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<document id="C869CFE8624F4C446B9AF2383525F91C" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F592020D0D15AD1EB8B1DEFD80" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona ederi Engel 2022, new species" docType="treatment" docUuid="3004F475-B4C8-48ED-8C9A-350A4EAA18D2" docUuidSource="ZooBank" docVersion="1" lastPageNumber="12" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="6" updateTime="1722456085954" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="CF19F3690AC72564480EB7D491CEAF9F">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="B5FC4850754D873EAF9C7BB6E563C397">Engel, Michael S.</mods:namePart>
<mods:affiliation id="A1D8E2C67C80703606E96E1C43535D92">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:title id="29945FED6CF840C44B574C5751599457">Journal of Melittology</mods:title>
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<mods:date id="9F0C7E89112F25FAAEF4CAEC62E4EE8E">2022</mods:date>
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<mods:number id="2334CD1708DDDE58B5169B6B2E13E8F1">2022-03-14</mods:number>
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<mods:number id="4E2EA0AAFD8B26908D9E93B0F138A719">110</mods:number>
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<paragraph id="1A7936E392020D0715AD1EB8B15FFB60" blockId="7.[433,895,1185,1209]" box="[433,895,1185,1209]" pageId="7" pageNumber="6">
<heading id="4131818F92020D0715AD1EB8B15FFB60" box="[433,895,1185,1209]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="2">
<taxonomicName id="DDC64D6092020D0715AD1EB8B0C7FB60" authority="Engel" authorityName="Engel" authorityYear="2022" box="[433,743,1185,1209]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF192020D0715AD1EB8B0BBFB60" bold="true" box="[433,667,1185,1209]" italics="true" pageId="7" pageNumber="6">Scaptotrigona ederi</emphasis>
Engel
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,
<taxonomicNameLabel id="3381578A92020D0716EE1EBBB15FFB60" box="[754,895,1185,1209]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="52DC656892020D0716491EF8B0FBFB20" box="[597,731,1249,1273]" pageId="7" pageNumber="6" type="description">
<paragraph id="1A7936E392020D0716491EF8B0FBFB20" blockId="7.[240,1087,1219,1273]" box="[597,731,1249,1273]" pageId="7" pageNumber="6">
(
<figureCitation id="82FD2A6692020D0716421EF8B0F2FB20" box="[606,722,1249,1273]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStart-2="Figures 711" captionStartId-0="8.[160,241,1212,1234]" captionStartId-1="10.[161,242,1457,1479]" captionStartId-2="12.[160,241,1357,1379]" captionTargetBox-0="[175,1153,237,1199]" captionTargetBox-1="[165,1151,227,1434]" captionTargetBox-2="[160,1156,223,1346]" captionTargetId-0="figure-201@8.[160,1168,224,1206]" captionTargetId-1="figure-91@10.[160,1168,224,1451]" captionTargetId-2="figure-180@12.[160,1168,223,1351]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Figures 12. Workers of Scaptotrigona ederi, new species. 1. Lateral habitus of individual on the darker end of the color spectrum. 2. Lateral habitus of individual on the lighter end of the spectrum." captionText-1="Figures 36. Workers of Scaptotrigona ederi, new species. 3. Facial view of individual on the lighter end of the color spectrum. 4. Facial view, darker worker. 5. Dorsal habitus, darker worker. 6. Dorsal habitus, lighter worker." captionText-2="Figures 711. Workers of Scaptotrigona ederi, new species. 7. Dorsal view of mesosoma, lighter worker. 8. Dorsal view of mesosoma, darker worker. 9. Mesoscutellum. 10. Prolateral surface of metatibia, metatarsus, and metapretarsus. 11. Apical view of metasoma." pageId="7" pageNumber="6">Figs. 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892020D0714CC1F38B1E2F900" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="1A7936E392020D0714CC1F38B1E2F900" blockId="7.[160,1168,1313,1754]" pageId="7" pageNumber="6">
DIAGNOSIS: This is a comparatively abundant and distinctive species in the western Andean region of
<collectingCountry id="62D1767392020D0715611F58B3D0FA80" box="[381,496,1345,1369]" name="Colombia" pageId="7" pageNumber="6">Colombia</collectingCountry>
around Cali and currently known to range in elevation from
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, making it one of the highest occurrences for
<taxonomicName id="DDC64D6092020D07178A1F7BB60CFAA0" box="[918,1068,1378,1401]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF192020D07178A1F7BB60CFAA0" box="[918,1068,1378,1401]" italics="true" pageId="7" pageNumber="6">Scaptotrigona</emphasis>
</taxonomicName>
(second only to
<taxonomicName id="DDC64D6092020D0714E71F9BB3B6FA40" authorityName="Engel" authorityYear="2022" box="[251,406,1409,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="magdalenae">
<emphasis id="28B2EAF192020D0714E71F9BB3B6FA40" box="[251,406,1409,1433]" italics="true" pageId="7" pageNumber="6">S. magdalenae</emphasis>
</taxonomicName>
,
<emphasis id="28B2EAF192020D0715B81F98B02CFA40" box="[420,524,1409,1433]" italics="true" pageId="7" pageNumber="6">vide infra</emphasis>
). Reports of
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<emphasis id="28B2EAF192020D0716B01F9BB0DDFA40" box="[684,765,1409,1433]" italics="true" pageId="7" pageNumber="6">S. ederi</emphasis>
</taxonomicName>
(as a
<emphasis id="28B2EAF192020D07175E1F9BB1C1FA40" box="[834,993,1409,1433]" italics="true" pageId="7" pageNumber="6">nomen nudum</emphasis>
) from the East of
<collectingCountry id="62D1767392020D0714A31FB8B312FA60" box="[191,306,1441,1465]" name="Colombia" pageId="7" pageNumber="6">Colombia</collectingCountry>
and elsewhere have generally been based on misidentifications. Among species in this subgroup,
<taxonomicName id="DDC64D6092020D0715CC1FDBB001FA00" authorityName="Engel" authorityYear="2022" box="[464,545,1473,1497]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192020D0715CC1FDBB001FA00" box="[464,545,1473,1497]" italics="true" pageId="7" pageNumber="6">S. ederi</emphasis>
</taxonomicName>
is most superficially similar to those species of sub- group B in relation to its generally dark coloration, although it still falls within sub- group A at the darker end of the spectrum. The species is one of a few that lacks a defined genal tooth projecting upward from the lower margin of the lateral concavity in the preoccipital carina. Among those without such a defined genal tooth, the dark coloration, black or at least dark fuscous bristles of the vertex and mesoscutal antero- lateral corners, and nearly entirely black bristles of the metabasitarsus (and frequently the metatibia), serve to distinguish this species most readily. In addition, the promi- nent bristles of metasomal terga
<collectionCode id="7CD7AE2692020D0716031CDBB01AF900" box="[543,570,1730,1753]" pageId="7" pageNumber="6">III</collectionCode>
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are yellow to fulvous in color.
</paragraph>
</subSubSection>
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<paragraph id="1A7936E3920D0D0814BC1EA5B32BFAD2" blockId="8.[160,1168,1212,1291]" pageId="8" pageNumber="7">
<emphasis id="28B2EAF1920D0D0814BC1EA5B304FB08" bold="true" box="[160,292,1212,1234]" pageId="8" pageNumber="7">Figures 12.</emphasis>
Workers of
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<emphasis id="28B2EAF1920D0D0815AA1EA4B055FB08" box="[438,629,1212,1234]" italics="true" pageId="8" pageNumber="7">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
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.
<emphasis id="28B2EAF1920D0D0817071EA5B10DFB08" bold="true" box="[795,813,1212,1233]" pageId="8" pageNumber="7">1.</emphasis>
Lateral habitus of individual on the darker end of the color spectrum.
<emphasis id="28B2EAF1920D0D0816201EC0B06EFB37" bold="true" box="[572,590,1241,1262]" pageId="8" pageNumber="7">2.</emphasis>
Lateral habitus of individual on the lighter end of the spectrum.
</paragraph>
</caption>
<subSubSection id="52DC6568920D0D0C14CC1F2FB30FFA3C" lastPageId="12" lastPageNumber="11" pageId="8" pageNumber="7" type="description">
<paragraph id="1A7936E3920D0D0814CC1F2FB194F937" blockId="8.[160,1175,1334,1774]" pageId="8" pageNumber="7">
DESCRIPTION: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite)
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. Head wider than long, width
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, length
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; compound eye length
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; upper interorbital distance
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, lower interorbital distance
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. Scape length
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, about as long as torulocellar distance, torulocellar distance
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; flagello- mere I longer than wide, longer than flagellomere II, remaining flagellomeres about as long as wide to slightly longer than wide except apical flagellomere longer than wide. Clypeus broader than long, approximately 1.71.75× as wide as long, length
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, width
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. Malar area long, length approximately 1.551.7× flagellar diameter or 0.750.82× basal mandibular width. Preoccipital ridge carinate, weakly lamellate dorsally, with bordering medial indentation shallower than lateral inden- tations, laterally concavity interrupting carina comparatively shallow, lower margin of concavity with carina forming rounded angle, sometimes slightly acute, without sharply acute lamellate projection (genal tooth) extending upward.
</paragraph>
<paragraph id="1A7936E3920C0D0914CD1AFBB0B3FA80" blockId="9.[161,1175,225,1785]" pageId="9" pageNumber="8">
Integument of labiomaxillary complex yellow orange except prementum typically brown; labrum yellow orange to dark orange; mandible yellow orange to ferruginous; clypeus largely yellow orange to orange except typically with dark brown mark at medioapical margin and apicolateral corners from apical margin to epistomal sulcus (sometimes absent or faint, and sometimes these all merged medioapically), often with mediolongitudinal brown mark from base to about midlength, epistomal sulcus well delineated by brown to black; supraclypeal area brown to dark brown; malar space often dark brown, sometimes light brown to orange; face below tangent of antennal toruli orange to light yellow brown, typically slightly darker toward lateral borders of antennal toruli and often extending along inner orbit as narrow strip to point slightly above upper tangent of antennal toruli; scape yellow orange except typically brown dorsoapically, sometimes brown extending along entire dorsal surface; pedicel and flagellum typically dark brown above, with narrow underside of flagellum orange to light brown; remainder of face dark brown to black, sometimes with small areas of light brown immediately above antennal toruli; posterior of head dark brown; upper gena dark brown, remainder of gena orange, although if malar space dark brown, then such dark brown extends onto gena ventroapically; postgena orange to yellow orange. Mesosoma typically dark brown to black, with ferruginous apex to mesoscutellum, except sometimes with areas of reddish orange to reddish brown on mesepisternum, typically as small areas posteriorly and ventrally, and sometimes ventrally on propo- deum, and mesoscutellum ranging from wholly orange to wholly black, with degrees of intermixing of colors between, V-shaped incision in mesoscutellum typically black; tegula wholly yellow orange and semi-translucent, sometimes with slight brownish darkening anteriorly; legs typically yellow orange on coxae, trochanters, basal por- tions of tibiae, distitarsomeres, and inferior and superior areas of metabasitarsus, with remainder of tibiae, femora, and tarsi dark brown to black, except sometimes meso- and metacoxae light brown. Wing membranes hyaline clear to lightly parchment col- ored; veins yellow orange. Metasoma generally dark brown to black except anterior-facing surface of tergum I typically reddish brown to light yellow brown, dorsal-facing disc of tergum I often medially lighter reddish brown to yellowish brown and with exceptionally narrow apical marginal zone of reddish brown to yellow and terga II
<collectionCode id="7CD7AE26920C0D0910681EBBB6B0FB60" box="[1140,1168,1186,1209]" pageId="9" pageNumber="8">IV</collectionCode>
with variably developed apical marginal zones of light brown to more frequently yel- low, sometimes quite broad on terga II and
<collectionCode id="7CD7AE26920C0D0916811EFBB09AFB20" box="[669,698,1250,1273]" pageId="9" pageNumber="8">III</collectionCode>
; tergum II sometimes with light reddish brown laterally and on disc and blending gradually to yellow apical marginal zone; yellow apical marginal zones of terga
<collectionCode id="7CD7AE26920C0D0916751F3BB0A4FAE0" box="[617,644,1314,1337]" pageId="9" pageNumber="8">III</collectionCode>
and
<collectionCode id="7CD7AE26920C0D0916DE1F3BB0FEFAE0" box="[706,734,1314,1337]" pageId="9" pageNumber="8">IV</collectionCode>
typically obscured by dense tomen- tum; sterna reddish brown to dark brown.
</paragraph>
<paragraph id="1A7936E3920C0D0B14CD1F7BB63FFE80" blockId="9.[161,1175,225,1785]" lastBlockId="11.[160,1168,225,1788]" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8">
Integument mostly smooth and shining between punctures, punctures of head difficult to see, those of mesosoma and metasoma clear; clypeus with small, shallow, nearly contiguous punctures, in some small areas punctures separated by up to 0.75× a puncture width, punctures somewhat faint; supraclypeal area with small, shallow, nearly contiguous punctures; lower face with small punctures separated less than a puncture width, slightly fainter and more spaced than on clypeus, and slightly sparser along orbital border and in malar space; upper face with small punctures nearly con- tiguous to contiguous medially, such punctures becoming shallower toward ocelli and giving surface a roughened appearance, laterally punctures smaller and more distinct, separated by a puncture width or less; ocellocular area with punctures of upper lat- eral face becoming denser and blending posteriorly to slightly larger punctures like those of medial face; vertex and posterior of head punctures with shallow, coarse, contiguous punctures giving surface a roughened appearance, such punctures blend- ing to minute, shallow, punctures separated by less than a puncture width on gena; postgena nearly impunctate and smooth. Mesoscutum and mesoscutellum with small contiguous punctures, punctures coarser and more irregular in shape along borders; pleura with punctures as on mesoscutum, those punctures ventrally a bit more sepa- rated, posteriorly punctures more irregularly shaped and elongate; metepisternum with small, nearly contiguous punctures; lateral surface of propodeum as on metepi- sternum; basal area of propodeum tessellate to microalveolate. Metasomal terga with minute contiguous punctures blending to imbricate integument in apical marginal zones, except anterior-facing surface of tergum I impunctate and smooth; punctures laterally blending to imbricate integument, punctures weaker on tergum
<collectionCode id="7CD7AE26920E0D0B17E41B3BB62CFEE0" box="[1016,1036,290,313]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
, tergum
<collectionCode id="7CD7AE26920E0D0B10681B3BB6B0FEE0" box="[1140,1168,290,313]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
largely impunctate and faintly imbricate; sterna finely and faintly imbricate.
</paragraph>
<caption id="4EB9666B920F0D0A14BD1FA8B07EF9D8" pageId="10" pageNumber="9" startId="10.[161,242,1457,1479]" targetBox="[165,1151,227,1434]" targetPageId="10" targetType="figure">
<paragraph id="1A7936E3920F0D0A14BD1FA8B07EF9D8" blockId="10.[161,1168,1457,1537]" pageId="10" pageNumber="9">
<emphasis id="28B2EAF1920F0D0A14BD1FA8B305FA1F" bold="true" box="[161,293,1457,1479]" pageId="10" pageNumber="9">Figures 36.</emphasis>
Workers of
<taxonomicName id="DDC64D60920F0D0A15AA1FABB054FA1F" authorityName="Engel" authorityYear="2022" box="[438,628,1457,1479]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF1920F0D0A15AA1FABB054FA1F" box="[438,628,1457,1479]" italics="true" pageId="10" pageNumber="9">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A920F0D0A169E1FABB126FA1E" box="[642,774,1457,1479]" pageId="10" pageNumber="9" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1920F0D0A17061FA8B10CFA1F" bold="true" box="[794,812,1457,1478]" pageId="10" pageNumber="9">3.</emphasis>
Facial view of individual on the lighter end of the color spectrum.
<emphasis id="28B2EAF1920F0D0A163A1FD7B018FA3A" bold="true" box="[550,568,1486,1507]" pageId="10" pageNumber="9">4.</emphasis>
Facial view, darker worker.
<emphasis id="28B2EAF1920F0D0A179D1FD7B1B3FA3A" bold="true" box="[897,915,1486,1507]" pageId="10" pageNumber="9">5.</emphasis>
Dorsal habitus, darker worker.
<emphasis id="28B2EAF1920F0D0A151C1FF2B332F9D9" bold="true" box="[256,274,1515,1536]" pageId="10" pageNumber="9">6.</emphasis>
Dorsal habitus, lighter worker.
</paragraph>
</caption>
<paragraph id="1A7936E3920E0D0B14CC1B78B126F900" blockId="11.[160,1168,225,1788]" pageId="11" pageNumber="10">
Fine pubescence generally consisting of sparse to scatered, minute to short, ap- pressed or decumbent yellow setae, such setae often simple but sometimes with min- ute branches or plumose, as well as areas of short, white, plumose setae such as dense on pronotal lobe, largely obscuring integument on metepisternum and lateral surface of propodeum; fine setae sparse on clypeus, more numerous on face and progressively longer on upper face and vertex; scape with sparse, minute, appressed setae and two erect to suberect, simple bristles at base (frequently obscured or even sometimes bro- ken off); terga I
<collectionCode id="7CD7AE26920E0D0B1542185BB35AFD80" box="[350,378,578,601]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
with plumose yellow tomentum, on tergum I such setae are diffuse, more suberect, and not obscuring integument, frequently worn such that in some spec- imens they are completely absent to only scatered in irregular places (most easily seen in profile), tergum II with such setae diffuse to sometimes lacking laterally (most easily seen in profile), but dense in apical marginal zone, terga
<collectionCode id="7CD7AE26920E0D0B174D18DBB14CFD00" box="[849,876,706,729]" pageId="11" pageNumber="10">III</collectionCode>
<collectionCode id="7CD7AE26920E0D0B176618DBB1ADFD00" box="[890,909,706,729]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
with such tomentum dense and obscuring integument, tomentum paler and sparser on tergum
<collectionCode id="7CD7AE26920E0D0B100018FBB61BFD20" box="[1052,1083,738,761]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
; sterna II
<collectionCode id="7CD7AE26920E0D0B14A3191BB2F2FCC0" box="[191,210,770,793]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
with scopa composed of elongate, erect, simple setae with characteristically wavy apices, such scopal setae yellowish. Bristles (thick, typically erect, often simple or with minute pectinate branches apically) typically sparse on body except in certain areas as noted. Labrum with sparse, erect, simple, yellow to fulvous bristles, those medially longest; vertex with numerous erect, simple, fulvous to sometimes fuscous bristles, longest medially and extending across vertex to just posterior to upper orbit; prono- tal dorsolateral margin near and anterior to pronotal lobe with bristles, sometimes arched apically, yellow to fulvous; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, bristles largely fulvous to sometimes fuscous along anterior margin, typically fuscous bristles intermixed with fulvous bristles in an- terolateral corner but sometimes intermixed along entire anterior margin, bristles on lateral border typically fuscous to black; tegula with short, suberect bristles anteriorly, such bristles typically yellow or fulvous, more uncommonly fuscous; mesoscutellum with abundant, elongate, erect bristles posteriorly, such bristles minutely pectinate apically, mesoscutellar bristles typically light fulvous, sometimes more darkly ful- vous; mesepisternum with scatered, subdecumbent fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, suberect to subdecumbent bristles extending ventrally from pronotal lobe, such bristles fulvous to dark fulvous; lateral surface of propodeum with scatered, long, erect, yellow to fulvous bristles emerging amid dense, appressed, white, plumose setae. Legs with abundant bristles, those on basal podites typically yellow to fulvous, those apically on tibiae and tarsi typically black, sometimes with black bristles apically on femora, particularly metafemur, distitarsi typically with fulvous bristles dominant; metatibia with elongate simple black corbicular bristles and margins with long, arched, largely simple, black bristles; penicillum black; rastellum fulvous to fuscous; metabasitarsus with scatered erect black bristles on prolateral surface and numerous black bristles on margins intermixed with fulvous bristles, those of inferior margin most numerous; a few black bristles on distitarsomeres. Metasoma with numerous, prominent, suberect to subdecumbent, yellow to fulvous bristles on terga
<collectionCode id="7CD7AE26920E0D0B17341C9BB163F940" box="[808,835,1666,1689]" pageId="11" pageNumber="10">III</collectionCode>
<collectionCode id="7CD7AE26920E0D0B174C1C9BB143F940" box="[848,867,1666,1689]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
arising amid tomentum, such bristles sparse to absent on tergum II, and more scatered on tergum
<collectionCode id="7CD7AE26920E0D0B10031CBBB61BF960" box="[1055,1083,1698,1721]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
except longer and more numerous laterally and on margin.
</paragraph>
<paragraph id="1A7936E3920E0D0B14CC1CFBB30FF920" blockId="11.[160,1168,225,1788]" box="[208,303,1762,1788]" pageId="11" pageNumber="10">
<specimenCount id="0CC0FD6A920E0D0B14CC1CFBB2C4F925" box="[208,228,1762,1788]" count="1" pageId="11" pageNumber="10" type="female"></specimenCount>
:
<emphasis id="28B2EAF1920E0D0B14ED1CFBB308F920" box="[241,296,1762,1785]" italics="true" pageId="11" pageNumber="10">Latet</emphasis>
.
</paragraph>
<caption id="4EB9666B92090D0C14BC1F54B19CFA45" pageId="12" pageNumber="11" startId="12.[160,241,1357,1379]" targetBox="[160,1156,223,1346]" targetPageId="12" targetType="figure">
<paragraph id="1A7936E392090D0C14BC1F54B19CFA45" blockId="12.[160,1168,1357,1437]" pageId="12" pageNumber="11">
<emphasis id="28B2EAF192090D0C14BC1F54B30EFABB" bold="true" box="[160,302,1357,1379]" pageId="12" pageNumber="11">Figures 711.</emphasis>
Workers of
<taxonomicName id="DDC64D6092090D0C15A41F54B054FABB" authorityName="Engel" authorityYear="2022" box="[440,628,1357,1378]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF192090D0C15A41F54B054FABB" box="[440,628,1357,1378]" italics="true" pageId="12" pageNumber="11">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92090D0C169C1F54B122FABA" box="[640,770,1357,1379]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192090D0C170E1F54B104FABB" bold="true" box="[786,804,1357,1378]" pageId="12" pageNumber="11">7.</emphasis>
Dorsal view of mesosoma, lighter worker.
<emphasis id="28B2EAF192090D0C151C1F73B332FAA6" bold="true" box="[256,274,1386,1407]" pageId="12" pageNumber="11">8.</emphasis>
Dorsal view of mesosoma, darker worker.
<emphasis id="28B2EAF192090D0C16F81F73B0D6FAA6" bold="true" box="[740,758,1386,1407]" pageId="12" pageNumber="11">9.</emphasis>
Mesoscutellum.
<emphasis id="28B2EAF192090D0C17AD1F73B1EFFAA6" bold="true" box="[945,975,1386,1407]" pageId="12" pageNumber="11">10.</emphasis>
Prolateral surface of metatibia, metatarsus, and metapretarsus.
<emphasis id="28B2EAF192090D0C16981F9FB082FA42" bold="true" box="[644,674,1414,1435]" pageId="12" pageNumber="11">11.</emphasis>
Apical view of metasoma.
</paragraph>
</caption>
<paragraph id="1A7936E392090D0C14CC1FD7B30FFA3C" blockId="12.[160,1168,1486,1797]" box="[208,303,1486,1512]" pageId="12" pageNumber="11">
<specimenCount id="0CC0FD6A92090D0C14CC1FD7B2C4FA31" box="[208,228,1486,1512]" count="1" pageId="12" pageNumber="11" type="male"></specimenCount>
:
<emphasis id="28B2EAF192090D0C14ED1FD7B308FA3C" box="[241,296,1486,1509]" italics="true" pageId="12" pageNumber="11">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892090D0D14CC1FF4B65FFE80" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="materials_examined">
<paragraph id="1A7936E392090D0C14CC1FF4B396F9FC" blockId="12.[160,1168,1486,1797]" pageId="12" pageNumber="11">
<materialsCitation id="AAAE3CBE92090D0C14CC1FF4B396F9FC" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" location="Dagua" pageId="12" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D884192090D0C14CC1FF4B368F9DC" box="[208,328,1517,1541]" pageId="12" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392090D0C15731FF4B3C5F9DC" box="[367,485,1517,1541]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
: Prov. Valle,
<quantity id="DD3E9B0692090D0C16631FF4B09AF9DC" box="[639,698,1517,1541]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="11" unit="km" value="5.0">5 km</quantity>
SE of
<location id="1F19603892090D0C171A1FF7B176F9DF" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C171A1FF7B176F9DF" box="[774,854,1518,1542]" country="Colombia" name="Dagua" pageId="12" pageNumber="11">Dagua</location>
,
<date id="6E78102392090D0C177F1FF4B1C1F9DC" box="[867,993,1517,1541]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C177F1FF4B1C1F9DC" box="[867,993,1517,1541]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
,
<collectorName id="B733533592090D0C17F11FF7B654F9DC" box="[1005,1140,1517,1541]" pageId="12" pageNumber="11">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C14BC1C14B36CF9FC" box="[160,332,1549,1573]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15461C14B38AF9FC" box="[346,426,1549,1573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392090D0D14CC1C34B65FFE80" blockId="12.[160,1168,1486,1797]" lastBlockId="13.[160,1168,225,601]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
<materialsCitation id="AAAE3CBE92090D0C14CC1C34B3F4F9BC" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" location="Prov. Valle" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<typeStatus id="C57D884192090D0C14CC1C34B36EF99C" box="[208,334,1581,1605]" pageId="12" pageNumber="11" type="paratype">PARATYPES</typeStatus>
: 14⚲⚲,
<collectingCountry id="62D1767392090D0C15871C34B031F99C" box="[411,529,1581,1605]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16071C37B0BDF99C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16071C37B0BDF99C" box="[539,669,1581,1605]" country="Colombia" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<locationDeviation id="0BCB2FC192090D0C16B41C34B159F99F" box="[680,889,1581,1606]" location="Dagua" pageId="12" pageNumber="11">
<quantity id="DD3E9B0692090D0C16B41C34B0C1F99C" box="[680,737,1581,1605]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="11" unit="km" value="5.0">5 km</quantity>
SE of Dagua
</locationDeviation>
,
<date id="6E78102392090D0C17981C34B620F99C" box="[900,1024,1581,1605]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C17981C34B620F99C" box="[900,1024,1581,1605]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
,
<collectorName id="B733533592090D0C10161C37B6B0F99C" box="[1034,1168,1581,1605]" pageId="12" pageNumber="11">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C14A21C54B34CF9BC" box="[190,364,1613,1637]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15611C54B3ECF9BC" box="[381,460,1613,1637]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16001C54B3B4F95C" collectingDate="1976-10-06" collectionCode="SEMC" collectorName="Breed" country="Colombia" location="Valle" municipality="Loboguerrera" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16001C54B0B2F9BC" box="[540,658,1613,1637]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16831C54B0FCF9BC" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16831C54B0FCF9BC" box="[671,732,1613,1637]" country="Colombia" municipality="Loboguerrera" name="Valle" pageId="12" pageNumber="11">Valle</location>
, near
<collectingMunicipality id="FA1DAC9992090D0C173A1C54B1EAF9BC" box="[806,970,1613,1637]" pageId="12" pageNumber="11">Loboguerrera</collectingMunicipality>
,
<date id="6E78102392090D0C17C41C54B679F9BC" box="[984,1113,1613,1637]" pageId="12" pageNumber="11" value="1976-10-06">
<collectingDate id="7E3CE9CB92090D0C17C41C54B679F9BC" box="[984,1113,1613,1637]" pageId="12" pageNumber="11" value="1976-10-06">6 Oct 1976</collectingDate>
</date>
, Mi- chael
<collectorName id="B733533592090D0C14FB1C74B30CF95C" box="[231,300,1645,1669]" pageId="12" pageNumber="11">Breed</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15201C74B3ABF95C" box="[316,395,1645,1669]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 14⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C15F51C74B078F97C" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="Breed &amp; C. D. Michener" country="Colombia" elevation="740" location="Prov. Valle" municipality="Loboguerrera" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C15F51C74B07FF95C" box="[489,607,1645,1669]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16701C77B0D1F95C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16701C77B0D1F95C" box="[620,753,1645,1669]" country="Colombia" municipality="Loboguerrera" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C16E31C74B183F95C" box="[767,931,1645,1669]" pageId="12" pageNumber="11">Loboguerrera</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C17AD1C74B1DCF95C" box="[945,1020,1645,1669]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="11" unit="m" value="740.0">
<elevation id="91EBD1D092090D0C17AD1C74B1DCF95C" box="[945,1020,1645,1669]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="11" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<date id="6E78102392090D0C10161C74B6ACF95C" box="[1034,1164,1645,1669]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C10161C74B6ACF95C" box="[1034,1164,1645,1669]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
, M.D.
<collectorName id="B733533592090D0C14FE1C94B307F97C" box="[226,295,1677,1701]" pageId="12" pageNumber="11">Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C15541C94B3D3F97C" box="[328,499,1677,1701]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C161D1C94B070F97C" box="[513,592,1677,1701]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 7⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16861C94B084F91C" collectingDate="1977-02-16" collectionCode="SEMC" collectorName="Breed &amp; C. D. Michener" country="Colombia" elevation="1000" location="Prov. Valle" municipality="Cali" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16861C94B130F97C" box="[666,784,1677,1701]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C17061C97B1BDF97C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C17061C97B1BDF97C" box="[794,925,1677,1701]" country="Colombia" municipality="Cali" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C17B41C94B1FAF97C" box="[936,986,1677,1701]" pageId="12" pageNumber="11">Cali</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C17F81C94B61AF97C" box="[996,1082,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="11" unit="m" value="1000.0">
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</quantity>
,
<date id="6E78102392090D0C10591C94B2F8F91C" pageId="12" pageNumber="11" value="1977-02-16">
<collectingDate id="7E3CE9CB92090D0C10591C94B2F8F91C" pageId="12" pageNumber="11" value="1977-02-16">16 Feb 1977</collectingDate>
</date>
, M.D.
<collectorName id="B733533592090D0C153B1CB4B34CF91C" box="[295,364,1709,1733]" pageId="12" pageNumber="11">Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C158C1CB4B01DF91C" box="[400,573,1709,1733]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C16511CB4B0BCF91C" box="[589,668,1709,1733]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16F51CB4B1BDF93C" collectingDate="1976-09-23" collectionCode="SEMC" collectorName="Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1500" location="Valle" municipality="Lago Calima" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16F51CB4B17FF91C" box="[745,863,1709,1733]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C17771CB4B188F91C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C17771CB4B188F91C" box="[875,936,1709,1733]" country="Colombia" municipality="Lago Calima" name="Valle" pageId="12" pageNumber="11">Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C17A81CB7B66EF91C" box="[948,1102,1709,1734]" pageId="12" pageNumber="11">Lago Calima</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C10471CB4B29AF93C" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="1500.0">
<elevation id="91EBD1D092090D0C10471CB4B29AF93C" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, IX-23-76 [
<date id="6E78102392090D0C152E1CD4B033F93C" box="[306,531,1741,1765]" pageId="12" pageNumber="11" value="1976-09-23">
<collectingDate id="7E3CE9CB92090D0C152E1CD4B033F93C" box="[306,531,1741,1765]" pageId="12" pageNumber="11" value="1976-09-23">23 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592090D0C163F1CD4B072F93C" box="[547,594,1741,1765]" pageId="12" pageNumber="11">Bell</collectorName>
,
<collectorName id="B733533592090D0C16401CD4B083F93C" box="[604,675,1741,1765]" pageId="12" pageNumber="11">Breed</collectorName>
, &amp;
<collectorName id="B733533592090D0C16D41CD4B118F93C" box="[712,824,1741,1765]" pageId="12" pageNumber="11">Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C175A1CD4B1B5F93C" box="[838,917,1741,1765]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 1⚲,
<materialsCitation id="AAAE3CBE92090D0D17D31CD4B356FF20" collectingDate="1976-09-27" collectionCode="SEMC" collectorName="Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1300" lastPageId="13" lastPageNumber="12" location="Valle" municipality="Buenaventura Rd." pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C17D31CD4B665F93C" box="[975,1093,1741,1765]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C10531CD4B6ACF93C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C10531CD4B6ACF93C" box="[1103,1164,1741,1765]" country="Colombia" municipality="Buenaventura Rd." name="Valle" pageId="12" pageNumber="11">Valle</location>
, km 33, old
<collectingMunicipality id="FA1DAC9992090D0C15371CF7B025F8DC" box="[299,517,1773,1797]" pageId="12" pageNumber="11">Buenaventura Rd.</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C16081CF4B04DF8DC" box="[532,621,1773,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="11" unit="m" value="1300.0">
<elevation id="91EBD1D092090D0C16081CF4B04DF8DC" box="[532,621,1773,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="11" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, IX-27-76 [
<date id="6E78102392090D0C16F11CF4B1F5F8DC" box="[749,981,1773,1797]" pageId="12" pageNumber="11" value="1976-09-27">
<collectingDate id="7E3CE9CB92090D0C16F11CF4B1F5F8DC" box="[749,981,1773,1797]" pageId="12" pageNumber="11" value="1976-09-27">27 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592090D0C17F51CF4B638F8DC" box="[1001,1048,1773,1797]" pageId="12" pageNumber="11">Bell</collectorName>
,
<collectorName id="B733533592090D0C103A1CF4B64DF8DC" box="[1062,1133,1773,1797]" pageId="12" pageNumber="11">Breed</collectorName>
, &amp;
<collectorName id="B733533592080D0D14BC1AF8B330FF20" box="[160,272,225,249]" pageId="13" pageNumber="12">Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D15031AF8B34EFF20" box="[287,366,225,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 21⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D15DB1AF8B044FEC0" collectingDate="1976-09-22" collectionCode="SEMC" collectorName="Valle &amp; El Carmen &amp; Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1370" location="Colombia" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D15DB1AF8B01DFF20" box="[455,573,225,249]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<collectorName id="B733533592080D0D16541AF8B0A5FF20" box="[584,645,225,249]" pageId="13" pageNumber="12">Valle</collectorName>
,
<collectorName id="B733533592080D0D168D1AF8B131FF20" box="[657,785,225,249]" pageId="13" pageNumber="12">El Carmen</collectorName>
,
<quantity id="DD3E9B0692080D0D17001AF8B153FF20" box="[796,883,225,249]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="13" pageNumber="12" unit="m" value="1370.0">
<elevation id="91EBD1D092080D0D17001AF8B153FF20" box="[796,883,225,249]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="13" pageNumber="12" unit="m" value="1370.0">1370 m</elevation>
</quantity>
, IX-22-76 [
<date id="6E78102392080D0D17F21AF8B2FAFEC0" pageId="13" pageNumber="12" value="1976-09-22">
<collectingDate id="7E3CE9CB92080D0D17F21AF8B2FAFEC0" pageId="13" pageNumber="12" value="1976-09-22">22 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592080D0D14F71B18B33AFEC0" box="[235,282,257,281]" pageId="13" pageNumber="12">Bell</collectorName>
,
<collectorName id="B733533592080D0D15381B18B34BFEC0" box="[292,363,257,281]" pageId="13" pageNumber="12">Breed</collectorName>
, &amp;
<collectorName id="B733533592080D0D158C1B18B020FEC0" box="[400,512,257,281]" pageId="13" pageNumber="12">Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D16111B18B07CFEC0" box="[525,604,257,281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 20⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D16AF1B18B1A9FEE0" collectingDate="1977-02-14" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="1200" location="Prov. Valle" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D16AF1B18B109FEC0" box="[691,809,257,281]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<location id="1F19603892080D0D172E1B1BB194FEC0" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892080D0D172E1B1BB194FEC0" box="[818,948,257,281]" country="Colombia" name="Prov. Valle" pageId="13" pageNumber="12">Prov. Valle</location>
,
<quantity id="DD3E9B0692080D0D17A21B18B625FEC0" box="[958,1029,257,281]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="12" unit="km" value="10.0">10 km</quantity>
SW Jamun- di,
<date id="6E78102392080D0D14DB1B38B375FEE0" box="[199,341,289,313]" pageId="13" pageNumber="12" value="1977-02-14">
<collectingDate id="7E3CE9CB92080D0D14DB1B38B375FEE0" box="[199,341,289,313]" pageId="13" pageNumber="12" value="1977-02-14">14 Feb 1977</collectingDate>
</date>
,
<quantity id="DD3E9B0692080D0D157E1B38B399FEE0" box="[354,441,289,313]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="12" unit="m" value="1200.0">
<elevation id="91EBD1D092080D0D157E1B38B399FEE0" box="[354,441,289,313]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="12" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<collectorName id="B733533592080D0D15DA1B3BB06FFEE0" box="[454,591,289,313]" pageId="13" pageNumber="12">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592080D0D16681B38B101FEE0" box="[628,801,289,313]" pageId="13" pageNumber="12">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D172D1B38B1A0FEE0" box="[817,896,289,313]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 5⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D17D31B38B65CFE80" collectingDate="1977-02-14" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="1100" location="Prov. Valle" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D17D31B38B665FEE0" box="[975,1093,289,313]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<location id="1F19603892080D0D104D1B3BB2FDFE80" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892080D0D104D1B3BB2FDFE80" country="Colombia" name="Prov. Valle" pageId="13" pageNumber="12">Prov. Valle</location>
,
<locationDeviation id="0BCB2FC192080D0D14F41B5BB3E0FE80" box="[232,448,321,345]" location="Jamundi" pageId="13" pageNumber="12">
<quantity id="DD3E9B0692080D0D14F41B5BB302FE80" box="[232,290,321,345]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="12" unit="km" value="7.0">7 km</quantity>
SW Jamundi
</locationDeviation>
,
<date id="6E78102392080D0D15D71B58B076FE80" box="[459,598,321,345]" pageId="13" pageNumber="12" value="1977-02-14">
<collectingDate id="7E3CE9CB92080D0D15D71B58B076FE80" box="[459,598,321,345]" pageId="13" pageNumber="12" value="1977-02-14">14 Feb 1977</collectingDate>
</date>
,
<quantity id="DD3E9B0692080D0D167C1B58B096FE80" box="[608,694,321,345]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="12" unit="m" value="1100.0">
<elevation id="91EBD1D092080D0D167C1B58B096FE80" box="[608,694,321,345]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="12" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<collectorName id="B733533592080D0D16DD1B5BB168FE80" box="[705,840,321,345]" pageId="13" pageNumber="12">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592080D0D17761B58B635FE80" box="[874,1045,321,345]" pageId="13" pageNumber="12">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D10381B58B654FE80" box="[1060,1140,321,345]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892080D0D14CC1B78B1DEFD80" pageId="13" pageNumber="12" type="etymology">
<paragraph id="1A7936E392080D0D14CC1B78B1DEFD80" blockId="13.[160,1168,225,601]" pageId="13" pageNumber="12">
ETYMOLOGY: This species has been frequently referenced in the literature but has never been described formally. In order to maintain some continuity the original name is maintained. The specific epithet was first proposed by Herbert F. Schwarz (1883 1960) for an unnamed member of the Eder family. Henry J. (18701949) and Walter Eder (
<emphasis id="28B2EAF192080D0D14F41BF8B30CFE20" box="[232,300,481,505]" italics="true" pageId="13" pageNumber="12">floruit</emphasis>
1927), son and grandson of the pioneering sugar industrialist James Mar- tin Eder (18381921), provided logistical assistance to Schwarz during his 1935 collect- ing visit to Cali (
<bibRefCitation id="7E574B1292080D0D157E1838B02DFDE0" author="Schwarz, H. P." box="[354,525,545,569]" pageId="13" pageNumber="12" pagination="1 - 546" refId="ref22362" refString="Schwarz, H. P. 1948. Stingless bees (Meliponidae) of the Western Hemisphere. Lestrimelita and the following subgenera of Trigona: Trigona, Paratrigona, Schwarziana, Parapartamona, Cephalotrigona, Oxytrigona, Scaura, and Mourella. Bulletin of the American Museum of Natural History 90: i - xviii, 1 - 546, + 8 pls." type="book chapter" year="1948">Schwarz, 1948</bibRefCitation>
). Herein, I take the presumption of proposing that the specific epithet honors the patriarch of the family in
<collectingCountry id="62D1767392080D0D17141858B15EFD80" box="[776,894,577,601]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
, J.M. Eder.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,225 @@
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<mods:identifier id="F311B22E3106856EB0AED8BDCDEA09EA" type="ZooBank">urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB</mods:identifier>
</mods:mods>
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<subSubSection id="52DC656892080D0D15891898B61CFD60" pageId="13" pageNumber="12" type="nomenclature">
<paragraph id="1A7936E392080D0D15891898B1BBFD40" blockId="13.[405,923,641,665]" box="[405,923,641,665]" pageId="13" pageNumber="12">
<heading id="4131818F92080D0D15891898B1BBFD40" box="[405,923,641,665]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="12" reason="2">
<taxonomicName id="DDC64D6092080D0D15891898B123FD40" authority="Engel" authorityName="Engel" authorityYear="2022" box="[405,771,641,665]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="gonzalezi" status="sp. nov.">
<emphasis id="28B2EAF192080D0D15891898B097FD40" bold="true" box="[405,695,641,665]" italics="true" pageId="13" pageNumber="12">Scaptotrigona gonzalezi</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92080D0D1712189BB1BBFD40" box="[782,923,641,665]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392080D0D14E818BAB61CFD60" blockId="13.[244,1084,675,729]" box="[244,1084,675,697]" pageId="13" pageNumber="12">
ZooBank:
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892080D0D165318D8B0C1FD00" box="[591,737,705,729]" pageId="13" pageNumber="12" type="description">
<paragraph id="1A7936E392080D0D165318D8B0C1FD00" blockId="13.[244,1084,675,729]" box="[591,737,705,729]" pageId="13" pageNumber="12">
(
<figureCitation id="82FD2A6692080D0D164B18D8B0FAFD00" box="[599,730,705,729]" captionStart-0="Figures 1214" captionStart-1="Figures 1517" captionStartId-0="14.[160,241,1457,1479]" captionStartId-1="15.[160,241,1316,1338]" captionTargetBox-0="[175,1138,234,1441]" captionTargetBox-1="[179,1159,241,1302]" captionTargetId-0="figure-116@14.[160,1168,224,1451]" captionTargetId-1="figure-167@15.[160,1168,224,1310]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Figures 1214. Worker of Scaptotrigona gonzalezi, new species. 12. Lateral habitus, darker work- er. 13. Facial view, darker worker. 14. Prolateral surface of metatibia, metatarsus, and metap- retarsus, darker worker." captionText-1="Figures 1517. Worker of Scaptotrigona gonzalezi, new species. 15. Lateral habitus, lighter work- er. 16. Facial view, lighter worker. 17. Mesosomal dorsum, lighter worker." pageId="13" pageNumber="12">Figs. 1217</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892080D0D14CC1918B600FC60" pageId="13" pageNumber="12" type="diagnosis">
<paragraph id="1A7936E392080D0D14CC1918B600FC60" blockId="13.[160,1168,769,1786]" pageId="13" pageNumber="12">
DIAGNOSIS: This is one of a few South American species in which the genal tooth is lacking. It is similar to
<taxonomicName id="DDC64D6092080D0D15B7193BB3DAFCE0" authorityName="Engel" authorityYear="2022" box="[427,506,801,825]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192080D0D15B7193BB3DAFCE0" box="[427,506,801,825]" italics="true" pageId="13" pageNumber="12">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192080D0D16151938B057FCE0" box="[521,631,801,825]" italics="true" pageId="13" pageNumber="12">vide supra</emphasis>
), but has a noticeably longer malar space (
<figureCitation id="82FD2A6692080D0D107F1938B29CFC80" captionStart="Figures 1214" captionStartId="14.[160,241,1457,1479]" captionTargetBox="[175,1138,234,1441]" captionTargetId="figure-116@14.[160,1168,224,1451]" captionTargetPageId="14" captionText="Figures 1214. Worker of Scaptotrigona gonzalezi, new species. 12. Lateral habitus, darker work- er. 13. Facial view, darker worker. 14. Prolateral surface of metatibia, metatarsus, and metap- retarsus, darker worker." pageId="13" pageNumber="12">Fig. 13</figureCitation>
), yellowish to fulvous bristles on the vertex and anterolateral areas of the mesoscu- tum, and nearly entirely fulvous bristles on the metabasitarsus (
<figureCitation id="82FD2A6692080D0D17841978B1CAFCA0" box="[920,1002,865,889]" captionStart="Figures 1214" captionStartId="14.[160,241,1457,1479]" captionTargetBox="[175,1138,234,1441]" captionTargetId="figure-116@14.[160,1168,224,1451]" captionTargetPageId="14" captionText="Figures 1214. Worker of Scaptotrigona gonzalezi, new species. 12. Lateral habitus, darker work- er. 13. Facial view, darker worker. 14. Prolateral surface of metatibia, metatarsus, and metap- retarsus, darker worker." pageId="13" pageNumber="12">Fig. 14</figureCitation>
). In addition, the concavity interrupting the preoccipital carina laterally is spanned by a continua- tion of the translucent lamella of the dorsal portion of the preoccipital ridge.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892080D1014CC19D8B30FFD20" lastPageId="16" lastPageNumber="15" pageId="13" pageNumber="12" type="description">
<paragraph id="1A7936E392080D0D14CC19D8B66AFAE0" blockId="13.[160,1168,769,1786]" pageId="13" pageNumber="12">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092080D0D165F19DBB0B6FC00" authorityName="Engel" authorityYear="2022" box="[579,662,961,985]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192080D0D165F19DBB0B6FC00" box="[579,662,961,985]" italics="true" pageId="13" pageNumber="12">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192080D0D16B519D8B13CFC00" box="[681,796,961,985]" italics="true" pageId="13" pageNumber="12">vide supra</emphasis>
), with the following modifica- tions: ⚲: Total body length approximately
<quantity id="DD3E9B0692080D0D168C19F8B138FC20" box="[656,792,993,1017]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="13" pageNumber="12" unit="mm" value="6.25" valueMax="6.4" valueMin="6.1">6.16.4 mm</quantity>
, forewing length (to base of hu- meral sclerite) 6.0
<quantity id="DD3E9B0692080D0D159C1E18B3FAFBC0" box="[384,474,1025,1049]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="13" pageNumber="12" unit="mm" value="6.1">6.1 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B0692080D0D174D1E18B1D5FBC0" box="[849,1013,1025,1049]" metricMagnitude="-3" metricUnit="m" metricValue="2.355" metricValueMax="2.38" metricValueMin="2.33" pageId="13" pageNumber="12" unit="mm" value="2.355" valueMax="2.38" valueMin="2.33">2.332.38 mm</quantity>
, length 2.00
<quantity id="DD3E9B0692080D0D14BC1E38B329FBE0" box="[160,265,1057,1081]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="13" pageNumber="12" unit="mm" value="2.08">2.08 mm</quantity>
; compound eye length
<quantity id="DD3E9B0692080D0D16341E38B0EDFBE0" box="[552,717,1057,1081]" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.42" metricValueMin="1.39" pageId="13" pageNumber="12" unit="mm" value="1.4049999999999998" valueMax="1.42" valueMin="1.39">1.391.42 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B0692080D0D10391E38B2F1FB80" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.65" metricValueMin="1.63" pageId="13" pageNumber="12" unit="mm" value="1.64" valueMax="1.65" valueMin="1.63">1.631.65 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B0692080D0D16061E58B09DFB80" box="[538,701,1089,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.53" metricValueMin="1.52" pageId="13" pageNumber="12" unit="mm" value="1.525" valueMax="1.53" valueMin="1.52">1.521.53 mm</quantity>
. Scape length
<quantity id="DD3E9B0692080D0D17761E58B1F0FB80" box="[874,976,1089,1113]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="13" pageNumber="12" unit="mm" value="0.91">0.91 mm</quantity>
, as long as toru- locellar distance, torulocellar distance
<quantity id="DD3E9B0692080D0D167A1E78B0EEFBA0" box="[614,718,1121,1145]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="13" pageNumber="12" unit="mm" value="0.91">0.91 mm</quantity>
. Clypeus approximately 1.7× as wide as long, length
<quantity id="DD3E9B0692080D0D15491E98B39CFB40" box="[341,444,1153,1177]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="13" pageNumber="12" unit="mm" value="0.61">0.61 mm</quantity>
, width
<quantity id="DD3E9B0692080D0D160A1E98B05EFB40" box="[534,638,1153,1177]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="13" pageNumber="12" unit="mm" value="1.06">1.06 mm</quantity>
. Malar area approximately 1.92.0× flagellar diameter or 0.880.92× basal mandibular width. Preoccipital ridge carinate, weakly lamellate dorsally, with bordering medial indentation shallower than lateral indenta- tions, laterally concavity interrupting carina comparatively shallow, lower margin of concavity with carina forming rounded angle, without sharply acute lamellate projec- tion (genal tooth) extending upward, concavity spanned by translucent lamella.
</paragraph>
<paragraph id="1A7936E392080D0F14CC1F5BB622F960" blockId="13.[160,1168,769,1786]" lastBlockId="15.[160,1168,1409,1785]" lastPageId="15" lastPageNumber="14" pageId="13" pageNumber="12">
Integument of labiomaxillary complex orange; clypeus largely orange to yellow orange except sometimes with small, triangular brown mark at mediobasal margin and apicolateral corners, epistomal sulcus well delineated by brown to black and sub- antennal areas sometimes infused with brown; supraclypeal area yellow orange to orange; malar space orange to yellow orange although sometimes blending to dark brown apically near mandibular base; face below tangent of antennal toruli orange to yellow orange, sometimes slightly darker toward lateral apical portion of episto- mal sulcus; scape yellow orange to orange except with brown dorsoapically; pedi- cel and flagellum typically dark brown above, with underside of flagellum orange; face above antennal toruli orange to yellow orange below, blending to dark brown or black above, orange area either extending across width of face or with areas of dark brown extending downward to antennal toruli remainder of face dark brown to black, sometimes with small areas of light brown immediately above antennal toruli; vertex, posterior of head, and uppermost gena dark brown to black; remainder of gena and postgena orange to yellow orange. Mesosoma dark brown to black except pronotum largely brown to orange and pronotal lobe orange, mesoscutellum either largely or- ange or black blending to fulvous or orange apical margin, mesepisternum with areas of orange to fulvous on hypoepimeral area and sometimes upper mesepisternum an- terior to hypoepimeral area, and sometimes as orange patch posteriorly along mese- pisternal-metepisternal suture; legs typically yellow orange to dark reddish brown, lighter morphs with legs almost wholly yellow orange except patches of dark brown on femora and centrally on prolateral surfaces of meso- and metatibiae, dark morphs with legs largely reddish brown except lighter on coxae, trochanters, femoral basally, protibia and extreme bases of meso- and metatibiae, and all tarsi. Wing membranes lightly parchment colored, faintly darker around marginal cell apex. Metasoma gener- ally dark brown to black except anterior-facing surface of tergum I typically reddish brown, dorsal-facing disc of tergum I medially lighter orange brown to orange, terga II
<collectionCode id="7CD7AE26920A0D0F14A31C7BB2FBF9A0" box="[191,219,1634,1657]" pageId="15" pageNumber="14">IV</collectionCode>
with variably developed apical marginal zones of orange brown to yellow or- ange, typically large on tergum II and narrow on remaining terga; sterna dark brown to reddish brown except orange in apical portions from which setae arise.
</paragraph>
<caption id="4EB9666B920B0D0E14BC1FA8B381F9D8" pageId="14" pageNumber="13" startId="14.[160,241,1457,1479]" targetBox="[175,1138,234,1441]" targetPageId="14" targetType="figure">
<paragraph id="1A7936E3920B0D0E14BC1FA8B381F9D8" blockId="14.[160,1168,1457,1537]" pageId="14" pageNumber="13">
<emphasis id="28B2EAF1920B0D0E14BC1FA8B318FA1F" bold="true" box="[160,312,1457,1479]" pageId="14" pageNumber="13">Figures 1214.</emphasis>
Worker of
<taxonomicName id="DDC64D60920B0D0E15A91FABB0BCFA1F" authorityName="Engel" authorityYear="2022" box="[437,668,1457,1479]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="gonzalezi" status="sp. nov.">
<emphasis id="28B2EAF1920B0D0E15A91FABB0BCFA1F" box="[437,668,1457,1479]" italics="true" pageId="14" pageNumber="13">Scaptotrigona gonzalezi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A920B0D0E16BB1FABB108FA1E" box="[679,808,1457,1479]" pageId="14" pageNumber="13" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1920B0D0E172B1FA8B175FA1F" bold="true" box="[823,853,1457,1478]" pageId="14" pageNumber="13">12.</emphasis>
Lateral habitus, darker work- er.
<emphasis id="28B2EAF1920B0D0E14D51FD7B2C7FA3A" bold="true" box="[201,231,1486,1507]" pageId="14" pageNumber="13">13.</emphasis>
Facial view, darker worker.
<emphasis id="28B2EAF1920B0D0E163D1FD7B01FFA3A" bold="true" box="[545,575,1486,1507]" pageId="14" pageNumber="13">14.</emphasis>
Prolateral surface of metatibia, metatarsus, and metap- retarsus, darker worker.
</paragraph>
</caption>
<caption id="4EB9666B920A0D0F14BC1F3DB19CFA8E" pageId="15" pageNumber="14" startId="15.[160,241,1316,1338]" targetBox="[179,1159,241,1302]" targetPageId="15" targetType="figure">
<paragraph id="1A7936E3920A0D0F14BC1F3DB19CFA8E" blockId="15.[160,1168,1316,1367]" pageId="15" pageNumber="14">
<emphasis id="28B2EAF1920A0D0F14BC1F3DB318FAE0" bold="true" box="[160,312,1316,1338]" pageId="15" pageNumber="14">Figures 1517.</emphasis>
Worker of
<taxonomicName id="DDC64D60920A0D0F15A91F3CB0BCFAE3" authorityName="Engel" authorityYear="2022" box="[437,668,1317,1338]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gonzalezi" status="sp. nov.">
<emphasis id="28B2EAF1920A0D0F15A91F3CB0BCFAE3" box="[437,668,1317,1338]" italics="true" pageId="15" pageNumber="14">Scaptotrigona gonzalezi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A920A0D0F16BB1F3CB108FAE2" box="[679,808,1317,1339]" pageId="15" pageNumber="14" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1920A0D0F172B1F3DB175FAE0" bold="true" box="[823,853,1316,1337]" pageId="15" pageNumber="14">15.</emphasis>
Lateral habitus, lighter work- er.
<emphasis id="28B2EAF1920A0D0F14DB1F58B2C5FA8F" bold="true" box="[199,229,1345,1366]" pageId="15" pageNumber="14">16.</emphasis>
Facial view, lighter worker.
<emphasis id="28B2EAF1920A0D0F16061F58B018FA8F" bold="true" box="[538,568,1345,1366]" pageId="15" pageNumber="14">17.</emphasis>
Mesosomal dorsum, lighter worker.
</paragraph>
</caption>
<paragraph id="1A7936E3920A0D1014CC1CD8B1ABFD60" blockId="15.[160,1168,1409,1785]" lastBlockId="16.[160,1168,225,953]" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
As in
<taxonomicName id="DDC64D60920A0D0F150A1CD8B347F900" authorityName="Engel" authorityYear="2022" box="[278,359,1729,1753]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1920A0D0F150A1CD8B347F900" box="[278,359,1729,1753]" italics="true" pageId="15" pageNumber="14">S. ederi</emphasis>
</taxonomicName>
except lacking areas of short, white, plumose setae, instead such se- tae yellow to pale yellow; scape with one suberect, simple bristle at base (frequently obscured or even sometimes broken off). Vertex with numerous erect, simple, yellow to fulvous bristles, sometimes a few lightly fuscous; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow bristles, sometimes arched apically; mesoscu- tum anterior and lateral borders with erect yellow to dark fulvous bristles; tegula with short, suberect bristles anteriorly, such bristles typically yellow or lightly fulvous; me- soscutellum with abundant, elongate, erect bristles posteriorly, such bristles minutely pectinate apically, mesoscutellar bristles typically yellow to fulvous; mesepisternum with scatered, subdecumbent yellow to light fulvous bristles. Legs with abundant bristles, those on basal podites typically yellow to fulvous, those on meso- and metati- biae intermixed with fuscous bristles, except those of metatibia mostly black to dark fuscous and intermixed with fewer yellow bristles, although in lighter morph black bristles confined to prolateral surface along superior margin and corbicular surface apicad fundus; penicillum dark to light fulvous; rastellum yellow to fulvous; metaba- sitarsus with bristles yellow to light fulvous, sometimes with a few dark fulvous to lightly fuscous bristles in dark morph along proventral margin.
</paragraph>
<paragraph id="1A7936E392150D1014CC18DBB30FFD00" blockId="16.[160,1168,225,953]" box="[208,303,706,732]" pageId="16" pageNumber="15">
<specimenCount id="0CC0FD6A92150D1014CC18DBB2C4FD05" box="[208,228,706,732]" count="1" pageId="16" pageNumber="15" type="female"></specimenCount>
:
<emphasis id="28B2EAF192150D1014ED18DBB308FD00" box="[241,296,706,729]" italics="true" pageId="16" pageNumber="15">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392150D1014CC18FBB30FFD20" blockId="16.[160,1168,225,953]" box="[208,303,738,764]" pageId="16" pageNumber="15">
<specimenCount id="0CC0FD6A92150D1014CC18FBB2C4FD25" box="[208,228,738,764]" count="1" pageId="16" pageNumber="15" type="male"></specimenCount>
:
<emphasis id="28B2EAF192150D1014ED18FBB308FD20" box="[241,296,738,761]" italics="true" pageId="16" pageNumber="15">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892150D1014CC1918B3F7FCA0" pageId="16" pageNumber="15" type="materials_examined">
<paragraph id="1A7936E392150D1014CC1918B323FCE0" blockId="16.[160,1168,225,953]" pageId="16" pageNumber="15">
<materialsCitation id="AAAE3CBE92150D1014CC1918B323FCE0" collectionCode="SEMC" collectorName="D. Gomez" country="Colombia" location="Vda. La Carbonera" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Cauca" typeStatus="holotype">
<typeStatus id="C57D884192150D1014CC1918B368FCC0" box="[208,328,769,793]" pageId="16" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392150D10156D1918B3C7FCC0" box="[369,487,769,793]" name="Colombia" pageId="16" pageNumber="15">Colombia</collectingCountry>
,
<collectingRegion id="D802F80192150D1015EF1918B01FFCC0" box="[499,575,769,793]" country="Colombia" name="Cauca" pageId="16" pageNumber="15">Cauca</collectingRegion>
, Bolivar,
<location id="1F19603892150D1016AC1918B1B5FCC0" LSID="urn:lsid:plazi:treatment:926F87F592080D1015891898B678FC60:1F19603892150D1016AC1918B1B5FCC0" box="[688,917,769,793]" country="Colombia" name="Vda. La Carbonera" pageId="16" pageNumber="15" stateProvince="Cauca">Vda. La Carbonera</location>
, 2010, col.
<collectorName id="B733533592150D101004191BB6B0FCC0" box="[1048,1168,769,793]" pageId="16" pageNumber="15">D. Gómez</collectorName>
(
<collectionCode id="7CD7AE2692150D1014B41938B2D8FCE0" box="[168,248,801,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="15" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392150D1014CC195BB3F7FCA0" blockId="16.[160,1168,225,953]" pageId="16" pageNumber="15">
<materialsCitation id="AAAE3CBE92150D1014CC195BB3F3FCA0" collectingDate="2010-06-04" collectionCode="SEMC" collectorName="El Patia &amp; G. Zambrano" country="Colombia" location="Carretera Bomba Chontural" municipality="El Patia" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Cauca" typeStatus="paratype">
<typeStatus id="C57D884192150D1014CC195BB363FC83" box="[208,323,834,858]" pageId="16" pageNumber="15" type="paratype">PARATYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392150D1015701958B3C2FC80" box="[364,482,833,857]" name="Colombia" pageId="16" pageNumber="15">Colombia</collectingCountry>
,
<collectingRegion id="D802F80192150D1015F21958B01AFC80" box="[494,570,833,857]" country="Colombia" name="Cauca" pageId="16" pageNumber="15">Cauca</collectingRegion>
,
<collectorName id="B733533592150D10165B1958B085FC80" box="[583,677,833,857]" pageId="16" pageNumber="15">
<collectingMunicipality id="FA1DAC9992150D10165B1958B085FC80" box="[583,677,833,857]" pageId="16" pageNumber="15">El Patia</collectingMunicipality>
</collectorName>
,
<location id="1F19603892150D1016AD1958B1DFFC80" LSID="urn:lsid:plazi:treatment:926F87F592080D1015891898B678FC60:1F19603892150D1016AD1958B1DFFC80" box="[689,1023,833,857]" country="Colombia" municipality="El Patia" name="Carretera Bomba Chontural" pageId="16" pageNumber="15" stateProvince="Cauca">Carretera Bomba Chontural</location>
,
<date id="6E78102392150D1010101958B6ACFC80" box="[1036,1164,833,857]" pageId="16" pageNumber="15" value="2010-06-04">
<collectingDate id="7E3CE9CB92150D1010101958B6ACFC80" box="[1036,1164,833,857]" pageId="16" pageNumber="15" value="2010-06-04">4-Jun-2010</collectingDate>
</date>
, col.
<collectorName id="B733533592150D1014D31978B34CFCA0" box="[207,364,865,889]" pageId="16" pageNumber="15">G. Zambrano</collectorName>
(
<collectionCode id="7CD7AE2692150D1015671978B3EBFCA0" box="[379,459,865,889]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="15" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892150D1014CC1998B678FC60" pageId="16" pageNumber="15" type="etymology">
<paragraph id="1A7936E392150D1014CC1998B678FC60" blockId="16.[160,1168,225,953]" pageId="16" pageNumber="15">ETYMOLOGY: The specific epithet honors Victor H. Gonzalez, good friend and prominent leader in melitological research, particularly of the Colombian fauna.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,221 @@
<document id="ED3D96C13AA5076C915DD0BEB9AF053F" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F592100D1715831E78B132F9C0" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona rosellae Engel 2022, new species" docType="treatment" docVersion="1" lastPageNumber="22" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="20" updateTime="1722456085954" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="C79BE19B7674A8E929EAAF55A7B1F0D1">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="3E9A609B8909ABD55F8F9FC632FC58DD">Engel, Michael S.</mods:namePart>
<mods:affiliation id="4B91F2BE4E6A68A8BEE48AFC53152258">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:title id="8C2B7D77D82BE18DBA28678B8E152D26">Journal of Melittology</mods:title>
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<subSubSection id="52DC656892100D1515831E78B618FB40" pageId="21" pageNumber="20" type="nomenclature">
<paragraph id="1A7936E392100D1515831E78B1B1FBA0" blockId="21.[415,913,1121,1145]" box="[415,913,1121,1145]" pageId="21" pageNumber="20">
<heading id="4131818F92100D1515831E78B1B1FBA0" box="[415,913,1121,1145]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="20" reason="2">
<taxonomicName id="DDC64D6092100D1515831E78B0D9FBA0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[415,761,1121,1145]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rosellae" status="sp. nov.">
<emphasis id="28B2EAF192100D1515831E78B08CFBA0" bold="true" box="[415,684,1121,1145]" italics="true" pageId="21" pageNumber="20">Scaptotrigona rosellae</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92100D1517181E7BB1B1FBA0" box="[772,913,1121,1145]" pageId="21" pageNumber="20" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="52DC656892100D1516531EB8B0C1FB60" box="[591,737,1185,1209]" pageId="21" pageNumber="20" type="description">
<paragraph id="1A7936E392100D1516531EB8B0C1FB60" blockId="21.[248,1080,1155,1209]" box="[591,737,1185,1209]" pageId="21" pageNumber="20">
(
<figureCitation id="82FD2A6692100D15164B1EB8B0F9FB60" box="[599,729,1185,1209]" captionStart="Figures 2628" captionStartId="22.[160,241,1387,1409]" captionTargetBox="[165,1159,241,1371]" captionTargetId="figure-139@22.[160,1168,224,1381]" captionTargetPageId="22" captionText="Figures 2628. Worker of Scaptotrigona rosellae, new species. 26. Lateral habitus. 27. Facial view. 28. Dorsal view of vertex and mesosoma." pageId="21" pageNumber="20">Figs. 2628</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1514CC1EF8B0DEFAA0" pageId="21" pageNumber="20" type="diagnosis">
<paragraph id="1A7936E392100D1514CC1EF8B0DEFAA0" blockId="21.[160,1175,1249,1785]" pageId="21" pageNumber="20">
DIAGNOSIS: This species is currently known only from the lower Anchicayá River Valley of
<collectingCountry id="62D1767392100D15150F1F18B3A9FAC0" box="[275,393,1281,1305]" name="Colombia" pageId="21" pageNumber="20">Colombia</collectingCountry>
, an area known for its considerable biodiversity and endemism. The species superficially resembles a lighter form of
<taxonomicName id="DDC64D6092100D1517161F3BB17AFAE0" authorityName="Engel" authorityYear="2022" box="[778,858,1313,1337]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192100D1517161F3BB17AFAE0" box="[778,858,1313,1337]" italics="true" pageId="21" pageNumber="20">S. ederi</emphasis>
</taxonomicName>
but is immediately distin- guished by the presence of a genal tooth, the presence of weak striping on the mesos- cutum, and the generally yellow to fulvous bristles.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1714CC1F98B30FFA80" lastPageId="23" lastPageNumber="22" pageId="21" pageNumber="20" type="description">
<paragraph id="1A7936E392100D1514CC1F98B1ABF920" blockId="21.[160,1175,1249,1785]" pageId="21" pageNumber="20">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092100D1516331F9BB05DFA40" authorityName="Engel" authorityYear="2022" box="[559,637,1409,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192100D1516331F9BB05DFA40" box="[559,637,1409,1433]" italics="true" pageId="21" pageNumber="20">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192100D1516971F98B0D9FA40" box="[651,761,1409,1433]" italics="true" pageId="21" pageNumber="20">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite) 6.0
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. Head wider than long, width
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, length
<quantity id="DD3E9B0692100D1510391FD8B2F1FA20" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.03" metricValueMin="1.97" pageId="21" pageNumber="20" unit="mm" value="2.0" valueMax="2.03" valueMin="1.97">1.972.03 mm</quantity>
; compound eye length
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; upper interorbital distance
<quantity id="DD3E9B0692100D1517FB1FF8B6ABFA20" box="[999,1163,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.58" metricValueMin="1.52" pageId="21" pageNumber="20" unit="mm" value="1.55" valueMax="1.58" valueMin="1.52">1.521.58 mm</quantity>
, lower interorbital distance
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. Scape length
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, about as long as toru- locellar distance, torulocellar distance
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. Clypeus approximately 1.71.8× as wide as long, length
<quantity id="DD3E9B0692100D1515A81C58B03BF980" box="[436,539,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="21" pageNumber="20" unit="mm" value="0.64">0.64 mm</quantity>
, width
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. Malar area approximately 1.8× flagellar diameter or 0.82× basal mandibular width. Preoccipital ridge carinate, lamel- late dorsally, with bordering medial and lateral indentations deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as a sharp, lamellate tooth, upper margin of concav- ity with lamella extending ventrally a third to half of concavity.
</paragraph>
<caption id="4EB9666B92130D1614BC1F72B073FA47" pageId="22" pageNumber="21" startId="22.[160,241,1387,1409]" targetBox="[165,1159,241,1371]" targetPageId="22" targetType="figure">
<paragraph id="1A7936E392130D1614BC1F72B073FA47" blockId="22.[160,1174,1387,1438]" pageId="22" pageNumber="21">
<emphasis id="28B2EAF192130D1614BC1F72B318FA58" bold="true" box="[160,312,1387,1409]" pageId="22" pageNumber="21">Figures 2628.</emphasis>
Worker of
<taxonomicName id="DDC64D6092130D1615AE1F75B0A6FA58" authorityName="Engel" authorityYear="2022" box="[434,646,1388,1409]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="rosellae" status="sp. nov.">
<emphasis id="28B2EAF192130D1615AE1F75B0A6FA58" box="[434,646,1388,1409]" italics="true" pageId="22" pageNumber="21">Scaptotrigona rosellae</emphasis>
</taxonomicName>
,
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.
<emphasis id="28B2EAF192130D1617031F75B11DFA58" bold="true" box="[799,829,1388,1409]" pageId="22" pageNumber="21">26.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192130D1617EC1F75B62EFA58" bold="true" box="[1008,1038,1388,1409]" pageId="22" pageNumber="21">27.</emphasis>
Facial view.
<emphasis id="28B2EAF192130D1614BC1F91B29EFA44" bold="true" box="[160,190,1416,1437]" pageId="22" pageNumber="21">28.</emphasis>
Dorsal view of vertex and mesosoma.
</paragraph>
</caption>
<paragraph id="1A7936E392130D1714CC1FD0B0DCFD40" blockId="22.[160,1168,1480,1793]" lastBlockId="23.[160,1168,225,1562]" lastPageId="23" lastPageNumber="22" pageId="22" pageNumber="21">
Integument of labiomaxillary complex yellow orange; labrum yellow orange; mandible yellow to reddish orange; clypeus yellow orange, epistomal sulcus well de- lineated by orange brown to brown; supraclypeal area yellow orange; malar space yellow orange; face below upper tangent of antennal toruli yellow orange; lower frons yellow orange blending to reddish orange to brown on upper frons, separated from in- ner and upper orbit by strip of yellow orange to reddish orange; vertex reddish brown to dark brown; ocellocular area reddish brown to dark brown except laterally where separated from upper orbit; posterior of head orange with strips of reddish brown to dark brown extending from ocellocular area to preoccipital carina, preoccipital carina dark brown to black; gena and postgena yellow orange to orange; scape yellow or- ange except sometimes with brown dorsoapically; pedicel and flagellum brown, with flagellum more orange ventrally. Mesosoma largely yellow orange to orange except mesoscutum black with reddish brown laterally and posteriorly, faint longitudinal strips of reddish brown from notauli to posterior border; mesepisternum with brown to dark brown patch ventrally; basal area and posterior surface of propodeum reddish brown to dark brown; metepisternum sometimes stained brown; tegula wholly yellow orange and semi-translucent; legs yellow orange. Metasoma generally brown to dark brown except anterior-facing surface of tergum I yellow orange, dorsal-facing disc of tergum I largely yellow orange with yellow brown laterally, sometimes lateral areas dark brown and more encroaching on yellow orange disc; tergum II largely yellow orange with small areas of brown laterally, although sometimes largely dark brown and blending to orange posteriorly before broad yellow marginal zone; terga
<collectionCode id="7CD7AE2692120D171057185BB646FD80" box="[1099,1126,578,601]" pageId="23" pageNumber="22">III</collectionCode>
<collectionCode id="7CD7AE2692120D171068185BB6B0FD80" box="[1140,1168,578,601]" pageId="23" pageNumber="22">IV</collectionCode>
brown to dark brown with broad yellow marginal zones, largely obscured by tomen- tum; sterna largely yellow orange to yellow brown.
</paragraph>
<paragraph id="1A7936E392120D1714CC18BBB31BFAC0" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
As for
<taxonomicName id="DDC64D6092120D17150218BBB34EFD60" authorityName="Engel" authorityYear="2022" box="[286,366,673,697]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192120D17150218BBB34EFD60" box="[286,366,673,697]" italics="true" pageId="23" pageNumber="22">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white; metepisternum and lateral surface of propodeum obscured by dense, plumose, yellow setae; dorsal-facing surface of tergum I with largely simple setae. Bristles of vertex erect, simple, yellow to fulvous; pronotal dorsolateral margin near and anterior to pronotal lobe with long, arched, yellow bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with largely fulvous bristles, sometimes intermixed with yellow to dark fulvous bristles; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elon- gate, erect bristles posteriorly, such bristles minutely pectinate apically, discal bristles shorter, mesoscutellar bristles yellow to fulvous; mesepisternal bristles as in
<taxonomicName id="DDC64D6092120D17102119DBB6B0FC00" authorityName="Engel" authorityYear="2022" box="[1085,1168,961,985]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192120D17102119DBB6B0FC00" box="[1085,1168,961,985]" italics="true" pageId="23" pageNumber="22">S. ederi</emphasis>
</taxonomicName>
except yellow to fulvous. Legs with abundant bristles, bristles typically yellow to ful- vous except sometimes noticeably intermixed with fuscous to black bristles on prolat- eral surface of mesotibia apically, prolateral surface of mesobasitarsus, metafemur at extreme apex; corbicular bristles fuscous to black, marginal bristles intermixed with some yellow to fulvous bristles; prolateral surface of metabasitarsus with scatered fine, simple, yellow setae (noticeably thinner than marginal bristles), but without black setae, margins with black bristles intermixed with yellow to fulvous bristles; penicil- lum dark fulvous to fuscous; rastellum fulvous. Metasoma with numerous, subde- cumbent, yellow bristles on terga
<collectionCode id="7CD7AE2692120D1716351EFBB064FB20" box="[553,580,1250,1273]" pageId="23" pageNumber="22">III</collectionCode>
<collectionCode id="7CD7AE2692120D17164D1EFBB044FB20" box="[593,612,1250,1273]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="23" pageNumber="22" type="Museum">V</collectionCode>
arising from tomentum, similar bristles sparser on tergum II.
</paragraph>
<paragraph id="1A7936E392120D1714CC1F3BB30FFAE0" blockId="23.[160,1168,225,1562]" box="[208,303,1314,1340]" pageId="23" pageNumber="22">
<specimenCount id="0CC0FD6A92120D1714CC1F3BB2C4FAE5" box="[208,228,1314,1340]" count="1" pageId="23" pageNumber="22" type="female"></specimenCount>
:
<emphasis id="28B2EAF192120D1714ED1F3BB308FAE0" box="[241,296,1314,1337]" italics="true" pageId="23" pageNumber="22">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392120D1714CC1F5BB30FFA80" blockId="23.[160,1168,225,1562]" box="[208,303,1346,1372]" pageId="23" pageNumber="22">
<specimenCount id="0CC0FD6A92120D1714CC1F5BB2C4FA85" box="[208,228,1346,1372]" count="1" pageId="23" pageNumber="22" type="male"></specimenCount>
:
<emphasis id="28B2EAF192120D1714ED1F5BB308FA80" box="[241,296,1346,1369]" italics="true" pageId="23" pageNumber="22">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D1714CC1F78B3F1FA00" pageId="23" pageNumber="22" type="materials_examined">
<paragraph id="1A7936E392120D1714CC1F78B03CFA40" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
<materialsCitation id="AAAE3CBE92120D1714CC1F78B03CFA40" collectingDate="1977-02-11" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="400" location="Rio Anchicaya" pageId="23" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D884192120D1714CC1F78B368FAA0" box="[208,328,1377,1401]" pageId="23" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392120D17156A1F78B3CCFAA0" box="[374,492,1377,1401]" name="Colombia" pageId="23" pageNumber="22">Colombia</collectingCountry>
: Prov. Valle,
<location id="1F19603892120D17168D1F78B161FAA0" LSID="urn:lsid:plazi:treatment:926F87F592100D1715831E78B132F9C0:1F19603892120D17168D1F78B161FAA0" box="[657,833,1377,1401]" country="Colombia" name="Rio Anchicaya" pageId="23" pageNumber="22">Rio Anchicaya</location>
,
<quantity id="DD3E9B0692120D17174C1F78B1BDFAA0" box="[848,925,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">
<elevation id="91EBD1D092120D17174C1F78B1BDFAA0" box="[848,925,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="6E78102392120D1717B01F78B666FAA0" box="[940,1094,1377,1401]" pageId="23" pageNumber="22" value="1977-02-11">
<collectingDate id="7E3CE9CB92120D1717B01F78B666FAA0" box="[940,1094,1377,1401]" pageId="23" pageNumber="22" value="1977-02-11">11 Feb. 1977</collectingDate>
</date>
,
<collectorName id="B733533592120D1710491F7BB2C5FA40" pageId="23" pageNumber="22">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592120D17151B1F98B392FA40" box="[263,434,1409,1433]" pageId="23" pageNumber="22">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692120D1715DD1F98B031FA40" box="[449,529,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="23" pageNumber="22" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392120D1714CC1FBBB3F1FA00" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
<materialsCitation id="AAAE3CBE92120D1714CC1FBBB3EDFA00" collectingDate="1977-02-10" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="400" location="Rio Anchicaya" pageId="23" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="C57D884192120D1714CC1FBBB363FA63" box="[208,323,1442,1466]" pageId="23" pageNumber="22" type="paratype">PARATYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392120D1715791FB8B3FBFA60" box="[357,475,1441,1465]" name="Colombia" pageId="23" pageNumber="22">Colombia</collectingCountry>
: Prov. Valle,
<location id="1F19603892120D1716731FB8B139FA60" LSID="urn:lsid:plazi:treatment:926F87F592100D1715831E78B132F9C0:1F19603892120D1716731FB8B139FA60" box="[623,793,1441,1465]" country="Colombia" name="Rio Anchicaya" pageId="23" pageNumber="22">Rio Anchicaya</location>
,
<quantity id="DD3E9B0692120D17173F1FB8B14AFA60" box="[803,874,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">
<elevation id="91EBD1D092120D17173F1FB8B14AFA60" box="[803,874,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="6E78102392120D1717681FB8B622FA60" box="[884,1026,1441,1465]" pageId="23" pageNumber="22" value="1977-02-10">
<collectingDate id="7E3CE9CB92120D1717681FB8B622FA60" box="[884,1026,1441,1465]" pageId="23" pageNumber="22" value="1977-02-10">10 Feb. 1977</collectingDate>
</date>
,
<collectorName id="B733533592120D1710171FBBB6B0FA60" box="[1035,1168,1441,1465]" pageId="23" pageNumber="22">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592120D1714A71FD8B347FA00" box="[187,359,1473,1497]" pageId="23" pageNumber="22">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692120D1715691FD8B3E5FA00" box="[373,453,1473,1497]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="23" pageNumber="22" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D1714CC1FF8B132F9C0" pageId="23" pageNumber="22" type="etymology">
<paragraph id="1A7936E392120D1714CC1FF8B132F9C0" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">ETYMOLOGY: The specific epithet honors my goddaughter, Ms. Rosella Amparo Be- tancourt, a brilliant young poet and budding scholar.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,691 @@
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<mods:title id="9CD54892D1E0D299C0F670B0098143CE">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:affiliation id="C8ADBE7638B86561B47BD3417DB48783">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<treatment id="926F87F592120D1E159B1C58B1D1FB00" LSID="urn:lsid:zoobank.org:act:A8381269-236C-42AB-897A-5B2B10E21A04" httpUri="http://treatment.plazi.org/id/926F87F592120D1E159B1C58B1D1FB00" lastPageId="30" lastPageNumber="29" pageId="23" pageNumber="22">
<subSubSection id="52DC656892120D17159B1C58B617F9A0" pageId="23" pageNumber="22" type="nomenclature">
<paragraph id="1A7936E392120D17159B1C58B189F980" blockId="23.[391,937,1601,1625]" box="[391,937,1601,1625]" pageId="23" pageNumber="22">
<heading id="4131818F92120D17159B1C58B189F980" box="[391,937,1601,1625]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="22" reason="2">
<taxonomicName id="DDC64D6092120D17159B1C58B132F980" authority="Engel" authorityName="Engel" authorityYear="2022" box="[391,786,1601,1625]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF192120D17159B1C58B0E5F980" bold="true" box="[391,709,1601,1625]" italics="true" pageId="23" pageNumber="22">Scaptotrigona magdalenae</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92120D1717001C5BB189F980" box="[796,937,1601,1625]" pageId="23" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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ZooBank:
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D17163B1C98B129F940" box="[551,777,1665,1689]" pageId="23" pageNumber="22" type="description">
<paragraph id="1A7936E392120D17163B1C98B129F940" blockId="23.[249,1079,1635,1689]" box="[551,777,1665,1689]" pageId="23" pageNumber="22">
(
<figureCitation id="82FD2A6692120D1716331C98B093F940" box="[559,691,1665,1689]" captionStart-0="Figures 2931" captionStart-1="Figures 3233" captionStart-2="Figures 3436" captionStart-3="Figures 3739" captionStart-4="Figures 4043" captionStartId-0="24.[160,241,1326,1348]" captionStartId-1="25.[160,241,918,940]" captionStartId-2="26.[160,241,1331,1353]" captionStartId-3="28.[160,241,1457,1479]" captionStartId-4="29.[160,241,723,745]" captionTargetBox-0="[189,1147,244,1308]" captionTargetBox-1="[160,1168,224,911]" captionTargetBox-2="[179,1156,249,1317]" captionTargetBox-3="[193,1134,245,1431]" captionTargetBox-4="[188,1133,239,692]" captionTargetId-0="figure-184@24.[160,1168,224,1320]" captionTargetId-1="figure-296@25.[160,1168,224,911]" captionTargetId-2="figure-148@26.[160,1168,223,1325]" captionTargetId-3="figure-105@28.[160,1168,224,1451]" captionTargetId-4="figure-376@29.[160,1168,224,717]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="28" captionTargetPageId-4="29" captionText-0="Figures 2931. Worker of Scaptotrigona magdalenae, new species, light morph. 29. Lateral habi- tus. 30. Dorsal habitus. 31. Facial view." captionText-1="Figures 3233. Worker of Scaptotrigona magdalenae, new species, light morph. 32. Dorsal view of vertex and mesosoma. 33. Prolateral surface of metatibia, metatarsus, and metapretarsus." captionText-2="Figures 3436. Worker of Scaptotrigona magdalenae, new species, dark morph. 34. Lateral habi- tus. 35. Dorsal habitus. 36. Facial view." captionText-3="Figures 3739. Drone of Scaptotrigona magdalenae, new species. 37. Lateral habitus. 38. Dorsal habitus. 39. Facial view." captionText-4="Figures 4043. Male terminalia of Scaptotrigona magdalenae, new species. 40. Sternum VI. 41. Sternum VII. 42. Sternum VIII. 43. Genital capsule, left is dorsal view, right is ventral view." pageId="23" pageNumber="22">Figs. 2943</figureCitation>
, cover)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D1914CC1CD8B1E0FBB3" lastPageId="25" lastPageNumber="24" pageId="23" pageNumber="22" type="diagnosis">
<paragraph id="1A7936E392120D1814CC1CD8B111F9BA" blockId="23.[160,1168,1729,1785]" lastBlockId="24.[160,1169,1419,1795]" lastPageId="24" lastPageNumber="23" pageId="23" pageNumber="22">
DIAGNOSIS: This is a smaller species of
<taxonomicName id="DDC64D6092120D1716841CDBB10EF900" box="[664,814,1730,1753]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF192120D1716841CDBB10EF900" box="[664,814,1730,1753]" italics="true" pageId="23" pageNumber="22">Scaptotrigona</emphasis>
</taxonomicName>
with mesoscutal striping cur- rently known only from the
<collectingRegion id="D802F80192120D1715EB1CF8B05BF920" box="[503,635,1761,1785]" country="Colombia" name="Magdalena" pageId="23" pageNumber="22">Magdalena</collectingRegion>
Valley, from
<collectingRegion id="D802F80192120D1717031CF8B152F920" box="[799,882,1761,1785]" country="Colombia" name="Tolima" pageId="23" pageNumber="22">Tolima</collectingRegion>
to
<collectingRegion id="D802F80192120D1717871CF8B66EF920" box="[923,1102,1761,1785]" country="Colombia" name="Cundinamarca" pageId="23" pageNumber="22">Cundinamarca</collectingRegion>
. It is somewhat similar to the neighboring
<taxonomicName id="DDC64D60921D0D1816421F92B0D0FA7B" authorityName="Engel" authorityYear="2022" box="[606,752,1419,1442]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="tatacoensis">
<emphasis id="28B2EAF1921D0D1816421F92B0D0FA7B" box="[606,752,1419,1442]" italics="true" pageId="24" pageNumber="23">S. tatacoensis</emphasis>
</taxonomicName>
, but lacks the distinctive color pat- terns of that species (
<emphasis id="28B2EAF1921D0D1815841FB2B3DFFA1A" box="[408,511,1451,1475]" italics="true" pageId="24" pageNumber="23">vide infra</emphasis>
). While most
<taxonomicName id="DDC64D60921D0D1816B61FB2B160FA1B" box="[682,832,1451,1474]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF1921D0D1816B61FB2B160FA1B" box="[682,832,1451,1474]" italics="true" pageId="24" pageNumber="23">Scaptotrigona</emphasis>
</taxonomicName>
have a faintly darker area at the apex of the marginal cell, usually merely a darkening of the background color of the wing membrane, this species has a more distinct infumate spot in this position, and appearing infused with black relative to the otherwise parchment color of the mem- brane (
<figureCitation id="82FD2A66921D0D1814EE1C32B36DF99A" box="[242,333,1579,1603]" captionStart="Figures 2931" captionStartId="24.[160,241,1326,1348]" captionTargetBox="[189,1147,244,1308]" captionTargetId="figure-184@24.[160,1168,224,1320]" captionTargetPageId="24" captionText="Figures 2931. Worker of Scaptotrigona magdalenae, new species, light morph. 29. Lateral habi- tus. 30. Dorsal habitus. 31. Facial view." pageId="24" pageNumber="23">Figs. 29</figureCitation>
,
<figureCitation id="82FD2A66921D0D1815441C32B356F99A" box="[344,374,1579,1603]" captionStart="Figures 3436" captionStartId="26.[160,241,1331,1353]" captionTargetBox="[179,1156,249,1317]" captionTargetId="figure-148@26.[160,1168,223,1325]" captionTargetPageId="26" captionText="Figures 3436. Worker of Scaptotrigona magdalenae, new species, dark morph. 34. Lateral habi- tus. 35. Dorsal habitus. 36. Facial view." pageId="24" pageNumber="23">34</figureCitation>
). This is not a feature unique to the genus, but is distinctive among the subset of species in subgroup A of the
<taxonomicName id="DDC64D60921D0D18168D1C52B0FCF9BB" box="[657,732,1611,1634]" class="Insecta" family="Apidae" genus="Nannotrigona" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="postica">
<emphasis id="28B2EAF1921D0D18168D1C52B0FCF9BB" box="[657,732,1611,1634]" italics="true" pageId="24" pageNumber="23">postica</emphasis>
</taxonomicName>
group.
</paragraph>
<caption id="4EB9666B921D0D1814BC1F37B067FAB8" pageId="24" pageNumber="23" startId="24.[160,241,1326,1348]" targetBox="[189,1147,244,1308]" targetPageId="24" targetType="figure">
<paragraph id="1A7936E3921D0D1814BC1F37B067FAB8" blockId="24.[160,1168,1326,1377]" pageId="24" pageNumber="23">
<emphasis id="28B2EAF1921D0D1814BC1F37B31AFA9A" bold="true" box="[160,314,1326,1348]" pageId="24" pageNumber="23">Figures 2931.</emphasis>
Worker of
<taxonomicName id="DDC64D60921D0D1815A61F36B099FA9A" authorityName="Engel" authorityYear="2022" box="[442,697,1326,1348]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF1921D0D1815A61F36B099FA9A" box="[442,697,1326,1348]" italics="true" pageId="24" pageNumber="23">Scaptotrigona magdalenae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A921D0D1816DA1F36B168FA9D" box="[710,840,1326,1348]" pageId="24" pageNumber="23" rank="species">new species</taxonomicNameLabel>
, light morph.
<emphasis id="28B2EAF1921D0D1817F91F37B623FA9A" bold="true" box="[997,1027,1326,1347]" pageId="24" pageNumber="23">29.</emphasis>
Lateral habi- tus.
<emphasis id="28B2EAF1921D0D1814CF1F52B2D0FAB9" bold="true" box="[211,240,1355,1376]" pageId="24" pageNumber="23">30.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF1921D0D1815B81F52B3E2FAB9" bold="true" box="[420,450,1355,1376]" pageId="24" pageNumber="23">31.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="1A7936E3921D0D1914CC1C72B1E0FBB3" blockId="24.[160,1169,1419,1795]" lastBlockId="25.[160,1169,1010,1771]" lastPageId="25" lastPageNumber="24" pageId="24" pageNumber="23">
This species occurs in two distinct color morphs. Two years ago, I had originally considered each to be a distinct species as they originally seemed to be biogeographi- cally distinct, albeit quite close to one another in the
<collectingRegion id="D802F801921D0D18170A1CB2B1BAF91A" box="[790,922,1707,1731]" country="Colombia" name="Magdalena" pageId="24" pageNumber="23">Magdalena</collectingRegion>
River Valley. Given the small distances involved, that both were found together in a single collecting event at one locality, and that two individuals of the dark morph showed some progression toward the lighter morph, I ultimately concluded that they represent extremes of a range of variation within a single taxon. Thankfully, observations made last year by Victor H. Gonzalez clarified the mater as he was able to confirm that these are one cohesive species, as he collected both color forms from a single nest.
</paragraph>
</subSubSection>
<caption id="4EB9666B921C0D1914BC198FB651FC11" pageId="25" pageNumber="24" startId="25.[160,241,918,940]" targetBox="[160,1168,224,911]" targetPageId="25" targetType="figure">
<paragraph id="1A7936E3921C0D1914BC198FB651FC11" blockId="25.[160,1168,918,969]" pageId="25" pageNumber="24">
<emphasis id="28B2EAF1921C0D1914BC198FB31AFC72" bold="true" box="[160,314,918,940]" pageId="25" pageNumber="24">Figures 3233.</emphasis>
Worker of
<taxonomicName id="DDC64D60921C0D1915A0198FB09BFC72" authorityName="Engel" authorityYear="2022" box="[444,699,918,939]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF1921C0D1915A0198FB09BFC72" box="[444,699,918,939]" italics="true" pageId="25" pageNumber="24">Scaptotrigona magdalenae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A921C0D1916D5198FB16BFC75" box="[713,843,918,940]" pageId="25" pageNumber="24" rank="species">new species</taxonomicNameLabel>
, light morph.
<emphasis id="28B2EAF1921C0D1917F6198FB627FC72" bold="true" box="[1002,1031,918,939]" pageId="25" pageNumber="24">32.</emphasis>
Dorsal view of vertex and mesosoma.
<emphasis id="28B2EAF1921C0D1915A819AAB3F2FC11" bold="true" box="[436,466,947,968]" pageId="25" pageNumber="24">33.</emphasis>
Prolateral surface of metatibia, metatarsus, and metapretarsus.
</paragraph>
</caption>
<subSubSection id="52DC6568921C0D1D14CC1E6AB670F9F1" lastPageId="29" lastPageNumber="28" pageId="25" pageNumber="24" type="description">
<paragraph id="1A7936E3921C0D1914CC1E6AB00EF9D3" blockId="25.[160,1169,1010,1771]" pageId="25" pageNumber="24">
DESCRIPTION: As described for
<taxonomicName id="DDC64D60921C0D1916331E6AB05DFB53" authorityName="Engel" authorityYear="2022" box="[559,637,1138,1162]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1921C0D1916331E6AB05DFB53" box="[559,637,1138,1162]" italics="true" pageId="25" pageNumber="24">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF1921C0D1916971E6BB0D9FB53" box="[651,761,1138,1162]" italics="true" pageId="25" pageNumber="24">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite) 5.06.0 mm. Head wider than long, width
<quantity id="DD3E9B06921C0D1917071EAAB1E1FB13" box="[795,961,1203,1227]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.42" metricValueMin="2.33" pageId="25" pageNumber="24" unit="mm" value="2.375" valueMax="2.42" valueMin="2.33">2.332.42 mm</quantity>
, length 1.912.00 mm; compound eye length
<quantity id="DD3E9B06921C0D1915F11ECAB0B1FB33" box="[493,657,1235,1259]" metricMagnitude="-3" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.39" metricValueMin="1.33" pageId="25" pageNumber="24" unit="mm" value="1.3599999999999999" valueMax="1.39" valueMin="1.33">1.331.39 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06921C0D1917FB1ECBB6ABFB33" box="[999,1163,1234,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" metricValueMax="1.55" metricValueMin="1.53" pageId="25" pageNumber="24" unit="mm" value="1.54" valueMax="1.55" valueMin="1.53">1.531.55 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06921C0D1915FF1EEBB0A8FAD3" box="[483,648,1266,1290]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" metricValueMax="1.52" metricValueMin="1.42" pageId="25" pageNumber="24" unit="mm" value="1.47" valueMax="1.52" valueMin="1.42">1.421.52 mm</quantity>
. Scape length
<quantity id="DD3E9B06921C0D1917211EEAB1C1FAD3" box="[829,993,1267,1291]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="25" pageNumber="24" unit="mm" value="0.895" valueMax="0.91" valueMin="0.88">0.880.91 mm</quantity>
, about as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B06921C0D1916A21F0BB144FAF3" box="[702,868,1298,1322]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="25" pageNumber="24" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.850.88 mm</quantity>
. Clypeus approximately 1.651.8× as wide as long, length
<quantity id="DD3E9B06921C0D1916301F2BB0F0FA93" box="[556,720,1330,1354]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.7" metricValueMin="6.4" pageId="25" pageNumber="24" unit="mm" value="0.655" valueMax="0.67" valueMin="0.64">0.640.67 mm</quantity>
, width
<quantity id="DD3E9B06921C0D1917361F2BB1EFFA93" box="[810,975,1330,1354]" metricMagnitude="-3" metricUnit="m" metricValue="1.135" metricValueMax="1.21" metricValueMin="1.06" pageId="25" pageNumber="24" unit="mm" value="1.135" valueMax="1.21" valueMin="1.06">1.061.21 mm</quantity>
. Malar area ap- proximately 1.8× flagellar diameter or 0.800.82× basal mandibular width. Preoccipi- tal ridge carinate, lamellate dorsally, with bordering medial and lateral indentations comparatively deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle, slightly projecting upward as a short lamellate tooth, upper margin with preoccipital lamella extending into concavity about half length, sometimes less.
</paragraph>
<paragraph id="1A7936E3921C0D1B14CC1C0BB34FFEE3" blockId="25.[160,1169,1010,1771]" lastBlockId="27.[160,1168,225,1786]" lastPageId="27" lastPageNumber="26" pageId="25" pageNumber="24">
<emphasis id="28B2EAF1921C0D1914CC1C0BB353F9F2" bold="true" box="[208,371,1554,1579]" pageId="25" pageNumber="24">Light morph.</emphasis>
Integument of labiomaxillary complex orange except prementum brown; labrum orange; mandible orange to reddish orange apically; clypeus orange, epistomal sulcus well delineated by orange brown to black; supraclypeal area orange; malar space orange; face below and above tangent of antennal toruli orange except reddish brown to black around ocelli and extending slightly onto upper frons, such marking not extending to orbits and sometimes medially divided by thin strip of or- ange from frons to median ocellus; scape orange, sometimes with brown dorsoapi- cally; pedicel and flagellum typically dark brown above, with narrow underside of flagellum orange to light brown; vertex and posterior of head orange except for mark- ing around ocelli; gena and postgena orange, hypostomal borders yellow. Mesosoma largely orange except mesoscutum black with orange along lateral borders and red- dish brown to reddish orange strips from notauli to reddish orange posterior border, and mesepisternum with reddish brown to dark brown area anteroventrally; tegula wholly orange and semi-translucent; legs wholly orange except lighter on corbicula and tarsi. Wing membranes hyaline clear posteriorly to lightly parchment colored anteriorly, with lightly infumate area at marginal cell apex; veins orange. Metasoma generally yellow orange to orange, with tergal discs largely yellow except brown to dark brown laterally on terga I and II, and at extreme base of tergum II; terga
<collectionCode id="7CD7AE26921F0D1A10481CC9B64FF93E" box="[1108,1135,1744,1767]" pageId="26" pageNumber="25">III</collectionCode>
<collectionCode id="7CD7AE26921F0D1A10611CC9B6B0F93E" box="[1149,1168,1744,1767]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="25" type="Museum">V</collectionCode>
with brown to dark brown at extreme bases, often obscured by overhanging tergal margins; tergum
<collectionCode id="7CD7AE26921E0D1B15721B1BB3AAFEC0" box="[366,394,258,281]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="26">VI</collectionCode>
brown basally and light brown to yellow apically; sterna yellow orange to orange.
</paragraph>
<caption id="4EB9666B921F0D1A14BC1F2AB066FABF" pageId="26" pageNumber="25" startId="26.[160,241,1331,1353]" targetBox="[179,1156,249,1317]" targetPageId="26" targetType="figure">
<paragraph id="1A7936E3921F0D1A14BC1F2AB066FABF" blockId="26.[160,1168,1331,1382]" pageId="26" pageNumber="25">
<emphasis id="28B2EAF1921F0D1A14BC1F2AB31AFA91" bold="true" box="[160,314,1331,1353]" pageId="26" pageNumber="25">Figures 3436.</emphasis>
Worker of
<taxonomicName id="DDC64D60921F0D1A15A61F2DB098FA91" authorityName="Engel" authorityYear="2022" box="[442,696,1331,1353]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF1921F0D1A15A61F2DB098FA91" box="[442,696,1331,1353]" italics="true" pageId="26" pageNumber="25">Scaptotrigona magdalenae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A921F0D1A16D91F2DB167FA90" box="[709,839,1331,1353]" pageId="26" pageNumber="25" rank="species">new species</taxonomicNameLabel>
, dark morph.
<emphasis id="28B2EAF1921F0D1A17F81F2AB622FA91" bold="true" box="[996,1026,1331,1352]" pageId="26" pageNumber="25">34.</emphasis>
Lateral habi- tus.
<emphasis id="28B2EAF1921F0D1A14CE1F49B2D1FABC" bold="true" box="[210,241,1360,1381]" pageId="26" pageNumber="25">35.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF1921F0D1A15BF1F49B3E1FABC" bold="true" box="[419,449,1360,1381]" pageId="26" pageNumber="25">36.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="1A7936E3921E0D1B14CC1B58B1DCFC00" blockId="27.[160,1168,225,1786]" pageId="27" pageNumber="26">
Fine pubescence generally consisting of scatered, minute to short, appressed or decumbent yellow setae, such setae often simple but sometimes with minute branches or plumose, such plumose setae sometimes blending to yellow or pale yellow when dense around pronotal lobe, and on metepisternum and lateral surface of propodeum; sterna II
<collectionCode id="7CD7AE26921E0D1B150C1BDBB303FE00" box="[272,291,450,473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="27" pageNumber="26" type="Museum">V</collectionCode>
with scopa composed elongate, erect, simple setae with characteristically wavy apices, such scopal setae yellow. Bristles as follow: Labrum with sparse, erect, simple, yellow bristles; vertex with numerous erect, simple, yellow to sometimes light fulvous bristles; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow to fulvous bristles; mesoscutum with anterior and lateral borders with erect bristles, those on lateral border short, bristles largely yellow to fulvous, although along anterior margin some with dark fuscous bases and some entirely black bristles antero- laterally; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elongate, erect, yellow to light fulvous bristles; mesepisternum with scat- tered, yellow to fulvous bristles, along rounded anterior border with preëpisternum a line of abundant, suberect yellow to fulvous bristles extending ventrally from pronotal lobe; lateral surface of propodeum with scatered, long, erect, yellow to light fulvous bristles. Legs with abundant yellow to fulvous bristles except some black bristles on prolateral surface of protibia, abundant on prolateral surfaces of mesotibia, pro- and mesobasitarsus, and apically on metafemur; metatibia and metabasitarsus with black bristles. Metasoma with numerous, prominent, suberect to subdecumbent, light ful- vous to sometimes yellow bristles on terga
<collectionCode id="7CD7AE26921E0D1B168019DBB097FC00" box="[668,695,962,985]" pageId="27" pageNumber="26">III</collectionCode>
<collectionCode id="7CD7AE26921E0D1B16D819DBB0F7FC00" box="[708,727,962,985]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="27" pageNumber="26" type="Museum">V</collectionCode>
arising amid tomentum.
</paragraph>
<paragraph id="1A7936E3921E0D1B14CC19F8B15EFAC0" blockId="27.[160,1168,225,1786]" pageId="27" pageNumber="26">
<emphasis id="28B2EAF1921E0D1B14CC19F8B34EFC20" bold="true" box="[208,366,993,1017]" pageId="27" pageNumber="26">Dark morph.</emphasis>
As for the light morph except as follows: upper two-thirds of frons dark brown to black, except often separated from inner orbit by thin strip of orange, rarely by broader strip that then extends into ocellocular area; vertex, posterior of head, and uppermost gena dark brown to black. Mesosoma largely dark brown to black except pronotal lobe, mesoscutellum, metanotum, and hypoepimeral area or- ange. Legs largely orange except prolateral surfaces of tibiae and upper surfaces of femora, except at joints reddish brown to brown. Metasoma generally dark brown to black except anterior-facing surface of tergum I brown, dorsal-facing surface of ter- gum I with orange brown apically and terga II
<collectionCode id="7CD7AE26921E0D1B16CB1EFBB0CAFB20" box="[727,746,1250,1273]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="27" pageNumber="26" type="Museum">V</collectionCode>
with yellow apical marginal zones; tergum
<collectionCode id="7CD7AE26921E0D1B14E11F1BB339FAC0" box="[253,281,1282,1305]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="26">VI</collectionCode>
brown to dark brown; sterna brown to dark brown.
</paragraph>
<paragraph id="1A7936E3921E0D1B14CC1F3BB30FFAE0" blockId="27.[160,1168,225,1786]" box="[208,303,1314,1340]" pageId="27" pageNumber="26">
<specimenCount id="0CC0FD6A921E0D1B14CC1F3BB2C4FAE5" box="[208,228,1314,1340]" count="1" pageId="27" pageNumber="26" type="female"></specimenCount>
:
<emphasis id="28B2EAF1921E0D1B14ED1F3BB308FAE0" box="[241,296,1314,1337]" italics="true" pageId="27" pageNumber="26">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E3921E0D1B14CC1F5BB09AF9A0" blockId="27.[160,1168,225,1786]" pageId="27" pageNumber="26">
<specimenCount id="0CC0FD6A921E0D1B14CC1F5BB2C4FA85" box="[208,228,1346,1372]" count="1" pageId="27" pageNumber="26" type="male"></specimenCount>
: As described for worker except as follows: Total body length approximately 5.56.0 mm, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B06921E0D1B17421F78B1C9FAA0" box="[862,1001,1377,1401]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="5.8" metricValueMin="5.6" pageId="27" pageNumber="26" unit="mm" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.65.8 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B06921E0D1B15611F98B005FA40" box="[381,549,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" metricValueMax="2.3" metricValueMin="2.24" pageId="27" pageNumber="26" unit="mm" value="2.27" valueMax="2.3" valueMin="2.24">2.242.30 mm</quantity>
, length
<quantity id="DD3E9B06921E0D1B16921F98B117FA40" box="[654,823,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="1.865" metricValueMax="1.88" metricValueMin="1.85" pageId="27" pageNumber="26" unit="mm" value="1.865" valueMax="1.88" valueMin="1.85">1.851.88 mm</quantity>
; compound eye length
<quantity id="DD3E9B06921E0D1B107D1F98B2F1FA60" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="27" pageNumber="26" unit="mm" value="1.48">1.48 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06921E0D1B16361FB8B0F0FA60" box="[554,720,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" metricValueMax="1.36" metricValueMin="1.3" pageId="27" pageNumber="26" unit="mm" value="1.33" valueMax="1.36" valueMin="1.3">1.301.36 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06921E0D1B10391FB8B2F2FA00" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.06" metricValueMin="0.97" pageId="27" pageNumber="26" unit="mm" value="1.015" valueMax="1.06" valueMin="0.97">0.971.06 mm</quantity>
. Scape length
<quantity id="DD3E9B06921E0D1B159B1FD8B00BFA00" box="[391,555,1473,1497]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" metricValueMax="6.7" metricValueMin="6.1" pageId="27" pageNumber="26" unit="mm" value="0.64" valueMax="0.67" valueMin="0.61">0.610.67 mm</quantity>
, much shorter than torulocellar distance, torulocel- lar distance
<quantity id="DD3E9B06921E0D1B15331FF8B3F3FA20" box="[303,467,1505,1529]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="7.9" metricValueMin="7.6" pageId="27" pageNumber="26" unit="mm" value="0.775" valueMax="0.79" valueMin="0.76">0.760.79 mm</quantity>
. Clypeus approximately 1.391.44× as wide as long, length
<quantity id="DD3E9B06921E0D1B14BC1C18B365F9C0" box="[160,325,1537,1561]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="7.0" metricValueMin="6.7" pageId="27" pageNumber="26" unit="mm" value="0.685" valueMax="0.7" valueMin="0.67">0.670.70 mm</quantity>
, width
<quantity id="DD3E9B06921E0D1B15BE1C18B02CF9C0" box="[418,524,1537,1561]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="27" pageNumber="26" unit="mm" value="0.97">0.97 mm</quantity>
. Malar area approximately 0.8× flagellar diameter or 0.4× basal mandibular width. Gena narrower than compound eye. Preoccipital ridge weakly carinate dorsally, sharply angled at best laterally, without concavity or border- ing indentations. Terminalia in figures 4043.
</paragraph>
<paragraph id="1A7936E3921E0D1D14CC1C9BB13DFAD1" blockId="27.[160,1168,225,1786]" lastBlockId="29.[160,1175,816,1768]" lastPageId="29" lastPageNumber="28" pageId="27" pageNumber="26">
Integument of labiomaxillary complex yellow to orange except prementum brown; labrum yellow to orange; mandible yellow to orange with reddish orange apically; clypeus yellow to orange, epistomal sulcus well delineated by dark brown to black; supraclypeal area yellow to orange; malar space yellow to orange; face below tan- gent of antennal toruli yellow to orange; frons largely brown to black except in broad yellow to orange triangular areas continuing from lower face on lower frons, such coloration extending as thin line along inner orbit to summit, sometimes with a small spot of yellow to orange anterior to median ocellus; vertex and posterior of head dark brown to black, uppermost margin of gena brown to black, remainder yellow to or- ange; postgena yellow to orange; scape orange; pedicel and flagellum typically brown to dark brown, flagellum lighter beneath. Mesosoma largely yellow to orange except mesoscutum black, sometimes with weak orange to reddish orange along lateral bor- ders, mesepisternum with dark brown to black ventrally, sometimes more restricted posteroventrally or even with mesepisternum largely black with orange to reddish orange posterodorsally; metepisternum sometimes dark brown to black; metanotum sometimes dark brown; mesoscutellum sometimes more reddish orange to reddish brown and lighter medioapically; propodeum dark brown to black; tegula wholly orange and semi-translucent; legs entirely yellow orange to orange except in darker forms with spots of dark brown to black on protibia, prolateral surface of meso- and metabasitarsi entirely dark brown to black, and metatibia with line of dark brown to black on superior margin of prolateral surface in apical two-thirds. Metasoma gener- ally yellow orange to black, either with terga dark brown to black with tergum I and tergum II to varying degrees broadly orange, tergum VII apically yellow, and sterna largely yellow brown, or in lighter individuals terga largely yellow to orange, with areas of brown to dark brown laterally on tergum I, basally and laterally on terga II
<collectionCode id="7CD7AE2692180D1D10641EC9B6ACFB3E" box="[1144,1164,1232,1255]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="28" type="Museum">V</collectionCode>
, with tergum
<collectionCode id="7CD7AE2692180D1D15261EE9B376FADE" box="[314,342,1264,1287]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="29" pageNumber="28">VI</collectionCode>
largely brown and tergum VII yellow.
</paragraph>
<caption id="4EB9666B92190D1C14BC1FA8B383FA3D" pageId="28" pageNumber="27" startId="28.[160,241,1457,1479]" targetBox="[193,1134,245,1431]" targetPageId="28" targetType="figure">
<paragraph id="1A7936E392190D1C14BC1FA8B383FA3D" blockId="28.[160,1168,1457,1508]" pageId="28" pageNumber="27">
<emphasis id="28B2EAF192190D1C14BC1FA8B31AFA1F" bold="true" box="[160,314,1457,1479]" pageId="28" pageNumber="27">Figures 3739.</emphasis>
Drone of
<taxonomicName id="DDC64D6092190D1C15B11FABB08CFA1F" authorityName="Engel" authorityYear="2022" box="[429,684,1457,1479]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF192190D1C15B11FABB08CFA1F" box="[429,684,1457,1479]" italics="true" pageId="28" pageNumber="27">Scaptotrigona magdalenae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92190D1C16A51FABB11BFA1E" box="[697,827,1457,1479]" pageId="28" pageNumber="27" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192190D1C17501FA8B14AFA1F" bold="true" box="[844,874,1457,1478]" pageId="28" pageNumber="27">37.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192190D1C10391FA8B662FA1F" bold="true" box="[1061,1090,1457,1478]" pageId="28" pageNumber="27">38.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF192190D1C151D1FD7B33EFA3A" bold="true" box="[257,286,1486,1507]" pageId="28" pageNumber="27">39.</emphasis>
Facial view.
</paragraph>
</caption>
<caption id="4EB9666B92180D1D14BC18CAB64DFCDF" pageId="29" pageNumber="28" startId="29.[160,241,723,745]" targetBox="[188,1133,239,692]" targetPageId="29" targetType="figure">
<paragraph id="1A7936E392180D1D14BC18CAB64DFCDF" blockId="29.[160,1168,723,774]" pageId="29" pageNumber="28">
<emphasis id="28B2EAF192180D1D14BC18CAB31BFD31" bold="true" box="[160,315,723,745]" pageId="29" pageNumber="28">Figures 4043.</emphasis>
Male terminalia of
<taxonomicName id="DDC64D6092180D1D160418CDB138FD31" authorityName="Engel" authorityYear="2022" box="[536,792,723,745]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="28B2EAF192180D1D160418CDB138FD31" box="[536,792,723,745]" italics="true" pageId="29" pageNumber="28">Scaptotrigona magdalenae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92180D1D173A18CDB189FD30" box="[806,937,723,745]" pageId="29" pageNumber="28" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192180D1D17A718CAB1F9FD31" bold="true" box="[955,985,723,744]" pageId="29" pageNumber="28">40.</emphasis>
Sternum VI.
<emphasis id="28B2EAF192180D1D106E18CAB6B0FD31" bold="true" box="[1138,1168,723,744]" pageId="29" pageNumber="28">41.</emphasis>
Sternum VII.
<emphasis id="28B2EAF192180D1D152918E9B372FCDC" bold="true" box="[309,338,752,773]" pageId="29" pageNumber="28">42.</emphasis>
Sternum VIII.
<emphasis id="28B2EAF192180D1D15E918E9B034FCDC" bold="true" box="[501,532,752,773]" pageId="29" pageNumber="28">43.</emphasis>
Genital capsule, left is dorsal view, right is ventral view.
</paragraph>
</caption>
<paragraph id="1A7936E392180D1D14CC1F09B670F9F1" blockId="29.[160,1175,816,1768]" pageId="29" pageNumber="28">
Appressed, simple, short, yellow setae of mesoscutum more abundant and more transversely directed; sterna without scopal setae as in worker but with dense, sub- decumbent to decumbent, mesial-posteriorly directed, simple, elongate, yellow setae. Bristles as follow: bristles of drone generally more elongate than those of worker;
<emphasis id="28B2EAF192180D1D10781F68B6A3FA51" box="[1124,1155,1393,1416]" italics="true" pageId="29" pageNumber="28">e.g</emphasis>
., bristles of vertex much more elongate than in worker, and gently arched. Legs with abundant yellow to fulvous bristles except numerous black bristles on prolateral sur- face of pro- and mesotibiae and pro- and mesobasitarsi; such black bristles apically on metafemur; black bristles dominant and scatered over prolateral surface of metatibia; a few on prolateral surface and dominant on superior margin of metabasitarsus.
</paragraph>
</subSubSection>
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<paragraph id="1A7936E392180D1D14CC1C29B1F8F9B1" blockId="29.[160,1175,816,1768]" pageId="29" pageNumber="28">
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: ⚲,
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:
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cruce a
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21-XII-
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[light morph] (
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<materialsCitation id="AAAE3CBE92180D1D14CC1C69B05AF97E" collectingDate="2011-12-21" collectionCode="V" collectorName="Boqueron &amp; El &amp; Pandi" country="Colombia" latitude="4.2600164" location="Cundinamarca" longLatPrecision="1" longitude="-74.54543" pageId="29" pageNumber="28" specimenCount="32" specimenCount-male="32" stateProvince="Cundinamarca" typeStatus="paratype">
<typeStatus id="C57D884192180D1D14CC1C69B36EF951" box="[208,334,1648,1672]" pageId="29" pageNumber="28" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="0CC0FD6A92180D1D15451C69B3BCF953" box="[345,412,1648,1674]" count="32" pageId="29" pageNumber="28" type="male">32♂♂</specimenCount>
,
<collectingCountry id="62D1767392180D1D15B71C69B001F951" box="[427,545,1648,1672]" name="Colombia" pageId="29" pageNumber="28">Colombia</collectingCountry>
:
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,
<collectorName id="B733533592180D1D16F01C69B142F95E" box="[748,866,1648,1671]" pageId="29" pageNumber="28">Boqueron</collectorName>
,
<collectorName id="B733533592180D1D17731C69B1A7F951" box="[879,903,1648,1672]" pageId="29" pageNumber="28">El</collectorName>
cruce a
<collectorName id="B733533592180D1D17F11C69B613F951" box="[1005,1075,1648,1672]" pageId="29" pageNumber="28">Pandi</collectorName>
,
<date id="6E78102392180D1D105C1C69B2F8F971" pageId="29" pageNumber="28" value="2011-12-21">
<collectingDate id="7E3CE9CB92180D1D105C1C69B2F8F971" pageId="29" pageNumber="28" value="2011-12-21">
21-XII-
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<collectorName id="B733533592180D1D16621C89B12FF971" box="[638,783,1680,1704]" pageId="29" pageNumber="28">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE2692180D1D17021C89B14EF971" box="[798,878,1680,1704]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="28" type="Museum">SEMC</collectionCode>
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<emphasis id="28B2EAF192180D1D17941C89B606F971" bold="true" box="[904,1062,1680,1704]" pageId="29" pageNumber="28">
<collectingCounty id="F3184E6F92180D1D17941C89B1E6F971" box="[904,966,1680,1704]" pageId="29" pageNumber="28">Dark</collectingCounty>
morph.
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4⚲⚲, Co- lombia:
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,
<collectingMunicipality id="FA1DAC9992180D1D15DE1CA9B018F91E" box="[450,568,1712,1735]" pageId="29" pageNumber="28">Boqueron</collectingMunicipality>
,
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cruce a
<collectorName id="B733533592180D1D16DB1CA9B12DF911" box="[711,781,1712,1736]" pageId="29" pageNumber="28">Pandi</collectorName>
,
<date id="6E78102392180D1D17071CA9B183F911" box="[795,931,1712,1736]" pageId="29" pageNumber="28" value="2011-12-21">
<collectingDate id="7E3CE9CB92180D1D17071CA9B183F911" box="[795,931,1712,1736]" pageId="29" pageNumber="28" value="2011-12-21">
21-XII-
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,
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,
<collectionCode id="7CD7AE2692180D1D155D1CC9B378F93E" box="[321,344,1744,1767]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="28" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE92180D1E15471CC9B635FF20" collectingDate="2011-12-21" collectionCode="SEMC, V" collectorName="H. Gonzalez &amp; Pandi" country="Colombia" county="Light" lastPageId="30" lastPageNumber="29" latitude="4.2600164" location="El" longLatPrecision="1" longitude="-74.54543" municipality="Boqueron" pageId="29" pageNumber="28" specimenCount="1" stateProvince="Cundinamarca" typeStatus="paratype">
<collectorName id="B733533592180D1D15471CC9B3CFF931" box="[347,495,1744,1768]" pageId="29" pageNumber="28">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE2692180D1D161C1CC9B070F931" box="[512,592,1744,1768]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="28" type="Museum">SEMC</collectionCode>
).
<emphasis id="28B2EAF192180D1D16721CC9B131F931" bold="true" box="[622,785,1744,1768]" pageId="29" pageNumber="28">
<collectingCounty id="F3184E6F92180D1D16721CC9B08FF931" box="[622,687,1744,1768]" pageId="29" pageNumber="28">Light</collectingCounty>
morph.
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6⚲⚲,
<collectingCountry id="62D1767392180D1D174A1CC9B1ECF931" box="[854,972,1744,1768]" name="Colombia" pageId="29" pageNumber="28">Colombia</collectingCountry>
:
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,
<collectingMunicipality id="FA1DAC99921B0D1E14BC1AFBB336FF20" box="[160,278,226,249]" pageId="30" pageNumber="29">Boqueron</collectingMunicipality>
,
<location id="1F196038921B0D1E153D1AF8B319FF20" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E153D1AF8B319FF20" box="[289,313,225,249]" country="Colombia" county="Light" latitude="4.2600164" longLatPrecision="1" longitude="-74.54543" municipality="Boqueron" name="El" pageId="30" pageNumber="29" stateProvince="Cundinamarca">El</location>
cruce a
<collectorName id="B7335335921B0D1E15861AF8B3C0FF20" box="[410,480,225,249]" pageId="30" pageNumber="29">Pandi</collectorName>
,
<date id="6E781023921B0D1E15F01AF8B054FF20" box="[492,628,225,249]" pageId="30" pageNumber="29" value="2011-12-21">
<collectingDate id="7E3CE9CB921B0D1E15F01AF8B054FF20" box="[492,628,225,249]" pageId="30" pageNumber="29" value="2011-12-21">
21-XII-
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</date>
,
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,
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,
<collectionCode id="7CD7AE26921B0D1E17E21AFBB635FF20" box="[1022,1045,226,249]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E10041AFBB0E7FEE0" collectingDate="2011-12-21" collectionCode="SEMC, V" collectorName="El Guamo Vereda Chontaduro &amp; La Guaraperia" country="Colombia" elevation="337" latitude="4.0061035" location="Tolima" longLatPrecision="1" longitude="-74.9493" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Tolima" typeStatus="paratype">
H. Gonza- lez (
<collectionCode id="7CD7AE26921B0D1E14CE1B18B301FEC0" box="[210,289,257,281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 15⚲⚲,
<collectingCountry id="62D17673921B0D1E15611B18B3D3FEC0" box="[381,499,257,281]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
:
<collectingRegion id="D802F801921B0D1E15E31B18B075FEC0" box="[511,597,257,281]" country="Colombia" name="Tolima" pageId="30" pageNumber="29">Tolima</collectingRegion>
,
<collectorName id="B7335335921B0D1E167E1B18B1F0FEC0" box="[610,976,257,281]" pageId="30" pageNumber="29">El Guamo Vereda Chontaduro</collectorName>
,
<collectorName id="B7335335921B0D1E17C11B1BB6ABFEC0" box="[989,1163,257,281]" pageId="30" pageNumber="29">La Guaraperia</collectorName>
,
<quantity id="DD3E9B06921B0D1E14BC1B38B2C9FEE0" box="[160,233,289,313]" metricMagnitude="2" metricUnit="m" metricValue="3.37" pageId="30" pageNumber="29" unit="m" value="337.0">
<elevation id="91EBD1D0921B0D1E14BC1B38B2C9FEE0" box="[160,233,289,313]" metricMagnitude="2" metricUnit="m" metricValue="3.37" pageId="30" pageNumber="29" unit="m" value="337.0">337 m</elevation>
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,
<geoCoordinate id="7FF25024921B0D1E14E81B38B355FEE0" box="[244,373,289,313]" degrees="4" direction="north" minutes="0.3662" orientation="latitude" pageId="30" pageNumber="29" precision="1" value="4.0061035">4°0.3662N</geoCoordinate>
,
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,
<date id="6E781023921B0D1E16011B38B086FEE0" box="[541,678,289,313]" pageId="30" pageNumber="29" value="2011-12-21">
<collectingDate id="7E3CE9CB921B0D1E16011B38B086FEE0" box="[541,678,289,313]" pageId="30" pageNumber="29" value="2011-12-21">21-XII-2011</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E16AC1B3BB0E7FEE0" box="[688,711,290,313]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E16D71B3BB1E2FEE0" box="[715,962,289,313]" collectionCode="SEMC" collectorName="Gonzalez" country="Colombia" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Tolima" typeStatus="paratype">
H.
<collectorName id="B7335335921B0D1E16F01B38B17BFEE0" box="[748,859,289,313]" pageId="30" pageNumber="29">Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E17761B38B19AFEE0" box="[874,954,289,313]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D1E14CC1B58B1C6FD60" pageId="30" pageNumber="29" type="description">
<paragraph id="1A7936E3921B0D1E14CC1B58B1C6FD60" blockId="30.[160,1175,225,1242]" pageId="30" pageNumber="29">
<materialsCitation id="AAAE3CBE921B0D1E14CC1B58B0BFFE40" collectingDate="2021-03-13" collectionCode="V" country="Colombia" elevation="1581" location="San Antonio de Tequendama" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
ADDITIONAL MATERIAL (each series from a different nest): 6⚲⚲ [dark], 3⚲⚲ [light] [from same nest],
<collectingCountry id="62D17673921B0D1E156D1B78B3C7FEA0" box="[369,487,353,377]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
,
<collectingRegion id="D802F801921B0D1E15EC1B78B082FEA0" box="[496,674,353,377]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
,
<location id="1F196038921B0D1E16B11B78B622FEA0" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E16B11B78B622FEA0" box="[685,1026,353,377]" country="Colombia" name="San Antonio de Tequendama" pageId="30" pageNumber="29" stateProvince="Cundinamarca">San Antonio de Tequendama</location>
, 4˚38.107N, 74˚21.331W,
<quantity id="DD3E9B06921B0D1E15581B98B383FE40" box="[324,419,385,409]" metricMagnitude="3" metricUnit="m" metricValue="1.581" pageId="30" pageNumber="29" unit="m" value="1581.0">
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,
<date id="6E781023921B0D1E15AB1B98B054FE40" box="[439,628,385,409]" pageId="30" pageNumber="29" value="2021-03-13">
<collectingDate id="7E3CE9CB921B0D1E15AB1B98B054FE40" box="[439,628,385,409]" pageId="30" pageNumber="29" value="2021-03-13">13 March 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E16941B9BB0BFFE40" box="[648,671,386,409]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E16BE1B9BB062FE00" collectingDate="2021-02-21" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
<collectorName id="B7335335921B0D1E16BE1B9BB11BFE40" box="[674,827,385,409]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
[examined three managed hives] (
<collectionCode id="7CD7AE26921B0D1E14E41BB8B367FE60" box="[248,327,417,441]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 4⚲⚲ [light], 1⚲ [dark],
<collectingCountry id="62D17673921B0D1E16791BB8B0FBFE60" box="[613,731,417,441]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
,
<collectingRegion id="D802F801921B0D1E16F41BB8B1BAFE60" box="[744,922,417,441]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
,
<location id="1F196038921B0D1E17B41BB8B1DFFE60" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E17B41BB8B1DFFE60" box="[936,1023,417,441]" country="Colombia" name="Beltran" pageId="30" pageNumber="29" stateProvince="Cundinamarca">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E15211BD8B3A7FE00" box="[317,391,449,473]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E15211BD8B3A7FE00" box="[317,391,449,473]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
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,
<date id="6E781023921B0D1E158F1BD8B03FFE00" box="[403,543,449,473]" pageId="30" pageNumber="29" value="2021-02-21">
<collectingDate id="7E3CE9CB921B0D1E158F1BD8B03FFE00" box="[403,543,449,473]" pageId="30" pageNumber="29" value="2021-02-21">21 Feb 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E16371BDBB062FE00" box="[555,578,450,473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E165A1BDBB1D5FE20" collectingDate="2021-02-21" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1">
<collectorName id="B7335335921B0D1E165A1BDBB0F7FE00" box="[582,727,449,473]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E16FB1BD8B116FE00" box="[743,822,449,473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 4⚲⚲ [light],
<collectingCountry id="62D17673921B0D1E17CF1BD8B669FE00" box="[979,1097,449,473]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
, Cun- dinamarca,
<location id="1F196038921B0D1E152E1BF8B3A9FE20" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E152E1BF8B3A9FE20" box="[306,393,481,505]" country="Colombia" name="Beltran" pageId="30" pageNumber="29">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E16CD1BF8B101FE20" box="[721,801,481,505]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E16CD1BF8B101FE20" box="[721,801,481,505]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
</quantity>
,
<date id="6E781023921B0D1E172F1BF8B1ECFE20" box="[819,972,481,505]" pageId="30" pageNumber="29" value="2021-02-21">
<collectingDate id="7E3CE9CB921B0D1E172F1BF8B1ECFE20" box="[819,972,481,505]" pageId="30" pageNumber="29" value="2021-02-21">21 Feb 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E17C21BFBB1D5FE20" box="[990,1013,482,505]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E17E41BFBB35FFDE0" collectingDate="2021-02-21" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
<collectorName id="B7335335921B0D1E17E41BFBB6B0FE20" box="[1016,1168,481,505]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E14B41818B2D7FDC0" box="[168,247,513,537]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 2⚲⚲ [light],
<collectingCountry id="62D17673921B0D1E158B1818B02DFDC0" box="[407,525,513,537]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
,
<collectingRegion id="D802F801921B0D1E16061818B0ECFDC0" box="[538,716,513,537]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
,
<location id="1F196038921B0D1E16C51818B110FDC0" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E16C51818B110FDC0" box="[729,816,513,537]" country="Colombia" name="Beltran" pageId="30" pageNumber="29" stateProvince="Cundinamarca">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E10741818B29AFDE0" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E10741818B29AFDE0" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
</quantity>
,
<date id="6E781023921B0D1E14D41838B37AFDE0" box="[200,346,545,569]" pageId="30" pageNumber="29" value="2021-02-21">
<collectingDate id="7E3CE9CB921B0D1E14D41838B37AFDE0" box="[200,346,545,569]" pageId="30" pageNumber="29" value="2021-02-21">21 Feb 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E1574183BB35FFDE0" box="[360,383,546,569]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E159F183BB1B5FD80" collectingDate="2021-03-28" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1">
<collectorName id="B7335335921B0D1E159F183BB037FDE0" box="[387,535,545,569]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E16341838B057FDE0" box="[552,631,545,569]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 2⚲⚲ [light], 1⚲ [dark],
<collectingCountry id="62D17673921B0D1E17801838B632FDE0" box="[924,1042,545,569]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
, Cundina- marca,
<location id="1F196038921B0D1E14E91858B36CFD80" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E14E91858B36CFD80" box="[245,332,577,601]" country="Colombia" name="Beltran" pageId="30" pageNumber="29">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E16651858B0E1FD80" box="[633,705,577,601]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E16651858B0E1FD80" box="[633,705,577,601]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
</quantity>
,
<date id="6E781023921B0D1E16D71858B155FD80" box="[715,885,577,601]" pageId="30" pageNumber="29" value="2021-03-28">
<collectingDate id="7E3CE9CB921B0D1E16D71858B155FD80" box="[715,885,577,601]" pageId="30" pageNumber="29" value="2021-03-28">28 March 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E1762185BB1B5FD80" box="[894,917,578,601]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E1785185BB374FD40" collectingDate="2021-03-30" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
<collectorName id="B7335335921B0D1E1785185BB608FD80" box="[921,1064,577,601]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E10291858B6A4FD80" box="[1077,1156,577,601]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 4⚲⚲ [dark],
<collectingCountry id="62D17673921B0D1E152F1878B389FDA0" box="[307,425,609,633]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
,
<collectingRegion id="D802F801921B0D1E15A51878B04BFDA0" box="[441,619,609,633]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
,
<location id="1F196038921B0D1E16601878B0F3FDA0" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E16601878B0F3FDA0" box="[636,723,609,633]" country="Colombia" name="Beltran" pageId="30" pageNumber="29" stateProvince="Cundinamarca">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E100A1878B645FDA0" box="[1046,1125,609,633]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E100A1878B645FDA0" box="[1046,1125,609,633]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
</quantity>
,
<date id="6E781023921B0D1E10691878B30EFD40" pageId="30" pageNumber="29" value="2021-03-30">
<collectingDate id="7E3CE9CB921B0D1E10691878B30EFD40" pageId="30" pageNumber="29" value="2021-03-30">30 March 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E1521189BB374FD40" box="[317,340,642,665]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E154B189BB0C8FD60" collectingDate="2021-03-30" collectionCode="SEMC, V" collectorName="H. Gonzalez" country="Colombia" elevation="237" location="Beltran" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
<collectorName id="B7335335921B0D1E154B189BB3CBFD40" box="[343,491,641,665]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E15E11898B06CFD40" box="[509,588,641,665]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
); 9⚲⚲ [dark],
<collectingCountry id="62D17673921B0D1E16EC1898B146FD40" box="[752,870,641,665]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
,
<collectingRegion id="D802F801921B0D1E17681898B606FD40" box="[884,1062,641,665]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
,
<location id="1F196038921B0D1E10281898B6ABFD40" LSID="urn:lsid:plazi:treatment:926F87F592120D1E159B1C58B1D1FB00:1F196038921B0D1E10281898B6ABFD40" box="[1076,1163,641,665]" country="Colombia" name="Beltran" pageId="30" pageNumber="29" stateProvince="Cundinamarca">Beltrán</location>
, 4˚48.020N, 74˚44.394W,
<quantity id="DD3E9B06921B0D1E15DA18B8B02FFD60" box="[454,527,673,697]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">
<elevation id="91EBD1D0921B0D1E15DA18B8B02FFD60" box="[454,527,673,697]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="30" pageNumber="29" unit="m" value="237.0">237 m</elevation>
</quantity>
,
<date id="6E781023921B0D1E160618B8B0E6FD60" box="[538,710,673,697]" pageId="30" pageNumber="29" value="2021-03-30">
<collectingDate id="7E3CE9CB921B0D1E160618B8B0E6FD60" box="[538,710,673,697]" pageId="30" pageNumber="29" value="2021-03-30">30 March 2021</collectingDate>
</date>
,
<collectionCode id="7CD7AE26921B0D1E16CD18BBB0C8FD60" box="[721,744,674,697]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="29" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE921B0D1E16F018BBB1C2FD60" box="[748,994,673,697]" collectionCode="SEMC" collectorName="H. Gonzalez" country="Colombia" pageId="30" pageNumber="29" specimenCount="1" stateProvince="Cundinamarca">
<collectorName id="B7335335921B0D1E16F018BBB15CFD60" box="[748,892,673,697]" pageId="30" pageNumber="29">H. Gonzalez</collectorName>
(
<collectionCode id="7CD7AE26921B0D1E179618B8B1FAFD60" box="[906,986,673,697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="29" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D1E14CC18D8B184FC80" pageId="30" pageNumber="29" type="etymology">
<paragraph id="1A7936E3921B0D1E14CC18D8B184FC80" blockId="30.[160,1175,225,1242]" pageId="30" pageNumber="29">
ETYMOLOGY: The specific epithet refers to the
<collectingRegion id="D802F801921B0D1E16FA18D8B14AFD00" box="[742,874,705,729]" country="Colombia" name="Magdalena" pageId="30" pageNumber="29">Magdalena</collectingRegion>
River Valley and the oc- currence of the species in area around the joining of the Saldaña River with the Magda- lena River and northward. The name is formed of
<collectingRegion id="D802F801921B0D1E16EF1918B157FCC0" box="[755,887,769,793]" country="Colombia" name="Magdalena" pageId="30" pageNumber="29">Magdalena</collectingRegion>
and the Latin suffix
<emphasis id="28B2EAF1921B0D1E106B191BB6B0FCC0" box="[1143,1168,770,793]" italics="true" pageId="30" pageNumber="29">us</emphasis>
(
<emphasis id="28B2EAF1921B0D1E14AA193BB2ECFCE0" box="[182,204,802,825]" italics="true" pageId="30" pageNumber="29">ae</emphasis>
, feminine genitive singular;
<emphasis id="28B2EAF1921B0D1E162E193BB019FCE0" box="[562,569,802,825]" italics="true" pageId="30" pageNumber="29">i</emphasis>
, masculine genitive singular), which forms adjec- tives from nouns and is used for commemoration and dedication.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D1E14CC1978B1D1FB00" pageId="30" pageNumber="29" type="discussion">
<paragraph id="1A7936E3921B0D1E14CC1978B1D1FB00" blockId="30.[160,1175,225,1242]" pageId="30" pageNumber="29">
COMMENTS: Victor H. Gonzalez examined several managed nests of this species in
<collectingRegion id="D802F801921B0D1E14BC1998B370FC40" box="[160,336,897,921]" country="Colombia" name="Cundinamarca" pageId="30" pageNumber="29">Cundinamarca</collectingRegion>
and made the following observations, which he has kindly shared with me: In Beltrán, I found nests inside living trees of
<taxonomicName id="DDC64D60921B0D1E16EB19B8B6B0FC60" authority="(Kunth) Dugand" authorityName="Dugand" baseAuthorityName="Kunth" box="[759,1168,929,953]" class="Magnoliopsida" family="Fabaceae" genus="Albizia" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="guachapele">
<emphasis id="28B2EAF1921B0D1E16EB19B8B1E3FC60" box="[759,963,929,953]" italics="true" pageId="30" pageNumber="29">Albizia guachapele</emphasis>
(Kunth) Dugand
</taxonomicName>
(
<taxonomicName id="DDC64D60921B0D1E14B419D8B324FC00" box="[168,260,961,985]" class="Magnoliopsida" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="order">Fabales</taxonomicName>
:
<taxonomicName id="DDC64D60921B0D1E151319D8B35DFC00" box="[271,381,961,985]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
: Caesalpinioideae: Mimosoid clade:
<taxonomicName id="DDC64D60921B0D1E173119DBB155FC00" box="[813,885,962,985]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="28B2EAF1921B0D1E173119DBB155FC00" box="[813,885,962,985]" italics="true" pageId="30" pageNumber="29">Acacia</emphasis>
</taxonomicName>
clade: Ingeae grade; lo- cal common name: Igua),
<taxonomicName id="DDC64D60921B0D1E15D019F8B170FC20" authority="(Roxb.) Benth." authorityName="Benth." baseAuthorityName="Roxb." box="[460,848,993,1017]" class="Magnoliopsida" family="Fabaceae" genus="Pithecellobium" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="duce">
<emphasis id="28B2EAF1921B0D1E15D019F8B082FC20" box="[460,674,993,1017]" italics="true" pageId="30" pageNumber="29">Pithecellobium duce</emphasis>
(Roxb.) Benth.
</taxonomicName>
(
<taxonomicName id="DDC64D60921B0D1E174119F8B199FC20" box="[861,953,993,1017]" class="Magnoliopsida" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="order">Fabales</taxonomicName>
:
<taxonomicName id="DDC64D60921B0D1E17DE19F8B610FC20" box="[962,1072,993,1017]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
: Caesal- pinioideae: Mimosoid clade:
<taxonomicName id="DDC64D60921B0D1E15E71E1BB063FBC0" box="[507,579,1026,1049]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="28B2EAF1921B0D1E15E71E1BB063FBC0" box="[507,579,1026,1049]" italics="true" pageId="30" pageNumber="29">Acacia</emphasis>
</taxonomicName>
clade:
<taxonomicName id="DDC64D60921B0D1E16851E18B119FBC0" box="[665,825,1025,1049]" class="Magnoliopsida" family="Fabaceae" genus="Pithecellobium" kingdom="Plantae" order="Fabales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="28B2EAF1921B0D1E16851E18B119FBC0" box="[665,825,1025,1049]" italics="true" pageId="30" pageNumber="29">Pithecellobium</emphasis>
</taxonomicName>
group; local common name: Payandé), and
<taxonomicName id="DDC64D60921B0D1E15521E3BB05BFBE0" authority="Jacq." authorityName="Jacq." box="[334,635,1057,1081]" class="Magnoliopsida" family="Anacardiaceae" genus="Astronium" kingdom="Plantae" order="Sapindales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="graveolens">
<emphasis id="28B2EAF1921B0D1E15521E3BB01DFBE0" box="[334,573,1057,1081]" italics="true" pageId="30" pageNumber="29">Astronium graveolens</emphasis>
Jacq.
</taxonomicName>
(
<taxonomicName id="DDC64D60921B0D1E16941E38B12CFBE0" box="[648,780,1057,1081]" class="Magnoliopsida" kingdom="Plantae" order="Sapindales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="order">Sapindales</taxonomicName>
:
<taxonomicName id="DDC64D60921B0D1E17091E38B1E6FBE0" box="[789,966,1057,1081]" class="Magnoliopsida" family="Anacardiaceae" kingdom="Plantae" order="Sapindales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
: Anacardioideae; locally known as Diomate). Nests are found from ground level to up to
<quantity id="DD3E9B06921B0D1E101B1E58B662FB80" box="[1031,1090,1089,1113]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="29" unit="m" value="12.0">12 m</quantity>
above ground. The nest entrance is a tube of brown wax of 2.53.0 cm long and a diameter of
<quantity id="DD3E9B06921B0D1E14BC1E98B2D0FB40" box="[160,240,1153,1177]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="29" unit="mm" value="12.0">12 mm</quantity>
×
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. According to our local informant, bees are known as “enreda” — liter- ally, “entangle” — because the bees are quite defensive and they get into peoples hair. Honey is sometimes used for human consumption (Gonzalez,
<emphasis id="28B2EAF1921B0D1E17631EDBB1C1FB00" box="[895,993,1217,1241]" italics="true" pageId="30" pageNumber="29">in literis</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,245 @@
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<mods:title id="03200E5A46879347A252C6E2F3F54070">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<subSubSection id="52DC656892150D1015B419F8B661FBC0" pageId="16" pageNumber="15" type="nomenclature">
<paragraph id="1A7936E392150D1015B419F8B1A8FC20" blockId="16.[424,904,993,1017]" box="[424,904,993,1017]" pageId="16" pageNumber="15">
<heading id="4131818F92150D1015B419F8B1A8FC20" box="[424,904,993,1017]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="15" reason="2">
<taxonomicName id="DDC64D6092150D1015B419F8B0D0FC20" authority="Engel" authorityName="Engel" authorityYear="2022" box="[424,752,993,1017]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="kuperi" status="sp. nov.">
<emphasis id="28B2EAF192150D1015B419F8B084FC20" bold="true" box="[424,676,993,1017]" italics="true" pageId="16" pageNumber="15">Scaptotrigona kuperi</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92150D1016E719FBB1A8FC20" box="[763,904,993,1017]" pageId="16" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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<paragraph id="1A7936E392150D1016531E38B0C2FBE0" blockId="16.[238,1089,1027,1081]" box="[591,738,1057,1081]" pageId="16" pageNumber="15">
(
<figureCitation id="82FD2A6692150D10164B1E38B0F9FBE0" box="[599,729,1057,1081]" captionStart="Figures 1820" captionStartId="17.[160,241,1381,1403]" captionTargetBox="[177,1155,242,1370]" captionTargetId="figure-133@17.[160,1168,224,1375]" captionTargetPageId="17" captionText="Figures 1820. Worker of Scaptotrigona kuperi, new species. 18. Lateral habitus. 19. Dorsal habi- tus. 20. Facial view." pageId="16" pageNumber="15">Figs. 1820</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892150D1014CC1E78B3D8FB00" pageId="16" pageNumber="15" type="diagnosis">
<paragraph id="1A7936E392150D1014CC1E78B3D8FB00" blockId="16.[160,1169,1121,1786]" pageId="16" pageNumber="15">
DIAGNOSIS: This species is superficially similar to
<taxonomicName id="DDC64D6092150D1017011E7BB14FFBA0" authorityName="Engel" authorityYear="2022" box="[797,879,1121,1145]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192150D1017011E7BB14FFBA0" box="[797,879,1121,1145]" italics="true" pageId="16" pageNumber="15">S. ederi</emphasis>
</taxonomicName>
, which also occurs west of the Cordillera Occidental but more northerly in
<collectingCountry id="62D1767392150D1016E21E98B151FB40" box="[766,881,1153,1177]" name="Colombia" pageId="16" pageNumber="15">Colombia</collectingCountry>
(versus in north-central
<collectingCountry id="62D1767392150D1014BC1EB8B323FB60" box="[160,259,1185,1209]" name="Ecuador" pageId="16" pageNumber="15">Ecuador</collectingCountry>
in
<taxonomicName id="DDC64D6092150D10153B1EBBB3A7FB60" authorityName="Engel" authorityYear="2022" box="[295,391,1185,1209]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="kuperi">
<emphasis id="28B2EAF192150D10153B1EBBB3A7FB60" box="[295,391,1185,1209]" italics="true" pageId="16" pageNumber="15">S. kuperi</emphasis>
</taxonomicName>
). It can most easily can be distinguished from
<taxonomicName id="DDC64D6092150D1017B31EBBB1DFFB60" authorityName="Engel" authorityYear="2022" box="[943,1023,1185,1209]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192150D1017B31EBBB1DFFB60" box="[943,1023,1185,1209]" italics="true" pageId="16" pageNumber="15">S. ederi</emphasis>
</taxonomicName>
by the pres- ence of a distinct genal tooth.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892150D1214CC1EF8B30FFC22" lastPageId="18" lastPageNumber="17" pageId="16" pageNumber="15" type="description">
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DESCRIPTION: As described for
<taxonomicName id="DDC64D6092150D1016331EFBB05DFB20" authorityName="Engel" authorityYear="2022" box="[559,637,1249,1273]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192150D1016331EFBB05DFB20" box="[559,637,1249,1273]" italics="true" pageId="16" pageNumber="15">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192150D1016971EF8B0D9FB20" box="[651,761,1249,1273]" italics="true" pageId="16" pageNumber="15">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite) 6.0
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. Head wider than long, width
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, length 1.912.00 mm; compound eye length
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; upper interorbital distance
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, lower interorbital distance
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. Scape length
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, about as long as torulocellar distance, torulocellar distance
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. Clypeus approximately 1.8× as wide as long, length
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, width
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. Malar area approximately 1.61.8× flagellar diameter or 0.730.81× basal mandibular width. Preoccipital ridge carinate, weakly lamellate dorsally, with bordering medial indentation as deep as lat- eral indentations, laterally concavity interrupting carina distinct and deep, lower mar- gin of concavity with carina forming short, upward-projecting, acute, lamellate tooth.
</paragraph>
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Integument of labiomaxillary complex yellow orange; labrum yellow orange; mandible yellow orange; clypeus yellow orange to orange except apical margin often stained with brown, epistomal sulcus well delineated by brown to black; supraclypeal area yellow orange to orange; malar space yellow orange to orange; face largely yellow orange to orange except upper half of frons, vertex, posterior of head, and sometimes uppermost portion of gena dark brown to black; remainder of gena and postgena yel- low orange to orange; scape yellow orange to orange, sometimes with stain of brown dorsoapically; pedicel and flagellum typically brown, with narrow underside of flagel- lum orange to light brown. Mesosoma largely yellow orange to orange except meso- scutum black, metanotum black, V-shaped incision in mesoscutellum dark reddish brown to black, mesepisternum either black on ventral half or anterior half; propode- um black to dark reddish brown; tegula wholly yellow orange and semi-translucent; legs typically yellow orange to orange except sometimes with brown to dark brown on prolateral surfaces of tibiae, metafemur apically, and centrally on prolateral surface of metabasitarsus. Metasoma generally dark brown to black except anterior-facing surface of tergum I typically light brown to yellow brown, dorsal-facing disc of tergum I often medially lighter yellow brown with exceptionally narrow apical marginal zone of yellow; tergum II with broad medial area of light orange blending to yellow brown on disc and narrow apical marginal zone of yellow; terga
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with apical marginal zones of yellow, typically obscured by tomentum; sterna dark reddish brown to dark brown.
</paragraph>
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<emphasis id="28B2EAF192140D1114BC1F7CB318FAA3" bold="true" box="[160,312,1381,1403]" pageId="17" pageNumber="16">Figures 1820.</emphasis>
Worker of
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<emphasis id="28B2EAF192140D1115A81F7FB05EFAA3" box="[436,638,1381,1403]" italics="true" pageId="17" pageNumber="16">Scaptotrigona kuperi</emphasis>
</taxonomicName>
,
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.
<emphasis id="28B2EAF192140D1117041F7CB116FAA3" bold="true" box="[792,822,1381,1402]" pageId="17" pageNumber="16">18.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192140D1117F71F7CB629FAA3" bold="true" box="[1003,1033,1381,1402]" pageId="17" pageNumber="16">19.</emphasis>
Dorsal habi- tus.
<emphasis id="28B2EAF192140D1114CE1F9BB2D0FA4E" bold="true" box="[210,240,1410,1431]" pageId="17" pageNumber="16">20.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="1A7936E392170D1214CC1B9DB39AFC62" blockId="18.[160,1175,228,1244]" pageId="18" pageNumber="17">
As for
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<emphasis id="28B2EAF192170D12153E1B9DB354FE45" box="[290,372,388,412]" italics="true" pageId="18" pageNumber="17">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white. Bristles of vertex erect, simple, yellow to fulvous, with those short bristles near upper orbit often fuscous and sometimes a few longer fuscous bristles intermixed posteriorly on vertex medially; pronotal dorsolateral margin near and anterior to pro- notal lobe with long, arched, yellow bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with mix of yellow and dark fulvous to black bristles, such bristles intermixed along entire margin; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elon- gate, erect bristles posteriorly, such bristles minutely pectinate apically, discal bristles shorter, mesoscutellar bristles yellow; mesepisternal bristles as in
<taxonomicName id="DDC64D6092170D1217A918BDB627FD65" authorityName="Engel" authorityYear="2022" box="[949,1031,676,700]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192170D1217A918BDB627FD65" box="[949,1031,676,700]" italics="true" pageId="18" pageNumber="17">S. ederi</emphasis>
</taxonomicName>
except yel- low. Legs with abundant bristles, bristles typically yellow except noticeably inter- mixed with black bristles on prolateral surface of protibia apically, prolateral surface of probasitarsus, prolateral and marginal surface of mesotibia, prolateral surface of mesobasitarsus, metafemur apically, metatibia, and metabasitarsus marginal bristles; prolateral surface of metabasitarsus with scatered fine, simple, black setae (notice- ably thinner than marginal bristles) intermixed with similar yellow setae and bristles. Metasoma with numerous, prominent, subdecumbent, yellow bristles on terga
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arising from tomentum.
</paragraph>
<paragraph id="1A7936E392170D1214CC19DDB30FFC02" blockId="18.[160,1175,228,1244]" box="[208,303,964,990]" pageId="18" pageNumber="17">
<specimenCount id="0CC0FD6A92170D1214CC19DDB2C4FC07" box="[208,228,964,990]" count="1" pageId="18" pageNumber="17" type="female"></specimenCount>
:
<emphasis id="28B2EAF192170D1214ED19DCB308FC05" box="[241,296,965,988]" italics="true" pageId="18" pageNumber="17">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392170D1214CC19FDB30FFC22" blockId="18.[160,1175,228,1244]" box="[208,303,996,1022]" pageId="18" pageNumber="17">
<specimenCount id="0CC0FD6A92170D1214CC19FDB2C4FC27" box="[208,228,996,1022]" count="1" pageId="18" pageNumber="17" type="male"></specimenCount>
:
<emphasis id="28B2EAF192170D1214ED19FCB308FC25" box="[241,296,997,1020]" italics="true" pageId="18" pageNumber="17">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892170D1214CC1E1DB13CFBA5" pageId="18" pageNumber="17" type="materials_examined">
<paragraph id="1A7936E392170D1214CC1E1DB13CFBE5" blockId="18.[160,1175,228,1244]" pageId="18" pageNumber="17">
<materialsCitation id="AAAE3CBE92170D1214CC1E1DB13CFBE5" collectingDate="1999-03-29" collectionCode="R, SEMC" country="Ecuador" elevation="530" latitude="0.112222224" location="Pedro Vicente Maldonado" longLatPrecision="21" longitude="-79.05584" pageId="18" pageNumber="17" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
<typeStatus id="C57D884192170D1214CC1E1DB368FBC5" box="[208,328,1028,1052]" pageId="18" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392170D1215691E1DB3FBFBC5" box="[373,475,1028,1052]" name="Ecuador" pageId="18" pageNumber="17">Ecuador</collectingCountry>
:
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,
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,
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N,
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,
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,
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,
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<collectingDate id="7E3CE9CB92170D1215B91E3DB016FBE5" box="[421,566,1060,1084]" pageId="18" pageNumber="17" value="1999-03-29">29 Mar 1999</collectingDate>
</date>
,
<collectionCode id="7CD7AE2692170D12165D1E3DB076FBE2" box="[577,598,1060,1083]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
. Brooks (
<collectionCode id="7CD7AE2692170D1216DC1E3DB130FBE5" box="[704,784,1060,1084]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="18" pageNumber="17" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392170D1214CC1E5DB13CFBA5" blockId="18.[160,1175,228,1244]" pageId="18" pageNumber="17">
<materialsCitation id="AAAE3CBE92170D1214CC1E5DB076FBA2" collectingDate="1999-03-29" collectionCode="R" country="Ecuador" elevation="530" latitude="0.112222224" location="Pedro Vicente Maldonado" longLatPrecision="21" longitude="-79.05584" pageId="18" pageNumber="17" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<typeStatus id="C57D884192170D1214CC1E5DB363FB85" box="[208,323,1092,1116]" pageId="18" pageNumber="17" type="paratype">PARATYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392170D12156D1E5DB3F7FB85" box="[369,471,1092,1116]" name="Ecuador" pageId="18" pageNumber="17">Ecuador</collectingCountry>
:
<collectingRegion id="D802F80192170D1215FA1E5DB07CFB85" box="[486,604,1092,1116]" country="Ecuador" name="Pichincha" pageId="18" pageNumber="17">Pichincha</collectingRegion>
,
<location id="1F19603892170D1216771E5DB189FB85" LSID="urn:lsid:plazi:treatment:926F87F592150D1215B419F8B303FB05:1F19603892170D1216771E5DB189FB85" box="[619,937,1092,1116]" country="Ecuador" latitude="0.112222224" longLatPrecision="21" longitude="-79.05584" name="Pedro Vicente Maldonado" pageId="18" pageNumber="17" stateProvince="Pichincha">Pedro Vicente Maldonado</location>
,
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N,
<quantity id="DD3E9B0692170D1210231E5DB6ACFB82" box="[1087,1164,1092,1116]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="18" pageNumber="17" unit="m" value="530.0">
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,
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,
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,
<date id="6E78102392170D1215B91E7DB016FBA5" box="[421,566,1124,1148]" pageId="18" pageNumber="17" value="1999-03-29">
<collectingDate id="7E3CE9CB92170D1215B91E7DB016FBA5" box="[421,566,1124,1148]" pageId="18" pageNumber="17" value="1999-03-29">29 Mar 1999</collectingDate>
</date>
,
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</materialsCitation>
.
<materialsCitation id="AAAE3CBE92170D12167C1E7DB138FBA5" box="[608,792,1124,1148]" collectionCode="SEMC" country="Ecuador" location="Brooks" pageId="18" pageNumber="17" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<location id="1F19603892170D12167C1E7DB092FBA5" LSID="urn:lsid:plazi:treatment:926F87F592150D1215B419F8B303FB05:1F19603892170D12167C1E7DB092FBA5" box="[608,690,1124,1148]" country="Ecuador" name="Brooks" pageId="18" pageNumber="17" stateProvince="Pichincha">Brooks</location>
(
<collectionCode id="7CD7AE2692170D1216DC1E7DB130FBA5" box="[704,784,1124,1148]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="18" pageNumber="17" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892170D1214CC1E9DB303FB05" pageId="18" pageNumber="17" type="etymology">
<paragraph id="1A7936E392170D1214CC1E9DB303FB05" blockId="18.[160,1175,228,1244]" pageId="18" pageNumber="17">ETYMOLOGY: The specific epithet honors Peter Kuper, acclaimed illustrator, thought-provoking visual artist, inspiring author, devoted and life-long entomologist, and friend.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,243 @@
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<mods:title id="2112B929FFB9053381F3F494F67F774D">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<treatment id="926F87F592170D1515871F1DB2C5FBE0" LSID="urn:lsid:plazi:treatment:926F87F592170D1515871F1DB2C5FBE0" httpUri="http://treatment.plazi.org/id/926F87F592170D1515871F1DB2C5FBE0" lastPageId="21" lastPageNumber="20" pageId="18" pageNumber="17">
<subSubSection id="52DC656892170D1215871F1DB611FAE5" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="1A7936E392170D1215871F1DB1B5FAC5" blockId="18.[411,917,1284,1308]" box="[411,917,1284,1308]" pageId="18" pageNumber="17">
<heading id="4131818F92170D1215871F1DB1B5FAC5" box="[411,917,1284,1308]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="17" reason="2">
<taxonomicName id="DDC64D6092170D1215871F1DB0DEFAC5" authority="Engel" authorityName="Engel" authorityYear="2022" box="[411,766,1284,1308]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="extranea" status="sp. nov.">
<emphasis id="28B2EAF192170D1215871F1DB091FAC2" bold="true" box="[411,689,1284,1308]" italics="true" pageId="18" pageNumber="17">Scaptotrigona extranea</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92170D1217141F1DB1B5FAC5" box="[776,917,1284,1308]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392170D1214E31F3FB611FAE5" blockId="18.[255,1073,1318,1372]" box="[255,1073,1318,1340]" pageId="18" pageNumber="17">
ZooBank:
<uri id="6E573AE192170D1215771F3FB611FAE5" box="[363,1073,1318,1340]" pageId="18" pageNumber="17">
urn:lsid:zoobank.org:act:
<uuid id="6E600C3692170D1216431F3FB611FAE5" box="[607,1073,1318,1340]" pageId="18" pageNumber="17">29C44161-56F9-4AE4-8326-D8A51217F494</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892170D1216531F5DB0C2FA85" box="[591,738,1348,1372]" pageId="18" pageNumber="17" type="description">
<paragraph id="1A7936E392170D1216531F5DB0C2FA85" blockId="18.[255,1073,1318,1372]" box="[591,738,1348,1372]" pageId="18" pageNumber="17">
(
<figureCitation id="82FD2A6692170D12164B1F5DB0F9FA85" box="[599,729,1348,1372]" captionStart-0="Figures 2123" captionStart-1="Figures 2425" captionStartId-0="19.[160,241,1454,1476]" captionStartId-1="20.[160,241,1035,1057]" captionTargetBox-0="[170,1158,244,1425]" captionTargetBox-1="[160,1160,224,1029]" captionTargetId-0="figure-99@19.[160,1168,221,1448]" captionTargetId-1="figure-260@20.[160,1168,224,1029]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Figures 2123. Worker of Scaptotrigona extranea, new species. 21. Lateral habitus. 22. Dorsal habitus. 23. Facial view." captionText-1="Figures 2425. Worker of Scaptotrigona extranea, new species. 24. Dorsal view, head and meso- soma. 25. Dorsal oblique view of vertex and upper gena." pageId="18" pageNumber="17">Figs. 2125</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892170D1214CC1F9DB073F965" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="1A7936E392170D1214CC1F9DB073F965" blockId="18.[160,1168,1412,1788]" pageId="18" pageNumber="17">
DIAGNOSIS: This is a rather unique species in that the concavity that typically in- terrupts the preoccipital carina laterally is exceptionally shallow and faint and does not disrupt the carina (
<figureCitation id="82FD2A6692170D1215A71FDDB02FFA05" box="[443,527,1476,1500]" captionStart="Figures 2425" captionStartId="20.[160,241,1035,1057]" captionTargetBox="[160,1160,224,1029]" captionTargetId="figure-260@20.[160,1168,224,1029]" captionTargetPageId="20" captionText="Figures 2425. Worker of Scaptotrigona extranea, new species. 24. Dorsal view, head and meso- soma. 25. Dorsal oblique view of vertex and upper gena." pageId="18" pageNumber="17">Fig. 25</figureCitation>
). In addition, the preoccipital ridge is faintly lamel- late dorsally and the bordering medial and lateral indentations are weak and shallow (
<figureCitation id="82FD2A6692170D1214B41C1DB2DAF9C5" box="[168,250,1540,1564]" captionStart="Figures 2425" captionStartId="20.[160,241,1035,1057]" captionTargetBox="[160,1160,224,1029]" captionTargetId="figure-260@20.[160,1168,224,1029]" captionTargetPageId="20" captionText="Figures 2425. Worker of Scaptotrigona extranea, new species. 24. Dorsal view, head and meso- soma. 25. Dorsal oblique view of vertex and upper gena." pageId="18" pageNumber="17">Fig. 25</figureCitation>
). It is similar in several respects to
<taxonomicName id="DDC64D6092170D1216BD1C1DB122F9C5" authorityName="Engel" authorityYear="2022" box="[673,770,1540,1564]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="kuperi">
<emphasis id="28B2EAF192170D1216BD1C1DB122F9C5" box="[673,770,1540,1564]" italics="true" pageId="18" pageNumber="17">S. kuperi</emphasis>
</taxonomicName>
, but can also be distinguished by the facial coloration in which the frons is largely dark brown to black, with yellow to yellow orange extending upward along the inner orbits (
<figureCitation id="82FD2A6692170D12175F1C5DB1B5F985" box="[835,917,1604,1628]" captionStart="Figures 2123" captionStartId="19.[160,241,1454,1476]" captionTargetBox="[170,1158,244,1425]" captionTargetId="figure-99@19.[160,1168,221,1448]" captionTargetPageId="19" captionText="Figures 2123. Worker of Scaptotrigona extranea, new species. 21. Lateral habitus. 22. Dorsal habitus. 23. Facial view." pageId="18" pageNumber="17">Fig. 23</figureCitation>
), while in the former the entire lower part of the frons orange (
<figureCitation id="82FD2A6692170D1216951C7DB0FBF9A5" box="[649,731,1636,1660]" captionStart="Figures 1820" captionStartId="17.[160,241,1381,1403]" captionTargetBox="[177,1155,242,1370]" captionTargetId="figure-133@17.[160,1168,224,1375]" captionTargetPageId="17" captionText="Figures 1820. Worker of Scaptotrigona kuperi, new species. 18. Lateral habitus. 19. Dorsal habi- tus. 20. Facial view." pageId="18" pageNumber="17">Fig. 20</figureCitation>
). In addition, the back of the head is entirely dark brown to black in
<taxonomicName id="DDC64D6092170D12160A1C9DB057F945" authorityName="Engel" authorityYear="2022" box="[534,631,1668,1692]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="kuperi">
<emphasis id="28B2EAF192170D12160A1C9DB057F945" box="[534,631,1668,1692]" italics="true" pageId="18" pageNumber="17">S. kuperi</emphasis>
</taxonomicName>
, versus orange and marked with dark brown stripes in
<taxonomicName id="DDC64D6092170D12150F1CBDB3ABF965" authorityName="Engel" authorityYear="2022" box="[275,395,1700,1724]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="extranea">
<emphasis id="28B2EAF192170D12150F1CBDB3ABF965" box="[275,395,1700,1724]" italics="true" pageId="18" pageNumber="17">S. extranea</emphasis>
</taxonomicName>
(
<figureCitation id="82FD2A6692170D1215861CBDB3D6F965" box="[410,502,1700,1724]" captionStart="Figures 2123" captionStartId="19.[160,241,1454,1476]" captionTargetBox="[170,1158,244,1425]" captionTargetId="figure-99@19.[160,1168,221,1448]" captionTargetPageId="19" captionText="Figures 2123. Worker of Scaptotrigona extranea, new species. 21. Lateral habitus. 22. Dorsal habitus. 23. Facial view." pageId="18" pageNumber="17">Figs. 22</figureCitation>
,
<figureCitation id="82FD2A6692170D12161D1CBDB067F965" box="[513,583,1700,1724]" captionStart="Figures 2425" captionStartId="20.[160,241,1035,1057]" captionTargetBox="[160,1160,224,1029]" captionTargetId="figure-260@20.[160,1168,224,1029]" captionTargetPageId="20" captionText="Figures 2425. Worker of Scaptotrigona extranea, new species. 24. Dorsal view, head and meso- soma. 25. Dorsal oblique view of vertex and upper gena." pageId="18" pageNumber="17">24, 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="52DC656892170D1514CC1CDDB30FFC40" lastPageId="21" lastPageNumber="20" pageId="18" pageNumber="17" type="description">
<paragraph id="1A7936E392170D1314CC1CDDB049F8DA" blockId="18.[160,1168,1412,1788]" lastBlockId="19.[160,1169,1547,1795]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092170D12165F1CDDB0B6F905" authorityName="Engel" authorityYear="2022" box="[579,662,1732,1756]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192170D12165F1CDDB0B6F905" box="[579,662,1732,1756]" italics="true" pageId="18" pageNumber="17">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192170D1216B51CDDB13CF905" box="[681,796,1732,1756]" italics="true" pageId="18" pageNumber="17">vide supra</emphasis>
), with the following modifica- tions: ⚲: Total body length approximately
<quantity id="DD3E9B0692170D1216961CFDB0C2F922" box="[650,738,1764,1788]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="18" pageNumber="17" unit="mm" value="6.3">6.3 mm</quantity>
, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B0692160D1315181C12B37FF9FB" box="[260,351,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="19" pageNumber="18" unit="mm" value="5.8">5.8 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B0692160D1316C91C12B11DF9FB" box="[725,829,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="18" unit="mm" value="2.3">2.30 mm</quantity>
, length
<quantity id="DD3E9B0692160D1317801C12B623F9FB" box="[924,1027,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="19" pageNumber="18" unit="mm" value="1.91">1.91 mm</quantity>
; compound eye length
<quantity id="DD3E9B0692160D1315381C32B3ACF99B" box="[292,396,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="19" pageNumber="18" unit="mm" value="1.33">1.33 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B0692160D1316FC1C32B168F99B" box="[736,840,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="19" pageNumber="18" unit="mm" value="1.48">1.48 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B0692160D1314BC1C52B327F9BB" box="[160,263,1611,1635]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="19" pageNumber="18" unit="mm" value="1.42">1.42 mm</quantity>
. Scape length
<quantity id="DD3E9B0692160D1315A91C52B03CF9BB" box="[437,540,1611,1635]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="19" pageNumber="18" unit="mm" value="0.88">0.88 mm</quantity>
, as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B0692160D1314BC1C72B32AF95B" box="[160,266,1643,1667]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="19" pageNumber="18" unit="mm" value="0.88">0.88 mm</quantity>
. Clypeus approximately 1.7× as wide as long, length
<quantity id="DD3E9B0692160D1317871C72B624F95B" box="[923,1028,1643,1667]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="19" pageNumber="18" unit="mm" value="0.58">0.58 mm</quantity>
, width 1.00 mm. Malar area approximately 1.6× flagellar diameter or 0.73× basal mandibular width. Preoccipital ridge carinate, faintly lamellate dorsally, with bordering medial and lateral indentations weak and shallow, carina not interrupted laterally, merely a faint concavity not interrupting carina.
</paragraph>
<caption id="4EB9666B92160D1314BC1FB7B383FA38" pageId="19" pageNumber="18" startId="19.[160,241,1454,1476]" targetBox="[170,1158,244,1425]" targetPageId="19" targetType="figure">
<paragraph id="1A7936E392160D1314BC1FB7B383FA38" blockId="19.[160,1168,1454,1505]" pageId="19" pageNumber="18">
<emphasis id="28B2EAF192160D1314BC1FB7B31BFA1A" bold="true" box="[160,315,1454,1476]" pageId="19" pageNumber="18">Figures 2123.</emphasis>
Worker of
<taxonomicName id="DDC64D6092160D1315DD1FB6B083FA1D" authorityName="Engel" authorityYear="2022" box="[449,675,1455,1476]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="extranea" status="sp. nov.">
<emphasis id="28B2EAF192160D1315DD1FB6B083FA1D" box="[449,675,1455,1476]" italics="true" pageId="19" pageNumber="18">Scaptotrigona extranea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92160D1316AC1FB6B114FA1D" box="[688,820,1454,1476]" pageId="19" pageNumber="18" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192160D13175B1FB7B145FA1A" bold="true" box="[839,869,1454,1475]" pageId="19" pageNumber="18">21.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192160D1310381FB7B661FA1A" bold="true" box="[1060,1089,1454,1475]" pageId="19" pageNumber="18">22.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF192160D13151D1FD2B33FFA39" bold="true" box="[257,287,1483,1504]" pageId="19" pageNumber="18">23.</emphasis>
Facial view.
</paragraph>
</caption>
<caption id="4EB9666B92110D1414BC1E12B0DCFBE7" pageId="20" pageNumber="19" startId="20.[160,241,1035,1057]" targetBox="[160,1160,224,1029]" targetPageId="20" targetType="figure">
<paragraph id="1A7936E392110D1414BC1E12B0DCFBE7" blockId="20.[160,1168,1035,1086]" pageId="20" pageNumber="19">
<emphasis id="28B2EAF192110D1414BC1E12B319FBF8" bold="true" box="[160,313,1035,1057]" pageId="20" pageNumber="19">Figures 2425.</emphasis>
Worker of
<taxonomicName id="DDC64D6092110D1415A41E15B0B9FBF8" authorityName="Engel" authorityYear="2022" box="[440,665,1036,1057]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="extranea" status="sp. nov.">
<emphasis id="28B2EAF192110D1415A41E15B0B9FBF8" box="[440,665,1036,1057]" italics="true" pageId="20" pageNumber="19">Scaptotrigona extranea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92110D1416B81E15B106FBFB" box="[676,806,1036,1058]" pageId="20" pageNumber="19" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192110D14172A1E15B173FBF8" bold="true" box="[822,851,1036,1057]" pageId="20" pageNumber="19">24.</emphasis>
Dorsal view, head and meso- soma.
<emphasis id="28B2EAF192110D1414F61E31B329FBE4" bold="true" box="[234,265,1064,1085]" pageId="20" pageNumber="19">25.</emphasis>
Dorsal oblique view of vertex and upper gena.
</paragraph>
</caption>
<paragraph id="1A7936E392110D1414CC1E70B116FB59" blockId="20.[160,1168,1128,1792]" box="[208,822,1128,1152]" pageId="20" pageNumber="19">
Punctures of face fainter and sparser than in
<taxonomicName id="DDC64D6092110D1416C31E70B10FFB59" authorityName="Engel" authorityYear="2022" box="[735,815,1128,1152]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192110D1416C31E70B10FFB59" box="[735,815,1128,1152]" italics="true" pageId="20" pageNumber="19">S. ederi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1A7936E392110D1414CC1E90B063F939" blockId="20.[160,1168,1128,1792]" pageId="20" pageNumber="19">Integument of labiomaxillary complex yellow orange; labrum yellow orange; mandible orange; clypeus yellow orange stained with areas of brown laterally and mediobasally, apical margin stained with brown, epistomal sulcus well delineated by dark brown to black; supraclypeal area yellow orange; malar space orange; face below lower tangent of antennal toruli yellow, with narrow yellow strip extending upward along inner orbit, frons and vertex dark brown to black albeit dark reddish brown above antennal toruli, separated from upper orbit by strip of orange to reddish brown, yellow orange mark anterior to median ocellus, most of ocellocular area orange; pos- terior of head largely orange except with oblique lateral strips of reddish brown ex- tending from ocellocular area to preoccipital carina, preoccipital carina black; gena and postgena orange to yellow orange; scape orange; pedicel and flagellum brown, with flagellum orange ventrally. Mesosoma largely orange except mesoscutum black, mesoscutellum largely black except dark fulvous to ferrugineous apically, metanotum black, basal area and posterior surface of propodeum black; mesepisternum with dark brown ventrally and metepisternum dark brown long borders; tegula wholly yellow orange and semi-translucent; legs orange. Metasoma generally dark brown to black except anterior-facing surface of tergum I reddish brown, dorsal-facing disc of tergum I dark orange to brown apically; tergum II with broad apical zone of brown; sterna dark reddish brown to dark brown.</paragraph>
<paragraph id="1A7936E392110D1514CC1CF1B638FC80" blockId="20.[160,1168,1128,1792]" lastBlockId="21.[160,1168,225,1081]" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19">
As for
<taxonomicName id="DDC64D6092110D1415021CF0B34EF8D9" authorityName="Engel" authorityYear="2022" box="[286,366,1768,1792]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192110D1415021CF0B34EF8D9" box="[286,366,1768,1792]" italics="true" pageId="20" pageNumber="19">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white; metepisternum and lateral surface of propodeum obscured by dense, plumose, yellow setae; dorsal-facing surface of tergum I densely covered with minute, decum- bent, yellow, plumose setae similar to tomentum of apical margin of tergum II and discs of terga
<collectionCode id="7CD7AE2692100D15155A1B5BB341FE80" box="[326,353,322,345]" pageId="21" pageNumber="20">III</collectionCode>
<collectionCode id="7CD7AE2692100D1515731B5BB3ACFE80" box="[367,396,322,345]" pageId="21" pageNumber="20">IV</collectionCode>
. Bristles of vertex erect, simple, yellow to fulvous; pronotal dor- solateral margin near and anterior to pronotal lobe with long, arched, yellow bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with mix of yellow and dark fulvous to black bristles, such bris- tles intermixed along entire margin; tegula with short, suberect, yellow bristles ante- riorly; mesoscutellum with abundant, elongate, erect bristles posteriorly, such bristles minutely pectinate apically, discal bristles shorter, mesoscutellar bristles largely yel- low except medioapically more fulvous to fuscous; mesepisternal bristles as in
<taxonomicName id="DDC64D6092100D15105C183BB6B0FDE0" authorityName="Engel" authorityYear="2022" box="[1088,1168,545,569]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192100D15105C183BB6B0FDE0" box="[1088,1168,545,569]" italics="true" pageId="21" pageNumber="20">S. ederi</emphasis>
</taxonomicName>
except yellow. Legs with abundant bristles, bristles typically yellow to orange except noticeably intermixed with fuscous to black bristles margin of protibia, prolateral sur- face of probasitarsus, prolateral and marginal surface of mesotibia, prolateral surface of mesobasitarsus, metafemur apically, metatibia, and a few metabasitarsus marginal bristles; corbicular bristles black, contrasting fulvous marginal bristles, some of later with fuscous bases; prolateral surface of metabasitarsus with scatered fine, simple, yellow to fulvous setae (noticeably thinner than marginal bristles), without black se- tae; penicillum black; rastellum yellow. Metasoma with numerous, prominent, suber- ect to subdecumbent, yellow bristles on terga
<collectionCode id="7CD7AE2692100D1516A7195BB0F6FC80" box="[699,726,834,857]" pageId="21" pageNumber="20">III</collectionCode>
<collectionCode id="7CD7AE2692100D1516FF195BB0D6FC80" box="[739,758,834,857]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="21" pageNumber="20" type="Museum">V</collectionCode>
arising from tomentum.
</paragraph>
<paragraph id="1A7936E392100D1514CC197BB30FFCA0" blockId="21.[160,1168,225,1081]" box="[208,303,866,892]" pageId="21" pageNumber="20">
<specimenCount id="0CC0FD6A92100D1514CC197BB2C4FCA5" box="[208,228,866,892]" count="1" pageId="21" pageNumber="20" type="female"></specimenCount>
:
<emphasis id="28B2EAF192100D1514ED197BB308FCA0" box="[241,296,866,889]" italics="true" pageId="21" pageNumber="20">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392100D1514CC199BB30FFC40" blockId="21.[160,1168,225,1081]" box="[208,303,898,924]" pageId="21" pageNumber="20">
<specimenCount id="0CC0FD6A92100D1514CC199BB2C4FC45" box="[208,228,898,924]" count="1" pageId="21" pageNumber="20" type="male"></specimenCount>
:
<emphasis id="28B2EAF192100D1514ED199BB308FC40" box="[241,296,898,921]" italics="true" pageId="21" pageNumber="20">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1514CC19B8B13CFC00" pageId="21" pageNumber="20" type="materials_examined">
<paragraph id="1A7936E392100D1514CC19B8B13CFC00" blockId="21.[160,1168,225,1081]" pageId="21" pageNumber="20">
<materialsCitation id="AAAE3CBE92100D1514CC19B8B13CFC00" collectingDate="1999-03-29" collectionCode="R, SEMC" country="Ecuador" elevation="530" latitude="0.112222224" location="Pedro Vicente Maldonado" longLatPrecision="21" longitude="-79.05584" pageId="21" pageNumber="20" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
<typeStatus id="C57D884192100D1514CC19B8B368FC60" box="[208,328,929,953]" pageId="21" pageNumber="20" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392100D15156919B8B3FBFC60" box="[373,475,929,953]" name="Ecuador" pageId="21" pageNumber="20">Ecuador</collectingCountry>
:
<collectingRegion id="D802F80192100D1515F519B8B07FFC60" box="[489,607,929,953]" country="Ecuador" name="Pichincha" pageId="21" pageNumber="20">Pichincha</collectingRegion>
,
<location id="1F19603892100D15167219B8B18CFC60" LSID="urn:lsid:plazi:treatment:926F87F592170D1515871F1DB2C5FBE0:1F19603892100D15167219B8B18CFC60" box="[622,940,929,953]" country="Ecuador" latitude="0.112222224" longLatPrecision="21" longitude="-79.05584" name="Pedro Vicente Maldonado" pageId="21" pageNumber="20" stateProvince="Pichincha">Pedro Vicente Maldonado</location>
,
<quantity id="DD3E9B0692100D1517A719B8B62DFC60" box="[955,1037,929,953]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="20" unit="km" value="3.5">3.5 km</quantity>
N,
<quantity id="DD3E9B0692100D15102319B8B6ACFC60" box="[1087,1164,929,953]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="21" pageNumber="20" unit="m" value="530.0">
<elevation id="91EBD1D092100D15102319B8B6ACFC60" box="[1087,1164,929,953]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="21" pageNumber="20" unit="m" value="530.0">530 m</elevation>
</quantity>
,
<geoCoordinate id="7FF2502492100D1514BC19D8B32EFC00" box="[160,270,961,985]" degrees="0" direction="north" minutes="6" orientation="latitude" pageId="21" pageNumber="20" precision="15" seconds="44" value="0.112222224">0°644N</geoCoordinate>
,
<geoCoordinate id="7FF2502492100D15150619D8B3BAFC00" box="[282,410,961,985]" degrees="79" direction="west" minutes="3" orientation="longitude" pageId="21" pageNumber="20" precision="15" seconds="21" value="-79.05584">79°321W</geoCoordinate>
,
<date id="6E78102392100D1515B919D8B016FC00" box="[421,566,961,985]" pageId="21" pageNumber="20" value="1999-03-29">
<collectingDate id="7E3CE9CB92100D1515B919D8B016FC00" box="[421,566,961,985]" pageId="21" pageNumber="20" value="1999-03-29">29 Mar 1999</collectingDate>
</date>
,
<collectionCode id="7CD7AE2692100D15165D19DBB076FC00" box="[577,598,962,985]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="20">R</collectionCode>
. Brooks (
<collectionCode id="7CD7AE2692100D1516DC19D8B130FC00" box="[704,784,961,985]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="21" pageNumber="20" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1514CC19F8B2C5FBE0" pageId="21" pageNumber="20" type="etymology">
<paragraph id="1A7936E392100D1514CC19F8B2C5FBE0" blockId="21.[160,1168,225,1081]" pageId="21" pageNumber="20">
ETYMOLOGY: The specific epithet is the Latin adjective
<emphasis id="28B2EAF192100D15172D19FBB1BAFC20" box="[817,922,994,1017]" italics="true" pageId="21" pageNumber="20">extrāneus</emphasis>
, meaning, “strange”, and refers to the near absence of a concavity in the lateral portion of the preoccipital ridge.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,272 @@
<document id="23DB8759AB01AAFEEE793590A1523DFA" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F5921B0D2115961F18B18AFA20" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona tatacoensis Engel 2022, new species" docType="treatment" docVersion="1" lastPageNumber="32" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="29" updateTime="1722456085954" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="240683DD355D1FB6953E7F8A079B3F4B">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="6536C91A332F502686CA3D4895EA66F6">Engel, Michael S.</mods:namePart>
<mods:affiliation id="B165E5A69A32FA40D8B09C2941CCD11A">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<treatment id="926F87F5921B0D2115961F18B18AFA20" LSID="urn:lsid:plazi:treatment:926F87F5921B0D2115961F18B18AFA20" httpUri="http://treatment.plazi.org/id/926F87F5921B0D2115961F18B18AFA20" lastPageId="33" lastPageNumber="32" pageId="30" pageNumber="29">
<subSubSection id="52DC6568921B0D1E15961F18B663FAE0" pageId="30" pageNumber="29" type="nomenclature">
<paragraph id="1A7936E3921B0D1E15961F18B186FAC0" blockId="30.[394,934,1281,1305]" box="[394,934,1281,1305]" pageId="30" pageNumber="29">
<heading id="4131818F921B0D1E15961F18B186FAC0" box="[394,934,1281,1305]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="29" reason="2">
<taxonomicName id="DDC64D60921B0D1E15961F18B12FFAC0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[394,783,1281,1305]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="tatacoensis" status="sp. nov.">
<emphasis id="28B2EAF1921B0D1E15961F18B0E2FAC0" bold="true" box="[394,706,1281,1305]" italics="true" pageId="30" pageNumber="29">Scaptotrigona tatacoensis</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A921B0D1E17051F1BB186FAC0" box="[793,934,1281,1305]" pageId="30" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D1E16531F58B0C1FA80" box="[591,737,1345,1369]" pageId="30" pageNumber="29" type="description">
<paragraph id="1A7936E3921B0D1E16531F58B0C1FA80" blockId="30.[237,1091,1315,1369]" box="[591,737,1345,1369]" pageId="30" pageNumber="29">
(
<figureCitation id="82FD2A66921B0D1E164B1F58B0FAFA80" box="[599,730,1345,1369]" captionStart-0="Figures 4446" captionStart-1="Figures 4748" captionStartId-0="31.[160,241,1457,1479]" captionStartId-1="32.[160,241,1288,1310]" captionTargetBox-0="[171,1149,238,1432]" captionTargetBox-1="[180,1143,224,1274]" captionTargetId-0="figure-88@31.[160,1168,224,1451]" captionTargetId-1="figure-177@32.[160,1168,224,1282]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Figures 4446. Worker of Scaptotrigona tatacoensis, new species. 44. Lateral habitus. 45. Dorsal habitus. 46. Facial view." captionText-1="Figures 4748. Worker of Scaptotrigona tatacoensis, new species. 47. Dorsal, slightly oblique, view of head. 48. Prolateral surface of metatibia, metatarsus, and metapretarsus." pageId="30" pageNumber="29">Figs. 4448</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D1E14CC1F98B16DF9E0" pageId="30" pageNumber="29" type="diagnosis">
<paragraph id="1A7936E3921B0D1E14CC1F98B16DF9E0" blockId="30.[160,1169,1409,1785]" pageId="30" pageNumber="29">
DIAGNOSIS: This is a distinctive, smaller species of
<taxonomicName id="DDC64D60921B0D1E17391F9BB19BFA40" box="[805,955,1410,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF1921B0D1E17391F9BB19BFA40" box="[805,955,1410,1433]" italics="true" pageId="30" pageNumber="29">Scaptotrigona</emphasis>
</taxonomicName>
, currently known only from the Tatacoa “desert” (tropical dry forest) of
<collectingCountry id="62D17673921B0D1E17241FB8B18EFA60" box="[824,942,1441,1465]" name="Colombia" pageId="30" pageNumber="29">Colombia</collectingCountry>
. Like some larger species, it has striped markings on the mesoscutum, but can be distinguished by the separation of the dark region of the frons from the inner orbits by a strip of yellow orange to reddish orange, by the striping on the posterior of the head, and the black bristles of the mesobasitarsus and metabasitarsal margins.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568921B0D2114CC1C58B30FFAC0" lastPageId="33" lastPageNumber="32" pageId="30" pageNumber="29" type="description">
<paragraph id="1A7936E3921B0D1F14CC1C58B63CF91F" blockId="30.[160,1169,1409,1785]" lastBlockId="31.[160,1168,1550,1766]" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29">
DESCRIPTION: As described for
<taxonomicName id="DDC64D60921B0D1E16331C5BB05DF980" authorityName="Engel" authorityYear="2022" box="[559,637,1601,1625]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1921B0D1E16331C5BB05DF980" box="[559,637,1601,1625]" italics="true" pageId="30" pageNumber="29">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF1921B0D1E16971C58B0D9F980" box="[651,761,1601,1625]" italics="true" pageId="30" pageNumber="29">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately 5.56.0 mm, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B06921B0D1E151B1C98B3B3F940" box="[263,403,1665,1689]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="30" pageNumber="29" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.45.8 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B06921B0D1E17071C98B1E1F940" box="[795,961,1665,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.315" metricValueMax="2.36" metricValueMin="2.27" pageId="30" pageNumber="29" unit="mm" value="2.315" valueMax="2.36" valueMin="2.27">2.272.36 mm</quantity>
, length
<quantity id="DD3E9B06921B0D1E10391C98B2F1F960" metricMagnitude="-3" metricUnit="m" metricValue="1.895" metricValueMax="1.91" metricValueMin="1.88" pageId="30" pageNumber="29" unit="mm" value="1.895" valueMax="1.91" valueMin="1.88">1.881.91 mm</quantity>
; compound eye length
<quantity id="DD3E9B06921B0D1E15FB1CB8B06EF960" box="[487,590,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="30" pageNumber="29" unit="mm" value="1.33">1.33 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06921B0D1E17821CB8B661F960" box="[926,1089,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.53" metricValueMin="1.52" pageId="30" pageNumber="29" unit="mm" value="1.525" valueMax="1.53" valueMin="1.52">1.521.53 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06921B0D1E158F1CD8B016F900" box="[403,566,1729,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" metricValueMax="1.44" metricValueMin="1.42" pageId="30" pageNumber="29" unit="mm" value="1.43" valueMax="1.44" valueMin="1.42">1.421.44 mm</quantity>
. Scape length
<quantity id="DD3E9B06921B0D1E16FE1CD8B168F900" box="[738,840,1729,1753]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="30" pageNumber="29" unit="mm" value="0.88">0.88 mm</quantity>
, slightly longer than torulo- cellar distance, torulocellar distance
<quantity id="DD3E9B06921B0D1E164C1CF8B0D5F920" box="[592,757,1761,1785]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="30" pageNumber="29" unit="mm" value="0.835" valueMax="0.85" valueMin="0.82">0.820.85 mm</quantity>
. Clypeus approximately 1.751.8× as wide as long, length
<quantity id="DD3E9B06921A0D1F15A81C17B03BF9FC" box="[436,539,1550,1574]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="31" pageNumber="30" unit="mm" value="0.61">0.61 mm</quantity>
, width
<quantity id="DD3E9B06921A0D1F166F1C17B137F9FC" box="[627,791,1550,1574]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.09" metricValueMin="1.06" pageId="31" pageNumber="30" unit="mm" value="1.075" valueMax="1.09" valueMin="1.06">1.061.09 mm</quantity>
. Malar area approximately 1.6× flagellar diameter or 0.730.75× basal mandibular width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, carina inter- rupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as a short lamellate tooth, upper margin of concavity with preoccipital lamella extending into upper third of concavity.
</paragraph>
<caption id="4EB9666B921A0D1F14BC1FA8B383FA3D" pageId="31" pageNumber="30" startId="31.[160,241,1457,1479]" targetBox="[171,1149,238,1432]" targetPageId="31" targetType="figure">
<paragraph id="1A7936E3921A0D1F14BC1FA8B383FA3D" blockId="31.[160,1168,1457,1508]" pageId="31" pageNumber="30">
<emphasis id="28B2EAF1921A0D1F14BC1FA8B319FA1F" bold="true" box="[160,313,1457,1479]" pageId="31" pageNumber="30">Figures 4446.</emphasis>
Worker of
<taxonomicName id="DDC64D60921A0D1F15A51FABB091FA1E" authorityName="Engel" authorityYear="2022" box="[441,689,1458,1479]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="species" species="tatacoensis" status="sp. nov.">
<emphasis id="28B2EAF1921A0D1F15A51FABB091FA1E" box="[441,689,1458,1479]" italics="true" pageId="31" pageNumber="30">Scaptotrigona tatacoensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A921A0D1F16A11FABB11FFA1E" box="[701,831,1457,1479]" pageId="31" pageNumber="30" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1921A0D1F17531FA8B14CFA1F" bold="true" box="[847,876,1457,1478]" pageId="31" pageNumber="30">44.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF1921A0D1F10391FA8B664FA1F" bold="true" box="[1061,1092,1457,1478]" pageId="31" pageNumber="30">45.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF1921A0D1F151D1FD7B33EFA3A" bold="true" box="[257,286,1486,1507]" pageId="31" pageNumber="30">46.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="1A7936E3921A0D2114CC1CD7B156FE20" blockId="31.[160,1168,1550,1766]" lastBlockId="33.[160,1168,225,1530]" lastPageId="33" lastPageNumber="32" pageId="31" pageNumber="30">
Integument of labiomaxillary complex yellow orange; labrum reddish orange; mandible reddish orange; clypeus orange, epistomal sulcus well delineated by dark brown to black; supraclypeal area orange to reddish orange; malar space orange; face below upper tangent of antennal toruli orange blending to reddish orange and red- dish brown on lower frons medially above antennal toruli, narrow strip of orange extending upward along inner and upper orbit, frons and vertex dark brown to black except typically small areas of orange anterior to median ocellus and lateral to lateral ocelli; ocellocular area dark brown except laterally where separated from upper orbit; posterior of head orange to reddish orange with strips of dark brown extending from ocellocular area to preoccipital carina, preoccipital carina black; gena and postgena or- ange to reddish orange; scape orange to reddish orange; pedicel and flagellum brown, with flagellum more orange ventrally. Mesosoma largely dark brown to black except orange to reddish orange on pronotum; pronotal lobe; mesoscutum except along ex- treme lateral border, posterior border, and with faint longitudinal strips from notauli to posterior border; mesoscutellum (except V-shaped incision); most of mesepister- num except large ventral area of dark brown; small area on metepisternum anteriorly; and lateral surface of propodeum centrally; tegula wholly yellow orange and semi-translucent; legs orange to reddish orange. Metasoma generally dark brown to black except anterior-facing surface of tergum I orange to reddish orange or reddish brown, dorsal-facing disc of tergum I posteriorly orange, sometimes with orange encompass- ing nearly entire dorsal surface; tergum II with broad areas of orange apically and narrow yellowish marginal zone; terga
<collectionCode id="7CD7AE2692240D21166D1BDBB0ACFE00" box="[625,652,450,473]" pageId="33" pageNumber="32">III</collectionCode>
<collectionCode id="7CD7AE2692240D2116861BDBB096FE00" box="[666,694,450,473]" pageId="33" pageNumber="32">IV</collectionCode>
with narrow yellow marginal zones ob- scured by tomentum; sterna largely orange to reddish brown.
</paragraph>
<caption id="4EB9666B92250D2014BC1F11B1D8FAE2" pageId="32" pageNumber="31" startId="32.[160,241,1288,1310]" targetBox="[180,1143,224,1274]" targetPageId="32" targetType="figure">
<paragraph id="1A7936E392250D2014BC1F11B1D8FAE2" blockId="32.[160,1168,1288,1339]" pageId="32" pageNumber="31">
<emphasis id="28B2EAF192250D2014BC1F11B31CFAC4" bold="true" box="[160,316,1288,1310]" pageId="32" pageNumber="31">Figures 4748.</emphasis>
Worker of
<taxonomicName id="DDC64D6092250D2015D91F10B09FFAC7" authorityName="Engel" authorityYear="2022" box="[453,703,1289,1310]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="tatacoensis" status="sp. nov.">
<emphasis id="28B2EAF192250D2015D91F10B09FFAC7" box="[453,703,1289,1310]" italics="true" pageId="32" pageNumber="31">Scaptotrigona tatacoensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92250D2016D11F10B171FAC7" box="[717,849,1288,1310]" pageId="32" pageNumber="31" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192250D20177A1F11B1A4FAC4" bold="true" box="[870,900,1288,1309]" pageId="32" pageNumber="31">47.</emphasis>
Dorsal, slightly oblique, view of head.
<emphasis id="28B2EAF192250D2015271F3CB37AFAE3" bold="true" box="[315,346,1317,1338]" pageId="32" pageNumber="31">48.</emphasis>
Prolateral surface of metatibia, metatarsus, and metapretarsus.
</paragraph>
</caption>
<paragraph id="1A7936E392240D2114CC181BB35FFB00" blockId="33.[160,1168,225,1530]" pageId="33" pageNumber="32">
As for
<taxonomicName id="DDC64D6092240D211502181BB34EFDC0" authorityName="Engel" authorityYear="2022" box="[286,366,513,537]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192240D211502181BB34EFDC0" box="[286,366,513,537]" italics="true" pageId="33" pageNumber="32">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white; metepisternum and lateral surface of propodeum obscured by dense, plumose, yellow setae; dorsal-facing surface of tergum I with largely simple setae, minute, de- cumbent, yellow, plumose setae sparse, typically in small areas laterally. Bristles of vertex erect, simple, yellow to fulvous, sometimes even dark fulvous; pronotal dorso- lateral margin near and anterior to pronotal lobe with long, arched, yellow to fulvous bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with largely fulvous bristles, sometimes intermixed with yellow or dark fulvous bristles; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elongate, erect bristles posteriorly, such bristles mi- nutely pectinate apically, discal bristles shorter, mesoscutellar bristles largely yellow fulvous to dark fulvous on disc, those on apical margin yellow to fulvous; mesepister- nal bristles as in
<taxonomicName id="DDC64D6092240D21157E199BB391FC40" authorityName="Engel" authorityYear="2022" box="[354,433,897,921]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192240D21157E199BB391FC40" box="[354,433,897,921]" italics="true" pageId="33" pageNumber="32">S. ederi</emphasis>
</taxonomicName>
except yellow to fulvous. Legs with abundant bristles, bristles typically yellow to fulvous except noticeably intermixed with fuscous to black bristles on prolateral surface of protibia apically, prolateral surface of probasitarsus, prolateral and marginal surfaces of mesotibia, prolateral surface of mesobasitarsus, metafemur apically, metatibia, along metabasitarsal margins; corbicular and marginal bristles black, marginal bristles intermixed with some yellow to fulvous bristles; prolateral surface of metabasitarsus with scatered fine, simple, yellow setae (noticeably thin- ner than marginal bristles), intermixed with some fine black setae; penicillum black; rastellum fulvous. Metasoma with numerous, prominent, suberect to subdecumbent, yellow bristles on terga
<collectionCode id="7CD7AE2692240D2115A31EBBB3FAFB60" box="[447,474,1186,1209]" pageId="33" pageNumber="32">III</collectionCode>
<collectionCode id="7CD7AE2692240D2115FB1EBBB3DAFB60" box="[487,506,1186,1209]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="33" pageNumber="32" type="Museum">V</collectionCode>
arising from tomentum, as well as similar bristles scat- tered on tergum II.
</paragraph>
<paragraph id="1A7936E392240D2114CC1EFBB30FFB20" blockId="33.[160,1168,225,1530]" box="[208,303,1250,1276]" pageId="33" pageNumber="32">
<specimenCount id="0CC0FD6A92240D2114CC1EFBB2C4FB25" box="[208,228,1250,1276]" count="1" pageId="33" pageNumber="32" type="female"></specimenCount>
:
<emphasis id="28B2EAF192240D2114ED1EFBB308FB20" box="[241,296,1250,1273]" italics="true" pageId="33" pageNumber="32">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392240D2114CC1F1BB30FFAC0" blockId="33.[160,1168,225,1530]" box="[208,303,1282,1308]" pageId="33" pageNumber="32">
<specimenCount id="0CC0FD6A92240D2114CC1F1BB2C4FAC5" box="[208,228,1282,1308]" count="1" pageId="33" pageNumber="32" type="male"></specimenCount>
:
<emphasis id="28B2EAF192240D2114ED1F1BB308FAC0" box="[241,296,1282,1305]" italics="true" pageId="33" pageNumber="32">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892240D2114CC1F38B156FA40" pageId="33" pageNumber="32" type="materials_examined">
<paragraph id="1A7936E392240D2114CC1F38B127FA80" blockId="33.[160,1168,225,1530]" pageId="33" pageNumber="32">
<materialsCitation id="AAAE3CBE92240D2114CC1F38B127FA80" collectingDate="2006-01-04" collectingDateMax="2006-01-08" collectingDateMin="2006-01-04" collectionCode="SEMC" collectorName="Desierto de la Tatacoa &amp; J. Florez" country="Colombia" latitude="3.2166667" location="Desierto de la Tatacoa" longLatPrecision="1307" longitude="-75.166664" municipality="Villavieja" pageId="33" pageNumber="32" specimenCount="1" stateProvince="Huila" typeStatus="holotype">
<typeStatus id="C57D884192240D2114CC1F38B368FAE0" box="[208,328,1313,1337]" pageId="33" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392240D2115711F38B3C3FAE0" box="[365,483,1313,1337]" name="Colombia" pageId="33" pageNumber="32">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192240D2115F21F38B013FAE0" box="[494,563,1313,1337]" country="Colombia" name="Huila" pageId="33" pageNumber="32">Huila</collectingRegion>
,
<collectingMunicipality id="FA1DAC9992240D2116221F38B08FFAE0" box="[574,687,1313,1337]" pageId="33" pageNumber="32">Villavieja</collectingMunicipality>
,
<collectorName id="B733533592240D2116A61F38B19FFAE0" box="[698,959,1313,1337]" pageId="33" pageNumber="32">
<location id="1F19603892240D2116A61F38B19FFAE0" LSID="urn:lsid:plazi:treatment:926F87F5921B0D2115961F18B18AFA20:1F19603892240D2116A61F38B19FFAE0" box="[698,959,1313,1337]" country="Colombia" latitude="3.2166667" longLatPrecision="1307" longitude="-75.166664" municipality="Villavieja" name="Desierto de la Tatacoa" pageId="33" pageNumber="32" stateProvince="Huila">Desierto de la Tatacoa</location>
</collectorName>
,
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,
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,
<date id="6E78102392240D2114BC1F5BB370FA80" box="[160,336,1345,1369]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">
<collectingDate id="7E3CE9CB92240D2114BC1F5BB370FA80" box="[160,336,1345,1369]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">Enero 48 2006</collectingDate>
</date>
[
<date id="6E78102392240D2115421F58B00CFA80" box="[350,556,1345,1370]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">
<collectingDate id="7E3CE9CB92240D2115421F58B00CFA80" box="[350,556,1345,1370]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">48 January 2006</collectingDate>
</date>
],
<collectorName id="B733533592240D2116211F5BB0BDFA80" box="[573,669,1345,1369]" pageId="33" pageNumber="32">J. Florez</collectorName>
(
<collectionCode id="7CD7AE2692240D2116B01F58B0DCFA80" box="[684,764,1345,1369]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="33" pageNumber="32" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392240D2114CC1F78B156FA40" blockId="33.[160,1168,225,1530]" pageId="33" pageNumber="32">
<materialsCitation id="AAAE3CBE92240D2114CC1F78B153FA40" collectingDate="2006-01-04" collectingDateMax="2006-01-08" collectingDateMin="2006-01-04" collectionCode="SEMC" collectorName="Desierto de la Tatacoa &amp; J. Florez" country="Colombia" latitude="3.2166667" location="Desierto de la Tatacoa" longLatPrecision="1307" longitude="-75.166664" municipality="Villavieja" pageId="33" pageNumber="32" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<typeStatus id="C57D884192240D2114CC1F78B36EFAA0" box="[208,334,1377,1401]" pageId="33" pageNumber="32" type="paratype">PARATYPES</typeStatus>
: 6⚲⚲,
<collectingCountry id="62D1767392240D2115BC1F78B036FAA0" box="[416,534,1377,1401]" name="Colombia" pageId="33" pageNumber="32">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192240D2116361F78B04FFAA0" box="[554,623,1377,1401]" country="Colombia" name="Huila" pageId="33" pageNumber="32">Huila</collectingRegion>
,
<collectingMunicipality id="FA1DAC9992240D21169F1F78B0D4FAA0" box="[643,756,1377,1401]" pageId="33" pageNumber="32">Villavieja</collectingMunicipality>
,
<collectorName id="B733533592240D21171B1F78B606FAA0" box="[775,1062,1377,1401]" pageId="33" pageNumber="32">
<location id="1F19603892240D21171B1F78B606FAA0" LSID="urn:lsid:plazi:treatment:926F87F5921B0D2115961F18B18AFA20:1F19603892240D21171B1F78B606FAA0" box="[775,1062,1377,1401]" country="Colombia" latitude="3.2166667" longLatPrecision="1307" longitude="-75.166664" municipality="Villavieja" name="Desierto de la Tatacoa" pageId="33" pageNumber="32" stateProvince="Huila">Desierto de la Tatacoa</location>
</collectorName>
,
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,
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,
<date id="6E78102392240D2115131F9BB39FFA40" box="[271,447,1409,1433]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">
<collectingDate id="7E3CE9CB92240D2115131F9BB39FFA40" box="[271,447,1409,1433]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">Enero 48 2006</collectingDate>
</date>
[
<date id="6E78102392240D2115D11F98B0BBFA40" box="[461,667,1409,1434]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">
<collectingDate id="7E3CE9CB92240D2115D11F98B0BBFA40" box="[461,667,1409,1434]" pageId="33" pageNumber="32" value="2006-01-04" valueMax="2006-01-08" valueMin="2006-01-04">48 January 2006</collectingDate>
</date>
],
<collectorName id="B733533592240D2116B01F9BB12CFA40" box="[684,780,1409,1433]" pageId="33" pageNumber="32">J. Florez</collectorName>
(
<collectionCode id="7CD7AE2692240D2117071F98B14BFA40" box="[795,875,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="33" pageNumber="32" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892240D2114CC1FB8B18AFA20" pageId="33" pageNumber="32" type="etymology">
<paragraph id="1A7936E392240D2114CC1FB8B18AFA20" blockId="33.[160,1168,225,1530]" pageId="33" pageNumber="32">
<materialsCitation id="AAAE3CBE92240D2114CC1FB8B186FA20" country="Colombia" county="Latin" location="Tatacoa Desert" municipality="Tatacoa" pageId="33" pageNumber="32" specimenCount="1" stateProvince="Huila">
ETYMOLOGY: The specific epithet is taken from the
<typeStatus id="C57D884192240D2117051FBBB16DFA63" box="[793,845,1442,1466]" pageId="33" pageNumber="32">type</typeStatus>
locality, the
<location id="1F19603892240D2117FF1FBBB6B0FA60" LSID="urn:lsid:plazi:treatment:926F87F5921B0D2115961F18B18AFA20:1F19603892240D2117FF1FBBB6B0FA60" box="[995,1168,1442,1465]" country="Colombia" county="Latin" municipality="Tatacoa" name="Tatacoa Desert" pageId="33" pageNumber="32" stateProvince="Huila">Tatacoa Desert</location>
of
<collectingRegion id="D802F80192240D2114A21FD8B3B9FA00" box="[190,409,1473,1497]" country="Colombia" name="Huila" pageId="33" pageNumber="32">Huila Department</collectingRegion>
,
<collectingCountry id="62D1767392240D2115B81FD8B03AFA00" box="[420,538,1473,1497]" name="Colombia" pageId="33" pageNumber="32">Colombia</collectingCountry>
.
<location id="1F19603892240D2116301FD8B079FA00" LSID="urn:lsid:plazi:treatment:926F87F5921B0D2115961F18B18AFA20:1F19603892240D2116301FD8B079FA00" box="[556,601,1473,1497]" country="Colombia" county="Latin" municipality="Tatacoa" name="The" pageId="33" pageNumber="32" stateProvince="Huila">The</location>
name is formed from
<collectingMunicipality id="FA1DAC9992240D2117431FDBB198FA00" box="[863,952,1474,1497]" pageId="33" pageNumber="32">Tatacoa</collectingMunicipality>
and the
<collectingCounty id="F3184E6F92240D2110021FD8B67CFA00" box="[1054,1116,1473,1497]" pageId="33" pageNumber="32">Latin</collectingCounty>
suf- fix
<emphasis id="28B2EAF192240D2114CE1FFBB327FA20" box="[210,263,1506,1529]" italics="true" pageId="33" pageNumber="32">ēnsis</emphasis>
, which is added to a toponym to indicate “from” or “of”
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,262 @@
<document id="6516C49708C6E69D7A30CBE20A5CFC1E" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F592200D2815811E78B2DEFA00" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona illescasi Engel 2022, new species" docType="treatment" docUuid="5C5ED6EA-EAEF-4564-9D13-DCBF02803F4F" docUuidSource="ZooBank" docVersion="1" lastPageNumber="39" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="36" updateTime="1722456085954" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:identifier id="396B2ED3C6897FA2BFE0065434C4233F" type="DOI">10.17161/jom.i110.17001</mods:identifier>
<mods:identifier id="61BADDB53D4E24B86E5BE4DB736737A1" type="ISSN">2325-4467</mods:identifier>
<mods:identifier id="DEF0922132DA15FC95CBF05CDED26C93" type="ZooBank">urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB</mods:identifier>
</mods:mods>
<treatment id="926F87F592200D2815811E78B2DEFA00" LSID="urn:lsid:zoobank.org:act:5C5ED6EA-EAEF-4564-9D13-DCBF02803F4F" httpUri="http://treatment.plazi.org/id/926F87F592200D2815811E78B2DEFA00" lastPageId="40" lastPageNumber="39" pageId="37" pageNumber="36">
<subSubSection id="52DC656892200D2515811E78B661FB40" pageId="37" pageNumber="36" type="nomenclature">
<paragraph id="1A7936E392200D2515811E78B1B3FBA0" blockId="37.[413,915,1121,1145]" box="[413,915,1121,1145]" pageId="37" pageNumber="36">
<heading id="4131818F92200D2515811E78B1B3FBA0" box="[413,915,1121,1145]" centered="true" fontSize="10" level="2" pageId="37" pageNumber="36" reason="2">
<taxonomicName id="DDC64D6092200D2515811E78B0DBFBA0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[413,763,1121,1145]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="illescasi" status="sp. nov.">
<emphasis id="28B2EAF192200D2515811E78B08FFBA0" bold="true" box="[413,687,1121,1145]" italics="true" pageId="37" pageNumber="36">Scaptotrigona illescasi</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92200D25171A1E7BB1B3FBA0" box="[774,915,1121,1145]" pageId="37" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392200D2514F31E9AB661FB40" blockId="37.[239,1089,1155,1209]" box="[239,1089,1155,1177]" pageId="37" pageNumber="36">
ZooBank:
<uri id="6E573AE192200D2515471E9AB661FB40" box="[347,1089,1155,1177]" pageId="37" pageNumber="36">
urn:lsid:zoobank.org:act:
<uuid id="6E600C3692200D2516521E9AB661FB40" box="[590,1089,1155,1177]" pageId="37" pageNumber="36">5C5ED6EA-EAEF-4564-9D13-DCBF02803F4F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892200D2516531EB8B0C2FB60" box="[591,738,1185,1209]" pageId="37" pageNumber="36" type="description">
<paragraph id="1A7936E392200D2516531EB8B0C2FB60" blockId="37.[239,1089,1155,1209]" box="[591,738,1185,1209]" pageId="37" pageNumber="36">
(
<figureCitation id="82FD2A6692200D25164B1EB8B0FAFB60" box="[599,730,1185,1209]" captionStart-0="Figures 5152" captionStart-1="Figures 5356" captionStartId-0="38.[160,241,1366,1388]" captionStartId-1="39.[160,241,1395,1417]" captionTargetBox-0="[173,1162,239,1337]" captionTargetBox-1="[160,1168,223,1390]" captionTargetId-0="figure-130@38.[160,1168,224,1359]" captionTargetId-1="figure-105@39.[160,1168,223,1390]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionText-0="Figures 5152. Workers of Scaptotrigona illescasi, new species. 51. Lateral habitus, darker indi- vidual. 52. Lateral habitus, lighter individual." captionText-1="Figures 5356. Workers of Scaptotrigona illescasi, new species. 53. Facial view, darker individual. 54. Facial view, lighter individual. 55. Mesosomal dorsum, darker individual. 56. Mesosomal dorsum, lighter individual." pageId="37" pageNumber="36">Figs. 5156</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892200D2514CC1EF8B01EFA40" pageId="37" pageNumber="36" type="diagnosis">
<paragraph id="1A7936E392200D2514CC1EF8B01EFA40" blockId="37.[160,1169,1249,1753]" pageId="37" pageNumber="36">
DIAGNOSIS: This western Ecuadorian species is comparatively dark, in this respect similar to
<taxonomicName id="DDC64D6092200D2515081F1BB343FAC0" authorityName="Engel" authorityYear="2022" box="[276,355,1281,1305]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192200D2515081F1BB343FAC0" box="[276,355,1281,1305]" italics="true" pageId="37" pageNumber="36">S. ederi</emphasis>
</taxonomicName>
, but possessing a distinct genal tooth and yellow to dark fulvous bris- tles along the entire mesoscutal margin, although sometimes with a few black bristles intermixed. The tegular bristles are yellow, the mesepisternum is marked with areas of yellow to reddish orange, and the metabasitarsal bristles are predominantly black, including on the prolateral surface.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892200D2814CC1FB8B30FFB60" lastPageId="40" lastPageNumber="39" pageId="37" pageNumber="36" type="description">
<paragraph id="1A7936E392200D2614CC1FB8B165FA33" blockId="37.[160,1169,1249,1753]" lastBlockId="38.[160,1168,1458,1771]" lastPageId="38" lastPageNumber="37" pageId="37" pageNumber="36">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092200D2516331FBBB05DFA60" authorityName="Engel" authorityYear="2022" box="[559,637,1441,1465]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192200D2516331FBBB05DFA60" box="[559,637,1441,1465]" italics="true" pageId="37" pageNumber="36">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192200D2516971FB8B0D9FA60" box="[651,761,1441,1465]" italics="true" pageId="37" pageNumber="36">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately 5.86.0 mm, forewing length (to base of humeral sclerite)
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. Head wider than long, width
<quantity id="DD3E9B0692200D2516F31FF8B1B0FA20" box="[751,912,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="2.3899999999999997" metricValueMax="2.42" metricValueMin="2.36" pageId="37" pageNumber="36" unit="mm" value="2.3899999999999997" valueMax="2.42" valueMin="2.36">2.362.42 mm</quantity>
, length
<quantity id="DD3E9B0692200D2517F61FF8B6ABFA20" box="[1002,1163,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.955" metricValueMax="1.97" metricValueMin="1.94" pageId="37" pageNumber="36" unit="mm" value="1.955" valueMax="1.97" valueMin="1.94">1.941.97 mm</quantity>
; compound eye length
<quantity id="DD3E9B0692200D2515B61C18B06DF9C0" box="[426,589,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.42" metricValueMin="1.39" pageId="37" pageNumber="36" unit="mm" value="1.4049999999999998" valueMax="1.42" valueMin="1.39">1.391.42 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B0692200D2517811C18B660F9C0" box="[925,1088,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.55" metricValueMin="1.52" pageId="37" pageNumber="36" unit="mm" value="1.535" valueMax="1.55" valueMin="1.52">1.521.55 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B0692200D2515881C38B018F9E0" box="[404,568,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="1.435" metricValueMax="1.45" metricValueMin="1.42" pageId="37" pageNumber="36" unit="mm" value="1.435" valueMax="1.45" valueMin="1.42">1.421.45 mm</quantity>
. Scape length
<quantity id="DD3E9B0692200D2516FB1C38B16DF9E0" box="[743,845,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="37" pageNumber="36" unit="mm" value="0.91">0.91 mm</quantity>
, about as long as torulocel- lar distance, torulocellar distance
<quantity id="DD3E9B0692200D25162C1C58B0F4F980" box="[560,724,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="9.1" metricValueMin="8.5" pageId="37" pageNumber="36" unit="mm" value="0.88" valueMax="0.91" valueMin="0.85">0.850.91 mm</quantity>
. Clypeus approximately 1.751.8× as wide as long, length
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, width
<quantity id="DD3E9B0692200D2516791C78B12BF9A0" box="[613,779,1633,1657]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.09" metricValueMin="1.06" pageId="37" pageNumber="36" unit="mm" value="1.075" valueMax="1.09" valueMin="1.06">1.061.09 mm</quantity>
. Malar area approximately 1.6× flagellar diameter or 0.730.75× basal mandibular width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, carina inter- rupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as short, lamellate tooth; upper margin with lamella extended ventrally about half way into concavity.
</paragraph>
<caption id="4EB9666B92230D2614BC1F4FB0A8FA50" pageId="38" pageNumber="37" startId="38.[160,241,1366,1388]" targetBox="[173,1162,239,1337]" targetPageId="38" targetType="figure">
<paragraph id="1A7936E392230D2614BC1F4FB0A8FA50" blockId="38.[160,1168,1366,1417]" pageId="38" pageNumber="37">
<emphasis id="28B2EAF192230D2614BC1F4FB31AFAB2" bold="true" box="[160,314,1366,1388]" pageId="38" pageNumber="37">Figures 5152.</emphasis>
Workers of
<taxonomicName id="DDC64D6092230D2615DA1F4FB0BEFAB2" authorityName="Engel" authorityYear="2022" box="[454,670,1366,1387]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="illescasi" status="sp. nov.">
<emphasis id="28B2EAF192230D2615DA1F4FB0BEFAB2" box="[454,670,1366,1387]" italics="true" pageId="38" pageNumber="37">Scaptotrigona illescasi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92230D2616B71F4FB10EFAB5" box="[683,814,1366,1388]" pageId="38" pageNumber="37" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192230D2617231F4FB17DFAB2" bold="true" box="[831,861,1366,1387]" pageId="38" pageNumber="37">51.</emphasis>
Lateral habitus, darker indi- vidual.
<emphasis id="28B2EAF192230D2614EB1F6AB335FA51" bold="true" box="[247,277,1395,1416]" pageId="38" pageNumber="37">52.</emphasis>
Lateral habitus, lighter individual.
</paragraph>
</caption>
<paragraph id="1A7936E392230D2814CC1FEAB35DFDC0" blockId="38.[160,1168,1458,1771]" lastBlockId="40.[160,1175,225,1497]" lastPageId="40" lastPageNumber="39" pageId="38" pageNumber="37">
Integument of labiomaxillary complex yellow orange; labrum orange; mandible reddish orange; clypeus largely yellow orange to orange with brown along apical mar- gin and sometimes with brownish stain laterally and at apicolateral corners, epistomal sulcus well delineated by brownish orange to black; supraclypeal area orange with stain of brown at epistomal sulcus; malar space often orange to reddish orange; face below tangent of antennal toruli orange, albeit blending to more reddish orange to- ward antennal toruli, lower frons orange to reddish orange, sometimes largely brown to dark brown except around antennal toruli; scape orange to brownish orange, some- times with brown dorsally; pedicel and flagellum brown to dark brown, flagellum orange underneath; vertex dark brown to black; posterior of head dark brown to black; upper third of gena dark brown to black, remainder of gena orange; postgena orange to yellow orange. Mesosoma black except pronotum orange to brown, mesoscutellum orange but sometimes with axilla black, mesepisternum reddish orange posteriorly but sometimes reddish orange anteriorly and orange posteriorly, sometimes propodeum orange with brown centrally on lateral surface; tegula yellow orange and semi-trans- lucent; legs largely yellow orange to orange except dark brown on femora apically, on prolateral surfaces of pro- and mesotibiae, prolateral surface of pro- and mesoba- sitarsus, prolateral surface of metatibia except extreme base of fundus and sometimes superior distal angle, and prolateral surface of metabasitarsus except margins, some- times dark brown areas reddish orange to reddish brown. Wing membranes lightly parchment colored; veins orange to orange brown. Metasoma generally dark brown to black except anterior-facing surface of tergum I orange to orange brown, dorsal-fac- ing disc of tergum I posteriorly coppery orange, tergum II posteriorly coppery orange, with yellow marginal zone; terga
<collectionCode id="7CD7AE26922D0D2816281BDBB06FFE00" box="[564,591,450,473]" pageId="40" pageNumber="39">III</collectionCode>
<collectionCode id="7CD7AE26922D0D2816401BDBB058FE00" box="[604,632,450,473]" pageId="40" pageNumber="39">IV</collectionCode>
with yellow apical marginal zones, typically obscured by tomentum; sterna dark brown basally and yellow to yellow brown api- cally in setal areas.
</paragraph>
<caption id="4EB9666B92220D2714BC1F6AB3E1FA1A" pageId="39" pageNumber="38" startId="39.[160,241,1395,1417]" targetBox="[160,1168,223,1390]" targetPageId="39" targetType="figure">
<paragraph id="1A7936E392220D2714BC1F6AB3E1FA1A" blockId="39.[160,1175,1395,1475]" pageId="39" pageNumber="38">
<emphasis id="28B2EAF192220D2714BC1F6AB318FA51" bold="true" box="[160,312,1395,1417]" pageId="39" pageNumber="38">Figures 5356.</emphasis>
Workers of
<taxonomicName id="DDC64D6092220D2715A01F6DB0B3FA50" authorityName="Engel" authorityYear="2022" box="[444,659,1396,1417]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="illescasi" status="sp. nov.">
<emphasis id="28B2EAF192220D2715A01F6DB0B3FA50" box="[444,659,1396,1417]" italics="true" pageId="39" pageNumber="38">Scaptotrigona illescasi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92220D2716821F6DB13EFA53" box="[670,798,1396,1418]" pageId="39" pageNumber="38" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192220D2717371F6AB169FA51" bold="true" box="[811,841,1395,1416]" pageId="39" pageNumber="38">53.</emphasis>
Facial view, darker individual.
<emphasis id="28B2EAF192220D2714BC1F89B29EFA7C" bold="true" box="[160,190,1424,1445]" pageId="39" pageNumber="38">54.</emphasis>
Facial view, lighter individual.
<emphasis id="28B2EAF192220D2716001F88B01AFA7F" bold="true" box="[540,570,1425,1446]" pageId="39" pageNumber="38">55.</emphasis>
Mesosomal dorsum, darker individual.
<emphasis id="28B2EAF192220D2717EE1F89B631FA7C" bold="true" box="[1010,1041,1424,1445]" pageId="39" pageNumber="38">56.</emphasis>
Mesosomal dorsum, lighter individual.
</paragraph>
</caption>
<paragraph id="1A7936E3922D0D2814CC183BB1F9FBA0" blockId="40.[160,1175,225,1497]" pageId="40" pageNumber="39">
Areas of short, white, plumose setae in
<taxonomicName id="DDC64D60922D0D281685183BB0C9FDE0" authorityName="Engel" authorityYear="2022" box="[665,745,545,569]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922D0D281685183BB0C9FDE0" box="[665,745,545,569]" italics="true" pageId="40" pageNumber="39">S. ederi</emphasis>
</taxonomicName>
instead yellow. Vertex with numer- ous erect, simple, yellow to dark fulvous bristles, sometimes with a few short black bristles laterally near upper orbit in darker forms; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow to fulvous bristles; mesoscutum anterior and lateral borders with largely yellow to dark fulvous bristles sometimes intermixed with a few black bristles, typically anterolaterally; tegula with suberect yellow bristles ante- riorly; mesoscutellum with abundant, elongate, erect yellow to fulvous bristles poste- riorly; mesepisternum with scatered, erect to subdecumbent yellow fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abun- dant, suberect to subdecumbent fulvous to dark fulvous bristles extending ventrally from pronotal lobe; lateral surface of propodeum with scatered, long, erect, yellow bristles emerging amid dense, appressed, yellow, plumose setae. Legs with abundant yellow to fulvous bristles except intermixed with dark fulvous to black bristles on protibia apically, prolateral surface of pro- and mesobasitarsi, and metafemur apically; metatibia with almost entirely black bristles; metabasitarsus with erect black bristles on margins intermixed with some yellow to fulvous bristles, prolateral surface with scatered fine, dark fulvous to black setae amid otherwise yellow to fulvous setae. Metasoma with numerous, prominent, suberect to subdecumbent, yellow bristles on terga
<collectionCode id="7CD7AE26922D0D2814FF1E7BB2DEFBA0" box="[227,254,1122,1145]" pageId="40" pageNumber="39">III</collectionCode>
<collectionCode id="7CD7AE26922D0D2815171E7BB33EFBA0" box="[267,286,1122,1145]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="39" type="Museum">V</collectionCode>
arising amid tomentum, such bristles sparser on tergum II.
</paragraph>
<paragraph id="1A7936E3922D0D2814CC1E9BB30FFB40" blockId="40.[160,1175,225,1497]" box="[208,303,1154,1180]" pageId="40" pageNumber="39">
<specimenCount id="0CC0FD6A922D0D2814CC1E9BB2C4FB45" box="[208,228,1154,1180]" count="1" pageId="40" pageNumber="39" type="female"></specimenCount>
:
<emphasis id="28B2EAF1922D0D2814ED1E9BB308FB40" box="[241,296,1154,1177]" italics="true" pageId="40" pageNumber="39">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E3922D0D2814CC1EBBB30FFB60" blockId="40.[160,1175,225,1497]" box="[208,303,1186,1212]" pageId="40" pageNumber="39">
<specimenCount id="0CC0FD6A922D0D2814CC1EBBB2C4FB65" box="[208,228,1186,1212]" count="1" pageId="40" pageNumber="39" type="male"></specimenCount>
:
<emphasis id="28B2EAF1922D0D2814ED1EBBB308FB60" box="[241,296,1186,1209]" italics="true" pageId="40" pageNumber="39">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922D0D2814CC1ED8B6A3FAA0" pageId="40" pageNumber="39" type="materials_examined">
<paragraph id="1A7936E3922D0D2814CC1ED8B38DFAC0" blockId="40.[160,1175,225,1497]" pageId="40" pageNumber="39">
<materialsCitation id="AAAE3CBE922D0D2814CC1ED8B38DFAC0" collectingDate="1956-10-09" collectionCode="SEMC" country="Ecuador" location="Foerster" pageId="40" pageNumber="39" specimenCount="1" stateProvince="Esmeraldas" typeStatus="holotype">
<typeStatus id="C57D8841922D0D2814CC1ED8B368FB00" box="[208,328,1217,1241]" pageId="40" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D17673922D0D28156F1ED8B3F9FB00" box="[371,473,1217,1241]" name="Ecuador" pageId="40" pageNumber="39">Ecuador</collectingCountry>
: Prov. San Mateo,
<collectingRegion id="D802F801922D0D2816A31ED8B167FB00" box="[703,839,1217,1241]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
[Province
<collectingRegion id="D802F801922D0D2817D01ED8B677FB00" box="[972,1111,1217,1241]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
, San Mateo, approximately
<quantity id="DD3E9B06922D0D2815AF1EF8B02AFB20" box="[435,522,1249,1273]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="40" pageNumber="39" unit="mi" value="2.0">2 miles</quantity>
south of the municipality of
<collectingRegion id="D802F801922D0D28176B1EF8B624FB20" box="[887,1028,1249,1273]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
],
<date id="6E781023922D0D2810061EF8B2F8FAC0" pageId="40" pageNumber="39" value="1956-10-09">
<collectingDate id="7E3CE9CB922D0D2810061EF8B2F8FAC0" pageId="40" pageNumber="39" value="1956-10-09">9 October 1956</collectingDate>
</date>
,
<location id="1F196038922D0D2814FF1F1BB363FAC0" LSID="urn:lsid:plazi:treatment:926F87F592200D2815811E78B2DEFA00:1F196038922D0D2814FF1F1BB363FAC0" box="[227,323,1282,1305]" country="Ecuador" name="Foerster" pageId="40" pageNumber="39" stateProvince="Esmeraldas">Foerster</location>
(
<collectionCode id="7CD7AE26922D0D28154D1F18B381FAC0" box="[337,417,1281,1305]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="40" pageNumber="39" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E3922D0D2814CC1F38B6A3FAA0" blockId="40.[160,1175,225,1497]" pageId="40" pageNumber="39">
<materialsCitation id="AAAE3CBE922D0D2814CC1F38B3CBFAA0" collectingDate="1956-10-09" collectionCode="SEMC" country="Ecuador" location="Foerster" pageId="40" pageNumber="39" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<typeStatus id="C57D8841922D0D2814CC1F38B36EFAE0" box="[208,334,1313,1337]" pageId="40" pageNumber="39" type="paratype">PARATYPES</typeStatus>
: 23⚲⚲,
<collectingCountry id="62D17673922D0D2815B81F38B02AFAE0" box="[420,522,1313,1337]" name="Ecuador" pageId="40" pageNumber="39">Ecuador</collectingCountry>
: Prov. San Mateo,
<collectingRegion id="D802F801922D0D2816EF1F38B15BFAE0" box="[755,891,1313,1337]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
[Province
<collectingRegion id="D802F801922D0D28101D1F38B6ACFAE0" box="[1025,1164,1313,1337]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
, San Mateo, approximately
<quantity id="DD3E9B06922D0D2815C01F58B00EFA80" box="[476,558,1345,1369]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="40" pageNumber="39" unit="mi" value="2.0">2 miles</quantity>
south of the municipality of
<collectingRegion id="D802F801922D0D28179C1F58B62DFA80" box="[896,1037,1345,1369]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
],
<date id="6E781023922D0D2810021F58B2F8FAA0" pageId="40" pageNumber="39" value="1956-10-09">
<collectingDate id="7E3CE9CB922D0D2810021F58B2F8FAA0" pageId="40" pageNumber="39" value="1956-10-09">9 October 1956</collectingDate>
</date>
,
<location id="1F196038922D0D2814FF1F7BB363FAA0" LSID="urn:lsid:plazi:treatment:926F87F592200D2815811E78B2DEFA00:1F196038922D0D2814FF1F7BB363FAA0" box="[227,323,1378,1401]" country="Ecuador" name="Foerster" pageId="40" pageNumber="39" stateProvince="Esmeraldas">Foerster</location>
(
<collectionCode id="7CD7AE26922D0D28154D1F78B380FAA0" box="[337,416,1377,1401]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="40" pageNumber="39" type="Museum">SEMC</collectionCode>
); 13⚲⚲
</materialsCitation>
,
<materialsCitation id="AAAE3CBE922D0D2815E41F78B65FFAA0" box="[504,1151,1377,1401]" collectingDate="1956-09-19" collectionCode="SEMC" country="Ecuador" location="San Mateo" pageId="40" pageNumber="39" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<collectingCountry id="62D17673922D0D2815E41F78B07EFAA0" box="[504,606,1377,1401]" name="Ecuador" pageId="40" pageNumber="39">Ecuador</collectingCountry>
:
<collectingRegion id="D802F801922D0D2816751F78B0D4FAA0" box="[617,756,1377,1401]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
,
<location id="1F196038922D0D2816E21F78B15DFAA0" LSID="urn:lsid:plazi:treatment:926F87F592200D2815811E78B2DEFA00:1F196038922D0D2816E21F78B15DFAA0" box="[766,893,1377,1401]" country="Ecuador" name="San Mateo" pageId="40" pageNumber="39" stateProvince="Esmeraldas">San Mateo</location>
,
<date id="6E781023922D0D2817941F78B638FAA0" box="[904,1048,1377,1401]" pageId="40" pageNumber="39" value="1956-09-19">
<collectingDate id="7E3CE9CB922D0D2817941F78B638FAA0" box="[904,1048,1377,1401]" pageId="40" pageNumber="39" value="1956-09-19">19 Sept 1956</collectingDate>
</date>
(
<collectionCode id="7CD7AE26922D0D28103B1F78B657FAA0" box="[1063,1143,1377,1401]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="40" pageNumber="39" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922D0D2814CC1F98B2DEFA00" pageId="40" pageNumber="39" type="etymology">
<paragraph id="1A7936E3922D0D2814CC1F98B2DEFA00" blockId="40.[160,1175,225,1497]" pageId="40" pageNumber="39">
<materialsCitation id="AAAE3CBE922D0D2814CC1F98B2DAFA00" collectorName="de Illescas" location="Alonso de Illescas" pageId="40" pageNumber="39" specimenCount="1" stateProvince="Esmeraldas">
ETYMOLOGY: The specific epithet honors
<location id="1F196038922D0D2816BC1F98B152FA40" LSID="urn:lsid:plazi:treatment:926F87F592200D2815811E78B2DEFA00:1F196038922D0D2816BC1F98B152FA40" box="[672,882,1409,1433]" name="Alonso de Illescas" pageId="40" pageNumber="39" stateProvince="Esmeraldas">
Alonso
<collectorName id="B7335335922D0D2816E61F98B152FA40" box="[762,882,1409,1433]" pageId="40" pageNumber="39">de Illescas</collectorName>
</location>
(
<emphasis id="28B2EAF1922D0D28179D1F98B1E5FA40" box="[897,965,1409,1433]" italics="true" pageId="40" pageNumber="39">floruit</emphasis>
1528circa 1596), leader of the African Maroons in the area of
<collectingRegion id="D802F801922D0D2816B51FB8B114FA60" box="[681,820,1441,1465]" country="Ecuador" name="Esmeraldas" pageId="40" pageNumber="39">Esmeraldas</collectingRegion>
, which encompasses the
<typeStatus id="C57D8841922D0D2810401FBBB6B0FA63" box="[1116,1168,1442,1466]" pageId="40" pageNumber="39">type</typeStatus>
locality
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,203 @@
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<mods:title id="38709AAD5F4EF9D0D9153F9F3AF484FC">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="5E2AACA2403ED404E46B5BC219D46BDB">Engel, Michael S.</mods:namePart>
<mods:affiliation id="542A6D879044D1AA61A34FFF77F4D5C6">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<subSubSection id="52DC656892210D2415851BD8B662FE20" pageId="36" pageNumber="35" type="nomenclature">
<paragraph id="1A7936E392210D2415851BD8B1B7FE00" blockId="36.[409,919,449,473]" box="[409,919,449,473]" pageId="36" pageNumber="35">
<heading id="4131818F92210D2415851BD8B1B7FE00" box="[409,919,449,473]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="35" reason="2">
<taxonomicName id="DDC64D6092210D2415851BD8B120FE00" authority="Engel" authorityName="Engel" authorityYear="2022" box="[409,768,449,473]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="caduceus" status="sp. nov.">
<emphasis id="28B2EAF192210D2415851BD8B093FE00" bold="true" box="[409,691,449,473]" italics="true" pageId="36" pageNumber="35">Scaptotrigona caduceus</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92210D2417161BDBB1B7FE00" box="[778,919,449,473]" pageId="36" pageNumber="35" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="52DC656892210D2416741818B0E8FDC0" box="[616,712,513,537]" pageId="36" pageNumber="35" type="description">
<paragraph id="1A7936E392210D2416741818B0E8FDC0" blockId="36.[238,1090,483,537]" box="[616,712,513,537]" pageId="36" pageNumber="35">
(
<figureCitation id="82FD2A6692210D24166D1818B0E0FDC0" box="[625,704,513,537]" captionStart="Figures 4950" captionStartId="34.[160,241,1430,1452]" captionTargetBox="[175,1155,233,1415]" captionTargetId="figure-116@34.[160,1168,224,1423]" captionTargetPageId="34" captionText="Figures 4950. Workers of Scaptotrigona Moure. 49. Lateral habitus of Scaptotrigona macarenensis, new species. 50. Lateral habitus of S. caduceus, new species." pageId="36" pageNumber="35">Fig. 50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892210D2414CC1858B654FD60" pageId="36" pageNumber="35" type="diagnosis">
<paragraph id="1A7936E392210D2414CC1858B654FD60" blockId="36.[160,1169,577,1786]" pageId="36" pageNumber="35">
DIAGNOSIS: This is a species easily confused with
<taxonomicName id="DDC64D6092210D24170D185BB141FD80" authorityName="Engel" authorityYear="2022" box="[785,865,577,601]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192210D24170D185BB141FD80" box="[785,865,577,601]" italics="true" pageId="36" pageNumber="35">S. ederi</emphasis>
</taxonomicName>
, but occurring to the East of the Andes. A distinct genal tooth is present and among such species it is perhaps most similar to
<taxonomicName id="DDC64D6092210D241544189BB3E7FD40" authorityName="Engel" authorityYear="2022" box="[344,455,641,665]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="illescasi">
<emphasis id="28B2EAF192210D241544189BB3E7FD40" box="[344,455,641,665]" italics="true" pageId="36" pageNumber="35">S. illescasi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192210D2415CB1898B01EFD40" box="[471,574,641,665]" italics="true" pageId="36" pageNumber="35">vide infra</emphasis>
) but differing in the wholly black mesepisternum, the black tegular bristles, and the bristles along the entire mesoscutal margin black.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892210D2514CC18D8B30FFCE0" lastPageId="37" lastPageNumber="36" pageId="36" pageNumber="35" type="description">
<paragraph id="1A7936E392210D2414CC18D8B1BBFBE0" blockId="36.[160,1169,577,1786]" pageId="36" pageNumber="35">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092210D24163318DBB05DFD00" authorityName="Engel" authorityYear="2022" box="[559,637,705,729]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192210D24163318DBB05DFD00" box="[559,637,705,729]" italics="true" pageId="36" pageNumber="35">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192210D24169718D8B0D9FD00" box="[651,761,705,729]" italics="true" pageId="36" pageNumber="35">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately 5.96.0 mm, forewing length (to base of humeral sclerite)
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. Head wider than long, width
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, length
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; compound eye length
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; upper interorbital distance
<quantity id="DD3E9B0692210D2417811938B660FCE0" box="[925,1088,801,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" metricValueMax="1.45" metricValueMin="1.39" pageId="36" pageNumber="35" unit="mm" value="1.42" valueMax="1.45" valueMin="1.39">1.391.45 mm</quantity>
, lower interorbital distance
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. Scape length
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, about as long as toru- locellar distance, torulocellar distance
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. Clypeus approximately 1.71.9× as wide as long, length
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, width
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. Malar area approximately 1.21.4× flagellar diameter or 0.550.65× basal mandibular width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, ca- rina interrupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as sharp, lamellate tooth; upper margin with lamella extended ventrally one third into concavity.
</paragraph>
<paragraph id="1A7936E392210D2514CC1E5BB6ADFF20" blockId="36.[160,1169,577,1786]" lastBlockId="37.[160,1168,225,1081]" lastPageId="37" lastPageNumber="36" pageId="36" pageNumber="35">
Integument of labiomaxillary complex yellow orange; labrum orange; mandible reddish orange; clypeus largely yellow orange to orange with brown along apical mar- gin except sometimes with brownish stain on disc, triangular brown stain mediobasal- ly, brown at apicolateral corners, epistomal sulcus well delineated by dark brown to black; supraclypeal area orange with stain of brown at epistomal sulcus; malar space often dark brown; face below tangent of antennal toruli orange, orange extending up- ward along inner orbit as narrow strip until about one antennal torular diameter above upper tangent of antennal toruli, remainder of face dark brown to black; scape orange, sometimes with brown dorsally; pedicel and flagellum brown above, flagellum orange underneath; vertex dark brown to black; posterior of head dark brown to black; upper third of gena dark brown to black, remainder of gena orange; postgena orange to yel- low orange. Mesosoma black except pronotum orange to brown, mesoscutellum dark fulvous to ferruginous; tegula orange and semi-translucent; legs largely yellow orange to orange except dark brown on prolateral surface of protibia, posterior surface and anterior and dorsal surfaces apically on mesofemur, prolateral surface of mesotibia, prolateral surface of mesobasitarsus, metafemur apically, prolateral surface of metati- bia except extreme base of fundus, superior and inferior margins of retrolateral surface of metatibia, and prolateral surface of metabasitarsus. Wing membranes hyaline clear to lightly parchment colored; veins light brown. Metasoma generally dark brown to black except anterior-facing surface of tergum I orange brown to brown, dorsal-facing disc of tergum I posteriorly coppery orange, tergum II posteriorly coppery orange, with narrow yellow marginal zone; terga
<collectionCode id="7CD7AE2692210D2416961CFBB085F920" box="[650,677,1762,1785]" pageId="36" pageNumber="35">III</collectionCode>
<collectionCode id="7CD7AE2692210D2416AF1CFBB0EFF920" box="[691,719,1762,1785]" pageId="36" pageNumber="35">IV</collectionCode>
with narrow apical marginal zones of light brown to yellow, typically obscured by tomentum; sterna dark brown to brown.
</paragraph>
<paragraph id="1A7936E392200D2514CC1B1BB31BFD20" blockId="37.[160,1168,225,1081]" pageId="37" pageNumber="36">
Areas of short, white, plumose setae in
<taxonomicName id="DDC64D6092200D2516851B1BB0C9FEC0" authorityName="Engel" authorityYear="2022" box="[665,745,257,281]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192200D2516851B1BB0C9FEC0" box="[665,745,257,281]" italics="true" pageId="37" pageNumber="36">S. ederi</emphasis>
</taxonomicName>
instead yellow. Vertex with numer- ous erect, simple, dark fulvous bristles, a few short black bristles laterally near upper orbit; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow and black bristles; mesoscutum anterior and lateral borders with largely black bristles, lat- erally black bristles intermingled with fulvous bristles; tegula with suberect black and fulvous bristles anteriorly; mesoscutellum with abundant, elongate, erect fulvous bris- tles posteriorly; mesepisternum with scatered, subdecumbent fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, su- berect to subdecumbent black to dark fulvous bristles extending ventrally from pro- notal lobe; lateral surface of propodeum with scatered, long, erect, fulvous bristles emerging amid dense, appressed, yellow, plumose setae. Legs with abundant black bristles except yellow to fulvous on coxae, trochanters, femora except metafemur api- cally, and distitarsomeres; metabasitarsus with erect black bristles on prolateral sur- face and margins. Metasoma with numerous, prominent, suberect to subdecumbent, yellow to fulvous bristles on terga
<collectionCode id="7CD7AE2692200D25162518DBB074FD00" box="[569,596,706,729]" pageId="37" pageNumber="36">III</collectionCode>
<collectionCode id="7CD7AE2692200D25167E18DBB055FD00" box="[610,629,706,729]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="37" pageNumber="36" type="Museum">V</collectionCode>
arising amid tomentum, such bristles sparser on tergum II.
</paragraph>
<paragraph id="1A7936E392200D2514CC191BB30FFCC0" blockId="37.[160,1168,225,1081]" box="[208,303,770,796]" pageId="37" pageNumber="36">
<specimenCount id="0CC0FD6A92200D2514CC191BB2C4FCC5" box="[208,228,770,796]" count="1" pageId="37" pageNumber="36" type="female"></specimenCount>
:
<emphasis id="28B2EAF192200D2514ED191BB308FCC0" box="[241,296,770,793]" italics="true" pageId="37" pageNumber="36">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392200D2514CC193BB30FFCE0" blockId="37.[160,1168,225,1081]" box="[208,303,802,828]" pageId="37" pageNumber="36">
<specimenCount id="0CC0FD6A92200D2514CC193BB2C4FCE5" box="[208,228,802,828]" count="1" pageId="37" pageNumber="36" type="male"></specimenCount>
:
<emphasis id="28B2EAF192200D2514ED193BB308FCE0" box="[241,296,802,825]" italics="true" pageId="37" pageNumber="36">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892200D2514CC1958B34DFC60" pageId="37" pageNumber="36" type="materials_examined">
<paragraph id="1A7936E392200D2514CC1958B34DFCA0" blockId="37.[160,1168,225,1081]" pageId="37" pageNumber="36">
<materialsCitation id="AAAE3CBE92200D2514CC1958B34DFCA0" collectingDate="1975-11-11" collectionCode="SEMC" country="Colombia" location="San Martin" pageId="37" pageNumber="36" specimenCount="1" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="C57D884192200D2514CC1958B368FC80" box="[208,328,833,857]" pageId="37" pageNumber="36" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392200D2515711958B3C3FC80" box="[365,483,833,857]" name="Colombia" pageId="37" pageNumber="36">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192200D2515F2195BB00CFC80" box="[494,556,834,857]" country="Colombia" name="Meta" pageId="37" pageNumber="36">Meta</collectingRegion>
,
<location id="1F19603892200D25162B1958B09BFC80" LSID="urn:lsid:plazi:treatment:926F87F592210D2515851BD8B322FBE0:1F19603892200D25162B1958B09BFC80" box="[567,699,833,857]" country="Colombia" name="San Martin" pageId="37" pageNumber="36" stateProvince="Meta">San Martin</location>
,
<date id="6E78102392200D2516DB1958B164FC80" box="[711,836,833,857]" pageId="37" pageNumber="36" value="1975-11-11">
<collectingDate id="7E3CE9CB92200D2516DB1958B164FC80" box="[711,836,833,857]" pageId="37" pageNumber="36" value="1975-11-11">11-XI-1975</collectingDate>
</date>
[
<date id="6E78102392200D25174E1958B612FC80" box="[850,1074,833,857]" pageId="37" pageNumber="36" value="1975-11-11">
<collectingDate id="7E3CE9CB92200D25174E1958B612FC80" box="[850,1074,833,857]" pageId="37" pageNumber="36" value="1975-11-11">11 November 1975</collectingDate>
</date>
], Monc- cuta leg. (
<collectionCode id="7CD7AE2692200D25150D1978B341FCA0" box="[273,353,865,889]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="37" pageNumber="36" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392200D2514CC199BB34DFC60" blockId="37.[160,1168,225,1081]" pageId="37" pageNumber="36">
<materialsCitation id="AAAE3CBE92200D2514CC199BB349FC60" collectingDate="1975-11-11" collectionCode="SEMC" country="Colombia" location="San Martin" pageId="37" pageNumber="36" specimenCount="1" stateProvince="Meta" typeStatus="paratype">
<typeStatus id="C57D884192200D2514CC199BB363FC43" box="[208,323,898,922]" pageId="37" pageNumber="36" type="paratype">PARATYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392200D2515751998B3FFFC40" box="[361,479,897,921]" name="Colombia" pageId="37" pageNumber="36">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192200D2515F6199BB008FC40" box="[490,552,898,921]" country="Colombia" name="Meta" pageId="37" pageNumber="36">Meta</collectingRegion>
,
<location id="1F19603892200D2516281998B099FC40" LSID="urn:lsid:plazi:treatment:926F87F592210D2515851BD8B322FBE0:1F19603892200D2516281998B099FC40" box="[564,697,897,921]" country="Colombia" name="San Martin" pageId="37" pageNumber="36" stateProvince="Meta">San Martin</location>
,
<date id="6E78102392200D2516D91998B162FC40" box="[709,834,897,921]" pageId="37" pageNumber="36" value="1975-11-11">
<collectingDate id="7E3CE9CB92200D2516D91998B162FC40" box="[709,834,897,921]" pageId="37" pageNumber="36" value="1975-11-11">11-XI-1975</collectingDate>
</date>
[
<date id="6E78102392200D25174C1998B612FC40" box="[848,1074,897,921]" pageId="37" pageNumber="36" value="1975-11-11">
<collectingDate id="7E3CE9CB92200D25174C1998B612FC40" box="[848,1074,897,921]" pageId="37" pageNumber="36" value="1975-11-11">11 November 1975</collectingDate>
</date>
], Monc- cuta leg. (
<collectionCode id="7CD7AE2692200D25150D19B8B341FC60" box="[273,353,929,953]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="37" pageNumber="36" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892200D2514CC19D8B322FBE0" pageId="37" pageNumber="36" type="etymology">
<paragraph id="1A7936E392200D2514CC19D8B322FBE0" blockId="37.[160,1168,225,1081]" pageId="37" pageNumber="36">
ETYMOLOGY: The specific epithet is the Latin noun
<taxonomicName id="DDC64D6092200D25172519D8B1BAFC00" authorityName="Engel" authorityYear="2022" box="[825,922,961,985]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="caduceus">
<emphasis id="28B2EAF192200D25172519D8B1BAFC00" box="[825,922,961,985]" italics="true" pageId="37" pageNumber="36">cādūceus</emphasis>
</taxonomicName>
, meaning, “heralds staff” and specifically the famed staff of the Roman god Mercury (Ancient Greek Her- mês / Ἑρμῆς), and from which the name of the neighboring Reserva El Caduceo is derived.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,273 @@
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<mods:title id="FD5E577B8F7954175C82A3BB57E78A0C">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="95E46A6FBAE0B2BB8EF5FEC76DA12CD4">Engel, Michael S.</mods:namePart>
<mods:affiliation id="7A2E910648C45D6755EE9B086D26DCA2">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:date id="02DBD1F4036DEF1EBCE7DB0272DBD9F5">2022</mods:date>
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<subSubSection id="52DC656892240D2115621C38B617F980" pageId="33" pageNumber="32" type="nomenclature">
<paragraph id="1A7936E392240D2115621C38B192F9E0" blockId="33.[382,946,1569,1593]" box="[382,946,1569,1593]" pageId="33" pageNumber="32">
<heading id="4131818F92240D2115621C38B192F9E0" box="[382,946,1569,1593]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="32" reason="2">
<taxonomicName id="DDC64D6092240D2115621C38B13AF9E0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[382,794,1569,1593]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="macarenensis" status="sp. nov.">
<emphasis id="28B2EAF192240D2115621C38B0EEF9E0" bold="true" box="[382,718,1569,1593]" italics="true" pageId="33" pageNumber="32">Scaptotrigona macarenensis</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92240D2117391C3BB192F9E0" box="[805,946,1569,1593]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392240D2114E51C5AB617F980" blockId="33.[249,1079,1603,1657]" box="[249,1079,1603,1625]" pageId="33" pageNumber="32">
ZooBank:
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="52DC656892240D2116741C78B0E8F9A0" box="[616,712,1633,1657]" pageId="33" pageNumber="32" type="description">
<paragraph id="1A7936E392240D2116741C78B0E8F9A0" blockId="33.[249,1079,1603,1657]" box="[616,712,1633,1657]" pageId="33" pageNumber="32">
(
<figureCitation id="82FD2A6692240D21166D1C78B0E0F9A0" box="[625,704,1633,1657]" captionStart="Figures 4950" captionStartId="34.[160,241,1430,1452]" captionTargetBox="[175,1155,233,1415]" captionTargetId="figure-116@34.[160,1168,224,1423]" captionTargetPageId="34" captionText="Figures 4950. Workers of Scaptotrigona Moure. 49. Lateral habitus of Scaptotrigona macarenensis, new species. 50. Lateral habitus of S. caduceus, new species." pageId="33" pageNumber="32">Fig. 49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892240D2214CC1CB8B1A5F9B3" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32" type="diagnosis">
<paragraph id="1A7936E392240D2214CC1CB8B1A5F9B3" blockId="33.[160,1168,1697,1785]" lastBlockId="34.[160,1169,1522,1771]" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32">
DIAGNOSIS: This is a larger species much like
<taxonomicName id="DDC64D6092240D2116F91CBBB1DAF960" authority="(Buysson)" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[741,1018,1697,1721]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192240D2116F91CBBB15BF960" box="[741,891,1697,1721]" italics="true" pageId="33" pageNumber="32">S. ochrotricha</emphasis>
(Buysson)
</taxonomicName>
,
<taxonomicName id="DDC64D6092240D21101A1CBBB3E8F900" authority="Nogueira &amp; Santos-Silva" authorityName="Nogueira &amp; Santos-Silva" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF192240D21101A1CBBB6B0F960" box="[1030,1168,1697,1721]" italics="true" pageId="33" pageNumber="32">S. nigrohirta</emphasis>
Nogueira &amp; Santos-Silva
</taxonomicName>
, and
<taxonomicName id="DDC64D6092240D2116161CDBB12BF900" authority="Moure" authorityName="Moure" authorityYear="1950" box="[522,779,1729,1753]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="xanthotricha">
<emphasis id="28B2EAF192240D2116161CDBB091F900" box="[522,689,1729,1753]" italics="true" pageId="33" pageNumber="32">S. xanthotricha</emphasis>
Moure
</taxonomicName>
, and, owing to the stripes of the mesoscutum most closely resembles the first of these. Unlike
<taxonomicName id="DDC64D6092240D2117631CFBB635F920" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[895,1045,1761,1785]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192240D2117631CFBB635F920" box="[895,1045,1761,1785]" italics="true" pageId="33" pageNumber="32">S. ochrotricha</emphasis>
</taxonomicName>
, however, the fore and mid-legs lack black bristles and the metatibia is dominated by yellow to fulvous bristles. There is the possibility that this is a much lighter form of
<taxonomicName id="DDC64D6092270D2217E01C0AB6B0F9F3" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[1020,1168,1554,1578]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192270D2217E01C0AB6B0F9F3" box="[1020,1168,1554,1578]" italics="true" pageId="34" pageNumber="33">S. ochrotricha</emphasis>
</taxonomicName>
(if this species exhibited color dimorphism), but until more sampling is made across the region and nests found it seems that they are quite distinct.
</paragraph>
</subSubSection>
<caption id="4EB9666B92270D2214BC1F8FB11DFA10" pageId="34" pageNumber="33" startId="34.[160,241,1430,1452]" targetBox="[175,1155,233,1415]" targetPageId="34" targetType="figure">
<paragraph id="1A7936E392270D2214BC1F8FB11DFA10" blockId="34.[160,1168,1430,1481]" pageId="34" pageNumber="33">
<emphasis id="28B2EAF192270D2214BC1F8FB31AFA72" bold="true" box="[160,314,1430,1452]" pageId="34" pageNumber="33">Figures 4950.</emphasis>
Workers of
<taxonomicName id="DDC64D6092270D2215D81F8FB080FA72" authority="Moure." authorityName="Moure." box="[452,672,1430,1451]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF192270D2215D81F8FB06BFA72" box="[452,587,1430,1451]" italics="true" pageId="34" pageNumber="33">Scaptotrigona</emphasis>
Moure.
</taxonomicName>
<emphasis id="28B2EAF192270D2216B01F8FB0EAFA72" bold="true" box="[684,714,1430,1451]" pageId="34" pageNumber="33">49.</emphasis>
Lateral habitus of
<taxonomicName id="DDC64D6092270D22178E1F8FB299FA11" authorityName="Engel" authorityYear="2022" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="macarenensis" status="sp. nov.">
<emphasis id="28B2EAF192270D22178E1F8FB299FA11" italics="true" pageId="34" pageNumber="33">Scaptotrigona macarenensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92270D2214D91FAAB367FA10" box="[197,327,1459,1481]" pageId="34" pageNumber="33" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192270D22154A1FAAB355FA11" bold="true" box="[342,373,1459,1480]" pageId="34" pageNumber="33">50.</emphasis>
Lateral habitus of
<taxonomicName id="DDC64D6092270D2216261FAAB08CFA11" authorityName="Engel" authorityYear="2022" box="[570,684,1459,1480]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="caduceus" status="new species">
<emphasis id="28B2EAF192270D2216261FAAB067FA11" box="[570,583,1459,1480]" italics="true" pageId="34" pageNumber="33">S</emphasis>
.
<emphasis id="28B2EAF192270D2216481FAAB08CFA11" box="[596,684,1459,1480]" italics="true" pageId="34" pageNumber="33">caduceus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92270D2216AB1FAAB119FA10" box="[695,825,1459,1481]" pageId="34" pageNumber="33" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<subSubSection id="52DC656892270D2314CC1C6AB30FF940" lastPageId="35" lastPageNumber="34" pageId="34" pageNumber="33" type="description">
<paragraph id="1A7936E392270D2314CC1C6AB653FE60" blockId="34.[160,1169,1522,1771]" lastBlockId="35.[160,1175,225,1785]" lastPageId="35" lastPageNumber="34" pageId="34" pageNumber="33">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092270D2216331C6AB05DF953" authorityName="Engel" authorityYear="2022" box="[559,637,1650,1674]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192270D2216331C6AB05DF953" box="[559,637,1650,1674]" italics="true" pageId="34" pageNumber="33">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192270D2216971C6BB0D9F953" box="[651,761,1650,1674]" italics="true" pageId="34" pageNumber="33">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately
<quantity id="DD3E9B0692270D2216521C8BB0F7F973" box="[590,727,1682,1706]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.8" metricValueMin="6.3" pageId="34" pageNumber="33" unit="mm" value="6.55" valueMax="6.8" valueMin="6.3">6.36.8 mm</quantity>
, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B0692270D22151B1CABB3B3F913" box="[263,403,1714,1738]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.5" metricValueMin="6.3" pageId="34" pageNumber="33" unit="mm" value="6.4" valueMax="6.5" valueMin="6.3">6.36.5 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B0692270D2217071CABB1E1F913" box="[795,961,1714,1738]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.73" metricValueMin="2.61" pageId="34" pageNumber="33" unit="mm" value="2.67" valueMax="2.73" valueMin="2.61">2.612.73 mm</quantity>
, length
<quantity id="DD3E9B0692270D2210391CAAB2F1F933" metricMagnitude="-3" metricUnit="m" metricValue="2.135" metricValueMax="2.18" metricValueMin="2.09" pageId="34" pageNumber="33" unit="mm" value="2.135" valueMax="2.18" valueMin="2.09">2.092.18 mm</quantity>
; compound eye length
<quantity id="DD3E9B0692270D2215F11CCBB0B1F933" box="[493,657,1746,1770]" metricMagnitude="-3" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.52" metricValueMin="1.45" pageId="34" pageNumber="33" unit="mm" value="1.4849999999999999" valueMax="1.52" valueMin="1.45">1.451.52 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B0692270D2217FB1CCBB6ABF933" box="[999,1163,1746,1770]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" metricValueMax="1.71" metricValueMin="1.67" pageId="34" pageNumber="33" unit="mm" value="1.69" valueMax="1.71" valueMin="1.67">1.671.71 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B0692260D2315FF1AF8B0A8FF20" box="[483,648,225,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.64" metricValueMin="1.55" pageId="35" pageNumber="34" unit="mm" value="1.595" valueMax="1.64" valueMin="1.55">1.551.64 mm</quantity>
. Scape length 0.971.00 mm, about as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B0692260D2316A21B18B144FEC0" box="[702,868,257,281]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.399999999999999" metricValueMin="9.1" pageId="35" pageNumber="34" unit="mm" value="0.925" valueMax="0.94" valueMin="0.91">0.910.94 mm</quantity>
. Clypeus approximately 1.71.75× as wide as long, length
<quantity id="DD3E9B0692260D2316301B38B0F0FEE0" box="[556,720,289,313]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.5" metricValueMin="6.7" pageId="35" pageNumber="34" unit="mm" value="0.71" valueMax="0.75" valueMin="0.67">0.670.75 mm</quantity>
, width
<quantity id="DD3E9B0692260D2317361B38B1EFFEE0" box="[810,975,289,313]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.27" metricValueMin="1.18" pageId="35" pageNumber="34" unit="mm" value="1.225" valueMax="1.27" valueMin="1.18">1.181.27 mm</quantity>
. Malar area ap- proximately 2.0× flagellar diameter or 0.830.85× basal mandibular width. Preoccipi- tal ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as a large lamellate tooth.
</paragraph>
<paragraph id="1A7936E392260D2314CC1BDBB0CFFBE0" blockId="35.[160,1175,225,1785]" pageId="35" pageNumber="34">
Integument of labiomaxillary complex yellow; labrum yellow; mandible yellow, sometimes blending to slightly orange apically; clypeus yellow, epistomal sulcus nearly concolorous with clypeus, delineated by light brown to orange; supraclypeal area yellow; malar space yellow; face below upper tangent of antennal toruli yellow; lower frons yellow and blending to brown or dark brown on upper frons, with brown extending slightly ventrally along inner orbit; ocellocular area and vertex brown to dark brown; uppermost gena brown; remainder of gena and postgena yellow; scape yellow, sometimes with patch of brown dorsoapically; pedicel and flagellum brown, with flagellum lighter ventrally. Mesosoma largely yellow except mesoscutum black and with longitudinal strips of reddish orange laterally, along posterior border, and from notauli to posterior border; metanotum sometimes dark brown to black; propo- deum sometimes dark brown to black, but always with lateral surface largely yellow; metepisternum sometimes with dark brown along posterior border; mesepisternum sometimes with dark brown patch ventrally; tegula yellow and semi-translucent; legs yellow except sometimes with brownish mark on metafemur dorsoapically. Meta- soma generally dark brown except anterior-facing surface of tergum I yellow, dorsal-facing disc of tergum I largely yellow except brown to dark brown laterally; tergum II yellow to yellow brown on disc, with broad areas of brown laterally and yellow apical marginal zone; terga
<collectionCode id="7CD7AE2692260D2315861E1BB395FBC0" box="[410,437,1026,1049]" pageId="35" pageNumber="34">III</collectionCode>
<collectionCode id="7CD7AE2692260D2315DF1E1BB3F6FBC0" box="[451,470,1026,1049]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="35" pageNumber="34" type="Museum">V</collectionCode>
dark brown with large yellow marginal zones obscured by tomentum; sterna largely yellow to yellow brown.
</paragraph>
<paragraph id="1A7936E392260D2314CC1E5BB336F980" blockId="35.[160,1175,225,1785]" pageId="35" pageNumber="34">
As for
<taxonomicName id="DDC64D6092260D2315021E5BB34EFB80" authorityName="Engel" authorityYear="2022" box="[286,366,1089,1113]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192260D2315021E5BB34EFB80" box="[286,366,1089,1113]" italics="true" pageId="35" pageNumber="34">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white; metepisternum and lateral surface of propodeum obscured by dense, plumose, yellow setae; dorsal-facing surface of tergum I with largely simple setae, minute, de- cumbent, yellow, plumose setae nearly absent. Bristles of vertex erect, simple, yellow to fulvous; pronotal dorsolateral margin near and anterior to pronotal lobe with long, arched, yellow bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with yellow to fulvous bristles; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elon- gate, erect bristles posteriorly, such bristles minutely pectinate apically, discal bristles shorter, mesoscutellar bristles yellow; mesepisternal bristles as in
<taxonomicName id="DDC64D6092260D2317A91F7BB627FAA0" authorityName="Engel" authorityYear="2022" box="[949,1031,1377,1401]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192260D2317A91F7BB627FAA0" box="[949,1031,1377,1401]" italics="true" pageId="35" pageNumber="34">S. ederi</emphasis>
</taxonomicName>
except yel- low. Legs with abundant bristles, bristles yellow to fulvous; corbicular bristles dark fulvous to fuscous, marginal bristles yellow to fulvous; prolateral surface of metaba- sitarsus with scatered fine, simple, yellow setae (noticeably thinner than marginal bristles), without black setae or bristles; penicillum dark fulvous; rastellum fulvous. Metasoma with numerous, prominent, suberect to subdecumbent, yellow to fulvous bristles on terga
<collectionCode id="7CD7AE2692260D23157B1C3BB3A2F9E0" box="[359,386,1570,1593]" pageId="35" pageNumber="34">III</collectionCode>
<collectionCode id="7CD7AE2692260D2315931C3BB382F9E0" box="[399,418,1570,1593]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="35" pageNumber="34" type="Museum">V</collectionCode>
arising from tomentum, as well as similar bristles scatered on tergum II.
</paragraph>
<paragraph id="1A7936E392260D2314CC1C7BB30FF9A0" blockId="35.[160,1175,225,1785]" box="[208,303,1634,1660]" pageId="35" pageNumber="34">
<specimenCount id="0CC0FD6A92260D2314CC1C7BB2C4F9A5" box="[208,228,1634,1660]" count="1" pageId="35" pageNumber="34" type="female"></specimenCount>
:
<emphasis id="28B2EAF192260D2314ED1C7BB308F9A0" box="[241,296,1634,1657]" italics="true" pageId="35" pageNumber="34">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392260D2314CC1C9BB30FF940" blockId="35.[160,1175,225,1785]" box="[208,303,1666,1692]" pageId="35" pageNumber="34">
<specimenCount id="0CC0FD6A92260D2314CC1C9BB2C4F945" box="[208,228,1666,1692]" count="1" pageId="35" pageNumber="34" type="male"></specimenCount>
:
<emphasis id="28B2EAF192260D2314ED1C9BB308F940" box="[241,296,1666,1689]" italics="true" pageId="35" pageNumber="34">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892260D2414CC1CB8B170FEC0" lastPageId="36" lastPageNumber="35" pageId="35" pageNumber="34" type="materials_examined">
<paragraph id="1A7936E392260D2314CC1CB8B0EEF900" blockId="35.[160,1175,225,1785]" pageId="35" pageNumber="34">
<materialsCitation id="AAAE3CBE92260D2314CC1CB8B0EEF900" collectingDate="1988-04-01" collectionCode="SEMC" collectorName="F. Fernandez" country="Colombia" location="Reserva La Macarena" pageId="35" pageNumber="34" specimenCount="1" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="C57D884192260D2314CC1CB8B368F960" box="[208,328,1697,1721]" pageId="35" pageNumber="34" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392260D23156D1CB8B3C7F960" box="[369,487,1697,1721]" name="Colombia" pageId="35" pageNumber="34">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192260D2315EF1CBBB011F960" box="[499,561,1698,1721]" country="Colombia" name="Meta" pageId="35" pageNumber="34">Meta</collectingRegion>
,
<location id="1F19603892260D2316221CBBB11FF960" LSID="urn:lsid:plazi:treatment:926F87F592240D2415621C38B374FE40:1F19603892260D2316221CBBB11FF960" box="[574,831,1698,1721]" country="Colombia" name="Reserva La Macarena" pageId="35" pageNumber="34" stateProvince="Meta">Reserva La Macarena</location>
,
<locationDeviation id="0BCB2FC192260D2317501CB8B304F900" firstLocation="Rio Duda" pageId="35" pageNumber="34" secondLocation="Rio Guayabero">between Rio Duda and Rio Guayabero</locationDeviation>
,
<date id="6E78102392260D2315331CD8B3E0F900" box="[303,448,1729,1753]" pageId="35" pageNumber="34" value="1988-04-01">
<collectingDate id="7E3CE9CB92260D2315331CD8B3E0F900" box="[303,448,1729,1753]" pageId="35" pageNumber="34" value="1988-04-01">1 April 1988</collectingDate>
</date>
,
<collectorName id="B733533592260D2315D61CDBB044F900" box="[458,612,1729,1753]" pageId="35" pageNumber="34">F. Fernández</collectorName>
(
<collectionCode id="7CD7AE2692260D23166E1CD8B0E2F900" box="[626,706,1729,1753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="35" pageNumber="34" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392260D2414CC1CF8B170FEC0" blockId="35.[160,1175,225,1785]" lastBlockId="36.[160,1168,225,409]" lastPageId="36" lastPageNumber="35" pageId="35" pageNumber="34">
<materialsCitation id="AAAE3CBE92260D2414CC1CF8B164FF20" collectingDate="1988-04-01" collectionCode="SEMC" collectorName="F. Fernandez" country="Colombia" lastPageId="36" lastPageNumber="35" location="Reserva La Macarena" pageId="35" pageNumber="34" specimenCount="1" stateProvince="Meta" typeStatus="paratype">
<typeStatus id="C57D884192260D2314CC1CF8B36EF920" box="[208,334,1761,1785]" pageId="35" pageNumber="34" type="paratype">PARATYPES</typeStatus>
: 2⚲⚲,
<collectingCountry id="62D1767392260D2315891CF8B02BF920" box="[405,523,1761,1785]" name="Colombia" pageId="35" pageNumber="34">Colombia</collectingCountry>
:
<collectingRegion id="D802F80192260D2316051CFBB077F920" box="[537,599,1762,1785]" country="Colombia" name="Meta" pageId="35" pageNumber="34">Meta</collectingRegion>
,
<location id="1F19603892260D23167A1CFBB14AF920" LSID="urn:lsid:plazi:treatment:926F87F592240D2415621C38B374FE40:1F19603892260D23167A1CFBB14AF920" box="[614,874,1762,1785]" country="Colombia" name="Reserva La Macarena" pageId="35" pageNumber="34" stateProvince="Meta">Reserva La Macarena</location>
,
<locationDeviation id="0BCB2FC192260D2417641CF8B376FF20" firstLocation="Rio Duda" lastPageId="36" lastPageNumber="35" pageId="35" pageNumber="34" secondLocation="Rio Guayabero">between Rio Duda and Rio Guayabero</locationDeviation>
,
<date id="6E78102392210D2415781AF8B3DAFF20" box="[356,506,225,249]" pageId="36" pageNumber="35" value="1988-04-01">
<collectingDate id="7E3CE9CB92210D2415781AF8B3DAFF20" box="[356,506,225,249]" pageId="36" pageNumber="35" value="1988-04-01">1 April 1988</collectingDate>
</date>
,
<collectorName id="B733533592210D2416141AFBB084FF20" box="[520,676,225,249]" pageId="36" pageNumber="35">F. Fernández</collectorName>
(
<collectionCode id="7CD7AE2692210D2416A91AF8B124FF20" box="[693,772,225,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="36" pageNumber="35" type="Museum">SEMC</collectionCode>
); 3⚲⚲
</materialsCitation>
,
<materialsCitation id="AAAE3CBE92210D2417491AF8B16CFEC0" collectingDate="1950-01-11" collectionCode="SEMC" collectorName="L. Richter" country="Colombia" elevation="575" location="Macarena Mts." pageId="36" pageNumber="35" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392210D2417491AF8B1EBFF20" box="[853,971,225,249]" name="Colombia" pageId="36" pageNumber="35">Colombia</collectingCountry>
:
<location id="1F19603892210D2417C41AFBB6ACFF20" LSID="urn:lsid:plazi:treatment:926F87F592240D2415621C38B374FE40:1F19603892210D2417C41AFBB6ACFF20" box="[984,1164,226,249]" country="Colombia" name="Macarena Mts." pageId="36" pageNumber="35">Macarena Mts.</location>
,
<date id="6E78102392210D2414BC1B18B32BFEC0" box="[160,267,257,281]" pageId="36" pageNumber="35" value="1950-01-11">
<collectingDate id="7E3CE9CB92210D2414BC1B18B32BFEC0" box="[160,267,257,281]" pageId="36" pageNumber="35" value="1950-01-11">I-11-1950</collectingDate>
</date>
[
<date id="6E78102392210D2415051B18B3F9FEC0" box="[281,473,257,282]" pageId="36" pageNumber="35" value="1950-01-11">
<collectingDate id="7E3CE9CB92210D2415051B18B3F9FEC0" box="[281,473,257,282]" pageId="36" pageNumber="35" value="1950-01-11">11 January 1950</collectingDate>
</date>
],
<quantity id="DD3E9B0692210D2415F61B18B048FEC0" box="[490,616,257,281]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="36" pageNumber="35" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">
<elevation id="91EBD1D092210D2415F61B18B048FEC0" box="[490,616,257,281]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="36" pageNumber="35" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500650 m</elevation>
</quantity>
,
<collectorName id="B733533592210D24166F1B1BB0C6FEC0" box="[627,742,257,281]" pageId="36" pageNumber="35">L. Richter</collectorName>
(
<collectionCode id="7CD7AE2692210D2416E81B18B164FEC0" box="[756,836,257,281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="36" pageNumber="35" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892210D2414CC1B38B374FE40" pageId="36" pageNumber="35" type="etymology">
<paragraph id="1A7936E392210D2414CC1B38B374FE40" blockId="36.[160,1168,225,409]" pageId="36" pageNumber="35">
ETYMOLOGY: The specific epithet refers to the Serranía de la Macarena and its asso- ciated national ecological reserve in
<collectingRegion id="D802F80192210D2416591B5BB138FE80" box="[581,792,322,345]" country="Colombia" name="Meta" pageId="36" pageNumber="35">Meta Department</collectingRegion>
,
<collectingCountry id="62D1767392210D24173E1B58B1B8FE80" box="[802,920,321,345]" name="Colombia" pageId="36" pageNumber="35">Colombia</collectingCountry>
. The name is formed from Macarena and the Latin suffix
<emphasis id="28B2EAF192210D24167D1B7BB0B6FEA0" box="[609,662,354,377]" italics="true" pageId="36" pageNumber="35">ēnsis</emphasis>
, which is added to a toponym to indicate “from” or “of”.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,288 @@
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<mods:title id="9DBBB65FBD8EE7E8AD90A91CC06E7170">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="CA08B58A3208DF13D8418482F3EFBB9D">Engel, Michael S.</mods:namePart>
<mods:affiliation id="506447EFD155D58FEC4D070065988E22">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0741A90CACE1B72438A25C147E2276C0">text</mods:typeOfResource>
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<mods:titleInfo id="4747F64525AF1DFCF2398C498C4E70CB">
<mods:title id="9DB09CBDFD331ACE4EDC65592C192021">Journal of Melittology</mods:title>
</mods:titleInfo>
<mods:part id="2BAB1DC33D0922552F1FCC0D8EED253D">
<mods:date id="C28DC41FBE80D4DE2A57CBDE6AF9A0DC">2022</mods:date>
<mods:detail id="C122D0D70F61D49F5A008A90D5A0D107" type="pubDate">
<mods:number id="985711A9450049F4A7C817139B789052">2022-03-14</mods:number>
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<mods:detail id="0FB2EAD029BF7E28B8A746563C691EA1" type="volume">
<mods:number id="F714E73AA9E10D0D29AE79DA9CE73D57">2022</mods:number>
</mods:detail>
<mods:detail id="53AC5D9B5EDE3AE0F7821B0ADB8B49F9" type="issue">
<mods:number id="3E245C031A87712C55AB2DAFA0948BAE">110</mods:number>
</mods:detail>
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<mods:start id="79390BBCB81E9B5A866CBDE8557D160D">1</mods:start>
<mods:end id="AD8022A82A2F6F462D3F49444ACB473A">51</mods:end>
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<mods:location id="F4915DCBC2154FB7DD29D622A9272881">
<mods:url id="B093902CD9D0CBC9086649D760703D35">http://dx.doi.org/10.17161/jom.i110.17001</mods:url>
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<mods:classification id="997A29658C7666A0DB95C8E0AE9E501D">journal article</mods:classification>
<mods:identifier id="A2C7DE7F34063AB24A811BF33BB337B4" type="DOI">10.17161/jom.i110.17001</mods:identifier>
<mods:identifier id="AC16DB9B032516E9AF6343667C8F7D03" type="ISSN">2325-4467</mods:identifier>
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<subSubSection id="52DC656892290D2C159A1898B61BFD60" pageId="44" pageNumber="43" type="nomenclature">
<paragraph id="1A7936E392290D2C159A1898B18AFD40" blockId="44.[390,938,641,665]" box="[390,938,641,665]" pageId="44" pageNumber="43">
<heading id="4131818F92290D2C159A1898B18AFD40" box="[390,938,641,665]" centered="true" fontSize="10" level="2" pageId="44" pageNumber="43" reason="2">
<taxonomicName id="DDC64D6092290D2C159A1898B132FD40" authority="Engel" authorityName="Engel" authorityYear="2022" box="[390,786,641,665]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="yungasensis" status="sp. nov.">
<emphasis id="28B2EAF192290D2C159A1898B0E6FD40" bold="true" box="[390,710,641,665]" italics="true" pageId="44" pageNumber="43">Scaptotrigona yungasensis</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92290D2C1701189BB18AFD40" box="[797,938,641,665]" pageId="44" pageNumber="43" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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ZooBank:
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<subSubSection id="52DC656892290D2C165318D8B0C2FD00" box="[591,738,705,729]" pageId="44" pageNumber="43" type="description">
<paragraph id="1A7936E392290D2C165318D8B0C2FD00" blockId="44.[245,1083,675,729]" box="[591,738,705,729]" pageId="44" pageNumber="43">
(
<figureCitation id="82FD2A6692290D2C164B18D8B0FAFD00" box="[599,730,705,729]" captionStart="Figures 5961" captionStartId="45.[160,241,1301,1323]" captionTargetBox="[189,1145,236,1280]" captionTargetId="figure-151@45.[160,1168,224,1295]" captionTargetPageId="45" captionText="Figures 5961. Worker of Scaptotrigona yungasensis, new species. 59. Lateral habitus. 60. Dorsal habitus. 61. Facial view." pageId="44" pageNumber="43">Figs. 5961</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892290D2C14CC1918B071FBC0" pageId="44" pageNumber="43" type="diagnosis">
<paragraph id="1A7936E392290D2C14CC1918B071FBC0" blockId="44.[160,1175,769,1786]" pageId="44" pageNumber="43">
DIAGNOSIS: In many respects this species is much like the Venezuelan
<taxonomicName id="DDC64D6092290D2C17E0191BB6B0FCC0" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[1020,1168,769,793]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192290D2C17E0191BB6B0FCC0" box="[1020,1168,769,793]" italics="true" pageId="44" pageNumber="43">S. ochrotricha</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192290D2C14B51938B330FCE0" box="[169,272,801,825]" italics="true" pageId="44" pageNumber="43">vide infra</emphasis>
), as both are comparatively large species with a generally similar coloration. However, in the new species the corbicular bristles differ in color between the inferior and superior margins and the fringes are typically yellow to light fulvous, while in
<taxonomicName id="DDC64D6092290D2C1067197BB339FC40" baseAuthorityName="Buysson" baseAuthorityYear="1892" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192290D2C1067197BB339FC40" italics="true" pageId="44" pageNumber="43">S. ochrotricha</emphasis>
</taxonomicName>
they are dark fuscous to black on the corbicula while the marginal fringes range from fulvous to more typically black. Additionally,
<taxonomicName id="DDC64D6092290D2C175719BBB1CBFC60" authorityName="Engel" authorityYear="2022" box="[843,1003,930,953]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="yungasensis">
<emphasis id="28B2EAF192290D2C175719BBB1CBFC60" box="[843,1003,930,953]" italics="true" pageId="44" pageNumber="43">S. yungasensis</emphasis>
</taxonomicName>
lacks the con- spicuous mesoscutal stripes present in the former species (
<emphasis id="28B2EAF192290D2C174319D8B151FC00" box="[863,881,961,985]" italics="true" pageId="44" pageNumber="43">cf</emphasis>
.
<figureCitation id="82FD2A6692290D2C179C19D8B1FDFC00" box="[896,989,961,985]" captionStart="Figures 5961" captionStartId="45.[160,241,1301,1323]" captionTargetBox="[189,1145,236,1280]" captionTargetId="figure-151@45.[160,1168,224,1295]" captionTargetPageId="45" captionText="Figures 5961. Worker of Scaptotrigona yungasensis, new species. 59. Lateral habitus. 60. Dorsal habitus. 61. Facial view." pageId="44" pageNumber="43">Figs. 60</figureCitation>
,
<figureCitation id="82FD2A6692290D2C17F519D8B626FC00" box="[1001,1030,961,985]" captionStart="Figures 6768" captionStartId="50.[160,241,935,957]" captionTargetBox="[160,1168,224,923]" captionTargetId="figure-266@50.[160,1168,224,929]" captionTargetPageId="50" captionText="Figures 6768. Worker of Scaptotrigona ochrotricha (Buysson). 67. Dorsal view of head and me- sosoma. 68. Dorsal view of metasoma." pageId="44" pageNumber="43">67</figureCitation>
), as well as the dark fulvous to black bristles otherwise present on the mesotibia and mesepister- num of
<taxonomicName id="DDC64D6092290D2C14E01E1BB3B1FBC0" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[252,401,1025,1049]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192290D2C14E01E1BB3B1FBC0" box="[252,401,1025,1049]" italics="true" pageId="44" pageNumber="43">S. ochrotricha</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192290D2C15BD1E18B393FBC0" box="[417,435,1025,1049]" italics="true" pageId="44" pageNumber="43">cf</emphasis>
.
<figureCitation id="82FD2A6692290D2C15DC1E18B03BFBC0" box="[448,539,1025,1049]" captionStart="Figures 5961" captionStartId="45.[160,241,1301,1323]" captionTargetBox="[189,1145,236,1280]" captionTargetId="figure-151@45.[160,1168,224,1295]" captionTargetPageId="45" captionText="Figures 5961. Worker of Scaptotrigona yungasensis, new species. 59. Lateral habitus. 60. Dorsal habitus. 61. Facial view." pageId="44" pageNumber="43">Figs. 59</figureCitation>
,
<figureCitation id="82FD2A6692290D2C163A1E18B064FBC0" box="[550,580,1025,1049]" captionStart="Figures 6466" captionStartId="49.[160,241,1434,1456]" captionTargetBox="[173,1148,233,1411]" captionTargetId="figure-61@49.[160,1168,223,1428]" captionTargetPageId="49" captionText="Figures 6466. Worker of Scaptotrigona ochrotricha (Buysson). 64. Lateral habitus. 65. Dorsal habitus. 66. Facial view." pageId="44" pageNumber="43">64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="52DC656892290D2E14CC1E38B30FFD40" lastPageId="46" lastPageNumber="45" pageId="44" pageNumber="43" type="description">
<paragraph id="1A7936E392290D2C14CC1E38B151FAA0" blockId="44.[160,1175,769,1786]" pageId="44" pageNumber="43">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092290D2C16331E3BB05DFBE0" authorityName="Engel" authorityYear="2022" box="[559,637,1057,1081]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192290D2C16331E3BB05DFBE0" box="[559,637,1057,1081]" italics="true" pageId="44" pageNumber="43">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192290D2C16971E38B0D9FBE0" box="[651,761,1057,1081]" italics="true" pageId="44" pageNumber="43">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately 6.47.0 mm, forewing length (to base of humeral sclerite)
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. Head wider than long, width
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, length
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; compound eye length
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; upper interorbital distance
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, lower interorbital distance
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. Scape length
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, about as long as torulocellar distance, torulocellar distance
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. Clypeus approximately 1.71.8× as wide as long, length
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, width
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. Malar area ap- proximately 1.8× flagellar diameter or 0.700.73× basal mandibular width. Preoccipi- tal ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming acute angle projecting upward as sharp tooth.
</paragraph>
<paragraph id="1A7936E392290D2D14CC1F9BB030F973" blockId="44.[160,1175,769,1786]" lastBlockId="45.[160,1168,1394,1770]" lastPageId="45" lastPageNumber="44" pageId="44" pageNumber="43">
Integument of labiomaxillary complex yellow orange except prementum typi- cally brown; labrum yellow orange to orange; mandible orange basally and reddish orange apically; clypeus largely yellow orange to orange, epistomal sulcus well delin- eated by brownish orange to black; supraclypeal area yellow orange to orange; malar space yellow orange to orange; face below tangent of antennal toruli yellow orange to orange; scape yellow except brown dorosapically; pedicel and flagellum brown to dark brown, flagellum orange underneath; face above antennal toruli reddish brown to dark brown, even black in some specimens, sometimes with areas of orange im- mediately above antennal toruli; vertex and posterior of head dark brown to black; uppermost portion of gena dark brown to black, remainder of gena orange to yel- low orange; postgena yellow orange to yellow. Mesosoma largely yellow orange to orange except mesoscutum black; propodeum orange to black, often reddish brown; mes- and metepisternum reddish brown to dark brown ventrally, mesepisternum also often with black above between pronotal lobe and hypoepimeral area; lateral surface of propodeum reddish brown to black except upper third to quarter typically orange; tegula yellow to yellow orange and semi-translucent; legs largely yellow orange to orange except reddish brown to dark brown medially on corbicula and sometimes on prolateral surface of metabasitarsus. Wing membranes hyaline clear to lightly parch- ment colored; veins orange. Metasoma generally dark brown to black except anterior-facing surface of tergum I yellow orange to orange, dorsal-facing surface of tergum I and terga II
<collectionCode id="7CD7AE2692280D2D152A1C6BB369F950" box="[310,329,1650,1673]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="45" pageNumber="44" type="Museum">V</collectionCode>
posteriorly brownish orange blending to yellow apical marginal zones; sterna yellow brown to orange.
</paragraph>
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<paragraph id="1A7936E392280D2D14BC1F0CB383FA91" blockId="45.[160,1168,1301,1352]" pageId="45" pageNumber="44">
<emphasis id="28B2EAF192280D2D14BC1F0CB318FAF3" bold="true" box="[160,312,1301,1323]" pageId="45" pageNumber="44">Figures 5961.</emphasis>
Worker of
<taxonomicName id="DDC64D6092280D2D15A91F0FB098FAF2" authorityName="Engel" authorityYear="2022" box="[437,696,1302,1323]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="44" phylum="Arthropoda" rank="species" species="yungasensis" status="sp. nov.">
<emphasis id="28B2EAF192280D2D15A91F0FB098FAF2" box="[437,696,1302,1323]" italics="true" pageId="45" pageNumber="44">Scaptotrigona yungasensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92280D2D16D81F0FB165FAF2" box="[708,837,1301,1323]" pageId="45" pageNumber="44" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192280D2D174F1F0CB152FAF3" bold="true" box="[851,882,1301,1322]" pageId="45" pageNumber="44">59.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192280D2D103A1F0CB665FAF3" bold="true" box="[1062,1093,1301,1322]" pageId="45" pageNumber="44">60.</emphasis>
Dorsal habitus.
<emphasis id="28B2EAF192280D2D151D1F2BB33FFA9E" bold="true" box="[257,287,1330,1351]" pageId="45" pageNumber="44">61.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="1A7936E392280D2E14CC1CABB0D6FD80" blockId="45.[160,1168,1394,1770]" lastBlockId="46.[160,1175,225,922]" lastPageId="46" lastPageNumber="45" pageId="45" pageNumber="44">
Areas of short, white, plumose setae in
<taxonomicName id="DDC64D6092280D2D16851CABB0C9F913" authorityName="Engel" authorityYear="2022" box="[665,745,1714,1738]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="44" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192280D2D16851CABB0C9F913" box="[665,745,1714,1738]" italics="true" pageId="45" pageNumber="44">S. ederi</emphasis>
</taxonomicName>
instead yellow. Vertex with numer- ous erect, simple, yellow to light fulvous bristles; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow to light fulvous bristles; mesoscutum ante- rior and lateral borders with largely yellow to sometimes light fulvous bristles; tegula with suberect yellow bristles anteriorly; mesoscutellum with abundant, elongate, erect yellow to sometimes light fulvous bristles; mesepisternum with scatered, erect to sub- decumbent yellow to light fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, suberect yellow to light fulvous bristles extending ventrally from pronotal lobe; lateral surface of propodeum with scatered, long, erect, yellow bristles. Legs with abundant yellow to sometimes light fulvous bristles except metatibial marginal fringes yellow to fulvous, inferior corbicular bris- tles light fulvous to fulvous, superior corbicular bristles dark fulvous to nearly black. Metasoma with numerous, prominent, suberect to subdecumbent, light fulvous to yel- low bristles on terga
<collectionCode id="7CD7AE26922B0D2E158A185BB391FD80" box="[406,433,578,601]" pageId="46" pageNumber="45">III</collectionCode>
<collectionCode id="7CD7AE26922B0D2E15A3185BB3F2FD80" box="[447,466,578,601]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="46" pageNumber="45" type="Museum">V</collectionCode>
arising amid tomentum.
</paragraph>
<paragraph id="1A7936E3922B0D2E14CC187BB30FFDA0" blockId="46.[160,1175,225,922]" box="[208,303,610,636]" pageId="46" pageNumber="45">
<specimenCount id="0CC0FD6A922B0D2E14CC187BB2C4FDA5" box="[208,228,610,636]" count="1" pageId="46" pageNumber="45" type="female"></specimenCount>
:
<emphasis id="28B2EAF1922B0D2E14ED187BB308FDA0" box="[241,296,610,633]" italics="true" pageId="46" pageNumber="45">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E3922B0D2E14CC189BB30FFD40" blockId="46.[160,1175,225,922]" box="[208,303,642,668]" pageId="46" pageNumber="45">
<specimenCount id="0CC0FD6A922B0D2E14CC189BB2C4FD45" box="[208,228,642,668]" count="1" pageId="46" pageNumber="45" type="male"></specimenCount>
:
<emphasis id="28B2EAF1922B0D2E14ED189BB308FD40" box="[241,296,642,665]" italics="true" pageId="46" pageNumber="45">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922B0D2E14CC18B8B0B7FCE0" pageId="46" pageNumber="45" type="materials_examined">
<paragraph id="1A7936E3922B0D2E14CC18B8B673FD60" blockId="46.[160,1175,225,922]" box="[208,1107,673,698]" pageId="46" pageNumber="45">
<materialsCitation id="AAAE3CBE922B0D2E14CC18B8B673FD60" box="[208,1107,673,698]" collectingDate="1950-10" collectionCode="SEMC" country="Bolivia" elevation="1000" location="Yungas del Palmar" pageId="46" pageNumber="45" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D8841922B0D2E14CC18B8B368FD60" box="[208,328,673,697]" pageId="46" pageNumber="45" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D17673922B0D2E157118B8B3E3FD60" box="[365,451,673,697]" name="Bolivia" pageId="46" pageNumber="45">Bolivia</collectingCountry>
:
<location id="1F196038922B0D2E15D118BBB08EFD60" LSID="urn:lsid:plazi:treatment:926F87F592290D2E159A1898B001FC40:1F196038922B0D2E15D118BBB08EFD60" box="[461,686,673,698]" country="Bolivia" name="Yungas del Palmar" pageId="46" pageNumber="45">Yungas del Palmar</location>
,
<quantity id="DD3E9B06922B0D2E16A518B8B129FD60" box="[697,777,673,697]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="45" unit="m" value="1000.0">
<elevation id="91EBD1D0922B0D2E16A518B8B129FD60" box="[697,777,673,697]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="45" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<date id="6E781023922B0D2E170F18B8B1A2FD60" box="[787,898,673,697]" pageId="46" pageNumber="45" value="1950-10">
<collectingDate id="7E3CE9CB922B0D2E170F18B8B1A2FD60" box="[787,898,673,697]" pageId="46" pageNumber="45" value="1950-10">Oct. 1950</collectingDate>
</date>
, Zischka (
<collectionCode id="7CD7AE26922B0D2E17E418B8B668FD60" box="[1016,1096,673,697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="45" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E3922B0D2E14CC18D8B0B7FCE0" blockId="46.[160,1175,225,922]" pageId="46" pageNumber="45">
<materialsCitation id="AAAE3CBE922B0D2E14CC18D8B6ACFD00" box="[208,1164,705,730]" collectingDate="1950-10" collectionCode="SEMC" collectorName="Zischka" country="Bolivia" elevation="1000" location="Yungas del Palmar" pageId="46" pageNumber="45" specimenCount="1" typeStatus="paratype">
<typeStatus id="C57D8841922B0D2E14CC18D8B36EFD00" box="[208,334,705,729]" pageId="46" pageNumber="45" type="paratype">PARATYPES</typeStatus>
: 116⚲⚲,
<collectingCountry id="62D17673922B0D2E15B618D8B020FD00" box="[426,512,705,729]" name="Bolivia" pageId="46" pageNumber="45">Bolivia</collectingCountry>
:
<location id="1F196038922B0D2E161618DBB0CBFD00" LSID="urn:lsid:plazi:treatment:926F87F592290D2E159A1898B001FC40:1F196038922B0D2E161618DBB0CBFD00" box="[522,747,705,730]" country="Bolivia" name="Yungas del Palmar" pageId="46" pageNumber="45">Yungas del Palmar</location>
,
<quantity id="DD3E9B06922B0D2E16EA18D8B166FD00" box="[758,838,705,729]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="45" unit="m" value="1000.0">
<elevation id="91EBD1D0922B0D2E16EA18D8B166FD00" box="[758,838,705,729]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="45" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<date id="6E781023922B0D2E174C18D8B19FFD00" box="[848,959,705,729]" pageId="46" pageNumber="45" value="1950-10">
<collectingDate id="7E3CE9CB922B0D2E174C18D8B19FFD00" box="[848,959,705,729]" pageId="46" pageNumber="45" value="1950-10">Oct. 1950</collectingDate>
</date>
,
<collectorName id="B7335335922B0D2E17D618D8B606FD00" box="[970,1062,705,729]" pageId="46" pageNumber="45">Zischka</collectorName>
(
<collectionCode id="7CD7AE26922B0D2E102918D8B6A4FD00" box="[1077,1156,705,729]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="45" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 1⚲,
<materialsCitation id="AAAE3CBE922B0D2E14D218F8B2DFFCC0" collectingDate="1966-11-10" collectionCode="SEMC" collectorName="L. Pena" country="Bolivia" location="Alto Marani" municipality="Depto." pageId="46" pageNumber="45" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
<collectingCountry id="62D17673922B0D2E14D218F8B304FD20" box="[206,292,737,761]" name="Bolivia" pageId="46" pageNumber="45">Bolivia</collectingCountry>
:
<collectingMunicipality id="FA1DAC99922B0D2E152E18FBB3A2FD20" box="[306,386,738,761]" pageId="46" pageNumber="45">Depto.</collectingMunicipality>
<collectingRegion id="D802F801922B0D2E159118FBB3C2FD20" box="[397,482,738,761]" country="Bolivia" name="La Paz" pageId="46" pageNumber="45">La Paz</collectingRegion>
,
<location id="1F196038922B0D2E15ED18F8B0A8FD20" LSID="urn:lsid:plazi:treatment:926F87F592290D2E159A1898B001FC40:1F196038922B0D2E15ED18F8B0A8FD20" box="[497,648,737,761]" country="Bolivia" municipality="Depto." name="Alto Marani" pageId="46" pageNumber="45" stateProvince="La Paz">Alto Marani</location>
,
<locationDeviation id="0BCB2FC1922B0D2E168418FBB159FD20" box="[664,889,737,761]" location="Rurrenbeque" pageId="46" pageNumber="45">N of Rurrenbeque</locationDeviation>
,
<date id="6E781023922B0D2E179418F8B606FD20" box="[904,1062,737,761]" pageId="46" pageNumber="45" value="1966-11-10">
<collectingDate id="7E3CE9CB922B0D2E179418F8B606FD20" box="[904,1062,737,761]" pageId="46" pageNumber="45" value="1966-11-10">10 Nov 1966</collectingDate>
</date>
,
<collectorName id="B7335335922B0D2E102918FBB6B0FD20" box="[1077,1168,738,761]" pageId="46" pageNumber="45">L. Peña</collectorName>
(
<collectionCode id="7CD7AE26922B0D2E14B41918B2D7FCC0" box="[168,247,769,793]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="45" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE922B0D2E155E1918B1D3FCC0" box="[322,1011,769,793]" collectingDate="1956-11-12" collectionCode="SEMC" collectorName="L. Pena" country="Bolivia" location="Pto. Fon" pageId="46" pageNumber="45" specimenCount="1" stateProvince="Beni" typeStatus="paratype">
<collectingCountry id="62D17673922B0D2E155E1918B3B8FCC0" box="[322,408,769,793]" name="Bolivia" pageId="46" pageNumber="45">Bolivia</collectingCountry>
:
<location id="1F196038922B0D2E15BF191BB025FCC0" LSID="urn:lsid:plazi:treatment:926F87F592290D2E159A1898B001FC40:1F196038922B0D2E15BF191BB025FCC0" box="[419,517,770,793]" country="Bolivia" name="Pto. Fon" pageId="46" pageNumber="45" stateProvince="Beni">Pto. Fon</location>
<collectingRegion id="D802F801922B0D2E16101918B063FCC0" box="[524,579,769,793]" country="Bolivia" name="Beni" pageId="46" pageNumber="45">Beni</collectingRegion>
,
<collectingRegion id="D802F801922B0D2E16521918B0A5FCC0" box="[590,645,769,793]" country="Bolivia" name="Beni" pageId="46" pageNumber="45">Beni</collectingRegion>
,
<date id="6E781023922B0D2E168C1918B10BFCC0" box="[656,811,769,793]" pageId="46" pageNumber="45" value="1956-11-12">
<collectingDate id="7E3CE9CB922B0D2E168C1918B10BFCC0" box="[656,811,769,793]" pageId="46" pageNumber="45" value="1956-11-12">12 Nov. 1956</collectingDate>
</date>
,
<collectorName id="B7335335922B0D2E172A191BB1ADFCC0" box="[822,909,770,793]" pageId="46" pageNumber="45">L. Peña</collectorName>
(
<collectionCode id="7CD7AE26922B0D2E17801918B1CBFCC0" box="[924,1003,769,793]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="45" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE922B0D2E102B1918B0B3FCE0" collectingDate="1956-11-30" collectionCode="SEMC" country="Bolivia" location="Reyes Chitihuara" pageId="46" pageNumber="45" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D17673922B0D2E102B1918B6ADFCC0" box="[1079,1165,769,793]" name="Bolivia" pageId="46" pageNumber="45">Bolivia</collectingCountry>
:
<location id="1F196038922B0D2E14BC193BB34EFCE0" LSID="urn:lsid:plazi:treatment:926F87F592290D2E159A1898B001FC40:1F196038922B0D2E14BC193BB34EFCE0" box="[160,366,801,826]" country="Bolivia" name="Reyes Chitihuara" pageId="46" pageNumber="45">Reyes Chitihuara</location>
, 14/
<date id="6E781023922B0D2E15871938B00CFCE0" box="[411,556,801,825]" pageId="46" pageNumber="45" value="1956-11-30">
<collectingDate id="7E3CE9CB922B0D2E15871938B00CFCE0" box="[411,556,801,825]" pageId="46" pageNumber="45" value="1956-11-30">30 Nov 1956</collectingDate>
</date>
(
<collectionCode id="7CD7AE26922B0D2E16271938B0ABFCE0" box="[571,651,801,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="45" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922B0D2E14CC1958B001FC40" pageId="46" pageNumber="45" type="etymology">
<paragraph id="1A7936E3922B0D2E14CC1958B001FC40" blockId="46.[160,1175,225,922]" pageId="46" pageNumber="45">
ETYMOLOGY: The specific epithet is taken from the
<typeStatus id="C57D8841922B0D2E170C195BB164FC83" box="[784,836,834,858]" pageId="46" pageNumber="45">type</typeStatus>
locality, Yungas del Palmar. The name is formed from Yungas and the Latin suffix
<emphasis id="28B2EAF1922B0D2E1732197BB143FCA0" box="[814,867,866,889]" italics="true" pageId="46" pageNumber="45">ēnsis</emphasis>
, which is added to a top- onym to indicate “from” or “of”.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,242 @@
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<paragraph id="1A7936E3922A0D2F15D31F3BB141FAE0" blockId="47.[463,865,1313,1337]" box="[463,865,1313,1337]" pageId="47" pageNumber="46">
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<taxonomicName id="DDC64D60922A0D2F15D31F3BB141FAE0" ID-CoL="6XVTC" authority="(Buysson)" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[463,865,1313,1337]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF1922A0D2F15D31F3BB0C5FAE0" box="[463,741,1313,1337]" italics="true" pageId="47" pageNumber="46">Scaptotrigona ochrotricha</emphasis>
(Buysson)
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="52DC6568922A0D2F15961F58B186FA80" box="[394,934,1345,1369]" pageId="47" pageNumber="46" type="description">
<paragraph id="1A7936E3922A0D2F15961F58B186FA80" blockId="47.[394,934,1345,1369]" box="[394,934,1345,1369]" pageId="47" pageNumber="46">
(
<figureCitation id="82FD2A66922A0D2F158E1F58B036FA80" box="[402,534,1345,1369]" captionStart-0="Figures 6466" captionStart-1="Figures 6768" captionStartId-0="49.[160,241,1434,1456]" captionStartId-1="50.[160,241,935,957]" captionTargetBox-0="[173,1148,233,1411]" captionTargetBox-1="[160,1168,224,923]" captionTargetId-0="figure-61@49.[160,1168,223,1428]" captionTargetId-1="figure-266@50.[160,1168,224,929]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionText-0="Figures 6466. Worker of Scaptotrigona ochrotricha (Buysson). 64. Lateral habitus. 65. Dorsal habitus. 66. Facial view." captionText-1="Figures 6768. Worker of Scaptotrigona ochrotricha (Buysson). 67. Dorsal view of head and me- sosoma. 68. Dorsal view of metasoma." pageId="47" pageNumber="46">Figs. 6468</figureCitation>
; additional figures in
<bibRefCitation id="7E574B12922A0D2F170B1F58B1BFFA80" author="Engel, M. S." box="[791,927,1345,1369]" pageId="47" pageNumber="46" pagination="41 - 59" refId="ref21775" refString="Engel, M. S. 2022. Notes on South American stingless bees of the genus Scaptotrigona (Hymenop- tera: Apidae), part I: short-bristle species, the tubiba species group. Entomologist's Monthly Magazine 158 (1): 41 - 59." type="journal article" year="2022">Engel, 2022</bibRefCitation>
)
</paragraph>
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<treatmentCitationGroup id="3AD611CD922A0D2F14BC1F9DB10BFA40" box="[160,811,1411,1433]" pageId="47" pageNumber="46">
<taxonomicName id="DDC64D60922A0D2F14BC1F9DB3ECFA40" ID-CoL="58QXC" authority="Buysson" authorityName="Buysson" authorityYear="1892" box="[160,460,1411,1433]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ochrotrichus">
<emphasis id="28B2EAF1922A0D2F14BC1F9DB34CFA41" box="[160,364,1411,1433]" italics="true" pageId="47" pageNumber="46">Trigona ochrotrichus</emphasis>
Buysson
</taxonomicName>
in
<treatmentCitation id="9B6710F2922A0D2F15F11F9DB107FA40" author="Buysson, R. &amp; T. A. Marshall" box="[493,807,1411,1433]" page="54" pageId="47" pageNumber="46" year="1892">
<bibRefCitation id="7E574B12922A0D2F15F11F9DB107FA40" author="Buysson, R. &amp; T. A. Marshall" box="[493,807,1411,1433]" pageId="47" pageNumber="46" pagination="53 - 76" refId="ref21650" refString="Buysson, R., du, &amp; T. A. Marshall. 1892. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 17 e memoire (1) - Hymenopteres. Annales de la Societe Entomologique de France 61 (1): 53 - 76, + 2 pls. [pls. III, IV]." type="book chapter" year="1892">Buysson &amp; Marshall, 1892: 54</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1A7936E3922A0D2F14BC1FBAB0F7FA60" blockId="47.[160,887,1411,1529]" box="[160,727,1443,1465]" pageId="47" pageNumber="46">
<treatmentCitationGroup id="3AD611CD922A0D2F14BC1FBAB0F7FA60" box="[160,727,1443,1465]" pageId="47" pageNumber="46">
<taxonomicName id="DDC64D60922A0D2F14BC1FBAB3C0FA60" authority="(Buysson)" baseAuthorityName="Buysson" box="[160,480,1443,1465]" class="Insecta" family="Apidae" genus="Melipona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF1922A0D2F14BC1FBAB34FFA61" box="[160,367,1443,1464]" italics="true" pageId="47" pageNumber="46">Melipona ochrotricha</emphasis>
(Buysson)
</taxonomicName>
;
<treatmentCitation id="9B6710F2922A0D2F15F71FBAB0F3FA60" author="Dalla Torre, C. G." box="[491,723,1443,1465]" page="582" pageId="47" pageNumber="46" year="1896">
<bibRefCitation id="7E574B12922A0D2F15F71FBAB0F3FA60" author="Dalla Torre, C. G." box="[491,723,1443,1465]" pageId="47" pageNumber="46" refId="ref21720" refString="Dalla Torre, C. G., de [K. W., von]. 1896. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Volumen X: Apidae (Anthophila). Engelmann; Lipsiae [Leipzig], Deutsches Reich [German Empire]; viii + 643 pp." type="book" year="1896">Dalla Torre, 1896: 582</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1A7936E3922A0D2F14BC1FDDB168FA01" blockId="47.[160,887,1411,1529]" box="[160,840,1475,1497]" pageId="47" pageNumber="46">
<treatmentCitationGroup id="3AD611CD922A0D2F14BC1FDDB168FA01" box="[160,840,1475,1497]" pageId="47" pageNumber="46">
<taxonomicName id="DDC64D60922A0D2F14BC1FDDB051FA00" authority="(Buysson)" baseAuthorityName="Buysson" box="[160,625,1475,1497]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ochrotricha" subGenus="Nannotrigona">
<emphasis id="28B2EAF1922A0D2F14BC1FDDB2CDFA00" box="[160,237,1476,1497]" italics="true" pageId="47" pageNumber="46">Trigona</emphasis>
(
<emphasis id="28B2EAF1922A0D2F14E71FDDB3A4FA00" box="[251,388,1476,1497]" italics="true" pageId="47" pageNumber="46">Nannotrigona</emphasis>
)
<emphasis id="28B2EAF1922A0D2F158F1FDAB020FA01" box="[403,512,1475,1496]" italics="true" pageId="47" pageNumber="46">ochrotricha</emphasis>
(Buysson)
</taxonomicName>
;
<treatmentCitation id="9B6710F2922A0D2F16671FDAB164FA01" author="Schwarz, H. F." box="[635,836,1475,1497]" page="482" pageId="47" pageNumber="46" year="1938">
<bibRefCitation id="7E574B12922A0D2F16671FDAB164FA01" author="Schwarz, H. F." box="[635,836,1475,1497]" pageId="47" pageNumber="46" pagination="437 - 508" refId="ref22313" refString="Schwarz, H. F. 1938. The stingless bees (Meliponidae) of British Guiana and some related forms. Bulletin of the American Museum of Natural History 74 (7): 437 - 508, + 11 pls. [lii - lxii]." type="book chapter" year="1938">Schwarz, 1938: 482</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1A7936E3922A0D2F14BC1FFAB157FA20" blockId="47.[160,887,1411,1529]" box="[160,887,1507,1529]" pageId="47" pageNumber="46">
<treatmentCitationGroup id="3AD611CD922A0D2F14BC1FFAB157FA20" box="[160,887,1507,1529]" pageId="47" pageNumber="46">
<taxonomicName id="DDC64D60922A0D2F14BC1FFAB3D6FA20" authority="(Buysson)" baseAuthorityName="Buysson" baseAuthorityYear="1892" box="[160,502,1507,1529]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ochrotica">
<emphasis id="28B2EAF1922A0D2F14BC1FFAB3A5FA21" box="[160,389,1507,1528]" italics="true" pageId="47" pageNumber="46">Scaptotrigona ochrotica</emphasis>
(Buysson)
</taxonomicName>
;
<treatmentCitation id="9B6710F2922A0D2F161C1FFAB0CBFA20" author="Vit, P. &amp; P. Pulcini" box="[512,747,1507,1529]" page="58" pageId="47" pageNumber="46" year="1996">
<bibRefCitation id="7E574B12922A0D2F161C1FFAB0CBFA20" author="Vit, P. &amp; P. Pulcini" box="[512,747,1507,1529]" pageId="47" pageNumber="46" pagination="57 - 62" refId="ref22427" refString="Vit, P., &amp; P. Pulcini. 1996. Diastase and invertase activities in Meliponini and Trigonini honeys from Venezuela. Journal of Apicultural Research 35 (2): 57 - 62." type="journal article" year="1996">Vit &amp; Pulcini, 1996: 58</bibRefCitation>
</treatmentCitation>
,
<emphasis id="28B2EAF1922A0D2F16E91FFAB151FA21" box="[757,881,1507,1528]" italics="true" pageId="47" pageNumber="46">lapsis calami</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922A0D3014CC1C38B65AF975" lastPageId="48" lastPageNumber="47" pageId="47" pageNumber="46" type="diagnosis">
<paragraph id="1A7936E3922A0D2F14CC1C38B17EF960" blockId="47.[160,1168,1569,1785]" pageId="47" pageNumber="46">
DIAGNOSIS: The identity of this species has been in question for quite some time given that the original series is lost. The material reported here is within the greater vicinity of the type locality, is of the same caste, and matches the original description quite well. Accordingly, I have selected
<specimenCount id="0CC0FD6A922A0D2F16901C9BB134F940" box="[652,788,1665,1689]" count="1" pageId="47" pageNumber="46" type="worker">one worker</specimenCount>
to serve as
<typeStatus id="C57D8841922A0D2F17BB1C9BB626F943" box="[935,1030,1666,1690]" pageId="47" pageNumber="46" type="neotype">neotype</typeStatus>
in order to stabilize the application and circumscription of this species.
</paragraph>
<paragraph id="1A7936E3922A0D3014CC1CD8B00CF995" blockId="47.[160,1168,1569,1785]" lastBlockId="48.[160,1168,1524,1772]" lastPageId="48" lastPageNumber="47" pageId="47" pageNumber="46">
This rather robust species is most similar to
<taxonomicName id="DDC64D60922A0D2F16CC1CDBB179F900" box="[720,857,1729,1753]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF1922A0D2F16CC1CDBB179F900" box="[720,857,1729,1753]" italics="true" pageId="47" pageNumber="46">S. nigrohirta</emphasis>
</taxonomicName>
, as well as to
<taxonomicName id="DDC64D60922A0D2F17E41CDBB29CF920" authorityName="Engel" authorityYear="2022" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="macarenensis">
<emphasis id="28B2EAF1922A0D2F17E41CDBB29CF920" italics="true" pageId="47" pageNumber="46">S. macarenensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DDC64D60922A0D2F14D51CFBB349F920" authorityName="Engel" authorityYear="2022" box="[201,361,1762,1785]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="yungasensis">
<emphasis id="28B2EAF1922A0D2F14D51CFBB349F920" box="[201,361,1762,1785]" italics="true" pageId="47" pageNumber="46">S. yungasensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="DDC64D60922A0D2F15B51CFBB06EF920" authorityName="Moure" authorityYear="1950" box="[425,590,1761,1785]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="xanthotricha">
<emphasis id="28B2EAF1922A0D2F15B51CFBB06EF920" box="[425,590,1761,1785]" italics="true" pageId="47" pageNumber="46">S. xanthotricha</emphasis>
</taxonomicName>
, the assemblage perhaps forming a distinct clade in the postica group. From
<taxonomicName id="DDC64D6092350D3015F91FECB04FF9D5" box="[485,623,1524,1548]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF192350D3015F91FECB04FF9D5" box="[485,623,1524,1548]" italics="true" pageId="48" pageNumber="47">S. nigrohirta</emphasis>
</taxonomicName>
,
<taxonomicName id="DDC64D6092350D3016601FECB131F9D5" box="[636,785,1524,1548]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="ochrotricha">
<emphasis id="28B2EAF192350D3016601FECB131F9D5" box="[636,785,1524,1548]" italics="true" pageId="48" pageNumber="47">S. ochrotricha</emphasis>
</taxonomicName>
differs most notably in the pres- ence of distinct striping on the mesoscutum and the absence of dark fuscous to black bristles on metasomal terga
<collectionCode id="7CD7AE2692350D3015F71C2CB026F995" box="[491,518,1589,1612]" pageId="48" pageNumber="47">III</collectionCode>
<collectionCode id="7CD7AE2692350D3016091C2CB009F995" box="[533,553,1589,1612]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="48" pageNumber="47" type="Museum">V</collectionCode>
.
</paragraph>
<caption id="4EB9666B92350D3014BC1F81B045FA12" pageId="48" pageNumber="47" startId="48.[160,241,1432,1454]" targetBox="[177,1159,232,1414]" targetPageId="48" targetType="figure">
<paragraph id="1A7936E392350D3014BC1F81B045FA12" blockId="48.[160,1168,1432,1483]" pageId="48" pageNumber="47">
<emphasis id="28B2EAF192350D3014BC1F81B31BFA74" bold="true" box="[160,315,1432,1454]" pageId="48" pageNumber="47">Figures 6263.</emphasis>
Worker of
<taxonomicName id="DDC64D6092350D3015DD1F81B0B8FA74" authorityName="Engel" authorityYear="2022" box="[449,664,1432,1453]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="faviziae" status="sp. nov.">
<emphasis id="28B2EAF192350D3015DD1F81B0B8FA74" box="[449,664,1432,1453]" italics="true" pageId="48" pageNumber="47">Scaptotrigona faviziae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92350D3016BB1F81B10AFA77" box="[679,810,1432,1454]" pageId="48" pageNumber="47" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192350D3017211F81B17BFA74" bold="true" box="[829,859,1432,1453]" pageId="48" pageNumber="47">62.</emphasis>
Lateral view of head, arrow indicating genal tooth.
<emphasis id="28B2EAF192350D3015871FADB399FA10" bold="true" box="[411,441,1460,1481]" pageId="48" pageNumber="47">63.</emphasis>
Lateral habitus.
</paragraph>
</caption>
<paragraph id="1A7936E392350D3014CC1C4DB65AF975" blockId="48.[160,1168,1524,1772]" pageId="48" pageNumber="47">
<materialsCitation id="AAAE3CBE92350D3014CC1C4DB65AF975" collectingDate="1990-02-05" collectingDateMax="1990-02-07" collectingDateMin="1990-02-05" collectionCode="SEMC" collectorName="C. D. Michener" country="Venezuela" elevation="500" latitude="6.3333335" location="Rancho Las" longLatPrecision="1304" longitude="-66.833336" pageId="48" pageNumber="47" specimenCount="2" stateProvince="Bolivar" typeStatus="neotype">
<typeStatus id="C57D884192350D3014CC1C4DB314F9B5" box="[208,308,1620,1644]" pageId="48" pageNumber="47" type="neotype">NEOTYPE</typeStatus>
(
<emphasis id="28B2EAF192350D3015511C4DB3D1F9B5" box="[333,497,1620,1644]" italics="true" pageId="48" pageNumber="47">hic designatus</emphasis>
): ⚲,
<collectingCountry id="62D1767392350D3016291C4DB091F9B5" box="[565,689,1620,1644]" name="Venezuela" pageId="48" pageNumber="47">Venezuela</collectingCountry>
,
<collectingRegion id="D802F80192350D3016D91C4DB13CF9B5" box="[709,796,1620,1644]" country="Venezuela" name="Bolivar" pageId="48" pageNumber="47">Bolivar</collectingRegion>
,
<location id="1F19603892350D30172C1C4DB1E1F9B5" LSID="urn:lsid:plazi:treatment:926F87F5922A0D3015D31F3BB17CF935:1F19603892350D30172C1C4DB1E1F9B5" box="[816,961,1620,1644]" country="Venezuela" latitude="6.3333335" longLatPrecision="1304" longitude="-66.833336" name="Rancho Las" pageId="48" pageNumber="47" stateProvince="Bolivar">Rancho Las</location>
Nieves,
<geoCoordinate id="7FF2502492350D3010251C4DB6ACF9B5" box="[1081,1164,1620,1644]" degrees="6" direction="north" minutes="20" orientation="latitude" pageId="48" pageNumber="47" precision="925" value="6.3333335">6°20N</geoCoordinate>
,
<geoCoordinate id="7FF2502492350D3014BC1C6DB324F955" box="[160,260,1652,1676]" degrees="66" direction="west" minutes="50" orientation="longitude" pageId="48" pageNumber="47" precision="925" value="-66.833336">66°50W</geoCoordinate>
,
<quantity id="DD3E9B0692350D30150D1C6DB37BF955" box="[273,347,1652,1676]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="47" unit="m" value="500.0">
<elevation id="91EBD1D092350D30150D1C6DB37BF955" box="[273,347,1652,1676]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="47" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="6E78102392350D3015741C6DB064F955" box="[360,580,1652,1676]" pageId="48" pageNumber="47" value="1990-02-05" valueMax="1990-02-07" valueMin="1990-02-05">
<collectingDate id="7E3CE9CB92350D3015741C6DB064F955" box="[360,580,1652,1676]" pageId="48" pageNumber="47" value="1990-02-05" valueMax="1990-02-07" valueMin="1990-02-05">57 February 1990</collectingDate>
</date>
,
<collectorName id="B733533592350D30164D1C6DB0DEF955" box="[593,766,1652,1676]" pageId="48" pageNumber="47">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692350D3017121C6DB17DF955" box="[782,861,1652,1676]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="48" pageNumber="47" type="Museum">SEMC</collectionCode>
); the
<typeStatus id="C57D884192350D3017BC1C6CB1DFF954" box="[928,1023,1653,1677]" pageId="48" pageNumber="47" type="neotype">neotype</typeStatus>
is here des- ignated for the express purpose of stabilizing the application of this specific epithet.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892350D3014CC1CADB17CF935" pageId="48" pageNumber="47" type="description">
<paragraph id="1A7936E392350D3014CC1CADB17CF935" blockId="48.[160,1168,1524,1772]" pageId="48" pageNumber="47">
<materialsCitation id="AAAE3CBE92350D3014CC1CADB178F935" collectingDate="1990-02-05" collectingDateMax="1990-02-07" collectingDateMin="1990-02-05" collectionCode="SEMC" collectorName="C. D. Michener" country="Venezuela" elevation="500" latitude="6.3333335" location="Rancho Las" longLatPrecision="1304" longitude="-66.833336" pageId="48" pageNumber="47" specimenCount="1" stateProvince="Bolivar">
ADDITIONAL MATERIAL: 24⚲⚲,
<collectingCountry id="62D1767392350D3016271CADB097F915" box="[571,695,1716,1740]" name="Venezuela" pageId="48" pageNumber="47">Venezuela</collectingCountry>
,
<collectingRegion id="D802F80192350D3016D61CADB101F915" box="[714,801,1716,1740]" country="Venezuela" name="Bolivar" pageId="48" pageNumber="47">Bolivar</collectingRegion>
,
<location id="1F19603892350D3017281CADB1E4F915" LSID="urn:lsid:plazi:treatment:926F87F5922A0D3015D31F3BB17CF935:1F19603892350D3017281CADB1E4F915" box="[820,964,1716,1740]" country="Venezuela" latitude="6.3333335" longLatPrecision="1304" longitude="-66.833336" name="Rancho Las" pageId="48" pageNumber="47" stateProvince="Bolivar">Rancho Las</location>
Nieves,
<geoCoordinate id="7FF2502492350D3010251CADB6ACF915" box="[1081,1164,1716,1740]" degrees="6" direction="north" minutes="20" orientation="latitude" pageId="48" pageNumber="47" precision="925" value="6.3333335">6°20N</geoCoordinate>
,
<geoCoordinate id="7FF2502492350D3014BC1CCDB324F935" box="[160,260,1748,1772]" degrees="66" direction="west" minutes="50" orientation="longitude" pageId="48" pageNumber="47" precision="925" value="-66.833336">66°50W</geoCoordinate>
,
<quantity id="DD3E9B0692350D3015131CCDB378F935" box="[271,344,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="47" unit="m" value="500.0">
<elevation id="91EBD1D092350D3015131CCDB378F935" box="[271,344,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="47" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="6E78102392350D30157E1CCDB01BF935" box="[354,571,1748,1772]" pageId="48" pageNumber="47" value="1990-02-05" valueMax="1990-02-07" valueMin="1990-02-05">
<collectingDate id="7E3CE9CB92350D30157E1CCDB01BF935" box="[354,571,1748,1772]" pageId="48" pageNumber="47" value="1990-02-05" valueMax="1990-02-07" valueMin="1990-02-05">57 February 1990</collectingDate>
</date>
,
<collectorName id="B733533592350D30165A1CCDB0D2F935" box="[582,754,1748,1772]" pageId="48" pageNumber="47">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692350D30171C1CCDB170F935" box="[768,848,1748,1772]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="48" pageNumber="47" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,180 @@
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<mods:title id="E597B822BE1CD0095400C6094AD867E5">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="FFAFA299B3A15EFD676FBDBD2073D38A">Engel, Michael S.</mods:namePart>
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<heading id="4131818F922B0D2E158219D8B1B2FC00" box="[414,914,961,985]" centered="true" fontSize="10" level="2" pageId="46" pageNumber="45" reason="2">
<taxonomicName id="DDC64D60922B0D2E158219D8B0DBFC00" authority="Engel" authorityName="Engel" authorityYear="2022" box="[414,763,961,985]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="faviziae" status="sp. nov.">
<emphasis id="28B2EAF1922B0D2E158219D8B08EFC00" bold="true" box="[414,686,961,985]" italics="true" pageId="46" pageNumber="45">Scaptotrigona faviziae</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A922B0D2E171919DBB1B2FC00" box="[773,914,961,985]" pageId="46" pageNumber="45" rank="species">new species</taxonomicNameLabel>
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ZooBank:
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urn:lsid:zoobank.org:act:
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<paragraph id="1A7936E3922B0D2E16531E18B0C1FBC0" blockId="46.[246,1082,995,1049]" box="[591,737,1025,1049]" pageId="46" pageNumber="45">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922B0D2E14CC1E58B36AFB20" pageId="46" pageNumber="45" type="diagnosis">
<paragraph id="1A7936E3922B0D2E14CC1E58B36AFB20" blockId="46.[160,1168,1089,1786]" pageId="46" pageNumber="45">
DIAGNOSIS: This species is noteworthy for the weaker development and coverage of the yellow tomentum on metasomal terga
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, with the tomentum widely inter- rupted medially and weaker laterally. In addition, the size of the concavity interrupt- ing the preoccipital carina laterally is exceptional in this species. Usually, the length of the concavity is about 4× its depth or a bit less, but in
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<emphasis id="28B2EAF1922B0D2E17081EDBB1A1FB00" box="[788,897,1217,1241]" italics="true" pageId="46" pageNumber="45">S. faviziae</emphasis>
</taxonomicName>
, the concavitys length is 5× its depth.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922B0D2F14CC1F18B30FFB80" lastPageId="47" lastPageNumber="46" pageId="46" pageNumber="45" type="description">
<paragraph id="1A7936E3922B0D2E14CC1F18B08FF980" blockId="46.[160,1168,1089,1786]" pageId="46" pageNumber="45">
DESCRIPTION: As described for
<taxonomicName id="DDC64D60922B0D2E165F1F1BB0B6FAC0" authorityName="Engel" authorityYear="2022" box="[579,662,1281,1305]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922B0D2E165F1F1BB0B6FAC0" box="[579,662,1281,1305]" italics="true" pageId="46" pageNumber="45">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF1922B0D2E16B51F18B13CFAC0" box="[681,796,1281,1305]" italics="true" pageId="46" pageNumber="45">vide supra</emphasis>
), with the following modifica- tions: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite) 6.0 mm. Head wider than long, width
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, length
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; compound eye length
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; upper interorbital distance
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, lower interorbital distance
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. Scape length
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, about as long as torulocellar distance, torulocellar distance
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. Clypeus approximately 1.7× as wide as long, length
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, width
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. Malar area approximately 1.8× flagellar diameter or 0.80× basal mandibu- lar width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, carina interrupted laterally by deep and unusually long concavity, lower margin of concavity with carina forming sharply acute angle, angle projecting upward as a large lamellate tooth.
</paragraph>
<paragraph id="1A7936E3922B0D2F14CC1C7BB6B0FE20" blockId="46.[160,1168,1089,1786]" lastBlockId="47.[160,1168,225,1273]" lastPageId="47" lastPageNumber="46" pageId="46" pageNumber="45">
Integument of labiomaxillary complex orange; labrum orange; mandible yellow orange basally, blending to reddish orange apically; clypeus yellow orange, epistomal sulcus delineated by orange; supraclypeal area yellow orange; malar space yellow or- ange; face below tangent of antennal toruli yellow orange; scape dark brown except yellow orange ventrally; pedicel and flagellum dark brown; face above tangent of an- tennal toruli reddish brown; vertex and posterior of head reddish brown; gena largely reddish brown except bordering malar space yellow to yellow orange; postgena yel- low. Mesosoma largely orange to reddish orange except mesoscutum black, mesepi- sternum anteriorly and propodeum reddish orange; tegula yellow orange and semi-translucent except brown on posterior quarter; legs largely reddish orange to brown except coxae, trochanters, underside of metafemur, and apicalmost tarsomeres lighter, yellow orange to yellow. Wing membranes clear to lightly parchment colored; veins orange. Metasoma generally reddish brown to dark brown except anterior-facing sur- face of tergum I orange, and terga I
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apically yellow orange to yellow; sterna brown.
</paragraph>
<paragraph id="1A7936E3922A0D2F14CC181BB153FBC0" blockId="47.[160,1168,225,1273]" pageId="47" pageNumber="46">
Areas of short, white, plumose setae in
<taxonomicName id="DDC64D60922A0D2F16AB181BB12AFDC0" authorityName="Engel" authorityYear="2022" box="[695,778,513,537]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922A0D2F16AB181BB12AFDC0" box="[695,778,513,537]" italics="true" pageId="47" pageNumber="46">S. ederi</emphasis>
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instead white to pale yellow or golden; metasomal terga II
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with apical fimbriae of minute, fine, simple, golden se- tae; terga
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with dense, plumose, appressed, yellow tomentum absent mediolon- gitudinally. Vertex with numerous erect, simple, black bristles; pronotal dorsolateral margin near and anterior to pronotal lobe with fulvous to black bristles; mesoscutum anterior and lateral borders with black bristles, disc with sparsely scatered, minute, erect, fulvous bristles; tegula with suberect fulvous to black bristles anteriorly; mesos- cutellum with abundant, elongate, erect yellow to fulvous bristles, except a few black bristles basolaterally; mesepisternum with scatered, erect to subdecumbent yellow to fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, suberect fulvous bristles extending ventrally from pronotal lobe, a few dorsally near lobe black; lateral surface of propodeum with scatered, long, erect, yellow fulvous bristles. Legs with abundant fulvous bristles except intermixed with black bristles on protibia apically, prolateral surface of mesotibia, pro- and mesobasi- tarsi, and metafemur apically; metatibia and metabasitarsus with almost entirely black bristles, some with fulvous apices. Metasoma with numerous, prominent, suberect to subdecumbent, dark fulvous to yellow bristles on terga
<collectionCode id="7CD7AE26922A0D2F172D1E1BB16CFBC0" box="[817,844,1026,1049]" pageId="47" pageNumber="46">III</collectionCode>
<collectionCode id="7CD7AE26922A0D2F17471E1BB14FFBC0" box="[859,879,1026,1049]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="47" pageNumber="46" type="Museum">V</collectionCode>
.
</paragraph>
<paragraph id="1A7936E3922A0D2F14CC1E3BB30FFBE0" blockId="47.[160,1168,225,1273]" box="[208,303,1058,1084]" pageId="47" pageNumber="46">
<specimenCount id="0CC0FD6A922A0D2F14CC1E3BB2C4FBE5" box="[208,228,1058,1084]" count="1" pageId="47" pageNumber="46" type="female"></specimenCount>
:
<emphasis id="28B2EAF1922A0D2F14ED1E3BB308FBE0" box="[241,296,1058,1081]" italics="true" pageId="47" pageNumber="46">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E3922A0D2F14CC1E5BB30FFB80" blockId="47.[160,1168,225,1273]" box="[208,303,1090,1116]" pageId="47" pageNumber="46">
<specimenCount id="0CC0FD6A922A0D2F14CC1E5BB2C4FB85" box="[208,228,1090,1116]" count="1" pageId="47" pageNumber="46" type="male"></specimenCount>
:
<emphasis id="28B2EAF1922A0D2F14ED1E5BB308FB80" box="[241,296,1090,1113]" italics="true" pageId="47" pageNumber="46">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922A0D2F14CC1E78B648FBA0" box="[208,1128,1121,1146]" pageId="47" pageNumber="46" type="materials_examined">
<paragraph id="1A7936E3922A0D2F14CC1E78B648FBA0" blockId="47.[160,1168,225,1273]" box="[208,1128,1121,1146]" pageId="47" pageNumber="46">
<materialsCitation id="AAAE3CBE922A0D2F14CC1E78B648FBA0" box="[208,1128,1121,1146]" collectionCode="INPA, SEMC" collectorName="D. Roubik" country="Brazil" location="Amazonas" municipality="Manaus" pageId="47" pageNumber="46" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D8841922A0D2F14CC1E78B368FBA0" box="[208,328,1121,1145]" pageId="47" pageNumber="46" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D17673922A0D2F15711E78B395FBA0" box="[365,437,1121,1145]" name="Brazil" pageId="47" pageNumber="46">Brazil</collectingCountry>
:
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,
<collectingMunicipality id="FA1DAC99922A0D2F16561E7BB08BFBA0" box="[586,683,1122,1145]" pageId="47" pageNumber="46">Manaus</collectingMunicipality>
,
<collectionCode id="7CD7AE26922A0D2F16AA1E7BB0DBFBA3" box="[694,763,1122,1146]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="47" pageNumber="46">INPA</collectionCode>
, 15 Aug 76,
<collectorName id="B7335335922A0D2F179A1E7BB1DEFBA0" box="[902,1022,1121,1145]" pageId="47" pageNumber="46">D. Roubik</collectorName>
(
<collectionCode id="7CD7AE26922A0D2F10101E78B67CFBA0" box="[1036,1116,1121,1145]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="47" pageNumber="46" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922A0D2F14CC1E98B65CFB20" pageId="47" pageNumber="46" type="etymology">
<paragraph id="1A7936E3922A0D2F14CC1E98B65CFB20" blockId="47.[160,1168,225,1273]" pageId="47" pageNumber="46">
ETYMOLOGY: The specific epithet honors Favízia Freitas de Oliveira, dear friend and renowned specialist of Meliponini. The name combines her forename and the Latin suffix
<emphasis id="28B2EAF1922A0D2F14EA1EDBB32FFB00" box="[246,271,1218,1241]" italics="true" pageId="47" pageNumber="46">us</emphasis>
(
<emphasis id="28B2EAF1922A0D2F15371EDBB361FB00" box="[299,321,1218,1241]" italics="true" pageId="47" pageNumber="46">ae</emphasis>
, feminine genitive singular;
<emphasis id="28B2EAF1922A0D2F16831EDBB086FB00" box="[671,678,1218,1241]" italics="true" pageId="47" pageNumber="46">i</emphasis>
, masculine and neuter genitive singular), which forms adjectives from nouns and is used for commemoration and dedication.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,280 @@
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<mods:title id="0C897C40176D090A25F5648824DCE073">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="F728073D821F7B3F83035A7ACB05002E">Engel, Michael S.</mods:namePart>
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<mods:title id="387ECF25B2B5D90CE2A27727E6CCEF62">Journal of Melittology</mods:title>
</mods:titleInfo>
<mods:part id="08B5371A67F628FFEA6090A1B27AABBC">
<mods:date id="85F4BD346B0C183DAFB76456D8718FB7">2022</mods:date>
<mods:detail id="D44FB10C26A91D9478892BE575B41004" type="pubDate">
<mods:number id="C13906F8E5316E1EA6106FF9AC72E391">2022-03-14</mods:number>
</mods:detail>
<mods:detail id="A2CFF492EEE96DE4BE7FF68754C02F79" type="volume">
<mods:number id="19A4B29699A3B602261A05CCBE45FE27">2022</mods:number>
</mods:detail>
<mods:detail id="7E182CBA766DA201900E655FD3621509" type="issue">
<mods:number id="129F137133C613674362A5CCBC998D6B">110</mods:number>
</mods:detail>
<mods:extent id="2C7A96B93D523FB5ACCAFE977930CE56" unit="page">
<mods:start id="64B50C719C21EA505A4181A3C056A690">1</mods:start>
<mods:end id="8B8CE7C51CEE6107699B7A4FE4484F0C">51</mods:end>
</mods:extent>
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<mods:location id="886BB2BC4C00EF3BA78D8DCB5F3E0F70">
<mods:url id="89432DED687F40D818CC997B281E1788">http://dx.doi.org/10.17161/jom.i110.17001</mods:url>
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<mods:classification id="E3D955E90B765BA76B485B61EB32E1BB">journal article</mods:classification>
<mods:identifier id="D44ED0E3E0C66A9AD9A2A7EEC9761BDE" type="DOI">10.17161/jom.i110.17001</mods:identifier>
<mods:identifier id="D40A329F7ACB94325F5E4FAC4F1EA506" type="ISSN">2325-4467</mods:identifier>
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<treatment id="926F87F5922D0D2B15841C18B1A8FEC0" LSID="urn:lsid:plazi:treatment:926F87F5922D0D2B15841C18B1A8FEC0" httpUri="http://treatment.plazi.org/id/926F87F5922D0D2B15841C18B1A8FEC0" lastPageId="43" lastPageNumber="41" pageId="40" pageNumber="39">
<subSubSection id="52DC6568922D0D2815841C18B665F9E0" pageId="40" pageNumber="39" type="nomenclature">
<paragraph id="1A7936E3922D0D2815841C18B1B8F9C0" blockId="40.[408,920,1537,1561]" box="[408,920,1537,1561]" pageId="40" pageNumber="39">
<heading id="4131818F922D0D2815841C18B1B8F9C0" box="[408,920,1537,1561]" centered="true" fontSize="10" level="2" pageId="40" pageNumber="39" reason="2">
<taxonomicName id="DDC64D60922D0D2815841C18B120F9C0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[408,768,1537,1561]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="baldwini" status="sp. nov.">
<emphasis id="28B2EAF1922D0D2815841C18B094F9C0" bold="true" box="[408,692,1537,1561]" italics="true" pageId="40" pageNumber="39">Scaptotrigona baldwini</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A922D0D2817171C1BB1B8F9C0" box="[779,920,1537,1561]" pageId="40" pageNumber="39" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E3922D0D2814F71C3AB665F9E0" blockId="40.[235,1093,1571,1625]" box="[235,1093,1571,1593]" pageId="40" pageNumber="39">
ZooBank:
<uri id="6E573AE1922D0D28154B1C3AB665F9E0" box="[343,1093,1571,1593]" pageId="40" pageNumber="39">
urn:lsid:zoobank.org:act:
<uuid id="6E600C36922D0D2816551C3AB665F9E0" box="[585,1093,1571,1593]" pageId="40" pageNumber="39">875A20FC-7ECC-46CA-BA97-924EADF05EB5</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922D0D2816741C58B0E8F980" box="[616,712,1601,1625]" pageId="40" pageNumber="39" type="description">
<paragraph id="1A7936E3922D0D2816741C58B0E8F980" blockId="40.[235,1093,1571,1625]" box="[616,712,1601,1625]" pageId="40" pageNumber="39">
(
<figureCitation id="82FD2A66922D0D28166D1C58B0E0F980" box="[625,704,1601,1625]" captionStart="Figures 5758" captionStartId="42.[160,241,1307,1329]" captionTargetBox="[184,1136,240,1284]" captionTargetId="figure-130@42.[160,1168,224,1301]" captionTargetPageId="42" captionText="Figures 5758. Workers of Scaptotrigona Moure. 57. Lateral habitus of Scaptotrigona baldwini, new species. 58. Lateral habitus of S. santiago, new species." pageId="40" pageNumber="39">Fig. 57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922D0D2914CC1C98B375FEE0" lastPageId="41" lastPageNumber="40" pageId="40" pageNumber="39" type="diagnosis">
<paragraph id="1A7936E3922D0D2914CC1C98B375FEE0" blockId="40.[160,1175,1665,1786]" lastBlockId="41.[160,1169,225,1786]" lastPageId="41" lastPageNumber="40" pageId="40" pageNumber="39">
DIAGNOSIS: This is a noticeably yellow species (
<figureCitation id="82FD2A66922D0D2817191C98B178F940" box="[773,856,1665,1689]" captionStart="Figures 5758" captionStartId="42.[160,241,1307,1329]" captionTargetBox="[184,1136,240,1284]" captionTargetId="figure-130@42.[160,1168,224,1301]" captionTargetPageId="42" captionText="Figures 5758. Workers of Scaptotrigona Moure. 57. Lateral habitus of Scaptotrigona baldwini, new species. 58. Lateral habitus of S. santiago, new species." pageId="40" pageNumber="39">Fig. 57</figureCitation>
), and generally resembles species such as
<taxonomicName id="DDC64D60922D0D28157D1CBBB3CEF960" box="[353,494,1697,1721]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF1922D0D28157D1CBBB3CEF960" box="[353,494,1697,1721]" italics="true" pageId="40" pageNumber="39">S. nigrohirta</emphasis>
</taxonomicName>
and
<taxonomicName id="DDC64D60922D0D2816331CBBB0F8F960" authorityName="Moure" authorityYear="1950" box="[559,728,1697,1721]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="xanthotricha">
<emphasis id="28B2EAF1922D0D2816331CBBB0F8F960" box="[559,728,1697,1721]" italics="true" pageId="40" pageNumber="39">S. xanthotricha</emphasis>
</taxonomicName>
, both of which are typically darker. From
<taxonomicName id="DDC64D60922D0D2814FB1CDBB352F900" box="[231,370,1729,1753]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF1922D0D2814FB1CDBB352F900" box="[231,370,1729,1753]" italics="true" pageId="40" pageNumber="39">S. nigrohirta</emphasis>
</taxonomicName>
it is most easily distinguished by the presence of yellow bristles on metasomal terga
<collectionCode id="7CD7AE26922D0D2815761CFBB3A5F920" box="[362,389,1762,1785]" pageId="40" pageNumber="39">III</collectionCode>
<collectionCode id="7CD7AE26922D0D28158F1CFBB386F920" box="[403,422,1762,1785]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="39" type="Museum">V</collectionCode>
(rather than the dark fuscous bristles of
<taxonomicName id="DDC64D60922D0D28179F1CFBB62DF920" box="[899,1037,1761,1785]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrohirta">
<emphasis id="28B2EAF1922D0D28179F1CFBB62DF920" box="[899,1037,1761,1785]" italics="true" pageId="40" pageNumber="39">S. nigrohirta</emphasis>
</taxonomicName>
), and from
<taxonomicName id="DDC64D60922C0D2914BC1AFBB368FF20" authorityName="Moure" authorityYear="1950" box="[160,328,225,249]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="xanthotricha">
<emphasis id="28B2EAF1922C0D2914BC1AFBB368FF20" box="[160,328,225,249]" italics="true" pageId="41" pageNumber="40">S. xanthotricha</emphasis>
</taxonomicName>
by the presence of yellow bristles on the tegula and metasomal terga
<collectionCode id="7CD7AE26922C0D2914BC1B1BB29BFEC0" box="[160,187,258,281]" pageId="41" pageNumber="40">III</collectionCode>
<collectionCode id="7CD7AE26922C0D2914D51B1BB2FCFEC0" box="[201,220,258,281]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="41" pageNumber="40" type="Museum">V</collectionCode>
(fuscous to black on the tegula, and fulvous to dark fulvous on the metasoma in
<taxonomicName id="DDC64D60922C0D2914BC1B3BB365FEE0" authorityName="Moure" authorityYear="1950" box="[160,325,289,313]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="xanthotricha">
<emphasis id="28B2EAF1922C0D2914BC1B3BB365FEE0" box="[160,325,289,313]" italics="true" pageId="41" pageNumber="40">S. xanthotricha</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922C0D2A14CC1B58B30FFA69" lastPageId="42" lastPageNumber="41" pageId="41" pageNumber="40" type="description">
<paragraph id="1A7936E3922C0D2914CC1B58B189FD40" blockId="41.[160,1169,225,1786]" pageId="41" pageNumber="40">
DESCRIPTION: As described for
<taxonomicName id="DDC64D60922C0D2916331B5BB05DFE80" authorityName="Engel" authorityYear="2022" box="[559,637,321,345]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922C0D2916331B5BB05DFE80" box="[559,637,321,345]" italics="true" pageId="41" pageNumber="40">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF1922C0D2916971B58B0D9FE80" box="[651,761,321,345]" italics="true" pageId="41" pageNumber="40">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately 6.0
<quantity id="DD3E9B06922C0D2916611B78B0F7FEA0" box="[637,727,353,377]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="41" pageNumber="40" unit="mm" value="6.5">6.5 mm</quantity>
, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B06922C0D29151B1B98B3B3FE40" box="[263,403,385,409]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.4" metricValueMin="5.9" pageId="41" pageNumber="40" unit="mm" value="6.15" valueMax="6.4" valueMin="5.9">5.96.4 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B06922C0D2917071B98B1E1FE40" box="[795,961,385,409]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" metricValueMax="2.67" metricValueMin="2.51" pageId="41" pageNumber="40" unit="mm" value="2.59" valueMax="2.67" valueMin="2.51">2.512.67 mm</quantity>
, length
<quantity id="DD3E9B06922C0D2910391B98B2F1FE60" metricMagnitude="-3" metricUnit="m" metricValue="2.195" metricValueMax="2.24" metricValueMin="2.15" pageId="41" pageNumber="40" unit="mm" value="2.195" valueMax="2.24" valueMin="2.15">2.152.24 mm</quantity>
; compound eye length
<quantity id="DD3E9B06922C0D2915F11BB8B0B1FE60" box="[493,657,417,441]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="41" pageNumber="40" unit="mm" value="1.5" valueMax="1.55" valueMin="1.45">1.451.55 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06922C0D2917FB1BB8B6ABFE60" box="[999,1163,417,441]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.79" metricValueMin="1.61" pageId="41" pageNumber="40" unit="mm" value="1.7" valueMax="1.79" valueMin="1.61">1.611.79 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06922C0D2915C71BD8B05EFE00" box="[475,638,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.6099999999999999" metricValueMax="1.7" metricValueMin="1.52" pageId="41" pageNumber="40" unit="mm" value="1.6099999999999999" valueMax="1.7" valueMin="1.52">1.521.70 mm</quantity>
. Scape length
<quantity id="DD3E9B06922C0D2917351BD8B1EAFE00" box="[809,970,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.03" metricValueMin="0.97" pageId="41" pageNumber="40" unit="mm" value="1.0" valueMax="1.03" valueMin="0.97">0.971.03 mm</quantity>
, about as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B06922C0D29168F1BF8B116FE20" box="[659,822,481,505]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" metricValueMax="10.3" metricValueMin="9.1" pageId="41" pageNumber="40" unit="mm" value="0.97" valueMax="1.03" valueMin="0.91">0.911.03 mm</quantity>
. Clypeus approximately 1.7× as wide as long, length
<quantity id="DD3E9B06922C0D2915AE1818B074FDC0" box="[434,596,513,537]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.3" metricValueMin="6.7" pageId="41" pageNumber="40" unit="mm" value="0.7" valueMax="0.73" valueMin="0.67">0.670.73 mm</quantity>
, width
<quantity id="DD3E9B06922C0D2916B71818B16EFDC0" box="[683,846,513,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.24" metricValueMin="1.16" pageId="41" pageNumber="40" unit="mm" value="1.2" valueMax="1.24" valueMin="1.16">1.161.24 mm</quantity>
. Malar area approximately 2.22.4× flagellar diameter or 0.960.98× basal mandibular width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lateral indentations deep, ca- rina interrupted laterally by deep concavity, lower margin of concavity with carina forming acute angle projecting upward as a sharp lamellate tooth.
</paragraph>
<paragraph id="1A7936E3922C0D2914CC18BBB12FFAC0" blockId="41.[160,1169,225,1786]" pageId="41" pageNumber="40">
Integument of labiomaxillary complex yellow; labrum yellow; mandible yellow orange to orange; clypeus largely yellow to yellow orange, epistomal sulcus well de- lineated by yellow brown to dark brown; supraclypeal area yellow to yellow orange; malar space yellow to yellow orange; face below tangent of antennal toruli yellow or- ange; scape yellow to yellow orange except with patch of brown dorsoapically; pedicel and flagellum brown, with narrow underside of flagellum orange; lowermost frons as on face below antennal toruli, blending above to reddish brown and brown, vertex and ocellocular as on upper frons, except sometimes with small spot of orange in ocel- locular area bordering upper orbit; posterior of as on upper frons; uppermost edge of gena as on vertex, remainder of gena orange; postgena orange to yellow orange. Me- sosoma largely orange except mesoscutum black, mesepisternum with large patch of dark brown ventroanteriorly, and sometimes brown on lateral margins of basal area of propodeum; tegula wholly yellow orange and semi-translucent; legs yellow orange to orange. Wing membranes lightly parchment colored except with noticeable infumate patch at apex of marginal cell; veins orange. Metasoma generally dark brown except anterior-facing surface of tergum I orange, dorsal-facing disc of tergum I largely yel- low medially and posteriorly, with lateral area dark brown; tergum II with large me- dial area of yellow and laterally with broad dark brown patches tapering mesally, api- cal marginal zone yellow; terga
<collectionCode id="7CD7AE26922C0D2916071EFBB016FB20" box="[539,566,1250,1273]" pageId="41" pageNumber="40">III</collectionCode>
<collectionCode id="7CD7AE26922C0D2916581EFBB077FB20" box="[580,599,1250,1273]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="41" pageNumber="40" type="Museum">V</collectionCode>
dark brown with yellow apical marginal zones; tergum
<collectionCode id="7CD7AE26922C0D2914E11F1BB339FAC0" box="[253,281,1282,1305]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="41" pageNumber="40">VI</collectionCode>
dark brown; sterna orange to light brown.
</paragraph>
<paragraph id="1A7936E3922C0D2914CC1F3BB62CF920" blockId="41.[160,1169,225,1786]" pageId="41" pageNumber="40">
Vertex with numerous erect, simple, yellow to fulvous bristles; pronotal dorsolat- eral margin near and anterior to pronotal lobe with yellow to fulvous bristles; mesos- cutum anterior and lateral borders with yellow to fulvous bristles; tegula with suber- ect yellow bristles anteriorly; mesoscutellum with abundant, elongate, erect yellow bristles posteriorly; mesepisternum with scatered, erect to subdecumbent yellow to fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, suberect to subdecumbent fulvous bristles extending ventrally from pronotal lobe; lateral surface of propodeum with scatered, long, erect, yellow bristles emerging amid dense, appressed, yellow, plumose setae. Legs with abundant yellow to fulvous bristles except corbicular bristles dark fuscous to black contrasting with yellow marginal bristles, sometimes with a few black bristles intermixed on margins; metabasitarsus with yellow bristles except sometimes with a few dark fulvous to black bristles intermixed on proventral margin, but prolateral surface uniformly with fine, erect, simple, yellow setae. Metasoma with numerous, prominent, suberect to subde- cumbent, yellow to fulvous bristles on terga
<collectionCode id="7CD7AE26922C0D2916B01CFBB0E7F920" box="[684,711,1762,1785]" pageId="41" pageNumber="40">III</collectionCode>
<collectionCode id="7CD7AE26922C0D2916C91CFBB0C8F920" box="[725,744,1762,1785]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="41" pageNumber="40" type="Museum">V</collectionCode>
arising amid tomentum.
</paragraph>
<caption id="4EB9666B922F0D2A14BC1F02B132FA97" pageId="42" pageNumber="41" startId="42.[160,241,1307,1329]" targetBox="[184,1136,240,1284]" targetPageId="42" targetType="figure">
<paragraph id="1A7936E3922F0D2A14BC1F02B132FA97" blockId="42.[160,1168,1307,1358]" pageId="42" pageNumber="41">
<emphasis id="28B2EAF1922F0D2A14BC1F02B31BFAE8" bold="true" box="[160,315,1307,1329]" pageId="42" pageNumber="41">Figures 5758.</emphasis>
Workers of
<taxonomicName id="DDC64D60922F0D2A15D01F05B08AFAE8" authority="Moure." authorityName="Moure." box="[460,682,1308,1329]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF1922F0D2A15D01F05B073FAE8" box="[460,595,1308,1329]" italics="true" pageId="42" pageNumber="41">Scaptotrigona</emphasis>
Moure.
</taxonomicName>
<emphasis id="28B2EAF1922F0D2A16A51F05B0F8FAE8" bold="true" box="[697,728,1308,1329]" pageId="42" pageNumber="41">57.</emphasis>
Lateral habitus of
<taxonomicName id="DDC64D60922F0D2A17BA1F05B6AAFAE8" authorityName="Engel" authorityYear="2022" box="[934,1162,1308,1329]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="baldwini" status="sp. nov.">
<emphasis id="28B2EAF1922F0D2A17BA1F05B6AAFAE8" box="[934,1162,1308,1329]" italics="true" pageId="42" pageNumber="41">Scaptotrigona baldwini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A922F0D2A14BC1F20B302FA97" box="[160,290,1336,1358]" pageId="42" pageNumber="41" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1922F0D2A152D1F21B36FFA94" bold="true" box="[305,335,1336,1357]" pageId="42" pageNumber="41">58.</emphasis>
Lateral habitus of
<taxonomicName id="DDC64D60922F0D2A16091F20B05FFA97" authorityName="Engel" authorityYear="2022" box="[533,639,1337,1358]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="santiago" status="new species">
<emphasis id="28B2EAF1922F0D2A16091F20B002FA97" box="[533,546,1337,1358]" italics="true" pageId="42" pageNumber="41">S</emphasis>
.
<emphasis id="28B2EAF1922F0D2A16321F20B05FFA97" box="[558,639,1337,1358]" italics="true" pageId="42" pageNumber="41">santiago</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A922F0D2A16901F20B12DFA97" box="[652,781,1336,1358]" pageId="42" pageNumber="41" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="1A7936E3922F0D2A14CC1F60B30FFA49" blockId="42.[160,1168,1401,1777]" box="[208,303,1401,1427]" pageId="42" pageNumber="41">
<specimenCount id="0CC0FD6A922F0D2A14CC1F60B2C4FA4A" box="[208,228,1401,1427]" count="1" pageId="42" pageNumber="41" type="female"></specimenCount>
:
<emphasis id="28B2EAF1922F0D2A14ED1F60B308FA49" box="[241,296,1401,1424]" italics="true" pageId="42" pageNumber="41">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E3922F0D2A14CC1F80B30FFA69" blockId="42.[160,1168,1401,1777]" box="[208,303,1433,1459]" pageId="42" pageNumber="41">
<specimenCount id="0CC0FD6A922F0D2A14CC1F80B2C4FA6A" box="[208,228,1433,1459]" count="1" pageId="42" pageNumber="41" type="male"></specimenCount>
:
<emphasis id="28B2EAF1922F0D2A14ED1F80B308FA69" box="[241,296,1433,1456]" italics="true" pageId="42" pageNumber="41">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922F0D2A14CC1FA1B37AF9A9" pageId="42" pageNumber="41" type="materials_examined">
<paragraph id="1A7936E3922F0D2A14CC1FA1B37AF9C9" blockId="42.[160,1168,1401,1777]" pageId="42" pageNumber="41">
<materialsCitation id="AAAE3CBE922F0D2A14CC1FA1B37AF9C9" collectingDate="2000-10-16" collectionCode="R, SEMC" country="Peru" county="Reserved Zone" elevation="317" latitude="-11.944722" location="Park" longLatPrecision="21" longitude="-71.28333" municipality="Manu" pageId="42" pageNumber="41" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<typeStatus id="C57D8841922F0D2A14CC1FA1B368FA09" box="[208,328,1464,1488]" pageId="42" pageNumber="41" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D17673922F0D2A156F1FA0B38DFA09" box="[371,429,1465,1488]" name="Peru" pageId="42" pageNumber="41">Peru</collectingCountry>
:
<collectingRegion id="D802F801922F0D2A15A61FA1B051FA09" box="[442,625,1464,1488]" country="Peru" name="Madre de Dios" pageId="42" pageNumber="41">Madre de Dios</collectingRegion>
, Pakita Bio. Stn.,
<collectingCounty id="F3184E6F922F0D2A174D1FA1B626FA09" box="[849,1030,1464,1488]" pageId="42" pageNumber="41">Reserved Zone</collectingCounty>
,
<collectingMunicipality id="FA1DAC99922F0D2A10081FA0B67AFA09" box="[1044,1114,1465,1488]" pageId="42" pageNumber="41">Manu</collectingMunicipality>
Na- tional
<location id="1F196038922F0D2A14F01FC1B304FA29" LSID="urn:lsid:plazi:treatment:926F87F5922D0D2B15841C18B1A8FEC0:1F196038922F0D2A14F01FC1B304FA29" box="[236,292,1496,1520]" country="Peru" county="Reserved Zone" latitude="-11.944722" longLatPrecision="21" longitude="-71.28333" municipality="Manu" name="Park" pageId="42" pageNumber="41" stateProvince="Madre de Dios">Park</location>
,
<quantity id="DD3E9B06922F0D2A152D1FC1B35BFA29" box="[305,379,1496,1520]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="42" pageNumber="41" unit="m" value="317.0">
<elevation id="91EBD1D0922F0D2A152D1FC1B35BFA29" box="[305,379,1496,1520]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="42" pageNumber="41" unit="m" value="317.0">317 m</elevation>
</quantity>
,
<geoCoordinate id="7FF25024922F0D2A15941FC1B029FA29" box="[392,521,1496,1520]" degrees="11" direction="south" minutes="56" orientation="latitude" pageId="42" pageNumber="41" precision="15" seconds="41" value="-11.944722">11°5641S</geoCoordinate>
,
<geoCoordinate id="7FF25024922F0D2A160A1FC1B0B6FA29" box="[534,662,1496,1520]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="42" pageNumber="41" precision="15" seconds="0" value="-71.28333">71°170W</geoCoordinate>
,
<date id="6E781023922F0D2A16BF1FC1B116FA29" box="[675,822,1496,1520]" pageId="42" pageNumber="41" value="2000-10-16">
<collectingDate id="7E3CE9CB922F0D2A16BF1FC1B116FA29" box="[675,822,1496,1520]" pageId="42" pageNumber="41" value="2000-10-16">16 Oct. 2000</collectingDate>
</date>
,
<collectionCode id="7CD7AE26922F0D2A175E1FC0B177FA29" box="[834,855,1497,1520]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="42" pageNumber="41">R</collectionCode>
. Brooks, ex: wet area near stream (
<collectionCode id="7CD7AE26922F0D2A14E31FE1B36FF9C9" box="[255,335,1528,1552]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="42" pageNumber="41" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E3922F0D2A14CC1C01B37AF9A9" blockId="42.[160,1168,1401,1777]" pageId="42" pageNumber="41">
<materialsCitation id="AAAE3CBE922F0D2A14CC1C01B177F989" collectingDate="2000-10-16" collectionCode="R" country="Peru" county="Reserved Zone" elevation="317" latitude="-11.944722" location="Park" longLatPrecision="21" longitude="-71.28333" municipality="Manu" pageId="42" pageNumber="41" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<typeStatus id="C57D8841922F0D2A14CC1C01B36EF9E9" box="[208,334,1560,1584]" pageId="42" pageNumber="41" type="paratype">PARATYPES</typeStatus>
: 2⚲⚲,
<collectingCountry id="62D17673922F0D2A15931C00B3E9F9E9" box="[399,457,1561,1584]" name="Peru" pageId="42" pageNumber="41">Peru</collectingCountry>
:
<collectingRegion id="D802F801922F0D2A15CF1C01B0A5F9E9" box="[467,645,1560,1584]" country="Peru" name="Madre de Dios" pageId="42" pageNumber="41">Madre de Dios</collectingRegion>
, Pakita Bio. Stn.,
<collectingCounty id="F3184E6F922F0D2A17451C01B62CF9E9" box="[857,1036,1560,1584]" pageId="42" pageNumber="41">Reserved Zone</collectingCounty>
,
<collectingMunicipality id="FA1DAC99922F0D2A100B1C00B67DF9E9" box="[1047,1117,1561,1584]" pageId="42" pageNumber="41">Manu</collectingMunicipality>
Na- tional
<location id="1F196038922F0D2A14F01C21B304F989" LSID="urn:lsid:plazi:treatment:926F87F5922D0D2B15841C18B1A8FEC0:1F196038922F0D2A14F01C21B304F989" box="[236,292,1592,1616]" country="Peru" county="Reserved Zone" latitude="-11.944722" longLatPrecision="21" longitude="-71.28333" municipality="Manu" name="Park" pageId="42" pageNumber="41" stateProvince="Madre de Dios">Park</location>
,
<quantity id="DD3E9B06922F0D2A152D1C21B35BF989" box="[305,379,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="42" pageNumber="41" unit="m" value="317.0">
<elevation id="91EBD1D0922F0D2A152D1C21B35BF989" box="[305,379,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="42" pageNumber="41" unit="m" value="317.0">317 m</elevation>
</quantity>
,
<geoCoordinate id="7FF25024922F0D2A15941C21B029F989" box="[392,521,1592,1616]" degrees="11" direction="south" minutes="56" orientation="latitude" pageId="42" pageNumber="41" precision="15" seconds="41" value="-11.944722">11°5641S</geoCoordinate>
,
<geoCoordinate id="7FF25024922F0D2A160A1C21B0B6F989" box="[534,662,1592,1616]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="42" pageNumber="41" precision="15" seconds="0" value="-71.28333">71°170W</geoCoordinate>
,
<date id="6E781023922F0D2A16BF1C21B116F989" box="[675,822,1592,1616]" pageId="42" pageNumber="41" value="2000-10-16">
<collectingDate id="7E3CE9CB922F0D2A16BF1C21B116F989" box="[675,822,1592,1616]" pageId="42" pageNumber="41" value="2000-10-16">16 Oct. 2000</collectingDate>
</date>
,
<collectionCode id="7CD7AE26922F0D2A175E1C20B177F989" box="[834,855,1593,1616]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="42" pageNumber="41">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="AAAE3CBE922F0D2A177F1C21B377F9A9" collectionCode="SEMC" country="Peru" location="Brooks" pageId="42" pageNumber="41" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<location id="1F196038922F0D2A177F1C21B198F989" LSID="urn:lsid:plazi:treatment:926F87F5922D0D2B15841C18B1A8FEC0:1F196038922F0D2A177F1C21B198F989" box="[867,952,1592,1616]" country="Peru" name="Brooks" pageId="42" pageNumber="41" stateProvince="Madre de Dios">Brooks</location>
, ex: wet area near stream (
<collectionCode id="7CD7AE26922F0D2A14E31C41B36FF9A9" box="[255,335,1624,1648]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="42" pageNumber="41" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922F0D2A14CC1C61B1FDF909" pageId="42" pageNumber="41" type="etymology">
<paragraph id="1A7936E3922F0D2A14CC1C61B1FDF909" blockId="42.[160,1168,1401,1777]" pageId="42" pageNumber="41">ETYMOLOGY: The specific epithet honors the inspiring writings and life of James A. Baldwin (19241987), whose energies and moral essays identified paths and placed crucial paving stones toward a more just, equitable, and healed world.</paragraph>
</subSubSection>
<subSubSection id="52DC6568922F0D2B14CC1CC1B1A8FEC0" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41" type="discussion">
<paragraph id="1A7936E3922F0D2B14CC1CC1B1A8FEC0" blockId="42.[160,1168,1401,1777]" lastBlockId="43.[160,1168,225,281]" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41">
COMMENT: One of the
<typeStatus id="C57D8841922F0D2A15C51CC0B06EF928" box="[473,590,1753,1777]" pageId="42" pageNumber="41" type="paratype">paratypes</typeStatus>
exhibits an interesting aberration in the form of the right mesotibial spur. Rather than being a slender serrate spur it is instead formed like the protibial spur, complete with a velum and short malus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,184 @@
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<mods:title id="D457DE250207966006186426D6D6B24F">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:namePart id="4BF9375AF4EE1B7712CB7247395B0795">Engel, Michael S.</mods:namePart>
<mods:affiliation id="103A7A195A2A680AECBC92A74410667D">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<subSubSection id="52DC6568922E0D2B15861B58B61AFEA0" pageId="43" pageNumber="42" type="nomenclature">
<paragraph id="1A7936E3922E0D2B15861B58B1B6FE80" blockId="43.[410,918,321,345]" box="[410,918,321,345]" pageId="43" pageNumber="42">
<heading id="4131818F922E0D2B15861B58B1B6FE80" box="[410,918,321,345]" centered="true" fontSize="10" level="2" pageId="43" pageNumber="42" reason="2">
<taxonomicName id="DDC64D60922E0D2B15861B58B0DEFE80" ID-CoL="7C9D" authority="Engel" authorityName="Engel" authorityYear="2022" box="[410,766,321,345]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="42" phylum="Arthropoda" rank="species" species="santiago" status="sp. nov.">
<emphasis id="28B2EAF1922E0D2B15861B58B092FE80" bold="true" box="[410,690,321,345]" italics="true" pageId="43" pageNumber="42">Scaptotrigona santiago</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A922E0D2B17151B5BB1B6FE80" box="[777,918,321,345]" pageId="43" pageNumber="42" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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ZooBank:
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</paragraph>
</subSubSection>
<subSubSection id="52DC6568922E0D2B16741B98B0E8FE40" box="[616,712,385,409]" pageId="43" pageNumber="42" type="description">
<paragraph id="1A7936E3922E0D2B16741B98B0E8FE40" blockId="43.[246,1082,355,409]" box="[616,712,385,409]" pageId="43" pageNumber="42">
(
<figureCitation id="82FD2A66922E0D2B166D1B98B0E0FE40" box="[625,704,385,409]" captionStart="Figures 5758" captionStartId="42.[160,241,1307,1329]" captionTargetBox="[184,1136,240,1284]" captionTargetId="figure-130@42.[160,1168,224,1301]" captionTargetPageId="42" captionText="Figures 5758. Workers of Scaptotrigona Moure. 57. Lateral habitus of Scaptotrigona baldwini, new species. 58. Lateral habitus of S. santiago, new species." pageId="43" pageNumber="42">Fig. 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922E0D2B14CC1BD8B60FFDE0" pageId="43" pageNumber="42" type="diagnosis">
<paragraph id="1A7936E3922E0D2B14CC1BD8B60FFDE0" blockId="43.[160,1175,449,1785]" pageId="43" pageNumber="42">
DIAGNOSIS: This is another species lacking a genal tooth and similar to
<taxonomicName id="DDC64D60922E0D2B10031BDBB651FE00" authorityName="Engel" authorityYear="2022" box="[1055,1137,449,473]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="42" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922E0D2B10031BDBB651FE00" box="[1055,1137,449,473]" italics="true" pageId="43" pageNumber="42">S. ederi</emphasis>
</taxonomicName>
in its overall darker coloration. It can be distinguished from
<taxonomicName id="DDC64D60922E0D2B175A1BFBB1B6FE20" authorityName="Engel" authorityYear="2022" box="[838,918,481,505]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="42" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF1922E0D2B175A1BFBB1B6FE20" box="[838,918,481,505]" italics="true" pageId="43" pageNumber="42">S. ederi</emphasis>
</taxonomicName>
by the wholly brown genae (
<figureCitation id="82FD2A66922E0D2B14EA1818B366FDC0" box="[246,326,513,537]" captionStart="Figures 5758" captionStartId="42.[160,241,1307,1329]" captionTargetBox="[184,1136,240,1284]" captionTargetId="figure-130@42.[160,1168,224,1301]" captionTargetPageId="42" captionText="Figures 5758. Workers of Scaptotrigona Moure. 57. Lateral habitus of Scaptotrigona baldwini, new species. 58. Lateral habitus of S. santiago, new species." pageId="43" pageNumber="42">Fig. 58</figureCitation>
) and the uniquely bicolored bristles of the mesoscutum and corbicula, where the majority of the rachis is black before blending to off white apically.
</paragraph>
</subSubSection>
<subSubSection id="52DC6568922E0D2C14CC1858B30FFE60" lastPageId="44" lastPageNumber="43" pageId="43" pageNumber="42" type="description">
<paragraph id="1A7936E3922E0D2B14CC1858B6A6FCA0" blockId="43.[160,1175,449,1785]" pageId="43" pageNumber="42">
DESCRIPTION: As described for
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<emphasis id="28B2EAF1922E0D2B165F185BB0B7FD80" box="[579,663,577,601]" italics="true" pageId="43" pageNumber="42">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF1922E0D2B16B51858B13CFD80" box="[681,796,577,601]" italics="true" pageId="43" pageNumber="42">vide supra</emphasis>
), with the following modifica- tions: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite)
<quantity id="DD3E9B06922E0D2B15181898B37FFD40" box="[260,351,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="43" pageNumber="42" unit="mm" value="6.1">6.1 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B06922E0D2B16C91898B11DFD40" box="[725,829,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="43" pageNumber="42" unit="mm" value="2.42">2.42 mm</quantity>
, length
<quantity id="DD3E9B06922E0D2B17801898B623FD40" box="[924,1027,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="1.94" pageId="43" pageNumber="42" unit="mm" value="1.94">1.94 mm</quantity>
; compound eye length
<quantity id="DD3E9B06922E0D2B153818B8B3ACFD60" box="[292,396,673,697]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="43" pageNumber="42" unit="mm" value="1.39">1.39 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06922E0D2B16FC18B8B168FD60" box="[736,840,673,697]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="43" pageNumber="42" unit="mm" value="1.52">1.52 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06922E0D2B14BC18D8B32AFD00" box="[160,266,705,729]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="43" pageNumber="42" unit="mm" value="1.42">1.42 mm</quantity>
. Scape length
<quantity id="DD3E9B06922E0D2B15DD18D8B00AFD00" box="[449,554,705,729]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="43" pageNumber="42" unit="mm" value="0.91">0.91 mm</quantity>
, about as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B06922E0D2B151B18F8B34EFD20" box="[263,366,737,761]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="43" pageNumber="42" unit="mm" value="0.88">0.88 mm</quantity>
. Clypeus approximately 1.6× as wide as long, length
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, width
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. Malar area approximately 1.8× flagellar diameter or 0.80× basal mandibular width. Preoccipital ridge carinate, lamellate dorsally, with bordering medial and lat- eral indentations deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming rounded angle, without sharply acute lamellate tooth.
</paragraph>
<paragraph id="1A7936E3922E0D2B14CC199BB385FAA0" blockId="43.[160,1175,449,1785]" pageId="43" pageNumber="42">
Integument of labiomaxillary complex orange; labrum yellow; mandible orange; clypeus largely yellow orange with stains of orange paramedially, epistomal sulcus delineated yellow orange; supraclypeal area orange; malar space yellow; face below tangent of antennal toruli yellow; scape yellow to yellow orange below, brown dor- sally; pedicel and flagellum brown, with narrow underside of flagellum orange; face above antennal toruli brown, vertex, ocellocular, and posterior of head as on upper frons; gena and postgena as on upper frons except area immediately bordering malar space yellow orange to orange. Mesosoma largely dark brown to black except mesos- cutellum, metanotum, and basal area and posterior surface of propodeum yellow; te- gula brownish orange and semi-translucent; legs largely dark brown except coxae and trochanters orange, apices of femora and tibiae orange including corbicular surface, and apical tarsomeres orange. Wing membranes parchment colored; veins orange. Metasoma generally dark brown except anterior-facing surface of tergum I brown, dorsal-facing disc of tergum I yellow posteriorly; tergum II yellow posteriorly; terga
<collectionCode id="7CD7AE26922E0D2B14BC1F5BB29BFA80" box="[160,187,1346,1369]" pageId="43" pageNumber="42">III</collectionCode>
<collectionCode id="7CD7AE26922E0D2B14D51F5BB2FCFA80" box="[201,220,1346,1369]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="43" pageNumber="42" type="Museum">V</collectionCode>
dark brown with yellow apical marginal zones; tergum
<collectionCode id="7CD7AE26922E0D2B179A1F5BB182FA80" box="[902,930,1346,1369]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="43" pageNumber="42">VI</collectionCode>
dark brown; sterna brown to dark brown.
</paragraph>
<paragraph id="1A7936E3922E0D2C14CC1F9BB67BFEA0" blockId="43.[160,1175,449,1785]" lastBlockId="44.[160,1168,225,602]" lastPageId="44" lastPageNumber="43" pageId="43" pageNumber="42">
Vertex with numerous erect, simple, fuscous bristles, typically with off white tips, some bristles near upper orbits black; pronotal dorsolateral margin near and anterior to pronotal lobe with yellow to dark fuscous bristles; mesoscutum anterior and lateral borders with fuscous bristles, typically with off white tips; tegula with suberect fus- cous bristles anteriorly; mesoscutellum with abundant, elongate, erect fuscous to dark fulvous bristles; mesepisternum with scatered, erect to subdecumbent pale yellow to off white bristles, longer ventrally, along rounded anterior border with preëpister- num a line of abundant, suberect to subdecumbent fuscous bristles with off white tips extending ventrally from pronotal lobe; lateral surface of propodeum with scatered, long, erect, yellow bristles emerging amid dense, appressed, white, plumose setae. Legs with abundant fuscous, dark fulvous, or fulvous bristles; corbicular bristles dark fulvous to black with pale yellow to off white apical halves, profundal bristles fuscous to black, marginal bristles dark fulvous to fuscous, some with off white tips; metaba- sitarsus with dark fulvous to fuscous bristles, sometimes with off white tips, prolateral surface with fine, erect, simple, fuscous to black setae. Metasoma with numerous, prominent, suberect to subdecumbent, yellow bristles on terga
<collectionCode id="7CD7AE2692290D2C17931B5BB18AFE80" box="[911,938,322,345]" pageId="44" pageNumber="43">III</collectionCode>
<collectionCode id="7CD7AE2692290D2C17A41B5BB1EBFE80" box="[952,971,322,345]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="44" pageNumber="43" type="Museum">V</collectionCode>
arising amid to- mentum, intermixed with a few fuscous to dark fulvous bristles with yellow tips.
</paragraph>
<paragraph id="1A7936E392290D2C14CC1B9BB30FFE40" blockId="44.[160,1168,225,602]" box="[208,303,386,412]" pageId="44" pageNumber="43">
<specimenCount id="0CC0FD6A92290D2C14CC1B9BB2C4FE45" box="[208,228,386,412]" count="1" pageId="44" pageNumber="43" type="female"></specimenCount>
:
<emphasis id="28B2EAF192290D2C14ED1B9BB308FE40" box="[241,296,386,409]" italics="true" pageId="44" pageNumber="43">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392290D2C14CC1BBBB30FFE60" blockId="44.[160,1168,225,602]" box="[208,303,418,444]" pageId="44" pageNumber="43">
<specimenCount id="0CC0FD6A92290D2C14CC1BBBB2C4FE65" box="[208,228,418,444]" count="1" pageId="44" pageNumber="43" type="male"></specimenCount>
:
<emphasis id="28B2EAF192290D2C14ED1BBBB308FE60" box="[241,296,418,441]" italics="true" pageId="44" pageNumber="43">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892290D2C14CC1BD8B347FE20" pageId="44" pageNumber="43" type="materials_examined">
<paragraph id="1A7936E392290D2C14CC1BD8B347FE20" blockId="44.[160,1168,225,602]" pageId="44" pageNumber="43">
<materialsCitation id="AAAE3CBE92290D2C14CC1BD8B347FE20" collectingDate="1986-02" collectionCode="SEMC" collectorName="L. Coloma" country="Ecuador" location="Santiago" pageId="44" pageNumber="43" specimenCode="II-86" specimenCount="1" stateProvince="Morona-Santiago" typeStatus="holotype">
<typeStatus id="C57D884192290D2C14CC1BD8B368FE00" box="[208,328,449,473]" pageId="44" pageNumber="43" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392290D2C156D1BD8B3F6FE00" box="[369,470,449,473]" name="Ecuador" pageId="44" pageNumber="43">Ecuador</collectingCountry>
,
<collectingRegion id="D802F80192290D2C15FF1BD8B092FE00" box="[483,690,449,473]" country="Ecuador" name="Morona-Santiago" pageId="44" pageNumber="43">Morona-Santiago</collectingRegion>
,
<location id="1F19603892290D2C16A31BD8B107FE00" LSID="urn:lsid:plazi:treatment:926F87F5922E0D2C15861B58B18BFD80:1F19603892290D2C16A31BD8B107FE00" box="[703,807,449,473]" country="Ecuador" name="Santiago" pageId="44" pageNumber="43" stateProvince="Morona-Santiago">Santiago</location>
,
<specimenCode id="4A609E9892290D2C17281BD8B14AFE00" box="[820,874,449,473]" collectionCode="II-" pageId="44" pageNumber="43">II-86</specimenCode>
[
<date id="6E78102392290D2C17651BD8B609FE00" box="[889,1065,449,473]" pageId="44" pageNumber="43" value="1986-02">
<collectingDate id="7E3CE9CB92290D2C17651BD8B609FE00" box="[889,1065,449,473]" pageId="44" pageNumber="43" value="1986-02">February 1986</collectingDate>
</date>
], legit
<collectorName id="B733533592290D2C10651BDBB2DDFE20" pageId="44" pageNumber="43">L. Coloma</collectorName>
(
<collectionCode id="7CD7AE2692290D2C15171BF8B37BFE20" box="[267,347,481,505]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="44" pageNumber="43" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892290D2C14CC1818B18BFD80" pageId="44" pageNumber="43" type="etymology">
<paragraph id="1A7936E392290D2C14CC1818B18BFD80" blockId="44.[160,1168,225,602]" pageId="44" pageNumber="43">
ETYMOLOGY: The specific epithet is taken from both the province in which the ho- lotype was collected (
<emphasis id="28B2EAF192290D2C15BC183BB398FDE0" box="[416,440,546,569]" italics="true" pageId="44" pageNumber="43">i.e</emphasis>
.,
<collectingRegion id="D802F80192290D2C15D11838B0BDFDE0" box="[461,669,545,569]" country="Ecuador" name="Morona-Santiago" pageId="44" pageNumber="43">Morona-Santiago</collectingRegion>
), as well as the middle name of Jesus San- tiago Moure (19122010), who established the genus
<taxonomicName id="DDC64D6092290D2C1712185BB184FD80" box="[782,932,578,601]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF192290D2C1712185BB184FD80" box="[782,932,578,601]" italics="true" pageId="44" pageNumber="43">Scaptotrigona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>