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<document id="8F7A95A08B6C238DE5E1CA1619F5D70B" ID-ISSN="1810-522X" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721915584095" checkinUser="felipe" docAuthor="Ng, Peter K. L., Lin, Chia-Wei &amp; Ho, Ping-Ho" docDate="2018" docId="039C87E62F10115EFC8477054691FE33" docLanguage="en" docName="ZoolStud.57.12.1-17.pdf" docOrigin="Zoological Studies 12" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Heteropilumnus satriai Yeo, Rahayu and Ng 2004" docType="treatment" docVersion="1" masterDocId="FFA5FF9E2F121158FFD67138457BFD25" masterDocTitle="On Three Species of Reef-dwelling Pilumnid Crabs from Taiwan, with Notes on Heteropilumnus De Man, 1895 (Crustacea: Brachyura)" updateTime="1721916044657" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
<mods:mods id="29E80FDBBCA3156C89F98A56FA85618A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B063823B4982B26C42F7323E6ABA0ED9">
<mods:title id="BFEA4025EAAF33D511275DD215D5257F">On Three Species of Reef-dwelling Pilumnid Crabs from Taiwan, with Notes on Heteropilumnus De Man, 1895 (Crustacea: Brachyura)</mods:title>
</mods:titleInfo>
<mods:name id="498927DAB008693EFC06C6CFD7FCDD29" type="personal">
<mods:role id="CCD7361A89C31CF065CABB29F4A67F0B">
<mods:roleTerm id="11CCEA1A4BD6677C7E2C365F4E47D654">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B0A25E39B5639CF451DB8DF1F7F0C38">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="52E1E774C0B62F7F2B676CE7B554394C">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive,</mods:affiliation>
</mods:name>
<mods:name id="26BCAF300D0E8D16647DE4905144AC34" type="personal">
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<mods:namePart id="53B92E7784E2108E78F23953052CC921">Lin, Chia-Wei</mods:namePart>
<mods:affiliation id="0DAB32A83C02D306F0BF8D1C69650B4B">National Museum of Marine Biology and Aquarium, 2 Houwan Road, Checheng, Pingtung, Taiwan 944, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="000EDBBA9229FD6B987EF2921531592C" type="personal">
<mods:role id="A0E6F78FE122DA1BFB0B057B31B1956A">
<mods:roleTerm id="C0F2EC76C5B4D0D05029F019833B8E3E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="01CA9CA23126AF3091A5A5BDF48EAD23">Ho, Ping-Ho</mods:namePart>
<mods:affiliation id="BB0D7B46777E3D8EE2EEDF19552F0D31">Department of Environmental Biology and Fisheries Science, National Taiwan Ocean University, 2 Pei-Ning Rd., Keelung 202, Taiwan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C6A95274A132F255EAD1F66812494E99">text</mods:typeOfResource>
<mods:relatedItem id="D5AE491DB234C0B43C5F95F990890C1B" type="host">
<mods:titleInfo id="003490A82B7A316ED12E47B654D1ED5E">
<mods:title id="3C38C20DC7742EF8E47458B058BBEF80">Zoological Studies</mods:title>
</mods:titleInfo>
<mods:part id="487F5A785519303733B22FD0BF4AA354">
<mods:date id="61ECF7C7A075B78FC6EF2B269D3ACF1C">2018</mods:date>
<mods:detail id="CACCED7B8AD76E22E7B65390AFD717AF" type="issue">
<mods:number id="C5048AE8D0257D5ADB55BA96AC441361">12</mods:number>
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<mods:classification id="2132F3B579DD2B254A1922C8489A6F1F">journal volume</mods:classification>
<mods:identifier id="EE20BC09ABC22786FA78A6F73F579C8C" type="ISSN">1810-522X</mods:identifier>
</mods:mods>
<treatment id="039C87E62F10115EFC8477054691FE33" LSID="urn:lsid:plazi:treatment:039C87E62F10115EFC8477054691FE33" httpUri="http://treatment.plazi.org/id/039C87E62F10115EFC8477054691FE33" lastPageId="6" pageId="2">
<subSubSection id="C32F657B2F10115AFC84770540D7FB73" box="[850,1452,1597,1622]" pageId="2" type="nomenclature">
<paragraph id="8B8A36F02F10115AFC84770540D7FB73" blockId="2.[850,1452,1597,1622]" box="[850,1452,1597,1622]" pageId="2">
<heading id="D0C2819C2F10115AFC84770540D7FB73" bold="true" box="[850,1452,1597,1622]" centered="true" fontSize="10" level="2" pageId="2" reason="3">
<taxonomicName id="4C354D732F10115AFC84770540D7FB73" ID-CoL="3LC4F" authority="Yeo, Rahayu and Ng, 2004" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[850,1452,1597,1622]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F10115AFC84770540D7FB73" bold="true" box="[850,1452,1597,1622]" pageId="2">
<emphasis id="B941EAE22F10115AFC847705411CFB73" bold="true" box="[850,1127,1597,1622]" italics="true" pageId="2">Heteropilumnus satriai</emphasis>
<bibRefCitation id="EFA44B012F10115AFBB8770540D7FB73" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[1134,1452,1597,1622]" pageId="2" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F657B2F10115AFBC9776541A4FB53" box="[1055,1247,1629,1654]" pageId="2" type="description">
<paragraph id="8B8A36F02F10115AFBC9776541A4FB53" blockId="2.[1055,1247,1629,1654]" box="[1055,1247,1629,1654]" pageId="2">
(
<figureCitation id="130E2A752F10115AFBF07765410AFB53" box="[1062,1137,1629,1654]" captionStart="Fig" captionStartId="3.[136,170,1869,1889]" captionTargetBox="[143,1406,608,1843]" captionTargetId="figure-256@3.[143,1406,608,1843]" captionTargetPageId="3" captionText="Fig. 1. Colour in life. A, Heteropilumnus decharmoyi (Bouvier, 1915), male (16.8 × 12.7 mm) (NMNST); B, H. decharmoyi (Bouvier, 1915), male (13.3 × 9.3 mm) (NMMBCD4084); C, H. satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043); D, H. satriai Yeo, Rahayu and Ng, 2004, female (16.1 × 11.9 mm) (NMMBCD4086); E, H. satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085); F, H. satriai Yeo, Rahayu and Ng, 2004, male (6.8 × 5.4 mm) (NMMBCD4086)." pageId="2">Figs. 1</figureCitation>
D-F, 5-9)
</paragraph>
</subSubSection>
<subSubSection id="C32F657B2F10115AFC8477A04138FBE3" pageId="2" type="reference_group">
<paragraph id="8B8A36F02F10115AFC8477A04138FBE3" blockId="2.[850,1451,1688,1734]" pageId="2">
<treatmentCitationGroup id="AB2511DE2F10115AFC8477A04138FBE3" pageId="2">
<treatmentCitation id="0A9410E12F10115AFC8477A04012FB89" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[850,1385,1688,1708]" page="80" pageId="2" year="2004">
<taxonomicName id="4C354D732F10115AFC8477A04012FB89" ID-CoL="3LC4F" authority="Yeo, Rahayu and Ng, 2004: 80" authorityName="Yeo, Rahayu and Ng" authorityPageNumber="80" authorityYear="2004" box="[850,1385,1688,1708]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F10115AFC8477A04157FB89" box="[850,1068,1688,1708]" italics="true" pageId="2">Heteropilumnus satriai</emphasis>
<bibRefCitation id="EFA44B012F10115AFBE277A04012FB89" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[1076,1385,1688,1708]" pageId="2" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004: 80</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0A9410E12F10115AFC51778A4144FBE3" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[903,1087,1714,1734]" page="140" pageId="2" year="2008">
<bibRefCitation id="EFA44B012F10115AFC51778A4144FBE3" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[903,1087,1714,1734]" pageId="2" pagination="1 - 286" refId="ref6452" refString="Ng PKL, Guinot D, Davie PJF. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles B Zool, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008: 140</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32F657B2F10115EFC5977D64691FE33" lastPageId="6" pageId="2" type="materials_examined">
<paragraph id="8B8A36F02F10115AFC5977D640D7FA82" blockId="2.[850,1452,1774,1959]" pageId="2">
<emphasis id="B941EAE22F10115AFC5977D64107FA22" box="[911,1148,1774,1799]" italics="true" pageId="2">Material examined</emphasis>
:
<materialsCitation id="3B5D3CAD2F10115AFB4677D640D7FA82" collectingDate="2002-03-16" collectionCode="MZB" collectorName="Anambas Expedition" country="Indonesia" location="Pulau Laut" municipality="Natuna Island" pageId="2" specimenCode="NMMBCD4085" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="548E88522F10115AFB4677D6407AFA22" box="[1168,1281,1774,1799]" pageId="2" type="holotype">Holotype</typeStatus>
- male (19.3 ×
<quantity id="4CCD9B152F10115AFCB9763646B7FA02" box="[879,972,1806,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" unit="mm" value="13.0">13 mm</quantity>
) (
<collectionCode id="ED24AE352F10115AFC317636415DFA02" box="[999,1062,1806,1831]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" type="Museum">MZB</collectionCode>
Cru 1668), station EA-D 11, rocky islet southeast of
<location id="8EEA602B2F10115AFB7476164041FA62" LSID="urn:lsid:plazi:treatment:039C87E62F10115EFC8477054691FE33:8EEA602B2F10115AFB7476164041FA62" box="[1186,1338,1838,1863]" country="Indonesia" municipality="Natuna Island" name="Pulau Laut" pageId="2">Pulau Laut</location>
,
<collectingMunicipality id="6BEEAC8A2F10115AFA9A761646DAFA42" pageId="2">Natuna Island</collectingMunicipality>
,
<collectingCountry id="F32276602F10115AFC6676764155FA42" box="[944,1070,1870,1895]" name="Indonesia" pageId="2">Indonesia</collectingCountry>
, coll.
<collectorName id="26C053262F10115AFBAE76764005FA42" box="[1144,1406,1870,1895]" pageId="2">Anambas Expedition</collectorName>
,
<date id="FF8B10302F10115AFA5B7676468AFAA2" pageId="2" value="2002-03-16">
<collectingDate id="EFCFE9D82F10115AFA5B7676468AFAA2" pageId="2" value="2002-03-16">16 March 2002</collectingDate>
</date>
. Others -
<specimenCount id="9D33FD792F10115AFBA0765641B5FAA2" box="[1142,1230,1902,1927]" count="1" pageId="2" type="male">1 male</specimenCount>
(11.1 ×
<quantity id="4CCD9B152F10115AFA96765640D8FAA2" box="[1344,1443,1902,1927]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="2" unit="mm" value="8.2">8.2 mm</quantity>
) (
<specimenCode id="DB939E8B2F10115AFC8F76B5416EFA82" box="[857,1045,1933,1959]" collectionCode="NMMBCD" pageId="2">NMMBCD4085</specimenCode>
), under rock with sandy substrate,
</materialsCitation>
</paragraph>
<paragraph id="8B8A36F02F11115BFFAB719044BDFD9A" blockId="3.[125,454,168,191]" box="[125,454,168,191]" pageId="3">
<bibCitation id="5E1F3E9C2F11115BFFAB719044BDFD9A" box="[125,454,168,191]" journal="Zoological Studies" page="12" pageId="3" score="5.2987992369807255" volume="57" year="2018">
<emphasis id="B941EAE22F11115BFFAB7190444CFD9A" box="[125,311,168,191]" italics="true" pageId="3">Zoological Studies</emphasis>
<emphasis id="B941EAE22F11115BFEEB71904425FD9A" bold="true" box="[317,350,168,191]" pageId="3">57:</emphasis>
12 (2018)
</bibCitation>
</paragraph>
<paragraph id="8B8A36F02F11115BFF5E71C540FEFD9A" blockId="3.[136,738,253,567]" lastBlockId="3.[1288,1413,168,191]" pageId="3">
Hojie, Kentin,
<geoCoordinate id="EE0150372F11115BFEE671C544BAFC33" box="[304,449,253,278]" degrees="21" direction="north" minutes="57.432" orientation="latitude" pageId="3" precision="1" value="21.9572">21°57.432'N</geoCoordinate>
<geoCoordinate id="EE0150372F11115BFE1C71C54716FC33" box="[458,621,253,278]" degrees="120" direction="east" minutes="42.631" orientation="longitude" pageId="3" precision="1" value="120.71052">120°42.631'E</geoCoordinate>
, southern
<collectingCountry id="F32276602F11115BFF5E70254599FC13" box="[136,226,285,310]" name="Taiwan" pageId="3">Taiwan</collectingCountry>
, by
<collectionCode id="ED24AE352F11115BFEC170254402FC12" box="[279,377,285,311]" pageId="3">SCUBA</collectionCode>
, coll. C.-W. Lin,
<date id="FF8B10302F11115BFD9E702547A5FC13" box="[584,734,285,310]" pageId="3" value="2013-07-08">8 July 2013</date>
;
<specimenCount id="9D33FD792F11115BFF5E700545A1FC73" box="[136,218,317,342]" count="1" pageId="3" type="male">1 male</specimenCount>
(18.3 × 13.0 mm) (
<collectionCode id="ED24AE352F11115BFE1570054487FC72" box="[451,508,317,343]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3">ZRC</collectionCode>
2017.1043), under rock with sand substrate, Houbihu, Kentin,
<collectingCountry id="F32276602F11115BFD51706547A5FC53" box="[647,734,349,374]" name="Taiwan" pageId="3">Taiwan</collectingCountry>
, by
<collectionCode id="ED24AE352F11115BFF7A70454477FCB2" box="[172,268,381,407]" pageId="3">SCUBA</collectionCode>
, coll. C.-W. Lin,
<date id="FF8B10302F11115BFE1D70454718FCB3" box="[459,611,381,406]" pageId="3" value="2016-06-08">8 June 2016</date>
;
<specimenCount id="9D33FD792F11115BFDBB7045479AFCB3" box="[621,737,381,406]" count="2" pageId="3" type="female">2 females</specimenCount>
(16.1 ×
<quantity id="4CCD9B152F11115BFF3D70A54421FC93" box="[235,346,413,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="3" unit="mm" value="11.9">11.9 mm</quantity>
, 6.8 ×
<quantity id="4CCD9B152F11115BFE6270A54776FC93" box="[436,525,413,438]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="3" unit="mm" value="5.4">5.4mm</quantity>
), Lutao (= Green Island), southern
<collectingCountry id="F32276602F11115BFEB9708544B6FCF3" box="[367,461,445,470]" name="Taiwan" pageId="3">Taiwan</collectingCountry>
, by
<collectionCode id="ED24AE352F11115BFDD070854717FCF2" box="[518,620,445,471]" pageId="3">SCUBA</collectionCode>
, coll. C.- W. Lin,
<date id="FF8B10302F11115BFF3E70E544ECFCD3" box="[232,407,477,502]" pageId="3" value="2017-06-24">24 June 2017</date>
;
<specimenCount id="9D33FD792F11115BFE7370E5476BFCD3" box="[421,528,477,502]" count="1" pageId="3" type="female">1 female</specimenCount>
(11.4 ×
<quantity id="4CCD9B152F11115BFDAD70E547A1FCD3" box="[635,730,477,502]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" unit="mm" value="8.2">8.2 mm</quantity>
) (
<collectionCode id="ED24AE352F11115BFF5870C54477FF32" box="[142,268,509,535]" pageId="3">NMMBCD</collectionCode>
4086), under large rock, sand substrate, Lutao (= Green Island), southern
<collectingCountry id="F32276602F11115BFD84732547C8FF13" box="[594,691,541,566]" name="Taiwan" pageId="3">Taiwan</collectingCountry>
, by page 4 of 17
</paragraph>
<paragraph id="8B8A36F02F11115BFCFB71C54077FC33" blockId="3.[813,1414,253,566]" box="[813,1292,253,279]" pageId="3">
<collectionCode id="ED24AE352F11115BFCFB71C546F6FC32" box="[813,909,253,279]" pageId="3">SCUBA</collectionCode>
, coll. C.-W. Lin,
<date id="FF8B10302F11115BFB9D71C54072FC33" box="[1099,1289,253,278]" pageId="3" value="2017-08-16">16 August 2017</date>
.
</paragraph>
<paragraph id="8B8A36F02F11115BFCBF702540FEFF13" blockId="3.[813,1414,253,566]" pageId="3">
<emphasis id="B941EAE22F11115BFCBF702546C3FC13" box="[873,952,285,310]" italics="true" pageId="3">Colour</emphasis>
: The colour and patterning in
<taxonomicName id="4C354D732F11115BFACA702540FEFC13" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[1308,1413,285,310]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F11115BFACA702540FEFC13" box="[1308,1413,285,310]" italics="true" pageId="3">H. satriai</emphasis>
</taxonomicName>
in life is similar to that of
<taxonomicName id="4C354D732F11115BFBA270054052FC73" baseAuthorityName="Bouvier" baseAuthorityYear="1915" box="[1140,1321,317,342]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F11115BFBA270054052FC73" box="[1140,1321,317,342]" italics="true" pageId="3">H. decharmoyi</emphasis>
</taxonomicName>
except that the red patches on the ambulatory legs are relatively smaller and more uneven (
<figureCitation id="130E2A752F11115BFADB70454638FC93" captionStart="Fig" captionStartId="3.[136,170,1869,1889]" captionTargetBox="[143,1406,608,1843]" captionTargetId="figure-256@3.[143,1406,608,1843]" captionTargetPageId="3" captionText="Fig. 1. Colour in life. A, Heteropilumnus decharmoyi (Bouvier, 1915), male (16.8 × 12.7 mm) (NMNST); B, H. decharmoyi (Bouvier, 1915), male (13.3 × 9.3 mm) (NMMBCD4084); C, H. satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043); D, H. satriai Yeo, Rahayu and Ng, 2004, female (16.1 × 11.9 mm) (NMMBCD4086); E, H. satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085); F, H. satriai Yeo, Rahayu and Ng, 2004, male (6.8 × 5.4 mm) (NMMBCD4086)." pageId="3">Figs. 1D, E</figureCitation>
). There is some variation in this species, from having the red colour cover most of the carapace (
<figureCitation id="130E2A752F11115BFCE270E546F0FCD3" box="[820,907,477,502]" captionStart="Fig" captionStartId="3.[136,170,1869,1889]" captionTargetBox="[143,1406,608,1843]" captionTargetId="figure-256@3.[143,1406,608,1843]" captionTargetPageId="3" captionText="Fig. 1. Colour in life. A, Heteropilumnus decharmoyi (Bouvier, 1915), male (16.8 × 12.7 mm) (NMNST); B, H. decharmoyi (Bouvier, 1915), male (13.3 × 9.3 mm) (NMMBCD4084); C, H. satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043); D, H. satriai Yeo, Rahayu and Ng, 2004, female (16.1 × 11.9 mm) (NMMBCD4086); E, H. satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085); F, H. satriai Yeo, Rahayu and Ng, 2004, male (6.8 × 5.4 mm) (NMMBCD4086)." pageId="3">Fig. 1D</figureCitation>
) to only covering less than half the surface (
<figureCitation id="130E2A752F11115BFCE270C546E8FF33" box="[820,915,509,534]" captionStart="Fig" captionStartId="3.[136,170,1869,1889]" captionTargetBox="[143,1406,608,1843]" captionTargetId="figure-256@3.[143,1406,608,1843]" captionTargetPageId="3" captionText="Fig. 1. Colour in life. A, Heteropilumnus decharmoyi (Bouvier, 1915), male (16.8 × 12.7 mm) (NMNST); B, H. decharmoyi (Bouvier, 1915), male (13.3 × 9.3 mm) (NMMBCD4084); C, H. satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043); D, H. satriai Yeo, Rahayu and Ng, 2004, female (16.1 × 11.9 mm) (NMMBCD4086); E, H. satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085); F, H. satriai Yeo, Rahayu and Ng, 2004, male (6.8 × 5.4 mm) (NMMBCD4086)." pageId="3">Fig. 1E</figureCitation>
), to the carapace and pereopods being mostly yellowish-white with only small patches of
</paragraph>
<caption id="DF4A66782F11115BFF5E76754117FA90" pageId="3" startId="3.[136,170,1869,1889]" targetBox="[143,1406,608,1843]" targetPageId="3" targetType="figure">
<paragraph id="8B8A36F02F11115BFF5E76754117FA90" blockId="3.[136,1415,1869,1973]" pageId="3">
<emphasis id="B941EAE22F11115BFF5E767545B3FA44" bold="true" box="[136,200,1869,1889]" pageId="3">Fig. 1.</emphasis>
Colour in life. A,
<taxonomicName id="4C354D732F11115BFEAA7675465CFA44" authority="(Bouvier, 1915)" baseAuthorityName="Bouvier" baseAuthorityYear="1915" box="[380,807,1869,1889]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F11115BFEAA767547F2FA44" box="[380,649,1869,1889]" italics="true" pageId="3">Heteropilumnus decharmoyi</emphasis>
(
<bibRefCitation id="EFA44B012F11115BFD437675465AFA44" author="Bouvier EL" box="[661,801,1869,1889]" pageId="3" pagination="178 - 318" refId="ref5756" refString="Bouvier EL. 1915. Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique 48 (3): 178 - 318." type="journal article" year="1915">Bouvier, 1915</bibRefCitation>
)
</taxonomicName>
, male (16.8 × 12.7 mm) (NMNST); B,
<taxonomicName id="4C354D732F11115BFB4B767545B9FA58" authority="(Bouvier, 1915)" baseAuthorityName="Bouvier" baseAuthorityYear="1915" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F11115BFB4B76754052FA44" box="[1181,1321,1869,1889]" italics="true" pageId="3">H. decharmoyi</emphasis>
(
<bibRefCitation id="EFA44B012F11115BFAE0767545C7FA58" author="Bouvier EL" pageId="3" pagination="178 - 318" refId="ref5756" refString="Bouvier EL. 1915. Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique 48 (3): 178 - 318." type="journal article" year="1915">Bouvier, 1915</bibRefCitation>
)
</taxonomicName>
, male (13.3 × 9.3 mm) (NMMBCD4084); C,
<taxonomicName id="4C354D732F11115BFDBD765146ABFA58" authority="Yeo, Rahayu and Ng, 2004" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[619,976,1897,1917]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F11115BFDBD765147BAFA58" box="[619,705,1897,1917]" italics="true" pageId="3">H. satriai</emphasis>
<bibRefCitation id="EFA44B012F11115BFD11765146ABFA58" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[711,976,1897,1917]" pageId="3" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004</bibRefCitation>
</taxonomicName>
, male (18.3 × 13.0 mm) (ZRC 2017.1043); D,
<taxonomicName id="4C354D732F11115BFF5E76BD4493FABC" authority="Yeo, Rahayu and Ng, 2004" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[136,488,1925,1945]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F11115BFF5E76BD45A6FABC" box="[136,221,1925,1945]" italics="true" pageId="3">H. satriai</emphasis>
<bibRefCitation id="EFA44B012F11115BFF3576BD4493FABC" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[227,488,1925,1945]" pageId="3" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004</bibRefCitation>
</taxonomicName>
, female (16.1 × 11.9 mm) (NMMBCD4086); E,
<taxonomicName id="4C354D732F11115BFC7076BD407DFABC" authority="Yeo, Rahayu and Ng, 2004" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[934,1286,1925,1945]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F11115BFC7076BD4680FABC" box="[934,1019,1925,1945]" italics="true" pageId="3">H. satriai</emphasis>
<bibRefCitation id="EFA44B012F11115BFBD776BD407DFABC" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[1025,1286,1925,1945]" pageId="3" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004</bibRefCitation>
</taxonomicName>
, male (11.1 × 8.2 mm) (NMMBCD4085); F,
<taxonomicName id="4C354D732F11115BFE4A76994782FA90" authority="Yeo, Rahayu and Ng, 2004" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[412,761,1953,1973]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F11115BFE4A7699448AFA90" box="[412,497,1953,1973]" italics="true" pageId="3">H. satriai</emphasis>
<bibRefCitation id="EFA44B012F11115BFE2076994782FA90" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[502,761,1953,1973]" pageId="3" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo, Rahayu and Ng, 2004</bibRefCitation>
</taxonomicName>
, male (6.8 × 5.4 mm) (NMMBCD4086).
</paragraph>
</caption>
<paragraph id="8B8A36F02F16115CFF787190448DFD9A" blockId="4.[174,502,168,191]" box="[174,502,168,191]" pageId="4">
<bibCitation id="5E1F3E9C2F16115CFF787190448DFD9A" box="[174,502,168,191]" journal="Zoological Studies" page="12" pageId="4" score="5.2987992369807255" volume="57" year="2018">
<emphasis id="B941EAE22F16115CFF787190441CFD9A" box="[174,359,168,191]" italics="true" pageId="4">Zoological Studies</emphasis>
<emphasis id="B941EAE22F16115CFEBB719044F5FD9A" bold="true" box="[365,398,168,191]" pageId="4">57:</emphasis>
12 (2018)
</bibCitation>
</paragraph>
<paragraph id="8B8A36F02F16115CFAFB719040D0FD9A" blockId="4.[1325,1451,168,191]" box="[1325,1451,168,191]" pageId="4">page 5 of 17</paragraph>
<caption id="DF4A66782F16115CFF7876504642FA91" pageId="4" startId="4.[174,210,1896,1916]" targetBox="[174,1451,258,1870]" targetPageId="4" targetType="figure">
<paragraph id="8B8A36F02F16115CFF7876504642FA91" blockId="4.[174,1452,1896,1972]" pageId="4">
<emphasis id="B941EAE22F16115CFF787650458AFA59" bold="true" box="[174,241,1896,1916]" pageId="4">Fig. 2.</emphasis>
<taxonomicName id="4C354D732F16115CFED5765047B2FA59" authority="(Bouvier, 1915)" baseAuthorityName="Bouvier" baseAuthorityYear="1915" box="[259,713,1896,1916]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F16115CFED576504764FA59" box="[259,543,1896,1916]" italics="true" pageId="4">Heteropilumnus decharmoyi</emphasis>
(
<bibRefCitation id="EFA44B012F16115CFDF8765047B9FA59" author="Bouvier EL" box="[558,706,1896,1916]" pageId="4" pagination="178 - 318" refId="ref5756" refString="Bouvier EL. 1915. Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique 48 (3): 178 - 318." type="journal article" year="1915">Bouvier, 1915</bibRefCitation>
)
</taxonomicName>
. A, B, E, F, male (16.8 × 12.7 mm) (NMNST); C, male (13.3 × 9.3 mm) (NMMBCD4084); D, female (8.6 × 5.9 mm) (ZRC 2017.1042). A, overall view; B-D, right side of carapace (denuded); E, frontal view of cephalothorax (right side denuded); F, left third maxilliped (denuded).
</paragraph>
</caption>
<paragraph id="8B8A36F02F17115DFFAB719044BDFD9A" blockId="5.[125,454,168,191]" box="[125,454,168,191]" pageId="5">
<bibCitation id="5E1F3E9C2F17115DFFAB719044BDFD9A" box="[125,454,168,191]" journal="Zoological Studies" page="12" pageId="5" score="5.2987992369807255" volume="57" year="2018">
<emphasis id="B941EAE22F17115DFFAB7190444CFD9A" box="[125,311,168,191]" italics="true" pageId="5">Zoological Studies</emphasis>
<emphasis id="B941EAE22F17115DFEEB71904425FD9A" bold="true" box="[317,350,168,191]" pageId="5">57:</emphasis>
12 (2018)
</bibCitation>
</paragraph>
<paragraph id="8B8A36F02F17115DFF5E71C54743FC33" blockId="5.[136,736,253,407]" box="[136,568,253,278]" pageId="5">
orange in small specimens (
<figureCitation id="130E2A752F17115DFE0571C54752FC33" box="[467,553,253,278]" captionStart="Fig" captionStartId="3.[136,170,1869,1889]" captionTargetBox="[143,1406,608,1843]" captionTargetId="figure-256@3.[143,1406,608,1843]" captionTargetPageId="3" captionText="Fig. 1. Colour in life. A, Heteropilumnus decharmoyi (Bouvier, 1915), male (16.8 × 12.7 mm) (NMNST); B, H. decharmoyi (Bouvier, 1915), male (13.3 × 9.3 mm) (NMMBCD4084); C, H. satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043); D, H. satriai Yeo, Rahayu and Ng, 2004, female (16.1 × 11.9 mm) (NMMBCD4086); E, H. satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085); F, H. satriai Yeo, Rahayu and Ng, 2004, male (6.8 × 5.4 mm) (NMMBCD4086)." pageId="5">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8B8A36F02F17115DFF13702540FEFD9A" blockId="5.[136,736,253,407]" lastBlockId="5.[1288,1413,168,191]" pageId="5">
<emphasis id="B941EAE22F17115DFF1370254445FC13" box="[197,318,285,310]" italics="true" pageId="5">Remarks</emphasis>
: This species was described from
<specimenCount id="9D33FD792F17115DFF077005442DFC73" box="[209,342,317,342]" count="1" pageId="5" type="male">one male</specimenCount>
, which was parasitized by a rhizocephalan and collected from the Indonesian Natuna Islands in the South
<collectingCountry id="F32276602F17115DFE2170454744FCB2" box="[503,575,381,407]" name="China" pageId="5">China</collectingCountry>
Sea (Yeo et page 6 of 17
</paragraph>
<paragraph id="8B8A36F02F17115DFCFB71C54129FC53" blockId="5.[813,1414,253,406]" pageId="5">
al. 2004). No colour notes were available at the time. The specimens from
<collectingCountry id="F32276602F17115DFBA7702541BFFC13" box="[1137,1220,285,310]" name="Taiwan" pageId="5">Taiwan</collectingCountry>
now allow us a better understanding of its colour, morphological characters and variation.
</paragraph>
<paragraph id="8B8A36F02F17115DFCBF704540F8FCB3" blockId="5.[813,1414,253,406]" box="[873,1411,381,406]" pageId="5">The teeth on the anterolateral margin</paragraph>
<caption id="DF4A66782F17115DFF5E76504788FA91" pageId="5" startId="5.[136,172,1896,1916]" targetBox="[136,1413,460,1870]" targetPageId="5" targetType="figure">
<paragraph id="8B8A36F02F17115DFF5E76504788FA91" blockId="5.[136,1413,1896,1972]" pageId="5">
<emphasis id="B941EAE22F17115DFF5E765045B7FA59" bold="true" box="[136,204,1896,1916]" pageId="5">Fig. 3.</emphasis>
<taxonomicName id="4C354D732F17115DFF08765047DDFA59" authority="(Bouvier, 1915)" baseAuthorityName="Bouvier" baseAuthorityYear="1915" box="[222,678,1896,1916]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F17115DFF0876504487FA59" box="[222,508,1896,1916]" italics="true" pageId="5">Heteropilumnus decharmoyi</emphasis>
(
<bibRefCitation id="EFA44B012F17115DFDDC765047DBFA59" author="Bouvier EL" box="[522,672,1896,1916]" pageId="5" pagination="178 - 318" refId="ref5756" refString="Bouvier EL. 1915. Decapodes marcheurs (Reptantia) et Stomatopodes recueillis a l'ile Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique 48 (3): 178 - 318." type="journal article" year="1915">Bouvier, 1915</bibRefCitation>
)
</taxonomicName>
. A, B, C-E, male (16.8 × 12.7 mm) (NMNST); F, male (13.3 × 9.3 mm) (NMMBCD4084); G, female (8.6 × 5.9 mm) (ZRC 2017.1042). A, anterior thoracic sternum and pleon; B, pleon; C, outer view of right chela; D, outer view of left chela; E-G, right fourth ambulatory leg.
</paragraph>
</caption>
<paragraph id="8B8A36F02F14115EFF787190448DFD9A" blockId="6.[174,502,168,191]" box="[174,502,168,191]" pageId="6">
<bibCitation id="5E1F3E9C2F14115EFF787190448DFD9A" box="[174,502,168,191]" journal="Zoological Studies" page="12" pageId="6" score="5.2987992369807255" volume="57" year="2018">
<emphasis id="B941EAE22F14115EFF787190441CFD9A" box="[174,359,168,191]" italics="true" pageId="6">Zoological Studies</emphasis>
<emphasis id="B941EAE22F14115EFEBB719044F5FD9A" bold="true" box="[365,398,168,191]" pageId="6">57:</emphasis>
12 (2018)
</bibCitation>
</paragraph>
<paragraph id="8B8A36F02F14115EFF7871C547DDFE53" blockId="6.[174,776,253,1047]" pageId="6">
of smaller specimens (
<emphasis id="B941EAE22F14115EFE3D71C54765FC33" box="[491,542,253,278]" italics="true" pageId="6">e.g.</emphasis>
, 11.1 ×
<quantity id="4CCD9B152F14115EFD4E71C5467AFC33" box="[664,769,253,278]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="6" unit="mm" value="8.2">8.2 mm</quantity>
,
<collectionCode id="ED24AE352F14115EFF787025444AFC13" box="[174,305,285,311]" pageId="6">NMMBCD</collectionCode>
4085) are less well defined, with the lobes relatively lower (
<figureCitation id="130E2A752F14115EFDC270054706FC73" box="[532,637,317,342]" captionStart="Fig" captionStartId="8.[174,209,1896,1916]" captionTargetBox="[180,1445,249,1869]" captionTargetId="figure-50@8.[180,1445,249,1869]" captionTargetPageId="8" captionText="Fig. 6. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004. A, B, E, male (11.1 × 8.2 mm) (NMMBCD4085); C, D, F, male (18.3 × 13.0 mm) (ZRC 2017.1043). A, C, overall view; B, D, right side of carapace (denuded); E, F, right fourth ambulatory leg." pageId="6">Fig. 6B</figureCitation>
). In larger specimens (
<emphasis id="B941EAE22F14115EFE927065440AFC53" box="[324,369,349,374]" italics="true" pageId="6">e.g.</emphasis>
,
<typeStatus id="548E88522F14115EFE5270654490FC53" box="[388,491,349,374]" pageId="6" type="holotype">holotype</typeStatus>
male and male 18.3 × 13.0 mm,
<collectionCode id="ED24AE352F14115EFEF97045441CFCB2" box="[303,359,381,407]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6">ZRC</collectionCode>
2017.1043); the teeth are better defined and separated by more distinct clefts (
<figureCitation id="130E2A752F14115EFF637085445DFCF3" box="[181,294,445,470]" captionStart="Fig" captionStartId="7.[136,170,1896,1916]" captionText="Fig. 5. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (19.3 × 13 mm) (MZB Cru 1668), Indonesia. A, overall view; B, right side of carapace (denuded); C, frontal view of cephalothorax (right side denuded); D, right third maxilliped (denuded); E, outer view of chelae; F, anterior thoracic sternum and pleon; G, right fourth ambulatory leg." pageId="6">Figs. 5B</figureCitation>
,
<figureCitation id="130E2A752F14115EFEE370854426FCF3" box="[309,349,445,470]" captionStart="Fig" captionStartId="8.[174,209,1896,1916]" captionTargetBox="[180,1445,249,1869]" captionTargetId="figure-50@8.[180,1445,249,1869]" captionTargetPageId="8" captionText="Fig. 6. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004. A, B, E, male (11.1 × 8.2 mm) (NMMBCD4085); C, D, F, male (18.3 × 13.0 mm) (ZRC 2017.1043). A, C, overall view; B, D, right side of carapace (denuded); E, F, right fourth ambulatory leg." pageId="6">6D</figureCitation>
). The male pleon of the
<typeStatus id="548E88522F14115EFD4A7085467DFCF3" box="[668,774,445,470]" pageId="6" type="holotype">holotype</typeStatus>
male is atypical as the specimen was infected with a rhizocephalan (
<figureCitation id="130E2A752F14115EFE1470C5475AFF33" box="[450,545,509,534]" captionStart="Fig" captionStartId="8.[174,209,1896,1916]" captionTargetBox="[180,1445,249,1869]" captionTargetId="figure-50@8.[180,1445,249,1869]" captionTargetPageId="8" captionText="Fig. 6. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004. A, B, E, male (11.1 × 8.2 mm) (NMMBCD4085); C, D, F, male (18.3 × 13.0 mm) (ZRC 2017.1043). A, C, overall view; B, D, right side of carapace (denuded); E, F, right fourth ambulatory leg." pageId="6">Fig. 6F</figureCitation>
). The pleon of the uninfected males is more triangular, with the telson semicircular in shape (
<figureCitation id="130E2A752F14115EFE6F7305473EFF73" box="[441,581,573,599]" captionStart="Fig" captionStartId="9.[136,170,1864,1884]" captionTargetBox="[136,1413,249,1838]" captionTargetId="figure-46@9.[136,1413,249,1838]" captionTargetPageId="9" captionText="Fig. 7. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085). A, frontal view of cephalothorax (right side denuded); B, left third maxilliped (denuded); C, anterior thoracic sternum and pleon; D, pleon; E, outer view of chelae." pageId="6">Figs. 7C, D</figureCitation>
,
<figureCitation id="130E2A752F14115EFD87730547E2FF73" box="[593,665,573,599]" captionStart="Fig" captionStartId="10.[174,208,1864,1884]" captionTargetBox="[174,1451,276,1838]" captionTargetId="figure-47@10.[174,1451,276,1838]" captionTargetPageId="10" captionText="Fig. 8. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (18.3 × 13.0 mm) (ZRC 2017.1043). A, frontal view of cephalothorax (right side denuded); B, left third maxilliped (denuded); C, anterior thoracic sternum and pleon; D, pleon; E, outer view of pleon." pageId="6">8C, D</figureCitation>
). The G1 of the present material varies slightly with size. In the larger male (18.3 × 13.0 mm,
<collectionCode id="ED24AE352F14115EFDEC7345470AFFB2" box="[570,625,637,663]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6">ZRC</collectionCode>
2017.1043), the distal elongated part of the G1 is gently curved to relatively straight (
<figureCitation id="130E2A752F14115EFE6273854771FFF2" box="[436,522,701,727]" captionStart="Fig" captionStartId="11.[136,170,1897,1917]" captionText="Fig. 9. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, gonopods. A-D, male (18.3 × 13.0 mm) (ZRC 2017.1043); E-H, male (11.1 × 8.2 mm) (NMMBCD4085). A, E, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsal view); D, H, left G2. Scale bars = 0.5 mm." pageId="6">Figs. 9</figureCitation>
A-C) whereas in the smaller male (11.1 × 8.2 mmm,
<collectionCode id="ED24AE352F14115EFDE973E547C4FFD2" box="[575,703,733,759]" pageId="6">NMMBCD</collectionCode>
4085), it is gently sinuous with the tip gently curved upwards (
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E-G). The form of the G1 of the smaller male agrees very well with that figured by
<bibRefCitation id="EFA44B012F14115EFF7872654417FE53" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[174,364,861,886]" pageId="6" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo et al. (2004</bibRefCitation>
: fig. 1e, f) for the
<typeStatus id="548E88522F14115EFDEF726547D9FE53" box="[569,674,861,886]" pageId="6" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8A36F02F14115EFF3C724540D0FD9A" blockId="6.[174,776,253,1047]" lastBlockId="6.[1325,1451,168,191]" pageId="6">
The differences between
<taxonomicName id="4C354D732F14115EFDE4724547DEFEB3" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[562,677,893,918]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F14115EFDE4724547DEFEB3" box="[562,677,893,918]" italics="true" pageId="6">H. satriai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C354D732F14115EFD3D72454442FE93" baseAuthorityName="Bouvier" baseAuthorityYear="1915" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="decharmoyi">
<emphasis id="B941EAE22F14115EFD3D72454442FE93" italics="true" pageId="6">H. decharmoyi</emphasis>
</taxonomicName>
have been discussed under the latter species.
<taxonomicName id="4C354D732F14115EFECB7285474BFEF3" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[285,560,957,982]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F14115EFECB7285474BFEF3" box="[285,560,957,982]" italics="true" pageId="6">Heteropilumnus satriai</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C354D732F14115EFF7872E54774FED3" authority="Ward, 1933" authorityName="Ward" authorityYear="1933" box="[174,527,989,1014]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B941EAE22F14115EFF7872E5440FFED3" box="[174,372,989,1014]" italics="true" pageId="6">H. granulimanus</emphasis>
<bibRefCitation id="EFA44B012F14115EFEA872E54774FED3" author="Ward M." box="[382,527,989,1014]" pageId="6" pagination="377 - 394" refId="ref6873" refString="Ward M. 1933. New genera and species of marine Decapod Brachyura from the coasts of New South Wales and Queensland. Australian Zoologist 7: 377 - 394." type="journal article" year="1933">Ward, 1933</bibRefCitation>
</taxonomicName>
, described from
<specimenCount id="9D33FD792F14115EFD0C72E54593F933" count="1" pageId="6" type="male">one male</specimenCount>
measuring 12.0 × 9.0 mm from the Capricorn page 7 of 17
</paragraph>
<paragraph id="8B8A36F02F14115EFC8471C54693FFB3" blockId="6.[850,1452,253,790]" pageId="6">
Group of islands in
<collectingRegion id="49F1F8122F14115EFB9D71C5419FFC32" box="[1099,1252,253,279]" country="Australia" name="Queensland" pageId="6">Queensland</collectingRegion>
,
<collectingCountry id="F32276602F14115EFB2771C5401AFC33" box="[1265,1377,253,278]" name="Australia" pageId="6">Australia</collectingCountry>
. In
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<emphasis id="B941EAE22F14115EFA4671C54689FC13" italics="true" pageId="6">H. granulimanus</emphasis>
</taxonomicName>
; however, the frontal margin is more truncate with the two lobes separated by a narrow fissure (
<bibRefCitation id="EFA44B012F14115EFC6670654141FC53" author="Ward M." box="[944,1082,349,374]" pageId="6" pagination="377 - 394" refId="ref6873" refString="Ward M. 1933. New genera and species of marine Decapod Brachyura from the coasts of New South Wales and Queensland. Australian Zoologist 7: 377 - 394." type="journal article" year="1933">Ward 1933</bibRefCitation>
: pl. 2 fig. 3) (vs. the two frontal lobes separated by a distinct V-shaped cleft in
<taxonomicName id="4C354D732F14115EFA47704546E2FC93" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F14115EFA47704546E2FC93" italics="true" pageId="6">H. satriai</emphasis>
</taxonomicName>
,
<figureCitation id="130E2A752F14115EFC7C70A5416AFC93" box="[938,1041,413,438]" captionStart="Fig" captionStartId="7.[136,170,1896,1916]" captionText="Fig. 5. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (19.3 × 13 mm) (MZB Cru 1668), Indonesia. A, overall view; B, right side of carapace (denuded); C, frontal view of cephalothorax (right side denuded); D, right third maxilliped (denuded); E, outer view of chelae; F, anterior thoracic sternum and pleon; G, right fourth ambulatory leg." pageId="6">Figs. 5B</figureCitation>
,
<figureCitation id="130E2A752F14115EFBC870A5411DFC93" box="[1054,1126,413,438]" captionStart="Fig" captionStartId="8.[174,209,1896,1916]" captionTargetBox="[180,1445,249,1869]" captionTargetId="figure-50@8.[180,1445,249,1869]" captionTargetPageId="8" captionText="Fig. 6. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004. A, B, E, male (11.1 × 8.2 mm) (NMMBCD4085); C, D, F, male (18.3 × 13.0 mm) (ZRC 2017.1043). A, C, overall view; B, D, right side of carapace (denuded); E, F, right fourth ambulatory leg." pageId="6">6B, D</figureCitation>
); the granules on the outer surface of the chela are more rounded, densely packed and arranged in rows (
<bibRefCitation id="EFA44B012F14115EFB0370E5401DFCD3" author="Ward M." box="[1237,1382,477,502]" pageId="6" pagination="377 - 394" refId="ref6873" refString="Ward M. 1933. New genera and species of marine Decapod Brachyura from the coasts of New South Wales and Queensland. Australian Zoologist 7: 377 - 394." type="journal article" year="1933">Ward 1933</bibRefCitation>
: pl. 2 fig. 4) (vs. the granules are rounded to conical and distributed evenly on surface in
<taxonomicName id="4C354D732F14115EFB1C73254048FF13" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[1226,1331,541,566]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F14115EFB1C73254048FF13" box="[1226,1331,541,566]" italics="true" pageId="6">H. satriai</emphasis>
</taxonomicName>
,
<figureCitation id="130E2A752F14115EFA94732540D3FF13" box="[1346,1448,541,566]" captionStart="Fig" captionStartId="7.[136,170,1896,1916]" captionText="Fig. 5. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (19.3 × 13 mm) (MZB Cru 1668), Indonesia. A, overall view; B, right side of carapace (denuded); C, frontal view of cephalothorax (right side denuded); D, right third maxilliped (denuded); E, outer view of chelae; F, anterior thoracic sternum and pleon; G, right fourth ambulatory leg." pageId="6">Figs. 5E</figureCitation>
,
<figureCitation id="130E2A752F14115EFC847305460CFF73" box="[850,887,573,598]" captionStart="Fig" captionStartId="9.[136,170,1864,1884]" captionTargetBox="[136,1413,249,1838]" captionTargetId="figure-46@9.[136,1413,249,1838]" captionTargetPageId="9" captionText="Fig. 7. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085). A, frontal view of cephalothorax (right side denuded); B, left third maxilliped (denuded); C, anterior thoracic sternum and pleon; D, pleon; E, outer view of chelae." pageId="6">7E</figureCitation>
); and the male telson is relatively wider (
<bibRefCitation id="EFA44B012F14115EFABF730546EFFF53" author="Ward M." pageId="6" pagination="377 - 394" refId="ref6873" refString="Ward M. 1933. New genera and species of marine Decapod Brachyura from the coasts of New South Wales and Queensland. Australian Zoologist 7: 377 - 394." type="journal article" year="1933">Ward 1933</bibRefCitation>
: pl. 2 fig. 4) (vs. less wide in in
<taxonomicName id="4C354D732F14115EFAE4736540D8FF53" authorityName="Yeo, Rahayu and Ng" authorityYear="2004" box="[1330,1443,605,630]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="satriai">
<emphasis id="B941EAE22F14115EFAE4736540D8FF53" box="[1330,1443,605,630]" italics="true" pageId="6">H. satriai</emphasis>
</taxonomicName>
,
<figureCitation id="130E2A752F14115EFC84734546A0FFB3" box="[850,987,637,663]" captionStart="Fig" captionStartId="9.[136,170,1864,1884]" captionTargetBox="[136,1413,249,1838]" captionTargetId="figure-46@9.[136,1413,249,1838]" captionTargetPageId="9" captionText="Fig. 7. Heteropilumnus satriai Yeo, Rahayu and Ng, 2004, male (11.1 × 8.2 mm) (NMMBCD4085). A, frontal view of cephalothorax (right side denuded); B, left third maxilliped (denuded); C, anterior thoracic sternum and pleon; D, pleon; E, outer view of chelae." pageId="6">Figs. 7C, D</figureCitation>
).
</paragraph>
<paragraph id="8B8A36F02F14115EFC5973A54691FE33" blockId="6.[850,1452,253,790]" pageId="6">
<emphasis id="B941EAE22F14115EFC5973A5416FFF93" box="[911,1044,669,694]" italics="true" pageId="6">Distribution</emphasis>
: The species was described from the Natunas Islands in
<collectingCountry id="F32276602F14115EFB567385407BFFF3" box="[1152,1280,701,726]" name="Indonesia" pageId="6">Indonesia</collectingCountry>
, in the South
<collectingCountry id="F32276602F14115EFC8473E546E3FFD2" box="[850,920,733,759]" name="China" pageId="6">China</collectingCountry>
Sea (
<bibRefCitation id="EFA44B012F14115EFC3773E541DAFFD3" author="Yeo DCJ &amp; Rahayu DL &amp; Ng PKL" box="[993,1185,733,758]" pageId="6" pagination="79 - 88" refId="ref6943" refString="Yeo DCJ, Rahayu DL, Ng PKL. 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles B Zool, Supplement 11: 79 - 88." type="journal article" year="2004">Yeo et al. 2004</bibRefCitation>
), and is now recorded from
<collectingCountry id="F32276602F14115EFC5973C5469DFE33" box="[911,998,765,790]" name="Taiwan" pageId="6">Taiwan</collectingCountry>
.
</paragraph>
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189
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Genus
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<emphasis id="1F89EA9358537E3AE6232019FE3EFE49" bold="true" box="[340,475,380,405]" italics="true" pageId="2">Mesocallis</emphasis>
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<taxonomicName id="EAFD4D0258537E3AE7D920B3FE06FE36" ID-CoL="8PQ9Q" authority="Matsumura, 1919: 104" authorityName="Matsumura" authorityPageNumber="104" authorityYear="1919" box="[174,483,470,490]" class="Insecta" family="Aphididae" genus="Neocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE7D920B3FEE0FE36" box="[174,261,470,490]" italics="true" pageId="2">Neocallis</emphasis>
<bibRefCitation id="496C4B7058537E3AE67D20B3FE06FE36" author="Matsumura S." box="[266,483,470,490]" pageId="2" pagination="99 - 114" refId="ref10297" refString="Matsumura S. 1919. New species and genera of Callipterinae (Aphididae) of Japan. Trans Sapporo Nat Hist Soc 7: 99 - 114." type="journal article" year="1919">Matsumura, 1919: 104</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D42368158537E3AE7D92094FDDDFDD9" blockId="2.[174,775,444,570]" box="[174,568,497,517]" pageId="2">
<treatmentCitationGroup id="0DED11AF58537E3AE7D92094FDDDFDD9" box="[174,568,497,517]" pageId="2">
<treatmentCitation id="AC5C109058537E3AE7D92094FDD1FDD9" author="Takahashi R." box="[174,564,497,517]" page="247" pageId="2" year="1961">
<taxonomicName id="EAFD4D0258537E3AE7D92094FDD1FDD9" ID-CoL="8PQCV" authority="Takahashi, 1961: 247" authorityName="Takahashi" authorityPageNumber="247" authorityYear="1961" box="[174,564,497,517]" class="Insecta" family="Aphididae" genus="Nippochaitophorus" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE7D92094FE85FDD9" box="[174,352,497,517]" italics="true" pageId="2">Nippochaitophorus</emphasis>
<bibRefCitation id="496C4B7058537E3AE6112094FDD1FDD9" author="Takahashi R." box="[358,564,497,517]" pageId="2" pagination="3 - 9" refId="ref10680" refString="Takahashi R. 1961. Four new species of Callipterinae from Japan (Aphididae, Homoptera). Mushi 35 (2): 3 - 9." type="journal article" year="1961">Takahashi, 1961: 247</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="65E7650A58537E3AE7D92369FE47FDE6" pageId="2" type="materials_examined">
<paragraph id="2D42368158537E3AE7D92369FE47FDE6" blockId="2.[174,775,444,570]" pageId="2">
<typeStatus id="F246882358537E3AE7D92369FF38FDFC" box="[174,221,524,544]" pageId="2">Type</typeStatus>
species:
<taxonomicName id="EAFD4D0258537E3AE6492369FD00FDFC" authority="(Matsumura, 1917)" baseAuthorityName="Matsumura" baseAuthorityYear="1917" box="[318,741,524,544]" class="Insecta" family="Aphididae" genus="Myzocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="species" species="sawashibae">
<emphasis id="1F89EA9358537E3AE6492369FDFBFDFC" box="[318,542,524,544]" italics="true" pageId="2">Myzocallis sawashibae</emphasis>
(
<bibRefCitation id="496C4B7058537E3AE55B2369FD05FDFC" author="Matsumura S." box="[556,736,524,544]" pageId="2" pagination="351 - 414" refId="ref10261" refString="Matsumura S. 1917. A list of the Aphididae of Japan, with description of new species and genera. J Coll Agric Hokkaido Imp Univ 7 (6): 351 - 414." type="journal article" year="1917">Matsumura, 1917</bibRefCitation>
)
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="65E7650A58537E3AE79D2304FBDAFE6B" pageId="2" type="diagnosis">
<paragraph id="2D42368158537E3AE79D2304FDC5FC67" blockId="2.[174,775,609,987]" pageId="2">
<emphasis id="1F89EA9358537E3AE79D2304FE30FDA7" box="[234,469,609,635]" italics="true" pageId="2">
<taxonomicName id="EAFD4D0258537E3AE79D2304FEA8FDA6" box="[234,333,609,634]" class="Insecta" family="Noctuidae" genus="Generic" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">Generic</taxonomicName>
Diagnosis
</emphasis>
: According to
<bibRefCitation id="496C4B7058537E3AE5E32304FF1AFD47" author="Quednau FW" pageId="2" refId="ref10469" refString="Quednau FW. 2003. Atlas of the drepanosiphine aphids of the world part II: Panaphidini Oestrund, 1923 - Panaphidina Oestrund, 1923 (Hemiptera: Aphidinae: Calaphidinae). The American Entomological Institute, Florida, 301 pp." type="book" year="2003">Quednau (2003)</bibRefCitation>
,
<taxonomicName id="EAFD4D0258537E3AE67D23E7FE63FD47" authorityName="Matsumura" authorityYear="1919" box="[266,390,642,667]" class="Insecta" family="Aphididae" genus="Mesocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE67D23E7FE63FD47" box="[266,390,642,667]" italics="true" pageId="2">Mesocallis</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="EAFD4D0258537E3AE53723E7FD5EFD47" authorityName="Passerini" authorityYear="1860" box="[576,699,642,667]" class="Insecta" family="Aphididae" genus="Pterocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE53723E7FD5EFD47" box="[576,699,642,667]" italics="true" pageId="2">Pterocallis</emphasis>
</taxonomicName>
group of genera. This genus is similar to
<taxonomicName id="EAFD4D0258537E3AE5F723C7FEF8FD07" authority="Passerini" authorityName="Passerini" authorityYear="1860" class="Insecta" family="Aphididae" genus="Pterocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE5F723C7FCE2FD67" box="[640,775,674,699]" italics="true" pageId="2">Pterocallis</emphasis>
Passerini
</taxonomicName>
but can be distinguished by having two setae on ABD VIII, black-bordered fore tibiae and Cu1b of forewing distinctly dark bordered. Representatives of
<taxonomicName id="EAFD4D0258537E3AE6B32247FDB1FCE7" authorityName="Matsumura" authorityYear="1919" box="[452,596,802,827]" class="Insecta" family="Aphididae" genus="Mesocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE6B32247FDB1FCE7" box="[452,596,802,827]" italics="true" pageId="2">Mesocallis</emphasis>
</taxonomicName>
can be also distinguished from
<taxonomicName id="EAFD4D0258537E3AE6E92227FDFAFC87" authorityName="Passerini" authorityYear="1860" box="[414,543,834,859]" class="Insecta" family="Aphididae" genus="Pterocallis" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="1F89EA9358537E3AE6E92227FDFAFC87" box="[414,543,834,859]" italics="true" pageId="2">Pterocallis</emphasis>
</taxonomicName>
by oval to narrow elliptical secondary rhinaria on ANT III, epicranial suture weakly developed but present on the head of apterous viviparous females.
</paragraph>
<paragraph id="2D42368158537E3AE79D22A7FA4EFF63" blockId="2.[174,775,609,987]" box="[234,1451,168,987]" lastBlockId="2.[1325,1451,168,191]" pageId="2">
<emphasis id="1F89EA9358537E3AE79D22A7FE8DFC07" box="[234,360,962,987]" italics="true" pageId="2">Host plant</emphasis>
: Species belonging to this genus page 3 of 20
</paragraph>
<paragraph id="2D42368158537E3AE4252198FBEBFE8A" blockId="2.[850,1452,253,439]" pageId="2">
are known to feed on
<taxonomicName id="EAFD4D0258537E3AE3122198FB09FECA" box="[1125,1260,253,278]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1F89EA9358537E3AE3122198FB4CFECA" box="[1125,1193,253,278]" italics="true" pageId="2">Alnus</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="EAFD4D0258537E3AE38E2198FA4DFECA" box="[1273,1448,253,278]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1F89EA9358537E3AE38E2198FA83FECA" box="[1273,1382,253,278]" italics="true" pageId="2">Carpinus</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="EAFD4D0258537E3AE4252078FC08FEEA" box="[850,1005,285,311]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1F89EA9358537E3AE4252078FC49FEEB" box="[850,940,285,311]" italics="true" pageId="2">Corylus</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="EAFD4D0258537E3AE48E2078FB66FEEA" box="[1017,1155,285,311]" class="Magnoliopsida" family="Betulaceae" genus="Ostrya" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1F89EA9358537E3AE48E2078FBADFEEB" box="[1017,1096,285,311]" italics="true" pageId="2">Ostrya</emphasis>
spp.
</taxonomicName>
(
<taxonomicName id="EAFD4D0258537E3AE3E52078FAF8FEEA" authorityName="S.F.Gray" authorityYear="1822" box="[1170,1309,285,310]" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
) and
<taxonomicName id="EAFD4D0258537E3AE2162078FC92FE8A" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1F89EA9358537E3AE2162078FA4EFEEA" box="[1377,1451,285,310]" italics="true" pageId="2">Fagus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="EAFD4D0258537E3AE4F32058FBE1FE8A" authorityName="Dumortier" authorityYear="1829" box="[900,1028,317,342]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="2" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
).
</paragraph>
<paragraph id="2D42368158537E3AE4F82038FBDAFE6B" blockId="2.[850,1452,253,439]" pageId="2">
<emphasis id="1F89EA9358537E3AE4F82038FBF1FEAA" box="[911,1044,349,374]" italics="true" pageId="2">Distribution</emphasis>
: This genus is known from South- East Asian countries (
<collectingCountry id="55EA761158537E3AE32B2018FB4CFE4B" box="[1116,1193,381,407]" name="China" pageId="2">China</collectingCountry>
,
<collectingCountry id="55EA761158537E3AE3C12018FB10FE4A" box="[1206,1269,381,406]" name="India" pageId="2">India</collectingCountry>
,
<collectingCountry id="55EA761158537E3AE2752018FAAAFE4A" box="[1282,1359,381,406]" name="Japan" pageId="2">Japan</collectingCountry>
,
<collectingCountry id="55EA761158537E3AE22B2018FA4DFE4A" box="[1372,1448,381,406]" name="South Korea" pageId="2">Korea</collectingCountry>
,
<collectingCountry id="55EA761158537E3AE42520F8FC79FE6A" box="[850,924,413,438]" name="Nepal" pageId="2">Nepal</collectingCountry>
, and Siberia).
</paragraph>
</subSubSection>
</treatment>
</document>