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<document id="BBC65D9D59F22D97BCE598AF2256976F" ID-DOI="10.3897/dez.72.144365" ID-ZooBank="E82AFF4A-F2BA-48AD-A246-D966CC7EF57D" ID-publisher-id="144365" URI-arpha="4549876C-BF70-5C59-800E-8482397039BE" URI-zoobank="http://zoobank.org/E82AFF4A-F2BA-48AD-A246-D966CC7EF57D" article-type="research-article" checkinTime="1744730459598" checkinUser="pensoft" docAuthor="Yin, Zi-Wei &amp; Niu, Xiao-Ling" docDate="2025" docId="4344351F000C5E0B8BA1AB45D8D697D3" docLanguage="en" docName="Deutsche Entomologische Zeitschrift 72(1): 9-14" docOrigin="Deutsche Entomologische Zeitschrift 72 (1)" docSource="https://dez.pensoft.net/article/144365/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Triartiger caiyujiei Yin &amp; Niu 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="14" masterDocId="4549876CBF705C59800E8482397039BE" masterDocTitle="Triartiger caiyujiei sp. nov., a remarkable clavigerite beetle (Coleoptera, Staphylinidae, Pselaphinae) associated with Pseudolasius ants (Hymenoptera, Formicidae) from East China" masterLastPageNumber="14" masterPageNumber="9" pageNumber="9" updateTime="1744730829051" updateUser="ExternalLinkService">
<mods:mods id="92132EA1AAF7162F0D44BB059824A597" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C9C7964134FE15BF3EF97F2DD2D5CD8B">
<mods:title id="BE86F6BF46F74D2428981EC81A518886">Triartiger caiyujiei sp. nov., a remarkable clavigerite beetle (Coleoptera, Staphylinidae, Pselaphinae) associated with Pseudolasius ants (Hymenoptera, Formicidae) from East China</mods:title>
</mods:titleInfo>
<mods:name id="908135B0BC445A8F29A9D443B494416F" type="personal">
<mods:role id="F01DE95AC562392C252C36547E6694D4">
<mods:roleTerm id="A80A8FD5B1C6CDBC01D06BBA3FDCE7C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="54BEE06CB588EC2F3D457E8A9F50292B">Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier id="AA06B4DE4DF90B0F22D853B01E971F08" type="ORCID">0000-0001-6659-9448</mods:nameIdentifier>
<mods:affiliation id="6073683F639DD3E142D9EF9C7D81FE32">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China</mods:affiliation>
</mods:name>
<mods:name id="85C3B19BEB23C1E33D831D4A73ECF2C4" type="personal">
<mods:role id="74A7F0A292540AC3D161411CF4F1AC89">
<mods:roleTerm id="29759AF6F9141F35626A1D2913FDF99E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02A9F8E711EE2B88DC48911033D5F773">Niu, Xiao-Ling</mods:namePart>
<mods:nameIdentifier id="3D4BA8B388CB229DE792EB5D5ED668C0" type="ORCID">https://orcid.org/0009-0002-7928-1492</mods:nameIdentifier>
<mods:affiliation id="A6916F9E79BF07E1E6E0370D87004D1D">Zhengjiang Tianmshan National Nature Reserve Management Bureau, Linan, Hangzhou 311300, Zhejiang, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0D5DB86D551A5F2A160B0BDEF388ED52">text</mods:typeOfResource>
<mods:relatedItem id="F770BF1B8923E3A472F6CC4D51082D8F" type="host">
<mods:titleInfo id="FF65D1DD6D2EE91ECCB3E88DB08B1D1C">
<mods:title id="09D83FBBF4EA156572F2B1D34111693C">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part id="314D8562E61F719384F885B3B63E8F46">
<mods:date id="E1B408EDA63A2D64CC939D891E536838">2025</mods:date>
<mods:detail id="1D9C3DCC385FD10266C79A0760AAEE44" type="pubDate">
<mods:number id="507D5A04AFDC03E83938F771BD51EB15">2025-04-15</mods:number>
</mods:detail>
<mods:detail id="62A7DE6F2E3081B05877F3356D7593DE" type="volume">
<mods:number id="FCB513C57F0C500110C10FC5F3C7F65D">72</mods:number>
</mods:detail>
<mods:detail id="099F467687FFCAF737D84FF2C3058066" type="issue">
<mods:number id="8FC3FBC20CADA7A1CC027732A04AD9B5">1</mods:number>
</mods:detail>
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<mods:start id="471DC27213FA0A274BCB02011A9A83A7">9</mods:start>
<mods:end id="09973568742868572A109EEEA92C8136">14</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="65C5051293AD564631863F57A1B09306">journal article</mods:classification>
<mods:identifier id="0C5C418FB50552E5524FCF5E878B9768" type="DOI">10.3897/dez.72.144365</mods:identifier>
<mods:identifier id="431FC4431E914C52D6636D6C9C73D701" type="ZooBank">E82AFF4A-F2BA-48AD-A246-D966CC7EF57D</mods:identifier>
</mods:mods>
<treatment id="4344351F000C5E0B8BA1AB45D8D697D3" ID-arpha="4344351F-000C-5E0B-8BA1-AB45D8D697D3" ID-zoobank="https://zoobank.org/C23E8A6A-E46B-4660-8AE5-4A77983CD7DD" LSID="urn:lsid:plazi:treatment:4344351F000C5E0B8BA1AB45D8D697D3" LSID-ZBK="urn:lsid:zoobank.org:act:C23E8A6A-E46B-4660-8AE5-4A77983CD7DD" httpUri="http://treatment.plazi.org/id/4344351F000C5E0B8BA1AB45D8D697D3">
<subSubSection id="210613C930B73661933B70DB00B4FF44" type="nomenclature">
<paragraph id="D5CD160645022FA362EF2FBF617B983E">
<taxonomicName id="7FF9B1B322DDE3FBFBADE5A86D0AC045" authority="Yin &amp; Niu" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="2FD5F6AA46BF98CFBD8F706AF014986C" italics="true">Triartiger caiyujiei</emphasis>
Yin &amp; Niu
</taxonomicName>
<taxonomicNameLabel id="E137E034132E448561D795E0477B3582" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D1EDEC84D3388B5717C1F7ACBF7E3BE8" type="description">
<paragraph id="B9FEC3E850E29AC254D4C150AD04A463">
<figureCitation id="8B008C9E0DCA9C2C9A5B6C936983605E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">Figs 1</figureCitation>
,
<figureCitation id="4A1AF45A2996BC8A3B9FF30ECE2D5ED6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution and biology of Triartiger caiyujiei sp. nov. A. Distribution of Triartiger species on Chinas mainland; B. An adult T. caiyujiei sp. nov. with a male Pseudolasius ant; C. General environment of the collecting site; D, E. Host ant worker, full-face view (D) and lateral habitus (E)." figureDoi="10.3897/dez.72.144365.figure2" httpUri="https://binary.pensoft.net/fig/1308359">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELIAC" type="type material">
<paragraph id="50897C6A1888193C2F6F0C51B1813052">
<heading id="811E30281FB4206DAFEF7D5C24C875BF" reason="title">Type material.</heading>
</paragraph>
<paragraph id="FE20851300315F722B0FC5A899780B6C">
<materialsCitation id="6FCF14E759075AE9F40327FDAA6B311D" collectingDate="2024-05-03" collectionCode="SNUC" collectorName="Linan City, W &amp; C. - Y. Jie" country="China" county="China" elevation="1245" latitude="30.342562" location="Luopansong" longLatPrecision="1" longitude="119.42811" municipality="Tianmushan Nature Reserve" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
<emphasis id="0FB23B4DBF248EFA565F3BC3FC8CB5DA" bold="true">
<emphasis id="2275EA84ACD51E98237213DD8F233300" bold="true" italics="true">
<typeStatus id="422C2CF33B0F086565516EC5E90EB2E7" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="78A5FBE2FA3319B70BC6897CE1B215C6" name="China">China</collectingCountry>
</emphasis>
: •
<specimenCount id="E8B8A7721FC4875D9FFCF0975CB84F89" count="1" type="male"></specimenCount>
,
<collectingCounty id="006CB0BB1A3836035F255593FA34B998">China</collectingCounty>
:
<collectingRegion id="E1B37F3C631DA3AAC73A7599E6CD496B" country="China" name="Zhejiang">Zhejiang</collectingRegion>
,
<collectorName id="A7991C4683C8A3E8C68D4623E48D8DFD">Linan City, W</collectorName>
.
<collectingMunicipality id="87DA4B8207B1E2842D7F7B09E23D3166">Tianmushan Nature Reserve</collectingMunicipality>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZIAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[119.428108,30.342561]}">
<geoCoordinate id="7B2F4302DABE7CE95BE365D1FC8AC935" degrees="30" direction="north" minutes="20" orientation="latitude" precision="1" seconds="33.22" value="30.342562">30 ° 20 ' 33.22 &quot; N</geoCoordinate>
,
<geoCoordinate id="D596F70E58396208E857B71C89C66B69" degrees="119" direction="east" minutes="25" orientation="longitude" precision="1" seconds="41.19" value="119.42811">119 ° 25 ' 41.19 &quot; E</geoCoordinate>
</named-content>
, pass to summit, nr.
<location id="2EBE4C638FB5BFA0A43C3BD89C029FFA" LSID="urn:lsid:plazi:treatment:4344351F000C5E0B8BA1AB45D8D697D3:2EBE4C638FB5BFA0A43C3BD89C029FFA" country="China" county="China" latitude="30.342562" longLatPrecision="1" longitude="119.42811" municipality="Tianmushan Nature Reserve" name="Luopansong" stateProvince="Zhejiang">Luopansong</location>
,
<quantity id="DC9C95838ADE6DA789106A6E63B8596F" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" unit="m" value="1245.0">
<elevation id="97CCA8CDCA1EDA8F48B03682FF4BAEC7" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" unit="m" value="1245.0">1245 m</elevation>
</quantity>
elev.,
<date id="61C7C33C17115C6DD1E7398EBA85D013" value="2024-05-03">
<collectingDate id="B345E5B23ED304EABD013CBF1F6C9344" value="2024-05-03">03. v. 2024</collectingDate>
</date>
,
<collectorName id="69A213ABC4E8D65FF1C853871753D95B">C. - Y. Jie</collectorName>
leg., 天目山近罗盘松石块下蚁巢, 蔡余杰’ (
<abbrev id="ABBRID0E5IAC" xlink_title="Insect Collection of Shanghai Normal University">
<collectionCode id="CDF461AD8C50A952248828AE95293FD8">SNUC</collectionCode>
</abbrev>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECJAC" type="diagnosis">
<paragraph id="D80C99568BF24D3FEACEC9C285B7898E">
<heading id="A6B8F9012204FA97BBE615EBD6BA7F3F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6053209B777AD2F0F0A99BF72C5A92FE">
<emphasis id="97537EDA802FD6B4039682176F6FE6D2" bold="true">Male.</emphasis>
Habitus moderately stout, with relatively long antennae and legs. Body length approximately
<quantity id="279EF5F6DB45E460048E4830E2C19AA4" metricMagnitude="-3" metricUnit="m" metricValue="2.3" unit="mm" value="2.3">2.3 mm</quantity>
. Head and pronotal base with distinct microsculpture cuticle and covered by suberect setae; antenna with short antennomeres 1 and 2 and exceptionally long antennomere 3; 3 approximately 13 × as long as 2. Elytra finely punctate, lacking linear sculpture, posterior margins with large trichomes. Mesofemur with large blade-like ventral spine, mesotibia with large triangular spine at approximately middle of mesal margin; metafemur simple. Abdomen suboval; composite tergite deeply and broadly concave at base, with pair of broad basolateral trichomes, disc glabrous and with long, sparse setae; paratergites 1 (
<collectionCode id="F279258D532D8D6611B71C2E644DD8B5">IV</collectionCode>
) with broadly curved longitudinal trichomes at bases; posterior margin of composite tergite and tergite 4 (
<collectionCode id="C47E2F0496A7423EC96AF9C67FF24ADB">VII</collectionCode>
) with row of long setae. Aedeagus dorso-ventrally symmetrical, divided into basal and apical parts; basal capsule with large ventral projection, apical lobe abruptly narrowed before apex.
<emphasis id="8A810BF7E4A3ABD831F29451F882BE53" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMJAC" type="description">
<paragraph id="D62DE58D570FEB87BE1951E6D277AE6E">
<heading id="9BCA334D916C41BF914803152EEC3956" reason="title">Description.</heading>
</paragraph>
<paragraph id="3EDE2CA52A45AE6F38491641CB109D0B">
<emphasis id="F3E607728183F7062EE0D70A0816F887" bold="true">Male.</emphasis>
Body (Fig.
<figureCitation id="4A647C7BFF81A856CF7576255D242FB4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 A</figureCitation>
) length
<quantity id="26A0325157F263AFA49FEF0FDA6B6521" metricMagnitude="-3" metricUnit="m" metricValue="2.33" unit="mm" value="2.33">2.33 mm</quantity>
; colour reddish-brown, tarsi and mouthparts lighter in colour. Dorsal surface of head, pronotum and elytral disc covered with suberect setae, those of head and pronotum distinctly longer than elytral disc; setae of abdominal disc sparse and long, those along posterior margin much denser than disc.
</paragraph>
<caption id="9F291D8E2654D0C6842D0DF09CC2B77A" ID-DOI="10.3897/dez.72.144365.figure1" ID-arpha="F0C0C448-8F38-5190-86C6-9018021FA788" httpUri="https://binary.pensoft.net/fig/1308358" startId="F1">
<paragraph id="7DAF93CDBC68E600A3455E198B2A7173">
<label id="EEB81DD0272FDFFCC627A2894A3F5A38">Figure 1.</label>
</paragraph>
<paragraph id="152EE1FB75E858A7514BA7ECA28858C9">
<taxonomicName id="9A8E4DA03E6224F2A2B52E4CF17D8B49" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="8B0DA7434A5761C37370A47F90A167C9" italics="true">Triartiger caiyujiei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31245BB083E5B32D892294276E86A1A9" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="C2CA6454F34D77E6004A5B111F4D4D1B" bold="true">A.</emphasis>
Dorsal habitus;
<emphasis id="C45D6D3F600233D33231671833E342D5" bold="true">B.</emphasis>
Head, lateral (vf = vertexal fovea);
<emphasis id="ED9AF8E9DA089BD83E2199093024FEF6" bold="true">C.</emphasis>
Head dorsum and pronotum;
<emphasis id="F62D376E2BBFA851A2E355EADBF259E8" bold="true">D.</emphasis>
Antenna;
<emphasis id="03AB4E465805C90C9462CAEED235A31F" bold="true">E.</emphasis>
Trichome of elytral apex and abdominal base;
<emphasis id="17D178492CB96E06B0A4512040932996" bold="true">F.</emphasis>
Middle leg;
<emphasis id="B1B5716208D5190E61845E25BCD1CC31" bold="true">G.</emphasis>
Hind leg;
<emphasis id="0509F065D8F3AE5009F838AF2F158B9A" bold="true">H J.</emphasis>
Aedeagus, dorsal (
<emphasis id="4E2B0AE9659BC80CF771628B4AE487EC" bold="true">H</emphasis>
), lateral (
<emphasis id="6E6FB283EAB273C3B25408223ACA8542" bold="true">I</emphasis>
) and ventral (
<emphasis id="701ADE16798947D27F01E88E4C9049B3" bold="true">J</emphasis>
). Scale bars: 0.5 mm (
<emphasis id="1DEBDC5CF1A93BA769E5F28CCF8FA8FC" bold="true">A</emphasis>
); 0.3 mm (
<emphasis id="41C6BEFDD97F040143BE277291F757D3" bold="true">B D, F, G</emphasis>
); 0.2 mm (
<emphasis id="040AB321B4A35B30AD56A8D3F59A4E19" bold="true">E, H J</emphasis>
).
</paragraph>
</caption>
<paragraph id="130AB7C039BAF2734394B257D827A237">
Head (Figs
<figureCitation id="31C2B36AFA3B2014416C7A9E926340F3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 B and C</figureCitation>
) dorsally subcylindrical, much longer than wide, length
<quantity id="C2CBBC2DC9DE9B621F933163C5B759B4" metricMagnitude="-4" metricUnit="m" metricValue="4.6" unit="mm" value="0.46">0.46 mm</quantity>
, width across eyes
<quantity id="F320E9AD95F697197D7C141087377327" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.30 mm</quantity>
; vertex slightly convex, foveae (dorsal tentorial pits) (Fig.
<figureCitation id="293FC23696CEA2FE624468B4FD577396" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 B</figureCitation>
;
<emphasis id="B31C8C9CBEBDBF870B321693BD050DD2" italics="true">vf</emphasis>
) laterally located just above eyes; frons forming narrow rostrum, extending anteriorly to form vertical keel at middle; vertex and lateral portions of head with distinct microsculpture, frons smooth, finely punctate; clypeus markedly short, anterior margin with short, sparse setae. Venter with rough surface, gular region broadly and moderately impressed, tiny gular foveae (posterior tentorial pits) widely separated, lacking median carina. Compound eyes weakly prominent, each composed of approximately 10 ommatidia. Cervical part sub-globose, surface with distinct reticulate microsculpture. Antenna (Fig.
<figureCitation id="E787C2651B9A273017A01B38CB4327A3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 D</figureCitation>
)
<quantity id="ADF928E88FB7F31F8C3D7E68B7129AAA" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" unit="mm" value="0.72">0.72 mm</quantity>
long, lacking modifications; antennomere 1 thick and short, dorsally completely covered by rostrum, invisible, 2 slightly shorter than 1, transverse, dorsally partially visible, antennomere 3,
<quantity id="34853B60AB24DE856E8EE4096393B585" metricMagnitude="-4" metricUnit="m" metricValue="6.7" unit="mm" value="0.67">0.67 mm</quantity>
long,
<quantity id="FC76F38A3E7E6E2078671F274D8DA300" metricMagnitude="-4" metricUnit="m" metricValue="1.5" unit="mm" value="0.15">0.15 mm</quantity>
wide, 13.4 × as long as 2, narrowing from base towards apex, broadest and truncate at apex.
</paragraph>
<paragraph id="63A176A09695A61926045EFAF8EA83F8">
Pronotum (Fig.
<figureCitation id="3D8F88AE7E0D9FD9C81F243DD966C11C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 C</figureCitation>
) approximately as long as wide, length
<quantity id="5556639E408EFE85221349F71CF6E840" metricMagnitude="-4" metricUnit="m" metricValue="4.4" unit="mm" value="0.44">0.44 mm</quantity>
, width
<quantity id="A36BF26D26F76DC9DD32F05DCC64A838" metricMagnitude="-4" metricUnit="m" metricValue="4.3" unit="mm" value="0.43">0.43 mm</quantity>
, widest at approximately anterior 1 / 3, basal 1 / 3 with microsculpture similar to that of head; lateral margins rounded at widest point, narrowing anteriorly and slightly so posteriorly; disc slightly impressed mediobasally, lacking foveae. Prosternum with basisternal (precoxal) portion longer than procoxal rests, lacking procoxal foveae; hypomera completely fused with sternum, lacking hypomeral grooves and carinae.
</paragraph>
<paragraph id="3A5F838D87D7795121886D5E8E9BA68B">
Elytra tightly joined, subtrapezoidal, much wider than long, length
<quantity id="ECE77A8BADDD508154CD49E675619018" metricMagnitude="-4" metricUnit="m" metricValue="4.9" unit="mm" value="0.49">0.49 mm</quantity>
, width
<quantity id="A2D3B0768C006C4297169DC7D4E054E2" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" unit="mm" value="0.94">0.94 mm</quantity>
, lacking foveae, anterior margin greatly oblique; each elytron with one complete adsutural carina and one short discal carina originating from base; posterolateral margins roundly convex; posteromedial portions with patches of dense, long setae, posterolateral portions with large, triangular trichomes (Fig.
<figureCitation id="398D6926640B49D4B1776C4A9F84C24F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 A</figureCitation>
). Metathoracic wings remnant.
</paragraph>
<paragraph id="80AABB7FD61077D3D904EBD7574066A1">Mesoventrite short, fused with metaventrite, with median mesoventral foveae anterior to median keel, the latter angularly projecting at middle; pair of small lateral mesoventral foveae present. Metaventrite greatly convex at middle, descending laterally, lacking foveae, broad metaventral intercoxal process widely separating coxae and broadly and slightly convex at middle; area anterior to process with golden, densely setose patch.</paragraph>
<paragraph id="75EB26FA12EE800DF0EE61B0359CE38C">
Legs long; mesofemur and mesotibia (Fig.
<figureCitation id="9AB8F7510923A047BA6AED00D8782C55" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 F</figureCitation>
) each with one large, blade-like and one much larger, subtriangular spine on ventral and middle of mesal margin, respectively; metafemur (Fig.
<figureCitation id="3422E5495DBDCE413B6C801755C07511" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 G</figureCitation>
) simple, lacking tubercles or spines.
</paragraph>
<paragraph id="F3423BFAE2AD854A35B3B55293539005">
Abdomen slightly broader than elytra, suboval, widest at basolateral margins of composite tergite, length
<quantity id="D02A947D61E4968328A03709C2BE7D8B" metricMagnitude="-4" metricUnit="m" metricValue="9.2" unit="mm" value="0.92">0.92 mm</quantity>
, width
<quantity id="86A6D6A2DCAA366D66BBDF7584B0EACC" metricMagnitude="-4" metricUnit="m" metricValue="9.8" unit="mm" value="0.98">0.98 mm</quantity>
. Tergite 1 (composite tergite formed by tergites 13 (IV
<collectionCode id="68C047F25EBC1857D7BDF464D170013E" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
)) deeply concave at base, with pair of large basolateral sockets, with large trichomes (Fig.
<figureCitation id="EAE57E5AC9DCF258DA22A2F4AF686A58" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 E</figureCitation>
) mesal and posterior to sockets, 2 (
<collectionCode id="1BDE4FB1FEC815A64EFA1A66AC722947">VII</collectionCode>
) short, transverse, with pair of small lateral nodules and foveae, 3 (
<collectionCode id="AE93886A26326E57CD4A9906AB289094">VIII</collectionCode>
) transverse, roundly triangular, with pair of small basolateral foveae, posterior margin rounded. Paratergites 1 (
<collectionCode id="E1B5F26705EE501EF9C5B6B40DC2D821">IV</collectionCode>
) with greatly curved, longitudinal trichomes (Fig.
<figureCitation id="61294EBC0026CA44307A27F8D139B07C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 E</figureCitation>
) at bases. Sternites finely punctate, 2 (
<collectionCode id="AEDA75FD19D25F3318E674A3C4D5964A">IV</collectionCode>
) as long as 3 (
<collectionCode id="CDAE34C67860A4458C46E88B0E237DA4" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
) 5 (
<collectionCode id="DA0E5AB2D43DFED969259EA472E15D01">VII</collectionCode>
) combined, with band of dense setae along base, with slightly prominent keel at middle, area lateral to keel weakly impressed, lacking foveae, 3 at middle as long as 4 (
<collectionCode id="1A173DC49C6886BE4323B3FA9D0F1FEB" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
) and 5 combined, 24 each with distinct microsculpture on posterior half; 6 (
<collectionCode id="9E208F9CCD94796D570A40EFDDE20E41">VIII</collectionCode>
) transverse, posterior margin broadly emarginate at middle, 7 (
<collectionCode id="BEA997DDF5015C066232FF8032AC6DD3">IX</collectionCode>
) not observed.
</paragraph>
<paragraph id="1A24A5C75C127928421565805F1786E9">
Aedeagus (Figs
<figureCitation id="6AECC14803B7AEE83550079AD0B29C63" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Triartiger caiyujiei sp. nov., male. A. Dorsal habitus; B. Head, lateral (vf = vertexal fovea); C. Head dorsum and pronotum; D. Antenna; E. Trichome of elytral apex and abdominal base; F. Middle leg; G. Hind leg; H J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm (A); 0.3 mm (B D, F, G); 0.2 mm (E, H J)." figureDoi="10.3897/dez.72.144365.figure1" httpUri="https://binary.pensoft.net/fig/1308358">1 H J</figureCitation>
)
<quantity id="9633546A8E6F8994A34F241DEBCADE89" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" unit="mm" value="0.47">0.47 mm</quantity>
long, dorsoventrally symmetrical; median lobe divided into basal capsule and apical lobe, lacking parameres; basal capsule elongate, broadened and rounded at base, with large ventral projection and small, transverse foramen, dorsally with large oval diaphragm, apical lobe narrowing apically and abruptly narrowed before apex, in lateral view pointed apically, with two short, tiny apical setae; endophallus armature composed of short sclerite at middle of capsule.
</paragraph>
<paragraph id="FF1604ED794A204A2BE7B3C9914FFA83">
<emphasis id="FEE317804DEE8CE8F705DC7920C69B81" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHNAC" type="comparisons">
<paragraph id="A182221CCA8E9BA093DE20C3116A51A1">
<heading id="EBD9B4ED4C51C0339BDBCFA11824B7AD" reason="title">Comparative notes.</heading>
</paragraph>
<paragraph id="6EE6CF0A738881AF9421F215EE58AD94">
The addition of the present species brings the total number of known species of the genus to seven. This species is relatively large within the genus and can be readily distinguished from
<taxonomicName id="85144CF6E8A58F0479EC50309DEDCBDD" authority="Nomura" authorityName="Nomura" authorityYear="1997" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="89534CDA453EAA16C7EAA4B35173710B" italics="true">T. reductus</emphasis>
Nomura
</taxonomicName>
,
<taxonomicName id="880474350BF31B44F6457FAE12E38D5D" authority="Yin &amp; Du" authorityName="Yin &amp; Du" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jiangxinongda">
<emphasis id="A72B384F4B8D251527B359D0FA9FB8BE" italics="true">T. jiangxinongda</emphasis>
Yin &amp; Du
</taxonomicName>
and
<taxonomicName id="5FF4D02E85E61A9122AB477D95EDAF7E" authority="Yin" authorityName="Yin" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="55F10B11C1C0FA66543C66643D0AC623" italics="true">T. spatulatus</emphasis>
Yin
</taxonomicName>
by its exceptionally elongate terminal antennal segment, whereas the latter three species possess considerably shorter antennae in proportion to their head size. The new species can be easily differentiated from
<taxonomicName id="B323F176EC9C2B5E37F2EDA457390831" authority="Kubota" authorityName="Kubota" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="urceus">
<emphasis id="C1F0121B7911427B403786A616D0AC1C" italics="true">T. urceus</emphasis>
Kubota
</taxonomicName>
and
<taxonomicName id="3DD6D98DFC8D9CC8709D68D871938D44" authority="Besuchet" authorityName="Besuchet" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="1D8508E633032A661143316316B33412" italics="true">T. nomurai</emphasis>
Besuchet
</taxonomicName>
by the nearly straight mesal and lateral margins of antennomere 3, in contrast to the slightly swollen mesal margins in the latter two species.
<taxonomicName id="56139367F312630222AA478593A856FD" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="353719DDFD93055D425DEB4CC1B682B5" italics="true">Triartiger caiyujiei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA395EC0ED8A8922B804F3CAF4F99F61" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="BA78B41AB6428BEBC037486EF4CC52FB" authority="Nomura" authorityName="Nomura" authorityYear="1997" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="klapperichorum">
<emphasis id="B81ABCC8A2D6262E04291049C2ADE234" italics="true">T. klapperichorum</emphasis>
Nomura
</taxonomicName>
in its relatively longer antennomere 3 (13 × vs. 8.5 × as long as antennomere 2) and the presence of additional setose patches on the posteromedial portions of the elytra. To facilitate identification, an updated key to all known species is provided below.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1PAC" type="distribution">
<paragraph id="7CB35130FE946B6A4445008D7A018398">
<heading id="482F357E1A5B2EF9D9181FCDC47C59DF" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1D1BE2879C070C2931BB51BC34EFC19A">
East
<collectingCountry id="3EBD40EAB0C8B5B20CA63B541407E3CA" name="China">China</collectingCountry>
:
<collectingRegion id="4ED7EDE122C84AA88047A3DC58C1D7D5" country="China" name="Zhejiang">Zhejiang</collectingRegion>
(Fig.
<figureCitation id="1096F577A3E6C0939168034182927689" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution and biology of Triartiger caiyujiei sp. nov. A. Distribution of Triartiger species on Chinas mainland; B. An adult T. caiyujiei sp. nov. with a male Pseudolasius ant; C. General environment of the collecting site; D, E. Host ant worker, full-face view (D) and lateral habitus (E)." figureDoi="10.3897/dez.72.144365.figure2" httpUri="https://binary.pensoft.net/fig/1308359">2 A</figureCitation>
).
</paragraph>
<caption id="9FF40FAF74C7C9655DA1BB8526F7169F" ID-DOI="10.3897/dez.72.144365.figure2" ID-arpha="266BED56-9252-53D4-B9D7-F584D57D2C8D" httpUri="https://binary.pensoft.net/fig/1308359" startId="F2">
<paragraph id="4768277BB6974F70463AB6094E826ABC">
<label id="58E3BD6797A78F752A486EAB4A2CC94F">Figure 2.</label>
</paragraph>
<paragraph id="342A626B93E650A1C0B267DECCF800E7">
Distribution and biology of
<taxonomicName id="5914EFE90647B8D28FB1C7920A7555E0" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="587551B3D20774007534B4B6D8B4A5D4" italics="true">Triartiger caiyujiei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9366968476B1DD0B452F541475EAD3F3" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="63231C15E2667BF3E3BCC5CB490D499D" bold="true">A.</emphasis>
Distribution of
<taxonomicName id="19BEA1BF6CC5C6B4F4DC5AAB1B697407">
<emphasis id="2017FA6EF9FA505F52B2E2328653EC48" italics="true">Triartiger</emphasis>
species
</taxonomicName>
on Chinas mainland;
<emphasis id="3A9E98568E9558B0BAB9957A2753DB5C" bold="true">B.</emphasis>
An adult
<taxonomicName id="BD02EC61FA4DB501202A8CF86850396B" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="BE13329843F105B5308D5BBA6B26FA4A" italics="true">T. caiyujiei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E40C460770D7AF228725E11A648E0150" rank="species">sp. nov.</taxonomicNameLabel>
with a male
<taxonomicName id="A1F55768F5BCACABCF2021DC72CA8BAE" authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="BEC14F3D2ADBEE4EEDF05BC62E003476" italics="true">Pseudolasius</emphasis>
</taxonomicName>
ant;
<emphasis id="229852B6E697B7B167622A332B776105" bold="true">C.</emphasis>
General environment of the collecting site;
<emphasis id="A35E0CF3AF6DBCA1E97D49945BF10800" bold="true">D, E.</emphasis>
Host ant worker, full-face view (
<emphasis id="3C4E6FE3597F481D8278F7BA8F804F17" bold="true">D</emphasis>
) and lateral habitus (
<emphasis id="CB8C31623F987D85A790D2B7102F5726" bold="true">E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ECCAE" type="biology">
<paragraph id="AC3DA258D14F2F6DBE166C25CB606E78">
<heading id="E76BAB71047AD46911F90EBB56360D1F" reason="title">Biology.</heading>
</paragraph>
<paragraph id="2A44E63F2D3AAD293ABD7C56BF9FFB03">
The single adult was collected from a small
<taxonomicName id="A2CB26ECC92F84884F711A17D2523F7F" authority="Emery" authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="7CC99375789906A539204AC7CCE4288F" italics="true">Pseudolasius</emphasis>
Emery
</taxonomicName>
ant colony nesting under a stone (Fig.
<figureCitation id="F6037CCF77EA622CDF39931649DE6610" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution and biology of Triartiger caiyujiei sp. nov. A. Distribution of Triartiger species on Chinas mainland; B. An adult T. caiyujiei sp. nov. with a male Pseudolasius ant; C. General environment of the collecting site; D, E. Host ant worker, full-face view (D) and lateral habitus (E)." figureDoi="10.3897/dez.72.144365.figure2" httpUri="https://binary.pensoft.net/fig/1308359">2 B</figureCitation>
) in a mixed coniferous-deciduous forest (Fig.
<figureCitation id="EB262B75B6C2783BE9A0975B50E9D086" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution and biology of Triartiger caiyujiei sp. nov. A. Distribution of Triartiger species on Chinas mainland; B. An adult T. caiyujiei sp. nov. with a male Pseudolasius ant; C. General environment of the collecting site; D, E. Host ant worker, full-face view (D) and lateral habitus (E)." figureDoi="10.3897/dez.72.144365.figure2" httpUri="https://binary.pensoft.net/fig/1308359">2 C</figureCitation>
). This represents only the third
<taxonomicName id="59C880D0A80F41B2E93745C5B2A60D6E">
<emphasis id="92C2EB588ED3E733B249644DCE72C748" italics="true">Triartiger</emphasis>
species
</taxonomicName>
with a known symbiotic host (Fig.
<figureCitation id="81BFE83E8B79161735ADAFCA67A2A6E9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution and biology of Triartiger caiyujiei sp. nov. A. Distribution of Triartiger species on Chinas mainland; B. An adult T. caiyujiei sp. nov. with a male Pseudolasius ant; C. General environment of the collecting site; D, E. Host ant worker, full-face view (D) and lateral habitus (E)." figureDoi="10.3897/dez.72.144365.figure2" httpUri="https://binary.pensoft.net/fig/1308359">2 D, E</figureCitation>
). Members of this genus are notably rare in both the field and museum collections and the biology and natural history of most species remain poorly understood. The current knowledge on the distribution, collecting methods, host associations and material availability of the genus is summarised in Table
<tableCitation id="5111BDBE194870DDA06CBBC8553B6465" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Current knowledge of the genus Triartiger." httpUri="http://table.plazi.org/id/5053A1B873E7D7078CEE803183B1F37A" tableUuid="5053A1B873E7D7078CEE803183B1F37A">1</tableCitation>
to facilitate future studies.
</paragraph>
<caption id="5053A1B873E7D7078CEE803183B1F37A" ID-Table-UUID="5053A1B873E7D7078CEE803183B1F37A" httpUri="http://table.plazi.org/id/5053A1B873E7D7078CEE803183B1F37A" startId="T1" targetIsTable="true">
<paragraph id="047E71D65E24CD85BD2A7D8083C3FF1B">
<label id="31DA4BCCEDDB1080AFB0C46A171B0ACF">Table 1.</label>
</paragraph>
<paragraph id="256CE366BE79C4F56CA5BDB4A7800E5F">
Current knowledge of the genus
<taxonomicName id="7049E7F005BAA88B9A598FBB368AAC27" authorityName="Kubota" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="FF34DDCEBE28C9AB0B8694363D2F697D" italics="true">Triartiger</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="710A721D81D7A4C27CEBE5AB52424E0C">
<table id="TID0EUTAE" rules="all">
<tbody id="409BF2DB1A5BF1DD2C0CB1B066D5B187">
<tr id="F90088316B35AE485D0F124AB6CA3ADE">
<th id="3657D39B7420A686458B8CA3B7814586" colspan="1" rowspan="1">Species</th>
<th id="142B5CCE313C700E8A7AFDD552F730CF" colspan="1" rowspan="1">Distribution</th>
<th id="29D5CA5020544AA94FF572F0F1E2E91A" colspan="1" rowspan="1">Collecting method</th>
<th id="B9A277ED6AB78978D788C0B82B222CDD" colspan="1" rowspan="1">Host</th>
<th id="634EF7BD67E1BED90E79126E813C881D" colspan="1" rowspan="1">Available material</th>
<th id="EBF6B2A6DE70F6C3AD8928A28BF56BB9" colspan="1" rowspan="1">Reference</th>
</tr>
<tr id="7E00E76BA1BA59BBF56EBE9CB5DDCF2D">
<td id="98898DB36E930E0B8D0F8F3D6950D6EA" colspan="1" rowspan="1">
<taxonomicName id="14B1AAC6C712DDDC06A4F1F19AD75F19" authorityName="Yin &amp; Niu" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="caiyujiei" status="sp. nov.">
<emphasis id="09BC85E57E5980B6596C77ACC74B32BA" italics="true">T. caiyujiei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="959651742E2C0CCAE83FDC2AE4811894" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="C8913AAE0CC1BA17171816915B78DF05" colspan="1" rowspan="1">Zhejiang</td>
<td id="92993F117333D056EF00E58C23395D3B" colspan="1" rowspan="1">by hand</td>
<td id="6CA0D023C672B4603D76CF373559A3F5" colspan="1" rowspan="1">
<taxonomicName id="375BC2045E6CF602A12481E2C338F536" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="33FD46F9EE679840826D87419B17D46F" italics="true">Pseudolasius</emphasis>
sp.
</taxonomicName>
</td>
<td id="F967A05D2232E2042C08E252CA064E0E" colspan="1" rowspan="1">Holotype ♂</td>
<td id="342F68EC229DED5ECCC57AD628C9A4E0" colspan="1" rowspan="1">present study</td>
</tr>
<tr id="E256C0B4CE214A04F274FF73C9541BB4">
<td id="C67A9B29B926A76D7D0F07D79651BF82" colspan="1" rowspan="1">
<taxonomicName id="3D22ACAC8E4F1D3B8F036F2AA8FCAEA3" authority="Yin &amp; Du, 2024" authorityName="Yin &amp; Du" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jiangxinongda">
<emphasis id="B0E9CA58AB718D8BA9860F90C69727AB" italics="true">T. jiangxinongda</emphasis>
<bibRefCitation id="E91EDDA4ABDDB3572B77DF8B432765B3" DOI="10.11646/zootaxa.5538.5.8" author="Yin Z-W &amp; Du J-Z" issue="5" journalOrPublisher="Zootaxa" pagination="479-484" refId="B12" refString="Yin Z-W, Du J-Z (2024) The second species of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) from China's mainland. Zootaxa 5538 (5): 479-484. https://doi.org/10.11646/zootaxa.5538.5.8" title="The second species of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) from Chinas mainland." volume="5538" year="2024">Yin &amp; Du, 2024</bibRefCitation>
</taxonomicName>
</td>
<td id="93C8E0AC1EB626FBAD95D300E82B1F28" colspan="1" rowspan="1">Jiangxi</td>
<td id="192E74E6CF8E65F7D2ADEE6E5808B45D" colspan="1" rowspan="1">
FIT
<superScript id="DA9308579F4617E0084CD6C5E9A83D9D">*</superScript>
</td>
<td id="DBA5B99268586990955CB4E9BAC985A8" colspan="1" rowspan="1">unknown</td>
<td id="8D498D5ACEAA78EE01975E72AD078C67" colspan="1" rowspan="1">Holotype ♂</td>
<td id="95A73CF5BA9CBE51595BAC25472CE50D" colspan="1" rowspan="1">
<bibRefCitation id="AD3555B0002FC34AB6E82A7069332D8B" DOI="10.11646/zootaxa.5538.5.8" author="Yin Z-W &amp; Du J-Z" firstAuthor="Yin" issue="5" journalOrPublisher="Zootaxa" pagination="479-484" refId="B12" refString="Yin Z-W, Du J-Z (2024) The second species of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) from Chinas mainland. Zootaxa 5538 (5): 479484. https://doi.org/10.11646/zootaxa.5538.5.8" title="The second species of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) from Chinas mainland." volume="5538" year="2024">Yin and Du (2024)</bibRefCitation>
</td>
</tr>
<tr id="6F021CD2026D0610BAEC3B14E2282FEF">
<td id="DC98F8706F01200AC34369A73B95A1A3" colspan="1" rowspan="1">
<taxonomicName id="C930D9588106DF00C18A9DCABD36705D" authority="Nomura, 1997" authorityName="Nomura" authorityYear="1997" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="klapperichorum">
<emphasis id="7576E8D817299C63CB007185BCABFFAC" italics="true">T. klapperichorum</emphasis>
<bibRefCitation id="82A3165031CE5FB3DE1FB3AE1D5669A6" author="Nomura S" issue="2" journalOrPublisher="Elytra" pagination="435-444" refId="B5" refString="Nomura S (1997) A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan. Elytra 25 (2): 435-444." title="A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan." volume="25" year="1997">Nomura, 1997</bibRefCitation>
</taxonomicName>
</td>
<td id="492ACFE4F87F84FB792E197D07F9C87D" colspan="1" rowspan="1">Taiwan</td>
<td id="5AE78325ADCDE57BCB521A3C26BB91B8" colspan="1" rowspan="1">
FIT
<superScript id="8E4E3BD030CEF6F1C7C3EE9BC5BDA193">*</superScript>
</td>
<td id="36CB4C5D7C133680C400D824E7D7EBAC" colspan="1" rowspan="1">unknown</td>
<td id="DCDD2755BFB055FC191A345D4149AA9B" colspan="1" rowspan="1">Holotype ♀ + 1 ♂, 6 ♀♀</td>
<td id="06BD0829507E94386F6A5C532301FE3F" colspan="1" rowspan="1">
<bibRefCitation id="B412D3067B8CE228206C05FF4544DD1B" author="Nomura S" firstAuthor="Nomura" issue="2" journalOrPublisher="Elytra" pagination="435-444" refId="B5" refString="Nomura S (1997) A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan. Elytra 25 (2): 435444." title="A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan." volume="25" year="1997">Nomura (1997)</bibRefCitation>
;
<bibRefCitation id="0413D3B2BA04713947768E3D6F9B1283" author="Sugaya H &amp; Maruyama M" firstAuthor="Sugaya" issue="1" journalOrPublisher="Entomological Review of Japan" pagination="11-12" refId="B7" refString="Sugaya H, Maruyama M (2003) Additional records of Triartiger klapperichorum from Taiwan, collected by use of flight interception traps (Coleoptera: Staphylinidae: Pselaphinae). Entomological Review of Japan 58 (1): 1112." title="Additional records of Triartiger klapperichorum from Taiwan, collected by use of flight interception traps (Coleoptera: Staphylinidae: Pselaphinae)." volume="58" year="2003">Sugaya and Maruyama (2003)</bibRefCitation>
</td>
</tr>
<tr id="6D91F61FE0CF0491709604A64201DA81">
<td id="6F265BC5F3E63560D6C87866BC826004" colspan="1" rowspan="1">
<taxonomicName id="5589C951EEB419FBB1EAF37D7608FF79" authority="Besuchet, 2008" authorityName="Besuchet" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="EDDD9CD04EE4618576467485A66F082E" italics="true">T. nomurai</emphasis>
<bibRefCitation id="2589719F4B613BA2506BD04004CC4486" author="Besuchet C" issue="1-2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="61-82" refId="B1" refString="Besuchet C (2008) Synonymies et combinaisons nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 81 (1 - 2): 61-82." title="Synonymies et combinaisons nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae)." volume="81" year="2008">Besuchet, 2008</bibRefCitation>
</taxonomicName>
</td>
<td id="45071C7DA3C94E397BB467A57FBF60CF" colspan="1" rowspan="1">Taiwan</td>
<td id="DC14FC5E45D09CD08B8FDFA3D63FF422" colspan="1" rowspan="1">unknown</td>
<td id="4019A30A960791CEE8C5C00970168DA8" colspan="1" rowspan="1">unknown</td>
<td id="BE0BADB3B2B6F269270457D640CF37A0" colspan="1" rowspan="1">Holotype ♂ + paratype 1 ♂, 2 ♀♀</td>
<td id="77A4B2F88BDC11D8FF743A042FFC25CA" colspan="1" rowspan="1">
<bibRefCitation id="EE896D8B16F34A2E1EED37FCCB140B5A" author="Besuchet C" firstAuthor="Besuchet" issue="1-2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="61-82" refId="B1" refString="Besuchet C (2008) Synonymies et combinaisons nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 81 (12): 6182." title="Synonymies et combinaisons nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae)." volume="81" year="2008">Besuchet (2008)</bibRefCitation>
</td>
</tr>
<tr id="70A7E391C908896C0F21CC844ADA153A">
<td id="D8A1EB3B13FE95183A41FA7F051B2216" colspan="1" rowspan="1">
<taxonomicName id="FC72121B23EC484BD07AE16F97D9CD7C" authority="Nomura, 1997" authorityName="Nomura" authorityYear="1997" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="1229A18EA39C7C24E12C252BE6436E58" italics="true">T. reductus</emphasis>
<bibRefCitation id="CFFCC479698C4907BC399EEB662B2857" author="Nomura S" issue="2" journalOrPublisher="Elytra" pagination="435-444" refId="B5" refString="Nomura S (1997) A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan. Elytra 25 (2): 435-444." title="A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan." volume="25" year="1997">Nomura, 1997</bibRefCitation>
</taxonomicName>
</td>
<td id="8F16F6FB38A49D9CAA9BBF1147F6A4D6" colspan="1" rowspan="1">Tsushima</td>
<td id="B2CB5190B104E928CA8A0D0B9BD0F2AD" colspan="1" rowspan="1">unknown</td>
<td id="BBB8D2F238FE976CBA9DBD2E453996C6" colspan="1" rowspan="1">
<taxonomicName id="D3498536E048F692EE48DD251FC093A0" authorityName="Forel" authorityYear="1900" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="osakensis">
<emphasis id="14955784F05D6F5E6F0FC15A910486D1" italics="true">Crematogaster osakensis</emphasis>
</taxonomicName>
</td>
<td id="498492CF101D44E0A8D7F9E4806DE2A9" colspan="1" rowspan="1">Holotype ♀ + 1 ♂, 1 ♀ + at least 13 exx.</td>
<td id="AE4CB7BC9BC565E1FAF7BF29120F20CF" colspan="1" rowspan="1">
<bibRefCitation id="1141080F978D2F8B11B8E00213FB9E85" author="Nomura" firstAuthor="Nomura" issue="2" journalOrPublisher="Elytra" pagination="435-444" refId="B5" refString="Nomura S (1997) A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan. Elytra 25 (2): 435444." title="A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan." volume="25" year="1997">Nomura (1997</bibRefCitation>
,
<bibRefCitation id="C5780FE168FC00D86E5C966C7AFD1DDD" author="Nomura" firstAuthor="Nomura" issue="2" journalOrPublisher="Elytra" pagination="499-501" refId="B6" refString="Nomura S (1999) Further note on the clavigerine species Triartiger reductus Nomura (Coleoptera, Staphylinidae, Pselaphinae). Elytra 27 (2): 499501." title="Further note on the clavigerine species Triartiger reductus Nomura (Coleoptera, Staphylinidae, Pselaphinae)." volume="27" year="1999">1999</bibRefCitation>
);
<bibRefCitation id="3992DAD5B476156B7BA83BF4C620181E" author="Maruyama M &amp; Komatsu T &amp; Kinomura K &amp; Shimada T" etAl="et al." firstAuthor="Maruyama" refId="B4" refString="Maruyama M, Komatsu T, Kinomura K, Shimada T (2013) The guests of Japanese ants. Tokai University Press, Minamiyana, Hadano-shi, Kanagawa, Japan, [xii +] 208 pp. [in Japanese and English]" year="2013">Maruyama et al. (2013)</bibRefCitation>
</td>
</tr>
<tr id="0368DE4CAB5D9534694A5472FFEA337E">
<td id="A63D27300A1A6E368DB8F4B859245214" colspan="1" rowspan="1">
<taxonomicName id="123AB9D67A31F8D7EBF635EFB71E8D4C" authority="Yin, 2023" authorityName="Yin" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="AF0F5DA3CE22EF41DF88D85146A5CC58" italics="true">T. spatulatus</emphasis>
Yin, 2023
</taxonomicName>
</td>
<td id="51C990B4A3405F1E086BF32E3B926AFD" colspan="1" rowspan="1">Fujian</td>
<td id="026C01CA15916791BEEA413EB38854EE" colspan="1" rowspan="1">by hand</td>
<td id="FC06A27EA7CD110B381257974AF67582" colspan="1" rowspan="1">
<taxonomicName id="D53F9C6B8398A27F1AA063A615A5BF1B" authorityName="Wheeler (Yin" authorityYear="2023" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="nipponense">
<emphasis id="AA587A49DC4731375EF772CCA9FD27F6" italics="true">Tetramorium nipponense</emphasis>
</taxonomicName>
</td>
<td id="3B4926185277AE767B588D9CB2563FB0" colspan="1" rowspan="1">Holotype ♂</td>
<td id="A00F8B6950B532C1361ABA839FD11577" colspan="1" rowspan="1">
<bibRefCitation id="B7724768030AA732020A8CCE9E8DE15C" DOI="10.1649/0010-065X-77.4.549" author="Yin Z-W" issue="4" journalOrPublisher="Coleopterists Bulletin" pagination="549-553" refId="B11" refString="Yin Z-W (2023 b) Occurrence of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) on Chinas Mainland. Coleopterists Bulletin 77 (4): 549553. https://doi.org/10.1649/0010-065X-77.4.549" title="Occurrence of Triartiger Kubota (Coleoptera: Staphylinidae: Pselaphinae) on Chinas Mainland." volume="77">Yin (2023 b</bibRefCitation>
)
</td>
</tr>
<tr id="89EB7E1E170864819FC374E1CE9F5065">
<td id="E28C332C0C863117C4F1DCEC90AC7D1E" colspan="1" rowspan="1">
<taxonomicName id="7AEAE4D5242256199A4AAF0B22076A32" authority="Kubota, 1944" authorityName="Kubota" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="urceus">
<emphasis id="80FF8B95293A73B1CB45F0C146B341CD" italics="true">T. urceus</emphasis>
<bibRefCitation id="BBD35F4907795D18A99E2FE48A3886BD" author="Kubota M" issue="1" journalOrPublisher="Transactions of the Kansai Entomological Society" pagination="6-11" refId="B3" refString="Kubota M (1944) New and little known Pselaphidae from Japan. Transactions of the Kansai Entomological Society 14 (1): 6-11." title="New and little known Pselaphidae from Japan." volume="14" year="1944">Kubota, 1944</bibRefCitation>
</taxonomicName>
</td>
<td id="3EBB7FA322375DADFB625903EFF5D3C4" colspan="1" rowspan="1">Taiwan</td>
<td id="772D1E90D1383A019A213159FB71A61B" colspan="1" rowspan="1">unknown</td>
<td id="B5DE36B9C496FB03768E7EB783191915" colspan="1" rowspan="1">unknown</td>
<td id="46078EB2F090E7346089973DAFEEB7EA" colspan="1" rowspan="1">Holotype ♀ + 2 ♂♂, 3 ♀♀</td>
<td id="CEC6D909611185067813FB8211FC077B" colspan="1" rowspan="1">
<bibRefCitation id="3DAACDF7601FB6321927706864CC6688" author="Kubota M" firstAuthor="Kubota" issue="1" journalOrPublisher="Transactions of the Kansai Entomological Society" pagination="6-11" refId="B3" refString="Kubota M (1944) New and little known Pselaphidae from Japan. Transactions of the Kansai Entomological Society 14 (1): 611." title="New and little known Pselaphidae from Japan." volume="14" year="1944">Kubota (1944)</bibRefCitation>
;
<bibRefCitation id="E0C483E57681232A69BE3C8A349E7747" author="Nomura S" firstAuthor="Nomura" issue="2" journalOrPublisher="Elytra" pagination="435-444" refId="B5" refString="Nomura S (1997) A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan. Elytra 25 (2): 435444." title="A revision of the clavigerine genus Triartiger Kubota (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan and Japan." volume="25" year="1997">Nomura (1997)</bibRefCitation>
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="1AAE983A535E43BA011AA85B4DE14F64">
<tableNote id="099108EAEEF1C97F49D74BB1148CC27F">* Note: FIT = flight intercept trap.</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYNAE" type="etymology">
<paragraph id="0EB3433FE1C579AB0EA5C3ABA55161A3">
<heading id="C73B109616B6F45065E52B185B575FDB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E63CD4D80E14B982549A0662401337DC">The species epithet is dedicated to Yu-Jie Cai (蔡余杰), who collected the single adult and provided it for study. The name is a noun in the genitive case.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,78 @@
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<mods:identifier id="BD1A45F4702CC526C4831316F6BFBBF2" type="DOI">10.3897/dez.72.144365</mods:identifier>
<mods:identifier id="F6FB3609302FAAA04DD74C1131706BF6" type="ZooBank">E82AFF4A-F2BA-48AD-A246-D966CC7EF57D</mods:identifier>
</mods:mods>
<treatment id="825E2BC1B762500C9E407B9A76CE1234" ID-arpha="825E2BC1-B762-500C-9E40-7B9A76CE1234" LSID="urn:lsid:plazi:treatment:825E2BC1B762500C9E407B9A76CE1234" httpUri="http://treatment.plazi.org/id/825E2BC1B762500C9E407B9A76CE1234">
<subSubSection id="BFE27B32228133CA0E85CD16C8A85F90" type="nomenclature">
<paragraph id="0D7751B893DCDF193C9F5527E5AA49CF">
<heading id="1FF27C2178248B2738442BD84E634403" reason="label">Genus</heading>
<taxonomicName id="F147E51CEDCEA2518BBEC1A28ACB01CC" ID-CoL="8W4B8" authority="Kubota, 1944" authorityName="Kubota" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="7D23F857ED1A72ED6EFBE86C3F967CF8" italics="true">Triartiger</emphasis>
<bibRefCitation id="768AD5A5BC85AB7DA42D7E404AB678DA" author="Kubota M" issue="1" journalOrPublisher="Transactions of the Kansai Entomological Society" pagination="6-11" refId="B3" refString="Kubota M (1944) New and little known Pselaphidae from Japan. Transactions of the Kansai Entomological Society 14 (1): 6-11." title="New and little known Pselaphidae from Japan." volume="14" year="1944">Kubota, 1944</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHGAC" type="type species">
<paragraph id="1DF454A3FD8A83C4BFE02A0C218E2B1C">
<heading id="C71358D90A27930EFA1F57CF37329CB6" reason="title">
<typeStatus id="4C7F14F21086D7C40185DA1D6481F485">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="9FCC2E89AB9175276D621DD303760CD7">
<treatmentCitationGroup id="785FF3A5A8DEFD5EC888E1E17E7E1DE9">
<treatmentCitation id="2924919C418EB5524C6DC15B7FE1822B" author="Kubota M" page="11" year="1944">
<taxonomicName id="A58833CB8FC8D1019DC2C3DA0C46F61C" ID-CoL="585NB" authority="Kubota, 1944: 11" authorityName="Kubota" authorityPageNumber="11" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Triartiger" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="urceus">
<emphasis id="453B32E80F889D1FD68B83C6C6079298" italics="true">Triartiger urceus</emphasis>
<bibRefCitation id="93BC0FE1264A956DAB721605E418295F" author="Kubota M" issue="1" journalOrPublisher="Transactions of the Kansai Entomological Society" pagination="6-11" refId="B3" refString="Kubota M (1944) New and little known Pselaphidae from Japan. Transactions of the Kansai Entomological Society 14 (1): 6-11." title="New and little known Pselaphidae from Japan." volume="14" year="1944">Kubota, 1944: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, by original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>