Add updates up until 2024-12-13 02:40:40
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<mods:title id="F93DAEC69DE879502C0E516BFF247565">Two new species of Trichodrilus (Annelida, Clitellata, Lumbriculidae) from Pacific drainages, western USA</mods:title>
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<mods:title id="6B581A1ADAA0F9A263BB6872BE97DA63">Two new species of Trichodrilus (Annelida, Clitellata, Lumbriculidae) from Pacific drainages, western USA</mods:title>
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<mods:namePart id="D4C359B924590A9FEC3C4ED8898877FE">Fend, Steven</mods:namePart>
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<mods:affiliation id="D81BC3B30165B54A79A7E3ED93DB8C10">756 W Vista Hermosa Dr., Green Valley, AZ 85614, USA.</mods:affiliation>
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<mods:namePart id="B1E7134C56D9EFEE96522BB29CBF43FE">Fend, Steven</mods:namePart>
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<mods:affiliation id="AF52BE04FE3793A94E303A6C92FB203B">756 W Vista Hermosa Dr., Green Valley, AZ 85614, USA.</mods:affiliation>
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<mods:namePart id="9186A629AE40E480F8FC296AD5E83D6E">Rodriguez, Pilar</mods:namePart>
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<mods:affiliation id="674825A12387E0C084911C9A4F79EF99">Zoology and Animal Cell Biology Dpt., Faculty of Science and Technology, University of the Basque Country UPV / EHU, Apdo. 644, Bilbao, 48080 Spain.</mods:affiliation>
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<mods:namePart id="14F831A648D986CF391E0BB85AF0D2EF">Rodriguez, Pilar</mods:namePart>
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<mods:affiliation id="658BCCDD9357A34F49EF2A8ED020C835">Zoology and Animal Cell Biology Dpt., Faculty of Science and Technology, University of the Basque Country UPV / EHU, Apdo. 644, Bilbao, 48080 Spain.</mods:affiliation>
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<mods:title id="0E2D3079475821C642202DD9B82425E9">Zootaxa</mods:title>
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<mods:title id="A320961FA5FBA73F544D6D3D176EB7F4">Zootaxa</mods:title>
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<mods:part id="CB2C483A6D6AA8782248C735AD766FA7">
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<mods:date id="0DBA3969EC0BDC3ED027550786757ACF">2024</mods:date>
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<mods:number id="596CACAAD5F194A957F63BFB41354E8A">2024-12-11</mods:number>
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<mods:number id="78F797EF0B6FFB18E3B7BD57762E3AB5">5551</mods:number>
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<treatment id="03F487C0FF937559D7E4C7DEC851FE28" LSID="urn:lsid:plazi:treatment:03F487C0FF937559D7E4C7DEC851FE28" httpUri="http://treatment.plazi.org/id/03F487C0FF937559D7E4C7DEC851FE28" lastPageId="10" lastPageNumber="151" pageId="5" pageNumber="146">
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<treatment id="03F487C0FF937559D7E4C7DEC851FE28" ID-DOI="http://doi.org/10.5281/zenodo.14390798" ID-Zenodo-Dep="14390798" LSID="urn:lsid:plazi:treatment:03F487C0FF937559D7E4C7DEC851FE28" httpUri="http://treatment.plazi.org/id/03F487C0FF937559D7E4C7DEC851FE28" lastPageId="10" lastPageNumber="151" pageId="5" pageNumber="146">
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<heading id="D0AA81BAFF937556D7E4C7DEC862F8A4" bold="true" box="[151,547,1770,1797]" fontSize="11" level="1" pageId="5" pageNumber="146" reason="1">
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|
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@ -71,35 +71,25 @@
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</paragraph>
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<subSubSection id="C347655DFF937554D7E4C663C97EFEE0" lastPageId="7" lastPageNumber="148" pageId="5" pageNumber="146" type="materials_examined">
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<paragraph id="8BE236D6FF937556D7E4C663CB63F834" blockId="5.[151,1436,1878,2013]" pageId="5" pageNumber="146">
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<paragraph id="8BE236D6FF937556D7E4C663CBB1F87C" blockId="5.[151,1436,1878,2013]" pageId="5" pageNumber="146">
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<emphasis id="B929EAC4FF937556D7E4C663CB78F8D1" bold="true" box="[151,313,1878,1905]" pageId="5" pageNumber="146">Type locality:</emphasis>
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||||
<materialsCitation id="3B353C8BFF937556D64CC663CB5FF834" collectorName="Grays Harbor Co. West Fork Humptulips River" county="Grays Harbor Co." latitude="47.4812" location="Harbor Co. West Fork Humptulips River" longLatPrecision="6" longitude="-123.6831" pageId="5" pageNumber="146" specimenCount="1" stateProvince="Washington">
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<materialsCitation id="3B353C8BFF937556D64CC663CBB1F87C" collectingDate="2003-06-02" collectionCode="USNM" country="Canada" latitude="47.4812" location="near Campbell Tree Grove" longLatPrecision="6" longitude="-123.6831" municipality="Grays Harbor Co. West Fork Humptulips River" pageId="5" pageNumber="146" specimenCode="USNM 1740731" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<collectingRegion id="4999F834FF937556D64CC663CB86F8D0" box="[319,455,1879,1905]" country="United States of America" name="Washington" pageId="5" pageNumber="146">Washington</collectingRegion>
|
||||
,
|
||||
<collectorName id="26A85300FF937556D6A1C663C9A5F8D0" box="[466,996,1879,1905]" pageId="5" pageNumber="146">
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<collectingCounty id="62834E5AFF937556D6A1C663C8DBF8D0" box="[466,666,1879,1905]" pageId="5" pageNumber="146">
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||||
Grays
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||||
<location id="8E82600DFF937556D56FC663C8DBF8D0" LSID="urn:lsid:plazi:treatment:03F487C0FF937559D7E4C7DEC851FE28:8E82600DFF937556D56FC663C8DBF8D0" box="[540,666,1879,1905]" county="Grays Harbor Co." latitude="47.4812" longLatPrecision="6" longitude="-123.6831" name="Harbor Co. West Fork Humptulips River" pageId="5" pageNumber="146" stateProvince="Washington">Harbor Co.</location>
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||||
</collectingCounty>
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||||
West Fork Humptulips River
|
||||
</collectorName>
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||||
, near Campbell Tree Grove,
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||||
<collectingMunicipality id="6B86ACACFF937556D6A1C663C9A5F8D0" box="[466,996,1879,1905]" pageId="5" pageNumber="146">Grays Harbor Co. West Fork Humptulips River</collectingMunicipality>
|
||||
,
|
||||
<location id="8E82600DFF937556D49CC663CF5AF8D0" LSID="urn:lsid:plazi:treatment:03F487C0FF937559D7E4C7DEC851FE28:8E82600DFF937556D49CC663CF5AF8D0" box="[1007,1307,1879,1905]" country="Canada" latitude="47.4812" longLatPrecision="6" longitude="-123.6831" municipality="Grays Harbor Co. West Fork Humptulips River" name="near Campbell Tree Grove" pageId="5" pageNumber="146" stateProvince="Washington">near Campbell Tree Grove</location>
|
||||
,
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||||
<geoCoordinate id="EE695011FF937556D255C663CFD9F8D0" box="[1318,1432,1879,1905]" degrees="47.4812" direction="north" orientation="latitude" pageId="5" pageNumber="146" precision="5" value="47.4812">N47.4812</geoCoordinate>
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,
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<geoCoordinate id="EE695011FF937556D7E4C64FCB5FF834" box="[151,286,1915,1941]" degrees="123.6831" direction="west" orientation="longitude" pageId="5" pageNumber="146" precision="5" value="-123.6831">W123.6831</geoCoordinate>
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</materialsCitation>
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||||
.
|
||||
</paragraph>
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||||
<paragraph id="8BE236D6FF937556D7B4C6ABCBB1F87C" blockId="5.[151,1436,1878,2013]" pageId="5" pageNumber="146">
|
||||
<materialsCitation id="3B353C8BFF937556D7B4C6ABCBB1F87C" collectingDate="2003-06-02" collectionCode="USNM" country="Canada" location="Canada" pageId="5" pageNumber="146" specimenCode="USNM 1740731" specimenCount="1" typeStatus="holotype">
|
||||
.
|
||||
<emphasis id="B929EAC4FF937556D7B4C6ABCB7CF818" bold="true" box="[199,317,1951,1977]" pageId="5" pageNumber="146">
|
||||
<typeStatus id="54E68874FF937556D7B4C6ABCB79F818" box="[199,312,1951,1977]" pageId="5" pageNumber="146" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCode id="DBFB9EADFF937556D630C6ABC841F818" box="[323,512,1951,1977]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="146" type="Museum">USNM 1740731</specimenCode>
|
||||
: A whole mounted worm, stained with borax carmine, and slide-mounted in
|
||||
<collectingCountry id="F34A7646FF937556D23BC6ABCFDDF818" box="[1352,1436,1951,1977]" name="Canada" pageId="5" pageNumber="146">Canada</collectingCountry>
|
||||
balsam; collected
|
||||
: A whole mounted worm, stained with borax carmine, and slide-mounted in Canada balsam; collected
|
||||
<date id="FFE31016FF937556D612C6F7CBACF87C" box="[353,493,1987,2013]" pageId="5" pageNumber="146" value="2003-06-02">
|
||||
<collectingDate id="EFA7E9FEFF937556D612C6F7CBACF87C" box="[353,493,1987,2013]" pageId="5" pageNumber="146" value="2003-06-02">2 June 2003</collectingDate>
|
||||
</date>
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||||
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@ -131,15 +121,13 @@ Reproductive segments of a partially mature worm.
|
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</paragraph>
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</caption>
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||||
<paragraph id="8BE236D6FF917554D7B4C1A3C9E3FF74" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="148">
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<materialsCitation id="3B353C8BFF917554D7B4C1A3C9DFFF74" collectingDate="2003-06-02" collectingDateMax="2022-04-29" collectingDateMin="2003-06-02" collectionCode="USNM" country="Canada" location="From" municipality="All" pageId="7" pageNumber="148" specimenCode="USNM 1740732-1740734" specimenCount="1" typeStatus="paratype">
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<materialsCitation id="3B353C8BFF917554D7B4C1A3C9DFFF74" collectingDate="2003-06-02" collectingDateMax="2022-04-29" collectingDateMin="2003-06-02" collectionCode="USNM" pageId="7" pageNumber="148" specimenCode="USNM 1740732-1740734" specimenCount="1" typeStatus="paratype">
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<emphasis id="B929EAC4FF917554D7B4C1A3CB09FF10" bold="true" box="[199,328,151,177]" pageId="7" pageNumber="148">
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<typeStatus id="54E68874FF917554D7B4C1A3CB02FF10" box="[199,323,151,177]" pageId="7" pageNumber="148" type="paratype">Paratypes</typeStatus>
|
||||
:
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||||
</emphasis>
|
||||
<specimenCode id="DBFB9EADFF917554D621C1A3C8C5FF10" box="[338,644,151,177]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="148" type="Museum">USNM 1740732–1740734</specimenCode>
|
||||
:
|
||||
<location id="8E82600DFF917554D5E2C1A3C88FFF10" LSID="urn:lsid:plazi:treatment:03F487C0FF937559D7E4C7DEC851FE28:8E82600DFF917554D5E2C1A3C88FFF10" box="[657,718,151,177]" country="Canada" municipality="All" name="From" pageId="7" pageNumber="148">From</location>
|
||||
the type locality, 2 dissected, collected
|
||||
: From the type locality, 2 dissected, collected
|
||||
<date id="FFE31016FF917554D3D3C1A3CF70FF10" box="[1184,1329,151,177]" pageId="7" pageNumber="148" value="2003-06-02">
|
||||
<collectingDate id="EFA7E9FEFF917554D3D3C1A3CF70FF10" box="[1184,1329,151,177]" pageId="7" pageNumber="148" value="2003-06-02">2 June 2003</collectingDate>
|
||||
</date>
|
||||
|
|
@ -147,23 +135,14 @@ the type locality, 2 dissected, collected
|
|||
<date id="FFE31016FF917554D62AC18FCBB9FF74" box="[345,504,187,213]" pageId="7" pageNumber="148" value="2022-04-29">
|
||||
<collectingDate id="EFA7E9FEFF917554D62AC18FCBB9FF74" box="[345,504,187,213]" pageId="7" pageNumber="148" value="2022-04-29">29 April 2022</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectingMunicipality id="6B86ACACFF917554D572C18FC864FF74" box="[513,549,187,213]" pageId="7" pageNumber="148">All</collectingMunicipality>
|
||||
slide-mounted in
|
||||
<collectingCountry id="F34A7646FF917554D583C18FC905FF74" box="[752,836,187,213]" name="Canada" pageId="7" pageNumber="148">Canada</collectingCountry>
|
||||
balsam
|
||||
. All slide-mounted in Canada balsam
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF917554D7B4C1EAC97EFEE0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="148">
|
||||
<materialsCitation id="3B353C8BFF917554D7B4C1EAC97AFEE0" collectingDate="2003-06-04" collectingDateMax="2022-04-29" collectingDateMin="2003-06-04" country="Canada" county="Jefferson Co." latitude="47.6782" location="Washington" longLatPrecision="6" longitude="-124.1194" municipality="All" pageId="7" pageNumber="148" specimenCount="2" specimenCount-immature="2">
|
||||
<emphasis id="B929EAC4FF917554D7B4C1EAC84BFF59" bold="true" box="[199,522,222,248]" pageId="7" pageNumber="148">
|
||||
<typeStatus id="54E68874FF917554D7B4C1EAC844FF59" box="[199,517,222,248]" isOtherTypeStatus="true" pageId="7" pageNumber="148">Other material examined</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
From the
|
||||
<typeStatus id="54E68874FF917554D5E2C1EBC880FF58" box="[657,705,223,249]" pageId="7" pageNumber="148">type</typeStatus>
|
||||
locality, 2 mature and
|
||||
<emphasis id="B929EAC4FF917554D7B4C1EAC84BFF59" bold="true" box="[199,522,222,248]" pageId="7" pageNumber="148">Other material examined:</emphasis>
|
||||
From the type locality, 2 mature and
|
||||
<specimenCount id="9D5BFD5FFF917554D493C1EBCE28FF58" box="[992,1129,223,249]" count="2" pageId="7" pageNumber="148" type="immature">2 immature</specimenCount>
|
||||
, collected
|
||||
<date id="FFE31016FF917554D39EC1EBCFD8FF58" box="[1261,1433,223,249]" pageId="7" pageNumber="148" value="2022-04-29">
|
||||
|
|
@ -185,11 +164,7 @@ at
|
|||
<date id="FFE31016FF917554D381C037CFC1FEBC" box="[1266,1408,259,285]" pageId="7" pageNumber="148" value="2003-06-04">
|
||||
<collectingDate id="EFA7E9FEFF917554D381C037CFC1FEBC" box="[1266,1408,259,285]" pageId="7" pageNumber="148" value="2003-06-04">4 June 2003</collectingDate>
|
||||
</date>
|
||||
; 1 partially mature worm.
|
||||
<collectingMunicipality id="6B86ACACFF917554D6EDC013CB83FEE0" box="[414,450,295,321]" pageId="7" pageNumber="148">All</collectingMunicipality>
|
||||
whole-mounts in
|
||||
<collectingCountry id="F34A7646FF917554D5FEC013C8A0FEE0" box="[653,737,295,321]" name="Canada" pageId="7" pageNumber="148">Canada</collectingCountry>
|
||||
balsam
|
||||
; 1 partially mature worm. All whole-mounts in Canada balsam
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
@ -313,18 +288,18 @@ with two spermathecal segments, two Palearctic species,
|
|||
<taxonomicName id="4C5D4D55FF9E755BD2FBC6B8CB8DF86B" authority="(Fauvel, 1903)" baseAuthorityName="Fauvel" baseAuthorityYear="1903" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="8" pageNumber="149" phylum="Annelida" rank="species" species="intermedius">
|
||||
<emphasis id="B929EAC4FF9E755BD2FBC6B8CB5AF868" italics="true" pageId="8" pageNumber="149">T. intermedius</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD65EC69BCB85F86B" author="Fauvel, P." box="[301,452,1967,1994]" pageId="8" pageNumber="149" refId="ref9560" refString="Fauvel, P. (1903) Un nouvel oligochete de puits (Trichodriloides intermedius n. g. n. sp.). Compte Rendu de la Association Francaise pour l'Avancement des Sciences, 32, 221." type="journal volume" year="1903">Fauvel, 1903</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD65EC69BCB85F86B" author="Fauvel" box="[301,452,1967,1994]" firstAuthor="Fauvel" pageId="8" pageNumber="149" refId="ref9560" refString="Fauvel, P. (1903) Un nouvel oligochete de puits (Trichodriloides intermedius n. g. n. sp.). Compte Rendu de la Association Francaise pour l'Avancement des Sciences, 32, 221." type="journal volume" year="1903">Fauvel, 1903</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5D4D55FF9E755BD574C684C958F868" authority="Hrabe, 1963" authorityName="Hrabe" authorityYear="1963" box="[519,793,1967,1993]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="8" pageNumber="149" phylum="Annelida" rank="species" species="tacensis">
|
||||
<emphasis id="B929EAC4FF9E755BD574C684C83EF868" box="[519,639,1968,1993]" italics="true" pageId="8" pageNumber="149">T. tacensis</emphasis>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD5FAC69BC958F868" author="Hrabe, S." box="[649,793,1967,1993]" pageId="8" pageNumber="149" pagination="67 - 75" refId="ref10038" refString="Hrabe, S. (1963) Oligochaeta limicola from Slovenija. Zoological Institute of J. E. Purkyne University in Brno, 11, 67 - 75." type="journal article" year="1963">Hrabě, 1963</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD5FAC69BC958F868" author="Hrabe" box="[649,793,1967,1993]" firstAuthor="Hrabe" pageId="8" pageNumber="149" pagination="67 - 75" refId="ref10038" refString="Hrabe, S. (1963) Oligochaeta limicola from Slovenija. Zoological Institute of J. E. Purkyne University in Brno, 11, 67 - 75." type="journal article" year="1963">Hrabě, 1963</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, also have tubular atria which are shorter than the body diameter.Another species,
|
||||
<taxonomicName id="4C5D4D55FF9E755BD6B3C6E0C8F4F84F" authority="Hrabe, 1937" authorityName="Hrabe" authorityYear="1937" box="[448,693,2003,2030]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="8" pageNumber="149" phylum="Annelida" rank="species" species="leruthi">
|
||||
<emphasis id="B929EAC4FF9E755BD6B3C6E0C864F84C" box="[448,549,2003,2029]" italics="true" pageId="8" pageNumber="149">T. leruthi</emphasis>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD55AC6E7C8F4F84F" author="Hrabe, S." box="[553,693,2003,2030]" pageId="8" pageNumber="149" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabĕ, 1937</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD55AC6E7C8F4F84F" author="Hrabe" box="[553,693,2003,2030]" firstAuthor="Hrabe" pageId="8" pageNumber="149" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabĕ, 1937</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, has a tube-like, elongate atrium (
|
||||
<figureCitation id="13662A53FF9E755BD357C6E7CE25F84F" box="[1060,1124,2003,2030]" captionStart="FIGURE 3" captionStartId="6.[151,250,1868,1892]" captionTargetBox="[226,1402,282,1811]" captionTargetId="figure-26@6.[151,1436,181,1844]" captionTargetPageId="6" captionText="FIGURE 3. Trichodrilus humptulips sp. nov. A–E from the type locality, F from Clearwater River. A, D from holotype, C, E from paratypes. A. Anterior segments, external view. B. Chaetae in mid-body, dorsal pair (above) and ventral pair (below). C. Atrium and male ducts. D. Reproductive segments, showing gonads, male and female ducts, and spermathecae. E. Reproductive segments, spermathecal ampullae extending anteriad into adjacent segment. F. Reproductive segments of a partially mature worm." figureDoi="http://doi.org/10.5281/zenodo.14390144" httpUri="https://zenodo.org/record/14390144/files/figure.png" pageId="8" pageNumber="149">
|
||||
|
|
@ -332,14 +307,14 @@ Fig.
|
|||
<quantity id="4CA59B33FF9E755BD325C6E0CE25F84F" box="[1110,1124,2004,2030]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="149" unit="in" value="3.0">3</quantity>
|
||||
</figureCitation>
|
||||
in
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD3F7C6E7CF4BF84F" author="Hrabe, S." box="[1156,1290,2003,2030]" pageId="8" pageNumber="149" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9E755BD3F7C6E7CF4BF84F" author="Hrabe" box="[1156,1290,2003,2030]" firstAuthor="Hrabe" pageId="8" pageNumber="149" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
||||
), terminating in a prominent penial bulb, with extruded lining cells forming elongate penes (
|
||||
<figureCitation id="13662A53FF9F755AD37EC1A3CE11FF10" box="[1037,1104,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1834,1858]" captionTargetBox="[174,1410,188,1796]" captionTargetId="figure-17@11.[151,1436,181,1810]" captionTargetPageId="11" captionText="FIGURE 5. Undetermined Trichodrilus species. A. Specimen from Jemez River, New Mexico, reproductive organs in X– XII. B. Specimen from Tar River, North Carolina, reproductive organs in X–XII. C. Specimen from Chinkapin Creek, North Carolina, atrium. D–E Specimens from North Carolina, lateral blood vessels in posterior segments; (D) dorsal view (Tar River), (E) lateral view (Chinkapin Creek)." figureDoi="http://doi.org/10.5281/zenodo.14390150" httpUri="https://zenodo.org/record/14390150/files/figure.png" pageId="9" pageNumber="150">
|
||||
Fig.
|
||||
<quantity id="4CA59B33FF9F755AD331C1A2CE11FF10" box="[1090,1104,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="150" unit="in" value="5.0">5</quantity>
|
||||
</figureCitation>
|
||||
in
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD306C1A3CFD1FF10" author="Rodriguez, P. & Giani, N." box="[1141,1424,151,177]" pageId="9" pageNumber="150" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD306C1A3CFD1FF10" author="Rodriguez & Giani" box="[1141,1424,151,177]" firstAuthor="Rodriguez" pageId="9" pageNumber="150" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
). These 3 European species are known primarily from groundwater. Two main reproductive characters distinguish
|
||||
<taxonomicName id="4C5D4D55FF9F755AD7E4C1EBCB71FF58" authorityName="Fend & Rodriguez" authorityYear="2024" box="[151,304,223,249]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="9" pageNumber="150" phylum="Annelida" rank="species" species="humptulips" status="sp. nov.">
|
||||
<emphasis id="B929EAC4FF9F755AD7E4C1EBCB71FF58" box="[151,304,223,249]" italics="true" pageId="9" pageNumber="150">T. humptulips</emphasis>
|
||||
|
|
@ -357,34 +332,31 @@ from
|
|||
</taxonomicName>
|
||||
do not penetrate the posterior septum, and join the atrial ampulla at the apical end (see Fig.
|
||||
<quantity id="4CA59B33FF9F755AD431C037C92CFEBC" box="[834,877,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="150" unit="in" value="6.0">6 in</quantity>
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD400C037C9BBFEBC" author="Hrabe, S." box="[883,1018,259,285]" pageId="9" pageNumber="150" pagination="67 - 75" refId="ref10038" refString="Hrabe, S. (1963) Oligochaeta limicola from Slovenija. Zoological Institute of J. E. Purkyne University in Brno, 11, 67 - 75." type="journal article" year="1963">Hrabĕ 1963</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD400C037C9BBFEBC" author="Hrabe" box="[883,1018,259,285]" firstAuthor="Hrabe" pageId="9" pageNumber="150" pagination="67 - 75" refId="ref10038" refString="Hrabe, S. (1963) Oligochaeta limicola from Slovenija. Zoological Institute of J. E. Purkyne University in Brno, 11, 67 - 75." type="journal article" year="1963">Hrabĕ 1963</bibRefCitation>
|
||||
);
|
||||
<taxonomicName id="4C5D4D55FF9F755AD37EC037CEC3FEBD" authorityName="Hrabe" authorityYear="1963" box="[1037,1154,259,284]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="9" pageNumber="150" phylum="Annelida" rank="species" species="tacensis">
|
||||
<emphasis id="B929EAC4FF9F755AD37EC037CEC3FEBD" box="[1037,1154,259,284]" italics="true" pageId="9" pageNumber="150">T. tacensis</emphasis>
|
||||
</taxonomicName>
|
||||
also has well-developed, cylindrical penes on porophores.
|
||||
</paragraph>
|
||||
<caption id="DF22665EFF9F755AD7EBC05FCFDDFE22" box="[152,1436,363,387]" pageId="9" pageNumber="150" startId="9.[152,244,363,387]" targetType="table">
|
||||
<paragraph id="8BE236D6FF9F755AD7EBC05FCFDDFE22" blockId="9.[151,1437,363,459]" box="[152,1436,363,387]" pageId="9" pageNumber="150">
|
||||
<caption id="DF22665EFF9F755AD7EBC05FC950FE6A" pageId="9" pageNumber="150" startId="9.[152,244,363,387]" targetBox="[159,1422,471,1941]" targetIsTable="true" targetPageId="9" targetType="table">
|
||||
<paragraph id="8BE236D6FF9F755AD7EBC05FC950FE6A" blockId="9.[151,1437,363,459]" pageId="9" pageNumber="150">
|
||||
<emphasis id="B929EAC4FF9F755AD7EBC05FCB53FE22" bold="true" box="[152,274,363,387]" pageId="9" pageNumber="150">TABLE 1.</emphasis>
|
||||
Comparison of the new species with similar
|
||||
<taxonomicName id="4C5D4D55FF9F755AD47FC05FC9AFFE22" box="[780,1006,363,387]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="9" pageNumber="150" phylum="Annelida" rank="species" species="undetermined">
|
||||
<emphasis id="B929EAC4FF9F755AD47FC05FC9D1FE22" box="[780,912,363,387]" italics="true" pageId="9" pageNumber="150">Trichodrilus</emphasis>
|
||||
species.
|
||||
</taxonomicName>
|
||||
Measurements of previously described
|
||||
</paragraph>
|
||||
</caption>
|
||||
Measurements of previously described
|
||||
<tableNote id="76BB3758FF9F755AD7E4C0BBC950FE6A" pageId="9" pageNumber="150" targetBox="[159,1422,471,1941]" targetPageId="9">
|
||||
<paragraph id="8BE236D6FF9F755AD7E4C0BBCFDCFE06" blockId="9.[151,1437,363,459]" box="[151,1437,399,423]" pageId="9" pageNumber="150">
|
||||
species are from the listed authorities; for
|
||||
<taxonomicName id="4C5D4D55FF9F755AD529C0A4C89DFE06" authorityName="Vejdovsky" authorityYear="1876" box="[602,732,400,423]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="9" pageNumber="150" phylum="Annelida" rank="species" species="pragensis">
|
||||
<emphasis id="B929EAC4FF9F755AD529C0A4C89DFE06" box="[602,732,400,423]" italics="true" pageId="9" pageNumber="150">T. pragensis</emphasis>
|
||||
</taxonomicName>
|
||||
these are supplemented by our measurements of specimens in the
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF9F755AD7E4C087C950FE6A" blockId="9.[151,1437,363,459]" box="[151,785,435,459]" pageId="9" pageNumber="150">Hrabĕ collection, National Museum of the Czech Republic.</paragraph>
|
||||
these are supplemented by our measurements of specimens in the Hrabĕ collection, National Museum of the Czech Republic.
|
||||
</tableNote>
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BE236D6FF9F755AD7A4C0E3CF5BF834" pageId="9" pageNumber="150">
|
||||
<table id="F95DC476FF9F8AACD7ECC0E3CFCFF834" box="[159,1422,471,1941]" gridcols="5" gridrows="39" pageId="9" pageNumber="150">
|
||||
<tr id="356D3494FF9F8AACD7ECC0E3CFCFFE51" box="[159,1422,471,496]" gridrow="0" pageId="9" pageNumber="150">
|
||||
|
|
@ -420,7 +392,7 @@ these are supplemented by our measurements of specimens in the
|
|||
<tr id="356D3494FF9F8AACD7ECC0C8CFCFFDB5" box="[159,1422,508,532]" gridrow="1" pageId="9" pageNumber="150" rowspan-0="1" rowspan-2="1">
|
||||
<td id="76BC5DE8FF9F8AACD6E8C0C8C8CAFDB5" box="[411,651,508,532]" gridcol="1" gridrow="1" pageId="9" pageNumber="150">Vejdovský, 1876;</td>
|
||||
<td id="76BC5DE8FF9F8AACD4D2C0C8CECEFDB5" box="[929,1167,508,532]" gridcol="3" gridrow="1" pageId="9" pageNumber="150">
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD4D2C0C8CE69FDB5" author="Fauvel, P." box="[929,1064,508,532]" pageId="9" pageNumber="150" refId="ref9560" refString="Fauvel, P. (1903) Un nouvel oligochete de puits (Trichodriloides intermedius n. g. n. sp.). Compte Rendu de la Association Francaise pour l'Avancement des Sciences, 32, 221." type="journal volume" year="1903">Fauvel, 1903</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9F755AD4D2C0C8CE69FDB5" author="Fauvel" box="[929,1064,508,532]" firstAuthor="Fauvel" pageId="9" pageNumber="150" refId="ref9560" refString="Fauvel, P. (1903) Un nouvel oligochete de puits (Trichodriloides intermedius n. g. n. sp.). Compte Rendu de la Association Francaise pour l'Avancement des Sciences, 32, 221." type="journal volume" year="1903">Fauvel, 1903</bibRefCitation>
|
||||
; Hrabĕ
|
||||
</td>
|
||||
<td id="76BC5DE8FF9F8AACD3D1C0C8CFCFFDB5" box="[1186,1422,508,532]" gridcol="4" gridrow="1" pageId="9" pageNumber="150">
|
||||
|
|
@ -432,10 +404,10 @@ these are supplemented by our measurements of specimens in the
|
|||
<emphasis id="B929EAC4FF9F755AD7ECC32BCB4FFD96" bold="true" box="[159,270,543,567]" pageId="9" pageNumber="150">Character</emphasis>
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</th>
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<td id="76BC5DE8FF9F8AACD6E8C32BC8CAFD99" box="[411,651,543,568]" gridcol="1" gridrow="2" pageId="9" pageNumber="150">
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<bibRefCitation id="EFCC4B27FF9F755AD6E8C314C853FD99" author="Hrabe, S." box="[411,530,544,568]" pageId="9" pageNumber="150" pagination="205 - 208" refId="ref10068" refString="Hrabe, S. (1971) On Trichodrilus pragensis Vejd. (Oligochaeta, Lumbriculidae). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 35, 205 - 208." type="journal article" year="1971">Hrabĕ 1971</bibRefCitation>
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<bibRefCitation id="EFCC4B27FF9F755AD6E8C314C853FD99" author="Hrabe" box="[411,530,544,568]" firstAuthor="Hrabe" pageId="9" pageNumber="150" pagination="205 - 208" refId="ref10068" refString="Hrabe, S. (1971) On Trichodrilus pragensis Vejd. (Oligochaeta, Lumbriculidae). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 35, 205 - 208." type="journal article" year="1971">Hrabĕ 1971</bibRefCitation>
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<td id="76BC5DE8FF9F8AACD4D2C32BCECEFD99" box="[929,1167,543,568]" gridcol="3" gridrow="2" pageId="9" pageNumber="150">
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<bibRefCitation id="EFCC4B27FF9F755AD4D2C314C995FD99" author="Hrabe, S." box="[929,980,544,568]" pageId="9" pageNumber="150" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">1937</bibRefCitation>
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<bibRefCitation id="EFCC4B27FF9F755AD4D2C314C995FD99" author="Hrabe" box="[929,980,544,568]" firstAuthor="Hrabe" pageId="9" pageNumber="150" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">1937</bibRefCitation>
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<tr id="356D3494FF9F8AACD7ECC370CFCFFDFE" box="[159,1422,580,607]" gridrow="3" pageId="9" pageNumber="150">
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|
|
@ -645,22 +617,22 @@ these are supplemented by our measurements of specimens in the
|
|||
</paragraph>
|
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<paragraph id="8BE236D6FF9C7559D7B4C1A3C851FE28" blockId="10.[151,1437,151,393]" pageId="10" pageNumber="151">
|
||||
Using the key by
|
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<bibRefCitation id="EFCC4B27FF9C7559D6FBC1A3C8EDFF10" author="Rodriguez, P. & Giani, N." box="[392,684,151,177]" pageId="10" pageNumber="151" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani (1994)</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF9C7559D6FBC1A3C8EDFF10" author="Rodriguez & Giani" box="[392,684,151,177]" firstAuthor="Rodriguez" pageId="10" pageNumber="151" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani (1994)</bibRefCitation>
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(see
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<tableCitation id="C6DF036DFF9C7559D597C1A3C974FF10" box="[740,821,151,177]" captionStart="TABLE 1" captionStartId="9.[152,244,363,387]" captionText="TABLE 1. Comparison of the new species with similar Trichodrilus species. Measurements of previously described" pageId="10" pageNumber="151">Table 1</tableCitation>
|
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<tableCitation id="C6DF036DFF9C7559D597C1A3C974FF10" box="[740,821,151,177]" captionStart="TABLE 1" captionStartId="9.[152,244,363,387]" captionTargetBox="[159,1422,471,1941]" captionTargetPageId="9" captionText="TABLE 1. Comparison of the new species with similar Trichodrilus species. Measurements of previously described species are from the listed authorities; for T. pragensis these are supplemented by our measurements of specimens in the Hrabĕ collection, National Museum of the Czech Republic." pageId="10" pageNumber="151">Table 1</tableCitation>
|
||||
), the closest match to the new species is
|
||||
<taxonomicName id="4C5D4D55FF9C7559D384C1A3CFD4FF10" baseAuthorityName="Fauvel" baseAuthorityYear="1903" box="[1271,1429,151,177]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="10" pageNumber="151" phylum="Annelida" rank="species" species="intermedius">
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<emphasis id="B929EAC4FF9C7559D384C1A3CFD4FF10" box="[1271,1429,151,177]" italics="true" pageId="10" pageNumber="151">T. intermedius</emphasis>
|
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</taxonomicName>
|
||||
, which also has posterior vasa deferentia penetrating the posterior septum (10/11), forming a loop in XI (Fig.
|
||||
<quantity id="4CA59B33FF9C7559D203C18FCFDDFF74" box="[1392,1436,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="151" unit="in" value="9.0">9 in</quantity>
|
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<bibRefCitation id="EFCC4B27FF9C7559D7E4C1EBCB60FF58" author="Hrabe, S." box="[151,289,223,249]" pageId="10" pageNumber="151" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
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<bibRefCitation id="EFCC4B27FF9C7559D7E4C1EBCB60FF58" author="Hrabe" box="[151,289,223,249]" firstAuthor="Hrabe" pageId="10" pageNumber="151" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
||||
). From the original description,
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||||
<taxonomicName id="4C5D4D55FF9C7559D5E0C1EBC974FF58" baseAuthorityName="Fauvel" baseAuthorityYear="1903" box="[659,821,223,249]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="10" pageNumber="151" phylum="Annelida" rank="species" species="intermedius">
|
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<emphasis id="B929EAC4FF9C7559D5E0C1EBC974FF58" box="[659,821,223,249]" italics="true" pageId="10" pageNumber="151">T. intermedius</emphasis>
|
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</taxonomicName>
|
||||
differs in having 2–6 pairs of lateral blood vessels in posterior segments.
|
||||
<bibRefCitation id="EFCC4B27FF9C7559D604C037C84CFEBC" author="Hrabe, S." box="[375,525,259,285]" pageId="10" pageNumber="151" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě (1937)</bibRefCitation>
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<bibRefCitation id="EFCC4B27FF9C7559D604C037C84CFEBC" author="Hrabe" box="[375,525,259,285]" firstAuthor="Hrabe" pageId="10" pageNumber="151" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě (1937)</bibRefCitation>
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||||
provided a detailed redescription, noting the very thick, muscular vasa deferentia (37–47 μm) and short-cylindrical penes. The muscle layer of the atrial ampulla in Hrabě's material appears somewhat thicker, at 9.5 μm, and both vasa deferentia join the atrial ampulla at the most ental (apical) end, in contrast to the basal junction of the new species.
|
||||
</paragraph>
|
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</subSubSection>
|
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|
|
|
|||
|
|
@ -1,52 +1,52 @@
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||||
</subSubSection>
|
||||
<subSubSection id="C347655DFF977550D7E4C740CB9BFEBC" lastPageId="3" lastPageNumber="144" pageId="1" pageNumber="142" type="materials_examined">
|
||||
<paragraph id="8BE236D6FF977552D7E4C740C80AF913" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="142">
|
||||
<paragraph id="8BE236D6FF977552D7E4C740C9F9F95B" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="142">
|
||||
<emphasis id="B929EAC4FF977552D7E4C740CB7CF92F" bold="true" box="[151,317,1652,1678]" pageId="1" pageNumber="142">Type locality:</emphasis>
|
||||
<materialsCitation id="3B353C8BFF977552D63AC740C807F913" county="Lane Co." latitude="43.972" location="Fall Creek" longLatPrecision="68" longitude="-122.541" municipality="Fall Creek" pageId="1" pageNumber="142" specimenCount="1" stateProvince="Oregon">
|
||||
<materialsCitation id="3B353C8BFF977552D63AC740C9F5F95B" collectingDate="2020-05-10" collectionCode="USNM" county="Lane Co." latitude="43.972" location="Fall Creek" longLatPrecision="68" longitude="-122.541" municipality="Fall Creek" pageId="1" pageNumber="142" specimenCode="USNM 1740726" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
|
||||
<collectingRegion id="4999F834FF977552D63AC740CBE1F92F" box="[329,416,1652,1678]" country="United States of America" name="Oregon" pageId="1" pageNumber="142">Oregon</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62834E5AFF977552D6DCC741C85DF92F" box="[431,540,1652,1679]" pageId="1" pageNumber="142">Lane Co.</collectingCounty>
|
||||
|
|
@ -85,12 +85,7 @@ reservoir, approximately
|
|||
<geoCoordinate id="EE695011FF977552D3DEC740CF50F92F" box="[1197,1297,1652,1678]" degrees="43.972" direction="north" orientation="latitude" pageId="1" pageNumber="142" precision="55" value="43.972">N43.972</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE695011FF977552D253C740CFD9F92E" box="[1312,1432,1652,1679]" degrees="122.541" direction="west" orientation="longitude" pageId="1" pageNumber="142" precision="55" value="-122.541">W122.541</geoCoordinate>
|
||||
, gravel-cobble streambed near margins
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF977552D7B4C788C9F9F95B" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="142">
|
||||
<materialsCitation id="3B353C8BFF977552D7B4C788C9F9F95B" collectingDate="2020-05-10" collectionCode="USNM" country="Canada" location="Canada" pageId="1" pageNumber="142" specimenCode="USNM 1740726" specimenCount="1" typeStatus="holotype">
|
||||
, gravel-cobble streambed near margins.
|
||||
<emphasis id="B929EAC4FF977552D7B4C788CB7CF977" bold="true" box="[199,317,1724,1750]" pageId="1" pageNumber="142">
|
||||
<typeStatus id="54E68874FF977552D7B4C788CB79F977" box="[199,312,1724,1750]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
|
|
@ -100,13 +95,12 @@ reservoir, approximately
|
|||
<date id="FFE31016FF977552D569C788C8FBF977" box="[538,698,1724,1751]" pageId="1" pageNumber="142" value="2020-05-10">
|
||||
<collectingDate id="EFA7E9FEFF977552D569C788C8FBF977" box="[538,698,1724,1751]" pageId="1" pageNumber="142" value="2020-05-10">10 May 2020</collectingDate>
|
||||
</date>
|
||||
, a whole mounted worm, thick epidermis of clitellum partially scraped off, stained with carmine and slide-mounted in
|
||||
<collectingCountry id="F34A7646FF977552D475C7D4C91BF95B" box="[774,858,1760,1786]" name="Canada" pageId="1" pageNumber="142">Canada</collectingCountry>
|
||||
balsam.
|
||||
, a whole mounted worm, thick epidermis of clitellum partially scraped off, stained with carmine and slide-mounted in Canada balsam
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF977552D7B4C630C985F82B" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="142">
|
||||
<materialsCitation id="3B353C8BFF977552D7B4C630C981F82B" collectingDate="2020-03-19" collectingDateMax="2020-05-10" collectingDateMin="2020-03-19" collectionCode="USNM, CASIZ, MNCN" country="Canada" location="All" pageId="1" pageNumber="142" specimenCode="USNM 1740727-1740730, CASIZ 242895-242899, MNCN 16.03" specimenCount="1" typeStatus="paratype">
|
||||
<materialsCitation id="3B353C8BFF977552D7B4C630C985F82B" collectingDate="2020-03-19" collectingDateMax="2020-05-10" collectingDateMin="2020-03-19" collectionCode="USNM, CASIZ, MNCN" pageId="1" pageNumber="142" specimenCode="USNM 1740727-1740730, CASIZ 242895-242899, MNCN 16.03" specimenCount="1" typeStatus="paratype">
|
||||
<emphasis id="B929EAC4FF977552D7B4C630CB09F8BF" bold="true" box="[199,328,1796,1822]" pageId="1" pageNumber="142">
|
||||
<typeStatus id="54E68874FF977552D7B4C630CB02F8BF" box="[199,323,1796,1822]" pageId="1" pageNumber="142" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
|
|
@ -145,13 +139,8 @@ All from the type locality.
|
|||
<date id="FFE31016FF977552D78BC644CBD1F82B" box="[248,400,1904,1931]" pageId="1" pageNumber="142" value="2020-05-10">
|
||||
<collectingDate id="EFA7E9FEFF977552D78BC644CBD1F82B" box="[248,400,1904,1931]" pageId="1" pageNumber="142" value="2020-05-10">10 May 2020</collectingDate>
|
||||
</date>
|
||||
, 1 dissected.
|
||||
<location id="8E82600DFF977552D550C644C806F82B" LSID="urn:lsid:plazi:treatment:03F487C0FF977556D7E4C733CB34F938:8E82600DFF977552D550C644C806F82B" box="[547,583,1904,1930]" country="Canada" name="All" pageId="1" pageNumber="142">All</location>
|
||||
slide-mounted in
|
||||
<collectingCountry id="F34A7646FF977552D461C644C927F82B" box="[786,870,1904,1930]" name="Canada" pageId="1" pageNumber="142">Canada</collectingCountry>
|
||||
balsam
|
||||
, 1 dissected. All slide-mounted in Canada balsam.
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF977552D7B4C6A0CF04F856" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="142">
|
||||
<emphasis id="B929EAC4FF977552D7B4C6A0CBBDF80F" bold="true" box="[199,508,1940,1966]" pageId="1" pageNumber="142">
|
||||
|
|
@ -196,7 +185,6 @@ Detail of male ducts, showing atrium, extended penis, and vasa deferentia.
|
|||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BE236D6FF957550D7B4C1A3CB9BFEBC" blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="144">
|
||||
<materialsCitation id="3B353C8BFF957550D7B4C1A3CB97FEBC" collectingDate="2000-01-30" collectingDateMax="2021-04-20" collectingDateMin="2000-01-30" county="Lane Co." latitude="45.2408" location="Dolly Varden Campground" longLatPrecision="6" longitude="-123.3862" municipality="Fall Creek" pageId="3" pageNumber="144" specimenCount="1" stateProvince="Oregon">
|
||||
<collectingRegion id="4999F834FF957550D7B4C1A3CB5FFF10" box="[199,286,151,177]" country="United States of America" name="Oregon" pageId="3" pageNumber="144">Oregon</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62834E5AFF957550D65EC1A3CBDBFF10" box="[301,410,151,177]" pageId="3" pageNumber="144">Lane Co.</collectingCounty>
|
||||
|
|
@ -228,9 +216,7 @@ below
|
|||
<date id="FFE31016FF957550D4AAC1EBCED6FF58" box="[985,1175,223,249]" pageId="3" pageNumber="144" value="2000-01-30">
|
||||
<collectingDate id="EFA7E9FEFF957550D4AAC1EBCED6FF58" box="[985,1175,223,249]" pageId="3" pageNumber="144" value="2000-01-30">30 January 2000</collectingDate>
|
||||
</date>
|
||||
, 1 dissected (mated), 5 whole mounts (all unmated)
|
||||
</materialsCitation>
|
||||
.
|
||||
, 1 dissected (mated), 5 whole mounts (all unmated).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C347655DFF957550D7B4C013CBBFFEC4" pageId="3" pageNumber="144" type="etymology">
|
||||
|
|
@ -279,7 +265,7 @@ Male pores in X, midway between chaetae and posterior septum, on ventral chaetal
|
|||
<figureCitation id="13662A53FF957550D423C59FC9E1FB64" box="[848,928,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1558,1582]" captionTargetBox="[167,1426,183,1524]" captionTargetId="figure-26@4.[151,1436,181,1535]" captionTargetPageId="4" captionText="FIGURE 2. Trichodrilus baylesi sp. nov..from Fall Creek, Oregon. All from paratypes, except B and C from non-types. A. Prostomium, anterior body region and pharynx, sagittal section. B. Anterior end of whole worm (fixed and stained), showing clitellum and penis (lower is detail of upper). C. Extended penis, from whole mount. D, E. Atrium and partially extended penis, sagittal section showing associated musculature (D is from the same specimen as E). F. Atrium and extended penis, sagittal section showing penial muscle bands surrounding atrial duct. G. Atrial ampulla, with multicellular prostates, and vas deferens junction, sagittal section. H. Atrium and vas deferens, sagittal section. I. Spermathecal duct, sagittal section. J–K. Ental part of spermathecal duct, showing sphincter at junction with ampulla (sagittal section above, whole mount below). L–M. Spermathecal ampulla, sagittal section (lower is detail of upper)." figureDoi="http://doi.org/10.5281/zenodo.14390142" httpUri="https://zenodo.org/record/14390142/files/figure.png" pageId="3" pageNumber="144">Fig. 2I</figureCitation>
|
||||
). Clitellum distinctly glandular, from mid-IX or X (at 9/10) through XII or XIII, wholly surrounding the body (
|
||||
<figureCitation id="13662A53FF957550D4F5C5FBC9A1FB48" box="[902,992,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[151,250,1558,1582]" captionTargetBox="[167,1426,183,1524]" captionTargetId="figure-26@4.[151,1436,181,1535]" captionTargetPageId="4" captionText="FIGURE 2. Trichodrilus baylesi sp. nov..from Fall Creek, Oregon. All from paratypes, except B and C from non-types. A. Prostomium, anterior body region and pharynx, sagittal section. B. Anterior end of whole worm (fixed and stained), showing clitellum and penis (lower is detail of upper). C. Extended penis, from whole mount. D, E. Atrium and partially extended penis, sagittal section showing associated musculature (D is from the same specimen as E). F. Atrium and extended penis, sagittal section showing penial muscle bands surrounding atrial duct. G. Atrial ampulla, with multicellular prostates, and vas deferens junction, sagittal section. H. Atrium and vas deferens, sagittal section. I. Spermathecal duct, sagittal section. J–K. Ental part of spermathecal duct, showing sphincter at junction with ampulla (sagittal section above, whole mount below). L–M. Spermathecal ampulla, sagittal section (lower is detail of upper)." figureDoi="http://doi.org/10.5281/zenodo.14390142" httpUri="https://zenodo.org/record/14390142/files/figure.png" pageId="3" pageNumber="144">Fig. 2B</figureCitation>
|
||||
); 20–46 µm thick. No obvious internal glands around genital pores. Testes small (not extending as far as chaetae) in IX and X; ovaries extend beyond mid- XI, usually to 11/12. Sperm sacs extend anteriad to VII or VIII and backward through XIII to XV; egg sacs extend 1–3 segments behind posterior sperm sacs.
|
||||
); 20–46 µm thick. No obvious internal glands around genital pores. Testes small (not extending as far as chaetae) in IX and X; ovaries extend beyond mid-XI, usually to 11/12. Sperm sacs extend anteriad to VII or VIII and backward through XIII to XV; egg sacs extend 1–3 segments behind posterior sperm sacs.
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF957550D7B4C46BCB55F938" blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="144">
|
||||
Spermathecal duct distinct, tubular, in two parts (
|
||||
|
|
@ -393,7 +379,7 @@ species
|
|||
<taxonomicNameLabel id="A21A57BFFF937556D574C3CFC825FCB4" box="[519,612,763,789]" pageId="5" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is a relatively small, unpigmented worm. Using the key by
|
||||
<bibRefCitation id="EFCC4B27FF937556D25BC3CFCB0AFC98" author="Rodriguez, P. & Giani, N." pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani (1994)</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D25BC3CFCB0AFC98" author="Rodriguez & Giani" firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani (1994)</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="4C5D4D55FF937556D62AC22BC87FFC98" box="[345,574,799,825]" pageId="5" pageNumber="146">
|
||||
<emphasis id="B929EAC4FF937556D62AC22BCBA2FC98" box="[345,483,799,825]" italics="true" pageId="5" pageNumber="146">Trichodrilus</emphasis>
|
||||
|
|
@ -413,19 +399,19 @@ Vejdovský, 1876
|
|||
<emphasis id="B929EAC4FF937556D4F3C253CE1EFC20" box="[896,1119,871,897]" italics="true" pageId="5" pageNumber="146">Trichodrilus culveri</emphasis>
|
||||
</taxonomicName>
|
||||
differs in lacking any penial structure, and in having a very short, conical spermathecal duct (
|
||||
<bibRefCitation id="EFCC4B27FF937556D417C2BFC9A4FC04" author="Cook, D. G." box="[868,997,906,933]" pageId="5" pageNumber="146" pagination="24 - 37" refId="ref9463" refString="Cook, D. G. (1975) Cave-dwelling aquatic Oligochaeta (Annelida) from the Eastern United States. Transactions of the American Microscopical Society, 94, 24 - 37. https: // doi. org / 10.2307 / 3225529" type="journal article" year="1975">Cook 1975</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D417C2BFC9A4FC04" author="Cook" box="[868,997,906,933]" firstAuthor="Cook" pageId="5" pageNumber="146" pagination="24 - 37" refId="ref9463" refString="Cook, D. G. (1975) Cave-dwelling aquatic Oligochaeta (Annelida) from the Eastern United States. Transactions of the American Microscopical Society, 94, 24 - 37. https: // doi. org / 10.2307 / 3225529" type="journal article" year="1975">Cook 1975</bibRefCitation>
|
||||
). The small penes (opening in a conical porophore and without a penial sac) of
|
||||
<taxonomicName id="4C5D4D55FF937556D520C29BC89DFC69" authorityName="Vejdovsky" authorityYear="1876" box="[595,732,943,968]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="pragensis">
|
||||
<emphasis id="B929EAC4FF937556D520C29BC89DFC69" box="[595,732,943,968]" italics="true" pageId="5" pageNumber="146">T. pragensis</emphasis>
|
||||
</taxonomicName>
|
||||
appear quite different from the large penes of the new species (see
|
||||
<tableCitation id="C6DF036DFF937556D7BEC2E7CB63FC4C" box="[205,290,979,1005]" captionStart="TABLE 1" captionStartId="9.[152,244,363,387]" captionText="TABLE 1. Comparison of the new species with similar Trichodrilus species. Measurements of previously described" pageId="5" pageNumber="146">Table 1</tableCitation>
|
||||
<tableCitation id="C6DF036DFF937556D7BEC2E7CB63FC4C" box="[205,290,979,1005]" captionStart="TABLE 1" captionStartId="9.[152,244,363,387]" captionTargetBox="[159,1422,471,1941]" captionTargetPageId="9" captionText="TABLE 1. Comparison of the new species with similar Trichodrilus species. Measurements of previously described species are from the listed authorities; for T. pragensis these are supplemented by our measurements of specimens in the Hrabĕ collection, National Museum of the Czech Republic." pageId="5" pageNumber="146">Table 1</tableCitation>
|
||||
below). The lateral blood vessels described in posterior segments of
|
||||
<taxonomicName id="4C5D4D55FF937556D337C2E7CE8EFC4D" authorityName="Vejdovsky" authorityYear="1876" box="[1092,1231,979,1004]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="pragensis">
|
||||
<emphasis id="B929EAC4FF937556D337C2E7CE8EFC4D" box="[1092,1231,979,1004]" italics="true" pageId="5" pageNumber="146">T. pragensis</emphasis>
|
||||
</taxonomicName>
|
||||
and several other species (
|
||||
<bibRefCitation id="EFCC4B27FF937556D784C2C3CB3CFBB0" author="Hrabe, S." box="[247,381,1015,1041]" pageId="5" pageNumber="146" pagination="205 - 208" refId="ref10068" refString="Hrabe, S. (1971) On Trichodrilus pragensis Vejd. (Oligochaeta, Lumbriculidae). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 35, 205 - 208." type="journal article" year="1971">Hrabĕ 1971</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D784C2C3CB3CFBB0" author="Hrabe" box="[247,381,1015,1041]" firstAuthor="Hrabe" pageId="5" pageNumber="146" pagination="205 - 208" refId="ref10068" refString="Hrabe, S. (1971) On Trichodrilus pragensis Vejd. (Oligochaeta, Lumbriculidae). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 35, 205 - 208." type="journal article" year="1971">Hrabĕ 1971</bibRefCitation>
|
||||
) were also not seen in
|
||||
<taxonomicName id="4C5D4D55FF937556D5F3C2C3C8AAFBB0" authorityName="Fend & Rodriguez" authorityYear="2024" box="[640,747,1015,1041]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="baylesi" status="sp. nov.">
|
||||
<emphasis id="B929EAC4FF937556D5F3C2C3C8AAFBB0" box="[640,747,1015,1041]" italics="true" pageId="5" pageNumber="146">T. baylesi</emphasis>
|
||||
|
|
@ -434,35 +420,35 @@ and several other species (
|
|||
<taxonomicNameLabel id="A21A57BFFF937556D580C2C3C90CFBB0" box="[755,845,1015,1041]" pageId="5" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, although this character may be difficult to define in preserved material (
|
||||
<bibRefCitation id="EFCC4B27FF937556D606C52FC8D0FB94" author="Rodriguez, P. & Giani, N." box="[373,657,1051,1077]" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D606C52FC8D0FB94" author="Rodriguez & Giani" box="[373,657,1051,1077]" firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE236D6FF937556D7B4C50BCBD2FA60" blockId="5.[151,1437,151,1689]" pageId="5" pageNumber="146">
|
||||
Unlike the new species, penes are not highly developed in most congeners. Two exceptions,
|
||||
<taxonomicName id="4C5D4D55FF937556D3BBC50BCB2EFBDC" authority="Giani & Rodriguez, 1994" authorityName="Giani & Rodriguez" authorityYear="1994" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="longipenis">
|
||||
<emphasis id="B929EAC4FF937556D3BBC50BCF19FBF8" box="[1224,1368,1087,1113]" italics="true" pageId="5" pageNumber="146">T. longipenis</emphasis>
|
||||
<bibRefCitation id="EFCC4B27FF937556D22DC50BCB2EFBDC" author="Rodriguez, P. & Giani, N." pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Giani & Rodriguez, 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D22DC50BCB2EFBDC" author="Giani & Rodriguez" firstAuthor="Giani" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Giani & Rodriguez, 1994</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5D4D55FF937556D6D6C557C8DCFBDC" authority="Hrabe, 1937" authorityName="Hrabe" authorityYear="1937" box="[421,669,1123,1149]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="leruthi">
|
||||
<emphasis id="B929EAC4FF937556D6D6C557C84DFBDC" box="[421,524,1123,1149]" italics="true" pageId="5" pageNumber="146">T. leruthi</emphasis>
|
||||
<bibRefCitation id="EFCC4B27FF937556D561C557C8DCFBDC" author="Hrabe, S." box="[530,669,1123,1149]" pageId="5" pageNumber="146" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabĕ, 1937</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D561C557C8DCFBDC" author="Hrabe" box="[530,669,1123,1149]" firstAuthor="Hrabe" pageId="5" pageNumber="146" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabĕ, 1937</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(both of which have two spermathecal segments), have large, “
|
||||
<typeStatus id="54E68874FF937556D211C557CFDDFBDC" box="[1378,1436,1123,1149]" pageId="5" pageNumber="146">Type</typeStatus>
|
||||
2” penes, formed by extruding elongated lining cells of the atrial duct (
|
||||
<figureCitation id="13662A53FF937556D4B1C5B3CE54FB00" box="[962,1045,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[200,1356,257,1770]" captionTargetId="figure-26@2.[151,1436,179,1842]" captionTargetPageId="2" captionText="FIGURE 1. Trichodrilus baylesi sp. nov. from Fall Creek, Oregon (A–C, E–F) and from Little Deer Creek, Oregon (D). A is from holotype, B, C, E from paratypes. A. Anterior end of a whole mounted worm, showing pharyngeal region and reproductive segments. B. Chaetae in VIII, ventral pair (below) and dorsal pair (above). C, D. Reproductive segments X–XII (D, unmated individual). E, F. Detail of male ducts, showing atrium, extended penis, and vasa deferentia." figureDoi="http://doi.org/10.5281/zenodo.14390138" httpUri="https://zenodo.org/record/14390138/files/figure.png" pageId="5" pageNumber="146">Fig. 1F</figureCitation>
|
||||
in
|
||||
<bibRefCitation id="EFCC4B27FF937556D34FC5B3CF18FB00" author="Rodriguez, P. & Giani, N." box="[1084,1369,1159,1185]" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Giani & Rodriguez 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D34FC5B3CF18FB00" author="Giani & Rodriguez" box="[1084,1369,1159,1185]" firstAuthor="Giani" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Giani & Rodriguez 1994</bibRefCitation>
|
||||
, Figs. 17D–G in
|
||||
<bibRefCitation id="EFCC4B27FF937556D663C59FC86CFB64" author="Rodriguez, P. & Giani, N." box="[272,557,1195,1221]" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D663C59FC86CFB64" author="Rodriguez & Giani" box="[272,557,1195,1221]" firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
,
|
||||
<figureCitation id="13662A53FF937556D54AC59FC83DFB64" box="[569,636,1195,1221]" captionStart="FIGURE 4" captionStartId="8.[151,250,1728,1752]" captionTargetBox="[162,1427,200,1690]" captionTargetId="figure-26@8.[151,1436,181,1704]" captionTargetPageId="8" captionText="FIGURE 4. Trichodrilus humptulips sp. nov. (All from type locality; A, D, H from holotype, others from paratypes. E–G from dissections, others from whole mounts). A. Prostomium and partially everted pharynx. B. Male pore, external view. C. Ventral chaetae in XI. D. Clitellum and male pore in X. E, F. Atrium. G. Spermathecal pore in XI. H. Spermathecal duct and part of ampulla." figureDoi="http://doi.org/10.5281/zenodo.14390148" httpUri="https://zenodo.org/record/14390148/files/figure.png" pageId="5" pageNumber="146">
|
||||
Fig.
|
||||
<quantity id="4CA59B33FF937556D51DC59FC83DFB64" box="[622,636,1195,1221]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="146" unit="in" value="4.0">4</quantity>
|
||||
</figureCitation>
|
||||
in
|
||||
<bibRefCitation id="EFCC4B27FF937556D5D1C59FC96AFB64" author="Hrabe, S." box="[674,811,1195,1221]" pageId="5" pageNumber="146" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D5D1C59FC96AFB64" author="Hrabe" box="[674,811,1195,1221]" firstAuthor="Hrabe" pageId="5" pageNumber="146" pagination="1 - 23" refId="ref9961" refString="Hrabe, S. (1937) Etudes biospeologiques. V. Contribution a l'etude du genre Trichodrilus (Oligoch., Lumbriculidae) et description de deux especes nouvelles. Bulletin du Musee royal d'Histoire naturelle de Belgique, 13 (32), 1 - 23." type="journal article" year="1937">Hrabě 1937</bibRefCitation>
|
||||
). In contrast,
|
||||
<taxonomicName id="4C5D4D55FF937556D4B4C59FCE72FB64" authorityName="Fend & Rodriguez" authorityYear="2024" box="[967,1075,1195,1221]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="baylesi" status="sp. nov.">
|
||||
<emphasis id="B929EAC4FF937556D4B4C59FCE72FB64" box="[967,1075,1195,1221]" italics="true" pageId="5" pageNumber="146">T. baylesi</emphasis>
|
||||
|
|
@ -473,11 +459,11 @@ in
|
|||
has large penes within deep, eversible sacs; although clearly elongated when the sacs are everted, the penes are “
|
||||
<typeStatus id="54E68874FF937556D3F9C5FBCE85FB48" box="[1162,1220,1231,1257]" pageId="5" pageNumber="146">Type</typeStatus>
|
||||
1”, with epidermis surrounding internal musculature and duct (Fig. 17A, C in
|
||||
<bibRefCitation id="EFCC4B27FF937556D430C5C7CE24FAAC" author="Rodriguez, P. & Giani, N." box="[835,1125,1267,1293]" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D430C5C7CE24FAAC" author="Rodriguez & Giani" box="[835,1125,1267,1293]" firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
). The sacs surrounding the retracted penes appear to be simple epidermal folds, thus differing from more histologically differentiated penial sacs described in other lumbriculid genera, e.g.,
|
||||
<taxonomicName id="4C5D4D55FF937556D5C6C40FCEC7FAF4" authority="Rodriguez et al., 2014" authorityName="Rodriguez" authorityYear="2014" box="[693,1158,1339,1365]" class="Clitellata" family="Lumbriculidae" genus="Sylphella" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="puccoon">
|
||||
<emphasis id="B929EAC4FF937556D5C6C40FC9C2FAF5" box="[693,899,1339,1365]" italics="true" pageId="5" pageNumber="146">Sylphella puccoon</emphasis>
|
||||
<bibRefCitation id="EFCC4B27FF937556D4FAC40FCEC7FAF4" author="Rodriguez, P. & Fend, S. V. & Lenat, D. R." box="[905,1158,1339,1365]" pageId="5" pageNumber="146" pagination="1 - 32" refId="ref10519" refString="Rodriguez, P., Fend, S. V. & Lenat, D. R. (2014) Sylphella puccoon gen. n., sp. n. and two additional new species of aquatic oligochaetes (Lumbriculidae, Clitellata) from poorly-known lotic habitats in North Carolina (USA). ZooKeys, 451, 1 - 32. https: // doi. org / 10.3897 / zookeys. 451.7304" type="journal article" year="2014">
|
||||
<bibRefCitation id="EFCC4B27FF937556D4FAC40FCEC7FAF4" author="Rodriguez" box="[905,1158,1339,1365]" etAl="et al." firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="1 - 32" refId="ref10519" refString="Rodriguez, P., Fend, S. V. & Lenat, D. R. (2014) Sylphella puccoon gen. n., sp. n. and two additional new species of aquatic oligochaetes (Lumbriculidae, Clitellata) from poorly-known lotic habitats in North Carolina (USA). ZooKeys, 451, 1 - 32. https: // doi. org / 10.3897 / zookeys. 451.7304" type="journal article" year="2014">
|
||||
Rodriguez
|
||||
<emphasis id="B929EAC4FF937556D376C40FCE7CFAF4" box="[1029,1085,1339,1365]" italics="true" pageId="5" pageNumber="146">et al.</emphasis>
|
||||
, 2014
|
||||
|
|
@ -487,14 +473,14 @@ Rodriguez
|
|||
<taxonomicName id="4C5D4D55FF937556D3DCC40FCB76FAD8" authority="(Smith, 1900)" baseAuthorityName="Smith" baseAuthorityYear="1900" class="Clitellata" family="Lumbriculidae" genus="Eclipidrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="palustris">
|
||||
<emphasis id="B929EAC4FF937556D3DCC40FCFDDFAF4" box="[1199,1436,1339,1365]" italics="true" pageId="5" pageNumber="146">Eclipidrilus palustris</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFCC4B27FF937556D7ECC46BCB6EFAD8" author="Smith, F." box="[159,303,1375,1401]" pageId="5" pageNumber="146" pagination="459 - 479" refId="ref10665" refString="Smith, F. (1900) Notes on species of North American Oligochaeta. IV. On a new lumbriculid genus from Florida, with additional notes on the nephridial and circulatory systems of Mesoporodrilus asymmetricus Smith. Bulletin of the Illinois State Laboratory of Natural History, 5, 459 - 479. https: // doi. org / 10.21900 / j. inhs. v 5.422" type="journal article" year="1900">Smith, 1900</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D7ECC46BCB6EFAD8" author="Smith" box="[159,303,1375,1401]" firstAuthor="Smith" pageId="5" pageNumber="146" pagination="459 - 479" refId="ref10665" refString="Smith, F. (1900) Notes on species of North American Oligochaeta. IV. On a new lumbriculid genus from Florida, with additional notes on the nephridial and circulatory systems of Mesoporodrilus asymmetricus Smith. Bulletin of the Illinois State Laboratory of Natural History, 5, 459 - 479. https: // doi. org / 10.21900 / j. inhs. v 5.422" type="journal article" year="1900">Smith, 1900</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Among congeners, the penes somewhat resemble those of
|
||||
<taxonomicName id="4C5D4D55FF937556D4AFC46BCF26FAD8" authority="(Beddard, 1908)" baseAuthorityName="Beddard" baseAuthorityYear="1908" box="[988,1383,1375,1401]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="cantabrigiensis">
|
||||
<emphasis id="B929EAC4FF937556D4AFC46BCEE7FAD8" box="[988,1190,1375,1401]" italics="true" pageId="5" pageNumber="146">T. cantabrigiensis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFCC4B27FF937556D3C5C46BCF1FFAD8" author="Beddard, F. E." box="[1206,1374,1375,1401]" pageId="5" pageNumber="146" pagination="365 - 369" refId="ref9179" refString="Beddard, F. E. (1908) A note on the occurrence of a species of Phreatothrix (Vejdovsky) in England, and on some points in its structure. Proceedings of the Zoological Society of London, 1908, 365 - 369. https: // doi. org / 10.1111 / j. 1096 - 3642.1908. tb 01848. x" type="book chapter" year="1908">Beddard, 1908</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D3C5C46BCF1FFAD8" author="Beddard" box="[1206,1374,1375,1401]" firstAuthor="Beddard" pageId="5" pageNumber="146" pagination="365 - 369" refId="ref9179" refString="Beddard, F. E. (1908) A note on the occurrence of a species of Phreatothrix (Vejdovsky) in England, and on some points in its structure. Proceedings of the Zoological Society of London, 1908, 365 - 369. https: // doi. org / 10.1111 / j. 1096 - 3642.1908. tb 01848. x" type="book chapter" year="1908">Beddard, 1908</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(see
|
||||
|
|
@ -503,7 +489,7 @@ Fig.
|
|||
<quantity id="4CA59B33FF937556D7B9C4B7CA99FA3C" box="[202,216,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="146" unit="in" value="1.0">1</quantity>
|
||||
</figureCitation>
|
||||
in
|
||||
<bibRefCitation id="EFCC4B27FF937556D788C4B7C852FA3C" author="Rodriguez, P. & Giani, N." box="[251,531,1411,1437]" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D788C4B7C852FA3C" author="Rodriguez & Giani" box="[251,531,1411,1437]" firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="35 - 51" refId="ref10466" refString="Rodriguez, P. & Giani, N. (1994) A preliminary review of the taxonomic characters used for the systematics of the genus Trichodrilus Claparede (Oligochaeta, Lumbriculidae). Hydrobiologia, 278, 35 - 51. https: // doi. org / 10.1007 / BF 00142310" type="journal article" year="1994">Rodriguez & Giani 1994</bibRefCitation>
|
||||
); however, the
|
||||
<taxonomicName id="4C5D4D55FF937556D5CFC4B7C9C5FA3C" baseAuthorityName="Beddard" baseAuthorityYear="1908" box="[700,900,1411,1437]" class="Clitellata" family="Lumbriculidae" genus="Trichodrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="cantabrigiensis">
|
||||
<emphasis id="B929EAC4FF937556D5CFC4B7C9C5FA3C" box="[700,900,1411,1437]" italics="true" pageId="5" pageNumber="146">T. cantabrigiensis</emphasis>
|
||||
|
|
@ -515,7 +501,7 @@ Muscular sphincters near the spermathecal pore have been described for species i
|
|||
<taxonomicName id="4C5D4D55FF937556D67AC4DBC826F9A8" authority="Eisen (Fend 2005)" authorityName="Eisen (Fend" authorityYear="2005" box="[265,615,1518,1545]" class="Clitellata" family="Lumbriculidae" genus="Eclipidrilus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B929EAC4FF937556D67AC4DBCBCCF9A8" box="[265,397,1519,1545]" italics="true" pageId="5" pageNumber="146">Eclipidrilus</emphasis>
|
||||
Eisen (
|
||||
<bibRefCitation id="EFCC4B27FF937556D691C4DBC81EF9A8" author="Fend, S. V." box="[482,607,1518,1545]" pageId="5" pageNumber="146" pagination="1 - 42" refId="ref9600" refString="Fend, S. V. (2005) A review of the genus Eclipidrilus (Annelida: Clitellata: Lumbriculidae), with description of a new species from western North America. Zootaxa, 969 (1), 1 - 42. https: // doi. org / 10.11646 / zootaxa. 969.1.1" type="journal article" year="2005">Fend 2005</bibRefCitation>
|
||||
<bibRefCitation id="EFCC4B27FF937556D691C4DBC81EF9A8" author="Fend" box="[482,607,1518,1545]" firstAuthor="Fend" pageId="5" pageNumber="146" pagination="1 - 42" refId="ref9600" refString="Fend, S. V. (2005) A review of the genus Eclipidrilus (Annelida: Clitellata: Lumbriculidae), with description of a new species from western North America. Zootaxa, 969 (1), 1 - 42. https: // doi. org / 10.11646 / zootaxa. 969.1.1" type="journal article" year="2005">Fend 2005</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
|
|
@ -527,7 +513,7 @@ Altman
|
|||
<taxonomicName id="4C5D4D55FF937556D41BC4DBCF29F9A8" authority="Michaelsen (Fend et al. 2017)" box="[872,1384,1519,1545]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B929EAC4FF937556D41BC4DBCE4BF9A8" box="[872,1034,1519,1545]" italics="true" pageId="5" pageNumber="146">Guestphalinus</emphasis>
|
||||
Michaelsen (
|
||||
<bibRefCitation id="EFCC4B27FF937556D3D1C4DBCF21F9A8" author="Fend S. V. & Rodriguez P. & Achurra A. & Erseus C." box="[1186,1376,1519,1545]" pageId="5" pageNumber="146" pagination="1 - 46" refId="ref9719" refString="Fend S. V., Rodriguez P., Achurra A. & Erseus C. (2017) On Kincaidiana Altman, 1936 and Guestphalinus Michaelsen, 1933 (Annelida, Clitellata, Lumbriculidae), with the descriptions of three new species. European Journal of Taxonomy, 361, 1 - 46. https: // doi. org / 10.5852 / ejt. 2017.361" type="journal article" year="2017">
|
||||
<bibRefCitation id="EFCC4B27FF937556D3D1C4DBCF21F9A8" author="Fend" box="[1186,1376,1519,1545]" etAl="et al." firstAuthor="Fend" pageId="5" pageNumber="146" pagination="1 - 46" refId="ref9719" refString="Fend S. V., Rodriguez P., Achurra A. & Erseus C. (2017) On Kincaidiana Altman, 1936 and Guestphalinus Michaelsen, 1933 (Annelida, Clitellata, Lumbriculidae), with the descriptions of three new species. European Journal of Taxonomy, 361, 1 - 46. https: // doi. org / 10.5852 / ejt. 2017.361" type="journal article" year="2017">
|
||||
Fend
|
||||
<emphasis id="B929EAC4FF937556D390C4DBCF5DF9A8" box="[1251,1308,1519,1545]" italics="true" pageId="5" pageNumber="146">et al.</emphasis>
|
||||
2017
|
||||
|
|
@ -537,7 +523,7 @@ Fend
|
|||
, and
|
||||
<taxonomicName id="4C5D4D55FF937556D7E4C727C832F98C" authority="Rodriguez et al., 2014" authorityName="Rodriguez" authorityYear="2014" box="[151,627,1555,1581]" class="Clitellata" family="Lumbriculidae" genus="Sylphella" kingdom="Animalia" order="Lumbriculida" pageId="5" pageNumber="146" phylum="Annelida" rank="species" species="puccoon">
|
||||
<emphasis id="B929EAC4FF937556D7E4C727CB26F98D" box="[151,359,1555,1581]" italics="true" pageId="5" pageNumber="146">Sylphella puccoon</emphasis>
|
||||
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|
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<bibRefCitation id="EFCC4B27FF937556D603C727C832F98C" author="Rodriguez" box="[368,627,1555,1581]" etAl="et al." firstAuthor="Rodriguez" pageId="5" pageNumber="146" pagination="1 - 32" refId="ref10519" refString="Rodriguez, P., Fend, S. V. & Lenat, D. R. (2014) Sylphella puccoon gen. n., sp. n. and two additional new species of aquatic oligochaetes (Lumbriculidae, Clitellata) from poorly-known lotic habitats in North Carolina (USA). ZooKeys, 451, 1 - 32. https: // doi. org / 10.3897 / zookeys. 451.7304" type="journal article" year="2014">
|
||||
Rodriguez
|
||||
<emphasis id="B929EAC4FF937556D69EC727C866F98C" box="[493,551,1555,1581]" italics="true" pageId="5" pageNumber="146">et al.</emphasis>
|
||||
, 2014
|
||||
|
|
@ -554,7 +540,7 @@ Claparède
|
|||
</taxonomicName>
|
||||
in Lake Baikal (e.g., Fig.
|
||||
<quantity id="4CA59B33FF937556D201C76FCAECF938" metricMagnitude="0" metricUnit="m" metricValue="5.3086" pageId="5" pageNumber="146" unit="in" value="209.0">209 in</quantity>
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<bibRefCitation id="EFCC4B27FF937556D7C7C74BCB2BF938" author="Semernoy, V. P." box="[180,362,1663,1689]" pageId="5" pageNumber="146" refId="ref10595" refString="Semernoy, V. P. (2004) Oligochaeta of Lake Baikal. Guides and Keys to Identification of Fauna and Flora of Lake Baikal. Russian Academy of Sciences. Nauka, Novosibirsk, 528 pp." type="book" year="2004">Semernoy 2004</bibRefCitation>
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<bibRefCitation id="EFCC4B27FF937556D7C7C74BCB2BF938" author="Semernoy" box="[180,362,1663,1689]" firstAuthor="Semernoy" pageId="5" pageNumber="146" refId="ref10595" refString="Semernoy, V. P. (2004) Oligochaeta of Lake Baikal. Guides and Keys to Identification of Fauna and Flora of Lake Baikal. Russian Academy of Sciences. Nauka, Novosibirsk, 528 pp." type="book" year="2004">Semernoy 2004</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
163
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|
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</taxonomicName>
|
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[
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFE461050FDF5FE98" authorityName="Busck" authorityYear="1932" box="[489,602,356,382]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="77" phylum="Arthropoda" rank="tribe" tribe="Atteriini">Atteriini</taxonomicName>
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<emphasis id="D8B2081F1860FF8DFF38109CFEB5FE26" box="[151,282,424,448]" italics="true" pageId="26" pageNumber="77">Archipimima</emphasis>
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<bibRefCitation id="8E57A9FC1860FF8DFE8E109CFE78FE26" author="Powell, J. A." box="[289,471,424,448]" pageId="26" pageNumber="77" pagination="372 - 398" refId="ref27669" refString="Powell, J. A. (1986) Synopsis of the classification of Neotropical Tortricinae, with descriptions of new genera and species (Lepidoptera: Tortricidae). Pan - Pacific Entomologist, 62, 372 - 398." type="journal article" year="1986">Powell 1986: 385</bibRefCitation>
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|
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</treatmentCitation>
|
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;
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<treatmentCitation id="6B67F21C1860FF8DFE4D109CFD7EFE27" author="Powell, J. A. & Razowski, J. & Brown, J. W." box="[482,721,424,449]" page="148" pageId="26" pageNumber="77" year="1995">
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<bibRefCitation id="8E57A9FC1860FF8DFE4D109CFD7EFE27" author="Powell, J. A. & Razowski, J. & Brown, J. W." box="[482,721,424,449]" pageId="26" pageNumber="77" pagination="138 - 151" refId="ref27710" refString="Powell, J. A., Razowski, J. & Brown, J. W. (1995) Tortricidae: Tortricinae, Chlidanotinae. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera, Checklist Part II: Hyblaeoidea - Pyraloidea - Tortricoidea. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida, pp. 138 - 151." type="book chapter" year="1995">
|
||||
Powell
|
||||
<emphasis id="D8B2081F1860FF8DFD80109DFDCDFE26" box="[559,610,424,448]" italics="true" pageId="26" pageNumber="77">et al.</emphasis>
|
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1995: 148
|
||||
</bibRefCitation>
|
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</treatmentCitation>
|
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; Razowski 2004: 349;
|
||||
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<bibRefCitation id="8E57A9FC1860FF8DFC15109CFB75FE27" author="Razowski, J. & Pelz, V." box="[954,1242,424,449]" pageId="26" pageNumber="77" pagination="131 - 144" refId="ref28690" refString="Razowski, J. & Pelz, V. (2004) Tortricidae collected in Ecuador in the years 1996 - 1999: Archipini and Atteriini (Lepidoptera). Nachrichten des Entomologischen Vereins Apollo, N. F., 25 (3), 131 - 144." type="journal article" year="2004">Razowski & Pelz 2004: 140</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<treatmentCitation id="6B67F21C1860FF8DFB4B109DFA37FE27" author="Brown, J. W." box="[1252,1432,424,449]" page="117" pageId="26" pageNumber="77" year="2005">
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<bibRefCitation id="8E57A9FC1860FF8DFB4B109DFA37FE27" author="Brown, J. W." box="[1252,1432,424,449]" pageId="26" pageNumber="77" refId="ref25469" refString="Brown, J. W. (2005) World catalogue of insects. Vol. 5. Tortricidae (Lepidoptera). Apollo Books, Stenstrup, 741 pp." type="book" year="2005">Brown 2005: 117</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<bibRefCitation id="8E57A9FC1860FF8DFF6810F3FDBCFE39" author="Razowski, J. & Wojtusiak, J." box="[199,531,455,479]" pageId="26" pageNumber="77" pagination="17 - 53" refId="ref28850" refString="Razowski, J. & Wojtusiak, J. (2006) Tortricidae (Lepidoptera) from the Valley of Rio Gualaceo, East Cordillera in Ecuador, with descriptions of new taxa. Acta Zoologica Cracoviensia, 49 (B), 17 - 53. https: // doi. org / 10.3409 / 000000006783995346" type="journal article" year="2006">Razowski & Wojtusiak 2006: 33</bibRefCitation>
|
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</treatmentCitation>
|
||||
, 2010: 113, 2013: 22;
|
||||
<treatmentCitation id="6B67F21C1860FF8DFD5910F3FB9FFE39" author="Razowski, J. & Becker, V. O." box="[758,1072,455,479]" page="167" pageId="26" pageNumber="77" year="2011">
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<bibRefCitation id="8E57A9FC1860FF8DFD5910F3FB9FFE39" author="Razowski, J. & Becker, V. O." box="[758,1072,455,479]" pageId="26" pageNumber="77" pagination="161 - 181" refId="ref28557" refString="Razowski, J. & Becker, V. O. (2011) Systematic and faunistic data on Neotropical Tortricidae: Phricanthini, Tortricini, Atteriini, Polyorthini, Chlidanotini (Lepidoptera: Tortricidae). SHILAP Revista de Lepidopterologia, 39, 161 - 181." type="journal article" year="2011">Razowski & Becker 2011: 167</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="A2DC87861860FF8DFF381339FD13FDC1" box="[151,700,525,552]" pageId="26" pageNumber="77" type="type_taxon">
|
||||
<paragraph id="EA79D40D1860FF8DFF381339FD13FDC1" blockId="26.[151,1436,525,804]" box="[151,700,525,552]" pageId="26" pageNumber="77">
|
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<emphasis id="D8B2081F1860FF8DFF381339FE84FDC1" bold="true" box="[151,299,525,551]" pageId="26" pageNumber="77">
|
||||
<typeStatus id="357D6AAF1860FF8DFF381339FF7DFDC1" box="[151,210,525,551]" pageId="26" pageNumber="77">Type</typeStatus>
|
||||
species
|
||||
</emphasis>
|
||||
:
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFE95133AFD17FDC1" ID-CoL="98LLY" authority="Dognin, 1908" authorityName="Dognin" authorityYear="1908" box="[314,696,525,551]" class="Insecta" family="Tortricidae" genus="Tortrix" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="77" phylum="Arthropoda" rank="species" species="flexicostalis">
|
||||
<emphasis id="D8B2081F1860FF8DFE95133AFDBBFDC1" box="[314,532,525,551]" italics="true" pageId="26" pageNumber="77">Tortrix flexicostalis</emphasis>
|
||||
Dognin, 1908
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="A2DC87861860FF8DFF681305FC35FCC5" pageId="26" pageNumber="77" type="diagnosis">
|
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|
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<emphasis id="D8B2081F1860FF8DFF681305FE71FDAD" bold="true" box="[199,478,561,587]" pageId="26" pageNumber="77">Diagnosis and remarks</emphasis>
|
||||
.
|
||||
<bibRefCitation id="8E57A9FC1860FF8DFE421305FD21FDAD" author="Powell, J. A." box="[493,654,561,587]" pageId="26" pageNumber="77" pagination="372 - 398" refId="ref27669" refString="Powell, J. A. (1986) Synopsis of the classification of Neotropical Tortricinae, with descriptions of new genera and species (Lepidoptera: Tortricidae). Pan - Pacific Entomologist, 62, 372 - 398." type="journal article" year="1986">Powell (1986)</bibRefCitation>
|
||||
proposed the genus
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFCD41305FBA3FDAD" authorityName="Powell" authorityYear="1986" box="[891,1036,561,587]" class="Insecta" family="Tortricidae" genus="Archipimima" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="D8B2081F1860FF8DFCD41305FBA3FDAD" box="[891,1036,561,587]" italics="true" pageId="26" pageNumber="77">Archipimima</emphasis>
|
||||
</taxonomicName>
|
||||
for three Meyrick and one Dognin species. Additional species were added by Razowski (2004),
|
||||
<bibRefCitation id="8E57A9FC1860FF8DFCCE1361FB2CFD89" author="Razowski, J. & Pelz, V." box="[865,1155,597,623]" pageId="26" pageNumber="77" pagination="131 - 144" refId="ref28690" refString="Razowski, J. & Pelz, V. (2004) Tortricidae collected in Ecuador in the years 1996 - 1999: Archipini and Atteriini (Lepidoptera). Nachrichten des Entomologischen Vereins Apollo, N. F., 25 (3), 131 - 144." type="journal article" year="2004">Razowski & Pelz (2004)</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="8E57A9FC1860FF8DFB3D1361FF74FD75" author="Razowski, J. & Wojtusiak, J." pageId="26" pageNumber="77" pagination="17 - 53" refId="ref28850" refString="Razowski, J. & Wojtusiak, J. (2006) Tortricidae (Lepidoptera) from the Valley of Rio Gualaceo, East Cordillera in Ecuador, with descriptions of new taxa. Acta Zoologica Cracoviensia, 49 (B), 17 - 53. https: // doi. org / 10.3409 / 000000006783995346" type="journal article" year="2006">Razowski & Wojtusiak (2006</bibRefCitation>
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, 2010, 2013), and
|
||||
<bibRefCitation id="8E57A9FC1860FF8DFE18134DFD5BFD75" author="Razowski, J. & Becker, V. O." box="[439,756,633,659]" pageId="26" pageNumber="77" pagination="161 - 181" refId="ref28557" refString="Razowski, J. & Becker, V. O. (2011) Systematic and faunistic data on Neotropical Tortricidae: Phricanthini, Tortricini, Atteriini, Polyorthini, Chlidanotini (Lepidoptera: Tortricidae). SHILAP Revista de Lepidopterologia, 39, 161 - 181." type="journal article" year="2011">Razowski & Becker (2011)</bibRefCitation>
|
||||
, resulting in a total of 12 species currently assigned to the genus. Species of
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFECD13A9FE5CFD51" authorityName="Powell" authorityYear="1986" box="[354,499,669,695]" class="Insecta" family="Tortricidae" genus="Archipimima" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="D8B2081F1860FF8DFECD13A9FE5CFD51" box="[354,499,669,695]" italics="true" pageId="26" pageNumber="77">Archipimima</emphasis>
|
||||
</taxonomicName>
|
||||
are recorded from
|
||||
<collectingCountry id="92D1949D1860FF8DFD6313A9FC95FD51" box="[716,826,669,695]" name="Colombia" pageId="26" pageNumber="77">Colombia</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="92D1949D1860FF8DFCDE13A9FC4BFD51" box="[881,996,669,695]" name="Venezuela" pageId="26" pageNumber="77">Venezuela</collectingCountry>
|
||||
to
|
||||
<collectingCountry id="92D1949D1860FF8DFBA713AAFB94FD5E" box="[1032,1083,670,696]" name="Peru" pageId="26" pageNumber="77">Peru</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="92D1949D1860FF8DFBDC13A9FB18FD51" box="[1139,1207,669,695]" name="Brazil" pageId="26" pageNumber="77">Brazil</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="28021AEF1860FF8DFB6913A9FAC2FD51" box="[1222,1389,669,695]" country="Brazil" name="Santa Catarina" pageId="26" pageNumber="77">Santa Catarina</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="28021AEF1860FF8DFF3813F5FF47FD3D" box="[151,232,705,731]" country="Brazil" name="Parana" pageId="26" pageNumber="77">Paraná</collectingRegion>
|
||||
). The new species described below extends the geographic range of the genus considerably southward to
|
||||
<collectingCountry id="92D1949D1860FF8DFF3813D1FEA5FD19" box="[151,266,741,767]" name="Argentina" pageId="26" pageNumber="77">Argentina</collectingCountry>
|
||||
. An undetermined species of
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFDFE13D1FCE2FD19" authority="(USNM)" baseAuthorityName="USNM" box="[593,845,741,767]" class="Insecta" family="Tortricidae" genus="Archipimima" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="D8B2081F1860FF8DFDFE13D1FD4DFD19" box="[593,738,741,767]" italics="true" pageId="26" pageNumber="77">Archipimima</emphasis>
|
||||
(USNM)
|
||||
</taxonomicName>
|
||||
was reared from
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFBA113D2FAA3FD19" authority="Thunb." authorityName="Thunb." box="[1038,1292,741,767]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="26" pageNumber="77" phylum="Tracheophyta" rank="species" species="kaki">
|
||||
<emphasis id="D8B2081F1860FF8DFBA113D2FB19FD19" box="[1038,1206,741,767]" italics="true" pageId="26" pageNumber="77">Diospyros kaki</emphasis>
|
||||
Thunb.
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="2DC6AF8E1860FF8DFAB613D1FA3BFD19" box="[1305,1428,741,767]" class="Magnoliopsida" family="Ebenaceae" kingdom="Plantae" order="Ericales" pageId="26" pageNumber="77" phylum="Tracheophyta" rank="family">Ebenaceae</taxonomicName>
|
||||
) in
|
||||
<collectingCountry id="92D1949D1860FF8DFF1B123DFF53FCC5" box="[180,252,777,803]" name="Brazil" pageId="26" pageNumber="77">Brazil</collectingCountry>
|
||||
. Otherwise, life history features for the genus are unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue