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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.103943" ID-Pensoft-Pub="1313-2970-1184-361" ID-Pensoft-UUID="98C058ACEB7C5834AB8CCD60BF77DF1E" ID-ZooBank="CD7838BA079A4A77A9999CAB5A10F879" ModsDocID="1313-2970-1184-361" checkinTime="1700831865335" checkinUser="pensoft" docAuthor="Huang, Xiu-Dong, Yang, Lin, Chen, Xiang-Sheng &amp; Long, Jian-Kun" docDate="2023" docId="9FD0136EEADD55FDBA5FC6C34C7D6087" docLanguage="en" docName="ZooKeys 1184: 361-375" docOrigin="ZooKeys 1184" docPubDate="2023-11-23" docSource="http://dx.doi.org/10.3897/zookeys.1184.103943" docTitle="Usana Distant 1906" docType="treatment" docVersion="1" id="98C058ACEB7C5834AB8CCD60BF77DF1E" lastPageNumber="361" masterDocId="98C058ACEB7C5834AB8CCD60BF77DF1E" masterDocTitle="Two new species of the planthopper genus Usana Distant, 1906 (Hemiptera, Fulgoromorpha, Achilidae) from China" masterLastPageNumber="375" masterPageNumber="361" pageNumber="361" updateTime="1700831865335" updateUser="pensoft">
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<mods:namePart id="6CBE17D079BD61292D8CE8C09C9498F3">Huang, Xiu-Dong</mods:namePart>
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<mods:affiliation id="9E097EDE6D68F8316A5C2CD7312B4698">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, China &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, Guizhou 550025, China</mods:affiliation>
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<taxonomicName id="428E36C6AC6C08C9E147FD6253125501" ID-CoL="647JD" ID-ENA="1518960" LSID="9FD0136E-EADD-55FD-BA5F-C6C34C7D6087" authority="Distant, 1906" authorityName="Distant" authorityYear="1906" class="Insecta" family="Achilidae" genus="Usana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Usana" order="Hemiptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="genus">Usana Distant, 1906</taxonomicName>
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<subSubSection id="83E75A59F44C49F082939A0E54DB883C" pageId="0" pageNumber="361" type="reference_group">
<paragraph id="B6451E6E891B677815E15855B219165B" pageId="0" pageNumber="361">
<taxonomicName id="42747A6009B70BE8A670DEB78B3C7196" authorityName="Distant" authorityYear="1906" class="Insecta" family="Achilidae" genus="Usana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Usana" order="Hemiptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="genus">Usana</taxonomicName>
Distant, 1906: 293;
<bibRefCitation id="4091020C8C98F400D5E0A14C7B1B726B" DOI="https://doi.org/10.5962/bhl.part.27228" author="Fennah, RG" journalOrPublisher="Entomology" pageId="0" pageNumber="361" pagination="1 - 170" refId="B8" refString="Fennah, RG, 1950. A generic revision of the Achilidae (Homopter, Fulgoroidea). Bulletin of the British Museum (Natural History). Entomology 1: 1 - 170, DOI: https://doi.org/10.5962/bhl.part.27228" title="A generic revision of the Achilidae (Homopter, Fulgoroidea). Bulletin of the British Museum (Natural History)." url="https://doi.org/10.5962/bhl.part.27228" volume="1" year="1950">Fennah 1950</bibRefCitation>
: 132;
<bibRefCitation id="4C76144C9BD369C423628DF58EA622BB" author="Fennah, RG" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="361" pagination="249 - 251" refId="B9" refString="Fennah, RG, 1978. Fulgoroidea (Homoptera) from Viet-nam. Annales Zoologici 34 (9): 249 - 251" title="Fulgoroidea (Homoptera) from Viet-nam." volume="34" year="1978">Fennah 1978</bibRefCitation>
: 249;
<bibRefCitation id="ADD8CD0C67C781E66B8FE1A1CBAECD71" author="Chen, CL" journalOrPublisher="Taiwan Museum Special Publication Series" pageId="0" pageNumber="361" pagination="23 - 26" refId="B5" refString="Chen, CL, Yang, CT, Wilson, MR, 1989. Achilidae of Taiwan (Homoptera: Fulgoroidea). Taiwan Museum Special Publication Series 8: 23 - 26" title="Achilidae of Taiwan (Homoptera: Fulgoroidea)." volume="8" year="1989">Chen et al. 1989</bibRefCitation>
: 23.
</paragraph>
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<subSubSection id="C8639FF6B98F99B2626DF168AC930DEB" pageId="0" pageNumber="361" type="type species">
<paragraph id="20C757E8643CA702D30F39526F39131C" pageId="0" pageNumber="361">Type species.</paragraph>
<paragraph id="04049A732E2C8E3D46AD9F1F9DB3585F" pageId="0" pageNumber="361">
<taxonomicName id="EFF25CCFEAA0858E2590074D14FF796C" authorityName="Distant" authorityYear="1906" class="Insecta" family="Achilidae" genus="Usana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Usana lineolalis" order="Hemiptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="lineolalis">
<emphasis id="D81B8AB6C485F055A53CCBE31BE5E734" italics="true" pageId="0" pageNumber="361">Usana lineolalis</emphasis>
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Distant, 1906, by original designation.
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<paragraph id="827CA49BAF38262090ACD05DE45606D8" pageId="0" pageNumber="361">Diagnosis.</paragraph>
<paragraph id="6FCAF9D7B1F9D0C079A24A622DD79DAE" pageId="0" pageNumber="361">
<emphasis id="1E3E281366A99AEAC2334DB5841BED13" bold="true" pageId="0" pageNumber="361">
<emphasis id="2391E744DF9AC20AFA0E188342F2209C" italics="true" pageId="0" pageNumber="361">Head</emphasis>
.
</emphasis>
Width of head at eyes 0.8-0.9 times wider than pronotum. Vertex not declivous, broader at base than long in middle line, median carina distinct, anterior margin carinate, triangular areolets at lateroapical angles of head distinct, posterior margin broadly concave. Frons longer at mid-line than at widest part, basal margin truncate, median carina distinct, lateral margins carinate. Rostrum almost or just reaching post-trochanter, with subapical segment shorter than apex.
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<emphasis id="68E7D16F48B489246BB21597FFB2053E" italics="true" pageId="0" pageNumber="361">Thorax</emphasis>
.
</emphasis>
Pronotum lateral lobe not inclined anteroventrally, with a small longitudinal carina between eye and tegula. Mesonotum length at midline longer than vertex and pronotum combined. Fore-wing longer than widest part by ~2.7-3.3: 1; vein ScP+R with fork distally or as level of CuA fork; vein MP with fork clearly after CuA fork, with 3 terminals. Hind wing with MP with 3 terminals (MP1, MP2 and MP3+4), vein CuA with 2 terminals. Post-tibiae with a lateral spine between basal 1/3 to near basal 1/2, spinal formula 7-7 (6)-7 (6).
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<emphasis id="8A602C1BE590A8F69B54132B6AA011E7" bold="true" pageId="0" pageNumber="361">
<emphasis id="0A72BC86FF82C4A14E0D51A533504031" italics="true" pageId="0" pageNumber="361">Male terminalia</emphasis>
.
</emphasis>
Length of anal segment in dorsal view at least equal to its width; apical margin of anal segment distinctly excavated at midline; pygofer in lateral view with dorsal margin distinctly shorter than ventral margin, medioventral process entire or with apex divided into 2 branches. Genital style with 3 processes arising from its dorsal margin, inner surface near anterior margin with a long, outwardly directed process. Phallobase sheathed, generally asymmetrical, with apical 1/2 divided into a dorsal, 2 lateral, and a ventral lobe: dorsal lobe relatively short, lateral lobes valviform, and ventral lobe with apical margin incised at midline and with subapical surface in middle giving rise to a long process, directed basad. Each phallic appendage generally not exceeding apical margin of phallobase, with a protrusion between basal 1/4 to 1/3.
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<paragraph id="9FC91DDF44EF27E93C6EFC58177A3868" pageId="0" pageNumber="361">Distribution.</paragraph>
<paragraph id="5D2BEC90EB6C455F3A2651D5BA6CB1DE" pageId="0" pageNumber="361">Palaearctic, Oriental, and Sino-Japanese realms.</paragraph>
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Schenck, 1853
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<paragraph id="B17188163A7CCCD19007DAF19D22CAFD" pageId="0" pageNumber="8050">England, Wales</paragraph>
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<mods:title id="089B683FD4707EF6A118232D3B826714">Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</mods:title>
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<taxonomicName id="D72E6BD725CD9E5A536A272695781733" ID-CoL="6XPHW" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Hypericaceae" genus="Sarothra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="gentianoides" status="spec. nov.">Sarothra gentianoides</taxonomicName>
,
<taxonomicNameLabel id="9A8F290266CA1219FA9D63DA9AD9FDE6" rank="species">spec. nov.</taxonomicNameLabel>
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1. Sarothra.
<emphasis id="B721510872F73CB352D06A05B91E8012" italics="true">Gen. nov. 1075.</emphasis>
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<paragraph id="066C94C46489FA4461BC6682CE33A18A" pageNumber="272">
Gentiana caule ramisque ramosissimis, foliis subulatis minimis.
<emphasis id="69A2BB6CBDBBB5D21CE704626D9C0EB4" italics="true">Gron. virg.29.</emphasis>
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Centaurium minus spicatum, angustissimo folio, s. scoparium marilandicum novum.
<emphasis id="9C0A00238B0E5AC559428B359A1A209F" italics="true">Pluk. mant. 43. t.342. f.4.</emphasis>
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<paragraph id="1468E8654CC5EE6EF618E2F8C805CF51" pageNumber="272">
<emphasis id="439A69927D3BE89A4EF73E7A07927D4A" italics="true">Habitat in</emphasis>
Virginiae
<emphasis id="EB4C90F5AF7E7EEDFF5D77DCF5DB06F6" italics="true">,</emphasis>
Pensylvaniae
<emphasis id="9646B7B38304297FBD19B9AD93124EB0" italics="true">apricis glareosis.</emphasis>
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<paragraph id="CD5E2AB1A8852E7AE14B36889EE0A286" pageNumber="273">
<heading id="BB7C76CA4C7A5715B4BC45411B0E79BD" level="4">
<pageBreakToken id="B7CB8620E5C9688871681BCDCF898572" pageNumber="273">TETRAGYNIA</pageBreakToken>
.
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<mods:namePart>Caldara, Roberto</mods:namePart>
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<mods:namePart>Tosevski, Ivo</mods:namePart>
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<mods:namePart>Letowski, Jacek</mods:namePart>
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<taxonomicName authority="H. Brisout, 1863" authorityName="H. Brisout" authorityYear="1863" class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis (H. Brisout, 1863)</taxonomicName>
Figures 41, 42, 43-44, 45-46, 47, 48-50
</paragraph>
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<subSubSection pageId="21" pageNumber="44" type="materials_examined">
<paragraph pageId="21" pageNumber="44">Material examined.</paragraph>
<paragraph pageId="21" pageNumber="44">
10 L3 larvae: south-eastern France, Castellar (Menton), Juin 2005, ex seed capsules of
<taxonomicName class="Magnoliopsida" family="Campanulaceae" genus="Campanula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Campanula rapunculus" order="Asterales" pageId="21" pageNumber="44" phylum="Tracheophyta" rank="species" species="rapunculus">Campanula rapunculus</taxonomicName>
L., leg. and det. R. Caldara all collected in association with adults.
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="47" pageId="21" pageNumber="44" type="description">
<paragraph pageId="21" pageNumber="44">Description.</paragraph>
<paragraph pageId="21" pageNumber="44">
Measurements (in mm). Body length: 2.20-3.15 (mean 2.8). Body width (metathorax or abdominal segments
<normalizedToken originalValue="I–II">I-II</normalizedToken>
) up to 0.73. Head width: 0.35-0.51 (mean 0.45).
</paragraph>
<paragraph pageId="21" pageNumber="44">General. Body elongated, slender, curved, rounded in cross section (Fig. 41).</paragraph>
<caption pageId="21" pageNumber="44">
<paragraph pageId="21" pageNumber="44">
Figure 41.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
mature larva habitus.
</paragraph>
</caption>
<paragraph pageId="21" pageNumber="44">Colouration. Pale brown or almost yellow head (Fig. 41). All thoracic and abdominal segments from distinctly white to slightly yellow (Fig. 41).</paragraph>
<paragraph pageId="21" pageNumber="44">Vestiture. Setae on body thin, slightly from orange to pale brown, distinctly different in length (minute to very short or long to very long). Cuticle distinctly asperate.</paragraph>
<paragraph pageId="21" pageNumber="44">Head capsule (Fig. 42). Head suboval, distinctly flattened laterally. Frontal sutures narrow, but distinct. Two stemmata (st), anterior one in the form of a small pigmented spot; and posterior one in form of a very small pigmented spot, located on each side close des5. Des1-3 and des5 very long; des4 relatively long (Fig. 42). Fs1 long to very long; fs2 absent; fs3 long medium, laterally to fs4; fs4 very long; and fs5 very long (Fig. 42). Les1 and les2 as long as des5; both ves medium size. Epicranial area with two sensilla and three pes in line with des2.</paragraph>
<caption pageId="21" pageNumber="44">
<paragraph pageId="21" pageNumber="44">
Figure 42.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
mature larva head, frontal view. Abbreviations: des - dorsal epicranial s., fs - frontal epicranial s., les - lateral epicranial s., pes - postepicranial s., ves - ventral s., at - antenna, st - stemmata.
</paragraph>
</caption>
<paragraph pageId="21" pageNumber="44">Antennae bearing one very long conical sensorium, and basal membranous article with three sensilla different in length, two behind conical sensorium, and one ahead of it (Fig. 43).</paragraph>
<caption pageId="21" pageNumber="44">
<paragraph pageId="21" pageNumber="44">
Figures 43-44.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
mature larva. 43 Antenna 44 Right mandible. Abbreviation: mds - mandible dorsal s.
</paragraph>
</caption>
<paragraph pageId="21" pageNumber="44">Clypeus (Fig. 45) approximately three times as wide as long with two medium size cls, cls1 distinctly longer than cls2, and one sensillum; anterior margin sinuate.</paragraph>
<caption pageId="21" pageNumber="44">
<paragraph pageId="21" pageNumber="44">
Figures 45-46.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
mature larva, mouthparts. 45 Labrum and clypeus 46 Epipharynx. Abbreviations: als - anteriolateral s., ams - anteromedial s., cls - clypeal s., lms - labral s., mes - median s., clss - clypeal sensillum, lr - labral rods.
</paragraph>
</caption>
<paragraph lastPageId="22" lastPageNumber="45" pageId="21" pageNumber="44">
Mouthparts. Labrum (Fig. 45) less than 2.5 times as wide as long, with three piliform lms, different in the length; lms1 located posteromedially, very close to margin of clypeus, lms2 located in the middle, and lms3 located laterally; lms1 very long and reaches distinctly the labrum margin, lms2 long, and lms3 medium size, more than twice times as short as lms1. Epipharynx (Fig. 46) with three medium sized finger-like als, two als of identical in length, and the third one distinctly shorter and also located close to labral rods (lr); with three short ams in different shape, ams1 and ams2 piliform, finger-like ams3 and enlarged in middle, and also located more close to lr; without mes; labral rods (lr) distinct, elongated, oval. Mandibles (Fig. 44) bifid; bearing with two setae in short to medium size, piliform, and aligned longitudinally., mds1 located basally; mds2, located distinctly apically. Maxilla (Fig. 47) stipes with very long stps and pfs2, medium size pfs1, very short to minute mbs, and sensillum close to mbs; mala with six medium sized finger-like dms; five vms, different in length, four setae medium size, and one seta very short. Maxillary palpi: basal palpomere with one short mxps and two sensilla; distal palpomere with short, cuticular apical processes; length ratio of basal and distal palpomeres 1:0.8. Prelabium (Fig. 47) with one very short prms; ligula with two very short to minute ligs; premental sclerite broad, ring-shaped. Labial palpi with two palpomeres; length ratio of basal and distal palpomeres 1:0.9;
<pageBreakToken pageId="22" pageNumber="45" start="start">each</pageBreakToken>
of the palpomeres with one sensillum, distal palpomere with short, cuticular apical processes. Postlabium (Fig. 47) with short pms1 located basally, long pms2 located medially and short pms3 located apically; membranous area basolaterally only a partly and finely asperate.
</paragraph>
<caption pageId="22" pageNumber="45">
<paragraph pageId="22" pageNumber="45">
Figure 47.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="22" pageNumber="45" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
larval mouthparts, maxillolabial complex, ventral view right maxilla. Abbreviations: dms - dorsal malar s., vms - ventral malar s., mpxs - maxillary palps s., mbs - basioventral s., pfs - palpiferal s., stps - stipital s., prms - premental s., pms - postmental s., ligs - ligular s.
</paragraph>
</caption>
<paragraph lastPageId="23" lastPageNumber="46" pageId="22" pageNumber="45">
Thorax. Prothorax (Fig. 48) with nine very long and one very short to minute prns, small pigmented dorsal sclerite present with four long prns, this sclerite subdivided in two triangular plates medially; two very long to long ps; and one short eus. Meso- and metathorax (Fig. 48) with one long prs, three very long to long pds; one
<pageBreakToken pageId="23" pageNumber="46" start="start">long</pageBreakToken>
as; two very long and one very short to minute ss; one long eps; one very long to long ps; and one short to very short eus. Each pedal area of the thoracic segments with 5-6 very long pda.
</paragraph>
<caption pageId="23" pageNumber="46">
<paragraph pageId="23" pageNumber="46">
Figures 48-50.
<taxonomicName class="Insecta" family="Curculionidae" genus="Cleopomiarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleopomiarus meridionalis" order="Coleoptera" pageId="23" pageNumber="46" phylum="Arthropoda" rank="species" species="meridionalis">Cleopomiarus meridionalis</taxonomicName>
mature larva, habitus. 48 Lateral view of thoracic segments 49 Lateral view of abdominal segment I 50 Lateral view of abdominal segments
<normalizedToken originalValue="VI–X">VI-X</normalizedToken>
. Abbreviations: as - alar s., ds - dorsal s., eps - epipleural s., eus - eusternal s., lsts - laterosternal s., pda - pedal s., pds - postdorsal s., prns - pronotal s., prs - prodorsal s., ss - spiracular s., ps - pleural s., sts - sternal s., ts - terminal s., Th1-3 - number of thoracic segments, Ab1-10 - number of abdominal seg.
</paragraph>
</caption>
<paragraph lastPageId="24" lastPageNumber="47" pageId="23" pageNumber="46">
Abdomen. Abdominal segments
<normalizedToken originalValue="I–VII">I-VII</normalizedToken>
(Figs 49-50) with one long prs; two relatively long to short and one very long to long pds (order: relatively long, very long, short); one very long to long and one minute ss; two very long eps; one very long to long ps; one relatively long to short lsts; and one short to very short and one relatively long eus. Abdominal segment VIII (Fig. 50) with sometimes one very short to minute
<pageBreakToken pageId="24" pageNumber="47" start="start">prs</pageBreakToken>
; one short and one long to relatively long pds (order: short, long); one long and one minute ss; two very long eps; one very long to long ps; one relatively long to short lsts; and one short to very short and one relatively long eus. Abdominal segment IX (Fig. 50) with two relatively long and two short to very short ds; one relatively long and one minute ps; and one relatively long sts. Abdominal segment X (Fig. 50) with one very short seta (ts).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="47" type="biology_ecology">
<paragraph pageId="24" pageNumber="47">Biology.</paragraph>
<paragraph pageId="24" pageNumber="47">
Adults of this species are usually collected on the flowers of
<taxonomicName class="Magnoliopsida" family="Campanulaceae" genus="Campanula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Campanula rapunculus" order="Asterales" pageId="24" pageNumber="47" phylum="Tracheophyta" rank="species" species="rapunculus">Campanula rapunculus</taxonomicName>
L., and we can confirm that larvae feed on seeds of this plant as previously reported by
<bibRefCitation pageId="24" pageNumber="47">Hoffmann (1958)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="48" type="remarks">
<paragraph pageId="25" pageNumber="48">
<pageBreakToken pageId="25" pageNumber="48" start="start">Remarks</pageBreakToken>
and comparative notes.
</paragraph>
<paragraph pageId="25" pageNumber="48">
This species is widely distributed and common in southern Europe, whereas it appears rare in North Africa and the Middle East. Adults can be confused with some related species such as
<taxonomicName lsidName="C. plantarum" pageId="25" pageNumber="48" rank="species" species="plantarum">C. plantarum</taxonomicName>
(Germar, 1823),
<taxonomicName lsidName="C. micros" pageId="25" pageNumber="48" rank="species" species="micros">C. micros</taxonomicName>
(Germar, 1821) and
<taxonomicName lsidName="C. reitteri" pageId="25" pageNumber="48" rank="species" species="reitteri">C. reitteri</taxonomicName>
(Caldara &amp; Legalov, 2016), from which they differ by some external characters and the shape of their genitalia (
<bibRefCitation author="Caldara, R" journalOrPublisher="Journal of Insect Biodiversity" pageId="54" pageNumber="77" pagination="1 - 47" title="Systematics of the Holarctic species of the weevil genus Cleopomiarus Pierce (Coleoptera: Curculionidae)." url="https://doi.org/10.12976/jib/2016.4.6" volume="4" year="2016">Caldara and Legalov 2016</bibRefCitation>
). In contrast, this species is poorly related morphologically to the other species of
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studied here. This difference is confirmed also by the larval morphology, which differs from all of the other species mainly by a longer fs3 that is almost as long as fs4.
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Genus
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Clairville, 1806: 46. Type species:
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