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data/8D/1E/40/8D1E409B9912598BA8C27167C8601D77.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" LSID="urn:lsid:zoobank.org:act:1AF0EE48-7F21-4744-961F-86BCEE7A9029" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="8D1E409B9912598BA8C27167C8601D77" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus bemaheva Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="1AF0EE48-7F21-4744-961F-86BCEE7A9029" docUuidSource="ZooBank" docVersion="1" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1651618226429" updateUser="pensoft">
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<mods:title>Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:title id="C24FFA6F781D2D5FD27CCA1DD63266BD">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="CE32A438866D4CFBD27ABBA611ECB35C" type="personal">
<mods:role id="3EED7A5220D9257B2591BE3E7A3E95BC">
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<mods:namePart>Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
<mods:namePart id="02EBAB7C7EB9F72EF0B0791357CEA3FF">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="019223A3964996FF574CC96714398BD1">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier id="FD153D49D3DB1884BABEA8D40800787D" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation>Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
<mods:namePart id="3B7DE2CFC9F0A79A5A9A2DB4A7CDD2E5">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="B8FC6192E50658E5D41CF5FE931058E1">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
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<treatment LSID="urn:lsid:zoobank.org:act:1AF0EE48-7F21-4744-961F-86BCEE7A9029" httpUri="http://treatment.plazi.org/id/8D1E409B9912598BA8C27167C8601D77" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/1AF0EE48-7F21-4744-961F-86BCEE7A9029" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva" status="sp. nov.">Camponotus bemaheva</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<treatment id="8D1E409B9912598BA8C27167C8601D77" ID-GBIF-Taxon="195190289" LSID="urn:lsid:zoobank.org:act:1AF0EE48-7F21-4744-961F-86BCEE7A9029" httpUri="http://treatment.plazi.org/id/8D1E409B9912598BA8C27167C8601D77" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="6ADB3D736E60383EF381A3D80B8C2BD3" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="B3E35EBE50E68399BCD4DDE70665D3C4" pageId="0" pageNumber="1">
<taxonomicName id="F4B07D8419BE8767438E50366608798D" LSID="http://zoobank.org/1AF0EE48-7F21-4744-961F-86BCEE7A9029" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva" status="sp. nov.">Camponotus bemaheva</taxonomicName>
<taxonomicNameLabel id="0C1E0C2D3D0A8F82A5DF74058143CAAC" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mandibles in frontal view A C. niavo (CASENT 0300086) B C. bemaheva (CASENT 0153778)." figureDoi="10.3897/zookeys.1098.73223.figure7" httpUri="https://binary.pensoft.net/fig/678066" pageId="0" pageNumber="1">Figs 7B</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Head in full-face view A C. bemaheva (CASENT 0154158) B C. roeseli (CASENT 0492473)." figureDoi="10.3897/zookeys.1098.73223.figure13" httpUri="https://binary.pensoft.net/fig/678072" pageId="0" pageNumber="1">, 13A</figureCitation>
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Camponotus bemaheva A lateral view B head in full-face view C dorsal view of minor worker CASENT 0154158 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1">, 46</figureCitation>
<subSubSection id="90366FFB02A6C153AD2BB561919EB207" pageId="0" pageNumber="1" type="description">
<paragraph id="AD639907221DE12E5001ACAFBF1182C3" pageId="0" pageNumber="1">
<figureCitation id="CBC84DD006523C910FB7D257FE1752E0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mandibles in frontal view A C. niavo (CASENT 0300086) B C. bemaheva (CASENT 0153778)." figureDoi="10.3897/zookeys.1098.73223.figure7" httpUri="https://binary.pensoft.net/fig/678066" pageId="0" pageNumber="1">Figs 7B</figureCitation>
<figureCitation id="CEBBE37D996750595380CC78DFABC028" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Head in full-face view A C. bemaheva (CASENT 0154158) B C. roeseli (CASENT 0492473)." figureDoi="10.3897/zookeys.1098.73223.figure13" httpUri="https://binary.pensoft.net/fig/678072" pageId="0" pageNumber="1">, 13A</figureCitation>
<figureCitation id="822B02AB79F494F3AB43FCA24C690B11" captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Camponotus bemaheva A lateral view B head in full-face view C dorsal view of minor worker CASENT 0154158 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1">, 46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype worker">
<paragraph pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
<subSubSection id="89F1220C17B050B4AC8FCFE26A792F59" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="5F6356856F530590BA32B243672B4215" pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph id="17C38E47345F7F9702F9EEA840B13D59" pageId="0" pageNumber="1">
<emphasis id="5E17F41338A9C2D71E3D5AF0BC72ED69" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
<emphasis id="A569350FE023C56C60C58009771EB649" bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
: PN Zombitse, 19.8 km 84° E Sakaraha, -22.84333, 44.71, 770 m, tropical dry forest, pitfall trap (Fisher, Griswold et al.) 05-09 Feb 2003, collection code: BLF07507, specimen code: CASENT0050097 (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<subSubSection id="4277782A6D1D4341E9A1D2398BBD70E8" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="114104FF597131F33B399DD9D3D797A3" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="0F7FD6DB1630D7D97DEF736EEFC298B7" pageId="0" pageNumber="1">
<emphasis id="9177BBAD835306F9FAB2262E4A4945C9" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="121CA033F4C21D5D5BA235A8AFB88358" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
<collectingRegion id="73E3DD290B06B193695B04FE2CB4AB4C" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
</emphasis>
: RS Ankarana,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" unit="km" value="13.6">13.6 km</quantity>
<quantity id="55999409601C569C77EC7F3C337FAE8C" metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" unit="km" value="13.6">13.6 km</quantity>
192° SSW Anivorano Nord,
<geoCoordinate degrees="12.86361" direction="south" orientation="latitude" precision="1" value="-12.86361">-12.86361</geoCoordinate>
<geoCoordinate id="97B135D533CA43A20CBC26F6CBE87840" degrees="12.86361" direction="south" orientation="latitude" precision="1" value="-12.86361">-12.86361</geoCoordinate>
,
<geoCoordinate degrees="49.22583" direction="east" orientation="longitude" precision="1" value="49.22583">49.22583</geoCoordinate>
<geoCoordinate id="B0E78E3C53038B6C1FAA21475B232371" degrees="49.22583" direction="east" orientation="longitude" precision="1" value="49.22583">49.22583</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">210 m</quantity>
<quantity id="696C16ACFBCB967F0C4F1C52FA4A8183" metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">210 m</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
<emphasis id="9C56D436035B66B4B565CC998919990C" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="330B77FEF839F6FD0B756AB99FCF26D5" country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
</emphasis>
:
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="d’Analalava">d'Analalava</normalizedToken>
<normalizedToken id="D9565FBB44D9D66F5B30DCC2C24525C5" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="93C27821CCE1679EF85ED21BFFB776FE" originalValue="d’Analalava">d'Analalava</normalizedToken>
,
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.96" unit="km" value="29.6">29.6 km</quantity>
<quantity id="8C88F54D289E5ADB445C102D6602E2CE" metricMagnitude="4" metricUnit="m" metricValue="2.96" unit="km" value="29.6">29.6 km</quantity>
280° W Ranohira,
<geoCoordinate degrees="22.59167" direction="south" orientation="latitude" precision="1" value="-22.59167">-22.59167</geoCoordinate>
<geoCoordinate id="2CF5DE85246F03867DB371BA158FDE2D" degrees="22.59167" direction="south" orientation="latitude" precision="1" value="-22.59167">-22.59167</geoCoordinate>
,
<geoCoordinate degrees="45.12833" direction="east" orientation="longitude" precision="1" value="45.12833">45.12833</geoCoordinate>
<geoCoordinate id="6A2CC22F2EB796CE32367A3D1254295C" degrees="45.12833" direction="east" orientation="longitude" precision="1" value="45.12833">45.12833</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</quantity>
<quantity id="6D07E40499B48AA0480F0A75A729F77D" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS); PN Isalo, Sahanafa River,
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.92" unit="km" value="29.2">29.2 km</quantity>
<quantity id="D8533BC8A87B92B9E55C7E05ACEA778E" metricMagnitude="4" metricUnit="m" metricValue="2.92" unit="km" value="29.2">29.2 km</quantity>
351° N Ranohira,
<geoCoordinate degrees="22.31333" direction="south" orientation="latitude" precision="1" value="-22.31333">-22.31333</geoCoordinate>
<geoCoordinate id="BD932C4625371D17DA5BF8AE57DC2249" degrees="22.31333" direction="south" orientation="latitude" precision="1" value="-22.31333">-22.31333</geoCoordinate>
,
<geoCoordinate degrees="45.29167" direction="east" orientation="longitude" precision="1" value="45.29167">45.29167</geoCoordinate>
<geoCoordinate id="5BFC6F2DBA23E892F1C484059584262F" degrees="45.29167" direction="east" orientation="longitude" precision="1" value="45.29167">45.29167</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
<quantity id="DD43FC0392E0B175CAE10F5B9248CD3B" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
, gallery forest (Fisher, Griswold et al.) (CAS).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Mahajanga">Mahajanga</collectingRegion>
<emphasis id="A6C58D4F6F7213DFFD8AA053ADA44774" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="6B75677C65D07235E283EDA4C032F2D5" country="Madagascar" name="Mahajanga">Mahajanga</collectingRegion>
</emphasis>
: PN Ankarafantsika,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="AC9F640612865E1D4006B0A35A9D998D" originalValue="Forêt">Foret</normalizedToken>
de Tsimaloto,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.83" unit="km" value="18.3">18.3 km</quantity>
<quantity id="6439AC27B47BCF56A93BB06805676EF8" metricMagnitude="4" metricUnit="m" metricValue="1.83" unit="km" value="18.3">18.3 km</quantity>
46° NE de Tsaramandroso,
<geoCoordinate degrees="16.22806" direction="south" orientation="latitude" precision="1" value="-16.22806">-16.22806</geoCoordinate>
<geoCoordinate id="A0174BAF6F3E5EE5ADA83DA1B6EF0D2C" degrees="16.22806" direction="south" orientation="latitude" precision="1" value="-16.22806">-16.22806</geoCoordinate>
,
<geoCoordinate degrees="47.14361" direction="east" orientation="longitude" precision="1" value="47.14361">47.14361</geoCoordinate>
<geoCoordinate id="92B4BD12ADFA17DF2546D57C1C5CD4B6" degrees="47.14361" direction="east" orientation="longitude" precision="1" value="47.14361">47.14361</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">135 m</quantity>
<quantity id="72D17E226DC31C7D0E15A7DF42A29D8E" metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">135 m</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Toliara">Toliara</collectingRegion>
<emphasis id="3A2793E05838FAA9FF4CFAAD43BFF6D9" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="855B63BC292F34A1AF8E074D78CAFAFB" country="Madagascar" name="Toliara">Toliara</collectingRegion>
</emphasis>
:
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.8" unit="km" value="48.0">48 km</quantity>
<quantity id="C5CF7EFC0399903C5553B252084AB9F0" metricMagnitude="4" metricUnit="m" metricValue="4.8" unit="km" value="48.0">48 km</quantity>
ENE Morondava, -20.06667, 44.65,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
<quantity id="494756F700ABD6887DEB23A9DB3F78AF" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
, tropical dry forest (D.M. Olson) (PSWC), Androy Region, District of Tsihombe,
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.4" unit="km" value="74.0">74 km</quantity>
<quantity id="3C3E16F5897F2D57094D6FD00FD5532F" metricMagnitude="4" metricUnit="m" metricValue="7.4" unit="km" value="74.0">74 km</quantity>
S of Tsihombe, RS Cap Ste Marie,
<geoCoordinate degrees="25.58767" direction="south" orientation="latitude" precision="1" value="-25.58767">-25.58767</geoCoordinate>
<geoCoordinate id="47BF7C1CFF117A1EC7AE486A86A9B3E7" degrees="25.58767" direction="south" orientation="latitude" precision="1" value="-25.58767">-25.58767</geoCoordinate>
,
<geoCoordinate degrees="45.163" direction="east" orientation="longitude" precision="55" value="45.163">45.163</geoCoordinate>
<geoCoordinate id="B79FBF3970CBD41A485ACFAC30EEBE58" degrees="45.163" direction="east" orientation="longitude" precision="55" value="45.163">45.163</geoCoordinate>
, 36 m, spiny bush (
<normalizedToken originalValue="Rin’Ha">Rin'Ha</normalizedToken>
<normalizedToken id="FA395E05A393E5AE9C6B06A45FD674D8" originalValue="Rin’Ha">Rin'Ha</normalizedToken>
, Mike) (CAS); Atsimo Andrefana Region, District of Betioky; RS Beza Mahafaly, Parcelle Belle vue, 07 km W of Research Station,
<geoCoordinate degrees="23.68983" direction="south" orientation="latitude" precision="1" value="-23.68983">-23.68983</geoCoordinate>
<geoCoordinate id="39FC92401A957E5B3D0E516624D66C5C" degrees="23.68983" direction="south" orientation="latitude" precision="1" value="-23.68983">-23.68983</geoCoordinate>
,
<geoCoordinate degrees="44.5755" direction="east" orientation="longitude" precision="5" value="44.5755">44.5755</geoCoordinate>
<geoCoordinate id="AC213C7433A4A9E5A0FE41D1922B99C2" degrees="44.5755" direction="east" orientation="longitude" precision="5" value="44.5755">44.5755</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.77" unit="m" value="177.0">177 m</quantity>
<quantity id="7A3ED50EA33232214D6CC9A54D99B8FA" metricMagnitude="2" metricUnit="m" metricValue="1.77" unit="m" value="177.0">177 m</quantity>
, spiny forest, (
<normalizedToken originalValue="Rin’ha">Rin'ha</normalizedToken>
<normalizedToken id="97EE56A69002665FFC6847A4C03FAB32" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS); Atsimo Andrefana Region, District of Betioky,
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
<quantity id="7A90668DA430A60FCAC7977BAFFFADD6" metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
E Betioky, RS Beza Mahafaly (Around Research Station),
<geoCoordinate degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
<geoCoordinate id="9A7B8FC761A71C0B52314D3BD533CE82" degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
,
<geoCoordinate degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
<geoCoordinate id="79E7A888BBEC3F3E6797531DE2CEEEF7" degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
, 165 m, Galery dry deciduous forest (
<normalizedToken originalValue="Rin’Ha">Rin'Ha</normalizedToken>
<normalizedToken id="C8EF540F7B8843006BAAA442A9209693" originalValue="Rin’Ha">Rin'Ha</normalizedToken>
, Mike) (CAS); Atsimo Andrefana Region, District of Sakaraha, PN Zombitse;
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
<quantity id="121A59BEE56ABD95291154C738FFB386" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
E of ANGAP Entrance office,
<geoCoordinate degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
<geoCoordinate id="1ACE98EE7E659E88EF5932E6047DC385" degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
,
<geoCoordinate degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
<geoCoordinate id="0541B8338F6A17C33E49AB4529BD21CB" degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" unit="m" value="838.0">838 m</quantity>
<quantity id="EA1C6E4851ECA67D764218C0062C1850" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" unit="m" value="838.0">838 m</quantity>
, deciduous spiny forest (
<normalizedToken originalValue="Rin’Ha">Rin'Ha</normalizedToken>
<normalizedToken id="78D71E6B77DF3FB475550846007586B6" originalValue="Rin’Ha">Rin'Ha</normalizedToken>
, Mike) (CAS); Atsimo Andrefana Region, District of Sakaraha, PN Zombitse;
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
<quantity id="EB4D2F03A1B58DA0254162109BDF05A8" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
N from ANGAP Entrance office,
<geoCoordinate degrees="22.8405" direction="south" orientation="latitude" precision="5" value="-22.8405">-22.8405</geoCoordinate>
<geoCoordinate id="5AA2D130CA3ACECA7A8615039763E3CE" degrees="22.8405" direction="south" orientation="latitude" precision="5" value="-22.8405">-22.8405</geoCoordinate>
,
<geoCoordinate degrees="44.73117" direction="east" orientation="longitude" precision="1" value="44.73117">44.73117</geoCoordinate>
<geoCoordinate id="2E7938AF7785321B74AB12CB68DA43B3" degrees="44.73117" direction="east" orientation="longitude" precision="1" value="44.73117">44.73117</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.23" unit="m" value="823.0">823 m</quantity>
<quantity id="91FCFC70E44793D1126BF7E2A558F4AE" metricMagnitude="2" metricUnit="m" metricValue="8.23" unit="m" value="823.0">823 m</quantity>
, spiny deciduous forest (
<normalizedToken originalValue="Rin’Ha">Rin'Ha</normalizedToken>
<normalizedToken id="C77B71F852D95FE8DC20D4DF96BD8220" originalValue="Rin’Ha">Rin'Ha</normalizedToken>
, Mike) (CAS); FC Analavelona,
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.92" unit="km" value="29.2">29.2 km</quantity>
<quantity id="3D47CB607E57CFC185A2A8D6BC148053" metricMagnitude="4" metricUnit="m" metricValue="2.92" unit="km" value="29.2">29.2 km</quantity>
343° NNW Mahaboboka, -22.675, 44.19,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</quantity>
<quantity id="10991756E576B0D4C2B470C623B3E753" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</quantity>
, montane rainforest (Fisher, Griswold et al.) (CAS);
<normalizedToken originalValue="Forét">Foret</normalizedToken>
<normalizedToken id="82D15BF49A88F123BC50DD772B3ED59A" originalValue="Forét">Foret</normalizedToken>
de Kirindy,
<geoCoordinate degrees="20.0671" direction="south" orientation="latitude" precision="5" value="-20.0671">-20.0671</geoCoordinate>
<geoCoordinate id="058AE7E37C912FC43430675E2A77A4EE" degrees="20.0671" direction="south" orientation="latitude" precision="5" value="-20.0671">-20.0671</geoCoordinate>
,
<geoCoordinate degrees="44.65723" direction="east" orientation="longitude" precision="1" value="44.65723">44.65723</geoCoordinate>
<geoCoordinate id="4F4E13B990B50F80D4C50416CF4A4E0B" degrees="44.65723" direction="east" orientation="longitude" precision="1" value="44.65723">44.65723</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
<quantity id="EBE40CC233BDD5313A3B164FEB18F25E" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
(H. Wood &amp; J. Miller) (CAS);
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="4EC11BD9D8FFE6889D1FE5A312D7D008" originalValue="Forêt">Foret</normalizedToken>
de Kirindy,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.55" unit="km" value="15.5">15.5 km</quantity>
<quantity id="5453BB1924C6AB8FD646A1858EB95D13" metricMagnitude="4" metricUnit="m" metricValue="1.55" unit="km" value="15.5">15.5 km</quantity>
64° ENE Marofandilia,
<geoCoordinate degrees="20.06915" direction="south" orientation="latitude" precision="1" value="-20.06915">-20.06915</geoCoordinate>
<geoCoordinate id="3B605DED3844456173D209C8680E186D" degrees="20.06915" direction="south" orientation="latitude" precision="1" value="-20.06915">-20.06915</geoCoordinate>
,
<geoCoordinate degrees="44.66042" direction="east" orientation="longitude" precision="1" value="44.66042">44.66042</geoCoordinate>
<geoCoordinate id="417FA9245B29F672F668E6474C33A5B0" degrees="44.66042" direction="east" orientation="longitude" precision="1" value="44.66042">44.66042</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
<quantity id="E47D33672C71077B2F7225630B3652B3" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
, tropical dry forest (B.L. Fisher et al.) (CAS); Makay Mts.,
<geoCoordinate degrees="21.227" direction="south" orientation="latitude" precision="55" value="-21.227">-21.227</geoCoordinate>
<geoCoordinate id="103ACD6D0CBDD725614AF132D9173C7D" degrees="21.227" direction="south" orientation="latitude" precision="55" value="-21.227">-21.227</geoCoordinate>
,
<geoCoordinate degrees="45.33222" direction="east" orientation="longitude" precision="1" value="45.33222">45.33222</geoCoordinate>
<geoCoordinate id="5B65E6E681A5818755DBEACF86EB0909" degrees="45.33222" direction="east" orientation="longitude" precision="1" value="45.33222">45.33222</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.75" unit="m" value="475.0">475 m</quantity>
<quantity id="0032D3EC1F54251CB04035BA4AB709AD" metricMagnitude="2" metricUnit="m" metricValue="4.75" unit="m" value="475.0">475 m</quantity>
, Gallery forest on sandy soil (B.L. Fisher et al.) (CAS); Manombo,
<geoCoordinate degrees="22.80707" direction="south" orientation="latitude" precision="1" value="-22.80707">-22.80707</geoCoordinate>
<geoCoordinate id="30EDBC713FC55F7508138806D5E21F90" degrees="22.80707" direction="south" orientation="latitude" precision="1" value="-22.80707">-22.80707</geoCoordinate>
,
<geoCoordinate degrees="43.76375" direction="east" orientation="longitude" precision="1" value="43.76375">43.76375</geoCoordinate>
<geoCoordinate id="0FA7C8B04B8ACB68AAD7C6D581994971" degrees="43.76375" direction="east" orientation="longitude" precision="1" value="43.76375">43.76375</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" unit="m" value="222.0">222 m</quantity>
<quantity id="1587A5CA53C977BBCDD20BC2FAA80B5C" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" unit="m" value="222.0">222 m</quantity>
, gallery forest, TS1 (Frontier Wilderness Project) (CAS); Menabe Region, District of Morondava, Beroboka village
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
<quantity id="A9B19DE939F3048D8651CB9822C63FAD" metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
NE of Morondava, Antsarongaza dry forest 07,5 Km E of Beroboka,
<geoCoordinate degrees="19.9775" direction="south" orientation="latitude" precision="5" value="-19.9775">-19.9775</geoCoordinate>
<geoCoordinate id="F92FC3B075151880E1471EC97BE057D7" degrees="19.9775" direction="south" orientation="latitude" precision="5" value="-19.9775">-19.9775</geoCoordinate>
,
<geoCoordinate degrees="44.66633" direction="east" orientation="longitude" precision="1" value="44.66633">44.66633</geoCoordinate>
<geoCoordinate id="1D002806EF1FC8E93CDDC1536B4E7100" degrees="44.66633" direction="east" orientation="longitude" precision="1" value="44.66633">44.66633</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
<quantity id="CDC7C6922845F98046B3953949AD6675" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
, dry forest (M. Irwin,
<normalizedToken originalValue="Rin’ha">Rin'ha</normalizedToken>
<normalizedToken id="8037DE98707D9EEF50705F046CE322C5" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS); Menabe Region, District of Morondava, Beroboka village,
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
<quantity id="5C9AD35F8145A60B32A10BCA8E3D42B2" metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
NE of Morondava, Antsarongaza galery forest, 07 km E of Beroboka,
<geoCoordinate degrees="19.9775" direction="south" orientation="latitude" precision="5" value="-19.9775">-19.9775</geoCoordinate>
<geoCoordinate id="8843123F088B57CB31800601CD4F8F9A" degrees="19.9775" direction="south" orientation="latitude" precision="5" value="-19.9775">-19.9775</geoCoordinate>
,
<geoCoordinate degrees="44.66533" direction="east" orientation="longitude" precision="1" value="44.66533">44.66533</geoCoordinate>
<geoCoordinate id="7B1805DF3902A24C31D8CEFE668E35BF" degrees="44.66533" direction="east" orientation="longitude" precision="1" value="44.66533">44.66533</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="4.5" unit="m" value="45.0">45 m</quantity>
<quantity id="9018658766A9FA518AA9C651E0975E04" metricMagnitude="1" metricUnit="m" metricValue="4.5" unit="m" value="45.0">45 m</quantity>
, Galery forest (M. Irwin,
<normalizedToken originalValue="Rin’ha">Rin'ha</normalizedToken>
<normalizedToken id="5DDFEDC31006ADF5C0E46E2E14C280DE" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS); PN Zombitse,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.98" unit="km" value="19.8">19.8 km</quantity>
<geoCoordinate degrees="84" direction="east" orientation="longitude" precision="55555" value="84.0">84° E</geoCoordinate>
<quantity id="11FE52A7E5093704960FAD0D708FF515" metricMagnitude="4" metricUnit="m" metricValue="1.98" unit="km" value="19.8">19.8 km</quantity>
<geoCoordinate id="AA2C3EBA36B1458FED87C001A9FA14FF" degrees="84" direction="east" orientation="longitude" precision="55555" value="84.0">84° E</geoCoordinate>
Sakaraha, -22.84333, 44.71,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</quantity>
<quantity id="A8CA9488E2BC86D906D19F2F7AC04FB6" metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS); Vohibasia Forest,
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.9" unit="km" value="59.0">59 km</quantity>
<quantity id="07312481239ED08D47C7DC94D8A981F7" metricMagnitude="4" metricUnit="m" metricValue="5.9" unit="km" value="59.0">59 km</quantity>
NE Sakaraha, -22.46667, 44.85,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">780 m</quantity>
<quantity id="118CE98662CE2EB485EA97BA1B83557A" metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">780 m</quantity>
(Sylvain) (CAS); near ANGAP office, PN Zombitse,
<geoCoordinate degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
<geoCoordinate id="A13E9759B01C2969D7882A5A5FE82784" degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
,
<geoCoordinate degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
<geoCoordinate id="7171B6C33DD61CA3141EB4F44955DB3B" degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">840 m</quantity>
<quantity id="21EF0BEDDAB820E28F25C42D84B7E4B5" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">840 m</quantity>
, deciduous spiny forest (R.
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
<normalizedToken id="8E5278E2EFBE466DF7BB4DDDE9FA41CB" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
) (CAS); Parcel I, RS Beza Mahafaly, near research station,
<geoCoordinate degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
<geoCoordinate id="63DD5B1CBA7A5BC868DD4C183F6F2FC2" degrees="23.6865" direction="south" orientation="latitude" precision="5" value="-23.6865">-23.6865</geoCoordinate>
,
<geoCoordinate degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
<geoCoordinate id="856C9A740B2064570CB86B239B83E383" degrees="44.591" direction="east" orientation="longitude" precision="55" value="44.591">44.591</geoCoordinate>
, 165 m, dry deciduous forest (R.
<normalizedToken originalValue="Harin’Hala">Harin'Hala</normalizedToken>
<normalizedToken id="F8A4F356E37A7302B4FA001C9C1EFE9C" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
) (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">In full-face view, lateral cephalic margins converge posteriorly towards eye level and without erect hairs posterior to eye level; two apical teeth of mandible positioned close to each other.</paragraph>
<subSubSection id="DDCD6ACDB65293EDCA743106A2A2F6EC" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="AAA20C4A444AC37CB10AD6FFA9E18CAB" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="FD3DA734F1A55B4DD78122DC220377EF" pageId="0" pageNumber="1">In full-face view, lateral cephalic margins converge posteriorly towards eye level and without erect hairs posterior to eye level; two apical teeth of mandible positioned close to each other.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides converging progressively towards posterior margin; eye convex and large (EL/CL: 0.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.29-0.34), breaking lateral cephalic borders, level of its posterior margin generally situated at posterior 1/3 to 1/4 of head (PoOc/CL: 0.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.24-0.28); frontal carinae posteriorly parallel to each other (FR/CS: 0.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 0.25-0.29); clypeus without well-defined anterolateral angle and with broadly convex anteromedian margin; mandible with six teeth, of which the two apical are close together; antennal scape relatively long (SL/CS: 1.63
<normalizedToken originalValue="±">+/-</normalizedToken>
<subSubSection id="AA60CDB54635C28D379B5CF35F682277" pageId="0" pageNumber="1" type="description">
<paragraph id="D10FC47913DD6C4D7175B8346B910FF7" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="E23E2E11F3DFDB923420FA991C8B2C3E" pageId="0" pageNumber="1">
<emphasis id="8C919F7AC85B71631CB4888CD48E0403" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides converging progressively towards posterior margin; eye convex and large (EL/CL: 0.30
<normalizedToken id="418A1D8AD9716006EDDCDD18E2DB5A9B" originalValue="±">+/-</normalizedToken>
0.01; 0.29-0.34), breaking lateral cephalic borders, level of its posterior margin generally situated at posterior 1/3 to 1/4 of head (PoOc/CL: 0.26
<normalizedToken id="5CA5345A89FB97AF6679F7CAEDE26AE6" originalValue="±">+/-</normalizedToken>
0.01; 0.24-0.28); frontal carinae posteriorly parallel to each other (FR/CS: 0.26
<normalizedToken id="5E956DECF4B6384E652780B4E38B7A8F" originalValue="±">+/-</normalizedToken>
0.01 0.25-0.29); clypeus without well-defined anterolateral angle and with broadly convex anteromedian margin; mandible with six teeth, of which the two apical are close together; antennal scape relatively long (SL/CS: 1.63
<normalizedToken id="D9DEDF60C697DA352F14475878663B27" originalValue="±">+/-</normalizedToken>
0.10; 1.47-1.87) with suberect hairs inclined 30° and pubescence. Promesonotum weakly convex, mesopropodeum almost flat, mesonotum flat immediately anterior to metanotal groove; metanotal groove weakly visible; propodeal dorsum anteriorly convex and posteriorly flat; junction of dorsal margin to declivity rounded; propodeal declivity height 1/2 length of propodeal dorsum; petiole nodiform, its dorsal margin rounding to anterior margin; anterior face 1/2 height of posterior face; tibia of hind leg rounded, not twisted basally.
</paragraph>
<paragraph pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; erect hairs covering lateral margin of head anterior to eye level but absent behind eyes; posterior margin of head with more than six erect hairs; antennal scape with suberect hairs inclined at ca. 30° and pubescence; posterodorsal angle of propodeum with a pair of erect hairs.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
<paragraph id="43593D73ACBCF8C4BCBFFC09F724DA88" pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; erect hairs covering lateral margin of head anterior to eye level but absent behind eyes; posterior margin of head with more than six erect hairs; antennal scape with suberect hairs inclined at ca. 30° and pubescence; posterodorsal angle of propodeum with a pair of erect hairs.</paragraph>
<paragraph id="056206928F2EA2F1FA2AF895A49DF069" pageId="0" pageNumber="1">
<emphasis id="5D243A50245D6C5163F33D5DF79BBA42" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
<subSubSection id="5182617F0606B0E168440F06D3F93807" pageId="0" pageNumber="1" type="distribution">
<paragraph id="2320D8CA94280D2991D9F30A516D1839" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="3FF362529FE186E113BB5EB0EF8ED44E" pageId="0" pageNumber="1">
<taxonomicName id="78E6085EE06FDC471536DED888128734" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis id="CBA52F79FE3A5EEDF3BC6BF10C297DFA" italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
</taxonomicName>
is restricted to the spiny bush and thicket of the southern region and the dry forest habitats of western Madagascar (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Camponotus bemaheva A lateral view B head in full-face view C dorsal view of minor worker CASENT 0154158 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1">46D</figureCitation>
<figureCitation id="7C24E79FA71A88D1C47BCFFF57F79DD9" captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Camponotus bemaheva A lateral view B head in full-face view C dorsal view of minor worker CASENT 0154158 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1">46D</figureCitation>
). The few colony nests that have been found were collected from rotten logs and under stones, but workers have been collected while foraging on the ground or on low vegetation.
</paragraph>
<caption doi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
<caption id="B731B71304B2DF1390D48C9457A8B638" doi="10.3897/zookeys.1098.73223.figure46" httpUri="https://binary.pensoft.net/fig/678105" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
<paragraph id="A7689B5D9DFC95A0CC03E1E454E44452" pageId="0" pageNumber="1">
<emphasis id="3B4913DA2402286B9086A064E2A8AD20" bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
<taxonomicName id="7BC59C9A4374D989AE7A4B291B1F8ADC" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis id="F80D8AEB685C1060179A17C6CA9FCE64" italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="A5D4230CE2A15AB8167D6EACE7B41A14" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="B605D8BAEECD233378966F9B35866122" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="CFEA3C287561CCA98E933D3C4D521046" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of minor worker CASENT0154158
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="66A7310ECEDA39D810D3F7CBBAA6F4AD" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="EFE8CFA8CE5602702F2146CDC5F6423A" pageId="0" pageNumber="1" type="discussion">
<paragraph id="83B6F7A55EF2B96216586FF37354936B" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="AC12BF94D21FA45108E2586DA7B67C4B" pageId="0" pageNumber="1">
Workers of
<taxonomicName genus="C." lsidName="C. bemaheva" pageId="0" pageNumber="1" rank="species" species="bemaheva">
<emphasis italics="true" pageId="0" pageNumber="1">C. bemaheva</emphasis>
<taxonomicName id="70B31BD0C940575FC5B45393EB7438DC" lsidName="C. bemaheva" pageId="0" pageNumber="1" rank="species" species="bemaheva">
<emphasis id="26B32AC566E0CC753A0F47C733619AC0" italics="true" pageId="0" pageNumber="1">C. bemaheva</emphasis>
</taxonomicName>
look very similar to those of
<taxonomicName genus="C." lsidName="C. daraina" pageId="0" pageNumber="1" rank="species" species="daraina">
<emphasis italics="true" pageId="0" pageNumber="1">C. daraina</emphasis>
<taxonomicName id="A393F472B104E75CBB34354186BB339D" lsidName="C. daraina" pageId="0" pageNumber="1" rank="species" species="daraina">
<emphasis id="3A2A462B2E2E6B4E4D3AB320CFAB618E" italics="true" pageId="0" pageNumber="1">C. daraina</emphasis>
</taxonomicName>
in that they have closely spaced apical teeth on the mandibles and the lateral margins of their head converge posteriorly towards eye level, but the latter is characterized by the presence of erect hairs on the lateral cephalic margin posterior to eye level.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="A7B785D5BE7D4B11C4CFE58254D2540B" pageId="0" pageNumber="1">
The alignment of
<taxonomicName genus="C." lsidName="C. bemaheva" pageId="0" pageNumber="1" rank="species" species="bemaheva">
<emphasis italics="true" pageId="0" pageNumber="1">C. bemaheva</emphasis>
<taxonomicName id="E08A07353F7EC58904D29A56236C1C4F" lsidName="C. bemaheva" pageId="0" pageNumber="1" rank="species" species="bemaheva">
<emphasis id="9AD0BFC90AB48FFE48FF558B84FCD79A" italics="true" pageId="0" pageNumber="1">C. bemaheva</emphasis>
</taxonomicName>
in the same cluster displayed by the dendrogram of multivariate morphometric method is confirmed by the cumulative LDA at 100% identification success, validating the taxonomy hypothesized by qualitative morphology.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="3A9B15D1F7559963B1797524053BFE50" pageId="0" pageNumber="1" type="etymology">
<paragraph id="D5E1614A3407B84DFE892C94B54A9E4B" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="15FE1AF754442D74A3BF9785804CDB00" pageId="0" pageNumber="1">
The new species
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
<taxonomicName id="790499B9356C59EC459D6A53A875937B" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus bemaheva" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemaheva">
<emphasis id="90DF7544D45F797F64E828765262F884" italics="true" pageId="0" pageNumber="1">Camponotus bemaheva</emphasis>
</taxonomicName>
is a non-Latin proper noun used in apposition and is named in honor of Fidelis Bemaheva for his help collecting ants in the Malagasy region.
</paragraph>