pageId="12" pageNumber="48">
<subSubSection id="D3CE94ADA327DB30FF56EB3DFEE9FBAC" pageId="12" pageNumber="48" type="nomenclature">
<paragraph id="9B6BC726A327DB30FF56EB3DFD63FBC8" blockId="12.[151,643,1040,1102]" box="[151,643,1040,1067]" pageId="12" pageNumber="48">
<heading id="C023704AA327DB30FF56EB3DFD63FBC8" bold="true" box="[151,643,1040,1067]" fontSize="11" level="1" pageId="12" pageNumber="48" reason="1">
<emphasis id="A9A01B34A327DB30FF56EB3DFD63FBC8" bold="true" box="[151,643,1040,1067]" pageId="12" pageNumber="48">
<taxonomicName id="5CD4BCA5A327DB30FF56EB3DFDD5FBC8" ID-CoL="4Z8QL" authority="Mwabvu" authorityName="Mwabvu" box="[151,565,1040,1067]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="batokensis" status="sp. nov.">
<emphasis id="A9A01B34A327DB30FF56EB3DFE5DFBC9" bold="true" box="[151,445,1041,1067]" italics="true" pageId="12" pageNumber="48">Spirostreptus batokensis</emphasis>
Mwabvu
</taxonomicName>
,
<taxonomicNameLabel id="B293A64FA327DB30FD80EB3CFD63FBC8" box="[577,643,1040,1066]" pageId="12" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B6BC726A327DB30FF56EB18FEE9FBAC" blockId="12.[151,643,1040,1102]" box="[151,265,1076,1102]" pageId="12" pageNumber="48">
<figureCitation id="03EFDBA3A327DB30FF56EB18FF02FBAC" box="[151,226,1076,1102]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Figs 6</figureCitation>
,
<figureCitation id="03EFDBA3A327DB30FF2FEB18FF1CFBAC" box="[238,252,1076,1102]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="12" pageNumber="48">7</figureCitation>
e
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA327DB30FF56EBACFC2AFB20" pageId="12" pageNumber="48" type="materials_examined">
<paragraph id="9B6BC726A327DB30FF56EBACFA73FB79" blockId="12.[151,1437,1152,2033]" box="[151,1427,1152,1179]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF56EBACFEADFB78" bold="true" box="[151,333,1152,1178]" pageId="12" pageNumber="48">
<typeStatus id="446F7984A327DB30FF56EBACFF32FB78" box="[151,210,1152,1178]" pageId="12" pageNumber="48">Type</typeStatus>
material:
</emphasis>
<typeStatus id="446F7984A327DB30FE97EBADFE28FB79" box="[342,456,1153,1179]" pageId="12" pageNumber="48" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="E3C387B6A327DB30FE12EBADFD97FB79" box="[467,631,1153,1179]" name="Zimbabwe" pageId="12" pageNumber="48">ZIMBABWE</collectingCountry>
: 1 3, Batoka Gorge [1726C3],
<date id="EF6AE1E6A327DB30FC27EBADFB8EFB79" box="[998,1134,1153,1179]" pageId="12" pageNumber="48" value="1995-07-28">28.vii.1995</date>
. F. Nyathi (NMZ/D835).
</paragraph>
<paragraph id="9B6BC726A327DB30FF07EB84FC2AFB20" blockId="12.[151,1437,1152,2033]" box="[198,970,1192,1218]" pageId="12" pageNumber="48">
Etymology: Refers to the
<typeStatus id="446F7984A327DB30FE34EB84FDC7FB20" box="[501,551,1192,1218]" pageId="12" pageNumber="48">type</typeStatus>
locality, Batoka Gorge,
<collectingCountry id="E3C387B6A327DB30FC84EB84FC26FB20" box="[837,966,1192,1218]" name="Zimbabwe" pageId="12" pageNumber="48">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA327DB30FF04EBE2FD03FB0A" box="[197,739,1230,1256]" pageId="12" pageNumber="48" type="diagnosis">
<paragraph id="9B6BC726A327DB30FF04EBE2FD03FB0A" blockId="12.[151,1437,1152,2033]" box="[197,739,1230,1256]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EBE2FEA7FB0A" bold="true" box="[197,327,1230,1256]" pageId="12" pageNumber="48">Diagnosis:</emphasis>
Distal lobe subtriangular (
<figureCitation id="03EFDBA3A327DB30FD41EBE2FD26FB0A" box="[640,710,1230,1256]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA327DB30FF04EBD9FCC2F813" pageId="12" pageNumber="48" type="description">
<paragraph id="9B6BC726A327DB30FF04EBD9FEA0FAD4" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EBD9FEBFFAED" bold="true" box="[197,351,1269,1295]" pageId="12" pageNumber="48">Description:</emphasis>
<emphasis id="A9A01B34A327DB30FEA6EBD9FE76FAEC" box="[359,406,1269,1294]" italics="true" pageId="12" pageNumber="48">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A327DB30FDFDEBD9FD41FAED" box="[572,673,1269,1295]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="12" pageNumber="48" unit="mm" value="215.0">215 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A327DB30FBA3EBD9FB46FAED" box="[1122,1190,1269,1295]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="48" unit="mm" value="6.0">6 mm</quantity>
and
<quantity id="5C2C6AC3A327DB30FB1EEBD9FAD4FAED" box="[1247,1332,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="48" unit="mm" value="13.0">13 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A327DB30FF26EA30FEDCFAD4" box="[231,316,1308,1334]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="48" unit="mm" value="11.0">11 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EA6FFE1AFABE" blockId="12.[151,1437,1152,2033]" box="[197,506,1346,1372]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EA6FFE25FABE" box="[197,453,1347,1372]" italics="true" pageId="12" pageNumber="48">Number of body rings</emphasis>
: 60.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EA45FD3CFA61" blockId="12.[151,1437,1152,2033]" box="[197,732,1385,1411]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EA45FEF8FA60" box="[197,280,1385,1410]" italics="true" pageId="12" pageNumber="48">Colour</emphasis>
: Body black; legs and antennae black.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EABCFE30FA32" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EABCFEC1FA4B" box="[197,289,1424,1449]" italics="true" pageId="12" pageNumber="48">Collum</emphasis>
: Anterior slightly produced into a short rounded lateral lobe, with 4 complete folds and 3 incomplete folds (
<figureCitation id="03EFDBA3A327DB30FEAAEA9AFE52FA32" box="[363,434,1462,1488]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
d).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04EACCFDA4F9C3" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04EACCFD3BFA1B" box="[197,731,1501,1529]" italics="true" pageId="12" pageNumber="48">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally rounded, medial nipples distally narrow and strongly deflected laterally (
<figureCitation id="03EFDBA3A327DB30FE20E92BFDC8F9C3" box="[481,552,1543,1569]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E902FC80F9A5" blockId="12.[151,1437,1152,2033]" box="[197,864,1581,1607]" pageId="12" pageNumber="48">
<emphasis id="A9A01B34A327DB30FF04E902FED2F9A5" box="[197,306,1582,1607]" italics="true" pageId="12" pageNumber="48">Gonopod</emphasis>
:
<quantity id="5C2C6AC3A327DB30FE80E901FE65F9A5" box="[321,389,1581,1607]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="48" unit="mm" value="6.0">6 mm</quantity>
long; telocoxites parallel, but separated.
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E978FCAAF98C" blockId="12.[151,1437,1152,2033]" box="[197,842,1620,1646]" pageId="12" pageNumber="48">Sternite apex not reaching the level of paracoxite apex.</paragraph>
<paragraph id="9B6BC726A327DB30FF04E956FD37F959" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
Proplica wider subapically and tapering apically. Apically proplica concealing the proximal lateral lobe. Proplica not overlapping the distal lobe (
<figureCitation id="03EFDBA3A327DB30FDB5E98DFD5BF959" box="[628,699,1697,1723]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
a).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E9E4FC3CF900" blockId="12.[151,1437,1152,2033]" box="[197,988,1736,1762]" pageId="12" pageNumber="48">
Metaplica with humps, wider proximal to telopodite knee (
<figureCitation id="03EFDBA3A327DB30FCB7E9E4FC5DF900" box="[886,957,1736,1762]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
b).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E9C3FBD6F8CD" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
Lateral lobe slanting steeply, not extending beyond the lateral edge of the distal lobe or widest part of proplica. Aborally, lateral lobe proximally wide; rounded distally (
<figureCitation id="03EFDBA3A327DB30FC63E839FC0DF8CD" box="[930,1005,1813,1839]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Figs 6</figureCitation>
a, 6b).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E810FE00F841" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
Distal lobe rounded and narrow apically, proximally wider than apex (subtriangular), surface of distal telocoxite lobe smooth with a depression; proximal distal lobe width not extending beyond lateral edge of subapical proplica (
<figureCitation id="03EFDBA3A327DB30FEBCE8A5FE23F841" box="[381,451,1929,1955]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Fig. 6</figureCitation>
a).
</paragraph>
<paragraph id="9B6BC726A327DB30FF04E89CFCC2F813" blockId="12.[151,1437,1152,2033]" pageId="12" pageNumber="48">
Telopodite with short crescent-shaped antetorsal process; antetorsal process not reaching paracoxite apex; post knee telopodite with an open coil (
<figureCitation id="03EFDBA3A327DB30FDA5E8FBFD4FF813" box="[612,687,2007,2033]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="12" pageNumber="48">Figs 6</figureCitation>
a, 6b, 7e).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA326DB31FF04EFBBFB52FF50" box="[197,1202,151,178]" pageId="13" pageNumber="49" type="distribution">
<paragraph id="9B6BC726A326DB31FF04EFBBFB52FF50" blockId="13.[151,1437,151,332]" box="[197,1202,151,178]" pageId="13" pageNumber="49">
<emphasis id="A9A01B34A326DB31FF04EFBBFE87FF53" bold="true" box="[197,359,151,177]" pageId="13" pageNumber="49">Distribution:</emphasis>
Known only from
<typeStatus id="446F7984A326DB31FD82EFB4FD94FF50" box="[579,628,152,178]" pageId="13" pageNumber="49">type</typeStatus>
locality near Victoria Falls in western
<collectingCountry id="E3C387B6A326DB31FBEFEFB4FB4DFF50" box="[1070,1197,152,178]" name="Zimbabwe" pageId="13" pageNumber="49">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA326DB31FF04EF92FB27FEAE" pageId="13" pageNumber="49" type="discussion">
<paragraph id="9B6BC726A326DB31FF04EF92FB27FEAE" blockId="13.[151,1437,151,332]" pageId="13" pageNumber="49">
<emphasis id="A9A01B34A326DB31FF04EF92FEA6FF3A" bold="true" box="[197,326,190,216]" pageId="13" pageNumber="49">Remarks:</emphasis>
Like congeners this species has a laterally rounded distal lobe. Although the gonopod is smaller (approximately the same size as that of
<taxonomicName id="5CD4BCA5A326DB31FD32EFC9FCDAFF1C" box="[755,826,229,254]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A326DB31FD32EFC9FCDAFF1C" box="[755,826,229,254]" italics="true" pageId="13" pageNumber="49">sebae</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A326DB31FCB9EFC9FC18FF1C" box="[888,1016,229,254]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A326DB31FCB9EFC9FC18FF1C" box="[888,1016,229,254]" italics="true" pageId="13" pageNumber="49">tripartitus</emphasis>
</taxonomicName>
), the surface texture of the distal telocoxite lobe resembles more closely that of
<emphasis id="A9A01B34A326DB31FD0FEE20FC9BFEC7" box="[718,891,268,293]" italics="true" pageId="13" pageNumber="49">
<taxonomicName id="5CD4BCA5A326DB31FD0FEE20FCF3FEC7" box="[718,787,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="heros">heros</taxonomicName>
,
<taxonomicName id="5CD4BCA5A326DB31FCDEEE20FC9BFEC7" box="[799,891,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="unciger">unciger</taxonomicName>
</emphasis>
and
<taxonomicName id="5CD4BCA5A326DB31FC79EE20FBFEFEC7" box="[952,1054,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A326DB31FC79EE20FBFEFEC7" box="[952,1054,268,293]" italics="true" pageId="13" pageNumber="49">kruegeri</emphasis>
</taxonomicName>
than
<taxonomicName id="5CD4BCA5A326DB31FBA3EE20FB47FEC7" box="[1122,1191,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A326DB31FBA3EE20FB47FEC7" box="[1122,1191,268,293]" italics="true" pageId="13" pageNumber="49">sebae</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A326DB31FB22EE20FABEFEC7" box="[1251,1374,268,293]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A326DB31FB22EE20FABEFEC7" box="[1251,1374,268,293]" italics="true" pageId="13" pageNumber="49">tripartitus</emphasis>
</taxonomicName>
. The subtriangular shape of the distal lobe distinguishes
<taxonomicName id="5CD4BCA5A326DB31FD2FEE1EFC8AFEA9" box="[750,874,306,331]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="batokensis">
<emphasis id="A9A01B34A326DB31FD2FEE1EFC8AFEA9" box="[750,874,306,331]" italics="true" pageId="13" pageNumber="49">batokensis</emphasis>
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from the congeners (
<figureCitation id="03EFDBA3A326DB31FBA5EE1EFB4BFEAE" box="[1124,1195,306,332]" captionStart="FIGURE 6" captionStartId="13.[151,255,1457,1481]" captionTargetBox="[158,1428,384,1433]" captionTargetId="figure@13.[151,1436,358,1463]" captionTargetPageId="13" captionText="FIGURE 6. Spirostreptus batokensis (NMZ / D 585). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189864/files/figure.png" pageId="13" pageNumber="49">Fig. 6</figureCitation>
a).
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<mods:title id="9BD3EC54D93EF9A11C007389B771CB9E">A revision of the taxonomy and distribution of Spirostreptus Brandt 1833 (Diplopoda, Spirostreptida, Spirostreptidae) with descriptions of a new species and a new genus of spirostreptid millipede</mods:title>
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<treatment id="137D7630A328DB39FF56EA5CFAE0FC55" ID-DOI="http://doi.org/10.5281/zenodo.6220848" ID-GBIF-Taxon="119623021" ID-Zenodo-Dep="6220848" LSID="urn:lsid:plazi:treatment:137D7630A328DB39FF56EA5CFAE0FC55" httpUri="http://treatment.plazi.org/id/137D7630A328DB39FF56EA5CFAE0FC55" lastPageId="5" lastPageNumber="41" pageId="3" pageNumber="39">
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<paragraph id="9B6BC726A328DB3FFF56EA5CFDF0FA68" blockId="3.[151,528,1392,1452]" box="[151,528,1392,1418]" pageId="3" pageNumber="39">
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<taxonomicName id="5CD4BCA5A328DB3FFF56EA5CFDF0FA68" ID-CoL="4Z8T6" authority="Porat 1872" authorityName="Porat" authorityYear="1872" box="[151,528,1392,1418]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A328DB3FFF56EA5CFDF0FA68" bold="true" box="[151,528,1392,1418]" pageId="3" pageNumber="39">
<emphasis id="A9A01B34A328DB3FFF56EA5CFE61FA68" bold="true" box="[151,385,1392,1418]" italics="true" pageId="3" pageNumber="39">Spirostreptus heros</emphasis>
<bibRefCitation id="FF45BAD7A328DB3FFE49EA5CFDF0FA68" author="Porat" box="[392,528,1392,1418]" pageId="3" pageNumber="56" refString="Porat, C. O. von (1872) Myriapoda Africae australis. Ofversigt af Konglicher Vetenskaps-Akademies Forhandlingar, 29 (5), 1 - 46. Stockholm." type="journal article" year="1872">Porat 1872</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="9B6BC726A328DB3FFF56EABEFEE9FA4E" blockId="3.[151,528,1392,1452]" box="[151,265,1426,1452]" pageId="3" pageNumber="39">
<figureCitation id="03EFDBA3A328DB3FFF56EABEFF00FA4E" box="[151,224,1426,1452]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="3" pageNumber="39">Figs 1</figureCitation>
,
<figureCitation id="03EFDBA3A328DB3FFF2FEABEFF1CFA4E" box="[238,252,1426,1452]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="3" pageNumber="39">7</figureCitation>
a
</paragraph>
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<paragraph id="9B6BC726A328DB3FFF56EACCFD2DF9F5" blockId="3.[151,1314,1504,1591]" pageId="3" pageNumber="39">
<taxonomicName id="5CD4BCA5A328DB3FFF56EACCFE02FA15" ID-CoL="4Z8T6" authority="Porat 1872" authorityName="Porat" authorityYear="1872" box="[151,482,1504,1527]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A328DB3FFF56EACCFE85FA15" box="[151,357,1504,1527]" italics="true" pageId="3" pageNumber="39">Spirostreptus heros</emphasis>
<bibRefCitation id="FF45BAD7A328DB3FFEAAEACCFE02FA15" author="Porat" box="[363,482,1504,1527]" pageId="3" pageNumber="56" refString="Porat, C. O. von (1872) Myriapoda Africae australis. Ofversigt af Konglicher Vetenskaps-Akademies Forhandlingar, 29 (5), 1 - 46. Stockholm." type="journal article" year="1872">Porat 1872</bibRefCitation>
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, p. 29;
<bibRefCitation id="FF45BAD7A328DB3FFDEEEACCFCE9FA15" author="Hoffman" box="[559,777,1504,1527]" pageId="3" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">
Hoffman
<emphasis id="A9A01B34A328DB3FFD57EACCFD23FA15" box="[662,707,1504,1527]" italics="true" pageId="3" pageNumber="39">et al</emphasis>
. 2001
</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A328DB3FFCD2EACCFC5EFA15" author="Mwabvu" box="[787,958,1504,1527]" pageId="3" pageNumber="56" refString="Mwabvu, T. (2005 a) New records and localities of spirostreptid millipedes (Myriapoda: Diplopoda) of Zimbabwe with notes on nomenclatural changes and habitat. Arnoldia, Zimbabwe, 10 (25), 249 - 256." type="journal article" year="2005" yearSuffix="a">Mwabvu 2005a</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A328DB3FFC09EACCFB84FA15" author="Hoffman" box="[968,1124,1504,1527]" pageId="3" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
<taxonomicName id="5CD4BCA5A328DB3FFF56E92CFDE7F9F5" ID-CoL="4Z8XW" authority="Attems 1909" authorityName="Attems" authorityYear="1909" box="[151,519,1536,1559]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="triodus">
<emphasis id="A9A01B34A328DB3FFF56E92CFE96F9F5" box="[151,374,1536,1559]" italics="true" pageId="3" pageNumber="39">Spirostreptus triodus</emphasis>
<bibRefCitation id="FF45BAD7A328DB3FFEBDE92CFDE7F9F5" author="Attems" box="[380,519,1536,1559]" pageId="3" pageNumber="55" refString="Attems, C. (1909) Myriapoda. In: Schultze, L. (Ed.), Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905. Denkschriften der medizinischnaturwissenschaftlichen Gesellschaft zu Jena, 14, 1 - 52." type="journal article" year="1909">Attems 1909</bibRefCitation>
</taxonomicName>
, p. 46, figs 7679.
</paragraph>
<paragraph id="9B6BC726A328DB3FFF56E90CFAC2F9D5" blockId="3.[151,1314,1504,1591]" box="[151,1314,1568,1591]" pageId="3" pageNumber="39">
<taxonomicName id="5CD4BCA5A328DB3FFF56E90CFDC9F9D5" ID-CoL="5857B" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,553,1568,1591]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="triodus">
<emphasis id="A9A01B34A328DB3FFF56E90CFE70F9D5" box="[151,400,1568,1591]" italics="true" pageId="3" pageNumber="39">Triaenostreptus triodus</emphasis>
:
<bibRefCitation id="FF45BAD7A328DB3FFE5CE90CFDC9F9D5" author="Attems" box="[413,553,1568,1591]" pageId="3" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="FF45BAD7A328DB3FFDF5E90CFD32F9D5" author="Hoffman" box="[564,722,1568,1591]" pageId="3" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A328DB3FFD1CE90CFCF6F9D5" author="Hoffman" box="[733,790,1568,1591]" pageId="3" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">2008</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A328DB3FFCE0E90CFC4EF9D5" author="Krabbe" box="[801,942,1568,1591]" pageId="3" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A328DB3FFC78E90CFBA1F9D5" author="Hamer" box="[953,1089,1568,1591]" pageId="3" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A328DB3FFB8DE90CFAC2F9D5" author="Hoffman" box="[1100,1314,1568,1591]" pageId="3" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">
Hoffman
<emphasis id="A9A01B34A328DB3FFB73E90CFB00F9D5" box="[1202,1248,1568,1591]" italics="true" pageId="3" pageNumber="39">et al</emphasis>
. 2001
</bibRefCitation>
</paragraph>
</subSubSection>
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<typeStatus id="446F7984A328DB3FFF56E945FF32F961" box="[151,210,1641,1667]" pageId="3" pageNumber="39">Type</typeStatus>
material:
</emphasis>
<typeStatus id="446F7984A328DB3FFE97E946FE28F966" box="[342,456,1642,1668]" pageId="3" pageNumber="39" type="holotype">Holotype</typeStatus>
:
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: 1 3, Kalahari [22
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4968).
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<paragraph id="9B6BC726A328DB3FFF04E9BFFD46F845" blockId="3.[151,1437,1638,1959]" pageId="3" pageNumber="39">
<emphasis id="A9A01B34A328DB3FFF04E9BFFDB1F94F" bold="true" box="[197,593,1683,1709]" pageId="3" pageNumber="39">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A328DB3FFD9BE9BFFCE5F94F" box="[602,773,1683,1709]" name="Botswana" pageId="3" pageNumber="39">BOTSWANA</collectingCountry>
: 1 3, 1 Ƥ, Gemsbok National Park, Kalahari [22
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, 23
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],
<date id="EF6AE1E6A328DB3FFED1E991FE75F935" box="[272,405,1725,1751]" pageId="3" pageNumber="39" value="1970-04-20">20.iv.1970</date>
, B. Lamoral (
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16152); 1 3, 1 Ƥ, Kalahari [22
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, 23
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], 2005, (
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20570).
<collectingCountry id="E3C387B6A328DB3FFF3BE9CAFE36F8E2" box="[250,470,1766,1792]" name="South Africa" pageId="3" pageNumber="39">SOUTH AFRICA</collectingCountry>
:
<emphasis id="A9A01B34A328DB3FFE23E9CAFD15F8E2" bold="true" box="[482,757,1766,1792]" pageId="3" pageNumber="39">North West Province:</emphasis>
2 3, Malopo [25
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, 24
<geoCoordinate id="FEE0A1E1A328DB3FFBEEE9CFFB90F8E2" box="[1071,1136,1763,1792]" direction="east" orientation="longitude" pageId="3" pageNumber="39" precision="925" value="0.28333333">0 17'E</geoCoordinate>
],
<date id="EF6AE1E6A328DB3FFB47E9CAFB18F8E2" box="[1158,1272,1766,1792]" pageId="3" pageNumber="39" value="2001-01-13">13.i.2001</date>
, R. Slotow &amp; M. Hamer (
<collectionCode id="FDC55FE3A328DB3FFEDFE83CFE95F8C8" LSID="urn:lsid:biocol.org:col:34985" box="[286,373,1808,1834]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="39">NMSA</collectionCode>
20495, 20498).
<emphasis id="A9A01B34A328DB3FFDF5E83CFC90F8C8" bold="true" box="[564,880,1808,1834]" pageId="3" pageNumber="39">Northern Cape Province:</emphasis>
1 3, 1 Ƥ, Northern Cape [27.697 0S, 21.355 0E], (
<collectionCode id="FDC55FE3A328DB3FFF5EE816FF16F8B6" LSID="urn:lsid:biocol.org:col:34985" box="[159,246,1850,1876]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="39">NMSA</collectionCode>
20551); 1 3, Northern Cape [27.697 0S, 21.355 0E], (
<collectionCode id="FDC55FE3A328DB3FFC98E816FC50F8B6" LSID="urn:lsid:biocol.org:col:34985" box="[857,944,1850,1876]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="39">NMSA</collectionCode>
20495).
<collectingCountry id="E3C387B6A328DB3FFBD9E816FB7EF8B6" box="[1048,1182,1850,1876]" name="Namibia" pageId="3" pageNumber="39">NAMIBIA</collectingCountry>
: 2 3, 5 Ƥ, Okawango [18
<geoCoordinate id="FEE0A1E1A328DB3FFF00E84CFEE3F89F" box="[193,259,1888,1917]" direction="south" orientation="latitude" pageId="3" pageNumber="39" precision="925" value="-0.3">0 18'S</geoCoordinate>
, 19
<geoCoordinate id="FEE0A1E1A328DB3FFEF0E84CFE98F89F" box="[305,376,1888,1917]" direction="east" orientation="longitude" pageId="3" pageNumber="39" precision="925" value="0.25">0 15'E</geoCoordinate>
],
<date id="EF6AE1E6A328DB3FFE4EE84FFDFDF89F" box="[399,541,1891,1917]" pageId="3" pageNumber="39" value="2002-03-29">29.iii.2002</date>
, G. Plotner (
<collectionCode id="FDC55FE3A328DB3FFD7DE84FFCE1F89F" box="[700,769,1891,1917]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="39">ZMB</collectionCode>
13737, 13738, 13739 &amp; 13742).
<collectingCountry id="E3C387B6A328DB3FFB6AE84FFAB8F89F" box="[1195,1368,1891,1917]" name="Zimbabwe" pageId="3" pageNumber="39">ZIMBABWE</collectingCountry>
: 1 3, Beitbridge [22
<geoCoordinate id="FEE0A1E1A328DB3FFE80E8A6FE9EF845" box="[321,382,1930,1959]" direction="south" orientation="latitude" pageId="3" pageNumber="39" precision="925" value="-0.033333335">0 02'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A328DB3FFE69E8A6FE0DF845" box="[424,493,1930,1959]" direction="east" orientation="longitude" pageId="3" pageNumber="39" precision="925" value="0.0">0 00'E</geoCoordinate>
], (NMZ/D835).
</paragraph>
</subSubSection>
<caption id="CFAB97AEA32FDB38FF56EA17FDE8FA59" httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="4" pageNumber="40" targetBox="[151,1427,209,1300]" targetPageId="4">
<paragraph id="9B6BC726A32FDB38FF56EA17FDE8FA59" blockId="4.[151,1436,1339,1467]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF56EA17FEF9FAB1" bold="true" box="[151,281,1339,1363]" pageId="4" pageNumber="40">FIGURE 1.</emphasis>
<taxonomicName id="5CD4BCA5A32FDB38FEE0EA10FE0EFAB1" box="[289,494,1340,1363]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32FDB38FEE0EA10FE0EFAB1" box="[289,494,1340,1363]" italics="true" pageId="4" pageNumber="40">Spirostreptus heros</emphasis>
</taxonomicName>
(NMSA 20495).
<emphasis id="A9A01B34A32FDB38FD6AEA17FD20FAB1" bold="true" box="[683,704,1339,1363]" pageId="4" pageNumber="40">a,</emphasis>
oral view of gonopods;
<emphasis id="A9A01B34A32FDB38FC04EA17FC3BFAB1" bold="true" box="[965,987,1339,1363]" pageId="4" pageNumber="40">b,</emphasis>
aboral view of gonopods;
<emphasis id="A9A01B34A32FDB38FB3BEA17FAECFAB1" bold="true" box="[1274,1292,1339,1363]" pageId="4" pageNumber="40">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="A9A01B34A32FDB38FD9BEA72FD8FFA94" bold="true" box="[602,623,1374,1398]" pageId="4" pageNumber="40">d,</emphasis>
lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe.
</paragraph>
</caption>
<subSubSection id="D3CE94ADA32FDB38FF04EAC5FEFAF9B2" pageId="4" pageNumber="40" type="diagnosis">
<paragraph id="9B6BC726A32FDB38FF04EAC5FEFAF9B2" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04EAC5FEA8F9E1" bold="true" box="[197,328,1513,1539]" pageId="4" pageNumber="40">Diagnosis:</emphasis>
Lateral and medial edges of proplica parallel; proplica without abrupt widening subapically; angle between lateral lobe and distal lobe equal to or less than a right angle; antetorsal process S-shaped (
<figureCitation id="03EFDBA3A32FDB38FAA9E93CFF43F9B2" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="4" pageNumber="40">Figs 1</figureCitation>
a, 1b, 7a).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32FDB39FF04E971FD50FDEC" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40" type="description">
<paragraph id="9B6BC726A32FDB38FF04E971FDD1F97C" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E971FEBFF995" bold="true" box="[197,351,1629,1655]" pageId="4" pageNumber="40">Description:</emphasis>
<emphasis id="A9A01B34A32FDB38FEA6E971FE76F994" box="[359,406,1629,1654]" italics="true" pageId="4" pageNumber="40">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A32FDB38FDFDE971FD3BF995" box="[572,731,1629,1655]" metricMagnitude="-1" metricUnit="m" metricValue="2.6399999999999997" metricValueMax="3.44" metricValueMin="1.8399999999999999" pageId="4" pageNumber="40" unit="mm" value="264.0" valueMax="344.0" valueMin="184.0">184344 mm</quantity>
; minimum and maximum body width 814.5 mm and 13.5
<quantity id="5C2C6AC3A32FDB38FF56E9A8FF0CF97C" box="[151,236,1668,1694]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="40" unit="mm" value="19.0">19 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A32FDB38FE6AE9A8FDCDF97C" box="[427,557,1668,1694]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="4" pageNumber="40" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E987FDC6F926" blockId="4.[151,1436,1513,2006]" box="[197,550,1706,1732]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E987FE25F926" box="[197,453,1707,1732]" italics="true" pageId="4" pageNumber="40">Number of body rings</emphasis>
: 5968.
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E9FDFC69F909" blockId="4.[151,1436,1513,2006]" box="[197,905,1745,1771]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E9FDFEF8F908" box="[197,280,1745,1770]" italics="true" pageId="4" pageNumber="40">Colour</emphasis>
: Body black; legs and antennae black or dark brown.
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E9D4FEE5F8DA" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E9D4FEFBF8F3" box="[197,283,1784,1809]" italics="true" pageId="4" pageNumber="40">Collum</emphasis>
: Anterior produced laterally into a narrow lobe, with 25 complete folds and 03 incomplete folds (
<figureCitation id="03EFDBA3A32FDB38FF5EE832FF02F8DA" box="[159,226,1822,1848]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="4" pageNumber="40">Fig. 1</figureCitation>
d).
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E864FD30F86B" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E864FD39F883" box="[197,729,1861,1889]" italics="true" pageId="4" pageNumber="40">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally parallel and rounded, medial nipples distally narrow and strongly deflected laterally (
<figureCitation id="03EFDBA3A32FDB38FDACE843FD4FF86B" box="[621,687,1903,1929]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="4" pageNumber="40">Fig. 1</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E8BAFBAAF84D" blockId="4.[151,1436,1513,2006]" box="[197,1098,1941,1967]" pageId="4" pageNumber="40">
<emphasis id="A9A01B34A32FDB38FF04E8BAFED2F84D" box="[197,306,1942,1967]" italics="true" pageId="4" pageNumber="40">Gonopod</emphasis>
: (
<figureCitation id="03EFDBA3A32FDB38FE88E8B9FE71F84D" box="[329,401,1941,1967]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="4" pageNumber="40">Figs 1</figureCitation>
a, 1b)
<quantity id="5C2C6AC3A32FDB38FE1CE8B9FDDFF84D" box="[477,575,1941,1967]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="4" pageNumber="40" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
long; telocoxites parallel, but not in contact.
</paragraph>
<paragraph id="9B6BC726A32FDB38FF04E890FB84F834" blockId="4.[151,1436,1513,2006]" box="[197,1124,1980,2006]" pageId="4" pageNumber="40">Sternite short, wide and apex not extending distally to level of paracoxite apex.</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EFB4FE64FF3A" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
Proplica tapering apically. Apex of proplica conceals proximal lateral lobe. Proplica does not overlap distal lobe (
<figureCitation id="03EFDBA3A32EDB39FEE1EF92FE83FF3A" box="[288,355,190,216]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Fig. 1</figureCitation>
a).
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EFC9FB86FF1D" blockId="5.[151,1437,152,951]" box="[197,1126,229,255]" pageId="5" pageNumber="41">
Metaplica without lobes or humps, wider proximal to telopodite knee (
<figureCitation id="03EFDBA3A32EDB39FBC0EFC9FBA3FF1D" box="[1025,1091,229,255]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Fig. 1</figureCitation>
b).
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EE20FDB8FEAE" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
Lateral lobe extends beyond lateral edges of proplica and distal lobe. Aborally, lateral lobe proximally narrow, rounded distally (
<figureCitation id="03EFDBA3A32EDB39FE02EE1EFDEBFEAE" box="[451,523,306,332]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Figs 1</figureCitation>
a, 1b).
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EE75FB9FFE22" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
Distal lobe laterally and apically rounded; medial edge of distal lobe concave, forming a shallow invagination. Surface of distal telocoxite lobe smooth; distal lobe not extending beyond lateral edge of proplica apex (
<figureCitation id="03EFDBA3A32EDB39FE84EE8AFE68FE22" box="[325,392,422,448]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Fig. 1</figureCitation>
a). Orally, the distal lobe may have a shallow central depression.
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EEE1FD50FDEC" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
Telopodite with antetorsal process which extends past paracoxite, post-knee telopodite with an open coil. Antetorsal process S-shaped (
<figureCitation id="03EFDBA3A32EDB39FE33EED8FDD9FDEC" box="[498,569,500,526]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Figs 1</figureCitation>
a, 1b, 7a).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32EDB39FF04ED36FEF9FDB9" pageId="5" pageNumber="41" type="distribution">
<paragraph id="9B6BC726A32EDB39FF04ED36FEF9FDB9" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
<emphasis id="A9A01B34A32EDB39FF04ED36FE87FDD6" bold="true" box="[197,359,538,564]" pageId="5" pageNumber="41">Distribution:</emphasis>
Known from southern
<collectingCountry id="E3C387B6A32EDB39FDB3ED36FD06FDD6" box="[626,742,538,564]" name="Botswana" pageId="5" pageNumber="41">Botswana</collectingCountry>
, north-western
<collectingCountry id="E3C387B6A32EDB39FC56ED36FBCDFDD6" box="[919,1069,538,564]" name="South Africa" pageId="5" pageNumber="41">South Africa</collectingCountry>
, northern
<collectingCountry id="E3C387B6A32EDB39FB5EED36FAE2FDD6" box="[1183,1282,538,564]" name="Namibia" pageId="5" pageNumber="41">Namibia</collectingCountry>
and southern
<collectingCountry id="E3C387B6A32EDB39FF56ED6DFEF6FDB9" box="[151,278,577,603]" name="Zimbabwe" pageId="5" pageNumber="41">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32EDB39FF04ED4BFAE0FC55" pageId="5" pageNumber="41" type="discussion">
<paragraph id="9B6BC726A32EDB39FF04ED4BFC7DFD4A" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
<emphasis id="A9A01B34A32EDB39FF04ED4BFEA0FD63" bold="true" box="[197,320,615,641]" pageId="5" pageNumber="41">Remarks:</emphasis>
The smooth surface texture and shape of the distal lobe of the gonopods resemble the form in
<taxonomicName id="5CD4BCA5A32EDB39FF56EDA3FEF3FD4A" box="[151,275,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="batokensis">
<emphasis id="A9A01B34A32EDB39FF56EDA3FEF3FD4A" box="[151,275,655,680]" italics="true" pageId="5" pageNumber="41">batokensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5CD4BCA5A32EDB39FEE0EDA3FE9AFD4A" box="[289,378,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A32EDB39FEE0EDA3FE9AFD4A" box="[289,378,655,680]" italics="true" pageId="5" pageNumber="41">unciger</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32EDB39FE75EDA3FDF7FD4A" box="[436,535,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FE75EDA3FDF7FD4A" box="[436,535,655,680]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
rather than
<taxonomicName id="5CD4BCA5A32EDB39FD65EDA3FD07FD4A" box="[676,743,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A32EDB39FD65EDA3FD07FD4A" box="[676,743,655,680]" italics="true" pageId="5" pageNumber="41">sebae</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32EDB39FCDEEDA3FC76FD4A" box="[799,918,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A32EDB39FCDEEDA3FC76FD4A" box="[799,918,655,680]" italics="true" pageId="5" pageNumber="41">tripartitus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04ED99FAE0FC55" blockId="5.[151,1437,152,951]" pageId="5" pageNumber="41">
The gonopods of
<taxonomicName id="5CD4BCA5A32EDB39FE52ED99FE33FD2C" box="[403,467,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FE52ED99FE33FD2C" box="[403,467,693,718]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32EDB39FDCDED99FD8FFD2C" box="[524,623,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FDCDED99FD8FFD2C" box="[524,623,693,718]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
are the most similar. However, in
<taxonomicName id="5CD4BCA5A32EDB39FBC5ED99FBA4FD2C" box="[1028,1092,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FBC5ED99FBA4FD2C" box="[1028,1092,693,718]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
the angle between the lateral lobe and distal lobe is equal to or less than a right angle while in
<taxonomicName id="5CD4BCA5A32EDB39FC53EDF0FBF2FD17" box="[914,1042,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FC53EDF0FBF2FD17" box="[914,1042,732,757]" italics="true" pageId="5" pageNumber="41">S. kruegeri</emphasis>
</taxonomicName>
it is wider. Additionally, in
<taxonomicName id="5CD4BCA5A32EDB39FA9DEDF0FA7CFD17" box="[1372,1436,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FA9DEDF0FA7CFD17" box="[1372,1436,732,757]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
the lateral edge of the proplica is straight, without abrupt widening subapically while in
<taxonomicName id="5CD4BCA5A32EDB39FB64EC2EFAE8FCF9" box="[1189,1288,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FB64EC2EFAE8FCF9" box="[1189,1288,770,795]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
the proplica is wide subapically. The antetorsal process in
<taxonomicName id="5CD4BCA5A32EDB39FD7BEC05FD1BFCA0" box="[698,763,809,834]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FD7BEC05FD1BFCA0" box="[698,763,809,834]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
is S-shaped unlike in
<taxonomicName id="5CD4BCA5A32EDB39FBC4EC05FB89FCA0" box="[1029,1129,809,834]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FBC4EC05FB89FCA0" box="[1029,1129,809,834]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
where it is concave (
<figureCitation id="03EFDBA3A32EDB39FAA6EC05FF47FC88" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="5" pageNumber="41">Figs 7</figureCitation>
a, 7b). The distal nipple on the lateral lobe in some specimens of
<taxonomicName id="5CD4BCA5A32EDB39FC60EC7CFBE4FC8B" box="[929,1028,848,873]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FC60EC7CFBE4FC8B" box="[929,1028,848,873]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
has not been observed in
<taxonomicName id="5CD4BCA5A32EDB39FAF4EC7CFA95FC8B" box="[1333,1397,848,873]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FAF4EC7CFA95FC8B" box="[1333,1397,848,873]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
. In addition, the medial edges of the telocoxites are straight and largely parallel along the whole length in
<taxonomicName id="5CD4BCA5A32EDB39FAFCEC5BFA9DFC72" box="[1341,1405,887,912]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32EDB39FAFCEC5BFA9DFC72" box="[1341,1405,887,912]" italics="true" pageId="5" pageNumber="41">heros</emphasis>
</taxonomicName>
as opposed to
<taxonomicName id="5CD4BCA5A32EDB39FEDFECB1FE61FC54" box="[286,385,925,950]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FEDFECB1FE61FC54" box="[286,385,925,950]" italics="true" pageId="5" pageNumber="41">kruegeri</emphasis>
</taxonomicName>
where they are widely separated at the level of the lateral lobe (
<figureCitation id="03EFDBA3A32EDB39FBAFECB1FB56FC55" box="[1134,1206,925,951]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="5" pageNumber="41">Figs 1</figureCitation>
a, 2a).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,228 @@
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Genus
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<emphasis id="A9A01B34A329DB3EFF2FE9A5FE6EF941" bold="true" box="[238,398,1673,1699]" italics="true" pageId="2" pageNumber="38">Spirostreptus</emphasis>
<bibRefCitation id="FF45BAD7A329DB3EFE54E9A4FDD0F940" author="Brandt" box="[405,560,1672,1698]" pageId="2" pageNumber="55" refString="Brandt, J. F. (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognathi Latreillii spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 6, 194 - 209." type="journal article" year="1833">Brandt 1833</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
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<paragraph id="9B6BC726A329DB3EFF56E9E3FC4CF8C7" blockId="2.[151,1436,1742,1925]" pageId="2" pageNumber="38">
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<emphasis id="A9A01B34A329DB3EFF56E9E3FEC3F904" box="[151,291,1743,1766]" italics="true" pageId="2" pageNumber="38">Spirostreptus</emphasis>
<bibRefCitation id="FF45BAD7A329DB3EFEEBE9E2FE51F907" author="Brandt" box="[298,433,1742,1765]" pageId="2" pageNumber="55" refString="Brandt, J. F. (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognathi Latreillii spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 6, 194 - 209." type="journal article" year="1833">Brandt 1833</bibRefCitation>
</taxonomicName>
, p. 203 (
<typeStatus id="446F7984A329DB3EFDCCE9E2FDDCF907" box="[525,572,1742,1765]" pageId="2" pageNumber="38">type</typeStatus>
species:
<taxonomicName id="5CD4BCA5A329DB3EFD5EE9E3FC1DF907" ID-CoL="4Z8WQ" authority="Brandt 1833" authorityName="Brandt" authorityYear="1833" box="[671,1021,1742,1766]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="38" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A329DB3EFD5EE9E3FC8EF904" box="[671,878,1743,1766]" italics="true" pageId="2" pageNumber="38">Spirostreptus sebae</emphasis>
<bibRefCitation id="FF45BAD7A329DB3EFCB7E9E2FC1DF907" author="Brandt" box="[886,1021,1742,1765]" pageId="2" pageNumber="55" refString="Brandt, J. F. (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognathi Latreillii spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 6, 194 - 209." type="journal article" year="1833">Brandt 1833</bibRefCitation>
</taxonomicName>
; by subsequent designation of
<bibRefCitation id="FF45BAD7A329DB3EFA86E9E2FF1FF8E7" author="Pocock" pageId="2" pageNumber="56" refString="Pocock, R. I. (1894) Chilopoda Symphyla and Diplopoda from the Malay Archipelago. Pp. 307 - 404. In: Weber M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 3. E. J. Brill, Leiden." type="journal article" year="1894">Pocock, 1894</bibRefCitation>
, p. 388);
<taxonomicName id="5CD4BCA5A329DB3EFEA7E9C3FD32F8E7" ID-CoL="7Q2ML" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[358,722,1774,1798]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="A9A01B34A329DB3EFEA7E9C3FE12F8E4" box="[358,498,1775,1798]" italics="true" pageId="2" pageNumber="38">Spirostreptus</emphasis>
:
<bibRefCitation id="FF45BAD7A329DB3EFDC3E9C2FD6EF8E7" author="Attems" box="[514,654,1774,1797]" pageId="2" pageNumber="55" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25 (65 / 66), 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFD5BE9C2FD32F8E7" author="Attems" box="[666,722,1774,1797]" pageId="2" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">1928</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="FF45BAD7A329DB3EFD1EE9C2FC9EF8E7" author="Hoffman" box="[735,894,1774,1797]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (1965) A new species in the diplopod genus Archispirostreptus from south western Arabia. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 3 (51), 17 - 23." type="journal article" year="1965">Hoffman 1965</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFC4BE9C2FC22F8E7" author="Hoffman" box="[906,962,1774,1797]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (1979) Classification of the Diplopoda. Geneve: Museum d'Histoire Naturelle, 237 pp." type="book" year="1979">1979</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFC0FE9C2FBE7F8E7" author="Hoffman" box="[974,1031,1774,1797]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">2008</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFBD2E9C2FB42F8E7" author="Krabbe" box="[1043,1186,1774,1797]" pageId="2" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFB6FE9C2FF1FF8C7" author="Golovatch" pageId="2" pageNumber="55" refString="Golovatch, S. I. &amp; Hoffman, R. L. (2000) On the diplopod taxa and type material of J. F. Brandt, with some new descriptions and identities (Diplopoda). Pp. 229 - 249. In: Wytwer, J. &amp; Golovatch, S. (Eds.), Progress in Studies on Myriapoda and Onychophora. Fragmenta Faunistica, 43 (Supplement)." type="journal volume" year="2000">Golovatch &amp; Hoffman 2000</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFEC8E822FE70F8C7" author="Hamer" box="[265,400,1806,1829]" pageId="2" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFE5AE822FE33F8C7" author="Hamer" box="[411,467,1806,1829]" pageId="2" pageNumber="55" refString="Hamer, M. L. (1999) An illustrated key to the spirostreptidan (Diplopoda: Spirostreptida) genera of Southern Africa. Annals of the Natal Museum, 40, 1 - 22." type="journal article" year="1999">1999</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFE1FE822FD57F8C7" author="Hoffman" box="[478,695,1806,1830]" pageId="2" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">
Hoffman
<emphasis id="A9A01B34A329DB3EFD85E823FD91F8C4" box="[580,625,1807,1830]" italics="true" pageId="2" pageNumber="38">et al</emphasis>
. 2001
</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFD03E822FC8DF8C7" author="Mwabvu" box="[706,877,1806,1829]" pageId="2" pageNumber="56" refString="Mwabvu, T. (2005 b) The density and distribution of millipedes on termite mounds in miombo woodland, Zimbabwe. African Journal of Ecology, 43 (4), 400 - 402." type="journal article" year="2005" yearSuffix="b">Mwabvu 2005b</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFCB6E822FC4CF8C7" author="Mwabvu" box="[887,940,1806,1829]" pageId="2" pageNumber="56" refString="Mwabvu, T. (2006) Spirostreptid millipedes (Diplopoda, Spirostreptida) of urban and peri-urban habitats in Zimbabwe. African Journal of Ecology, 45 (3), 311 - 314." type="journal article" year="2006">2006</bibRefCitation>
</paragraph>
<paragraph id="9B6BC726A329DB3EFF56E803FF1CF867" blockId="2.[151,1436,1742,1925]" pageId="2" pageNumber="38">
<taxonomicName id="5CD4BCA5A329DB3EFF56E803FE37F8A7" ID-CoL="b683b7ee-ceea-4e63-8d8d-d00896a28b07" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[151,471,1838,1862]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="A9A01B34A329DB3EFF56E803FEDDF8A4" box="[151,317,1839,1862]" italics="true" pageId="2" pageNumber="38">Triaenostreptus</emphasis>
<bibRefCitation id="FF45BAD7A329DB3EFE89E802FE37F8A7" author="Attems" box="[328,471,1838,1861]" pageId="2" pageNumber="55" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25 (65 / 66), 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
</taxonomicName>
, p. 149 (
<typeStatus id="446F7984A329DB3EFDFFE802FD8DF8A7" box="[574,621,1838,1861]" pageId="2" pageNumber="38">type</typeStatus>
species
<taxonomicName id="5CD4BCA5A329DB3EFD0EE803FBABF8A7" ID-CoL="4Z8XW" authority="Attems 1909" authorityName="Attems" authorityYear="1909" box="[719,1099,1838,1862]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="38" phylum="Arthropoda" rank="species" species="triodus">
<emphasis id="A9A01B34A329DB3EFD0EE803FC52F8A4" box="[719,946,1839,1862]" italics="true" pageId="2" pageNumber="38">Spirostreptus triodus</emphasis>
<bibRefCitation id="FF45BAD7A329DB3EFC7DE802FBABF8A7" author="Attems" box="[956,1099,1838,1861]" pageId="2" pageNumber="55" refString="Attems, C. (1909) Myriapoda. In: Schultze, L. (Ed.), Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905. Denkschriften der medizinischnaturwissenschaftlichen Gesellschaft zu Jena, 14, 1 - 52." type="journal article" year="1909">Attems 1909</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation id="FF45BAD7A329DB3EFF06E862FEB0F887" author="Jeekel" box="[199,336,1870,1893]" pageId="2" pageNumber="56" refString="Jeekel, C. A. W. (1970) Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging, 5, xii + 412 pp." type="book" year="1970">Jeekel, 1970</bibRefCitation>
, p. 139);
<bibRefCitation id="FF45BAD7A329DB3EFE77E862FDA4F887" author="Attems" box="[438,580,1870,1893]" pageId="2" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFD8EE862FC9BF887" author="Golovatch" box="[591,891,1870,1893]" pageId="2" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">Golovatch &amp; Hoffman 2001</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFC47E862FBC5F887" author="Hoffman" box="[902,1061,1870,1893]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFBF0E862FB89F887" author="Hoffman" box="[1073,1129,1870,1893]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (1979) Classification of the Diplopoda. Geneve: Museum d'Histoire Naturelle, 237 pp." type="book" year="1979">1979</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A329DB3EFBB4E862FB4EF887" author="Hoffman" box="[1141,1198,1870,1893]" pageId="2" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">2008</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFB78E862FAA8F887" author="Krabbe" box="[1209,1352,1870,1893]" pageId="2" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A329DB3EFA92E862FF1CF867" author="Hamer" pageId="2" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA329DB3FFF56E899FE44FC03" lastPageId="3" lastPageNumber="39" pageId="2" pageNumber="38" type="diagnosis">
<paragraph id="9B6BC726A329DB3EFF56E899FC89F814" blockId="2.[151,1436,1973,2038]" pageId="2" pageNumber="38">
<emphasis id="A9A01B34A329DB3EFF56E899FEFFF82D" bold="true" box="[151,287,1973,1999]" pageId="2" pageNumber="38">Diagnosis:</emphasis>
Large spirostreptid millipedes, body length
<quantity id="5C2C6AC3A329DB3EFC85E899FC0AF82D" box="[836,1002,1973,1999]" metricMagnitude="-1" metricUnit="m" metricValue="2.53" metricValueMax="3.44" metricValueMin="1.6199999999999999" pageId="2" pageNumber="38" unit="mm" value="253.0" valueMax="344.0" valueMin="162.0">162344 mm</quantity>
; antennae
<quantity id="5C2C6AC3A329DB3EFBADE899FB05F82D" box="[1132,1253,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="2" pageNumber="38" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">913 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A329DB3EFE60E8F0FDF1F814" box="[417,529,2012,2038]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="2" pageNumber="38" unit="mm" value="9.5" valueMax="15.0" valueMin="4.0">415 mm</quantity>
and
<quantity id="5C2C6AC3A329DB3EFD88E8F0FD2AF814" box="[585,714,2012,2038]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="2" pageNumber="38" unit="mm" value="15.0" valueMax="19.0" valueMin="11.0">1119 mm</quantity>
, respectively.
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EFB4FCA7FF50" blockId="3.[151,1437,152,1302]" box="[197,839,152,178]" pageId="3" pageNumber="39">
Prozonite
<quantity id="5C2C6AC3A328DB3FFEFDEFB4FE7DFF50" box="[316,413,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="39" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
wide and metazonite
<quantity id="5C2C6AC3A328DB3FFD5FEFB4FD1FFF50" box="[670,767,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="39" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
wide.
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EF92FDC1FF3A" blockId="3.[151,1437,152,1302]" box="[197,545,190,216]" pageId="3" pageNumber="39">Number of body rings 6073.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EFC9FBF8FF1D" blockId="3.[151,1437,152,1302]" box="[197,1048,229,255]" pageId="3" pageNumber="39">Three separate sclerites form the prebasilar plate of the gnathochilarium.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EE20FF39FEAE" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">Collum with an anterior lobe, black or dark brown in colour, with 25 complete folds and 07 incomplete folds.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EE75FE22FE78" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">Body black or brown in colour. Legs, antennae and anal valve colour generally lighter shades of body colour, often light brown.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EE85FA75FE21" blockId="3.[151,1437,152,1302]" box="[197,1429,422,451]" pageId="3" pageNumber="39">Prozonite yellowish to brown; metazonite with one row of sagilla. Ozopores begin on 5th or 6th body ring.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EEFCFB4AFDF3" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
Gonopod
<quantity id="5C2C6AC3A328DB3FFEFBEEFCFE7DFE08" box="[314,413,464,490]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="3" pageNumber="39" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">69 mm</quantity>
long; sternite triangular; paracoxite fused tightly to the metaplica, apically horizontal. Telocoxites approximately parallel to each other, medial edge straight and close together.
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04ED31FC4DFDBC" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">Proplica approximately 2/3 of gonopod length; proplica broad sub-apically, setose apically. Proplica with a central groove flanked by sharply raised lateral and medial edges.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04ED46FCB0FD49" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">Thumb-like lateral lobe present between the distal telocoxite lobe and apical proplica, lateral lobe directed towards paracoxite at an angle to lateral edge of distal lobe.</paragraph>
<paragraph id="9B6BC726A328DB3FFF04ED94FF0EFD1B" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
Distal telocoxite enlarged and laterally produced into rounded golf club-shaped lobe (
<figureCitation id="03EFDBA3A328DB3FFB7AED94FAE3FD30" box="[1211,1283,696,722]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="3" pageNumber="39">Figs 1</figureCitation>
a, 2a, 3a, 4a, 5a, 6a).
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EC29FF11FCA4" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
Metaplica with a small window through which part of pre-knee telopodite is visible (
<figureCitation id="03EFDBA3A328DB3FFB75EC29FB1CFCFD" box="[1204,1276,773,799]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="3" pageNumber="39">Figs 1</figureCitation>
b, 2b, 3b, 4b, 5b, 6b).
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EC7EFE44FC03" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
Telopodite knee and origin of antetorsal process concealed under apex of proplica. Antetorsal process tapering; post knee telopodite coils or spirals proximally, becoming lamellate before narrowing distally into a long thin extension with a trifurcate ending. Antetorsal process straight, S-shaped or crescent-shaped, with terminal spine (
<figureCitation id="03EFDBA3A328DB3FFE8CECEBFE75FC03" box="[333,405,967,993]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="3" pageNumber="39">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA328DB3FFF04ECC1FC9AFBCC" pageId="3" pageNumber="39" type="distribution">
<paragraph id="9B6BC726A328DB3FFF04ECC1FC9AFBCC" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
<emphasis id="A9A01B34A328DB3FFF04ECC1FE97FBE5" bold="true" box="[197,375,1005,1031]" pageId="3" pageNumber="39">Distribution:</emphasis>
Known from southern Africa (
<collectingCountry id="E3C387B6A328DB3FFCC4ECC1FC88FBE5" box="[773,872,1005,1031]" name="Angola" pageId="3" pageNumber="39">Angola</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFCB7ECC1FC16FBE5" box="[886,1014,1005,1031]" name="Botswana" pageId="3" pageNumber="39">Botswana</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFBC5ECC1FA77FBE5" box="[1028,1431,1005,1031]" name="Democratic Republic of the Congo" pageId="3" pageNumber="39">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFF56EB38FED5FBCC" box="[151,309,1044,1070]" name="Mozambique" pageId="3" pageNumber="39">Mozambique</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFE81EB38FE48FBCC" box="[320,424,1044,1070]" name="Namibia" pageId="3" pageNumber="39">Namibia</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFE75EB38FDADFBCC" box="[436,589,1044,1070]" name="South Africa" pageId="3" pageNumber="39">South Africa</collectingCountry>
,
<collectingCountry id="E3C387B6A328DB3FFD98EB38FD53FBCC" box="[601,691,1044,1070]" name="Zambia" pageId="3" pageNumber="39">Zambia</collectingCountry>
and
<collectingCountry id="E3C387B6A328DB3FFD2AEB38FC8EFBCC" box="[747,878,1044,1070]" name="Zimbabwe" pageId="3" pageNumber="39">Zimbabwe</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA328DB3FFF04EB16FC09FAF4" pageId="3" pageNumber="39" type="discussion">
<paragraph id="9B6BC726A328DB3FFF04EB16FB56FB40" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">
<emphasis id="A9A01B34A328DB3FFF04EB16FED5FBB6" bold="true" box="[197,309,1082,1108]" pageId="3" pageNumber="39">Remark:</emphasis>
Although
<bibRefCitation id="FF45BAD7A328DB3FFE73EB16FD91FBB6" author="Hoffman" box="[434,625,1082,1108]" pageId="3" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman (2008)</bibRefCitation>
did not use the presence of the distal lobe and the slanting digitiform lateral lobe to define
<taxonomicName id="5CD4BCA5A328DB3FFE51EB4DFDCAFB98" box="[400,554,1121,1146]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="A9A01B34A328DB3FFE51EB4DFDCAFB98" box="[400,554,1121,1146]" italics="true" pageId="3" pageNumber="39">Spirostreptus</emphasis>
</taxonomicName>
in his key to the genera of Spirostreptini, we think that these characters are taxonomically more informative at the genus level than the form of the antetorsal process.
</paragraph>
<paragraph id="9B6BC726A328DB3FFF04EB83FC09FAF4" blockId="3.[151,1437,152,1302]" pageId="3" pageNumber="39">The distribution of the genus shows large gaps even in habitats similar to those from which the genus is known. These gaps are mainly in less accessible regions which suggest that current understanding of the distribution is a result of collecting bias rather than a true fragmentation.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,379 @@
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<mods:title id="9D02BA6E645D1C07D1A1633B4F06E721">A revision of the taxonomy and distribution of Spirostreptus Brandt 1833 (Diplopoda, Spirostreptida, Spirostreptidae) with descriptions of a new species and a new genus of spirostreptid millipede</mods:title>
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<mods:namePart id="BC445463E7BCC6D4BBBB4C9E98650280">Mwabvu, Tarombera</mods:namePart>
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<mods:namePart id="7910B21A62670ACC5935F06BB347F6F9">Hamer, Michelle</mods:namePart>
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<mods:namePart id="5065FCFF0E51F6CAE4360E4ED9FAEC0E">Slotow, Robert</mods:namePart>
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<mods:namePart id="2A01D75CC6A5308F74B18FC5B1C826E9">Barraclough, David</mods:namePart>
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<mods:typeOfResource id="BC34975E1E92F497298068714942FC55">text</mods:typeOfResource>
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<mods:title id="8E862835E300686C2F2BF02E5FCDC730">Zootaxa</mods:title>
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<mods:date id="D911EEEEC81689535F9E2234A5A8108C">2009</mods:date>
<mods:detail id="B0A3D0F20CE2B40FDCE488C52B86AC87" type="volume">
<mods:number id="3AA8802D37AF8DF6A08DCDC112D554D8">2211</mods:number>
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<mods:identifier id="1E1EAFD644E8E35181DEB7B86EF14067" type="DOI">10.5281/zenodo.189858</mods:identifier>
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<treatment id="137D7630A32CDB35FF56EB3DFD8CFD14" ID-DOI="http://doi.org/10.5281/zenodo.6220852" ID-GBIF-Taxon="119623016" ID-Zenodo-Dep="6220852" LSID="urn:lsid:plazi:treatment:137D7630A32CDB35FF56EB3DFD8CFD14" httpUri="http://treatment.plazi.org/id/137D7630A32CDB35FF56EB3DFD8CFD14" lastPageId="9" lastPageNumber="45" pageId="7" pageNumber="43">
<subSubSection id="D3CE94ADA32CDB3BFF56EB3DFEE9FBAC" pageId="7" pageNumber="43" type="nomenclature">
<paragraph id="9B6BC726A32CDB3BFF56EB3DFDC4FBC8" blockId="7.[151,548,1040,1102]" box="[151,548,1040,1067]" pageId="7" pageNumber="43">
<heading id="C023704AA32CDB3BFF56EB3DFDC4FBC8" bold="true" box="[151,548,1040,1067]" fontSize="11" level="1" pageId="7" pageNumber="43" reason="1">
<taxonomicName id="5CD4BCA5A32CDB3BFF56EB3DFDC4FBC8" ID-CoL="4Z8WQ" authority="Brandt 1833" authorityName="Brandt" authorityYear="1833" box="[151,548,1040,1067]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A32CDB3BFF56EB3DFDC4FBC8" bold="true" box="[151,548,1040,1067]" pageId="7" pageNumber="43">
<emphasis id="A9A01B34A32CDB3BFF56EB3DFE61FBC9" bold="true" box="[151,385,1041,1067]" italics="true" pageId="7" pageNumber="43">Spirostreptus sebae</emphasis>
<bibRefCitation id="FF45BAD7A32CDB3BFE49EB3CFDC4FBC8" author="Brandt" box="[392,548,1040,1066]" pageId="7" pageNumber="55" refString="Brandt, J. F. (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognathi Latreillii spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 6, 194 - 209." type="journal article" year="1833">Brandt 1833</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B6BC726A32CDB3BFF56EB18FEE9FBAC" blockId="7.[151,548,1040,1102]" box="[151,265,1076,1102]" pageId="7" pageNumber="43">
<figureCitation id="03EFDBA3A32CDB3BFF56EB18FF01FBAC" box="[151,225,1076,1102]" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="7" pageNumber="43">Figs 3</figureCitation>
,
<figureCitation id="03EFDBA3A32CDB3BFF2FEB18FF1DFBAC" box="[238,253,1076,1102]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="7" pageNumber="43">7</figureCitation>
c
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32CDB3BFF56EBADFA87FB3A" pageId="7" pageNumber="43" type="reference_group">
<paragraph id="9B6BC726A32CDB3BFF56EBADFE82FB5A" blockId="7.[151,1436,1153,1240]" pageId="7" pageNumber="43">
<taxonomicName id="5CD4BCA5A32CDB3BFF56EBADFE17FB7A" ID-CoL="4Z8WQ" authority="Brandt 1833" authorityName="Brandt" authorityYear="1833" box="[151,503,1153,1176]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A32CDB3BFF56EBADFE88FB7A" box="[151,360,1153,1176]" italics="true" pageId="7" pageNumber="43">Spirostreptus sebae</emphasis>
<bibRefCitation id="FF45BAD7A32CDB3BFEAFEBADFE17FB7A" author="Brandt" box="[366,503,1153,1176]" pageId="7" pageNumber="55" refString="Brandt, J. F. (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognathi Latreillii spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 6, 194 - 209." type="journal article" year="1833">Brandt 1833</bibRefCitation>
</taxonomicName>
, p.203, figs 4045;
<bibRefCitation id="FF45BAD7A32CDB3BFD0FEBADFC1AFB7A" author="Golovatch" box="[718,1018,1153,1176]" pageId="7" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">Golovatch &amp; Hoffman 2001</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFBC4EBADFB00FB7A" author="Hoffman" box="[1029,1248,1153,1176]" pageId="7" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">
Hoffman
<emphasis id="A9A01B34A32CDB3BFBADEBADFB7AFB7A" box="[1132,1178,1153,1176]" italics="true" pageId="7" pageNumber="43">et al</emphasis>
. 2001
</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFB2CEBADFA78FB7A" author="Mwabvu" box="[1261,1432,1153,1176]" pageId="7" pageNumber="56" refString="Mwabvu, T. (2005 a) New records and localities of spirostreptid millipedes (Myriapoda: Diplopoda) of Zimbabwe with notes on nomenclatural changes and habitat. Arnoldia, Zimbabwe, 10 (25), 249 - 256." type="journal article" year="2005" yearSuffix="a">Mwabvu 2005a</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFF06EB8DFE82FB5A" author="Hoffman" box="[199,354,1185,1208]" pageId="7" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
</paragraph>
<paragraph id="9B6BC726A32CDB3BFF56EBEDFA87FB3A" blockId="7.[151,1436,1153,1240]" box="[151,1383,1217,1240]" pageId="7" pageNumber="43">
<taxonomicName id="5CD4BCA5A32CDB3BFF56EBEDFDFEFB3A" ID-CoL="58579" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,542,1217,1240]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="petersi">
<emphasis id="A9A01B34A32CDB3BFF56EBEDFE6CFB3A" box="[151,396,1217,1240]" italics="true" pageId="7" pageNumber="43">Triaenostreptus petersi</emphasis>
<bibRefCitation id="FF45BAD7A32CDB3BFE52EBEDFDFEFB3A" author="Attems" box="[403,542,1217,1240]" pageId="7" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</taxonomicName>
, p. 367;
<bibRefCitation id="FF45BAD7A32CDB3BFDB8EBEDFCF8FB3A" author="Hoffman" box="[633,792,1217,1240]" pageId="7" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
,
<bibRefCitation id="FF45BAD7A32CDB3BFCE2EBEDFCBBFB3A" author="Hoffman" box="[803,859,1217,1240]" pageId="7" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">2008</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFCA7EBEDFC14FB3A" author="Krabbe" box="[870,1012,1217,1240]" pageId="7" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFC3EEBEDFB66FB3A" author="Hamer" box="[1023,1158,1217,1240]" pageId="7" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32CDB3BFB50EBEDFA87FB3A" author="Hoffman" box="[1169,1383,1217,1240]" pageId="7" pageNumber="56" refString="Hoffman, R. L., Golovatch, S. I. &amp; Hamer, M. (2001) Identities of millipede genera Spirostreptus Brandt, 1833 and Spiropoeus Brandt, 1833 (Diplopoda, Spirostreptida, Spirostreptidae). Myriapodologica, 7 (5), 35 - 47." type="journal article" year="2001">
Hoffman
<emphasis id="A9A01B34A32CDB3BFB39EBEDFAC5FB3A" box="[1272,1317,1217,1240]" italics="true" pageId="7" pageNumber="43">et al</emphasis>
. 2001
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32CDB3BFF56EA2BFEB7F83E" pageId="7" pageNumber="43" type="materials_examined">
<paragraph id="9B6BC726A32CDB3BFF56EA2BFBE4FAC0" blockId="7.[151,1437,1287,2012]" box="[151,1028,1287,1314]" pageId="7" pageNumber="43">
<emphasis id="A9A01B34A32CDB3BFF56EA2BFEADFAC3" bold="true" box="[151,333,1287,1313]" pageId="7" pageNumber="43">
<typeStatus id="446F7984A32CDB3BFF56EA2BFF32FAC3" box="[151,210,1287,1313]" pageId="7" pageNumber="43">Type</typeStatus>
material:
</emphasis>
<typeStatus id="446F7984A32CDB3BFE97EA24FE28FAC0" box="[342,456,1288,1314]" pageId="7" pageNumber="43" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="E3C387B6A32CDB3BFE12EA24FD42FAC0" box="[467,674,1288,1314]" name="Mozambique" pageId="7" pageNumber="43">MOZAMBIQUE</collectingCountry>
: 1 3, Tete, Peters (
<collectionCode id="FDC55FE3A32CDB3BFCBFEA24FC5FFAC0" box="[894,959,1288,1314]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="7" pageNumber="43">ZMB</collectionCode>
477).
</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EA02FEB7F83E" blockId="7.[151,1437,1287,2012]" pageId="7" pageNumber="43">
<emphasis id="A9A01B34A32CDB3BFF04EA02FDB0FAAA" bold="true" box="[197,592,1326,1352]" pageId="7" pageNumber="43">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A32CDB3BFD98EA02FCCBFAAA" box="[601,811,1326,1352]" name="Mozambique" pageId="7" pageNumber="43">MOZAMBIQUE</collectingCountry>
: 1 3, Tete, Peters (
<collectionCode id="FDC55FE3A32CDB3BFBD2EA02FBB5FAAA" box="[1043,1109,1326,1352]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="7" pageNumber="43">ZMB</collectionCode>
2179).
<collectingCountry id="E3C387B6A32CDB3BFB72EA02FABAFAAA" box="[1203,1370,1326,1352]" name="Zimbabwe" pageId="7" pageNumber="43">ZIMBABWE</collectingCountry>
: 1 3, Tobwe School, Kamativi [18
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, 27
<geoCoordinate id="FEE0A1E1A32CDB3BFDB8EA79FD21FA90" box="[633,705,1365,1394]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.033333335">0 02'E</geoCoordinate>
],
<date id="EF6AE1E6A32CDB3BFD16EA74FC85FA90" box="[727,869,1368,1394]" pageId="7" pageNumber="43" value="1985-08-03">3.viii.1985</date>
, J. M. Sango (NMZ/D248); 1 3, Rifa Camp [1628B2],
<date id="EF6AE1E6A32CDB3BFED9EA53FE72FA7B" box="[280,402,1407,1433]" pageId="7" pageNumber="43" value="1995-12-08">8.xii.1995</date>
, (NMZ/D799); 1 3, Chapoto Village [1530C2],
<date id="EF6AE1E6A32CDB3BFC14EA53FBB9FA7B" box="[981,1113,1407,1433]" pageId="7" pageNumber="43" value="1997-03-21">21.iii.1997</date>
, S. Nyathi (NMZ/D820); 1 3, Mzola Camp [1827C2],
<date id="EF6AE1E6A32CDB3BFE1BEA84FDB3FA20" box="[474,595,1448,1474]" pageId="7" pageNumber="43" value="1997-12-04">4.xii.1997</date>
, (NMZ/D847); 1 3, Gweru [19
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, 29
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],
<date id="EF6AE1E6A32CDB3BFB40EA84FAE9FA20" box="[1153,1289,1448,1474]" pageId="7" pageNumber="43" value="2002-12-17">17.xii.2002</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFF5EEAFEFF19FA0E" LSID="urn:lsid:biocol.org:col:34985" box="[159,249,1490,1516]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
20500); 1 3, Mount Darwin [16
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, 31
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], 1998, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFB8AEAFEFB45FA0E" LSID="urn:lsid:biocol.org:col:34985" box="[1099,1189,1490,1516]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
22050); 1 3, Karoi [16
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, 29
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], 1999, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFDA9EAD7FD20F9F7" LSID="urn:lsid:biocol.org:col:34985" box="[616,704,1531,1557]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
22048); 1 3, Gokwe [18
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, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFB9CEAD4FB40F9F7" box="[1117,1184,1528,1557]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.13333334">0 08'E</geoCoordinate>
],
<date id="EF6AE1E6A32CDB3BFB74EAD7FAE4F9F7" box="[1205,1284,1531,1557]" pageId="7" pageNumber="43" value="1999-01">i.1999</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFF5EE909FF16F9DD" LSID="urn:lsid:biocol.org:col:34985" box="[159,246,1573,1599]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
20488); 1 3, Chipangali [20
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, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFD71E90EFD12F9DD" box="[688,754,1570,1599]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.6666667">0 40'E</geoCoordinate>
], 2001, M. Perrin (
<collectionCode id="FDC55FE3A32CDB3BFC0FE909FBC5F9DD" LSID="urn:lsid:biocol.org:col:34985" box="[974,1061,1573,1599]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
20416); 2 3, Chihota [18
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, 31
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],
<date id="EF6AE1E6A32CDB3BFECAE963FE64F98B" box="[267,388,1615,1641]" pageId="7" pageNumber="43" value="2006-02-11">11.ii.2006</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFDEBE963FD61F98B" LSID="urn:lsid:biocol.org:col:34985" box="[554,641,1615,1641]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
21938); 2 3, Chegutu [18
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, 30
<geoCoordinate id="FEE0A1E1A32CDB3BFBE7E967FB89F98B" box="[1062,1129,1611,1641]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.33333334">0 20'E</geoCoordinate>
],
<date id="EF6AE1E6A32CDB3BFBBCE963FAE7F98B" box="[1149,1287,1615,1641]" pageId="7" pageNumber="43" value="2008-12-19">19.xii.2008</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFF5EE954FF16F970" LSID="urn:lsid:biocol.org:col:34985" box="[159,246,1656,1682]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
21940); 2 3, Marange [19
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, 32
<geoCoordinate id="FEE0A1E1A32CDB3BFD5EE959FD02F970" box="[671,738,1653,1682]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.6666667">0 40'E</geoCoordinate>
],
<date id="EF6AE1E6A32CDB3BFD37E954FC9FF970" box="[758,895,1656,1682]" pageId="7" pageNumber="43" value="2008-12-23">23.xii.2008</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFBE5E954FB9BF970" LSID="urn:lsid:biocol.org:col:34985" box="[1060,1147,1656,1682]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
21967); 1 3, Bulawayo [20
<geoCoordinate id="FEE0A1E1A32CDB3BFF7FE9B3FF1BF95E" box="[190,251,1695,1724]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.16666667">0 10'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFEE5E9B3FE86F95E" box="[292,358,1695,1724]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.6666667">0 40'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Selous [18
<geoCoordinate id="FEE0A1E1A32CDB3BFBCCE9B3FBAAF95E" box="[1037,1098,1695,1724]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.06666667">0 04'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32CDB3BFBB5E9B3FB59F95E" box="[1140,1209,1695,1724]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.43333334">0 26'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Gokwe [18
<geoCoordinate id="FEE0A1E1A32CDB3BFDA9E9E4FD47F907" box="[616,679,1736,1765]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.033333335">0 02'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFD12E9E4FCF3F907" box="[723,787,1736,1765]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.13333334">0 08'E</geoCoordinate>
],1999, T. Mwabvu (NMZ uncatalogued);1 3, Guruve [16
<geoCoordinate id="FEE0A1E1A32CDB3BFF7FE9DEFF1BF8ED" box="[190,251,1778,1807]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.71666664">0 43'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32CDB3BFEE5E9DEFE85F8ED" box="[292,357,1778,1807]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.3">0 18'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Mushumbi Pools [16
<geoCoordinate id="FEE0A1E1A32CDB3BFB43E9DEFB5FF8ED" box="[1154,1215,1778,1807]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.16666667">0 10'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32CDB3BFB29E9DEFAC8F8ED" box="[1256,1320,1778,1807]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.55">0 33'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Centenary [16
<geoCoordinate id="FEE0A1E1A32CDB3BFD30E837FCCEF8DB" box="[753,814,1819,1849]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.73333335">0 44'S</geoCoordinate>
, 31
<geoCoordinate id="FEE0A1E1A32CDB3BFC99E837FC79F8DB" box="[856,921,1819,1849]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.06666667">0 04'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Raffingora [17
<geoCoordinate id="FEE0A1E1A32CDB3BFEADE869FE4AF880" box="[364,426,1861,1890]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.033333335">0 02'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32CDB3BFE14E869FDF8F880" box="[469,536,1861,1890]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.43333334">0 26'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Mutoko [17
<geoCoordinate id="FEE0A1E1A32CDB3BFB1EE869FAFEF880" box="[1247,1310,1861,1890]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.5">0 30'S</geoCoordinate>
, 32
<geoCoordinate id="FEE0A1E1A32CDB3BFA88E869FA6FF880" box="[1353,1423,1861,1890]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.33333334">0 20'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued); 1 3, Esigodini [20
<geoCoordinate id="FEE0A1E1A32CDB3BFC53E843FC35F86E" box="[914,981,1903,1932]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.28333333">0 17'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFBC5E843FBACF86E" box="[1028,1100,1903,1932]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.93333334">0 56'E</geoCoordinate>
], 1998, T. Mwabvu (NMZ uncatalogued);
<collectingCountry id="E3C387B6A32CDB3BFE97E8B7FE32F857" box="[342,466,1947,1973]" name="Zambia" pageId="7" pageNumber="43">ZAMBIA</collectingCountry>
: 2 3, 5 Ƥ, Lusaka [15
<geoCoordinate id="FEE0A1E1A32CDB3BFD24E8B4FCCAF857" box="[741,810,1944,1973]" direction="south" orientation="latitude" pageId="7" pageNumber="43" precision="925" value="-0.4">0 24'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32CDB3BFC97E8B4FC7AF857" box="[854,922,1944,1973]" direction="east" orientation="longitude" pageId="7" pageNumber="43" precision="925" value="0.25">0 15'E</geoCoordinate>
],
<date id="EF6AE1E6A32CDB3BFC70E8B7FBDFF857" box="[945,1087,1947,1973]" pageId="7" pageNumber="43" value="2008-12-12">12.xii.2008</date>
, M. Bingham &amp; T. Mwabvu (
<collectionCode id="FDC55FE3A32CDB3BFF5EE8EEFF16F83E" LSID="urn:lsid:biocol.org:col:34985" box="[159,246,1986,2012]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="43">NMSA</collectionCode>
22048).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA323DB34FF04EFBBFCEBFEC4" pageId="8" pageNumber="44" type="diagnosis">
<paragraph id="9B6BC726A323DB34FF04EFBBFCEBFEC4" blockId="8.[151,1436,151,294]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04EFBBFEA7FF53" bold="true" box="[197,327,151,177]" pageId="8" pageNumber="44">Diagnosis:</emphasis>
Apical medial edge of proplica extended into a narrow medial tip which touches distal lobe of telocoxite proximally; distal lobe tilted anteriad, lateral width of distal lobe 0.5 times greater than length, apex of distal lobe horizontal (
<figureCitation id="03EFDBA3A323DB34FE0DEFC9FDF6FF1D" box="[460,534,229,255]" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="8" pageNumber="44">Fig. 3</figureCitation>
a). Apical metaplica with horizontal hump, medial hump and central cone reaching apex giving two prong appearance (
<figureCitation id="03EFDBA3A323DB34FD67EE20FD0CFEC4" box="[678,748,268,294]" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="8" pageNumber="44">Fig. 3</figureCitation>
b).
</paragraph>
</subSubSection>
<caption id="CFAB97AEA323DB34FF56EAA3FDC9F9EC" httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="8" pageNumber="44" targetBox="[159,1422,366,1383]" targetPageId="8">
<paragraph id="9B6BC726A323DB34FF56EAA3FDC9F9EC" blockId="8.[151,1436,1423,1550]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF56EAA3FEFCFA45" bold="true" box="[151,284,1423,1447]" pageId="8" pageNumber="44">FIGURE 3.</emphasis>
<taxonomicName id="5CD4BCA5A323DB34FEE2EAA3FE18FA44" box="[291,504,1423,1446]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A323DB34FEE2EAA3FE18FA44" box="[291,504,1423,1446]" italics="true" pageId="8" pageNumber="44">Spirostreptus sebae</emphasis>
</taxonomicName>
(NMZ/D820).
<emphasis id="A9A01B34A323DB34FD61EAA3FD54FA45" bold="true" box="[672,692,1423,1447]" pageId="8" pageNumber="44">a,</emphasis>
oral view of gonopods;
<emphasis id="A9A01B34A323DB34FC00EAA3FC37FA45" bold="true" box="[961,983,1423,1447]" pageId="8" pageNumber="44">b,</emphasis>
aboral view of gonopods;
<emphasis id="A9A01B34A323DB34FB3CEAA3FAF0FA45" bold="true" box="[1277,1296,1423,1447]" pageId="8" pageNumber="44">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="A9A01B34A323DB34FD9BEA9EFD8FFA28" bold="true" box="[602,623,1458,1482]" pageId="8" pageNumber="44">d,</emphasis>
lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe.
</paragraph>
</caption>
<subSubSection id="D3CE94ADA323DB35FF04E910FEFAFDEC" lastPageId="9" lastPageNumber="45" pageId="8" pageNumber="44" type="description">
<paragraph id="9B6BC726A323DB34FF04E910FE1EF99F" blockId="8.[151,1436,1596,2012]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E910FEBFF9B4" bold="true" box="[197,351,1596,1622]" pageId="8" pageNumber="44">Description:</emphasis>
<emphasis id="A9A01B34A323DB34FEA7E911FE75F9B4" box="[358,405,1597,1622]" italics="true" pageId="8" pageNumber="44">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A323DB34FDFAE911FD39F9B5" box="[571,729,1597,1623]" metricMagnitude="-1" metricUnit="m" metricValue="1.885" metricValueMax="2.15" metricValueMin="1.6199999999999999" pageId="8" pageNumber="44" unit="mm" value="188.5" valueMax="215.0" valueMin="162.0">162215 mm</quantity>
; minimum and maximum body width 711.5 mm and
<quantity id="5C2C6AC3A323DB34FA95E911FF28F99F" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="8" pageNumber="44" unit="mm" value="12.5" valueMax="14.0" valueMin="11.0">1114 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A323DB34FE46E94FFE1BF99F" box="[391,507,1635,1661]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="8" pageNumber="44" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E9A6FDC6F946" blockId="8.[151,1436,1596,2012]" box="[197,550,1674,1700]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E9A6FE25F941" box="[197,453,1674,1699]" italics="true" pageId="8" pageNumber="44">Number of body rings</emphasis>
: 5763.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E99DFC1BF929" blockId="8.[151,1436,1596,2012]" box="[197,1019,1713,1739]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E99DFEF8F928" box="[197,280,1713,1738]" italics="true" pageId="8" pageNumber="44">Colour</emphasis>
: Body black or dark brown; legs and antennae brown or black.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E9F4FAC9F913" blockId="8.[151,1436,1596,2012]" box="[197,1321,1751,1777]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E9F4FEFBF913" box="[197,283,1752,1777]" italics="true" pageId="8" pageNumber="44">Collum</emphasis>
: Anterior lobe slightly produced ventrally, with 34 complete and 12 incomplete folds.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E82DFF08F8A0" blockId="8.[151,1436,1596,2012]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E82DFD2DF8F8" box="[197,717,1790,1818]" italics="true" pageId="8" pageNumber="44">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally oval, with a strongly laterally deflected medial nipple.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E863FAA4F88A" blockId="8.[151,1436,1596,2012]" box="[197,1348,1870,1896]" pageId="8" pageNumber="44">
<emphasis id="A9A01B34A323DB34FF04E863FED2F88A" box="[197,306,1871,1896]" italics="true" pageId="8" pageNumber="44">Gonopod</emphasis>
: (
<figureCitation id="03EFDBA3A323DB34FE88E862FE74F88A" box="[329,404,1870,1896]" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="8" pageNumber="44">Figs 3</figureCitation>
a, 3b)
<quantity id="5C2C6AC3A323DB34FE1CE862FDC1F88A" box="[477,545,1870,1896]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="44" unit="mm" value="6.0">6 mm</quantity>
long. Sternite apex extended distally to the level of paracoxite apex.
</paragraph>
<paragraph id="9B6BC726A323DB34FF04E859FC17F86D" blockId="8.[151,1436,1596,2012]" box="[197,1015,1909,1935]" pageId="8" pageNumber="44">Telocoxites approximately parallel, wider apart distal to sternite apex.</paragraph>
<paragraph id="9B6BC726A323DB34FF04E8B7FDD8F83E" blockId="8.[151,1436,1596,2012]" pageId="8" pageNumber="44">Proplica wide sub-apically, medial edge extended apically into a narrow medial tip which may touch the distal lobe of telocoxite proximally.</paragraph>
<paragraph id="9B6BC726A322DB35FF04EFB4FB00FF1D" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">
Metaplica wider proximal to telopodite knee, with humps along the length. Distally metaplica with horizontal hump, raised medial edge and central cone giving a two prong appearance; central cone extending distally to the apex of the distal lobe, the central cone may have perpendicular folds (
<figureCitation id="03EFDBA3A322DB35FBBAEFC9FB21FF1D" box="[1147,1217,229,255]" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="9" pageNumber="45">Fig. 3</figureCitation>
b).
</paragraph>
<paragraph id="9B6BC726A322DB35FF04EE20FE30FEAE" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">Finger-like lateral lobe folded and rounded apically. Lateral lobe extends slightly beyond lateral margins of proplica and distal lobe.</paragraph>
<paragraph id="9B6BC726A322DB35FF04EE75FAFCFE78" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">Lateral edge of distal lobe extending to or beyond the lateral margin of proplica apex. Surface of distal lobe rough, with folds extending to edges, lateral width of distal lobe 0.5 times greater than length.</paragraph>
<paragraph id="9B6BC726A322DB35FF04EE8AFEFAFDEC" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">
Telopodite with antetorsal process extending past paracoxite or may bend medially and crossing the metaplica. Antetorsal process strongly curved into a crescent shape; post knee telopodite with loose coil (
<figureCitation id="03EFDBA3A322DB35FAA6EEE1FF46FDEC" captionStart="FIGURE 3" captionStartId="8.[151,256,1423,1447]" captionTargetBox="[159,1422,366,1383]" captionTargetId="figure@8.[151,1436,349,1399]" captionTargetPageId="8" captionText="FIGURE 3. Spirostreptus sebae (NMZ / D 820). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; dlf, distal lobe fold; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189861/files/figure.png" pageId="9" pageNumber="45">Figs 3</figureCitation>
a, 3b, 7c).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF04ED36FD4DFDB9" pageId="9" pageNumber="45" type="distribution">
<paragraph id="9B6BC726A322DB35FF04ED36FD4DFDB9" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04ED36FE89FDD6" bold="true" box="[197,361,538,564]" pageId="9" pageNumber="45">Distribution:</emphasis>
Known from central, northern, north-eastern, eastern and western parts of
<collectingCountry id="E3C387B6A322DB35FB13ED36FAB1FDD6" box="[1234,1361,538,564]" name="Zimbabwe" pageId="9" pageNumber="45">Zimbabwe</collectingCountry>
, Beira and Tete (
<collectingCountry id="E3C387B6A322DB35FEC6ED6DFE42FDB9" box="[263,418,577,603]" name="Mozambique" pageId="9" pageNumber="45">Mozambique</collectingCountry>
) and Lusaka (
<collectingCountry id="E3C387B6A322DB35FD82ED6DFD41FDB9" box="[579,673,577,603]" name="Zambia" pageId="9" pageNumber="45">Zambia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA322DB35FF04ED4BFD8CFD14" pageId="9" pageNumber="45" type="discussion">
<paragraph id="9B6BC726A322DB35FF04ED4BFD8CFD14" blockId="9.[151,1437,152,758]" pageId="9" pageNumber="45">
<emphasis id="A9A01B34A322DB35FF04ED4BFEA2FD63" bold="true" box="[197,322,615,641]" pageId="9" pageNumber="45">Remarks:</emphasis>
The gonopods of
<taxonomicName id="5CD4BCA5A322DB35FDD9ED44FDBAFD63" box="[536,602,616,641]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A322DB35FDD9ED44FDBAFD63" box="[536,602,616,641]" italics="true" pageId="9" pageNumber="45">sebae</emphasis>
</taxonomicName>
closely resemble those of
<taxonomicName id="5CD4BCA5A322DB35FC50ED44FBE6FD63" box="[913,1030,616,641]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FC50ED44FBE6FD63" box="[913,1030,616,641]" italics="true" pageId="9" pageNumber="45">tripartitus</emphasis>
</taxonomicName>
. In these species the gonopods are smaller and the surface of the distal lobe is rough with several folds, unlike in
<emphasis id="A9A01B34A322DB35FBF9EDA3FA89FD4A" box="[1080,1385,655,680]" italics="true" pageId="9" pageNumber="45">
<taxonomicName id="5CD4BCA5A322DB35FBF9EDA3FB71FD4A" box="[1080,1169,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="unciger">unciger</taxonomicName>
,
<taxonomicName id="5CD4BCA5A322DB35FB5FEDA3FAFDFD4A" box="[1182,1309,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="batokensis">batokensis</taxonomicName>
,
<taxonomicName id="5CD4BCA5A322DB35FAE8EDA3FA89FD4A" box="[1321,1385,655,680]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="heros">heros</taxonomicName>
</emphasis>
and
<taxonomicName id="5CD4BCA5A322DB35FF56ED99FF1BFD2C" box="[151,251,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A322DB35FF56ED99FF1BFD2C" box="[151,251,693,718]" italics="true" pageId="9" pageNumber="45">kruegeri</emphasis>
</taxonomicName>
. Despite the similarities, the distal lobe of
<taxonomicName id="5CD4BCA5A322DB35FD3DED99FC94FD2C" box="[764,884,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FD3DED99FC94FD2C" box="[764,884,693,718]" italics="true" pageId="9" pageNumber="45">tripartitus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5CD4BCA5A322DB35FBD2ED99FBB6FD2C" box="[1043,1110,693,718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="sebae">
<emphasis id="A9A01B34A322DB35FBD2ED99FBB6FD2C" box="[1043,1110,693,718]" italics="true" pageId="9" pageNumber="45">sebae</emphasis>
</taxonomicName>
by being smaller and more rounded (see remarks under
<taxonomicName id="5CD4BCA5A322DB35FE25EDF0FDBBFD17" box="[484,603,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="A9A01B34A322DB35FE25EDF0FDBBFD17" box="[484,603,732,757]" italics="true" pageId="9" pageNumber="45">tripartitus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,409 @@
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<mods:title id="A5E5A36A5FD1F9E3CE2D1733B2C7BA47">A revision of the taxonomy and distribution of Spirostreptus Brandt 1833 (Diplopoda, Spirostreptida, Spirostreptidae) with descriptions of a new species and a new genus of spirostreptid millipede</mods:title>
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<treatment id="137D7630A32EDB3BFF56EB3DFE4FFC55" ID-DOI="http://doi.org/10.5281/zenodo.6220850" ID-GBIF-Taxon="119623015" ID-Zenodo-Dep="6220850" LSID="urn:lsid:plazi:treatment:137D7630A32EDB3BFF56EB3DFE4FFC55" httpUri="http://treatment.plazi.org/id/137D7630A32EDB3BFF56EB3DFE4FFC55" lastPageId="7" lastPageNumber="43" pageId="5" pageNumber="41">
<subSubSection id="D3CE94ADA32EDB39FF56EB3DFEEBFBAC" pageId="5" pageNumber="41" type="nomenclature">
<paragraph id="9B6BC726A32EDB39FF56EB3DFDBAFBC8" blockId="5.[151,602,1040,1102]" box="[151,602,1040,1067]" pageId="5" pageNumber="41">
<heading id="C023704AA32EDB39FF56EB3DFDBAFBC8" bold="true" box="[151,602,1040,1067]" fontSize="11" level="1" pageId="5" pageNumber="41" reason="1">
<taxonomicName id="5CD4BCA5A32EDB39FF56EB3DFDBAFBC8" ID-CoL="4Z8TT" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,602,1040,1067]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FF56EB3DFDBAFBC8" bold="true" box="[151,602,1040,1067]" pageId="5" pageNumber="41">
<emphasis id="A9A01B34A32EDB39FF56EB3DFE44FBC9" bold="true" box="[151,420,1041,1067]" italics="true" pageId="5" pageNumber="41">Spirostreptus kruegeri</emphasis>
(
<bibRefCitation id="FF45BAD7A32EDB39FE72EB3CFDB2FBC8" author="Attems" box="[435,594,1040,1066]" pageId="5" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B6BC726A32EDB39FF56EB18FEEBFBAC" blockId="5.[151,602,1040,1102]" box="[151,267,1076,1102]" pageId="5" pageNumber="41">
<figureCitation id="03EFDBA3A32EDB39FF56EB18FF02FBAC" box="[151,226,1076,1102]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="5" pageNumber="41">Figs 2</figureCitation>
,
<figureCitation id="03EFDBA3A32EDB39FF2FEB18FF1CFBAC" box="[238,252,1076,1102]" captionStart="FIGURE 7" captionStartId="14.[151,255,1491,1515]" captionTargetBox="[191,1393,187,1475]" captionTargetId="figure@14.[185,1407,185,1483]" captionTargetPageId="14" captionText="FIGURE 7. Shape of antetorsal process of the telopodites of Spirostreptus species. a, S. heros; b, S. kruegeri; c, S. sebae; d, S. unciger; e, S. batokensis; f, S. tripartitus. Abbreviations: tp, telopodite; tk, telopodite knee; at, antetorsal process." httpUri="https://zenodo.org/record/189865/files/figure.png" pageId="5" pageNumber="41">7</figureCitation>
b
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32EDB39FF56EBADFD02FB5A" pageId="5" pageNumber="41" type="reference_group">
<paragraph id="9B6BC726A32EDB39FF56EBADFD02FB5A" blockId="5.[151,1259,1153,1208]" pageId="5" pageNumber="41">
<taxonomicName id="5CD4BCA5A32EDB39FF56EBADFDD0FB7A" ID-CoL="58575" authority="Attems 1928" authorityName="Attems" authorityYear="1928" box="[151,560,1153,1176]" class="Diplopoda" family="Spirostreptidae" genus="Triaenostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FF56EBADFE7EFB7A" box="[151,414,1153,1176]" italics="true" pageId="5" pageNumber="41">Triaenostreptus kruegeri</emphasis>
<bibRefCitation id="FF45BAD7A32EDB39FE64EBADFDD0FB7A" author="Attems" box="[421,560,1153,1176]" pageId="5" pageNumber="55" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</taxonomicName>
, p. 367, figs 315317;
<bibRefCitation id="FF45BAD7A32EDB39FCE4EBADFC23FB7A" author="Hoffman" box="[805,963,1153,1176]" pageId="5" pageNumber="56" refString="Hoffman, R. L. (1971) Studies on spirostreptid millipedes. X. Some taxonomic notes on the tribe Triaenostreptini, specific and generic synonomy, and a new species of Triaenostreptus. Revue de Zooogie et de Botanique Africaines, 83 (3 - 4), 207 - 225." type="journal article" year="1971">Hoffman 1971</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32EDB39FC0FEBADFBBDFB7A" author="Krabbe" box="[974,1117,1153,1176]" pageId="5" pageNumber="56" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey, Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="FF45BAD7A32EDB39FBA6EBADFB0BFB7A" author="Hamer" box="[1127,1259,1153,1176]" pageId="5" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
<taxonomicName id="5CD4BCA5A32EDB39FF56EB8DFDD0FB5A" ID-CoL="4Z8TT" authority="Mwabvu 2005" authorityName="Mwabvu" authorityYear="2005" box="[151,560,1185,1208]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32EDB39FF56EB8DFE63FB5A" box="[151,387,1185,1208]" italics="true" pageId="5" pageNumber="41">Spirostreptus kruegeri</emphasis>
:
<bibRefCitation id="FF45BAD7A32EDB39FE53EB8DFDD0FB5A" author="Mwabvu" box="[402,560,1185,1208]" pageId="5" pageNumber="56" refString="Mwabvu, T. (2005 a) New records and localities of spirostreptid millipedes (Myriapoda: Diplopoda) of Zimbabwe with notes on nomenclatural changes and habitat. Arnoldia, Zimbabwe, 10 (25), 249 - 256." type="journal article" year="2005" yearSuffix="a">Mwabvu 2005</bibRefCitation>
</taxonomicName>
a;
<bibRefCitation id="FF45BAD7A32EDB39FD86EB8DFD02FB5A" author="Hoffman" box="[583,738,1185,1208]" pageId="5" pageNumber="56" refString="Hoffman, R. L. (2008) Two new genera of spirostreptid millipeds from central Africa, with revised terminology for male gonopod structures (Diplopoda: Spirostreptida). Tropical Zoology, 21 (2), 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32EDB39FF56EBCBFDADF85B" pageId="5" pageNumber="41" type="materials_examined">
<paragraph id="9B6BC726A32EDB39FF56EBCBFB38FAE0" blockId="5.[151,1437,1255,1977]" box="[151,1240,1255,1282]" pageId="5" pageNumber="41">
<emphasis id="A9A01B34A32EDB39FF56EBCBFEA6FAE3" bold="true" box="[151,326,1255,1281]" pageId="5" pageNumber="41">
<typeStatus id="446F7984A32EDB39FF56EBCBFF32FAE3" box="[151,210,1255,1281]" pageId="5" pageNumber="41">Type</typeStatus>
material
</emphasis>
(not examined):
<collectingCountry id="E3C387B6A32EDB39FDCFEBC4FD07FAE0" box="[526,743,1256,1282]" name="South Africa" pageId="5" pageNumber="41">SOUTH AFRICA</collectingCountry>
:
<emphasis id="A9A01B34A32EDB39FD33EBCBFC33FAE3" bold="true" box="[754,979,1255,1281]" pageId="5" pageNumber="41">Gauteng Province</emphasis>
: Krugersdorp (
<collectionCode id="FDC55FE3A32EDB39FB47EBC4FB2CFAE0" box="[1158,1228,1256,1282]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="5" pageNumber="41">SAM</collectionCode>
).
</paragraph>
<paragraph id="9B6BC726A32EDB39FF04EA3DFDADF85B" blockId="5.[151,1437,1255,1977]" pageId="5" pageNumber="41">
<emphasis id="A9A01B34A32EDB39FF04EA3DFDA5FAC9" bold="true" box="[197,581,1297,1323]" pageId="5" pageNumber="41">Additional material examined:</emphasis>
<collectingCountry id="E3C387B6A32EDB39FD8DEA3DFCC5FAC9" box="[588,805,1297,1323]" name="South Africa" pageId="5" pageNumber="41">SOUTH AFRICA</collectingCountry>
:
<emphasis id="A9A01B34A32EDB39FCF1EA3DFBFBFAC9" bold="true" box="[816,1051,1297,1323]" pageId="5" pageNumber="41">Gauteng Province:</emphasis>
1 3, Pretoria [25
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, 28
<geoCoordinate id="FEE0A1E1A32EDB39FA8EEA22FA6EFAC9" box="[1359,1422,1294,1323]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.18333334">0 11'E</geoCoordinate>
], 1961, (
<collectionCode id="FDC55FE3A32EDB39FF2CEA17FEA7FAB7" LSID="urn:lsid:biocol.org:col:34985" box="[237,327,1339,1365]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
8896).
<emphasis id="A9A01B34A32EDB39FE66EA16FD44FAB6" bold="true" box="[423,676,1338,1364]" pageId="5" pageNumber="41">Limpopo Province:</emphasis>
1 3, Makalali Game Reserve [24
<geoCoordinate id="FEE0A1E1A32EDB39FB89EA14FB68FAB7" box="[1096,1160,1336,1365]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.18333334">0 11'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32EDB39FB74EA14FB1EFAB7" box="[1205,1278,1336,1365]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.6166667">0 37'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FAD5EA17FA8CFAB7" box="[1300,1388,1339,1365]" pageId="5" pageNumber="41" value="1998-10">x.1998</date>
, M. Hamer (
<collectionCode id="FDC55FE3A32EDB39FF36EA4EFEAEFA9E" LSID="urn:lsid:biocol.org:col:34985" box="[247,334,1378,1404]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
22051); 1 3, Klipfontein, N. Transvaal [2428A2],
<date id="EF6AE1E6A32EDB39FC72EA4EFBCBFA9E" box="[947,1067,1378,1404]" pageId="5" pageNumber="41" value="1979-12-04">4.xii.1979</date>
(NMZ/D97); 1 3, Phalaborwa [23
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, 31
<geoCoordinate id="FEE0A1E1A32EDB39FEE6EAA4FE85FA47" box="[295,357,1416,1445]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.2">0 12'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FE46EAA7FE34FA47" box="[391,468,1419,1445]" pageId="5" pageNumber="41" value="1998-01">i.1998</date>
, S. Piper (
<collectionCode id="FDC55FE3A32EDB39FD90EAA7FD4CFA47" LSID="urn:lsid:biocol.org:col:34985" box="[593,684,1419,1445]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
16196).
<emphasis id="A9A01B34A32EDB39FCCDEAA7FBFFFA47" bold="true" box="[780,1055,1419,1445]" pageId="5" pageNumber="41">North West Province:</emphasis>
1 3, Pilanesberg National Park [25
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, 27
<geoCoordinate id="FEE0A1E1A32EDB39FEE4EA9EFE8AFA2D" box="[293,362,1458,1487]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.1">0 06'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FEBFEA99FE15FA2D" box="[382,501,1461,1487]" pageId="5" pageNumber="41" value="2000-10-30">30.x.2000</date>
, R. Slotow (
<collectionCode id="FDC55FE3A32EDB39FD46EA99FD3EFA2D" LSID="urn:lsid:biocol.org:col:34985" box="[647,734,1461,1487]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
15816); 1 3, Pilanesberg National Park [25
<geoCoordinate id="FEE0A1E1A32EDB39FB22EA9EFAC0FA2D" box="[1251,1312,1458,1487]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.25">0 15'S</geoCoordinate>
, 27
<geoCoordinate id="FEE0A1E1A32EDB39FA8BEA9EFA70FA2D" box="[1354,1424,1458,1487]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.1">0 06'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FF56EAF2FEC2FA1A" box="[151,290,1502,1528]" pageId="5" pageNumber="41" value="1999-12-12">12.xii.1999</date>
, R. Slotow (
<collectionCode id="FDC55FE3A32EDB39FE79EAF2FDF0FA1A" LSID="urn:lsid:biocol.org:col:34985" box="[440,528,1502,1528]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20478); 1 3, Pilanesberg National Park [25
<geoCoordinate id="FEE0A1E1A32EDB39FBE7EAF7FB85FA1A" box="[1062,1125,1499,1528]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.25">0 15'S</geoCoordinate>
, 27
<geoCoordinate id="FEE0A1E1A32EDB39FB50EAF7FB38FA1A" box="[1169,1240,1499,1528]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.1">0 06'E</geoCoordinate>
], 1999, J. Londt (
<collectionCode id="FDC55FE3A32EDB39FF5EE924FF19F9C0" LSID="urn:lsid:biocol.org:col:34985" box="[159,249,1544,1570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20513); 1 3, Pilanesberg [25
<geoCoordinate id="FEE0A1E1A32EDB39FDAEE929FD51F9C0" box="[623,689,1541,1570]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.25">0 15'S</geoCoordinate>
, 27
<geoCoordinate id="FEE0A1E1A32EDB39FD1FE929FCC4F9C0" box="[734,804,1541,1570]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.1">0 06'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FCFAE924FC54F9C0" box="[827,948,1544,1570]" pageId="5" pageNumber="41" value="1999-01-21">21.i.1999</date>
, (
<collectionCode id="FDC55FE3A32EDB39FC0BE924FBC9F9C0" LSID="urn:lsid:biocol.org:col:34985" box="[970,1065,1544,1570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
15816); 1 3, Botsalano Game Reserve [25
<geoCoordinate id="FEE0A1E1A32EDB39FEEAE902FE8BF9AE" box="[299,363,1582,1612]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.56666666">0 34'S</geoCoordinate>
, 25
<geoCoordinate id="FEE0A1E1A32EDB39FE59E902FE02F9AE" box="[408,482,1582,1612]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.7">0 42'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FE36E91EFDBFF9AE" box="[503,607,1586,1612]" pageId="5" pageNumber="41" value="2000-01-07">7.i.2000</date>
, (
<collectionCode id="FDC55FE3A32EDB39FDB4E91EFD2EF9AE" LSID="urn:lsid:biocol.org:col:34985" box="[629,718,1586,1612]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
22053); 1 3, Madikwe Game Reserve [25
<geoCoordinate id="FEE0A1E1A32EDB39FB1FE902FAFEF9AE" box="[1246,1310,1582,1612]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.35">0 21'S</geoCoordinate>
, 26
<geoCoordinate id="FEE0A1E1A32EDB39FA8AE902FA6FF9AE" box="[1355,1423,1582,1612]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.53333336">0 32'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FF56E977FF16F997" box="[151,246,1627,1653]" pageId="5" pageNumber="41" value="2000-03">iii.2000</date>
, R. Slotow (
<collectionCode id="FDC55FE3A32EDB39FE4DE977FE04F997" LSID="urn:lsid:biocol.org:col:34985" box="[396,484,1627,1653]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20497); 1 3, Madikwe [25
<geoCoordinate id="FEE0A1E1A32EDB39FCF1E974FC8FF997" box="[816,879,1624,1653]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.35">0 21'S</geoCoordinate>
, 26
<geoCoordinate id="FEE0A1E1A32EDB39FC5AE974FC3DF997" box="[923,989,1624,1653]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.53333336">0 32'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FC32E977FBAAF997" box="[1011,1098,1627,1653]" pageId="5" pageNumber="41" value="2000-02">ii.2000</date>
, R. Slotow (
<collectionCode id="FDC55FE3A32EDB39FB21E977FAD8F997" LSID="urn:lsid:biocol.org:col:34985" box="[1248,1336,1627,1653]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20479).
<emphasis id="A9A01B34A32EDB39FF56E9A8FE35F97C" bold="true" box="[151,469,1668,1694]" pageId="5" pageNumber="41">Northern Cape Province:</emphasis>
1 3, Northern Cape [28.577 0S, 22.489 0E], (
<collectionCode id="FDC55FE3A32EDB39FC1AE9A9FBD2F97D" LSID="urn:lsid:biocol.org:col:34985" box="[987,1074,1669,1695]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
22052).
<collectingCountry id="E3C387B6A32EDB39FB5DE9A9FAC1F97D" box="[1180,1313,1669,1695]" name="Namibia" pageId="5" pageNumber="41">NAMIBIA</collectingCountry>
: 1 3, 1 Ƥ, Palmfontain Farm [1914CB],
<date id="EF6AE1E6A32EDB39FE3BE987FD94F927" box="[506,628,1707,1733]" pageId="5" pageNumber="41" value="1969-02-25">25.ii.1969</date>
, B. Lamoral (
<collectionCode id="FDC55FE3A32EDB39FCD9E987FC8FF927" LSID="urn:lsid:biocol.org:col:34985" box="[792,879,1707,1733]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
9965).
<collectingCountry id="E3C387B6A32EDB39FC0AE987FB90F927" box="[971,1136,1707,1733]" name="Zimbabwe" pageId="5" pageNumber="41">ZIMBABWE</collectingCountry>
: 1 3, Humani Ranch HQ [2032A3],
<date id="EF6AE1E6A32EDB39FED6E9F9FE72F90D" box="[279,402,1749,1775]" pageId="5" pageNumber="41" value="1987-04-13">13.iv.1987</date>
(NMZ/D502); 1 3, Maphisa [21
<geoCoordinate id="FEE0A1E1A32EDB39FCD3E9FEFCAFF90D" box="[786,847,1746,1775]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.0">0 00'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32EDB39FCB9E9FEFC59F90D" box="[888,953,1746,1775]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.45">0 27'E</geoCoordinate>
], 1998, T. Mwabvu (
<collectionCode id="FDC55FE3A32EDB39FB6BE9F9FAE1F90D" LSID="urn:lsid:biocol.org:col:34985" box="[1194,1281,1749,1775]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20539); 1 3, Doddieburn Ranch, West Nicholson [2129A4],
<date id="EF6AE1E6A32EDB39FD0BE9D0FCD5F8F4" box="[714,821,1788,1814]" pageId="5" pageNumber="41" value="1985-12-18">18.xii.85</date>
, (NMZ/D362, NM/D381); 1 3, Doddieburn Ranch, West Nicholson [2129A4],
<date id="EF6AE1E6A32EDB39FE31E809FDAAF8DD" box="[496,586,1829,1855]" pageId="5" pageNumber="41" value="1986-09-05">5.ix.86</date>
, F. Nyathi (NMZ/D387); 1 3, Manana Hospital [20
<geoCoordinate id="FEE0A1E1A32EDB39FB1BE80EFAFBF8DD" box="[1242,1307,1826,1855]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.93333334">0 56'S</geoCoordinate>
, 29
<geoCoordinate id="FEE0A1E1A32EDB39FA88E80EFA6FF8DD" box="[1353,1423,1826,1855]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.0">0 00'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FF56E863FEC3F88B" box="[151,291,1871,1897]" pageId="5" pageNumber="41" value="1996-12-14">14.12.1996</date>
, F. Nyathi (NMZ/D838); 1 3, Hippo Valley [21
<geoCoordinate id="FEE0A1E1A32EDB39FCB2E860FC53F88B" box="[883,947,1868,1897]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.16666667">0 10'S</geoCoordinate>
, 31
<geoCoordinate id="FEE0A1E1A32EDB39FC1EE860FBC4F88B" box="[991,1060,1868,1897]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.53333336">0 32'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FBF8E863FB79F88B" box="[1081,1177,1871,1897]" pageId="5" pageNumber="41" value="1998-11">xi.1998</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32EDB39FA82E863FA7CF88B" LSID="urn:lsid:biocol.org:col:34985" box="[1347,1436,1871,1897]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
20543); 1 3, West Nicholson [21
<geoCoordinate id="FEE0A1E1A32EDB39FDE6E859FD86F871" box="[551,614,1909,1939]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.05">0 03'S</geoCoordinate>
, 28
<geoCoordinate id="FEE0A1E1A32EDB39FD50E859FD33F871" box="[657,723,1909,1939]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.35">0 21'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FD28E855FC92F871" box="[745,882,1913,1939]" pageId="5" pageNumber="41" value="1974-12-17">17.xii.1974</date>
(NMZ/D574); 1 3, Chivi [20
<geoCoordinate id="FEE0A1E1A32EDB39FB1AE859FAFAF871" box="[1243,1306,1909,1939]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="925" value="-0.5">0 30'S</geoCoordinate>
, 30
<geoCoordinate id="FEE0A1E1A32EDB39FA85E859FA62F871" box="[1348,1410,1909,1939]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="925" value="0.8333333">0 50'E</geoCoordinate>
],
<date id="EF6AE1E6A32EDB39FF56E8B3FF12F85B" box="[151,242,1951,1977]" pageId="5" pageNumber="41" value="1998-11">xi.1998</date>
, T. Mwabvu (
<collectionCode id="FDC55FE3A32EDB39FE55E8B3FE0BF85B" LSID="urn:lsid:biocol.org:col:34985" box="[404,491,1951,1977]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="5" pageNumber="41">NMSA</collectionCode>
16196).
</paragraph>
</subSubSection>
<caption id="CFAB97AEA32DDB3AFF56EBF6FDE8FABB" httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="6" pageNumber="42" targetBox="[172,1428,204,1223]" targetPageId="6">
<paragraph id="9B6BC726A32DDB3AFF56EBF6FDE8FABB" blockId="6.[151,1436,1242,1369]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF56EBF6FEF9FB10" bold="true" box="[151,281,1242,1266]" pageId="6" pageNumber="42">FIGURE 2.</emphasis>
<taxonomicName id="5CD4BCA5A32DDB3AFEE0EBF6FDEEFB13" box="[289,526,1242,1265]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32DDB3AFEE0EBF6FDEEFB13" box="[289,526,1242,1265]" italics="true" pageId="6" pageNumber="42">Spirostreptus kruegeri</emphasis>
</taxonomicName>
(NMZ/D362).
<emphasis id="A9A01B34A32DDB3AFD72EBF6FD27FB10" bold="true" box="[691,711,1242,1266]" pageId="6" pageNumber="42">a,</emphasis>
oral view of gonopods;
<emphasis id="A9A01B34A32DDB3AFC0DEBF6FC02FB10" bold="true" box="[972,994,1242,1266]" pageId="6" pageNumber="42">b,</emphasis>
aboral view of gonopods;
<emphasis id="A9A01B34A32DDB3AFAC0EBF6FAF3FB10" bold="true" box="[1281,1299,1242,1266]" pageId="6" pageNumber="42">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="A9A01B34A32DDB3AFD9BEBD0FD8FFAF6" bold="true" box="[602,623,1276,1300]" pageId="6" pageNumber="42">d,</emphasis>
lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe.
</paragraph>
</caption>
<subSubSection id="D3CE94ADA32DDB3AFF04EAABFCB7F9F7" pageId="6" pageNumber="42" type="diagnosis">
<paragraph id="9B6BC726A32DDB3AFF04EAABFCB7F9F7" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04EAABFEAFFA43" bold="true" box="[197,335,1415,1441]" pageId="6" pageNumber="42">Diagnosis:</emphasis>
Telocoxites wider apart at the level of the apex of the proplica; proplica and metaplica abruptly widen subapically; lateral lobe may have an apical nipple; lateral lobe strongly slanting towards paracoxite away from the distal lobe, at greater than right angle; apically lateral lobe deflected aborally; antetorsal process crescent-shaped distally (
<figureCitation id="03EFDBA3A32DDB3AFD56EAD7FD3EF9F7" box="[663,734,1531,1557]" captionStart="FIGURE 1" captionStartId="4.[151,255,1339,1363]" captionTargetBox="[151,1427,209,1300]" captionTargetId="figure@4.[151,1436,197,1314]" captionTargetPageId="4" captionText="FIGURE 1. Spirostreptus heros (NMSA 20495). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189859/files/figure.png" pageId="6" pageNumber="42">Figs 1</figureCitation>
a, 1b, 7b).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32DDB3BFF04E90EFED3FE78" lastPageId="7" lastPageNumber="43" pageId="6" pageNumber="42" type="description">
<paragraph id="9B6BC726A32DDB3AFF04E90EFE1EF981" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E90EFE82F9DE" bold="true" box="[197,354,1570,1596]" pageId="6" pageNumber="42">Description:</emphasis>
<emphasis id="A9A01B34A32DDB3AFEABE90EFE7AF9D9" box="[362,410,1570,1595]" italics="true" pageId="6" pageNumber="42">Size</emphasis>
: Body length
<quantity id="5C2C6AC3A32DDB3AFD87E90EFD09F9DE" box="[582,745,1570,1596]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="6" pageNumber="42" unit="mm" value="230.0" valueMax="240.0" valueMin="220.0">220240 mm</quantity>
; minimum and maximum body width
<quantity id="5C2C6AC3A32DDB3AFB75E90EFAF7F9DE" box="[1204,1303,1570,1596]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="6" pageNumber="42" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">59 mm</quantity>
and
<quantity id="5C2C6AC3A32DDB3AFA92E90EFF28F981" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="6" pageNumber="42" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">1115 mm</quantity>
; antenna length
<quantity id="5C2C6AC3A32DDB3AFE46E965FE1BF981" box="[391,507,1609,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="6" pageNumber="42" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">912 mm</quantity>
.
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E95CFDC6F96B" blockId="6.[151,1437,1415,2024]" box="[197,550,1647,1673]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E95CFE25F96B" box="[197,453,1648,1673]" italics="true" pageId="6" pageNumber="42">Number of body rings</emphasis>
: 6373.
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E9BAFB83F952" blockId="6.[151,1437,1415,2024]" box="[197,1123,1686,1712]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E9BAFEF8F94D" box="[197,280,1686,1711]" italics="true" pageId="6" pageNumber="42">Colour</emphasis>
: Body brown and black; legs reddish brown or brown; antennae brown.
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E991FEE5F91F" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E991FEFCF934" box="[197,284,1725,1750]" italics="true" pageId="6" pageNumber="42">Collum</emphasis>
: Rounded anterior lobe slightly produced ventrally, with 34 complete and 07 incomplete folds (
<figureCitation id="03EFDBA3A32DDB3AFF5EE9CFFF06F91F" box="[159,230,1763,1789]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="6" pageNumber="42">Fig. 2</figureCitation>
d).
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E821FE7AF8AC" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E821FD5DF8C4" box="[197,701,1802,1830]" italics="true" pageId="6" pageNumber="42">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally parallel with a short and broad, laterally deflected distal nipple (
<figureCitation id="03EFDBA3A32DDB3AFEF6E818FE9FF8AC" box="[311,383,1844,1870]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="6" pageNumber="42">Fig. 2</figureCitation>
c).
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E877FDB2F879" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">
<emphasis id="A9A01B34A32DDB3AFF04E877FED2F896" box="[197,306,1883,1908]" italics="true" pageId="6" pageNumber="42">Gonopod</emphasis>
: (
<figureCitation id="03EFDBA3A32DDB3AFE88E876FE74F896" box="[329,404,1882,1908]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="6" pageNumber="42">Figs 2</figureCitation>
a, 2b)
<quantity id="5C2C6AC3A32DDB3AFE1CE876FDDEF896" box="[477,574,1882,1908]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="42" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
long. Sternite long, reaching level of paracoxite apex. Telocoxites further apart at the level of the proplica apex.
</paragraph>
<paragraph id="9B6BC726A32DDB3AFF04E884FBDFF80A" blockId="6.[151,1437,1415,2024]" pageId="6" pageNumber="42">Proplica widens abruptly subapically and tapers apically. Raised lateral edge of proplica pronounced and extending further than medial edge. Proplica concealing lateral lobe proximally.</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EFB4FA9DFF50" blockId="7.[151,1437,152,951]" box="[197,1405,152,178]" pageId="7" pageNumber="43">Metaplica without lobes, wider with humps proximal to lateral lobe, and with a prominent distal hump.</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EF92FB89FEC4" blockId="7.[151,1437,152,951]" pageId="7" pageNumber="43">
Lateral lobe strongly slanting towards paracoxite (away from distal lobe) and extending beyond lateral edges of proplica and distal lobe, proximally constricted and rounded distally (
<figureCitation id="03EFDBA3A32CDB3BFBF1EFC9FB96FF1D" box="[1072,1142,229,255]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="7" pageNumber="43">Fig. 2</figureCitation>
b). Lateral lobe deflected aborally, may end in a prominent apical nipple which is clearly seen in aboral view.
</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EE1EFACBFEAE" blockId="7.[151,1437,152,951]" box="[197,1323,306,332]" pageId="7" pageNumber="43">Lateral and medial edges of distal lobe rounded distally, with shallow depression on the surface.</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EE75FED3FE78" blockId="7.[151,1437,152,951]" pageId="7" pageNumber="43">
Telopodite with open coil or spiral, antetorsal process crescent-shaped distally, may reach paracoxite apex (
<figureCitation id="03EFDBA3A32CDB3BFF5EEEACFF0AFE78" box="[159,234,384,410]" captionStart="FIGURE 2" captionStartId="6.[151,255,1242,1266]" captionTargetBox="[172,1428,204,1223]" captionTargetId="figure@6.[151,1436,194,1240]" captionTargetPageId="6" captionText="FIGURE 2. Spirostreptus kruegeri (NMZ / D 362). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: p, proplica; m, metaplica; llb, lateral lobe; dl, distal lobe; d, depression; tp, telopodite; at, antetorsal process; pc, paracoxite; st, sternite; cf, collum fold; al, anterior lobe; pfl, prefemoral lobe." httpUri="https://zenodo.org/record/189860/files/figure.png" pageId="7" pageNumber="43">Figs 2</figureCitation>
a, 2b).
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32CDB3BFF04EE8AFE7AFE05" pageId="7" pageNumber="43" type="distribution">
<paragraph id="9B6BC726A32CDB3BFF04EE8AFE7AFE05" blockId="7.[151,1437,152,951]" pageId="7" pageNumber="43">
<emphasis id="A9A01B34A32CDB3BFF04EE8AFE87FE22" bold="true" box="[197,359,422,448]" pageId="7" pageNumber="43">Distribution:</emphasis>
Known from north and north-central
<collectingCountry id="E3C387B6A32CDB3BFCD9EE8AFC4EFE22" box="[792,942,422,448]" name="South Africa" pageId="7" pageNumber="43">South Africa</collectingCountry>
, southern
<collectingCountry id="E3C387B6A32CDB3BFBE5EE8AFB42FE22" box="[1060,1186,422,448]" name="Zimbabwe" pageId="7" pageNumber="43">Zimbabwe</collectingCountry>
, Gaberone,
<collectingCountry id="E3C387B6A32CDB3BFAEBEE8AFA7BFE22" box="[1322,1435,422,448]" name="Botswana" pageId="7" pageNumber="43">Botswana</collectingCountry>
and northern
<collectingCountry id="E3C387B6A32CDB3BFEEEEEE1FE76FE05" box="[303,406,461,487]" name="Namibia" pageId="7" pageNumber="43">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D3CE94ADA32CDB3BFF04EEDFFE4FFC55" pageId="7" pageNumber="43" type="discussion">
<paragraph id="9B6BC726A32CDB3BFF04EEDFFB84FCFE" blockId="7.[151,1437,152,951]" pageId="7" pageNumber="43">
<emphasis id="A9A01B34A32CDB3BFF04EEDFFEA0FDEF" bold="true" box="[197,320,499,525]" pageId="7" pageNumber="43">Remarks:</emphasis>
The texture of the distal lobe of the gonopod is similar to that of
<taxonomicName id="5CD4BCA5A32CDB3BFBF8EED8FB54FDEF" box="[1081,1204,500,525]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="batokensis">
<emphasis id="A9A01B34A32CDB3BFBF8EED8FB54FDEF" box="[1081,1204,500,525]" italics="true" pageId="7" pageNumber="43">batokensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5CD4BCA5A32CDB3BFB02EED8FAFCFDEF" box="[1219,1308,500,525]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="A9A01B34A32CDB3BFB02EED8FAFCFDEF" box="[1219,1308,500,525]" italics="true" pageId="7" pageNumber="43">unciger</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32CDB3BFA95EED8FA74FDEF" box="[1364,1428,500,525]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFA95EED8FA74FDEF" box="[1364,1428,500,525]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
, however the resemblance to
<taxonomicName id="5CD4BCA5A32CDB3BFE33ED36FDD4FDD1" box="[498,564,538,563]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFE33ED36FDD4FDD1" box="[498,564,538,563]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
is greater. Besides having overlapping distributions,
<taxonomicName id="5CD4BCA5A32CDB3BFB78ED36FB1BFDD1" box="[1209,1275,538,563]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFB78ED36FB1BFDD1" box="[1209,1275,538,563]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32CDB3BFAF7ED36FA7CFDD1" box="[1334,1436,538,563]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32CDB3BFAF7ED36FA7CFDD1" box="[1334,1436,538,563]" italics="true" pageId="7" pageNumber="43">kruegeri</emphasis>
</taxonomicName>
have similar body and gonopod characteristics, particularly the shape and surface texture of the distal telocoxite lobe. The major differences in the gonopods of
<taxonomicName id="5CD4BCA5A32CDB3BFCB8ED44FC5EFD63" box="[889,958,616,641]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFCB8ED44FC5EFD63" box="[889,958,616,641]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD4BCA5A32CDB3BFC3DED44FB87FD63" box="[1020,1127,616,641]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32CDB3BFC3DED44FB87FD63" box="[1020,1127,616,641]" italics="true" pageId="7" pageNumber="43">kruegeri</emphasis>
</taxonomicName>
are the proximity of the telocoxites, the shapes of the subapical proplica and antetorsal process, the angle between the lateral and distal lobes, the height of the sternite in relation to the paracoxite apex, and the nature of the telopodite coil. Subapically the proplica is abruptly wider and angular in
<taxonomicName id="5CD4BCA5A32CDB3BFC81EDF0FC43FD17" box="[832,931,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32CDB3BFC81EDF0FC43FD17" box="[832,931,732,757]" italics="true" pageId="7" pageNumber="43">kruegeri</emphasis>
</taxonomicName>
, unlike in
<taxonomicName id="5CD4BCA5A32CDB3BFBE2EDF0FB83FD17" box="[1059,1123,732,757]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFBE2EDF0FB83FD17" box="[1059,1123,732,757]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
. A difference in the width of the metaplica is also evident between the two species (see remarks under
<taxonomicName id="5CD4BCA5A32CDB3BFBD2EC2EFBB3FCF9" box="[1043,1107,770,795]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="A9A01B34A32CDB3BFBD2EC2EFBB3FCF9" box="[1043,1107,770,795]" italics="true" pageId="7" pageNumber="43">heros</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9B6BC726A32CDB3BFF04EC05FE4FFC55" blockId="7.[151,1437,152,951]" pageId="7" pageNumber="43">
According to the literature (
<bibRefCitation id="FF45BAD7A32CDB3BFDD0EC05FD49FCA1" author="Hamer" box="[529,681,809,835]" pageId="7" pageNumber="55" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
), the
<typeStatus id="446F7984A32CDB3BFD2CEC05FCC0FCA1" box="[749,800,809,835]" pageId="7" pageNumber="43">type</typeStatus>
material of
<taxonomicName id="5CD4BCA5A32CDB3BFC72EC05FBD7FCA0" box="[947,1079,809,834]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="A9A01B34A32CDB3BFC72EC05FBD7FCA0" box="[947,1079,809,834]" italics="true" pageId="7" pageNumber="43">S. kruegeri</emphasis>
</taxonomicName>
is in the SAM. However, this material is missing, presumed lost, from the SAM collection, and is not in other collections in the southern African region. Suggestions that the material could be in the Naturhistorisches Museum, Vienna,
<collectingCountry id="E3C387B6A32CDB3BFA82EC5AFA7CFC72" box="[1347,1436,886,912]" name="Austria" pageId="7" pageNumber="43">Austria</collectingCountry>
could not be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Thomson, 1895" authorityName="Thomson" authorityYear="1895" class="Insecta" family="Braconidae" genus="Exotela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exotela melanocera" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="melanocera">Exotela melanocera (Thomson, 1895)</taxonomicName>
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<taxonomicName class="Insecta" family="Braconidae" genus="Dacnusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dacnusa melanocera" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="melanocera">Dacnusa melanocera</taxonomicName>
Thomson, 1895
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(Nixon, 1954,
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)
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<mods:namePart id="814B75058E5EB7B747B1224EC2F14C42">Landolt, Elias</mods:namePart>
</mods:name>
<mods:name id="7C90CE7BA2266336303297DC133A0B01" type="personal">
<mods:role id="BF33215FEA40452268D262C10568B50C">
<mods:roleTerm id="10BB2B43269DE3031AEA03B74EB1FDCD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="038E0DC36918CCE4E162D8CD73B507DE">Hirzel, Rosmarie</mods:namePart>
</mods:name>
<mods:typeOfResource id="CEDD5F9AD7018749E04B164684BE83A0">text</mods:typeOfResource>
<mods:originInfo id="E27D1D4B6FF42049619CF60E9D70DF1C">
<mods:dateIssued id="21DEFDE51168DD3B2EF128623AB937B2">1972</mods:dateIssued>
<mods:publisher id="52C0FEACBF860673338282901F5D9A3E">Birkhaeuser Verlag</mods:publisher>
</mods:originInfo>
<mods:location id="9363CE90F51B36E66CB0E2E68C8742A4">
<mods:url id="9B4120D75DD4BDD1D2DD43440918F7FE">https://doi.org/10.5281/zenodo.291815</mods:url>
</mods:location>
<mods:classification id="2099D45BE70A90A6AF70FDE7CFB556D1">book</mods:classification>
<mods:identifier id="FC564B3884BD03D9C8355A4E40CFD708" type="Zenodo-Dep">291815</mods:identifier>
<mods:identifier id="8D2744D5619E0161E37E926163169FA0" type="DOI">10.5281/zenodo.291815</mods:identifier>
<mods:identifier id="69B40A9F7244BBFD6BB8A23B9218C633" type="ISBN">3-7643-0843-5</mods:identifier>
</mods:mods>
<treatment id="137DBEF8D6B81EA5E3B5EC52180989FF" LSID="urn:lsid:plazi:treatment:137DBEF8D6B81EA5E3B5EC52180989FF" httpUri="http://treatment.plazi.org/id/137DBEF8D6B81EA5E3B5EC52180989FF" lastPageNumber="440" pageId="null" pageNumber="440" sec="registerzuband1.s742">
<title id="60B9CC061207C99804B8C9F63DD1A48C" pageId="null" pageNumber="440">
<subSubSection id="329FF9F97CF0638B976D3F7995E0569E" pageId="null" pageNumber="440" type="nomenclature">
<paragraph id="78209A7D8322F5B7A39EEC4830201EF4" pageId="null" pageNumber="440">
<taxonomicName id="F5126B6764EF7529D8FB5E274F6B136A" ID-CoL="R9M6" ID-ENA="240690" authority="Ehrh." class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="null" pageNumber="440" phylum="Tracheophyta" rank="species" species="heleonastes">
<pageBreakToken id="307A3D34B24C4A4742E2ACBBC33A0A3F" pageId="null" pageNumber="440" start="start">Carex</pageBreakToken>
<normalizedToken id="67EE8345F455E048652E78A9AE0FE5E1" originalValue="Heleonástes" pageId="null" pageNumber="440">Heleonastes</normalizedToken>
Ehrh.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="81C2D72F900EE9BE6C534BE574F6117B" pageId="null" pageNumber="440" type="vernacular_names">
<paragraph id="C2DC82C9CA6EBBB25A5EC6D905B4F4E9" pageId="null" pageNumber="440">Sumpfbewohnende Segge</paragraph>
</subSubSection>
</title>
<subSubSection id="006B36D91AD01D050197FC7BFA89A6DE" pageId="null" pageNumber="440" type="description">
<paragraph id="20DF2E7CA65E2BFD2CDADD8763926C49" pageId="null" pageNumber="440">
10-30 cm hoch; lockere Horste mit kurzen
<normalizedToken id="02C921B8D42788348CDF99662728A44E" originalValue="Ausläufern" pageId="null" pageNumber="440">Auslaeufern</normalizedToken>
bildend.
<normalizedToken id="4B9E93C766D5C367E1156980FBDF986D" originalValue="Grundständige" pageId="null" pageNumber="440">Grundstaendige</normalizedToken>
Scheiden braun.
<normalizedToken id="336E3C0E6E4337C359C251FFF7F6489B" originalValue="Blätter" pageId="null" pageNumber="440">Blaetter</normalizedToken>
1,5-2 mm breit,
<emphasis id="1E1582DEB068C6B1607E013F934CB368" italics="true" pageId="null" pageNumber="440">rinnig gefaltet</emphasis>
, steif,
<emphasis id="B5794D54DE5FAF4ECB571D72C24E4A88" italics="true" pageId="null" pageNumber="440">
<normalizedToken id="43C79E12076D9FFEFE14FE3C81523C40" originalValue="graugrün" pageId="null" pageNumber="440">graugruen</normalizedToken>
</emphasis>
, den
<normalizedToken id="3FE1E29B415862D4A4800640C15EFA9B" originalValue="Blütenstand" pageId="null" pageNumber="440">Bluetenstand</normalizedToken>
meist nicht erreichend. Stengel 3kantig,
<emphasis id="59571F856A7808B77FA78EE6285AF174" italics="true" pageId="null" pageNumber="440">
in der obern
<normalizedToken id="D4A88FD93F47437E37679EE58015ACA1" originalValue="Hälfte" pageId="null" pageNumber="440">Haelfte</normalizedToken>
stark rauh
</emphasis>
, steif aufrecht.
<normalizedToken id="B39EB647BA62B37240D3723900B5C7D5" originalValue="Blütenstand" pageId="null" pageNumber="440">Bluetenstand</normalizedToken>
1-2 cm lang, meist kopfig, aus 3-5
<normalizedToken id="F90C5111DB3B48E7B13130A9DC65E20C" originalValue="Ähren" pageId="null" pageNumber="440">Aehren</normalizedToken>
bestehend;
<normalizedToken id="52351BCC7CEF7EDE44AB0DFE93697694" originalValue="Ähren" pageId="null" pageNumber="440">Aehren</normalizedToken>
3-7 mm lang, alle unten ♂, oben ♀.
<normalizedToken id="93A9CC5FDC031FE8F6E4758CA0DCEE01" originalValue="Hochblätter" pageId="null" pageNumber="440">Hochblaetter</normalizedToken>
<normalizedToken id="C26942D3C6FD7AB14B19D0AE01D0719F" originalValue="tragblattähnlich" pageId="null" pageNumber="440">tragblattaehnlich</normalizedToken>
, auch
<normalizedToken id="D58A75F03B0C67EFC87FEE5BF18B0C37" originalValue="blattähnlich" pageId="null" pageNumber="440">blattaehnlich</normalizedToken>
und dann den
<normalizedToken id="F7B440AA5B4C47A03EF50FF5F7116B38" originalValue="Blütenstand" pageId="null" pageNumber="440">Bluetenstand</normalizedToken>
<normalizedToken id="D338D6D8464E27D01F4E25EB094A2393" originalValue="überragend" pageId="null" pageNumber="440">ueberragend</normalizedToken>
(oft beide Typen an derselben Pflanze).
<normalizedToken id="5E9851D34ADE9B5191C8D4E12064B5DD" originalValue="Tragblätter" pageId="null" pageNumber="440">Tragblaetter</normalizedToken>
⅔-
<normalizedToken id="B2B377CB6258631FFC67726F0446AD8E" originalValue="¾" pageId="null" pageNumber="440">3/4</normalizedToken>
so lang wie die reifen
<normalizedToken id="AEFB9B17A3E2F1595A4F46A989311F67" originalValue="Fruchtschläuche" pageId="null" pageNumber="440">Fruchtschlaeuche</normalizedToken>
, mit stumpfer Spitze, braun, mit breitem,
<normalizedToken id="936430BD9967F90F1C6205FEB4EB7B57" originalValue="weißem" pageId="null" pageNumber="440">weissem</normalizedToken>
Rand.
<normalizedToken id="42F087F3FC39B23F4AA5A52778031E72" originalValue="Fruchtschläuche" pageId="null" pageNumber="440">Fruchtschlaeuche</normalizedToken>
2,5-3,5 mm lang, in der Mitte am breitesten (1,2-1,6 mm), innerseits flach,
<normalizedToken id="9504DE65649535D345435F34C01D51C2" originalValue="außerseits" pageId="null" pageNumber="440">ausserseits</normalizedToken>
<normalizedToken id="F6F7E76FE48F5978D32B03E9A9486B10" originalValue="gewölbt" pageId="null" pageNumber="440">gewoelbt</normalizedToken>
, beiderseits undeutlich
<normalizedToken id="A275BEA65C17BF85239C6AFB7DC994F3" originalValue="längsnervig" pageId="null" pageNumber="440">laengsnervig</normalizedToken>
, nicht
<normalizedToken id="D8B2285791096375072136EA04EE5BB2" originalValue="geflügelt" pageId="null" pageNumber="440">gefluegelt</normalizedToken>
, braun;
<normalizedToken id="E58DF4D8BB99C78BEBD7E8765AC47CD3" originalValue="Oberflächenstruktur" pageId="null" pageNumber="440">Oberflaechenstruktur</normalizedToken>
wie bei
<taxonomicName id="290A55D5447F0CAC17EFA1F62BCC9F62" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="null" pageNumber="440" phylum="Tracheophyta" rank="species" species="vulpina">
<emphasis id="08DAF6F5E7A3A98A34C7E013B9068B46" italics="true" pageId="null" pageNumber="440">C. vulpina</emphasis>
</taxonomicName>
(Nr. 18);
<normalizedToken id="76CBB5774B532AA87C27DBA679E92E37" originalValue="allmählich" pageId="null" pageNumber="440">allmaehlich</normalizedToken>
in den kurzen Schnabel
<normalizedToken id="AC56E4B2611960D4C0583EBE8B323D23" originalValue="verschmälert" pageId="null" pageNumber="440">verschmaelert</normalizedToken>
, Schnabel
<normalizedToken id="D6B2279092DAA81C173A550A5A8A8916" originalValue="außerseits" pageId="null" pageNumber="440">ausserseits</normalizedToken>
aufgeschlitzt. Narben 2.
</paragraph>
<paragraph id="B2110970D3CB7D602786E63728762EF8" pageId="null" pageNumber="440">
<emphasis id="7DF6AB042A576CFDF6E076EF0B1DA1B0" italics="true" pageId="null" pageNumber="440">Zytologische Angaben. 2n = 56:</emphasis>
Material aus Island und Kanada (
<normalizedToken id="47CE0A1B2D1D81419E77C39215BF2E87" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
und
<normalizedToken id="4436A5723A89BB987B35E5D329F903C8" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
1961,
<normalizedToken id="D2947FE8E7BFCFA545DABCD5C13BF20F" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
und Solbrig 1965,
<normalizedToken id="D1EED0C840D14BB9FE85DD71ADEB18B9" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
und Ritchie 1966).
<emphasis id="213200F7F97D9658BE6B86BB6D3A6E59" italics="true" pageId="null" pageNumber="440">2n ca. 64:</emphasis>
Material aus Island (
<normalizedToken id="804DEF379AEAACC751D7D4FA239B9E44" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
und
<normalizedToken id="8ABDD7DA101E5AC6F3B2B39EC7590480" originalValue="Löve" pageId="null" pageNumber="440">Loeve</normalizedToken>
1956b).
</paragraph>
<paragraph id="2F91A68BB031B7F3A320DCF04DC8A9A4" pageId="null" pageNumber="440">
<emphasis id="DDF8EB040A5D138852FA8C83E906008B" italics="true" pageId="null" pageNumber="440">Standort.</emphasis>
Kollin, montan und subalpin.
<normalizedToken id="98C923E9458A1B5EA23367A38551CFDC" originalValue="Torfschlammböden" pageId="null" pageNumber="440">Torfschlammboeden</normalizedToken>
, die zeitweise austrocknen, oder
<normalizedToken id="46B79511D9EEF770E3708549BE5721BF" originalValue="ständig" pageId="null" pageNumber="440">staendig</normalizedToken>
<normalizedToken id="E01068F1C8D1474AF95D6D5176194C54" originalValue="überschwemmte" pageId="null" pageNumber="440">ueberschwemmte</normalizedToken>
Schlenken; nur in Hochmooren. Sehr seltene, nordische Reliktpflanze.
<emphasis id="8E376F013BA26B1AF4855CFC103425BA" italics="true" pageId="null" pageNumber="440">Caricetum Heleonastis</emphasis>
Paul et Lutz 1941.
</paragraph>
<paragraph id="6C416341A0E2464F6EC01CF015B83EB6" pageId="null" pageNumber="440">
<emphasis id="5B634D47F294B60257763B4FAED68F53" italics="true" pageId="null" pageNumber="440">Verbreitung. Eurosibirisch-nordamerikanische Pflanze:</emphasis>
Island, Skandinavien (zwischen 57 und 71° NB), Finnland, Norddeutschland (erloschen?), Baltikum, Polen,
<normalizedToken id="49142F60CF35499043528A94F10A590A" originalValue="nördliches" pageId="null" pageNumber="440">noerdliches</normalizedToken>
<normalizedToken id="5815F0CB7371681F59125C3A799B223C" originalValue="Rußland" pageId="null" pageNumber="440">Russland</normalizedToken>
; Jura,
<normalizedToken id="98BC280A700FF1FAA4A20C812D859721" originalValue="Süddeutschland" pageId="null" pageNumber="440">Sueddeutschland</normalizedToken>
(wenige Fundstellen), Alpen (Alpi marittime in Piemont [Bono 1964], zerstreut von den Berner Alpen bis Steiermark), Grenzgebirge zwischen Jugoslawien und Bulgarien; Sibirien (vom Ural bis Transbaikalien zwischen 55 und 69° NB), Kaukasus, Altai; in Nordamerika zwischen 53 und 65° NB. Verbreitungskarten von
<normalizedToken id="60442CC62CD198FFF0AF1AB5412BCC69" originalValue="Hultén" pageId="null" pageNumber="440">Hulten</normalizedToken>
(1962) und Meusel (1964). - Im Gebiet:
<normalizedToken id="D9ECA2E0E6439F81464637DADB6DD837" originalValue="Französischer" pageId="null" pageNumber="440">Franzoesischer</normalizedToken>
Jura (neue Angaben von Moreau 1965), wenige Fundorte im
<normalizedToken id="202D8285D9A66968D9536B9FC0F3F637" originalValue="Waadtländer" pageId="null" pageNumber="440">Waadtlaender</normalizedToken>
und Neuenburger Jura, deutsches Bodenseegebiet (Tiefenweiher bei Klosterwald und Gegend von Lindau),
<pageBreakToken id="075B7B80BF8C4F6477F9ADF875D973E6" pageId="null" pageNumber="440">Mittelland</pageBreakToken>
(Wetzikon), Alpenvorland und Nordalpen in Bern (Amsoldingen, Schwarzenegg, Haslital), Luzern (Ballmoos bei Entlebuch), Zug (
<normalizedToken id="C669740F8ABC70818A1619C72CF77261" originalValue="Geißboden" pageId="null" pageNumber="440">Geissboden</normalizedToken>
erloschen), St. Gallen (
<normalizedToken id="D5B4CDFB9E79D2754D7E2B67AD8C2350" originalValue="Gräppelensee" pageId="null" pageNumber="440">Graeppelensee</normalizedToken>
, Wideralp,
<normalizedToken id="AEFC0EB4544FFD7542B2541510F186AE" originalValue="Älpli" pageId="null" pageNumber="440">Aelpli</normalizedToken>
bei Wildhaus), Innerrhoden (Potersalp) und
<normalizedToken id="94FF81B5B6EB6465AD24897CDBA0D799" originalValue="Graubünden" pageId="null" pageNumber="440">Graubuenden</normalizedToken>
(
<normalizedToken id="2C566E77CCB756AAA1495EDE7DC7C668" originalValue="Prättigau" pageId="null" pageNumber="440">Praettigau</normalizedToken>
). Karte der zirkumalpinen Verbreitung von Bresinsky (1965).
</paragraph>
<paragraph id="7A7F85A3AA2DED105A09A3A2B90A569E" pageId="null" pageNumber="440">
<emphasis id="D3493B65C9255F67765D7953C25BB797" italics="true" pageId="null" pageNumber="440">
Bemerkungen.
<taxonomicName id="57FBDEF3CF9474D74F10513737EBB6EF" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="null" pageNumber="440" phylum="Tracheophyta" rank="species" species="heleonastes">C. Heleonastes</taxonomicName>
</emphasis>
bildet eine
<emphasis id="EEF061C905724D16CC06481F2CA67C02" italics="true" pageId="null" pageNumber="440">Artengruppe mit eurosibirisch-nordamerikanischer Verbreitung.</emphasis>
Sie ist von
<normalizedToken id="0586EB1336C494F3FF85E51B9BBC1413" originalValue="Böcher" pageId="null" pageNumber="440">Boecher</normalizedToken>
(1952) in 2 Arten mit je 1 Unterart gegliedert worden. In
<taxonomicName id="018F864CA40854F2554C5620902AF6EC" class="Insecta" family="Cerambycidae" genus="Europa" kingdom="Animalia" order="Coleoptera" pageId="null" pageNumber="440" phylum="Arthropoda" rank="genus">Europa</taxonomicName>
kommt nur
<taxonomicName id="02B5761EF078ECD25280E3DC2D48140A" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="null" pageNumber="440" phylum="Tracheophyta" rank="species" species="heleonastes">
<emphasis id="7764F6EDB2DB7B4DBAC17C438EBD28A1" italics="true" pageId="null" pageNumber="440">C. Heleonastes</emphasis>
</taxonomicName>
vor, in Nordamerika
<taxonomicName id="144A86AE6AEBE5736DEDE188277F1717" authority="(Holm) Boecher, in Ostsibirien und Nordkanada C." authorityName="Boecher, in Ostsibirien und Nordkanada C." baseAuthorityName="Holm" pageId="null" pageNumber="440" rank="subSpecies" subGenus="Yukon" subSpecies="neurochlaena">
(Yukon) ssp.
<emphasis id="F06A19EA2278ED4F794D4192E09FB8BB" italics="true" pageId="null" pageNumber="440">neurochlaena</emphasis>
(Holm)
<normalizedToken id="E3550011EECF4BD2A657B826BA1A2ACE" originalValue="Böcher" pageId="null" pageNumber="440">Boecher</normalizedToken>
, in Ostsibirien und Nordkanada
<emphasis id="C6159B83DC218E374828845218ECE835" italics="true" pageId="null" pageNumber="440">C. ambylorhyncha</emphasis>
</taxonomicName>
Krecz., in
<normalizedToken id="E2C57BE489D58B46285F87CB3F151960" originalValue="Grönland" pageId="null" pageNumber="440">Groenland</normalizedToken>
<taxonomicName id="B1FDDBAAE40DA12E3B0FFD54A35D643B" authority="(Soerensen) Boecher." authorityName="Boecher." baseAuthorityName="Soerensen" pageId="null" pageNumber="440" rank="subSpecies" subSpecies="pseudolagopina">
ssp.
<emphasis id="0071FC09F3D290F230E4315DFFF92E35" italics="true" pageId="null" pageNumber="440">pseudolagopina</emphasis>
(
<normalizedToken id="BA815C8CB4A7B88B15AC6E67F861B2C0" originalValue="Sörensen" pageId="null" pageNumber="440">Soerensen</normalizedToken>
)
<normalizedToken id="E69156133B6FA136E8EA3CF127FF88C0" originalValue="Böcher" pageId="null" pageNumber="440">Boecher</normalizedToken>
.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
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