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<document id="ECA35499601A0AA509D76C6CC317E196" ID-CLB-Dataset="8170" ID-DOI="10.11646/zootaxa.4926.3.4" ID-GBIF-Dataset="72fbd015-d727-4e48-b8e1-54496db6c932" ID-ISSN="1175-5326" ID-Zenodo-Dep="4529361" ID-ZooBank="3AF255EA-2C21-4DCD-AD73-B8CC6CFED3BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612952550026" checkinUser="plazi" docAuthor="Henry, Shasta C., Cameron, Stephen L., Smolenski, Adam &amp; Mcquillan, Peter" docDate="2021" docId="130C843D9C33FF9CFF34FA0EFC78F9FB" docLanguage="en" docName="zootaxa.4926.3.4.pdf" docOrigin="Zootaxa 4926 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyzosteria yingina Henry &amp; Cameron &amp; Smolenski &amp; Mcquillan 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="389" masterDocId="EF35FC459C37FF99FFA3FFE9FF80FFD6" masterDocTitle="Polyzosteria cockroaches in Tasmania (Blattodea: Blattidae: Polyzosteriinae) represent a new, endemic species, with allopatric alpine and coastal sub-populations" masterLastPageNumber="400" masterPageNumber="384" pageNumber="388" updateTime="1698928618637" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BEB4C3CA9083446F6B0A18B1E8DE5913">Polyzosteria cockroaches in Tasmania (Blattodea: Blattidae: Polyzosteriinae) represent a new, endemic species, with allopatric alpine and coastal sub-populations</mods:title>
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<mods:namePart id="D829A513EAA6711F29443D50EDC109D4">Henry, Shasta C.</mods:namePart>
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<mods:affiliation id="B27129987A1FFED36541863B08D5EE1A">University of Tasmania, Discipline of Geography and Spatial Science, Churchill Ave, Sandy Bay TAS 7005. Shasta. Henry @ utas. edu. au; https: // orcid. org / 0000 - 0002 - 1001 - 3433</mods:affiliation>
<mods:nameIdentifier id="12CF5BA0547794DCEF13AF9CE32F75EB" type="email">enry@utas.edu.au</mods:nameIdentifier>
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<mods:namePart id="BE9E144AF9DE441DC0DBBDF33AC4D74A">Cameron, Stephen L.</mods:namePart>
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<mods:affiliation id="A42E418FA57EC9E010AD08A69D6196B5">610 Purdue Mall, West Lafayette, IN 47907, USA. cameros @ purdue. edu; https: // orcid. org / 0000 - 0002 - 6694 - 4130</mods:affiliation>
<mods:nameIdentifier id="7FAF16A548B92801637F86CCEEFC008C" type="email">cameros@purdue.edu</mods:nameIdentifier>
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<mods:namePart id="6100B6CEEB34390C3E0B5C8ACF4BC85C">Smolenski, Adam</mods:namePart>
<mods:affiliation id="81AE992CBD5558C487F52F6A5F4812AF">University of Tasmania, Central Science Laboratories, Churchill Ave, Sandy Bay TAS 7005.</mods:affiliation>
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<mods:namePart id="88EC1713766CDCB2B9B936E685D1BCBD">Mcquillan, Peter</mods:namePart>
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<mods:affiliation id="FB9A8F2799A2A3E14FE9569E41A0D491">University of Tasmania, Discipline of Geography and Spatial Science, Churchill Ave, Sandy Bay TAS 7005. Shasta. Henry @ utas. edu. au; https: // orcid. org / 0000 - 0002 - 1001 - 3433 &amp; University of Tasmania, Discipline of Geography and Spatial Science, Churchill Ave, Sandy Bay TAS 7005. p. b. mcquillan @ utas. edu. au; https: // orcid. org / 0000 - 0001 - 6334 - 372 X</mods:affiliation>
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<treatment id="130C843D9C33FF9CFF34FA0EFC78F9FB" ID-DOI="http://doi.org/10.5281/zenodo.4545992" ID-GBIF-Taxon="177420953" ID-Zenodo-Dep="4545992" LSID="urn:lsid:plazi:treatment:130C843D9C33FF9CFF34FA0EFC78F9FB" httpUri="http://treatment.plazi.org/id/130C843D9C33FF9CFF34FA0EFC78F9FB" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388">
<subSubSection id="D3BF66A09C33FF9DFF34FA0EFEEEF9F0" pageId="4" pageNumber="388" type="nomenclature">
<paragraph id="9B1A352B9C33FF9DFF34FA0EFD1EF9D7" blockId="4.[151,670,1511,1611]" box="[151,670,1511,1538]" pageId="4" pageNumber="388">
<heading id="C05282479C33FF9DFF34FA0EFD1EF9D7" bold="true" box="[151,670,1511,1538]" fontSize="11" level="1" pageId="4" pageNumber="388" reason="1">
<emphasis id="A9D1E9399C33FF9DFF34FA0EFD1EF9D7" bold="true" box="[151,670,1511,1538]" pageId="4" pageNumber="388">
<taxonomicName id="5CA54EA89C33FF9DFF34FA0EFE0CF9D4" authority="Henry &amp; Cameron &amp; Smolenski &amp; Mcquillan, 2021" authorityName="Henry &amp; Cameron &amp; Smolenski &amp; Mcquillan" authorityYear="2021" box="[151,396,1511,1538]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="yingina" status="sp. nov.">
<emphasis id="A9D1E9399C33FF9DFF34FA0EFE0CF9D4" bold="true" box="[151,396,1511,1538]" italics="true" pageId="4" pageNumber="388">Polyzosteria yingina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2E254429C33FF9DFE37FA01FE71F9D4" box="[404,497,1512,1538]" pageId="4" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
Henry (2020)
</emphasis>
</heading>
</paragraph>
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<emphasis id="A9D1E9399C33FF9DFF34F9E5FEEEF9F0" box="[151,366,1548,1574]" italics="true" pageId="4" pageNumber="388">Zoobank name link</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="D3BF66A09C33FF9DFF34F9D9FEA4F99D" box="[151,292,1584,1611]" pageId="4" pageNumber="388" type="description">
<paragraph id="9B1A352B9C33FF9DFF34F9D9FEA4F99D" blockId="4.[151,670,1511,1611]" box="[151,292,1584,1611]" pageId="4" pageNumber="388">
(
<figureCitation id="039E29AE9C33FF9DFF3CF9D9FE9BF99D" box="[159,283,1584,1611]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="9.[151,250,1971,1995]" captionStartId-1="10.[151,250,1807,1831]" captionStartId-2="11.[151,250,1970,1994]" captionTargetBox-0="[302,1285,184,1943]" captionTargetBox-1="[151,1436,187,1778]" captionTargetBox-2="[180,1404,189,1920]" captionTargetId-0="figure-24@9.[279,1309,181,1954]" captionTargetId-1="figure-20@10.[151,1436,181,1784]" captionTargetId-2="figure-24@11.[179,1408,181,1946]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." captionText-1="FIGURE 5. A–D Specimen QVM:2018:12:0916 E. QVM:2015:12:1503 A,B male genitalia in dorsal and ventral view; C,D terminalia; E close up of pronotum and T 1–4 (RHS) showing texture and translucent pale scales. Scale bars 5 mm." captionText-2="FIGURE 6. A–B Polyzosteria yingina sp. nov. holotype, male. A alive in its natural habitat at Lake Augusta prior to collection (photo Simon Grove, TMAG). B Pinned type specimen, (TMAG:F57626) scale bar 5mm." figureDoi-0="http://doi.org/10.5281/zenodo.4529369" figureDoi-1="http://doi.org/10.5281/zenodo.4529373" figureDoi-2="http://doi.org/10.5281/zenodo.4529379" httpUri-0="https://zenodo.org/record/4529369/files/figure.png" httpUri-1="https://zenodo.org/record/4529373/files/figure.png" httpUri-2="https://zenodo.org/record/4529379/files/figure.png" pageId="4" pageNumber="388">Figure 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D3BF66A09C33FF9DFF34F991FBE1F90C" pageId="4" pageNumber="388" type="diagnosis">
<paragraph id="9B1A352B9C33FF9DFF34F991FBE1F90C" blockId="4.[151,1437,1656,2042]" pageId="4" pageNumber="388">
<emphasis id="A9D1E9399C33FF9DFF34F991FE92F944" bold="true" box="[151,274,1656,1682]" pageId="4" pageNumber="388">Diagnosis.</emphasis>
Medium-sized, oval, convex, apterous; macroscopically smooth and uniformly coloured above excepting a non-metallic orange margin to the cerci. Upper surfaces dull metallic bronze but sometimes dark and matt. Ventral surface shining bronze and reddish brown with pale coxal border and process’.
</paragraph>
</subSubSection>
<subSubSection id="D3BF66A09C33FF9CFF64F90DFEDDFBC7" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388" type="description">
<paragraph id="9B1A352B9C33FF9DFF64F90DFDE5F891" blockId="4.[151,1437,1656,2042]" pageId="4" pageNumber="388">
<emphasis id="A9D1E9399C33FF9DFF64F90DFB6FF928" bold="true" box="[199,1263,1763,1790]" pageId="4" pageNumber="388">Description. Measurements (mm). Length, Pronotum Length/Width, Width, Hind Tibiae.</emphasis>
♂(n=20) L 21.4–30.3;
<materialsCitation id="2BCD3F769C33FF9DFE61F8E1FDC0F8F4" ID-GBIF-Occurrence="3037445302" box="[450,576,1800,1826]" collectionCode="PL" pageId="4" pageNumber="388" specimenCode="PL 6.8, 8.9" specimenCount="1">
<specimenCode id="CB039D509C33FF9DFE61F8E1FD8CF8F4" box="[450,524,1800,1826]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="388" type="Museum">PL 6.8</specimenCode>
–
<specimenCode id="CB039D509C33FF9DFDBFF8E1FDC0F8F4" box="[540,576,1800,1826]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="388" type="Museum">8.9</specimenCode>
</materialsCitation>
;
<materialsCitation id="2BCD3F769C33FF9DFDE9F8E1FD58F8F5" ID-GBIF-Occurrence="3037445307" box="[586,728,1800,1827]" collectionCode="PW" pageId="4" pageNumber="388" specimenCode="PW 10, 16.5" specimenCount="1">
<specimenCode id="CB039D509C33FF9DFDE9F8E1FD15F8F5" box="[586,661,1800,1827]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="388">PW 10</specimenCode>
–
<specimenCode id="CB039D509C33FF9DFD06F8E1FD58F8F5" box="[677,728,1800,1827]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="388">16.5</specimenCode>
</materialsCitation>
; W 16.5–19.1; HT 9.1–10.1;
<specimenCount id="8DA3FEA29C33FF9DFBB9F8E1FBAFF8F4" box="[1050,1071,1800,1826]" pageId="4" pageNumber="388" type="female">♀</specimenCount>
(n=25) L 26.9–36.5;
<materialsCitation id="2BCD3F769C33FF9DFAB8F8E1FA18F8F4" ID-GBIF-Occurrence="3037445306" box="[1307,1432,1800,1826]" collectionCode="PL" pageId="4" pageNumber="388" specimenCode="PL 7.3, 9.1" specimenCount="1">
<specimenCode id="CB039D509C33FF9DFAB8F8E1FAE5F8F4" box="[1307,1381,1800,1826]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="388" type="Museum">PL 7.3</specimenCode>
–
<specimenCode id="CB039D509C33FF9DFAD6F8E1FA18F8F4" box="[1397,1432,1800,1826]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="388" type="Museum">9.1</specimenCode>
</materialsCitation>
;
<materialsCitation id="2BCD3F769C33FF9DFF34F8C5FEA6F890" ID-GBIF-Occurrence="3037445308" box="[151,294,1836,1862]" collectionCode="PW" pageId="4" pageNumber="388" specimenCode="PW 14.6, 17" specimenCount="1">
<specimenCode id="CB039D509C33FF9DFF34F8C5FF7BF890" box="[151,251,1836,1862]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="388">PW 14.6</specimenCode>
–
<specimenCode id="CB039D509C33FF9DFEA8F8C5FEA6F890" box="[267,294,1836,1862]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="388">17</specimenCode>
</materialsCitation>
; W 18.2–21.5; HT 9.5–11.1.
</paragraph>
<paragraph id="9B1A352B9C33FF9CFF64F8B9FEE6FC73" blockId="4.[151,1437,1656,2042]" lastBlockId="5.[151,1437,151,1581]" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388">
Exoskeleton is microscopically pitted throughout, pits with minute translucent scales above (
<figureCitation id="039E29AE9C33FF9DFB4DF8B9FAE2F8BC" box="[1262,1378,1872,1898]" captionStart="FIGURE 3" captionStartId="4.[151,250,1009,1033]" captionTargetBox="[155,1425,260,954]" captionTargetId="figure-54@4.[155,1433,237,986]" captionTargetPageId="4" captionText="FIGURE 3. Evolutionary analysis by Maximum Likelihood method using CO1 (673BP) of Polyzosteria yingina sp. nov. included in a Blast search in MEGA incorporating P. limbata and P. viridissima specimens from the BOLD database." figureDoi="http://doi.org/10.5281/zenodo.4529367" httpUri="https://zenodo.org/record/4529367/files/figure.png" pageId="4" pageNumber="388">Figure 3E</figureCitation>
) and hairs below; scales are very loosely attached so wear and tear may explain their absence on many specimens. Vertex almost hidden from above, shining bronze. Ocelli indistinguishable. Vertex broadly flat, frons domed between concave antennal pits. Antennae are dark, bases brownish red, segments minutely haired with a ring of setae around the widest part. Clypeus and labrum are dark, each with a pale distal border. Labrum is deeply lobed around the notch. Palps are dark and densely haired (
<figureCitation id="039E29AE9C32FF9CFDD3FF7EFC8AFF67" box="[624,778,151,177]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4B, D</figureCitation>
). Dorsal surface colour is dull metallic bronze, sometimes with green highlights though can also be entirely dark and matt, always excepting the rim of the cerci which are bright orange and matt (
<figureCitation id="039E29AE9C32FF9CFE05FF36FE7AFF2F" box="[422,506,222,249]" captionStart="FIGURE 5" captionStartId="10.[151,250,1807,1831]" captionTargetBox="[151,1436,187,1778]" captionTargetId="figure-20@10.[151,1436,181,1784]" captionTargetPageId="10" captionText="FIGURE 5. A–D Specimen QVM:2018:12:0916 E. QVM:2015:12:1503 A,B male genitalia in dorsal and ventral view; C,D terminalia; E close up of pronotum and T 1–4 (RHS) showing texture and translucent pale scales. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529373" httpUri="https://zenodo.org/record/4529373/files/figure.png" pageId="5" pageNumber="389">Fig 5D</figureCitation>
). Texture is microscopically pitted and finely, unevenly nodular. These high points (and overlaps of tergites) are often smoother and shinier. Peripheral edges of dorsal segments are rolled and smooth. Paired, large and smooth stigmatic dots are obvious on each thoracic and abdominal segment, excepting T1,T8,T9 and T10 (
<figureCitation id="039E29AE9C32FF9CFEA4FEA2FE24FEB3" box="[263,420,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4A, C</figureCitation>
). A small, brush-like structure on the medial anterior margin of T1, usually obscured by the metanotum, has been observed in living males (
<bibRefCitation id="FF3448DA9C32FF9CFD78FE86FB97FE5F" author="Richards, K. &amp; Spencer, C." box="[731,1047,367,393]" pageId="5" pageNumber="389" pagination="27 - 33" refId="ref12087" refString="Richards, K. &amp; Spencer, C. (2019) Notes on the ecology of the Tasmanian alpine cockroach Polyzosteria sp. Burmeister, 1838 (Blattodea: Polyzosteriinae) including parasitism by Gordian worms (Nematomorpha: Gordioida). The Tasmanian Naturalist, 141, 27 - 33." type="journal article" year="2019">Richards and Spencer 2019</bibRefCitation>
). This ‘hair tuft’ is presumed to be involved in evaporation of pheromone secretions (
<bibRefCitation id="FF3448DA9C32FF9CFD74FE7AFC15FE7B" author="Mackerras, M. J." box="[727,917,402,429]" pageId="5" pageNumber="389" pagination="841 - 882" refId="ref11660" refString="Mackerras, M. J. (1965) Australian Blattidae (Blattodea). I. General remarks, and revision of the genus Polyzosteria Burmeister. Australian Journal of Zoology, 13 (5), 841 - 882. https: // doi. org / 10.1071 / ZO 9650841" type="journal article" year="1965">Mackerras 1965</bibRefCitation>
). Tergite T8, T9 and T10 are bounded by the projected angles of T7 and T10 also by the flattened, oblanceolate cerci. This species is sexually dimorphic. In males T10 is roughly oblong though slightly concave with slightly acute distal angles (
<figureCitation id="039E29AE9C32FF9CFBB6FE32FB0BFE23" box="[1045,1163,475,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4A</figureCitation>
,
<figureCitation id="039E29AE9C32FF9CFB35FE32FB3CFE23" box="[1174,1212,475,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1009,1033]" captionTargetBox="[155,1425,260,954]" captionTargetId="figure-54@4.[155,1433,237,986]" captionTargetPageId="4" captionText="FIGURE 3. Evolutionary analysis by Maximum Likelihood method using CO1 (673BP) of Polyzosteria yingina sp. nov. included in a Blast search in MEGA incorporating P. limbata and P. viridissima specimens from the BOLD database." figureDoi="http://doi.org/10.5281/zenodo.4529367" httpUri="https://zenodo.org/record/4529367/files/figure.png" pageId="5" pageNumber="389">3D</figureCitation>
). In the female, T10 is half-oval and convex with a peaked midline (
<figureCitation id="039E29AE9C32FF9CFD00FE16FC98FDCF" box="[675,792,511,537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4C</figureCitation>
), adapted for clamping the ootheca (
<bibRefCitation id="FF3448DA9C32FF9CFB08FE16FAAFFDCF" author="Rentz, D." box="[1195,1327,511,537]" pageId="5" pageNumber="389" pagination="167 - 175" refId="ref12010" refString="Rentz, D. (2014) A Guide To The Cockroaches Of Australia. CSIRO Publishing, Collingwood, pp. 167 - 175. https: // doi. org / 10.1071 / 9781486300372" type="book chapter" year="2014">Rentz 2014</bibRefCitation>
,
<bibRefCitation id="FF3448DA9C32FF9CFA9AFE16FEE4FDEB" author="Richards, K. &amp; Spencer, C." pageId="5" pageNumber="389" pagination="27 - 33" refId="ref12087" refString="Richards, K. &amp; Spencer, C. (2019) Notes on the ecology of the Tasmanian alpine cockroach Polyzosteria sp. Burmeister, 1838 (Blattodea: Polyzosteriinae) including parasitism by Gordian worms (Nematomorpha: Gordioida). The Tasmanian Naturalist, 141, 27 - 33." type="journal article" year="2019">Richards and Spencer 2019</bibRefCitation>
). Legs; tibiae, tops of tarsi and coxae-proximally are shining bronze; coxa-distally, the trochanter and femur are shining reddish brown. Coxal border and processes pale and femora with pale knee. Tibiae flat, biseriately spined with a fringe of hairs on the proximal edge. Tarsal claws light brown with dark tips. All spines yellow with dark points. Underside of thoracic segments shining bronze to greenish-blue. Anterior sternites are shiny metallic bronze, posterior sternites are shining red-brown; this colour transition beginning around the middle of S5. Smooth, stigmatic dots apparent from S2–S7. The opening of the scent gland is indicated by the rounded, sclerotised edge protruding onto S7 medially. Underside of cerci are bright orange and banded with tiny hairs (
<figureCitation id="039E29AE9C32FF9CFB55FD12FA10FCC3" box="[1270,1424,763,789]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4B, D</figureCitation>
). In males S9 bears paired anal styles in notches, protruding one third of their length past the posterior margin (
<figureCitation id="039E29AE9C32FF9CFAF2FCF6FF3CFC8B" captionStart="FIGURE 5" captionStartId="10.[151,250,1807,1831]" captionTargetBox="[151,1436,187,1778]" captionTargetId="figure-20@10.[151,1436,181,1784]" captionTargetPageId="10" captionText="FIGURE 5. A–D Specimen QVM:2018:12:0916 E. QVM:2015:12:1503 A,B male genitalia in dorsal and ventral view; C,D terminalia; E close up of pronotum and T 1–4 (RHS) showing texture and translucent pale scales. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529373" httpUri="https://zenodo.org/record/4529373/files/figure.png" pageId="5" pageNumber="389">Figure 5C</figureCitation>
). The subgenital lamina of the female (S7) is large and sculptural, plateaued medially, coming to a produced point where its posterior part is modified into the bivalvular hypogynum (
<figureCitation id="039E29AE9C32FF9CFC40FC8EFBDAFC57" box="[995,1114,871,897]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Figure 4D</figureCitation>
) characteristic of the family (
<bibRefCitation id="FF3448DA9C32FF9CFF3CFC62FEDAFC73" author="Mackerras, M. J." box="[159,346,906,933]" pageId="5" pageNumber="389" pagination="841 - 882" refId="ref11660" refString="Mackerras, M. J. (1965) Australian Blattidae (Blattodea). I. General remarks, and revision of the genus Polyzosteria Burmeister. Australian Journal of Zoology, 13 (5), 841 - 882. https: // doi. org / 10.1071 / ZO 9650841" type="journal article" year="1965">Mackerras 1965</bibRefCitation>
).
</paragraph>
<paragraph id="9B1A352B9C32FF9CFF64FC46FEDDFBC7" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="389">
The male genitalia conform to the general plan of
<taxonomicName id="5CA54EA89C32FF9CFD57FC46FCE5FC1F" authorityName="Burmeister" authorityYear="1838" box="[756,869,943,969]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="limbata">
<emphasis id="A9D1E9399C32FF9CFD57FC46FCE5FC1F" box="[756,869,943,969]" italics="true" pageId="5" pageNumber="389">P. limbata</emphasis>
</taxonomicName>
described by
<bibRefCitation id="FF3448DA9C32FF9CFC5CFC46FB4AFC1F" author="Mackerras, M. J." box="[1023,1226,942,969]" pageId="5" pageNumber="389" pagination="841 - 882" refId="ref11660" refString="Mackerras, M. J. (1965) Australian Blattidae (Blattodea). I. General remarks, and revision of the genus Polyzosteria Burmeister. Australian Journal of Zoology, 13 (5), 841 - 882. https: // doi. org / 10.1071 / ZO 9650841" type="journal article" year="1965">Mackerras (1965)</bibRefCitation>
. Male genitalia are most similar to
<taxonomicName id="5CA54EA89C32FF9CFEE6FC3AFE5DFC3B" authorityName="Shelford" authorityYear="1909" box="[325,477,979,1005]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="viridissima">
<emphasis id="A9D1E9399C32FF9CFEE6FC3AFE5DFC3B" box="[325,477,979,1005]" italics="true" pageId="5" pageNumber="389">P. viridissima</emphasis>
</taxonomicName>
, but differentiated by the process of R2 (R2b) having a more strongly defined hooked apex (
<figureCitation id="039E29AE9C32FF9CFF78FC1EFED0FBC7" box="[219,336,1014,1041]" captionStart="FIGURE 5" captionStartId="10.[151,250,1807,1831]" captionTargetBox="[151,1436,187,1778]" captionTargetId="figure-20@10.[151,1436,181,1784]" captionTargetPageId="10" captionText="FIGURE 5. A–D Specimen QVM:2018:12:0916 E. QVM:2015:12:1503 A,B male genitalia in dorsal and ventral view; C,D terminalia; E close up of pronotum and T 1–4 (RHS) showing texture and translucent pale scales. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529373" httpUri="https://zenodo.org/record/4529373/files/figure.png" pageId="5" pageNumber="389">Figure 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D3BF66A09C32FF9CFF64FBF3FC78F9FB" pageId="5" pageNumber="389" type="materials_examined">
<paragraph id="9B1A352B9C32FF9CFF64FBF3FB8DFB8F" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="389">
<emphasis id="A9D1E9399C32FF9CFF64FBF3FE37FBE2" bold="true" box="[199,439,1050,1076]" pageId="5" pageNumber="389">Material Examined.</emphasis>
*asterisk indicates where latitude and longitude (and altitude) have been calculated from coordinates on labels. Otherwise estimated according to place names on labels.
</paragraph>
<paragraph id="9B1A352B9C32FF9CFF64FB8AFDA8FB3F" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="389">
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<emphasis id="A9D1E9399C32FF9CFF64FB8AFEC7FBAA" bold="true" box="[199,327,1122,1149]" pageId="5" pageNumber="389">
<typeStatus id="441E8B899C32FF9CFF64FB8AFEB4FBAB" box="[199,308,1123,1149]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
(
</emphasis>
<figureCitation id="039E29AE9C32FF9CFEEBFB8AFE64FBAB" box="[328,484,1123,1149]" captionStart="FIGURE 6" captionStartId="11.[151,250,1970,1994]" captionTargetBox="[180,1404,189,1920]" captionTargetId="figure-24@11.[179,1408,181,1946]" captionTargetPageId="11" captionText="FIGURE 6. A–B Polyzosteria yingina sp. nov. holotype, male. A alive in its natural habitat at Lake Augusta prior to collection (photo Simon Grove, TMAG). B Pinned type specimen, (TMAG:F57626) scale bar 5mm." figureDoi="http://doi.org/10.5281/zenodo.4529379" httpUri="https://zenodo.org/record/4529379/files/figure.png" pageId="5" pageNumber="389">Figure 6A, B</figureCitation>
).
<collectingCountry id="E3B275BB9C32FF9CFE59FB8AFD17FBAB" box="[506,663,1123,1149]" name="Australia" pageId="5" pageNumber="389">AUSTRALIA</collectingCountry>
• 1♂, (Size mm: L 23,
<specimenCode id="CB039D509C32FF9CFC17FB8AFB81FBAB" box="[948,1025,1123,1149]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="389" type="Museum">PL 7.4</specimenCode>
x
<specimenCode id="CB039D509C32FF9CFB80FB8AFB0AFBAB" box="[1059,1162,1123,1149]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="389">PW 14.8</specimenCode>
, W 18.7, HT 9.2); Tasmania,
<location id="9E7A63F09C32FF9CFF48FB6EFE0DFB77" LSID="urn:lsid:plazi:treatment:130C843D9C33FF9CFF34FA0EFC78F9FB:9E7A63F09C32FF9CFF48FB6EFE0DFB77" box="[235,397,1159,1185]" country="Australia" county="Photographed" latitude="-41.870167" longLatPrecision="1" longitude="146.51271" municipality="Central Plateau" name="Lake Augusta" pageId="5" pageNumber="389">Lake Augusta</location>
: dunes,
<collectingMunicipality id="7B7EAF519C32FF9CFE49FB6EFD17FB77" box="[490,663,1159,1185]" pageId="5" pageNumber="389">Central Plateau</collectingMunicipality>
*
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;
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;
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asl
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;
<date id="EF1B13EB9C32FF9CFB0BFB6EFAC2FB77" box="[1192,1346,1159,1185]" pageId="5" pageNumber="389" value="2018-02-06">
<collectingDate id="FF5FEA039C32FF9CFB0BFB6EFAC2FB77" box="[1192,1346,1159,1185]" pageId="5" pageNumber="389" value="2018-02-06">06 Feb. 2018</collectingDate>
</date>
;
<collectorName id="365050FD9C32FF9CFAECFB6EFF3FFB13" pageId="5" pageNumber="389">Grove, S.J.</collectorName>
leg;
<collectingCounty id="727B4DA79C32FF9CFF5EFB42FE19FB13" box="[253,409,1195,1221]" pageId="5" pageNumber="389">Photographed</collectingCounty>
alive before hand-collection; DNA (table 1); https://www.flickr.com/photos/simongrovetmag/39415278624;
<collectionCode id="FDB4ADEE9C32FF9CFED4FB27FE4CFB3F" box="[375,460,1230,1257]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="5" pageNumber="389">TMAG</collectionCode>
:
<accessionNumber id="84F6A8C89C32FF9CFE73FB27FDA8FB3F" box="[464,552,1230,1257]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F57626" pageId="5" pageNumber="389" type="EnaNcbi">F57626</accessionNumber>
</materialsCitation>
</paragraph>
<paragraph id="9B1A352B9C32FF9CFF64FB1AFC3DFA83" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="389">
<emphasis id="A9D1E9399C32FF9CFF64FB1AFEF1FADB" bold="true" box="[199,369,1267,1293]" pageId="5" pageNumber="389">Photographed</emphasis>
<materialsCitation id="2BCD3F769C32FF9CFED4FB1AFC3DFA83" ID-GBIF-Occurrence="3037445301" accessionNumber="F13361" collectingDate="2019-02-16" collectingDateMax="2019-02-17" collectingDateMin="2019-02-16" collectionCode="TMAG" collectorName="C. Spencer &amp; K. Richards" country="Australia" elevation="1156" latitude="-41.861614" location="Central Plateau" longLatPrecision="1" longitude="146.52684" municipality="Carter Lakes" pageId="5" pageNumber="389" specimenCode="PL 7.8, PW 14.7" specimenCount="1" stateProvince="Tasmania" typeStatus="paratype">
<emphasis id="A9D1E9399C32FF9CFED4FB1AFD0CFADA" bold="true" box="[375,652,1266,1293]" pageId="5" pageNumber="389">
<typeStatus id="441E8B899C32FF9CFED4FB1AFE6DFADB" box="[375,493,1267,1293]" pageId="5" pageNumber="389" type="paratype">paratypes</typeStatus>
(
<figureCitation id="039E29AE9C32FF9CFE5FFB1BFDBAFADA" box="[508,570,1266,1292]" captionStart="FIGURE 4" captionStartId="9.[151,250,1971,1995]" captionTargetBox="[302,1285,184,1943]" captionTargetId="figure-24@9.[279,1309,181,1954]" captionTargetPageId="9" captionText="FIGURE 4. A–D Polyzosteria yingina sp. nov. ‘Photo paratypes’ A–B Male TMAG:F13361. C–D Female TMAG:F13354. A,C dorsal view. B,D ventral view. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4529369" httpUri="https://zenodo.org/record/4529369/files/figure.png" pageId="5" pageNumber="389">Fig 4</figureCitation>
. A–D).
</emphasis>
<collectingCountry id="E3B275BB9C32FF9CFD31FB1AFCAFFADB" box="[658,815,1267,1293]" name="Australia" pageId="5" pageNumber="389">AUSTRALIA</collectingCountry>
• 1♂ (Size mm:L 28.6,
<specimenCode id="CB039D509C32FF9CFB9AFB1AFB02FADB" box="[1081,1154,1267,1293]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="389" type="Museum">PL 7.8</specimenCode>
x
<specimenCode id="CB039D509C32FF9CFB3EFB1AFA80FADB" box="[1181,1280,1267,1293]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="389">PW 14.7</specimenCode>
, W 17.11, HT 10);
<collectingRegion id="5961FBC99C32FF9CFF6EFAFEFEBBFAE7" box="[205,315,1303,1329]" country="Australia" name="Tasmania" pageId="5" pageNumber="389">Tasmania</collectingRegion>
,
<collectingMunicipality id="7B7EAF519C32FF9CFEEAFAFEFE5EFAE7" box="[329,478,1303,1329]" pageId="5" pageNumber="389">Carter Lakes</collectingMunicipality>
,
<location id="9E7A63F09C32FF9CFE4FFAFEFD1AFAE7" LSID="urn:lsid:plazi:treatment:130C843D9C33FF9CFF34FA0EFC78F9FB:9E7A63F09C32FF9CFE4FFAFEFD1AFAE7" box="[492,666,1303,1329]" country="Australia" latitude="-41.861614" longLatPrecision="1" longitude="146.52684" municipality="Carter Lakes" name="Central Plateau" pageId="5" pageNumber="389" stateProvince="Tasmania">Central Plateau</location>
*
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;
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;
<elevation id="1088D2189C32FF9CFB8BFAFFFB21FAE7" box="[1064,1185,1302,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.156" pageId="5" pageNumber="389" unit="m" value="1156.0">
<quantity id="5C5D98CE9C32FF9CFB8BFAFFFBF5FAE7" box="[1064,1141,1302,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.156" pageId="5" pageNumber="389" unit="m" value="1156.0">1156m</quantity>
asl
</elevation>
;
<date id="EF1B13EB9C32FF9CFB0CFAFEFAF4FAE7" box="[1199,1396,1303,1329]" pageId="5" pageNumber="389" value="2019-02-16" valueMax="2019-02-17" valueMin="2019-02-16">
<collectingDate id="FF5FEA039C32FF9CFB0CFAFEFAF4FAE7" box="[1199,1396,1303,1329]" pageId="5" pageNumber="389" value="2019-02-16" valueMax="2019-02-17" valueMin="2019-02-16">16–17 Feb. 2019</collectingDate>
</date>
;
<collectorName id="365050FD9C32FF9CFA20FAFEFF71FA83" pageId="5" pageNumber="389">C. Spencer</collectorName>
and
<collectorName id="365050FD9C32FF9CFE8BFAD2FE26FA83" box="[296,422,1339,1365]" pageId="5" pageNumber="389">K.Richards</collectorName>
leg; lake shore mass stranding;
<collectionCode id="FDB4ADEE9C32FF9CFCAEFAD2FCE5FA83" box="[781,869,1339,1365]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="5" pageNumber="389">TMAG</collectionCode>
:F13361
</materialsCitation>
</paragraph>
<paragraph id="9B1A352B9C32FF9CFF64FAB6FE3DFA17" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="389">
<materialsCitation id="2BCD3F769C32FF9CFF64FAB6FE3DFA17" ID-GBIF-Occurrence="3037445304" accessionNumber="F13354" collectingDate="2019-02-16" collectingDateMax="2019-02-17" collectingDateMin="2019-02-16" collectionCode="TMAG" collectorName="C. Spencer &amp; K. Richards" country="Australia" elevation="1156" latitude="-41.861614" location="Central Plateau" longLatPrecision="1" longitude="146.52684" municipality="Carter Lakes" pageId="5" pageNumber="389" specimenCode="PL 8, PW 15.4" specimenCount="1" specimenCount-female="1" stateProvince="Tasmania">
<collectingCountry id="E3B275BB9C32FF9CFF64FAB6FEE4FAAF" box="[199,356,1375,1401]" name="Australia" pageId="5" pageNumber="389">AUSTRALIA</collectingCountry>
•
<specimenCount id="8DA3FEA29C32FF9CFED9FAB6FE1DFAAF" box="[378,413,1375,1401]" pageId="5" pageNumber="389" type="female">1♀</specimenCount>
, (Size mm:L 34,
<specimenCode id="CB039D509C32FF9CFDF8FAB6FD0FFAAF" box="[603,655,1375,1401]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="389" type="Museum">PL 8</specimenCode>
x
<specimenCode id="CB039D509C32FF9CFD0AFAB6FC8CFAAF" box="[681,780,1374,1401]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="389">PW 15.4</specimenCode>
, W 18.02, HT 10);
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,
<collectingMunicipality id="7B7EAF519C32FF9CFBF5FAB6FB68FAAF" box="[1110,1256,1375,1401]" pageId="5" pageNumber="389">Carter Lakes</collectingMunicipality>
,
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*
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asl
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;
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and
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leg; lake shore mass stranding;
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<emphasis id="A9D1E9399C32FF9CFF64FA23FE1BFA32" bold="true" box="[199,411,1482,1508]" pageId="5" pageNumber="389">Genital dissection</emphasis>
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<emphasis id="A9D1E9399C32FF9CFE02FA22FD2CFA32" bold="true" box="[417,684,1482,1509]" pageId="5" pageNumber="389">
<typeStatus id="441E8B899C32FF9CFE02FA22FD8CFA33" box="[417,524,1483,1509]" pageId="5" pageNumber="389" type="paratype">paratype</typeStatus>
(
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. A–D):
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<collectingCountry id="E3B275BB9C32FF9CFD12FA22FCCEFA33" box="[689,846,1483,1509]" name="Australia" pageId="5" pageNumber="389">AUSTRALIA</collectingCountry>
• 1♂, (Size mm:L 30.3,
<specimenCode id="CB039D509C32FF9CFBFEFA22FB26FA33" box="[1117,1190,1483,1509]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="389" type="Museum">PL 7.8</specimenCode>
x
<specimenCode id="CB039D509C32FF9CFB63FA22FA8EFA33" box="[1216,1294,1482,1509]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="389">PW 15</specimenCode>
, W 17.2, HT 9.8);
<collectingRegion id="5961FBC99C32FF9CFF72FA06FEBFF9DF" box="[209,319,1519,1545]" country="Australia" name="Tasmania" pageId="5" pageNumber="389">Tasmania</collectingRegion>
,
<collectingMunicipality id="7B7EAF519C32FF9CFEE9FA06FE6DF9DF" box="[330,493,1519,1545]" pageId="5" pageNumber="389">Lake Botsford</collectingMunicipality>
,
<location id="9E7A63F09C32FF9CFE5BFA06FD23F9DF" LSID="urn:lsid:plazi:treatment:130C843D9C33FF9CFF34FA0EFC78F9FB:9E7A63F09C32FF9CFE5BFA06FD23F9DF" box="[504,675,1519,1545]" country="Australia" latitude="-41.878902" longLatPrecision="1" longitude="146.50842" municipality="Lake Botsford" name="Central Plateau" pageId="5" pageNumber="389" stateProvince="Tasmania">Central Plateau</location>
*
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;
<geoCoordinate id="FE9153EC9C32FF9CFCC4FA06FB9BF9DF" box="[871,1051,1519,1545]" degrees="146" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="389" precision="1" seconds="30.33" value="146.50842">146°30'30.33''E</geoCoordinate>
;
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asl
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;
<date id="EF1B13EB9C32FF9CFB05FA06FABFF9DF" box="[1190,1343,1519,1545]" pageId="5" pageNumber="389" value="2016-12-30">
<collectingDate id="FF5FEA039C32FF9CFB05FA06FABFF9DF" box="[1190,1343,1519,1545]" pageId="5" pageNumber="389" value="2016-12-30">30 Dec. 2016</collectingDate>
</date>
; Among low alpine herbage. Genitalia dissected. DNA (table 1);
<collectionCode id="FDB4ADEE9C32FF9CFCA8F9FAFCD3F9FB" box="[779,851,1555,1581]" country="Australia" httpUri="http://grbio.org/cool/2s8k-2m7w" name="Queen Victoria Museum" pageId="5" pageNumber="389">QVM</collectionCode>
</materialsCitation>
:2018:12:0916.
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<mods:title id="C4DD4860CE566120DAB8CCF509234C61">Polyzosteria cockroaches in Tasmania (Blattodea: Blattidae: Polyzosteriinae) represent a new, endemic species, with allopatric alpine and coastal sub-populations</mods:title>
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<mods:namePart id="75FBC66B69B9AF40331E2A6FCB6CE27C">Cameron, Stephen L.</mods:namePart>
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<mods:affiliation id="74D966F2634F1E3F43DD2371CAD4637F">610 Purdue Mall, West Lafayette, IN 47907, USA. cameros @ purdue. edu; https: // orcid. org / 0000 - 0002 - 6694 - 4130</mods:affiliation>
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<mods:affiliation id="FBA490A65A1EB00DC359936C32CBE6D2">University of Tasmania, Discipline of Geography and Spatial Science, Churchill Ave, Sandy Bay TAS 7005. Shasta. Henry @ utas. edu. au; https: // orcid. org / 0000 - 0002 - 1001 - 3433 &amp; University of Tasmania, Discipline of Geography and Spatial Science, Churchill Ave, Sandy Bay TAS 7005. p. b. mcquillan @ utas. edu. au; https: // orcid. org / 0000 - 0001 - 6334 - 372 X</mods:affiliation>
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Genus
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<emphasis id="A9D1E9399C33FF9DFF4DFB2EFE01FB34" bold="true" box="[238,385,1223,1250]" italics="true" pageId="4" pageNumber="388">Polyzosteria</emphasis>
Burmeister, 1838
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<taxonomicName id="5CA54EA89C33FF9DFF34FAF9FE01FAFC" authorityName="Henry &amp; Cameron &amp; Smolenski &amp; Mcquillan" authorityYear="2021" box="[151,385,1296,1322]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="yingina">
<emphasis id="A9D1E9399C33FF9DFF34FAF9FE01FAFC" box="[151,385,1296,1322]" italics="true" pageId="4" pageNumber="388">Polyzosteria yingina</emphasis>
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is a conventional member of this genus because it is: large, apterous and diurnal; with the margins of the pronotum forming low, rolled edges; the angles of T7 are acute; the hind tarsal segments are bare; the dorsal surfaces of the tibiae are flattened with the spines in two rows; and the cerci and styles are very short (
<bibRefCitation id="FF3448DA9C33FF9DFF3CFA95FED9FA41" author="Mackerras, M. J." box="[159,345,1404,1431]" pageId="4" pageNumber="388" pagination="841 - 882" refId="ref11660" refString="Mackerras, M. J. (1965) Australian Blattidae (Blattodea). I. General remarks, and revision of the genus Polyzosteria Burmeister. Australian Journal of Zoology, 13 (5), 841 - 882. https: // doi. org / 10.1071 / ZO 9650841" type="journal article" year="1965">Mackerras 1965</bibRefCitation>
,
<bibRefCitation id="FF3448DA9C33FF9DFEC7FA95FE69FA40" author="Rentz, D." box="[356,489,1404,1430]" pageId="4" pageNumber="388" pagination="167 - 175" refId="ref12010" refString="Rentz, D. (2014) A Guide To The Cockroaches Of Australia. CSIRO Publishing, Collingwood, pp. 167 - 175. https: // doi. org / 10.1071 / 9781486300372" type="book chapter" year="2014">Rentz 2014</bibRefCitation>
).
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<paragraph id="0C509D2C57DD5E56C08A20A39323151E" pageId="104" pageNumber="105">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<taxonomicName id="829A48C098C871F86BFC739DE8626861" ID-CoL="SDCR" LSID="http://species-id.net/wiki/Cephaloleia_cylindrica" authority="Staines, 1996" authorityName="Staines" authorityYear="1996" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia cylindrica" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="cylindrica">Cephaloleia cylindrica Staines, 1996</taxonomicName>
Fig. 110
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<bibRefCitation id="1D1E07D0D268FEC9DEF9C8E7F542A35E" pageId="104" pageNumber="105">Staines 1996</bibRefCitation>
: 25.
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<paragraph id="5216F12CC243DBE1BF47B9CC687CEC01" pageId="104" pageNumber="105">Description.</paragraph>
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Elongate; narrow; convex reddish-brown; antennae, eyes, and legs black. Head: vertex smooth, medial sulcus absent; frons not projecting; not depressed between eyes. Antenna: reaches past humerus; slender; antennomeres 1-2 subglobular, short; 3-4 elongate, subequal in length, each
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x length of 1 and 2 combined; longest; 5-10 elongate, each shorter than 4; 11 subequal to 10, pointed at apex; 1-4 punctate with scattered setae; 5-11 setose. Pronotum: transverse; lateral margin bisinuate,
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; anterior angle pointed, slightly produced; posterior angle acute; anterior margin emarginate behind head; disc subconvex; surface impunctate; basal impression absent; pronotal length 1.1 mm; pronotal width 1.1-1.3 mm. Scutellum: triangular; impunctate. Elytron: lateral margin straight, smooth, margined; apex rounded; sutural angle without tooth; humerus rounded, not produced; slightly constricted behind humerus; shallowly punctate-striate, punctures large; elytral length 3.3-3.6 mm; elytral width 1.7 mm. Venter: pro- and metasterna impunctate; mesosternum impunctate medially, punctate laterally; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete. Leg: slender; femur punctate at base; tibia with fringe of setae on inner margin of apex. Total length: 4.7-5.3 mm.
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<paragraph id="B1436CBA71DFA9419662153C899D25B9" pageId="105" pageNumber="106">Diagnosis.</paragraph>
<paragraph id="EC017FA5480BB6B0CE889F7DD74EAE49" pageId="105" pageNumber="106">
This species is similar to
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and
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. It can be distinguished by antennomeres 1 and 2 being subglobose.
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<paragraph id="19DFAA6705CC024DEFE330A25C843399" pageId="105" pageNumber="106">Distribution.</paragraph>
<paragraph id="654CA3C5BA57BC3B45410CC9B949B9DE" pageId="105" pageNumber="106">Costa Rica, Panama.</paragraph>
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<paragraph id="4D709156FD8209C51CC3B6343C5B79CF" pageId="105" pageNumber="106">Type material examined.</paragraph>
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Holotype: X X Plantation, Rep Panama, Feb. 11, 1930 [blue-green label]/ Blackwelder Collection [blue-green label]/ Holotype
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Staines, Des. C. L. Staines 1994 [red label] (USNM).
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<paragraph id="2A3FBE68967654AE46F269DFCADE36B4" pageId="105" pageNumber="106">Specimens examined.</paragraph>
<paragraph id="E558FCAF0F0F9B9BBA3C0C95A98BC62E" pageId="105" pageNumber="106">COSTA RICA: Puntarenas- Est. Sirena, 0-100 m, May 1992 (INBIO). PANAMA: 11 February 1930 (USNM). Total: 2.</paragraph>
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<paragraph id="3D732E5057C76FE4EABFA8751BBC938D" pageNumber="513">&quot;Habitat in arenosis Belgii, Hispaniae, Italiae.&quot; RCN: 3534.</paragraph>
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Original material:
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Herb. Linn. No. 630.42 (
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; [icon] in
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, Hist. General. Pl. 2: 1652. 1586.
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Current name:
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</mods:mods>
<treatment ID-GBIF-Taxon="152051587" LSID="urn:lsid:plazi:treatment:130CD4EB8C589FD9349E62D0A0305617" httpUri="http://treatment.plazi.org/id/130CD4EB8C589FD9349E62D0A0305617" lastPageId="9" lastPageNumber="82" pageId="8" pageNumber="81">
<subSubSection pageId="8" pageNumber="81" type="nomenclature">
<paragraph pageId="8" pageNumber="81">
<taxonomicName LSID="http://species-id.net/wiki/Metriocnemus_tristellus" authority="Edwards" class="Insecta" family="Chironomidae" genus="Metriocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metriocnemus (Metriocnemus) tristellus" order="Diptera" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="tristellus" subGenus="Metriocnemus">Metriocnemus (Metriocnemus) tristellus Edwards</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="81" type="reference_group">
<paragraph pageId="8" pageNumber="81">
<taxonomicName class="Insecta" family="Chironomidae" genus="Metriocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metriocnemus tristellus" order="Diptera" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="tristellus">Metriocnemus tristellus</taxonomicName>
Edwards, 1929: 312
</paragraph>
<paragraph pageId="8" pageNumber="81">
<taxonomicName class="Insecta" family="Chironomidae" genus="Metriocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metriocnemus gracei" order="Diptera" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="gracei">Metriocnemus gracei</taxonomicName>
Edwards, 1929: 312 sensu
<bibRefCitation author="Wang, X" editor="Hoffrichter, O" journalOrPublisher="Shaker Verlag, Aachen" pageId="13" pageNumber="86" pagination="629 - 652" title="A revised checklist of Chironomidae from China (Diptera)." volumeTitle="Late 20 th century research on Chironomidae: an anthology from the 13 th International Symposium on Chironomidae, Freiburg, 5 - 9 September 1997" year="2000">Wang 2000</bibRefCitation>
: 637.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="81" type="material examined">
<paragraph pageId="8" pageNumber="81">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="81">
CHINA: Zhejiang Province, Qingyuan City, Baishanzu County,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="27.730833">27°43'51&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.12583">100°7'33&quot;E</geoCoordinate>
, 2 males, 18-22.iv.1994, light trap, H. Wu.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="82" type="remarks">
<paragraph pageId="9" pageNumber="82">
<pageBreakToken pageId="9" pageNumber="82" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="9" pageNumber="82">
The species has an AR = 1.2, very low inferior volsella, and weak anal point. Basal half of the wing membrane is bare, costa is strongly produced, and the distance
<normalizedToken originalValue="C–M">C-M</normalizedToken>
is rather less than
<normalizedToken originalValue="M–Cu">M-Cu</normalizedToken>
. The palp is unusually short, palpomeres 3 and 4 are less than three times as long as broad, palpomere 2 is rather longer.
<bibRefCitation author="Wang, X" editor="Hoffrichter, O" journalOrPublisher="Shaker Verlag, Aachen" pageId="13" pageNumber="86" pagination="629 - 652" title="A revised checklist of Chironomidae from China (Diptera)." volumeTitle="Late 20 th century research on Chironomidae: an anthology from the 13 th International Symposium on Chironomidae, Freiburg, 5 - 9 September 1997" year="2000">Wang (2000)</bibRefCitation>
recorded
<taxonomicName class="Insecta" family="Chironomidae" genus="Metriocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metriocnemus gracei" order="Diptera" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="gracei">Metriocnemus gracei</taxonomicName>
Edwards, 1929 from China, which should be
<taxonomicName class="Insecta" family="Chironomidae" genus="Metriocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metriocnemus (Metriocnemus) tristellus" order="Diptera" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="tristellus" subGenus="Metriocnemus">Metriocnemus (Metriocnemus) tristellus</taxonomicName>
Edwards after we re-examined the specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="82" type="distribution">
<paragraph pageId="9" pageNumber="82">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="82">
The species is recorded from the Nearctic and Palaearctic regions (
<bibRefCitation author="Ashe, P" journalOrPublisher="Irish Biogeographical Society &amp; National Museum of Ireland, Dublin" pageId="11" pageNumber="84" title="A World Catalogue of Chironomidae (Diptera)" year="2012">
Ashe and
<normalizedToken originalValue="O’Connor">O'Connor</normalizedToken>
2012
</bibRefCitation>
).The species has been collected in Oriental China for first time.
</paragraph>
</subSubSection>
</treatment>
</document>