Add updates up until 2024-11-13 12:45:04

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<document id="942AD7C45141A5EE620215C6F72AC56F" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666301A5226EF08FE4574260246" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coprophanaeus telamon" docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="65" updateTime="1731501843003" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:affiliation id="AB35E856802EA660155C54C5A9442058">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<emphasis id="A78D1B62301A5224EF08FE45744400C2" bold="true" box="[543,840,482,509]" italics="true" pageId="64" pageNumber="65">Coprophanaeus telamon</emphasis>
(Erichson, 1847)
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<figureCitation id="0DC2DBF5301A5224EFF1FDA27441031F" box="[742,845,517,544]" captionStart="Fig" captionStartId="65.[189,232,1963,1989]" captionTargetBox="[279,1304,277,1880]" captionTargetId="figure-12@65.[268,1319,264,1913]" captionTargetPageId="65" captionText="Fig. 25. Species distribution. A. Coprophanaeus parvulus (dOlsoufieff, 1924). B. Coprophanaeus telamon (Erichson, 1847). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945529" httpUri="https://zenodo.org/record/13945529/files/figure.png" pageId="64" pageNumber="65">Fig. 25B</figureCitation>
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<emphasis id="A78D1B62301A5224EDAAFDE0769A035E" box="[189,406,583,609]" italics="true" pageId="64" pageNumber="65">Phanaeus telamon</emphasis>
<bibRefCitation id="F168BA81301A5224EC8AFDE0759A035E" author="Erichson W. F." box="[413,662,583,609]" pageId="64" pageNumber="65" pagination="67 - 185" refId="ref117813" refString="Erichson W. F. 1847 a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185. Available from https: // www. biodiversitylibrary. org / page / 7251438 [accessed 14 Feb. 2024]." type="journal article" year="1847">Erichson, 1847a: 106</bibRefCitation>
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(original description). Type locality:
<collectingCountry id="EDEE87E0301A5224E95EFDE0738F035E" box="[1097,1155,583,609]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
. Name-bearing type:
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(MFNB), not examined.
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<treatmentCitationGroup id="B5E9E05E301A5224EDAAFD0C7582026D" pageId="64" pageNumber="65">
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<emphasis id="A78D1B62301A5224EDAAFD0C769B03FA" box="[189,407,683,709]" italics="true" pageId="64" pageNumber="65">Phanaeus telamon</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161301A5224ECA3FD0C75AE03FA" author="Harold E." box="[436,674,683,710]" page="1019" pageId="64" pageNumber="65" year="1869">
<bibRefCitation id="F168BA81301A5224ECA3FD0C75AE03FA" author="Harold E." box="[436,674,683,710]" pageId="64" pageNumber="65" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1019</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E0301A5224EE47FD0C748703FA" box="[848,907,683,709]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EED4FD0C738303FA" author="Kirsch T." box="[963,1167,683,709]" page="341" pageId="64" pageNumber="65" year="1873">
<bibRefCitation id="F168BA81301A5224EED4FD0C738303FA" author="Kirsch T." box="[963,1167,683,709]" pageId="64" pageNumber="65" pagination="339 - 418" refId="ref125736" refString="Kirsch T. 1873. Beitrage zur Kenntniss der peruanische Kaferfauna I - II. Berliner entomologische Zeitschrift 17 (121 - 152): 339 - 418. Available from https: // www. biodiversitylibrary. org / page / 8341419 [accessed 16 Feb. 2024]." type="journal article" year="1873">Kirsch 1873: 341</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E0301A5224E81CFD0C724A03FA" box="[1291,1350,683,709]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EDFCFD68769503D6" author="Bates H. W." box="[235,409,718,745]" page="56" pageId="64" pageNumber="65" year="1887">
<bibRefCitation id="F168BA81301A5224EDFCFD68769503D6" author="Bates H. W." box="[235,409,718,745]" pageId="64" pageNumber="65" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1887">Bates 1887: 56</bibRefCitation>
</treatmentCitation>
(diagnosis). —
<treatmentCitation id="1458E161301A5224EF42FD69742903D6" author="Nevinson B. G." box="[597,805,718,745]" page="7" pageId="64" pageNumber="65" year="1892">
<bibRefCitation id="F168BA81301A5224EF42FD69742903D6" author="Nevinson B. G." box="[597,805,718,745]" pageId="64" pageNumber="65" refId="ref131974" refString="Nevinson B. G. 1892. Revised Synonymic List of Species in the Genera Phanӕus Macleay and Oxysternon Castelnau. Privately published, London. https: // doi. org / 10.5962 / bhl. title. 9416" type="book" year="1892">Nevinson 1892: 7</bibRefCitation>
</treatmentCitation>
(list, revision, cited for
<collectingCountry id="EDEE87E0301A5224E92AFD69738903D7" box="[1085,1157,718,744]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
and
<collectingCountry id="EDEE87E0301A5224E9AAFD6873F403D6" box="[1213,1272,719,745]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E838FD6976530232" author="Kolbe H. J." page="581" pageId="64" pageNumber="65" year="1905">
<bibRefCitation id="F168BA81301A5224E838FD6976530232" author="Kolbe H. J." pageId="64" pageNumber="65" pagination="475 - 594" refId="ref126331" refString="Kolbe H. J. 1905. Uber die Lebensweise und die geographische Verbreitung der coprophagen Lamellicornier. Zoologischen Jahrbucher Supplementheff 8: 475 - 594. Available from https: // www. biodiversitylibrary. org / page / 14202040 [accessed 14 Feb. 2024]." type="journal article" year="1905">Kolbe 1905: 581</bibRefCitation>
</treatmentCitation>
(comments).
<emphasis id="A78D1B62301A5224ECEAFD54751B0233" box="[509,535,755,780]" italics="true" pageId="64" pageNumber="65"></emphasis>
<treatmentCitation id="1458E161301A5224EF08FD5575D70233" author="Gillet J. J. E." box="[543,731,754,780]" page="86" pageId="64" pageNumber="65" year="1911">
<bibRefCitation id="F168BA81301A5224EF08FD5575D70233" author="Gillet J. J. E." box="[543,731,754,780]" pageId="64" pageNumber="65" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 86</bibRefCitation>
</treatmentCitation>
(catalogue, cited for
<collectingCountry id="EDEE87E0301A5224EEC7FD55730A0233" box="[976,1030,754,780]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
and
<collectingCountry id="EDEE87E0301A5224E92AFD5573860233" box="[1085,1162,754,780]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E9A8FD557619020F" author="Balthasar V." page="350" pageId="64" pageNumber="65" year="1941">
<bibRefCitation id="F168BA81301A5224E9A8FD557619020F" author="Balthasar V." pageId="64" pageNumber="65" pagination="337 - 358" refId="ref110399" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="book chapter" year="1941">Balthasar 1941: 350</bibRefCitation>
</treatmentCitation>
(distribution);
<treatmentCitation id="1458E161301A5224ECDAFCB2754B0210" author="Balthasar V." box="[461,583,789,816]" page="336" pageId="64" pageNumber="65" year="1951">
<bibRefCitation id="F168BA81301A5224ECDAFCB2754B0210" author="Balthasar V." box="[461,583,789,816]" pageId="64" pageNumber="65" pagination="322 - 344" refId="ref110452" refString="Balthasar V. 1951. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 2: 322 - 344." type="book chapter" year="1951">1951: 336</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E161301A5224EE30FCB273330210" author="Blackwelder R. E." box="[807,1087,789,815]" page="210" pageId="64" pageNumber="65" year="1944">
<bibRefCitation id="F168BA81301A5224EE30FCB273330210" author="Blackwelder R. E." box="[807,1087,789,815]" pageId="64" pageNumber="65" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 210</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E0301A5224E9EDFCB2724E0210" box="[1274,1346,789,815]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
and
<collectingCountry id="EDEE87E0301A5224EDFDFC9F7629026D" box="[234,293,824,850]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EC4AFC9F7522026D" author="Pereira F. S." box="[349,558,824,851]" page="391" pageId="64" pageNumber="65" year="1953">
<bibRefCitation id="F168BA81301A5224EC4AFC9F7522026D" author="Pereira F. S." box="[349,558,824,851]" pageId="64" pageNumber="65" pagination="387 - 402" refId="ref135282" refString="Pereira F. S. 1953. Notas sinonimicas. Dusenia 4 (5 - 6): 387 - 402." type="journal article" year="1953">Pereira 1953: 391</bibRefCitation>
</treatmentCitation>
(notes).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770301A5224EDAAFCFC734A0283" blockId="64.[189,1399,683,1696]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="B5E9E05E301A5224EDAAFCFC734A0283" pageId="64" pageNumber="65">
<taxonomicName id="52F9BCF3301A5224EDAAFCFC7566024A" ID-CoL="7Y4M7" authorityName="Erichson" authorityYear="1847" box="[189,618,859,885]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="telamon" subGenus="Coprophanaeus">
<emphasis id="A78D1B62301A5224EDAAFCFC763D024A" box="[189,305,859,885]" italics="true" pageId="64" pageNumber="65">Phanaeus</emphasis>
(
<emphasis id="A78D1B62301A5224EC56FCFC76F7024A" box="[321,507,859,885]" italics="true" pageId="64" pageNumber="65">Coprophanaeus</emphasis>
)
<emphasis id="A78D1B62301A5224EF1CFCFC7566024A" box="[523,618,859,885]" italics="true" pageId="64" pageNumber="65">telamon</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161301A5224EF92FCFC748F024A" author="d'Olsoufieff G." box="[645,899,859,885]" page="26" pageId="64" pageNumber="65" year="1924">
<bibRefCitation id="F168BA81301A5224EF92FCFC748F024A" author="d'Olsoufieff G." box="[645,899,859,885]" pageId="64" pageNumber="65" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff 1924: 26</bibRefCitation>
</treatmentCitation>
, 68, 135, 142 (key, distribution). —
<treatmentCitation id="1458E161301A5224E830FCFB766A02A7" author="Pessoa S. B." page="296" pageId="64" pageNumber="65" year="1934">
<bibRefCitation id="F168BA81301A5224E830FCFB766A02A7" author="Pessoa S. B." pageId="64" pageNumber="65" pagination="279 - 314" refId="ref135808" refString="Pessoa S. B. 1934. Contribuicao para o conhecimento das especies brasileiras do genero Phanaeus. Annales da Faculdade de Medicina da Universidade de Sao Paulo 10 (3): 279 - 314. https: // doi. org / 10.5962 / bhl. title. 98039" type="journal article" year="1934">Pessôa 1934: 296</bibRefCitation>
</treatmentCitation>
, 299 (key, redescription). —
<treatmentCitation id="1458E161301A5224EFD1FCD874B302A6" author="Martinez A." box="[710,959,894,921]" page="114" pageId="64" pageNumber="65" year="1947">
<bibRefCitation id="F168BA81301A5224EFD1FCD874B302A6" author="Martinez A." box="[710,959,894,921]" pageId="64" pageNumber="65" pagination="109 - 114" refId="ref129500" refString="Martinez A. 1947 b. Addenda y corrigenda al trabajo de Blackwelder &quot; Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America &quot; (Scarabaeidae, Coprinae). Arthropoda 1 (1): 109 - 114." type="journal article" year="1947">Martínez 1947b: 114</bibRefCitation>
</treatmentCitation>
(cited). —
<treatmentCitation id="1458E161301A5224E95BFCD976140282" author="Vulcano M. A. &amp; Pereira F. S." page="572" pageId="64" pageNumber="65" year="1967">
<bibRefCitation id="F168BA81301A5224E95BFCD976140282" author="Vulcano M. A. &amp; Pereira F. S." pageId="64" pageNumber="65" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 572</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161301A5224EC86FC05756A0283" author="Krajcik M." box="[401,614,930,956]" page="204" pageId="64" pageNumber="65" year="2012">
<bibRefCitation id="F168BA81301A5224EC86FC05756A0283" author="Krajcik M." box="[401,614,930,956]" pageId="64" pageNumber="65" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 204</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161301A5224EFCFFC0574F30283" author="Gamez J. &amp; Mora E." box="[728,1023,930,956]" page="17" pageId="64" pageNumber="65" year="2000">
<bibRefCitation id="F168BA81301A5224EFCFFC0574F30283" author="Gamez J. &amp; Mora E." box="[728,1023,930,956]" pageId="64" pageNumber="65" pagination="3 - 16" refId="ref120147" refString="Gamez J. &amp; Mora E. 2000. Aspectos autoecologicos y distribucion local de Phanaeus (Phanaeus) prasinus Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae) en Venezuela. Memorias de la Fundacion La Salle de Ciencias Naturales 60 (153): 3 - 16." type="journal article" year="2000">Gámez &amp; Mora 2000: 17</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770301A5224EDAAFC6272450553" blockId="64.[189,1399,683,1696]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="B5E9E05E301A5224EDAAFC6272450553" pageId="64" pageNumber="65">
<taxonomicName id="52F9BCF3301A5224EDAAFC62744802E0" ID-CoL="7ZMQ3" box="[189,836,965,991]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subSpecies" species="telamon" subGenus="Coprophanaeus" subSpecies="telamon">
<emphasis id="A78D1B62301A5224EDAAFC62767B02E0" box="[189,375,965,991]" italics="true" pageId="64" pageNumber="65">Coprophanaeus</emphasis>
(
<emphasis id="A78D1B62301A5224EC80FC62755D02E0" box="[407,593,965,991]" italics="true" pageId="64" pageNumber="65">Coprophanaeus</emphasis>
)
<emphasis id="A78D1B62301A5224EF66FC62744802E0" box="[625,836,965,991]" italics="true" pageId="64" pageNumber="65">telamon telamon</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161301A5224EE69FC62722A02E0" author="Pereira F. S. &amp; Martinez A." box="[894,1318,965,992]" page="233" pageId="64" pageNumber="65" year="1956">
<bibRefCitation id="F168BA81301A5224EE69FC62722A02E0" author="Pereira F. S. &amp; Martinez A." box="[894,1318,965,992]" pageId="64" pageNumber="65" pagination="229 - 240" refId="ref135426" refString="Pereira F. S. &amp; Martinez A. 1956 b. Algunas notas sinonimicas en Phanaeini. Revista Brasileira de Entomologia 6: 229 - 240." type="journal article" year="1956">Pereira &amp; Martínez 1956b: 233</bibRefCitation>
</treatmentCitation>
(new combination). —
<treatmentCitation id="1458E161301A5224ECAEFC4F7576053D" author="Blanco J." box="[441,634,1000,1026]" page="43" pageId="64" pageNumber="65" year="1988">
<bibRefCitation id="F168BA81301A5224ECAEFC4F7576053D" author="Blanco J." box="[441,634,1000,1026]" pageId="64" pageNumber="65" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 43</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161301A5224EE2EFC4F7314053D" author="Havranek D." box="[825,1048,1000,1026]" page="61" pageId="64" pageNumber="65" year="1989">
<bibRefCitation id="F168BA81301A5224EE2EFC4F7314053D" author="Havranek D." box="[825,1048,1000,1026]" pageId="64" pageNumber="65" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161301A5224E99DFC4F76490519" author="Martinez A. &amp; Martinez A." page="7" pageId="64" pageNumber="65" year="1990">
<bibRefCitation id="F168BA81301A5224E99DFC4F76490519" author="Martinez A. &amp; Martinez A." pageId="64" pageNumber="65" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo, 1990: 7</bibRefCitation>
</treatmentCitation>
(notes of Phanaeina). —
<treatmentCitation id="1458E161301A5224EF6DFBAC745F0519" author="Arnaud P." box="[634,851,1035,1062]" page="35" pageId="64" pageNumber="65" year="2002">
<bibRefCitation id="F168BA81301A5224EF6DFBAC745F0519" author="Arnaud P." box="[634,851,1035,1062]" pageId="64" pageNumber="65" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud 2002b: 35</bibRefCitation>
</treatmentCitation>
(monograph, redescription). —
<treatmentCitation id="1458E161301A5224E9C0FBAC76050576" author="Gamez J." page="59" pageId="64" pageNumber="65" year="2004">
<bibRefCitation id="F168BA81301A5224E9C0FBAC76050576" author="Gamez J." pageId="64" pageNumber="65" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004: 59</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E161301A5224ECFDFB89745D0576" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[490,849,1070,1097]" page="17" pageId="64" pageNumber="65" year="2006">
<bibRefCitation id="F168BA81301A5224ECFDFB89745D0576" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[490,849,1070,1097]" pageId="64" pageNumber="65" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B62301A5224EF8AFB8875D70577" box="[669,731,1070,1096]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2006: 17
</bibRefCitation>
</treatmentCitation>
(list);
<treatmentCitation id="1458E161301A5224EEB0FB88731E0576" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[935,1042,1070,1097]" page="57" pageId="64" pageNumber="65" year="2009">
<bibRefCitation id="F168BA81301A5224EEB0FB88731E0576" author="Hamel-Leigue A. &amp; Herzog S. K. &amp; Mann D. J. &amp; Larsen T. H. &amp; Gill B. D. &amp; Edmonds W. D. &amp; Spector S." box="[935,1042,1070,1097]" pageId="64" pageNumber="65" pagination="43 - 95" refId="ref122950" refString="Hamel-Leigue A., Herzog S. K., Mann D. J., Larsen T. H., Gill B. D., Edmonds W. D., Spector S. 2009. Distribucion e historia natural de escarabajos coprofagos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">2009: 57</bibRefCitation>
</treatmentCitation>
(distribution for
<collectingCountry id="EDEE87E0301A5224E9F0FB89724F0577" box="[1255,1347,1070,1096]" name="Bolivia" pageId="64" pageNumber="65">Bolivia</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EDFDFBF575420553" author="Gamez J. &amp; Acconcia R." box="[234,590,1106,1132]" page="394" pageId="64" pageNumber="65" year="2009">
<bibRefCitation id="F168BA81301A5224EDFDFBF575420553" author="Gamez J. &amp; Acconcia R." box="[234,590,1106,1132]" pageId="64" pageNumber="65" pagination="387 - 396" refId="ref119925" refString="Gamez J. &amp; Acconcia R. 2009. Informaciones ecologicas sobre Coprophanaeus (Coprophanaeus) gamezi Arnaud (Coleoptera: Scarabaeinae: Phanaeini) en un sistema agropastoril en la depresion de Maracaibo, estado Zulia, Venezuela. Acta Zoologica Mexicana (n. s.) 25 (2): 387 - 396. https: // doi. org / 10.21829 / azm. 2009.252645" type="journal article" year="2009">Gámez &amp; Acconcia 2009: 394</bibRefCitation>
</treatmentCitation>
(ecological information). —
<treatmentCitation id="1458E161301A5224EEB1FBF573C70553" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[934,1227,1106,1132]" page="318" pageId="64" pageNumber="65" year="2011">
<bibRefCitation id="F168BA81301A5224EEB1FBF573C70553" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[934,1227,1106,1132]" pageId="64" pageNumber="65" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B62301A5224E906FBF473400553" box="[1041,1100,1106,1132]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2011: 318
</bibRefCitation>
</treatmentCitation>
, 320 (list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770301A5224EDAAFBD2733A07A0" blockId="64.[189,1399,683,1696]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="B5E9E05E301A5224EDAAFBD2733A07A0" pageId="64" pageNumber="65">
<taxonomicName id="52F9BCF3301A5224EDAAFBD275BD05B0" ID-CoL="7ZMQ3" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[189,689,1141,1167]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="telamon" subGenus="Coprophanaeus">
<emphasis id="A78D1B62301A5224EDAAFBD2767B05B0" box="[189,375,1141,1167]" italics="true" pageId="64" pageNumber="65">Coprophanaeus</emphasis>
(
<emphasis id="A78D1B62301A5224EC9FFBD2754E05B0" box="[392,578,1141,1167]" italics="true" pageId="64" pageNumber="65">Coprophanaeus</emphasis>
)
<emphasis id="A78D1B62301A5224EF45FBD275BD05B0" box="[594,689,1141,1167]" italics="true" pageId="64" pageNumber="65">telamon</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161301A5224EFDAFBD274EF05B0" author="Forsyth A. B. &amp; Spector S. &amp; Gill B. &amp; Guerra F. &amp; Ayzama S." box="[717,995,1141,1167]" page="370" pageId="64" pageNumber="65" year="1998">
<bibRefCitation id="F168BA81301A5224EFDAFBD274EF05B0" author="Forsyth A. B. &amp; Spector S. &amp; Gill B. &amp; Guerra F. &amp; Ayzama S." box="[717,995,1141,1167]" pageId="64" pageNumber="65" refId="ref119575" refString="Forsyth A. B., Spector S., Gill B., Guerra F. &amp; Ayzama S. 1998. Dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Parque Nacinal Noel Kempff Mercado / Escarabajos (Coleoptera: Scarabaeidae: Scarabaeinae) del Parque Nacional Noel Kempff Mercado. In: Killeen T. J. &amp; Schulenberg T. S. (eds) Rapid Assessment Program. A Biological Assessment of Parque Nacional Noel Kempff Mercado, Bolivia: 181 - 200 + appendix 6. RAP working papers, Conservation International, Washington, D. C." type="journal volume" year="1998">
Forsyth
<emphasis id="A78D1B62301A5224EE3AFBD1746B05B0" box="[813,871,1141,1167]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
1998: 370
</bibRefCitation>
</treatmentCitation>
(conservation, list). —
<treatmentCitation id="1458E161301A5224E9E2FBD276A6058D" author="Amezquita S. J. &amp; Forsyth A. &amp; Lopera A. &amp; Camacho A." page="119" pageId="64" pageNumber="65" year="1999">
<bibRefCitation id="F168BA81301A5224E9E2FBD276A6058D" author="Amezquita S. J. &amp; Forsyth A. &amp; Lopera A. &amp; Camacho A." pageId="64" pageNumber="65" pagination="113 - 126" refId="ref109271" refString="Amezquita S. J., Forsyth A., Lopera A. &amp; Camacho A. 1999. Comparacion de la composicion y riqueza de especies de escarabajos coprofagos (Coleoptera: Scarabaeidae) en remanentes de bosque de la Orinoquia Colombiana. Acta Zoologica Mexicana (n. s.) 76: 113 - 126. https: // doi. org / 10.21829 / azm. 1999.76761703" type="journal article" year="1999">
Amézquita
<emphasis id="A78D1B62301A5224EDFDFB3E7624058D" box="[234,296,1176,1202]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
1999: 119
</bibRefCitation>
</treatmentCitation>
(biodiversity). —
<treatmentCitation id="1458E161301A5224EF9BFB3F7461058D" author="Escobar F." box="[652,877,1176,1202]" page="207" pageId="64" pageNumber="65" year="2000">
<bibRefCitation id="F168BA81301A5224EF9BFB3F7461058D" author="Escobar F." box="[652,877,1176,1202]" pageId="64" pageNumber="65" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 207</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0301A5224E937FB3F7395058D" box="[1056,1177,1176,1202]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E9C2FB3F766905EA" author="Vaz-de-Mello F. Z." page="192" pageId="64" pageNumber="65" year="2000">
<bibRefCitation id="F168BA81301A5224E9C2FB3F766905EA" author="Vaz-de-Mello F. Z." pageId="64" pageNumber="65" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 192</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0301A5224EF0BFB1C756505EA" box="[540,617,1211,1237]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EFBFFB1C747505E9" author="Vitolo A. L." box="[680,889,1211,1238]" page="599" pageId="64" pageNumber="65" year="2000">
<bibRefCitation id="F168BA81301A5224EFBFFB1C747505E9" author="Vitolo A. L." box="[680,889,1211,1238]" pageId="64" pageNumber="65" pagination="591 - 601" refId="ref141219" refString="Vitolo A. L. 2000. Clave para la identificacion de los generos y especies Phanaeinas de Colombia. Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales 24 (93): 591 - 601." type="journal article" year="2000">Vítolo 2000: 599</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="1458E161301A5224EEC1FB1C735D05EA" author="Vitolo A." box="[982,1105,1211,1237]" page="290" pageId="64" pageNumber="65" year="2004">
<bibRefCitation id="F168BA81301A5224EEC1FB1C735D05EA" author="Vitolo A." box="[982,1105,1211,1237]" pageId="64" pageNumber="65" pagination="277 - 318" refId="ref141259" refString="Vitolo A. 2004. Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae). In: Fernandez F., Andrade M. G. &amp; Amat G. D. (eds) Insectos de Colombia 3: 277 - 318. Facultad de Ciencias, Universidad Nacional de Colombia, Bogota." type="book chapter" year="2004">2004: 290</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution, cited for
<collectingCountry id="EDEE87E0301A5224EC44FB7976C005C7" box="[339,460,1246,1272]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EF14FB7975CC05C7" author="Barbero E." box="[515,704,1246,1273]" page="6" pageId="64" pageNumber="65" year="2001">
<bibRefCitation id="F168BA81301A5224EF14FB7975CC05C7" author="Barbero E." box="[515,704,1246,1273]" pageId="64" pageNumber="65" pagination="11 - 21" refId="ref110583" refString="Barbero E. 2001. Scarabaeidae (Coleoptera) copronecrofagos interesantes del Departamento de Rio San Juan, Nicaragua. Revista Nicaraguense de Entomologia 55 - 58: 11 - 21." type="book chapter" year="2001">Barbero 2001: 6</bibRefCitation>
</treatmentCitation>
(note for
<collectingCountry id="EDEE87E0301A5224EE24FB7974BD05C7" box="[819,945,1246,1272]" name="Nicaragua" pageId="64" pageNumber="65">Nicaragua</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EEF0FB7973F205C6" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[999,1278,1246,1273]" page="140" pageId="64" pageNumber="65" year="2001">
<bibRefCitation id="F168BA81301A5224EEF0FB7973F205C6" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[999,1278,1246,1273]" pageId="64" pageNumber="65" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B62301A5224E950FB78738D05C7" box="[1095,1153,1246,1272]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2001: 140
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0301A5224EC0FFAA5769D0423" box="[280,401,1282,1308]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224ECC5FAA574450423" author="Halffter G. &amp; Arellano L." box="[466,841,1282,1308]" page="148" pageId="64" pageNumber="65" year="2002">
<bibRefCitation id="F168BA81301A5224ECC5FAA574450423" author="Halffter G. &amp; Arellano L." box="[466,841,1282,1308]" pageId="64" pageNumber="65" pagination="144 - 154" refId="ref122390" refString="Halffter G. &amp; Arellano L. 2002. Response of dung beetle diversity to human-induced changes in a tropical landscape. Biotropica 34 (1): 144 - 154. https: // doi. org / 10.1111 / j. 1744 - 7429.2002. tb 00250. x" type="journal article" year="2002">Halffter &amp; Arellano 2002: 148</bibRefCitation>
</treatmentCitation>
(ecology). —
<treatmentCitation id="1458E161301A5224E914FAA5727A0422" author="Pulido-Herrera L. A. &amp; Canas R. A. R. &amp; Harders F. G. &amp; von Hildebrand P." box="[1027,1398,1282,1309]" page="54" pageId="64" pageNumber="65" year="2003">
<bibRefCitation id="F168BA81301A5224E914FAA5727A0422" author="Pulido-Herrera L. A. &amp; Canas R. A. R. &amp; Harders F. G. &amp; von Hildebrand P." box="[1027,1398,1282,1309]" pageId="64" pageNumber="65" refId="ref136416" refString="Pulido-Herrera L. A., Canas R. A. R., Harders F. G. &amp; von Hildebrand P. 2003. Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) del Parque Nacional Natural &quot; Serrania de Chiribiquete &quot;, Caqueta, Colombia (Parte I). In: Onore G., Reyes-Castillo P. &amp; Zunino M. (eds) Escarabeidos de Latinoamerica: Estado del conocimiento: 51 - 58. m 3 m: Monografias Tercer Milenio Vol. 3, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2003">
Pulido-Herrera
<emphasis id="A78D1B62301A5224E9A8FAA473F20423" box="[1215,1278,1282,1308]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2003: 54
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</treatmentCitation>
(list for
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,
<collectingCountry id="EDEE87E0301A5224ECD7FA8275350400" box="[448,569,1317,1343]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
);
<treatmentCitation id="1458E161301A5224EF47FA8275C50400" author="Pulido-Herrera L. A. &amp; Canas R. A. R. &amp; Harders F. G. &amp; von Hildebrand P." box="[592,713,1317,1343]" page="307" pageId="64" pageNumber="65" year="2007">
<bibRefCitation id="F168BA81301A5224EF47FA8275C50400" author="Pulido-Herrera L. A. &amp; Medina C. A. &amp; Riveros R. A." box="[592,713,1317,1343]" pageId="64" pageNumber="65" pagination="305 - 310" refId="ref136514" refString="Pulido-Herrera L. A., Medina C. A. &amp; Riveros R. A. 2007. Nuevos registros de escarabajos coprofagos (Scarabaeidae: Scarabaeinae) para la region andina de Colombia. Parte I. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 31: 305 - 310. https: // doi. org / 10.18257 / raccefyn. 31 (119). 2007.2338" type="journal article" year="2007">2007: 307</bibRefCitation>
</treatmentCitation>
(cited for Andean region of
<collectingCountry id="EDEE87E0301A5224E93FFA8273AD0400" box="[1064,1185,1317,1343]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E9C8FA827668045D" author="Carpio C. &amp; Donoso D. A. &amp; Ramon G. &amp; Dangles O." page="469" pageId="64" pageNumber="65" year="2009">
<bibRefCitation id="F168BA81301A5224E9C8FA827668045D" author="Carpio C. &amp; Donoso D. A. &amp; Ramon G. &amp; Dangles O." pageId="64" pageNumber="65" pagination="455 - 469" refId="ref113707" refString="Carpio C., Donoso D. A., Ramon G. &amp; Dangles O. 2009. Short term response of dung beetle communities to disturbance by road construction in the Ecuadorian Amazon. Annales de la Societe entomologique de France 45 (4): 455 - 469. https: // doi. org / 10.1080 / 00379271.2009.10697629" type="journal article" year="2009">
Carpio
<emphasis id="A78D1B62301A5224E82EFA81727B0400" box="[1337,1399,1317,1343]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2009: 469
</bibRefCitation>
</treatmentCitation>
(ecology). —
<treatmentCitation id="1458E161301A5224EF0FFAEF74CD045D" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[536,961,1352,1378]" page="61" pageId="64" pageNumber="65" year="2009">
<bibRefCitation id="F168BA81301A5224EF0FFAEF74CD045D" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[536,961,1352,1378]" pageId="64" pageNumber="65" pagination="55 - 62" refId="ref130761" refString="Medina C. A. &amp; Pulido-Herrera L. A. 2009. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de la Orinoquia Colombiana. Biota Colombiana 10 (1 - 2): 55 - 62." type="journal article" year="2009">Medina &amp; Pulido-Herrera 2009: 61</bibRefCitation>
</treatmentCitation>
(diversity). —
<treatmentCitation id="1458E161301A5224E968FAEF7246045D" author="Gamez J." box="[1151,1354,1352,1378]" page="16" pageId="64" pageNumber="65" year="2010">
<bibRefCitation id="F168BA81301A5224E968FAEF7246045D" author="Gamez J." box="[1151,1354,1352,1378]" pageId="64" pageNumber="65" pagination="15 - 19" refId="ref119876" refString="Gamez J. 2010. Escarabajos necrofagos (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini y Coleoptera: Silphidae: Necrophorinae y Silphinae) de la Cordillera de Los Andes, Depresion de Maracaibo y Llanos de Venezuela. Kempffiana 6 (2): 15 - 19." type="journal article" year="2010">Gámez 2010: 16</bibRefCitation>
</treatmentCitation>
, 19 (comments). —
<treatmentCitation id="1458E161301A5224ECBCFACC75FB04B9" author="Edmonds W. D. &amp; Zidek J." box="[427,759,1387,1414]" page="77" pageId="64" pageNumber="65" year="2010">
<bibRefCitation id="F168BA81301A5224ECBCFACC75FB04B9" author="Edmonds W. D. &amp; Zidek J." box="[427,759,1387,1414]" pageId="64" pageNumber="65" pagination="1 - 111" refId="ref117660" refString="Edmonds W. D. &amp; Zidek J. 2010. A taxonomic review of the Neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae). Insecta Mundi 129: 1 - 111." type="journal article" year="2010">Edmonds &amp; Zídek 2010: 77</bibRefCitation>
</treatmentCitation>
, 91 (key, diagnosis). —
<treatmentCitation id="1458E161301A5224E900FACC723204BA" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[1047,1342,1387,1413]" page="318" pageId="64" pageNumber="65" year="2011">
<bibRefCitation id="F168BA81301A5224E900FACC723204BA" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[1047,1342,1387,1413]" pageId="64" pageNumber="65" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B62301A5224E994FACB73B204BA" box="[1155,1214,1387,1413]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2011: 318
</bibRefCitation>
</treatmentCitation>
, 320 (list). —
<treatmentCitation id="1458E161301A5224EC45FA2975A70497" author="Vaz-de-Mello F. Z. &amp; Silva R. L. R. &amp; Nunes L. G. O. A. &amp; Correa P. R. O. A." box="[338,683,1422,1448]" page="86" pageId="64" pageNumber="65" year="2011">
<bibRefCitation id="F168BA81301A5224EC45FA2975A70497" author="Vaz-de-Mello F. Z. &amp; Silva R. L. R. &amp; Nunes L. G. O. A. &amp; Correa P. R. O. A." box="[338,683,1422,1448]" pageId="64" pageNumber="65" pagination="77 - 102" refId="ref140870" refString="Vaz-de-Mello F. Z., Silva R. L. R., Nunes L. G. O. A. &amp; Correa P. R. O. A. 2011 b. Os besouros rola-bosta (Insecta: Coleoptera: Scarabaeidae: Scarabaeinae) da Fazenda Sao Nicolau. In: Rodrigues D. J., Izzo T. J. &amp; Battirola L. D. (eds.) Descobrindo a Amazonica Meridional: Biodiversidade da Fazenda Sao Nicolau: 77 - 102. Pau e Prosa Comunicacao, Cuiaba." type="book chapter" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B62301A5224ECE0FA28753C0497" box="[503,560,1422,1448]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2011b: 86
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161301A5224EE0DFA29737D0496" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[794,1137,1422,1449]" page="57" pageId="64" pageNumber="65" year="2012">
<bibRefCitation id="F168BA81301A5224EE0DFA29737D0496" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[794,1137,1422,1449]" pageId="64" pageNumber="65" refId="ref114949" refString="Cultid-Medina C. A., Medina C. A. U., Martinez-Quintero B. G., Escobar A. F. V., Constantino L. M. C. &amp; Betancur N. J. 2012. Escarabajos Coprofagos (Scarabaeinae) del Eje Cafetero: Guia para el Estudio Ecologico. WCS Colombia, Cenicafe, Villa Maria, Caldas." type="book" year="2012">
Cultid-Medina
<emphasis id="A78D1B62301A5224EEDCFA2873080497" box="[971,1028,1422,1448]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2012: 57
</bibRefCitation>
</treatmentCitation>
(guide). —
<treatmentCitation id="1458E161301A5224E9EAFA29751504F2" ID-DOI="http://doi.org/10.5281/zenodo.6146741" ID-Zenodo-Dep="6146741" author="Cupello M. &amp; Vaz-de-Mello F. Z." httpUri="http://treatment.plazi.org/id/627E87F5196FA45AFF49FDA4FEEAFD28" page="358" pageId="64" pageNumber="65" year="2013">
<bibRefCitation id="F168BA81301A5224E9EAFA29751504F2" author="Cupello M. &amp; Vaz-de-Mello F. Z." pageId="64" pageNumber="65" pagination="359 - 368" refId="ref115382" refString="Cupello M. &amp; Vaz-de-Mello F. Z. 2013 a. New evidence for the validity of Coprophanaeus (C.) terrali Arnaud, 2002 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini), a dung beetle from Brazil. Zootaxa, 3717 (3): 359 - 368. https: // doi. org / 10.11646 / zootaxa. 3717.3.5" type="journal article" year="2013">Cupello &amp; Vaz-de-Mello 2013a: 358</bibRefCitation>
</treatmentCitation>
, 361364, 367 (comments, distribution). —
<treatmentCitation id="1458E161301A5224E93EFA15727A04F3" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1065,1398,1458,1484]" page="109" pageId="64" pageNumber="65" year="2013">
<bibRefCitation id="F168BA81301A5224E93EFA15727A04F3" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1065,1398,1458,1484]" pageId="64" pageNumber="65" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62301A5224E9AAFA1473F404F3" box="[1213,1272,1458,1484]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161301A5224EC40FA72758E04D0" author="Figueroa L. &amp; Edmonds W. D. &amp; Martinez N. L." box="[343,642,1493,1519]" page="127" pageId="64" pageNumber="65" year="2014">
<bibRefCitation id="F168BA81301A5224EC40FA72758E04D0" author="Figueroa L. &amp; Edmonds W. D. &amp; Martinez N. L." box="[343,642,1493,1519]" pageId="64" pageNumber="65" pagination="125 - 138" refId="ref119373" refString="Figueroa L., Edmonds W. D., Martinez N. L. 2014. La tribu Phanaeini (Coleoptera, Scarabaeidae, Scarabaeinae) en el Peru. Revista Peruana de Biologia 21 (2): 125 - 138. https: // doi. org / 10.15381 / rpb. v 21 i 2.9815" type="journal article" year="2014">
Figueroa
<emphasis id="A78D1B62301A5224ECDFFA71750F04D0" box="[456,515,1493,1519]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2014: 127
</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<treatmentCitation id="1458E161301A5224EE99FA72724D04D0" author="Pardo-Locarno L. C. &amp; Camero E." box="[910,1345,1493,1519]" page="211" pageId="64" pageNumber="65" year="2014">
<bibRefCitation id="F168BA81301A5224EE99FA72724D04D0" author="Pardo-Locarno L. C. &amp; Camero E." box="[910,1345,1493,1519]" pageId="64" pageNumber="65" pagination="207 - 224" refId="ref134721" refString="Pardo-Locarno L. C. &amp; Camero E. 2014. Escarabajos coprofagos (Coleptera Scarabaeinae) en siete microcuencas del rio Dagua, Choco biogeografico de Colombia. Boletin de la SEA 54: 207 - 224." type="journal article" year="2014">Pardo-Locarno &amp; Camero 2014: 211</bibRefCitation>
</treatmentCitation>
(list for Chocó,
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). —
<treatmentCitation id="1458E161301A5224EF3EFA5F7421072D" author="Silva R. J. &amp; Coletti F. &amp; Costa D. A. &amp; Vaz-De-Mello F. Z." box="[553,813,1528,1554]" page="348" pageId="64" pageNumber="65" year="2014">
<bibRefCitation id="F168BA81301A5224EF3EFA5F7421072D" author="Silva R. J. &amp; Coletti F. &amp; Costa D. A. &amp; Vaz-De-Mello F. Z." box="[553,813,1528,1554]" pageId="64" pageNumber="65" refId="ref138928" refString="Silva R. J., Coletti F., Costa D. A. &amp; Vaz-De-Mello F. Z. 2014. Rola-bostas (Coleoptera: Scarabaeidae: Scarabaeinae) de florestas e pastagens no sudoeste da Amazonia brasileira: Levantamento de especies e guildas alimentares. Acta Amazonica 44 (3): 345 - 352. https: // doi. org / 10.1590 / 1809 - 4392201304472 Silva R. J., Storck-Tonon D. &amp; Vaz-de-Mello F. Z. 2016. Dung beetle (Coleoptera: Scarabaeinae) persistence in Amazonian forest fragments and adjacent pastures: biogeographic implications for alpha and beta diversity. Journal of Insect Conservation 20 (4): 549 - 564. https: // doi. org / 10.1007 / s 10841 - 016 - 9885 - 7" type="book" year="2014">
Silva
<emphasis id="A78D1B62301A5224EF66FA5E75A0072D" box="[625,684,1528,1554]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2014: 348
</bibRefCitation>
</treatmentCitation>
(diversity);
<treatmentCitation id="1458E161301A5224EED4FA5F7312072C" author="Silva R. J. &amp; Pelissari T. D. &amp; Krinski D. &amp; Canale G. &amp; Vaz-de-Mello F. Z." box="[963,1054,1528,1555]" page="7" pageId="64" pageNumber="65" year="2016">
<bibRefCitation id="F168BA81301A5224EED4FA5F7312072C" author="Silva R. J. &amp; Storck-Tonon D. &amp; Vaz-de-Mello F. Z." box="[963,1054,1528,1555]" journalOrPublisher="Journal of Insect Conservation" pageId="64" pageNumber="65" pagination="549 - 564" part="20" publicationUrl="https://doi.org/10.1007/s10841-016-9885-7" refId="ref138928" refString="Silva R. J., Storck-Tonon D. &amp; Vaz-de-Mello F. Z. 2016. Dung beetle (Coleoptera: Scarabaeinae) persistence in Amazonian forest fragments and adjacent pastures: biogeographic implications for alpha and beta diversity. Journal of Insect Conservation 20 (4): 549 - 564. https: // doi. org / 10.1007 / s 10841 - 016 - 9885 - 7" title="Dung beetle (Coleoptera: Scarabaeinae) persistence in Amazonian forest fragments and adjacent pastures: biogeographic implications for alpha and beta diversity" type="journal article" year="2016">2016: 7</bibRefCitation>
</treatmentCitation>
(ecology, biogeography). —
<treatmentCitation id="1458E161301A5224EDFAF9BC7503070A" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[237,527,1563,1590]" page="197" pageId="64" pageNumber="65" year="2015">
<bibRefCitation id="F168BA81301A5224EDFAF9BC7503070A" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[237,527,1563,1590]" pageId="64" pageNumber="65" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B62301A5224EC4FF9BB769E070A" box="[344,402,1563,1589]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2015: 197
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0301A5224EFAEF9BC75F8070A" box="[697,756,1563,1589]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224EE3BF9BC735F070A" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[812,1107,1563,1589]" page="93" pageId="64" pageNumber="65" year="2018">
<bibRefCitation id="F168BA81301A5224EE3BF9BC735F070A" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[812,1107,1563,1589]" pageId="64" pageNumber="65" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B62301A5224EEBDF9BB74E9070A" box="[938,997,1563,1589]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2018: 93
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E0301A5224E9AEF9BC722C070A" box="[1209,1312,1563,1589]" name="Ecuador" pageId="64" pageNumber="65">Ecuador</collectingCountry>
);
<treatmentCitation id="1458E161301A5224E823F9BC760B0766" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." httpUri="http://treatment.plazi.org/id/A5BF9328A0E831B5D875FC48E5D5CDB5" page="77" pageId="64" pageNumber="65" year="2019">
<bibRefCitation id="F168BA81301A5224E823F9BC760B0766" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." pageId="64" pageNumber="65" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 77</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161301A5224ECD3F99874140766" author="Gamez J. &amp; Acconcia R." box="[452,792,1598,1625]" page="65" pageId="64" pageNumber="65" year="2018">
<bibRefCitation id="F168BA81301A5224ECD3F99874140766" author="Gamez J. &amp; Acconcia R." box="[452,792,1598,1625]" pageId="64" pageNumber="65" pagination="65 - 75" refId="ref120046" refString="Gamez J. &amp; Acconcia R. 2018. Escarabajos coprofagos y necrofagos (Scarabaeidae: Scarabaeinae: Phanaeini) del flanco andino llanero, piedemonte y llanos, com el reporte de nuevos registros para los estados de Barina y Merida (Venezuela). Revista Gaditana de Entomologia 9 (1): 65 - 75." type="journal article" year="2018">Gámez &amp; Acconcia 2018: 65</bibRefCitation>
</treatmentCitation>
, 69, 73 (comments, cited for
<collectingCountry id="EDEE87E0301A5224E97CF99873AA0766" box="[1131,1190,1599,1625]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E9CBF999765F0743" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." page="70" pageId="64" pageNumber="65" year="2018">
<bibRefCitation id="F168BA81301A5224E9CBF999765F0743" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." pageId="64" pageNumber="65" refId="ref121556" refString="Giraldo C., Montoya S. &amp; Escobar F. 2018. Escarabajos del Estiercol en Paisajes Ganaderos de Colombia. Fundacion CIPAV, Cali, Colombia." type="book" year="2018">
Giraldo
<emphasis id="A78D1B62301A5224E82BF998727A0767" box="[1340,1398,1598,1624]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2018: 70
</bibRefCitation>
</treatmentCitation>
(guide). —
<treatmentCitation id="1458E161301A5224ECFFF9C575E20743" author="Santos R. &amp; Sutil W. P. &amp; de Oliveira J. F. A." box="[488,750,1634,1660]" page="46" pageId="64" pageNumber="65" year="2018">
<bibRefCitation id="F168BA81301A5224ECFFF9C575E20743" author="Santos R. &amp; Sutil W. P. &amp; de Oliveira J. F. A." box="[488,750,1634,1660]" pageId="64" pageNumber="65" pagination="43 - 48" refId="ref138029" refString="Santos R. S, Sutil W. P. &amp; de Oliveira J. F. A. 2018. Besouros escarabeideos (Coleoptera: Scarabaeidae) coletados em remanescente florestal em Rio Branco, Acre, Brasil. Agrotropica 30 (1): 43 - 48. https: // doi. org / 10.21757 / 0103 - 3816.2018 v 30 n 1 p 43 - 48. pdf" type="journal article" year="2018">
Santos
<emphasis id="A78D1B62301A5224EF55F9C475720743" box="[578,638,1634,1660]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2018: 46
</bibRefCitation>
</treatmentCitation>
(list for
<collectingRegion id="573D0992301A5224EE4FF9C574D80743" box="[856,980,1634,1660]" country="Brazil" name="Acre" pageId="64" pageNumber="65">Acre state</collectingRegion>
,
<collectingCountry id="EDEE87E0301A5224EEF6F9C573220743" box="[993,1070,1634,1660]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E161301A5224E97EF9C5765F079F" ID-DOI="http://doi.org/10.5281/zenodo.3683815" ID-Zenodo-Dep="3683815" author="Hielkema A. J. &amp; Hielkema M. A." httpUri="http://treatment.plazi.org/id/03D7FF1CFFABFF8CFF48FDB53395FDF6" page="95" pageId="64" pageNumber="65" year="2019">
<bibRefCitation id="F168BA81301A5224E97EF9C5765F079F" author="Hielkema A. J. &amp; Hielkema M. A." pageId="64" pageNumber="65" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 95</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas). —
<treatmentCitation id="1458E161301A5224EFD9F92274C107A0" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[718,973,1669,1695]" page="136" pageId="64" pageNumber="65" year="2020">
<bibRefCitation id="F168BA81301A5224EFD9F92274C107A0" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[718,973,1669,1695]" pageId="64" pageNumber="65" pagination="134 - 141" refId="ref132145" refString="Nieto M. J. A., Giraldo E. C., Quevedo V. C. J., Chara J. &amp; Medina A. C. 2020. Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Biota Colombiana 21: 134 - 141. https: // doi. org / 10.21068 / c 2020. v 21 n 02 a 09" type="journal article" year="2020">
Nieto
<emphasis id="A78D1B62301A5224EE02F921745C07A0" box="[789,848,1669,1695]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2020: 136
</bibRefCitation>
</treatmentCitation>
(report).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB301A5224EDAAF96276CC064C" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="9546C770301A5224EDAAF96276A607E0" blockId="64.[189,426,1733,1759]" box="[189,426,1733,1759]" pageId="64" pageNumber="65">
<heading id="CE0E701C301A5224EDAAF96276A607E0" bold="true" box="[189,426,1733,1759]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A78D1B62301A5224EDAAF96276A607E0" bold="true" box="[189,426,1733,1759]" pageId="64" pageNumber="65">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770301A5224EDAAF95476CC064C" blockId="64.[187,1398,1778,1907]" pageId="64" pageNumber="65">
<materialsCitation id="2591CD2D301A5224EDAAF95473E90632" box="[189,1253,1778,1806]" collectingDate="2004-07-06" collectionCode="CEMT" collectorName="D. Garcia" country="Venezuela" elevation="50" location="Cuyagua" pageId="64" pageNumber="65" specimenCount="1" stateProvince="Aragua">
<collectingCountry id="EDEE87E0301A5224EDAAF95476670632" box="[189,363,1779,1805]" name="Venezuela" pageId="64" pageNumber="65">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62301A5224EC92F95576E80633" bold="true" box="[389,484,1778,1804]" pageId="64" pageNumber="65">
<collectingRegion id="573D0992301A5224EC92F95576E80633" box="[389,484,1778,1804]" country="Venezuela" name="Aragua" pageId="64" pageNumber="65">Aragua</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9301A5224ECEDF95475410632" box="[506,589,1779,1805]" count="1" pageId="64" pageNumber="65" type="generic">1 spec.</specimenCount>
;
<location id="902691AB301A5224EF4FF95475C80632" LSID="urn:lsid:plazi:treatment:1D507666301A5226EF08FE4574260246:902691AB301A5224EF4FF95475C80632" box="[600,708,1779,1805]" country="Venezuela" name="Cuyagua" pageId="64" pageNumber="65" stateProvince="Aragua">Cuyagua</location>
; 50 m a.s.l.;
<date id="E147E1B0301A5224EE43F95474DA0632" box="[852,982,1779,1805]" pageId="64" pageNumber="65" value="2004-07-06">
<collectingDate id="F1031858301A5224EE43F95474DA0632" box="[852,982,1779,1805]" pageId="64" pageNumber="65" value="2004-07-06">6 Jul. 2004</collectingDate>
</date>
;
<collectorName id="380CA2A6301A5224EEF6F954735E0632" box="[993,1106,1779,1805]" pageId="64" pageNumber="65">D. García</collectorName>
leg.;
<collectionCode id="F3E85FB5301A5224E985F95473E90632" box="[1170,1253,1779,1805]" pageId="64" pageNumber="65">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D301A5224E814F95573FC0610" collectingDate="1993-07" collectionCode="CEMT" collectorName="J. Blanco" country="Venezuela" location="Parque Nacional Natural Paramillo" municipality="San Cristobal" pageId="64" pageNumber="65" specimenCount="2" stateProvince="Tachira">
<emphasis id="A78D1B62301A5224E814F955726A0632" bold="true" box="[1283,1382,1778,1805]" pageId="64" pageNumber="65">
<collectingRegion id="573D0992301A5224E814F955726A0632" box="[1283,1382,1778,1805]" country="Venezuela" name="Tachira" pageId="64" pageNumber="65">Táchira</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9301A5224EDAAF8B276190610" box="[189,277,1813,1839]" count="2" pageId="64" pageNumber="65" type="generic">2 specs</specimenCount>
;
<collectingMunicipality id="75225D0A301A5224EC36F8B276CE0610" box="[289,450,1813,1839]" pageId="64" pageNumber="65">San Cristóbal</collectingMunicipality>
,
<location id="902691AB301A5224ECDAF8B274660610" LSID="urn:lsid:plazi:treatment:1D507666301A5226EF08FE4574260246:902691AB301A5224ECDAF8B274660610" box="[461,874,1813,1839]" country="Venezuela" municipality="San Cristobal" name="Parque Nacional Natural Paramillo" pageId="64" pageNumber="65" stateProvince="Tachira">Parque Nacional Natural Paramillo</location>
;
<date id="E147E1B0301A5224EE62F8B274E80610" box="[885,996,1813,1839]" pageId="64" pageNumber="65" value="1993-07">
<collectingDate id="F1031858301A5224EE62F8B274E80610" box="[885,996,1813,1839]" pageId="64" pageNumber="65" value="1993-07">Jul. 1993</collectingDate>
</date>
;
<collectorName id="380CA2A6301A5224EEF8F8B273500610" box="[1007,1116,1813,1839]" pageId="64" pageNumber="65">J. Blanco</collectorName>
leg.;
<collectionCode id="F3E85FB5301A5224E98AF8B273FC0610" box="[1181,1264,1813,1839]" pageId="64" pageNumber="65">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D301A5224E819F8B276B0064C" collectingDate="2009-07" collectionCode="CEMT" collectorName="Asmussen &amp; Colmenares &amp; Martinez" country="Venezuela" elevation="1380" latitude="10.3867" location="Aroa" longLatPrecision="7" longitude="-68.6471" pageId="64" pageNumber="65" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B62301A5224E819F8B2727A0610" bold="true" box="[1294,1398,1813,1839]" pageId="64" pageNumber="65">
<collectingRegion id="573D0992301A5224E819F8B2727A0610" box="[1294,1398,1813,1839]" country="Venezuela" name="Yaracuy" pageId="64" pageNumber="65">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9301A5224EDD8F8907628066D" box="[207,292,1847,1874]" count="1" pageId="64" pageNumber="65" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="573D0992301A5224EC27F8907686066E" box="[304,394,1847,1873]" country="Venezuela" name="Bolivar" pageId="64" pageNumber="65">Bolívar</collectingRegion>
,
<location id="902691AB301A5224EC83F89076DE066E" LSID="urn:lsid:plazi:treatment:1D507666301A5226EF08FE4574260246:902691AB301A5224EC83F89076DE066E" box="[404,466,1847,1873]" country="Venezuela" latitude="10.3867" longLatPrecision="7" longitude="-68.6471" name="Aroa" pageId="64" pageNumber="65" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B7301A5224ECC9F8907565066D" box="[478,617,1847,1874]" degrees="10.3867" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="5" value="10.3867">10.3867° N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B7301A5224EF62F8907409066D" box="[629,773,1847,1874]" degrees="68.6471" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="5" value="-68.6471">68.6471° W</geoCoordinate>
; 1380 m a.s.l.;
<date id="E147E1B0301A5224EEA0F890732A066E" box="[951,1062,1847,1873]" pageId="64" pageNumber="65" value="2009-07">
<collectingDate id="F1031858301A5224EEA0F890732A066E" box="[951,1062,1847,1873]" pageId="64" pageNumber="65" value="2009-07">Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A6301A5224E926F89073A1066E" box="[1073,1197,1847,1873]" pageId="64" pageNumber="65">Asmüssen</collectorName>
,
<collectorName id="380CA2A6301A5224E9AFF8907248066E" box="[1208,1348,1847,1873]" pageId="64" pageNumber="65">Colmenares</collectorName>
and
<collectorName id="380CA2A6301A5224EDAAF8FE762B064C" box="[189,295,1881,1907]" pageId="64" pageNumber="65">Martínez</collectorName>
leg.;
<collectionCode id="F3E85FB5301A5224EC7EF8FE76B0064C" box="[361,444,1881,1907]" pageId="64" pageNumber="65">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB301A5226EDAAF83E76FB00AB" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="distribution">
<paragraph id="9546C770301A5224EDAAF83E765A068C" blockId="64.[189,342,1945,1971]" box="[189,342,1945,1971]" pageId="64" pageNumber="65">
<heading id="CE0E701C301A5224EDAAF83E765A068C" bold="true" box="[189,342,1945,1971]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A78D1B62301A5224EDAAF83E765A068C" bold="true" box="[189,342,1945,1971]" pageId="64" pageNumber="65">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770301A5224EDAAF860746C06DE" blockId="64.[189,864,1991,2018]" box="[189,864,1991,2018]" pageId="64" pageNumber="65">
<collectingCountry id="EDEE87E0301A5224EDAAF860763806DE" box="[189,308,1991,2017]" name="Colombia" pageId="64" pageNumber="65">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E0301A5224EC28F86076B706DE" box="[319,443,1991,2017]" name="Venezuela" pageId="64" pageNumber="65">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0301A5224ECD1F860751D06DE" box="[454,529,1991,2017]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0301A5224EF0BF860758C06DE" box="[540,640,1991,2017]" name="Ecuador" pageId="64" pageNumber="65">Ecuador</collectingCountry>
,
<collectingCountry id="EDEE87E0301A5224EF9BF86F75C906DD" box="[652,709,1992,2018]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
, and
<collectingCountry id="EDEE87E0301A5224EE15F860745006DE" box="[770,860,1991,2017]" name="Bolivia" pageId="64" pageNumber="65">Bolivia</collectingCountry>
.
</paragraph>
<caption id="C18697F8301B5225EDAAF80C723C06D7" ID-DOI="http://doi.org/10.5281/zenodo.13945529" ID-Zenodo-Dep="13945529" httpUri="https://zenodo.org/record/13945529/files/figure.png" pageId="65" pageNumber="66" startId="65.[189,232,1963,1989]" targetBox="[279,1304,277,1880]" targetPageId="65">
<paragraph id="9546C770301B5225EDAAF80C723C06D7" blockId="65.[189,1398,1963,2025]" pageId="65" pageNumber="66">
<emphasis id="A78D1B62301B5225EDAAF80C762C06FA" bold="true" box="[189,288,1963,1989]" pageId="65" pageNumber="66">Fig. 25.</emphasis>
Species distribution.
<emphasis id="A78D1B62301B5225EF3AF80C754E06FA" bold="true" box="[557,578,1963,1989]" pageId="65" pageNumber="66">A</emphasis>
.
<taxonomicName id="52F9BCF3301B5225EF41F80C738906FA" authority="(d'Olsoufieff, 1924)" baseAuthorityName="d'Olsoufieff" baseAuthorityYear="1924" box="[598,1157,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="A78D1B62301B5225EF41F80C748806FA" box="[598,900,1963,1989]" italics="true" pageId="65" pageNumber="66">Coprophanaeus parvulus</emphasis>
(
<bibRefCitation id="F168BA81301B5225EE8FF80C737706FA" author="d'Olsoufieff G." box="[920,1147,1963,1989]" pageId="65" pageNumber="66" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff, 1924</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="A78D1B62301B5225E982F80C73A506FA" bold="true" box="[1173,1193,1963,1989]" pageId="65" pageNumber="66">B</emphasis>
.
<taxonomicName id="52F9BCF3301B5225E9ABF80C76E006D7" authority="(Erichson, 1847)" baseAuthorityName="Erichson" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="telamon">
<emphasis id="A78D1B62301B5225E9ABF80C761006D7" italics="true" pageId="65" pageNumber="66">Coprophanaeus telamon</emphasis>
(Erichson, 1847)
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C77030185226EDAAFEAF76E1001C" blockId="66.[189,493,264,291]" box="[189,493,264,291]" pageId="66" pageNumber="67">
<heading id="CE0E701C30185226EDAAFEAF76E1001C" bold="true" box="[189,493,264,291]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="A78D1B6230185226EDAAFEAF76E1001C" bold="true" box="[189,493,264,291]" pageId="66" pageNumber="67">
Subregions of
<collectingCountry id="EDEE87E030185226EC78FEAE76E1001C" box="[367,493,265,291]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030185226EDAAFE9176FB00AB" blockId="66.[189,1400,310,405]" pageId="66" pageNumber="67">
Maracaibo Depression, Plains, System of hills and low sierras Lara-Falcón, System of low mountains and hills Imataca-Cuyuní of Northeast
<collectingCountry id="EDEE87E030185226EFB2FEFF740C004D" box="[677,768,344,370]" name="Guyana" pageId="66" pageNumber="67">Guyana</collectingCountry>
Shield, Andes mountains, Central Coast Mountain Range, and Guiana Shield.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30185226EDAAFE1D74260246" pageId="66" pageNumber="67" type="reference_group">
<paragraph id="9546C77030185226EDAAFE1D76A800EA" blockId="66.[189,420,442,469]" box="[189,420,442,469]" pageId="66" pageNumber="67">
<heading id="CE0E701C30185226EDAAFE1D76A800EA" bold="true" box="[189,420,442,469]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="A78D1B6230185226EDAAFE1D76A800EA" bold="true" box="[189,420,442,469]" pageId="66" pageNumber="67">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030185226EDAAFE4F74260246" blockId="66.[188,1399,488,890]" pageId="66" pageNumber="67">
<bibRefCitation id="F168BA8130185226EDAAFE4F7672033D" author="Blanco J." box="[189,382,488,514]" pageId="66" pageNumber="67" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 43</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EC99FE4F7501033D" box="[398,525,488,514]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226EF00FE4F7579033D" box="[535,629,488,514]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
: Carretera
<collectingCountry id="EDEE87E030185226EFE3FE4F749E033D" box="[756,914,488,514]" name="Saint Kitts and Nevis" pageId="66" pageNumber="67">San Cristóbal</collectingCountry>
Chorro del Indio, Río Negro, Paramillo, Palo Gordo and Morretón). —
<bibRefCitation id="F168BA8130185226EF5DFDAD7434031B" author="Havranek D." box="[586,824,522,548]" pageId="66" pageNumber="67" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EE58FDAD74C2031B" box="[847,974,522,548]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226EEC8FDAD7332031B" box="[991,1086,522,548]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA8130185226E994FDAD76150378" author="Martinez A. &amp; Martinez A." pageId="66" pageNumber="67" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo 1990: 7</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EC3BFD8B76A70379" box="[300,427,556,582]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226ECA0FD8A751E0378" box="[439,530,557,583]" country="Venezuela" name="Aragua" pageId="66" pageNumber="67">Aragua</collectingRegion>
: Parque Nacional Henri Pittier, Estación Biológica Rancho Grande). —
<bibRefCitation id="F168BA8130185226EDAAFDE876EA0356" author="Gamez J. &amp; Mora E." box="[189,486,591,617]" pageId="66" pageNumber="67" pagination="3 - 16" refId="ref120147" refString="Gamez J. &amp; Mora E. 2000. Aspectos autoecologicos y distribucion local de Phanaeus (Phanaeus) prasinus Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae) en Venezuela. Memorias de la Fundacion La Salle de Ciencias Naturales 60 (153): 3 - 16." type="journal article" year="2000">Gámez &amp; Mora 2000: 17</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226ECE0FDE9757B0357" box="[503,631,590,616]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130185226EFB9FDE974880356" author="Arnaud P." box="[686,900,590,617]" pageId="66" pageNumber="67" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud 2002b: 35</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EE83FDE9731F0357" box="[916,1043,590,616]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226E909FDE973710357" box="[1054,1149,590,616]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA8130185226E9A1FDE9727B0357" author="Barbero E." box="[1206,1399,590,617]" pageId="66" pageNumber="67" pagination="11 - 21" refId="ref110583" refString="Barbero E. 2001. Scarabaeidae (Coleoptera) copronecrofagos interesantes del Departamento de Rio San Juan, Nicaragua. Revista Nicaraguense de Entomologia 55 - 58: 11 - 21." type="book chapter" year="2001">Barbero 2001: 6</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EDD1FDD6764A03B4" box="[198,326,625,651]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130185226EC68FDD6754F03B4" author="Gamez J." box="[383,579,624,651]" pageId="66" pageNumber="67" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004: 59</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EF43FDD675DF03B4" box="[596,723,625,651]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226EFC8FDD6743403B4" box="[735,824,625,651]" country="Venezuela" name="Merida" pageId="66" pageNumber="67">Mérida</collectingRegion>
: Guaraque: Mesa de Quintero; and
<collectingRegion id="573D099230185226E9F7FDD6723203B4" box="[1248,1342,625,651]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
:
<collectingCountry id="EDEE87E030185226E85DFDD676220392" name="Saint Kitts and Nevis" pageId="66" pageNumber="67">San Cristóbal</collectingCountry>
: Hacienda Pánaga); 2010: 16, 19 (
<collectingCountry id="EDEE87E030185226EFD9FD3474410392" box="[718,845,659,685]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
: Andean plains flank). —
<bibRefCitation id="F168BA8130185226E99BFD34763803F0" author="Gamez J. &amp; Acconcia R." pageId="66" pageNumber="67" pagination="387 - 396" refId="ref119925" refString="Gamez J. &amp; Acconcia R. 2009. Informaciones ecologicas sobre Coprophanaeus (Coprophanaeus) gamezi Arnaud (Coleoptera: Scarabaeinae: Phanaeini) en un sistema agropastoril en la depresion de Maracaibo, estado Zulia, Venezuela. Acta Zoologica Mexicana (n. s.) 25 (2): 387 - 396. https: // doi. org / 10.21829 / azm. 2009.252645" type="journal article" year="2009">Gámez &amp; Acconcia 2009: 394</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EC52FD1276C803F0" box="[325,452,693,719]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226ECC7FD12751E03F0" box="[464,530,693,719]" country="Venezuela" name="Zulia" pageId="66" pageNumber="67">Zulia</collectingRegion>
: Colón: Depresión de Maracaibo); 2018: 65, 69, 73 (
<collectingCountry id="EDEE87E030185226E99EFD12720403F0" box="[1161,1288,693,719]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226E802FD12727F03F0" box="[1301,1395,693,719]" country="Venezuela" name="Barinas" pageId="66" pageNumber="67">Barinas</collectingRegion>
:
<collectingRegion id="573D099230185226EDA9FD70761A03CE" box="[190,278,727,753]" country="Venezuela" name="Bolivar" pageId="66" pageNumber="67">Bolívar</collectingRegion>
and Andrés Eloy Blanco). —
<bibRefCitation id="F168BA8130185226EF6CFD7074D903CD" author="Hamel-Leigue A. &amp; Herzog S. K. &amp; Mann D. J. &amp; Larsen T. H. &amp; Gill B. D. &amp; Edmonds W. D. &amp; Spector S." box="[635,981,727,754]" pageId="66" pageNumber="67" pagination="43 - 95" refId="ref122950" refString="Hamel-Leigue A., Herzog S. K., Mann D. J., Larsen T. H., Gill B. D., Edmonds W. D., Spector S. 2009. Distribucion e historia natural de escarabajos coprofagos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
<emphasis id="A78D1B6230185226EE3CFD7F745203CE" box="[811,862,727,753]" italics="true" pageId="66" pageNumber="67">et al</emphasis>
. 2009: 57
</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EEF2FD70736903CE" box="[997,1125,727,753]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130185226E98AFD70762B022B" author="Edmonds W. D. &amp; Zidek J." pageId="66" pageNumber="67" pagination="1 - 111" refId="ref117660" refString="Edmonds W. D. &amp; Zidek J. 2010. A taxonomic review of the Neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae). Insecta Mundi 129: 1 - 111." type="journal article" year="2010">Edmonds &amp; Zídek 2010: 77</bibRefCitation>
, 91 (
<collectingCountry id="EDEE87E030185226EC49FD5E76D1022C" box="[350,477,761,787]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226ECF1FD5E756E022C" box="[486,610,761,787]" country="Venezuela" name="Amazonas" pageId="66" pageNumber="67">Amazonas</collectingRegion>
[
<collectingRegion id="573D099230185226EF67FD5E75D9022C" box="[624,725,761,787]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
?]: La Tuquerena Rubio;
<collectingRegion id="573D099230185226EEF8FD5E7345022C" box="[1007,1097,761,787]" country="Venezuela" name="Aragua" pageId="66" pageNumber="67">Aragua</collectingRegion>
;
<collectingRegion id="573D099230185226E943FD5E73BC022C" box="[1108,1200,761,787]" country="Venezuela" name="Bolivar" pageId="66" pageNumber="67">Bolívar</collectingRegion>
;
<collectingRegion id="573D099230185226E9ACFD5E7203022C" box="[1211,1295,761,787]" country="Venezuela" name="Falcon" pageId="66" pageNumber="67">Falcón</collectingRegion>
;
<collectingRegion id="573D099230185226E80DFD5E727F022C" box="[1306,1395,761,787]" country="Venezuela" name="Merida" pageId="66" pageNumber="67">Mérida</collectingRegion>
; and
<collectingRegion id="573D099230185226EDE5FCBC765D020A" box="[242,337,795,821]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA8130185226EC84FCBC745F020A" author="Cupello M. &amp; Vaz-de-Mello F. Z." box="[403,851,795,821]" pageId="66" pageNumber="67" pagination="359 - 368" refId="ref115382" refString="Cupello M. &amp; Vaz-de-Mello F. Z. 2013 a. New evidence for the validity of Coprophanaeus (C.) terrali Arnaud, 2002 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini), a dung beetle from Brazil. Zootaxa, 3717 (3): 359 - 368. https: // doi. org / 10.11646 / zootaxa. 3717.3.5" type="journal article" year="2013">Cupello &amp; Vaz-de-Mello 2013a: 367</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EE71FCBC74E9020A" box="[870,997,795,821]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226EEE4FCBC7341020A" box="[1011,1101,795,821]" country="Venezuela" name="Aragua" pageId="66" pageNumber="67">Aragua</collectingRegion>
;
<collectingRegion id="573D099230185226E94AFCBC73B7020A" box="[1117,1211,795,821]" country="Venezuela" name="Tachira" pageId="66" pageNumber="67">Táchira</collectingRegion>
:
<collectingCountry id="EDEE87E030185226E9DCFCBC727E020A" box="[1227,1394,795,821]" name="Saint Kitts and Nevis" pageId="66" pageNumber="67">San Cristóbal</collectingCountry>
; La Guaira; and Yaracuy: Bolívar). —
<bibRefCitation id="F168BA8130185226EF83FC9A74FE0267" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[660,1010,829,856]" pageId="66" pageNumber="67" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230185226EE3BFC9974680268" box="[812,868,829,855]" italics="true" pageId="66" pageNumber="67">et al</emphasis>
. 2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226E911FC9A73890268" box="[1030,1157,829,855]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230185226E984FC9973F40267" box="[1171,1272,830,856]" country="Venezuela" name="Yaracuy" pageId="66" pageNumber="67">Yaracuy</collectingRegion>
: Hacienda Guáquira). —
<bibRefCitation id="F168BA8130185226EC70FCF875830245" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[359,655,863,890]" pageId="66" pageNumber="67" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230185226ECF2FCC7752C0246" box="[485,544,863,889]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
2019: 77
</bibRefCitation>
(
<collectingCountry id="EDEE87E030185226EF89FCF874120246" box="[670,798,863,889]" name="Venezuela" pageId="66" pageNumber="67">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,339 @@
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Genus
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<emphasis id="A78D1B62305C5262EF8FF899742F0667" bold="true" box="[664,803,1854,1880]" italics="true" pageId="6" pageNumber="7">Anomiopus</emphasis>
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(original description).
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species:
<taxonomicName id="52F9BCF3305C5262EEE2F82776240682" ID-CoL="676N2" authority="Westwood, 1842" authorityName="Westwood" authorityYear="1842" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="A78D1B62305C5262EEE2F82773E106A5" box="[1013,1261,1920,1946]" italics="true" pageId="6" pageNumber="7">Anomiopus virescens</emphasis>
<bibRefCitation id="F168BA81305C5262E9E3F82776240682" author="Westwood J. O." pageId="6" pageNumber="7" pagination="58 - 59" refId="ref141897" refString="Westwood J. O. 1842. Description of new exotic genera belonging to the family of the sacred beetles. Proceedings of the Entomological Society of London 1842: 58 - 59." type="book chapter" year="1842">Westwood, 1842</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation id="F168BA81305C5262EF86F804745F0682" author="Canhedo V. L." box="[657,851,1955,1981]" pageId="6" pageNumber="7" pagination="349 - 502" refId="ref113649" refString="Canhedo V. L. 2006. Revisao taxonomica do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Arquivos de Zoologia 37 (4): 349 - 502. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 37 i 4 p 349 - 502" type="journal article" year="2006">Canhedo (2006)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<caption id="C18697F8305D5263EDAAF80C734C06D7" ID-DOI="http://doi.org/10.5281/zenodo.13945472" ID-Zenodo-Dep="13945472" httpUri="https://zenodo.org/record/13945472/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1963,1989]" targetBox="[279,1301,286,1895]" targetPageId="7">
<paragraph id="9546C770305D5263EDAAF80C734C06D7" blockId="7.[189,1398,1963,2025]" pageId="7" pageNumber="8">
<emphasis id="A78D1B62305D5263EDAAF80C760206FA" bold="true" box="[189,270,1963,1989]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
Species distribution.
<emphasis id="A78D1B62305D5263EF01F80C752706FA" bold="true" box="[534,555,1963,1989]" pageId="7" pageNumber="8">A</emphasis>
.
<taxonomicName id="52F9BCF3305D5263EF2AF80C74B206FA" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[573,958,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Agamopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lampros">
<emphasis id="A78D1B62305D5263EF2AF80C742E06FA" box="[573,802,1963,1989]" italics="true" pageId="7" pageNumber="8">Agamopus lampros</emphasis>
<bibRefCitation id="F168BA81305D5263EE3AF80B74B206FA" author="Bates H. W." box="[813,958,1963,1990]" pageId="7" pageNumber="8" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1887">Bates, 1887</bibRefCitation>
</taxonomicName>
.
<emphasis id="A78D1B62305D5263EEDBF80C74EC06FA" bold="true" box="[972,992,1963,1989]" pageId="7" pageNumber="8">B</emphasis>
.
<taxonomicName id="52F9BCF3305D5263EEE5F80C77F706D7" authority="Canhedo, 2006" authorityName="Canhedo" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alexandrei">
<emphasis id="A78D1B62305D5263EEE5F80C73F006FA" box="[1010,1276,1963,1989]" italics="true" pageId="7" pageNumber="8">Anomiopus alexandrei</emphasis>
<bibRefCitation id="F168BA81305D5263E810F80C77F706D7" author="Canhedo V. L." pageId="7" pageNumber="8" pagination="349 - 502" refId="ref113649" refString="Canhedo V. L. 2006. Revisao taxonomica do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Arquivos de Zoologia 37 (4): 349 - 502. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 37 i 4 p 349 - 502" type="journal article" year="2006">Canhedo, 2006</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C7703052526CEDAAFEAE75BA00B2" blockId="8.[189,1400,265,397]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="B5E9E05E3052526CEDAAFEAE74FA0078" pageId="8" pageNumber="9">
<treatmentCitation id="1458E1613052526CEDAAFEAE7565001C" author="Erichson W. F." box="[189,617,265,291]" page="189" pageId="8" pageNumber="9" year="1847">
<taxonomicName id="52F9BCF33052526CEDAAFEAE7565001C" ID-CoL="68DM" authority="Erichson, 1847 b: 189" authorityName="Erichson" authorityPageNumber="189" authorityYear="1847" box="[189,617,265,291]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEDAAFEAE765C001C" box="[189,336,265,291]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
<bibRefCitation id="F168BA813052526CEC49FEAE7565001C" author="Erichson W. F." box="[350,617,265,291]" pageId="8" pageNumber="9" pagination="1 - 968" refId="ref117862" refString="Erichson W. F. 1847 b. Naturgeschichte des Insecten Deutschland. I. Coleoptera, Scaphidilia, Scarabaeides. Erste Antheilung, Dritter Band, Erste Lieferung. Nicolaischen Buchhandlung, Berlin. 1 - 968. https: // doi. org / 10.5962 / bhl. title. 50882" type="book chapter" year="1847">Erichson, 1847b: 189</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D23052526CEE99FEAD74C4001B" box="[910,968,266,292]" pageId="8" pageNumber="9">Type</typeStatus>
species:
<taxonomicName id="52F9BCF33052526CE955FEAE727B001C" authority="Westwood, 1842" box="[1090,1399,265,291]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="smaragdinus">
<emphasis id="A78D1B623052526CE955FEAE727B001C" box="[1090,1399,265,291]" italics="true" pageId="8" pageNumber="9">Scatonomus smaragdinus</emphasis>
</taxonomicName>
<bibRefCitation id="F168BA813052526CEDFDFE8B76BD0079" author="Westwood J. O." box="[234,433,300,326]" pageId="8" pageNumber="9" pagination="58 - 59" refId="ref141897" refString="Westwood J. O. 1842. Description of new exotic genera belonging to the family of the sacred beetles. Proceedings of the Entomological Society of London 1842: 58 - 59." type="book chapter" year="1842">Westwood, 1842</bibRefCitation>
, by subsequent designation of
<bibRefCitation id="F168BA813052526CEE0DFE8B74FE0078" author="Lacordaire J. T." box="[794,1010,300,327]" pageId="8" pageNumber="9" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire (1855)</bibRefCitation>
.
</treatmentCitationGroup>
<treatmentCitationGroup id="B5E9E05E3052526CEDAAFEE875BA00B2" pageId="8" pageNumber="9">
<treatmentCitation id="1458E1613052526CEDAAFEE875730055" author="Balthasar V." box="[189,639,335,362]" page="214" pageId="8" pageNumber="9" year="1938">
<taxonomicName id="52F9BCF33052526CEDAAFEE875730055" ID-CoL="52MH" authority="Balthasar, 1938: 214" authorityName="Balthasar" authorityPageNumber="214" authorityYear="1938" box="[189,639,335,362]" class="Insecta" family="Scarabaeidae" genus="Hypocanthidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEDAAFEE8768D0056" box="[189,385,335,361]" italics="true" pageId="8" pageNumber="9">Hypocanthidium</emphasis>
<bibRefCitation id="F168BA813052526CEC9EFEE875730055" author="Balthasar V." box="[393,639,335,362]" pageId="8" pageNumber="9" pagination="210 - 223" refId="ref110121" refString="Balthasar V. 1938. Neue Gattungen und Arten der sudamerikanischen Coprophagen. (4. Beitrag zur Kenntnis der Scarabaeiden der neotropischen Region). Entomologische Blatter 34 (4): 210 - 223." type="journal article" year="1938">Balthasar, 1938: 214</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D23052526CEE85FEF774C00055" box="[914,972,336,362]" pageId="8" pageNumber="9">Type</typeStatus>
species:
<taxonomicName id="52F9BCF33052526CE92AFEE872740056" ID-CoL="3NTH9" authorityName="Balthasar" authorityYear="1938" box="[1085,1400,335,361]" class="Insecta" family="Scarabaeidae" genus="Hypocanthidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="globulum">
<emphasis id="A78D1B623052526CE92AFEE872740056" box="[1085,1400,335,361]" italics="true" pageId="8" pageNumber="9">Hypocanthidium globulum</emphasis>
</taxonomicName>
<bibRefCitation id="F168BA813052526CEDFBFED576A900B2" author="Balthasar V." box="[236,421,370,397]" pageId="8" pageNumber="9" pagination="210 - 223" refId="ref110121" refString="Balthasar V. 1938. Neue Gattungen und Arten der sudamerikanischen Coprophagen. (4. Beitrag zur Kenntnis der Scarabaeiden der neotropischen Region). Entomologische Blatter 34 (4): 210 - 223." type="journal article" year="1938">Balthasar, 1938</bibRefCitation>
, by original monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703052526CEDAAFE1374160211" blockId="8.[189,1399,436,920]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="B5E9E05E3052526CEDAAFE1374160211" pageId="8" pageNumber="9">
<emphasis id="A78D1B623052526CEDAAFE13765500F1" box="[189,345,436,462]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="52F9BCF33052526CEDAAFE13764D00F1" ID-CoL="62BHN" authorityName="Westwood" authorityYear="1842" box="[189,321,436,462]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Anomiopus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E1613052526CEC76FE13754500F1" author="Erichson W. F." box="[353,585,436,462]" page="189" pageId="8" pageNumber="9" year="1843">
<bibRefCitation id="F168BA813052526CEC76FE13754500F1" author="Erichson W. F." box="[353,585,436,462]" pageId="8" pageNumber="9" pagination="149 - 288" refId="ref117750" refString="Erichson W. F. 1843. Bericht uber die wissenschaftlichen Leistung in der Naturgeschichte der Insecten, Arachniden, Crustaceen u. Entomostraceen wahrend des Jahres 1842. Archiv fur Naturgeschichte 9 (2): 149 - 288. Available from https: // www. biodiversitylibrary. org / page / 13703576 [accessed 14 Feb. 2024]." type="journal article" year="1843">Erichson 1843: 189</bibRefCitation>
</treatmentCitation>
(list); 1847b: 761 (cited as synonym of
<taxonomicName id="52F9BCF33052526CE93CFE1373B000F1" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[1067,1212,436,462]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CE93CFE1373B000F1" box="[1067,1212,436,462]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613052526CE9ECFE13765800CE" author="Westwood J. O." page="62" pageId="8" pageNumber="9" year="1843">
<bibRefCitation id="F168BA813052526CE9ECFE13765800CE" author="Westwood J. O." pageId="8" pageNumber="9" pagination="61 - 62" refId="ref141931" refString="Westwood J. O. 1843. Description of some new exotic genera belonging to the family of the sacred beetles. The Annals and Magazine of Natural History, Zoology, Botany, and Geology 11: 61 - 62. Available from https: // www. biodiversitylibrary. org / page / 2326216 [accessed 14 Feb. 2024]." type="journal article" year="1843">Westwood 1843: 62</bibRefCitation>
</treatmentCitation>
(redescription); 1847: 231 (redescription).
<emphasis id="A78D1B623052526CEE4AFE7F747B00CE" box="[861,887,472,497]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEE68FE70737D00CE" author="Lacordaire J. T." box="[895,1137,471,498]" page="94" pageId="8" pageNumber="9" year="1855">
<bibRefCitation id="F168BA813052526CEE68FE70737D00CE" author="Lacordaire J. T." box="[895,1137,471,498]" pageId="8" pageNumber="9" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 94</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33052526CEDFAFE5D7671032B" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[237,381,506,532]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEDFAFE5D7671032B" box="[237,381,506,532]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
, see footnote 2).
<emphasis id="A78D1B623052526CEF4CFE5C7579032B" box="[603,629,507,532]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEF96FE5D747B032A" author="Harold E." box="[641,887,506,533]" page="1002" pageId="8" pageNumber="9" year="1869">
<bibRefCitation id="F168BA813052526CEF96FE5D747B032A" author="Harold E." box="[641,887,506,533]" pageId="8" pageNumber="9" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1002</bibRefCitation>
</treatmentCitation>
(catalogue, synonym of
<taxonomicName id="52F9BCF33052526CE9BAFE5D724C032B" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[1197,1344,506,532]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CE9BAFE5D724C032B" box="[1197,1344,506,532]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623052526CE84BFE5C727A032B" box="[1372,1398,507,532]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEDFDFDB976A40307" author="Gillet J. J. E." box="[234,424,541,568]" page="50" pageId="8" pageNumber="9" year="1911">
<bibRefCitation id="F168BA813052526CEDFDFDB976A40307" author="Gillet J. J. E." box="[234,424,541,568]" pageId="8" pageNumber="9" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 50</bibRefCitation>
</treatmentCitation>
(catalogue, synonym of
<taxonomicName id="52F9BCF33052526CEFDCFDB974520307" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[715,862,542,568]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEFDCFDB974520307" box="[715,862,542,568]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623052526CEE61FDB9749C0308" box="[886,912,542,567]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEE80FDB973570307" author="Lucas R." box="[919,1115,542,568]" page="100" pageId="8" pageNumber="9" year="1920">
<bibRefCitation id="F168BA813052526CEE80FDB973570307" author="Lucas R." box="[919,1115,542,568]" pageId="8" pageNumber="9" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 100</bibRefCitation>
</treatmentCitation>
(catalogue, synonym of
<taxonomicName id="52F9BCF33052526CEDFDFDE676710364" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[234,381,577,603]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEDFDFDE676710364" box="[234,381,577,603]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623052526CEC81FDE576BC0364" box="[406,432,578,603]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CECA0FDE675C60364" author="Blackwelder R. E." box="[439,714,577,603]" page="204" pageId="8" pageNumber="9" year="1944">
<bibRefCitation id="F168BA813052526CECA0FDE675C60364" author="Blackwelder R. E." box="[439,714,577,603]" pageId="8" pageNumber="9" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 204</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33052526CEEC7FDE6736E0364" ID-CoL="68DM" authorityName="Erichson" authorityYear="1847" box="[976,1122,577,603]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEEC7FDE6736E0364" box="[976,1122,577,603]" italics="true" pageId="8" pageNumber="9">Onthocharis</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623052526CE96CFDE573990364" box="[1147,1173,578,603]" italics="true" pageId="8" pageNumber="9"></emphasis>
Medina &amp; Lopera, 2000: 306 (key).
<emphasis id="A78D1B623052526CECD6FDC276D70341" box="[449,475,613,638]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CECF1FDC374010341" author="Vaz-de-Mello F. Z." box="[486,781,612,638]" page="190" pageId="8" pageNumber="9" year="2000">
<bibRefCitation id="F168BA813052526CECF1FDC374010341" author="Vaz-de-Mello F. Z." box="[486,781,612,638]" pageId="8" pageNumber="9" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 190</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03052526CEED4FDC3731C0341" box="[963,1040,612,638]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
).
<emphasis id="A78D1B623052526CE93FFDC2734E0341" box="[1064,1090,613,638]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CE95AFDC3727A0341" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[1101,1398,612,638]" page="137" pageId="8" pageNumber="9" year="2001">
<bibRefCitation id="F168BA813052526CE95AFDC3727A0341" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[1101,1398,612,638]" pageId="8" pageNumber="9" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B623052526CE9A5FDC273FD0341" box="[1202,1265,612,638]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2001: 137
</bibRefCitation>
</treatmentCitation>
(list).
<emphasis id="A78D1B623052526CEC24FD2F7641039E" box="[307,333,648,673]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEC4FFD207548039D" author="Canhedo V. L." box="[344,580,647,674]" page="354" pageId="8" pageNumber="9" year="2006">
<bibRefCitation id="F168BA813052526CEC4FFD207548039D" author="Canhedo V. L." box="[344,580,647,674]" pageId="8" pageNumber="9" pagination="349 - 502" refId="ref113649" refString="Canhedo V. L. 2006. Revisao taxonomica do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Arquivos de Zoologia 37 (4): 349 - 502. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 37 i 4 p 349 - 502" type="journal article" year="2006">Canhedo 2006: 354</bibRefCitation>
</treatmentCitation>
(revision).
<emphasis id="A78D1B623052526CEFC5FD2F75E0039E" box="[722,748,648,673]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEFE0FD207351039E" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[759,1117,647,673]" page="12" pageId="8" pageNumber="9" year="2006">
<bibRefCitation id="F168BA813052526CEFE0FD207351039E" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[759,1117,647,673]" pageId="8" pageNumber="9" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B623052526CEEBDFD2F74E4039E" box="[938,1000,647,673]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2006: 12
</bibRefCitation>
</treatmentCitation>
(list).
<emphasis id="A78D1B623052526CE9A6FD2F73C7039E" box="[1201,1227,648,673]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CE9C2FD2076A603FA" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." page="25" pageId="8" pageNumber="9" year="2011">
<bibRefCitation id="F168BA813052526CE9C2FD2076A603FA" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." pageId="8" pageNumber="9" pagination="1 - 73" refId="ref140802" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C., Schoolmeesters P. 2011 a. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B623052526CEDFDFD0C762503FB" box="[234,297,682,708]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2011a: 25
</bibRefCitation>
</treatmentCitation>
(key).
<emphasis id="A78D1B623052526CEF11FD0C752C03FB" box="[518,544,683,708]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEF3CFD0D745203FA" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[555,862,682,709]" page="124" pageId="8" pageNumber="9" year="2011">
<bibRefCitation id="F168BA813052526CEF3CFD0D745203FA" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[555,862,682,709]" pageId="8" pageNumber="9" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B623052526CEF8EFD0C75D403FB" box="[665,728,682,708]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2011: 124
</bibRefCitation>
</treatmentCitation>
, 318 (description, list).
<emphasis id="A78D1B623052526CE994FD0C739103FB" box="[1155,1181,683,708]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CE9BFFD0D727A03FA" author="Krajcik M." box="[1192,1398,682,709]" page="28" pageId="8" pageNumber="9" year="2012">
<bibRefCitation id="F168BA813052526CE9BFFD0D727A03FA" author="Krajcik M." box="[1192,1398,682,709]" pageId="8" pageNumber="9" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 28</bibRefCitation>
</treatmentCitation>
(list).
<emphasis id="A78D1B623052526CEC27FD69764603D8" box="[304,330,718,743]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEC46FD69758703D7" author="Solis A. &amp; Kohlmann B." box="[337,651,718,744]" page="2" pageId="8" pageNumber="9" year="2012">
<bibRefCitation id="F168BA813052526CEC46FD69758703D7" author="Solis A. &amp; Kohlmann B." box="[337,651,718,744]" pageId="8" pageNumber="9" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 2</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03052526CEE24FD6974B403D7" box="[819,952,718,744]" name="Costa Rica" pageId="8" pageNumber="9">Costa Rica</collectingCountry>
).
<emphasis id="A78D1B623052526CEEDDFD6974E803D8" box="[970,996,718,743]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEEFBFD69727A03D7" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[1004,1398,718,744]" page="107" pageId="8" pageNumber="9" year="2013">
<bibRefCitation id="F168BA813052526CEEFBFD69727A03D7" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[1004,1398,718,744]" pageId="8" pageNumber="9" pagination="103 - 112" refId="ref111991" refString="Boilly O. &amp; Vaz-de-Mello F. Z. 2013. Les Scarabaeinae de Guyane: cle illustree des genres. ACOREP- France 7: 103 - 112." type="journal article" year="2013">Boilly &amp; Vaz-de-Mello 2013: 107</bibRefCitation>
</treatmentCitation>
(key).
<emphasis id="A78D1B623052526CEC20FD55765D0234" box="[311,337,754,779]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEC4FFD56758C0233" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[344,640,753,780]" page="75" pageId="8" pageNumber="9" year="2018">
<bibRefCitation id="F168BA813052526CEC4FFD56758C0233" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[344,640,753,780]" pageId="8" pageNumber="9" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B623052526CECC0FD55751E0234" box="[471,530,753,779]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2018: 75
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03052526CEE3CFD56749E0234" box="[811,914,753,779]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
); 2019: 910 (catalogue).
<emphasis id="A78D1B623052526CE9D2FD5573D30234" box="[1221,1247,754,779]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CE9F1FD5676C70210" ID-DOI="http://doi.org/10.5281/zenodo.3684665" ID-Zenodo-Dep="3684665" author="Hielkema A. J. &amp; Hielkema M. A." httpUri="http://treatment.plazi.org/id/03D7FF1CFFF0FFD6FF48FF97308BFAAE" page="58" pageId="8" pageNumber="9" year="2019">
<bibRefCitation id="F168BA813052526CE9F1FD5676C70210" author="Hielkema A. J. &amp; Hielkema M. A." pageId="8" pageNumber="9" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 58</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703052526CEDAAFC90751002A7" blockId="8.[189,1399,436,920]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="B5E9E05E3052526CEDAAFC90751002A7" pageId="8" pageNumber="9">
<emphasis id="A78D1B623052526CEDAAFC907699026E" box="[189,405,823,849]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="52F9BCF33052526CEDAAFC90768D026E" ID-CoL="52MH" authorityName="Balthasar" authorityYear="1938" box="[189,385,823,849]" class="Insecta" family="Scarabaeidae" genus="Hypocanthidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Hypocanthidium</taxonomicName>
</emphasis>
<treatmentCitation id="1458E1613052526CEC8EFC90756F026D" author="Pereira F. S." box="[409,611,823,850]" page="56" pageId="8" pageNumber="9" year="1954">
<bibRefCitation id="F168BA813052526CEC8EFC90756F026D" author="Pereira F. S." box="[409,611,823,850]" pageId="8" pageNumber="9" pagination="53 - 78" refId="ref135304" refString="Pereira F. S. 1954 a. O genero Scatonomus Er. (Coleoptera, Scarabaeidae). Revista Brasileira de Entomologia 1: 53 - 78." type="journal article" year="1954">Pereira 1954a: 56</bibRefCitation>
</treatmentCitation>
(key).
<emphasis id="A78D1B623052526CEFA7FC9F75C6026E" box="[688,714,824,849]" italics="true" pageId="8" pageNumber="9"></emphasis>
Halffter &amp; Mathews 1966: 256 (catalogue, distribution).
<emphasis id="A78D1B623052526CE84DFC9F7278026E" box="[1370,1396,824,849]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CEDFDFCFD755D024A" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[234,593,858,885]" page="3" pageId="8" pageNumber="9" year="2011">
<bibRefCitation id="F168BA813052526CEDFDFCFD755D024A" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[234,593,858,885]" pageId="8" pageNumber="9" pagination="1 - 73" refId="ref140802" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C., Schoolmeesters P. 2011 a. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B623052526CEC8EFCFC76D5024B" box="[409,473,858,884]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2011a: 3
</bibRefCitation>
</treatmentCitation>
(junior synonym of
<taxonomicName id="52F9BCF33052526CEE4EFCFC74D1024A" ID-CoL="62BHN" authorityName="Westwood" authorityYear="1842" box="[857,989,859,885]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEE4EFCFC74D1024A" box="[857,989,859,885]" italics="true" pageId="8" pageNumber="9">Anomiopus</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623052526CEEEBFCFC731A024B" box="[1020,1046,859,884]" italics="true" pageId="8" pageNumber="9"></emphasis>
<treatmentCitation id="1458E1613052526CE934FCFD727B024A" author="Solis A. &amp; Kohlmann B." box="[1059,1399,858,885]" page="2" pageId="8" pageNumber="9" year="2012">
<bibRefCitation id="F168BA813052526CE934FCFD727B024A" author="Solis A. &amp; Kohlmann B." box="[1059,1399,858,885]" pageId="8" pageNumber="9" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 2</bibRefCitation>
</treatmentCitation>
(synonym of
<taxonomicName id="52F9BCF33052526CEC90FCD9750702A7" ID-CoL="62BHN" authorityName="Westwood" authorityYear="1842" box="[391,523,894,920]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623052526CEC90FCD9750702A7" box="[391,523,894,920]" italics="true" pageId="8" pageNumber="9">Anomiopus</emphasis>
</taxonomicName>
).
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View file

@ -0,0 +1,377 @@
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<heading id="CE0E701C30765248EF4EFDE974D70357" box="[601,987,590,617]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="45" reason="2">
<taxonomicName id="52F9BCF330765248EF4EFDE974D70357" ID-CoL="QP8G" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" box="[601,987,590,617]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="A78D1B6230765248EF4EFDE974350356" bold="true" box="[601,825,590,617]" italics="true" pageId="44" pageNumber="45">Canthon steinheili</emphasis>
<bibRefCitation id="F168BA8130765248EE57FDE974D70357" author="Harold E." box="[832,987,590,616]" pageId="44" pageNumber="45" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30765248EFF1FDD6744103B4" box="[742,845,625,651]" pageId="44" pageNumber="45" type="description">
<paragraph id="9546C77030765248EFF1FDD6744103B4" blockId="44.[601,987,590,651]" box="[742,845,625,651]" pageId="44" pageNumber="45">
<figureCitation id="0DC2DBF530765248EFF1FDD6744103B4" box="[742,845,625,651]" captionStart="Fig" captionStartId="45.[189,232,1963,1989]" captionTargetBox="[279,1311,278,1899]" captionTargetId="figure-11@45.[267,1321,265,1919]" captionTargetPageId="45" captionText="Fig. 18. Species distribution. A. Canthon simulans (Martínez, 1950). B. Canthon steinheili Harold, 1880. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945506" httpUri="https://zenodo.org/record/13945506/files/figure.png" pageId="44" pageNumber="45">Fig. 18B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30765248EDAAFD1574AD02C0" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="9546C77030765248EDAAFD15736603D0" blockId="44.[189,1399,690,751]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="B5E9E05E30765248EDAAFD15736603D0" pageId="44" pageNumber="45">
<treatmentCitation id="1458E16130765248EDAAFD15755203F3" author="Harold E." box="[189,606,690,716]" page="16" pageId="44" pageNumber="45" year="1880">
<taxonomicName id="52F9BCF330765248EDAAFD15755203F3" ID-CoL="QP8G" authority="Harold, 1880: 16" authorityName="Harold" authorityPageNumber="16" authorityYear="1880" box="[189,606,690,716]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="A78D1B6230765248EDAAFD15769803F3" box="[189,404,690,716]" italics="true" pageId="44" pageNumber="45">Canthon steinheili</emphasis>
<bibRefCitation id="F168BA8130765248EC8EFD15755203F3" author="Harold E." box="[409,606,690,716]" pageId="44" pageNumber="45" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880: 16</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030765248E919FD15738A03F3" box="[1038,1158,690,716]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
:
<collectingRegion id="573D099230765248E998FD1573EA03F3" box="[1167,1254,690,716]" country="Colombia" name="Tolima" pageId="44" pageNumber="45">Tolima</collectingRegion>
:Ambalema. Name-bearing type: a single known
<typeStatus id="4A4279D230765248EF83FD7275E303D0" box="[660,751,725,751]" pageId="44" pageNumber="45" type="syntype">syntype</typeStatus>
(MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030765248EDAAFCB2724702E2" blockId="44.[189,1399,789,1024]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="B5E9E05E30765248EDAAFCB2724702E2" pageId="44" pageNumber="45">
<emphasis id="A78D1B6230765248EDAAFCB276A10210" box="[189,429,789,815]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="52F9BCF330765248EDAAFCB2769A0210" ID-CoL="QP8G" authorityName="Harold" authorityYear="1880" box="[189,406,789,815]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="steinheili">Canthon steinheili</taxonomicName>
</emphasis>
<treatmentCitation id="1458E16130765248ECA1FCB2757A020F" author="Gillet J. J. E." box="[438,630,789,816]" page="33" pageId="44" pageNumber="45" year="1911">
<bibRefCitation id="F168BA8130765248ECA1FCB2757A020F" author="Gillet J. J. E." box="[438,630,789,816]" pageId="44" pageNumber="45" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 33</bibRefCitation>
</treatmentCitation>
(catalogue, cited for
<collectingCountry id="EDEE87E030765248EE63FCB274E10210" box="[884,1005,789,815]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130765248E931FCB273E1020F" author="Bruch C." box="[1062,1261,789,816]" page="540" pageId="44" pageNumber="45" year="1915">
<bibRefCitation id="F168BA8130765248E931FCB273E1020F" author="Bruch C." box="[1062,1261,789,816]" pageId="44" pageNumber="45" pagination="538 - 573" refId="ref112550" refString="Bruch C. 1915. Suplemento al catalogo sistematico de los coleopteros de la Republica Argentina. I (addenda, corrigenda y resumen). Revista del Museo de La Plata 19 (2): 538 - 573. Available from http: // sedici. unlp. edu. ar / handle / 10915 / 155145 [accessed 14 Feb. 2024]." type="journal article" year="1915">Bruch 1915: 540</bibRefCitation>
</treatmentCitation>
(catalogue, cited for Misiones,
<collectingCountry id="EDEE87E030765248ECC8FC9F7555026D" box="[479,601,824,850]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130765248EF8AFC9F74AE026C" author="Balthasar V." box="[669,930,824,851]" page="225" pageId="44" pageNumber="45" year="1939">
<bibRefCitation id="F168BA8130765248EF8AFC9F74AE026C" author="Balthasar V." box="[669,930,824,851]" pageId="44" pageNumber="45" pagination="179 - 238" refId="ref110158" refString="Balthasar V. 1939 a. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939a: 225</bibRefCitation>
</treatmentCitation>
(monograph, cited for
<collectingCountry id="EDEE87E030765248E9DFFC9F724D026D" box="[1224,1345,824,850]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130765248EDFDFCFC76F5024A" author="Blackwelder R. E." box="[234,505,859,885]" page="202" pageId="44" pageNumber="45" year="1944">
<bibRefCitation id="F168BA8130765248EDFDFCFC76F5024A" author="Blackwelder R. E." box="[234,505,859,885]" pageId="44" pageNumber="45" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 202</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130765248EF71FCFC745D024A" author="Gacharna C." box="[614,849,859,886]" page="221" pageId="44" pageNumber="45" year="1951">
<bibRefCitation id="F168BA8130765248EF71FCFC745D024A" author="Gacharna C." box="[614,849,859,886]" pageId="44" pageNumber="45" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 221</bibRefCitation>
</treatmentCitation>
(catalogue, cited for
<collectingCountry id="EDEE87E030765248E953FCFC73B1024A" box="[1092,1213,859,885]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130765248E9E5FCFC760A02A7" author="Roze J. A." page="43" pageId="44" pageNumber="45" year="1955">
<bibRefCitation id="F168BA8130765248E9E5FCFC760A02A7" author="Roze J. A." pageId="44" pageNumber="45" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 43</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030765248ECB8FCDA752102A8" box="[431,557,893,919]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E16130765248EF72FCD9743102A7" author="Martinez A." box="[613,829,893,920]" page="43" pageId="44" pageNumber="45" year="1959">
<bibRefCitation id="F168BA8130765248EF72FCD9743102A7" author="Martinez A." box="[613,829,893,920]" pageId="44" pageNumber="45" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 43</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E030765248EEFAFCDA736B02A8" box="[1005,1127,893,919]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130765248E98AFCDA766C0285" author="Vulcano M. A. &amp; Pereira F. S." page="630" pageId="44" pageNumber="45" year="1964">
<bibRefCitation id="F168BA8130765248E98AFCDA766C0285" author="Vulcano M. A. &amp; Pereira F. S." pageId="44" pageNumber="45" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 630</bibRefCitation>
</treatmentCitation>
(catalogue, distribution). —
<treatmentCitation id="1458E16130765248EFA5FC0773060285" author="Halffter G. &amp; Martinez A." box="[690,1034,928,954]" page="89" pageId="44" pageNumber="45" year="1977">
<bibRefCitation id="F168BA8130765248EFA5FC0773060285" author="Halffter G. &amp; Martinez A." box="[690,1034,928,954]" pageId="44" pageNumber="45" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 89</bibRefCitation>
</treatmentCitation>
(list of species group). —
<treatmentCitation id="1458E16130765248E856FC0676DF02E2" author="Vaz-de-Mello F. Z." page="191" pageId="44" pageNumber="45" year="2000">
<bibRefCitation id="F168BA8130765248E856FC0676DF02E2" author="Vaz-de-Mello F. Z." pageId="44" pageNumber="45" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 191</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030765248EF69FC6475C702E2" box="[638,715,963,989]" name="Brazil" pageId="44" pageNumber="45">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E16130765248EE15FC64731702E2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[770,1051,963,989]" page="139" pageId="44" pageNumber="45" year="2001">
<bibRefCitation id="F168BA8130765248EE15FC64731702E2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[770,1051,963,989]" pageId="44" pageNumber="45" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230765248EE74FC63749202E2" box="[867,926,963,989]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2001: 139
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030765248E9D1FC64723302E2" box="[1222,1343,963,989]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030765248EDAAFC4274AD02C0" blockId="44.[189,1399,789,1024]" box="[189,929,997,1024]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="B5E9E05E30765248EDAAFC4274AD02C0" box="[189,929,997,1024]" pageId="44" pageNumber="45">
<emphasis id="A78D1B6230765248EDAAFC42752C02C0" box="[189,544,997,1023]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="52F9BCF330765248EDAAFC42750602C0" ID-CoL="QP8G" box="[189,522,997,1023]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="mutabilis" subSpecies="steinheili">Canthon mutabilis steinheili</taxonomicName>
</emphasis>
<treatmentCitation id="1458E16130765248EF30FC4275FB053F" author="Schmidt A." box="[551,759,997,1024]" page="77" pageId="44" pageNumber="45" year="1922">
<bibRefCitation id="F168BA8130765248EF30FC4275FB053F" author="Schmidt A." box="[551,759,997,1024]" pageId="44" pageNumber="45" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 77</bibRefCitation>
</treatmentCitation>
(distribution).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30765248EDAAFB8176FA045E" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="9546C77030765248EDAAFB8176A6057F" blockId="44.[189,426,1062,1088]" box="[189,426,1062,1088]" pageId="44" pageNumber="45">
<heading id="CE0E701C30765248EDAAFB8176A6057F" bold="true" box="[189,426,1062,1088]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A78D1B6230765248EDAAFB8176A6057F" bold="true" box="[189,426,1062,1088]" pageId="44" pageNumber="45">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030765248EDAAFBF276FA045E" blockId="44.[188,1399,1107,1378]" pageId="44" pageNumber="45">
<materialsCitation id="2591CD2D30765248EDAAFBF273F205AE" collectedFrom="human faeces" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="H. Martinez &amp; P. Cely &amp; M. Cordova &amp; M. Nunez" country="Venezuela" elevation="17" latitude="10.597169" location="La Guaira" longLatPrecision="1" longitude="-66.2703" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Vargas">
<collectingCountry id="EDEE87E030765248EDAAFBF276670550" box="[189,363,1109,1135]" name="Venezuela" pageId="44" pageNumber="45">VENEZUELA</collectingCountry>
<emphasis id="A78D1B6230765248EC91FBF375050551" bold="true" box="[390,521,1108,1134]" pageId="44" pageNumber="45">
<location id="902691AB30765248EC91FBF375050551" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EC91FBF375050551" box="[390,521,1108,1134]" country="Venezuela" latitude="10.597169" longLatPrecision="1" longitude="-66.2703" name="La Guaira" pageId="44" pageNumber="45" stateProvince="Vargas">La Guaira</location>
</emphasis>
<specimenCount id="83FF0CF930765248EF37FBF375790550" box="[544,629,1108,1135]" count="1" pageId="44" pageNumber="45" type="generic">1 spec.</specimenCount>
;
<location id="902691AB30765248EF68FBF275DD0550" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EF68FBF275DD0550" box="[639,721,1109,1135]" country="Venezuela" latitude="10.597169" longLatPrecision="1" longitude="-66.2703" name="Vargas" pageId="44" pageNumber="45" stateProvince="Vargas">Vargas</location>
,
<location id="902691AB30765248EFCAFBF274570551" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EFCAFBF274570551" box="[733,859,1108,1135]" country="Venezuela" latitude="10.597169" longLatPrecision="1" longitude="-66.2703" name="La Sabana" pageId="44" pageNumber="45" stateProvince="Vargas">La Sabana</location>
;
<geoCoordinate id="F0CDA1B730765248EE71FBF473110550" box="[870,1053,1107,1135]" degrees="10" direction="north" minutes="35" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="49.81" value="10.597169">10°3549.81″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730765248E93CFBF473E70550" box="[1067,1259,1107,1135]" degrees="66" direction="west" minutes="16" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="13.08" value="-66.2703">66°1613.08″ W</geoCoordinate>
; 17 m a.s.l.;
<date id="E147E1B030765248EDAAFBD0765905AE" box="[189,341,1143,1169]" pageId="44" pageNumber="45" value="2009-07-19">
<collectingDate id="F103185830765248EDAAFBD0765905AE" box="[189,341,1143,1169]" pageId="44" pageNumber="45" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630765248EC74FBDF76FA05AE" box="[355,502,1143,1170]" pageId="44" pageNumber="45">H. Martínez</collectorName>
,
<collectorName id="380CA2A630765248EF14FBDF755505AE" box="[515,601,1143,1170]" pageId="44" pageNumber="45">P. Cely</collectorName>
,
<collectorName id="380CA2A630765248EF70FBDF75F605AE" box="[615,762,1143,1170]" pageId="44" pageNumber="45">M. Córdova</collectorName>
, and
<collectorName id="380CA2A630765248EE2CFBDF74BF05AD" box="[827,947,1144,1170]" pageId="44" pageNumber="45">M. Nuñez</collectorName>
leg.;
<collectedFrom id="26E5848D30765248EEEEFBD0739305AE" box="[1017,1183,1143,1169]" pageId="44" pageNumber="45">human faeces</collectedFrom>
;
<collectionCode id="F3E85FB530765248E9BAFBD073F205AE" box="[1197,1278,1143,1169]" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30765248E80CFBD0731A058B" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="La Sabana &amp; H. Martinez &amp; P. Cely &amp; M. Cordova &amp; M. Nunez" country="Venezuela" elevation="31" latitude="10.597169" location="Vargas" longLatPrecision="1" longitude="-66.2703" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Yaracuy">
<specimenCount id="83FF0CF930765248E80CFBD0727E05AD" box="[1307,1394,1143,1170]" count="1" pageId="44" pageNumber="45" type="generic">1 spec.</specimenCount>
; same locality data as for the preceding; 31 m a.s.l.; minced meat;
<collectionCode id="F3E85FB530765248EEA9FB3D731D058B" box="[958,1041,1178,1204]" pageId="44" pageNumber="45">CEMT</collectionCode>
.
</materialsCitation>
<materialsCitation id="2591CD2D30765248E90BFB3D76F705C6" collectedFrom="human faeces" collectingDate="2009-07-20" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="1376" latitude="10.597169" location="Aroa" longLatPrecision="1" longitude="-66.2703" pageId="44" pageNumber="45" specimenCount="4" stateProvince="Yaracuy">
<emphasis id="A78D1B6230765248E925FB3D7396058B" bold="true" box="[1074,1178,1178,1204]" pageId="44" pageNumber="45">
<collectingRegion id="573D099230765248E925FB3D7396058B" box="[1074,1178,1178,1204]" country="Venezuela" name="Yaracuy" pageId="44" pageNumber="45">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930765248E9A4FB3D7200058B" box="[1203,1292,1178,1204]" pageId="44" pageNumber="45">4 specs</specimenCount>
;
<collectingRegion id="573D099230765248E80FFB3D727E058B" box="[1304,1394,1178,1204]" country="Venezuela" name="Bolivar" pageId="44" pageNumber="45">Bolívar</collectingRegion>
,
<location id="902691AB30765248EDAAFB1A77F705E8" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EDAAFB1A77F705E8" box="[189,251,1213,1239]" country="Venezuela" latitude="10.597169" longLatPrecision="1" longitude="-66.2703" name="Aroa" pageId="44" pageNumber="45" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B730765248EC10FB1C767705E8" box="[263,379,1211,1239]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730765248EC9DFB1C750B05E8" box="[394,519,1211,1239]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
; 1376 m a.s.l.;
<date id="E147E1B030765248EFA0FB1B744605E9" box="[695,842,1212,1239]" pageId="44" pageNumber="45" value="2009-07-20">
<collectingDate id="F103185830765248EFA0FB1B744605E9" box="[695,842,1212,1239]" pageId="44" pageNumber="45" value="2009-07-20">20 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630765248EE42FB1A74F405E8" box="[853,1016,1213,1239]" pageId="44" pageNumber="45">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A630765248E914FB1A73A105E9" box="[1027,1197,1212,1239]" pageId="44" pageNumber="45">P. Colmenares</collectorName>
, and
<collectorName id="380CA2A630765248E9FEFB1A727B05E9" box="[1257,1399,1212,1239]" pageId="44" pageNumber="45">H. Martínez</collectorName>
leg.;
<collectedFrom id="26E5848D30765248EDEEFB78769205C6" box="[249,414,1247,1273]" pageId="44" pageNumber="45">human faeces</collectedFrom>
;
<collectionCode id="F3E85FB530765248ECBDFB7876F705C6" box="[426,507,1247,1273]" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30765248EF01FB7873930423" collectedFrom="human faeces" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="1373" latitude="10.340096" location="Aroa" longLatPrecision="1" longitude="-68.835175" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Bolivar">
<specimenCount id="83FF0CF930765248EF01FB78756005C5" box="[534,620,1247,1274]" count="1" pageId="44" pageNumber="45" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="573D099230765248EF6EFB7875DF05C6" box="[633,723,1247,1273]" country="Venezuela" name="Bolivar" pageId="44" pageNumber="45">Bolívar</collectingRegion>
,
<location id="902691AB30765248EFC9FB78741005C6" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EFC9FB78741005C6" box="[734,796,1247,1273]" country="Venezuela" latitude="10.340096" longLatPrecision="1" longitude="-68.835175" name="Aroa" pageId="44" pageNumber="45" stateProvince="Bolivar">Aroa</location>
;
<geoCoordinate id="F0CDA1B730765248EE3EFB7A74ED05C5" box="[809,993,1245,1274]" degrees="10" direction="north" minutes="20" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="24.35" value="10.340096">10°2024.35″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730765248EEE6FB7A73AF05C5" box="[1009,1187,1245,1274]" degrees="68" direction="west" minutes="50" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="6.61" value="-68.835175">68°506.61″ W</geoCoordinate>
; 1373 m a.s.l.;
<date id="E147E1B030765248E84EFB7876200423" pageId="44" pageNumber="45" value="2009-07-19">
<collectingDate id="F103185830765248E84EFB7876200423" pageId="44" pageNumber="45" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630765248EC2FFAA576D70423" box="[312,475,1282,1308]" pageId="44" pageNumber="45">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A630765248ECF1FAA5759D0423" box="[486,657,1282,1308]" pageId="44" pageNumber="45">P. Colmenares</collectorName>
, and
<collectorName id="380CA2A630765248EFD9FAA574500423" box="[718,860,1282,1308]" pageId="44" pageNumber="45">H. Martínez</collectorName>
leg.;
<collectedFrom id="26E5848D30765248EE89FAA5734E0423" box="[926,1090,1282,1308]" pageId="44" pageNumber="45">human faeces</collectedFrom>
;
<collectionCode id="F3E85FB530765248E959FAA573930423" box="[1102,1183,1282,1308]" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30765248E9A0FAA576FE045E" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="97" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Bolivar">
<specimenCount id="83FF0CF930765248E9A0FAA572000423" box="[1207,1292,1282,1308]" count="1" pageId="44" pageNumber="45" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="573D099230765248E80FFAA5727E0423" box="[1304,1394,1282,1308]" country="Venezuela" name="Bolivar" pageId="44" pageNumber="45">Bolívar</collectingRegion>
,
<location id="902691AB30765248EDAAFA8277F70400" LSID="urn:lsid:plazi:treatment:1D50766630765248EF4EFDE9752B07CC:902691AB30765248EDAAFA8277F70400" box="[189,251,1317,1343]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" name="Aroa" pageId="44" pageNumber="45" stateProvince="Bolivar">Aroa</location>
;
<geoCoordinate id="F0CDA1B730765248EC1FFA8476730400" box="[264,383,1315,1343]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730765248EC98FA8475020400" box="[399,526,1315,1343]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
; 97 m a.s.l.;
<date id="E147E1B030765248EFBFFA8374320401" box="[680,830,1316,1343]" pageId="44" pageNumber="45" value="2009-07-19">
<collectingDate id="F103185830765248EFBFFA8374320401" box="[680,830,1316,1343]" pageId="44" pageNumber="45" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630765248EE5CFA8274E30400" box="[843,1007,1317,1343]" pageId="44" pageNumber="45">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A630765248EEEBFA8273A40401" box="[1020,1192,1316,1343]" pageId="44" pageNumber="45">P. Colmenares</collectorName>
, and
<collectorName id="380CA2A630765248E9FFFA82727B0401" box="[1256,1399,1316,1343]" pageId="44" pageNumber="45">H. Martínez</collectorName>
leg.; minced meat;
<collectionCode id="F3E85FB530765248EC88FAE076FE045E" box="[415,498,1351,1377]" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30765248EDAAFA2F753D0700" pageId="44" pageNumber="45" type="distribution">
<paragraph id="9546C77030765248EDAAFA2F765A049D" blockId="44.[189,342,1416,1442]" box="[189,342,1416,1442]" pageId="44" pageNumber="45">
<heading id="CE0E701C30765248EDAAFA2F765A049D" bold="true" box="[189,342,1416,1442]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A78D1B6230765248EDAAFA2F765A049D" bold="true" box="[189,342,1416,1442]" pageId="44" pageNumber="45">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030765248EDAAFA1175C504EF" blockId="44.[189,713,1462,1488]" box="[189,713,1462,1488]" pageId="44" pageNumber="45">
<collectingCountry id="EDEE87E030765248EDAAFA11763804EF" box="[189,308,1462,1488]" name="Colombia" pageId="44" pageNumber="45">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030765248EC28FA1176B704EF" box="[319,443,1462,1488]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E030765248ECD1FA11751D04EF" box="[454,529,1462,1488]" name="Brazil" pageId="44" pageNumber="45">Brazil</collectingCountry>
, and
<collectingCountry id="EDEE87E030765248EF5BFA1175C904EF" box="[588,709,1462,1488]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030765248EDAAFA5176EE072E" blockId="44.[189,482,1526,1553]" box="[189,482,1526,1553]" pageId="44" pageNumber="45">
<heading id="CE0E701C30765248EDAAFA5176EE072E" bold="true" box="[189,482,1526,1553]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A78D1B6230765248EDAAFA5176EE072E" bold="true" box="[189,482,1526,1553]" pageId="44" pageNumber="45">
Subregion of
<collectingCountry id="EDEE87E030765248EC73FA5076EE072E" box="[356,482,1527,1553]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030765248EDAAF982753D0700" blockId="44.[189,561,1573,1599]" box="[189,561,1573,1599]" pageId="44" pageNumber="45">
Central
<collectingRegion id="573D099230765248EC0CF98276520700" box="[283,350,1573,1599]" country="New Zealand" name="West Coast" pageId="44" pageNumber="45">Coast</collectingRegion>
Mountain Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30765248EDAAF9C2752B07CC" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="9546C77030765248EDAAF9C276A807BF" blockId="44.[189,420,1637,1664]" box="[189,420,1637,1664]" pageId="44" pageNumber="45">
<heading id="CE0E701C30765248EDAAF9C276A807BF" bold="true" box="[189,420,1637,1664]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="A78D1B6230765248EDAAF9C276A807BF" bold="true" box="[189,420,1637,1664]" pageId="44" pageNumber="45">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030765248EDAAF933752B07CC" blockId="44.[187,1399,1684,1779]" pageId="44" pageNumber="45">
<bibRefCitation id="F168BA8130765248EDAAF93376860791" author="Schmidt A." box="[189,394,1684,1710]" pageId="44" pageNumber="45" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 77</bibRefCitation>
(
<collectingCountry id="EDEE87E030765248EC8FF933751B0791" box="[408,535,1684,1710]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230765248EF37F93375D40791" box="[544,728,1684,1710]" country="Venezuela" name="Distrito Capital" pageId="44" pageNumber="45">Distrito Capital</collectingRegion>
: Caracas, and
<collectingRegion id="573D099230765248EE6CF93374FE0791" box="[891,1010,1684,1710]" country="Venezuela" name="Carabobo" pageId="44" pageNumber="45">Carabobo</collectingRegion>
: Puerto Cabello). —
<bibRefCitation id="F168BA8130765248E9F5F933763C07EE" author="Blackwelder R. E." pageId="44" pageNumber="45" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 202</bibRefCitation>
(
<collectingCountry id="EDEE87E030765248EC29F91176B207EF" box="[318,446,1718,1744]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130765248ECE2F910759107EE" author="Roze J. A." box="[501,669,1718,1745]" pageId="44" pageNumber="45" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 43</bibRefCitation>
(
<collectingCountry id="EDEE87E030765248EFBCF911742707EF" box="[683,811,1718,1744]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130765248EE77F910733907EE" author="Martinez A." box="[864,1077,1718,1745]" pageId="44" pageNumber="45" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 43</bibRefCitation>
(
<collectingCountry id="EDEE87E030765248E953F91173C807EF" box="[1092,1220,1718,1744]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130765248E9EFF911768007CC" author="Vulcano M. A. &amp; Pereira F. S." pageId="44" pageNumber="45" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 630</bibRefCitation>
(
<collectingCountry id="EDEE87E030765248EC8BF97E751007CC" box="[412,540,1753,1779]" name="Venezuela" pageId="44" pageNumber="45">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,573 @@
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<mods:title id="7D83C0B2023C484F58D92B62053AB2F7">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:identifier id="90CF6C38AE3C8FCB3B4B435F8A36E28F" type="Zenodo-Dep">13945464</mods:identifier>
<mods:identifier id="11D8766B7EF0CAD318CA14E8F6C3C909" type="ZooBank">30872C13-516F-42FA-AFA7-30ADC6BF1BAF</mods:identifier>
</mods:mods>
<treatment id="1D507666307F5242EF4DF985733F0796" LSID="urn:lsid:plazi:treatment:1D507666307F5242EF4DF985733F0796" httpUri="http://treatment.plazi.org/id/1D507666307F5242EF4DF985733F0796" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="DDE394FB307F5241EF4DF98574D60702" box="[602,986,1570,1597]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="9546C770307F5241EF4DF98574D60702" blockId="37.[602,986,1570,1631]" box="[602,986,1570,1597]" pageId="37" pageNumber="38">
<heading id="CE0E701C307F5241EF4DF98574D60702" box="[602,986,1570,1597]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="52F9BCF3307F5241EF4DF98574D60702" ID-CoL="QP7M" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[602,986,1570,1597]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="juvencus">
<emphasis id="A78D1B62307F5241EF4DF98574340702" bold="true" box="[602,824,1570,1597]" italics="true" pageId="37" pageNumber="38">Canthon juvencus</emphasis>
Harold, 1868
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB307F5241EFF1F9E274430760" box="[742,847,1605,1631]" pageId="37" pageNumber="38" type="description">
<paragraph id="9546C770307F5241EFF1F9E274430760" blockId="37.[602,986,1570,1631]" box="[742,847,1605,1631]" pageId="37" pageNumber="38">
<figureCitation id="0DC2DBF5307F5241EFF1F9E274430760" box="[742,847,1605,1631]" captionStart="Fig" captionStartId="39.[189,232,1963,1989]" captionTargetBox="[281,1312,278,1897]" captionTargetId="figure-11@39.[270,1317,265,1919]" captionTargetPageId="39" captionText="Fig. 16. Species distribution. A. Canthon juvencus Harold, 1868. B. Canthon rufocoeruleus (Martínez, 1948). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945498" httpUri="https://zenodo.org/record/13945498/files/figure.png" pageId="37" pageNumber="38">Fig. 16A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB307F5242EDAAF921741602AC" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="9546C770307F5241EDAAF92173CE07FB" blockId="37.[189,1399,1670,1836]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="B5E9E05E307F5241EDAAF92173CE07FB" pageId="37" pageNumber="38">
<treatmentCitation id="1458E161307F5241EDAAF9217563079E" author="Harold E." box="[189,623,1670,1697]" page="111" pageId="37" pageNumber="38" year="1868">
<taxonomicName id="52F9BCF3307F5241EDAAF9217563079E" ID-CoL="QP7M" authority="Harold, 1868 a: 111" authorityName="Harold" authorityPageNumber="111" authorityYear="1868" box="[189,623,1670,1697]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="juvencus">
<emphasis id="A78D1B62307F5241EDAAF9217682079F" box="[189,398,1670,1696]" italics="true" pageId="37" pageNumber="38">Canthon juvencus</emphasis>
<bibRefCitation id="F168BA81307F5241EC85F9217563079E" author="Harold E." box="[402,623,1670,1697]" pageId="37" pageNumber="38" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold, 1868a: 111</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0307F5241E90DF921739E079F" box="[1050,1170,1670,1696]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
: Bolívar: Cartagena de Indias. Name-bearing type:
<typeStatus id="4A4279D2307F5241EF45F90D75B507FB" box="[594,697,1706,1732]" pageId="37" pageNumber="38" type="syntype">syntypes</typeStatus>
(MFNB and MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307F5241EDAAF96B74510613" blockId="37.[189,1399,1670,1836]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="B5E9E05E307F5241EDAAF96B74510613" pageId="37" pageNumber="38">
<treatmentCitation id="1458E161307F5241EDAAF96B754907D9" author="Bates H. W." box="[189,581,1740,1766]" page="30" pageId="37" pageNumber="38" year="1887">
<taxonomicName id="52F9BCF3307F5241EDAAF96B754907D9" ID-CoL="5WXGQ" authority="Bates 1887: 30" authorityName="Bates" authorityPageNumber="30" authorityYear="1887" box="[189,581,1740,1766]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="raripilus">
<emphasis id="A78D1B62307F5241EDAAF96B769C07D9" box="[189,400,1740,1766]" italics="true" pageId="37" pageNumber="38">Canthon raripilus</emphasis>
<bibRefCitation id="F168BA81307F5241EC81F96B754907D9" author="Bates H. W." box="[406,581,1740,1766]" pageId="37" pageNumber="38" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1887">Bates 1887: 30</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0307F5241EEEDF96B735007D9" box="[1018,1116,1740,1766]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
:
<collectingCountry id="EDEE87E0307F5241E972F96B73CB07D9" box="[1125,1223,1740,1766]" name="Panama" pageId="37" pageNumber="38">Panamá</collectingCountry>
: in the districts of Balboa (Pearl Islands:
<collectingCountry id="EDEE87E0307F5241EF00F949754D0637" box="[535,577,1774,1800]" name="Iceland" pageId="37" pageNumber="38">Isla</collectingCountry>
del Rey: San Miguel) and Taboga (
<collectingCountry id="EDEE87E0307F5241EEFEF94973190637" box="[1001,1045,1774,1800]" name="Iceland" pageId="37" pageNumber="38">Isla</collectingCountry>
Taboga). Name-bearing type:
<typeStatus id="4A4279D2307F5241EDFCF8B5765E0613" box="[235,338,1810,1836]" pageId="37" pageNumber="38" type="syntype">syntypes</typeStatus>
(BMNH and MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307F5242EDAAF8F576730329" blockId="37.[189,1399,1874,2005]" lastBlockId="38.[189,1400,265,915]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<treatmentCitationGroup id="B5E9E05E307F5242EDAAF8F576730329" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<taxonomicName id="52F9BCF3307F5241EDAAF8F5769C0653" ID-CoL="QP7M" authorityName="Harold" authorityYear="1868" box="[189,400,1874,1900]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="juvencus">
<emphasis id="A78D1B62307F5241EDAAF8F5769C0653" box="[189,400,1874,1900]" italics="true" pageId="37" pageNumber="38">Canthon juvencus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307F5241ECBBF8F5759C0653" author="Harold E." box="[428,656,1874,1900]" page="991" pageId="37" pageNumber="38" year="1869">
<bibRefCitation id="F168BA81307F5241ECBBF8F5759C0653" author="Harold E." box="[428,656,1874,1900]" pageId="37" pageNumber="38" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold, 1869d: 991</bibRefCitation>
</treatmentCitation>
(cited for Cartagena);
<treatmentCitation id="1458E161307F5241EE8EF8F5730C0653" author="Harold E." box="[921,1024,1874,1900]" page="17" pageId="37" pageNumber="38" year="1880">
<bibRefCitation id="F168BA81307F5241EE8EF8F5730C0653" author="Harold E." box="[921,1024,1874,1900]" pageId="37" pageNumber="38" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">1880: 17</bibRefCitation>
</treatmentCitation>
(cited for Muzo,
<collectingCountry id="EDEE87E0307F5241E9DAF8F5724A0653" box="[1229,1350,1874,1900]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161307F5241EDFDF8D276DF06B0" author="Candeze E." box="[234,467,1909,1935]" page="329" pageId="37" pageNumber="38" year="1891">
<bibRefCitation id="F168BA81307F5241EDFDF8D276DF06B0" author="Candeze E." box="[234,467,1909,1935]" pageId="37" pageNumber="38" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891: 329</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161307F5241EF59F8D2742806B0" author="Schmidt A." box="[590,804,1909,1935]" page="76" pageId="37" pageNumber="38" year="1922">
<bibRefCitation id="F168BA81307F5241EF59F8D2742806B0" author="Schmidt A." box="[590,804,1909,1935]" pageId="37" pageNumber="38" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 76</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<treatmentCitation id="1458E161307F5241E92FF8D273F106B0" author="Gillet J. J. E." box="[1080,1277,1909,1935]" page="30" pageId="37" pageNumber="38" year="1911">
<bibRefCitation id="F168BA81307F5241E92FF8D273F106B0" author="Gillet J. J. E." box="[1080,1277,1909,1935]" pageId="37" pageNumber="38" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 30</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E0307F5241EDFDF830766F068E" box="[234,355,1943,1969]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161307F5241EC8AF8307595068E" author="Balthasar V." box="[413,665,1943,1969]" page="206" pageId="37" pageNumber="38" year="1939">
<bibRefCitation id="F168BA81307F5241EC8AF8307595068E" author="Balthasar V." box="[413,665,1943,1969]" pageId="37" pageNumber="38" pagination="179 - 238" refId="ref110158" refString="Balthasar V. 1939 a. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939a: 206</bibRefCitation>
</treatmentCitation>
(key, cited for
<collectingCountry id="EDEE87E0307F5241EE47F83074C8068E" box="[848,964,1943,1969]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
and
<collectingCountry id="EDEE87E0307F5241EEE8F8307352068E" box="[1023,1118,1943,1969]" name="Mexico" pageId="37" pageNumber="38">Mexico</collectingCountry>
). —
<treatmentCitation id="1458E161307F5241E98FF830765E06EB" author="Howden H. F. &amp; Young O. P." page="28" pageId="37" pageNumber="38" year="1981">
<bibRefCitation id="F168BA81307F5241E98FF830765E06EB" author="Howden H. F. &amp; Young O. P." pageId="37" pageNumber="38" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981: 28</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="1458E161307F5241ECE8F81D75D706EB" author="Escobar F." box="[511,731,1978,2004]" page="423" pageId="37" pageNumber="38" year="1997">
<bibRefCitation id="F168BA81307F5241ECE8F81D75D706EB" author="Escobar F." box="[511,731,1978,2004]" pageId="37" pageNumber="38" pagination="19 - 430" refId="ref118000" refString="Escobar F. 1997. Estudio de la comunidad de coleopteros Coprofagos (Scarabaeidae) en un remanente de bosque seco al Norte del Tolima, Colombia. Caldasia 19 (3): 4 19 - 430." type="journal article" year="1997">Escobar 1997: 423</bibRefCitation>
</treatmentCitation>
(list);
<treatmentCitation id="1458E161307F5241EE3FF81D749206EB" author="Escobar F." box="[808,926,1978,2004]" page="207" pageId="37" pageNumber="38" year="2000">
<bibRefCitation id="F168BA81307F5241EE3FF81D749206EB" author="Escobar F." box="[808,926,1978,2004]" pageId="37" pageNumber="38" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">2000: 207</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0307F5241E95FF81D73CD06EB" box="[1096,1217,1978,2004]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161307F5242E9E1F81D76AB001C" author="Amezquita S. J. &amp; Forsyth A. &amp; Lopera A. &amp; Camacho A." lastPageId="38" lastPageNumber="39" page="119" pageId="37" pageNumber="38" year="1999">
<bibRefCitation id="F168BA81307F5242E9E1F81D76AB001C" author="Amezquita S. J. &amp; Forsyth A. &amp; Lopera A. &amp; Camacho A." lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" pagination="113 - 126" refId="ref109271" refString="Amezquita S. J., Forsyth A., Lopera A. &amp; Camacho A. 1999. Comparacion de la composicion y riqueza de especies de escarabajos coprofagos (Coleoptera: Scarabaeidae) en remanentes de bosque de la Orinoquia Colombiana. Acta Zoologica Mexicana (n. s.) 76: 113 - 126. https: // doi. org / 10.21829 / azm. 1999.76761703" type="journal article" year="1999">
Amézquita
<emphasis id="A78D1B62307C5242EDFDFEAD762B001C" box="[234,295,265,291]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
1999: 119
</bibRefCitation>
</treatmentCitation>
(biodiversity). —
<treatmentCitation id="1458E161307C5242EF90FEAE74D5001C" author="Solis A. &amp; Kohlmann B." box="[647,985,265,291]" page="27" pageId="38" pageNumber="39" year="2002">
<bibRefCitation id="F168BA81307C5242EF90FEAE74D5001C" author="Solis A. &amp; Kohlmann B." box="[647,985,265,291]" pageId="38" pageNumber="39" pagination="1 - 68" refId="ref139141" refString="Solis A. &amp; Kohlmann B. 2002. El genero Canthon en Costa Rica. Giornale Italiano di Entomologia 10: 1 - 68." type="journal article" year="2002">Solís &amp; Kohlmann 2002: 27</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="1458E161307C5242E9B6FEAE73F0001B" author="Solis A. &amp; Kohlmann B." box="[1185,1276,265,292]" page="7" pageId="38" pageNumber="39" year="2012">
<bibRefCitation id="F168BA81307C5242E9B6FEAE73F0001B" author="Solis A. &amp; Kohlmann B." box="[1185,1276,265,292]" pageId="38" pageNumber="39" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">2012: 7</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0307C5242EC00FE8B76910079" box="[279,413,300,326]" name="Costa Rica" pageId="38" pageNumber="39">Costa Rica</collectingCountry>
). —
<treatmentCitation id="1458E161307C5242ECC2FE8B74A30079" author="Bustos-Gomez L. &amp; Lopera-Toro A." box="[469,943,300,326]" page="61" pageId="38" pageNumber="39" year="2003">
<bibRefCitation id="F168BA81307C5242ECC2FE8B74A30079" author="Bustos-Gomez L. &amp; Lopera-Toro A." box="[469,943,300,326]" pageId="38" pageNumber="39" refId="ref113213" refString="Bustos-Gomez L. &amp; Lopera-Toro A. 2003. Preferencia por cebo de los escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de un remanente de bosque seco tropical al norte del Tolima (Colombia). In: Onore G., Reyes-Castillo P. &amp; Zunino M. (eds) Escarabeidos de Latinoamerica: Estado del Conocimiento; 59 - 65. m 3 m: Monografias Tercer Milenio, vol. 3. Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2003">Bustos-Gómez &amp; Lopera-Toro 2003: 61</bibRefCitation>
</treatmentCitation>
(diet). —
<treatmentCitation id="1458E161307C5242E93AFE8B73F00079" author="Noriega-Alvarado J. A." box="[1069,1276,300,326]" page="40" pageId="38" pageNumber="39" year="2004">
<bibRefCitation id="F168BA81307C5242E93AFE8B73F00079" author="Noriega-Alvarado J. A." box="[1069,1276,300,326]" firstAuthor="Noriega" pageId="38" pageNumber="39" pagination="37 - 44" refId="ref132303" refString="Noriega-Alvarado J. A. 2004. Preliminary checklist of the scarab community (Coleoptera: Scarabaeidae) at CIEM, Tinigua National Park, Meta-Colombia. Field Studies of Fauna and Flora, La Macarena, Colombia 14: 37 - 44." type="journal article" year="2004">Noriega 2004: 40</bibRefCitation>
</treatmentCitation>
(checklist for Tinigua Park,
<collectingCountry id="EDEE87E0307C5242ECA8FEE975340057" box="[447,568,334,360]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
).
<emphasis id="A78D1B62307C5242EF5AFEE8756B0057" box="[589,615,335,360]" italics="true" pageId="38" pageNumber="39"></emphasis>
<treatmentCitation id="1458E161307C5242EF79FEE974C10057" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[622,973,334,360]" page="13" pageId="38" pageNumber="39" year="2006">
<bibRefCitation id="F168BA81307C5242EF79FEE974C10057" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[622,973,334,360]" pageId="38" pageNumber="39" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B62307C5242EE37FEE874500057" box="[800,860,334,360]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2006: 13
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161307C5242E953FEE9727A0057" author="Kohlmann B. &amp; Solis A. &amp; Ortwin E. &amp; Soto X. &amp; Russo R." box="[1092,1398,334,360]" page="29" pageId="38" pageNumber="39" year="2007">
<bibRefCitation id="F168BA81307C5242E953FEE9727A0057" author="Kohlmann B. &amp; Solis A. &amp; Ortwin E. &amp; Soto X. &amp; Russo R." box="[1092,1398,334,360]" pageId="38" pageNumber="39" pagination="1 - 34" refId="ref126221" refString="Kohlmann B., Solis A., Ortwin E., Soto X. &amp; Russo R. 2007. Biodiversity, conservation, and hotspot atlas of Costa Rica: a dung beetle perspective (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 1457: 1 - 34. https: // doi. org / 10.11646 / zootaxa. 1457.1.1" type="journal article" year="2007">
Kohlmann
<emphasis id="A78D1B62307C5242E9DDFEE8720A0057" box="[1226,1286,334,360]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2007: 29
</bibRefCitation>
</treatmentCitation>
(atlas). —
<treatmentCitation id="1458E161307C5242EC7DFED675D000B4" author="Pulido-Herrera L. A. &amp; Canas R. A. R. &amp; Harders F. G. &amp; von Hildebrand P." box="[362,732,369,395]" page="307" pageId="38" pageNumber="39" year="2007">
<bibRefCitation id="F168BA81307C5242EC7DFED675D000B4" author="Pulido-Herrera L. A. &amp; Zunino M." box="[362,732,369,395]" firstAuthor="Pulido-Herrera" pageId="38" pageNumber="39" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">
Pulido-Herrera
<emphasis id="A78D1B62307C5242EF35FED5755200B4" box="[546,606,369,395]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2007: 307
</bibRefCitation>
</treatmentCitation>
(cited for Andean region of
<collectingCountry id="EDEE87E0307C5242E927FED673A500B4" box="[1072,1193,369,395]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161307C5242E9F4FED5767D0091" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." page="1294" pageId="38" pageNumber="39" year="2008">
<bibRefCitation id="F168BA81307C5242E9F4FED5767D0091" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." pageId="38" pageNumber="39" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B62307C5242E82BFED5727400B4" box="[1340,1400,369,395]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2008: 1294
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161307C5242ECF1FE3374850091" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[486,905,403,430]" page="58" pageId="38" pageNumber="39" year="2009">
<bibRefCitation id="F168BA81307C5242ECF1FE3374850091" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[486,905,403,430]" pageId="38" pageNumber="39" pagination="55 - 62" refId="ref130761" refString="Medina C. A. &amp; Pulido-Herrera L. A. 2009. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de la Orinoquia Colombiana. Biota Colombiana 10 (1 - 2): 55 - 62." type="journal article" year="2009">Medina &amp; Pulido-Herrera 2009: 58</bibRefCitation>
</treatmentCitation>
(diversity). —
<treatmentCitation id="1458E161307C5242E957FE33761A00EF" author="Arias-Buritica J. A. &amp; Delgado-Gomez P. &amp; F. A &amp; Vaz-de-Mello F. Z." page="876" pageId="38" pageNumber="39" year="2011">
<bibRefCitation id="F168BA81307C5242E957FE33761A00EF" author="Arias-Buritica J. A. &amp; Delgado-Gomez P. &amp; F. A &amp; Vaz-de-Mello F. Z." pageId="38" pageNumber="39" pagination="875 - 878" refId="ref109558" refString="Arias-Buritica J. A., Delgado-Gomez P., Gonzalez-A F. A. &amp; Vaz-de-Mello F. Z. 2011. Nuevos registros de escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) para el departamento de Choco (Colombia). Acta Zoologica Mexicana (n. s.) 27 (3): 875 - 878. https: // doi. org / 10.21829 / azm. 2011.273792" type="journal article" year="2011">
Arias-Buriticá
<emphasis id="A78D1B62307C5242E9E6FE3372210091" box="[1265,1325,404,430]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2011: 876
</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E0307C5242EC84FE1176D200EF" box="[403,478,438,464]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242ECFDFE11755200EF" box="[490,606,438,464]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
and
<collectingCountry id="EDEE87E0307C5242EF8EFE1175F700EF" box="[665,763,438,464]" name="Panama" pageId="38" pageNumber="39">Panama</collectingCountry>
). —
<treatmentCitation id="1458E161307C5242EE23FE11739800EF" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[820,1172,438,464]" page="67" pageId="38" pageNumber="39" year="2012">
<bibRefCitation id="F168BA81307C5242EE23FE11739800EF" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[820,1172,438,464]" pageId="38" pageNumber="39" refId="ref114949" refString="Cultid-Medina C. A., Medina C. A. U., Martinez-Quintero B. G., Escobar A. F. V., Constantino L. M. C. &amp; Betancur N. J. 2012. Escarabajos Coprofagos (Scarabaeinae) del Eje Cafetero: Guia para el Estudio Ecologico. WCS Colombia, Cenicafe, Villa Maria, Caldas." type="book" year="2012">
Cultid-Medina
<emphasis id="A78D1B62307C5242EEFEFE10732800EF" box="[1001,1060,438,464]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2012: 67
</bibRefCitation>
</treatmentCitation>
(guide). —
<treatmentCitation id="1458E161307C5242E831FE1076E400CC" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." page="108" pageId="38" pageNumber="39" year="2013">
<bibRefCitation id="F168BA81307C5242E831FE1076E400CC" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." pageId="38" pageNumber="39" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62307C5242EC3AFE7D766500CC" box="[301,361,473,499]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2013: 108
</bibRefCitation>
</treatmentCitation>
(list).
<emphasis id="A78D1B62307C5242EF2EFE7D755F00CC" box="[569,595,474,499]" italics="true" pageId="38" pageNumber="39"></emphasis>
<treatmentCitation id="1458E161307C5242EF4DFE7E748800CB" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[602,900,473,500]" page="195" pageId="38" pageNumber="39" year="2015">
<bibRefCitation id="F168BA81307C5242EF4DFE7E748800CB" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[602,900,473,500]" pageId="38" pageNumber="39" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B62307C5242EFDEFE7D740900CC" box="[713,773,473,499]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2015: 195
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0307C5242E926FE7E736000CC" box="[1073,1132,473,499]" name="Peru" pageId="38" pageNumber="39">Peru</collectingCountry>
). —
<treatmentCitation id="1458E161307C5242E9B1FE7E761A0329" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." page="136" pageId="38" pageNumber="39" year="2020">
<bibRefCitation id="F168BA81307C5242E9B1FE7E761A0329" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." pageId="38" pageNumber="39" pagination="134 - 141" refId="ref132145" refString="Nieto M. J. A., Giraldo E. C., Quevedo V. C. J., Chara J. &amp; Medina A. C. 2020. Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Biota Colombiana 21: 134 - 141. https: // doi. org / 10.21068 / c 2020. v 21 n 02 a 09" type="journal article" year="2020">
Nieto
<emphasis id="A78D1B62307C5242E9E6FE7D722000CC" box="[1265,1324,473,499]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2020: 136
</bibRefCitation>
</treatmentCitation>
(report).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFDB975300364" blockId="38.[189,1400,265,915]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFDB975300364" pageId="38" pageNumber="39">
<emphasis id="A78D1B62307C5242EDAAFDB976A10307" box="[189,429,542,568]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFDB976980307" ID-CoL="5WXGQ" authorityName="Bates" authorityYear="1887" box="[189,404,542,568]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="raripilus">Canthon raripilus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E161307C5242ECA0FDB975700307" author="Gillet J. J. E." box="[439,636,542,568]" page="32" pageId="38" pageNumber="39" year="1911">
<bibRefCitation id="F168BA81307C5242ECA0FDB975700307" author="Gillet J. J. E." box="[439,636,542,568]" pageId="38" pageNumber="39" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 32</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E0307C5242EE16FDB9746F0307" box="[769,867,542,568]" name="Panama" pageId="38" pageNumber="39">Panama</collectingCountry>
). —
<treatmentCitation id="1458E161307C5242EEB6FDB9738B0306" author="Schmidt A." box="[929,1159,542,569]" page="125" pageId="38" pageNumber="39" year="1920">
<bibRefCitation id="F168BA81307C5242EEB6FDB9738B0306" author="Schmidt A." box="[929,1159,542,569]" pageId="38" pageNumber="39" pagination="114 - 147" refId="ref138313" refString="Schmidt A. 1920. Beitrag zur Kenntnis der Gattungen Canthon Hffsg., Sybax Boh., Aphodius Ill., Simogenius Har., Ataenius Har. Archiv fur Naturgeschichte Abteilung A 86 (9): 114 - 147. Available from https: // www. biodiversitylibrary. org / page / 13314274 [accessed 14 Feb. 2024]." type="journal article" year="1920">Schmidt 1920: 125</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="52F9BCF3307C5242EDFDFDE676B20364" ID-CoL="QP7M" authorityName="Harold" authorityYear="1868" box="[234,446,577,603]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvencus">
<emphasis id="A78D1B62307C5242EDFDFDE676B20364" box="[234,446,577,603]" italics="true" pageId="38" pageNumber="39">Canthon juvencus</emphasis>
</taxonomicName>
, revision).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFDC3769A039F" blockId="38.[189,1400,265,915]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFDC3769A039F" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFDC376960342" authorityName="Harold" authorityYear="1868" box="[189,410,612,638]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvencum">
<emphasis id="A78D1B62307C5242EDAAFDC376960342" box="[189,410,612,638]" italics="true" pageId="38" pageNumber="39">Canthon juvencum</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307C5242ECAEFDC375C00341" author="Blackwelder R. E." box="[441,716,612,638]" page="199" pageId="38" pageNumber="39" year="1944">
<bibRefCitation id="F168BA81307C5242ECAEFDC375C00341" author="Blackwelder R. E." box="[441,716,612,638]" pageId="38" pageNumber="39" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 199</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161307C5242EE29FDC374E60341" author="Roze J. A." box="[830,1002,611,638]" page="41" pageId="38" pageNumber="39" year="1955">
<bibRefCitation id="F168BA81307C5242EE29FDC374E60341" author="Roze J. A." box="[830,1002,611,638]" pageId="38" pageNumber="39" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 41</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0307C5242E981FDC3721E0341" box="[1174,1298,612,638]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
, cited as
<taxonomicName id="52F9BCF3307C5242EDFDFD20768A039F" authorityName="Harold" authorityYear="1868" box="[234,390,647,673]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvenicum">
<emphasis id="A78D1B62307C5242EDFDFD20768A039F" box="[234,390,647,673]" italics="true" pageId="38" pageNumber="39">C. juvenicum</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFD0E74AA03FC" blockId="38.[189,1400,265,915]" box="[189,934,681,708]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFD0E74AA03FC" box="[189,934,681,708]" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFD0E750403FC" ID-CoL="5WXGQ" box="[189,520,681,707]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="juvencum" subSpecies="raripilus">
<emphasis id="A78D1B62307C5242EDAAFD0E750403FC" box="[189,520,681,707]" italics="true" pageId="38" pageNumber="39">Canthon juvencum raripilus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307C5242EF32FD0E741F03FC" author="Gacharna C." box="[549,787,681,708]" page="221" pageId="38" pageNumber="39" year="1951">
<bibRefCitation id="F168BA81307C5242EF32FD0E741F03FC" author="Gacharna C." box="[549,787,681,708]" pageId="38" pageNumber="39" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 221</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFD6B762A0237" blockId="38.[189,1400,265,915]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFD6B762A0237" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFD6B75E603DA" box="[189,746,716,742]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvencus" subGenus="Glaphyrocanthon">
<emphasis id="A78D1B62307C5242EDAAFD6B768703D9" box="[189,395,716,742]" italics="true" pageId="38" pageNumber="39">Glaphyrocanthon</emphasis>
(
<emphasis id="A78D1B62307C5242ECB7FD6B756203D9" box="[416,622,716,742]" italics="true" pageId="38" pageNumber="39">Glaphyrocanthon</emphasis>
)
<emphasis id="A78D1B62307C5242EF94FD6B75E603DA" box="[643,746,716,741]" italics="true" pageId="38" pageNumber="39">juvencus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307C5242EE18FD6B738703D9" author="Pereira F. S. &amp; Martinez A." box="[783,1163,715,742]" page="127" pageId="38" pageNumber="39" year="1956">
<bibRefCitation id="F168BA81307C5242EE18FD6B738703D9" author="Pereira F. S. &amp; Martinez A." box="[783,1163,715,742]" pageId="38" pageNumber="39" pagination="91 - 192" refId="ref135398" refString="Pereira F. S. &amp; Martinez A. 1956 a. Os generos de Canthonini Americanos. Revista Brasileira de Entomologia 6: 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956a: 127</bibRefCitation>
</treatmentCitation>
(new combination, key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFCB676F80271" blockId="38.[189,1400,265,915]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFCB676F80271" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFCB676F00214" box="[189,508,785,811]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvencus">
<emphasis id="A78D1B62307C5242EDAAFCB676F00214" box="[189,508,785,811]" italics="true" pageId="38" pageNumber="39">Glaphyrocanthon juvencus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307C5242EF09FCB6745F0214" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[542,851,785,811]" page="10" pageId="38" pageNumber="39" year="1964">
<bibRefCitation id="F168BA81307C5242EF09FCB6745F0214" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[542,851,785,811]" pageId="38" pageNumber="39" pagination="1 - 42" refId="ref130276" refString="Martinez A., Halffter G. &amp; Halffter V. 1964 a. Notas sobre el genero Glaphyrocanthon. Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez
<emphasis id="A78D1B62307C5242EF85FCB575C30214" box="[658,719,785,811]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
1964a: 10
</bibRefCitation>
</treatmentCitation>
, 12 (distribution, key). —
<treatmentCitation id="1458E161307C5242E980FCB6766C0271" author="Vulcano M. A. &amp; Pereira F. S." page="662" pageId="38" pageNumber="39" year="1964">
<bibRefCitation id="F168BA81307C5242E980FCB6766C0271" author="Vulcano M. A. &amp; Pereira F. S." pageId="38" pageNumber="39" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 662</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770307C5242EDAAFCF1741602AC" blockId="38.[189,1400,265,915]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="B5E9E05E307C5242EDAAFCF1741602AC" pageId="38" pageNumber="39">
<taxonomicName id="52F9BCF3307C5242EDAAFCF17517024F" ID-CoL="QP7M" authorityName="Harold" authorityYear="1868" box="[189,539,854,880]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juvencus" subGenus="Canthon">
<emphasis id="A78D1B62307C5242EDAAFCF1762E024F" box="[189,290,854,880]" italics="true" pageId="38" pageNumber="39">Canthon</emphasis>
(
<emphasis id="A78D1B62307C5242EC2FFCF17691024F" box="[312,413,854,880]" italics="true" pageId="38" pageNumber="39">Canthon</emphasis>
)
<emphasis id="A78D1B62307C5242ECA3FCF07517024F" box="[436,539,855,880]" italics="true" pageId="38" pageNumber="39">juvencus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161307C5242EF46FCF174B3024F" author="Halffter G. &amp; Martinez A." box="[593,959,854,880]" page="90" pageId="38" pageNumber="39" year="1977">
<bibRefCitation id="F168BA81307C5242EF46FCF174B3024F" author="Halffter G. &amp; Martinez A." box="[593,959,854,880]" pageId="38" pageNumber="39" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 90</bibRefCitation>
</treatmentCitation>
(monograph, key). —
<treatmentCitation id="1458E161307C5242E9F6FCF176C702AC" ID-DOI="http://doi.org/10.5281/zenodo.3684707" ID-Zenodo-Dep="3684707" author="Hielkema A. J. &amp; Hielkema M. A." httpUri="http://treatment.plazi.org/id/03D7FF1CFFF5FFD2FF48FD5A330FFEE1" page="61" pageId="38" pageNumber="39" year="2019">
<bibRefCitation id="F168BA81307C5242E9F6FCF176C702AC" author="Hielkema A. J. &amp; Hielkema M. A." pageId="38" pageNumber="39" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 61</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB307C5242EDAAFC1D75A80555" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="9546C770307C5242EDAAFC1D76A602EB" blockId="38.[189,426,954,980]" box="[189,426,954,980]" pageId="38" pageNumber="39">
<heading id="CE0E701C307C5242EDAAFC1D76A602EB" bold="true" box="[189,426,954,980]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A78D1B62307C5242EDAAFC1D76A602EB" bold="true" box="[189,426,954,980]" pageId="38" pageNumber="39">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770307C5242EDAAFC4F75A80555" blockId="38.[189,1400,999,1130]" pageId="38" pageNumber="39">
<materialsCitation id="2591CD2D307C5242EDAAFC4F7278053D" box="[189,1396,999,1027]" collectingDate="1959-08-20" collectionCode="CEMT" collectorName="C. J. Rosales" country="Venezuela" location="near Paso Real Guarico" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Guarico">
<collectingCountry id="EDEE87E0307C5242EDAAFC4F7667053D" box="[189,363,1000,1026]" name="Venezuela" pageId="38" pageNumber="39">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62307C5242EC9DFC4076FD053D" bold="true" box="[394,497,999,1026]" pageId="38" pageNumber="39">
<collectingRegion id="573D0992307C5242EC9DFC4076FD053D" box="[394,497,999,1026]" country="Venezuela" name="Guarico" pageId="38" pageNumber="39">Guárico</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9307C5242EF1BFC4F756E053D" box="[524,610,1000,1026]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
;
<location id="902691AB307C5242EF67FC4F7480053D" LSID="urn:lsid:plazi:treatment:1D507666307F5242EF4DF985733F0796:902691AB307C5242EF67FC4F7480053D" box="[624,908,1000,1026]" country="Venezuela" name="near Paso Real Guarico" pageId="38" pageNumber="39" stateProvince="Guarico">near Paso Real Guárico</location>
;
<date id="E147E1B0307C5242EE8EFC4F7332053C" box="[921,1086,1000,1027]" pageId="38" pageNumber="39" value="1959-08-20">
<collectingDate id="F1031858307C5242EE8EFC4F7332053C" box="[921,1086,1000,1027]" pageId="38" pageNumber="39" value="1959-08-20">20 Aug. 1959</collectingDate>
</date>
;
<collectorName id="380CA2A6307C5242E95CFC4F73D0053D" box="[1099,1244,1000,1026]" pageId="38" pageNumber="39">C.J. Rosales</collectorName>
leg.;
<collectionCode id="F3E85FB5307C5242E836FC4F7278053D" box="[1313,1396,1000,1026]" pageId="38" pageNumber="39">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D307C5242EDC3FBAC749A0578" collectedFrom="in human faeces" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="702" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B62307C5242EDC3FBAC7630051A" bold="true" box="[212,316,1035,1061]" pageId="38" pageNumber="39">
<collectingRegion id="573D0992307C5242EDC3FBAC7630051A" box="[212,316,1035,1061]" country="Venezuela" name="Yaracuy" pageId="38" pageNumber="39">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9307C5242EC4DFBAC76BD051A" box="[346,433,1035,1061]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="573D0992307C5242ECA8FBAD7515051B" box="[447,537,1034,1060]" country="Venezuela" name="Bolivar" pageId="38" pageNumber="39">Bolívar</collectingRegion>
,
<location id="902691AB307C5242EF32FBAC756F051A" LSID="urn:lsid:plazi:treatment:1D507666307F5242EF4DF985733F0796:902691AB307C5242EF32FBAC756F051A" box="[549,611,1035,1061]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" name="Aroa" pageId="38" pageNumber="39" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B7307C5242EF65FBAE75E0051A" box="[626,748,1033,1061]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B7307C5242EFEDFBAE7476051A" box="[762,890,1033,1061]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
; 702 m a.s.l.;
<date id="E147E1B0307C5242E930FBAC73B3051A" box="[1063,1215,1034,1061]" pageId="38" pageNumber="39" value="2009-07-19">
<collectingDate id="F1031858307C5242E930FBAC73B3051A" box="[1063,1215,1034,1061]" pageId="38" pageNumber="39" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A6307C5242E9DAFBAC727F051A" box="[1229,1395,1035,1061]" pageId="38" pageNumber="39">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A6307C5242EDAAFB8976640578" box="[189,360,1069,1096]" pageId="38" pageNumber="39">P. Colmenares</collectorName>
, and
<collectorName id="380CA2A6307C5242ECB1FB8975380578" box="[422,564,1069,1096]" pageId="38" pageNumber="39">H. Martínez</collectorName>
leg.;
<collectedFrom id="26E5848D307C5242EF61FB8A74350578" box="[630,825,1069,1095]" pageId="38" pageNumber="39">in human faeces</collectedFrom>
;
<collectionCode id="F3E85FB5307C5242EE52FB8A749A0578" box="[837,918,1069,1095]" pageId="38" pageNumber="39">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D307C5242EEA7FB8A75AC0555" collectingDate="2009-07-19" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="710" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Yaracuy">
<specimenCount id="83FF0CF9307C5242EEA7FB8A73090577" box="[944,1029,1069,1096]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
; same data as for the preceding except for elevation; 710 m a.s.l.;
<collectionCode id="F3E85FB5307C5242EF5AFBF775AC0555" box="[589,672,1104,1130]" pageId="38" pageNumber="39">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB307C5242EDAAFB37761C04B1" pageId="38" pageNumber="39" type="distribution">
<paragraph id="9546C770307C5242EDAAFB37765A0595" blockId="38.[189,342,1168,1194]" box="[189,342,1168,1194]" pageId="38" pageNumber="39">
<heading id="CE0E701C307C5242EDAAFB37765A0595" bold="true" box="[189,342,1168,1194]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A78D1B62307C5242EDAAFB37765A0595" bold="true" box="[189,342,1168,1194]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770307C5242EDAAFB18736F05E6" blockId="38.[189,1123,1215,1241]" box="[189,1123,1215,1241]" pageId="38" pageNumber="39">
<collectingCountry id="EDEE87E0307C5242EDAAFB18764C05E6" box="[189,320,1215,1241]" name="Costa Rica" pageId="38" pageNumber="39">Costa Rica</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242EC5BFB1876A005E6" box="[332,428,1215,1241]" name="Panama" pageId="38" pageNumber="39">Panama</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242ECA0FB18752205E6" box="[439,558,1215,1241]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242EF2EFB1875B905E6" box="[569,693,1215,1241]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242EFD7FB18743F05E6" box="[704,819,1215,1241]" name="Suriname" pageId="38" pageNumber="39">Suriname</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242EE28FB18748605E6" box="[831,906,1215,1241]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0307C5242EE81FB1874C005E6" box="[918,972,1215,1241]" name="Peru" pageId="38" pageNumber="39">Peru</collectingCountry>
and
<collectingCountry id="EDEE87E0307C5242E913FB18735205E6" box="[1028,1118,1215,1241]" name="Bolivia" pageId="38" pageNumber="39">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="9546C770307C5242EDAAFB5876E10425" blockId="38.[189,493,1279,1306]" box="[189,493,1279,1306]" pageId="38" pageNumber="39">
<heading id="CE0E701C307C5242EDAAFB5876E10425" bold="true" box="[189,493,1279,1306]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A78D1B62307C5242EDAAFB5876E10425" bold="true" box="[189,493,1279,1306]" pageId="38" pageNumber="39">
Subregions of
<collectingCountry id="EDEE87E0307C5242EC78FAA776E10425" box="[367,493,1280,1306]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770307C5242EDAAFA89761C04B1" blockId="38.[189,1400,1326,1422]" pageId="38" pageNumber="39">
Maracaibo Depression, System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní of Escudo Northeast Guiana, and Central
<collectingRegion id="573D0992307C5242E9AFFAF773F70455" box="[1208,1275,1360,1386]" country="New Zealand" name="West Coast" pageId="38" pageNumber="39">Coast</collectingRegion>
Mountain Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB307C5242EDAAFA13733F0796" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="9546C770307C5242EDAAFA1376A804F1" blockId="38.[189,420,1460,1486]" box="[189,420,1460,1486]" pageId="38" pageNumber="39">
<heading id="CE0E701C307C5242EDAAFA1376A804F1" bold="true" box="[189,420,1460,1486]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A78D1B62307C5242EDAAFA1376A804F1" bold="true" box="[189,420,1460,1486]" pageId="38" pageNumber="39">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770307C5242EDAAFA45733F0796" blockId="38.[189,1399,1506,1706]" pageId="38" pageNumber="39">
<bibRefCitation id="F168BA81307C5242EDAAFA45769604C3" author="Candeze E." box="[189,410,1506,1532]" pageId="38" pageNumber="39" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891: 329</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242ECB0FA45752A04C3" box="[423,550,1506,1532]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
: [actually,
<collectingRegion id="573D0992307C5242EF89FA45741404C3" box="[670,792,1506,1532]" country="Venezuela" name="Carabobo" pageId="38" pageNumber="39">Carabobo</collectingRegion>
]: San-Esteban). —
<bibRefCitation id="F168BA81307C5242EEE6FA4573B004C3" author="Schmidt A." box="[1009,1212,1506,1532]" pageId="38" pageNumber="39" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 76</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242E9DEFA45724504C3" box="[1225,1353,1506,1532]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242EDAAF9A276C30720" author="Blackwelder R. E." box="[189,463,1541,1567]" pageId="38" pageNumber="39" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 199</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242ECC9F9A275520720" box="[478,606,1541,1567]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242EF81F9A2730F0720" author="Pereira F. S. &amp; Martinez A." box="[662,1027,1540,1567]" pageId="38" pageNumber="39" pagination="91 - 192" refId="ref135398" refString="Pereira F. S. &amp; Martinez A. 1956 a. Os generos de Canthonini Americanos. Revista Brasileira de Entomologia 6: 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956a: 127</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242E904F9A2739F0720" box="[1043,1171,1541,1567]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242E9DDF9A2763F077E" author="Martinez A. &amp; Halffter G. &amp; Halffter V." pageId="38" pageNumber="39" pagination="1 - 42" refId="ref130276" refString="Martinez A., Halffter G. &amp; Halffter V. 1964 a. Notas sobre el genero Glaphyrocanthon. Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez
<emphasis id="A78D1B62307C5242E82BF9A2727B0720" box="[1340,1399,1541,1567]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
1964a: 10
</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242EC52F98076C9077E" box="[325,453,1575,1601]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242ECE8F9807453077E" author="Vulcano M. A. &amp; Pereira F. S." box="[511,863,1575,1601]" pageId="38" pageNumber="39" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 662</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242EE67F98074FC077E" box="[880,1008,1575,1601]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242E93CF980727A077E" author="Howden H. F. &amp; Young O. P." box="[1067,1398,1575,1602]" pageId="38" pageNumber="39" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 28</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242EDD0F9ED764B075B" box="[199,327,1610,1636]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242EC93F9ED75D3075B" author="Solis A. &amp; Kohlmann B." box="[388,735,1610,1636]" pageId="38" pageNumber="39" pagination="1 - 68" refId="ref139141" refString="Solis A. &amp; Kohlmann B. 2002. El genero Canthon en Costa Rica. Giornale Italiano di Entomologia 10: 1 - 68." type="journal article" year="2002">Solís &amp; Kohlmann 2002: 27</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242EFE4F9ED747F075B" box="[755,883,1610,1636]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81307C5242EEA5F9ED73ED075B" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[946,1249,1610,1636]" pageId="38" pageNumber="39" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B62307C5242E919F9EC7349075B" box="[1038,1093,1610,1636]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2008: 1294
</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242E9E3F9ED727F075B" box="[1268,1395,1610,1636]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992307C5242EDAAF9CA765B07B8" box="[189,343,1645,1671]" country="Venezuela" name="Bolivar" pageId="38" pageNumber="39">Bolívar state</collectingRegion>
). —
<bibRefCitation id="F168BA81307C5242EC98F9CA75D507B8" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[399,729,1645,1671]" pageId="38" pageNumber="39" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62307C5242EF36F9CA755807B8" box="[545,596,1645,1671]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2013: 108
</bibRefCitation>
(
<collectingCountry id="EDEE87E0307C5242EFFFF9CA746B07B8" box="[744,871,1645,1671]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992307C5242EE66F9CA74C207B8" box="[881,974,1645,1671]" country="Venezuela" name="Bolivar" pageId="38" pageNumber="39">Bolívar</collectingRegion>
:
<collectingCountry id="EDEE87E0307C5242EECEF9CA730F07B8" box="[985,1027,1645,1671]" name="Iceland" pageId="38" pageNumber="39">Isla</collectingCountry>
de Anacoco; Miranda: Altos de Pipe; Sucre: Araya; Yaracuy: Hacienda Guáquira; Zulia: Rosario de Perijá).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,169 @@
<document id="BEA9A6A28B5D46E205DEB81588910DF8" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666308352BDEF07FE4F742202EB" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Canthon virens subsp. chalybaeus" docType="treatment" docVersion="7" lastPageNumber="218" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="218" updateTime="1731501843003" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="432834388D3ABD9ED0E51C7B4DBDD1F7">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="1AC00EFA9A50F4678E8934D5A4BAE75C">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="8569D8C352D9C7F831A1928E175699C6">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="A934B1676F5DB6ED187D07D1056A0C71" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AC61671542B96AAA872DC1D43B0362E1">Cupello, Mario</mods:namePart>
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<mods:affiliation id="87C128A2AE7D6D06205B370EEB768E70">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="CCDB444C8189F7C04D42CC1243738643" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="7D8ADD1310CA04401733DFF09A80D88A">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="6B37D304A8D7728139774CA2ABFDAADA">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="B6CF8B3DD6971A6C7BE672EE89C8D04C" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="C889297638E1EAB48C047431543925DB">2024</mods:date>
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<taxonomicName id="52F9BCF3308352BDEF07FE4F7455033C" ID-CoL="5G74T" authority="Blanchard, 1846" box="[528,857,488,515]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="virens" subSpecies="chalybaeus">
<emphasis id="A78D1B62308352BDEF07FE4F7455033C" bold="true" box="[528,857,488,515]" italics="true" pageId="217" pageNumber="218">Canthon virens chalybaeus</emphasis>
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<bibRefCitation id="F168BA81308352BDEE76FE4F732F033D" author="Blanchard E." box="[865,1059,488,514]" pageId="217" pageNumber="218" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846</bibRefCitation>
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<treatmentCitation id="1458E161308352BDEDAAFD8D75B9037B" author="Blanchard E." box="[189,693,554,580]" page="163" pageId="217" pageNumber="218" year="1846">
<taxonomicName id="52F9BCF3308352BDEDAAFD8D76A2037B" authority="Blanchard, 1846" box="[189,430,554,580]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="chalybaeus">
<emphasis id="A78D1B62308352BDEDAAFD8D76A2037B" box="[189,430,554,580]" italics="true" pageId="217" pageNumber="218">Canthon chalybaeus</emphasis>
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<bibRefCitation id="F168BA81308352BDECA1FD8D75B9037B" author="Blanchard E." box="[438,693,554,580]" pageId="217" pageNumber="218" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846: 163</bibRefCitation>
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(original description). Type locality:
<collectingCountry id="EDEE87E0308352BDE978FD8D73C6037B" box="[1135,1226,554,580]" name="Bolivia" pageId="217" pageNumber="218">Bolivia</collectingCountry>
:
<collectingRegion id="573D0992308352BDE9C2FD8D727F037B" box="[1237,1395,554,580]" country="Bolivia" name="Cochabamba" pageId="217" pageNumber="218">Cochabamba</collectingRegion>
: Carrasco: Totora. Name-bearing type: a single known
<typeStatus id="4A4279D2308352BDEE70FDE974CE0357" box="[871,962,590,616]" pageId="217" pageNumber="218" type="syntype">syntype</typeStatus>
(MNHN), examined by FZVM.
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<treatmentCitationGroup id="B5E9E05E308352BDEDAAFDD772230391" pageId="217" pageNumber="218">
<treatmentCitation id="1458E161308352BDEDAAFDD7757503B5" author="Harold E." box="[189,633,624,650]" page="14" pageId="217" pageNumber="218" year="1868">
<taxonomicName id="52F9BCF3308352BDEDAAFDD7757503B5" ID-CoL="QPBS" authority="Harold, 1868 a: 14" authorityName="Harold" authorityPageNumber="14" authorityYear="1868" box="[189,633,624,650]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="conformis">
<emphasis id="A78D1B62308352BDEDAAFDD7769103B5" box="[189,413,624,650]" italics="true" pageId="217" pageNumber="218">Canthon conformis</emphasis>
<bibRefCitation id="F168BA81308352BDECB3FDD7757503B5" author="Harold E." box="[420,633,624,650]" pageId="217" pageNumber="218" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold, 1868a: 14</bibRefCitation>
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, 86 (original description, key). Type locality:
<collectingCountry id="EDEE87E0308352BDE99EFDD773DD03B5" box="[1161,1233,624,650]" name="Brazil" pageId="217" pageNumber="218">Brazil</collectingCountry>
and
<collectingCountry id="EDEE87E0308352BDE81FFDD6727F03B4" box="[1288,1395,625,651]" name="Uruguay" pageId="217" pageNumber="218">Uruguay</collectingCountry>
:
<collectingRegion id="573D0992308352BDEDFEFD3476770392" box="[233,379,659,685]" country="Uruguay" name="Montevideo" pageId="217" pageNumber="218">Montevideo</collectingRegion>
. Name-bearing type: two known
<typeStatus id="4A4279D2308352BDEE15FD3374650391" box="[770,873,660,686]" pageId="217" pageNumber="218" type="syntype">syntypes</typeStatus>
(MNHN, ZSM), examined by FZVM.
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<heading id="CE0E701C308352BDEDAAFD72765A03D0" bold="true" box="[189,342,725,751]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A78D1B62308352BDEDAAFD72765A03D0" bold="true" box="[189,342,725,751]" pageId="217" pageNumber="218">Distribution</emphasis>
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Supposedly
<collectingCountry id="EDEE87E0308352BDEC42FCA476DD0222" box="[341,465,771,797]" name="Venezuela" pageId="217" pageNumber="218">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDECF3FCA475910222" box="[484,669,771,797]" name="French Guiana" pageId="217" pageNumber="218">French Guiana</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEFA6FCA475F00222" box="[689,764,771,797]" name="Brazil" pageId="217" pageNumber="218">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEE07FCA374450221" box="[784,841,772,798]" name="Peru" pageId="217" pageNumber="218">Peru</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEE4BFCA474B90222" box="[860,949,771,797]" name="Bolivia" pageId="217" pageNumber="218">Bolivia</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEEDFFCA4734C0222" box="[968,1088,771,797]" name="Argentina" pageId="217" pageNumber="218">Argentina</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDE943FCA373CE0221" box="[1108,1218,772,798]" name="Paraguay" pageId="217" pageNumber="218">Paraguay</collectingCountry>
, and
<collectingCountry id="EDEE87E0308352BDE807FCA3727B0221" box="[1296,1399,772,798]" name="Uruguay" pageId="217" pageNumber="218">Uruguay</collectingCountry>
(
<bibRefCitation id="F168BA81308352BDEDD2FC8076C8027E" author="Schoolmeesters P." box="[197,452,807,833]" pageId="217" pageNumber="218" refId="ref138584" refString="Schoolmeesters P. 2023. World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Vandepitte L., Hobern D., Remsen D., Schalk P., DeWalt R. E., Keping M., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., Adriaenssens E. M., Aedo C., Aescht E., Akkari N. et al. (eds) Catalogue of Life Checklist (Version 2023 - 03 - 06). Available from https: // www. catalogueoflife. org / data / dataset / 1027 [accessed 3 Apr. 2023]." type="url" year="2023">Schoolmeesters 2023</bibRefCitation>
; but see
<bibRefCitation id="F168BA81308352BDEF3DFC807477027E" author="Hielkema A. J. &amp; Hielkema M. A." box="[554,891,807,833]" pageId="217" pageNumber="218" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB308352BDEDAAFCCF742202EB" pageId="217" pageNumber="218" type="reference_group">
<paragraph id="9546C770308352BDEDAAFCCF76A802BD" blockId="217.[189,420,872,898]" box="[189,420,872,898]" pageId="217" pageNumber="218">
<heading id="CE0E701C308352BDEDAAFCCF76A802BD" bold="true" box="[189,420,872,898]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A78D1B62308352BDEDAAFCCF76A802BD" bold="true" box="[189,420,872,898]" pageId="217" pageNumber="218">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308352BDEDAAFC30742202EB" blockId="217.[189,1398,919,980]" pageId="217" pageNumber="218">
<bibRefCitation id="F168BA81308352BDEDAAFC307695028E" author="Schmidt A." box="[189,409,919,945]" pageId="217" pageNumber="218" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 74</bibRefCitation>
(cited as
<taxonomicName id="52F9BCF3308352BDEF0AFC3075B6028E" authorityName="Harold" authorityYear="1868" box="[541,698,919,945]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="conformis">
<emphasis id="A78D1B62308352BDEF0AFC3075B6028E" box="[541,698,919,945]" italics="true" pageId="217" pageNumber="218">C. conformis</emphasis>
</taxonomicName>
). —
<bibRefCitation id="F168BA81308352BDEE14FC30732D028E" author="Blackwelder R. E." box="[771,1057,919,945]" pageId="217" pageNumber="218" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 198</bibRefCitation>
(cited as
<emphasis id="A78D1B62308352BDE9B3FC307237028E" box="[1188,1339,919,945]" italics="true" pageId="217" pageNumber="218">C. conforme</emphasis>
). —
<bibRefCitation id="F168BA81308352BDEDAAFC1D751902EB" author="Vulcano M. A. &amp; Pereira F. S." box="[189,533,954,980]" pageId="217" pageNumber="218" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 608</bibRefCitation>
(cited as
<taxonomicName id="52F9BCF3308352BDEF91FC1D741102EB" authorityName="Harold" authorityYear="1868" box="[646,797,954,980]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="conformis">
<emphasis id="A78D1B62308352BDEF91FC1D741102EB" box="[646,797,954,980]" italics="true" pageId="217" pageNumber="218">C. conformis</emphasis>
</taxonomicName>
).
</paragraph>
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<document id="95C0D29FBD775A9DFBA3483B14095A80" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666308352BEEF0AFA6075C700A8" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum carinatum" docType="treatment" docVersion="8" lastPageNumber="219" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="218" updateTime="1731501843003" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EDC575E59FBEDACB1BF828258B9E37F8">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="358E617E7A7157768BECF7A9900E86B7">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="F426442142A95DE94EF037C1A0E741DE">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="A4B1ABB24C5336FDFEDFC0481B0F4D1F">Cupello, Mario</mods:namePart>
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<mods:affiliation id="5593582B228ACEC8A5289A91A49C5A65">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="83B43775FBDA9F49F48C443A592FED6F" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="A29041D57A5C55CF7ACCBC4A58E5E724">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="30F7EDB7D3CF0ECB3A82D69E7EE8739F">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="E4B04A1810C7FD13BBD937FDEA4E563C" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<paragraph id="9546C770308352BDEF0AFA60731A04DD" blockId="217.[541,1046,1479,1506]" box="[541,1046,1479,1506]" pageId="217" pageNumber="218">
<heading id="CE0E701C308352BDEF0AFA60731A04DD" box="[541,1046,1479,1506]" centered="true" fontSize="11" level="2" pageId="217" pageNumber="218" reason="2">
<taxonomicName id="52F9BCF3308352BDEF0AFA60731A04DD" ID-CoL="34KBB" authority="(Westwood, 1837)" baseAuthorityName="Westwood" baseAuthorityYear="1837" box="[541,1046,1479,1506]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="A78D1B62308352BDEF0AFA60743404DD" bold="true" box="[541,824,1479,1506]" italics="true" pageId="217" pageNumber="218">Deltochilum carinatum</emphasis>
(
<bibRefCitation id="F168BA81308352BDEE50FA6F730204DD" author="Westwood J. O." box="[839,1038,1480,1506]" pageId="217" pageNumber="218" pagination="251 - 257" refId="ref141840" refString="Westwood J. O. 1837. Descriptions of some new species of exotic coleopterus insects from the collection of Sir Patrick Walker. Magazine of Zoology and Botany 1: 251 - 257. Available from https: // www. biodiversitylibrary. org / page / 40029670 [accessed 14 Feb. 2024]." type="journal article" year="1837">Westwood, 1837</bibRefCitation>
)
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<treatmentCitation id="1458E161308352BDEDAAF9AE7591071B" author="Westwood J. O." box="[189,669,1545,1572]" page="256" pageId="217" pageNumber="218" year="1837">
<taxonomicName id="52F9BCF3308352BDEDAAF9AE7682071C" ID-CoL="3MY7X" authority="Westwood, 1837" box="[189,398,1545,1571]" class="Insecta" family="Scarabaeidae" genus="Hyboma" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="A78D1B62308352BDEDAAF9AE7682071C" box="[189,398,1545,1571]" italics="true" pageId="217" pageNumber="218">Hyboma carinata</emphasis>
</taxonomicName>
<bibRefCitation id="F168BA81308352BDEC8FF9AE7591071B" author="Westwood J. O." box="[408,669,1545,1572]" pageId="217" pageNumber="218" pagination="251 - 257" refId="ref141840" refString="Westwood J. O. 1837. Descriptions of some new species of exotic coleopterus insects from the collection of Sir Patrick Walker. Magazine of Zoology and Botany 1: 251 - 257. Available from https: // www. biodiversitylibrary. org / page / 40029670 [accessed 14 Feb. 2024]." type="journal article" year="1837">Westwood, 1837: 256</bibRefCitation>
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(original description). Type locality: South America. Name-bearing type: a single known
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(OUMNH), examined by FZVM.
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</paragraph>
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<treatmentCitationGroup id="B5E9E05E308352BDEDAAF9E876B507EB" pageId="217" pageNumber="218">
<treatmentCitation id="1458E161308352BDEDAAF9E875B20755" author="Castelnau" box="[189,702,1615,1642]" page="74" pageId="217" pageNumber="218" year="1840">
<taxonomicName id="52F9BCF3308352BDEDAAF9E875B20755" ID-CoL="3MY82" authority="Castelnau, 1840: 74" authorityName="Castelnau" authorityPageNumber="74" authorityYear="1840" box="[189,702,1615,1642]" class="Insecta" family="Scarabaeidae" genus="Hyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="dromedarius">
<emphasis id="A78D1B62308352BDEDAAF9E876CC0756" box="[189,448,1615,1641]" italics="true" pageId="217" pageNumber="218">Hyboma dromedarius</emphasis>
<bibRefCitation id="F168BA81308352BDECDDF9E875B20755" author="Castelnau" box="[458,702,1615,1642]" pageId="217" pageNumber="218" refId="ref113936" refString="Castelnau, Comte de 1840. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des insectes coleopteres. Tome deuxieme. Dumenil, Paris. https: // doi. org / 10.5962 / bhl. title. 47104" type="book" year="1840">Castelnau, 1840: 74</bibRefCitation>
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.
<typeStatus id="4A4279D2308352BDEFDCF9F774090755" box="[715,773,1616,1642]" pageId="217" pageNumber="218">Type</typeStatus>
locality:
<collectingCountry id="EDEE87E0308352BDEE6DF9E874C90756" box="[890,965,1615,1641]" name="Brazil" pageId="217" pageNumber="218">Brazil</collectingCountry>
. Name-bearing
<typeStatus id="4A4279D2308352BDE991F9F773B10755" box="[1158,1213,1616,1642]" pageId="217" pageNumber="218">type</typeStatus>
: originally said to be in the Gory collection, whose material was eventually scattered across European museums (
<treatmentCitation id="1458E161308352BDEDE5F93176FC078F" author="Maldaner M. E. &amp; Cupello M. &amp; Ferreira D. C. &amp; Vaz-de-Mello F. Z." box="[242,496,1686,1712]" pageId="217" pageNumber="218" year="2017">
<bibRefCitation id="F168BA81308352BDEDE5F93176FC078F" author="Maldaner M. E. &amp; Cupello M. &amp; Ferreira D. C. &amp; Vaz-de-Mello F. Z." box="[242,496,1686,1712]" pageId="217" pageNumber="218" pagination="83 - 102" refId="ref129157" refString="Maldaner M. E., Cupello M., Ferreira D. C. &amp; Vaz-de-Mello F. Z. 2017. Type specimens and names assigned to Coprophanaeus (Megaphanaeus) d'Olsoufieff, 1924, the largest New World dung beetles (Coleoptera: Scarabaeidae: Phanaeini). Zootaxa 4272 (1): 83 - 102. https: // doi. org / 10.11646 / zootaxa. 4272.1.4" type="journal article" year="2017">
Maldaner
<emphasis id="A78D1B62308352BDEC7BF93076A4078F" box="[364,424,1686,1712]" italics="true" pageId="217" pageNumber="218">et al.</emphasis>
2017
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). The particular whereabouts of the
<typeStatus id="4A4279D2308352BDEE8EF93174C0078F" box="[921,972,1686,1712]" pageId="217" pageNumber="218">type</typeStatus>
specimens of
<taxonomicName id="52F9BCF3308352BDE960F931727B078F" ID-CoL="3MY82" authorityName="Castelnau" authorityYear="1840" box="[1143,1399,1686,1712]" class="Insecta" family="Scarabaeidae" genus="Hyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="dromedarius">
<emphasis id="A78D1B62308352BDE960F931727B078F" box="[1143,1399,1686,1712]" italics="true" pageId="217" pageNumber="218">Hyboma dromedarius</emphasis>
</taxonomicName>
is unknown to us.
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<paragraph id="9546C770308352BDEDAAF95D765A062B" blockId="217.[189,342,1786,1812]" box="[189,342,1786,1812]" pageId="217" pageNumber="218">
<heading id="CE0E701C308352BDEDAAF95D765A062B" bold="true" box="[189,342,1786,1812]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A78D1B62308352BDEDAAF95D765A062B" bold="true" box="[189,342,1786,1812]" pageId="217" pageNumber="218">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308352BDEDAAF88E753C0659" blockId="217.[189,1398,1833,1894]" pageId="217" pageNumber="218">
Supposedly
<collectingCountry id="EDEE87E0308352BDEC43F88E76C7067C" box="[340,459,1833,1859]" name="Colombia" pageId="217" pageNumber="218">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDECCAF88E7555067C" box="[477,601,1833,1859]" name="Venezuela" pageId="217" pageNumber="218">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEF7CF88E75BA067C" box="[619,694,1833,1859]" name="Brazil" pageId="217" pageNumber="218">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEFDEF88E742B067C" box="[713,807,1833,1859]" name="Guyana" pageId="217" pageNumber="218">Guyana</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDEE2DF88E74A1067C" box="[826,941,1833,1859]" name="Suriname" pageId="217" pageNumber="218">Suriname</collectingCountry>
, French Guiana,
<collectingCountry id="EDEE87E0308352BDE99DF88E73E2067C" box="[1162,1262,1833,1859]" name="Ecuador" pageId="217" pageNumber="218">Ecuador</collectingCountry>
,
<collectingCountry id="EDEE87E0308352BDE816F88D7236067B" box="[1281,1338,1834,1860]" name="Peru" pageId="217" pageNumber="218">Peru</collectingCountry>
, and
<collectingCountry id="EDEE87E0308352BDEDAAF8EB761F0659" box="[189,275,1868,1894]" name="Bolivia" pageId="217" pageNumber="218">Bolivia</collectingCountry>
(
<bibRefCitation id="F168BA81308352BDEC35F8EB75280659" author="Schoolmeesters P." box="[290,548,1868,1894]" pageId="217" pageNumber="218" refId="ref138584" refString="Schoolmeesters P. 2023. World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Vandepitte L., Hobern D., Remsen D., Schalk P., DeWalt R. E., Keping M., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., Adriaenssens E. M., Aedo C., Aescht E., Akkari N. et al. (eds) Catalogue of Life Checklist (Version 2023 - 03 - 06). Available from https: // www. catalogueoflife. org / data / dataset / 1027 [accessed 3 Apr. 2023]." type="url" year="2023">Schoolmeesters 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB308052BEEDAAFEAF75C700A8" pageId="218" pageNumber="219" type="reference_group">
<paragraph id="9546C770308052BEEDAAFEAF76A8001C" blockId="218.[189,420,264,291]" box="[189,420,264,291]" pageId="218" pageNumber="219">
<heading id="CE0E701C308052BEEDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" fontSize="11" level="3" pageId="218" pageNumber="219" reason="3">
<emphasis id="A78D1B62308052BEEDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" pageId="218" pageNumber="219">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308052BEEDAAFE9075C700A8" blockId="218.[189,1398,311,408]" pageId="218" pageNumber="219">
<bibRefCitation id="F168BA81308052BEEDAAFE907529006E" author="Pereira F. S. &amp; D'Andretta M. A. V." box="[189,549,311,338]" pageId="218" pageNumber="219" pagination="7 - 50" refId="ref135363" refString="Pereira F. S. &amp; D'Andretta M. A. V. 1955. The species of Deltochilum of the subgenus Calhyboma Kolbe. Revista Brasileira de Entomologia 4: 7 - 50." type="book chapter" year="1955">Pereira &amp; dAndretta 1955: 8</bibRefCitation>
, 22 (
<collectingCountry id="EDEE87E0308052BEEF70FE9075EA006E" box="[615,742,311,337]" name="Venezuela" pageId="218" pageNumber="219">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992308052BEEFE4FE90747F006E" box="[755,883,311,337]" country="Venezuela" name="Amazonas" pageId="218" pageNumber="219">Amazonas</collectingRegion>
: Yavita). —
<bibRefCitation id="F168BA81308052BEE907FE90727A006E" author="Vulcano M. A. &amp; Pereira F. S." box="[1040,1398,311,337]" pageId="218" pageNumber="219" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 643</bibRefCitation>
(
<collectingCountry id="EDEE87E0308052BEEDD2FEFD7649004B" box="[197,325,346,372]" name="Venezuela" pageId="218" pageNumber="219">Venezuela</collectingCountry>
). — González
<emphasis id="A78D1B62308052BEECEDFEFC7522004B" box="[506,558,346,372]" italics="true" pageId="218" pageNumber="219">et al</emphasis>
. 2009: 263 (
<collectingCountry id="EDEE87E0308052BEEFDFFEFD7444004B" box="[712,840,346,372]" name="Venezuela" pageId="218" pageNumber="219">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81308052BEEE94FEFD73BF004B" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[899,1203,346,373]" pageId="218" pageNumber="219" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B62308052BEE911FEFC7335004B" box="[1030,1081,346,372]" italics="true" pageId="218" pageNumber="219">et al</emphasis>
. 2019: 86
</bibRefCitation>
(
<collectingCountry id="EDEE87E0308052BEE9D3FEFD7248004B" box="[1220,1348,346,372]" name="Venezuela" pageId="218" pageNumber="219">Venezuela</collectingCountry>
). —
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(
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(original description). Type locality:
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Presumably
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and
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(
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;
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).
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<emphasis id="A78D1B62308552BBEDAAFBF276950550" bold="true" box="[189,409,1109,1135]" pageId="223" pageNumber="224">Literature record</emphasis>
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<bibRefCitation id="F168BA81308552BBEDAAFB24751405A1" author="Vulcano M. A. &amp; Pereira F. S." box="[189,536,1155,1182]" pageId="223" pageNumber="224" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 579</bibRefCitation>
, 581 (
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<emphasis id="A78D1B62308552BBEDAAFB6276E905E0" bold="true" box="[189,485,1221,1247]" pageId="223" pageNumber="224">III Erroneous records</emphasis>
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View file

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<heading id="CE0E701C308652B8EF0AFAA1731A041E" box="[541,1046,1286,1313]" centered="true" fontSize="11" level="2" pageId="220" pageNumber="221" reason="2">
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<emphasis id="A78D1B62308652B8EF0AFAA17432041E" bold="true" box="[541,830,1286,1313]" italics="true" pageId="220" pageNumber="221">Dichotomius subaeneus</emphasis>
(
<bibRefCitation id="F168BA81308652B8EE59FAA17302041E" author="Castelnau" box="[846,1038,1286,1313]" pageId="220" pageNumber="221" refId="ref113936" refString="Castelnau, Comte de 1840. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des insectes coleopteres. Tome deuxieme. Dumenil, Paris. https: // doi. org / 10.5962 / bhl. title. 47104" type="book" year="1840">Castelnau, 1840</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB308652B8EDAAFAEF76480496" pageId="220" pageNumber="221" type="reference_group">
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<emphasis id="A78D1B62308652B8EDAAFAEF768D045D" box="[189,385,1352,1378]" italics="true" pageId="220" pageNumber="221">Copris subaenea</emphasis>
</taxonomicName>
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</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0308652B8E900FAEF73CB045D" box="[1047,1223,1352,1378]" name="French Guiana" pageId="220" pageNumber="221">French Guiana</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D2308652B8EC39FACC769604BA" box="[302,410,1387,1413]" pageId="220" pageNumber="221" type="lectotype">lectotype</typeStatus>
(BMNH), designated by
<bibRefCitation id="F168BA81308652B8EFC1FACC73C004BA" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." box="[726,1228,1387,1413]" pageId="220" pageNumber="221" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello (2021)</bibRefCitation>
, examined by FZVM.
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<paragraph id="9546C770308652B8EDAAFA68765A04D6" blockId="220.[189,342,1487,1513]" box="[189,342,1487,1513]" pageId="220" pageNumber="221">
<heading id="CE0E701C308652B8EDAAFA68765A04D6" bold="true" box="[189,342,1487,1513]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="A78D1B62308652B8EDAAFA68765A04D6" bold="true" box="[189,342,1487,1513]" pageId="220" pageNumber="221">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308652B8EDAAFA5974FA0704" blockId="220.[188,1399,1534,1596]" pageId="220" pageNumber="221">
<bibRefCitation id="F168BA81308652B8EDAAFA5975910727" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." box="[189,669,1534,1560]" pageId="220" pageNumber="221" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello (2021)</bibRefCitation>
recorded the species from
<collectingCountry id="EDEE87E0308652B8EEC3FA59733E0727" box="[980,1074,1534,1560]" name="Guyana" pageId="220" pageNumber="221">Guyana</collectingCountry>
, French Guiana, and
<collectingCountry id="EDEE87E0308652B8E830FA59727E0727" box="[1319,1394,1534,1560]" name="Brazil" pageId="220" pageNumber="221">Brazil</collectingCountry>
.
<bibRefCitation id="F168BA81308652B8EDABF986750C0704" author="Vulcano M. A. &amp; Pereira F. S." box="[188,512,1569,1596]" pageId="220" pageNumber="221" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereiras (1967)</bibRefCitation>
record from
<collectingCountry id="EDEE87E0308652B8EF8FF986741D0704" box="[664,785,1569,1595]" name="Venezuela" pageId="220" pageNumber="221">Venezuela</collectingCountry>
was not addressed.
</paragraph>
<paragraph id="9546C770308652B8EDAAF9C476950742" blockId="220.[189,409,1635,1661]" box="[189,409,1635,1661]" pageId="220" pageNumber="221">
<heading id="CE0E701C308652B8EDAAF9C476950742" bold="true" box="[189,409,1635,1661]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="A78D1B62308652B8EDAAF9C476950742" bold="true" box="[189,409,1635,1661]" pageId="220" pageNumber="221">Literature record</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308652B8EDAAF93675BC0794" blockId="220.[189,688,1681,1708]" box="[189,688,1681,1708]" pageId="220" pageNumber="221">
<bibRefCitation id="F168BA81308652B8EDAAF93675190793" author="Vulcano M. A. &amp; Pereira F. S." box="[189,533,1681,1708]" pageId="220" pageNumber="221" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 587</bibRefCitation>
(
<collectingCountry id="EDEE87E0308652B8EF33F93675A80794" box="[548,676,1681,1707]" name="Venezuela" pageId="220" pageNumber="221">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,249 @@
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<taxonomicName id="52F9BCF3308B52B5EF27FEAF730F001C" ID-CoL="6742F" authority="(Harold, 1868)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[560,1027,264,291]" class="Insecta" family="Scarabaeidae" genus="Anisocanthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="A78D1B62308B52B5EF27FEAF7442001C" bold="true" box="[560,846,264,291]" italics="true" pageId="209" pageNumber="210">Anisocanthon sericinus</emphasis>
(Harold, 1868)
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<subSubSection id="DDE394FB308B52B5EDAAFEED733803DC" pageId="209" pageNumber="210" type="reference_group">
<paragraph id="9546C770308B52B5EDAAFEED75C00094" blockId="209.[189,1399,330,427]" pageId="209" pageNumber="210">
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<treatmentCitation id="1458E161308B52B5EDAAFEED7579005A" author="Harold E." box="[189,629,330,357]" page="18" pageId="209" pageNumber="210" year="1868">
<taxonomicName id="52F9BCF3308B52B5EDAAFEED7579005A" ID-CoL="68XH9" authority="Harold, 1868 a: 18" authorityName="Harold" authorityPageNumber="18" authorityYear="1868" box="[189,629,330,357]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="A78D1B62308B52B5EDAAFEED7699005B" box="[189,405,330,356]" italics="true" pageId="209" pageNumber="210">Canthon sericinus</emphasis>
<bibRefCitation id="F168BA81308B52B5EC89FEED7579005A" author="Harold E." box="[414,629,330,357]" pageId="209" pageNumber="210" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold, 1868a: 18</bibRefCitation>
</taxonomicName>
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(original description). Type locality:
<collectingCountry id="EDEE87E0308B52B5E924FEED7372005B" box="[1075,1150,330,356]" name="Brazil" pageId="209" pageNumber="210">Brazil</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D2308B52B5EDFBFEC9765F00B7" box="[236,339,366,392]" pageId="209" pageNumber="210" type="holotype">holotype</typeStatus>
(unknown whereabouts), said by
<treatmentCitation id="1458E161308B52B5EFF6FEC9749900B8" author="Harold E." box="[737,917,365,392]" pageId="209" pageNumber="210" year="1868">
<bibRefCitation id="F168BA81308B52B5EFF6FEC9749900B8" author="Harold E." box="[737,917,365,392]" pageId="209" pageNumber="210" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold (1868a)</bibRefCitation>
</treatmentCitation>
to be a female and belong to the MFNB, not found there by FZVM in 20132014.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFE757414032F" blockId="209.[189,1399,466,739]" pageId="209" pageNumber="210">
<treatmentCitationGroup id="B5E9E05E308B52B5EDAAFE757414032F" pageId="209" pageNumber="210">
<taxonomicName id="52F9BCF3308B52B5EDAAFE75766000D3" ID-CoL="QPBB" authorityName="Harold" authorityYear="1868" box="[189,364,466,492]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="A78D1B62308B52B5EDAAFE75766000D3" box="[189,364,466,492]" italics="true" pageId="209" pageNumber="210">Canthon asper</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161308B52B5EC9AFE75755300D3" author="Harold E." box="[397,607,466,493]" page="11" pageId="209" pageNumber="210" year="1868">
<bibRefCitation id="F168BA81308B52B5EC9AFE75755300D3" author="Harold E." box="[397,607,466,493]" pageId="209" pageNumber="210" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold 1868a: 11</bibRefCitation>
</treatmentCitation>
, 29 (original description). Type locality:
<collectingCountry id="EDEE87E0308B52B5E95CFE7573CF00D3" box="[1099,1219,466,492]" name="Colombia" pageId="209" pageNumber="210">Colombia</collectingCountry>
. Name-bearing type: a
<typeStatus id="4A4279D2308B52B5EC55FE517691032F" box="[322,413,502,528]" pageId="209" pageNumber="210" type="syntype">syntype</typeStatus>
(MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFDBE726E030C" blockId="209.[189,1399,466,739]" box="[189,1378,536,563]" pageId="209" pageNumber="210">
<treatmentCitationGroup id="B5E9E05E308B52B5EDAAFDBE726E030C" box="[189,1378,536,563]" pageId="209" pageNumber="210">
<taxonomicName id="52F9BCF3308B52B5EDAAFDBE7691030D" ID-CoL="62L63" authorityName="Hoffmannsegg" authorityYear="1817" box="[189,413,537,563]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="sericinum">
<emphasis id="A78D1B62308B52B5EDAAFDBE7691030D" box="[189,413,537,563]" italics="true" pageId="209" pageNumber="210">Canthon sericinum</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161308B52B5ECADFDBE75C0030C" author="Blackwelder R. E." box="[442,716,537,563]" page="201" pageId="209" pageNumber="210" year="1944">
<bibRefCitation id="F168BA81308B52B5ECADFDBE75C0030C" author="Blackwelder R. E." box="[442,716,537,563]" pageId="209" pageNumber="210" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 201</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="1458E161308B52B5EE94FDBE7321030C" author="Roze J. A." box="[899,1069,536,563]" page="41" pageId="209" pageNumber="210" year="1955">
<bibRefCitation id="F168BA81308B52B5EE94FDBE7321030C" author="Roze J. A." box="[899,1069,536,563]" pageId="209" pageNumber="210" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 41</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0308B52B5E9C0FDBE7259030C" box="[1239,1365,537,563]" name="Venezuela" pageId="209" pageNumber="210">Venezuela</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFD9B72090346" blockId="209.[189,1399,466,739]" pageId="209" pageNumber="210">
<treatmentCitationGroup id="B5E9E05E308B52B5EDAAFD9B72090346" pageId="209" pageNumber="210">
<emphasis id="A78D1B62308B52B5EDAAFD9B76BC0369" box="[189,432,572,598]" italics="true" pageId="209" pageNumber="210">
<taxonomicName id="52F9BCF3308B52B5EDAAFD9B769B0369" ID-CoL="68XH9" authorityName="Harold" authorityYear="1868" box="[189,407,572,598]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="sericinus">Canthon sericinus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E161308B52B5ECADFD9B759D0369" author="Harold E." box="[442,657,572,598]" page="11" pageId="209" pageNumber="210" year="1869">
<bibRefCitation id="F168BA81308B52B5ECADFD9B759D0369" author="Harold E." box="[442,657,572,598]" pageId="209" pageNumber="210" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 11</bibRefCitation>
</treatmentCitation>
, 29 (catalogue). —
<treatmentCitation id="1458E161308B52B5EE90FD9B73400369" author="Gillet J. J. E." box="[903,1100,572,598]" page="33" pageId="209" pageNumber="210" year="1911">
<bibRefCitation id="F168BA81308B52B5EE90FD9B73400369" author="Gillet J. J. E." box="[903,1100,572,598]" pageId="209" pageNumber="210" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 33</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161308B52B5E802FD9B76580346" author="Schmidt A." page="63" pageId="209" pageNumber="210" year="1922">
<bibRefCitation id="F168BA81308B52B5E802FD9B76580346" author="Schmidt A." pageId="209" pageNumber="210" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 63</bibRefCitation>
</treatmentCitation>
, 80 (cited). —
<treatmentCitation id="1458E161308B52B5EF12FDF875F20346" author="Balthasar V." box="[517,766,607,633]" page="184" pageId="209" pageNumber="210" year="1939">
<bibRefCitation id="F168BA81308B52B5EF12FDF875F20346" author="Balthasar V." box="[517,766,607,633]" pageId="209" pageNumber="210" pagination="179 - 238" refId="ref110158" refString="Balthasar V. 1939 a. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939a: 184</bibRefCitation>
</treatmentCitation>
(key).
<emphasis id="A78D1B62308B52B5EE45FDC774600346" box="[850,876,608,633]" italics="true" pageId="209" pageNumber="210"></emphasis>
<treatmentCitation id="1458E161308B52B5EE63FDF873660346" author="Martinez A." box="[884,1130,607,633]" page="187" pageId="209" pageNumber="210" year="1949">
<bibRefCitation id="F168BA81308B52B5EE63FDF873660346" author="Martinez A." box="[884,1130,607,633]" pageId="209" pageNumber="210" pagination="175 - 193" refId="ref129675" refString="Martinez A. 1949 b. Insectos nuevos o poco conocidos VII. Revista de la Sociedad Entomologica Argentina 14 (4): 175 - 193." type="journal article" year="1949">Martínez 1949b: 187</bibRefCitation>
</treatmentCitation>
(comments).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFD25733803DC" blockId="209.[189,1399,466,739]" pageId="209" pageNumber="210">
<treatmentCitationGroup id="B5E9E05E308B52B5EDAAFD25733803DC" pageId="209" pageNumber="210">
<taxonomicName id="52F9BCF3308B52B5EDAAFD2576DC03A3" ID-CoL="6742F" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[189,464,642,668]" class="Insecta" family="Scarabaeidae" genus="Anisocanthon" kingdom="Animalia" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="A78D1B62308B52B5EDAAFD2576DC03A3" box="[189,464,642,668]" italics="true" pageId="209" pageNumber="210">Anisocanthon sericinus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161308B52B5ECE7FD25745A03A3" author="Martinez A. &amp; Pereira F. S." box="[496,854,642,669]" page="366" pageId="209" pageNumber="210" year="1956">
<bibRefCitation id="F168BA81308B52B5ECE7FD25745A03A3" author="Martinez A. &amp; Pereira F. S." box="[496,854,642,669]" pageId="209" pageNumber="210" pagination="363 - 388" refId="ref130174" refString="Martinez A. &amp; Pereira F. S. 1956. Dois generos novos de Canthonini americanos (Col. Scarabaeoidea, Scarabaeidae). Papeis Avulsos do Departemente de Zoologia 12: 363 - 388." type="journal article" year="1956">Martínez &amp; Pereira 1956: 366</bibRefCitation>
</treatmentCitation>
, 376, 386 (key, diagnosis). —
<treatmentCitation id="1458E161308B52B5E9D5FD25760403FF" author="Martinez A." page="49" pageId="209" pageNumber="210" year="1959">
<bibRefCitation id="F168BA81308B52B5E9D5FD25760403FF" author="Martinez A." pageId="209" pageNumber="210" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 49</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161308B52B5ECD9FD0275AB0380" author="Schmidt A." box="[462,679,677,704]" page="63" pageId="209" pageNumber="210" year="1922">
<bibRefCitation id="F168BA81308B52B5ECD9FD0275AB0380" author="Schmidt A." box="[462,679,677,704]" pageId="209" pageNumber="210" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 63</bibRefCitation>
</treatmentCitation>
, 72 (cited). —
<treatmentCitation id="1458E161308B52B5EE74FD02736D03FF" author="Balthasar V." box="[867,1121,677,704]" page="184" pageId="209" pageNumber="210" year="1939">
<bibRefCitation id="F168BA81308B52B5EE74FD02736D03FF" author="Balthasar V." box="[867,1121,677,704]" pageId="209" pageNumber="210" pagination="179 - 238" refId="ref110158" refString="Balthasar V. 1939 a. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939a: 184</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161308B52B5E9F5FD02766C03DC" author="Blackwelder R. E." page="201" pageId="209" pageNumber="210" year="1944">
<bibRefCitation id="F168BA81308B52B5E9F5FD02766C03DC" author="Blackwelder R. E." pageId="209" pageNumber="210" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 201</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="1458E161308B52B5EF00FD6E740903DC" author="Gacharna C." box="[535,773,712,739]" page="221" pageId="209" pageNumber="210" year="1951">
<bibRefCitation id="F168BA81308B52B5EF00FD6E740903DC" author="Gacharna C." box="[535,773,712,739]" pageId="209" pageNumber="210" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 221</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0308B52B5EEB8FD6E732403DC" box="[943,1064,713,739]" name="Colombia" pageId="209" pageNumber="210">Colombia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB308B52B5EDAAFCAD725E026C" box="[189,1362,778,851]" pageId="209" pageNumber="210" type="distribution">
<paragraph id="9546C770308B52B5EDAAFCAD765A021B" blockId="209.[189,342,778,804]" box="[189,342,778,804]" pageId="209" pageNumber="210">
<heading id="CE0E701C308B52B5EDAAFCAD765A021B" bold="true" box="[189,342,778,804]" fontSize="11" level="3" pageId="209" pageNumber="210" reason="3">
<emphasis id="A78D1B62308B52B5EDAAFCAD765A021B" bold="true" box="[189,342,778,804]" pageId="209" pageNumber="210">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFC9E725E026C" blockId="209.[189,1362,825,851]" box="[189,1362,825,851]" pageId="209" pageNumber="210">
Supposedly
<collectingCountry id="EDEE87E0308B52B5EC5AFC9E76C8026C" box="[333,452,825,851]" name="Colombia" pageId="209" pageNumber="210">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E0308B52B5ECD8FC9E7547026C" box="[463,587,825,851]" name="Venezuela" pageId="209" pageNumber="210">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0308B52B5EF41FC9E75C5026C" box="[598,713,825,851]" name="Suriname" pageId="209" pageNumber="210">Suriname</collectingCountry>
,
<collectingCountry id="EDEE87E0308B52B5EFC2FC9E742C026C" box="[725,800,825,851]" name="Brazil" pageId="209" pageNumber="210">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E0308B52B5EE3BFC9E7489026C" box="[812,901,825,851]" name="Bolivia" pageId="209" pageNumber="210">Bolivia</collectingCountry>
, and
<taxonomicName id="52F9BCF3308B52B5EED6FC9E7243026C" authority="(Schoolmeesters 2023)" baseAuthorityName="Schoolmeesters" baseAuthorityYear="2023" box="[961,1359,825,851]" class="Arachnida" family="Diarthrophallidae" genus="Megatopha" kingdom="Animalia" order="Mesostigmata" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="argentina">
Argentina (
<bibRefCitation id="F168BA81308B52B5E952FC9E724A026C" author="Schoolmeesters P." box="[1093,1350,825,851]" pageId="209" pageNumber="210" refId="ref138584" refString="Schoolmeesters P. 2023. World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Vandepitte L., Hobern D., Remsen D., Schalk P., DeWalt R. E., Keping M., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., Adriaenssens E. M., Aedo C., Aescht E., Akkari N. et al. (eds) Catalogue of Life Checklist (Version 2023 - 03 - 06). Available from https: // www. catalogueoflife. org / data / dataset / 1027 [accessed 3 Apr. 2023]." type="url" year="2023">Schoolmeesters 2023</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB308B52B5EDAAFCDD740D02D9" pageId="209" pageNumber="210" type="reference_group">
<paragraph id="9546C770308B52B5EDAAFCDD76A802AB" blockId="209.[189,420,890,916]" box="[189,420,890,916]" pageId="209" pageNumber="210">
<heading id="CE0E701C308B52B5EDAAFCDD76A802AB" bold="true" box="[189,420,890,916]" fontSize="11" level="3" pageId="209" pageNumber="210" reason="3">
<emphasis id="A78D1B62308B52B5EDAAFCDD76A802AB" bold="true" box="[189,420,890,916]" pageId="209" pageNumber="210">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770308B52B5EDAAFC0E740D02D9" blockId="209.[189,1399,936,998]" pageId="209" pageNumber="210">
<bibRefCitation id="F168BA81308B52B5EDAAFC0E769002FC" author="Martinez A." box="[189,412,936,963]" pageId="209" pageNumber="210" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 49</bibRefCitation>
(
<collectingCountry id="EDEE87E0308B52B5ECB8FC0F752302FD" box="[431,559,936,962]" name="Venezuela" pageId="209" pageNumber="210">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81308B52B5EF7AFC0F74DE02FC" author="Vulcano M. A. &amp; Pereira F. S." box="[621,978,936,963]" pageId="209" pageNumber="210" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 588</bibRefCitation>
(
<collectingCountry id="EDEE87E0308B52B5EEF3FC0F736802FD" box="[996,1124,936,962]" name="Venezuela" pageId="209" pageNumber="210">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81308B52B5E9B5FC0F763F02D9" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." pageId="209" pageNumber="210" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62308B52B5E82FFC0E727B02FD" box="[1336,1399,936,962]" italics="true" pageId="209" pageNumber="210">et al.</emphasis>
2013: 108
</bibRefCitation>
(
<collectingCountry id="EDEE87E0308B52B5EC55FC6B76CD02D9" box="[322,449,972,998]" name="Venezuela" pageId="209" pageNumber="210">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992308B52B5ECDBFC6B752502D9" box="[460,553,972,998]" country="Venezuela" name="Bolivar" pageId="209" pageNumber="210">Bolívar</collectingRegion>
:
<collectingCountry id="EDEE87E0308B52B5EF23FC6B755202D9" box="[564,606,972,998]" name="Iceland" pageId="209" pageNumber="210">Isla</collectingCountry>
de Anacoco).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,65 +1,65 @@
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@ -110,10 +108,9 @@ Noriega
2008: 132
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<mods:name id="DE94ACCF9CED045A8ACEDB0F222F11C1" type="personal">
<mods:role id="D621DED08C1C07247E72E89779A7606C">
<mods:roleTerm id="ACFCC6C984EC897E2475315E6DB4DD3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D96D7E15DEF9905934F98217848D2D9E">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="52560B778B3D84CE97FE33794D1F780F" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="F2B78B2535C06870AF07DB69FA8C105E">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D220829E438E05F7E81AD7733B947DD3" type="email">vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="699AEEA52550052C27DB8A93B95806C7">text</mods:typeOfResource>
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<mods:titleInfo id="65106CE2E5F7F3D2561DFE0041A990F3">
<mods:title id="93B989D43D3856DDD9B3EEE11E3FD8EE">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="74592EFE8B3FF1796139B7C5D368F934">
<mods:date id="D5863CD598E976A395F348C5E66EB851">2024</mods:date>
<mods:detail id="620735730C86FED01F1C338422D55AFF" type="pubDate">
<mods:number id="0C184C781AB0152858079066A22A29D1">2024-10-11</mods:number>
</mods:detail>
<mods:detail id="222DB0EED74F345F7B5543E1EFBE7D90" type="volume">
<mods:number id="24F77AB879660736D52FCD21F20C7F2D">959</mods:number>
</mods:detail>
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<mods:number id="AA0B0BD092107F0D164C45A2D1F6403D">1</mods:number>
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<mods:start id="2A0245C25FF9B3A682DA7D33E8ABC6AD">1</mods:start>
<mods:end id="6EC2E9BE7AAF8AAC9EBB16BB15A98D89">272</mods:end>
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<mods:location id="CF6023B77E1D11DEE0CC4E0F30ED8EFE">
<mods:url id="54376F6EC669F48753EEB3294D88FE21">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401</mods:url>
</mods:location>
<mods:classification id="1734DA2C0ACA5CA14A538DD16520F3A5">journal article</mods:classification>
<mods:identifier id="9998569AA6EF80932E9DF9B8557FC14B" type="DOI">10.5852/ejt.2024.959.2677</mods:identifier>
<mods:identifier id="D86708FA069560FD5A83E8FAD66E4773" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="83D9230880AA4E95DDB97FEDF85A3966" type="Zenodo-Dep">13945464</mods:identifier>
<mods:identifier id="0CBEB82B8782233411DEE28298718420" type="ZooBank">30872C13-516F-42FA-AFA7-30ADC6BF1BAF</mods:identifier>
</mods:mods>
<treatment id="1D50766630CD52FCEF53F9F6769504BA" LSID="urn:lsid:plazi:treatment:1D50766630CD52FCEF53F9F6769504BA" httpUri="http://treatment.plazi.org/id/1D50766630CD52FCEF53F9F6769504BA" lastPageId="152" lastPageNumber="153" pageId="151" pageNumber="152">
<subSubSection id="DDE394FB30CD52F3EF53F9F674FC0754" box="[580,1008,1616,1643]" pageId="151" pageNumber="152" type="nomenclature">
<paragraph id="9546C77030CD52F3EF53F9F674FC0754" blockId="151.[580,1008,1616,1643]" box="[580,1008,1616,1643]" pageId="151" pageNumber="152">
<heading id="CE0E701C30CD52F3EF53F9F674FC0754" box="[580,1008,1616,1643]" centered="true" fontSize="11" level="2" pageId="151" pageNumber="152" reason="2">
Genus
<taxonomicName id="52F9BCF330CD52F3EF80F9F774FC0754" ID-CoL="68DQ" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[663,1008,1616,1643]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230CD52F3EF80F9F774300754" bold="true" box="[663,828,1616,1643]" italics="true" pageId="151" pageNumber="152">Onthophagus</emphasis>
<bibRefCitation id="F168BA8130CD52F3EE54F9F674FC0754" author="Latreille P. A." box="[835,1008,1617,1643]" pageId="151" pageNumber="152" refId="ref127095" refString="Latreille P. A. 1802. Histoire naturelle des Fourmis, et Recueil de Memoires et d'Observations sur les Abeilles, les Araignees, les Faucheurs, et autres Insectes. Avec figures. Barrois, Paris. https: // doi. org / 10.5962 / bhl. title. 11138" type="book" year="1802">Latreille, 1802</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CD52FCEDAAF935769504BA" lastPageId="152" lastPageNumber="153" pageId="151" pageNumber="152" type="reference_group">
<paragraph id="9546C77030CD52F3EDAAF935753407EF" blockId="151.[189,1399,1682,1955]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAF935753407EF" pageId="151" pageNumber="152">
<treatmentCitation id="1458E16130CD52F3EDAAF935755B0793" author="Latreille P. A." box="[189,599,1682,1708]" page="141" pageId="151" pageNumber="152" year="1802">
<taxonomicName id="52F9BCF330CD52F3EDAAF935755B0793" ID-CoL="68DQ" authority="Latreille, 1802: 141" authorityName="Latreille" authorityPageNumber="141" authorityYear="1802" box="[189,599,1682,1708]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230CD52F3EDAAF93576570793" box="[189,347,1682,1708]" italics="true" pageId="151" pageNumber="152">Onthophagus</emphasis>
<bibRefCitation id="F168BA8130CD52F3EC70F935755B0793" author="Latreille P. A." box="[359,599,1682,1708]" pageId="151" pageNumber="152" refId="ref127095" refString="Latreille P. A. 1802. Histoire naturelle des Fourmis, et Recueil de Memoires et d'Observations sur les Abeilles, les Araignees, les Faucheurs, et autres Insectes. Avec figures. Barrois, Paris. https: // doi. org / 10.5962 / bhl. title. 11138" type="book" year="1802">Latreille, 1802: 141</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D230CD52F3EE67F93474A60792" box="[880,938,1683,1709]" pageId="151" pageNumber="152">Type</typeStatus>
species:
<taxonomicName id="52F9BCF330CD52F3E909F935762407EF" ID-CoL="7YY9F" authority="Schreber, 1759" authorityName="Schreber" authorityYear="1759" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="taurus">
<emphasis id="A78D1B6230CD52F3E909F93573F10793" box="[1054,1277,1682,1708]" italics="true" pageId="151" pageNumber="152">Scarabaeus taurus</emphasis>
Schreber, 1759
</taxonomicName>
, by original monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAF97F722A07CC" blockId="151.[189,1399,1682,1955]" box="[189,1318,1752,1779]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAF97F722A07CC" box="[189,1318,1752,1779]" pageId="151" pageNumber="152">
<treatmentCitation id="1458E16130CD52F3EDAAF97F755A07CD" author="Erichson W. F." box="[189,598,1752,1778]" page="763" pageId="151" pageNumber="152" year="1847">
<taxonomicName id="52F9BCF330CD52F3EDAAF97F755A07CD" ID-CoL="BX75W" authority="Erichson, 1847b: 763" authorityName="Erichson" authorityPageNumber="763" authorityYear="1847" box="[189,598,1752,1778]" class="Insecta" family="Scarabaeidae" genus="Chalcoderus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230CD52F3EDAAF97F765F07CD" box="[189,339,1752,1778]" italics="true" pageId="151" pageNumber="152">Chalcoderus</emphasis>
<bibRefCitation id="F168BA8130CD52F3EC4DF97F755A07CD" author="Erichson W. F." box="[346,598,1752,1778]" pageId="151" pageNumber="152" pagination="1 - 968" refId="ref117862" refString="Erichson W. F. 1847 b. Naturgeschichte des Insecten Deutschland. I. Coleoptera, Scaphidilia, Scarabaeides. Erste Antheilung, Dritter Band, Erste Lieferung. Nicolaischen Buchhandlung, Berlin. 1 - 968. https: // doi. org / 10.5962 / bhl. title. 50882" type="book chapter" year="1847">Erichson, 1847b: 763</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D230CD52F3EE73F97E749207CC" box="[868,926,1753,1779]" pageId="151" pageNumber="152">Type</typeStatus>
species: no fixation known to us.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAF95A72140662" blockId="151.[189,1399,1682,1955]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAF95A72140662" pageId="151" pageNumber="152">
<treatmentCitation id="1458E16130CD52F3EDAAF95A75250629" author="Erichson W. F." box="[189,553,1788,1814]" page="763" pageId="151" pageNumber="152" year="1847">
<taxonomicName id="52F9BCF330CD52F3EDAAF95A75250629" ID-CoL="BX7K7" authority="Erichson, 1847 b: 763" authorityName="Erichson" authorityPageNumber="763" authorityYear="1847" box="[189,553,1788,1814]" class="Insecta" family="Scarabaeidae" genus="Monapus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230CD52F3EDAAF95A76260629" box="[189,298,1789,1814]" italics="true" pageId="151" pageNumber="152">Monapus</emphasis>
<bibRefCitation id="F168BA8130CD52F3EC27F95B75250629" author="Erichson W. F." box="[304,553,1788,1814]" pageId="151" pageNumber="152" pagination="1 - 968" refId="ref117862" refString="Erichson W. F. 1847 b. Naturgeschichte des Insecten Deutschland. I. Coleoptera, Scaphidilia, Scarabaeides. Erste Antheilung, Dritter Band, Erste Lieferung. Nicolaischen Buchhandlung, Berlin. 1 - 968. https: // doi. org / 10.5962 / bhl. title. 50882" type="book chapter" year="1847">Erichson, 1847b: 763</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D230CD52F3EE24F95B74610629" box="[819,877,1788,1814]" pageId="151" pageNumber="152">Type</typeStatus>
species: no fixation known to us; no nominal species seem to have been originally included.
<treatmentCitation id="1458E16130CD52F3EE0BF8B873220606" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[796,1070,1823,1849]" pageId="151" pageNumber="152" year="2019">
<bibRefCitation id="F168BA8130CD52F3EE0BF8B873220606" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[796,1070,1823,1849]" pageId="151" pageNumber="152" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230CD52F3EE8AF88774D40606" box="[925,984,1823,1849]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
(2019)
</bibRefCitation>
</treatmentCitation>
stated the
<typeStatus id="4A4279D230CD52F3E9A6F8B873E80606" box="[1201,1252,1823,1849]" pageId="151" pageNumber="152">type</typeStatus>
species was “probably”
<taxonomicName id="52F9BCF330CD52F3EC63F8E5743E0663" ID-CoL="7XNKM" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[372,818,1858,1884]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="mniszechi">
<emphasis id="A78D1B6230CD52F3EC63F8E575820663" box="[372,654,1858,1884]" italics="true" pageId="151" pageNumber="152">Onthophagus mniszechi</emphasis>
Harold, 1869
</taxonomicName>
, but did not explain why they thought so.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAF8C276E3069C" blockId="151.[189,1399,1682,1955]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAF8C276E3069C" pageId="151" pageNumber="152">
<treatmentCitation id="1458E16130CD52F3EDAAF8C275060640" author="Erichson W. F." box="[189,522,1893,1919]" page="764" pageId="151" pageNumber="152" year="1847">
<taxonomicName id="52F9BCF330CD52F3EDAAF8C275060640" ID-CoL="BX7PD" authority="Erichson, 1847b: 764" authorityName="Erichson" authorityPageNumber="764" authorityYear="1847" box="[189,522,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Psilax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230CD52F3EDAAF8C2760A0640" box="[189,262,1893,1919]" italics="true" pageId="151" pageNumber="152">Psilax</emphasis>
<bibRefCitation id="F168BA8130CD52F3EC19F8C275060640" author="Erichson W. F." box="[270,522,1893,1919]" pageId="151" pageNumber="152" pagination="1 - 968" refId="ref117862" refString="Erichson W. F. 1847 b. Naturgeschichte des Insecten Deutschland. I. Coleoptera, Scaphidilia, Scarabaeides. Erste Antheilung, Dritter Band, Erste Lieferung. Nicolaischen Buchhandlung, Berlin. 1 - 968. https: // doi. org / 10.5962 / bhl. title. 50882" type="book chapter" year="1847">Erichson, 1847b: 764</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D230CD52F3EE0EF8C1745F06BF" box="[793,851,1894,1920]" pageId="151" pageNumber="152">Type</typeStatus>
species:
<taxonomicName id="52F9BCF330CD52F3EED7F8C2727E0640" ID-CoL="7XNXZ" authority="Erichson, 1842" authorityName="Erichson" authorityYear="1842" box="[960,1394,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pronus">
<emphasis id="A78D1B6230CD52F3EED7F8C273BA0640" box="[960,1206,1893,1919]" italics="true" pageId="151" pageNumber="152">Onthophagus pronus</emphasis>
Erichson, 1842
</taxonomicName>
, by original monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C252FCEDAAFEAD74B60079" blockId="152.[189,1399,265,326]" pageId="152" pageNumber="153">
<treatmentCitationGroup id="B5E9E05E30C252FCEDAAFEAD74B60079" pageId="152" pageNumber="153">
<treatmentCitation id="1458E16130C252FCEDAAFEAD7550001B" author="Arrow G. J." box="[189,604,265,292]" page="435" pageId="152" pageNumber="153" year="1920">
<taxonomicName id="52F9BCF330C252FCEDAAFEAD7550001B" ID-CoL="5JKR" authority="Arrow, 1920: 435" authorityName="Arrow" authorityPageNumber="435" authorityYear="1920" box="[189,604,265,292]" class="Insecta" family="Scarabaeidae" genus="Macropocopris" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFEAD767D001C" box="[189,369,266,291]" italics="true" pageId="152" pageNumber="153">Macropocopris</emphasis>
<bibRefCitation id="F168BA8130C252FCEC69FEAE7550001B" author="Arrow G. J." box="[382,604,265,292]" pageId="152" pageNumber="153" pagination="434 - 437" refId="ref109949" refString="Arrow G. J. 1920. A peculiar new genus of Australian beetles. The Annals and Magazine of Natural History, Series 9 6 (34): 434 - 437. https: // doi. org / 10.1080 / 00222932008632460" type="journal article" year="1920">Arrow, 1920: 435</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D230C252FCEE97FEAD74B6001B" box="[896,954,266,292]" pageId="152" pageNumber="153">Type</typeStatus>
species:
<treatmentCitation id="1458E16130C252FCE923FEAD768D0079" author="Arrow G. J." pageId="152" pageNumber="153" year="1920">
<taxonomicName id="52F9BCF330C252FCE923FEAD768D0079" ID-CoL="7X3SN" authority="Arrow, 1920" authorityName="Arrow" authorityYear="1920" class="Insecta" family="Scarabaeidae" genus="Macropocopris" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="prehensilis">
<emphasis id="A78D1B6230C252FCE923FEAD727B001C" box="[1076,1399,265,291]" italics="true" pageId="152" pageNumber="153">Macropocopris prehensilis</emphasis>
<bibRefCitation id="F168BA8130C252FCEDFDFE8B768D0079" author="Arrow G. J." box="[234,385,300,326]" pageId="152" pageNumber="153" pagination="434 - 437" refId="ref109949" refString="Arrow G. J. 1920. A peculiar new genus of Australian beetles. The Annals and Magazine of Natural History, Series 9 6 (34): 434 - 437. https: // doi. org / 10.1080 / 00222932008632460" type="journal article" year="1920">Arrow, 1920</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, by subsequent designation of
<treatmentCitation id="1458E16130C252FCEFFDFE8B74BB0079" author="Matthews E. G." box="[746,951,300,326]" pageId="152" pageNumber="153" year="1972">
<bibRefCitation id="F168BA8130C252FCEFFDFE8B74BB0079" author="Matthews E. G." box="[746,951,300,326]" pageId="152" pageNumber="153" pagination="1 - 330" refId="ref130633" refString="Matthews E. G. 1972. A revision of the scarabaeine dung beetles of Australia. I. Tribe Onthophagini. Australian Journal of Zoology, Supplementary Series 9: 1 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews (1972)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C252FCEDAAFEC973EA02A7" blockId="152.[189,1400,365,1413]" pageId="152" pageNumber="153">
<treatmentCitationGroup id="B5E9E05E30C252FCEDAAFEC973EA02A7" pageId="152" pageNumber="153">
<taxonomicName id="52F9BCF330C252FCEDAAFEC9765700B7" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[189,347,366,392]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFEC9765700B7" box="[189,347,366,392]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C252FCEC6DFEC9733E00B7" author="Le Peletier de Saint-Fargeau &amp; Audinet-Serville" box="[378,1074,365,392]" page="353" pageId="152" pageNumber="153" year="1828">
<bibRefCitation id="F168BA8130C252FCEC6DFEC9733E00B7" author="Le Peletier de Saint-Fargeau &amp; Audinet-Serville" box="[378,1074,365,392]" editor="Latreille &amp; Audinet-] Serville &amp; Guerin-Meneville" firstAuthor="Le" journalOrPublisher="Publication date following Bousquet" pageId="152" pageNumber="153" part="2016" refId="ref127845" refString="Le Peletier de Saint-Fargeau [A. L. M.] &amp; Audinet-Serville [J. G.] 1828. Scarabe. In: Latreille [P. A.], Le Peletier [A. L. M.], [Audinet-] Serville [J. G.] &amp; Guerin-Meneville [F. E.] 1825 - 1828. Encyclopedie methodique, ou par ordre de matieres; par une societe de gens de lettres, de savans et d'artistes; precedee d'un vocabulaire universel, servant de table pour tout l'ouvrage, ornee des portraits de Mm. Diderot &amp; d'Alembert, premiers editeurs de l'Encyclopedie. Histoire Naturelle. Entomologie, ou histoire naturelle des crustaces, des arachnides et des insectes. Tome dixieme: 346 - 382. Mme Veuve Agasse, Paris. [Publication date following Bousquet (2016).]" title="Encyclopedie methodique, ou par ordre de matieres; par une societe de gens de lettres, de savans et d'artistes; precedee d'un vocabulaire universel, servant de table pour tout l'ouvrage, ornee des portraits de Mm. Diderot &amp; d'Alembert, premiers editeurs de l'Encyclopedie" type="journal volume" volumeTitle="Histoire Naturelle. Entomologie, ou histoire naturelle des crustaces, des arachnides et des insectes">Le Peletier de Saint-Fargeau &amp; Audinet-Serville 1828: 353</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCE80DFEC976530094" author="Agassiz L." page="749" pageId="152" pageNumber="153" year="1846">
<bibRefCitation id="F168BA8130C252FCE80DFEC976530094" author="Agassiz L." pageId="152" pageNumber="153" refId="ref108975" refString="Agassiz L. 1846. Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilum, secundum ordinem alphabeticum unicum deposita, adjectis homonymus platarum, nec non variis adnotationibus et emendationibus. Jent and Gassman, Solothurn [Soloduri]. https: // doi. org / 10.5962 / bhl. title. 1819" type="book" year="1846">Agassiz 1846: 749</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C252FCEF0FFE3675F90093" author="Latreille P. A." box="[536,757,401,428]" page="536" pageId="152" pageNumber="153" year="1829">
<bibRefCitation id="F168BA8130C252FCEF0FFE3675F90093" author="Par M. le" box="[536,757,401,428]" firstAuthor="Latreille" pageId="152" pageNumber="153" refId="ref127536" refString="Latreille [P. A.] 1829. Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee. Par M. le Baron Cuvier. Avec figures, dessinees d'apres nature. Nouvelle edition, revue et augmentee. Tome IV. Crustaces, arachnides et partie des insectes. Deterville, Paris. https: // doi. org / 10.5962 / bhl. title. 49223" type="book" year="1829">Latreille 1829: 536</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCEEC0FE3673960094" author="Brulle A." box="[983,1178,401,427]" page="300" pageId="152" pageNumber="153" year="1838">
<bibRefCitation id="F168BA8130C252FCEEC0FE3673960094" author="Brulle A." box="[983,1178,401,427]" pageId="152" pageNumber="153" pagination="225 - 448" refId="ref112853" refString="Brulle A. 1838. [livraison 8: pp. 225 - 448]. In: Brulle A. 1837 - 1838. Histoire naturelle des insectes, traitant de leur organisation et de leurs moeurs en general, par M. V. Audouin, et comprenant leur classification et la description des especes, par M. A. Brulle. Le tout accompagne de planches gravees sur acier, d'apres des peintures executees pour cette edition sur la collection du Museum de Paris. Tome VI. Coleopteres III. F. D. Pillot, Paris. [Publication date and authorship based on Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 34228" type="journal article" year="1838">Brullé 1838: 300</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCEDFEFE1376C400F1" author="Castelnau" box="[233,456,436,462]" page="83" pageId="152" pageNumber="153" year="1840">
<bibRefCitation id="F168BA8130C252FCEDFEFE1376C400F1" author="Castelnau" box="[233,456,436,462]" pageId="152" pageNumber="153" refId="ref113936" refString="Castelnau, Comte de 1840. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des insectes coleopteres. Tome deuxieme. Dumenil, Paris. https: // doi. org / 10.5962 / bhl. title. 47104" type="book" year="1840">Castelnau 1840: 83</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCEFB9FE1374A500F1" author="Lacordaire J. T." box="[686,937,436,463]" page="107" pageId="152" pageNumber="153" year="1855">
<bibRefCitation id="F168BA8130C252FCEFB9FE1374A500F1" author="Lacordaire J. T." box="[686,937,436,463]" pageId="152" pageNumber="153" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 107</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCE99BFE13727B00F1" author="Harold E." box="[1164,1399,436,463]" page="1024" pageId="152" pageNumber="153" year="1869">
<bibRefCitation id="F168BA8130C252FCE99BFE13727B00F1" author="Harold E." box="[1164,1399,436,463]" pageId="152" pageNumber="153" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1024</bibRefCitation>
</treatmentCitation>
(catalogue).
<emphasis id="A78D1B6230C252FCEC97FE7F769600CE" box="[384,410,472,497]" italics="true" pageId="152" pageNumber="153"></emphasis>
<treatmentCitation id="1458E16130C252FCECB3FE70757B00CE" author="Gillet J. J. E." box="[420,631,471,497]" page="118" pageId="152" pageNumber="153" year="1911">
<bibRefCitation id="F168BA8130C252FCECB3FE70757B00CE" author="Gillet J. J. E." box="[420,631,471,497]" pageId="152" pageNumber="153" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 118</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C252FCEE28FE7073B500CD" author="Gillet J. J. E. &amp; Boucomont A." box="[831,1209,471,498]" page="205" pageId="152" pageNumber="153" year="1927">
<bibRefCitation id="F168BA8130C252FCEE28FE7073B500CD" author="Gillet J. J. E. &amp; Boucomont A." box="[831,1209,471,498]" pageId="152" pageNumber="153" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 205</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C252FCEDFDFE5C76CC032B" author="Dawson R. W." box="[234,448,506,533]" page="61" pageId="152" pageNumber="153" year="1922">
<bibRefCitation id="F168BA8130C252FCEDFDFE5C76CC032B" author="Dawson R. W." box="[234,448,506,533]" pageId="152" pageNumber="153" pagination="163 - 244" refId="ref116280" refString="Dawson R. W. 1922. A synopsis of the Scarabaeidae of Nebraska. University Studies of the University of Nebraska 22: 163 - 244." type="journal article" year="1922">Dawson 1922: 61</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEF53FE5D742D032A" author="Paulian R." box="[580,801,506,533]" page="232" pageId="152" pageNumber="153" year="1938">
<bibRefCitation id="F168BA8130C252FCEF53FE5D742D032A" author="Paulian R." box="[580,801,506,533]" pageId="152" pageNumber="153" pagination="213 - 296" refId="ref134801" refString="Paulian R. 1938. Contribution a l'etude des Canthonides americans [Coleopt. Lamellic.]. Annales de la Societe entomologique de France 107: 213 - 296." type="journal article" year="1938">Paulian 1938: 232</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEEB2FE5D73B1032A" author="Blackwelder R. E." box="[933,1213,506,533]" page="211" pageId="152" pageNumber="153" year="1944">
<bibRefCitation id="F168BA8130C252FCEEB2FE5D73B1032A" author="Blackwelder R. E." box="[933,1213,506,533]" pageId="152" pageNumber="153" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C252FCE82DFE5C765D0307" author="Roze J. A." page="45" pageId="152" pageNumber="153" year="1955">
<bibRefCitation id="F168BA8130C252FCE82DFE5C765D0307" author="Roze J. A." pageId="152" pageNumber="153" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 45</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C252FCECEDFDB975740307" box="[506,632,542,568]" name="Venezuela" pageId="152" pageNumber="153">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCEFB9FDB974990307" author="Martinez A." box="[686,917,541,568]" page="108" pageId="152" pageNumber="153" year="1959">
<bibRefCitation id="F168BA8130C252FCEFB9FDB974990307" author="Martinez A." box="[686,917,541,568]" pageId="152" pageNumber="153" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 108</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E030C252FCE953FDB973B20307" box="[1092,1214,542,568]" name="Argentina" pageId="152" pageNumber="153">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCE9E2FDB976C30364" author="Howden H. F. &amp; Cartwright O. L." page="6" pageId="152" pageNumber="153" year="1963">
<bibRefCitation id="F168BA8130C252FCE9E2FDB976C30364" author="Howden H. F. &amp; Cartwright O. L." pageId="152" pageNumber="153" pagination="1 - 133" refId="ref124770" refString="Howden H. F. &amp; Cartwright O. L. 1963. Scarab beetles of the genus Onthophagus Latreille North of Mexico. Proceedings of the United States National Museum 114 (3467): 1 - 133. https: // doi. org / 10.5479 / si. 00963801.114 - 3467.1" type="journal article" year="1963">Howden &amp; Cartwright 1963: 6</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCEFA9FDE673300363" author="Halffter G. &amp; Matthews E. G." box="[702,1084,577,604]" page="254" pageId="152" pageNumber="153" year="1966">
<bibRefCitation id="F168BA8130C252FCEFA9FDE673300363" author="Halffter G. &amp; Matthews E. G." box="[702,1084,577,604]" pageId="152" pageNumber="153" pagination="1 - 312" refId="ref122681" refString="Halffter G. &amp; Matthews E. G. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966: 254</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E16130C252FCEDFDFDC3755C0340" author="Vulcano M. A. &amp; Pereira F. S." box="[234,592,612,639]" page="562" pageId="152" pageNumber="153" year="1967">
<bibRefCitation id="F168BA8130C252FCEDFDFDC3755C0340" author="Vulcano M. A. &amp; Pereira F. S." box="[234,592,612,639]" pageId="152" pageNumber="153" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 562</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEFC3FDC3732B0341" author="Howden H. F. &amp; Young O. P." box="[724,1063,612,639]" page="11" pageId="152" pageNumber="153" year="1981">
<bibRefCitation id="F168BA8130C252FCEFC3FDC3732B0341" author="Howden H. F. &amp; Young O. P." box="[724,1063,612,639]" pageId="152" pageNumber="153" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 11</bibRefCitation>
</treatmentCitation>
, 93 (key, redescription). —
<treatmentCitation id="1458E16130C252FCEDFDFD20756E039D" author="Halffter G. &amp; Edmonds W. D." box="[234,610,647,674]" page="135" pageId="152" pageNumber="153" year="1982">
<bibRefCitation id="F168BA8130C252FCEDFDFD20756E039D" author="Halffter G. &amp; Edmonds W. D." box="[234,610,647,674]" pageId="152" pageNumber="153" refId="ref122441" refString="Halffter G. &amp; Edmonds W. D. 1982. The Nesting Behavior of Dung Beetles (Scarabaeinae). An Ecological and Evolutive Approach. Instituto de Ecologia, Mexico, D. F." type="book" year="1982">Halffter &amp; Edmonds 1982: 135</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E16130C252FCEEB2FD207370039E" author="Huchet J. B." box="[933,1148,647,673]" page="300" pageId="152" pageNumber="153" year="1992">
<bibRefCitation id="F168BA8130C252FCEEB2FD207370039E" author="Huchet J. B." box="[933,1148,647,673]" pageId="152" pageNumber="153" pagination="297 - 303" refId="ref124933" refString="Huchet J. B. 1992. Un scarabee nouveau pour les Petites Antilles: Digitonthophagus gazella (Fabricius) [Col. Scarabaeidae, Coprinae, Onthophagini]. L'Entomologiste 48 (6): 297 - 303." type="journal article" year="1992">Huchet 1992: 300</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCE9EBFD2076F303FA" author="Medina C. A. &amp; Lopera-Toro A." page="301" pageId="152" pageNumber="153" year="2000">
<bibRefCitation id="F168BA8130C252FCE9EBFD2076F303FA" author="Medina C. A. &amp; Lopera-Toro A." pageId="152" pageNumber="153" pagination="299 - 315" refId="ref130725" refString="Medina C. A. &amp; Lopera-Toro A. 2000. Clave ilustrada para la identificacion de generos de escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Caldasia 22: 299 - 315." type="journal article" year="2000">Medina &amp; Lopera-Toro 2000: 301</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEF6AFD0D749303FA" author="Vaz-de-Mello F. Z." box="[637,927,682,709]" page="194" pageId="152" pageNumber="153" year="2000">
<bibRefCitation id="F168BA8130C252FCEF6AFD0D749303FA" author="Vaz-de-Mello F. Z." box="[637,927,682,709]" pageId="152" pageNumber="153" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C252FCE958FD0D739003FB" box="[1103,1180,682,708]" name="Brazil" pageId="152" pageNumber="153">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCE9C0FD0D766D03D7" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." page="139" pageId="152" pageNumber="153" year="2001">
<bibRefCitation id="F168BA8130C252FCE9C0FD0D766D03D7" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." pageId="152" pageNumber="153" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230C252FCE82BFD0C727403FB" box="[1340,1400,682,708]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2001: 139
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030C252FCECDDFD69754F03D7" box="[458,579,718,744]" name="Colombia" pageId="152" pageNumber="153">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCEF6AFD6974D703D8" author="Kohlmann B. &amp; Solis A." box="[637,987,717,744]" page="160" pageId="152" pageNumber="153" year="2001">
<bibRefCitation id="F168BA8130C252FCEF6AFD6974D703D8" author="Kohlmann B. &amp; Solis A." box="[637,987,717,744]" pageId="152" pageNumber="153" pagination="159 - 261" refId="ref126127" refString="Kohlmann B. &amp; Solis A. 2001. El genero Onthophagus en Costa Rica. Giornale Italiano di Entomologia 49 (9): 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís 2001: 160</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C252FCE9D2FD6976040234" author="Ratcliffe B. C." page="17" pageId="152" pageNumber="153" year="2002">
<bibRefCitation id="F168BA8130C252FCE9D2FD6976040234" author="Ratcliffe B. C." firstAuthor="Ratcliffe" pageId="152" pageNumber="153" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 17</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C252FCECA2FD56751B0234" box="[437,535,753,779]" name="Panama" pageId="152" pageNumber="153">Panama</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCEF47FD5674650234" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Smith A. B. T." box="[592,873,753,779]" page="49" pageId="152" pageNumber="153" year="2002">
<bibRefCitation id="F168BA8130C252FCEF47FD5674650234" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Smith A. B. T." box="[592,873,753,779]" pageId="152" pageNumber="153" pagination="39 - 81" refId="ref136930" refString="Ratcliffe B. C., Jameson M. L. &amp; Smith A. B. T. 2002. Scarabaeidae Latreille 1802. In: Arnett Jr R. H., Thomas M. C., Skelley P. E. &amp; Frank J. H. (eds) American Beetles. Polyphaga: Scarabaeoidea through Curculionoidea. Volume 2: 39 - 81. CRC Press, Boca Raton, London, New York, and Washington, D. C." type="book chapter" year="2002">
Ratcliffe
<emphasis id="A78D1B6230C252FCEFA9FD5575F60234" box="[702,762,753,779]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2002: 49
</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEEF3FD56724D0234" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[996,1345,753,779]" page="16" pageId="152" pageNumber="153" year="2006">
<bibRefCitation id="F168BA8130C252FCEEF3FD56724D0234" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[996,1345,753,779]" pageId="152" pageNumber="153" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B6230C252FCE982FD5573DD0234" box="[1173,1233,753,779]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2006: 16
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030C252FCEC01FCB3767E0211" box="[278,370,788,814]" name="Bolivia" pageId="152" pageNumber="153">Bolivia</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCECBAFCB374410211" author="Pulido-Herrera L. A. &amp; Zunino M." box="[429,845,788,814]" page="94" pageId="152" pageNumber="153" year="2007">
<bibRefCitation id="F168BA8130C252FCECBAFCB374410211" author="Pulido-Herrera L. A. &amp; Zunino M." box="[429,845,788,814]" pageId="152" pageNumber="153" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 94</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C252FCE906FCB3727A0211" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[1041,1398,788,814]" page="23" pageId="152" pageNumber="153" year="2011">
<bibRefCitation id="F168BA8130C252FCE906FCB3727A0211" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[1041,1398,788,814]" pageId="152" pageNumber="153" pagination="1 - 73" refId="ref140802" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C., Schoolmeesters P. 2011 a. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B6230C252FCE9ACFCB273F40211" box="[1211,1272,788,814]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2011a: 23
</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C252FCEC74FC90759C026D" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[355,656,823,850]" page="145" pageId="152" pageNumber="153" year="2011">
<bibRefCitation id="F168BA8130C252FCEC74FC90759C026D" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[355,656,823,850]" pageId="152" pageNumber="153" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B6230C252FCECD9FC9F7500026E" box="[462,524,823,849]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2011: 145
</bibRefCitation>
</treatmentCitation>
, 322 (diagnosis, list). —
<treatmentCitation id="1458E16130C252FCEED1FC9073AC026E" author="Krajcik M." box="[966,1184,823,849]" page="174" pageId="152" pageNumber="153" year="2012">
<bibRefCitation id="F168BA8130C252FCEED1FC9073AC026E" author="Krajcik M." box="[966,1184,823,849]" pageId="152" pageNumber="153" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 174</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C252FCE80CFC9076C9024A" author="Solis A. &amp; Kohlmann B." page="8" pageId="152" pageNumber="153" year="2012">
<bibRefCitation id="F168BA8130C252FCE80CFC9076C9024A" author="Solis A. &amp; Kohlmann B." pageId="152" pageNumber="153" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 8</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C252FCEF7CFCFC75FC024B" box="[619,752,858,885]" name="Costa Rica" pageId="152" pageNumber="153">Costa Rica</collectingCountry>
). —
<treatmentCitation id="1458E16130C252FCEE32FCFD7340024A" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[805,1100,858,885]" page="74" pageId="152" pageNumber="153" year="2018">
<bibRefCitation id="F168BA8130C252FCEE32FCFD7340024A" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[805,1100,858,885]" pageId="152" pageNumber="153" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B6230C252FCEEB5FCFC74D7024B" box="[930,987,858,884]" italics="true" pageId="152" pageNumber="153">et al.</emphasis>
2018: 74
</bibRefCitation>
</treatmentCitation>
, 9697 (list for
<collectingCountry id="EDEE87E030C252FCE814FCFD7266024B" box="[1283,1386,858,884]" name="Ecuador" pageId="152" pageNumber="153">Ecuador</collectingCountry>
);
<treatmentCitation id="1458E16130C252FCEDFEFCD9765302A7" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[233,351,894,920]" page="179" pageId="152" pageNumber="153" year="2019">
<bibRefCitation id="F168BA8130C252FCEDFEFCD9765302A7" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[233,351,894,920]" pageId="152" pageNumber="153" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 179</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C252FCEF09FCD9749A02A7" ID-DOI="http://doi.org/10.5281/zenodo.3683552" ID-Zenodo-Dep="3683552" author="Hielkema A. J. &amp; Hielkema M. A." box="[542,918,894,920]" httpUri="http://treatment.plazi.org/id/03D7FF1CFF90FFB6FF48F9FF3353FCC6" page="90" pageId="152" pageNumber="153" year="2019">
<bibRefCitation id="F168BA8130C252FCEF09FCD9749A02A7" author="Hielkema A. J. &amp; Hielkema M. A." box="[542,918,894,920]" pageId="152" pageNumber="153" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 90</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C252FCEDAAFC067695051A" blockId="152.[189,1400,365,1413]" pageId="152" pageNumber="153">
<treatmentCitationGroup id="B5E9E05E30C252FCEDAAFC067695051A" pageId="152" pageNumber="153">
<taxonomicName id="52F9BCF330C252FCEDAAFC06765F0284" ID-CoL="BX75W" box="[189,339,929,955]" class="Insecta" family="Scarabaeidae" genus="Chalcoderus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFC06765F0284" box="[189,339,929,955]" italics="true" pageId="152" pageNumber="153">Chalcoderus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C252FCEC69FC0675840284" author="Lacordaire J. T." box="[382,648,929,956]" page="109" pageId="152" pageNumber="153" year="1855">
<bibRefCitation id="F168BA8130C252FCEC69FC0675840284" author="Lacordaire J. T." box="[382,648,929,956]" pageId="152" pageNumber="153" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 109</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130C252FCEE73FC0673520284" author="Harold E." box="[868,1118,929,955]" page="1024" pageId="152" pageNumber="153" year="1869">
<bibRefCitation id="F168BA8130C252FCEE73FC0673520284" author="Harold E." box="[868,1118,929,955]" pageId="152" pageNumber="153" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1024</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEDFDFC63768402E1" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[234,392,964,990]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDFDFC63768402E1" box="[234,392,964,990]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B6230C252FCECB3FC6276B202E1" box="[420,446,965,990]" italics="true" pageId="152" pageNumber="153"></emphasis>
<treatmentCitation id="1458E16130C252FCECDEFC63759102E1" author="Gillet J. J. E." box="[457,669,964,990]" page="118" pageId="152" pageNumber="153" year="1911">
<bibRefCitation id="F168BA8130C252FCECDEFC63759102E1" author="Gillet J. J. E." box="[457,669,964,990]" pageId="152" pageNumber="153" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 118</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEEA5FC63735C02E1" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[946,1104,964,990]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEEA5FC63735C02E1" box="[946,1104,964,990]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCE982FC637619053E" author="Blackwelder R. E." page="211" pageId="152" pageNumber="153" year="1944">
<bibRefCitation id="F168BA8130C252FCE982FC637619053E" author="Blackwelder R. E." pageId="152" pageNumber="153" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEF25FC4075DC053E" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[562,720,999,1025]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEF25FC4075DC053E" box="[562,720,999,1025]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCEE00FC40736B053E" author="Solis A. &amp; Kohlmann B." box="[791,1127,999,1025]" page="8" pageId="152" pageNumber="153" year="2012">
<bibRefCitation id="F168BA8130C252FCEE00FC40736B053E" author="Solis A. &amp; Kohlmann B." box="[791,1127,999,1025]" pageId="152" pageNumber="153" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 8</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEDFDFBAD7684051B" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[234,392,1034,1060]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDFDFBAD7684051B" box="[234,392,1034,1060]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C252FCEDAAFB897695058E" blockId="152.[189,1400,365,1413]" pageId="152" pageNumber="153">
<treatmentCitationGroup id="B5E9E05E30C252FCEDAAFB897695058E" pageId="152" pageNumber="153">
<taxonomicName id="52F9BCF330C252FCEDAAFB8976260578" ID-CoL="BX7K7" authorityName="Erichson" authorityYear="1847" box="[189,298,1070,1095]" class="Insecta" family="Scarabaeidae" genus="Monapus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFB8976260578" box="[189,298,1070,1095]" italics="true" pageId="152" pageNumber="153">Monapus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C252FCEC4BFB8975600577" author="Lacordaire J. T." box="[348,620,1069,1096]" page="109" pageId="152" pageNumber="153" year="1855">
<bibRefCitation id="F168BA8130C252FCEC4BFB8975600577" author="Lacordaire J. T." box="[348,620,1069,1096]" pageId="152" pageNumber="153" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 109</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130C252FCEE46FB89735E0577" author="Harold E." box="[849,1106,1069,1096]" page="1024" pageId="152" pageNumber="153" year="1869">
<bibRefCitation id="F168BA8130C252FCEE46FB89735E0577" author="Harold E." box="[849,1106,1069,1096]" pageId="152" pageNumber="153" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1024</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEDFDFBF676840554" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[234,392,1105,1131]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDFDFBF676840554" box="[234,392,1105,1131]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B6230C252FCECB3FBF576B20554" box="[420,446,1106,1131]" italics="true" pageId="152" pageNumber="153"></emphasis>
<treatmentCitation id="1458E16130C252FCECDEFBF675910554" author="Gillet J. J. E." box="[457,669,1105,1131]" page="118" pageId="152" pageNumber="153" year="1911">
<bibRefCitation id="F168BA8130C252FCECDEFBF675910554" author="Gillet J. J. E." box="[457,669,1105,1131]" pageId="152" pageNumber="153" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 118</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEEA5FBF6735C0554" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[946,1104,1105,1131]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEEA5FBF6735C0554" box="[946,1104,1105,1131]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCE982FBF6761905B1" author="Blackwelder R. E." page="211" pageId="152" pageNumber="153" year="1944">
<bibRefCitation id="F168BA8130C252FCE982FBF6761905B1" author="Blackwelder R. E." pageId="152" pageNumber="153" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEF25FBD375DC05B1" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[562,720,1140,1166]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEF25FBD375DC05B1" box="[562,720,1140,1166]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCEE00FBD3736B05B1" author="Solis A. &amp; Kohlmann B." box="[791,1127,1140,1166]" page="8" pageId="152" pageNumber="153" year="2012">
<bibRefCitation id="F168BA8130C252FCEE00FBD3736B05B1" author="Solis A. &amp; Kohlmann B." box="[791,1127,1140,1166]" pageId="152" pageNumber="153" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 8</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEDFDFB307684058E" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[234,392,1175,1201]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDFDFB307684058E" box="[234,392,1175,1201]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C252FCEDAAFB1D73D80424" blockId="152.[189,1400,365,1413]" pageId="152" pageNumber="153">
<treatmentCitationGroup id="B5E9E05E30C252FCEDAAFB1D73D80424" pageId="152" pageNumber="153">
<taxonomicName id="52F9BCF330C252FCEDAAFB1D760A05EB" ID-CoL="BX7PD" authorityName="Erichson" authorityYear="1847" box="[189,262,1210,1236]" class="Insecta" family="Scarabaeidae" genus="Psilax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFB1D760A05EB" box="[189,262,1210,1236]" italics="true" pageId="152" pageNumber="153">Psilax</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C252FCEC0AFB1D751F05EA" author="Lacordaire J. T." box="[285,531,1210,1237]" page="109" pageId="152" pageNumber="153" year="1855">
<bibRefCitation id="F168BA8130C252FCEC0AFB1D751F05EA" author="Lacordaire J. T." box="[285,531,1210,1237]" pageId="152" pageNumber="153" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 109</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130C252FCEFC7FB1D74BA05EA" author="Harold E." box="[720,950,1210,1237]" page="1024" pageId="152" pageNumber="153" year="1869">
<bibRefCitation id="F168BA8130C252FCEFC7FB1D74BA05EA" author="Harold E." box="[720,950,1210,1237]" pageId="152" pageNumber="153" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1024</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCE9B1FB1D724805EB" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[1190,1348,1210,1236]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCE9B1FB1D724805EB" box="[1190,1348,1210,1236]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B6230C252FCE84BFB1C727A05EB" box="[1372,1398,1211,1236]" italics="true" pageId="152" pageNumber="153"></emphasis>
<treatmentCitation id="1458E16130C252FCEDFDFB7976B905C7" author="Gillet J. J. E." box="[234,437,1246,1272]" page="118" pageId="152" pageNumber="153" year="1911">
<bibRefCitation id="F168BA8130C252FCEDFDFB7976B905C7" author="Gillet J. J. E." box="[234,437,1246,1272]" pageId="152" pageNumber="153" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 118</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEFA4FB79745D05C7" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[691,849,1246,1272]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEFA4FB79745D05C7" box="[691,849,1246,1272]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCEE99FB79739105C7" author="Blackwelder R. E." box="[910,1181,1246,1272]" page="211" pageId="152" pageNumber="153" year="1944">
<bibRefCitation id="F168BA8130C252FCEE99FB79739105C7" author="Blackwelder R. E." box="[910,1181,1246,1272]" pageId="152" pageNumber="153" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEC1DFAA676A40424" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[266,424,1281,1307]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEC1DFAA676A40424" box="[266,424,1281,1307]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
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<bibRefCitation id="F168BA8130C252FCECF2FAA6742E0424" author="Solis A. &amp; Kohlmann B." box="[485,802,1281,1307]" pageId="152" pageNumber="153" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 8</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCE932FAA673CF0424" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[1061,1219,1281,1307]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCE932FAA673CF0424" box="[1061,1219,1281,1307]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
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).
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<paragraph id="9546C77030C252FCEDAAFA82769504BA" blockId="152.[189,1400,365,1413]" pageId="152" pageNumber="153">
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<taxonomicName id="52F9BCF330C252FCEDAAFA82767D0401" ID-CoL="5JKR" authorityName="Arrow" authorityYear="1920" box="[189,369,1317,1342]" class="Insecta" family="Scarabaeidae" genus="Macropocopris" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDAAFA82767D0401" box="[189,369,1317,1342]" italics="true" pageId="152" pageNumber="153">Macropocopris</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C252FCEC8FFA83742C0400" author="Halffter G. &amp; Matthews E. G." box="[408,800,1316,1343]" page="254" pageId="152" pageNumber="153" year="1966">
<bibRefCitation id="F168BA8130C252FCEC8FFA83742C0400" author="Halffter G. &amp; Matthews E. G." box="[408,800,1316,1343]" pageId="152" pageNumber="153" pagination="1 - 312" refId="ref122681" refString="Halffter G. &amp; Matthews E. G. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966: 254</bibRefCitation>
</treatmentCitation>
(cited as genus). —
<treatmentCitation id="1458E16130C252FCE93BFA83761A045D" author="Halffter G. &amp; Edmonds W. D." page="135" pageId="152" pageNumber="153" year="1982">
<bibRefCitation id="F168BA8130C252FCE93BFA83761A045D" author="Halffter G. &amp; Edmonds W. D." pageId="152" pageNumber="153" refId="ref122441" refString="Halffter G. &amp; Edmonds W. D. 1982. The Nesting Behavior of Dung Beetles (Scarabaeinae). An Ecological and Evolutive Approach. Instituto de Ecologia, Mexico, D. F." type="book" year="1982">Halffter &amp; Edmonds 1982: 135</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEF25FAE075DC045E" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[562,720,1351,1377]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEF25FAE075DC045E" box="[562,720,1351,1377]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C252FCEE00FAE0736B045E" author="Solis A. &amp; Kohlmann B." box="[791,1127,1351,1377]" page="8" pageId="152" pageNumber="153" year="2012">
<bibRefCitation id="F168BA8130C252FCEE00FAE0736B045E" author="Solis A. &amp; Kohlmann B." box="[791,1127,1351,1377]" pageId="152" pageNumber="153" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 8</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C252FCEDFDFACD768404BB" ID-CoL="68DQ" authorityName="Latreille" authorityYear="1802" box="[234,392,1386,1412]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B6230C252FCEDFDFACD768404BB" box="[234,392,1386,1412]" italics="true" pageId="152" pageNumber="153">Onthophagus</emphasis>
</taxonomicName>
).
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