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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.827.31365" ID-GBIF-Dataset="85f74389-c078-44de-b0bd-8b7e4c3b9404" ID-PMC="PMC6472301" ID-Pensoft-Pub="1313-2970-827-31" ID-PubMed="31114425" ID-ZBK="10B663895E424E5287D8F49E2405D651" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-827-31" ModsDocOrigin="ZooKeys 827" ModsDocTitle="Mollusc species from the Pontocaspian region – an expert opinion list" checkinTime="1551875342560" checkinUser="pensoft" docAuthor="Wesselingh, Frank, Poorten, Jan Johan ter, Kijashko, Pavel, Albrecht, Christian, Anistratenko, Olga Yu, Frolov, Pavel, Gándara, Alberto Martinez, Gittenberger, Arjan, Gogaladze, Aleksandre, Mikhail Karpinsky, Popa, Luis, Sands, Arthur F, Vandendorpe, Justine &amp; Wilke, Thomas" docDate="2019" docId="62DD080E5F88CD46DA53BFCEFAED20F2" docLanguage="en" docName="ZooKeys 827: 31-124" docOrigin="ZooKeys 827" docSource="http://dx.doi.org/10.3897/zookeys.827.31365" docTitle="Clessiniola variabilis" docType="treatment" docVersion="5" lastPageNumber="74" masterDocId="FFA2FA62FFEC4705FFBBFB79FFAAFFFE" masterDocTitle="Mollusc species from the Pontocaspian region - an expert opinion list" masterLastPageNumber="124" masterPageNumber="31" pageNumber="73" updateTime="1668166983143" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Mollusc species from the Pontocaspian region - an expert opinion list</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesselingh, Frank</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poorten, Jan Johan ter</mods:namePart>
<mods:affiliation>Field Museum of Natural History, Chicago, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kijashko, Pavel</mods:namePart>
<mods:affiliation>Moscow State University, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Albrecht, Christian</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anistratenko, Olga Yu</mods:namePart>
<mods:affiliation>Schmalhausen Instite of Zoology, National Academy of Sciences of Ukraine, Kiev, Ukraine &amp; Institute of Geological Sciences, National Academy of Sciences of Ukraine, KievUkraine</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Pavel</mods:namePart>
<mods:affiliation>Saint-Petersburg State University, Saint Petersburg, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gándara, Alberto Martinez</mods:namePart>
<mods:affiliation>Grigore Artipa National Museum of Natural History, Bucharest, Romania</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gittenberger, Arjan</mods:namePart>
<mods:affiliation>Gittenberger Marine Research, Inventory &amp; Strategy, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gogaladze, Aleksandre</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mikhail Karpinsky</mods:namePart>
<mods:affiliation>Russian Federal Research Institute of Fisheries and Oceanography, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Popa, Luis</mods:namePart>
<mods:affiliation>Grigore Antirpa National Museum of Natural History, Bucharest, Romania</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sands, Arthur F</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vandendorpe, Justine</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wilke, Thomas</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>827</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>124</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.827.31365</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.827.31365</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-827-31</mods:identifier>
<mods:identifier type="ZBK">10B663895E424E5287D8F49E2405D651</mods:identifier>
<mods:identifier type="ZooBank">10B663895E424E5287D8F49E2405D651</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183410471" LSID="urn:lsid:plazi:treatment:62DD080E5F88CD46DA53BFCEFAED20F2" httpUri="http://treatment.plazi.org/id/62DD080E5F88CD46DA53BFCEFAED20F2" lastPageNumber="74" pageNumber="73">
<subSubSection type="nomenclature">
<paragraph pageId="42" pageNumber="73">
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Clessiniola variabilis</taxonomicName>
(Eichwald, 1838)
</paragraph>
</subSubSection>
<subSubSection type="reference_group">
<paragraph pageId="42" pageNumber="73">
*1838
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Paludina variabilis</taxonomicName>
Eichwald: 151-152.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1838
<taxonomicName class="Gastropoda" family="Amphimelaniidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Paludina</taxonomicName>
Triton Eichwald: 152.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1874
<taxonomicName class="Gastropoda" family="Bithyniidae" genus="Bithynia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bithynia" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Bithynia</taxonomicName>
?
<taxonomicName genus="Eichwaldi" lsidName="Eichwaldi" pageId="42" pageNumber="73" rank="genus">Eichwaldi</taxonomicName>
Martens: 81.
</paragraph>
<paragraph pageId="42" pageNumber="73">
?1887
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Caspia</taxonomicName>
<taxonomicName genus="Grimmi" lsidName="Grimmi" pageId="42" pageNumber="73" rank="genus">Grimmi</taxonomicName>
Clessin and Dybowski in Dybowski: 39
</paragraph>
<paragraph pageId="42" pageNumber="73">
?1888 [
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Caspia</taxonomicName>
]
<taxonomicName genus="Grimmi" lsidName="Grimmi" pageId="42" pageNumber="73" rank="genus">Grimmi</taxonomicName>
n. sp. - Dybowski: 79, pl. 3, fig. 5a, b.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1887
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Clessinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessinia" order="Stylommatophora" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Clessinia</taxonomicName>
Martensii Clessin and Dybowski in Dybowski: 43.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1888
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Clessinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessinia" order="Stylommatophora" pageId="42" pageNumber="73" phylum="Mollusca" rank="genus">Clessinia</taxonomicName>
Martensii n. sp. - Dybowski: 79, pl. 2, fig. 5.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1902a
<taxonomicName class="Gastropoda" family="Odontostomidae" genus="Clessinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessinia ahngeri" order="Stylommatophora" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="ahngeri">Clessinia ahngeri</taxonomicName>
Westerlund: 45-46.
</paragraph>
<paragraph pageId="42" pageNumber="73">1966 P.[yrgula] (Clessiniola) pseudotriton Golikov and Starobogatov: 356-357, fig. 2(3</paragraph>
<paragraph pageId="42" pageNumber="73">
?1969
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgula (Caspiella) derbentina" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="derbentina" subGenus="Caspiella">Pyrgula (Caspiella) derbentina</taxonomicName>
Logvinenko and Starobogatov: 374, fig. 366(8).
</paragraph>
<paragraph pageId="42" pageNumber="73">
1969
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgula (Caspiella) ovum" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="ovum" subGenus="Caspiella">Pyrgula (Caspiella) ovum</taxonomicName>
Logvinenko and Starobogatov: 374, fig. 366(9).
</paragraph>
<paragraph pageId="42" pageNumber="73">
1969
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgula (Caspiella) trivialis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="trivialis" subGenus="Caspiella">Pyrgula (Caspiella) trivialis</taxonomicName>
Logvinenko and Starobogatov: 374-375, fig. 366(10).
</paragraph>
<paragraph pageId="42" pageNumber="73">
1987
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia (Clessiniola) variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis" subGenus="Clessiniola">Turricaspia (Clessiniola) variabilis</taxonomicName>
(Eichwald, 1838). - Alexenko and Starobogatov: 34, text fig. 5.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1987
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia (Clessiniola) triton" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="triton" subGenus="Clessiniola">Turricaspia (Clessiniola) triton</taxonomicName>
(Eichwald, 1838). - Alexenko and Starobogatov: 34, text fig. 3.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1987
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia (Clessiniola) martensii" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="martensii" subGenus="Clessiniola">Turricaspia (Clessiniola) martensii</taxonomicName>
(Clessin &amp; Dybowski in Dybowski, 1888). - Alexenko and Starobogatov: 34, text fig. 4.
</paragraph>
<paragraph pageId="42" pageNumber="73">
1987
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia (Clessiniola) bogensis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="bogensis" subGenus="Clessiniola">Turricaspia (Clessiniola) bogensis</taxonomicName>
(
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
, 1852). - Alexenko and Starobogatov: 34.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2006
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Turricaspia variabilis</taxonomicName>
(Eichwald, 1838). - Kantor and Sysoev: 111, pl. 49, fig. J.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2011
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia martensii" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="martensii">Turricaspia martensii</taxonomicName>
(Clessin &amp; W. Dybowski in W. Dybowski, 1888). - Anistratenko et al.: 86, fig. 3(17).
</paragraph>
<paragraph pageId="42" pageNumber="73">
2011
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia triton" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="triton">Turricaspia triton</taxonomicName>
(Eichwald, 1838). - Anistratenko et al.: 85-86, fig. 3(16).
</paragraph>
<paragraph pageId="42" pageNumber="73">
2011
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Turricaspia variabilis</taxonomicName>
(Eichwald, 1838). - Anistratenko et al.: 85, fig. 3(15).
</paragraph>
<paragraph pageId="42" pageNumber="73">
2014
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Turricaspia variabilis</taxonomicName>
. - Taviani et al.: 4, fig. 3b.
</paragraph>
<paragraph pageId="42" pageNumber="73">
?2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia derbentina" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="derbentina">Turricaspia derbentina</taxonomicName>
(Logvinenko &amp; Starobogatov, 1968). - Vinarski and Kantor: 247.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia martensii" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="martensii">Turricaspia martensii</taxonomicName>
(Clessin &amp; W. Dybowski in W. Dybowski, 1888). - Vinarski and Kantor: 248.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia ovum" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="ovum">Turricaspia ovum</taxonomicName>
(Logvinenko &amp; Starobogatov, 1968). - Vinarski and Kantor: 248-249.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia pseudotriton" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="pseudotriton">Turricaspia pseudotriton</taxonomicName>
(Golikov &amp; Starobogatov, 1966). - Vinarski and Kantor: 249.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia triton" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="triton">Turricaspia triton</taxonomicName>
(Eichwald, 1838). - Vinarski and Kantor: 250.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia trivialis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="trivialis">Turricaspia trivialis</taxonomicName>
(Logvinenko &amp; Starobogatov, 1968). - Vinarski and Kantor: 250-251.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Turricaspia variabilis</taxonomicName>
(Eichwald, 1838).- Vinarski and Kantor: 251.
</paragraph>
<paragraph pageId="42" pageNumber="73">
2018
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola variabilis" order="Littorinimorpha" pageId="42" pageNumber="73" phylum="Mollusca" rank="species" species="variabilis">Clessiniola variabilis</taxonomicName>
(Eichwald, 1838). - Neubauer et al.: 60-63, fig. 7
<normalizedToken originalValue="A–I">A-I</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="43" pageNumber="74">
<pageBreakToken pageId="43" pageNumber="74" start="start">Status</pageBreakToken>
. Accepted Pontocaspian species.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageId="43" pageNumber="74">Type locality. At the Volga River mouth near Astrakhan, and towards the Caspian Sea; also in recently lithified fossil limestone at the shores of Dagestan, Russia.</paragraph>
<paragraph pageId="43" pageNumber="74">
Distribution. Caspian Sea, Azov Sea, and northern Black Sea region. This species was mentioned in the South Caspian Basin of Azerbaijan (
<bibRefCitation author="Mirzoev, GS" journalOrPublisher="International Journal of Zoological Studies" pageId="79" pageNumber="110" pagination="43 - 48" title="Zoobenthos distribution patterns in the deepwater horizon of Azerbaijan sector of the Caspian Sea." url="http://www.zoologyjournals.com/download/74/1-7-29-696.pdf" volume="2" year="2017">Mirzoev and Alekperov 2017</bibRefCitation>
, who reported the species as
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia variabilis" order="Littorinimorpha" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="variabilis">Turricaspia variabilis</taxonomicName>
,
<taxonomicName lsidName="T. derbentica" pageId="43" pageNumber="74" rank="species" species="derbentica">T. derbentica</taxonomicName>
, and
<taxonomicName lsidName="T. trivialis" pageId="43" pageNumber="74" rank="species" species="trivialis">T. trivialis</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="43" pageNumber="74">
Taxonomic notes.
<bibRefCitation author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="80" pageNumber="111" pagination="43 - 103" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. (2018)</bibRefCitation>
recently demonstrated the high variability of this species. Comparison of available illustrations and descriptions of the species listed in the synonymy list indicates that all of them range within this
<normalizedToken originalValue="species’">species'</normalizedToken>
variability. Consequently, we consider all of them as junior synonyms of
<taxonomicName lsidName="C. variabilis" pageId="43" pageNumber="74" rank="species" species="variabilis">C. variabilis</taxonomicName>
. A more in-depth review of the type material of the species involved is required to confirm this approach.
</paragraph>
<paragraph pageId="43" pageNumber="74">
The status of
<taxonomicName class="Gastropoda" family="Amphimelaniidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina bogensis" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="bogensis">Paludina bogensis</taxonomicName>
Dubois in
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
, 1852, which was listed as a valid species of
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="43" pageNumber="74" phylum="Mollusca" rank="genus">Turricaspia</taxonomicName>
by
<bibRefCitation author="Anistratenko, VV" journalOrPublisher="Naukova dumka, Kiev" pageId="72" pageNumber="103" title="Fauna Ukraine. Vol. 29: Mollusca. Fasc. 1. B. 2: Orders Littoriniformes, Rissoiformes." year="1995">Anistratenko and Stadnichenko (1995)</bibRefCitation>
, is still unclear. That species was described from the Zapadnyi Bug River in Poland and closely resembles
<taxonomicName lsidName="C. variabilis" pageId="43" pageNumber="74" rank="species" species="variabilis">C. variabilis</taxonomicName>
. It is, however, unlikely that a Pontocaspian species typical of oligohaline conditions occurs so far away in a pure freshwater environment. &quot;
<taxonomicName class="Gastropoda" family="Amphimelaniidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina eichwaldi" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="eichwaldi">Paludina eichwaldi</taxonomicName>
Krynicki, 1837&quot; found in the literature is a nomen nudum.
<bibRefCitation author="Martens, Ev" journalOrPublisher="Supplement" pageId="78" pageNumber="109" pagination="1 - 127" title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates conchologicae." url="https://biodiversitylibrary.org/page/12980992" volume="5" year="1874">Martens (1874)</bibRefCitation>
provided measurements and made the name available, but he listed
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina variabilis" order="Littorinimorpha" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="variabilis">Paludina variabilis</taxonomicName>
Eichwald, 1838 in synonymy, which has priority. Dybowski (1887) obviously overlooked this and considered
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Nematurella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nematurella eichwaldi" order="Littorinimorpha" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="eichwaldi">Nematurella eichwaldi</taxonomicName>
Krynicki a valid species. We follow
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="86" pageNumber="117" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor (2016)</bibRefCitation>
and consider the species as a junior synonym of
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola variabilis" order="Littorinimorpha" pageId="43" pageNumber="74" phylum="Mollusca" rank="species" species="variabilis">Clessiniola variabilis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection type="conservation">
<paragraph pageId="43" pageNumber="74">
Conservation status. Least Concern (
<bibRefCitation author="Cioboiu, O" journalOrPublisher="Bollettino della Societa Malacologica Italiana" pageId="73" pageNumber="104" url="https://doi.org/10.2305/IUCN.UK.2011-2.RLTS.T155608A4807675.en" year="2011">Cioboiu et al. 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.384.6197" ID-GBIF-Dataset="f8902ab2-4591-440e-93f1-fb2d2f850112" ID-PMC="PMC3950428" ID-Pensoft-Pub="1313-2970-384-1" ID-PubMed="24624022" ID-ZBK="832EF8274642416895252C2AD202EB9B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-384-1" ModsDocOrigin="ZooKeys 384" ModsDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" checkinTime="1451246226561" checkinUser="pensoft" docAuthor="Hopkins, Heidi" docDate="2014" docId="62DD8A48D5D05D68EF16BFFC909A12BB" docLanguage="en" docName="ZooKeys 384: 1-256" docOrigin="ZooKeys 384" docSource="http://dx.doi.org/10.3897/zookeys.384.6197" docTitle="Arenivaga rothi Hopkins, 2014, sp. n." docType="treatment" docUuid="AE40A344-BA80-414C-BA4A-16A469412DBF" docUuidSource="ZooBank" docVersion="4" lastPageNumber="105" masterDocId="FF86641EFFB8B30DFFC9F429FFD4FFA1" masterDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" masterLastPageNumber="256" masterPageNumber="1" pageNumber="104" updateTime="1668157791821" updateUser="ExternalLinkService">
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<mods:title>A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hopkins, Heidi</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
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<mods:number>384</mods:number>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.384.6197</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.384.6197</mods:identifier>
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<treatment ID-GBIF-Taxon="152051405" LSID="urn:lsid:zoobank.org:act:AE40A344-BA80-414C-BA4A-16A469412DBF" httpUri="http://treatment.plazi.org/id/62DD8A48D5D05D68EF16BFFC909A12BB" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104">
<subSubSection pageId="103" pageNumber="104" type="nomenclature">
<paragraph pageId="103" pageNumber="104">
<taxonomicName LSID="http://zoobank.org/AE40A344-BA80-414C-BA4A-16A469412DBF" class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="rothi">
<pageBreakToken pageId="103" pageNumber="104" start="start">Arenivaga</pageBreakToken>
rothi
</taxonomicName>
<taxonomicNameLabel pageId="103" pageNumber="104">sp. n.</taxonomicNameLabel>
Figures 138-140
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="type locality">
<paragraph pageId="103" pageNumber="104">Type locality.</paragraph>
<paragraph pageId="103" pageNumber="104">MEXICO, Coahuila, 8 mi N Viesca.</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="material examined">
<paragraph pageId="103" pageNumber="104">Material examined.</paragraph>
<paragraph pageId="103" pageNumber="104">
Holotype: ♂ in EMEC labeled &quot;MEX: Coah., sand dunes at Bilbao, 8 mi. N. Viesca, V-30/31-1981, J. Doyen, J. Liebherr, at blacklight&quot; &quot;HOLOTYPE
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
Hopkins, 2012&quot; [red label with black border].
</paragraph>
<paragraph pageId="103" pageNumber="104">
Paratypes (13): USA: TX, Presidio, May-June 1953, 53-10102, Presidio-3268L, at lights (8, USNM); TX, Presidio, 6/?-8/?/1955, JH Russell, at electric light (1, USNM); TX, Presidio, 6/17/1954, JH Russell, at light, 54-6842, Int. 3399L, (1, HEH); TX, Presidio, 5/14/1944, Presidio 1334, Lot No. 44-16214 (1, LACM). MEXICO: Coahuila, Bilbao 8 mi N of Viesca, 5/30-31/1981, Doyen &amp; Liebherr, on sand at night (2, EMEC); Coahuila, 10 mi E of San Pedro de las Colonias, 7/3/1959, ER Tinkham, low dunes (1, USNM). All paratypes labeled &quot;Paratype
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
Hopkins 2012&quot; [blue label with black border].
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="etymology">
<paragraph pageId="103" pageNumber="104">Etymology.</paragraph>
<paragraph pageId="103" pageNumber="104">The name is a noun in the genitive case. This species is named for the late Louis M. Roth who devoted his career to the description of new cockroach species and co-authored Cockroaches: Ecology, Behavior and Natural History.</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="distribution">
<paragraph pageId="103" pageNumber="104">Distribution.</paragraph>
<paragraph pageId="103" pageNumber="104">This species is found in north central Mexico in the state of Coahuila up to Presidio County, Texas. See Fig. 140.</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="diagnosis">
<paragraph pageId="103" pageNumber="104">Diagnosis.</paragraph>
<paragraph pageId="103" pageNumber="104">
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
may be distinguished by the very wide gap on the right ventral phallomere and the unusually narrow, pointed head on the genital hook. See Fig. 139.
</paragraph>
</subSubSection>
<subSubSection lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104" type="description">
<paragraph pageId="103" pageNumber="104">Description.</paragraph>
<paragraph pageId="103" pageNumber="104">Male.Measurements. Holotype TL = 21.0 mm, GW = 10.2 mm, PW = 6.64 mm, PL = 4.65 mm, TL/GW = 2.06, PL/PW = 0.70. EW = 0.20 mm; OW = 0.40 mm. Size range among paratypes: TL 17.9-21.8 mm, GW 8.2-10.4 mm, PW 5.70-7.00 mm, PW 4.40-4.82 mm.</paragraph>
<paragraph pageId="103" pageNumber="104">
Head. Two ocelli large, ovoid and protruding (0.50
<normalizedToken originalValue="×">x</normalizedToken>
0.40 mm); vertex dark brown, with small ridges between apices of eyes and extending onto ocellar tubercles; interocellar space concave, dark brown, light brown medially, with two pale spots medial to base of ocelli. Frons light brown; posterior concave; anterior portion of frons bulbous, light brown; wide light brown anteclypeus. See Fig. 138d.
</paragraph>
<paragraph pageId="103" pageNumber="104">Pronotum. Pronotum translucent waxy beige; dorsal surface of pronotum with dense very short light orange-brown setae that are thicker and longer laterally; pronotal pattern light orange-brown &quot;hippo face&quot; with little discernible detail; no aura. See Fig. 138c.</paragraph>
<paragraph pageId="103" pageNumber="104">Body. Wing brace present. Two tarsal claws present. Legs and body light brown; sternites with small pale maculations and fine lines laterally on each; subgenital plate light orange-brown with darker border; asymmetrical with pointed apices. See Fig. 138b.</paragraph>
<paragraph pageId="103" pageNumber="104">Forewings. Wings extended well beyond abdominal apex (up to ~35% of wing length); very pale translucent white-beige or brown; surface translucent and matte or with dull sheen. See Fig. 138a.</paragraph>
<paragraph pageId="104" pageNumber="105">
<pageBreakToken pageId="104" pageNumber="105" start="start">Genitalia</pageBreakToken>
. Right dorsal phallomere composed of lightly sclerotized, bulbous hook-shaped lobe, articulated with right ventral phallomere on lateral side; central field lightly sclerotized, punctate and deeply foreshortened; medial edge rolled outwards from central field, heavily sclerotized, shagreened to toothed. Small central sclerite lightly sclerotized, finely punctate, concave with posterior rim attaching to dorsal posterior point of dorsal phallomere, anterior end folded posteriorly into wide punctate cup. Right ventral phallomere arises from articulation to form smooth rounded lobe becoming sclerotized and shagreened medially and anteriorly; small shagreened fold in moderate gap followed by wide dorsally curving shagreened lip with central convexity. Folded anterior portion of left phallomere setose, otherwise unmodified. Genital hook with long extension to pointed head; short hook with slight dimple near point; arm smoothly curved and relatively short. See Fig. 139.
</paragraph>
<caption pageId="104" pageNumber="105">
<paragraph pageId="104" pageNumber="105">
Figure 138.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
a dorsal habitus b ventral habitus c pronotum d head.
</paragraph>
</caption>
<caption pageId="104" pageNumber="105">
<paragraph pageId="104" pageNumber="105">
Figure 139.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
, genitalia: a) right dorsal phallomere b right ventral phallomere c small central sclerite d genital hook. Arrow(s) indicate diagnostic characters (see text).
</paragraph>
</caption>
<caption pageId="104" pageNumber="105">
<paragraph pageId="104" pageNumber="105">
Figure 140.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga rothi" order="Blattodea" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="rothi">Arenivaga rothi</taxonomicName>
, distribution.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="habitat and natural history">
<paragraph pageId="104" pageNumber="105">Habitat and natural history.</paragraph>
<paragraph pageId="104" pageNumber="105">All life history elements remain unobserved.</paragraph>
</subSubSection>
</treatment>
</document>