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(
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,
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Key to Genera of
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1 Hind wings obsolete. Mesosternum narrow between mesocoxae (
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)
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2
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19 Hind wings fully developed. Mesosternum wide between mesocoxae (
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) - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -
<taxonomicName id="D084F4CF6A23FFFCFCA7FC567CCCBE92" ID-CoL="4BXH" authority="Lansverge" authorityName="Lansverge" box="[771,1005,898,919]" class="Insecta" family="Scarabaeidae" genus="Ennearabdus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
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Lansverge
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2 Mesocoxae separated at base (
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). Mesotarsus longer than metatarsus
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3
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29 Mesocoxae nearly contiguous at base (
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). Mesotarsus shorter than metatarsus - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -
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Brullé
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<heading id="4C7338206A29FFF6FF0BFBDE7D44B91A" box="[175,613,1034,1055]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="303" reason="2">
<taxonomicName id="D084F4CF6A29FFF6FF0BFBDE7D41B91A" ID-CoL="6KM8F" authority="Burmeister, 1861: 67" authorityName="Burmeister" authorityPageNumber="67" authorityYear="1861" box="[175,608,1034,1055]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="25F0535E6A29FFF6FF0BFBDE7EB0B91A" box="[175,401,1034,1055]" italics="true" pageId="8" pageNumber="303">Glyphoderus monticola</emphasis>
Burmeister, 1861:67
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF63FBE57EC3B804" pageId="8" pageNumber="303" type="diagnosis">
<paragraph id="173B8F4C6A29FFF6FF63FBE57EC3B804" blockId="8.[175,1005,1073,1387]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FBE57E0EB943" bold="true" box="[199,303,1073,1094]" pageId="8" pageNumber="303">Diagnosis.</emphasis>
Males. This species is separated from other
<taxonomicName id="D084F4CF6A29FFF6FD47FBE67C88B942" box="[739,937,1074,1095]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="25F0535E6A29FFF6FD47FBE67C7FB942" box="[739,862,1074,1095]" italics="true" pageId="8" pageNumber="303">Glyphoderus</emphasis>
species
</taxonomicName>
by the shape and placement of the pronotal horns and tubercles (
<figureCitation id="8FBF93C96A29FFF6FCB2FB987C79B964" box="[790,856,1100,1121]" captionStart="Fig" captionStartId="5.[169,200,1175,1194]" captionTargetBox="[151,980,199,1162]" captionTargetId="figure-166@5.[148,980,194,1162]" captionTargetPageId="5" captionText="Fig. 4. Dorsal habitus of Glyphoderus monticola Burmeister." figureDoi="http://doi.org/10.5281/zenodo.10104691" httpUri="https://zenodo.org/record/10104691/files/figure.png" pageId="8" pageNumber="303">Fig. 4</figureCitation>
).
<taxonomicName id="D084F4CF6A29FFF6FCD6FB987E2FB979" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="25F0535E6A29FFF6FCD6FB987E2FB979" italics="true" pageId="8" pageNumber="303">Glyphoderus monticola</emphasis>
</taxonomicName>
has the pronotum with two horns in middle and two transversally elongated tubercles (sometimes developed as horns) on the basal half; horns and tubercles on margin of medial depression; anteromedial horn with ‘‘arms’’ elongated, with two anteromedial teeth; distance between teeth as long as tooth base. No morphological characters were found to diagnose females of this species, females can be identified only by association with males.
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF63FAD27E1BB86E" pageId="8" pageNumber="303" type="distribution">
<paragraph id="173B8F4C6A29FFF6FF63FAD27C68B833" blockId="8.[175,1005,1073,1387]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FAD27E65B81E" bold="true" box="[199,324,1286,1307]" pageId="8" pageNumber="303">Distribution</emphasis>
(
<figureCitation id="8FBF93C96A29FFF6FEF0FAD27EB3B81E" box="[340,402,1286,1307]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="8" pageNumber="303">Fig. 6</figureCitation>
).
<emphasis id="25F0535E6A29FFF6FE0EFAD27DE7B81E" bold="true" box="[426,710,1286,1307]" pageId="8" pageNumber="303">
<collectingCountry id="6F93CFDC6A29FFF6FE0EFAD27D1EB81E" box="[426,575,1286,1307]" name="Argentina" pageId="8" pageNumber="303">ARGENTINA</collectingCountry>
:
<collectingRegion id="D54041AE6A29FFF6FDE9FAD27DE2B81E" box="[589,707,1286,1307]" country="Argentina" name="Catamarca" pageId="8" pageNumber="303">Catamarca</collectingRegion>
:
</emphasis>
El Recreo (3).
<emphasis id="25F0535E6A29FFF6FCC1FAD27CEBB81E" bold="true" box="[869,970,1286,1307]" pageId="8" pageNumber="303">
<collectingRegion id="D54041AE6A29FFF6FCC1FAD27CE4B81E" box="[869,965,1286,1307]" country="Argentina" name="La Rioja" pageId="8" pageNumber="303">La Rioja</collectingRegion>
:
</emphasis>
La Rioja (3); La Rioja (
<quantity id="D07C22A96A29FFF6FED3FAF57E97B833" box="[375,438,1313,1334]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="303" unit="km" value="20.0">20 km</quantity>
N) (4); Mascasin (33); no more data (2).
</paragraph>
<paragraph id="173B8F4C6A29FFF6FF63FAEF7E1BB86E" blockId="8.[175,1005,1073,1387]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FAEF7E9AB855" bold="true" box="[199,443,1339,1360]" pageId="8" pageNumber="303">Temporal Distribution.</emphasis>
January (31), February (4), March (3), November (2), December (1).
</paragraph>
</subSubSection>
</treatment>
</document>

135
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<mods:title id="FB4B25E34219CCC616E98E662DD4FB51">Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini)</mods:title>
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<paragraph id="173B8F4C6A29FFF6FF0BFE787D44BCC4" blockId="8.[175,613,428,449]" box="[175,613,428,449]" pageId="8" pageNumber="303">
<heading id="4C7338206A29FFF6FF0BFE787D44BCC4" box="[175,613,428,449]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="303" reason="2">
<taxonomicName id="D084F4CF6A29FFF6FF0BFE787D40BCC4" ID-CoL="3GGNT" authority="Burmeister, 1873: 407" authorityName="Burmeister" authorityPageNumber="407" authorityYear="1873" box="[175,609,428,449]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="25F0535E6A29FFF6FF0BFE787EA4BCC4" box="[175,389,428,449]" italics="true" pageId="8" pageNumber="303">Glyphoderus centralis</emphasis>
Burmeister, 1873:407
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF63FE007EB1BF8D" pageId="8" pageNumber="303" type="diagnosis">
<paragraph id="173B8F4C6A29FFF6FF63FE007EB1BF8D" blockId="8.[175,1005,468,914]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FE007E0EBCEC" bold="true" box="[199,303,468,489]" pageId="8" pageNumber="303">Diagnosis.</emphasis>
Males. This species is separated from other
<taxonomicName id="D084F4CF6A29FFF6FD47FE007C88BCEC" box="[739,937,468,489]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="25F0535E6A29FFF6FD47FE007C7FBCEC" box="[739,862,468,489]" italics="true" pageId="8" pageNumber="303">Glyphoderus</emphasis>
species
</taxonomicName>
by the shape and placement of the pronotal horns and tubercles (
<figureCitation id="8FBF93C96A29FFF6FD6AFE3B7C26BF01" box="[718,775,495,516]" captionStart="Fig" captionStartId="4.[197,228,693,712]" captionTargetBox="[196,983,194,679]" captionTargetId="figure-385@4.[196,983,194,679]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of the pronotum of Glyphoderus centralis Burmeister." figureDoi="http://doi.org/10.5281/zenodo.10104688" httpUri="https://zenodo.org/record/10104688/files/figure.png" pageId="8" pageNumber="303">Fig. 3</figureCitation>
).
<taxonomicName id="D084F4CF6A29FFF6FCBDFE3B7CCCBF01" box="[793,1005,495,516]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="25F0535E6A29FFF6FCBDFE3B7CCCBF01" box="[793,1005,495,516]" italics="true" pageId="8" pageNumber="303">Glyphoderus centralis</emphasis>
</taxonomicName>
has the pronotum with two transversally elongated tubercles on apical half and two horns in middle; horns and tubercles on margin of central depression; anteromedial horn with ‘‘arms’’ robust, with medial anterior notch. No morphological characters were found to diagnose females of this species, females can be identified only by association with males.
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF63FD5A7D30BE2D" pageId="8" pageNumber="303" type="distribution">
<paragraph id="173B8F4C6A29FFF6FF63FD5A7CCCBFB8" blockId="8.[175,1005,468,914]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FD5A7E65BFA6" bold="true" box="[199,324,654,675]" pageId="8" pageNumber="303">Distribution</emphasis>
(
<figureCitation id="8FBF93C96A29FFF6FEFEFD5A7EBFBFA6" box="[346,414,654,675]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="8" pageNumber="303">Fig. 6</figureCitation>
).
<emphasis id="25F0535E6A29FFF6FE11FD5A7D86BFA6" bold="true" box="[437,679,654,675]" pageId="8" pageNumber="303">
<collectingCountry id="6F93CFDC6A29FFF6FE11FD5A7D6BBFA6" box="[437,586,654,675]" name="Argentina" pageId="8" pageNumber="303">ARGENTINA</collectingCountry>
:
<collectingRegion id="D54041AE6A29FFF6FDFAFD5A7D82BFA6" box="[606,675,654,675]" country="Argentina" name="Chaco" pageId="8" pageNumber="303">Chaco</collectingRegion>
:
</emphasis>
Taco Pozo (1);
<emphasis id="25F0535E6A29FFF6FCC1FD5A7C81BFA6" bold="true" box="[869,928,654,675]" pageId="8" pageNumber="303">
<collectingRegion id="D54041AE6A29FFF6FCC1FD5A7CBDBFA6" box="[869,924,654,675]" country="Argentina" name="Salta" pageId="8" pageNumber="303">Salta</collectingRegion>
:
</emphasis>
Chaco Occidental, Los Colorados (
<quantity id="D07C22A96A29FFF6FE6FFD7D7D3BBFBB" box="[459,538,681,702]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="303" unit="km" value="100.0">100 km</quantity>
NE) (1);
<emphasis id="25F0535E6A29FFF6FD22FD7C7DCCBFB8" bold="true" box="[646,749,680,701]" pageId="8" pageNumber="303">
<collectingRegion id="D54041AE6A29FFF6FD22FD7C7DC8BFB8" box="[646,745,680,701]" country="Argentina" name="San Luis" pageId="8" pageNumber="303">San Luis</collectingRegion>
:
</emphasis>
Talita (1).
<emphasis id="25F0535E6A29FFF6FCCDFD7C7CCCBFB8" bold="true" box="[873,1005,680,701]" pageId="8" pageNumber="303">
<collectingRegion id="D54041AE6A29FFF6FCCDFD7C7CCCBFB8" box="[873,1005,680,701]" country="Argentina" name="Santiago del Estero" pageId="8" pageNumber="303">Santiago del</collectingRegion>
</emphasis>
</paragraph>
<paragraph id="173B8F4C6A29FFF6FF0BFD177D78BFF6" blockId="8.[175,1005,468,914]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF0BFD177FD8BFDD" bold="true" box="[175,249,707,728]" pageId="8" pageNumber="303">Estero:</emphasis>
Beltrán (2); Choya (21); El Charro (13); Fernández (5); Río Salado (4); San Gregorio (4); Puna (15); no more data (13).
</paragraph>
<paragraph id="173B8F4C6A29FFF6FF63FD2C7D30BE2D" blockId="8.[175,1005,468,914]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FD2C7E94BE08" bold="true" box="[199,437,760,781]" pageId="8" pageNumber="303">Temporal Distribution.</emphasis>
January (3); February (11), March (14), April (1); October (21), November (2), December (17).
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF63FCF97EFFBE97" pageId="8" pageNumber="303" type="discussion">
<paragraph id="173B8F4C6A29FFF6FF63FCF97EFFBE97" blockId="8.[175,1005,468,914]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FF63FCF97E0BBE47" bold="true" box="[199,298,813,834]" pageId="8" pageNumber="303">Remarks.</emphasis>
<bibRefCitation id="7315F2BD6A29FFF6FE92FCF97ED9BE47" author="Zunino" box="[310,504,813,834]" pageId="8" pageNumber="303" pagination="235 - 240" refId="ref6427" refString="Zunino, M., E. Barbero, and M. Luzzatto. 1989. Food relocation behavior in Eucraniina beetles (Scarabaeidae) and the constraints of xeric environment. Tropical Zoology 2: 235 - 240." type="journal article" year="1989">
Zunino
<emphasis id="25F0535E6A29FFF6FE26FCF97E92BE47" box="[386,435,813,834]" italics="true" pageId="8" pageNumber="303">et al.</emphasis>
(1989)
</bibRefCitation>
and
<bibRefCitation id="7315F2BD6A29FFF6FD8CFCF97DE2BE47" author="Martinez" box="[552,707,813,834]" pageId="8" pageNumber="303" pagination="1 - 130" refId="ref6061" refString="Martinez, A. 1956. Catalogo de los Scarabaeidae argentinos (Coleoptera). Revista del Museo Argentino de Ciencias Naturales Bernardino Rivadavia 5: 1 - 130." type="journal article" year="1956">Martínez (1956)</bibRefCitation>
recorded
<taxonomicName id="D084F4CF6A29FFF6FC87FCF97CB5BE47" box="[803,916,813,834]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="25F0535E6A29FFF6FC87FCF97CB5BE47" box="[803,916,813,834]" italics="true" pageId="8" pageNumber="303">G. centralis</emphasis>
</taxonomicName>
from the Argentinean provinces of
<collectingRegion id="D54041AE6A29FFF6FE0EFC9C7D30BE58" box="[426,529,840,861]" country="Argentina" name="Catamarca" pageId="8" pageNumber="303">Catamarca</collectingRegion>
,
<collectingRegion id="D54041AE6A29FFF6FDB8FC9C7D51BE58" box="[540,624,840,861]" country="Argentina" name="Cordoba" pageId="8" pageNumber="303">Córdoba</collectingRegion>
,
<collectingRegion id="D54041AE6A29FFF6FDD8FC9C7DF3BE58" box="[636,722,840,861]" country="Argentina" name="La Rioja" pageId="8" pageNumber="303">La Rioja</collectingRegion>
, and
<collectingRegion id="D54041AE6A29FFF6FCA2FC9C7C42BE58" box="[774,867,840,861]" country="Argentina" name="Tucuman" pageId="8" pageNumber="303">Tucumán</collectingRegion>
. Though I did not study specimens from these provinces I believe that they should be considered as part of the distributional range.
</paragraph>
</subSubSection>
</treatment>
</document>

168
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<mods:title id="D69C5D868E50FAA79CCFD19FFE32B0E0">Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini)</mods:title>
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<mods:namePart id="BCDA9D29E1E7CBC5066A4C81BA6A0315">Ocampo, Federico C.</mods:namePart>
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<mods:title id="CAEA83FB2D8BC5069C6618507FE58B31">The Coleopterists Bulletin</mods:title>
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<mods:date id="BF84D93C582AC285E861A7452A4966FC">2004</mods:date>
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<paragraph id="173B8F4C6A29FFF6FED5FA647C0AB8C0" blockId="8.[369,811,1456,1504]" box="[369,811,1456,1477]" pageId="8" pageNumber="303">
<heading id="4C7338206A29FFF6FED5FA647C0AB8C0" box="[369,811,1456,1477]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="303" reason="2">
<taxonomicName id="D084F4CF6A29FFF6FED5FA647C0AB8C0" ID-CoL="3GGNV" authority="(Westwood, 1837)" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[369,811,1456,1477]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FED5FA647C0AB8C0" bold="true" box="[369,811,1456,1477]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FED5FA647D4DB8C0" bold="true" box="[369,620,1456,1477]" italics="true" pageId="8" pageNumber="303">Glyphoderus sterquilinus</emphasis>
(
<bibRefCitation id="7315F2BD6A29FFF6FDDEFA647C05B8C0" author="Westwood" box="[634,804,1456,1477]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FDBCFA1F7DA5B8E5" box="[536,644,1483,1504]" pageId="8" pageNumber="303" type="description">
<paragraph id="173B8F4C6A29FFF6FDBCFA1F7DA5B8E5" blockId="8.[369,811,1456,1504]" box="[536,644,1483,1504]" pageId="8" pageNumber="303">
(
<figureCitation id="8FBF93C96A29FFF6FDBBFA1F7D42B8E5" box="[543,611,1483,1504]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="8" pageNumber="303">Figs. 5</figureCitation>
,
<figureCitation id="8FBF93C96A29FFF6FDD4FA1F7D5DB8E5" box="[624,636,1483,1504]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="8" pageNumber="303">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF0BFA277C63BB27" pageId="8" pageNumber="303" type="reference_group">
<paragraph id="173B8F4C6A29FFF6FF0BFA277DA3BB0D" blockId="8.[175,834,1523,1570]" box="[175,642,1523,1544]" pageId="8" pageNumber="303">
<taxonomicName id="D084F4CF6A29FFF6FF0BFA277D43BB0D" ID-CoL="3GGNV" authority="(Westwood, 1837)" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[175,610,1523,1544]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FF0BFA277E83BB0D" box="[175,418,1523,1544]" italics="true" pageId="8" pageNumber="303">Glyphoderus sterquilinus</emphasis>
(
<bibRefCitation id="7315F2BD6A29FFF6FE14FA277D7ABB0D" author="Westwood" box="[432,603,1523,1544]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837</bibRefCitation>
)
</taxonomicName>
:13.
</paragraph>
<paragraph id="173B8F4C6A29FFF6FF0BF9D97C63BB27" blockId="8.[175,834,1523,1570]" box="[175,834,1549,1570]" pageId="8" pageNumber="303">
<treatmentCitationGroup id="3794A8626A29FFF6FF0BF9D97C63BB27" box="[175,834,1549,1570]" pageId="8" pageNumber="303">
<treatmentCitation id="9625A95D6A29FFF6FF0BF9D97D43BB27" author="Westwood" box="[175,610,1549,1570]" page="13" pageId="8" pageNumber="303" year="1837">
<taxonomicName id="D084F4CF6A29FFF6FF0BF9D97D43BB27" ID-CoL="5V6LJ" authority="Westwood, 1837: 13" authorityName="Westwood" authorityPageNumber="13" authorityYear="1837" box="[175,610,1549,1570]" class="Insecta" family="Scarabaeidae" genus="Anomiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FF0BF9D97EB7BB27" box="[175,406,1549,1570]" italics="true" pageId="8" pageNumber="303">Anomiopsis sterquilinus</emphasis>
<bibRefCitation id="7315F2BD6A29FFF6FE3AF9D97D43BB27" author="Westwood" box="[414,610,1549,1570]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837:13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A28FFF7FF08FF167C88BCC3" pageId="9" pageNumber="304" type="diagnosis">
<paragraph id="173B8F4C6A28FFF7FF08FF167C88BCC3" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<emphasis id="25F0535E6A28FFF7FF08FF167E35BDD2" bold="true" box="[172,276,194,215]" pageId="9" pageNumber="304">Diagnosis.</emphasis>
Males. This species is separated from other
<taxonomicName id="D084F4CF6A28FFF7FD63FF167CACBDD2" box="[711,909,194,215]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="25F0535E6A28FFF7FD63FF167C63BDD2" box="[711,834,194,215]" italics="true" pageId="9" pageNumber="304">Glyphoderus</emphasis>
species
</taxonomicName>
by the shape and placement of the pronotal horns and tubercles (
<figureCitation id="8FBF93C96A28FFF7FD5FFF097C1DBDF7" box="[763,828,221,242]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="9" pageNumber="304">Fig. 5</figureCitation>
).
<taxonomicName id="D084F4CF6A28FFF7FCF2FF097E25BC08" baseAuthorityName="Westwood" baseAuthorityYear="1838" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A28FFF7FCF2FF097E25BC08" italics="true" pageId="9" pageNumber="304">Glyphoderus sterquilinus</emphasis>
</taxonomicName>
has the pronotum with two horns in middle and two transversally elongated tubercles (sometimes developed as horns) on basal half; horns on margin of central depression; tubercles of basal half included in central depression; apical medial horn with ‘‘arms’’ robust, with anteromedial notch. Some specimens of
<taxonomicName id="D084F4CF6A28FFF7FCB4FE937CBFBC59" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[784,926,327,348]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A28FFF7FCB4FE937CBFBC59" box="[784,926,327,348]" italics="true" pageId="9" pageNumber="304">G. sterquilinus</emphasis>
</taxonomicName>
have the posterior pronotal tubercles on the central depression margin (observed particularly in minor males), in this case, the shape of the medial horn will separate this species from
<taxonomicName id="D084F4CF6A28FFF7FF6EFE437E69BCA9" box="[202,328,407,428]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="25F0535E6A28FFF7FF6EFE437E69BCA9" box="[202,328,407,428]" italics="true" pageId="9" pageNumber="304">G. monticola</emphasis>
</taxonomicName>
(
<figureCitation id="8FBF93C96A28FFF7FEFCFE437EBCBCA9" box="[344,413,407,428]" captionStart="Fig" captionStartId="5.[169,200,1175,1194]" captionTargetBox="[151,980,199,1162]" captionTargetId="figure-166@5.[148,980,194,1162]" captionTargetPageId="5" captionText="Fig. 4. Dorsal habitus of Glyphoderus monticola Burmeister." figureDoi="http://doi.org/10.5281/zenodo.10104691" httpUri="https://zenodo.org/record/10104691/files/figure.png" pageId="9" pageNumber="304">Figs. 4</figureCitation>
,
<figureCitation id="8FBF93C96A28FFF7FE0EFE437E96BCA9" box="[426,439,407,428]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="9" pageNumber="304">5</figureCitation>
). No morphological characters were found to diagnose females of this species, females can be identified only by association with males.
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A28FFF7FF08FE187D70BF4E" pageId="9" pageNumber="304" type="distribution">
<paragraph id="173B8F4C6A28FFF7FF08FE187CF1BF13" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<emphasis id="25F0535E6A28FFF7FF08FE187E08BCE4" bold="true" box="[172,297,460,481]" pageId="9" pageNumber="304">Distribution</emphasis>
(
<figureCitation id="8FBF93C96A28FFF7FE93FE187E52BCE4" box="[311,371,460,481]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="9" pageNumber="304">Fig. 6</figureCitation>
).
<emphasis id="25F0535E6A28FFF7FE2FFE187DB2BCE4" bold="true" box="[395,659,460,481]" pageId="9" pageNumber="304">
<collectingCountry id="6F93CFDC6A28FFF7FE2FFE187D01BCE4" box="[395,544,460,481]" name="Argentina" pageId="9" pageNumber="304">ARGENTINA</collectingCountry>
:
<collectingRegion id="D54041AE6A28FFF7FD88FE187DAFBCE4" box="[556,654,460,481]" country="Argentina" name="Mendoza" pageId="9" pageNumber="304">Mendoza</collectingRegion>
:
</emphasis>
Las Catitias (
<quantity id="D07C22A96A28FFF7FCBFFE187C6FBCE4" box="[795,846,460,481]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="304" unit="km" value="7.0">7 km</quantity>
S) (6); Santa Rosa (1); Villavicencio (1).
<emphasis id="25F0535E6A28FFF7FE06FE327D24BCFE" bold="true" box="[418,517,486,507]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FE06FE327D21BCFE" box="[418,512,486,507]" country="Argentina" name="La Rioja" pageId="9" pageNumber="304">La Rioja</collectingRegion>
:
</emphasis>
Chepes (2); Ulapes (
<quantity id="D07C22A96A28FFF7FD76FE337C31BCF9" box="[722,784,487,508]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="304" unit="km" value="17.0">17 km</quantity>
S) (28); Ulapes (
<quantity id="D07C22A96A28FFF7FC1CFE337F93BF13" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="304" unit="km" value="10.0">10 km</quantity>
N) (8).
<emphasis id="25F0535E6A28FFF7FEA5FDD57E49BF13" bold="true" box="[257,360,513,534]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FEA5FDD57E45BF13" box="[257,356,513,534]" country="Argentina" name="San Juan" pageId="9" pageNumber="304">San Juan</collectingRegion>
:
</emphasis>
Encón (
<quantity id="D07C22A96A28FFF7FE1EFDD57ED9BF13" box="[442,504,513,534]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="304" unit="km" value="12.0">12 km</quantity>
E) (1); Nikizanga (2); Morayes –Astica road (1).
</paragraph>
<paragraph id="173B8F4C6A28FFF7FF30FDCF7D70BF4E" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<materialsCitation id="A7EC85116A28FFF7FF30FDCF7F9CBF4E" collectorName="La Chanarienta" location="San Luis" pageId="9" pageNumber="304" specimenCount="1" stateProvince="San Luis">
<emphasis id="25F0535E6A28FFF7FF30FDCF7FD9BF35" bold="true" box="[148,248,539,560]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FF30FDCF7FD5BF35" box="[148,244,539,560]" country="Argentina" name="San Luis" pageId="9" pageNumber="304">San Luis</collectingRegion>
:
</emphasis>
Jarilla (1);
<collectorName id="BA71EA9A6A28FFF7FECAFDC87D23BF34" box="[366,514,540,561]" pageId="9" pageNumber="304">La Chañarienta</collectorName>
(2); San Antonio (5); Talita (8); no more data (11)
</materialsCitation>
.
<emphasis id="25F0535E6A28FFF7FF6DFDE27EBBBF4E" bold="true" box="[201,410,566,587]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FF6DFDE27EB7BF4E" box="[201,406,566,587]" country="Argentina" name="Santiago del Estero" pageId="9" pageNumber="304">Santiago del Estero</collectingRegion>
:
</emphasis>
No more data (6).
</paragraph>
</subSubSection>
</treatment>
</document>