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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1049.66063" ID-GBIF-Dataset="5606bde8-a7da-4058-a3fc-f69a02ecbd0b" ID-PMC="PMC8298371" ID-Pensoft-Pub="1313-2970-1049-15" ID-Pensoft-UUID="F43B77514DD55AC69BA1E334AE61ABF2" ID-PubMed="34321953" ID-ZooBank="E0AEB4DB6C32407697542AA74386C517" ModsDocID="1313-2970-1049-15" checkinTime="1626449947631" checkinUser="pensoft" docAuthor="Jezek, Jan, Manko, Peter &amp; Obona, Jozef" docDate="2021" docId="C1A8651D80BF55CABA3DAAF352278776" docLanguage="en" docName="ZooKeys 1049: 15-42" docOrigin="ZooKeys 1049" docPubDate="2021-06-15" docSource="http://dx.doi.org/10.3897/zookeys.1049.66063" docTitle="Trichomyia urbica Haliday in Curtis 1839" docType="treatment" docVersion="4" id="F43B77514DD55AC69BA1E334AE61ABF2" lastPageNumber="15" masterDocId="F43B77514DD55AC69BA1E334AE61ABF2" masterDocTitle="Psychodidae (Diptera) of Azerbaijan and Georgia - faunistics with biodiversity notes" masterLastPageNumber="42" masterPageNumber="15" pageNumber="15" updateTime="1668150595893" updateUser="ExternalLinkService">
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<mods:title>Psychodidae (Diptera) of Azerbaijan and Georgia - faunistics with biodiversity notes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jezek, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, National Museum, Cirkusova 1740, CZ - 193 00 Praha 9 - Horni Pocernice, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Manko, Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1862-9117</mods:nameIdentifier>
<mods:affiliation>Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK - 081 16 Presov, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">peter.manko@unipo.sk</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Obona, Jozef</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1185-658X</mods:nameIdentifier>
<mods:affiliation>Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK - 081 16 Presov, Slovakia</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-15</mods:number>
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<mods:detail type="volume">
<mods:number>1049</mods:number>
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<mods:start>15</mods:start>
<mods:end>42</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1049.66063</mods:url>
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<taxonomicName LSID="C1A8651D-80BF-55CA-BA3D-AAF352278776" authority="Haliday in Curtis, 1839" authorityName="Haliday in Curtis" authorityYear="1839" class="Insecta" family="Psychodidae" genus="Trichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichomyia urbica" order="Diptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="urbica">Trichomyia urbica Haliday in Curtis, 1839</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="materials_examined">
<paragraph pageId="0" pageNumber="15">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="15">
<materialsCitation collectingDate="2019-05-09" collectorName="Inv. No." country="Azerbaijan" location="Azerbaijan" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" pageId="0" pageNumber="15">
<collectingCountry name="Azerbaijan">Azerbaijan</collectingCountry>
</emphasis>
: A 02,
<collectingDate value="2019-05-09">9.5.2019</collectingDate>
,
<specimenCount type="male">1♂</specimenCount>
, slide
<collectorName>Inv. No.</collectorName>
25663, leg. JO (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Occurrence of individual species on a map of the area of interest. Different colours (altitude) indicate the elevation in m a. s. l. and different symbols mark the sampling sites with occurrence of the species. When species co-occurred at the locality, an offset is shown in the legend and described in the Materials and methods. The number of sites where the species is present is given in brackets." figureDoi="10.3897/zookeys.1049.66063.figure2" httpUri="https://binary.pensoft.net/fig/567174" pageId="0" pageNumber="15">2</figureCitation>
)
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.
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<paragraph pageId="0" pageNumber="15">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="15">
Rare species, known from Austria, Belgium, Czech Republic, Denmark, France, Germany, Great Britain, Greece, Hungary, Ireland, the Netherlands, Norway, Poland, Romania, Slovakia, and Sweden; larvae are xylophagous and occur in shaded slope spring areas and some other habitats with decaying organic matter (
<bibRefCitation author="Jezek, J" journalOrPublisher="Journal of the National Museum (Prague) Natural History Series" pageId="0" pageNumber="15" pagination="121 - 132" refId="B20" refString="Jezek, J, 2003. New faunistic data and check list of non Phlebotomine moth flies (Diptera, Psychodidae) from the Czech and Slovak republics. Journal of the National Museum (Prague) Natural History Series 172 (1-4): 121 - 132" title="New faunistic data and check list of non Phlebotomine moth flies (Diptera, Psychodidae) from the Czech and Slovak republics." volume="172" year="2003">
<normalizedToken originalValue="Ježek">Jezek</normalizedToken>
2003
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;
<bibRefCitation author="Jezek, J" journalOrPublisher="Dipterologica Bohemoslovaca" pageId="0" pageNumber="15" refId="B34" refString="Jezek, J, Omelkova, M, 2012. Moth flies (Diptera: Psychodidae) of the Bile Karpaty Protected Landscape Area and Biosphere Reserve (Czech Republic). Acta Musei Moraviae, Scientiae Biologicae (Brno) (2011) 96(2): 763-802." title="Moth flies (Diptera: Psychodidae) of the Bile Karpaty Protected Landscape Area and Biosphere Reserve (Czech Republic). Acta Musei Moraviae, Scientiae Biologicae (Brno) (2011) 96 (2): 763 - 802." year="2012">
<normalizedToken originalValue="Ježek">Jezek</normalizedToken>
and
<normalizedToken originalValue="Omelková">Omelkova</normalizedToken>
2012
</bibRefCitation>
).
</paragraph>
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<subSubSection pageId="0" pageNumber="15" type="notes">
<paragraph pageId="0" pageNumber="15">Note.</paragraph>
<paragraph pageId="0" pageNumber="15">First record for Azerbaijan and Transcaucasia.</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="C6BF873770C9CB95BE38E7DA9683F96C" ID-CoL="4RF6J" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="B33ACCE512E148FE29E181AC46060D30" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
(Linnaeus 1758)
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<taxonomicName id="F1935846D5478D0766FB4E07C00B59EC" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cervidae" genus="Cervus" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="FBB0AEED7793CD4B1726C4B8840F7EB4" italics="true" pageNumber="660">[Cervus] tarandus</emphasis>
<bibRefCitation id="A23295F5BC9E0D3BB0410B728FFDEB88" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
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,
<bibCitation id="74D3FC158450114BFE318A6B105ED355" author="Linnaeus" pageNumber="660" pagination="67" title="Cervus tarandus" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">Syst. Nat., 10th ed., Vol. 1: 67</bibCitation>
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.
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<paragraph id="DC66B0D36DA9C73E337065AD79A9CCC0" pageNumber="660">
<emphasis id="7D1E1574C8E33A476BCF52DD83893DA7" inLineHeading="true" pageNumber="660">Type Locality:</emphasis>
<materialsCitation id="616AFCFA2BA9EE20B77A925659FEDA77" ID-GBIF-Occurrence="4867329660" country="Sweden" pageNumber="660">
&quot;Habitat in Alpibus Europae et Asiae maxime septentrionalibus&quot;; identified as
<collectingCountry id="A17BC6C2E466173E3F9BA1DE8DE1D912" name="Sweden">Sweden</collectingCountry>
, Alpine Lapland by
<bibRefCitation id="00ED9DA560DC7EB98ACAF3FAA8CB763C" author="Thomas, O." refId="ref353417" refString="Thomas, O. 1911 a. The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London, 1911: 120 - 158." year="1911">
Thomas (1911
<emphasis id="BFBB332AEB9DD07CF4C5089B698F098E" italics="true" pageNumber="660">a</emphasis>
:151)
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; based on domesticated stock
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.
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<subSubSection id="169428F24D8D8F065DC7C03ED3DB9CEB" pageNumber="660" type="vernacular_names">
<paragraph id="F23D250B8B79BEF5131068EEF2402476" pageNumber="660">
<emphasis id="0EB1F0276BA450BA3E48293475C28FC9" inLineHeading="true" pageNumber="660">Vernacular Names:</emphasis>
<vernacularName id="A83ED0DF7AB0F90BF6696FC1323AE270" language="eng" pageNumber="660">Reindeer</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="CDC046BD94D265BAD39A804CA1087EAB" pageNumber="660" type="child_taxa">
<paragraph id="4F76B7180C8AAA182CBE2F44ADDA3411" pageNumber="660">
<emphasis id="75D229AD677E4BDB4D419EEF6E37D0A6" inLineHeading="true" pageNumber="660">Subspecies:</emphasis>
:
</paragraph>
<paragraph id="03D1460C5C5BBEC38DCFD7B1FAD96C4D" pageNumber="660">
Subspecies
<taxonomicName id="0ED7CA4222EE3A4C02B629FDE3388745" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="tarandus">
<emphasis id="ECDBB728895E8B52804D74414107609A" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="571B46C0DD9CDC2815757E287687108C" italics="true" pageNumber="660">tarandus</emphasis>
<bibRefCitation id="9E6EB27907999B3C8CFCCA2EF4A813FF" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="FD66DB14E87BFC1E01781255AB35206C" pageNumber="660">
Subspecies
<taxonomicName id="B9B83B94BEA6F321BC9BFCA35E568F49" authority="Millais 1915" authorityName="Millais" authorityYear="1915" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="buskensis">
<emphasis id="2E7879CE6CEEFD15184A47D8193CC104" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="DA93B7E4B763375F160E4EFB568321F8" italics="true" pageNumber="660">buskensis</emphasis>
Millais 1915
</taxonomicName>
</paragraph>
<paragraph id="8E782BB9C8D1054B9D194DCFA425FA90" pageNumber="660">
Subspecies
<taxonomicName id="0DE201412F04231D91908AB2B21B4B6A" authority="G. M. Allen 1914" authorityName="G. M. Allen" authorityYear="1914" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="caboti">
<emphasis id="BDA5D097068BF3A77ECB241CC57F93A8" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="E187A81432214094B50286CCFF9C743A" italics="true" pageNumber="660">caboti</emphasis>
G. M. Allen 1914
</taxonomicName>
</paragraph>
<paragraph id="B0D109899C24A9A1EE84738132CCD810" pageNumber="660">
Subspecies
<taxonomicName id="CC1E435BE39B78411DA6E7A426173227" authority="Gmelin 1788" authorityName="Gmelin" authorityYear="1788" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="caribou">
<emphasis id="D3585AEF397B11131ADE22C5BE2097E1" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="891FC0EF20E9A16B18C82511CAED4656" italics="true" pageNumber="660">caribou</emphasis>
Gmelin 1788
</taxonomicName>
</paragraph>
<paragraph id="E166DB49D2647A1F63A1D41CCCDA6BF3" pageNumber="660">
Subspecies
<taxonomicName id="7D1A827352A1D545BC076F47DC357A2C" authority="Thompson-Seton 1900" authorityName="Thompson-Seton" authorityYear="1900" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="dawsoni">
<emphasis id="13789409A5A17E292F00BFBC35823CA9" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="5DF04BDDC77B0852C3057D1BE0384ED7" italics="true" pageNumber="660">dawsoni</emphasis>
Thompson-Seton 1900
</taxonomicName>
</paragraph>
<paragraph id="7424F95841CCD4E4C95CA0A595D14809" pageNumber="660">
Subspecies
<taxonomicName id="9E0ADF28DFE835EBEC0EEFC3D24B6F12" authority="Lönnberg 1909" authorityName="Lönnberg" authorityYear="1909" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="fennicus">
<emphasis id="792B1DCEC919C875F889A8A571D7242C" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="C08D303FCE220BB76C43ACE145E858B1" italics="true" pageNumber="660">fennicus</emphasis>
Lönnberg 1909
</taxonomicName>
</paragraph>
<paragraph id="38C28722F50335A72B98B082AD487B03" pageNumber="660">
Subspecies
<taxonomicName id="02D3783EBA3648C3D274F5196EF538BC" authority="Borowski 1780" authorityName="Borowski" authorityYear="1780" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="groenlandicus">
<emphasis id="D42DC8C69ADD45397024196A2FE87C6A" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="FB1A4E96CE97324AA0A040C91A165512" italics="true" pageNumber="660">groenlandicus</emphasis>
Borowski 1780
</taxonomicName>
</paragraph>
<paragraph id="F0954983B2404B3596231483C6274393" pageNumber="660">
Subspecies
<taxonomicName id="81E796F59E6AB726D9CF7E5EDD4A9FC8" authority="J. A. Allen 1902" authorityName="J. A. Allen" authorityYear="1902" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="osborni">
<emphasis id="5722D8FD1E6879692B3D45C28ED1ACC9" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="87DDB8ABF9E4A02E9D7F847174896243" italics="true" pageNumber="660">osborni</emphasis>
J. A.
<bibRefCitation id="D629BE6C5311EC7759E33F427010526A" author="Allen, J. A." refId="ref5572" refString="Allen, J. A. 1902. Mammal names proposed by Oken in his &quot; Lehrbuch der Zoologie &quot;. Bulletin of the American Museum of Natural History, 16: 373 - 379." year="1902">Allen 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1561FA35BB6951E736A5F6EA86F0587E" pageNumber="660">
Subspecies
<taxonomicName id="AE23EC753A08B15CE9CA644AA9218A9D" authority="Lydekker 1903" authorityName="Lydekker" authorityYear="1903" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="pearsoni">
<emphasis id="B6E307F4705506A4DE754FBF04C8D8B6" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="08D146BE5743E93BF040007E71B94781" italics="true" pageNumber="660">pearsoni</emphasis>
Lydekker 1903
</taxonomicName>
</paragraph>
<paragraph id="712A98384738331F5F93487832FD45BE" pageNumber="660">
Subspecies
<taxonomicName id="DB2FD68CB91A5A2768CF5FDEC3EE07F1" authority="J. A. Allen 1902" authorityName="J. A. Allen" authorityYear="1902" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="pearyi">
<emphasis id="5AD3C386CC049D2C01124B56DCDBB73C" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="703809AC1C06883155E9A560E6D0E5D1" italics="true" pageNumber="660">pearyi</emphasis>
J. A.
<bibRefCitation id="2848E303CB865AF5214B7F2CDEBAA682" author="Allen, J. A." refId="ref5572" refString="Allen, J. A. 1902. Mammal names proposed by Oken in his &quot; Lehrbuch der Zoologie &quot;. Bulletin of the American Museum of Natural History, 16: 373 - 379." year="1902">Allen 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="0708353D335BE9C1EC21BA781CC25E94" pageNumber="660">
Subspecies
<taxonomicName id="E94F7D1BE57C1A304016EF235873BCEB" authority="Hollister 1912" authorityName="Hollister" authorityYear="1912" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="phylarchus">
<emphasis id="1E3429D40A3B72E51174CEE6BDB63F9C" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="DA2B1A199C717AF95BE0231E3EAB1DE4" italics="true" pageNumber="660">phylarchus</emphasis>
<bibRefCitation id="64DAFFA73FD4BD202052718DBF6D0866" author="Hollister, N." refId="ref156830" refString="Hollister, N. 1912. The names of the Rocky Mountain goat. Proceedings of the Biological Society of Washington, 25: 185 - 186." year="1912">Hollister 1912</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="CCF9C0106F95ABAC853785C56111C4A9" pageNumber="660">
Subspecies
<taxonomicName id="5D989CE31FF7B91066424D5C99D6762F" authority="Vrolik 1829" authorityName="Vrolik" authorityYear="1829" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="platyrhynchus">
<emphasis id="2272422F18C4BF49FA8390C9B960AAEC" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="972F449D55A283D74199EE21FFDFAAE8" italics="true" pageNumber="660">platyrhynchus</emphasis>
Vrolik 1829
</taxonomicName>
</paragraph>
<paragraph id="B6B04C01A1E55835475C7ED66B5B0425" pageNumber="660">
Subspecies
<taxonomicName id="1BF6FF9765D756E8090A532B08CA6C73" authority="Murray 1866" authorityName="Murray" authorityYear="1866" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="sibiricus">
<emphasis id="327DF301EB18BC661A2691D666044AAB" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="E7C512EB90718FAB9B4770FF7BBA7266" italics="true" pageNumber="660">sibiricus</emphasis>
<bibRefCitation id="172CC74DD302374927391EF71158045E" author="Murray, A." refId="ref250505" refString="Murray, A. 1866. The geographical distribution of mammals. Day and Son, Ltd., London, 420 pp." year="1866">Murray 1866</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="9169555083A3ABA149B0F6C7C2F0C4C7" pageNumber="660">
Subspecies
<taxonomicName id="B083F46687CE06CC4A965A707FD8867E" authority="Bangs 1896" authorityName="Bangs" authorityYear="1896" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageNumber="660" phylum="Chordata" rank="subSpecies" species="tarandus" subSpecies="terraenovae">
<emphasis id="371434B963F8ACD768B154F37FABB461" italics="true" pageNumber="660">Rangifer tarandus</emphasis>
subsp.
<emphasis id="E8FBB1FD1D53C1293593EB422641C91D" italics="true" pageNumber="660">terraenovae</emphasis>
Bangs 1896
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="30BD829577A2B79F85D8E966A1451003" pageNumber="660" type="distribution">
<paragraph id="938C3553CCB45163919A402ACCA0BFD9" pageNumber="660">
<emphasis id="3C1B01B3B81366BD4156A3979304C233" inLineHeading="true" pageNumber="660">Distribution:</emphasis>
Circumboreal in tundra and taiga from
<collectingCountry id="2E008EADC67F1EDC2D6E6D5143D6559F" name="Svalbard and Jan Mayen">Svalbard</collectingCountry>
,
<collectingCountry id="97FB5401D5A517B5BAC993BC69EEB60E" name="Norway">Norway</collectingCountry>
,
<collectingCountry id="0DCD85E814A0244D3BD881815653E775" name="Finland">Finland</collectingCountry>
,
<collectingCountry id="347A016733C0E58BD8A97F02250968C8" name="Russia">Russia</collectingCountry>
,
<collectingRegion id="7C37E1AF14C935D80E7EA80ABF575DC9" country="United States of America" name="Alaska">Alaska</collectingRegion>
(
<collectingCountry id="F263BFC18B24981F0B0B381028A269EA" name="United States of America">USA</collectingCountry>
) and
<collectingCountry id="DC3ACEE62C4F5A2AC7C348E82DA934BA" name="Canada">Canada</collectingCountry>
including most arctic islands, and
<collectingCountry id="454D34A12F6822DF132691F2594C0B6A" name="Greenland">Greenland</collectingCountry>
, south to N
<collectingCountry id="447B0873CAD7B31CD59F32D939F8F8C1" name="Mongolia">Mongolia</collectingCountry>
,
<collectingCountry id="1753E3363615EB2B20CDD4C1F705481F" name="China">China</collectingCountry>
(
<collectingRegion id="D94272AE458514EA3EB9D5FA0B42D336" country="China" name="Nei Mongol">Inner Mongolia</collectingRegion>
; now only domesticated or feral?), Sakhalin Isl, and
<collectingCountry id="CD13851299B4F8F8AC792623571CA608" name="United States of America">USA</collectingCountry>
(N
<collectingRegion id="4FE20EBE095BF36CE7204834B94AAC89" country="United States of America" name="Idaho">Idaho</collectingRegion>
and Great Lakes region). Introduced to, and feral in,
<collectingCountry id="B630D22352E05326D53CDE57D8D1EA02" name="Iceland">Iceland</collectingCountry>
, Kerguelen Isls,
<collectingCountry id="59D6EE45B3203EF2E0430D62A811E463" name="South Georgia and the South Sandwich Islands">South Georgia</collectingCountry>
Isl, Pribilof Isls, St. Matthew Isl. Extinct in
<collectingCountry id="455D8EB312E552FEE6E4F8D7C20252B5" name="Sweden">Sweden</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3203353EA93AA111BA487D33173A2CDA" pageNumber="660" type="conservation">
<paragraph id="6C716329D95620C1A0F3F83BD134AC19" pageNumber="660">
<emphasis id="A2DD9595E09D35570FB4B3F53B23B216" inLineHeading="true" pageNumber="660">Conservation:</emphasis>
<collectingCountry id="21B0BCE314D9F6F0F540D03B914A1B6D" name="United States of America">U.S.</collectingCountry>
<collectionCode id="80D95D04964D18D5E46CFDE73791BE32" country="Brazil" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" type="Herbarium">ESA</collectionCode>
– Endangered as
<emphasis id="86EC8322C9A9E3BA4B9E5F5B3123E883" italics="true" pageNumber="660">
<collectionCode id="70592D7EA5CA3250C7E522EA263329DF" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. t. caribou
</emphasis>
in
<collectingCountry id="3AF2287E3CDCDCD20C05AC27BFE25929" name="Canada">Canada</collectingCountry>
(
<collectionCode id="697A9B802E54C89AAC04958F1242475C">SE</collectionCode>
<collectingRegion id="112612C6B1C56B2146D4E0BE318EC6CC" country="Canada" name="British Columbia">British Columbia</collectingRegion>
at the Canadian-USA border, Columbia River, Kootenay River, Kootenay Lake, and Kootenai River) and
<collectingCountry id="DBE03D852A453E73D155DFE270F28972" name="United States of America">USA</collectingCountry>
(
<collectingRegion id="96E96FD1C91B2DAE8AD9A254AA9D22AD" country="United States of America" name="Idaho">Idaho</collectingRegion>
,
<collectingRegion id="C957FA6DFEC46ED111D69D1024AE7551" country="United States of America" name="Washington">Washington</collectingRegion>
);
<collectionCode id="D3525E17335C5645C00F513C0FF3CAAD">IUCN</collectionCode>
– Endangered as
<emphasis id="A2BEB6B12B78F36A9609E0A53427F352" italics="true" pageNumber="660">
<collectionCode id="D68889844598903730F339EE696B64D6" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. t. pearyi
</emphasis>
, otherwise Lower Risk (lc). The woodland
<emphasis id="6F48AA9584F4A0F9158BB290573C0C32" italics="true" pageNumber="660">caribou</emphasis>
is highly endangered throughout its distribution right into Ontario (
<collectionCode id="19E1B31528773EF586991B03613CC4EB" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. Geist, in litt.).
</paragraph>
</subSubSection>
<subSubSection id="568A1382CF17E13BB548D04DBC596002" pageNumber="660" type="discussion">
<paragraph id="3DABFAFBFD2A55EC05D2E07F2C456085" pageNumber="660">
<emphasis id="E353D41EECA3B1BD9F024EBF83496DFC" inLineHeading="true" pageNumber="660">Discussion:</emphasis>
Subspecies have been placed in two divisions, compressicornis or Woodland Reindeer, and cylindricornis or Tundra Reindeer (
<bibRefCitation id="4C8EB3571F7C85C262ED3D0BFBC2AFDD" author="Jacobi, A." refId="ref170634" refString="Jacobi, A. 1931. Das Rentier; eine zoologische Monographie der Gattung Rangifer. Zoologischer Anzeiger, Erganzungsband zu Band 96, 264 pp." year="1931">Jacobi, 1931</bibRefCitation>
). These names of divisions are non-Linnean;
<emphasis id="D6D768F518904F0D3E89AA7E62C65575" italics="true" pageNumber="660">cilindricornis</emphasis>
Camerano, 1902 is a lapsus for cylindricornis and is not a Linnean name. An additional category has since been recognised for the Peary Caribou, due to marginal or seasonal sympatry between caribou in Arctic America, following
<bibRefCitation id="2A2C34D08ACFAF2B44F4EB5BFEFD6A61" author="Banfield, A. W. F." refId="ref19636" refString="Banfield, A. W. F. 1963. The post glacial dispersal of American caribou. Proceedings of the 16 th International Congress of Zoology, Washington D. C., 1: 206." year="1963">Banfield (1963)</bibRefCitation>
. Subspecies here considered valid are based on
<bibRefCitation id="E5B1C0809F0C6A7948B959EE73F3B336" author="Banfield, A. W. F." refId="ref19606" refString="Banfield, A. W. F. 1961. A revision of the reindeer and caribou. National Museum of Canada Bulletin, 177: 1 - 137." year="1961">Banfield (1961)</bibRefCitation>
, considerably modified by
<bibRefCitation id="9CE5B56D0138B977816469D9C5BDA684" author="Geist, V." refId="ref116300" refString="Geist, V. 1998. Deer of the world. Their evolution, behavior and ecology. Stackpole Books, Mechanicsburg, Pensylvania, 421 pp." year="1998">Geist (1998)</bibRefCitation>
:
<emphasis id="0D0159C13EBD29BEE31CF13CD5D521F8" italics="true" pageNumber="660">caribou</emphasis>
division or Woodland Caribou (includes also
<emphasis id="1F92EAE106DC41E4707361775D8FA7DD" italics="true" pageNumber="660">buskensis, valentinae, dawsoni, fennicus</emphasis>
, and
<emphasis id="14C0093DA672AA4FFC9812C627A702ED" italics="true" pageNumber="660">phylarchus</emphasis>
); populations transitional between
<emphasis id="2456C0696E472B19C323D6F60866F926" italics="true" pageNumber="660">caribou</emphasis>
and
<taxonomicName id="EB75582CB3D774677E314D4F8B349C4D" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="5033D8E2289D6DC7DF63D7312F4ED53C" italics="true" pageNumber="660">tarandus</emphasis>
</taxonomicName>
divisions (includes
<emphasis id="CE4AF80831791DA35AABF77473D4D535" italics="true" pageNumber="660">osborni</emphasis>
);
<taxonomicName id="A5108D08CEF48C67F4F73D7B5865BF89" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="326B66DB3CA54458D02049E38608BAB0" italics="true" pageNumber="660">tarandus</emphasis>
</taxonomicName>
division, Barren-ground Caribou or Reindeer (includes also
<emphasis id="FE73977BCF5B7003E3D0C2CB451D63BD" italics="true" pageNumber="660">caboti, groenlandicus, pearsoni, sibiricus</emphasis>
, and
<emphasis id="BFA19CFD8DA1DD320EFCED47FC209456" italics="true" pageNumber="660">terraenovae</emphasis>
); and
<emphasis id="28390C299DD1F494CFBD97ED4B5441DA" italics="true" pageNumber="660">platyrhynchus</emphasis>
division (including
<emphasis id="A4B4270A8EE3E9421E8A4FF612FDC322" italics="true" pageNumber="660">pearyi</emphasis>
or Peary Caribou and
<emphasis id="C421C887605529F5283D08BCCA9A4104" italics="true" pageNumber="660">platyrhynchus</emphasis>
or
<collectingCountry id="E8D492F5E28FE24107C1CE66C0006034" name="Svalbard and Jan Mayen">Svalbard</collectingCountry>
Reindeer). The extinct insular
<emphasis id="6B6F406684E6A484AF997C904DCFA46D" italics="true" pageNumber="660">dawsoni</emphasis>
has been treated as a distinct species (
<bibRefCitation id="3878A8BA6664D5AA4FA9575BFF19E544" author="Cowan, I. McT. &amp; C. J. Guiguet" refId="ref70157" refString="Cowan, I. McT., and C. J. Guiguet. 1965. The mammals of British Columbia. Third Ed. British Columbia Provincial Museum, Handbook, 11: 1 - 414." year="1965">Cowan and Guiguet, 1965</bibRefCitation>
) but does not differ from
<emphasis id="33D53363FB7190812579AB2B4E7E0856" italics="true" pageNumber="660">caribou</emphasis>
or
<taxonomicName id="5208CA029246343B5F81645089244190" baseAuthorityName="Brooke" baseAuthorityYear="1872" class="Mammalia" family="Bovidae" genus="Nanger" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="species" species="granti">
<emphasis id="84899E47E39C92E07B19595449DE6747" italics="true" pageNumber="660">granti</emphasis>
</taxonomicName>
(=
<emphasis id="F06C4E2C462A91DCEE399C7E147FD52A" italics="true" pageNumber="660">groenlandicus</emphasis>
) in mtDNA sequences (
<bibRefCitation id="FEAA551ABB5392E15B522452FBAAC4E3" author="Byun, S. A. &amp; B. F. Koop &amp; T. E. Reimchen" refId="ref48266" refString="Byun, S. A., B. F. Koop, and T. E. Reimchen. 2002. Evolution of the Dawson caribou (Rangifer tarandus dawsoni). Canadian Journal of Zoology, 80: 956 - 960." year="2002">Byun et al., 2002</bibRefCitation>
). Grouping the
<collectingCountry id="0E71A9B35BCB204A0E9BCB3708F50A39" name="Svalbard and Jan Mayen">Svalbard</collectingCountry>
Reindeer with the Peary Caribou is provisional (
<bibRefCitation id="BA8A472C8CD3C7E3F061C911CB5BA4CB" author="Groves, C. P. &amp; P. Grubb" refId="ref133049" refString="Groves, C. P., and P. Grubb. 1987. Relationships of living deer. Pp. 2 l- 59, in Biology and management of the Cervidae (C. M. Wemmer, ed.). Smithsonian Institution Press, Washington, D. C., 577 pp." year="1987">Groves and Grubb, 1987</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C02F942DBD19475599C675C557996EEC">A revision of the parasitoid wasp genus Alphomelon Mason with the description of 30 new species (Hymenoptera, Braconidae)</mods:title>
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<treatment id="C1A875C9524D52658141EAF9EDFCABDF" LSID="urn:lsid:plazi:treatment:C1A875C9524D52658141EAF9EDFCABDF" httpUri="http://treatment.plazi.org/id/C1A875C9524D52658141EAF9EDFCABDF" lastPageNumber="5" pageId="0" pageNumber="5">
<subSubSection id="C91A5AF2517905ADE65430C2AB9D2277" pageId="0" pageNumber="5" type="nomenclature">
<paragraph id="883284E6F91451E702DF95B5FA57C8AA" pageId="0" pageNumber="5">
<taxonomicName id="66B2D0C7ED38D4372A4B21BD516E7612" ID-CoL="C69G" ID-ENA="496401" LSID="C1A875C9-524D-5265-8141-EAF9EDFCABDF" authority="Deans, 2003" authorityName="Deans" authorityYear="2003" class="Insecta" family="Braconidae" genus="Alphomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alphomelon xestopyga" order="Hymenoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="xestopyga">Alphomelon xestopyga Deans, 2003</taxonomicName>
</paragraph>
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<paragraph id="2D382F54EFE5F5864B737F0572A15E91" pageId="0" pageNumber="5">
<figureCitation id="CB1C1CB78BDCF3EED1EED785CE79D5FA" captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Alphomelon xestopyga Deans female DHJPAR 0031632 A habitus, lateral B metasoma, lateral C head, frontal D wings E head, lateral F metasoma, dorsal G mesosoma, dorsal." figureDoi="10.3897/zookeys.1175.105068.figure47" httpUri="https://binary.pensoft.net/fig/893149" pageId="0" pageNumber="5">Figs 47A-G</figureCitation>
<figureCitation id="EC45F14509BF4238568C8A7A6E4D0057" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Alphomelon xestopyga Deans female CNC 308749 A habitus, lateral B head, frontal C head, lateral D wings E metasoma, dorsal F mesosoma, dorsal." figureDoi="10.3897/zookeys.1175.105068.figure86" httpUri="https://binary.pensoft.net/fig/893188" pageId="0" pageNumber="5">, 86A-F</figureCitation>
<figureCitation id="2754CAF54E3B06C3F21DD3DF73D5DA45" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Alphomelon xestopyga Deans female CNC 721122 A habitus, lateral B head, frontal C antenna, lateral D wings E propodeum and metasoma, dorsal F mesosoma, dorsal." figureDoi="10.3897/zookeys.1175.105068.figure87" httpUri="https://binary.pensoft.net/fig/893189" pageId="0" pageNumber="5">, 87A-F</figureCitation>
<figureCitation id="3098BD6EA05ACBB8EA96AC0489ACFA29" captionStart="Figure 88" captionStartId="F88" captionText="Figure 88. Alphomelon xestopyga Deans female CNC 1620777 A habitus, lateral B head, lateral C head, frontal D wings E metasoma, dorsal F head and mesosoma, dorsal." figureDoi="10.3897/zookeys.1175.105068.figure88" httpUri="https://binary.pensoft.net/fig/893190" pageId="0" pageNumber="5">, 88A-F</figureCitation>
<figureCitation id="7405CF0B6D53755D8D4F737E97A6A7E5" captionStart="Figure 89" captionStartId="F89" captionText="Figure 89. Alphomelon xestopyga Deans holotype female USNMENT 00828302 A habitus, lateral B habitus, dorsal C close-up of habitus, lateral." figureDoi="10.3897/zookeys.1175.105068.figure89" httpUri="https://binary.pensoft.net/fig/893191" pageId="0" pageNumber="5">, 89A-C</figureCitation>
<figureCitation id="A8B3E6CF0CBEA4BFEDDCC2F8673B6B6D" captionStart="Figure 103" captionStartId="F103" captionText="Figure 103. Parasite cocoons and caterpillars A Alphomelon talidicida 17 - SRNP- 30054 - DHJ 735829 B Alphomelon xestopyga 16 - SRNP- 55580 - DHJ 492508." figureDoi="10.3897/zookeys.1175.105068.figure103" httpUri="https://binary.pensoft.net/fig/893205" pageId="0" pageNumber="5">, 103B</figureCitation>
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<subSubSection id="DF601433640F4A984B2532A5EBD55D10" pageId="0" pageNumber="5" type="distribution">
<paragraph id="AA3EDFE486DB9356F5C1588E6A1CBD76" pageId="0" pageNumber="5">Distribution.</paragraph>
<paragraph id="9FD756E7E6A2C18929EE234142994BAF" pageId="0" pageNumber="5">Costa Rica (ACG).</paragraph>
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<subSubSection id="45D08BA7821440168E9C24AC954A87E1" pageId="0" pageNumber="5" type="biology_ecology">
<paragraph id="63D45AF6C90931EFF26557164D009160" pageId="0" pageNumber="5">Biology.</paragraph>
<paragraph id="D9FD5CF597AE093C6E71648962AA9754" pageId="0" pageNumber="5">
Gregarious, reared from
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<emphasis id="59650C78C0993C284E9B7F1D51D0E32D" italics="true" pageId="0" pageNumber="5">Calpodes esperi</emphasis>
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,
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<emphasis id="CD3E6BB85A3B570CB94583A18DD5AD4F" italics="true" pageId="0" pageNumber="5">C. ethlius</emphasis>
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,
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<emphasis id="4C948E0BFEB8C9954689C35E80D702B6" italics="true" pageId="0" pageNumber="5">C. fusta</emphasis>
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,
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<emphasis id="0D241CE3C06769238635A1A9C7470342" italics="true" pageId="0" pageNumber="5">C. severus</emphasis>
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,
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<emphasis id="26273D6158BF22895CDA165CB56E74A5" italics="true" pageId="0" pageNumber="5">Cobalopsis nero</emphasis>
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,
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<emphasis id="4A34F4F98D52947E324F7A9AD20B51F6" italics="true" pageId="0" pageNumber="5">Cynea irma</emphasis>
</taxonomicName>
,
<taxonomicName id="81C4796650D274E93C6B7267DDFEB2CF" baseAuthorityName="Mabille" baseAuthorityYear="1891" class="Insecta" family="Hesperiidae" genus="Cymaenes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cymaenes odilia subsp. trebius" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthopoda" rank="subSpecies" species="odilia" subSpecies="trebius">
<emphasis id="BE31871099C9CB0F2D3FDE8FA5E98C44" italics="true" pageId="0" pageNumber="5">Cymaenes odilia trebius</emphasis>
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,
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<emphasis id="4A1AE2A554B73330F40925AC4EA19BB6" italics="true" pageId="0" pageNumber="5">C.</emphasis>
</taxonomicName>
Burns01,
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<emphasis id="F97F7EE916402496A8963B0DD17E8448" italics="true" pageId="0" pageNumber="5">Joanna joanna</emphasis>
</taxonomicName>
,
<taxonomicName id="FA5B5EE64B5B810462BE5FCE9217396D" authorityName="Evans" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Justinia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Justinia norda" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="norda">
<emphasis id="2A4E89EC4BC1E757FE4D68F3377C650C" italics="true" pageId="0" pageNumber="5">Justinia norda</emphasis>
</taxonomicName>
,
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<emphasis id="16DA4437DB190C2FF6306BCC6382C10C" italics="true" pageId="0" pageNumber="5">Morys lyde</emphasis>
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,
<taxonomicName id="E12C5793BE3D2F55F1F9E6CE22F90E33" lsidName="M. micythus" pageId="0" pageNumber="5" rank="species" species="micythus">
<emphasis id="AE38AE7132FF4089B51FA854B1A752B5" italics="true" pageId="0" pageNumber="5">M. micythus</emphasis>
</taxonomicName>
,
<taxonomicName id="699C81A867869519B10FFDDF2BFE8269" lsidName="M. valerius subsp. valda" pageId="0" pageNumber="5" rank="subSpecies" species="valerius" subSpecies="valda">
<emphasis id="3C45A6001D19B8D2950979DE56AF52D6" italics="true" pageId="0" pageNumber="5">M. valerius valda</emphasis>
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,
<taxonomicName id="BA26E95579BAECAD65AEAFEF6886EDFD" genus="M." lsidName="M." pageId="0" pageNumber="5" rank="genus">
<emphasis id="A8FD93608F30CC18AF974052563304D3" italics="true" pageId="0" pageNumber="5">M.</emphasis>
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lydeDHJ01,
<taxonomicName id="F5CACD886EFF246C4F077A141F497C20" genus="M." lsidName="M." pageId="0" pageNumber="5" rank="genus">
<emphasis id="443F0743E810D99ABD0F1FB5D85DB811" italics="true" pageId="0" pageNumber="5">M.</emphasis>
</taxonomicName>
lydeDHJ02,
<taxonomicName id="88066A34B0B76900CE2E6365BA454FD8" authorityName="Austin" authorityYear="1997" class="Insecta" family="Hesperiidae" genus="Niconiades" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Niconiades incomptus" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="3F959D75A3A9058A3F1BCF8495342325" italics="true" pageId="0" pageNumber="5">Niconiades incomptus</emphasis>
</taxonomicName>
,
<taxonomicName id="CB03833A0E1BF407005FF160EBC6DF93" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" class="Insecta" family="Hesperiidae" genus="Parphorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parphorus decora" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="E31626AE1FEFF6D149720608F9276B31" italics="true" pageId="0" pageNumber="5">Parphorus decora</emphasis>
</taxonomicName>
,
<taxonomicName id="E9AE10F7B73CEAD40CF4614407FCCD8B" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" class="Insecta" family="Hesperiidae" genus="Quinta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quinta cannae" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="cannae">
<emphasis id="164579D67E0C3A1468D13116D7A5C62D" italics="true" pageId="0" pageNumber="5">Quinta cannae</emphasis>
</taxonomicName>
,
<taxonomicName id="7BC5519956E7023C2C85C4CE4D476AD8" lsidName="R. osca" pageId="0" pageNumber="5" rank="species" species="osca">
<emphasis id="4EE271E3B775412AC204AE6E66F14A51" italics="true" pageId="0" pageNumber="5">R. osca</emphasis>
</taxonomicName>
,
<taxonomicName id="6432857DB6B01C347B8372A33DE9B0F3" baseAuthorityName="Mabille" baseAuthorityYear="1883" class="Insecta" family="Hesperiidae" genus="Synapte" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Synapte salenus" order="Lepidoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="salenus">
<emphasis id="F528C8F98A2E4AD83C549E29EA10D73B" italics="true" pageId="0" pageNumber="5">Synapte salenus</emphasis>
</taxonomicName>
,
<taxonomicName id="9DF86ACFEEEB51426BBB5A0B959C3047" lsidName="S. silius" pageId="0" pageNumber="5" rank="species" species="silius">
<emphasis id="657789CBCB87B800B3CBC83D35224C3B" italics="true" pageId="0" pageNumber="5">S. silius</emphasis>
</taxonomicName>
, and
<taxonomicName id="C619322B1BA99AF1C2E65FB61431AFC2" class="Insecta" family="Poaceae" genus="Vettius" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vettius aurelius" order="Poales" pageId="0" pageNumber="5" phylum="Liliopsida" rank="species" species="aurelius">
<emphasis id="0F00D9FC9FD4E1D118846B0266BC4FAE" italics="true" pageId="0" pageNumber="5">Vettius aurelius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2F6A9D831725F100FAF172B056452747" pageId="0" pageNumber="5" type="dna barcoding">
<paragraph id="E2856C4316AE9F44A62D43809D24FD6C" pageId="0" pageNumber="5">DNA barcoding.</paragraph>
<paragraph id="97974DC8C1D51DD0F334E1A4ABE947E0" pageId="0" pageNumber="5">BINBOLD: AAA1634.</paragraph>
</subSubSection>
<subSubSection id="11EA7EC9246D71C66967F5953F9EB4D5" pageId="0" pageNumber="5" type="materials_examined">
<paragraph id="AB81730DAF0535F4052799E1374C2168" pageId="0" pageNumber="5">Other specimens examined</paragraph>
<paragraph id="E603707EB66D1D2332B78F2F2C3B6FF2" pageId="0" pageNumber="5">
(
<specimenCount id="F745F3D7F2B7EB01947D42AF3A03D684" type="female">93 females</specimenCount>
,
<specimenCount id="840E160442B35DBA0470B31CD2B1A87D" type="male">32 males</specimenCount>
): CNC308749, CNC308837, CNC308838, CNC308839, CNC308840 (additional specimens in a gel capsule associated with that specimen), DHJPAR0049842, CNC1620806, CNC1620807, CNC1620808, CNC1620809, CNC1620810, CNC1620811, DHJPAR0051223, CNC1620802, CNC1620803, CNC1620804, CNC1620805, DHJPAR0031632, DHJPAR0053792, DHJPAR0053789, DHJPAR0053751, CNC1620772, CNC1620773, CNC1620774, CNC1620775, CNC1620776, DHJPAR0053767, CNC1620777, DHJPAR0020278, CNC958681, CNC958682 (additional specimens in a gel capsule associated with that specimen), DHJPAR0053760, CNC1620778, CNC1620779, CNC1620780, CNC1620781, CNC1620782 (additional specimens in a gel capsule associated with that specimen), CNC1620783, CNC1620784, CNC1620785, CNC1620786, CNC1620787, CNC1620788, CNC1620789, CNC1620790, DHJPAR0031616, DHJPAR0031735, DHJPAR0031620, DHJPAR0052957, CNC1620791, CNC1620792, CNC1620793, DHJPAR0053758, CNC1620794, CNC1620795, CNC1620796, CNC1620797, CNC1620798, CNC1620812, CNC1620813, CNC1620814, CNC1620815, CNC1620816, CNC1620817, CNC1620818, CNC1620819, CNC1620820, CNC1620821, CNC1620822, CNC1620823, CNC1620824, CNC1620825, CNC1620826, CNC1620827, CNC1620828, CNC1620829, CNC1620830, CNC1802047, CNC1802048, DHJPAR0053755, CNC1620837, CNC1620831, CNC1620832, CNC1620833, CNC1620834, CNC1620835, CNC1620836, CNC1620799, CNC1620800, CNC1620838, CNC1620839, CNC1620840, CNC1620801, CNC1620841, CNC1620842, CNC1620843, CNC1620844, CNC721102, CNC721114, CNC721118, CNC721119, CNC721122, CNC721123, CNC721125, CNCHYM 00061, CNC721103, CNC721104, CNC721106, CNC721107, CNC721108, CNC721112, CNC721101, CNC721113, CNC721115, CNC721116, CNC721117, CNC721120, CNC721121, CNC721124, CNC721126, CNC280525, CNC721105, CNC721109, CNC721110, CNC721111.
</paragraph>
<caption id="29F138ACFF3E68F2E7C9EA5AF574AEDD" doi="10.3897/zookeys.1175.105068.figure86" httpUri="https://binary.pensoft.net/fig/893188" pageId="0" pageNumber="5" start="Figure 86" startId="F86">
<paragraph id="D74E68F7EC7A82F25DF6F52BB8E6CDBA" pageId="0" pageNumber="5">
<emphasis id="84E0895D393181FAE128F318F1E20F70" bold="true" pageId="0" pageNumber="5">Figure 86.</emphasis>
<taxonomicName id="3F1969D91D147BC73AC2EC1CA9C825A8" authorityName="Deans" authorityYear="2003" class="Insecta" family="Braconidae" genus="Alphomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alphomelon xestopyga" order="Hymenoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="xestopyga">
<emphasis id="7116BC9A120279D323320E371F1E993D" italics="true" pageId="0" pageNumber="5">Alphomelon xestopyga</emphasis>
</taxonomicName>
Deans female CNC308749
<emphasis id="4FEECD4F4A2DEF64025E4E7D53F0AA04" bold="true" pageId="0" pageNumber="5">A</emphasis>
habitus, lateral
<emphasis id="66D22D1EA038976348B6AF5475FFE311" bold="true" pageId="0" pageNumber="5">B</emphasis>
head, frontal
<emphasis id="73B2F8804F9E4E25C2C61561C9C42A99" bold="true" pageId="0" pageNumber="5">C</emphasis>
head, lateral
<emphasis id="F2D2F31A3BB5171416E9551C1FB6F08D" bold="true" pageId="0" pageNumber="5">D</emphasis>
wings
<emphasis id="354CEFF05748DC246A95AA179CEA0659" bold="true" pageId="0" pageNumber="5">E</emphasis>
metasoma, dorsal
<emphasis id="602DD25F995D911BCB766AB826400084" bold="true" pageId="0" pageNumber="5">F</emphasis>
mesosoma, dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9EEAA9C130A362E61D37A27E64D613DA" pageId="0" pageNumber="5" type="notes">
<paragraph id="D395D2F39F0CB801D2E5CDF9EAD55039" pageId="0" pageNumber="5">Notes.</paragraph>
<paragraph id="32728D0F5C47B312D2555A0E08F78B43" pageId="0" pageNumber="5">
There are some differences between
<taxonomicName id="2A9F0DE892C919C050F83FEDC5A1D52F" lsidName="A. xestopyga" pageId="0" pageNumber="5" rank="species" species="xestopyga">
<emphasis id="81DAF904FB191D3A9EF7763ED661858E" italics="true" pageId="0" pageNumber="5">A. xestopyga</emphasis>
</taxonomicName>
in the sense of
<bibRefCitation id="0FB616252459F6B2F844E7E6AC2C7D8D" author="Deans, AR" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="5" pagination="1 - 41" publicationUrl="https://www.biodiversitylibrary.org/part/32897" refId="B3" refString="Deans, AR, Whitfield, JB, Janzen, DH, 2003. Taxonomy and natural history of the Microgastrinae genus Alphomelon Mason (Hymenoptera: Braconidae). Journal of Hymenoptera Research 12 (1): 1 - 41, https://www.biodiversitylibrary.org/part/32897" title="Taxonomy and natural history of the Microgastrinae genus Alphomelon Mason (Hymenoptera: Braconidae)." url="https://www.biodiversitylibrary.org/part/32897" volume="12" year="2003">Deans et al. (2003)</bibRefCitation>
and the ACG specimens identified as such that we have been able to examine. The species clearly comprises a complex of species, with some specimens we have seen in the CNC collection from other countries besides Costa Rica. Solving this potential complex will require obtaining more DNA barcodes, especially from areas outside of ACG.
</paragraph>
<caption id="069197C03A8EFD057352FFBD631097D3" doi="10.3897/zookeys.1175.105068.figure87" httpUri="https://binary.pensoft.net/fig/893189" pageId="0" pageNumber="5" start="Figure 87" startId="F87">
<paragraph id="0B141602166AFAC189B297002F2579A6" pageId="0" pageNumber="5">
<emphasis id="8C02700C43885C916D63A58F3BFD5801" bold="true" pageId="0" pageNumber="5">Figure 87.</emphasis>
<taxonomicName id="41B40250C3C884211F6C8E454278E18D" authorityName="Deans" authorityYear="2003" class="Insecta" family="Braconidae" genus="Alphomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alphomelon xestopyga" order="Hymenoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="xestopyga">
<emphasis id="3FF1636DA29AADCC53504EC24341BF7E" italics="true" pageId="0" pageNumber="5">Alphomelon xestopyga</emphasis>
</taxonomicName>
Deans female CNC721122
<emphasis id="1D3BE6F99926B7DE0DB816F80DEC06D1" bold="true" pageId="0" pageNumber="5">A</emphasis>
habitus, lateral
<emphasis id="1A7AC4675F86BE3B00966BC2E0D86380" bold="true" pageId="0" pageNumber="5">B</emphasis>
head, frontal
<emphasis id="D589190E537559E695542D1FD02F823B" bold="true" pageId="0" pageNumber="5">C</emphasis>
antenna, lateral
<emphasis id="982CB6AB77697189C9DF642A53ED4DD9" bold="true" pageId="0" pageNumber="5">D</emphasis>
wings
<emphasis id="C5EA143E4D4C3FFA2E9216BF2AFEDD3C" bold="true" pageId="0" pageNumber="5">E</emphasis>
propodeum and metasoma, dorsal
<emphasis id="665A1A518422B07AE911403D22C85E7B" bold="true" pageId="0" pageNumber="5">F</emphasis>
mesosoma, dorsal.
</paragraph>
</caption>
<caption id="CA20887A590C69908D8A1BF34B383DB7" doi="10.3897/zookeys.1175.105068.figure88" httpUri="https://binary.pensoft.net/fig/893190" pageId="0" pageNumber="5" start="Figure 88" startId="F88">
<paragraph id="B7F8BC2C49DACB67F219FDB173922D8B" pageId="0" pageNumber="5">
<emphasis id="882785EA579A389FDB35C46830ABD58C" bold="true" pageId="0" pageNumber="5">Figure 88.</emphasis>
<taxonomicName id="FFB43828DFE5344AC7F741EB41506595" authorityName="Deans" authorityYear="2003" class="Insecta" family="Braconidae" genus="Alphomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alphomelon xestopyga" order="Hymenoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="xestopyga">
<emphasis id="32AFE2B30072374BE6381B77E42C3CF1" italics="true" pageId="0" pageNumber="5">Alphomelon xestopyga</emphasis>
</taxonomicName>
Deans female CNC1620777
<emphasis id="7DA329EBFB1BBB12A7F18977655F68D8" bold="true" pageId="0" pageNumber="5">A</emphasis>
habitus, lateral
<emphasis id="81DF30BA0F41F4926614C23135F57D43" bold="true" pageId="0" pageNumber="5">B</emphasis>
head, lateral
<emphasis id="78A6533DCEA3EE8D20AB5596526FC210" bold="true" pageId="0" pageNumber="5">C</emphasis>
head, frontal
<emphasis id="35066F8A1B7D5BDE7C340618179F3C6A" bold="true" pageId="0" pageNumber="5">D</emphasis>
wings
<emphasis id="679670355B0B48191EACD8C28152B0BE" bold="true" pageId="0" pageNumber="5">E</emphasis>
metasoma, dorsal
<emphasis id="B8B9F6E819C8044F1AAD9965786EEE02" bold="true" pageId="0" pageNumber="5">F</emphasis>
head and mesosoma, dorsal.
</paragraph>
</caption>
<caption id="BE90C7E367767DE4E3A0CB3F01D77D67" doi="10.3897/zookeys.1175.105068.figure89" httpUri="https://binary.pensoft.net/fig/893191" pageId="0" pageNumber="5" start="Figure 89" startId="F89">
<paragraph id="3795CA70E1D3005341C9364FF0FF972C" pageId="0" pageNumber="5">
<emphasis id="618CA8CBF4F48783E677A9A4F89F9DDB" bold="true" pageId="0" pageNumber="5">Figure 89.</emphasis>
<taxonomicName id="8275C47BCA1472B237599BAA0A0F3265" authorityName="Deans" authorityYear="2003" class="Insecta" family="Braconidae" genus="Alphomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alphomelon xestopyga" order="Hymenoptera" pageId="0" pageNumber="5" phylum="Arthropoda" rank="species" species="xestopyga">
<emphasis id="95438A682C3D58D03565AA632E659E14" italics="true" pageId="0" pageNumber="5">Alphomelon xestopyga</emphasis>
</taxonomicName>
Deans holotype female USNMENT00828302
<emphasis id="6DF3F3FFE609CAD2F68EFE88859AB7E6" bold="true" pageId="0" pageNumber="5">A</emphasis>
habitus, lateral
<emphasis id="CAB83F27F3F10C09AEF1B7E6307FB885" bold="true" pageId="0" pageNumber="5">B</emphasis>
habitus, dorsal
<emphasis id="BC6F9E6ECABA6360316082DFECBE9F1D" bold="true" pageId="0" pageNumber="5">C</emphasis>
close-up of habitus, lateral.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

317
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@ -0,0 +1,317 @@
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<mods:title>A widespread new genus of Baetidae (Baetidae, Ephemeroptera) from Southeast Asia</mods:title>
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<mods:namePart>Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:namePart>Kluge, Nikita J.</mods:namePart>
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<mods:affiliation>Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
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<mods:role>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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4.
<taxonomicName LSID="https://zoobank.org/4C505602-E896-4CA6-99BE-EFF79C82DBDF" authority="Kaltenbach &amp; Kluge &amp; Gattolliat, 2022" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Branchiobaetis hamatus" order="Ephemeroptera" pageId="0" pageNumber="1" rank="species" species="hamatus" status="sp. nov.">Branchiobaetis hamatus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">Figs 16</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">, 17</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Habitus, larvae, dorsal view a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. (Sumatra, volcano Sago) d Branchiobaetis joachimi sp. nov. (Sumatra, Bukit Barisan, above Padang)." figureDoi="10.3897/zookeys.1135.93800.figure24" httpUri="https://binary.pensoft.net/fig/780354" pageId="0" pageNumber="1">, 24b</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Larva, tergalii IV a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. d Branchiobaetis minangkabau sp. nov. e Branchiobaetis jhoanae sp. nov." figureDoi="10.3897/zookeys.1135.93800.figure26" httpUri="https://binary.pensoft.net/fig/780356" pageId="0" pageNumber="1">, 26b</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution maps a, b new species of Branchiobaetis gen. nov. in Sumatra." figureDoi="10.3897/zookeys.1135.93800.figure28" httpUri="https://binary.pensoft.net/fig/780358" pageId="0" pageNumber="1">, 28a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Indonesia • Sumatra, volcano Talamau; River Pularian;
<geoCoordinate degrees="00" direction="north" minutes="00" orientation="latitude" precision="15" seconds="60" value="0.016666668">00°00'60&quot;N</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="00" orientation="longitude" precision="15" seconds="01" value="100.000275">100°00'01&quot;E</geoCoordinate>
; 960 m; 01.IV.2014; leg. M. Gueuning: larva on slide; GBIFCH00422261; MZL.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Same data as holotype; 2 larvae on slides; GBIFCH00422231, GBIFCH00422242; 20 larvae in alcohol; GBIFCH00422233, GBIFCH00422252, GBIFCH00422267, GBIFCH00422276, GBIFCH00422355, GBIFCH00422359, GBIFCH00422445, GBIFCH00422748, GBIFCH00422753, GBIFCH00422798, GBIFCH00422843, GBIFCH00423022, GBIFCH00975634, GBIFCH00975635; MZL. Indonesia • Sumatra, volcano Singgalang, River Sianok;
<geoCoordinate degrees="00" direction="south" minutes="19" orientation="latitude" precision="15" seconds="57" value="-0.33249998">00°19'57&quot;S</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="19" orientation="longitude" precision="15" seconds="19" value="100.321945">100°19'19&quot;E</geoCoordinate>
; 1150 m; 24.III.2014; leg. M. Gueuning; 2 larvae on slides; GBIFCH00422184, GBIFCH00423074; 13 larvae in alcohol; GBIFCH00422123, GBIFCH00422167, GBIFCH00422206, GBIFCH004208, GBIFCH00422215, GBIFCH00422216, GBIFCH00422224, GBIFCH00422797, GBIFCH00422889, GBIFCH00422938, GBIFCH01115975, GBIFCH01116020; MZL.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters distinguish
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="B. hamatus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis italics="true" pageId="0" pageNumber="1">B. hamatus</emphasis>
</taxonomicName>
sp. nov. from other species of
<taxonomicName authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" family="Baetidae" genus="Branchiobaetis" kingdom="Animalia" lsidName="Branchiobaetis" order="Ephemeroptera" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Branchiobaetis</emphasis>
</taxonomicName>
gen. nov.: A) labial palp segment II with medium, rounded protuberance, segment III apically slightly pointed (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16h</figureCitation>
); B) incisor of right mandible with ventral denticle (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16b</figureCitation>
); C) dorsal margin of femur with row of medium, spine-like setae, basally longer and clavate; additional row of short, hook-like setae along margin (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17a, b</figureCitation>
); D) dorsal margin of tibia and tarsus with row of short, hook-like setae (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17a, b</figureCitation>
); E) posterior margin of tergites: I smooth, without spines; II-IX triangular, narrower and longer towards last segment (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17e</figureCitation>
); posterior margin of sternites: I-VII smooth, without spines; VIII-IX with small, spaced, triangular spines; F) tergalius IV apically slightly concave (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17f</figureCitation>
); G) paraproct with short, stout, apically rounded setae along posterior margin (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17g</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" family="Baetidae" genus="Branchiobaetis" kingdom="Animalia" lsidName="Branchiobaetis hamatus" order="Ephemeroptera" pageId="0" pageNumber="1" rank="species" species="hamatus">
<emphasis italics="true" pageId="0" pageNumber="1">Branchiobaetis hamatus</emphasis>
</taxonomicName>
sp. nov., larva
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
labrum (left: ventral view, right: dorsal view)
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
right mandible
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
hypopharynx and superlinguae
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
maxilla
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
labium (left: ventral view, right: dorsal view).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(Figs
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16</figureCitation>
,
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17</figureCitation>
,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Habitus, larvae, dorsal view a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. (Sumatra, volcano Sago) d Branchiobaetis joachimi sp. nov. (Sumatra, Bukit Barisan, above Padang)." figureDoi="10.3897/zookeys.1135.93800.figure24" httpUri="https://binary.pensoft.net/fig/780354" pageId="0" pageNumber="1">24b</figureCitation>
,
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Larva, tergalii IV a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. d Branchiobaetis minangkabau sp. nov. e Branchiobaetis jhoanae sp. nov." figureDoi="10.3897/zookeys.1135.93800.figure26" httpUri="https://binary.pensoft.net/fig/780356" pageId="0" pageNumber="1">26b</figureCitation>
). Body length 6.8-8.5 mm. Cerci: broken. Paracercus: ca. 0.4
<normalizedToken originalValue="×">x</normalizedToken>
body length. Antenna: ca. 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as head length.
</paragraph>
<caption doi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" family="Baetidae" genus="Branchiobaetis" kingdom="Animalia" lsidName="Branchiobaetis hamatus" order="Ephemeroptera" pageId="0" pageNumber="1" rank="species" species="hamatus">
<emphasis italics="true" pageId="0" pageNumber="1">Branchiobaetis hamatus</emphasis>
</taxonomicName>
sp. nov., larva
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
hook-like seta of leg dorsal margins
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
fore femur apex, posterior view
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
fore claw
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
tergalius IV
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
paraproct
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
base of antenna.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Habitus, larvae, dorsal view a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. (Sumatra, volcano Sago) d Branchiobaetis joachimi sp. nov. (Sumatra, Bukit Barisan, above Padang)." figureDoi="10.3897/zookeys.1135.93800.figure24" httpUri="https://binary.pensoft.net/fig/780354" pageId="0" pageNumber="1">24b</figureCitation>
). Head, thorax and abdomen dorsally brown, abdominal segment X light brown; head, thorax and abdomen ventrally light brown, scape with dark brown spot at inner lateral side. Femur light brown, apically and dorsally along margin dark brown, with large, medial, dark brown spot; tibia light brown, tarsus dark brown in distal half. Caudalii light brown, cerci darker brown in area of ca.
<normalizedToken originalValue="½">1/2</normalizedToken>
of length, paracercus darker brown parallel to cerci; primary swimming setae dark brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Precursors of turbinate eyes</emphasis>
(Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Habitus, larvae, dorsal view a Branchiobaetis minangkabau sp. nov. b Branchiobaetis jhoanae sp. nov. Precursors of turbinate eyes developing in male last instar larvae c Branchiobaetis hamatus sp. nov. d Branchiobaetis joachimi sp. nov. e Branchiobaetis minangkabau sp. nov." figureDoi="10.3897/zookeys.1135.93800.figure25" httpUri="https://binary.pensoft.net/fig/780355" pageId="0" pageNumber="1">25c</figureCitation>
) in male last instar larvae representing a pair of subtriangular maculae; in the middle of this macula, a smaller, round, elevated area with well-expressed facets, approx. ten facets in diameter; peripheral area of the macula with indistinct facets.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17h</figureCitation>
). Scape distally and outside distolaterally with short, stout, apically rounded setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16a</figureCitation>
). Length 0.6
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Submarginal arc of setae composed of 7-10 long, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16b, c</figureCitation>
). Incisor blade-like with three denticles and a ventral denticle; kinetodontium with four denticles. Margin between prostheca and mola straight, with minute denticles.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16d, e</figureCitation>
). Incisor blade-like with four denticles; kinetodontium with three denticles. Margin between prostheca and mola straight, with minute denticles towards subtriangular process.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins slightly convex.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16f</figureCitation>
). Lingua as long as superlinguae. Lingua longer than broad; medial tuft of stout setae well developed. Superlinguae distally rounded; lateral margins rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16g</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae under canines. Medially with one pectinate, spine-like seta and five or six medium, simple setae. Maxillary palp approx. as long as galea-lacinia; palp segment II approx. as long as segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Branchiobaetis hamatus sp. nov., larva a labrum (left: ventral view, right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1135.93800.figure16" httpUri="https://binary.pensoft.net/fig/780346" pageId="0" pageNumber="1">16h</figureCitation>
). Inner margin of glossa with 10-12 spine-like setae, increasing in length distally; apex with two long and one medium, robust setae; outer margin with six or seven spine-like setae; Paraglossa with two short, simple setae in anteromedial area and one in posterolateral area; dorsally with three long, spine-like setae near inner margin. Labial palp with segment I approx. as long as length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with medium, rounded, distomedial protuberance; distomedial protuberance 0.3
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with 4-8 spine-like setae near outer margin. Segment III apically slightly pointed; length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
maximum width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17a-d</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.5:0.2.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Femur</emphasis>
. Length ca. 3
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with row of 7-9 medium, curved, spine-like setae and basally 10-15 longer, clavate setae. Additional row of short, stout, hook-like setae along dorsal margin. Apex rounded, with pair of spine-like setae; short, stout, hook-like setae on anterior and posterior side. Short, stout, apically rounded setae scattered along ventral margin.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
. Dorsal margin with two irregular rows of short, stout, hook-like setae. On surface short, stout, hook-like setae along patella-tibial suture. Ventral margin with row of short, curved, spine-like setae, on apex a tuft of fine, simple setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
. Dorsal margin with row of short, stout, hook-like setae and row of fine, simple setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Claw</emphasis>
with one row of ten or eleven denticles, distal denticle much longer than other denticles.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Terga</emphasis>
(Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17e</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergites: I smooth, without spines; II-IX triangular, narrower and longer towards last segment. Posterior margin of sternites: I-VII smooth, without spines; VIII-IX with small, spaced, triangular spines.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Tergalii</emphasis>
(Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17f</figureCitation>
,
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Larva, tergalii IV a Branchiobaetis aduncus sp. nov. b Branchiobaetis hamatus sp. nov. c Branchiobaetis joachimi sp. nov. d Branchiobaetis minangkabau sp. nov. e Branchiobaetis jhoanae sp. nov." figureDoi="10.3897/zookeys.1135.93800.figure26" httpUri="https://binary.pensoft.net/fig/780356" pageId="0" pageNumber="1">26b</figureCitation>
). Tracheae extending from main trunk to inner and outer margins; with light brown band along main trunk of tracheae on anal side. Tergalius I 2/3 as long as segment II, tergalius IV as long as length of segments V and 1/2 VI combined, tergalius VII as long as length of segment VIII.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Branchiobaetis hamatus sp. nov., larva a foreleg b hook-like seta of leg dorsal margins c fore femur apex, posterior view d fore claw e tergum IV f tergalius IV g paraproct h base of antenna." figureDoi="10.3897/zookeys.1135.93800.figure17" httpUri="https://binary.pensoft.net/fig/780347" pageId="0" pageNumber="1">17g</figureCitation>
). Posterior margin with 11-16 stout spines. Short, stout, apically rounded setae near posterior margin. Surface scattered with scale bases, micropores and fine, simple setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on the Latin word
<emphasis italics="true" pageId="0" pageNumber="1">hamatus</emphasis>
, meaning hooked, with reference to the hook-like setae on the legs.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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Indonesia: Sumatra (Fig.
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105
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Supra-antennal elevations 0.5
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as long as scapus and outer side strongly steep (angle about 70°), upper half thickened and backward, parallel-sided in anterior view, area between elevations with a small protuberance (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B, C</figureCitation>
); tyloids of male antenna linear (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10A</figureCitation>
); occipital carina widened, lamelliform and smooth (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); supra-antennal elevations largely yellow; head with brownish-yellow oblique patches postero-laterally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); vertex largely coarsely punctate with transversely rugose and becoming sparsely punctate posteriorly (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); mesoscutum coarsely punctate-rugose in dorsal view, with pair of large triangular yellow patches on middle lobe and pair of large oblong yellow patches on scutellum axilla (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); notauli narrow anteriorly, widened posteriorly and coarsely crenulate (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); scutellum coarsely reticulate-punctate and shallow-slightly concave medially, with pair of small brown-yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); metanotum slightly convex, rugose, with four elliptical yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); posterior propodeal carina distinctly arched, narrow lamelliform, foramen comparatively wide (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); first tergite with yellow stripe posteriorly; second tergite widely yellow posteriorly, third tergite with weak and slim yellow patches medio-posteriorly; fourth-sixth tergites with large yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10F</figureCitation>
); first and second sternites with wavy yellow stripe posteriorly; third and fourth sternites with small yellow patches postero-laterally (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10G</figureCitation>
); second sternite slightly concave medio-anteriorly (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9A</figureCitation>
); third sternite without depression (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10G</figureCitation>
).
</paragraph>
<caption doi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Figure 9.</emphasis>
<taxonomicName authorityName="Zhang &amp; Chen" authorityYear="2022" class="Insecta" family="Trigonalyidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taeniogonalos paradoxica" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="species" species="paradoxica">
<emphasis italics="true" pageId="0" pageNumber="101">Taeniogonalos paradoxica</emphasis>
</taxonomicName>
Zhang &amp; Chen, sp. nov., holotype, male (En-419024)
<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
habitus, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
head, anterior aspect
<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
head, dorsal aspect.
</paragraph>
</caption>
<caption doi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Figure 10.</emphasis>
<taxonomicName authorityName="Zhang &amp; Chen" authorityYear="2022" class="Insecta" family="Trigonalyidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taeniogonalos paradoxica" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="species" species="paradoxica">
<emphasis italics="true" pageId="0" pageNumber="101">Taeniogonalos paradoxica</emphasis>
</taxonomicName>
Zhang &amp; Chen, sp. nov., holotype, male (En-419024)
<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
tyloids on 10th-16th segments of antenna
<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
wings
<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
head, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
mesosoma, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="101">E</emphasis>
mesosoma, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="101">F</emphasis>
metasoma, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="101">G</emphasis>
metasoma, ventral aspect
<emphasis bold="true" pageId="0" pageNumber="101">H</emphasis>
legs, ventral aspect.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="description">
<paragraph pageId="0" pageNumber="101">Description.</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Female.</emphasis>
Holotype, length of body 8.6 mm (of fore wing 7.1 mm).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">
<emphasis italics="true" pageId="0" pageNumber="101">Head</emphasis>
.
</emphasis>
Antenna with 23 segments; tyloids linear, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as antennal segment on 11th-13th segments, about half as long on 14th segment, 0.3
<normalizedToken originalValue="×">x</normalizedToken>
as long on 15th segment and 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long on 10th segment (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10A</figureCitation>
); frons densely and coarsely punctate with rugae near clypeus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B</figureCitation>
); vertex largely coarsely punctate with transversely rugae and becoming sparsely punctate posteriorly (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); temple largely smooth with few punctures near orbita (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10C</figureCitation>
); head gradually narrowed behind eyes, eye in dorsal view 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as temple (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); occipital carina widened, lamelliform and smooth (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); supra-antennal elevations smooth, outer side very steep (angle about 70°), upper half thickened and backward, parallel-sided in anterior view, 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as scapus, with a small protuberance between them (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B, C</figureCitation>
); clypeus slightly concave and thick medio-ventrally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">
<emphasis italics="true" pageId="0" pageNumber="101">Mesosoma</emphasis>
.
</emphasis>
Length of mesosoma 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as its height (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10E</figureCitation>
); transverse mesopleural groove narrow, crenulate; mesopleuron above transverse mesopleural groove reticulate-punctate anteriorly and coarsely punctate posteriorly, below groove rugulose-punctate anteriorly and smooth posteriorly (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10E</figureCitation>
); notauli narrow anteriorly, widened posteriorly and coarsely crenulate; middle lobe of mesoscutum coarsely transverely punctate-rugose anteriorly and reticulate-punctate posteriorly, lateral lobes largely punctate with longitudinal rugae (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); scutellar sulcus complete, moderately narrow and crenulate; scutellum coarsely reticulate-punctate and slightly concave medially, anteriorly protruding above level of mesoscutum; metanotum slightly convex, rugose (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); propodeum punctate-rugose (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); posterior propodeal carina distinctly arched, narrow lamelliform, foramen approximately 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high medially.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">
<emphasis italics="true" pageId="0" pageNumber="101">Wings</emphasis>
.
</emphasis>
Fore wing: vein 1-M 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as vein 1-SR; second submarginal cell 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as third submarginal cell (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">
<emphasis italics="true" pageId="0" pageNumber="101">Metasoma</emphasis>
.
</emphasis>
First tergite 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as its apical width, smooth and with shallow but wide depression medially (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10F</figureCitation>
); following tergites densely and coarsely punctate (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10F</figureCitation>
); first sternite sparse punctate and concave laterally; following sternites densely punctate (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10G</figureCitation>
); second sternite slightly concave medio-anteriorly, rather flat media-posteriorly and slightly depressed near middle of posterior margin (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9A</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Colour.</emphasis>
Black; apical half of antenna and part of scapus dark brown; remainder of antenna mainly yellow; orbita yellow, narrow but widened near antennal sockets; supra-antennal elevations largely yellow; clypeus with pair of yellow patches; lateral corner of clypeus dark brown; mandible teeth dark brown to dark (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B</figureCitation>
); head with brownish-yellow oblique patches postero-laterally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); pronotum yellow dorsally (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); middle lobe of mesoscutum with pair of large triangular yellow patches antero-laterally; scutellum axilla with pair of large oblong yellow patches; scutellum with pair of small brown-yellow patches media-laterally; metanotum with four elliptical yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); palpi dark brown (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10C</figureCitation>
); tegulae brown (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); first tergite with yellow stripe posteriorly; second tergite widely yellow posteriorly, third tergite with weak and slim yellow patches media-posteriorly; fourth-sixth tergites with large yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10F</figureCitation>
); first and second sternites with wavy yellow stripe posteriorly; third and fourth sternites with small yellow patches postero-laterally (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10G</figureCitation>
); apex of fore and middle femora yellow; tibiae and basal half of tarsi of fore leg largely yellow; remainder of tibiae and tarsi dark brown (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10H</figureCitation>
); pterostigma yellow brown; anterior half of fore wing dark brown; remainder of wing subhyaline (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="etymology">
<paragraph pageId="0" pageNumber="101">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="101">
Name after the unusual supra-antennal elevations: from &quot;
<emphasis italics="true" pageId="0" pageNumber="101">paradoxus</emphasis>
&quot; (Latin for &quot;strange, contrary to expectation&quot;).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="materials_examined">
<paragraph pageId="0" pageNumber="101">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="101">
<materialsCitation accessionNumber="OM057987" collectingDate="2020-09-20" collectorName="Huayan Chen, GenBank" country="China" county="Diqing Tibetan Autonomous Prefecture" elevation="2639" latitude="27.526667" location="Jiantang town" longLatPrecision="20" longitude="100.092224" municipality="City" specimenCount="1" stateProvince="Yunnan" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">
<typeStatus>Holotype</typeStatus>
</emphasis>
, female,
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingCounty>Diqing Tibetan Autonomous Prefecture</collectingCounty>
, Shangri-La
<collectingMunicipality>City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C1A8CC692EF151A28F692CD43416C178:A451513C980BCCDBE353C6330A0F35E0" country="China" county="Diqing Tibetan Autonomous Prefecture" latitude="27.526667" longLatPrecision="20" longitude="100.092224" municipality="City" name="Jiantang town" stateProvince="Yunnan">Jiantang town</location>
, apple orchard,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.639" unit="m" value="2639.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.639" unit="m" value="2639.0">2639 m</elevation>
</quantity>
,
<geoCoordinate degrees="27" direction="north" minutes="31" orientation="latitude" precision="15" seconds="36" value="27.526667">27°31'36&quot;N</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="5" orientation="longitude" precision="15" seconds="32" value="100.092224">100°5'32&quot;E</geoCoordinate>
,
<collectingDate value="2020-09-20">20.IX.2020</collectingDate>
,
<collectorName>Huayan Chen</collectorName>
leg., YPT, SYSBM En-419024 (deposited in SYSBM);
<collectorName>GenBank</collectorName>
:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OM057987">OM057987</accessionNumber>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="distribution">
<paragraph pageId="0" pageNumber="101">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="101">China (Yunnan). Collected at 2639 m.</paragraph>
<paragraph pageId="0" pageNumber="101">
The key to species of
<taxonomicName authorityName="Schulz" authorityYear="1906" class="Insecta" family="Trigonalyidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taeniogonalos" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="101">Taeniogonalos</emphasis>
</taxonomicName>
published by
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.385.6560" author="Chen, H-Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 207" refId="B6" refString="Chen, H-Y, van Achterberg, C, He, J-H, Xu, Z-F, 2014. A revision of the Chinese Trigonalyidae (Hymenoptera, Trigonalyoidea). ZooKeys 385: 1 - 207, DOI: https://doi.org/10.3897/zookeys.385.6560" title="A revision of the Chinese Trigonalyidae (Hymenoptera, Trigonalyoidea)." url="https://doi.org/10.3897/zookeys.385.6560" volume="385" year="2014">Chen et al. (2014)</bibRefCitation>
could be updated to accommodate
<taxonomicName lsidName="T. paradoxica" pageId="0" pageNumber="101" rank="species" species="paradoxica">
<emphasis italics="true" pageId="0" pageNumber="101">T. paradoxica</emphasis>
</taxonomicName>
by replacing couplet 12 as follows:
</paragraph>
<paragraph pageId="0" pageNumber="101">
<table inLine="true" pageId="0" pageNumber="101">
<tr pageId="0" pageNumber="101">
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">Protuberance of third sternite of ♀ subtruncate medio-apically; pronotum, metanotum medially and head of ♀ posteriorly with pale pattern; hind tibia black or dark brown; outer sides of of supra-antennal elevations oblique</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="101">12</emphasis>
'
</td>
</tr>
<tr pageId="0" pageNumber="101">
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">Protuberance of third sternite of ♀ acutely protruding medio-apically; metanotum, apex of supra-antennal elevations and head posteriorly in dorsal view black; hind tibia brown; outer side of supra-antennal elevations subvertical</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="101">13</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="101">
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">12'</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">Pronotum black dorsally; metanotum ivory medially; apex of supra-antennal elevations pale yellowish and remainder black; head of ♀ posteriorly with pair of ivory patches in dorsal view (black in ♂); hind tibia black; outer side of supra-antennal elevations oblique; metanotum with pair of yellowish patches</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="101">
<taxonomicName lsidName="T. subtruncata" pageId="0" pageNumber="101" rank="species" species="subtruncata">
<emphasis italics="true" pageId="0" pageNumber="101">T. subtruncata</emphasis>
</taxonomicName>
Chen, van Achterberg, He &amp; Xu, 2014
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="101">
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
Pronotum yellow dorsally (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
); supra-antennal elevations largely yellow (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9B, C</figureCitation>
); head of ♂ posteriorly with pair of brownish-yellow oblique patches in dorsal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); hind tibia dark brown (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10H</figureCitation>
); outer side of supra-antennal elevations strongly steep (angle about 70°; Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure9" httpUri="https://binary.pensoft.net/fig/677746" pageId="0" pageNumber="101">9C</figureCitation>
); metanotum with four elliptical yellow patches (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Taeniogonalos paradoxica Zhang &amp; Chen, sp. nov., holotype, male (En- 419024) A tyloids on 10 th- 16 th segments of antenna B wings C head, lateral aspect D mesosoma, dorsal aspect E mesosoma, lateral aspect F metasoma, dorsal aspect G metasoma, ventral aspect H legs, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure10" httpUri="https://binary.pensoft.net/fig/677747" pageId="0" pageNumber="101">10D</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="101">
<taxonomicName lsidName="T. paradoxica" pageId="0" pageNumber="101" rank="species" species="paradoxica">
<emphasis italics="true" pageId="0" pageNumber="101">T. paradoxica</emphasis>
</taxonomicName>
Zhang &amp; Chen, sp. nov.
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>