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<document id="B4F6A7C4412CDBC84637960646505B55" ID-DOI="10.3897/subtbiol.49.135303" ID-ZooBank="F3B1718A-795E-4946-ACC1-9585C414053F" ID-publisher-id="135303" URI-arpha="0ADA8BE1-21BD-5DA4-AEFA-DEAB2B8E3C5D" URI-zoobank="http://zoobank.org/F3B1718A-795E-4946-ACC1-9585C414053F" article-type="research-article" checkinTime="1730214910008" checkinUser="pensoft" docAuthor="Watiroyram, Santi &amp; Koompoot, Kamonwan" docDate="2024" docId="DA81B40A1B2754A1820780809F840989" docLanguage="en" docName="Subterranean Biology 49: 185-200" docOrigin="Subterranean Biology 49" docSource="https://subtbiol.pensoft.net/article/135303/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Siamcyclops isanus Watiroyram &amp; Koompoot, 2024, sp. nov." docType="treatment" docUuid="7B285B72-0665-4AE7-8F0F-0F40F5452BDB" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="200" masterDocId="0ADA8BE121BD5DA4AEFADEAB2B8E3C5D" masterDocTitle="Siamcyclops isanus sp. nov. (Copepoda, Cyclopoida, Cyclopidae), a new cave-dwelling species from northeastern Thailand" masterLastPageNumber="200" masterPageNumber="185" pageNumber="185" updateTime="1730215272374" updateUser="ExternalLinkService">
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<mods:title id="A923A51DBDDA96212D16C0BBF2F3F330">Siamcyclops isanus sp. nov. (Copepoda, Cyclopoida, Cyclopidae), a new cave-dwelling species from northeastern Thailand</mods:title>
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<mods:namePart id="A300E103C8938E1EEF6A2F9386C1C3C7">Watiroyram, Santi</mods:namePart>
<mods:nameIdentifier id="6D9DFE7324A57F0BC743B5A047499365" type="ORCID">0000-0003-4920-3979</mods:nameIdentifier>
<mods:affiliation id="BAF6CFE3C6ABF65BC4659500C96EEEBC">Division of Biology, Faculty of Science, Nakhon Phanom University, Nakhon Phanom 48000, Thailand</mods:affiliation>
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<mods:namePart id="8367641080409AD76087D9A43E9C7157">Koompoot, Kamonwan</mods:namePart>
<mods:nameIdentifier id="2E6E561630FFA316F9CA444FB79C659C" type="ORCID">https://orcid.org/0009-0005-2116-5191</mods:nameIdentifier>
<mods:affiliation id="C8D55CA0714088368FDE372E726FC761">Diversity of Family Zingibeaceae and Vascular Plant of Its Applications Research Unit, Walai Rukhavej Botanical Research Institute, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand</mods:affiliation>
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<mods:title id="90FB4B3F24B5C46DBA179B9E1A9B19D9">Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part id="C93C23B9BE9FDF5D99D22B85C61D6BBF">
<mods:date id="F7CBDE8060637A66BCE6510CBFC8F7C0">2024</mods:date>
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<mods:number id="EEBC689C6809C0AF2071B1E1CC16C7F4">2024-10-29</mods:number>
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<subSubSection id="1F3188D8D8129EDAEAABD6228DC641A1" type="nomenclature">
<paragraph id="3FA8DFD396A8BDD23B9A93B50FE9CF79">
<taxonomicName id="E4F5B992817314257C664FAF2755E4D2" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="A86D59D8F9CF6E5679751845FF8B49F8" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="126394BD03D13F96DABEC60049C4C41F" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="73E5B522EA208AAAFD4E458E1823C86E" type="description">
<paragraph id="6297D607D20C6F450EB9710D40ABA730">
<figureCitation id="C67A64DF8C61C19CE39F07C6DDA83ED1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">Figs 2</figureCitation>
,
<figureCitation id="A1B35DCD56B4A6461D8963E31EE1B010" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3</figureCitation>
,
<figureCitation id="7CCD0FC8119C99997E65F0D9F73A3919" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4</figureCitation>
,
<figureCitation id="22947C4DD29B6C156131EA24A3BBA135" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5</figureCitation>
,
<figureCitation id="55064AEC4E6DB7323437EB6176B9988B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siamcyclops isanus sp. nov., allotype male A P 3 B P 4." figureDoi="10.3897/subtbiol.49.135303.figure6" httpUri="https://binary.pensoft.net/fig/1163358">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWLAC" type="type locality">
<paragraph id="358A4A24BDFA2165C1A46B08D0A21903">
<heading id="01C07514008F2B3D8E8A4A7B38C669AF" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="25E1BDD4A45E19E6BABE01E43DA43F7E">
The Maholarn cave, Nonghin Subdistrict, Nonghin District,
<collectingRegion id="5C75F298710996B4EBEDC1AB8570465C" country="Thailand" name="Loei">Loei Province</collectingRegion>
, northeastern
<collectingCountry id="0793E1BC289FF2683C476F61E30FBA41" name="Thailand">Thailand</collectingCountry>
:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6LAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.880150,17.106400]}">
<geoCoordinate id="3752C3C0E4A1CE9E6FFD2E999DF2FD22" degrees="17" direction="north" minutes="06" orientation="latitude" precision="1" seconds="23.04" value="17.1064">17 ° 06 ' 23.04 &quot; N</geoCoordinate>
,
<geoCoordinate id="40403D5021D7422753AC605C9CAD5C87" degrees="101" direction="east" minutes="52" orientation="longitude" precision="1" seconds="48.54" value="101.88015">101 ° 52 ' 48.54 &quot; E</geoCoordinate>
</named-content>
,
<quantity id="813BF5A08A7DA2F44F8A59E0E72A82F7" metricMagnitude="2" metricUnit="m" metricValue="3.15" unit="m" value="315.0">315 m</quantity>
altitude. Sample collected from a rimstone pool in the innermost zone, filled with dripping water.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEMAC" type="etymology">
<paragraph id="D57B93BC90191BED59D15E14C0E50292">
<heading id="9E6E504F22BC8C47083D47E5ED77AA87" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="16BA5043F0ED16DD25DD3D5591E5A37D">
<taxonomicName id="7D4C85EC9C5BD8AC50873CAA87AEEBD2" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="C47D766B0FF58C55A65D54DE1EBFAB2B" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFFAF0274A4B51F77A8DF17852E68FCF" rank="species">sp. nov.</taxonomicNameLabel>
is named after Isan, the local name for northeastern
<collectingCountry id="402E19C0750A5CE44AFA1DAB95371424" name="Thailand">Thailand</collectingCountry>
. The species epithet is a masculine, singular adjective.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVMAC" type="Type specimens">
<paragraph id="4DCD3E3B8A8D0BAC02D5FCF9DF7EF56B">
<heading id="50993B992730124F485E8FEE9237B43D" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="51EE5068A9391EE8E4DB4925F73B8E76">
<emphasis id="CE9E2171C1D860A05447DA32AC03E682" bold="true" italics="true">
<typeStatus id="8923C1A66B73968C3AC879452C36C391" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="BE67214D80A42941265B539B008E6E3A" count="1" type="adult">one adult</specimenCount>
female dissected and mounted on one slide, THNHM-IV- 20465.
<emphasis id="9C5568EC6F3F7D38DB4C2D0EAC3783B8" bold="true" italics="true">
<typeStatus id="8BDB8CD76AB174EEDEA445351AEA66EC" type="allotype">Allotype</typeStatus>
</emphasis>
<specimenCount id="AD34A79ECBC10E69A7D19BB44073D952" count="1" type="adult">one adult</specimenCount>
male dissected and mounted on one slide, THNHM-IV- 20466.
<emphasis id="A2EB3B7A1E510A27D4651EB8E90172D7" bold="true" italics="true">
<typeStatus id="0E29A3D3A84FC20F36AACFE5159163EF" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="1F037E14067C06EBF037C96772089BEF" count="1" type="adult">one adult</specimenCount>
female and male dissected and mounted on two separate slides, (NPU
<date id="E8504319047E539885C7AD1C914F1EB4" value="2024-02-01">2024-01 02</date>
) •
<specimenCount id="3BE1B92BFADDEE34BF669F93BBA0FFD0" count="4" type="adult">four adult</specimenCount>
females and
<specimenCount id="67C81247ABEEB14E188629E69081F349" count="9" type="adult">nine adult</specimenCount>
males stored in a 1.5 ml microtube with 70 % ethanol, (NPU
<date id="8A98EBCBF1548F53E870FC210F69E8EC" value="2024-02-01">2024-01 02</date>
) •
<specimenCount id="DC493922C8FF14E8E61348275D6A12D3" count="1" type="adult">one adult</specimenCount>
female and male dissected and mounted on two separate slides, NPU 2020-003 •
<specimenCount id="CA831A15AC01D0EC5F6A55654D29136A" count="1" type="adult">one adult</specimenCount>
male dissected and mounted on one slide, THNHM-IV- 20467. All samples were collected by the first author on
<date id="879B89473AEF1E14A9CB5D6EB7F10A99" value="2013-07-05">5 July 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGNAC" type="description">
<paragraph id="CB159B84F14E3359B358EA329ACC6A55">
<heading id="CDB99FD1B1FB66BDA51B71D9F3F91053" reason="title">Description of adult female</heading>
</paragraph>
<paragraph id="F8DED1B0CDA1D5D5D819216A3D68A846">
<emphasis id="9023020E6E7D441D89B81668BEE12325" bold="true">
(
<typeStatus id="9F79124421C86849A52113E2A283C725" type="holotype">holotype</typeStatus>
).
</emphasis>
Habitus (Fig.
<figureCitation id="C13166BBBDC8D4EE34C7FB6CAE6C1383" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 A</figureCitation>
) cyclopiform. Body length, excluding caudal setae, 550 μm, with prosome / urosome ratio of 1.5. Body surface ornamented with refractile points (not figured). Nauplius eye indiscernible. Posterior margin of prosomites with smooth free margin of hyaline fringe. Cephalosome completely fused with first thoracic somite, forming cephalothorax. Urosome length 212 μm; length / width ratio of 1.1. Pseudosegment present between pediger 5 and genital double-somite. Genital double-somite (Fig.
<figureCitation id="00BB11DB18890AD9CC1DDA6B597751C2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 B D</figureCitation>
) enlarged; anterior part slightly wider than posterior part; 1.2 times as wide as long; with pair of sclerotised semi-rounded structures laterally and
<abbrev id="ABBRID0EWNAC" xlink_title="swimming legs 16">P 6</abbrev>
dorsolaterally; posterior margin with irregularly serrated free hyaline fringe. Urosomites 34 (Fig.
<figureCitation id="24722715EEFDFA89EB297932241E4C13" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 B D</figureCitation>
) with about 1.6 times as long as wide; posterior margin with irregularly serrated free hyaline fringe. Anal somite (Fig.
<figureCitation id="C48E3B556AEF161FF19D9914635B7C35" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 A D</figureCitation>
) short, 1.6 times as long as wide; with a pair of dorsal sensilla at base of anal operculum. Anal operculum (Fig.
<figureCitation id="E8A33D7120B92BA2B46AFAE17ACC63D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 B, D</figureCitation>
) reaching slightly beyond distal end of anal somite; semicircular; free margin smooth. Caudal rami slightly divergent (Fig.
<figureCitation id="2CF72A1DA3800DBA6983D19F33D6A1BB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 A C</figureCitation>
). Caudal ramus rectangular, about 2.5 times as long as wide, without dorsal longitudinal keel. Anterolateral accessory seta (I) reduced. Lateral seta (
<collectionCode id="92B04CA4075DCCE94A40E8129F0ED559">II</collectionCode>
) pinnate, slightly shorter than 1 / 2 length of caudal ramus, inserted at 1 / 3 length of caudal ramus. Posterolateral seta (
<collectionCode id="D5D01617582EF9B02F962103892E0CDC">III</collectionCode>
) pinnate, strong, spine like, shorter than caudal ramus, with spinules at insertion point on ventrolateral side. Outer terminal seta (
<collectionCode id="B2EAE81CE6A56BB7BE3B11F2AB2DC9B5">IV</collectionCode>
) pinnate, about 2.5 times as long as caudal ramus, without fracture plane. Inner terminal seta (
<collectionCode id="5A02343F0D2A20BA023CA4240B1CFF76" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
) pinnate, about 4.0 times as long as caudal ramus, without fracture plane. Innermost terminal seta (
<collectionCode id="CECA18C78B7E0B46C2161A9BCAA9E6CC" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
) bare; shortest. Dorsal seta (
<collectionCode id="EA9A3014CE79CBA885188B5C25C160EB">VII</collectionCode>
) pinnate, articulated, longer than caudal ramus.
</paragraph>
<caption id="FE8AE356935E04A2B93F8174E5D27749" ID-DOI="10.3897/subtbiol.49.135303.figure2" ID-arpha="31184733-910A-548E-8640-14D95ED971AA" httpUri="https://binary.pensoft.net/fig/1163354" startId="F2">
<paragraph id="7C02C761E6A9231BA4E7E7B79642D815">
<label id="D5B1F19BE0E64D59C642DAAC9B098805">Figure 2.</label>
</paragraph>
<paragraph id="B7FF59ADDD9DB2CDB57E995D9DE3CCD4">
<taxonomicName id="BC823FADBCC152CE8F5DF8208C690395" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="40F9B52EB73A2BC1571E71DD2DA5A348" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DF41695DBE0AE89C552CF7E1957BC0C7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female (
<emphasis id="34BA0CC2C0C886DC340C3181B84EDBFD" bold="true">A, B</emphasis>
) and paratype female (
<emphasis id="69E61CFD569DB76A0CB0452CEA03FCA8" bold="true">C, D</emphasis>
)
<emphasis id="EC23314A221E962C320BA69C9012F87C" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="8A2FE7477124CAF277BD297A861DEB94" bold="true">B</emphasis>
urosome with a pair of egg sacs, dorsal view
<emphasis id="BCD25AF663530AB0387ABA2F412356C5" bold="true">C</emphasis>
urosome with spermatophore, ventral view
<emphasis id="7BC4FFCD40449190A11C1D963E3AD37D" bold="true">C</emphasis>
urosome with spermatophore, lateral view.
</paragraph>
</caption>
<paragraph id="0BB9D55E9B1CD472091FFE6D38769D86">
Antennule (Fig.
<figureCitation id="4BB6823E0E8AC961ADE26FDB0B9DE741" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 A</figureCitation>
). 11 - segmented, not reaching posterior margin of cephalothorax. Armature formula as follows: 7.2. 5.2. 0 + S. 2.3. 2 + A. 2.2 + A. 7 + A. Aesthetascs cylindrical.
</paragraph>
<caption id="98FFD7C922C5556DC795EF54DB08DEF7" ID-DOI="10.3897/subtbiol.49.135303.figure3" ID-arpha="D70DA324-E1F2-5293-ABB2-F79D0AAA78B9" httpUri="https://binary.pensoft.net/fig/1163355" startId="F3">
<paragraph id="71AE92D3133CB012C7384B346A7950A5">
<label id="908848918B50838F89EEB2DE5DCAE67F">Figure 3.</label>
</paragraph>
<paragraph id="10CC1F491DD836543328278C74B04171">
<taxonomicName id="61EEEE02DC6F1FA4E05ED9AD3215A71A" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="7B8B7081217F07948533FCB0A18924D2" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50CF370600525B8956FF88C281D18CFD" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female
<emphasis id="78A15884DA36FB524C4FAB495770CFED" bold="true">A</emphasis>
antennule
<emphasis id="2AEF4F0199F85DC97E834805411CEB29" bold="true">B</emphasis>
antenna
<emphasis id="74D7FD4159D77F7023FCB4588EE470B7" bold="true">C</emphasis>
mandible
<emphasis id="D2366A54F7F79212D53888AE998DDA01" bold="true">D</emphasis>
maxillule
<emphasis id="F0CF4C60A3576C3EE821B25FBD428719" bold="true">E</emphasis>
maxilla
<emphasis id="656C5D9D7B233D536DD541EC1A0D253C" bold="true">F</emphasis>
maxilliped.
</paragraph>
</caption>
<paragraph id="B32E9AA1D05200872E2006DA778A6977">
Antenna (Fig.
<figureCitation id="6B1BD4E929864D8E59A91D7D38937178" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 B</figureCitation>
). 4 - segmented; coxobasis smooth, with seta on distal inner corner.
<abbrev id="ABBRID0E6AAE" xlink_title="endopod">Enp</abbrev>
13 with small spinular rows on outer margin,
<abbrev id="ABBRID0EDBAE" xlink_title="endopod">Enp</abbrev>
- 1 with seta on distomedial margin;
<abbrev id="ABBRID0EHBAE" xlink_title="endopod">Enp</abbrev>
- 2 with medial seta and four distal outer setae;
<abbrev id="ABBRID0ELBAE" xlink_title="endopod">Enp</abbrev>
- 3 with seven apical setae, two of them geniculated. All setae smooth.
</paragraph>
<paragraph id="57BD4B1A4C082F5FA97D1841A125D10D">
Mandible (Fig.
<figureCitation id="8C60DBE5E6366AA4E6BABB54625ABC08" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 C</figureCitation>
). Gnathobase with eight strongly chitinized teeth, and a small seta dorsally. Basis very reduced, with two long bipinnate setae and a short smooth seta, representing
<abbrev id="ABBRID0EVBAE" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0EZBAE" xlink_title="endopod">Enp</abbrev>
.
</paragraph>
<paragraph id="70D2EAC6672E14D8E504FCF6F05D5992">
Maxillule (Fig.
<figureCitation id="BB991EA7EEA3D5223918300CADF1E804" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 D</figureCitation>
). Precoxal arthrite with strongly chitinized teeth, one tricuspidate and two bicuspidate; four strong smooth setae and one spinulate seta on inner margin. Coxobasis with spinulate seta and two smooth setae distally.
<abbrev id="ABBRID0EDCAE" xlink_title="exopod">Exp</abbrev>
completely reduced, represented by a smooth seta.
<abbrev id="ABBRID0EHCAE" xlink_title="endopod">Enp</abbrev>
with three smooth setae.
</paragraph>
<paragraph id="E016202F7966851C64C3A396A0C21FCA">
Maxilla (Fig.
<figureCitation id="B3201FD40BC337E75A37100F245646E1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 E</figureCitation>
). Precoxal endite with two pinnate setae. Coxa with two endites: proximal endite with smooth seta; distal endite with smooth seta and pinnate seta. Basis drawn out into claw-like expansions, with slender seta on proximal inner margin. Two-segmented
<abbrev id="ABBRID0ERCAE" xlink_title="endopod">Enp</abbrev>
:
<abbrev id="ABBRID0EVCAE" xlink_title="endopod">Enp</abbrev>
- 1 with two smooth setae;
<abbrev id="ABBRID0EZCAE" xlink_title="endopod">Enp</abbrev>
- 2 with three smooth setae.
</paragraph>
<paragraph id="956120EC3E932DDF1A96D6F6B4E154B1">
Maxilliped (Fig.
<figureCitation id="3078BA397C3D9FDA4B7A4B453E388F53" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 F</figureCitation>
). Syncoxa with two spinulate setae on distal inner margin. Basis with two spinulate setae on distal inner margin, accompanied by two spinule rows on its surface.
<abbrev id="ABBRID0EDDAE" xlink_title="endopod">Enp</abbrev>
- 1 with inner seta;
<abbrev id="ABBRID0EHDAE" xlink_title="endopod">Enp</abbrev>
- 2 with three apical setae; all setae smooth.
</paragraph>
<paragraph id="61DC3974DC2D684BC5EBAF978913114D">
<abbrev id="ABBRID0ENDAE" xlink_title="swimming legs 16">P 1</abbrev>
P 4 (Fig.
<figureCitation id="DD946009A383A5549BCFC3580FBCDAD4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 A D</figureCitation>
).
<abbrev id="ABBRID0EVDAE" xlink_title="swimming legs 16">P 1</abbrev>
P 4 with 2 - segmented
<abbrev id="ABBRID0EZDAE" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0E4DAE" xlink_title="endopod">Enp</abbrev>
.
<abbrev id="ABBRID0EBEAE" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0EFEAE" xlink_title="endopod">Enp</abbrev>
ornamented with setules on its inner and outer margin, respectively. No blunt seta on
<abbrev id="ABBRID0EJEAE" xlink_title="exopod">Exp</abbrev>
- 2, all setae being equal. Armature formula (setae in Arabic numerals and spines in Roman numerals from outer-inner or outer-apical-inner margins) as follows:
</paragraph>
<paragraph id="E346CDD6BCC27E6C6E81B601DB74F171">
<table id="TID0EJGAG" rules="all">
<tbody id="FA825CBC2CC768971ED45EA0D8B5B0E0">
<table-wrap id="T1" orientation="portrait" position="float">
<tr id="433994E510242BF90A58D725A45BE547">
<td id="5DF3A2147F9A6E8DE2AA4381EBEB3583" colspan="1" isEmpty="true" rowspan="1">-</td>
<emphasis id="FA239218DCD5DEB06E8EED2DE2BDC301" bold="true">
<td id="88B7379075A684804709B67CF372BD44" colspan="1" rowspan="1">Coxa</td>
<td id="B509B1B338F6EDBE8B9DAF3266357C96" colspan="1" rowspan="1">Basis</td>
<td id="3D9935FEF9D0A2158803888D11E65333" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5TAE" xlink_title="exopod">Exp</abbrev>
</td>
<td id="C435E0B233AB26DCE4B8ECBE56FC58F8" colspan="1" rowspan="1">
<abbrev id="ABBRID0EHUAE" xlink_title="endopod">Enp</abbrev>
</td>
</emphasis>
</tr>
<tr id="06CAFED63C9C64D92DA94F82017ACB88">
<td id="D3AC516F5E0C7EC7CD17A5DE13F6F30A" colspan="1" rowspan="1">
<abbrev id="ABBRID0EQUAE" xlink_title="swimming legs 16">P 1</abbrev>
</td>
<td id="A001F817704DC71C8B42551A9AF4A0C0" colspan="1" rowspan="1">0-1</td>
<td id="4951D5E3A97B4460FE0D635ACC0B8EC1" colspan="1" rowspan="1">1 - I</td>
<td id="B86965E9F066E22A6D420E635CD675B5" colspan="1" rowspan="1">I- 0; III- 2-3</td>
<td id="75BC041B48ABB6B57FC3B1BCDC8AFDF5" colspan="1" rowspan="1">0-1; 1 - I- 2</td>
</tr>
<tr id="7BEE8DAA53C2B380F22DD796EF6BF2B6">
<td id="C69CAFD93E4AF30104753C6EE3B1BB08" colspan="1" rowspan="1">P 2</td>
<td id="DFA88FE0C0DCDB0AE68BC204C88CBB23" colspan="1" rowspan="1">0-1</td>
<td id="F393AEC70BA794159598BB06C903DD49" colspan="1" rowspan="1">1-0</td>
<td id="DEE42306D590E79FE5323CF90B3E9650" colspan="1" rowspan="1">I- 0; III- 2-3</td>
<td id="5F8C8E71397AD13B17914AF6BEFEB104" colspan="1" rowspan="1">0-1; 1 - I + 1-2</td>
</tr>
<tr id="63E4B44CFA36A328899D178E1C97D071">
<td id="FE103783420C3A72CE57217135D4D057" colspan="1" rowspan="1">P 3</td>
<td id="6DD0837587344C0AE89FB88B3B2F7EBE" colspan="1" rowspan="1">0-1</td>
<td id="FC91085A1BE1433AAEE66CAAE48D834C" colspan="1" rowspan="1">1-0</td>
<td id="67D82D3C6B4042A011D341F7F7C349E1" colspan="1" rowspan="1">I- 0; III- 2-3</td>
<td id="A04F53A3448908694E172404931D654E" colspan="1" rowspan="1">0-1; 1 - I + 1-3</td>
</tr>
<tr id="4A69E681D664094DCED53E8E34D9CD67">
<td id="1E19FC0394B254D5E6BDF922F3080504" colspan="1" rowspan="1">P 4</td>
<td id="CCE0E946BFDB9D14C51D502ADC7309C4" colspan="1" rowspan="1">0-0</td>
<td id="02E8E33F5762390432E792C2E77ECBC1" colspan="1" rowspan="1">1-0</td>
<td id="266CA3EBEE170D12BD0236AE8431ED68" colspan="1" rowspan="1">I- 0; II- 2-2</td>
<td id="F58C7880D8B7B7BC9C0E2A760A9879C7" colspan="1" rowspan="1">0-1; 0 - I + 1-2</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<caption id="B7675EC394969B9FA7DB5D076D62A6BD" ID-DOI="10.3897/subtbiol.49.135303.figure4" ID-arpha="BD29152B-C408-51D1-8C40-F07CFDE9D55B" httpUri="https://binary.pensoft.net/fig/1163356" startId="F4">
<paragraph id="8FA789B672D255A276AAE7848E8D692D">
<label id="73D6B3E84DD5B2353C8EFBF713912E6E">Figure 4.</label>
</paragraph>
<paragraph id="99A35A4DD3B07AC9834AFE6AEBB0DA3F">
<taxonomicName id="9962C0F3F1BAF6E925D055E982A7E252" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="518A5AC42FB9A15B4CC855A35A023642" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74DFEB7B8EDB1DEAAA9C48DC0AC21523" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female
<emphasis id="0123F2995FC29CA62D41DD84BBFD5575" bold="true">A</emphasis>
<abbrev id="ABBRID0ECFAE" xlink_title="swimming legs 16">P 1</abbrev>
<emphasis id="9861595B18114634D6294847E57F6399" bold="true">B</emphasis>
P 2
<emphasis id="D945BF0B82CC2A7F5166302EE6F12057" bold="true">C</emphasis>
P 3
<emphasis id="577ACD030BA86DC940C84B95867E21FC" bold="true">D</emphasis>
P 4
<emphasis id="B2ECE3037C5EA500960ACF867247F3A4" bold="true">E</emphasis>
P 5.
</paragraph>
</caption>
<paragraph id="A40FD330394EB098262FB81E17221287">
<abbrev id="ABBRID0EVFAE" xlink_title="swimming legs 16">P 1</abbrev>
(Fig.
<figureCitation id="5C54015F62BD660936ECBE399A3B7FB9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 A</figureCitation>
). Basis with small spinules near its insertion.
<abbrev id="ABBRID0E4FAE" xlink_title="exopod">Exp</abbrev>
slightly larger than
<abbrev id="ABBRID0EBGAE" xlink_title="endopod">Enp</abbrev>
.
<abbrev id="ABBRID0EFGAE" xlink_title="exopod">Exp</abbrev>
- 1 much smaller than
<abbrev id="ABBRID0EJGAE" xlink_title="exopod">Exp</abbrev>
- 2.
<abbrev id="ABBRID0ENGAE" xlink_title="exopod">Exp</abbrev>
- 2 about twice as long as wide.
<abbrev id="ABBRID0ERGAE" xlink_title="endopod">Enp</abbrev>
- 2 larger than
<abbrev id="ABBRID0EVGAE" xlink_title="endopod">Enp</abbrev>
- 1, about 2.0 times.
<abbrev id="ABBRID0EZGAE" xlink_title="endopod">Enp</abbrev>
- 2 with robust apical seta, as long as
<abbrev id="ABBRID0E4GAE" xlink_title="endopod">Enp</abbrev>
- 1 and
<abbrev id="ABBRID0EBHAE" xlink_title="endopod">Enp</abbrev>
- 2 combined, its tip bent outward; distal inner seta as long as apical spine, the two remaining setae shorter.
</paragraph>
<paragraph id="DFA890D09E653EAFDA370E08F26EAB78">
P 2 (Fig.
<figureCitation id="052F8246AB5A5581F14339C494CD4C1E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 B</figureCitation>
). Similar to
<abbrev id="ABBRID0ELHAE" xlink_title="swimming legs 16">P 1</abbrev>
but lateral seta of basis shorter.
<abbrev id="ABBRID0EPHAE" xlink_title="endopod">Enp</abbrev>
- 2 with strong spinulated apical spine, with strengthened tip; two setae as long as apical spine but two remaining setae shorter.
</paragraph>
<paragraph id="2EF0EB0A0CBF4DB150D04781E31F4657">
P 3 (Fig.
<figureCitation id="495E787901FAE17C0176152A70EDF8E6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 C</figureCitation>
). Similar to P 2 but lateral seta on basis longer,
<abbrev id="ABBRID0EZHAE" xlink_title="endopod">Enp</abbrev>
- 2 with slightly less robust apical spine.
<abbrev id="ABBRID0E4HAE" xlink_title="endopod">Enp</abbrev>
- 2 with two setae longer than apical spine, three remaining setae shorter.
</paragraph>
<paragraph id="E90BC30CF4DA5E9AD5D90FDF28A7E6B0">
P 4 (Fig.
<figureCitation id="E99BB9DB0FB39E81CB169BB1B8AAB3BB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 D</figureCitation>
).
<abbrev id="ABBRID0EHIAE" xlink_title="exopod">Exp</abbrev>
- 1 and
<abbrev id="ABBRID0ELIAE" xlink_title="exopod">Exp</abbrev>
- 2 with weakly spinulated outer spines.
<abbrev id="ABBRID0EPIAE" xlink_title="exopod">Exp</abbrev>
- 2 twice as long as wide; three inner setae longer than
<abbrev id="ABBRID0ETIAE" xlink_title="exopod">Exp</abbrev>
- 1 and 2 combined, outermost seta shortest.
<abbrev id="ABBRID0EXIAE" xlink_title="endopod">Enp</abbrev>
- 1 slightly shorter than wide,
<abbrev id="ABBRID0E2IAE" xlink_title="endopod">Enp</abbrev>
- 2 about 1.3 times as long as wide.
<abbrev id="ABBRID0E6IAE" xlink_title="endopod">Enp</abbrev>
- 2 with slender apical spine, short as long as segment bearing it; outer and innermost seta shorter than apical spine, two remaining setae longer than apical spine.
</paragraph>
<paragraph id="C3B09628D74035398C784504C290CCAE">
P 5 (Fig.
<figureCitation id="4FBB7ABAB506B43C50DD6882F56F9DB5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Siamcyclops isanus sp. nov., holotype female A P 1 B P 2 C P 3 D P 4 E P 5." figureDoi="10.3897/subtbiol.49.135303.figure4" httpUri="https://binary.pensoft.net/fig/1163356">4 E</figureCitation>
). Completely fused to pediger 5, proximal segment represented by lateral seta, and distal segment retains two pinnate apical setae on a small lobe. Lateral seta slightly longer than apical inner seta; apical inner seta about 1.3 times as long as apical outer one.
</paragraph>
<paragraph id="9AA6A913A1745FF28AAE7FD24E97E8E8">
<abbrev id="ABBRID0ELJAE" xlink_title="swimming legs 16">P 6</abbrev>
(Fig.
<figureCitation id="596F5A1BA24A8C4CDFBDB884FDF287A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Siamcyclops isanus sp. nov., holotype female (A, B) and paratype female (C, D) A habitus, dorsal view B urosome with a pair of egg sacs, dorsal view C urosome with spermatophore, ventral view C urosome with spermatophore, lateral view." figureDoi="10.3897/subtbiol.49.135303.figure2" httpUri="https://binary.pensoft.net/fig/1163354">2 D</figureCitation>
). Reduced to a semi-circular plate, with three short elements: dorsal seta thin and short, two ventrally positioned elements spiniform.
</paragraph>
<paragraph id="432D89C077B9B92209942133D66EB71D">
Egg sac (Fig.
<figureCitation id="8E2B724A1CB310F1DDE9268744EA4F7E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 A, B</figureCitation>
). Each sac with two large eggs; each egg about 70 µm in diameter.
</paragraph>
<paragraph id="A4D4BC66CC2B50BD8AA0D8C976D9BBFA">
Spermatophore (Fig.
<figureCitation id="E2DF83A688C5F1D4EB0B073EB45B6C37" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Siamcyclops isanus sp. nov., holotype female A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/subtbiol.49.135303.figure3" httpUri="https://binary.pensoft.net/fig/1163355">3 C, D</figureCitation>
). Three-dimensional C-shaped.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6JAE" type="description">
<paragraph id="5CEBBC3498690C8B121157348951CE54">
<heading id="851A3A13534AA15A97E39C2FA10A2797" reason="title">Description of adult male</heading>
</paragraph>
<paragraph id="35B73B2EEA6C31191308D1F5E19934D0">
<emphasis id="8BB7A220AB40D40AD27BB2EAEA414A5C" bold="true">
(
<typeStatus id="52D51F6DB7AC6BEA75D6EB04DA4F2D43" type="allotype">allotype</typeStatus>
).
</emphasis>
Body length (Fig.
<figureCitation id="B532888832711196A287498B2CAAB4D0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 A</figureCitation>
), excluding caudal rami, 540 µm; smaller than female. General segmentation and ornamentation (Fig.
<figureCitation id="C40160B94203CCB0672CB4A0F9F7D296" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 A C</figureCitation>
) similar to female except genital somite and antennules. Genital somite (Fig.
<figureCitation id="4E5EFC6FB85A9144C0F5FCCDFAAA5803" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 B, C</figureCitation>
) not fused with urosomite 3 like in female, enlarged laterally; 1.3 times as wide as long; with pair of sclerotised rounded structures dorso-laterally; posterior margin with irregularly serrated free hyaline fringe. Urosomites 35 (Fig.
<figureCitation id="6207B084B778D328F727D9FAEF91CF37" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 A C</figureCitation>
) with about 1.5 times as long as wide; posterior margin with irregularly serrated free hyaline fringe.
</paragraph>
<caption id="1F99C0170AE70A31824373A8D3153D1B" ID-DOI="10.3897/subtbiol.49.135303.figure5" ID-arpha="C8B925AE-BFAF-5045-BA27-19FD6D89538A" httpUri="https://binary.pensoft.net/fig/1163357" startId="F5">
<paragraph id="356B8167C27AC6797E9B48543A1D1AC3">
<label id="6BE8AC913531248D5878EC38153A0DC5">Figure 5.</label>
</paragraph>
<paragraph id="AC77D854F92B5A54BE51B994FD3CB7E5">
<taxonomicName id="D143D027662024BE05A550BAFD4976EF" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="C0BE4F1C1F3070A4E2F094D7622BA03C" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B918A2B09B6BDD610E68553DC372244E" rank="species">sp. nov.</taxonomicNameLabel>
, allotype male
<emphasis id="65789C0F93678599FD1E20DF49F6A224" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="58805B6DD087E3A7527F17C6F7C37C59" bold="true">B</emphasis>
urosome, ventral view
<emphasis id="586AC86C1BF0FD8BEE7816351F837955" bold="true">C</emphasis>
urosome with urosomite 1, lateral view
<emphasis id="BDB0927DD70D0090DA792B6CEA40046A" bold="true">D</emphasis>
antennule.
</paragraph>
</caption>
<paragraph id="45D16D895AB40C26FB1EF2EA072F67E0">
Antennule (Fig.
<figureCitation id="9676145035F2BB1A10142336A7F45EA3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 D</figureCitation>
). 15 - segmented, geniculate. Armature formula as follows: 6 + 3 A. 4.2. 2 + A. 1.2.2.2 + A. 0.1. 1 + S. 1 + A. 0.1. 8 + 2 A.
</paragraph>
<paragraph id="990B4534DD62C1D865B96BFE7BA7B649">
Antenna, mouthparts,
<abbrev id="ABBRID0E6LAE" xlink_title="swimming legs 16">P 1</abbrev>
, P 2 and P 5 similar to female.
</paragraph>
<paragraph id="14AD93AF120ACA37CA6C148BE707BC77">
P 3 (Fig.
<figureCitation id="C01FB136A13671E10B3D00F969964640" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siamcyclops isanus sp. nov., allotype male A P 3 B P 4." figureDoi="10.3897/subtbiol.49.135303.figure6" httpUri="https://binary.pensoft.net/fig/1163358">6 A</figureCitation>
).
<abbrev id="ABBRID0EJMAE" xlink_title="exopod">Exp</abbrev>
similar to female.
<abbrev id="ABBRID0ENMAE" xlink_title="endopod">Enp</abbrev>
- 2 shorter than wide,
<abbrev id="ABBRID0ERMAE" xlink_title="endopod">Enp</abbrev>
- 2 about 1.5 times as long as wide.
<abbrev id="ABBRID0EVMAE" xlink_title="endopod">Enp</abbrev>
- 2 with transformed spine and smooth seta apically: transformed spine bent inward, with round tip bent outward, serrated by two rows of spinules on distal half of it. Inner subterminal seta transformed to spine-like seta, strong, bare, with blunt tip.
</paragraph>
<caption id="945B8935A69942DEE0B9B452A478A423" ID-DOI="10.3897/subtbiol.49.135303.figure6" ID-arpha="9FE8FB3D-08C2-5C23-B6E4-72B440E12BF2" httpUri="https://binary.pensoft.net/fig/1163358" startId="F6">
<paragraph id="EEC412F827130A00262751B0A0622C1D">
<label id="86D1D4835EF4C9120C64F6D78A07085C">Figure 6.</label>
</paragraph>
<paragraph id="30AD13F88F2950B88360A3751DDCA4D4">
<taxonomicName id="497A3B49CA96B9629E23944E8E7BD2E7" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="6E6C573CDE1551CEEE793840CAAFC13C" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C3E7BEB5A45C2BA20F91C1BDDC6ACED" rank="species">sp. nov.</taxonomicNameLabel>
, allotype male
<emphasis id="BE6367FC88D9E91227453229C307F826" bold="true">A</emphasis>
P 3
<emphasis id="0A6DC52C6D8C1967FE579D8F1F469F78" bold="true">B</emphasis>
P 4.
</paragraph>
</caption>
<paragraph id="7BF754A434B0AD310EA990F76815E68C">
P 4 (Fig.
<figureCitation id="E137E15601015055F45CE021F42F2ACD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siamcyclops isanus sp. nov., allotype male A P 3 B P 4." figureDoi="10.3897/subtbiol.49.135303.figure6" httpUri="https://binary.pensoft.net/fig/1163358">6 B</figureCitation>
).
<abbrev id="ABBRID0E2NAE" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0E6NAE" xlink_title="endopod">Enp</abbrev>
similar to female but
<abbrev id="ABBRID0EDOAE" xlink_title="endopod">Enp</abbrev>
with slightly longer, about 1.5 times as long as wide.
</paragraph>
<paragraph id="DEA15109BA946FFD662C6D5D8E9536F5">
<abbrev id="ABBRID0EJOAE" xlink_title="swimming legs 16">P 6</abbrev>
(Fig.
<figureCitation id="7829473C3FAAE35AD6636311000BAB3F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siamcyclops isanus sp. nov., allotype male A habitus, dorsal view B urosome, ventral view C urosome with urosomite 1, lateral view D antennule." figureDoi="10.3897/subtbiol.49.135303.figure5" httpUri="https://binary.pensoft.net/fig/1163357">5 B, C</figureCitation>
). Reduced to simple plate, represented by three subequal pinnate setae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EROAE" type="variability">
<paragraph id="8335BD05EB5371D02E45B0D64C3C1815">
<heading id="61EA61C64DD172E046F1E9C45CEACFB8" reason="title">Variability.</heading>
</paragraph>
<paragraph id="11680C2DE2565B5C05D3418F7C5A0134">
One of female (out of five) has
<abbrev id="ABBRID0EXOAE" xlink_title="swimming legs 16">P 6</abbrev>
with two elements.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2OAE" type="distribution">
<paragraph id="8FB238B43F9E149244AEDD837DCFD762">
<heading id="6508178A0310CCE9576DADAE92A22DD0" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="128F302BA2E084D3347BE1CE4B669E03">
Known only from the
<typeStatus id="0CA79B025F26E721251934150598F7A2">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAPAE" type="Differential diagnosis">
<paragraph id="4B2161D16D7B0C3257D1954BAB22F6AA">
<heading id="BAF51ED3C03A0AE36A6F8B30121D54DA" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="F3B065F84B195790E37157B0EF2CD5C9">
The new species is similar to
<taxonomicName id="15FADF7B24B33E3583644011B0A2F339" authority="Boonyanusith, Sanoamuang &amp; Brnacelj, 2018" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="CEAD10CE5423C531619E17EEAF89EEAF" italics="true">S. cavernicolus</emphasis>
Boonyanusith, Sanoamuang &amp; Brnacelj,
<quantity id="2F49B7E73AFF3A63D3918B51CC88E02A" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" unit="in" value="2018.0">2018 in</quantity>
</taxonomicName>
the following characteristics: 1) mandibular palp with three setae; 2)
<abbrev id="ABBRID0ERPAE" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0EVPAE" xlink_title="endopod">Enp</abbrev>
of
<abbrev id="ABBRID0EZPAE" xlink_title="swimming legs 16">P 1</abbrev>
P 4 are subequal; 3) spine formula of
<abbrev id="ABBRID0E4PAE" xlink_title="swimming legs 16">P 1</abbrev>
P 4
<abbrev id="ABBRID0EBQAE" xlink_title="exopod">Exp</abbrev>
- 2 is 3.3.3.2; 4) P 4 lacks coxal seta; 5) P 4 intercoxal sclerite with round distal margin; 6) P 5 fused to pediger 5, remaining small unsegmented lobe; 7)
<abbrev id="ABBRID0EFQAE" xlink_title="swimming legs 16">P 6</abbrev>
with three elements (see Table
<tableCitation id="176ED4BA78570CF3AECE8DF562B9EA7D" captionStart="Table 1" captionStartId="T2" captionText="Table 1. Morphological differences between the genus Siamcyclops and the closely related genera." httpUri="http://table.plazi.org/id/A39350D05EA0790BCCFB1E133D019671" tableUuid="A39350D05EA0790BCCFB1E133D019671">1</tableCitation>
). However, the new species differs from
<taxonomicName id="A25A63BF855E97EEAC23B56480EED9F2" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="2F5E7A65F0F713F2CF7378BC61378A1A" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
in: 1) anal operculum smooth and unproduced over segment bearing it (serrated free margin and well-developed in
<taxonomicName id="D0F049FE36DBC2BF6FA293901AF1EA3F" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="DE716CA04271CA2038C453790C6756DF" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
); 2)
<abbrev id="ABBRID0EDRAE" xlink_title="swimming legs 16">P 1</abbrev>
<abbrev id="ABBRID0EHRAE" xlink_title="endopod">Enp</abbrev>
- 2 with four elements (five elements in
<taxonomicName id="BB48A3E4DAD6E9BC49F43F16FBF795CB" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="9FFC2502411CB8103B26CC3BE6CFE2B3" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
); 3) P 3
<abbrev id="ABBRID0EWRAE" xlink_title="endopod">Enp</abbrev>
- 2 with six elements in male (five elements in
<taxonomicName id="C0A20377F3A40819169A5E241E938757" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="F7D35807BAB08EF882EFF46F44AF4426" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
); 4) P 4
<abbrev id="ABBRID0EFSAE" xlink_title="exopod">Exp</abbrev>
- 2 with four setae (five setae in
<taxonomicName id="41B9308371BB5C0D0E6E2320F048DBAF" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="540E7536CC010F12125EC9498BDE358C" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
); 5) spermatophore is three-dimensional C-shaped (L-shaped in
<taxonomicName id="8A37DF92B6CC24BBDF89056D9BAF48E4" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="49A399F46AD5CE48B0864947E5D88241" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="2CD750B5B1677436E4BBD8886DE2E66C">
The SEM photos of P 5 show the degree of reduction in
<taxonomicName id="7DAF517F54DA8213D8D786634106DD4C" authorityName="Boonyanusith, Sanoamuang &amp; Brancelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="genus">
<emphasis id="568CF7D99682E514CE552FC0CFC9B4C5" italics="true">Siamcyclops</emphasis>
</taxonomicName>
and
<taxonomicName id="0C9B5737589C20660BCDD459CC162172" class="Copepoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F462985C54084C63F21986C01AFC4CE4" italics="true">Bryocyclops</emphasis>
species.
</taxonomicName>
The distal segment of P 5 is reduced to a small lobe on pediger
<quantity id="24A4F03C113CF7BAA1768D1CB9EAD6C4" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="D0E189395E938EA8DF55A7A96828CE0C" authorityName="Boonyanusith, Sanoamuang &amp; Brancelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="genus">
<emphasis id="74151B8F13CF44AED35FA8781971ED24" italics="true">Siamcyclops</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="DF6618634C5F7D841F346672AE1C86E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photographs taken using scanning electron microscope A Bryocyclops asetus Watiroyram, 2018 B B. jayabhumi Watiroyram, 2021 C Siamcyclops cavernicolus, male D S. cavernicolus, female." figureDoi="10.3897/subtbiol.49.135303.figure7" httpUri="https://binary.pensoft.net/fig/1163359">7 C, D</figureCitation>
), while it is more reduced in
<taxonomicName id="C3AC87FEF88355EFA4C8DAD942629A28" authorityName="Kiefer" authorityYear="1927" class="Copepoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="genus">
<emphasis id="4749344323C22748023CF441656FCC27" italics="true">Bryocyclops</emphasis>
</taxonomicName>
, where it is represented by a small prominent knob (Fig.
<figureCitation id="699AB9242FC7C3BB27D4CCE2D3290C9A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photographs taken using scanning electron microscope A Bryocyclops asetus Watiroyram, 2018 B B. jayabhumi Watiroyram, 2021 C Siamcyclops cavernicolus, male D S. cavernicolus, female." figureDoi="10.3897/subtbiol.49.135303.figure7" httpUri="https://binary.pensoft.net/fig/1163359">7 A, B</figureCitation>
). In addition, the setae tend to be shorter and stronger, resembling spiniform setae in some species (Fig.
<figureCitation id="05904027D1776DCD73BFA2141CF20C7F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photographs taken using scanning electron microscope A Bryocyclops asetus Watiroyram, 2018 B B. jayabhumi Watiroyram, 2021 C Siamcyclops cavernicolus, male D S. cavernicolus, female." figureDoi="10.3897/subtbiol.49.135303.figure7" httpUri="https://binary.pensoft.net/fig/1163359">7 B</figureCitation>
), which is advantageous for species living in the confined spaces of the epikarst.
</paragraph>
<caption id="CC1655A578F779DC040258654492AC95" ID-DOI="10.3897/subtbiol.49.135303.figure7" ID-arpha="6A29DA00-CC7E-5C3A-80D9-8E9C26D37C84" httpUri="https://binary.pensoft.net/fig/1163359" startId="F7">
<paragraph id="A66E34D109CDFAF8A6E23BFC26411371">
<label id="964C9E16FCCBF81D88CEF0ABE77FB181">Figure 7.</label>
</paragraph>
<paragraph id="9E59545966519ED4CA29EB88FB087D46">
Photographs taken using scanning electron microscope
<emphasis id="600B78A7A55056986A17B97715A97D60" bold="true">A</emphasis>
<taxonomicName id="ED7EDCF2E064DDF4BE638C1436910F9C" authority="Watiroyram, 2018" authorityName="Watiroyram" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="asetus">
<emphasis id="100B7A7C9E8039067D1F12B4313252F0" italics="true">Bryocyclops asetus</emphasis>
Watiroyram, 2018
</taxonomicName>
<emphasis id="C0B5E68B3DB17E9FBA166A58BCCAF43F" bold="true">B</emphasis>
<taxonomicName id="5B6B82286044270D473EA756D606A677" authority="Watiroyram, 2021" authorityName="Watiroyram" authorityYear="2021" class="Copepoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="jayabhumi">
<emphasis id="DEF4EFCC008C62681B88C612DF03C507" italics="true">B. jayabhumi</emphasis>
Watiroyram, 2021
</taxonomicName>
<emphasis id="4BB3F16F0230FF5890FC441130B3B6B9" bold="true">C</emphasis>
<taxonomicName id="B9783CFC935D4EDF11C107108088A215" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="EAA050AB78C16F997E6CF01F854D340B" italics="true">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
, male
<emphasis id="962401BEABE57C7AEF755B14F6A1AAA6" bold="true">D</emphasis>
<taxonomicName id="3B50F1709A4B024CB25BC2E5B1D74AE0" authorityName="Boonyanusith, Sanoamuang &amp; Brnacelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="D4AEA8A375B3EF670FF9D01F5CAAF0F2" italics="true">S. cavernicolus</emphasis>
</taxonomicName>
, female.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,109 @@
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<mods:title id="38BD5521BEB925106F57AE916DB31DB4">Siamcyclops isanus sp. nov. (Copepoda, Cyclopoida, Cyclopidae), a new cave-dwelling species from northeastern Thailand</mods:title>
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<mods:title id="E0DB890EE4F3F4B5137405CCD25B7BA1">Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part id="B250ECD009841CDEC39DB5CD92F13F2F">
<mods:date id="802F3D72E946670445A2C4CF4B8FC351">2024</mods:date>
<mods:detail id="15724E11FEA28B738FFCEA9E1822E326" type="pubDate">
<mods:number id="01C8DA765EB5CDF66DC80FA100CE4E2B">2024-10-29</mods:number>
</mods:detail>
<mods:detail id="7CC74330FD798104B37D5773AC416D50" type="volume">
<mods:number id="B577CD0EAACD758CBF08CE34C371F515">49</mods:number>
</mods:detail>
<mods:extent id="207C48D6E8704C1C0BF77DEB95EC9C6E" unit="page">
<mods:start id="692DC254C0C0CB762DC39D184B7F6821">185</mods:start>
<mods:end id="7B191F80A61BB6F782F142A7F99A10B1">200</mods:end>
</mods:extent>
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<mods:classification id="4C396162598B7B9E9183016F092C01E9">journal article</mods:classification>
<mods:identifier id="B0EBDB28DEBEF3F9A61506AEE4EA8A40" type="DOI">10.3897/subtbiol.49.135303</mods:identifier>
<mods:identifier id="BA4E2218703024B6756822B7C32A139C" type="ZooBank">F3B1718A-795E-4946-ACC1-9585C414053F</mods:identifier>
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<subSubSection id="2CCAA4A35948D3D133875707F150902B" type="nomenclature">
<paragraph id="D3B3F83C008C885263AE7A83DEB1CD9A">
<heading id="223FFA9598C22E60D2539032026E3EC2" reason="label">Genus</heading>
<taxonomicName id="23F892EFF5773BD207EA19E557BA9C36" ID-CoL="7PZMD" authority="Boonyanusith, Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith, Sanoamuang &amp; Brancelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="genus">
<emphasis id="25E772CFCACFA74BB1AEE5F54B8278C8" italics="true">Siamcyclops</emphasis>
<bibRefCitation id="A372836347B4F49B64BA542A96544057" DOI="10.5852/ejt.2018.431" author="Boonyanusith C &amp; Sanoamuang L &amp; Brancelj A" journalOrPublisher="European Journal of Taxonomy" pagination="1-30" refId="B1" refString="Boonyanusith C, Sanoamuang L, Brancelj A (2018) A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenerathe Bryocyclops and Microcyclops groups. European Journal of Taxonomy 431: 1 - 30. https: // doi. org / 10.5852 / ejt. 2018.431" title="A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenerathe Bryocyclops and Microcyclops groups." volume="431" year="2018">Boonyanusith, Sanoamuang &amp; Brancelj, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJHAC" type="type species">
<paragraph id="513A94D7BD824D2571CE28BBA4D42E93">
<heading id="1599CA1736E49EB827FD9CF9D8DD0E2A" reason="title">
<typeStatus id="8591FADD014E51B3665A1D4A722CD02C">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="E4B985CB25423C3FD01D2D525BBF9832">
<taxonomicName id="97D9558FC6970C3730CD9B30D9935379" authority="Boonyanusith, Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith, Sanoamuang &amp; Brancelj" authorityYear="2018" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="B87ECD97DEB01C92EF224291906CBE02" italics="true">Siamcyclops cavernicolus</emphasis>
<bibRefCitation id="BA6A494BB76D0153834026AB238E2B9B" DOI="10.5852/ejt.2018.431" author="Boonyanusith C &amp; Sanoamuang L &amp; Brancelj A" journalOrPublisher="European Journal of Taxonomy" pagination="1-30" refId="B1" refString="Boonyanusith C, Sanoamuang L, Brancelj A (2018) A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenerathe Bryocyclops and Microcyclops groups. European Journal of Taxonomy 431: 1 - 30. https: // doi. org / 10.5852 / ejt. 2018.431" title="A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenerathe Bryocyclops and Microcyclops groups." volume="431" year="2018">Boonyanusith, Sanoamuang &amp; Brancelj, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1HAC" type="Other species">
<paragraph id="7564FC2EAD809FE99B93C49CB019B357">
<heading id="EC27766878B90C3CBDC69787FBB3FADE" reason="title">Other species.</heading>
</paragraph>
<paragraph id="F00C441F8FDE60382BEA76507F46F067">
<taxonomicName id="C91613D474789B1420173ADB035FEE88" authorityName="Watiroyram &amp; Koompoot" authorityYear="2024" class="Copepoda" family="Cyclopidae" genus="Siamcyclops" kingdom="Animalia" order="Cyclopoida" phylum="Arthropoda" rank="species" species="isanus" status="sp. nov.">
<emphasis id="1FC4477E9C717FC66A1AB52B08B710AF" italics="true">Siamcyclops isanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB6C732FB778B7290B419E230F28ED49" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELIAC" type="Emended diagnosis of the genus">
<paragraph id="2E38DB91B3C705BCE29A96AFD469D087">
<heading id="F354EEC8242F37AF3705E6F103192CE4" reason="title">Emended diagnosis of the genus.</heading>
</paragraph>
<paragraph id="C108997CFF797FEAF03E9472D086ED07">
Body cyclopiform, small size, with the greatest width at the cephalothorax. Pseudosegment present anterior to the genital double somite. Genital double-somite enlarged laterally, shorter than wide. Genital pore located near middle of somite length. Seminal receptacle small; anterior expansion longer and wider than posterior part. Anal operculum well developed or not, with a smooth or serrated distal margin. Antennule 11 - segmented in female, 15 - segmented in male. Antenna without
<abbrev id="ABBRID0ERIAC" xlink_title="exopod">Exp</abbrev>
. Mandibular palp reduced, represented by three setae.
<abbrev id="ABBRID0EVIAC" xlink_title="swimming legs 16">P 1</abbrev>
P 3 coxa with inner seta but absent on P 4.
<abbrev id="ABBRID0EZIAC" xlink_title="swimming legs 16">P 1</abbrev>
P 4 intercoxal sclerite with round prominences.
<abbrev id="ABBRID0E4IAC" xlink_title="swimming legs 16">P 1</abbrev>
basis with inner seta.
<abbrev id="ABBRID0EBJAC" xlink_title="swimming legs 16">P 1</abbrev>
P 4 with 2 - segmented
<abbrev id="ABBRID0EFJAC" xlink_title="exopod">Exp</abbrev>
and
<abbrev id="ABBRID0EJJAC" xlink_title="endopod">Enp</abbrev>
, subequal in size; setal and spines formula of
<abbrev id="ABBRID0ENJAC" xlink_title="exopod">Exp</abbrev>
- 2 as 5.5.5.4 5 and 3.3.3.2, respectively. P 5 fused to pediger 5, with lateral seta and two ventral setae on small lobe.
<abbrev id="ABBRID0ERJAC" xlink_title="swimming legs 16">P 6</abbrev>
of female with slender seta and two spiniform setae, located dorso-laterally on the simple plate of the genital double-somite; three slender setae in male. P 3
<abbrev id="ABBRID0EVJAC" xlink_title="endopod">Enp</abbrev>
- 2 of male with a transformed apical spine and inner subterminal seta.
</paragraph>
</subSubSection>
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