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<mods:title id="3866BF42611449E5A44A5709BE29067E">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
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<mods:affiliation id="32121387DE4253A8ABA7B33F7BF0830B">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<treatment LSID="urn:lsid:zoobank.org:act:3C81BF3C-27C7-476A-889D-F2DD66EE7E55" httpUri="http://treatment.plazi.org/id/D937B665284D52DCB4C9EBA8B74FDF9F" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/3C81BF3C-27C7-476A-889D-F2DD66EE7E55" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aro" status="sp. nov.">Camponotus aro</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<treatment id="D937B665284D52DCB4C9EBA8B74FDF9F" ID-GBIF-Taxon="195190281" LSID="urn:lsid:zoobank.org:act:3C81BF3C-27C7-476A-889D-F2DD66EE7E55" httpUri="http://treatment.plazi.org/id/D937B665284D52DCB4C9EBA8B74FDF9F" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Head in full-face view A C. tapia (CASENT 0217310) B C. aro (CASENT 0489917)." figureDoi="10.3897/zookeys.1098.73223.figure25" httpUri="https://binary.pensoft.net/fig/678084" pageId="0" pageNumber="1">Figs 25B</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Head in full-face view A C. gouldi (CASENT 0121617) B C. aro (CASENT 0489917)." figureDoi="10.3897/zookeys.1098.73223.figure31" httpUri="https://binary.pensoft.net/fig/678090" pageId="0" pageNumber="1">, 31B</figureCitation>
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Camponotus aro A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0489917 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1">, 41</figureCitation>
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<figureCitation id="6F6ECB44418E23AE291E803C01EB8E22" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Head in full-face view A C. tapia (CASENT 0217310) B C. aro (CASENT 0489917)." figureDoi="10.3897/zookeys.1098.73223.figure25" httpUri="https://binary.pensoft.net/fig/678084" pageId="0" pageNumber="1">Figs 25B</figureCitation>
<figureCitation id="88C2706C7C6693E09277AD1ED4838B9E" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Head in full-face view A C. gouldi (CASENT 0121617) B C. aro (CASENT 0489917)." figureDoi="10.3897/zookeys.1098.73223.figure31" httpUri="https://binary.pensoft.net/fig/678090" pageId="0" pageNumber="1">, 31B</figureCitation>
<figureCitation id="54337BE9029983F2B882A48DD3C03FBB" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Camponotus aro A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0489917 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1">, 41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype worker">
<paragraph pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
<subSubSection id="E879D313E7B95E60DA2FF7F8CE279E88" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="7600C56D63150CEB6AB06B2DA27D4721" pageId="0" pageNumber="1">Holotype worker.</paragraph>
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<emphasis id="2C3268961CA02A327689A3AF34A5CEA7" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis bold="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
<emphasis id="99C8C3866EB7B2659A746973E0FF33B8" bold="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
: PN de Namoroka, 17.8 km 329° WNW Vilanandro,
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,
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, 100 m, dry forest, under stone, 08 Nov 2002 (Fisher, Griswold et al.) collection code: BLF06542, specimen code: CASENT0489917 (CAS).
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">2 minor workers and 1 major worker with same data as holotype but with specimen codes: CASENT0837586, CASENT0837587, CASENT0489918 (NHMUK, MHNG, CAS).</paragraph>
<subSubSection id="5DAA3BDC53E1DB217F4E8E49D6424D51" pageId="0" pageNumber="1" type="paratypes">
<paragraph id="2F33D4BE407F454BFB94C2DB016E2B15" pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph id="E3A698B97085A7A904D04614E89BEC6C" pageId="0" pageNumber="1">2 minor workers and 1 major worker with same data as holotype but with specimen codes: CASENT0837586, CASENT0837587, CASENT0489918 (NHMUK, MHNG, CAS).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="PN Namoroka" country="Madagascar" elevation="100" latitude="-16.37667" location="Vilanandro" longLatPrecision="1" longitude="45.32667" specimenCount="1" stateProvince="Mahajanga">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
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<paragraph id="4D45667C4F4549C9F51E0BC13F538BC5" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="C6C59200329E55099229BEC06F3AA502" pageId="0" pageNumber="1">
<materialsCitation id="1A957A9359768BF37F6696C20B417C4C" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="PN Namoroka" country="Madagascar" elevation="100" latitude="-16.37667" location="Vilanandro" longLatPrecision="1" longitude="45.32667" specimenCount="1" stateProvince="Mahajanga">
<emphasis id="6B8690160F783E60075350C05B2108C1" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="C50D2398D1CCFC167AA59F7C1C09A1A1" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion country="Madagascar" name="Mahajanga">Mahajanga</collectingRegion>
<collectingRegion id="5EC47F54DA046917C0122D4524541DB5" country="Madagascar" name="Mahajanga">Mahajanga</collectingRegion>
</emphasis>
:
<collectorName>PN Namoroka</collectorName>
<collectorName id="DC1E16CFCB7861638F5F0E865928BC2B">PN Namoroka</collectorName>
,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.69" unit="km" value="16.9">16.9 km</quantity>
<quantity id="E4EEBE015867AB638E30103B7FE90379" metricMagnitude="4" metricUnit="m" metricValue="1.69" unit="km" value="16.9">16.9 km</quantity>
317° NW
<location LSID="urn:lsid:plazi:treatment:D937B665284D52DCB4C9EBA8B74FDF9F:7408B2A4DD9C8F969C037A607C80A294" country="Madagascar" latitude="-16.37667" longLatPrecision="1" longitude="45.32667" name="Vilanandro" stateProvince="Mahajanga">Vilanandro</location>
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, -16.40667, 45.31,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
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<elevation id="694F33A89A85DC76C0EFBB3AB1FBC6E3" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS);
<collectorName>PN Namoroka</collectorName>
<collectorName id="928B7645B24C74728BCE1BA206BD20FA">PN Namoroka</collectorName>
,
<locationDeviation location="Vilanandro">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.78" unit="km" value="17.8">17.8 km</quantity>
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<quantity id="AE97B7220AF8293C86EBA457B4F431EE" metricMagnitude="4" metricUnit="m" metricValue="1.78" unit="km" value="17.8">17.8 km</quantity>
329° WNW Vilanandro
</locationDeviation>
,
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<geoCoordinate id="B6824ABE32BA50F408354DAB54C3BD60" degrees="16.37667" direction="south" orientation="latitude" precision="1" value="-16.37667">-16.37667</geoCoordinate>
,
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<geoCoordinate id="30946520AB2D3D1741004A89F9A7CC14" degrees="45.32667" direction="east" orientation="longitude" precision="1" value="45.32667">45.32667</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
<quantity id="995FF0EF15E4218368A43A61F39A0788" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="4431379E346C0331271F582353E592C0" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
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, tropical dry forest (Fisher, Griswold et al.) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">With head in full-face view, lateral margins of head anterior to eye level parallel, lacking erect hairs, lateral cephalic margin rounding to posterior margin, anteromedian clypeal margin continuously forming broad convexity; propodeal dorsum immediately posterior to metanotal groove convex, then concave medially and rounding to declivity surface.</paragraph>
<subSubSection id="82DE84A60641C6F209638DCB3D702250" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="EAE8C096B709E52DA62C86C47D736275" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="F92A660DA96DC9B01EBB7E8E40D5C4FF" pageId="0" pageNumber="1">With head in full-face view, lateral margins of head anterior to eye level parallel, lacking erect hairs, lateral cephalic margin rounding to posterior margin, anteromedian clypeal margin continuously forming broad convexity; propodeal dorsum immediately posterior to metanotal groove convex, then concave medially and rounding to declivity surface.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head widest at midlength; lateral margins posterior to level of eye, rounding evenly to posterior margin; eye protruding and large (EL/CS: 0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.26), not breaking lateral cephalic margin; level of its posterior border located at ca. posterior 1/3 of head (PoOc/CL: 0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.27-0.30); frontal carinae posteriorly parallel (FR/CS: 0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.24-0.27); clypeus without anterolateral angle, its anteromedian margin broadly convex; mandible with six teeth, its two apical teeth distantly spaced; antennal scape relatively long (SL/CS: 1.59
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08; 1.46-1.71). Mesosoma long and low (MPH/ML: 0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
<subSubSection id="C3E16366E2EAE4C3CA789F425D758650" pageId="0" pageNumber="1" type="description">
<paragraph id="AE55779CC7925887D705413BB66B2325" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="1917BAF73D2721A78703512A0F19DB8D" pageId="0" pageNumber="1">
<emphasis id="A8BD1107A0C18F5E8B2E1F411531CED5" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head widest at midlength; lateral margins posterior to level of eye, rounding evenly to posterior margin; eye protruding and large (EL/CS: 0.25
<normalizedToken id="93D65ADB971B5B5E6A0341C49F5EE467" originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.26), not breaking lateral cephalic margin; level of its posterior border located at ca. posterior 1/3 of head (PoOc/CL: 0.28
<normalizedToken id="ED11EA3F3BC33B7BA1D58676E89C7A4A" originalValue="±">+/-</normalizedToken>
0.01; 0.27-0.30); frontal carinae posteriorly parallel (FR/CS: 0.25
<normalizedToken id="90984E4FDFEBD080683A9613A53F6F9D" originalValue="±">+/-</normalizedToken>
0.01; 0.24-0.27); clypeus without anterolateral angle, its anteromedian margin broadly convex; mandible with six teeth, its two apical teeth distantly spaced; antennal scape relatively long (SL/CS: 1.59
<normalizedToken id="4AF9468A2328238DDCE03E2B4C29A67D" originalValue="±">+/-</normalizedToken>
0.08; 1.46-1.71). Mesosoma long and low (MPH/ML: 0.28
<normalizedToken id="2A319CAF8F7B4E0EB699B790F147E459" originalValue="±">+/-</normalizedToken>
0.01; 0.26-0.30), promesonotum weakly convex, mesonotum with posterior portion flat immediately anterior to a weakly visible metanotal groove; propodeal dorsum anteriorly convex, with feeble concavity medially, dorsal margin of propodeum rounding to declivity; propodeal dorsum 2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="32BB22120D0A7E76BCF8FF3BC3463683" originalValue="×">x</normalizedToken>
as long as declivity. Petiolar node nodiform; with dorsal margin inclined posteriorly and forming a blunt angle to anterior face; anterior face of petiolar node 1/2 height of posterior face; femur of hind leg rounded axially and twisted basally.
</paragraph>
<paragraph pageId="0" pageNumber="1">First and second gastral tergites with a pair of white spots; erect hairs on lateral margin of head absent; two erect hairs present near posterior margin of head; antennal scape only covered with appressed hairs; pronotum with a pair of erect hairs; posterodorsal angle of propodeum with a pair of erect hairs.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: enlarged head (CS: 3.10
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18; 2.82-3.29; CWb/CL: 0.92
<normalizedToken originalValue="±">+/-</normalizedToken>
<paragraph id="73242D782098FEA48B6925F381D0DE58" pageId="0" pageNumber="1">First and second gastral tergites with a pair of white spots; erect hairs on lateral margin of head absent; two erect hairs present near posterior margin of head; antennal scape only covered with appressed hairs; pronotum with a pair of erect hairs; posterodorsal angle of propodeum with a pair of erect hairs.</paragraph>
<paragraph id="A860D15CF2B1C2844E97D72D4D219F4E" pageId="0" pageNumber="1">
<emphasis id="F9DF2AED707CA1DD3154C70A332AC5F5" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: enlarged head (CS: 3.10
<normalizedToken id="5285B3A9505386FBDF3DF8BF3CD3F279" originalValue="±">+/-</normalizedToken>
0.18; 2.82-3.29; CWb/CL: 0.92
<normalizedToken id="B2F0866A279FDC5BA376C50230B35616" originalValue="±">+/-</normalizedToken>
0.01; 0.90-0.94) with markedly concave posterior margin; apical 1/3 of antennal scape extending beyond posterior cephalic margin; robust mesosoma with propodeal dorsum slightly concave and its length ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B86AC2069BD4E8E969D9805A9B6A6DFC" originalValue="×">x</normalizedToken>
height of declivity; petiole tapering dorsally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="1774CDF256AD5905C4F69B6633B39EC9" pageId="0" pageNumber="1" type="distribution">
<paragraph id="17508F3C47DE1C21815DFFCF4A705C98" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="8309A9E637B72D266D827DB457D33CA2" pageId="0" pageNumber="1">
Geographically restricted to the PN Namoroka in the western dry forest of Madagascar (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Camponotus aro A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0489917 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1">41D</figureCitation>
<figureCitation id="1148F95393B736D183CB80DC1638C424" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Camponotus aro A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0489917 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1">41D</figureCitation>
),
<taxonomicName genus="C." lsidName="C. aro" pageId="0" pageNumber="1" rank="species" species="aro">
<emphasis italics="true" pageId="0" pageNumber="1">C. aro</emphasis>
<taxonomicName id="667F6043ACBBD97A7A253D8C50EA28D5" lsidName="C. aro" pageId="0" pageNumber="1" rank="species" species="aro">
<emphasis id="C8C2BF19D5DD440960840080024EA4EB" italics="true" pageId="0" pageNumber="1">C. aro</emphasis>
</taxonomicName>
has been found nesting under stones while workers forage on the ground or through leaf litter and on lower vegetation.
</paragraph>
<caption doi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aro">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus aro</emphasis>
<caption id="6A02727144843A8B4628F61C8B8DC734" doi="10.3897/zookeys.1098.73223.figure41" httpUri="https://binary.pensoft.net/fig/678100" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
<paragraph id="9A058715CB689C903DB09C80509A8FC8" pageId="0" pageNumber="1">
<emphasis id="3A61D98F87090B51C3FA4DF5BD848D9E" bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
<taxonomicName id="7D858141FB597846B8510B574AFF9148" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aro">
<emphasis id="EEA8DAD45783CFE982CB9400096CAB2B" italics="true" pageId="0" pageNumber="1">Camponotus aro</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="9569332E0D2FB17CA6B70105CD50EF51" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="3A671B28C23ADA192DD8758A4AE1BC97" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="39A042A41C954C501680368F599F0C86" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of holotype minor worker CASENT0489917
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="DE7EC5BD5283A39F01F4A5433341D70F" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aro">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus aro</emphasis>
<subSubSection id="DECF3FDD4E8CB6B1BF3F77022E7E0E1F" pageId="0" pageNumber="1" type="discussion">
<paragraph id="B1AECEFC23A0DF0257EC7FC203C6FD00" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="D3640752E94CD0D04A33FCDD7C005EC2" pageId="0" pageNumber="1">
<taxonomicName id="26F8BF7142C1E7FF7E354A1F189FF138" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aro">
<emphasis id="E965834606C263669D4DD4F9F2C249D5" italics="true" pageId="0" pageNumber="1">Camponotus aro</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName genus="C." lsidName="C. gouldi" pageId="0" pageNumber="1" rank="species" species="gouldi">
<emphasis italics="true" pageId="0" pageNumber="1">C. gouldi</emphasis>
<taxonomicName id="49970FCED9B96CE105B828CDE5500397" lsidName="C. gouldi" pageId="0" pageNumber="1" rank="species" species="gouldi">
<emphasis id="8D5E6A47015F9715BB6C629AEBC0BAFE" italics="true" pageId="0" pageNumber="1">C. gouldi</emphasis>
</taxonomicName>
in that both species are characterized by a broadly convex anteromedian margin of clypeus, but the latter species is diagnosed as having a head that extends posteriorly into a broad, short neck and its propodeal dorsum is straight.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="B9A5A7CF136F1E9835A935D99A02B956" pageId="0" pageNumber="1">
The definition for
<taxonomicName genus="C." lsidName="C. aro" pageId="0" pageNumber="1" rank="species" species="aro">
<emphasis italics="true" pageId="0" pageNumber="1">C. aro</emphasis>
<taxonomicName id="F84A30DD7186105273981638B0631462" lsidName="C. aro" pageId="0" pageNumber="1" rank="species" species="aro">
<emphasis id="18B4D75FC268E556D088E61C2D0F612D" italics="true" pageId="0" pageNumber="1">C. aro</emphasis>
</taxonomicName>
is supported by the congruence between the results of traditional qualitative morphology and the NC-clustering technique. The classification of the samples for the species is at 100% success.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="B8612F454E1BAC67CF63D50C55227F85" pageId="0" pageNumber="1" type="etymology">
<paragraph id="AE7E84B14251E267DF743AE9E02FF5C3" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="F8AFF05CF158FA3EA1D273CB951004D1" pageId="0" pageNumber="1">
The species name
<emphasis italics="true" pageId="0" pageNumber="1">aro</emphasis>
<emphasis id="B8BEF2F984C959D5266BE69F441BE9B1" italics="true" pageId="0" pageNumber="1">aro</emphasis>
is a non-Latin singular noun used in apposition.
</paragraph>
</subSubSection>