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<mods:title id="ABDA7CC85CFBDD5C580EEC48EAB7516C">The ant tribe Dacetini. With a revision of the Strumigenys species of the Malgasy Region by Brian L. Fisher, and a revision of the Austral epopostrumiform genera by Steven O. Shattuck.</mods:title>
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<paragraph id="9CA5E8CCA94F7814E175CC0E454CFB73" pageNumber="641">
<taxonomicName id="AF19246A9FF590F4A6001461AB42B7D1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="641" phylum="Arthropoda" rank="species" species="dexis">Strumigenys dexis Fisher</taxonomicName>
<taxonomicNameLabel id="4D921D6ED43C4BF7EEF287AA56080004" rank="species">sp. n.</taxonomicNameLabel>
(Figs 387, 412)
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<paragraph id="EECE76FC0ADF2838B51440FFEEA706A5" pageNumber="641">HOLOTYPE WORKER. TL 2.4, HL 0.66, HW 0.57, CI 88, ML 0.12, MI 18, SL 0.29, SI 51, PW 0.36, AL 0.68. Characters of &lt;fem-complex. Transverse edge of the depressed proximodorsal area of mandible with a short row of curved spoon-shaped hairs approximately the same as those fringing the upper scrobe margin. Each labral lobe terminates anteriorly in a broad fan-shaped lamella of thin translucent cuticle and a short trigger-hair mesad of fan-like lamella; trigger-hairs often difficult to see. Clypeal dorsum with minute appressed hairs, the lateral margins with a short row of anteriorly curved large spoon-shaped hairs. Upper scrobe margin in full-face view with a continuous row of anteriorly curved spoon-shaped hairs that are similar, to those on the leading edge of the scape; the row ends at the apex of the scrobe and does not extend onto the side of the occipital lobe. Eye with 7 ommatidia in total. Cephalic dorsum without standing hairs of any description but with scattered minute flattened appressed hairs present. Area of cuticle within antennal scrobe densely reticulate-punctate except for extreme posterior portion which is superficially reticulate to smooth. Dorsum of head behind clypeus with separated shallow punctures, the spaces between them shining; ventral surface of head unsculptured. Pronotum in dorsal view with strongly convex sides, appearing very short and broad. Flattened dorsum of pronotum sculptured as vertex; pleurae and sides of propodeum smooth and shining; propodeal declivity with a band of fine punctate sculpture between the teeth. Lamella of propodeum broad and sinuate, apices of propodeal teeth free. Dorsal surfaces of alitrunk, waist segments and first gastral tergite without standing hairs, but minute appressed flattened hairs are sparsely present everywhere except on first gastral tergite. Base of first gastral sternite and posterior margins of petiole and postpetiole, with a row of flat, broad, spatulate appressed hairs. Petiole node in dorsal view broader than long, with extensive lateral spongiform lobes. Disc of postpetiole completely surrounded by spongiform tissue in dorsal view; sides and posterior margin of the disc forming a single convex arc and the disc with scattered but easily visible punctures upon a smooth surface. In profile all spongiform lobes of the waist segments well developed. Legs with spatulate to scale-like hairs present, but without standing pilosity.</paragraph>
<paragraph id="F77516D772C25B850FF4734EA5501174" pageNumber="642">PARATYPE WORKERS. TL 2.4 - 2.5, HL 0.65 - 0.66, HW 0.58 - 0.59, CI 88 - 90, ML 0.12 - 0.13, MI 18 - 20, SL 0.29, SI 49 - 50, PW 0.36 - 0.37, AL 0.68 (2 measured). As holotype.</paragraph>
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<subSubSection id="14BADD811C164439E3BE5FB28B718437" type="materials_examined">
<paragraph id="B87509FF18D98AAD3EDCE1946BA4D7E6" pageNumber="642">Holotype worker, Madagascar: 45 km. S Ambalavao, 22 ° 13 ' S, 47 ° 01 ' E, 785 m., 25. ix. 1993, sifted litter (leaf mold, rotten wood), rainforest, # 696 (40) - 6 (B. L. Fisher) (MCZ).</paragraph>
<paragraph id="DF8B5DAC593149BBCD12B54D62D2BD04" pageNumber="642">Paratypes. 1 worker with same data as holotype; 1 worker with same data but # 696 (20) - 9 (SAM, BMNH).</paragraph>
<paragraph id="84F05488E508207A71C655B1BF72A1B2" pageNumber="642">NON-PARATYPIC MATERIAL EXAMINED. Madagascar: 6.5 km. SSW Befingotra, Ris. Anjanaharibe-Sud 875 m. (B. L. Fisher); 6.9 km. NE Ambanizana, Ambohitsitondroina 825 m. (B. L. Fisher); S. F. Tampolo, 10 km. NNE Fenoarivo Atn. 10 m. (B. L. Fisher); Rte. d'Anosibe, km. 33 (A. Peyrieras); F. C. Sandranantitra 450 m. (H. I. Ratsirarson); F. C. Andriantantely 530 m. (H. J. Ratsirarson); Ranomafana Nat. Park, Vohiparara forest 1160 m. (E. Rajeriarison); 43 km. S Ambalavao, Ris. Andringitra 825 m. (B. L. Fisher); 9.0 km. NE Ivohibe 900 m. (S. Razqfimandimby); R. S. Ivohibe 8.0 km. E Ivohibe 1200 m. (5. Razafimandimby); 11 km. NW Enakara, Ris. Andohahela 800 m. (B. L. Fisher).</paragraph>
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<subSubSection id="E7FC2BFDCBC65C055251464CC1B1F949" type="description">
<paragraph id="A11F5E0C223F4E0300ED756AFCBF715C" pageNumber="642">Members of these series tend to be a little larger than the type-series and to have the head fractionally broader, HL 0.68 - 0.70, HW 0.63 - 0.64, CI 90 - 93, ML 0.12 - 0.13, MI 17 - 19, but otherwise match the types perfectly.</paragraph>
</subSubSection>
<subSubSection id="8521403AAE5BB9BAE5B3522B648BA8C0" type="diagnosis">
<paragraph id="55143AE2EC91157DA475BC8F52E9FA39" pageNumber="642">
Within the group
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is related to
<taxonomicName id="EDC97D3CF8E5D2F1CCB0975B1794C66C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155822" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys coveri Fisher" lsidName-HNS="Strumigenys coveri Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="coveri">coveri</taxonomicName>
,
<taxonomicName id="C961B154B93EFA3CBBA9564EFD8D53B7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155824" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys gorgon Fisher" lsidName-HNS="Strumigenys gorgon Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="gorgon">gorgon</taxonomicName>
, peyrierasi and rabesoni, a complex of five species that share a distinctive form of sculpture in which cephalic and promesonotal punctures are spaced out and separated by areas of smooth cuticle, rather than having the entire surface opaque and blanketed with reticulate- punctate sculpture (promesonotal punctures absent in
<taxonomicName id="0B299B3C799BA9FCE76BB1C7D690E1E1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155821" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys carolinae Fisher" lsidName-HNS="Strumigenys carolinae Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="carolinae">carolinae</taxonomicName>
). Of these five species
<taxonomicName id="98A398A72F4A05851EAC4313F14B70AD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
is identified by its relatively short mandibles, broad head and unique labrum, where both anterior lobes sport a single fan-like lamella of very thin translucent cuticle that projects anteriorly. In addition:
</paragraph>
<paragraph id="BB176A14A03B58811238FBEA9EC48A41" pageNumber="642">
1 The petiole node in
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is as long as or longer than broad in dorsal view (distinctly broader than long in
<taxonomicName id="E1792E1C3C18EE6BB9B7E3510C1C63F3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
).
</paragraph>
<paragraph id="E05DF335DD17D43BED024EE961200D9B" pageNumber="642">
2 The head has large spoon-shaped hairs on the dorsum of the occipital lobes in
<taxonomicName id="6D47BFEDF603EA61A5E041558E6CB410" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155824" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys gorgon Fisher" lsidName-HNS="Strumigenys gorgon Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="gorgon">gorgon</taxonomicName>
(absent in
<taxonomicName id="ED450C28F29FBB68FC99AF87B3DD2D85" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
).
</paragraph>
<paragraph id="DA8D467BA8ED3B83F34E7E7D6D601F61" pageNumber="642">
3 The punctate sculpture of the vertex is much denser and more crowded in rabesoni than in
<taxonomicName id="87C5AB1B288F6626DDB49445D46B2C77" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
.
</paragraph>
<paragraph id="817A062F5794A82579896AD109636E5A" pageNumber="642">
4 The mesonotum has a single pair of short straight standing hairs in peyrierasi (absent in
<taxonomicName id="82D2DE6BA1BD63774CBD419F29FC26C5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
).
</paragraph>
<paragraph id="326BC5023F3019841E4F4B37D988D60F" pageNumber="642">
The presence of dense blanketing reticulate-punctate sculpture on the dorsum of the head distinguishes the sylvaini-complex from
<taxonomicName id="1C72C7F1D7AA09069EDA737BC04123C2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155823" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys dexis Fisher" lsidName-HNS="Strumigenys dexis Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="dexis">dexis</taxonomicName>
and its immediate allies discussed above. Beside the sculpture
<taxonomicName id="9EEF439687184C8695DE9BCDA67A4632" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155820" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys alperti Fisher" lsidName-HNS="Strumigenys alperti Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="alperti">alperti</taxonomicName>
and
<taxonomicName id="BD43909E88AFC66128B46A0510D4D63C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:155821" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strumigenys carolinae Fisher" lsidName-HNS="Strumigenys carolinae Fisher" order="Hymenoptera" pageNumber="642" phylum="Arthropoda" rank="species" species="carolinae">carolinae</taxonomicName>
have a unique distribution of scale-like hairs anteriorly on the cephalic dorsum, and sylvaini and wardi have standing hairs on the head and body, neither of which occurs in the four species of the (feus-complex.
</paragraph>
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<subSubSection id="5F816568EE48FFDF45A9F9FCFDF44439" box="[151,601,1581,1608]" pageId="0" pageNumber="191" type="nomenclature">
<paragraph id="172436E3EE48FFDF45A9F9FCFDF44439" blockId="0.[151,1437,1581,2039]" box="[151,601,1581,1608]" pageId="0" pageNumber="191">
<heading id="4C6C818FEE48FFDF45A9F9FCFDF44439" bold="true" box="[151,601,1581,1608]" fontSize="11" level="1" pageId="0" pageNumber="191" reason="1">
<emphasis id="25EFEAF1EE48FFDF45A9F9FCFDF44439" bold="true" box="[151,601,1581,1608]" pageId="0" pageNumber="191">
<taxonomicName id="D09B4D60EE48FFDF45A9F9FCFED34436" ID-CoL="7DZPG" authority="Lu &amp; Wang, 2019" authorityName="Lu &amp; Wang" authorityYear="2019" box="[151,382,1581,1608]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="0" pageNumber="191" phylum="Arthropoda" rank="species" species="probolus" status="sp. nov.">
<emphasis id="25EFEAF1EE48FFDF45A9F9FCFED34436" bold="true" box="[151,382,1581,1608]" italics="true" pageId="0" pageNumber="191">Urothrips probolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EDC578AEE48FFDF44BBF9FFFE6A4436" box="[389,455,1582,1608]" pageId="0" pageNumber="191" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="8FA02A66EE48FFDF44E8F9FCFDE24439" box="[470,591,1581,1607]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 610" captionStartId-0="1.[151,258,2003,2025]" captionStartId-1="2.[151,258,1864,1886]" captionTargetBox-0="[159,1429,182,1982]" captionTargetBox-1="[151,1435,597,1844]" captionTargetId-0="figure@1.[159,1429,182,1982]" captionTargetId-1="figure@2.[151,1435,597,1844]" captionTargetPageId-0="1" captionText-0="FIGURES 15. Urothrips humpus sp. n. (female): (1) Habitus, (2) head, (3) antenna, (4) abdominal tergites IV, (5) fore legs" captionText-1="FIGURES 610. Urothrips humpus sp. n. (female): (6) pronotum, (7) prosternum, (8) meso- and metanotum, (9) meso- and metaster- num, (10) abdominal sternum IV" httpUri-0="https://zenodo.org/record/3241083/files/figure.png" httpUri-1="https://zenodo.org/record/3404906/files/figure.png" pageId="0" pageNumber="191">Figs 110</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5F816568EE48FFDC45A9F982FED14360" lastPageId="3" lastPageNumber="194" pageId="0" pageNumber="191" type="description">
<paragraph id="172436E3EE48FFDF45A9F982FB8E4561" blockId="0.[151,1437,1581,2039]" pageId="0" pageNumber="191">
<emphasis id="25EFEAF1EE48FFDF45A9F982FE904415" bold="true" box="[151,317,1619,1643]" pageId="0" pageNumber="191">Female aptera</emphasis>
. Body (
<figureCitation id="8FA02A66EE48FFDF44A7F982FE744415" box="[409,473,1619,1643]" captionStart="FIGURES 15" captionStartId="1.[151,258,2003,2025]" captionTargetBox="[159,1429,182,1982]" captionTargetId="figure@1.[159,1429,182,1982]" captionTargetPageId="1" captionText="FIGURES 15. Urothrips humpus sp. n. (female): (1) Habitus, (2) head, (3) antenna, (4) abdominal tergites IV, (5) fore legs" httpUri="https://zenodo.org/record/3241083/files/figure.png" pageId="0" pageNumber="191">Fig. 1</figureCitation>
) bicolored, brown and yellowish white; head, pro- and mesothorax brown, but anterior half of prothorax yellowish white, metathorax and abdominal segments IVIII yellowish white and with brown shadings laterally and medially, segment IX brownish with longitudinal brown shadings on both lateral and anterior a third yellowish, tube yellow with apex brown; antennal segments IIII yellow, segment IV brownish yellow and gradually darkened to apex, segments VVI brown; fore legs yellowish white but coxae brown, mid legs yellowish white but shaded with brownish and coxae brown, hind leg brownish, all tarsi yellow with brown markings.
</paragraph>
<paragraph id="172436E3EE48FFDD45F9F8FAFE9243F7" blockId="0.[151,1437,1581,2039]" lastBlockId="2.[151,1437,154,574]" lastPageId="2" lastPageNumber="193" pageId="0" pageNumber="191">
Head (
<figureCitation id="8FA02A66EE48FFDF442FF8FAFEFE453D" box="[273,339,1835,1859]" captionStart="FIGURES 15" captionStartId="1.[151,258,2003,2025]" captionTargetBox="[159,1429,182,1982]" captionTargetId="figure@1.[159,1429,182,1982]" captionTargetPageId="1" captionText="FIGURES 15. Urothrips humpus sp. n. (female): (1) Habitus, (2) head, (3) antenna, (4) abdominal tergites IV, (5) fore legs" httpUri="https://zenodo.org/record/3241083/files/figure.png" pageId="0" pageNumber="191">Fig. 2</figureCitation>
) longer than wide, gradually widened towards base, cheeks setae small and blunt, vertex prolonged over antennal bases as a prominent protuberance, dorsal surface and cheeks with numerous small tubercles, without long anterior cephalic setae; eyes each with 6 ommatidia dorsally, ocelli absent. Antennae (
<figureCitation id="8FA02A66EE48FFDF4107F8A2FBD545F5" box="[1081,1144,1907,1931]" captionStart="FIGURES 15" captionStartId="1.[151,258,2003,2025]" captionTargetBox="[159,1429,182,1982]" captionTargetId="figure@1.[159,1429,182,1982]" captionTargetPageId="1" captionText="FIGURES 15. Urothrips humpus sp. n. (female): (1) Habitus, (2) head, (3) antenna, (4) abdominal tergites IV, (5) fore legs" httpUri="https://zenodo.org/record/3241083/files/figure.png" pageId="0" pageNumber="191">Fig. 3</figureCitation>
) 6-segmented, segment IV longest and with faint sutures between the morphological segments, VI without sutures; maxillary stylets retracted to compound eyes, subparallel and their nearest distance about one tenth of the head width, without maxillary bridge; mouth cone rounded, reaching prosternum medially Pronotum (Fig. 6) about 1.7 times as wide as long, dorsal surface with small tubercles and each with a setae blunt at apex (see a red arrow in Fig. 6); epimeral setae well-developed and somewhat dilated at apex; epimeral suture absent. Basantra reduced to two small triangle plates laterally, each with 2 pairs of setae; ferna present (Fig. 7), mesopresternum narrow and boat-shaped. Meso- and metanotum (Fig. 8) with many small tubercles, each with a small setae blunt at apex (see red arrows in Fig. 8); mesothoracic furcae incompletely fused together medially, but metathoracic furcae widely separated (Fig. 9). Legs short with some tubercles, each with minute setae; fore tarsus with a hook-like hamus on external margin (Fig. 5).
</paragraph>
<paragraph id="172436E3EE48FFDF4749F80EFDD64589" blockId="0.[151,1437,1581,2039]" box="[631,635,2015,2039]" pageId="0" pageNumber="191">.</paragraph>
<caption id="43E4666BEE49FFDE45A9F802FAFD4597" box="[151,1360,2003,2025]" httpUri="https://zenodo.org/record/3241083/files/figure.png" pageId="1" pageNumber="192" startId="1.[151,258,2003,2025]" targetBox="[159,1429,182,1982]" targetPageId="1">
<paragraph id="172436E3EE49FFDE45A9F802FAFD4597" blockId="1.[151,1360,2003,2025]" box="[151,1360,2003,2025]" pageId="1" pageNumber="192">
<emphasis id="25EFEAF1EE49FFDE45A9F802FE814596" bold="true" box="[151,300,2003,2025]" pageId="1" pageNumber="192">FIGURES 15</emphasis>
.
<taxonomicName id="D09B4D60EE49FFDE4406F802FE4A4596" authority="Lu &amp; Wang, 2019" authorityName="Lu &amp; Wang" authorityYear="2019" box="[312,487,2003,2024]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="1" pageNumber="192" phylum="Arthropoda" rank="species" species="humpus" status="sp. nov.">
<emphasis id="25EFEAF1EE49FFDE4406F802FE4A4596" box="[312,487,2003,2024]" italics="true" pageId="1" pageNumber="192">Urothrips humpus</emphasis>
</taxonomicName>
<emphasis id="25EFEAF1EE49FFDE44D2F802FD9D4597" bold="true" box="[492,560,2003,2025]" pageId="1" pageNumber="192">
<taxonomicNameLabel id="3EDC578AEE49FFDE44D2F802FD8E4596" box="[492,547,2003,2024]" pageId="1" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
(
</emphasis>
female): (1) Habitus, (2) head, (3) antenna, (4) abdominal tergites IV, (5) fore legs
</paragraph>
</caption>
<paragraph id="172436E3EE4AFFDC45F9FE47FD094284" blockId="2.[151,1437,154,574]" lastBlockId="3.[151,1437,153,790]" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193">
Abdominal tergite I with few irregular transverse striate, closely fused to tergite II (Fig. 4); tergites IIVIII each with a pair of posterolateral setae, blunt at apex, with a transverse row of 1820 short setae; tergite II with two pairs of posteromarginal setae (Fig. 4), IIIVIII with three pairs. Abdominal sternites (Fig. 10) IIVIII each with 57 pairs of pointed setae medially, and 2 pairs of posteromarginal setae placed in front of posterior margin on IIVI, but at margin on VIIVIII; tube as long as head; anal setae more than three times length of tube Measurements (
<typeStatus id="C8208841EE4AFFDD444CF862FE7A45B5" box="[370,471,1971,1995]" pageId="2" pageNumber="193" type="holotype">Holotype</typeStatus>
in microns). Body length 1341. Head length 205, maximum width 155. Pronotum length 109, median width 180. Abdominal tergite VIII length 71, tergite IX length 191, width near base 101, distal 53. Tube length 207, width basal 29, apical 31. Antennae segments I to VI length (width) as follows: 22 (28), 22 (27), 26 (21), 58 (28), 30 (17), 38 (13). Length of setae: pronotum epimeral setae 14; abdominal tergites discal setae 4-5, posterolateral setae 21, posteromarginal setae 8; anal setae 662.
</paragraph>
<paragraph id="172436E3EE4AFFDD46E1FDF7FC4E4040" blockId="2.[151,1437,154,574]" box="[991,995,550,574]" pageId="2" pageNumber="193">.</paragraph>
<caption id="43E4666BEE4AFFDD45A9F899FE7245FC" httpUri="https://zenodo.org/record/3404906/files/figure.png" pageId="2" pageNumber="193" startId="2.[151,258,1864,1886]" targetBox="[151,1435,597,1844]" targetPageId="2">
<paragraph id="172436E3EE4AFFDD45A9F899FE7245FC" pageId="2" pageNumber="193">
<emphasis id="25EFEAF1EE4AFFDD45A9F899FE954523" bold="true" box="[151,312,1864,1886]" pageId="2" pageNumber="193">FIGURES 610</emphasis>
.
<emphasis id="25EFEAF1EE4AFFDD447DF899FE5F4523" box="[323,498,1864,1885]" italics="true" pageId="2" pageNumber="193">Urothrips humpus</emphasis>
<emphasis id="25EFEAF1EE4AFFDD44C9F899FD964520" bold="true" box="[503,571,1864,1886]" pageId="2" pageNumber="193">sp. n. (</emphasis>
female): (6) pronotum, (7) prosternum, (8) meso- and metanotum, (9) meso- and metaster- num, (10) abdominal sternum IV
</paragraph>
</caption>
<paragraph id="172436E3EE4BFFDC45F9FED7FED14360" blockId="3.[151,1437,153,790]" box="[199,380,262,286]" pageId="3" pageNumber="194">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="5F816568EE4BFFDC45F9FEFBFDB243AC" pageId="3" pageNumber="194" type="materials_examined">
<paragraph id="172436E3EE4BFFDC45F9FEFBFDB243AC" blockId="3.[151,1437,153,790]" pageId="3" pageNumber="194">
Specimens examined.
<materialsCitation id="A7F33CBEEE4BFFDC448FFEFBFF4543F4" ID-GBIF-Occurrence="2412513772" collectedFrom="monsoon forest canopy" collectingDate="2012-08-05" country="China" county="Mengla County" elevation="706" latitude="21.96" location="Bakaxiaozhai Reserve" longLatPrecision="757" longitude="101.2" municipality="Xishuangbanna Dai Autonomous Prefecture" pageId="3" pageNumber="194" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan province" typeStatus="holotype">
<typeStatus id="C8208841EE4BFFDC448FFEFBFDB9433C" box="[433,532,298,322]" pageId="3" pageNumber="194" type="holotype">Holotype</typeStatus>
<specimenCount id="019DFD6AEE4BFFDC4726FEFBFDCE433C" box="[536,611,298,322]" pageId="3" pageNumber="194" type="female">female</specimenCount>
,
<collectingCountry id="6F8C7673EE4BFFDC4755FEFBFD00433C" box="[619,685,298,322]" name="China" pageId="3" pageNumber="194">China</collectingCountry>
,
<collectingRegion id="D55FF801EE4BFFDC478BFEFBFCC7433C" box="[693,874,298,322]" country="China" name="Yunnan" pageId="3" pageNumber="194">Yunnan province</collectingRegion>
,
<collectingMunicipality id="F740AC99EE4BFFDC464CFEFBFAEE433C" box="[882,1347,298,322]" pageId="3" pageNumber="194">Xishuangbanna Dai Autonomous Prefecture</collectingMunicipality>
,
<collectingCounty id="FE454E6FEE4BFFDC4075FEFBFF4A4318" pageId="3" pageNumber="194">Mengla County</collectingCounty>
,
<collectingMunicipality id="F740AC99EE4BFFDC45CEFE9FFE38431B" box="[240,405,334,358]" pageId="3" pageNumber="194">Menglun Town</collectingMunicipality>
,
<location id="12446038EE4BFFDC44A1FE9FFD23431B" LSID="urn:lsid:plazi:treatment:9F3287F5EE48FFDC45A9F9FCFF554168:12446038EE4BFFDC44A1FE9FFD23431B" box="[415,654,334,358]" country="China" county="Mengla County" latitude="21.96" longLatPrecision="757" longitude="101.2" municipality="Xishuangbanna Dai Autonomous Prefecture" name="Bakaxiaozhai Reserve" pageId="3" pageNumber="194" stateProvince="Yunnan province">Bakaxiaozhai Reserve</location>
,
<collectedFrom id="A487751EEE4BFFDC47A6FE9FFC224318" box="[664,911,334,358]" pageId="3" pageNumber="194">monsoon forest canopy</collectedFrom>
(
<geoCoordinate id="72AF5024EE4BFFDC46A2FE9FFC574318" box="[924,1018,334,358]" degrees="21.96" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="555" value="21.96">21.96°N</geoCoordinate>
,
<geoCoordinate id="72AF5024EE4BFFDC413AFE9FFBC74318" box="[1028,1130,334,358]" degrees="101.20" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="555" value="101.2">101.20°E</geoCoordinate>
, alt
<quantity id="D0639B06EE4BFFDC41ABFE9FFB774318" box="[1173,1242,334,358]" metricMagnitude="2" metricUnit="m" metricValue="7.06" pageId="3" pageNumber="194" unit="m" value="706.0">
<elevation id="9CB6D1D0EE4BFFDC41ABFE9FFB774318" box="[1173,1242,334,358]" metricMagnitude="2" metricUnit="m" metricValue="7.06" pageId="3" pageNumber="194" unit="m" value="706.0">706 m</elevation>
</quantity>
),
<date id="63251023EE4BFFDC41D2FE9CFAF2431B" box="[1260,1375,333,357]" pageId="3" pageNumber="194" value="2012-08-05">
<collectingDate id="7361E9CBEE4BFFDC41D2FE9CFAF2431B" box="[1260,1375,333,357]" pageId="3" pageNumber="194" value="2012-08-05">5.viii.2012</collectingDate>
</date>
(
<location id="12446038EE4BFFDC4053FE9FFF4D43F4" LSID="urn:lsid:plazi:treatment:9F3287F5EE48FFDC45A9F9FCFF554168:12446038EE4BFFDC4053FE9FFF4D43F4" country="China" county="Mengla County" latitude="21.96" longLatPrecision="757" longitude="101.2" municipality="Xishuangbanna Dai Autonomous Prefecture" name="Guo Zheng" pageId="3" pageNumber="194" stateProvince="Yunnan province">Guo Zheng</location>
)
</materialsCitation>
.
<materialsCitation id="A7F33CBEEE4BFFDC45CEFEA3FDB243AC" ID-GBIF-Occurrence="2412513771" collectedFrom="monsoon forest canopy" collectingDate="2012-08-12" collectionCode="JLU" country="China" county="Mengla County" elevation="706" latitude="21.96" location="Bakaxiaozhai Reserve" longLatPrecision="757" longitude="101.2" municipality="Xishuangbanna Dai Autonomous Prefecture" pageId="3" pageNumber="194" specimenCount="4" specimenCount-female="4" stateProvince="Jilin" typeStatus="paratype">
<typeStatus id="C8208841EE4BFFDC45CEFEA3FEFF43F4" box="[240,338,370,394]" pageId="3" pageNumber="194" type="paratype">Paratype</typeStatus>
:
<specimenCount id="019DFD6AEE4BFFDC4465FEA3FE6943F4" box="[347,452,370,394]" pageId="3" pageNumber="194" type="female">2 females</specimenCount>
, collected with holotype,
<specimenCount id="019DFD6AEE4BFFDC47EFFEA3FC9743F4" box="[721,826,370,394]" pageId="3" pageNumber="194" type="female">2 females</specimenCount>
, same data with holotype except
<date id="63251023EE4BFFDC41ABFEA3FAB743F4" box="[1173,1306,370,394]" pageId="3" pageNumber="194" value="2012-08-12">
<collectingDate id="7361E9CBEE4BFFDC41ABFEA3FAB743F4" box="[1173,1306,370,394]" pageId="3" pageNumber="194" value="2012-08-12">12.viii.2012</collectingDate>
</date>
. Type specimens deposited in the Insect Collection of Jilin University (
<collectionCode id="718AAE26EE4BFFDC461FFE47FCF943D0" box="[801,852,406,430]" pageId="3" pageNumber="194">JLU</collectionCode>
); 2 paratypes are deposited in the Australian National Insect Collection, CSIRO, Canberra.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5F816568EE4BFFDC45F9FE0FFB56438B" box="[199,1275,478,502]" pageId="3" pageNumber="194" type="etymology">
<paragraph id="172436E3EE4BFFDC45F9FE0FFB56438B" blockId="3.[151,1437,153,790]" box="[199,1275,478,502]" pageId="3" pageNumber="194">
<emphasis id="25EFEAF1EE4BFFDC45F9FE0FFEE54388" bold="true" box="[199,328,478,502]" pageId="3" pageNumber="194">Etymology.</emphasis>
The Greek word
<taxonomicName id="D09B4D60EE4BFFDC473DFE0FFDCC4388" authorityName="Lu &amp; Wang" authorityYear="2019" box="[515,609,478,502]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="probolus">
<emphasis id="25EFEAF1EE4BFFDC473DFE0FFDCC4388" box="[515,609,478,502]" italics="true" pageId="3" pageNumber="194">probolus</emphasis>
</taxonomicName>
refers to the produced, hump-like ocellar region of the vertex.
</paragraph>
</subSubSection>
<subSubSection id="5F816568EE4BFFDC45F9FDD0FF554168" pageId="3" pageNumber="194" type="diagnosis">
<paragraph id="172436E3EE4BFFDC45F9FDD0FF554168" blockId="3.[151,1437,153,790]" pageId="3" pageNumber="194">
<emphasis id="25EFEAF1EE4BFFDC45F9FDD0FE6F4067" bold="true" box="[199,450,513,537]" pageId="3" pageNumber="194">Differential diagnosis.</emphasis>
This new species is the fourth species of
<taxonomicName id="D09B4D60EE4BFFDC46B8FDD3FC434064" authorityName="Bagnall" authorityYear="1909" box="[902,1006,514,538]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="genus">
<emphasis id="25EFEAF1EE4BFFDC46B8FDD3FC434064" box="[902,1006,514,538]" italics="true" pageId="3" pageNumber="194">Urothrips</emphasis>
</taxonomicName>
in which the ocellar region is produced as a hump over the basal antennal segments. However both
<taxonomicName id="D09B4D60EE4BFFDC4617FDF7FC1B4040" box="[809,950,550,574]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="gibberosa">
<emphasis id="25EFEAF1EE4BFFDC4617FDF7FC1B4040" box="[809,950,550,574]" italics="true" pageId="3" pageNumber="194">U. gibberosa</emphasis>
</taxonomicName>
and
<taxonomicName id="D09B4D60EE4BFFDC46D0FDF7FB2C4040" baseAuthorityName="Bournier" baseAuthorityYear="1963" box="[1006,1153,550,574]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="25EFEAF1EE4BFFDC46D0FDF7FB2C4040" box="[1006,1153,550,574]" italics="true" pageId="3" pageNumber="194">U. reticulatus</emphasis>
</taxonomicName>
have antennal segment V constricted at the base and distinct from IV. Moreover, in
<taxonomicName id="D09B4D60EE4BFFDC463EFD9BFCDC401C" baseAuthorityName="Bournier" baseAuthorityYear="1963" box="[768,881,586,610]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="25EFEAF1EE4BFFDC463EFD9BFCDC401C" box="[768,881,586,610]" italics="true" pageId="3" pageNumber="194">reticulatus</emphasis>
</taxonomicName>
antennal segment VIII is distinguished from VII by a clear suture, although these two segments are completely fused in
<taxonomicName id="D09B4D60EE4BFFDC4644FDBFFC4940F8" box="[890,996,622,646]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="gibberosa">
<emphasis id="25EFEAF1EE4BFFDC4644FDBFFC4940F8" box="[890,996,622,646]" italics="true" pageId="3" pageNumber="194">gibberosa</emphasis>
</taxonomicName>
. In contrast, both
<taxonomicName id="D09B4D60EE4BFFDC4194FDBFFABF40FB" box="[1194,1298,622,645]" class="Insecta" family="Phlaeothripidae" genus="Urothrips" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="194" phylum="Arthropoda" rank="species" species="juncus">
<emphasis id="25EFEAF1EE4BFFDC4194FDBFFABF40FB" box="[1194,1298,622,645]" italics="true" pageId="3" pageNumber="194">U. juncus</emphasis>
</taxonomicName>
and the new species have antennal segment V broad at the base, although in
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there is a clear suture indicating the apex of IV, whereas in the new species these segments are completely fused with no obvious suture. Moreover, the maxillary stylets of
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are about one third of head width apart, whereas in the new species the stylets are close together medially in the head.
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<paragraph id="BB2E1A44835730820BB1521ACE9A1CFC" pageId="271" pageNumber="272">Original source.</paragraph>
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<bibRefCitation id="AA702CC6DD68264F8140E1327BECED9A" author="Jekelius, E" journalOrPublisher="Memoriile Institutului geologic al Romaniei" pageId="315" pageNumber="316" pagination="1 - 167" refId="B246" refString="Jekelius, E, 1944. Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei 5: 1 - 167" title="Sarmat und Pont von Soceni (Banat)." volume="5" year="1944">Jekelius 1944</bibRefCitation>
: 136, pl. 55, figs 15-19.
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<paragraph id="ED11D235DF64825958582F5B3E7E61B7" pageId="271" pageNumber="272">Type horizon.</paragraph>
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<paragraph id="8C087DAC19BD505EBB65AC180F73CEF6" pageId="271" pageNumber="272">Type locality.</paragraph>
<paragraph id="94145E35DB8BB0AC18B29781E83BE597" pageId="271" pageNumber="272">&quot;Turislav-Tal bei Soceni&quot; [Turislav valley near Soceni], Romania.</paragraph>
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<paragraph id="856172F229A69EC2F47899BA19C83A98" pageId="0" pageNumber="79255">Atlanto-Mediterranean</paragraph>
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<paragraph id="7F91EE5CB6985F901585DEFF72BC96E4" pageId="0" pageNumber="79255">Biological data: Univoltine. Flight period: IX. First record in Murcia Region.</paragraph>
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