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<document ID-CLB-Dataset="297075" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="7e8b8d63-711e-4456-bdce-b4c77868fda7" ID-ISBN="0-8018-8221-4" checkinTime="1670905182117" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="C17C0D7B1BA7B345C06C127E27AD346E" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Molossidae.432-451.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Mops (Xiphonycteris) nanulus J. A. Allen 1917" docType="treatment" docVersion="4" id="3B127064EB7DDB182ED4ED54593B25D0" lastPageNumber="443" masterDocId="3B127064EB7DDB182ED4ED54593B25D0" masterDocTitle="Order Chiroptera - Family Molossidae" masterLastPageNumber="451" masterPageNumber="432" pageNumber="443" updateTime="1716777726396" updateUser="ExternalLinkService">
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<mods:titleInfo id="CD5A9C0DBD9AAB2B50A5D66EDC7DFE3D">
<mods:title id="BFDB389FE7F931761557E86AD45B9BCD">Order Chiroptera - Family Molossidae</mods:title>
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<mods:namePart id="41337A3C3FE7C783C55F96517F701641">Reeder, DeeAnn</mods:namePart>
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<mods:title id="42710DFE0D59E519022FD7FB546393A6">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1</mods:title>
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<subSubSection id="F25881266B7C833A4D2A2E7136F3664F" pageNumber="443" type="nomenclature">
<paragraph id="286810D1D902C66A840D7528BEF41320" pageNumber="443">
<taxonomicName id="DC4C9CF5FD6B5650A51FC0D9A20B5939" ID-CoL="7VW4N" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="443" phylum="Chordata" rank="species" species="nanulus" subGenus="Xiphonycteris">
<emphasis id="A7696F419ED4D2D1521E7AF5F781E502" italics="true" pageNumber="443">Mops (Xiphonycteris) nanulus</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C6B4B98C3E5F13FB372605A3218B501F" pageNumber="443" type="reference_group">
<paragraph id="A341D92324B5D64F1B0CB6CF55B62E3E" pageNumber="443">
<treatmentCitation id="889474190B9FA62D65CEB548655DA7BD" author="J. A. Allen" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" journal="Bull. Am. Mus. Nat. Hist." kingdom="Animalia" order="Chiroptera" page="477" pageNumber="443" phylum="Chordata" rank="species" species="nanulus" subGenus="Xiphonycteris" title="Mops (Xiphonycteris) nanulus" volume="37" year="1917">
<taxonomicName id="087A64F4A99815AFB965F5ED9F09AC14" ID-CoL="7VW4N" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="443" phylum="Chordata" rank="species" species="nanulus" subGenus="Xiphonycteris">
<emphasis id="60F300A5E7869BB9A9B0D1D73ED5BD03" italics="true" pageNumber="443">Mops (Xiphonycteris) nanulus</emphasis>
J. A. Allen 1917
</taxonomicName>
,
<bibCitation id="1DC4C82C05E27544648B12E1B1B020BF" author="J. A. Allen" journal="Bull. Am. Mus. Nat. Hist." pageNumber="443" pagination="477" title="Mops (Xiphonycteris) nanulus" volume="37" year="1917">Bull. Am. Mus. Nat. Hist., 37: 477</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC5C9F257629D2D7AE076C7A32B9C72E" pageNumber="443" type="materials_examined">
<paragraph id="91C9DE978AB4FCE9B575D0A80EC0A21E" pageNumber="443">
<emphasis id="F6DF05F40F01FC5973CA0304DE20F448" inLineHeading="true" pageNumber="443">Type Locality:</emphasis>
<materialsCitation id="88DB5BB775D2E2CEE061855E6C84D6F1" ID-GBIF-Occurrence="4867302431" country="Democratic Republic of the Congo" pageNumber="443" stateProvince="Orientale">
Dem. Rep.
<collectingCountry id="E6C5D58C5E169C0E0718C99774F031C0" name="Democratic Republic of the Congo">Congo</collectingCountry>
,
<collectingRegion id="A2F26C01AD2486EF7061BE26DD9E087E" country="Democratic Republic of the Congo" name="Orientale">Oriental</collectingRegion>
, Niangara.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3FF06387834B3A8B654A296A29B789D0" pageNumber="443" type="vernacular_names">
<paragraph id="181314AAB4CB1BADD296B6F49D4DB0D8" pageNumber="443">
<emphasis id="40956F107A486A16CC52BD6D0D23BCB7" inLineHeading="true" pageNumber="443">Vernacular Names:</emphasis>
<vernacularName id="875BE8492F5C2C70B328F6DDC4F4BC32" language="eng" pageNumber="443">Dwarf Free-tailed Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCD12D1BA468C20E53CF9978338BBDAA" pageNumber="443" type="synonym_list">
<paragraph id="1DD8DF9FF267FAA13DC3946B6053B971" pageNumber="443">
<emphasis id="A9544D144654B7FD7DF6FBAF1242B6B9" inLineHeading="true" pageNumber="443">Synonyms:</emphasis>
<taxonomicName id="38EF0971125F63C27180432DEF2CEDBB" authority="Hayman 1940" authorityName="Hayman" authorityYear="1940" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="443" phylum="Chordata" rank="species" species="calabarensis" subGenus="Xiphonycteris">
<emphasis id="6E6F44D3DCDA551437F6163E4298B74A" italics="true" pageNumber="443">Mops (Xiphonycteris) calabarensis</emphasis>
Hayman 1940
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EA249060B4C35A111766CD05430BBBB6" pageNumber="443" type="distribution">
<paragraph id="514F8F05DAC56966F4267DC82FCCCB68" pageNumber="443">
<emphasis id="8B07E132B5D4BF57F9E61CF36F6685C4" inLineHeading="true" pageNumber="443">Distribution:</emphasis>
<collectingCountry id="577F7AD55C3D008BD9DE179A4B121FC8" name="Sierra Leone">Sierra Leone</collectingCountry>
to
<collectingCountry id="A694AC73583853A2440C6F3BB18C2B87" name="Ethiopia">Ethiopia</collectingCountry>
and
<collectingCountry id="0974EF649389329E9E5A803A22CCB50D" name="Kenya">Kenya</collectingCountry>
. A previous report of this species from
<collectingCountry id="38B5883A08B3F87EEA3F1E90EE784CC1" name="Gambia">The Gambia</collectingCountry>
is in error, probably based on a specimen of
<taxonomicName id="B2D59825686E2113397D59DF717786A1" authorityName="Peters" authorityYear="1852" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="brachypterus">
<emphasis id="EC77B1A0C1914A3BF3BB6B1F6A2ABB60" italics="true" pageNumber="443">brachypterus</emphasis>
</taxonomicName>
(J. Fahr, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="107FCA99148801FABB0844E923F1BA46" pageNumber="443" type="conservation">
<paragraph id="6081F13A16060871A9030521304030B0" pageNumber="443">
<emphasis id="0625772AFCC552232D9B464DCA25A349" inLineHeading="true" pageNumber="443">Conservation:</emphasis>
<collectionCode id="99FBE1CA04A1B6BCA4AE29DB3961AEB3">IUCN</collectionCode>
2003 and
<collectionCode id="659994E410F42B84DCD95138D1508804">IUCN</collectionCode>
/
<collectionCode id="DF16DD60770357A74960E0B21903BB2B" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="6626D65EF1B84C005B2227CFFF423D57" pageNumber="443" type="discussion">
<paragraph id="747D35E2FCA410412A04BA3A8BCE726A" pageNumber="443">
<emphasis id="2A7FE9CD326A417D74555D993B8D8265" inLineHeading="true" pageNumber="443">Discussion:</emphasis>
Subgenus
<taxonomicName id="F6F8B9D30188F454EDD55F5DF0DEE63F" authorityName="Dollman" authorityYear="1911" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="subGenus" subGenus="Xiphonycteris">
<emphasis id="27B95BC3F71F90862D22ABC0620071F9" italics="true" pageNumber="443">Xiphonycteris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. &amp; Nuss, Matthias" docDate="2020" docId="C17C0F5F4055574B88F96843DAF341BB" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa niveofascia Leger &amp; Nuss 2020, sp. nov." docType="treatment" docUuid="4112C427-AF2B-4CAE-BDB3-E94033B8E8C2" docUuidSource="ZooBank" docVersion="4" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1668168367651" updateUser="ExternalLinkService">
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<mods:title>Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
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<treatment ID-GBIF-Taxon="161926043" LSID="urn:lsid:zoobank.org:act:4112C427-AF2B-4CAE-BDB3-E94033B8E8C2" httpUri="http://treatment.plazi.org/id/C17C0F5F4055574B88F96843DAF341BB" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="C17C0F5F-4055-574B-88F9-6843DAF341BB" authority="Léger &amp; Nuss" authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa niveofascia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveofascia" status="sp. nov.">
Hoploscopa niveofascia
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; Nuss
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 418" captionStartId="F2" captionText="Figures 4 - 18. Forewing of Hoploscopa species. 4 Hoploscopa albipuncta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu Trail starting point, near Liwagu River, 6 ° 1 ' 40 &quot; N, 116 ° 32 ' 59 &quot; E, 1760 m, 18. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 335 ♂) 5 Hoploscopa sepanggi sp. nov., holotype, ♂, Malaysia, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 &quot; N, 116 ° 35 ' 48.03 ' E, 1925 m, at lamps, 30. v. 2015 (T. Leger &amp; R. Mally) 6 Hoploscopa cynodonta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, junction Kiau View- and Pandanus Trail, 6 ° 0 ' 32.84 &quot; N, 116 ° 32 ' 14.94 &quot; E, 1690 m, UV light, 07. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 327 ♂) 7 Hoploscopa parvimacula sp. nov., paratype, ♀, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 &quot; N, 116 ° 35 ' 48.03 &quot; E, 1925 m, at lamps, 30. v. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 312 ♀. 8 Hoploscopa kinabaluensis sp. nov., paratype, ♀, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu trail starting point, near Liwagu River, 6 ° 1 ' 40 &quot; N, 116 ° 32 ' 59 &quot; E, 1760 m, UV light, 18. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 326 ♀) 9 Hoploscopa luteomacula Nuss, paratype, ♀, Indonesia, Sumatra, Barat, N-Padangpanjang, Mt Singgalang, 2100 m, 10 - 11. ii. 1996 (A. Kallies) (genitalia on slide GU prep. Nuss 743) 10 Hoploscopa obliqua Rotschild, abdomen missing, Papua New Guinea, Madang Province, Wanang village, 05 ° 15 ' S, 145 ° 17 ' E, reared from Diplazium esculentum, 12. vi. 2007 11 Hoploscopa niveofascia sp. nov., holotype, ♂, Papua New Guinea, Morobe Prov., nr. Bulolo, Mt. Susu Nat. Res., 975 m, 27 - 28. viii. 1983, Araucaria For. (S. Miller) (genitalia on slide TL 442 ♂) 12 Hoploscopa gombongi sp. nov., paratype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (B Gewa, J. Kua, S Sau, A. Kinibel) 13 Hoploscopa tonsepi sp. nov., holotype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (J. Valeba, J. Auga, M. Dilu, F. Philip, R. Lilip) (genitalia on slide TL 655 ♂) 14 Hoploscopa marijoweissae sp. nov., ♂, Papua New Guinea: Morobe Province, Mount Kaindi, 2350 m, 11. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 437 ♂) 15 Hoploscopa titika sp. nov., holotype, ♂, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, Strasse nach Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 505 ♂) 16 Hoploscopa pangrangoensis sp. nov., paratype, ♀, Indonesia, Java, Mt. Pangrange, SE Bogor, 6.30 S 107.10 E, 1625 m, primary forest, 16 - 20. ii. 1996 (Siniaev &amp; Afonin) (genitalia on slide TL 627 ♀) 17 Hoploscopa isarogensis sp. nov., paratype, ♀, Philippines, South Luzon, Mt Isarog, 13 ° 40 ' N, 123 ° 20 ' E, 530 m, submontane forest, at light, 22. iii. 2000 (M. Nuss) (genitalia on slide TL 523 ♀) 18 Hoploscopa ypsilon sp. nov., paratype, ♀, Philippines, Luzon, Mountain Province, Chatol, 2100 m, 16 - 18. xi. 1997 (Mey, Ebert &amp; Nuss)." figureDoi="10.3897/zookeys.907.36563.figures4-18" httpUri="https://binary.pensoft.net/fig/375757" pageId="0" pageNumber="1">Figs 11</figureCitation>
<figureCitation captionStart="Figures 5257" captionStartId="F6" captionText="Figures 52 - 57. Male genitalia features of Hoploscopa. 52 Hoploscopa luteomacula Nuss, holotype, GU prep. Nuss 744 ♂ 53 Hoploscopa obliqua, holotype, Pyralidae Brit. Mus. Slide N ° BMNH 20252 ♂ (phallus with juxta attached) 54 Hoploscopa gombongi sp. nov., holotype, TL 653 ♂ 55 Hoploscopa tonsepi sp. nov., paratype, TL 661 ♂ 56 Hoploscopa niveofascia sp. nov., holotype, TL 442 ♂ 57 Hoploscopa marijoweissae sp. nov., holotype, TL 710 ♂." figureDoi="10.3897/zookeys.907.36563.figures52-57" httpUri="https://binary.pensoft.net/fig/375761" pageId="0" pageNumber="1">, 56</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂, with labels: &quot;Papua New Guinea | Morobe Prov[ince]., n[ea]r. Bulolo | Mt. Susu Nat[ional]. Res[erve]., 975m | 27-28 Aug.1983, S. Miller | UV Lite, Araucaria For[est]&quot;; &quot;DNA voucher | Lepidoptera | MTD2016 | [vertically written:] no. 3162&quot;; &quot;TL442 | ♂&quot;. Deposited in USNM.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa niveofascia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveofascia">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa niveofascia</emphasis>
</taxonomicName>
sp. nov. displays snow white postmedian patch, subterminal line and well-marked marginal spots on the forewing. In male genitalia, the uncus is broad, with straight lateral margin and a blunt apex. Female genitalia not known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1919" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa diffusa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa diffusa</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa niveofascia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveofascia">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa niveofascia</emphasis>
</taxonomicName>
sp. nov. shares with
<taxonomicName lsidName="H. diffusa" pageId="0" pageNumber="1" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="1">H. diffusa</emphasis>
</taxonomicName>
the white markings of the forewing. However, postmedian patch of
<taxonomicName lsidName="H. diffusa" pageId="0" pageNumber="1" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="1">H. diffusa</emphasis>
</taxonomicName>
is broadly marked with brown, marginal spots are reduced to small dots, and fringes exhibit white spots, while fringes are brown in
<taxonomicName lsidName="H. niveofascia" pageId="0" pageNumber="1" rank="species" species="niveofascia">
<emphasis italics="true" pageId="0" pageNumber="1">H. niveofascia</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Antennae dorsally dark brown. Proboscis white. Maxillary palpi dark brown, basally pale brown. Labial palpi dark brown, ventro-basally white.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
(Fig.
<figureCitation captionStart="Figures 418" captionStartId="F2" captionText="Figures 4 - 18. Forewing of Hoploscopa species. 4 Hoploscopa albipuncta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu Trail starting point, near Liwagu River, 6 ° 1 ' 40 &quot; N, 116 ° 32 ' 59 &quot; E, 1760 m, 18. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 335 ♂) 5 Hoploscopa sepanggi sp. nov., holotype, ♂, Malaysia, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 &quot; N, 116 ° 35 ' 48.03 ' E, 1925 m, at lamps, 30. v. 2015 (T. Leger &amp; R. Mally) 6 Hoploscopa cynodonta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, junction Kiau View- and Pandanus Trail, 6 ° 0 ' 32.84 &quot; N, 116 ° 32 ' 14.94 &quot; E, 1690 m, UV light, 07. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 327 ♂) 7 Hoploscopa parvimacula sp. nov., paratype, ♀, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 &quot; N, 116 ° 35 ' 48.03 &quot; E, 1925 m, at lamps, 30. v. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 312 ♀. 8 Hoploscopa kinabaluensis sp. nov., paratype, ♀, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu trail starting point, near Liwagu River, 6 ° 1 ' 40 &quot; N, 116 ° 32 ' 59 &quot; E, 1760 m, UV light, 18. vi. 2015 (T. Leger &amp; R. Mally) (genitalia on slide TL 326 ♀) 9 Hoploscopa luteomacula Nuss, paratype, ♀, Indonesia, Sumatra, Barat, N-Padangpanjang, Mt Singgalang, 2100 m, 10 - 11. ii. 1996 (A. Kallies) (genitalia on slide GU prep. Nuss 743) 10 Hoploscopa obliqua Rotschild, abdomen missing, Papua New Guinea, Madang Province, Wanang village, 05 ° 15 ' S, 145 ° 17 ' E, reared from Diplazium esculentum, 12. vi. 2007 11 Hoploscopa niveofascia sp. nov., holotype, ♂, Papua New Guinea, Morobe Prov., nr. Bulolo, Mt. Susu Nat. Res., 975 m, 27 - 28. viii. 1983, Araucaria For. (S. Miller) (genitalia on slide TL 442 ♂) 12 Hoploscopa gombongi sp. nov., paratype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (B Gewa, J. Kua, S Sau, A. Kinibel) 13 Hoploscopa tonsepi sp. nov., holotype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (J. Valeba, J. Auga, M. Dilu, F. Philip, R. Lilip) (genitalia on slide TL 655 ♂) 14 Hoploscopa marijoweissae sp. nov., ♂, Papua New Guinea: Morobe Province, Mount Kaindi, 2350 m, 11. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 437 ♂) 15 Hoploscopa titika sp. nov., holotype, ♂, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, Strasse nach Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 505 ♂) 16 Hoploscopa pangrangoensis sp. nov., paratype, ♀, Indonesia, Java, Mt. Pangrange, SE Bogor, 6.30 S 107.10 E, 1625 m, primary forest, 16 - 20. ii. 1996 (Siniaev &amp; Afonin) (genitalia on slide TL 627 ♀) 17 Hoploscopa isarogensis sp. nov., paratype, ♀, Philippines, South Luzon, Mt Isarog, 13 ° 40 ' N, 123 ° 20 ' E, 530 m, submontane forest, at light, 22. iii. 2000 (M. Nuss) (genitalia on slide TL 523 ♀) 18 Hoploscopa ypsilon sp. nov., paratype, ♀, Philippines, Luzon, Mountain Province, Chatol, 2100 m, 16 - 18. xi. 1997 (Mey, Ebert &amp; Nuss)." figureDoi="10.3897/zookeys.907.36563.figures4-18" httpUri="https://binary.pensoft.net/fig/375757" pageId="0" pageNumber="1">11</figureCitation>
). Thorax dark brown with mesodorsal transversal white line. Collar white. Forewing length: 9.5 mm; forewing ground colour dark brown; basal area without marked patches, white scales scattered near dorsum; median discoidal stigma darker coloured, rectangular, basally and distally thinly edged with white; postmedian patch diffuse, white; postmedian line marked on costal half; subterminal line thick, zigzagging, disrupted distally at M1; margin with large white spots; fringes bronze. Fore- and midlegs dark brown, with tibia and tarsi segments distally white. Hindlegs brown to bronze.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen.</emphasis>
Male sternum A8 posterior margin straight.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1) (Fig.
<figureCitation captionStart="Figures 5257" captionStartId="F6" captionText="Figures 52 - 57. Male genitalia features of Hoploscopa. 52 Hoploscopa luteomacula Nuss, holotype, GU prep. Nuss 744 ♂ 53 Hoploscopa obliqua, holotype, Pyralidae Brit. Mus. Slide N ° BMNH 20252 ♂ (phallus with juxta attached) 54 Hoploscopa gombongi sp. nov., holotype, TL 653 ♂ 55 Hoploscopa tonsepi sp. nov., paratype, TL 661 ♂ 56 Hoploscopa niveofascia sp. nov., holotype, TL 442 ♂ 57 Hoploscopa marijoweissae sp. nov., holotype, TL 710 ♂." figureDoi="10.3897/zookeys.907.36563.figures52-57" httpUri="https://binary.pensoft.net/fig/375761" pageId="0" pageNumber="1">56</figureCitation>
). Uncus broad with straight lateral margin, apex truncate, medially slightly incurved. Gnathos reduced to thin band without posterior projection. Valva ventral margin bent dorsad on distal half, dorsal margin slightly convex, apex pointed. Juxta elongated, with base rounded, medially narrowing, apex blunt. Saccus small, pointing dorsad.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known from Mount Susu (975 m) in the Morobe Province (Papua New Guinea).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phylogenetic relationships">
<paragraph pageId="0" pageNumber="1">Phylogenetic relationships.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Rothschild" baseAuthorityYear="1915" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa obliqua" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa obliqua</emphasis>
</taxonomicName>
is recovered as sister group in the ML analysis of the COI barcode (BS = 95).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
From the Latin
<emphasis italics="true" pageId="0" pageNumber="1">niveus</emphasis>
, snowy, and
<emphasis italics="true" pageId="0" pageNumber="1">fascia</emphasis>
, band, referring to the snow-white markings on the forewing.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="426E4744322880EDDAC1945DADB0EA47">Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia</mods:title>
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<heading id="122281A09908716DFF66E67890A9F8ED" box="[130,498,1795,1817]" fontSize="9" level="2" pageId="1" pageNumber="92" reason="7">
Suborder
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<emphasis id="7BA1EADE9908716DFF00E6789034F8ED" bold="true" box="[228,367,1795,1817]" pageId="1" pageNumber="92">Micrasterina</emphasis>
Fisher, 1966
</taxonomicName>
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<paragraph id="496A36CC9908716DFF66E6549005F8B1" blockId="1.[130,350,1839,1861]" box="[130,350,1839,1861]" pageId="1" pageNumber="92">
Family
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sedis
</emphasis>
</paragraph>
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<emphasis id="7BA1EADE9908716DFF66E6219182F884" box="[130,217,1882,1904]" italics="true" pageId="1" pageNumber="92">Remarks</emphasis>
. The combination of generic features, particularly the presence of a marginal peripetalous fasciole, intermittent horizontal fasciole bands and a rudimentary non re-entrant peripetalous fasciole clear of the distal ends of paired petals, clearly distinguish the new genus
<taxonomicName id="8ED54D4F9908716DFB9FE03C9446FEA9" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[1147,1309,327,349]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFB9FE03C9446FEA9" box="[1147,1309,327,349]" italics="true" pageId="1" pageNumber="92">Murraypneustes</emphasis>
</taxonomicName>
from virtually all other taxa assigned to the
<taxonomicName id="8ED54D4F9908716DFB88E01F95B6FE8E" authorityName="Fisher" authorityYear="1966" box="[1132,1261,356,378]" class="Echinoidea" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="subOrder" subOrder="Micrasterina">Micrasterina</taxonomicName>
, in particular the 22 genera included in the
<taxonomicName id="8ED54D4F9908716DFBC7E0FA9402FE63" authority="Fisher, 1966" authorityName="Fisher" authorityYear="1966" box="[1059,1369,385,407]" class="Echinoidea" family="Asterostomatidae" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="family">Asterostomatidae Fisher, 1966</taxonomicName>
. Only one of these genera,
<taxonomicName id="8ED54D4F9908716DFBE7E0E59444FE40" authority="Agassiz, 1847" authorityName="Agassiz" authorityYear="1847" box="[1027,1311,414,436]" class="Echinoidea" family="Asterostomatidae" genus="Asterostoma" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFBE7E0E59524FE40" box="[1027,1151,414,436]" italics="true" pageId="1" pageNumber="92">Asterostoma</emphasis>
<bibRefCitation id="2D444B3D9908716DFB6DE0E59444FE40" author="Agassiz, L. in &amp; Agassiz, L. &amp; Desor, E." box="[1161,1311,414,436]" pageId="1" pageNumber="92" pagination="129 - 168" refId="ref4999" refString="Agassiz, L. in: Agassiz, L., and Desor, E. 1847. Catologue raisonne des families, des genres et des especes de la classe des echinoderms. Annales des Sciences Naturelles 7: 129 - 168." type="journal article" year="1847">Agassiz, 1847</bibRefCitation>
</taxonomicName>
, has tentatively been retained in the
<taxonomicName id="8ED54D4F9908716DFBA2E0C095AEFE25" authorityName="Fisher" authorityYear="1966" box="[1094,1269,443,465]" class="Echinoidea" family="Asterostomatidae" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="family">Asterostomatidae</taxonomicName>
(together with
<taxonomicName id="8ED54D4F9908716DFCFAE0A39574FE1A" authority="Cotteau, 1888" authorityName="Cotteau" authorityYear="1888" box="[798,1071,472,494]" class="Echinoidea" family="Asterostomatidae" genus="Stomaporus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFCFAE0A392CEFE1A" box="[798,917,472,494]" italics="true" pageId="1" pageNumber="92">Stomaporus</emphasis>
<bibRefCitation id="2D444B3D9908716DFC7AE0A39574FE1A" author="Cotteau, G." box="[926,1071,472,494]" pageId="1" pageNumber="92" pagination="976 - 978" refId="ref5173" refString="Cotteau, G. 1888. Echinides eocenes de la province d' Alicante (Espagne). Compte Rendu de l'Academie des Sciences Paris 107 (25): 976 - 978." type="journal article" year="1888">Cotteau, 1888</bibRefCitation>
</taxonomicName>
, a genus originally assigned to the
<taxonomicName id="8ED54D4F9908716DFCFAE08E92A9FDFF" authority="Gray, 1855" authorityName="Gray" authorityYear="1855" box="[798,1010,501,523]" class="Echinoidea" family="Brissidae" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="family">
Brissidae
<bibRefCitation id="2D444B3D9908716DFC66E08E92A9FDFF" author="Gray, J. E." box="[898,1010,501,523]" pageId="1" pageNumber="92" refId="ref5330" refString="Gray, J. E. 1855. Catalogue of the Recent Echinida, or sea eggs, in the collection of the British Museum 1: Echinida Irregularia. Woodfall and Kinder: London. 69 pp., 6 pls." type="book" year="1855">Gray, 1855</bibRefCitation>
</taxonomicName>
) by
<bibRefCitation id="2D444B3D9908716DFBC4E08E9584FDFF" author="Smith, A. B. &amp; Stockley, B. &amp; Godfrey, D." box="[1056,1247,501,523]" pageId="1" pageNumber="92" refId="ref5974" refString="Smith, A. B., Stockley, B., and Godfrey, D. 2003. Spatangoida. In: Smith, A. B. (ed.), The echinoid directory. http: / www. nhm. ac. uk / palaeontology / echinoids [accessed 19 Jun 2005]." type="book" year="2003">Smith et al. (2003)</bibRefCitation>
. Of the remaining 21 genera, Smith et al. have conditionally reassigned 13 to other families, only six of which are included in the
<taxonomicName id="8ED54D4F9908716DFCFAE33092FBFD95" authorityName="Fisher" authorityYear="1966" box="[798,928,587,609]" class="Echinoidea" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="subOrder" subOrder="Micrasterina">Micrasterina</taxonomicName>
. The remaining eight are listed as
<emphasis id="7BA1EADE9908716DFB14E33794CEFD96" box="[1264,1429,588,610]" italics="true" pageId="1" pageNumber="92">incertae sedis or</emphasis>
as belonging to an unnamed taxon. Until a detailed revision of these latter genera is published, it is considered imprudent to assign the new genus to a specific family.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="40827913B1A0EA16656AF963D5506C9D">Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia</mods:title>
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<subSubSection id="01CF65479908716DFCFAE38B9572FCF2" box="[798,1065,752,774]" pageId="1" pageNumber="92" type="nomenclature">
<paragraph id="496A36CC9908716DFCFAE38B9572FCF2" blockId="1.[798,1065,752,774]" box="[798,1065,752,774]" pageId="1" pageNumber="92">
<heading id="122281A09908716DFCFAE38B9572FCF2" box="[798,1065,752,774]" fontSize="9" level="2" pageId="1" pageNumber="92" reason="5">
<taxonomicName id="8ED54D4F9908716DFCFAE38B929DFCF1" ID-CoL="7PPKN" authority="Holmes &amp; Yee &amp; Krause, 2005" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[798,966,752,773]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="7BA1EADE9908716DFCFAE38B929DFCF1" bold="true" box="[798,966,752,773]" italics="true" pageId="1" pageNumber="92">Murraypneustes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609257A59908716DFC29E38B9572FCF2" box="[973,1065,752,774]" pageId="1" pageNumber="92" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01CF65479908716DFCFAE2609212FCBA" pageId="1" pageNumber="92" type="type_taxon">
<paragraph id="496A36CC9908716DFCFAE2609212FCBA" blockId="1.[798,1429,795,846]" pageId="1" pageNumber="92">
<emphasis id="7BA1EADE9908716DFCFAE260942AFCC5" box="[798,1393,795,817]" italics="true" pageId="1" pageNumber="92">
<typeStatus id="966E886E9908716DFCFAE2609216FCC5" box="[798,845,795,817]" pageId="1" pageNumber="92">Type</typeStatus>
and only known species.
<taxonomicName id="8ED54D4F9908716DFBB6E260942AFCC5" authority="Holmes &amp; Yee &amp; Krause, 2005" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[1106,1393,795,817]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="biannulatus" status="sp. nov.">Murraypneustes biannulatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="609257A59908716DFA9CE2609212FCBA" pageId="1" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="01CF65479908716DFCFAE21F947CFB6F" pageId="1" pageNumber="92" type="diagnosis">
<paragraph id="496A36CC9908716DFCFAE21F947CFB6F" blockId="1.[798,1430,867,1179]" pageId="1" pageNumber="92">
<emphasis id="7BA1EADE9908716DFCFAE21F92D8FC8E" box="[798,899,868,890]" italics="true" pageId="1" pageNumber="92">Diagnosis</emphasis>
. Large ovoid spatangoid with centrally depressed adapical surface, apex well posterior of centre. Apical system ethmolytic with 4 gonopores. Aboral primary tubercles of 2 distinct sizes, small and randomly spaced, the larger proximal to the ambitus. Labrum long, narrow, partially tuberculate, extending to third ambulacral plate. Two peripetalous fascioles present; one marginal, passing above the periproct (pseudolateral); the second, rudimentary, non re-entrant and clear of the distal ends of paired petals. Intermittent horizontal fasciole bands also occur between the 2 peripetalous fascioles. Subanal fasciole in contact with marginal periproct.
</paragraph>
</subSubSection>
<subSubSection id="01CF65479908716DFCFAE5CB9513FAF4" pageId="1" pageNumber="92" type="etymology">
<paragraph id="496A36CC9908716DFCFAE5CB9513FAF4" blockId="1.[798,1429,1200,1280]" pageId="1" pageNumber="92">
<emphasis id="7BA1EADE9908716DFCFAE5CB92D2FB32" box="[798,905,1200,1222]" italics="true" pageId="1" pageNumber="92">Etymology</emphasis>
. For the Murray River cliffs, the origin of the fossils, and “pneustes”, a common suffix used for spatangoid echinoids. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="01CF65479908716EFCFAE46C93C1F8F0" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92" type="discussion">
<paragraph id="496A36CC9908716DFCFAE46C95B3F99F" blockId="1.[798,1429,1303,1961]" pageId="1" pageNumber="92">
<emphasis id="7BA1EADE9908716DFCFAE46C922EFAD9" box="[798,885,1303,1325]" italics="true" pageId="1" pageNumber="92">Remarks</emphasis>
. The following analysis of morphological features of genera similar to
<taxonomicName id="8ED54D4F9908716DFC35E44F9528FABE" authority="Holmes &amp; Yee &amp; Krause, 2005" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[977,1139,1332,1354]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="7BA1EADE9908716DFC35E44F9528FABE" box="[977,1139,1332,1354]" italics="true" pageId="1" pageNumber="92">Murraypneustes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609257A59908716DFB9FE4489583FABD" box="[1147,1240,1331,1353]" pageId="1" pageNumber="92" rank="genus">gen. nov.</taxonomicNameLabel>
is based primarily on
<bibRefCitation id="2D444B3D9908716DFCA2E42B9556FA92" author="Mortensen, T." box="[838,1037,1360,1382]" pageId="1" pageNumber="92" refId="ref5846" refString="Mortensen, T. 1950 a. A monograph of the Echinoidea 5 (1). Spatangoida 1. Protosternata, Meriodosternata, Amphisternata 1. Palaeopneustidae, Palaeostomaridae, Aeropsidae, Toxasteridae, Micrasteridae, Hemiasteridae. C. A. Reitzel: Copenhagen. 432 pp., 25 pls." type="book" year="1950">Mortensen (1950a)</bibRefCitation>
and
<bibRefCitation id="2D444B3D9908716DFBAAE42B947DFA92" author="Smith, A. B. &amp; Stockley, B. &amp; Godfrey, D." box="[1102,1318,1360,1382]" pageId="1" pageNumber="92" refId="ref5974" refString="Smith, A. B., Stockley, B., and Godfrey, D. 2003. Spatangoida. In: Smith, A. B. (ed.), The echinoid directory. http: / www. nhm. ac. uk / palaeontology / echinoids [accessed 19 Jun 2005]." type="book" year="2003">Smith et al. (2003)</bibRefCitation>
. Although approximately 30 genera within the
<taxonomicName id="8ED54D4F9908716DFB46E416947BFA77" authorityName="Fisher" authorityYear="1966" box="[1186,1312,1389,1411]" class="Echinoidea" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="subOrder" subOrder="Micrasterina">Micrasterina</taxonomicName>
have been investigated, only eight warranted further scrutiny; four currently unassigned to a family by
<bibRefCitation id="2D444B3D9908716DFB4AE4DC94CAFA49" author="Smith, A. B. &amp; Stockley, B. &amp; Godfrey, D." box="[1198,1425,1447,1469]" pageId="1" pageNumber="92" refId="ref5974" refString="Smith, A. B., Stockley, B., and Godfrey, D. 2003. Spatangoida. In: Smith, A. B. (ed.), The echinoid directory. http: / www. nhm. ac. uk / palaeontology / echinoids [accessed 19 Jun 2005]." type="book" year="2003">Smith et al. (2003)</bibRefCitation>
,
<taxonomicName id="8ED54D4F9908716DFCFAE4BF9561FA2E" authority="Koehler, 1914" authorityName="Koehler" authorityYear="1914" box="[798,1082,1476,1498]" class="Echinoidea" family="Eurypatagidae" genus="Elipneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFCFAE4BF92CAFA2E" box="[798,913,1476,1498]" italics="true" pageId="1" pageNumber="92">Elipneustes</emphasis>
<bibRefCitation id="2D444B3D9908716DFC45E4BF9561FA2E" author="Koehler, R." box="[929,1082,1476,1498]" pageId="1" pageNumber="92" pagination="191" refId="ref5434" refString="Koehler, R. 1914. Rectification. Zoologischer Anzeiger 44: 191." type="journal article" year="1914">Koehler, 1914</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFBA9E4BF94CAFA2E" authority="Mortensen, 1948" authorityName="Mortensen" authorityYear="1948" box="[1101,1425,1476,1498]" class="Echinoidea" family="Eurypatagidae" genus="Eurypatagus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFBA9E4BF9597FA2E" box="[1101,1228,1476,1498]" italics="true" pageId="1" pageNumber="92">Eurypatagus</emphasis>
<bibRefCitation id="2D444B3D9908716DFB38E4BF94CAFA2E" author="Mortensen, T." box="[1244,1425,1476,1498]" pageId="1" pageNumber="92" pagination="93 - 140" refId="ref5771" refString="Mortensen, T. 1948. Report on the Echinoidea collected by the United States Fisheries Steamer &quot; Albatross &quot; during the Philippine Expedition, 1907 - 1910. Part 3. The Echinoneidae, Echinolampadidae, Clypeasteridae, Arachnoididae, Laganidae, Fibulariidae, Urchinidae, Echinocorythidae, Palaeostomatidae, Micrasteridae, Palaeopneustidae, Hemiasteridae and Spatanidae. Bulletin of the United States National Museum 100 (14): 93 - 140." type="journal article" year="1948">Mortensen, 1948</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFCFAE49A950CFA03" authority="A. Agassiz, 1881" authorityName="A. Agassiz" authorityYear="1881" box="[798,1111,1505,1527]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFCFAE49A92F9FA03" box="[798,930,1505,1527]" italics="true" pageId="1" pageNumber="92">Linopneustes</emphasis>
A.
<bibRefCitation id="2D444B3D9908716DFC22E49A950CFA03" author="Agassiz, A." box="[966,1111,1505,1527]" pageId="1" pageNumber="92" pagination="1 - 321" refId="ref4952" refString="Agassiz, A. 1881. Report on the Echinoidea. Reports on the scientific results of the voyage of H. M. S. Challenger during the years 1873 - 76, Zoology 3 (9): 1 - 321, 45 pls." type="book chapter" year="1881">Agassiz, 1881</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="8ED54D4F9908716DFB69E49A94CAFA03" authority="Grube, 1865" authorityName="Grube" authorityYear="1865" box="[1165,1425,1505,1527]" class="Echinoidea" family="Eurypatagidae" genus="Platybrissus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFB69E49A9453FA03" box="[1165,1288,1505,1527]" italics="true" pageId="1" pageNumber="92">Platybrissus</emphasis>
<bibRefCitation id="2D444B3D9908716DFAF4E49A94CAFA03" author="Grube, A. E." box="[1296,1425,1505,1527]" pageId="1" pageNumber="92" pagination="35 - 37" refId="ref5373" refString="Grube, A. E. 1865. Einige neue Seesterne des hiesigen zoologischen Museums. Jahresbericht der Schlesischen Gesellschaft fur vaterlandische Cultur 1865: 35 - 37." type="journal article" year="1865">Grube, 1865</bibRefCitation>
</taxonomicName>
; three belonging to the
<taxonomicName id="8ED54D4F9908716DFBE0E4859445F9E0" authority="(Lambert, 1905)" baseAuthorityName="Lambert" baseAuthorityYear="1905" box="[1028,1310,1534,1556]" class="Echinoidea" family="Maretiidae" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="family">
Maretiidae (
<bibRefCitation id="2D444B3D9908716DFB9AE485944CF9E0" author="Lambert, J." box="[1150,1303,1534,1556]" pageId="1" pageNumber="92" pagination="129 - 164" refId="ref5448" refString="Lambert, J. 1905. Echinides eoceniques de l'Aude et de l'Herault. Pp. 129 - 164 in: Doncieux, L. (ed.), Catalogue descriptif des fossils nummuliques de l'Aude et de l'Herault. Premiere partie: Montagne Noire et Minervous. Annales de l'Universite de Lyon 17: 1 - 184,5 pls." type="journal article" year="1905">Lambert, 1905</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFACEE4859288F9C5" authority="L. Agassiz, 1847" authorityName="L. Agassiz" authorityYear="1847" class="Echinoidea" family="Eupatagidae" genus="Eupatagus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFACEE48594CEF9E0" box="[1322,1429,1534,1556]" italics="true" pageId="1" pageNumber="92">Eupatagus</emphasis>
L.
<bibRefCitation id="2D444B3D9908716DFCDAE7609288F9C5" author="Agassiz, L. in &amp; Agassiz, L. &amp; Desor, E." box="[830,979,1563,1585]" pageId="1" pageNumber="92" pagination="129 - 168" refId="ref4999" refString="Agassiz, L. in: Agassiz, L., and Desor, E. 1847. Catologue raisonne des families, des genres et des especes de la classe des echinoderms. Annales des Sciences Naturelles 7: 129 - 168." type="journal article" year="1847">Agassiz, 1847</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFC07E760943EF9C5" authority="Lambert and Thiery, 1915" authorityName="Lambert and Thiery" authorityYear="1915" box="[995,1381,1563,1585]" class="Magnoliopsida" family="Asteraceae" genus="Mazzettia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="92" phylum="Tracheophyta" rank="genus">
<emphasis id="7BA1EADE9908716DFC07E760951FF9C5" box="[995,1092,1563,1585]" italics="true" pageId="1" pageNumber="92">Mazzettia</emphasis>
<bibRefCitation id="2D444B3D9908716DFBB4E760943EF9C5" author="Lambert, J. &amp; Thiery, P." box="[1104,1381,1563,1585]" pageId="1" pageNumber="92" pagination="155 - 240" refId="ref5553" refString="Lambert, J., and Thiery, P. 1915. Descriptions des echinides des Terrains Neogenes du Basin du Rhone. Memoires de la Societe Paleontologique Suisse (Abhandlungen der Schweizerischen Palaontologischen Gesellschaft) 41: 155 - 240, pls 13 - 17." type="journal article" year="1915">Lambert and Thiéry, 1915</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8ED54D4F9908716DFCFAE7439529F9BA" authority="H. L. Clark, 1923" authorityName="H. L. Clark" authorityYear="1923" box="[798,1138,1592,1614]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFCFAE74392EDF9BA" box="[798,950,1592,1614]" italics="true" pageId="1" pageNumber="92">Spatagobrissus</emphasis>
H. L.
<bibRefCitation id="2D444B3D9908716DFC1DE7439529F9BA" author="Clark, H. L." box="[1017,1138,1592,1614]" pageId="1" pageNumber="92" pagination="221 - 435" refId="ref5090" refString="Clark, H. L. 1923. The echinoderm fauna of South Africa. Annals of the South African Museum 113: 221 - 435." type="journal article" year="1923">Clark, 1923</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="8ED54D4F9908716DFB4CE743940FF9BA" authorityName="Lambert" authorityYear="1905" box="[1192,1364,1592,1614]" class="Echinoidea" family="Macropneustidae" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="family">Macropneustidae</taxonomicName>
genus
<taxonomicName id="8ED54D4F9908716DFCFAE72E95BFF99F" authority="Lambert and Sanchez Roig, 1924" authorityName="Lambert and Sanchez Roig" authorityYear="1924" box="[798,1252,1621,1643]" class="Echinoidea" family="Macropneustidae" genus="Lajanaster" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFCFAE72E92D1F99F" box="[798,906,1621,1643]" italics="true" pageId="1" pageNumber="92">Lajanaster</emphasis>
<bibRefCitation id="2D444B3D9908716DFC75E72E95BFF99F" author="Lambert, J. &amp; Sanchez Roig, M." box="[913,1252,1621,1643]" pageId="1" pageNumber="92" pagination="7" refId="ref5509" refString="Lambert, J., and Sanchez Roig, M. 1924. P. 449 in: Lambert J., and Thiery, P. Essai de nomenclature raisonnee des echinides 6 and 7. Libraire L. Ferriere: Chaumont." type="book chapter" year="1924">Lambert and Sanchez Roig, 1924</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="496A36CC9908716EFCDAE70990DFFDB1" blockId="1.[798,1429,1303,1961]" lastBlockId="2.[138,770,299,1796]" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
Using a broad comparison of 38 features,
<taxonomicName id="8ED54D4F9908716DFAF5E70994CEF97C" authorityName="A. Agassiz" authorityYear="1881" box="[1297,1429,1650,1672]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFAF5E70994CEF97C" box="[1297,1429,1650,1672]" italics="true" pageId="1" pageNumber="92">Linopneustes</emphasis>
</taxonomicName>
stands out from the other seven genera as having closest affinity with
<taxonomicName id="8ED54D4F9908716DFC64E7D79579F936" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[896,1058,1708,1730]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE9908716DFC64E7D79579F936" box="[896,1058,1708,1730]" italics="true" pageId="1" pageNumber="92">Murraypneustes</emphasis>
</taxonomicName>
; four of its recorded six species,
<taxonomicName id="8ED54D4F9908716DFCFAE7B295D7F92A" authority="(A. Agassiz, 1878)" baseAuthorityName="A. Agassiz" baseAuthorityYear="1878" box="[798,1164,1736,1759]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="longispinus">
<emphasis id="7BA1EADE9908716DFCFAE7B292E8F92B" box="[798,947,1737,1759]" italics="true" pageId="1" pageNumber="92">L. longispinus</emphasis>
(A.
<bibRefCitation id="2D444B3D9908716DFC09E7B395DFF92A" author="Agassiz, A." box="[1005,1156,1736,1758]" pageId="1" pageNumber="92" pagination="185 - 196" refId="ref4866" refString="Agassiz, A. 1878. Report on the Echini in No. 9 (2) - Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, by the United States Coast Survey Steamer &quot; Blake &quot;, Lieutenant-Commander C. D. Sigsbee, U. S. N., commanding. Bulletin of the Museum of Comparative Zoology, Harvard University 5: 185 - 196, pls 1 - 5." type="journal article" year="1878">Agassiz, 1878</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFB79E7B29201F90F" authority="(de Meijere, 1903)" baseAuthorityName="de Meijere" baseAuthorityYear="1903" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="fragilis">
<emphasis id="7BA1EADE9908716DFB79E7B2945CF92B" box="[1181,1287,1737,1759]" italics="true" pageId="1" pageNumber="92">L. fragilis</emphasis>
(de
<bibRefCitation id="2D444B3D9908716DFAA5E7B39208F90F" author="Meijere, J. C. H. de" pageId="1" pageNumber="92" pagination="1 - 16" refId="ref5711" refString="Meijere, J. C. H. de 1903. Vorlaufige Beschreibung der neuen, durch die Siboga-Expedition gesamelten Echiniden. Tijdschrift der Nederlandsche Dierkundige Vereeniging (ser. 2) 8: 1 - 16." type="journal article" year="1903">Meijere, 1903</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="8ED54D4F9908716DFC83E79E95ECF90F" authority="(de Meijere, 1903)" baseAuthorityName="de Meijere" baseAuthorityYear="1903" box="[871,1207,1765,1787]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="spectabilis">
<emphasis id="7BA1EADE9908716DFC83E79E92B5F90F" box="[871,1006,1765,1787]" italics="true" pageId="1" pageNumber="92">L. spectabilis</emphasis>
(de
<bibRefCitation id="2D444B3D9908716DFBFBE79E95EBF90F" author="Meijere, J. C. H. de" box="[1055,1200,1765,1787]" pageId="1" pageNumber="92" pagination="1 - 16" refId="ref5711" refString="Meijere, J. C. H. de 1903. Vorlaufige Beschreibung der neuen, durch die Siboga-Expedition gesamelten Echiniden. Tijdschrift der Nederlandsche Dierkundige Vereeniging (ser. 2) 8: 1 - 16." type="journal article" year="1903">Meijere, 1903</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="8ED54D4F9908716DFB14E79E92BEF8EC" authority="Mortensen, 1950 b" authorityName="Mortensen" authorityYear="1950" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="brachipetalus">
<emphasis id="7BA1EADE9908716DFB14E79E94CEF90F" box="[1264,1429,1765,1787]" italics="true" pageId="1" pageNumber="92">L. brachipetalus</emphasis>
<bibRefCitation id="2D444B3D9908716DFCFAE67992BEF8EC" author="Mortensen, T." box="[798,997,1794,1816]" pageId="1" pageNumber="92" pagination="157 - 163" refId="ref5900" refString="Mortensen, T. 1950 b. New Echinoidea (Spatangoida). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kjobenhavn 112: 157 - 163." type="journal article" year="1950">Mortensen, 1950b</bibRefCitation>
</taxonomicName>
having distinct marginal peripetalous fascioles passing above the periproct. In the other two,
<taxonomicName id="8ED54D4F9908716DFCFAE647950FF8A6" authority="(A. Agassiz, 1879)" baseAuthorityName="A. Agassiz" baseAuthorityYear="1879" box="[798,1108,1852,1874]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="murrayi">
<emphasis id="7BA1EADE9908716DFCFAE64792D0F8A6" box="[798,907,1852,1874]" italics="true" pageId="1" pageNumber="92">L. murrayi</emphasis>
(A. Agassiz, 1879)
</taxonomicName>
and
<taxonomicName id="8ED54D4F9908716DFB6EE6479209F89B" authority="de Meijere, 1903" authorityName="de Meijere" authorityYear="1903" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="excentricus">
<emphasis id="7BA1EADE9908716DFB6EE6479443F8A6" box="[1162,1304,1852,1874]" italics="true" pageId="1" pageNumber="92">L. excentricus</emphasis>
de
<bibRefCitation id="2D444B3D9908716DFAA5E6479209F89B" author="Meijere, J. C. H. de" pageId="1" pageNumber="92" pagination="1 - 16" refId="ref5711" refString="Meijere, J. C. H. de 1903. Vorlaufige Beschreibung der neuen, durch die Siboga-Expedition gesamelten Echiniden. Tijdschrift der Nederlandsche Dierkundige Vereeniging (ser. 2) 8: 1 - 16." type="journal article" year="1903">Meijere, 1903</bibRefCitation>
</taxonomicName>
, the peripetalous fasciole is not marginal but somewhat higher up on the test (
<bibRefCitation id="2D444B3D9908716DFBC5E60D9445F878" author="Mortensen, T." box="[1057,1310,1910,1932]" pageId="1" pageNumber="92" refId="ref5846" refString="Mortensen, T. 1950 a. A monograph of the Echinoidea 5 (1). Spatangoida 1. Protosternata, Meriodosternata, Amphisternata 1. Palaeopneustidae, Palaeostomaridae, Aeropsidae, Toxasteridae, Micrasteridae, Hemiasteridae. C. A. Reitzel: Copenhagen. 432 pp., 25 pls." type="book" year="1950">Mortensen, 1950a: 221</bibRefCitation>
). Although
<taxonomicName id="8ED54D4F9908716DFCFAE6E892DCF85D" baseAuthorityName="de Meijere" baseAuthorityYear="1903" box="[798,903,1939,1961]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="92" phylum="Echinodermata" rank="species" species="fragilis">
<emphasis id="7BA1EADE9908716DFCFAE6E892DCF85D" box="[798,903,1939,1961]" italics="true" pageId="1" pageNumber="92">L. fragilis</emphasis>
</taxonomicName>
has multiple fasciole bands around the ambitus,
<taxonomicName id="8ED54D4F990B716EFF6EE05091A1FEB5" baseAuthorityName="A. Agassiz" baseAuthorityYear="1879" box="[138,250,299,321]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="species" species="murrayi">
<emphasis id="7BA1EADE990B716EFF6EE05091A1FEB5" box="[138,250,299,321]" italics="true" pageId="2" pageNumber="93">L. murrayi</emphasis>
</taxonomicName>
a rounded margin (Smith, pers. com. 2005) and
<taxonomicName id="8ED54D4F990B716EFF6EE0339041FEAA" baseAuthorityName="A. Agassiz" baseAuthorityYear="1878" box="[138,282,328,350]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="species" species="longispinus">
<emphasis id="7BA1EADE990B716EFF6EE0339041FEAA" box="[138,282,328,350]" italics="true" pageId="2" pageNumber="93">L. longispinus</emphasis>
</taxonomicName>
relatively short closing petals;
<taxonomicName id="8ED54D4F990B716EFDBBE033925AFEAA" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[607,769,328,350]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFDBBE033925AFEAA" box="[607,769,328,350]" italics="true" pageId="2" pageNumber="93">Murraypneustes</emphasis>
</taxonomicName>
is distinguished by its centrally depressed aboral surface forming four apices (domed on
<taxonomicName id="8ED54D4F990B716EFE72E0F99341FE6C" authorityName="A. Agassiz" authorityYear="1881" box="[406,538,386,408]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFE72E0F99341FE6C" box="[406,538,386,408]" italics="true" pageId="2" pageNumber="93">Linopneustes</emphasis>
</taxonomicName>
) with the highest point of the test posterior to the apical disk (anterior in
<taxonomicName id="8ED54D4F990B716EFD8BE0E493A8FE41" authorityName="A. Agassiz" authorityYear="1881" box="[623,755,415,437]" class="Echinoidea" family="Eurypatagidae" genus="Linopneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFD8BE0E493A8FE41" box="[623,755,415,437]" italics="true" pageId="2" pageNumber="93">Linopneustes</emphasis>
</taxonomicName>
), the presence of two peripetalous fascioles and intermittent horizontal fasciole bands, two distinct sizes of small randomly spaced aboral primary tubercles, the larger restricted to the area outside the upper peripetalous fasciole, and a prominent angular subanal fasciole.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE337933EFD68" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
Based on the same criteria, three other genera show a moderate degree of morphological similarity to
<taxonomicName id="8ED54D4F990B716EFDBDE31293A0FD8B" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[601,763,617,639]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFDBDE31293A0FD8B" box="[601,763,617,639]" italics="true" pageId="2" pageNumber="93">Murraypneustes</emphasis>
</taxonomicName>
, namely
<taxonomicName id="8ED54D4F990B716EFF3FE3FD9015FD68" authorityName="Koehler" authorityYear="1914" box="[219,334,646,668]" class="Echinoidea" family="Eurypatagidae" genus="Elipneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFF3FE3FD9015FD68" box="[219,334,646,668]" italics="true" pageId="2" pageNumber="93">Elipneustes</emphasis>
</taxonomicName>
,
<taxonomicName id="8ED54D4F990B716EFEBFE3FD90E7FD68" authorityName="Lambert and Thiery" authorityYear="1915" box="[347,444,646,668]" class="Magnoliopsida" family="Asteraceae" genus="Mazzettia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="7BA1EADE990B716EFEBFE3FD90E7FD68" box="[347,444,646,668]" italics="true" pageId="2" pageNumber="93">Mazzettia</emphasis>
</taxonomicName>
, and
<taxonomicName id="8ED54D4F990B716EFE10E3FD9304FD68" authorityName="L. Agassiz" authorityYear="1847" box="[500,607,646,668]" class="Echinoidea" family="Eupatagidae" genus="Eupatagus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFE10E3FD9304FD68" box="[500,607,646,668]" italics="true" pageId="2" pageNumber="93">Eupatagus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE3D891A2FC92" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
<taxonomicName id="8ED54D4F990B716EFF4EE3D89046FD4D" authorityName="Koehler" authorityYear="1914" box="[170,285,675,697]" class="Echinoidea" family="Eurypatagidae" genus="Elipneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFF4EE3D89046FD4D" box="[170,285,675,697]" italics="true" pageId="2" pageNumber="93">Elipneustes</emphasis>
</taxonomicName>
, however, can be distinguished by its very long, parallel-sided, open-ended, flush petals with conjugate pore pairs; single, narrow, marginally situated peripetalous fasciole; peristome situated immediately below the apical disk; small thickset plastron with minimal posterior swelling; and much larger primary tubercles scattered over all the aboral interambulacra.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE216939CFBE0" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
<taxonomicName id="8ED54D4F990B716EFF4EE2169050FC77" authorityName="Lambert and Thiery" authorityYear="1915" box="[170,267,877,899]" class="Magnoliopsida" family="Asteraceae" genus="Mazzettia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="7BA1EADE990B716EFF4EE2169050FC77" box="[170,267,877,899]" italics="true" pageId="2" pageNumber="93">Mazzettia</emphasis>
</taxonomicName>
, a poorly known fossil genus, has an elongated heart-shaped test with a low sharp margin, very long weaklybowed petals closing distally, no recorded peripetalous fasciole and only occasionally a weak shield-shaped subanal fasciole, an elongated labrum just reaching the relatively small plastron, and random but closely spaced coarse tubercles aborally.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE56090F3FA78" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
<taxonomicName id="8ED54D4F990B716EFF4EE5609043FBC5" authorityName="L. Agassiz" authorityYear="1847" box="[170,280,1051,1073]" class="Echinoidea" family="Eupatagidae" genus="Eupatagus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFF4EE5609043FBC5" box="[170,280,1051,1073]" italics="true" pageId="2" pageNumber="93">Eupatagus</emphasis>
</taxonomicName>
, although possessing well developed peripetalous and subanal fascioles is easily categorised by its lack of an anterior sulcus, predominately short lanceolate closed petals, primary tubercles of varying density restricted to the area within the peripetalous fasciole in aboral interlambracra 14, and very small evenly spaced tubercles distally over the remainder of the aboral surface. Comparison of other features is difficult because of variability among the very large number of described species. In particular, the eight Australian fossil species vary considerably in profile, are generally more rounded, and have an extremely wide range of primary tubercle densities when compared with the
<typeStatus id="966E886E990B716EFE10E4229344FA9B" box="[500,543,1369,1391]" pageId="2" pageNumber="93">type</typeStatus>
species on which the generic description is based.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE4E99387FA16" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
Two other genera,
<taxonomicName id="8ED54D4F990B716EFE81E4E890BBFA5D" authorityName="Grube" authorityYear="1865" box="[357,480,1427,1449]" class="Echinoidea" family="Eurypatagidae" genus="Platybrissus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFE81E4E890BBFA5D" box="[357,480,1427,1449]" italics="true" pageId="2" pageNumber="93">Platybrissus</emphasis>
</taxonomicName>
and
<taxonomicName id="8ED54D4F990B716EFDF5E4E893CBFA5D" authorityName="Mortensen" authorityYear="1948" box="[529,656,1427,1449]" class="Echinoidea" family="Eurypatagidae" genus="Eurypatagus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFDF5E4E893CBFA5D" box="[529,656,1427,1449]" italics="true" pageId="2" pageNumber="93">Eurypatagus</emphasis>
</taxonomicName>
, are distinguished by their complete lack of fascioles, although the former may have a subanal fasciole present in juvenile specimens.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE4929184F987" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
<taxonomicName id="8ED54D4F990B716EFF4EE492904DFA0B" authorityName="Lambert and Sanchez Roig" authorityYear="1924" box="[170,278,1513,1535]" class="Echinoidea" family="Macropneustidae" genus="Lajanaster" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFF4EE492904DFA0B" box="[170,278,1513,1535]" italics="true" pageId="2" pageNumber="93">Lajanaster</emphasis>
</taxonomicName>
, although depressed aborally, has no anterior sulcus, a relatively sharp ambitus, a markedly narrow plaston, and primary tubercles confined to the posterior column of anterior and lateral interambulacra within the peripetalous fasciole.
</paragraph>
<paragraph id="496A36CC990B716EFF4EE70193C1F8F0" blockId="2.[138,770,299,1796]" pageId="2" pageNumber="93">
<taxonomicName id="8ED54D4F990B716EFF4EE7019019F964" authorityName="H. L. Clark" authorityYear="1923" box="[170,322,1658,1680]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFF4EE7019019F964" box="[170,322,1658,1680]" italics="true" pageId="2" pageNumber="93">Spatagobrissus</emphasis>
</taxonomicName>
was included in the comparative analysis, primarily as a new species of the genus occurs at the same locality (
<collectionCode id="2FC4AE09990B716EFF07E7CF9059F93E" box="[227,258,1716,1738]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="93" type="Museum">PL</collectionCode>
3203) as
<taxonomicName id="8ED54D4F990B716EFEBAE7CF935BF93E" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[350,512,1716,1738]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990B716EFEBAE7CF935BF93E" box="[350,512,1716,1738]" italics="true" pageId="2" pageNumber="93">Murraypneustes</emphasis>
</taxonomicName>
. In most respects the former is very similar to
<taxonomicName id="8ED54D4F990B716EFE80E7AA9097F913" authorityName="L. Agassiz" authorityYear="1847" box="[356,460,1745,1767]" class="Echinoidea" family="Eupatagidae" genus="Eupatagus" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="95" phylum="Echinodermata" rank="genus">Eupatagus</taxonomicName>
but has shorter petals and only small tubercles over the whole of the aboral surface.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="5A3342145285DC894314A95B77661C47" ID-DOI="10.24199/j.mmv.2005.62.3" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665587" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1708004635701" checkinUser="felipe" docAuthor="Holmes, Francis C., Yee, Christopher Ah &amp; Krause, Janice" docDate="2005" docId="C17C87DA990A7168FF66E49591B6FBE7" docLanguage="en" docName="MemMusVictoria.62.1.91-99.pdf" docOrigin="Memoirs of Museum Victoria 62 (1)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-62-issue-1-2005/pages-91-99/" docStyle="DocumentStyle:99484BB52C5EA01CDE4EA5EECB5782C2.2:MemMusVictoria.2003-.journal_article.type2" docStyleId="99484BB52C5EA01CDE4EA5EECB5782C2" docStyleName="MemMusVictoria.2003-.journal_article.type2" docStyleVersion="2" docTitle="Spatagobrissus dermodyorum Holmes &amp; Yee &amp; Krause 2005, sp. nov" docType="treatment" docVersion="3" lastPageNumber="95" masterDocId="3D45FFA29909716CFFE4E17B915BFFF4" masterDocTitle="Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia" masterLastPageNumber="99" masterPageNumber="91" pageNumber="94" updateTime="1711397301711" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="66914D04A1C4D2DD6687DB5075C2B52F">Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia</mods:title>
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<mods:namePart id="F3DC9A9B63BF82C414E35E557E799FDF">Holmes, Francis C.</mods:namePart>
<mods:affiliation id="459B4C3470E2CCC5FADB8B4477AAB404">Kenbry Road, Heathmont, Victoria 3135, Australia, and Invertebrate Palaeontology, Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia (fholmes @ bigpond. net. au)</mods:affiliation>
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<mods:namePart id="321D6AC48E81AC1A4109F5B97EE8F580">Yee, Christopher Ah</mods:namePart>
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<mods:namePart id="F70B601D142688A801F6A79A83781F9C">Krause, Janice</mods:namePart>
<mods:affiliation id="CDCFF33DCFC5CEFF90B0AD3D51A05099">P. O. Box 581, Hamilton, Victoria 3300, Australia</mods:affiliation>
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<paragraph id="496A36CC990A716FFF66E495935FF9F0" blockId="3.[130,516,1518,1540]" box="[130,516,1518,1540]" pageId="3" pageNumber="94">
<heading id="122281A0990A716FFF66E495935FF9F0" box="[130,516,1518,1540]" fontSize="9" level="2" pageId="3" pageNumber="94" reason="5">
<taxonomicName id="8ED54D4F990A716FFF66E49590EFF9F0" ID-CoL="4YFGS" authority="Holmes &amp; Yee &amp; Krause, 2005" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[130,436,1518,1540]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="dermodyorum" status="sp. nov.">
<emphasis id="7BA1EADE990A716FFF66E49590EFF9F0" bold="true" box="[130,436,1518,1540]" italics="true" pageId="3" pageNumber="94">Spatagobrissus dermodyorum</emphasis>
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<taxonomicNameLabel id="609257A5990A716FFE5FE495935FF9F0" box="[443,516,1518,1540]" pageId="3" pageNumber="94" rank="species">sp. nov</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="01CF6547990A716FFF66E76190C5F9C4" box="[130,414,1562,1584]" pageId="3" pageNumber="94" type="description">
<paragraph id="496A36CC990A716FFF66E76190C5F9C4" blockId="3.[130,414,1562,1584]" box="[130,414,1562,1584]" pageId="3" pageNumber="94">
<figureCitation id="D1EE2A49990A716FFF66E761904BF9C4" box="[130,272,1562,1584]" captionStart="Figure 6" captionStartId="7.[130,188,1789,1809]" captionTargetBox="[127,1427,1011,1764]" captionTargetId="figure-97@7.[126,1428,1011,1764]" captionTargetPageId="7" captionText="Figure 6. Spatagobrissus dermodyorum sp. nov. A, B, D, adapical, adoral, and left lateral views of holotype, NMV P312570; C, adoral view of paratype, NMV P312571; E, posterior view of paratype, NMV P312572 All specimens from the early Middle Miocene Glenforslan Formation in the vicinity of locality NMV PL3203, north of Blanchetown, South Australia. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10665599" httpUri="https://zenodo.org/record/10665599/files/figure.png" pageId="3" pageNumber="94">Figures 6AE</figureCitation>
,
<figureCitation id="D1EE2A49990A716FFEFFE7619002F9C4" box="[283,345,1562,1584]" captionStart="Figure 7" captionStartId="8.[138,196,768,788]" captionTargetBox="[138,1440,295,742]" captionTargetId="figure-11@8.[138,1441,295,743]" captionTargetPageId="8" captionText="Figure 7. Spatagobrissus dermodyorum sp. nov. A, detail of petals and apical disk of holotype, NMV P312570; B, peristome and labrum tuberculation of paratype, NMV P312571; C, subanal fasciole and pore pairs of paratype, NMV P312573. Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.10665601" httpUri="https://zenodo.org/record/10665601/files/figure.png" pageId="3" pageNumber="94">7AC</figureCitation>
,
<figureCitation id="D1EE2A49990A716FFE80E76190C5F9C4" box="[356,414,1562,1584]" captionStart="Figure 8" captionStartId="8.[138,196,1349,1369]" captionTargetBox="[152,1440,929,1322]" captionTargetId="figure-57@8.[130,1460,914,1331]" captionTargetPageId="8" captionText="Figure 8. Comparative biometric data on specimens of Spatagobrissus laubei (Duncan, 1877) from the Middle Miocene Port Campbell Limestone, Port Campbell, Victoria (•), and S. dermodyorum sp. nov. from the early Middle Miocene Glenforslan Formation, Blanchetown district, South Australia (o)." figureDoi="http://doi.org/10.5281/zenodo.10665603" httpUri="https://zenodo.org/record/10665603/files/figure.png" pageId="3" pageNumber="94">8AE</figureCitation>
</paragraph>
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<subSubSection id="01CF6547990A716FFF66E73F900FF900" pageId="3" pageNumber="94" type="materials_examined">
<paragraph id="496A36CC990A716FFF66E73F90CDF952" blockId="3.[130,762,1603,1780]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFF66E73F905AF9A2" box="[130,257,1603,1622]" italics="true" pageId="3" pageNumber="94">Type Material</emphasis>
.
<materialsCitation id="F9BD3C91990A716FFEE9E73890CDF952" collectionCode="NMV" county="Middle Miocene Glenforslan Formation" location="Blanchetown" municipality="Morgan Group" pageId="3" pageNumber="94" specimenCode="P312570, PL3203" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<typeStatus id="966E886E990A716FFEE9E7389038F9A3" box="[269,355,1603,1623]" pageId="3" pageNumber="94" type="holotype">Holotype</typeStatus>
.
<collectionCode id="2FC4AE09990A716FFE89E73890FAF9A2" box="[365,417,1603,1622]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">NMV</collectionCode>
<specimenCode id="19739EB7990A716FFE43E73890ADF9A3" box="[423,502,1603,1623]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">P312570</specimenCode>
from early
<collectingCounty id="A00B4E40990A716FFD87E738900BF985" pageId="3" pageNumber="94">Middle Miocene Glenforslan Formation</collectingCounty>
(Batesfordian),
<collectingMunicipality id="A90EACB6990A716FFE01E7259335F985" box="[485,622,1629,1649]" pageId="3" pageNumber="94">Morgan Group</collectingMunicipality>
, in the vicinity of
<collectionCode id="2FC4AE09990A716FFF45E703918EF97F" box="[161,213,1656,1675]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">NMV</collectionCode>
locality
<specimenCode id="19739EB7990A716FFED4E70C9022F97F" box="[304,377,1655,1675]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="94" type="Museum">PL3203</specimenCode>
,
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<quantity id="8E2D9B29990A716FFE6CE70390E6F97F" box="[392,445,1655,1675]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="94" unit="km" value="7.0">7 km</quantity>
NNE of Murray River Lock
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1,
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,
<collectingRegion id="8B11F82E990A716FFEE0E7E990C9F952" box="[260,402,1682,1702]" country="Australia" name="South Australia" pageId="3" pageNumber="94">South Australia</collectingRegion>
.
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</paragraph>
<paragraph id="496A36CC990A716FFF78E7D6900FF900" blockId="3.[130,762,1603,1780]" pageId="3" pageNumber="94">
<materialsCitation id="F9BD3C91990A716FFF78E7D690D5F934" box="[156,398,1708,1728]" collectionCode="NMV" pageId="3" pageNumber="94" specimenCode="P312571" specimenCount="1" typeStatus="paratype">
<typeStatus id="966E886E990A716FFF78E7D691AEF934" box="[156,245,1709,1728]" pageId="3" pageNumber="94" type="paratype">Paratypes</typeStatus>
,
<collectionCode id="2FC4AE09990A716FFEE6E7D6906DF934" box="[258,310,1709,1728]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">NMV</collectionCode>
<specimenCode id="19739EB7990A716FFEDBE7D790D5F934" box="[319,398,1708,1728]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">P312571</specimenCode>
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<materialsCitation id="F9BD3C91990A716FFE7DE7D790B2F934" box="[409,489,1708,1728]" collectionCode="NMV" pageId="3" pageNumber="94" specimenCode="P312373" specimenCount="1" typeStatus="paratype">
<specimenCode id="19739EB7990A716FFE7DE7D790B2F934" box="[409,489,1708,1728]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="94" type="Museum">P312373</specimenCode>
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from the same general area. Other material used for statistical purposes is held in Museum
<collectingRegion id="8B11F82E990A716FFD55E7BD93A2F92E" box="[689,761,1734,1754]" country="Australia" name="Victoria" pageId="3" pageNumber="94">Victoria</collectingRegion>
and private collections.
</paragraph>
</subSubSection>
<subSubSection id="01CF6547990A716FFF66E67192E6F987" pageId="3" pageNumber="94" type="description">
<paragraph id="496A36CC990A716FFF66E67192A0FE40" blockId="3.[130,761,1802,1969]" lastBlockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFF66E67191A3F8D4" box="[130,248,1802,1824]" italics="true" pageId="3" pageNumber="94">Description</emphasis>
. Test small, subcircular in outline with minimal posterior truncation and no anterior sulcus. Specimens range 30.0
<quantity id="8E2D9B29990A716FFF5EE63F9041F8AE" box="[186,282,1860,1882]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="3" pageNumber="94" unit="mm" value="45.5">45.5 mm</quantity>
in length with maximum width 8290%TL occurring 5158 %TL from anterior ambitus. Maximum height 49.560%TL at 52.665.5%TL from anterior ambitus. Adapical surface moderately inflated, evenly curved transversely above well-rounded margin with ambitus situated at about 30%TH. Adoral surface very mildly convex but with prominent posterior keel caused by sharp rise of ambulacra I and
<collectionCode id="2FC4AE09990A716FFCAEE0FA9207FE63" box="[842,860,385,407]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="94" type="Museum">V</collectionCode>
posterior to centre. In lateral view, posterior truncation covers about 35%TH.
</paragraph>
<paragraph id="496A36CC990A716FFCDAE0C09226FCBD" blockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">
Apical system ethmolytic with 4 genital pores, centre 31.338.6%TL from anterior ambitus (
<figureCitation id="D1EE2A49990A716FFB48E0A395A4FE1A" box="[1196,1279,472,494]" captionStart="Figure 7" captionStartId="8.[138,196,768,788]" captionTargetBox="[138,1440,295,742]" captionTargetId="figure-11@8.[138,1441,295,743]" captionTargetPageId="8" captionText="Figure 7. Spatagobrissus dermodyorum sp. nov. A, detail of petals and apical disk of holotype, NMV P312570; B, peristome and labrum tuberculation of paratype, NMV P312571; C, subanal fasciole and pore pairs of paratype, NMV P312573. Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.10665601" httpUri="https://zenodo.org/record/10665601/files/figure.png" pageId="3" pageNumber="94">Fig. 7A</figureCitation>
). Paired petals short (26.532.5%TL measured along surface of perradial suture from centre of apical disk), narrow (7.09.0%TL), lanceolate, closing to closed, posterior pair marginally longer than anterior pair. Anterior pair diverge at about 135°, posterior pair 310°. Pore pairs conjugate, inner pores oval, outer-tear shaped. Anterior row of pore pairs in anterior paired petals distinctly narrower than posterior row and atrophied adapically. Interporiferous zone up to 1.5 times width of poriferous zone at widest point. Ambulacrum III flush adapially, with 2 rows of indistinct longitudinally orientated pore pairs and interporiferous zone containing few secondary tubercles and numerous miliaries.
</paragraph>
<paragraph id="496A36CC990A716FFCDAE22B9234FC02" blockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">Peripetalous fasciole subcircular, not indented. Numerous small, randomly spaced perforate, crenulate, primary tubercles occur in interambulacra within fasciole. Outside fasciole, tubercles in posterior half of test very small, but anterior to centre, gradually increasing in size towards interambulacra 2 and 3.</paragraph>
<paragraph id="496A36CC990A716FFCDAE28692DAFB0F" blockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">
Peristome reniform, mildly sunken, width 16.019.3%TL, length 8.910.3%TL with anterior border 23.426.8%TL from anterior ambitus. Phyllodes moderately developed with slotshaped pores in circular depressions. Labrum short and wedgeshaped extending only to centre of second pair of adjacent ambulacral plates. Anterior edge raised above surrounding ambulacra and slightly projecting over peristome. Miliary tubercles present with few secondary tubercles in anterior half (
<figureCitation id="D1EE2A49990A716FFCC1E59E922FFB0F" box="[805,884,1253,1275]" captionStart="Figure 7" captionStartId="8.[138,196,768,788]" captionTargetBox="[138,1440,295,742]" captionTargetId="figure-11@8.[138,1441,295,743]" captionTargetPageId="8" captionText="Figure 7. Spatagobrissus dermodyorum sp. nov. A, detail of petals and apical disk of holotype, NMV P312570; B, peristome and labrum tuberculation of paratype, NMV P312571; C, subanal fasciole and pore pairs of paratype, NMV P312573. Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.10665601" httpUri="https://zenodo.org/record/10665601/files/figure.png" pageId="3" pageNumber="94">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="496A36CC990A716FFCDAE479951AFA0B" blockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">
Plastron long, width about 5560% length, with ambulacral plates indenting posteriorly. Subanal fasciole circular to transversely oval, enclosing 3 pore pairs each side of interradial suture, and with slight anterior projection at posterior end of prominent plastronal keel. Posterior edge of fasciole marginally clear of periproct opening. Adorally, ambulacra I and
<collectionCode id="2FC4AE09990A716FFA67E4E994CEFA5C" box="[1411,1429,1426,1448]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="94" type="Museum">V</collectionCode>
relatively wide, covered with very fine, randomly spaced miliary tubercles up to sixth plate, then by tubercles of similar size to rest of adoral surface.
</paragraph>
<paragraph id="496A36CC990A716FFCDAE77D92E6F987" blockId="3.[798,1430,298,1651]" pageId="3" pageNumber="94">Periproct, tear-drop shaped, generally positioned vertically on truncated posterior margin but slightly visible from above on some specimens; height 17.119.1%TL, width 13.216.0%TL.</paragraph>
</subSubSection>
<subSubSection id="01CF6547990A716FFCFAE7F392B2F94F" pageId="3" pageNumber="94" type="etymology">
<paragraph id="496A36CC990A716FFCFAE7F392B2F94F" blockId="3.[798,1429,1672,1723]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFCFAE7F392D2F96A" box="[798,905,1672,1694]" italics="true" pageId="3" pageNumber="94">Etymology</emphasis>
. Named for Michael and Marie Dermody, owners of Glenforslan Station.
</paragraph>
</subSubSection>
<subSubSection id="01CF6547990A7168FCFAE7AB91B6FBE7" lastPageId="4" lastPageNumber="95" pageId="3" pageNumber="94" type="discussion">
<paragraph id="496A36CC990A7168FCFAE7AB9316FE63" blockId="3.[798,1429,1744,1969]" lastBlockId="4.[138,769,298,1043]" lastPageId="4" lastPageNumber="95" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFCFAE7AB922EF912" box="[798,885,1744,1766]" italics="true" pageId="3" pageNumber="94">Remarks</emphasis>
.
<taxonomicName id="8ED54D4F990A716FFC6CE7AB95E2F912" authority="Holmes &amp; Yee &amp; Krause, 2005" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[904,1209,1744,1766]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="dermodyorum" status="sp. nov.">
<emphasis id="7BA1EADE990A716FFC6CE7AB95E2F912" box="[904,1209,1744,1766]" italics="true" pageId="3" pageNumber="94">Spatagobrissus dermodyorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609257A5990A716FFB21E7AB9442F912" box="[1221,1305,1744,1766]" pageId="3" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
differs primarily from the Middle Miocene Port Campbell Limestone species,
<taxonomicName id="8ED54D4F990A716FFC98E6719441F8D4" authority="(Duncan, 1877)" baseAuthorityName="Duncan" baseAuthorityYear="1877" box="[892,1306,1802,1824]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="laubei">
<emphasis id="7BA1EADE990A716FFC98E6719539F8D4" box="[892,1122,1802,1824]" italics="true" pageId="3" pageNumber="94">Spatagobrissus laubei</emphasis>
(
<bibRefCitation id="2D444B3D990A716FFB9DE6719448F8D4" author="Duncan, P. M." box="[1145,1299,1802,1824]" pageId="3" pageNumber="94" pagination="42 - 73" refId="ref5206" refString="Duncan, P. M., 1877. On the Echinodermata of the Australian Cainozoic (Tertiary) deposits. Quarterly Journal of the Geological Society of London 33 (1): 42 - 73, pls 3, 4." type="book chapter" year="1877">Duncan, 1877</bibRefCitation>
)
</taxonomicName>
, in having narrower test with more posterior maximum width (
<figureCitation id="D1EE2A49990A716FFAD3E65C94D2F8C9" box="[1335,1417,1831,1853]" captionStart="Figure 8" captionStartId="8.[138,196,1349,1369]" captionTargetBox="[152,1440,929,1322]" captionTargetId="figure-57@8.[130,1460,914,1331]" captionTargetPageId="8" captionText="Figure 8. Comparative biometric data on specimens of Spatagobrissus laubei (Duncan, 1877) from the Middle Miocene Port Campbell Limestone, Port Campbell, Victoria (•), and S. dermodyorum sp. nov. from the early Middle Miocene Glenforslan Formation, Blanchetown district, South Australia (o)." figureDoi="http://doi.org/10.5281/zenodo.10665603" httpUri="https://zenodo.org/record/10665603/files/figure.png" pageId="3" pageNumber="94">Fig. 8A</figureCitation>
), markedly larger peristome and periproct (
<figureCitation id="D1EE2A49990A716FFB5BE63F9477F8AE" box="[1215,1324,1860,1882]" captionStart="Figure 8" captionStartId="8.[138,196,1349,1369]" captionTargetBox="[152,1440,929,1322]" captionTargetId="figure-57@8.[130,1460,914,1331]" captionTargetPageId="8" captionText="Figure 8. Comparative biometric data on specimens of Spatagobrissus laubei (Duncan, 1877) from the Middle Miocene Port Campbell Limestone, Port Campbell, Victoria (•), and S. dermodyorum sp. nov. from the early Middle Miocene Glenforslan Formation, Blanchetown district, South Australia (o)." figureDoi="http://doi.org/10.5281/zenodo.10665603" httpUri="https://zenodo.org/record/10665603/files/figure.png" pageId="3" pageNumber="94">Fig. 8D, E</figureCitation>
), and very much shorter labrum (
<figureCitation id="D1EE2A49990A716FFBE9E61A953AF883" box="[1037,1121,1889,1911]" captionStart="Figure 8" captionStartId="8.[138,196,1349,1369]" captionTargetBox="[152,1440,929,1322]" captionTargetId="figure-57@8.[130,1460,914,1331]" captionTargetPageId="8" captionText="Figure 8. Comparative biometric data on specimens of Spatagobrissus laubei (Duncan, 1877) from the Middle Miocene Port Campbell Limestone, Port Campbell, Victoria (•), and S. dermodyorum sp. nov. from the early Middle Miocene Glenforslan Formation, Blanchetown district, South Australia (o)." figureDoi="http://doi.org/10.5281/zenodo.10665603" httpUri="https://zenodo.org/record/10665603/files/figure.png" pageId="3" pageNumber="94">Fig. 8B</figureCitation>
). In addition, anterior paired petals are shorter and posterior petals longer (
<figureCitation id="D1EE2A49990A716FFAE4E6059409F860" box="[1280,1362,1918,1940]" captionStart="Figure 8" captionStartId="8.[138,196,1349,1369]" captionTargetBox="[152,1440,929,1322]" captionTargetId="figure-57@8.[130,1460,914,1331]" captionTargetPageId="8" captionText="Figure 8. Comparative biometric data on specimens of Spatagobrissus laubei (Duncan, 1877) from the Middle Miocene Port Campbell Limestone, Port Campbell, Victoria (•), and S. dermodyorum sp. nov. from the early Middle Miocene Glenforslan Formation, Blanchetown district, South Australia (o)." figureDoi="http://doi.org/10.5281/zenodo.10665603" httpUri="https://zenodo.org/record/10665603/files/figure.png" pageId="3" pageNumber="94">Fig. 8C</figureCitation>
), with divergent angle of latter greater than
<taxonomicName id="8ED54D4F990A716FFB71E6E095B6F845" baseAuthorityName="Duncan" baseAuthorityYear="1877" box="[1173,1261,1947,1969]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="laubei">
<emphasis id="7BA1EADE990A716FFB71E6E095B6F845" box="[1173,1261,1947,1969]" italics="true" pageId="3" pageNumber="94">S. laubei</emphasis>
</taxonomicName>
. Aboral primary tubercles larger within peripetalous fasciole and interambulacra 2 and 3, while fine tubercles outside fasciole on posterior half of test are much smaller in diameter. Adorally, plastron wider and longer, and ambulacra I and
<collectionCode id="2FC4AE09990D7168FE30E0FA90BDFE63" box="[468,486,385,407]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="95" type="Museum">V</collectionCode>
narrower.
</paragraph>
<paragraph id="496A36CC990D7168FF4EE0E59341FD8A" blockId="4.[138,769,298,1043]" pageId="4" pageNumber="95">
The extant
<typeStatus id="966E886E990D7168FEF8E0E5901CFE40" box="[284,327,414,436]" pageId="4" pageNumber="95">type</typeStatus>
species
<taxonomicName id="8ED54D4F990D7168FE7AE0E593C3FE40" authorityName="H. L. Clark" authorityYear="1923" box="[414,664,414,436]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="7BA1EADE990D7168FE7AE0E593C3FE40" box="[414,664,414,436]" italics="true" pageId="4" pageNumber="95">Spatagobrissus mirabilis</emphasis>
</taxonomicName>
is characterised by a larger (up to
<quantity id="8E2D9B29990D7168FE75E0C090BFFE25" box="[401,484,443,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="95" unit="mm" value="110.0">110 mm</quantity>
long) and less inflated test, more posteriorly located apical system in line with maximum width, greater area enclosed by peripetalous fasciole, shorter peristome, and periproct situated on an obliquely truncated surface below the ambitus. Primary tubercles within the peripetalous fasciole of
<taxonomicName id="8ED54D4F990D7168FE67E33790A1FD96" authorityName="H. L. Clark" authorityYear="1923" box="[387,506,588,610]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="7BA1EADE990D7168FE67E33790A1FD96" box="[387,506,588,610]" italics="true" pageId="4" pageNumber="95">S. mirabilis</emphasis>
</taxonomicName>
are also larger and more closely spaced than in
<taxonomicName id="8ED54D4F990D7168FE8BE312934FFD8B" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[367,532,617,639]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="dermodyorum">
<emphasis id="7BA1EADE990D7168FE8BE312934FFD8B" box="[367,532,617,639]" italics="true" pageId="4" pageNumber="95">S. dermodyorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="496A36CC990D7168FF4EE3FE9184FCBD" blockId="4.[138,769,298,1043]" pageId="4" pageNumber="95">
<taxonomicName id="8ED54D4F990D7168FF4EE3FE93D5FD6F" ID-CoL="6Z79C" authority="Baker and Rowe, 1990" authorityName="Baker and Rowe" authorityYear="1990" box="[170,654,645,667]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="incus">
<emphasis id="7BA1EADE990D7168FF4EE3FE90DFFD6F" box="[170,388,645,667]" italics="true" pageId="4" pageNumber="95">Spatagobrissus incus</emphasis>
<bibRefCitation id="2D444B3D990D7168FE75E3FE93D5FD6F" author="Baker, A. L. &amp; Rowe, F. W. E." box="[401,654,645,667]" pageId="4" pageNumber="95" pagination="281 - 316" refId="ref5043" refString="Baker, A. L., and Rowe, F. W. E. 1990. Atelostomatid sea urchins from Australia and New Zealand waters (Echinoidea: Cassiduloida, Holasteroida, Spatangoida, Neolampadoida). Invertebrate Taxonomy 4: 281 - 316." type="journal article" year="1990">Baker and Rowe, 1990</bibRefCitation>
</taxonomicName>
, an extant species endemic to southeast Australian waters, particularly between Flinders Island (
<collectingRegion id="8B11F82E990D7168FE63E3C490B1FD21" box="[391,490,703,725]" country="Australia" name="Tasmania" pageId="4" pageNumber="95">Tasmania</collectingRegion>
) and western Spencer Gulf,
<collectingRegion id="8B11F82E990D7168FF6EE3A79072FD06" box="[138,297,732,754]" country="Australia" name="South Australia" pageId="4" pageNumber="95">South Australia</collectingRegion>
, has a larger and more rounded test (up to
<quantity id="8E2D9B29990D7168FD0CE3A791EAFCFB" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="95" unit="mm" value="80.0">80 mm</quantity>
long) and, similar to
<taxonomicName id="8ED54D4F990D7168FE6FE38290A4FCFB" ID-CoL="6YV7F" authorityName="H. L. Clark" authorityYear="1923" box="[395,511,761,783]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="7BA1EADE990D7168FE6FE38290A4FCFB" box="[395,511,761,783]" italics="true" pageId="4" pageNumber="95">S. mirabilis</emphasis>
</taxonomicName>
, more posteriorly located apical system and greater area enclosed by peripetalous fasciole.
</paragraph>
<paragraph id="496A36CC990D7168FF4EE22B91B6FBE7" blockId="4.[138,769,298,1043]" pageId="4" pageNumber="95">
Compared with
<taxonomicName id="8ED54D4F990D7168FEBBE22B934BFC92" ID-CoL="4YFGS" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[351,528,848,870]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="dermodyorum">
<emphasis id="7BA1EADE990D7168FEBBE22B934BFC92" box="[351,528,848,870]" italics="true" pageId="4" pageNumber="95">S. dermodyorum</emphasis>
</taxonomicName>
, it also has a much wider and longer plastron and narrower adoral ambulacra I and
<collectionCode id="2FC4AE09990D7168FF32E2F191B3FC54" box="[214,232,906,928]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="95" type="Museum">V</collectionCode>
.
<bibRefCitation id="2D444B3D990D7168FF1EE2F190B8FC54" author="Miskelly, A." box="[250,483,906,928]" pageId="4" pageNumber="95" refId="ref5749" refString="Miskelly, A. 2002. Sea urchins of Australia and the Indo-Pacific. Capricornia Publications: Sydney. 179 pp." type="book" year="2002">Miskelly (2002: 156)</bibRefCitation>
noted two pairs of pore pairs occur in each side of the subanal fasciole, a feature also recorded for
<taxonomicName id="8ED54D4F990D7168FEFEE2BF939FFC2D" authority="(McNamara et al., 1986: 80)" baseAuthorityName="McNamara" baseAuthorityPageNumber="80" baseAuthorityYear="1986" box="[282,708,963,986]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="laubei">
<emphasis id="7BA1EADE990D7168FEFEE2BF9022FC2E" box="[282,377,964,986]" italics="true" pageId="4" pageNumber="95">S. laubei</emphasis>
(
<bibRefCitation id="2D444B3D990D7168FE69E2B893E7FC2D" author="McNamara, K. J. &amp; Philip, G. M. &amp; Kruse, P. D." box="[397,700,963,985]" pageId="4" pageNumber="95" pagination="55 - 84" refId="ref5674" refString="McNamara, K. J., Philip, G. M., and Kruse, P. D. 1986. Tertiary brissid echinoids of southern Australia. Alcheringa 10: 55 - 84." type="journal article" year="1986">McNamara et al., 1986: 80</bibRefCitation>
)
</taxonomicName>
. This contrasts with the three pairs found on
<taxonomicName id="8ED54D4F990D7168FDB6E29A925BFC03" ID-CoL="4YFGS" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[594,768,993,1015]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="95" phylum="Echinodermata" rank="species" species="dermodyorum">
<emphasis id="7BA1EADE990D7168FDB6E29A925BFC03" box="[594,768,993,1015]" italics="true" pageId="4" pageNumber="95">S. dermodyorum</emphasis>
</taxonomicName>
(
<figureCitation id="D1EE2A49990D7168FF76E28691BAFBE7" box="[146,225,1021,1043]" captionStart="Figure 7" captionStartId="8.[138,196,768,788]" captionTargetBox="[138,1440,295,742]" captionTargetId="figure-11@8.[138,1441,295,743]" captionTargetPageId="8" captionText="Figure 7. Spatagobrissus dermodyorum sp. nov. A, detail of petals and apical disk of holotype, NMV P312570; B, peristome and labrum tuberculation of paratype, NMV P312571; C, subanal fasciole and pore pairs of paratype, NMV P312573. Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.10665601" httpUri="https://zenodo.org/record/10665601/files/figure.png" pageId="4" pageNumber="95">Fig. 7C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,86 @@
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<heading id="122281A0990A716FFF66E431908DFA94" box="[130,470,1354,1376]" fontSize="9" level="2" pageId="3" pageNumber="94" reason="5">
<taxonomicName id="8ED54D4F990A716FFF66E431908DFA94" ID-CoL="87KNW" authority="H. L. Clark, 1923" authorityName="H. L. Clark" authorityYear="1923" box="[130,470,1354,1376]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="genus">
<emphasis id="7BA1EADE990A716FFF66E4319046FA94" bold="true" box="[130,285,1354,1376]" italics="true" pageId="3" pageNumber="94">Spatagobrissus</emphasis>
H. L.
<bibRefCitation id="2D444B3D990A716FFE84E431908DFA94" author="Clark, H. L." box="[352,470,1354,1376]" pageId="3" pageNumber="94" pagination="221 - 435" refId="ref5090" refString="Clark, H. L. 1923. The echinoderm fauna of South Africa. Annals of the South African Museum 113: 221 - 435." type="journal article" year="1923">Clark, 1923</bibRefCitation>
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<subSubSection id="01CF6547990A716FFF66E40E9354FA27" pageId="3" pageNumber="94" type="materials_examined">
<paragraph id="496A36CC990A716FFF66E40E900AFA5C" blockId="3.[130,761,1397,1448]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFF66E40E9052FA7F" box="[130,265,1397,1419]" italics="true" pageId="3" pageNumber="94">
<typeStatus id="966E886E990A716FFF66E40E91EAFA7F" box="[130,177,1397,1419]" pageId="3" pageNumber="94">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8ED54D4F990A716FFEF6E40E9389FA7F" authority="H. L. Clark, 1923" authorityName="H. L. Clark" authorityYear="1923" box="[274,722,1397,1419]" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="7BA1EADE990A716FFEF6E40E9356FA7F" box="[274,525,1397,1419]" italics="true" pageId="3" pageNumber="94">Spatagobrissus mirabilis</emphasis>
H. L.
<bibRefCitation id="2D444B3D990A716FFDB3E40E9389FA7F" author="Clark, H. L." box="[599,722,1397,1419]" pageId="3" pageNumber="94" pagination="221 - 435" refId="ref5090" refString="Clark, H. L. 1923. The echinoderm fauna of South Africa. Annals of the South African Museum 113: 221 - 435." type="journal article" year="1923">Clark, 1923</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="496A36CC990A716FFF66E4C59354FA27" blockId="3.[130,527,1469,1492]" box="[130,527,1469,1492]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFF66E4C591BCFA20" box="[130,231,1470,1492]" italics="true" pageId="3" pageNumber="94">Diagnosis</emphasis>
. See H. L.
<bibRefCitation id="2D444B3D990A716FFEBFE4C69354FA27" author="Clark, H. L." box="[347,527,1469,1491]" pageId="3" pageNumber="94" pagination="221 - 435" refId="ref5090" refString="Clark, H. L. 1923. The echinoderm fauna of South Africa. Annals of the South African Museum 113: 221 - 435." type="journal article" year="1923">Clark (1923: 402)</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,170 @@
<document id="6C94C12E5774BB9DA357D055B0213682" ID-DOI="10.24199/j.mmv.2005.62.3" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665587" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708004635701" checkinUser="felipe" docAuthor="Holmes, Francis C., Yee, Christopher Ah &amp; Krause, Janice" docDate="2005" docId="C17C87DA990B716FFF6EE65C9347FB18" docLanguage="en" docName="MemMusVictoria.62.1.91-99.pdf" docOrigin="Memoirs of Museum Victoria 62 (1)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-62-issue-1-2005/pages-91-99/" docStyle="DocumentStyle:99484BB52C5EA01CDE4EA5EECB5782C2.2:MemMusVictoria.2003-.journal_article.type2" docStyleId="99484BB52C5EA01CDE4EA5EECB5782C2" docStyleName="MemMusVictoria.2003-.journal_article.type2" docStyleVersion="2" docTitle="Murraypneustes biannulatus Holmes &amp; Yee &amp; Krause 2005" docType="treatment" docVersion="2" lastPageNumber="94" masterDocId="3D45FFA29909716CFFE4E17B915BFFF4" masterDocTitle="Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia" masterLastPageNumber="99" masterPageNumber="91" pageNumber="93" updateTime="1708004957363" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B4DE5FF251AFA19CC4A0D958E61B18B8">Two new Middle Miocene spatangoids (Echinoidea) from the Murray Basin, South Australia</mods:title>
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<mods:namePart id="1F331D9FDFF7C324F75295F5460A6A91">Holmes, Francis C.</mods:namePart>
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<mods:namePart id="15660EE71A4A6BEB4CB894F9A0D5A902">Yee, Christopher Ah</mods:namePart>
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<mods:namePart id="A36D09B48EBB37265105565C3985973E">Krause, Janice</mods:namePart>
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<subSubSection id="01CF6547990B716EFF6EE65C934BF8C8" box="[138,528,1830,1852]" pageId="2" pageNumber="93" type="nomenclature">
<paragraph id="496A36CC990B716EFF6EE65C934BF8C8" blockId="2.[138,528,1830,1852]" box="[138,528,1830,1852]" pageId="2" pageNumber="93">
<heading id="122281A0990B716EFF6EE65C934BF8C8" box="[138,528,1830,1852]" fontSize="9" level="2" pageId="2" pageNumber="93" reason="5">
<taxonomicName id="8ED54D4F990B716EFF6EE65C90EEF8C8" ID-CoL="44N7H" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[138,437,1830,1852]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="2" pageNumber="93" phylum="Echinodermata" rank="species" species="biannulatus">
<emphasis id="7BA1EADE990B716EFF6EE65C90EEF8C8" bold="true" box="[138,437,1830,1852]" italics="true" pageId="2" pageNumber="93">Murraypneustes biannulatus</emphasis>
</taxonomicName>
. sp. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01CF6547990B716FFF6EE62A9347FB18" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93" type="description">
<paragraph id="496A36CC990B716EFF6EE62A90E5F893" blockId="2.[138,446,1873,1895]" box="[138,446,1873,1895]" pageId="2" pageNumber="93">
<figureCitation id="D1EE2A49990B716EFF6EE62A904EF893" box="[138,277,1873,1895]" captionStart="Figure 2" captionStartId="6.[138,196,1066,1086]" captionTargetBox="[136,1440,295,1040]" captionTargetId="figure-11@6.[136,1440,295,1040]" captionTargetPageId="6" captionText="Figure 2. Murraypneustes biannulatus gen et sp. nov. AD, adapical, posterior, adoral and left lateral views of holotype NMV P312370; E, adapical view of paratype NMV P312371; F, left lateral view of paratype, NMV P312372. All from the early Middle Miocene Glenforslan Formation in the vicinity of NMV locality PL3203, north of Blanchetown, South Australia." figureDoi="http://doi.org/10.5281/zenodo.10665591" httpUri="https://zenodo.org/record/10665591/files/figure.png" pageId="2" pageNumber="93">Figures 2AF</figureCitation>
,
<figureCitation id="D1EE2A49990B716EFEC4E62A9005F893" box="[288,350,1873,1895]" captionStart="Figure 3" captionStartId="6.[138,196,1752,1772]" captionTargetBox="[136,1440,1338,1726]" captionTargetId="figure-71@6.[136,1441,1338,1726]" captionTargetPageId="6" captionText="Figure 3. Fasciole details of Murraypneustes biannulatus gen. et sp. nov. A, B, oblique lateral view and ambulacrum IV detail above margin of paratype, NMV P312371; C, oblique posterior view of paratype, NMV P312372. Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10665593" httpUri="https://zenodo.org/record/10665593/files/figure.png" pageId="2" pageNumber="93">3AC</figureCitation>
,
<figureCitation id="D1EE2A49990B716EFE8DE62A90FDF893" box="[361,422,1873,1895]" captionStart="Figure 4" captionStartId="7.[130,188,746,766]" captionTargetBox="[131,747,297,718]" captionTargetId="figure-15@7.[131,761,297,718]" captionTargetPageId="7" captionText="Figure 4. Apical disk details of Murraypneustes biannulatus gen. et sp. nov. A, holotype NMV P312370; B, drawing of paratype, NMV P312372, showing extent of hydropores (black circles), tubercles (white circles), and gonopores and oculars (black or stippled). Location of posterior oculars (marked *) is assumed. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10665597" httpUri="https://zenodo.org/record/10665597/files/figure.png" pageId="2" pageNumber="93">4A, B</figureCitation>
,
<figureCitation id="D1EE2A49990B716EFE56E62A90E5F893" box="[434,446,1873,1895]" captionStart="Figure 5" captionStartId="7.[801,859,649,669]" captionTargetBox="[798,1431,297,621]" captionTargetId="figure-69@7.[798,1432,297,621]" captionTargetPageId="7" captionText="Figure 5. Murraypneustes biannulatus gen. et sp. nov. Detail of pore pairs and tubercles in and adjacent to ambulacrum V of paratype, NMV P312372." figureDoi="http://doi.org/10.5281/zenodo.10665595" httpUri="https://zenodo.org/record/10665595/files/figure.png" pageId="2" pageNumber="93">5</figureCitation>
</paragraph>
<paragraph id="496A36CC990B716EFF6FE600925AF85D" blockId="2.[138,769,1915,1961]" pageId="2" pageNumber="93">
<emphasis id="7BA1EADE990B716EFF6FE600905CF87A" box="[139,263,1915,1934]" italics="true" pageId="2" pageNumber="93">Type material</emphasis>
.
<materialsCitation id="F9BD3C91990B716EFEF7E600925AF85D" collectionCode="NMV" pageId="2" pageNumber="93" specimenCode="P312370" specimenCount="1" typeStatus="holotype">
<typeStatus id="966E886E990B716EFEF7E6009033F87B" box="[275,360,1915,1935]" pageId="2" pageNumber="93" type="holotype">Holotype</typeStatus>
,
<collectionCode id="2FC4AE09990B716EFE97E60090FCF87A" box="[371,423,1915,1934]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="93" type="Museum">NMV</collectionCode>
<specimenCode id="19739EB7990B716EFE4AE60090A6F87B" box="[430,509,1915,1935]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="93" type="Museum">P312370</specimenCode>
from early Middle Miocene Glenforslan Formation (Batesfordian), Morgan Group,
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NNE of
</materialsCitation>
</paragraph>
<paragraph id="496A36CC990B716EFCC2E0509220FEAC" blockId="2.[806,1437,298,371]" pageId="2" pageNumber="93">
Murray River Lock 1, Blanchetown,
<collectingRegion id="8B11F82E990B716EFB64E0519455FECA" box="[1152,1294,298,318]" country="Australia" name="South Australia" pageId="2" pageNumber="93">South Australia</collectingRegion>
[
<collectionCode id="2FC4AE09990B716EFAFFE0509409FECA" box="[1307,1362,299,318]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="93" type="Museum">NMV</collectionCode>
locality
<collectionCode id="2FC4AE09990B716EFCC2E03F9218FEAC" box="[806,835,324,344]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="93" type="Museum">PL</collectionCode>
3203].
</paragraph>
<paragraph id="496A36CC990B716EFCA5E02494DEFE87" blockId="2.[806,1437,298,371]" box="[833,1413,351,371]" pageId="2" pageNumber="93">
<materialsCitation id="F9BD3C91990B716EFCA5E02494DAFE87" box="[833,1409,351,371]" collectionCode="NMV" pageId="2" pageNumber="93" specimenCode="P312371" specimenCount="1" typeStatus="paratype">
<typeStatus id="966E886E990B716EFCA5E02492C1FE86" box="[833,922,351,370]" pageId="2" pageNumber="93" type="paratype">Paratypes</typeStatus>
,
<collectionCode id="2FC4AE09990B716EFC47E024928CFE86" box="[931,983,351,370]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="93" type="Museum">NMV</collectionCode>
<specimenCode id="19739EB7990B716EFC39E0249577FE87" box="[989,1068,351,371]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="93" type="Museum">P312371</specimenCode>
and P312372 from the same location
</materialsCitation>
.
</paragraph>
<paragraph id="496A36CC990B716EFCC2E0F29544FE6A" blockId="2.[806,1055,392,415]" box="[806,1055,392,415]" pageId="2" pageNumber="93">
<emphasis id="7BA1EADE990B716EFCC2E0F292D0FE6B" box="[806,907,393,415]" italics="true" pageId="2" pageNumber="93">Diagnosis</emphasis>
. As for genus.
</paragraph>
<paragraph id="496A36CC990B716EFCC2E0CF9226FD83" blockId="2.[806,1438,436,1962]" pageId="2" pageNumber="93">
<emphasis id="7BA1EADE990B716EFCC2E0CF92C7FE3E" box="[806,924,436,458]" italics="true" pageId="2" pageNumber="93">Description</emphasis>
. Test moderately large, ovoid in outline with shallow anterior sulcus and pointed, slightly truncated posterior. Specimens range from
<quantity id="8E2D9B29990B716EFBF5E09595CEFDF0" box="[1041,1173,494,516]" metricMagnitude="-2" metricUnit="m" metricValue="7.65" metricValueMax="8.1" metricValueMin="7.2" pageId="2" pageNumber="93" unit="mm" value="76.5" valueMax="81.0" valueMin="72.0">72 to 81 mm</quantity>
in length, with maximum width 8186%TL occurring at 45%TL from anterior ambitus. Test 44.1%TL high (uncompressed specimen) with apex of all specimens well posterior of centre, 6367%TL from anterior ambitus.
</paragraph>
<paragraph id="496A36CC990B716EFCA2E30592AFFCAB" blockId="2.[806,1438,436,1962]" pageId="2" pageNumber="93">Centre of test on adapical surface depressed around apical disk and proximal end of paired petals to form minor apices in interambulacra 1, 4 and 5, and conjointly, a raised area across ambulacrum III and interambulcra 2 and 3. Adoral surface mildly concave in vicinity of peristome with ambulacrum III slightly recessed anteriorly and the plastron forming a fairly pronounced keel posteriorly, terminating at the anterior edge of the subanal fasciole.</paragraph>
<paragraph id="496A36CC990B716EFCA2E21D92B2FB00" blockId="2.[806,1438,436,1962]" pageId="2" pageNumber="93">
Primary tubercles on adapical surface of 2 distinct sizes, both widely and randomly spaced. Larger of the two restricted to the distal 35% of the radius on interambulacrum 5, and 25% elsewhere. Adorally, larger primary tubercles closely and evenly spaced throughout, with exception of naked areas in ambulacra I and
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and phyllode plates of ambulacra II, III and IV. Overlapping scrobicules on adoral surface form distinct diagonal ridges. Small tubercles closely spaced immediately below ambitus increase in size to that of larger primary tubercles below curvature of margin. Large primary tubercles, perforate with undercut mamelon and crenulated platform, and maximum scrobicular diameter of approximately
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, twice size of smaller counterparts. Ring of scrobicular tubercles not always present.
</paragraph>
<paragraph id="496A36CC990B716EFCA2E58194C6FA0C" blockId="2.[806,1438,436,1962]" pageId="2" pageNumber="93">
Peripetalous and subanal fascioles present, the former, though not continuously visible on any specimen clearly passes above periproct, while latter is distinctly angular, forming hexagonal outline where in contact with lower edge of periproct. Intermittent horizontal fasciole bands also present laterally above and parallel to marginal peripetalous fasciole. Uppermost very narrow and more continuous fasciole, although clear of distal ends of petals, appears to be a rudimentary peripetalus fasciole not indented interradially (
<figureCitation id="D1EE2A49990B716EFAB7E49994CBFA0C" box="[1363,1424,1506,1528]" captionStart="Figure 3" captionStartId="6.[138,196,1752,1772]" captionTargetBox="[136,1440,1338,1726]" captionTargetId="figure-71@6.[136,1441,1338,1726]" captionTargetPageId="6" captionText="Figure 3. Fasciole details of Murraypneustes biannulatus gen. et sp. nov. A, B, oblique lateral view and ambulacrum IV detail above margin of paratype, NMV P312371; C, oblique posterior view of paratype, NMV P312372. Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10665593" httpUri="https://zenodo.org/record/10665593/files/figure.png" pageId="2" pageNumber="93">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="496A36CC990B716EFCA2E48492A1F908" blockId="2.[806,1438,436,1962]" pageId="2" pageNumber="93">
<materialsCitation id="F9BD3C91990B716EFCA2E48492ADF908" collectionCode="V" pageId="2" pageNumber="93" specimenCount="1">
Apical system anterior of centre, 39.542.2%TL from anterior ambitus to centre of disk, level or slightly below proximal end of paired petals. Ethmolytic, with 4 small closely spaced gonopores approximately
<quantity id="8E2D9B29990B716EFBD3E72E95D2F99F" box="[1079,1161,1621,1643]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="93" unit="mm" value="0.3">0.3 mm</quantity>
in diameter, anterior pair closer together than posterior pair. Detail of ocular plates indeterminate. Hydropores numerous, approximately 70 visible in
<specimenCount id="5FD3FD45990B716EFC79E7D79568F936" box="[925,1075,1708,1730]" count="1" pageId="2" pageNumber="93" type="generic">one specimen</specimenCount>
, centrally located but extending between posterior pair of gonopores and possibly ocular plates I and
<collectionCode id="2FC4AE09990B716EFC84E79D9229F908" box="[864,882,1766,1788]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="93" type="Museum">V</collectionCode>
(
<figureCitation id="D1EE2A49990B716EFC64E79D92B5F908" box="[896,1006,1766,1788]" captionStart="Figure 4" captionStartId="7.[130,188,746,766]" captionTargetBox="[131,747,297,718]" captionTargetId="figure-15@7.[131,761,297,718]" captionTargetPageId="7" captionText="Figure 4. Apical disk details of Murraypneustes biannulatus gen. et sp. nov. A, holotype NMV P312370; B, drawing of paratype, NMV P312372, showing extent of hydropores (black circles), tubercles (white circles), and gonopores and oculars (black or stippled). Location of posterior oculars (marked *) is assumed. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10665597" httpUri="https://zenodo.org/record/10665597/files/figure.png" pageId="2" pageNumber="93">Fig. 4A, B</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="496A36CC990B716FFCA2E67893FDFD4C" blockId="2.[806,1438,436,1962]" lastBlockId="3.[130,761,298,1188]" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
Petals lanceolate, moderately wide at midlength, closed distally, situated in gentle concave depressions incorporating adradial edges of adjacent interambulacra and continuing proximally across apical disk. Anterior paired petals shorter than posterior pair, extending on average 60% of the radius measured along the surface of the perradial suture from centre of apical disk to ambitus; posterior pair about 56% radius. Inner pores of petals oval, outer pores slot-like, slightly curved and 50% wider. Pore pairs not conjugate but linked by a fine ridge which extends along both sides of each pore (
<figureCitation id="D1EE2A49990A716FFDA6E0FA9325FE63" box="[578,638,385,407]" captionStart="Figure 5" captionStartId="7.[801,859,649,669]" captionTargetBox="[798,1431,297,621]" captionTargetId="figure-69@7.[798,1432,297,621]" captionTargetPageId="7" captionText="Figure 5. Murraypneustes biannulatus gen. et sp. nov. Detail of pore pairs and tubercles in and adjacent to ambulacrum V of paratype, NMV P312372." figureDoi="http://doi.org/10.5281/zenodo.10665595" httpUri="https://zenodo.org/record/10665595/files/figure.png" pageId="3" pageNumber="94">Fig. 5</figureCitation>
). Maximum width of interporiferous zone slightly more than twice width of poriferous zone. Anterior paired petals diverge at approximately 135° and contain on average 23 pore pairs, posterior petals 297° and 26 pairs. Secondary tubercles extend randomly across interporiferous and poriferous zones and for a distance outside petals without primary tubercles. Ambulacrum III not petaloid, basically flush with adjoining interambulacra for about 50% radius, then gradually becoming concave towards anterior sulcus. Other details unknown, no sign of pores or regularly spaced tubercles being visible on specimens.
</paragraph>
<paragraph id="496A36CC990A716FFF46E3C490C7FCD8" blockId="3.[130,761,298,1188]" pageId="3" pageNumber="94">Peristome reniform, centre situated 2730%TL from anterior ambitus, longitudinal dimension approximately 6.5%TL, transverse dimension 12%TL. Phyllodes short and not particularly well developed.</paragraph>
<paragraph id="496A36CC990A716FFF46E24890B0FC48" blockId="3.[130,761,298,1188]" pageId="3" pageNumber="94">
Labrum long and narrow, averaging 20.5%TL, slightly curved at junction with peristome and abutting the plastron at centre of third pair of adjacent ambulacral plates. Numerous small tubercles adjacent to peristome with a few larger ones towards the posterior end (
<figureCitation id="D1EE2A49990A716FFE6BE2DD9084FC48" box="[399,479,934,956]" captionStart="Figure 2" captionStartId="6.[138,196,1066,1086]" captionTargetBox="[136,1440,295,1040]" captionTargetId="figure-11@6.[136,1440,295,1040]" captionTargetPageId="6" captionText="Figure 2. Murraypneustes biannulatus gen et sp. nov. AD, adapical, posterior, adoral and left lateral views of holotype NMV P312370; E, adapical view of paratype NMV P312371; F, left lateral view of paratype, NMV P312372. All from the early Middle Miocene Glenforslan Formation in the vicinity of NMV locality PL3203, north of Blanchetown, South Australia." figureDoi="http://doi.org/10.5281/zenodo.10665591" httpUri="https://zenodo.org/record/10665591/files/figure.png" pageId="3" pageNumber="94">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="496A36CC990A716FFF46E2B89070FBB9" blockId="3.[130,761,298,1188]" pageId="3" pageNumber="94">
Plastron closely tuberculate, width approximately 75% length measured from posterior edge of labrum to anterior edge of subanal fasciole. Strong posterior taper suggests sixth and subsequent plates of ambulacra I and
<collectionCode id="2FC4AE09990A716FFDEFE5619346FBC4" box="[523,541,1050,1072]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="94" type="Museum">V</collectionCode>
indent behind paired episternal plates.
</paragraph>
<paragraph id="496A36CC990A716FFF46E52F90B4FB50" blockId="3.[130,761,298,1188]" pageId="3" pageNumber="94">Periproct opening marginal, not visible from above, tear shaped, slightly wider than high, set in a slight truncation approximately 65° to the horizontal.</paragraph>
<paragraph id="496A36CC990A716FFF66E5C29347FB18" blockId="3.[130,761,1209,1260]" pageId="3" pageNumber="94">
<emphasis id="7BA1EADE990A716FFF66E5C29035FB3B" box="[130,366,1209,1231]" italics="true" pageId="3" pageNumber="94">
Etymology.
<taxonomicName id="8ED54D4F990A716FFF1CE5C29035FB3B" authorityName="Holmes &amp; Yee &amp; Krause" authorityYear="2005" box="[248,366,1209,1231]" class="Echinoidea" family="Maretiidae" genus="Murraypneustes" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="94" phylum="Echinodermata" rank="species" species="biannulatus">biannulatus</taxonomicName>
</emphasis>
(L) two-ringed, referring to the presence of two peripetalous fasciole rings.
</paragraph>
</subSubSection>
</treatment>
</document>