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<mods:title id="B7AA6F7B1083859767A7669C3784303D">Another piece in the puzzle of mantis shrimp evolution - fossils from the Early Jurassic Osteno Lagerstätte of Northern Italy</mods:title>
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<mods:namePart id="8FB2BAF0F5FD7EA063058A4EC05BBFDD">Braig, Florian</mods:namePart>
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<mods:namePart id="63F350DAA36078320E701288AB08429B">Haug, Joachim T.</mods:namePart>
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<mods:affiliation id="C3AAE9D821DC50B7AF601E91985D1957">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center of the LMU Munich, Richard-Wagner-Strasse 10, 80333 München (Germany) joachim. haug @ palaeo-evo-devo. info</mods:affiliation>
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<mods:nameIdentifier id="F45EAE83A47F3433C562B1B7DA2713FB" type="email">joachim.haug@palaeo-evo-devo.info</mods:nameIdentifier>
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<mods:namePart id="E207E270546188C09205C37653E461AF">Ahyong, Shane T.</mods:namePart>
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<mods:affiliation id="AAFDB148DBCF196EAFAA03E53B7319D0">Australian Museum Research Institute, 1 William Strasse, Sydney, NSW 2010 (Australia) and School of Biological, Earth & Environmental Sciences, University of New South Wales, Kensington, NSW 2052 (Australia) shane. ahyong @ austmus. gov. au</mods:affiliation>
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<mods:nameIdentifier id="D08B4484EBFB5354FEF0A8078DDAFD29" type="email">shane.ahyong@austmus.gov.au</mods:nameIdentifier>
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<mods:namePart id="522A81AFF0AE3ECE201C82A0260C214A">Garassino, Alessandro</mods:namePart>
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<mods:affiliation id="FFA454D8291B8C4DD6B750B602B31FC8">Department of Earth and Biological Sciences, Loma Linda University, Loma Linda, 92350 CA (United States) alegarassino @ gmail. com</mods:affiliation>
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<mods:namePart id="7D0CF89AB06EDFC1B1C6BC2EE0E83E0D">Schädel, Mario</mods:namePart>
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<mods:affiliation id="A9F1F0F4153D3504B3321DF4CF975736">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) mario. schaedel @ gmail. com</mods:affiliation>
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<mods:namePart id="F7FD58225022E2F6DE276EC9B84E0C5B">Haug, Carolin</mods:namePart>
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<mods:affiliation id="108D683012152F829A98B4C616601686">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center of the LMU Munich, Richard-Wagner-Strasse 10, 80333 München (Germany) carolin. haug @ palaeo-evo-devo. info (corresponding author)</mods:affiliation>
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<mods:nameIdentifier id="486B28CB51CB1926DEC3130F3F2C0D54" type="email">carolin.haug@palaeo-evo-devo.info</mods:nameIdentifier>
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<mods:title id="2F386D8F02AD20EDBD32824C6217AA03">Comptes Rendus Palevol</mods:title>
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<mods:date id="0EB942BDD60A6EE87DA51F0F44ABCA81">2023</mods:date>
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<treatment id="89766D40FFD34875F43ECBB3FA86FC54" ID-GBIF-Taxon="247569220" LSID="urn:lsid:zoobank.org:act:30E1D19D-C038-45DC-A85E-D971F27DB86D" httpUri="http://treatment.plazi.org/id/89766D40FFD34875F43ECBB3FA86FC54" lastPageNumber="20" pageId="5" pageNumber="20" scope_order="Stomatopoda" scope_subClass="Hoplocarida">
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<taxonomicName id="C6DFA7D5FFD34875F43ECBB3FB0DFE9B" authority="Braig & Haug & Ahyong & Garassino & Schädel & Haug, 2023" authorityName="Braig & Haug & Ahyong & Garassino & Schädel & Haug" authorityYear="2023" box="[1026,1160,1491,1517]" class="Malacostraca" genus="Ostenosculda" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="33AB0044FFD34875F43ECBB3FB0DFE9B" box="[1026,1160,1491,1517]" italics="true" pageId="5" pageNumber="20">Ostenosculda</emphasis>
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<taxonomicNameLabel id="2898BD3FFFD34875F4B3CBB4FB5FFE98" box="[1167,1242,1492,1518]" pageId="5" pageNumber="20" rank="genus">n. gen.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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. —
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<taxonomicName id="C6DFA7D5FFD34875F3F2C804FB14FD0A" authority="Braig & Haug & Ahyong & Garassino & Schädel & Haug, 2023" authorityName="Braig & Haug & Ahyong & Garassino & Schädel & Haug" authorityYear="2023" box="[974,1169,1636,1660]" class="Malacostraca" genus="Ostenosculda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="teruzzii" status="sp. nov.">
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<emphasis id="33AB0044FFD34875F3F2C804FB14FD0A" box="[974,1169,1636,1660]" italics="true" pageId="5" pageNumber="20">Ostenosculda teruzzii</emphasis>
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<taxonomicNameLabel id="2898BD3FFFD34875F4AAC804FB4EFD0D" box="[1174,1227,1636,1659]" pageId="5" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
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, by present designation.
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. — As for the species.
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. — Referring to the Lagerstätte from which the studied specimens originate and the superficial similarities to some specimens of
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<taxonomicName id="C6DFA7D5FFD34875F43FC96AFBC7FC54" authorityName="Munster" authorityYear="1840" box="[1027,1090,1802,1826]" class="Malacostraca" genus="Sculda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="33AB0044FFD34875F43FC96AFBC7FC54" box="[1027,1090,1802,1826]" italics="true" pageId="5" pageNumber="20">Sculda</emphasis>
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</taxonomicName>
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. Gender: feminine.
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</paragraph>
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</treatment>
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data/89/76/6D/89766D40FFD34879F391C913FA34FE78.xml
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data/89/76/6D/89766D40FFD34879F391C913FA34FE78.xml
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<mods:title id="C0091917D2D1F49D3197AD77AC167B45">Another piece in the puzzle of mantis shrimp evolution - fossils from the Early Jurassic Osteno Lagerstätte of Northern Italy</mods:title>
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<mods:namePart id="6ED8E18DEA52B9483A2B72E2669F2685">Braig, Florian</mods:namePart>
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<mods:affiliation id="8DBD94216C83E8F78E5E0DE5CC131B9A">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) florianbraig @ t-online. de</mods:affiliation>
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<mods:namePart id="85DA6225B0BA968D0CA34FF6464C8DB6">Haug, Joachim T.</mods:namePart>
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<mods:affiliation id="21A3FF5930E6FEC7DE5EA589342E1F9A">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center of the LMU Munich, Richard-Wagner-Strasse 10, 80333 München (Germany) joachim. haug @ palaeo-evo-devo. info</mods:affiliation>
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<mods:namePart id="8E358D389E3A8AAA34E26E59C2E5DC71">Ahyong, Shane T.</mods:namePart>
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<mods:affiliation id="77B5553DF384D379E11360EBDEFED3B0">Australian Museum Research Institute, 1 William Strasse, Sydney, NSW 2010 (Australia) and School of Biological, Earth & Environmental Sciences, University of New South Wales, Kensington, NSW 2052 (Australia) shane. ahyong @ austmus. gov. au</mods:affiliation>
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<mods:affiliation id="243533D4DB8FE26107178A073E95E6B3">Department of Earth and Biological Sciences, Loma Linda University, Loma Linda, 92350 CA (United States) alegarassino @ gmail. com</mods:affiliation>
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<mods:affiliation id="AF255BF29380979EF4229E7C7EF842D9">Faculty of Biology, Ludwig-Maximilians-Universität München (LMU Munich), Biocenter, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center of the LMU Munich, Richard-Wagner-Strasse 10, 80333 München (Germany) carolin. haug @ palaeo-evo-devo. info (corresponding author)</mods:affiliation>
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<treatment id="89766D40FFD34879F391C913FA34FE78" ID-GBIF-Taxon="247569221" LSID="urn:lsid:zoobank.org:act:7C2C839B-81D7-4C97-9280-3DA3F62FB6BC" httpUri="http://treatment.plazi.org/id/89766D40FFD34879F391C913FA34FE78" lastPageId="9" lastPageNumber="24" pageId="5" pageNumber="20" scope_order="Stomatopoda" scope_subClass="Hoplocarida">
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<emphasis id="33AB0044FFD34875F391C913FB10FCFA" bold="true" box="[941,1173,1906,1933]" italics="true" pageId="5" pageNumber="20">Ostenosculda teruzzii</emphasis>
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(
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<figureCitation id="99E4C0D3FFD34875F47EC9F2FB1CFCDA" box="[1090,1177,1938,1964]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStartId-0="4.[132,143,1106,1123]" captionStartId-1="6.[132,143,1580,1597]" captionStartId-2="7.[134,143,1581,1598]" captionStartId-3="8.[132,143,1581,1598]" captionStartId-4="10.[132,143,1268,1285]" captionStartId-5="11.[131,142,998,1015]" captionStartId-6="12.[132,143,667,684]" captionTargetBox-0="[132,1455,215,1066]" captionTargetBox-1="[132,1455,214,1537]" captionTargetBox-2="[132,1455,193,1556]" captionTargetBox-3="[132,1455,215,1538]" captionTargetBox-4="[132,1455,215,1222]" captionTargetBox-5="[132,1422,93,955]" captionTargetBox-6="[132,1456,185,638]" captionTargetId-0="figure-474@4.[698,1267,213,783]" captionTargetId-1="figure-340@6.[698,1267,780,1349]" captionTargetId-2="figure-273@7.[699,1268,781,1349]" captionTargetId-3="figure-320@8.[698,1267,781,1349]" captionTargetId-4="figure-485@10.[698,1267,213,765]" captionTargetId-5="figure-662@11.[698,1267,103,676]" captionTargetId-6="figure-789@12.[131,889,202,640]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIG. 1. — A-B, Comparison of part (B) and counterpart (A) of the fossil Ostenosculda teruzzii n. gen., n. sp., ventral view, MSNM i13561: B flipped vertically. Scale bars: 3 mm." captionText-1="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." captionText-2="FIG. 3. — Comparison of part (A) and counterpart (B) of the lateral fossil of Ostenosculda teruzzii n. gen.,n. sp.,MSNM i13562:B, flipped horizontally.Scale bars:3 mm." captionText-3="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." captionText-4="FIG. 5. — Comparison of ventral morphology of an extant mantis shrimp and Ostenosculda teruzzii n. gen., n. sp.: A, thorax and pleon of an extant “spearing” mantis shrimp (Squillidae); B, C, reconstruction of O. teruzzii n. gen., n. sp.: B, ventral view, third to fifth maxillipeds and walking appendages are blurred out, since they are not fully preserved and their morphology can only be estimated; C, lateral view. Scale bar: 8 mm." captionText-5="FIG. 6. — Simplified evolutionary reconstruction and possible phylogenetic affiliation of Ostenosculda teruzzii n. gen., n. sp. indicated by red ellipse; phylogeny based on Haug et al. (2010): evolutionary appearance of characters: 1, further decrease in size of maxillipeds 4 and 5; 2, distinct V-shaped arrangement of maxilliped insertion areas; A, B, schematic reconstruction of arrangement of insertion areas of maxillipeds 2–5 (post-ocular segments 7-10) in mantis shrimps; A, extant species; B, O. teruzzii n. gen., n. sp." captionText-6="FIG. 7. — Reconstruction of the three different schemes of maxilliped arrangement in mantis shrimps: A, Ostenosculda teruzzii n. gen., n. sp., note that the exact size of maxillipeds 3-5 is mostly inferred; B, mantis shrimps from Solnhofen limestones; C, extant mantis shrimps." pageId="5" pageNumber="20">Figs 1-7</figureCitation>
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)
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. 2. —
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<taxonomicName id="C6DFA7D5FFD04876F0E9C84CFE00FD4B" authority="Braig & Haug & Ahyong & Garassino & Schädel & Haug, 2023" authorityName="Braig & Haug & Ahyong & Garassino & Schädel & Haug" authorityYear="2023" box="[213,389,1580,1597]" class="Malacostraca" genus="Ostenosculda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stomatopoda" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="teruzzii" status="n. gen., n. sp.">
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<emphasis id="33AB0044FFD04876F0E9C84CFE00FD4B" box="[213,389,1580,1597]" italics="true" pageId="6" pageNumber="21">Ostenosculda teruzzii</emphasis>
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, MSNM i 13561, composite images of part and counterpart:
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<emphasis id="33AB0044FFD04876F3C9C84CFB87FD4B" bold="true" box="[1013,1026,1580,1597]" pageId="6" pageNumber="21">A</emphasis>
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, overview;
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, colour-coded version of
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, with body segments in different colours for easier identification of segment borders;
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<emphasis id="33AB0044FFD04876F2C5C822FC82FD25" bold="true" box="[761,775,1602,1619]" pageId="6" pageNumber="21">C</emphasis>
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, detailed image of
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, focus on the head and thorax;
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<emphasis id="33AB0044FFD04876F4F0C822FB5FFD25" bold="true" box="[1228,1242,1602,1619]" pageId="6" pageNumber="21">D</emphasis>
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, colour-coded version of
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<emphasis id="33AB0044FFD04876F0B8C839FF17FD1C" bold="true" box="[132,146,1625,1642]" pageId="6" pageNumber="21">C</emphasis>
|
||||
, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations:
|
||||
<emphasis id="33AB0044FFD04876F465C839FBEAFD1C" bold="true" box="[1113,1135,1625,1642]" pageId="6" pageNumber="21">ba</emphasis>
|
||||
, basipod;
|
||||
<emphasis id="33AB0044FFD04876F4F4C839FB58FD1C" bold="true" box="[1224,1245,1625,1642]" pageId="6" pageNumber="21">cx</emphasis>
|
||||
, coxa;
|
||||
<emphasis id="33AB0044FFD04876F527C839FAC7FD1C" bold="true" box="[1307,1346,1625,1642]" pageId="6" pageNumber="21">e1-5</emphasis>
|
||||
, elements of the appendage 1-5;
|
||||
<emphasis id="33AB0044FFD04876F113C810FEDEFDF7" bold="true" box="[303,347,1648,1665]" pageId="6" pageNumber="21">i7-10</emphasis>
|
||||
, insertion areas of the appendages of post-ocular segments 7-10;
|
||||
<emphasis id="33AB0044FFD04876F3B1C810FC2CFDF7" bold="true" box="[909,937,1648,1665]" pageId="6" pageNumber="21">ma</emphasis>
|
||||
, major raptorial appendage;
|
||||
<emphasis id="33AB0044FFD04876F4A0C810FB3DFDF7" bold="true" box="[1180,1208,1648,1665]" pageId="6" pageNumber="21">md</emphasis>
|
||||
, mandible;
|
||||
<emphasis id="33AB0044FFD04876F526C810FAB7FDF7" bold="true" box="[1306,1330,1648,1665]" pageId="6" pageNumber="21">rst</emphasis>
|
||||
, rostrum;
|
||||
<emphasis id="33AB0044FFD04876F5B5C810FF1CFDE1" bold="true" pageId="6" pageNumber="21">s10- 19</emphasis>
|
||||
, sternites of the post-ocular segments 10-19;
|
||||
<emphasis id="33AB0044FFD04876F220C8E6FDD6FDE1" bold="true" box="[540,595,1670,1687]" pageId="6" pageNumber="21">t11-19</emphasis>
|
||||
, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="49C58FDDFFD04876F0B8C8BDFE77FC5F" pageId="6" pageNumber="21" type="materials_examined">
|
||||
<paragraph id="0160DC56FFD04876F0B8C8BDFF6BFC79" blockId="6.[131,775,1756,1833]" pageId="6" pageNumber="21">
|
||||
<smallCapsWord id="07864A8AFFD04876F0B8C8BDFF30FD84" baselines="1774,1774" box="[132,181,1757,1780]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="21">TYPE</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F085C882FE9BFD85" baselines="1775" box="[185,286,1762,1779]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="specimens" pageId="6" pageNumber="21">SPECIMENS</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F11EC882FECEFD85" baselines="1775" box="[290,331,1762,1779]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="6" pageNumber="21">AND</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F173C882FE04FD85" baselines="1775" box="[335,385,1762,1779]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="only" pageId="6" pageNumber="21">ONLY</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F1BAC882FE67FD85" baselines="1775" box="[390,482,1762,1779]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="6" pageNumber="21">MATERIAL</smallCapsWord>
|
||||
. —
|
||||
<materialsCitation id="B1B7D60BFFD04876F234C8BCFF6BFC79" ID-GBIF-Occurrence="4985212301" collectionCode="TYPE, SPECIMENS, AND, ONLY, MATERIAL" pageId="6" pageNumber="21" specimenCode="MSNM i13561" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="33AB0044FFD04876F234C8BCFDE0FD82" bold="true" box="[520,613,1756,1780]" pageId="6" pageNumber="21">
|
||||
<typeStatus id="DE6462F4FFD04876F234C8BCFDE0FD82" box="[520,613,1756,1780]" pageId="6" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
<specimenCode id="5179742DFFD04876F24DC8BCFC82FD82" box="[625,775,1756,1780]" pageId="6" pageNumber="21">MSNM i13561</specimenCode>
|
||||
(
|
||||
<figureCitation id="99E4C0D3FFD04876F0B0C897FF43FC79" box="[140,198,1783,1807]" captionStart="FIG" captionStartId="4.[132,143,1106,1123]" captionTargetBox="[132,1455,215,1066]" captionTargetId="figure-474@4.[698,1267,213,783]" captionTargetPageId="4" captionText="FIG. 1. — A-B, Comparison of part (B) and counterpart (A) of the fossil Ostenosculda teruzzii n. gen., n. sp., ventral view, MSNM i13561: B flipped vertically. Scale bars: 3 mm." pageId="6" pageNumber="21">Figs 1</figureCitation>
|
||||
;
|
||||
<figureCitation id="99E4C0D3FFD04876F0EFC897FF65FC79" box="[211,224,1783,1807]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="6" pageNumber="21">2</figureCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFD04876F0B8C971FE77FC5F" blockId="6.[131,775,1756,1833]" box="[132,498,1809,1833]" pageId="6" pageNumber="21">
|
||||
<materialsCitation id="B1B7D60BFFD04876F0B8C971FE77FC5F" ID-GBIF-Occurrence="4985212302" box="[132,498,1809,1833]" collectionCode="MSNM" pageId="6" pageNumber="21" specimenCode="MSNM i13562" specimenCount="1" typeStatus="paratype">
|
||||
<emphasis id="33AB0044FFD04876F0B8C971FF5EFC5F" bold="true" box="[132,219,1809,1833]" pageId="6" pageNumber="21">
|
||||
<typeStatus id="DE6462F4FFD04876F0B8C971FF5EFC5F" box="[132,219,1809,1833]" pageId="6" pageNumber="21" type="paratype">Paratype</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
<specimenCode id="5179742DFFD04876F0D4C971FE07FC5F" box="[232,386,1809,1833]" pageId="6" pageNumber="21">MSNM i13562</specimenCode>
|
||||
(
|
||||
<figureCitation id="99E4C0D3FFD04876F1ADC971FE4EFC5F" box="[401,459,1809,1833]" captionStart="FIG" captionStartId="7.[134,143,1581,1598]" captionTargetBox="[132,1455,193,1556]" captionTargetId="figure-273@7.[699,1268,781,1349]" captionTargetPageId="7" captionText="FIG. 3. — Comparison of part (A) and counterpart (B) of the lateral fossil of Ostenosculda teruzzii n. gen.,n. sp.,MSNM i13562:B, flipped horizontally.Scale bars:3 mm." pageId="6" pageNumber="21">Figs 3</figureCitation>
|
||||
;
|
||||
<figureCitation id="99E4C0D3FFD04876F1EBC971FE61FC5F" box="[471,484,1809,1833]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="6" pageNumber="21">4</figureCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="49C58FDDFFD04876F0B8C92AFA35FC35" pageId="6" pageNumber="21" type="diagnosis">
|
||||
<paragraph id="0160DC56FFD04876F0B8C92AFA35FC35" blockId="6.[130,778,1866,2022]" lastBlockId="6.[811,1456,1756,1859]" pageId="6" pageNumber="21">
|
||||
<smallCapsWord id="07864A8AFFD04876F0B8C92AFF68FC16" baselines="1883,1884" box="[132,237,1866,1889]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="6" pageNumber="21">DIAGNOSIS</smallCapsWord>
|
||||
. — Compound eyes without apparent midband, on short peduncles. Short tapering rostrum. Shield rectangular in ventral and lateral view with postero-lateral corners elongated. Major raptorial appendages with thick ovate carpopropodi and spineless scythe-shaped dactyli. Eight circular insertion areas (foramina) of posterior four maxilliped pairs arranged evenly spaced in anterior-posterior axis. The first pair of insertion areas positioned further laterally, the following three pairs arranged successively medially in two anterior-posterior running, almost straight lines. Insertion area of each pleopod subdivided into three individual areas separated by bar-shaped sclerites.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="49C58FDDFFD04876F311C904FA0BFCE0" pageId="6" pageNumber="21" type="etymology">
|
||||
<paragraph id="0160DC56FFD04876F311C904FA0BFCE0" blockId="6.[813,1455,1892,1942]" pageId="6" pageNumber="21">
|
||||
<smallCapsWord id="07864A8AFFD04876F311C904FC22FC0C" baselines="1909,1910" box="[813,935,1892,1915]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Derivation" pageId="6" pageNumber="21">DERIVATION</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F393C909FC4DFC0C" baselines="1910" box="[943,968,1897,1914]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="of" pageId="6" pageNumber="21">OF</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F3ECC909FB82FC0C" baselines="1910" box="[976,1031,1897,1914]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="name" pageId="6" pageNumber="21">NAME</smallCapsWord>
|
||||
. — Named after Giorgio Teruzzi, Milano, in honour of his scientific work including fossils from Osteno.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="49C58FDDFFD04876F311C9D7FB2FFC9F" pageId="6" pageNumber="21" type="distribution">
|
||||
<paragraph id="0160DC56FFD04876F311C9D7FB2FFC9F" blockId="6.[813,1456,1975,2025]" pageId="6" pageNumber="21">
|
||||
<smallCapsWord id="07864A8AFFD04876F311C9D7FCD8FCBB" baselines="1992,1993" box="[813,861,1975,1998]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="21">
|
||||
<collectionCode id="67CE4493FFD04876F311C9D7FCD8FCBB" box="[813,861,1975,1998]" pageId="6" pageNumber="21">TYPE</collectionCode>
|
||||
</smallCapsWord>
|
||||
<smallCapsWord id="07864A8AFFD04876F359C9DCFC45FCBB" baselines="1993" box="[869,960,1980,1997]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="6" pageNumber="21">
|
||||
<collectionCode id="67CE4493FFD04876F359C9DCFC45FCBB" box="[869,960,1980,1997]" pageId="6" pageNumber="21">LOCALITY</collectionCode>
|
||||
</smallCapsWord>
|
||||
. — Early Jurassic (Sinemurian), limestone quarry, Osteno,
|
||||
<collectingRegion id="C31B12B4FFD04876F3BEC9B1FC6DFC9F" box="[898,1000,2001,2025]" country="Italy" name="Lombardia" pageId="6" pageNumber="21">Lombardy</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="79C89CC6FFD04876F3CDC9B1FB99FC9F" box="[1009,1052,2001,2025]" name="Italy" pageId="6" pageNumber="21">Italy</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="654EA1A7FFD04876F416C9B1FB18FC9F" author="PINNA G." box="[1066,1181,2001,2025]" pageId="6" pageNumber="21" pagination="91 - 136" refId="ref11297" refString="PINNA G. 2000. - Die Fossillagerstatte im Sinemurium (Lias) von Osteno, Italien, in PINNA G. & MEISCHNER D. (eds), Europaische Fossillagerstatten. Springer-Verlag, Berlin Heidelberg: 91 - 136." type="book chapter" year="2000">Pinna 2000</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="55A08CDEFFD14877F0BAC84DFA34FD48" box="[134,1457,1581,1598]" pageId="7" pageNumber="22" startId="7.[134,143,1581,1598]" targetBox="[132,1455,193,1556]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="0160DC56FFD14877F0BAC84DFA34FD48" blockId="7.[134,1457,1581,1598]" box="[134,1457,1581,1598]" pageId="7" pageNumber="22">
|
||||
<smallCapsWord id="07864A8AFFD14877F0BAC84DFF18FD4B" baselines="1594,1594" box="[134,157,1581,1598]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="7" pageNumber="22">FIG</smallCapsWord>
|
||||
. 3. — Comparison of part (
|
||||
<emphasis id="33AB0044FFD14877F14FC84DFE05FD48" bold="true" box="[371,384,1581,1598]" pageId="7" pageNumber="22">A</emphasis>
|
||||
) and counterpart (
|
||||
<emphasis id="33AB0044FFD14877F22DC84DFD9BFD48" bold="true" box="[529,542,1581,1598]" pageId="7" pageNumber="22">B</emphasis>
|
||||
) of the lateral fossil of
|
||||
<taxonomicName id="C6DFA7D5FFD14877F2F0C84DFCFDFD48" authority="Braig & Haug & Ahyong & Garassino & Schädel & Haug, 2023" authorityName="Braig & Haug & Ahyong & Garassino & Schädel & Haug" authorityYear="2023" box="[716,888,1581,1598]" class="Malacostraca" genus="Ostenosculda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="teruzzii" status="n. gen., n. sp.">
|
||||
<emphasis id="33AB0044FFD14877F2F0C84DFCFDFD48" box="[716,888,1581,1598]" italics="true" pageId="7" pageNumber="22">Ostenosculda teruzzii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="2898BD3FFFD14877F347C84DFC6CFD48" box="[891,1001,1581,1598]" pageId="7" pageNumber="22" rank="species">n. gen.,n. sp.</taxonomicNameLabel>
|
||||
,MSNM i13562:
|
||||
<emphasis id="33AB0044FFD14877F456C84DFBF2FD48" bold="true" box="[1130,1143,1581,1598]" pageId="7" pageNumber="22">B</emphasis>
|
||||
, flipped horizontally.Scale bars:3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="49C58FDDFFD14879F0B8C813FA34FE78" lastPageId="9" lastPageNumber="24" pageId="7" pageNumber="22" type="description">
|
||||
<paragraph id="0160DC56FFD14877F0B8C813FE90FDFD" blockId="7.[130,775,1651,2028]" box="[132,277,1651,1677]" pageId="7" pageNumber="22">
|
||||
<smallCapsWord id="07864A8AFFD14877F0B8C813FE90FDFD" baselines="1670,1670" box="[132,277,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="22">DESCRIPTION</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFD14877F0B8C8F3FC5EFD9B" blockId="7.[130,775,1651,2028]" lastBlockId="7.[811,1458,1651,2028]" pageId="7" pageNumber="22">
|
||||
Body apparently organised into six functional units (
|
||||
<bibRefCitation id="654EA1A7FFD14877F2F5C8F3FF7EFDBB" author="HAUG C. & SALLAM W. S. & MAAS A. & WALOSZEK D. & KUTSCHERA V. & HAUG J. T." pageId="7" pageNumber="22" refId="ref10037" refString="HAUG C., SALLAM W. S., MAAS A., WALOSZEK D., KUTSCHERA V. & HAUG J. T. 2012. - Tagmatization in Stomatopoda - reconsidering functional units of modern-day mantis shrimps (Verunipeltata, Hoplocarida) and implications for the interpretation of fossils. Frontiers in Zoology 9 (31). https: // doi. org / 10.1186 / 1742 - 9994 - 9 - 31" type="journal volume" year="2012">
|
||||
Haug
|
||||
<emphasis id="33AB0044FFD14877F0B8C8D3FF3FFDBA" box="[132,186,1714,1740]" italics="true" pageId="7" pageNumber="22">et al.</emphasis>
|
||||
2012
|
||||
</bibRefCitation>
|
||||
): 1) sensorial unit with ocular and post-ocular segments 1-2 (anterior head); 2) anterior food-processing unit with post-ocular segments 3-5 (posterior head); 3) posterior food-processing unit with post-ocular segments 6-10 (thoracic segments 1-5); 4) walking apparatus with post-ocular segments 11-13 (thoracic segments 6-8); 5) swimming and respiration unit with post-ocular segments 14-18 (anterior pleon); 6) tailfan with post-ocular segment 19 (pleomere 6) and telson (seeFig. 5 for comparison with extant mantis shrimp). The body length is
|
||||
<quantity id="C62771B3FFD14877F1FCC9B1FDAFFC9A" box="[448,554,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="4.66" pageId="7" pageNumber="22" unit="mm" value="46.6">46.6 mm</quantity>
|
||||
, from the tip of the rostrum to the end of the telson for the ventrally accessible specimen (
|
||||
<figureCitation id="99E4C0D3FFD14877F3EDC8F3FB97FDDB" box="[977,1042,1683,1709]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="7" pageNumber="22">Fig. 2</figureCitation>
|
||||
); for the laterally accessible one this measurement is difficult due to the spaces between the body segments.
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFD14877F378C892FB9AFCDA" blockId="7.[811,1458,1651,2028]" pageId="7" pageNumber="22">
|
||||
The only clearly visible structure of the sensorial unit is one (presumably the left) large compound eye on a short peduncle, protruding near the anterior end of the shield. The cornea is shorter but wider than the stalk; an ommatidial midband is not determinable. Antennula and antenna are not preserved (
|
||||
<figureCitation id="99E4C0D3FFD14877F3ECC9F2FB8AFCDA" box="[976,1039,1938,1964]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="7" pageNumber="22">Fig. 4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFD14878F37FC9D2FB9CFC9D" blockId="7.[811,1458,1651,2028]" lastBlockId="8.[811,1457,1714,2027]" lastPageId="8" lastPageNumber="23" pageId="7" pageNumber="22">
|
||||
The only accessible feature of the food-processing unit (mandible, maxillula, maxilla) are the mandibles of the ventrally accessible specimen (
|
||||
<figureCitation id="99E4C0D3FFDE4878F1E8C8D2FD91FDBB" box="[468,532,1714,1741]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="8" pageNumber="23">Fig. 2</figureCitation>
|
||||
). They insert anterior to the major raptorial appendages, but more laterally to them on both sides, on the lateral edges of the body. The mandibles appear to be roughly triangular in ventral view, though their proximal parts at the lateral sides of the body are very round and their distal parts at the median areas of the body are very rough and serrated, indicating grinding/cutting structures. They measure 2.7 and
|
||||
<quantity id="C62771B3FFDE4878F2B1C9F2FD62FCDA" box="[653,743,1938,1964]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="23" unit="mm" value="3.1">3.1 mm</quantity>
|
||||
in length, left and right mandible respectively. The segments to which the appendages of the food-processing unit belong are indirectly observable as they dorsally form the prominent shield. The shield is about two times as long as wide, about
|
||||
<quantity id="C62771B3FFDE4878F348C892FC40FC7B" box="[884,965,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="23" unit="mm" value="12.0">12 mm</quantity>
|
||||
long and
|
||||
<quantity id="C62771B3FFDE4878F404C892FBFFFC7B" box="[1080,1146,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="23" unit="mm" value="6.0">6 mm</quantity>
|
||||
wide in the ventrally accessible specimen (
|
||||
<figureCitation id="99E4C0D3FFDE4878F3DDC972FBA7FC5A" box="[993,1058,1810,1836]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="8" pageNumber="23">Fig. 2</figureCitation>
|
||||
), appearing subrectangular both in ventral and lateral view, covering the post-ocular segments
|
||||
<quantity id="C62771B3FFDE4878F311C932FCF5FC1A" box="[813,880,1874,1901]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="8" pageNumber="23" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1-5 in</quantity>
|
||||
ventral view and post-ocular segments
|
||||
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|
||||
lateral view. A short tapering rostrum bridging the gap between the eye peduncles is present at the anteromedial margin of the shield. The posterolateral corners of the shield are slightly produced posteriorly.
|
||||
</paragraph>
|
||||
<caption id="55A08CDEFFDE4878F0B8C84DFC08FDF4" pageId="8" pageNumber="23" startId="8.[132,143,1581,1598]" targetBox="[132,1455,215,1538]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="0160DC56FFDE4878F0B8C84DFC08FDF4" blockId="8.[132,1457,1581,1666]" pageId="8" pageNumber="23">
|
||||
<smallCapsWord id="07864A8AFFDE4878F0B8C84DFF18FD4B" baselines="1594,1594" box="[132,157,1581,1598]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="23">FIG</smallCapsWord>
|
||||
. 4. —
|
||||
<taxonomicName id="C6DFA7D5FFDE4878F0EAC84DFE02FD48" authority="Braig & Haug & Ahyong & Garassino & Schädel & Haug, 2023" authorityName="Braig & Haug & Ahyong & Garassino & Schädel & Haug" authorityYear="2023" box="[214,391,1581,1598]" class="Malacostraca" genus="Ostenosculda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="teruzzii" status="n. gen., n. sp.">
|
||||
<emphasis id="33AB0044FFDE4878F0EAC84DFE02FD48" box="[214,391,1581,1598]" italics="true" pageId="8" pageNumber="23">Ostenosculda teruzzii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="2898BD3FFFDE4878F1B2C84DFD83FD48" box="[398,518,1581,1598]" pageId="8" pageNumber="23" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||||
, MSNM i13562, composite images of part and counterpart:
|
||||
<emphasis id="33AB0044FFDE4878F43CC84DFB88FD48" bold="true" box="[1024,1037,1581,1598]" pageId="8" pageNumber="23">A</emphasis>
|
||||
, overview;
|
||||
<emphasis id="33AB0044FFDE4878F451C84DFBFFFD48" bold="true" box="[1133,1146,1581,1598]" pageId="8" pageNumber="23">B</emphasis>
|
||||
, colour-coded version of
|
||||
<emphasis id="33AB0044FFDE4878F568C84DFAE4FD48" bold="true" box="[1364,1377,1581,1598]" pageId="8" pageNumber="23">A</emphasis>
|
||||
, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations:
|
||||
<emphasis id="33AB0044FFDE4878F3C0C824FB94FD23" bold="true" box="[1020,1041,1604,1621]" pageId="8" pageNumber="23">ce</emphasis>
|
||||
, compound eye;
|
||||
<emphasis id="33AB0044FFDE4878F499C824FB44FD23" bold="true" box="[1189,1217,1604,1621]" pageId="8" pageNumber="23">ma</emphasis>
|
||||
, major raptorial appendage;
|
||||
<emphasis id="33AB0044FFDE4878F0B8C83AFF2FFD1D" bold="true" box="[132,170,1626,1643]" pageId="8" pageNumber="23">mx?</emphasis>
|
||||
, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);
|
||||
<emphasis id="33AB0044FFDE4878F391C83AFC6DFD1D" bold="true" box="[941,1000,1626,1643]" pageId="8" pageNumber="23">s12-19</emphasis>
|
||||
, sternites of the post-ocular segments 12-19;
|
||||
<emphasis id="33AB0044FFDE4878F55CC83AFAF0FD1D" bold="true" box="[1376,1397,1626,1643]" pageId="8" pageNumber="23">sh</emphasis>
|
||||
, shield (carapace);
|
||||
<emphasis id="33AB0044FFDE4878F0DAC811FE98FDF4" bold="true" box="[230,285,1649,1666]" pageId="8" pageNumber="23">t11-19</emphasis>
|
||||
, tergites of the post-ocular segments 11-19;
|
||||
<emphasis id="33AB0044FFDE4878F2ABC811FD28FDF4" bold="true" box="[663,685,1649,1666]" pageId="8" pageNumber="23">tel</emphasis>
|
||||
, telson. Scale bars: 3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="0160DC56FFDF4879F0A7CEB8FD8DFE39" blockId="9.[130,777,216,2028]" pageId="9" pageNumber="24">
|
||||
The posterior food-processing unit is mainly represented through the major raptorial appendage (maxilliped 2). The exact number of appendage elements of these is not observable. Only the two most distal elements are recognizable: the thick and bulky, in lateral view oval-shaped carpopropodus (propodus of other authors), and the long and spineless scythe-shaped dactylus (
|
||||
<figureCitation id="99E4C0D3FFDF4879F1B3CFF7FE4AFAC7" box="[399,463,407,433]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
). The dactylus is arcuate, at least three-fourths as long as the carpopropodus, evenly tapering to a spiniform tip; the outer margin is smooth, lacking a proximal notch or swelling. The carpopropodus is twice as long as high, about
|
||||
<quantity id="C62771B3FFDF4879F161CC77FE31F947" box="[349,436,535,561]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="9" pageNumber="24" unit="mm" value="8.9">8.9 mm</quantity>
|
||||
long in the ventrally accessible specimen and about
|
||||
<quantity id="C62771B3FFDF4879F15ACC57FE38F927" box="[358,445,567,593]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="24" unit="mm" value="4.3">4.3 mm</quantity>
|
||||
wide in the laterally accessible specimen, and is deepest at the midlength. Other appendages are not fully preserved, but two additional dactyli are visible between the posterior portion of the maxillipeds 2; these small dactyli are about one fourth of the length of the dactyli of maxilliped 2 (
|
||||
<figureCitation id="99E4C0D3FFDF4879F109CCB6FEF7F986" box="[309,370,726,752]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="9" pageNumber="24">Fig. 2</figureCitation>
|
||||
) and possibly belong to maxillipeds 3 or 4. In lateral view, darkened areas are visible ventral to the shield, which may represent proximal maxilliped elements stacked dorsoventrally on top of each other (
|
||||
<figureCitation id="99E4C0D3FFDF4879F246CD56FD3EF826" box="[634,699,822,848]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
: mx?). Furthermore, the insertions of the posterior four maxillipeds (appendages of post-ocular segments 7-10) are clearly recognizable. These are evenly arranged in anterior-posterior axis, slightly decreasing in size from anterior to posterior. The first pair of insertion areas is positioned further laterally, the following three pairs are arranged successively medially in an almost straight line. The most anterior pair of insertions belongs presumably to the major raptorial appendages (maxilliped 2) since the bulging cuticle rings surrounding the insertions are thicker than those of the more posterior maxillipeds and the diameter is significantly larger (
|
||||
<figureCitation id="99E4C0D3FFDF4879F284CAF4FD72FFD9" box="[696,759,1172,1199]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="9" pageNumber="24">Fig. 2</figureCitation>
|
||||
). The next posterior insertion is smaller, but slightly larger in diameter and with a thicker bulging ring than the next two insertions. The last segment of this tagma has a distinct short dorsal sclerotisation (tergite) and a longer (anterior-posterior axis) ventral sclerotisation (sternite).
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFDF4879F0A7CB34FB35FA04" blockId="9.[130,777,216,2028]" lastBlockId="9.[811,1457,216,1295]" pageId="9" pageNumber="24">
|
||||
The thoracic segments 6-8, which form the walking apparatus, are clearly recognizable. Each segment seems to have an oval cross-section and is about four times as high as long (
|
||||
<figureCitation id="99E4C0D3FFDF4879F286CBF3FD72FED8" box="[698,759,1427,1454]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
). Each segment has two cuticle plates, a dorsal tergite and a ventral sternite; in lateral view, the tergite occupies two-thirds of the segment height, the sternite only one-third, possibly as a result of compressional displacement (
|
||||
<figureCitation id="99E4C0D3FFDF4879F275C873FD03FD5B" box="[585,646,1555,1581]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
). While the tergites are continuous and broad rectangular-shaped cuticle plates, the sternites are interspersed with softer structures. Most anteriorly on each sternite, and probably marking its anterior edge, is a continuous crescent-like shaped sclerotised bar stretching over the full width of the segment and continuing in posterior direction (after a kink with roughly 90°) at both of its ends to the posterior end of the segment. Posterior to this bar, the seventh thoracic segment shows structures of hardened cuticle. These form two broadly connected ovate forms around the possible insertion areas of the walking appendages. Posterior to the sclerotised bar of the eighth thoracic segment is a broad rectangular cuticle plate with small anterior spines pointing laterally. Posterior to this is a structure mirrored on an anterior-posterior running middle line of the segment. The structure consists of a walking-stick-shaped cuticle bar (curved end pointing medially) above a rough triangle pointing posterior and has its anterior corners elongated laterally. These triangles reach into the gap between the eighth thoracic and the first pleon segment (
|
||||
<figureCitation id="99E4C0D3FFDF4879F45ECF37FB25FA07" box="[1122,1184,343,369]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="9" pageNumber="24">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFDF4879F37FCF17FB72F846" blockId="9.[811,1457,216,1295]" pageId="9" pageNumber="24">
|
||||
The anterior pleon consists of a narrow anterior segment, followed by four broader segments, each two times higher than long (
|
||||
<figureCitation id="99E4C0D3FFDF4879F39DCFD7FC58FAA7" box="[929,989,439,465]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
). Dorsally, the tergite of each of the five segments is a broad continuous plate covering the whole segment (
|
||||
<figureCitation id="99E4C0D3FFDF4879F30ACF97FCF7F967" box="[822,882,503,529]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="9" pageNumber="24">Fig. 2</figureCitation>
|
||||
). The sternites show multiple structures and soft spots that are best described by the letter “W”, but with the three anterior tips being connected and by overall broader lines. This leads to two triangular gaps, which form the insertion areas (foramina) of the pleopods. Within these gaps, there are two cuticle bars, one running from the middle of the lateral side towards the midline and bending posterior after two thirds of the way, and one running from posterior towards the spot where the first line starts to bend. This pattern continues with slight variation for the other four sternites.
|
||||
</paragraph>
|
||||
<paragraph id="0160DC56FFDF4879F37FCD56FA34FE78" blockId="9.[811,1457,216,1295]" pageId="9" pageNumber="24">
|
||||
The tailfan is formed by the telson and the appendages (uropods) of pleomere 6, which is slightly narrower compared to the other pleon segments. Pleomere 6 again consists of a continuous broad cuticular plate that is the tergite, but in contrast to further anterior sternites, the sternite of pleomere 6 is broad and almost continuous, only with spaces postero-laterally for the uropods. The uropods (at least the parts that are preserved) are bent in ventral view and curved around the lateral edges of the telson (
|
||||
<figureCitation id="99E4C0D3FFDF4879F4ECCA55FA8BFF39" box="[1232,1294,1077,1103]" captionStart="FIG" captionStartId="6.[132,143,1580,1597]" captionTargetBox="[132,1455,214,1537]" captionTargetId="figure-340@6.[698,1267,780,1349]" captionTargetPageId="6" captionText="FIG. 2. — Ostenosculda teruzzii n. gen., n. sp., MSNM i 13561, composite images of part and counterpart: A, overview; B, colour-coded version of A, with body segments in different colours for easier identification of segment borders; C, detailed image of A, focus on the head and thorax; D, colour-coded version of C, with the numerous appendage elements and details in different colours for an easier identification. Abbreviations: ba, basipod; cx, coxa; e1-5, elements of the appendage 1-5; i7-10, insertion areas of the appendages of post-ocular segments 7-10; ma, major raptorial appendage; md, mandible; rst, rostrum; s10- 19, sternites of the post-ocular segments 10-19; t11-19, tergites of the post-ocular segments 11-19. Scale bars: A, B, 3 mm; C, D, 2 mm." pageId="9" pageNumber="24">Fig. 2</figureCitation>
|
||||
). The telson is flatter than the body segments and flattens out even further towards its posterior end (
|
||||
<figureCitation id="99E4C0D3FFDF4879F47ECA14FBFBFFF9" box="[1090,1150,1140,1167]" captionStart="FIG" captionStartId="8.[132,143,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-320@8.[698,1267,781,1349]" captionTargetPageId="8" captionText="FIG. 4. — Ostenosculda teruzzii n. gen., n. sp., MSNM i13562, composite images of part and counterpart: A, overview; B, colour-coded version of A, with the numerous body segments in different colours for easier identification of segment borders. Abbreviations: ce, compound eye; ma, major raptorial appendage; mx?, possible maxillipeds (proximal parts either of maxillipeds 2 or of further posterior maxillipeds);s12-19, sternites of the post-ocular segments 12-19;sh, shield (carapace); t11-19, tergites of the post-ocular segments 11-19; tel, telson. Scale bars: 3 mm." pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||||
). It is about
|
||||
<quantity id="C62771B3FFDF4879F539CA14FADFFFF9" box="[1285,1370,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="24" unit="mm" value="6.9">6.9 mm</quantity>
|
||||
wide in the ventrally accessible specimen and about
|
||||
<quantity id="C62771B3FFDF4879F539CAF4FADEFFD9" box="[1285,1371,1172,1199]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="24" unit="mm" value="5.2">5.2 mm</quantity>
|
||||
long in the laterally accessible specimen, yet it is neither in lateral nor in ventral view fully preserved, therefore we could draw no further information from it. Telson spines are not preserved.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Add table
Add a link
Reference in a new issue