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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.666.9191" ID-PMC="PMC5534527" ID-Pensoft-Pub="1313-2970-666-1" ID-Pensoft-UUID="C43EEB41A94B930FFE439D1FAD29FF9C" ID-PubMed="28769631" ID-Zenodo-Dep="579453" ID-ZooBank="B3C377E8BBB14F328AECA2C22D1E625A" ModsDocID="1313-2970-666-1" checkinTime="1491551283800" checkinUser="pensoft" docAuthor="Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. &amp; Seidel, Matthias" docDate="2017" docId="9F716EDA36CEA8316521C9C12BC7B1E3" docLanguage="en" docName="ZooKeys 666: 1-349" docOrigin="ZooKeys 666" docPubDate="2017-04-06" docSource="http://dx.doi.org/10.3897/zookeys.666.9191" docTitle="Chrysina chrysopedila" docType="treatment" docVersion="6" id="C43EEB41A94B930FFE439D1FAD29FF9C" lastPageNumber="1" masterDocId="C43EEB41A94B930FFE439D1FAD29FF9C" masterDocTitle="Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies" masterLastPageNumber="349" masterPageNumber="1" pageNumber="1" updateTime="1668164364999" updateUser="ExternalLinkService">
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<mods:title>Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jameson, Mary L.</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
<mods:nameIdentifier type="email">maryliz.jameson@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garner, Beulah H.</mods:namePart>
<mods:affiliation>Natural History Museum, Insects Division, Department of Life Sciences, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Audibert, Cedric</mods:namePart>
<mods:affiliation>Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew B. T.</mods:namePart>
<mods:affiliation>Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
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<treatment ID-GBIF-Taxon="128344536" LSID="urn:lsid:plazi:treatment:9F716EDA36CEA8316521C9C12BC7B1E3" httpUri="http://treatment.plazi.org/id/9F716EDA36CEA8316521C9C12BC7B1E3" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="9F716EDA-36CE-A831-6521-C9C12BC7B1E3" authority="(H. W. Bates, 1888)" baseAuthorityName="H. W. Bates" baseAuthorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysina chrysopedila" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chrysopedila">Chrysina chrysopedila (H. W. Bates, 1888)</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Plusiotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusiotis aurora subsp. var. var. chrysopedila" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="aurora" subSpecies="var." variety="chrysopedila">Plusiotis aurora var. chrysopedila</taxonomicName>
H. W. Bates, 1888: 277 [original combination].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bates" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Plusiotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusiotis chrysopedila" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chrysopedila">Plusiotis chrysopedila</taxonomicName>
H.W. Bates [new species status by
<bibRefCitation author="Ohaus, F" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="1" pagination="273 - 319" refId="B273" refString="Ohaus, F, 1912. Beitraege zur Kenntnis der Ruteliden, X. Stettiner Entomologische Zeitung 73: 273 - 319" title="Beitraege zur Kenntnis der Ruteliden, X." volume="73" year="1912">Ohaus 1912</bibRefCitation>
: 307].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="H. W. Bates" baseAuthorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysina chrysopedila" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chrysopedila">Chrysina chrysopedila</taxonomicName>
(H. W. Bates) [new combination by
<bibRefCitation author="Hawks, DC" journalOrPublisher="Occasional Papers of the Consortium Coleopterorum" pageId="0" pageNumber="1" pagination="1 - 8" refId="B116" refString="Hawks, DC, 2001. Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 4: 1 - 8" title="Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae)." volume="4" year="2001">Hawks 2001</bibRefCitation>
: 4].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
COSTA RICA (
<bibRefCitation author="Moron, MA" journalOrPublisher="Sciences Nat, Venette" pageId="0" pageNumber="1" refId="B233" refString="Moron, MA, 1990. The Beetles of the World. Volume 10: Rutelini 1. Sciences Nat, Venette" title="The Beetles of the World. Volume 10: Rutelini 1." year="1990">
<normalizedToken originalValue="Morón">Moron</normalizedToken>
1990
</bibRefCitation>
, Thomas et al. 2006). NICARAGUA: Chontales (H. W. Bates 1888,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.27703" author="Nonfried, AF" journalOrPublisher="Wiener Entomologische Zeitung" pageId="0" pageNumber="1" pagination="127 - 130" refId="B259" refString="Nonfried, AF, 1892. Beitrag zu einer Monographie der Gattung Plusiotis Burm. Wiener Entomologische Zeitung 11: 127 - 130, DOI: https://doi.org/10.5962/bhl.part.27703" title="Beitrag zu einer Monographie der Gattung Plusiotis Burm." url="https://doi.org/10.5962/bhl.part.27703" volume="11" year="1892">Nonfried 1892</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="1" pagination="1 - 241" refId="B278" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">Ohaus 1918</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
,
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="189 - 265" refId="B14" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185: 189 - 265" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
,
<bibRefCitation author="Maes, J-M" journalOrPublisher="Revista Nicaragueense de Entomologia" pageId="0" pageNumber="1" pagination="27 - 60" refId="B192" refString="Maes, J-M, 1987. Catalogo de los Scarabaeidae (Coleoptera) de Nicaragua. Revista Nicaragueense de Entomologia 1: 27 - 60" title="Catalogo de los Scarabaeidae (Coleoptera) de Nicaragua." volume="1" year="1987">Maes 1987</bibRefCitation>
,
<bibRefCitation author="Moron, MA" journalOrPublisher="Sciences Nat, Venette" pageId="0" pageNumber="1" refId="B233" refString="Moron, MA, 1990. The Beetles of the World. Volume 10: Rutelini 1. Sciences Nat, Venette" title="The Beetles of the World. Volume 10: Rutelini 1." year="1990">
<normalizedToken originalValue="Morón">Moron</normalizedToken>
1990
</bibRefCitation>
, Thomas et al. 2006). PANAMA:
<normalizedToken originalValue="Chiriquí">Chiriqui</normalizedToken>
(H. W. Bates 1888,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.27703" author="Nonfried, AF" journalOrPublisher="Wiener Entomologische Zeitung" pageId="0" pageNumber="1" pagination="127 - 130" refId="B259" refString="Nonfried, AF, 1892. Beitrag zu einer Monographie der Gattung Plusiotis Burm. Wiener Entomologische Zeitung 11: 127 - 130, DOI: https://doi.org/10.5962/bhl.part.27703" title="Beitrag zu einer Monographie der Gattung Plusiotis Burm." url="https://doi.org/10.5962/bhl.part.27703" volume="11" year="1892">Nonfried 1892</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="1" pagination="1 - 241" refId="B278" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">Ohaus 1918</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
,
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="189 - 265" refId="B14" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185: 189 - 265" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
,
<bibRefCitation author="Maes, J-M" journalOrPublisher="Revista Nicaragueense de Entomologia" pageId="0" pageNumber="1" pagination="27 - 60" refId="B192" refString="Maes, J-M, 1987. Catalogo de los Scarabaeidae (Coleoptera) de Nicaragua. Revista Nicaragueense de Entomologia 1: 27 - 60" title="Catalogo de los Scarabaeidae (Coleoptera) de Nicaragua." volume="1" year="1987">Maes 1987</bibRefCitation>
,
<bibRefCitation author="Moron, MA" journalOrPublisher="Sciences Nat, Venette" pageId="0" pageNumber="1" refId="B233" refString="Moron, MA, 1990. The Beetles of the World. Volume 10: Rutelini 1. Sciences Nat, Venette" title="The Beetles of the World. Volume 10: Rutelini 1." year="1990">
<normalizedToken originalValue="Morón">Moron</normalizedToken>
1990
</bibRefCitation>
,
<bibRefCitation author="Hawks, DC" journalOrPublisher="Occasional Papers of the Consortium Coleopterorum" pageId="0" pageNumber="1" pagination="1 - 8" refId="B116" refString="Hawks, DC, 2001. Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 4: 1 - 8" title="Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae)." volume="4" year="2001">Hawks 2001</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.32.1.1" author="Ratcliffe, BC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 48" refId="B317" refString="Ratcliffe, BC, 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1 - 48, DOI: https://doi.org/10.11646/zootaxa.32.1.1" title="A checklist of the Scarabaeoidea (Coleoptera) of Panama." url="https://doi.org/10.11646/zootaxa.32.1.1" volume="32" year="2002">Ratcliffe 2002</bibRefCitation>
, Thomas et al. 2006,
<bibRefCitation author="Krajcik, M" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B155" refString="Krajcik, M, 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." title="Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." year="2008">Krajcik 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
1 ♀ lectotype and 8 paralectotypes of
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Plusiotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusiotis aurora subsp. chrysopedila" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="aurora" subSpecies="chrysopedila">
<emphasis italics="true" pageId="0" pageNumber="1">Plusiotis aurora chrysopedila</emphasis>
</taxonomicName>
at BMNH (
<bibRefCitation author="Hawks, DC" journalOrPublisher="Occasional Papers of the Consortium Coleopterorum" pageId="0" pageNumber="1" pagination="1 - 8" refId="B116" refString="Hawks, DC, 2001. Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 4: 1 - 8" title="Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae)." volume="4" year="2001">Hawks 2001</bibRefCitation>
); 1 paralectotype at CNC (
<bibRefCitation author="Hawks, DC" journalOrPublisher="Occasional Papers of the Consortium Coleopterorum" pageId="0" pageNumber="1" pagination="1 - 8" refId="B116" refString="Hawks, DC, 2001. Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 4: 1 - 8" title="Taxonomic and nomenclatural changes in Chrysina and synonymic checklist of species (Scarabaeidae: Rutelinae)." volume="4" year="2001">Hawks 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<subSubSection id="D0CBACC1E39811AB1046DAF9AE0D0405" pageNumber="608" type="nomenclature">
<paragraph id="19609C3F10B1ADED69DDAF77E4EB2E8E" pageNumber="608">
<taxonomicName id="606C13129BF51678F2A8A03DF76755F9" ID-CoL="5LMD" authority="Pinel 1792" authorityName="Pinel" authorityYear="1792" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="2DEB2EBCC42F144EB0D869C541683895" italics="true" pageNumber="608">Martes</emphasis>
Pinel 1792
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D8A40BD8F7624D86958CF3CE73C66B80" pageNumber="608" type="reference_group">
<paragraph id="1360061A622D91EAC5AB793074AF679E" pageNumber="608">
<treatmentCitation id="CC57A44E7C8BA2D745A1940154A3A741" author="Pinel" authority="Pinel 1792" authorityName="Pinel" authorityYear="1792" class="Mammalia" family="Mustelidae" genus="Martes" journal="Actes Soc. Hist. Nat. Paris" kingdom="Animalia" order="Carnivora" page="55" pageNumber="608" phylum="Chordata" rank="genus" title="Martes" volume="1" year="1792">
<taxonomicName id="49DE8819A7780B22C04B04A7EBC1E15A" ID-CoL="5LMD" authority="Pinel 1792" authorityName="Pinel" authorityYear="1792" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="A7BF04D6BE8CF00191F6B6A565DC73C1" italics="true" pageNumber="608">Martes</emphasis>
Pinel 1792
</taxonomicName>
,
<bibCitation id="2DAC04E75674823CC8480C0C253ABDBA" author="Pinel" journal="Actes Soc. Hist. Nat. Paris" pageNumber="608" pagination="55" title="Martes" volume="1" year="1792">Actes Soc. Hist. Nat. Paris, 1: 55</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5641DAFCE271329356E22F563E76161B" pageNumber="608" type="type_taxon">
<paragraph id="048E955B74C47619089004B59553689F" pageNumber="608">
<emphasis id="0FBC151A56EF4C1E2484698EBC807CE2" inLineHeading="true" pageNumber="608">Type Species:</emphasis>
<taxonomicName id="B8B6583E50A835A4B23B9F619DBB12BE" authority="Pinel 1792" authorityName="Pinel" authorityYear="1792" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="domestica">
<emphasis id="8B6D0D587822893DB52204270F6F1FB4" italics="true" pageNumber="608">Martes domestica</emphasis>
Pinel 1792
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F3F3C713A9589423A4AA7B7353C8A621" pageNumber="608" type="synonym_list">
<paragraph id="EC12AE9AD82735AAED059212519C2922" pageNumber="608">
<emphasis id="D526F0865275F6A0C701024F0BFE2038" inLineHeading="true" pageNumber="608">Synonyms:</emphasis>
<taxonomicName id="91BB5707434CEA7DF95E7F64DB432659" authority="Gray 1865" authorityName="Gray" authorityYear="1865" class="Mammalia" family="Mustelidae" genus="Charronia" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="FD5257BFCB2691CF889A8553C6688D2D" italics="true" pageNumber="608">Charronia</emphasis>
Gray 1865
</taxonomicName>
;
<taxonomicName id="A8B3C47092BDFB0F1BECA72ECD27847E" authority="Gray 1865" authorityName="Gray" authorityYear="1865" class="Mammalia" family="Mustelidae" genus="Foiana" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="EC3B8F0C02A9C4D4A355C86AE02FECB2" italics="true" pageNumber="608">Foiana</emphasis>
Gray 1865
</taxonomicName>
;
<taxonomicName id="4E7E5805B865753241BAD2D972161158" authority="Ognev 1928" authorityName="Ognev" authorityYear="1928" class="Mammalia" family="Mustelidae" genus="Lamprogale" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="590B20D7CAE4007F0AFA6BB82CD0B59C" italics="true" pageNumber="608">Lamprogale</emphasis>
<bibRefCitation id="0085B7B23FAF8ABC42288E7DC5A8D8E0" author="Ognev, S. I." refId="ref264248" refString="Ognev, S. I. 1928. Zveri vostochnoi Evropy i severnoi Azii: Nasekomoyadnye i letychie myshi [Mammals of eastern Europe and northern Asia: Insectivora and Chiroptera]. Glavnauka, Moscow, 1: 1 - 631 (in Russian)." year="1928">Ognev 1928</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D298142382EE8C4D67589A43F43D9474" authority="Gray 1865" authorityName="Gray" authorityYear="1865" class="Mammalia" family="Mustelidae" genus="Pekania" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="D98C87408F5F671B074CBFA3421ACD1C" italics="true" pageNumber="608">Pekania</emphasis>
Gray 1865
</taxonomicName>
;
<taxonomicName id="F61C3AF6DF8D16B40564F986B7055456" authority="Kaup 1829" authorityName="Kaup" authorityYear="1829" class="Mammalia" family="Mustelidae" genus="Zibellina" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="genus">
<emphasis id="0420812F0EB048DB3119D98208D50FDC" italics="true" pageNumber="608">Zibellina</emphasis>
Kaup 1829
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5F339EC3B47CE14D3442B20CA47A09C6" pageNumber="608" type="child_taxa">
<paragraph id="458309D6773CC9D11B547E05030E89E0" pageNumber="608">
<emphasis id="6011FEB042FE1C01CBAA9626008EEF7E" inLineHeading="true" pageNumber="608">Species and subspecies:</emphasis>
8 species with 63 subspecies:
</paragraph>
<paragraph id="E34877A6886F3E83B12EB656F903BA49" pageNumber="608">
Species
<taxonomicName id="3E888A2C7105E7466D54894F68BA73C5" authority="Turton 1806" authorityName="Turton" authorityYear="1806" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="americana">
<emphasis id="E42687BD99400C0B923924E4FDB57FE1" italics="true" pageNumber="608">Martes americana</emphasis>
Turton 1806
</taxonomicName>
</paragraph>
<paragraph id="A986AD3F1CC19878357EBCEFC22E9D80" pageNumber="608">
Subspecies
<taxonomicName id="CEEFD580FA4F907B199397D35A15F7A0" authority="Turton 1806" authorityName="Turton" authorityYear="1806" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="americana">
<emphasis id="79AF07F66195D868BAB7BF9020A33B5A" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="B1A9A84CC0A51AD2DCDE4FAB3E117558" italics="true" pageNumber="608">americana</emphasis>
Turton 1806
</taxonomicName>
</paragraph>
<paragraph id="62CF1C1AF5DF06A00C972E8EFB3D77AC" pageNumber="608">
Subspecies
<taxonomicName id="EE4E3389BCFBCB6572C0E286DBEE9D58" authority="Preble 1902" authorityName="Preble" authorityYear="1902" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="abieticola">
<emphasis id="BE7F8EEA9D970DC421B48461204D5520" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="E0029C68C9C6715E1EE62FA45281E3AC" italics="true" pageNumber="608">abieticola</emphasis>
<bibRefCitation id="A043912EF94093468C4AD72AC5D0E283" author="Preble, E. A." refId="ref293139" refString="Preble, E. A. 1902. A biological investigation of the Hudson Bay region. North American Fauna, 22: 1 - 140." year="1902">Preble 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="19DF1361A6DBFE08D9D425536D57C0E5" pageNumber="608">
Subspecies
<taxonomicName id="AA3033770DB3D25E1862E7B9699DD36C" authority="Gray 1865" authorityName="Gray" authorityYear="1865" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="abietinoides">
<emphasis id="0474677D6002CD6054022612F58C9233" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="F5794EB2DFFB7E4B646E54C145C1364D" italics="true" pageNumber="608">abietinoides</emphasis>
Gray 1865
</taxonomicName>
</paragraph>
<paragraph id="8F47767F8199C7C235231270F661A716" pageNumber="608">
Subspecies
<taxonomicName id="14DCFDA68B857C538F61A6CBF4C1367B" authority="Osgood 1900" authorityName="Osgood" authorityYear="1900" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="actuosa">
<emphasis id="154CC5194D44DC86081B295C6B521B9B" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="9C39F0AF54822DD998BBA36AC1F17E8C" italics="true" pageNumber="608">actuosa</emphasis>
<bibRefCitation id="D81A3523BA08941B1C028FB9BAFDE060" author="Osgood, W. H." refId="ref268619" refString="Osgood, W. H. 1900. Revision of the pocket mice of the genus Perognathus. North American Fauna, 18: 1 - 73." year="1900">Osgood 1900</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="26098C96627E35D1206BC30851C6D848" pageNumber="608">
Subspecies
<taxonomicName id="76B6D43798D2CAF755112CA5BD7EEECC" authority="Bangs 1897" authorityName="Bangs" authorityYear="1897" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="atrata">
<emphasis id="AB8129F38A42C717A854D64BEE17F4ED" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="A2E664F18E591923BDECF743DC901C78" italics="true" pageNumber="608">atrata</emphasis>
Bangs 1897
</taxonomicName>
</paragraph>
<paragraph id="A46AC5E9FA94BFB0C385D233A2BDACC3" pageNumber="608">
Subspecies
<taxonomicName id="6A8D2412EEFFEB44B9B7798BC0632B62" authority="Merriam 1890" authorityName="Merriam" authorityYear="1890" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="caurina">
<emphasis id="0B995AF5D2F40E83EF4540C92AA0E0E6" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="AE2A06439328DA58E5AEF2F80B83DCB7" italics="true" pageNumber="608">caurina</emphasis>
<bibRefCitation id="53AD31FA2D1CB0144A83AA5762D24790" author="Merriam, C. H." refId="ref238216" refString="Merriam, C. H. 1890. Description of a new prairie dog from Wyoming. North American Fauna, 4: 33 - 35." year="1890">Merriam 1890</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="C5C37E46D2AD63C20929E80C478A6620" pageNumber="608">
Subspecies
<taxonomicName id="44FD39FA685A5D1991B92493E98AC0F6" authority="Grinnell and Dixon 1926" authorityName="Grinnell and Dixon" authorityYear="1926" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="humboldtensis">
<emphasis id="AE7690AB8A0D5DC0A5445E2435C09B05" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="1B4C4BD3A91C05F86AEB7D06C3140B5E" italics="true" pageNumber="608">humboldtensis</emphasis>
Grinnell and Dixon 1926
</taxonomicName>
</paragraph>
<paragraph id="1D583C6F8782CDD19CE545BB75C0036A" pageNumber="608">
Subspecies
<taxonomicName id="572BBAAB65BE13DBC0E5A04CEAA2A476" authority="Elliot 1903" authorityName="Elliot" authorityYear="1903" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="kenaiensis">
<emphasis id="7CAD65294197424AFED2A79DBD1CE627" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="93BDCA5A91CE490AEC8069863CF8FBCE" italics="true" pageNumber="608">kenaiensis</emphasis>
Elliot 1903
</taxonomicName>
</paragraph>
<paragraph id="CA21EE85BED74288CA0B1E7C1F832A34" pageNumber="608">
Subspecies
<taxonomicName id="956F301887F1E762DF952B53C80191DD" authority="Osgood 1901" authorityName="Osgood" authorityYear="1901" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="nesophila">
<emphasis id="1AD2FEF83C98EA181C0EC9F1C0E7F01B" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="D1AF1248DE222C5C88ECD0F5E84B858C" italics="true" pageNumber="608">nesophila</emphasis>
Osgood 1901
</taxonomicName>
</paragraph>
<paragraph id="7BA4A371559ECFE9BBA23DADA1C9A4B5" pageNumber="608">
Subspecies
<taxonomicName id="EABD5940CEC599A4BD8B46F4EEDAB17D" authority="Rhoads 1902" authorityName="Rhoads" authorityYear="1902" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="origensis">
<emphasis id="C6D0F8FC69B466982927E1E156AA8A44" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="15A2EAECDB2CFCB5DB0094010298F61C" italics="true" pageNumber="608">origensis</emphasis>
Rhoads 1902
</taxonomicName>
</paragraph>
<paragraph id="47C34151471642CD9D5F79D5BB9EF4DF" pageNumber="608">
Subspecies
<taxonomicName id="09EC6D4492790ADC52C5050AED3E7722" authority="Grinnell and Storer 1916" authorityName="Grinnell and Storer" authorityYear="1916" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="sierrae">
<emphasis id="4635876456CEF4AB9FCFC5BBF505B72E" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="CCFB6AA0980D581637998C52D9D7F7DA" italics="true" pageNumber="608">sierrae</emphasis>
Grinnell and Storer 1916
</taxonomicName>
</paragraph>
<paragraph id="D18FB8DDCCEE61581D8E74AE4E657C7D" pageNumber="608">
Subspecies
<taxonomicName id="DE122DA035DEF321C2B69DBFB2914790" authority="Grinnell and Dixon 1926" authorityName="Grinnell and Dixon" authorityYear="1926" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="vancourverensis">
<emphasis id="658A598E9DADC99829ACDBE9C86D0D27" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="E9926BF384DCDFF1AC48EA513771EDD4" italics="true" pageNumber="608">vancourverensis</emphasis>
Grinnell and Dixon 1926
</taxonomicName>
</paragraph>
<paragraph id="411B35D5386BEA78D00B28316718BA4B" pageNumber="608">
Subspecies
<taxonomicName id="D3A276B8E23E9DB2729784E39581AA55" authority="Rafinesque 1819" authorityName="Rafinesque" authorityYear="1819" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="americana" subSpecies="vulpina">
<emphasis id="88A0B8F64999E511A3CBFE7590F46DDF" italics="true" pageNumber="608">Martes americana</emphasis>
subsp.
<emphasis id="6A46DFD12A8090633F0B3FF6744344A6" italics="true" pageNumber="608">vulpina</emphasis>
Rafinesque 1819
</taxonomicName>
</paragraph>
<paragraph id="8C4CCC6A0483DA5CF22FC09F108E1C0F" pageNumber="608">
Species
<taxonomicName id="1412F9231815AF64D7C5A5ED3AB9EEE9" authority="(Boddaert 1785)" baseAuthorityName="Boddaert" baseAuthorityYear="1785" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="flavigula">
<emphasis id="CBE3651CB6006821C310EB9A890592FA" italics="true" pageNumber="608">Martes flavigula</emphasis>
(
<bibRefCitation id="1FF740065987ED820A746D21A82E9997" author="Boddaert, P." refId="ref34353" refString="Boddaert, P. 1785. Elenchus animalium, volumen 1: Sistens quadrupedia huc usque nota, erorumque varietates. C. R. Hake, Rotterdam, 174 pp." year="1785">Boddaert 1785</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="948E7CAE7B9A3C6FEEFDB892B59A3851" pageNumber="608">
Subspecies
<taxonomicName id="7619E9576204F6F98F5684F5DDDF358F" authority="Boddaert 1785" authorityName="Boddaert" authorityYear="1785" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="flavigula">
<emphasis id="4CF5FFAB3328E1FD11834624931C45D5" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="36B1AC716619F130D9A9C345D52CEFB1" italics="true" pageNumber="608">flavigula</emphasis>
<bibRefCitation id="AC302A59B981C0FBACC1EF201D0E54D8" author="Boddaert, P." refId="ref34353" refString="Boddaert, P. 1785. Elenchus animalium, volumen 1: Sistens quadrupedia huc usque nota, erorumque varietates. C. R. Hake, Rotterdam, 174 pp." year="1785">Boddaert 1785</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="5F0B4AB23E61AF36F607FD94D33E8F19" pageNumber="608">
Subspecies
<taxonomicName id="54DBBEA7019FA34A845EC2226C8DBD83" authority="Radde 1862" authorityName="Radde" authorityYear="1862" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="borealis">
<emphasis id="067AD9EE13349AE9AE0758D0F0F18F7B" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="7EA4423F55837CF73D8311A8FCD16767" italics="true" pageNumber="608">borealis</emphasis>
Radde 1862
</taxonomicName>
</paragraph>
<paragraph id="8D1E8CDF0F86F234CB934859666DC95B" pageNumber="608">
Subspecies
<taxonomicName id="B8948DBFFFFA00906698E46259573357" authority="Swinhoe 1866" authorityName="Swinhoe" authorityYear="1866" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="chrysospila">
<emphasis id="7A8E21BB62A2ABBBB4EF83435EE4C305" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="DBEB519341EB4C7DCFB6E6BFEC025947" italics="true" pageNumber="608">chrysospila</emphasis>
Swinhoe 1866
</taxonomicName>
</paragraph>
<paragraph id="FB71873F7E173877683E20F281DF4667" pageNumber="608">
Subspecies
<taxonomicName id="3B9625FDA6170B3B775C5F780B187B90" authority="Hsu and Wu 1981" authorityName="Hsu and Wu" authorityYear="1981" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="hainana">
<emphasis id="F98A89FCAEF25F10B8265184893B40A1" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="6E964D296DA8500D24FC7AD8FE2E80DB" italics="true" pageNumber="608">hainana</emphasis>
Hsu and Wu 1981
</taxonomicName>
</paragraph>
<paragraph id="9D93A20BC060EEDA4A8849F292CADE5F" pageNumber="608">
Subspecies
<taxonomicName id="DEA6DC5A0391BC10BF1B809D53081EE7" authority="Schinz 1845" authorityName="Schinz" authorityYear="1845" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="henrici">
<emphasis id="33D4E643C8B45751E230B8E15E9BC217" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="B4EDFC535B659736CF22E02491EB2BA6" italics="true" pageNumber="608">henrici</emphasis>
Schinz 1845
</taxonomicName>
</paragraph>
<paragraph id="AC8469D781CD7140BC8B5931EAE2710B" pageNumber="608">
Subspecies
<taxonomicName id="9016D9CED42EEE8F11CE4C4B62620EC1" authority="Kloss 1916" authorityName="Kloss" authorityYear="1916" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="indochinensis">
<emphasis id="30EB8971CDE8066A0C6677341E875428" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="92C144E802BB45C14776413BA5B743C3" italics="true" pageNumber="608">indochinensis</emphasis>
Kloss 1916
</taxonomicName>
</paragraph>
<paragraph id="8648ECBB49CCE4CFDAA87D9F735AE9B7" pageNumber="608">
Subspecies
<taxonomicName id="8119D653C73627D3E5155EF34E92F1BB" authority="Bonhote 1901" authorityName="Bonhote" authorityYear="1901" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="peninsularis">
<emphasis id="377C415F0D638A86D61EDF31D0E504E1" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="69BA3227FDD88DABFA3F2CA9FDB9AF75" italics="true" pageNumber="608">peninsularis</emphasis>
Bonhote 1901
</taxonomicName>
</paragraph>
<paragraph id="BE7AF47CFBB0B726DB3AF20808E93809" pageNumber="608">
Subspecies
<taxonomicName id="853E64F1E5B35174DEE891B2513B8963" authority="Pocock 1936" authorityName="Pocock" authorityYear="1936" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="robinsoni">
<emphasis id="26756F8A432D6FC73CE591AEA67DC6FA" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="C2AEB8A630202E20F32D6D74528685CE" italics="true" pageNumber="608">robinsoni</emphasis>
Pocock 1936
</taxonomicName>
</paragraph>
<paragraph id="DFDE9DB9EF8F4299174A9CDD00CABDE2" pageNumber="608">
Subspecies
<taxonomicName id="A00E588A96F5B10D30197B41978D0B78" authority="Chasen and Kloss 1931" authorityName="Chasen and Kloss" authorityYear="1931" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="flavigula" subSpecies="saba">
<emphasis id="BF15BEBCADE7D70435ADA3EDC27C0A2F" italics="true" pageNumber="608">Martes flavigula</emphasis>
subsp.
<emphasis id="07F2B73C65CFF775AAC8F0AD08DD983B" italics="true" pageNumber="608">saba</emphasis>
Chasen and Kloss 1931
</taxonomicName>
</paragraph>
<paragraph id="5EF1CC12242D176D543F5D0356BFB230" pageNumber="608">
Species
<taxonomicName id="952654388D08027B420FE4B4A2AE7A4E" authority="(Erxleben 1777)" baseAuthorityName="Erxleben" baseAuthorityYear="1777" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="foina">
<emphasis id="5FFCF5B77D2B817FC6FEB2E8ECF78899" italics="true" pageNumber="608">Martes foina</emphasis>
(
<bibRefCitation id="C4AF77D4A83F2EF02A030217F10CC3A3" author="Erxleben, J. C. P." refId="ref96470" refString="Erxleben, J. C. P. 1777. Systema regni animalis per classes, ordines, genera, species, varietates, cum synonymia et historia animalium. Classis I. Mammalia. Weygandianis, Lipsiae, 636 pp." year="1777">Erxleben 1777</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="D0A8B4E86C98D42274E537F4F271D12C" pageNumber="608">
Subspecies
<taxonomicName id="C7674D7EB4F69FFBFB76F0D8F1392129" authority="Erxleben 1777" authorityName="Erxleben" authorityYear="1777" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="foina">
<emphasis id="EB99B082E24395AC501EB90C574B6A10" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="B14466DD18EC419F242D6397AE6EC909" italics="true" pageNumber="608">foina</emphasis>
<bibRefCitation id="061A497329FB9BF9C1FA22A5C04F0773" author="Erxleben, J. C. P." refId="ref96470" refString="Erxleben, J. C. P. 1777. Systema regni animalis per classes, ordines, genera, species, varietates, cum synonymia et historia animalium. Classis I. Mammalia. Weygandianis, Lipsiae, 636 pp." year="1777">Erxleben 1777</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="3D16DBBE254ED6BD1018D90397A6E7A1" pageNumber="608">
Subspecies
<taxonomicName id="F401A1AC20F410A9E3D20425E8195AAF" authority="Brass 1911" authorityName="Brass" authorityYear="1911" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="bosniaca">
<emphasis id="6E2697A9392FB52B39E76DE1B44E663F" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="C18A7CDBC1DC50AC4988D03E544AD661" italics="true" pageNumber="608">bosniaca</emphasis>
Brass 1911
</taxonomicName>
</paragraph>
<paragraph id="E3F7B0CF0E7ECCF1D6258A3A8A51E7CE" pageNumber="608">
Subspecies
<taxonomicName id="5E74631DC4E92B21BA1C72A4B73BB0CD" authority="Bate 1906" authorityName="Bate" authorityYear="1906" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="bunites">
<emphasis id="BC3D225978EC110ED0471A451B11CAA0" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="1407E4550ACC098B998D62B65E62ABEF" italics="true" pageNumber="608">bunites</emphasis>
Bate 1906
</taxonomicName>
</paragraph>
<paragraph id="D0CA0913C816D7790D713C29418010B0" pageNumber="608">
Subspecies
<taxonomicName id="D2D4AE44C7DF3727A6F6984544E2369C" authority="Ognev 1931" authorityName="Ognev" authorityYear="1931" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="kozlovi">
<emphasis id="1B70AB688444052623D088836576343D" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="1FA0D28DAB0F8871FB179BB5D57ED6C7" italics="true" pageNumber="608">kozlovi</emphasis>
<bibRefCitation id="FF76D5E6241CA919BEC5EF9D8CD21738" author="Ognev, S. I." refId="ref264295" refString="Ognev, S. I. 1931. Zveri vostochnoi Evropy i severnoi Azii: Khishchnye mlekopitayushchie [Mammals of eastern Europe and northern Asia: Carnivorous mammals]. Glavnauka, Moscow, 2: 1 - 776 (in Russian)." year="1931">Ognev 1931</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1326BB551005871EDB77C018FE3886A8" pageNumber="608">
Subspecies
<taxonomicName id="7BE5299C1F501EA70F0647780091B636" authority="Severtzov 1873" authorityName="Severtzov" authorityYear="1873" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="intermedia">
<emphasis id="DB71374F2FB585AFE1C193C2F51656B4" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="BEF5E90D29D8AFA4F894471B0AD5CC1B" italics="true" pageNumber="608">intermedia</emphasis>
Severtzov 1873
</taxonomicName>
</paragraph>
<paragraph id="5F0E8F1BB7DFFEAB58387B9039385B14" pageNumber="608">
Subspecies
<taxonomicName id="EAE14F0A236BCB7D2E85690ADCDCED80" authority="Barrett-Hamilton 1898" authorityName="Barrett-Hamilton" authorityYear="1898" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="mediterranea">
<emphasis id="AEE5F19BE390B3C427F9D21415851E4D" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="11793655E2A3113260389D3940927498" italics="true" pageNumber="608">mediterranea</emphasis>
Barrett-Hamilton 1898
</taxonomicName>
</paragraph>
<paragraph id="3E3FE891137D99EC42ABA8EB19B968ED" pageNumber="608">
Subspecies
<taxonomicName id="D6F9C030CDAD44BB13273F98959B9E49" authority="Festa 1914" authorityName="Festa" authorityYear="1914" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="milleri">
<emphasis id="D538A752F71E36DD39E80B79A795E715" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="4D7ED381E2913DABE529D11EBB576CBF" italics="true" pageNumber="608">milleri</emphasis>
Festa 1914
</taxonomicName>
</paragraph>
<paragraph id="5285B172609F99A4D636A4D67536865A" pageNumber="608">
Subspecies
<taxonomicName id="0AE2B2BA5D7BFB6D3DD5001ED642A761" authority="Satunin 1906" authorityName="Satunin" authorityYear="1906" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="nehringi">
<emphasis id="956BBE5F44BFDB499680FDF0959548F6" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="03499EF31ECBF7DC41B88BDDBCBC7A3D" italics="true" pageNumber="608">nehringi</emphasis>
Satunin 1906
</taxonomicName>
</paragraph>
<paragraph id="290CFEF4A78E548A4116C8E2D8268B51" pageNumber="608">
Subspecies
<taxonomicName id="AF0074C63096FC1431DE586FCA58B57C" authority="Martino and Martino 1917" authorityName="Martino and Martino" authorityYear="1917" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="rosanowi">
<emphasis id="89E929FD7710F5A5EF29EE7D9098F8D8" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="512793FF83E9CD875A8DD4DCE6D2B647" italics="true" pageNumber="608">rosanowi</emphasis>
Martino and Martino 1917
</taxonomicName>
</paragraph>
<paragraph id="1EB2F4B094C26D403BD9530917C3AD2E" pageNumber="608">
Subspecies
<taxonomicName id="9F0324CBB79167547BA40B9720635BF1" authority="Nehring 1902" authorityName="Nehring" authorityYear="1902" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="syriaca">
<emphasis id="3D56DC1E8F080B597FB4873496B03EB0" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="887A3F945E4C951FE2DFE71A88A7ECF5" italics="true" pageNumber="608">syriaca</emphasis>
Nehring 1902
</taxonomicName>
</paragraph>
<paragraph id="D559648DCB7C8D5241CDDA3AECCCAF4C" pageNumber="608">
Subspecies
<taxonomicName id="C20CFBAE90A9A98344E703828DB58EC2" authority="Hodgson 1842" authorityName="Hodgson" authorityYear="1842" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="foina" subSpecies="toufoeus">
<emphasis id="986733D6231F401047498242C23880A6" italics="true" pageNumber="608">Martes foina</emphasis>
subsp.
<emphasis id="C3D0D1E89B865082019F737B53C5B19A" italics="true" pageNumber="608">toufoeus</emphasis>
<bibRefCitation id="B9E405F6359C26776DD03881B262A8C5" author="Hodgson, B. H." refId="ref154390" refString="Hodgson, B. H. 1842. Notice of the mammals of Tibet, with descriptions and plates of some new species. Journal of the Asiatic Society of Bengal, 11: 275 - 289." year="1842">Hodgson 1842</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="BA38ABA93171252AEA73CFE5174568A0" pageNumber="608">
Species
<taxonomicName id="6C9EF2F6FD9C6629612A165E6FE88877" authority="Horsfield 1851" authorityName="Horsfield" authorityYear="1851" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="gwatkinsii">
<emphasis id="A0ABE8C9D76C0833A25BA1F9E20E6325" italics="true" pageNumber="608">Martes gwatkinsii</emphasis>
Horsfield 1851
</taxonomicName>
</paragraph>
<paragraph id="EDF14D1E64BBF4B277B0D011FAB6A81C" pageNumber="608">
Species
<taxonomicName id="ABFC24361B6DF196D8DB45923F0E1FB3" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="martes">
<emphasis id="7B67B8B615DAF0C2E926782BB9108424" italics="true" pageNumber="608">Martes martes</emphasis>
(
<bibRefCitation id="A525F85882133C38EC9CEA82ABBE26BE" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="0867F38E8ECAA064C59D75D95ADCFDFA" pageNumber="608">
Subspecies
<taxonomicName id="FAC0FA2F7D25800E5E396B4A21C5718D" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="martes">
<emphasis id="5919C4A802CF031A0E7A751826803360" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="471C6EB45AB00FB4D42B449BAEE0F9EA" italics="true" pageNumber="608">martes</emphasis>
<bibRefCitation id="F8319ED4EC0166863BEC027B0DA3E5A2" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="0F5FF320D4F7C9F499EE6C9BD154A66E" pageNumber="608">
Subspecies
<taxonomicName id="46BA2F3D0D0CE754C32722ED1ACE8F1A" authority="Kuznetsov 1944" authorityName="Kuznetsov" authorityYear="1944" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="borealis">
<emphasis id="8D1B231ACC7F123638DA8FD16F7AAD23" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="DB217CCE40ACFA48D25F877C1B630CFE" italics="true" pageNumber="608">borealis</emphasis>
<bibRefCitation id="6D7439EF6D41C6F227EA31C532CEC3C3" author="Kuznetsov, B. A." refId="ref202214" refString="Kuznetsov, B. A. 1944. Ordo Rodentia. Pp. 262 - 362, in Opredelitel' mlekopitayushchikh SSSR [Guide to the mammals of the USSR] (Bobrinskii, N. A., Kuznetsov, B. A. and A. P. Kuzyakin, eds.). Sovetskaya Nauka, Moscow, 439 pp. (in Russian)." year="1944">Kuznetsov 1944</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="34AF8F4201B84C647951B3361D78CE5E" pageNumber="608">
Subspecies
<taxonomicName id="6070A00E61DDEC717B5837F2F5C88DB0" authority="Barrett-Hamilton 1904" authorityName="Barrett-Hamilton" authorityYear="1904" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="latinorum">
<emphasis id="66F9D6B9E839CE15FDCA4D7CDCD70CFF" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="552C5D1661217687C5A6E21C83D1B928" italics="true" pageNumber="608">latinorum</emphasis>
Barrett-Hamilton 1904
</taxonomicName>
</paragraph>
<paragraph id="399757EE94FCBBD74CDA60F106D77858" pageNumber="608">
Subspecies
<taxonomicName id="E4C72CB707F89D0D72910DFE4F31F2CC" authority="Ognev 1926" authorityName="Ognev" authorityYear="1926" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="lorenzi">
<emphasis id="7E673D71015D782C4274C2172657FEF3" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="A15F9C6EBD0D7A463A14ECA0A30B4CBA" italics="true" pageNumber="608">lorenzi</emphasis>
Ognev 1926
</taxonomicName>
</paragraph>
<paragraph id="F553A1BA4BFC04928BA5706849D1C0B0" pageNumber="608">
Subspecies
<taxonomicName id="71B63DAD989520C402E94430D1D3F5A6" authority="Alcover, Delibes, Gosálbez, and Nadal 1987" authorityName="Alcover, Delibes, Gosálbez, and Nadal" authorityYear="1987" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="minoricensis">
<emphasis id="C15C6EDF388601ACD5DFF65C1E8B5D31" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="5EDC58AA30F78E5AC1A02CD6F99A856D" italics="true" pageNumber="608">minoricensis</emphasis>
Alcover, Delibes, Gosálbez, and Nadal 1987
</taxonomicName>
</paragraph>
<paragraph id="79E9DF466F5D3B739AC2059665D9FB9F" pageNumber="608">
Subspecies
<taxonomicName id="7F050528DB73C437A33E4D090A3079D9" authority="Cavazza 1912" authorityName="Cavazza" authorityYear="1912" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="notialis">
<emphasis id="7160C4413E2CEE80B165C44C959808BE" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="75551BC281D756D710D639F986B52236" italics="true" pageNumber="608">notialis</emphasis>
Cavazza 1912
</taxonomicName>
</paragraph>
<paragraph id="09FBF7A7DA54336C0E2D80E1B4B8E85B" pageNumber="608">
Subspecies
<taxonomicName id="846F40FF9994BCCCFA51D35BFEB0F7B4" authority="Ognev 1926" authorityName="Ognev" authorityYear="1926" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="ruthena">
<emphasis id="31E59D5B66473D415A1FC0B3C4FA1708" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="DB9E93F682F6C0485C898D802A426A25" italics="true" pageNumber="608">ruthena</emphasis>
Ognev 1926
</taxonomicName>
</paragraph>
<paragraph id="5F9B19CB3AC457FD4ABBDF214ACC7932" pageNumber="608">
Subspecies
<taxonomicName id="2EC8140D39F649675F398C1F1186C00C" authority="Kuznetsov 1941" authorityName="Kuznetsov" authorityYear="1941" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="martes" subSpecies="uralensis">
<emphasis id="A792E26134F1D432E35BB1BAC444D482" italics="true" pageNumber="608">Martes martes</emphasis>
subsp.
<emphasis id="0F40A8FBD4CB51E9788580E22FD4283A" italics="true" pageNumber="608">uralensis</emphasis>
Kuznetsov 1941
</taxonomicName>
</paragraph>
<paragraph id="C41C8CE80A654CAD0911E6AE204CB0A0" pageNumber="608">
Species
<taxonomicName id="45DB8993BB59617D742284328F972AAE" authority="Wagner 1840" authorityName="Wagner" authorityYear="1840" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="melampus">
<emphasis id="2899C2E6E408F33E120F126A86E31C3B" italics="true" pageNumber="608">Martes melampus</emphasis>
Wagner 1840
</taxonomicName>
</paragraph>
<paragraph id="BD0A6E23BCE524E1373EF86782B69C43" pageNumber="608">
Subspecies
<taxonomicName id="8D2F78442087195974851041B5E747C6" authority="Wagner 1840" authorityName="Wagner" authorityYear="1840" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="melampus" subSpecies="melampus">
<emphasis id="C2C639FE638014014A50D39FDBD1FFFD" italics="true" pageNumber="608">Martes melampus</emphasis>
subsp.
<emphasis id="FE0831C161CE2AEC2147233BE2E0847C" italics="true" pageNumber="608">melampus</emphasis>
Wagner 1840
</taxonomicName>
</paragraph>
<paragraph id="1AA4718996C7007E8A893AEC0AE1E0B4" pageNumber="608">
Subspecies
<taxonomicName id="59F5C5D6EEB63F70044D447BE5499DD8" authority="Kuroda and Mori 1923" authorityName="Kuroda and Mori" authorityYear="1923" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="melampus" subSpecies="coreensis">
<emphasis id="8A17731503DAC34CC9AFE47A16FE6BE8" italics="true" pageNumber="608">Martes melampus</emphasis>
subsp.
<emphasis id="B61B9DF8488E585EA79B079920A1E060" italics="true" pageNumber="608">coreensis</emphasis>
Kuroda and Mori 1923
</taxonomicName>
</paragraph>
<paragraph id="7120E83C5DCE1B6CE54676B0B3094584" pageNumber="608">
Subspecies
<taxonomicName id="D0B67F5CEFE57C12C9A740BD32525725" authority="Thomas 1897" authorityName="Thomas" authorityYear="1897" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="melampus" subSpecies="tsuensis">
<emphasis id="4650A0555464F202B62510C7981DAB1D" italics="true" pageNumber="608">Martes melampus</emphasis>
subsp.
<emphasis id="078F68AC4F3D26481D9E707593AE966C" italics="true" pageNumber="608">tsuensis</emphasis>
Thomas 1897
</taxonomicName>
</paragraph>
<paragraph id="588E3764D01001F15E1A731F979A2000" pageNumber="608">
Species
<taxonomicName id="BD8ED5B8055A69EBDF53AD5A7DDB0296" authority="(Erxleben 1777)" baseAuthorityName="Erxleben" baseAuthorityYear="1777" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="pennanti">
<emphasis id="CFA48E759476F01A7C881C4095E47F1B" italics="true" pageNumber="608">Martes pennanti</emphasis>
(
<bibRefCitation id="E28C338176C29AACF85612DBB2274150" author="Erxleben, J. C. P." refId="ref96470" refString="Erxleben, J. C. P. 1777. Systema regni animalis per classes, ordines, genera, species, varietates, cum synonymia et historia animalium. Classis I. Mammalia. Weygandianis, Lipsiae, 636 pp." year="1777">Erxleben 1777</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="C5F19801B31B92E9855353C9BC6E4C2E" pageNumber="608">
Subspecies
<taxonomicName id="C215D0E178A3B30B39C67BBAC887B584" authority="Erxleben 1777" authorityName="Erxleben" authorityYear="1777" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="pennanti" subSpecies="pennanti">
<emphasis id="F740090D5CD20BB588FB8348709F5A22" italics="true" pageNumber="608">Martes pennanti</emphasis>
subsp.
<emphasis id="8A7F4E673155D9DE23593ADE842979E2" italics="true" pageNumber="608">pennanti</emphasis>
<bibRefCitation id="CB82BBE182E85A5287D1A229DE9FE6C3" author="Erxleben, J. C. P." refId="ref96470" refString="Erxleben, J. C. P. 1777. Systema regni animalis per classes, ordines, genera, species, varietates, cum synonymia et historia animalium. Classis I. Mammalia. Weygandianis, Lipsiae, 636 pp." year="1777">Erxleben 1777</bibRefCitation>
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</paragraph>
<paragraph id="66F4470A1A54FB0AB29B312272EF66D2" pageNumber="608">
Subspecies
<taxonomicName id="242C77DAE3B73B7A6D7D5D5A821CECFB" authority="Goldman 1935" authorityName="Goldman" authorityYear="1935" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="pennanti" subSpecies="columbiana">
<emphasis id="8464291D1474F706BB66D249212FB6FF" italics="true" pageNumber="608">Martes pennanti</emphasis>
subsp.
<emphasis id="0D9517039795CA9FFFF7231A595A624F" italics="true" pageNumber="608">columbiana</emphasis>
<bibRefCitation id="99F498955040254955D62E803E8D8153" author="Goldman, E. A." refId="ref122773" refString="Goldman, E. A. 1935. New American mustelids of the genera Martes, Gulo, and Lutra. Proceedings of the Biological Society of Washington, 48: 175 - 186." year="1935">Goldman 1935</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="98C052AF6FA729F3B4AA632F9AA2E0BE" pageNumber="608">
Subspecies
<taxonomicName id="E1633E977B22C77DE1396B002BA5064F" authority="Rhoads 1898" authorityName="Rhoads" authorityYear="1898" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="pennanti" subSpecies="pacifica">
<emphasis id="0FB121275B7D3A0F2FEB320A321457CB" italics="true" pageNumber="608">Martes pennanti</emphasis>
subsp.
<emphasis id="38F743B15D7BF476ECD10D3EBD09CD72" italics="true" pageNumber="608">pacifica</emphasis>
Rhoads 1898
</taxonomicName>
</paragraph>
<paragraph id="6944C39D3702EE66E07BFE47DF22F42E" pageNumber="608">
Species
<taxonomicName id="CAAAC61F3F5159235B182525C1A78821" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="species" species="zibellina">
<emphasis id="AE0AE17FCB01FDBF1FA02239BF95B037" italics="true" pageNumber="608">Martes zibellina</emphasis>
(
<bibRefCitation id="309DC6A18C549FD962F51C031E715641" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="3E22F01046611CF680971FAD9F676CEC" pageNumber="608">
Subspecies
<taxonomicName id="54775EAF5C71B5B703B04DE609EABE29" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="zibellina">
<emphasis id="42CA3B1DFCEBB054C2033C27D88CB188" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="16F4BF798F195F56BEEA0A5D92243B0A" italics="true" pageNumber="608">zibellina</emphasis>
<bibRefCitation id="3C26EDF510DD58902B59A39E217BEF16" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1916B0A49F13FEEFC66FC82BB9C89248" pageNumber="608">
Subspecies
<taxonomicName id="77A246B8CA6E21276EB61BF80E925B5E" authority="Timofeev and Nadeev 1955" authorityName="Timofeev and Nadeev" authorityYear="1955" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="angarensis">
<emphasis id="DE317E466A4E542BAC726742DBA4F2B8" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="666BA7D41A0541E00A6FE8E0A19EA945" italics="true" pageNumber="608">angarensis</emphasis>
Timofeev and Nadeev 1955
</taxonomicName>
</paragraph>
<paragraph id="FA5FE87E7336111FF722822978286754" pageNumber="608">
Subspecies
<taxonomicName id="779554E8110E628DF4CBED82C695EF65" authority="Kuznetsov 1944" authorityName="Kuznetsov" authorityYear="1944" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="arsenjevi">
<emphasis id="9479ECE4F60BDB4DF0E5C7FB60E5C232" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="2FAFCE25104D64AEF5084F176192A635" italics="true" pageNumber="608">arsenjevi</emphasis>
<bibRefCitation id="D24C2694379E1E9BDFC13036F84E8A37" author="Kuznetsov, B. A." refId="ref202214" refString="Kuznetsov, B. A. 1944. Ordo Rodentia. Pp. 262 - 362, in Opredelitel' mlekopitayushchikh SSSR [Guide to the mammals of the USSR] (Bobrinskii, N. A., Kuznetsov, B. A. and A. P. Kuzyakin, eds.). Sovetskaya Nauka, Moscow, 439 pp. (in Russian)." year="1944">Kuznetsov 1944</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="22323BE41BA831F3B3528A0ECBA75B7C" pageNumber="608">
Subspecies
<taxonomicName id="A29796A6EE2C925BAE70F0C629C5FD45" authority="Bashanov 1943" authorityName="Bashanov" authorityYear="1943" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="averini">
<emphasis id="CE1C4EEB13CF36F11D0D0781E6C3B2C3" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="611E593FEDE43ED276B42A4B567EB090" italics="true" pageNumber="608">averini</emphasis>
Bashanov 1943
</taxonomicName>
</paragraph>
<paragraph id="AC75A001A5E2B83E3EACFD8266104623" pageNumber="608">
Subspecies
<taxonomicName id="29536FAB7E9007DCAC493006B4F36AD8" authority="Temminck 1844" authorityName="Temminck" authorityYear="1844" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="brachyura">
<emphasis id="F7B447E498F7C6F4079D11477980BC3C" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="78FB8C9DE44A1F98FC89D9C874148892" italics="true" pageNumber="608">brachyura</emphasis>
Temminck 1844
</taxonomicName>
</paragraph>
<paragraph id="CDD98789945A68F33C16E8B8DE5BBDFE" pageNumber="608">
Subspecies
<taxonomicName id="FADFF2A973F7F9129D2D0D10B0F5D180" authority="Timofeev and Nadeev 1955" authorityName="Timofeev and Nadeev" authorityYear="1955" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="ilimpiensis">
<emphasis id="278A0219596C320FFCFE06D1FC271D4B" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="8B4BEECC71AE13FFB9A611E2C47AB310" italics="true" pageNumber="608">ilimpiensis</emphasis>
Timofeev and Nadeev 1955
</taxonomicName>
</paragraph>
<paragraph id="01E7AC6477E6D1DE6AE6C0B63E2A256D" pageNumber="608">
Subspecies
<taxonomicName id="2AD47E19D52F051AEEA980526585F1FB" authority="Novikov 1956" authorityName="Novikov" authorityYear="1956" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="jakutensis">
<emphasis id="F42BB6BEDD87E3112CFDE07C1A95325A" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="989FECD9310D113459250C03D88D9199" italics="true" pageNumber="608">jakutensis</emphasis>
<bibRefCitation id="08A73B613394701211581A3D34B28C60" author="Novikov, G. A." refId="ref262736" refString="Novikov, G. A. 1956. Khishchye mlekopitayushchie fauna SSSR [Carnivorous mammals of the fauna of the USSR]. Akademiya Nauk SSSR, Moscow-Leningrad, 293 pp. (in Russian)." year="1956">Novikov 1956</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="AA12F65CE21C8962E510CAA27852EBD6" pageNumber="608">
Subspecies
<taxonomicName id="C75C23ADBF6BEE9F98EE5E72BF422E7E" authority="Birula 1919" authorityName="Birula" authorityYear="1919" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="kamtschadalica">
<emphasis id="FBF9931BB0F4AB1CD9843B476475C161" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="B5EA7AB4635BE40B910E83B081248F8D" italics="true" pageNumber="608">kamtschadalica</emphasis>
Birula 1919
</taxonomicName>
</paragraph>
<paragraph id="9DCEFEC230B5D32F5E5DF008485683FE" pageNumber="608">
Subspecies
<taxonomicName id="2D320407A2344108335E7EBBD654775F" authority="Timofeev and Nadeev 1955" authorityName="Timofeev and Nadeev" authorityYear="1955" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="obscura">
<emphasis id="9DBB7AB5CA9408DDC644D3A011463AC9" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="855330A0B19C657C16470098514EE435" italics="true" pageNumber="608">obscura</emphasis>
Timofeev and Nadeev 1955
</taxonomicName>
</paragraph>
<paragraph id="2B9CB11BB1F129FCE495802AFAEF8F65" pageNumber="608">
Subspecies
<taxonomicName id="D7EF939F79927869A320DBE4317687F3" authority="Birula 1922" authorityName="Birula" authorityYear="1922" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="princeps">
<emphasis id="6231267B8D5B94C7696C2D2CED586F3F" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="8004659AA80F741A718AED9D699A69E2" italics="true" pageNumber="608">princeps</emphasis>
Birula 1922
</taxonomicName>
</paragraph>
<paragraph id="0B1E88CAA1F42E45A0BA5E449F8ABE9E" pageNumber="608">
Subspecies
<taxonomicName id="653DB0FD9C508967B17DB0039763AA7B" authority="Ognev 1925" authorityName="Ognev" authorityYear="1925" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="sahalinensis">
<emphasis id="96A620EEE239124A5D5BE793EFD73A91" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="90FFC6C36A48328C3551FCD90C550A00" italics="true" pageNumber="608">sahalinensis</emphasis>
Ognev 1925
</taxonomicName>
</paragraph>
<paragraph id="C02CEBB8812F7A0019341CD922CCE6A9" pageNumber="608">
Subspecies
<taxonomicName id="01B38FC2CE88538F1BEA1BABBEF07B2E" authority="Ognev 1925" authorityName="Ognev" authorityYear="1925" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="sajanensis">
<emphasis id="98801F4BEAF5A611934F4045F9C1A389" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="80F0EBCD20570B0B14CE58B17ACC3E39" italics="true" pageNumber="608">sajanensis</emphasis>
Ognev 1925
</taxonomicName>
</paragraph>
<paragraph id="EEE221FF43BAD95A750485FBD212CB4E" pageNumber="608">
Subspecies
<taxonomicName id="BE78BF156ACB9DD23BF61F8B344A4BCB" authority="Kuznetsov 1941" authorityName="Kuznetsov" authorityYear="1941" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="schantaricus">
<emphasis id="51D637B4CE14B13BC71A414B3DF4F0C3" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="B2FD685F9CEAF7D85BD821D59B902686" italics="true" pageNumber="608">schantaricus</emphasis>
Kuznetsov 1941
</taxonomicName>
</paragraph>
<paragraph id="20B46A96D9BE68723FF8109AC7F71E75" pageNumber="608">
Subspecies
<taxonomicName id="5EFCFD6E9E88BA2A398D8F08582ADA3E" authority="Timofeev and Nadeev 1955" authorityName="Timofeev and Nadeev" authorityYear="1955" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="tomensis">
<emphasis id="66FE745D1BB9D193B3D21D0BD227328C" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="B299DA86BC6C90BB47E0BA53E9B7B450" italics="true" pageNumber="608">tomensis</emphasis>
Timofeev and Nadeev 1955
</taxonomicName>
</paragraph>
<paragraph id="7A7DACF790D1160F2FDCC2C8C1D23485" pageNumber="608">
Subspecies
<taxonomicName id="06551EDFE8A6D29DD64929B273EE063F" authority="Kuznetsov 1944" authorityName="Kuznetsov" authorityYear="1944" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="tungussensis">
<emphasis id="952043B24B8686CC89843715E8F1C1B8" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="1255761AFD285D76C1E35D9B2D1B6A84" italics="true" pageNumber="608">tungussensis</emphasis>
<bibRefCitation id="4CA569C1D05D4C7961304962875888DB" author="Kuznetsov, B. A." refId="ref202214" refString="Kuznetsov, B. A. 1944. Ordo Rodentia. Pp. 262 - 362, in Opredelitel' mlekopitayushchikh SSSR [Guide to the mammals of the USSR] (Bobrinskii, N. A., Kuznetsov, B. A. and A. P. Kuzyakin, eds.). Sovetskaya Nauka, Moscow, 439 pp. (in Russian)." year="1944">Kuznetsov 1944</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="E39947CDCE5A0B5C6AAB0105DE39EDB7" pageNumber="608">
Subspecies
<taxonomicName id="CC19064A43552F1644B6388B7BE912F4" authority="Ognev 1925" authorityName="Ognev" authorityYear="1925" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" pageNumber="608" phylum="Chordata" rank="subSpecies" species="zibellina" subSpecies="yeniseensis">
<emphasis id="C1C94587D381FCBB23E659EE34C8337E" italics="true" pageNumber="608">Martes zibellina</emphasis>
subsp.
<emphasis id="1BB68D2BD06FED45F95C858B7BA432B4" italics="true" pageNumber="608">yeniseensis</emphasis>
Ognev 1925
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1DEB7C9BA2F478A6CDC1CA8BC4835477" pageNumber="608" type="discussion">
<paragraph id="E2FD8B5E6CE403691EB30195ED9B13D5" pageNumber="608">
<emphasis id="3B7EF53957F17A8E820899D7F4064253" inLineHeading="true" pageNumber="608">Discussion:</emphasis>
Stone and Cook’s (2002) cytochrome
<emphasis id="D001956824B2335560329C790381C729" italics="true" pageNumber="608">b</emphasis>
data suggested that the recognition of
<taxonomicName id="C241293A6C9EC96D1BA349807751A6B0" authorityName="Pinel" authorityYear="1792" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="genus">
<emphasis id="4EFBF868B6D0470FB60FE296B54B70B7" italics="true" pageNumber="608">Martes</emphasis>
</taxonomicName>
as here understood (to the exclusion of
<taxonomicName id="731CDDF6061F0DB31EFA54D97459CF13" authorityName="Frisch" authorityYear="1775" class="Mammalia" family="Mustelidae" genus="Gulo" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="genus">
<emphasis id="34AD4E1497D8A5ABFB56D80A6BB0C491" italics="true" pageNumber="608">Gulo</emphasis>
</taxonomicName>
) would make the genus paraphyletic.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="086DEF6909276D2426398111703F5CFF">Dunn, Robert R.</mods:namePart>
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<mods:typeOfResource id="B02172CD839D38F7444DD39D80FDAA8B">text</mods:typeOfResource>
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<mods:title id="E9D8E3DB5E1A10161DA03D1C3FBA4466">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C705273713AB281D4C40757FFE2C74AA">
<mods:date id="2B6126F3B47792CEC50E810DA786DA17">2010</mods:date>
<mods:detail id="FE5FE1EC820FFA70E6D3077FFD6BDAA4" type="volume">
<mods:number id="347B522512355D3CA00217E7D6CD690B">2685</mods:number>
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<mods:start id="B055E9F1E67E82829035CC9F11C05FB4">39</mods:start>
<mods:end id="DF38517A43DD6410BD3B3D7D920AB376">50</mods:end>
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<mods:classification id="03E0E1C8DBCDE8CC36D72D8CBF55C2B5">journal article</mods:classification>
<mods:identifier id="B9F55C875B7A54A6B143B9C7FBF9C3D9" type="DOI">10.5281/zenodo.199426</mods:identifier>
<mods:identifier id="530A6CEE49302325E0A8376E2D2974FC" type="GBIF-Dataset">37bf5700-ad0c-44f1-b035-da049614d7d9</mods:identifier>
<mods:identifier id="B2BD2FB016CCD9555C71F59E5103FD07" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8A4C1F2EB1F6AAAFE7A1086E551CF179" type="Zenodo-Dep">199426</mods:identifier>
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<treatment id="9F7187D2FFD35712D7BF94C0FDB2FA4D" ID-DOI="http://doi.org/10.5281/zenodo.6205795" ID-GBIF-Taxon="119405425" ID-Zenodo-Dep="6205795" LSID="urn:lsid:plazi:treatment:9F7187D2FFD35712D7BF94C0FDB2FA4D" httpUri="http://treatment.plazi.org/id/9F7187D2FFD35712D7BF94C0FDB2FA4D" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44">
<subSubSection id="5FC2654FFFD3571FD7BF94C0FEDAFEF0" pageId="5" pageNumber="44" type="nomenclature">
<paragraph id="176736C4FFD3571FD7BF94C0FDD6FF1D" blockId="5.[151,541,221,282]" box="[151,541,221,247]" pageId="5" pageNumber="44">
<heading id="4C2F81A8FFD3571FD7BF94C0FDD6FF1D" bold="true" box="[151,541,221,247]" fontSize="11" level="1" pageId="5" pageNumber="44" reason="1">
<emphasis id="25ACEAD6FFD3571FD7BF94C0FDD6FF1D" bold="true" box="[151,541,221,247]" pageId="5" pageNumber="44">
<taxonomicName id="D0D84D47FFD3571FD7BF94C0FE73FF1D" box="[151,440,221,247]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="nakasujii" status="sp. nov.">
<emphasis id="25ACEAD6FFD3571FD7BF94C0FE73FF1D" bold="true" box="[151,440,221,247]" italics="true" pageId="5" pageNumber="44">Pachycondyla nakasujii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E9F57ADFFD3571FD69794C0FDD6FF1D" box="[447,541,221,247]" pageId="5" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="176736C4FFD3571FD7BF951DFEDAFEF0" blockId="5.[151,541,221,282]" box="[151,273,256,282]" pageId="5" pageNumber="44">
(
<figureCitation id="8FE32A41FFD3571FD7B7951DFF20FEF0" box="[159,235,256,282]" captionStart="FIGURE 4" captionStartId="6.[151,255,1916,1940]" captionTargetBox="[163,1425,194,1894]" captionTargetId="figure@6.[163,1428,194,1896]" captionTargetPageId="6" captionText="FIGURE 4. Characters of worker propodeum and petiole. The left side of worker propodeum and petiole of P. nakasujii (A), P. chinensis (C) and P. luteipes (E), and the posterior face of worker petiole of P. nakasujii (B), P. chinensis (D) and P. luteipes (F) (Scanning electron microscopy photographs). Horizontal white arrow: PW. Also see Japanese Ant Database Group (2003)." httpUri="https://zenodo.org/record/199430/files/figure.png" pageId="5" pageNumber="44">Figs 4</figureCitation>
,
<figureCitation id="8FE32A41FFD3571FD7D0951DFECCFEF0" box="[248,263,256,282]" captionStart="FIGURE 5" captionStartId="7.[151,255,1975,1999]" captionTargetBox="[173,1391,194,1940]" captionTargetId="figure@7.[173,1409,194,1942]" captionTargetPageId="7" captionText="FIGURE 5. Characters of male coloration and mandible of P. nakasujii (A, B), P. chinensis (C, D) and P. luteipes (E, F). Black arrows indicate the mandible." httpUri="https://zenodo.org/record/199431/files/figure.png" pageId="5" pageNumber="44">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5FC2654FFFD35712D7BF9557FEDDFE86" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44" type="description">
<paragraph id="176736C4FFD3571FD7BF9557FD9EFB55" blockId="5.[151,1437,330,2036]" pageId="5" pageNumber="44">
<emphasis id="25ACEAD6FFD3571FD7BF9557FEC9FE8E" bold="true" box="[151,258,330,356]" pageId="5" pageNumber="44">Worker:</emphasis>
<emphasis id="25ACEAD6FFD3571FD6239557FE7DFE89" box="[267,438,330,355]" italics="true" pageId="5" pageNumber="44">Measurements</emphasis>
(mm)
<typeStatus id="C8638866FFD3571FD52F9557FDA5FE8E" box="[519,622,330,356]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
(
<typeStatus id="C8638866FFD3571FD5569557FD38FE8E" box="[638,755,330,356]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
, n = 15): HW 0.83 (0.79–0.85); HL 0.91 (0.88–0.94); SL 0.87 (0.83–0.92); EL 0.15 (0.14–0.18); WL 1.42 (1.36–1.47); MW 0.60 (0.59–0.64); PW 0.52 (0.49–0.55).
<emphasis id="25ACEAD6FFD3571FD7BF9588FECAFE44" box="[151,257,405,430]" italics="true" pageId="5" pageNumber="44">Structure</emphasis>
. Head slightly longer than wide, weakly round-sided behind eyes, with weakly emarginate posterior margin in full-face view. Mandible with 6 larger and 4 smaller teeth on the masticatory margin, and with distinct basal fovea. Antenna long, scape surpassing the posterior margin of head by less than 1/4 of its total length in full-face view; all funicular segments longer than wide. Pronotal dorsum bluntly demarcated from lateral face; seen from above anterolateral corner round. Mesonotum distinctly demarcated from pronotum and propodeum by deep furrows, posteriorly weakly emarginate. Mesopleuron with a somewhat vestigially transverse groove. In profile propodeum demarcated by a distinct obliquely running suture from metapleuron; posterior face well demarcated from lateral face by a rather sharp ridge, but separation from dorsal face indistinct. Petiole thin and high, with weakly convex dorsal margin in profile; in posterior view petiolar scale round and slightly higher than wide, with convex dorsal margin.
<emphasis id="25ACEAD6FFD3571FD48696F8FBEAFD14" box="[942,1057,741,766]" italics="true" pageId="5" pageNumber="44">Sculpture</emphasis>
. Dorsum of head minutely and densely punctate. Clypeus with much more superficial sculpture and shiny except for sculptured median portion. Mandible finely striatopunctate except for the area around apex and along mesal margin smooth with large sparse punctures. Dorsum of pronotum finely punctate; its lateral face and mesonotum with still finer punctation; mesopleuron almost smooth and shiny, with rugulae in lower area; upper portion of metapleuron punctate, with smooth interspaces, lower portion rugose, with smooth interspaces. Dorsolateral face of propodeum irregularly sculptured and matte, posterior face smooth except for margins and upper portion weakly sculputured. Petiole anteriorly micropunctate, posteriorly smooth. Gastral segments densely and weakly punctate evenly over the surface.
<emphasis id="25ACEAD6FFD3571FD5B2900DFD3DFBC3" box="[666,758,1040,1065]" italics="true" pageId="5" pageNumber="44">Pilosity</emphasis>
. Body covered with decumbent pubescence except for mesopleuron, metapleuron, and posterior faces of propodeum and petiole that are almost bare. Standing hairs sparse; gastral tergites 1 and 2 each with more than ten standing hairs.
<emphasis id="25ACEAD6FFD3571FD3249046FB58FB9E" box="[1036,1171,1115,1140]" italics="true" pageId="5" pageNumber="44">Coloration</emphasis>
. Body dark brown to blackish; gaster slightly lighter than head and mesosoma. Antennal funiculus, mandible, subpetiolar process and legs yellowish brown to orangish.
</paragraph>
<paragraph id="176736C4FFD3571FD7EE90D7FAE9FA00" blockId="5.[151,1437,330,2036]" pageId="5" pageNumber="44">
<emphasis id="25ACEAD6FFD3571FD7EE90D7FEEAFB0E" bold="true" box="[198,289,1226,1252]" pageId="5" pageNumber="44">Queen:</emphasis>
<emphasis id="25ACEAD6FFD3571FD60290D6FE5FFB0E" box="[298,404,1227,1252]" italics="true" pageId="5" pageNumber="44">Structure</emphasis>
. Body distinctly larger than the worker, with the head width ranging from
<quantity id="D0209B21FFD3571FD22690D7FF02FAE0" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.7" metricValueMin="8.6" pageId="5" pageNumber="44" unit="mm" value="0.865" valueMax="0.87" valueMin="0.86">0.86 to 0.87 mm</quantity>
. Eye much larger than the worker; EL =
<quantity id="D0209B21FFD3571FD58C90EDFC99FAE0" box="[676,850,1264,1290]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.5" metricValueMin="2.2" pageId="5" pageNumber="44" unit="mm" value="0.235" valueMax="0.25" valueMin="0.22">0.22–0.25 mm</quantity>
. Ocelli small, ca.
<quantity id="D0209B21FFD3571FD30A90EDFB41FAE0" box="[1058,1162,1264,1290]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="5" pageNumber="44" unit="mm" value="0.07">0.07 mm</quantity>
in maximum diameter; distance between posterior ocelli greater than that between anterior and posterior ocelli. Mesopleuron above always with a distinct transverse groove. Compared with the worker, petiole relatively thin, seen from above almost lacking dorsal flat portion.
<emphasis id="25ACEAD6FFD3571FD56A917DFD7DFA93" box="[578,694,1376,1401]" italics="true" pageId="5" pageNumber="44">Sculpture</emphasis>
. Body sculpture much as in the worker except for distinctly punctate metapleuron.
<emphasis id="25ACEAD6FFD3571FD69B9198FDDBFA74" box="[435,528,1413,1438]" italics="true" pageId="5" pageNumber="44">Pilosity</emphasis>
. Body covered with decumbent pubescence except for mesopleuron, and posterior faces of propodeum and petiole that are almost bare. Eye with short fine standing hairs. Mesosoma dorsally and gastral tergites with numerous long standing hairs.
<emphasis id="25ACEAD6FFD3571FD4AC91CDFBCEFA03" box="[900,1029,1488,1513]" italics="true" pageId="5" pageNumber="44">Coloration</emphasis>
. Much as in the worker.
</paragraph>
<paragraph id="176736C4FFD35712D7EE91E8FEDDFE86" blockId="5.[151,1437,330,2036]" lastBlockId="8.[151,1437,152,1447]" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44">
<emphasis id="25ACEAD6FFD3571FD7EE91E8FEC4F9E5" bold="true" box="[198,271,1525,1551]" pageId="5" pageNumber="44">Male:</emphasis>
<emphasis id="25ACEAD6FFD3571FD63291E8FE4FF9E4" box="[282,388,1525,1550]" italics="true" pageId="5" pageNumber="44">Structure</emphasis>
. Head including eyes much wider than long. Eye large; EL =
<quantity id="D0209B21FFD3571FD35591E8FAE6F9E5" box="[1149,1325,1525,1551]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="5" pageNumber="44" unit="mm" value="0.265" valueMax="0.27" valueMin="0.26">0.26–0.27 mm</quantity>
, strongly convex, mesally weakly emarginate. Clypeus transverse with anterior and posterior margins truncate; labrum anteriorly produced as a narrow lobe. Mandible long and fragile, not opposable, basally wide and suddenly narrowed toward bluntly pointed apex. Ocelli small, ca.
<quantity id="D0209B21FFD3571FD4689278FC60F995" box="[832,939,1637,1663]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="44" unit="mm" value="0.08">0.08 mm</quantity>
in maximum diameter; distance between posterior ocelli much greater than that between anterior and posterior ocelli. Antenna long; scape twice as long as pedicel and shorter than segment 3. Pronotum short; mesoscutum large, with weak notauli and sharp parapsidal lines; mesopleuron large, with a wide transverse furrow which is striate on the bottom; scutellum dorsally convex with an anterior transverse zone which is much lower than the main disc; axilla extensively concave; metanotum short. Propodeum rounded; dorsal, lateral and posterior faces not clearly differentiated. Petiole relatively low, in profile rather strongly narrowed above, seen from back almost as wide as high, with dorsal margin almost straight; subpetiolar process flat, with obtuse posterior tooth in profile.
<emphasis id="25ACEAD6FFD3571FD3C89376FA84F86E" box="[1248,1359,1899,1924]" italics="true" pageId="5" pageNumber="44">Sculpture</emphasis>
. Head superficially micropunctate. Pronotum, mesoscutum, metanotum and metapleuron weakly sculptured; mesopleuron extensively smooth and strongly shiny; axilla coarsely rugose. Propodeum more strongly and densely sculptured, but on its posterior face sculpture somewhat weaker. Anterior and posterior faces of petiole densely and irregularly sculptured; posterior face very weakly sculptured and somewhat shiny. Gaster only superficially sculptured.
<emphasis id="25ACEAD6FFDE5712D52994A0FD90FF3C" box="[513,603,189,214]" italics="true" pageId="8" pageNumber="47">Pilosity</emphasis>
. Body covered with decumbent pubescence except for mesopleuron, and posterior faces of propodeum and petiole that are almost bare. Petiole extensively with dense obliquely standing short hairs on its anterior and lateral faces; subpetiolar process below with standing hairs. Outer margin of forewing and hindwing with dense short fringe.
<emphasis id="25ACEAD6FFDE5712D4609530FC02FEAC" box="[840,969,301,326]" italics="true" pageId="8" pageNumber="47">Coloration</emphasis>
. Body brown to pale brown, with ivory white legs.
</paragraph>
</subSubSection>
<caption id="43A7664CFFD0571CD7BF9361FE53F819" httpUri="https://zenodo.org/record/199430/files/figure.png" pageId="6" pageNumber="45" targetBox="[163,1425,194,1894]" targetPageId="6">
<paragraph id="176736C4FFD0571CD7BF9361FE53F819" blockId="6.[151,1437,1916,2035]" pageId="6" pageNumber="45">
<emphasis id="25ACEAD6FFD0571CD7BF9361FED2F87E" bold="true" box="[151,281,1916,1940]" pageId="6" pageNumber="45">FIGURE 4.</emphasis>
Characters of worker propodeum and petiole. The left side of worker propodeum and petiole of
<taxonomicName id="D0D84D47FFD0571CD2379361FA57F879" box="[1311,1436,1916,1939]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFD0571CD2379361FAE7F879" box="[1311,1324,1916,1939]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD2129361FA57F879" box="[1338,1436,1916,1939]" italics="true" pageId="6" pageNumber="45">nakasujii</emphasis>
</taxonomicName>
(A),
<taxonomicName id="D0D84D47FFD0571CD7E19381FE8CF859" box="[201,327,1948,1971]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFD0571CD7E19381FF1DF859" box="[201,214,1948,1971]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD7CC9381FE8CF859" box="[228,327,1948,1971]" italics="true" pageId="6" pageNumber="45">chinensis</emphasis>
</taxonomicName>
(C) and
<taxonomicName id="D0D84D47FFD0571CD68D9381FDD8F859" box="[421,531,1948,1971]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFD0571CD68D9381FE79F859" box="[421,434,1948,1971]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD6E89381FDD8F859" box="[448,531,1948,1971]" italics="true" pageId="6" pageNumber="45">luteipes</emphasis>
</taxonomicName>
(E), and the posterior face of worker petiole of
<taxonomicName id="D0D84D47FFD0571CD3399381FB45F859" box="[1041,1166,1948,1971]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFD0571CD3399381FBD5F859" box="[1041,1054,1948,1971]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD3049381FB45F859" box="[1068,1166,1948,1971]" italics="true" pageId="6" pageNumber="45">nakasujii</emphasis>
</taxonomicName>
(B),
<taxonomicName id="D0D84D47FFD0571CD3ED9381FA88F859" box="[1221,1347,1948,1971]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFD0571CD3ED9381FB19F859" box="[1221,1234,1948,1971]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD3C89381FA88F859" box="[1248,1347,1948,1971]" italics="true" pageId="6" pageNumber="45">chinensis</emphasis>
</taxonomicName>
(D) and
<taxonomicName id="D0D84D47FFD0571CD7BF93A1FEC7F839" box="[151,268,1980,2003]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFD0571CD7BF93A1FF6EF839" box="[151,165,1980,2003]" italics="true" pageId="6" pageNumber="45">P</emphasis>
.
<emphasis id="25ACEAD6FFD0571CD79C93A1FEC7F839" box="[180,268,1980,2003]" italics="true" pageId="6" pageNumber="45">luteipes</emphasis>
</taxonomicName>
(F) (Scanning electron microscopy photographs). Horizontal white arrow: PW. Also see Japanese Ant Database Group (2003).
</paragraph>
</caption>
<caption id="43A7664CFFD1571DD7BF93AAFDD3F804" httpUri="https://zenodo.org/record/199431/files/figure.png" pageId="7" pageNumber="46" targetBox="[173,1391,194,1940]" targetPageId="7">
<paragraph id="176736C4FFD1571DD7BF93AAFDD3F804" blockId="7.[151,1436,1975,2030]" pageId="7" pageNumber="46">
<emphasis id="25ACEAD6FFD1571DD7BF93AAFED2F825" bold="true" box="[151,281,1975,1999]" pageId="7" pageNumber="46">FIGURE 5.</emphasis>
Characters of male coloration and mandible of
<taxonomicName id="D0D84D47FFD1571DD43C93AAFC5BF824" box="[788,912,1975,1998]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFD1571DD43C93AAFCEAF824" box="[788,801,1975,1998]" italics="true" pageId="7" pageNumber="46">P</emphasis>
.
<emphasis id="25ACEAD6FFD1571DD40793AAFC5BF824" box="[815,912,1975,1998]" italics="true" pageId="7" pageNumber="46">nakasujii</emphasis>
</taxonomicName>
(A, B),
<taxonomicName id="D0D84D47FFD1571DD4C093AAFBADF824" box="[1000,1126,1975,1998]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFD1571DD4C093AAFC3EF824" box="[1000,1013,1975,1998]" italics="true" pageId="7" pageNumber="46">P</emphasis>
.
<emphasis id="25ACEAD6FFD1571DD32B93AAFBADF824" box="[1027,1126,1975,1998]" italics="true" pageId="7" pageNumber="46">chinensis</emphasis>
</taxonomicName>
(C, D) and
<taxonomicName id="D0D84D47FFD1571DD3CC93AAFA9AF824" box="[1252,1361,1975,1998]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFD1571DD3CC93AAFB3AF824" box="[1252,1265,1975,1998]" italics="true" pageId="7" pageNumber="46">P</emphasis>
.
<emphasis id="25ACEAD6FFD1571DD3D693AAFA9AF824" box="[1278,1361,1975,1998]" italics="true" pageId="7" pageNumber="46">luteipes</emphasis>
</taxonomicName>
(E, F). Black arrows indicate the mandible.
</paragraph>
</caption>
<subSubSection id="5FC2654FFFDE5712D7EE956AFEAEFC9D" pageId="8" pageNumber="47" type="materials_examined">
<paragraph id="176736C4FFDE5712D7EE956AFBE8FDE8" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<emphasis id="25ACEAD6FFDE5712D7EE956AFDCBFE7B" bold="true" box="[198,512,375,401]" pageId="8" pageNumber="47">
<typeStatus id="C8638866FFDE5712D7EE956AFEC8FE7B" box="[198,259,375,401]" pageId="8" pageNumber="47">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="C8638866FFDE5712D5219565FDB0FE78" box="[521,635,376,402]" pageId="8" pageNumber="47" type="holotype">Holotype</typeStatus>
: worker,
<collectingCountry id="6FCF7654FFDE5712D5C19565FC8EFE78" box="[745,837,376,402]" name="Japan" pageId="8" pageNumber="47">JAPAN</collectingCountry>
, Okayama Prefecture, Okayama, Mt. Handayama,
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,
<geoCoordinate id="72EC5003FFDE5712D6109580FE13FE5D" box="[312,472,413,439]" direction="east" orientation="longitude" pageId="8" pageNumber="47" precision="15" value="133.92557">133°55’32”E</geoCoordinate>
,
<quantity id="D0209B21FFDE5712D6CD9580FDEEFE5D" box="[485,549,413,439]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="47" unit="m" value="22.0">22 m</quantity>
,
<date id="63661004FFDE5712D5199580FD6DFE5D" box="[561,678,413,439]" pageId="8" pageNumber="47" value="2007-06-05">5 vi 2007</date>
, (coll. T. Yashiro).
<typeStatus id="C8638866FFDE5712D4A69580FBCDFE5D" box="[910,1030,413,439]" pageId="8" pageNumber="47" type="paratype">Paratypes</typeStatus>
: 3 queens and
<specimenCount id="01DEFD4DFFDE5712D39F9580FAE5FE5D" box="[1207,1326,413,439]" pageId="8" pageNumber="47" type="worker">8 workers</specimenCount>
from the same locality,
<date id="63661004FFDE5712D66A95DFFE7EFE36" box="[322,437,450,476]" pageId="8" pageNumber="47" value="2007-06-05">5 vi 2007</date>
, (coll. T. Yashiro);
<specimenCount id="01DEFD4DFFDE5712D5B395DFFD3CFE36" box="[667,759,450,476]" pageId="8" pageNumber="47" type="male">3 males</specimenCount>
and
<specimenCount id="01DEFD4DFFDE5712D41995DFFC61FE36" box="[817,938,450,476]" pageId="8" pageNumber="47" type="worker">7 workers</specimenCount>
,
<collectingCountry id="6FCF7654FFDE5712D49E95DFFBD9FE36" box="[950,1042,450,476]" name="Japan" pageId="8" pageNumber="47">JAPAN</collectingCountry>
, Okayama Prefecture,
<collectingCountry id="6FCF7654FFDE5712D20895DFFAADFE36" box="[1312,1382,450,476]" name="Wake Island" pageId="8" pageNumber="47">Wake</collectingCountry>
, Mt. Tenjinyama,
<geoCoordinate id="72EC5003FFDE5712D60795F5FE0AFDE8" box="[303,449,488,514]" direction="north" orientation="latitude" pageId="8" pageNumber="47" precision="15" value="34.84889">34°50’56”N</geoCoordinate>
,
<geoCoordinate id="72EC5003FFDE5712D6E595F5FDA0FDE8" box="[461,619,488,514]" direction="east" orientation="longitude" pageId="8" pageNumber="47" precision="15" value="134.12973">134°07’47”E</geoCoordinate>
,
<quantity id="D0209B21FFDE5712D55F95F5FD0FFDE8" box="[631,708,488,514]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="8" pageNumber="47" unit="m" value="206.0">206 m</quantity>
,
<date id="63661004FFDE5712D5F895F5FC82FDE8" box="[720,841,488,514]" pageId="8" pageNumber="47" value="2008-07-04">4 vii 2008</date>
, (coll. T. Yashiro).
</paragraph>
<paragraph id="176736C4FFDE5712D7EE9610FD91FD56" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<emphasis id="25ACEAD6FFDE5712D7EE9610FE5CFDCD" bold="true" box="[198,407,525,551]" pageId="8" pageNumber="47">
<typeStatus id="C8638866FFDE5712D7EE9610FEC8FDCD" box="[198,259,525,551]" pageId="8" pageNumber="47">Type</typeStatus>
depository.
</emphasis>
The
<typeStatus id="C8638866FFDE5712D6F19610FD8BFDCD" box="[473,576,525,551]" pageId="8" pageNumber="47" type="holotype">holotype</typeStatus>
(NSMT-I-Hym 51990) is deposited in the National Museum of Nature and Science, Tokyo.
<typeStatus id="C8638866FFDE5712D6B9962FFDCFFDA6" box="[401,516,562,588]" pageId="8" pageNumber="47" type="paratype">Paratypes</typeStatus>
are deposited in the National Museum of Nature and Science, Tokyo,
<collectingCountry id="6FCF7654FFDE5712D278962FFA5CFDA6" box="[1360,1431,562,588]" name="Japan" pageId="8" pageNumber="47">Japan</collectingCountry>
; the Osaka Museum of Natural History, Osaka,
<collectingCountry id="6FCF7654FFDE5712D5E89645FCCDFD98" box="[704,774,600,626]" name="Japan" pageId="8" pageNumber="47">Japan</collectingCountry>
; the Museo Civico di Storia Naturale “Giacomo Doria”, Genova,
<collectingCountry id="6FCF7654FFDE5712D62C9660FEF4FD7D" box="[260,319,637,663]" name="Italy" pageId="8" pageNumber="47">Italy</collectingCountry>
; the Natural History Museum, London,
<collectingCountry id="6FCF7654FFDE5712D40D9660FC46FD7D" box="[805,909,637,663]" name="United Kingdom" pageId="8" pageNumber="47">England</collectingCountry>
; and the Museum of Comparative Zoology, Harvard University, Cambridge,
<collectingCountry id="6FCF7654FFDE5712D53F96BFFD9DFD56" box="[535,598,674,700]" name="United States of America" pageId="8" pageNumber="47">USA</collectingCountry>
.
</paragraph>
<paragraph id="176736C4FFDE5712D7EE96DAFEAEFC9D" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<emphasis id="25ACEAD6FFDE5712D7EE96DAFD96FD0B" bold="true" box="[198,605,711,737]" pageId="8" pageNumber="47">Additional material examined.</emphasis>
<specimenCount id="01DEFD4DFFDE5712D55996D5FD3EFD08" box="[625,757,712,738]" pageId="8" pageNumber="47" type="worker">5 workers</specimenCount>
,
<collectingCountry id="6FCF7654FFDE5712D42596D5FCA2FD08" box="[781,873,712,738]" name="Japan" pageId="8" pageNumber="47">JAPAN</collectingCountry>
, Kanagawa Prefecture, Yokosuka, Hayama,
<geoCoordinate id="72EC5003FFDE5712D7BF96F0FEE0FCED" box="[151,299,749,775]" direction="north" orientation="latitude" pageId="8" pageNumber="47" precision="15" value="35.260277">35°15’37”N</geoCoordinate>
,
<geoCoordinate id="72EC5003FFDE5712D61196F0FE12FCED" box="[313,473,749,775]" direction="east" orientation="longitude" pageId="8" pageNumber="47" precision="15" value="139.6064">139°36’23”E</geoCoordinate>
,
<quantity id="D0209B21FFDE5712D6CE96F0FDFDFCED" box="[486,566,749,775]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="47" unit="m" value="150.0">150 m</quantity>
,
<date id="63661004FFDE5712D56B96F0FD7BFCED" box="[579,688,749,775]" pageId="8" pageNumber="47" value="2008-05-07">7 v 2008</date>
, (coll. M. Yashiro); 1 queens and
<specimenCount id="01DEFD4DFFDE5712D36296F0FB0FFCED" box="[1098,1220,749,775]" pageId="8" pageNumber="47" type="worker">5 workers</specimenCount>
,
<collectingCountry id="6FCF7654FFDE5712D3F996F0FAE5FCED" box="[1233,1326,749,775]" name="Japan" pageId="8" pageNumber="47">JAPAN</collectingCountry>
, Kagawa Prefecture, Takamatsu, Nishiueta,
<geoCoordinate id="72EC5003FFDE5712D501970FFD70FCC6" box="[553,699,786,812]" direction="north" orientation="latitude" pageId="8" pageNumber="47" precision="15" value="34.231388">34°13’53”N</geoCoordinate>
,
<geoCoordinate id="72EC5003FFDE5712D5EF970FFC9DFCC6" box="[711,854,786,812]" direction="east" orientation="longitude" pageId="8" pageNumber="47" precision="15" value="134.07722">134°4’38”E</geoCoordinate>
,
<quantity id="D0209B21FFDE5712D44A970FFC6BFCC6" box="[866,928,786,812]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="47" unit="m" value="86.0">86 m</quantity>
,
<date id="63661004FFDE5712D483970FFBEFFCC6" box="[939,1060,786,812]" pageId="8" pageNumber="47" value="2007-10-14">14 x 2007</date>
, (coll. T. Yashiro); 1 queens and
<specimenCount id="01DEFD4DFFDE5712D7BF9725FEC5FCB8" box="[151,270,824,850]" pageId="8" pageNumber="47" type="worker">5 workers</specimenCount>
,
<collectingCountry id="6FCF7654FFDE5712D6329725FEBDFCB8" box="[282,374,824,850]" name="Japan" pageId="8" pageNumber="47">JAPAN</collectingCountry>
, Fukuoka Prefecture, Tagawa, Mt. Hikosan,
<geoCoordinate id="72EC5003FFDE5712D4A89725FBD9FCB8" box="[896,1042,824,850]" direction="north" orientation="latitude" pageId="8" pageNumber="47" precision="15" value="33.481113">33°28’52”N</geoCoordinate>
,
<geoCoordinate id="72EC5003FFDE5712D3369725FB77FCB8" box="[1054,1212,824,850]" direction="east" orientation="longitude" pageId="8" pageNumber="47" precision="15" value="130.91945">130°55’10”E</geoCoordinate>
,
<quantity id="D0209B21FFDE5712D3EF9725FADEFCB8" box="[1223,1301,824,850]" metricMagnitude="2" metricUnit="m" metricValue="9.94" pageId="8" pageNumber="47" unit="m" value="994.0">994 m</quantity>
,
<date id="63661004FFDE5712D2089725FA5CFCB8" box="[1312,1431,824,850]" pageId="8" pageNumber="47" value="2007-10-11">11 x 2007</date>
, (coll. T. Yashiro).
</paragraph>
</subSubSection>
<subSubSection id="5FC2654FFFDE5712D7EE979FFC2AFC0D" pageId="8" pageNumber="47" type="etymology">
<paragraph id="176736C4FFDE5712D7EE979FFC2AFC0D" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<emphasis id="25ACEAD6FFDE5712D7EE979FFE98FC76" bold="true" box="[198,339,898,924]" pageId="8" pageNumber="47">Etymology.</emphasis>
This species is named in honor of Professor Emeritus Fusao Nakasuji, a notable Japanese entomologist and an authority on integrated pest management, who supervised the undergraduate and master's theses of one of the authors (Toshihisa Yashiro) at Okayama University.
</paragraph>
</subSubSection>
<subSubSection id="5FC2654FFFDE5712D7EE97EFFDB2FA4D" pageId="8" pageNumber="47" type="discussion">
<paragraph id="176736C4FFDE5712D7EE97EFFD40FB06" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<emphasis id="25ACEAD6FFDE5712D7EE97EFFE8BFBE6" bold="true" box="[198,320,1010,1036]" pageId="8" pageNumber="47">Remarks.</emphasis>
This species belongs to the
<taxonomicName id="D0D84D47FFDE5712D5A497EFFC6CFBE1" box="[652,935,1010,1035]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D5A497EFFC6CFBE1" box="[652,935,1010,1035]" italics="true" pageId="8" pageNumber="47">Pachycondyla chinensis</emphasis>
</taxonomicName>
species complex. It is easily distinguished from the sympatric species
<taxonomicName id="D0D84D47FFDE5712D6F69005FDA1FBDB" box="[478,618,1048,1074]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D6F69005FE26FBDB" box="[478,493,1048,1073]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D6D49005FDA1FBDB" box="[508,618,1048,1073]" italics="true" pageId="8" pageNumber="47">chinensis</emphasis>
</taxonomicName>
in the worker by having a proportionally wider petiole (PI [Pronotal index: PW/HW x 100] =
<quantity id="D0209B21FFDE5712D6E09020FDF9FBBD" box="[456,562,1085,1111]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" metricValueMax="1.6256" metricValueMin="1.524" pageId="8" pageNumber="47" unit="in" value="62.0" valueMax="64.0" valueMin="60.0">60–64 in</quantity>
<taxonomicName id="D0D84D47FFDE5712D5139020FD00FBBC" box="[571,715,1085,1111]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFDE5712D5139020FD81FBBC" box="[571,586,1085,1110]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D5729020FD00FBBC" box="[602,715,1085,1110]" italics="true" pageId="8" pageNumber="47">nakasujii</emphasis>
</taxonomicName>
, PI =
<quantity id="D0209B21FFDE5712D43C9020FCB5FBBD" box="[788,894,1085,1111]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" metricValueMax="1.4731999999999998" metricValueMin="1.2192" pageId="8" pageNumber="47" unit="in" value="53.0" valueMax="58.0" valueMin="48.0">48–58 in</quantity>
<taxonomicName id="D0D84D47FFDE5712D4AF9020FBDEFBBC" box="[903,1045,1085,1111]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D4AF9020FC5DFBBC" box="[903,918,1085,1110]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D48D9020FBDEFBBC" box="[933,1045,1085,1110]" italics="true" pageId="8" pageNumber="47">chinensis</emphasis>
</taxonomicName>
) (
<figureCitation id="8FE32A41FFDE5712D3199020FBB1FBBD" box="[1073,1146,1085,1111]" captionStart="FIGURE 4" captionStartId="6.[151,255,1916,1940]" captionTargetBox="[163,1425,194,1894]" captionTargetId="figure@6.[163,1428,194,1896]" captionTargetPageId="6" captionText="FIGURE 4. Characters of worker propodeum and petiole. The left side of worker propodeum and petiole of P. nakasujii (A), P. chinensis (C) and P. luteipes (E), and the posterior face of worker petiole of P. nakasujii (B), P. chinensis (D) and P. luteipes (F) (Scanning electron microscopy photographs). Horizontal white arrow: PW. Also see Japanese Ant Database Group (2003)." httpUri="https://zenodo.org/record/199430/files/figure.png" pageId="8" pageNumber="47">Fig. 4</figureCitation>
B,D) and in the male by having much darker coloration and the well developed mandibles (
<figureCitation id="8FE32A41FFDE5712D486907FFC3FFB96" box="[942,1012,1122,1148]" captionStart="FIGURE 5" captionStartId="7.[151,255,1975,1999]" captionTargetBox="[173,1391,194,1940]" captionTargetId="figure@7.[173,1409,194,1942]" captionTargetPageId="7" captionText="FIGURE 5. Characters of male coloration and mandible of P. nakasujii (A, B), P. chinensis (C, D) and P. luteipes (E, F). Black arrows indicate the mandible." httpUri="https://zenodo.org/record/199431/files/figure.png" pageId="8" pageNumber="47">Fig. 5</figureCitation>
A–D). Moreover, the worker of this species is easily distinguished from another closely related species,
<taxonomicName id="D0D84D47FFDE5712D4E69095FB82FB4B" box="[974,1097,1160,1185]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFDE5712D4E69095FB82FB4B" box="[974,1097,1160,1185]" italics="true" pageId="8" pageNumber="47">P. luteipes</emphasis>
</taxonomicName>
, by having a proportionally wider petiole (PI =
<quantity id="D0209B21FFDE5712D65790B0FE2CFB2D" box="[383,487,1197,1223]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" metricValueMax="1.6256" metricValueMin="1.524" pageId="8" pageNumber="47" unit="in" value="62.0" valueMax="64.0" valueMin="60.0">60–64 in</quantity>
<taxonomicName id="D0D84D47FFDE5712D6C790B0FDB0FB2C" box="[495,635,1197,1223]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFDE5712D6C790B0FE35FB2C" box="[495,510,1197,1222]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D52590B0FDB0FB2C" box="[525,635,1197,1222]" italics="true" pageId="8" pageNumber="47">nakasujii</emphasis>
</taxonomicName>
, PI =
<quantity id="D0209B21FFDE5712D5EC90B0FCE7FB2D" box="[708,812,1197,1223]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.4731999999999998" metricValueMin="1.3208000000000002" pageId="8" pageNumber="47" unit="in" value="55.0" valueMax="58.0" valueMin="52.0">52–58 in</quantity>
<taxonomicName id="D0D84D47FFDE5712D41C90B0FC66FB2C" box="[820,941,1197,1223]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFDE5712D41C90B0FC88FB2C" box="[820,835,1197,1222]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D47A90B0FC66FB2C" box="[850,941,1197,1222]" italics="true" pageId="8" pageNumber="47">luteipes</emphasis>
</taxonomicName>
) (
<figureCitation id="8FE32A41FFDE5712D4E090B0FBDBFB2D" box="[968,1040,1197,1223]" captionStart="FIGURE 4" captionStartId="6.[151,255,1916,1940]" captionTargetBox="[163,1425,194,1894]" captionTargetId="figure@6.[163,1428,194,1896]" captionTargetPageId="6" captionText="FIGURE 4. Characters of worker propodeum and petiole. The left side of worker propodeum and petiole of P. nakasujii (A), P. chinensis (C) and P. luteipes (E), and the posterior face of worker petiole of P. nakasujii (B), P. chinensis (D) and P. luteipes (F) (Scanning electron microscopy photographs). Horizontal white arrow: PW. Also see Japanese Ant Database Group (2003)." httpUri="https://zenodo.org/record/199430/files/figure.png" pageId="8" pageNumber="47">Fig. 4</figureCitation>
B,F) and a sculptured propodeum (largely smooth in
<taxonomicName id="D0D84D47FFDE5712D65C90CEFE27FB06" box="[372,492,1235,1260]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="25ACEAD6FFDE5712D65C90CEFE27FB06" box="[372,492,1235,1260]" italics="true" pageId="8" pageNumber="47">P. luteipes</emphasis>
</taxonomicName>
) (
<figureCitation id="8FE32A41FFDE5712D52E90CFFD87FB06" box="[518,588,1234,1260]" captionStart="FIGURE 4" captionStartId="6.[151,255,1916,1940]" captionTargetBox="[163,1425,194,1894]" captionTargetId="figure@6.[163,1428,194,1896]" captionTargetPageId="6" captionText="FIGURE 4. Characters of worker propodeum and petiole. The left side of worker propodeum and petiole of P. nakasujii (A), P. chinensis (C) and P. luteipes (E), and the posterior face of worker petiole of P. nakasujii (B), P. chinensis (D) and P. luteipes (F) (Scanning electron microscopy photographs). Horizontal white arrow: PW. Also see Japanese Ant Database Group (2003)." httpUri="https://zenodo.org/record/199430/files/figure.png" pageId="8" pageNumber="47">Fig. 4</figureCitation>
A,E).
</paragraph>
<paragraph id="176736C4FFDE5712D7EE90E4FDB2FA4D" blockId="8.[151,1437,152,1447]" pageId="8" pageNumber="47">
<taxonomicName id="D0D84D47FFDE5712D7EE90E4FE15FAFB" box="[198,478,1272,1297]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFDE5712D7EE90E4FE15FAFB" box="[198,478,1272,1297]" italics="true" pageId="8" pageNumber="47">Pachycondyla nakasujii</emphasis>
</taxonomicName>
nests in rotten wood or leaf litter on the ground. It is common in Honshu, Shikoku, and Kyushu,
<collectingCountry id="6FCF7654FFDE5712D6839100FE39FADD" box="[427,498,1309,1335]" name="Japan" pageId="8" pageNumber="47">Japan</collectingCountry>
, where it is sympatric with
<taxonomicName id="D0D84D47FFDE5712D46A9100FC1AFADC" box="[834,977,1309,1335]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D46A9100FC9AFADC" box="[834,849,1309,1334]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D4499100FC1AFADC" box="[865,977,1309,1334]" italics="true" pageId="8" pageNumber="47">chinensis</emphasis>
</taxonomicName>
(
<figureCitation id="8FE32A41FFDE5712D4CA9100FBE7FADD" box="[994,1068,1309,1335]" captionStart="FIGURE 6" captionStartId="9.[151,255,984,1008]" captionTargetBox="[158,1428,313,956]" captionTargetId="figure@9.[156,1431,306,963]" captionTargetPageId="9" captionText="FIGURE 6. Currently known distribution of P. nakasujii (A) and P. chinensis (B) in Japan. Each dot may indicate more than one site if they are located less than 20 km apart." httpUri="https://zenodo.org/record/199432/files/figure.png" pageId="8" pageNumber="47">Fig. 6</figureCitation>
). Although
<taxonomicName id="D0D84D47FFDE5712D3959100FA81FADC" box="[1213,1354,1309,1335]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFDE5712D3959100FB07FADC" box="[1213,1228,1309,1334]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D3F49100FA81FADC" box="[1244,1354,1309,1334]" italics="true" pageId="8" pageNumber="47">nakasujii</emphasis>
</taxonomicName>
and
<taxonomicName id="D0D84D47FFDE5712D2AD9100FECFFAB6" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D2AD9100FA5FFADC" box="[1413,1428,1309,1334]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D7BF915EFECFFAB6" box="[151,260,1347,1372]" italics="true" pageId="8" pageNumber="47">chinensis</emphasis>
</taxonomicName>
are sympatrically distributed in temperate zones,
<taxonomicName id="D0D84D47FFDE5712D460915EFC1BFAB6" box="[840,976,1346,1372]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="nakasujii">
<emphasis id="25ACEAD6FFDE5712D460915EFC9DFAB6" box="[840,854,1347,1372]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D44C915EFC1BFAB6" box="[868,976,1347,1372]" italics="true" pageId="8" pageNumber="47">nakasujii</emphasis>
</taxonomicName>
is rare in dry and disturbed area, where
<taxonomicName id="D0D84D47FFDE5712D7BF9175FEE8FA6B" box="[151,291,1384,1410]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="25ACEAD6FFDE5712D7BF9175FF6DFA6B" box="[151,166,1384,1409]" italics="true" pageId="8" pageNumber="47">P</emphasis>
.
<emphasis id="25ACEAD6FFDE5712D79D9175FEE8FA6B" box="[181,291,1384,1409]" italics="true" pageId="8" pageNumber="47">chinensis</emphasis>
</taxonomicName>
remains common. Both species have been observed to nest near termite nests frequently and to carry termites back to their nests as food.
</paragraph>
</subSubSection>
</treatment>
</document>