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<document ID-CLB-Dataset="292499" ID-GBIF-Dataset="be1b0f9f-6964-4184-9e92-5243a5c744fd" checkinTime="1706626311646" checkinUser="plazi" docAuthor="Info Flora" docDate="2021" docId="C1521005503C6D96E9AF60181064DCC8" docLanguage="de" docName="InfoFlora.Asteraceae.xml" docOrigin="https://www.infoflora.ch/de/flora/asteraceae.html (accessed 2023-10-20)" docSource="https://www.infoflora.ch/de/flora/asteraceae.html" docTitle="Calendula officinalis L." docType="treatment" docVersion="10" id="858B0CC8E7E221B7CA651A71EE7C56E3" masterDocId="726559447A88CFA27342DA99178CB9AD" masterDocTitle="Info Flora Schweiz - Asteraceae" updateTime="1712196430618" updateUser="ExternalLinkService">
<mods:mods id="B2B7A9FD2AF7F380DE8E1EF76FAD4A74" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BFC6A8E5E8F48EA0CB047B724899BBDF">
<mods:title id="EA6713490180CB4D09767F480241248A">Info Flora Schweiz - Asteraceae</mods:title>
</mods:titleInfo>
<mods:name id="D9A8B8ED4DA78ECDDCCF5A0FBD69D35A" type="personal">
<mods:role id="473BD66F2209D52FCC58EB18AC474A01">
<mods:roleTerm id="038895E40111E1268AFCD165B486D5CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08E6CE89DBFA3597734112DC65D18B50">Info Flora</mods:namePart>
</mods:name>
<mods:typeOfResource id="C9B6F121692FD0F57DE947F062A135C6">text</mods:typeOfResource>
<mods:originInfo id="DB6B67E996E13DEEB34AE63A9D3A5A02">
<mods:dateIssued id="9EAFF22568DB4FFC39CB59B8570FA47F">2021</mods:dateIssued>
<mods:dateCaptured id="8B3FACB7A0EECDD44E2419BDA4FBF8D9">2023-10-20</mods:dateCaptured>
<mods:publisher id="54288A476988DED4A445CBB05700E0AB">Info Flora Schweiz</mods:publisher>
<mods:place id="2F7DF6E3DA38CBDBEA02D32C868746AF">
<mods:placeTerm id="2AB7442E964B873D66EC6434BA3F4297">Geneve</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:location id="9B4A446EF5BE7AAFFEEF4968832C3903">
<mods:url id="692D2C2621214D642FA61594B272BA73">https://www.infoflora.ch/de/flora/asteraceae.html</mods:url>
</mods:location>
<mods:classification id="27596B02493AAFB8FA7B64E7CBF55B3F">url</mods:classification>
</mods:mods>
<treatment id="C1521005503C6D96E9AF60181064DCC8" ID-DOI="http://doi.org/10.5281/zenodo.10915582" ID-GBIF-Taxon="224774677" ID-Zenodo-Dep="10915582" LSID="urn:lsid:plazi:treatment:C1521005503C6D96E9AF60181064DCC8" httpUri="http://treatment.plazi.org/id/C1521005503C6D96E9AF60181064DCC8">
<subSubSection id="5D83D51EF1066BCA4FDFAE26D1439126" type="nomenclature">
<paragraph id="393794034B522A24DBE93499F0BFC349">
<taxonomicName id="44CDCC62CE28A039FCEAE34F43C68541" ID-CoL="PJDZ" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="1004C9C605D930D1C23E355EDAF47FA8" italics="true">Calendula officinalis</emphasis>
L.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="40BABA479C3A9471BF767957E7B8105C" type="vernacular_names">
<paragraph id="0CB4DA9B94958F3DE9416C8B26F29BE3">
<vernacularName id="704F4F947B743233925BF7FA8B79793D">Garten-Ringelblume</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="685A7C6548E2912EBFCC48E30D1E5D1C" type="multiple">
<paragraph id="4984841D41D0E841E410120EB3170EB6">
Art ISFS: 72500 Checklist: 1008040
<taxonomicName id="6C162C3F3BB02441B0FE7E422AF87DB0" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
<taxonomicName id="C6DF85654378A3A0186E3F987F8805E0" class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="genus">Calendula</taxonomicName>
<taxonomicName id="C0969653CFA3CDF0C4073284CD31CAD5" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</paragraph>
<paragraph id="7313C4EC52D9D1A544B83ED123D09A69">
<heading id="D818DFE53B5970F1FAB17BE8201EB2D4">
<normalizedToken id="DBA27B34295647DA3EEE211F29081DF5" originalValue="Bestimmungsschlüssel">Bestimmungsschluessel</normalizedToken>
</heading>
</paragraph>
<paragraph id="1BEAC88F6CCE5EDEA7AC75E3A79510D5">
<heading id="338A3478A9A32EBB5953C901024AE395">Zusammenfassung</heading>
</paragraph>
</subSubSection>
<subSubSection id="C67CD7B4D6EA734035A571649F22C341" type="description">
<paragraph id="2EE93C1BACC54C3D634DFDF4B17E344F">
Artbeschreibung
<emphasis id="92E70B29FA3434E01818059323B2D9B0" italics="true">
(nach
<bibRefCitation id="60B3FBC44AB54B32C8737B437E0EB9BD" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="AB00BC0BB9A23A72DEEE05FF478E6775" originalValue="Ähnlich">Aehnlich</normalizedToken>
wie
<taxonomicName id="A361A337CF9FDC7EE6F29ED5FE7F0BC0" class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="7CEFAED2CC904F97A5DAA34171CF47BC" italics="true">C. arvensis</emphasis>
</taxonomicName>
, aber
<normalizedToken id="55ADADDBCED7CF1FF6289CE6F5E64E44" originalValue="Stängel">Staengel</normalizedToken>
aufsteigend bis aufrecht,
<normalizedToken id="614A838FDEE4172D587C3B14AA1F6033" originalValue="Blätter">Blaetter</normalizedToken>
<normalizedToken id="74491AF5045B7C0A168C5DBFC64F0C18" originalValue="eiförmig">eifoermig</normalizedToken>
bis lanzettlich, die unteren
<normalizedToken id="3AACFB65304327F984479CDFB1F62F54" originalValue="allmählich">allmaehlich</normalizedToken>
in einen kurzen,
<normalizedToken id="D9A16D9DF61022DC63D46A5278FCBBE1" originalValue="geflügelten">gefluegelten</normalizedToken>
Stiel
<normalizedToken id="77DEC832C37FE09CB12BC7B60C24B33A" originalValue="verschmälert">verschmaelert</normalizedToken>
,
<emphasis id="2C9FC95B8591145E22648CDC9D8A9C19" bold="true">
Kopfdurchmesser
<quantity id="2179DB6E6F186FEA7A0866CC53D4F60E" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">2-5 cm</quantity>
, Zungen- und
<normalizedToken id="4772C087E3ABC7DE60FD0A6E8100D322" originalValue="Röhrenblüten">Roehrenblueten</normalizedToken>
dunkelgelb bis orange
</emphasis>
, zur Fruchtzeit aufrecht (bei
<taxonomicName id="DC11693D5FECCE84DDD9915B83E2CD04" class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="1F58A950967C385EA7B09A6D7DBE0074" italics="true">C. arvensis</emphasis>
</taxonomicName>
zur Fruchtzeit nickend).
</paragraph>
</subSubSection>
<subSubSection id="07DF14B6DF06A6D682B092FF0F9F96F3" type="biology_ecology">
<paragraph id="76718E7E833526918D1B810B654AABDB">
<normalizedToken id="66F7FA5C6B90685DE372C187EDF4077A" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="E08D10E72C7E1CB44AE271B045113FC5" italics="true">
(nach
<bibRefCitation id="F2FD8DCE79F1034AA14EA534F7F04BBE" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: 6-8
</paragraph>
<paragraph id="E40CCE9796835132AA711E5306F97BAB">
Standort und Verbreitung in der Schweiz
<emphasis id="CB5A7FA8690109727208F8753BB015B5" italics="true">
(nach
<bibRefCitation id="3FAF24AC81A0C526E2FF98CFAEE58B89" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="64FF20997E6F319363FB440F24B05B11" originalValue="Schuttplätze">Schuttplaetze</normalizedToken>
, kultiviert und gelegentlich verwildert / kollin-montan /
</paragraph>
</subSubSection>
<subSubSection id="9E8260E4ECB33BA841D8EB293616316E" type="distribution">
<paragraph id="64A5F6AEFB4379F370E31CC09A84D4B2">
Verbreitung global
<emphasis id="42B9B7E81014E6F4B5EA8BC74575AE1B" italics="true">
(nach
<bibRefCitation id="CC9279D19986674E9870474E76F08BB5" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Mediterran
</paragraph>
</subSubSection>
<subSubSection id="7097953C422CE85F9795929D4FB955BA" type="biology_ecology">
<paragraph id="66E02C7B040F1D16FB914811F4600110">
<normalizedToken id="A26019F276B7002B45BED452CC7BFDAA" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte
<emphasis id="9550859A19724C0DB19522ECF5D0BCCB" italics="true">
(nach
<bibRefCitation id="CE41DF0549A20A21712A48509527B059" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. 2010</bibRefCitation>
)
</emphasis>
2 + 34-44 + 3.t.2n=28,32,36
</paragraph>
</subSubSection>
<subSubSection id="EF9760A90B13342E4FFB50C2490D51C5" type="multiple">
<paragraph id="47495073C69424092F1A5ED3A4D15439">
<heading id="5936E902A96136EA5E162F5244612310">
<normalizedToken id="025E35FF5E179AF12C282BE0BBF532C4" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="77D335616E32B6C22D727091609C77E0">Lebensform Therophyt</paragraph>
<paragraph id="48393EB3B965FC4D98748C3DA9128C96">
<heading id="B3170895163E4124CFF54E78B086AB10">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="1D65A40D5DDDE7E5A602A951372DA79B" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie Gefährdung Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie Gefährdung Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="270D6C6EDAC93FAD24CA884156BA833A">
<table id="E94539E351167F99B26E8100033B0451" class="table" inLine="true">
<tbody id="8719F1428FF18F365962804C7F1F2C38">
<tr id="320A1BDD9C69ADB635166E1A5AD70063">
<td id="25187FFF2E9A91FE74DB5679B96BA71F">KEINE ANGABE</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="1C078A666046DCC7C92630F27B52A2BB" type="biology_ecology">
<paragraph id="EDE1CDF53E8225F6E0D966D7F77FB9E5">
<normalizedToken id="0534D55E6A470406F15FCD66700350AD" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="191EAFFFE9BE2225333C468522E8827E" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="00CD5A1C7BC81B446686A838BF03E4DD">
<table id="3C95EB0CA5234B2BA6A8B865A8242AF5" class="table" inLine="true">
<tbody id="C1267933B535DD65918E03D07723493A">
<tr id="4B88B986B5845DE4820FE8C996FA80AF">
<th id="E8D08FABD52AF06B059B113022426E74" colspan="2">
<emphasis id="08BD9AF407AA1D26E588A2B16EC559A7" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="66BC4BD33D0AD20014929F7706F07EC8" colspan="2">
<emphasis id="50F6347CCF18D19791554CA72B660523" bold="true">Klimafaktoren</emphasis>
</th>
<th id="595F16C87E3049E0D6E1F75D58F29FE7" colspan="2">
<emphasis id="48FEA8A99DF4B5E1C4B01C13B99E9A04" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="41D727A9E0CA7F0FFF91C79AD9B030E5">
<td id="582F9B5687120C32AD61040673CBAFA2">Feuchtezahl F</td>
<td id="90181586955A0CE7159A474FA981B148" style="text-align:left;width:100px">frisch</td>
<td id="187E56DB1B23DA152113247188084E43">Lichtzahl L</td>
<td id="D11CFCFF81D5B4F9EB7CF3564AE3BE22" style="text-align:left;width:60px">hell</td>
<td id="A513C764520C8799996FBED668B52420">Salzzeichen</td>
<td id="08158E99DB688AC06E1D8B8E188ED246" style="text-align:left;width:60px">--</td>
</tr>
<tr id="F055417CB15EDED4ABFAF56BA85E4C02">
<td id="FBA79AC8FDE6B62D23E63517AF6FB68C">Reaktionszahl R</td>
<td id="0A8D48D840EC28B476B7D9B2E2053273" style="text-align:left">schwach sauer bis neutral (pH 4.5-7.5)</td>
<td id="8020F0A631D52A00F52AE6830D88CF2C">Temperaturzahl T</td>
<td id="81FECC441E946426BA908F858EDA6681" style="text-align:left">warm-kollin</td>
</tr>
<tr id="07BDA46A521CEAFD16CF82FF66B7B32B">
<td id="45888BD4C21DD4909C7E8992EA4B6312">
<normalizedToken id="01E612888260C4ED30394965F054BADC" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="B2CC9E3F424064A046782453457D9DD8" style="text-align:left">
<normalizedToken id="D27C348B66F88A79157D7C5124C67277" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="B8D94A722BD69A71E12824706B2E68CA">
<normalizedToken id="6A241C9F8F8D3CD6AED1681D3FEB1DC4" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="3858AA56BC2619FF9F5FDD9FF4EE7339" style="text-align:left">
subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
<normalizedToken id="C893ACD5055DB845E64A2BDBF56FB9D5" originalValue="mässige">maessige</normalizedToken>
Temperaturschwankungen und
<normalizedToken id="0A3CCBCC8F5BF2F4D44D06F1B2CCF77B" originalValue="mässig">maessig</normalizedToken>
tiefe Wintertemperaturen)
</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="F1E2DF152E21C26CB4D5EA38A1FF29DB" type="reference_group">
<paragraph id="55F944D9141A36C34D888196865AE223">
<heading id="1FEB32C674979778A778475F78697C0D">Nomenklatur</heading>
</paragraph>
<paragraph id="203C700A553F942169C69714521C7FD2">
<treatmentCitation id="17CC2F4AB703C244599CDFE70A1DF367">
<heading id="389D152C67AD532075D858C86B99739E">
<normalizedToken id="3B2DC608EABEF4C6A56773F3DE597E9D" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="161DA9149093E6B5D6446A7F15C4E373" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="D26159774159D27283006BFDD679C4CB" ID-CoL="PJDZ" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="012C096822A65A8C95C2E3D9D4515002" italics="true">Calendula officinalis</emphasis>
L.
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F7B15860608EB7A6170495622C0B002" type="vernacular_names">
<paragraph id="831639A0B34878DB8C2D942430D13874">
Volksname Deutscher Name:
<vernacularName id="D9737865ADC017B8B369D7A1EF91E912" language="deu">Garten-Ringelblume</vernacularName>
Nom
<normalizedToken id="131482438786C76C59D761EACF39997D" originalValue="français">francais</normalizedToken>
:
<vernacularName id="457F6F7107D7EE71008645C87918F777" language="fra">Souci des jardins</vernacularName>
Nome italiano:
<vernacularName id="54E29B6176410B62ED5A7879EDD54371" language="ita">Fiorrancio coltivato</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="2AF42F3E4A580BCD59931F6641F07CB9" type="reference_group">
<paragraph id="3CA75024F5EC93B3C3174546CCA4DBB3">
<normalizedToken id="B4B9F52C7E0A41CCE3AAF14C3F850DDB" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="A544209B1D47A5E86153864DC786A494">
<table id="F6BF96AA3818300EA39163050A468CE5" class="table table-condensed" inLine="true">
<tbody id="673233C38238D82D591DF7EC8CC133C3">
<tr id="D07BB33852619F742263EA40D2CC2F7A">
<th id="18E00EB7C7D055BC84F0672D50FAE44F" width="80px">Relation</th>
<th id="A5679E1293693E8AC4747D3E1CD69F85">Nom</th>
<th id="E191C61BEC684A947E222D852183113F">Referenzwerke</th>
<th id="7485549C76EF9BD3EB83E9208136E300" width="90px">No</th>
</tr>
<tr id="07C49EBFA8EEF38B82105B06B61A65BF">
<td id="241E526C72500FAF4010140B6302CFDF" class="taxon-relation">=</td>
<td id="B53E7432FE98E12CA6779C6D6931507D">
<taxonomicName id="21D46DEDEE0CAFFC4929E74D0C86E2E7" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="A85FEA108EA17805F627551B8D589F94">
<treatmentCitation id="1F5983263581119B7CBE6A2E2D1DC785">
<bibRefCitation id="75A37D49EAE4A26A1A9021783CDDB63E" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
</treatmentCitation>
</td>
<td id="70E25AF29E981FF7259BC19650F59D57">72500</td>
</tr>
<tr id="343FFB312EB6F547FBDF6329BD758DE8">
<td id="F15038A1AA764FEF2A20FE6A549CD993" class="taxon-relation">=</td>
<td id="61B4AB3819013437698BFAE5F71FCD46">
<taxonomicName id="8FF0B1BAE5541DCF088FE09B8898573E" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="345154CBBD5A6404237C8C3DCF0E8309">
<treatmentCitation id="A379CD6188887EC59308392E4A405E63">
<bibRefCitation id="6DAA84C8AFF9C8C350250DD38913666C">Flora Helvetica 2001</bibRefCitation>
</treatmentCitation>
</td>
<td id="9100008370F47FC076E6F2AF9FFDD8F6">2190</td>
</tr>
<tr id="D5D2AB54B7916088C57EFD9DEB03CF7A">
<td id="0C404F8BC3937A0CB467D9DD860725C6" class="taxon-relation">=</td>
<td id="C72AFEDF20054DA1E572D09C5CEF66ED">
<taxonomicName id="29B986910077F7F5EDD8DAD2473A0302" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="FBC53A10EC4EC8BDACFE6C5ECD385AEA">
<treatmentCitation id="E6D392447A77988F55D26FB4C9C614B7">
<bibRefCitation id="386AA2EC34A973976FAD179439C6DC32">Flora Helvetica 2012</bibRefCitation>
</treatmentCitation>
</td>
<td id="551FBB5CBCBFE115A19C6B6450B0865C">2182</td>
</tr>
<tr id="CFBCEB320B7BE193F1B87FF21177A4E3">
<td id="6D7B676F4EBA5BE0F82D5FD95D31FC64" class="taxon-relation">=</td>
<td id="CF6437A3289E5E320D6CC930D907EADB">
<taxonomicName id="1DACDA3A9AD69E4947D3A935C953717E" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="9B69C8AFE280F50414485E894C7F2D45">
<treatmentCitation id="523E83E7A35CCC99D6AFC1B23F7C5DFC" httpUri="http://treatment.plazi.org/id/EDDDD5F3E23A41D6CEAFC26814DFE656">
<bibRefCitation id="393AC8C8F703DA9B63FB0705D140E4A8">Flora Helvetica 2018</bibRefCitation>
</treatmentCitation>
</td>
<td id="AFF0A6109054FFA6793B58A213D473DC">2182</td>
</tr>
<tr id="25210C9AFDEF2BACFE1DC59E2F0C4716">
<td id="98AA197DC67D213B3B7EED1879CCB76F" class="taxon-relation">=</td>
<td id="EAE6533E99168967B22FC805129B73D4">
<taxonomicName id="D7219015BE7CBAEDC8E7F378E9159B26" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="B0BF1D4E7B17172BFFDC22BC3A978026">
<treatmentCitation id="B7D7B79B21645CB0A20C14533389437F">
<bibRefCitation id="3C1942666F7A2DF4093AB45CF4DA366C">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="DADFA7FB45AF66573C51CD1E584159A8">72500</td>
</tr>
<tr id="52F0EF1A7F3BF3E256C95E170B4F1AEC">
<td id="2BAC000F3B0996FB5C9F66FA054A7655" class="taxon-relation">=</td>
<td id="231255337F5FA16A2ABA49502FBB9EBE">
<taxonomicName id="E6E34DA1DBAC745B930A9E8DB98B5FFD" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="AD23C05DCCA83B53351C1E25CCD162BA">
<treatmentCitation id="3E747987AEDEF1BE044E548498CABFD3">
<bibRefCitation id="419E7B9DF2FF90ED38713FB82FE71299">Landolt 1977</bibRefCitation>
</treatmentCitation>
</td>
<td id="4D8B6C93C65CF3D87707B6493CBAA361">3063</td>
</tr>
<tr id="2A48C4EFB437DE395F6428EB35573F05">
<td id="A79D4B0307BC8C06EAF49F2E51EBE51F" class="taxon-relation">=</td>
<td id="FC21A390B910A8360F714FACF50F6972">
<taxonomicName id="9A9C3DDB9BB543CC9255412B61A15FC0" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Calendula" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="officinalis">Calendula officinalis L.</taxonomicName>
</td>
<td id="E9A900751B22721C52E9BBAC29A3F6D8">
<treatmentCitation id="511D6DE985E7159A0F2B426FC70A1A54">
<bibRefCitation id="248071D2622F6AFD44B80F2F512190B9">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="7A66E374645B128E55DC59BA730C3429">72500</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="B9BB381C2DECC9268675AD17BF5959F5">
<tableNote id="F9CE3D32AE5B3DCE659E87307C09DBE8">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="2AF68EF95ED62EDAA07143378505D0A1" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="DE2E6B90957CBF7AD816810BA8D73AD2" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="7082D6F53836CF196F894D991C1EB143" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="309AA668DCD61D7F8FEB04FF20E78CCB" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="D1972C07B92F2A0E0C63BBBBF8C3C3FD" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="2D14FE93CF14FA7BBAC8A3356311D771" type="conservation">
<paragraph id="F3B437063152BB32109A405F3DC09C64">
<heading id="293AB79971C497D13BC395D258E90CF3">Status Indigenat</heading>
: Kultivierte Pflanze, vor dem Jahr
<quantity id="779AA17CA1176C4FC80E6EFABD182C66" metricMagnitude="1" metricUnit="m" metricValue="3.81" unit="in" value="1500.0">1500 in</quantity>
der Schweiz aufgetreten
</paragraph>
<paragraph id="5DDAC88F54595777040976B163A33AFE">
<emphasis id="9153074ABE413C41736C4382C7DB4F74" bold="true">
<heading id="99C27EC8F39E3B1D61198306A716B428">
Liste der
<normalizedToken id="4DD73BB344EF920B9F973FC637C34A8A" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="1A334820155C939ACF06804641CF4C60" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,113 @@
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<mods:title id="6135655BAC727D4F9C7673AB9B13BB5A">Flora der Schweiz und angrenzender Gebiete. Band 2. Nymphaceae bis Primulaceae (2 nd edition) (p. 956): Papaveraceae</mods:title>
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<mods:namePart id="173D881BCE5BC7A1C0D1683370CEACA9">Hess, Hans Ernst</mods:namePart>
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<mods:namePart id="0F08EF28C457F7536FF9D665658FC9A7">Landolt, Elias</mods:namePart>
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</mods:role>
<mods:namePart id="798FB12D121E624AC25D1BC6A8896ED3">Hirzel, Rosmarie</mods:namePart>
</mods:name>
<mods:typeOfResource id="E2C995B792E9467510AECFCF38E86856">text</mods:typeOfResource>
<mods:originInfo id="3C06720081FB6F8DD469CC9D7B9B724E">
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<mods:location id="27F8FBBEB35E99BFD246C85C7C31986F">
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<treatment id="C152451BDAC024BFE11D1E1DC94AAE0A" LSID="urn:lsid:plazi:treatment:C152451BDAC024BFE11D1E1DC94AAE0A" httpUri="http://treatment.plazi.org/id/C152451BDAC024BFE11D1E1DC94AAE0A" lastPageNumber="114" pageId="null" pageNumber="114" sec="papaveraceae.s19">
<title id="8B897913E4C9815DA83D706E8C556B16" pageId="null" pageNumber="114">
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<paragraph id="7DEDABD42F382DEC155678CA2E63AE92" pageId="null" pageNumber="114">
<taxonomicName id="98B577A961144B533B806A4250C67414" authority="L." class="Magnoliopsida" family="Papaveraceae" genus="Papaver" kingdom="Plantae" order="Ranunculales" pageId="null" pageNumber="114" phylum="Tracheophyta" rank="species" species="argemone">
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<authorityName id="3FAC29B48570A822EF66F1C10A3377FE" pageId="null" pageNumber="114">L.</authorityName>
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</paragraph>
</subSubSection>
<subSubSection id="69FC299DA68FC243AA9C4ED334C0535F" pageId="null" pageNumber="114" type="vernacular_names">
<paragraph id="7DC7F9BC46D51C0C9C5C7E8E404EE7CD" pageId="null" pageNumber="114">Sand-Mohn</paragraph>
</subSubSection>
</title>
<subSubSection id="A5014BB315DC3319F63AAD942047A561" pageId="null" pageNumber="114" type="description">
<paragraph id="F88194F2C5C9A72E1EDA3D020541548D" pageId="null" pageNumber="114">
<emphasis id="7991EC9D2D796458D08F3F58598CC5AD" italics="true" pageId="null" pageNumber="114">Stengel mit 1,5-3 mm langen Haaren. Kelch zerstreut mit 1,5-3 mm langen Haaren</emphasis>
besetzt oder fast kahl.
<emphasis id="B50D437317DAB28C23F785D4EBA6D117" italics="true" pageId="null" pageNumber="114">
Frucht
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, 1,5-2 cm lang
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und 0,4-0,5 cm dick, mit einzelnen 1,5-3 mm langen und am Grund 0,2 mm dicken Haaren. Samen 0,9-1,1 mm lang. -
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:
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und
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Sommer.
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<pageBreakToken id="0CF018B8991DEB5A8B258AD4042011FC" pageId="null" pageNumber="114">,</pageBreakToken>
Kawatani und Ohno aus
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und
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1961), aus England (McNaughton 1960). Koopmans (aus
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und
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1961)
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2n = 40.
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<paragraph id="B1D00B3694B3F280B386F6FBE201A679" pageId="null" pageNumber="114">
<emphasis id="57393271013389AC03F67F46503CE6BF" italics="true" pageId="null" pageNumber="114">Standort.</emphasis>
Kollin bis subalpin. Lockere, kalkarme,
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, sandige
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in warmen Lagen. Getreidefelder (besonders Roggenfelder),
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.
<emphasis id="0DAD88863D7542FD0989C29A92602A31" italics="true" pageId="null" pageNumber="114">Papaveretum Argentone</emphasis>
(Libh.) Krusem. et Vlieg. (1939).
</paragraph>
<paragraph id="78C6B5B4F8F496CBFE81D048C88DD550" pageId="null" pageNumber="114">
<emphasis id="0986ADAC79036AAD95EDE1DED749CC99" italics="true" pageId="null" pageNumber="114">Verbreitung. Mediterrane Pflanze:</emphasis>
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verschleppt bis
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und
<normalizedToken id="ABE5984AD4768B249BC278604D199300" originalValue="Mittelrußland" pageId="null" pageNumber="114">Mittelrussland</normalizedToken>
; Westasien (
<normalizedToken id="7C7AB87209B8409FE8AB3BB817A3DB5A" originalValue="ostwärts" pageId="null" pageNumber="114">ostwaerts</normalizedToken>
bis Westpersien); Nordwestafrika. - Im Gebiet zerstreut (besonders in den
<normalizedToken id="48DE474946C4258EDE16330E87A98C50" originalValue="wärmeren" pageId="null" pageNumber="114">waermeren</normalizedToken>
Gebieten, aus dem Tessin jedoch keine sicheren Angaben), selten und oft
<normalizedToken id="5463C58A8DC036E47A9152F379FAFAEC" originalValue="unbeständig" pageId="null" pageNumber="114">unbestaendig</normalizedToken>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,177 @@
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?
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<subSubSection id="01E16563FFB2710584F8FD431CDBF9B3" pageId="5" type="materials_examined">
<paragraph id="494436E8FFB2710584F8FD431937FD0A" blockId="5.[145,761,657,740]" pageId="5">
<emphasis id="7B8FEAFAFFB2710584F8FD431964FD49" box="[145,369,657,679]" italics="true" pageId="5">Referred specimens:</emphasis>
<collectionCode id="2FEAAE2DFFB271058517FD4019AEFD49" box="[382,443,658,679]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198, upper left or lower right canine with complete crown (
<figureCitation id="D1C02A6DFFB271058676FD621A4AFD2B" box="[543,607,688,710]" captionStart="Figure 4" captionStartId="5.[146,229,1817,1839]" captionTargetBox="[150,756,1243,1777]" captionTargetId="figure-744@5.[149,757,1242,1778]" captionTargetPageId="5" captionText="Figure 4.?Phocidae, gen. &amp; sp. indet., NMV P16198, isolated upper left or lower right canine tooth, whitened with ammonium chloride. A, lingual view. B, labial view." figureDoi="http://doi.org/10.5281/zenodo.5300487" httpUri="https://zenodo.org/record/5300487/files/figure.png" pageId="5">Fig. 4</figureCitation>
), collected by J. M. Wilson.
</paragraph>
<paragraph id="494436E8FFB2710584F8FCDC1D87FC16" blockId="5.[145,761,781,1202]" lastBlockId="5.[809,1426,197,1629]" pageId="5">
<emphasis id="7B8FEAFAFFB2710584F8FCDC1938FCCD" box="[145,301,782,803]" italics="true" pageId="5">Description:</emphasis>
The tooth
<collectionCode id="2FEAAE2DFFB2710585A3FCDC1A1EFCCD" box="[458,523,782,803]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198 is nearly complete, with only the distal tip of the root missing (
<figureCitation id="D1C02A6DFFB2710584F3FC9818CAFC8E" box="[154,223,842,864]" captionStart="Figure 4" captionStartId="5.[146,229,1817,1839]" captionTargetBox="[150,756,1243,1777]" captionTargetId="figure-744@5.[149,757,1242,1778]" captionTargetPageId="5" captionText="Figure 4.?Phocidae, gen. &amp; sp. indet., NMV P16198, isolated upper left or lower right canine tooth, whitened with ammonium chloride. A, lingual view. B, labial view." figureDoi="http://doi.org/10.5281/zenodo.5300487" httpUri="https://zenodo.org/record/5300487/files/figure.png" pageId="5">Fig. 4</figureCitation>
).
<collectionCode id="2FEAAE2DFFB271058490FC99192DFC8E" box="[249,312,843,864]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198 was originally identified as a squalodontid cetacean tooth (
<bibRefCitation id="2D6A4B19FFB271058665FCBB1A94FC91" author="Gill E" box="[524,641,873,895]" pageId="5" pagination="135 - 203" refId="ref11366" refString="Gill E. 1957. The stratigraphical occurrence and palaeoecology of some Australian Tertiary marsupials. Memoirs of the National Museum of Victoria 21: 135 - 203." type="journal article" year="1957">Gill, 1957</bibRefCitation>
), but was subsequently identified as a pinniped, possibly representing a phocid incisor (
<bibRefCitation id="2D6A4B19FFB27105859BFC741A85FC52" author="Fordyce RE" box="[498,656,934,956]" pageId="5" pagination="43 - 58" refId="ref11153" refString="Fordyce RE. 1982. A review of Australian fossil Cetacea. Memoirs of the National Museum of Victoria 43: 43 - 58." type="journal article" year="1982">Fordyce, 1982</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFB2710586F7FC741965FC34" author="Fordyce RE &amp; Flannery T" pageId="5" pagination="99 - 100" refId="ref11204" refString="Fordyce RE, Flannery T. 1983. Fossil phocid seals from the late Tertiary of Victoria. Proceedings of the Royal Society of Victoria 95: 99 - 100." type="journal article" year="1983">Fordyce &amp; Flannery, 1983</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFB2710585EDFC141A4BFC34" author="Long JA &amp; Archer M &amp; Flannery T &amp; Hand S" box="[388,606,965,987]" pageId="5" refId="ref11679" refString="Long JA, Archer M, Flannery T, Hand S. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution. Sydney: JHU Press." type="book" year="2002">
Long
<emphasis id="7B8FEAFAFFB2710585A7FC141A12FC34" box="[462,519,965,987]" italics="true" pageId="5">et al</emphasis>
., 2002
</bibRefCitation>
). While the narrowing at the base of the crown is reminiscent of incisors,
<collectionCode id="2FEAAE2DFFB27105849FFBD11926FBF6" box="[246,307,1027,1048]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198 is interpreted here as a canine based on a relatively thick root, as opposed to the tapering, mesiodistally flattened nature of incisor roots. This thickening of the root is only present in the derived caniform incisors of Leopard seals,
<taxonomicName id="8EFB4D6BFFB2710586EEFBAF19F8FB5C" authority="(Blainville, 1820)" baseAuthorityName="Blainville" baseAuthorityYear="1820" class="Mammalia" family="Phocidae" genus="Hydrurga" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="leptonyx">
<emphasis id="7B8FEAFAFFB2710586EEFBAF18EEFB5F" italics="true" pageId="5">Hydrurga leptonyx</emphasis>
(Blainville, 1820)
</taxonomicName>
, which have a more exaggerated narrowing at the base of the crown, and is not present on the incisors of any other pinniped. The enamelocementum boundary allows for orientation, with a straight profile characteristic of the labial aspect of a canine and an angular enamelocementum boundary profile characteristic of the lingual aspect (
<figureCitation id="D1C02A6DFFB271058758FEAF1B7AFE7C" box="[817,879,381,403]" captionStart="Figure 4" captionStartId="5.[146,229,1817,1839]" captionTargetBox="[150,756,1243,1777]" captionTargetId="figure-744@5.[149,757,1242,1778]" captionTargetPageId="5" captionText="Figure 4.?Phocidae, gen. &amp; sp. indet., NMV P16198, isolated upper left or lower right canine tooth, whitened with ammonium chloride. A, lingual view. B, labial view." figureDoi="http://doi.org/10.5281/zenodo.5300487" httpUri="https://zenodo.org/record/5300487/files/figure.png" pageId="5">Fig. 4</figureCitation>
). The cingulum is prominent on the lingual side, being slightly smaller on the labial side. Apicobasal ridges are concentrated on the labial side of the canine, being more prominent distally. This specimen is interpreted as either a lower right or an upper left canine, on the basis that the root is recurved towards the labial side, a ridge is present on the lingual side and there is a strong curvature along the tooth as a whole. While the ridge and the strong curvature are more common in mandibular canines, they are also variable between canine positions. Specimens of
<taxonomicName id="8EFB4D6BFFB271058740FD1C1C16FD0D" baseAuthorityName="Blainville" baseAuthorityYear="1820" box="[809,1027,718,739]" class="Mammalia" family="Phocidae" genus="Hydrurga" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="leptonyx">
<emphasis id="7B8FEAFAFFB271058740FD1C1C16FD0D" box="[809,1027,718,739]" italics="true" pageId="5">Hydrurga leptonyx</emphasis>
</taxonomicName>
(
<collectionCode id="2FEAAE2DFFB271058078FD1C1C45FD0D" box="[1041,1104,718,739]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C23614 and
<collectionCode id="2FEAAE2DFFB271058082FD1D1D3CFD0A" box="[1259,1321,719,740]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C31561) demonstrate that the lingual ridge can be present on the lingual surface of upper canines, as well as the lowers. Upper canines of Ross seal,
<taxonomicName id="8EFB4D6BFFB27105808AFCF81BE9FCB1" authority="(Gray, 1844)" baseAuthorityName="Gray" baseAuthorityYear="1844" class="Mammalia" family="Phocidae" genus="Ommatophoca" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="rossii">
<emphasis id="7B8FEAFAFFB27105808AFCF81B7DFCB1" italics="true" pageId="5">Ommatophoca rossii</emphasis>
(Gray, 1844)
</taxonomicName>
(
<collectionCode id="2FEAAE2DFFB271058062FC9B1C5FFCB0" box="[1035,1098,841,862]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C7354.1), and Weddell seal,
<taxonomicName id="8EFB4D6BFFB271058740FCBA1CFEFC90" authority="(Lesson, 1826)" baseAuthorityName="Lesson" baseAuthorityYear="1826" box="[809,1259,872,894]" class="Mammalia" family="Phocidae" genus="Leptonychotes" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="weddellii">
<emphasis id="7B8FEAFAFFB271058740FCBA1C29FC93" box="[809,1084,872,893]" italics="true" pageId="5">Leptonychotes weddellii</emphasis>
(Lesson, 1826)
</taxonomicName>
(
<collectionCode id="2FEAAE2DFFB271058093FCBA1D2CFC93" box="[1274,1337,872,893]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C7297,
<collectionCode id="2FEAAE2DFFB271058740FC551B87FC72" box="[809,914,903,924]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" type="Museum">NHMUK</collectionCode>
1951.5.1.4), also possess strong curvature, demonstrating that it is not exclusively a mandibular character. Hence these characters cannot be used to ascertain between the upper or lower canine positions.
</paragraph>
<paragraph id="494436E8FFB271058728FBD31CDBF9B3" blockId="5.[809,1426,197,1629]" pageId="5">
The relatively short, curved crown and the apiciobasal ridges (an aquatic adaptation;
<bibRefCitation id="2D6A4B19FFB27105814EFBF21BBDFBBD" author="McCurry MR &amp; Evans AR &amp; Fitzgerald EMG &amp; McHenry CR &amp; Bevitt J &amp; Pyenson ND" pageId="5" pagination="245 - 259" refId="ref11827" refString="McCurry MR, Evans AR, Fitzgerald EMG, McHenry CR, Bevitt J, Pyenson ND. 2019. The repeated evolution of dental apicobasal ridges in aquatic-feeding mammals and reptiles. Biological Journal of the Linnean Society 127: 245 - 259." type="journal article" year="2019">
McCurry
<emphasis id="7B8FEAFAFFB271058740FBED1B4EFBBD" box="[809,859,1086,1108]" italics="true" pageId="5">et al</emphasis>
., 2019
</bibRefCitation>
) rule out possible carnivorous marsupials or terrestrial placentals. The presence of strong apiciobasal ridges and the robust nature of the crown (as opposed to the slender nature of otariids) suggest
<collectionCode id="2FEAAE2DFFB271058740FB6B1B72FB20" box="[809,871,1209,1230]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198 may be a phocid. While some phocids do have strong apiciobasal ridges (such as
<taxonomicName id="8EFB4D6BFFB271058177FB0A1B84FAE5" baseAuthorityName="Blainville" baseAuthorityYear="1820" class="Mammalia" family="Phocidae" genus="Hydrurga" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="leptonyx">
<emphasis id="7B8FEAFAFFB271058177FB0A1B84FAE5" italics="true" pageId="5">Hydrurga leptonyx</emphasis>
</taxonomicName>
;
<collectionCode id="2FEAAE2DFFB2710587CDFB251BF1FAE2" box="[932,996,1271,1292]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C23614 and
<collectionCode id="2FEAAE2DFFB2710580FBFB251CC7FAE2" box="[1170,1234,1271,1292]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
C31561), such ridges are highly variable in pinnipeds, and so this feature cannot be used to confidently assign
<collectionCode id="2FEAAE2DFFB27105813BFAE61D85FAA7" box="[1362,1424,1332,1353]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P16198 to
<taxonomicName id="8EFB4D6BFFB2710587CFFA801C07FA86" box="[934,1042,1362,1384]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="family">Phocidae</taxonomicName>
. The relatively small size is only slightly larger than the size of canines in the phocine seals
<taxonomicName id="8EFB4D6BFFB271058704FA5D1C61FA4B" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[877,1140,1423,1445]" class="Mammalia" family="Phocidae" genus="Phoca" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFB271058704FA5D1BA1FA4A" box="[877,948,1423,1444]" italics="true" pageId="5">Phoca</emphasis>
Linnaeus, 1758
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFB2710580D8FA421D85FA4B" authority="Scopoli, 1771" authorityName="Scopoli" authorityYear="1771" box="[1201,1424,1423,1445]" class="Mammalia" family="Phocidae" genus="Pusa" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFB2710580D8FA421CFEFA4B" box="[1201,1259,1424,1445]" italics="true" pageId="5">Pusa</emphasis>
Scopoli, 1771
</taxonomicName>
(Supporting Information, Appendix S6). However, it should be noted that canines can be sexually dimorphic. The only monachine that approaches the size of these small seals is
<taxonomicName id="8EFB4D6BFFB2710587A7F9D81D14F9F1" box="[974,1281,1546,1567]" class="Mammalia" family="Phocidae" genus="Australophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="species" species="changorum">
<emphasis id="7B8FEAFAFFB2710587A7F9D81D14F9F1" box="[974,1281,1546,1567]" italics="true" pageId="5">Australophoca changorum</emphasis>
</taxonomicName>
Valenzuela- Toro
<emphasis id="7B8FEAFAFFB27105870AF9FB1B80F9D0" box="[867,917,1577,1598]" italics="true" pageId="5">et al</emphasis>
., 2016, for which no craniodental fossils are known (
<bibRefCitation id="2D6A4B19FFB2710587EDF9951CAAF9B3" author="Valenzuela-Toro AM &amp; Pyenson ND &amp; Gutstein CS &amp; Suarez ME" box="[900,1215,1607,1629]" pageId="5" pagination="101 - 115" refId="ref12703" refString="Valenzuela-Toro AM, Pyenson ND, Gutstein CS, Suarez ME. 2016. A new dwarf seal from the late Neogene of South America and the evolution of pinnipeds in the southern hemisphere. Papers in Palaeontology 2: 101 - 115." type="journal article" year="2016">
Valenzuela-Toro
<emphasis id="7B8FEAFAFFB27105802AF99A1C61F9B2" box="[1091,1140,1607,1629]" italics="true" pageId="5">et al</emphasis>
., 2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,314 @@
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<subSubSection id="01E16563FFB2710587C6F9401D1EF944" box="[943,1291,1682,1706]" pageId="5" type="nomenclature">
<paragraph id="494436E8FFB2710587C6F9401D1EF944" blockId="5.[943,1291,1682,1706]" box="[943,1291,1682,1706]" pageId="5">
<heading id="120C8184FFB2710587C6F9401D1EF944" box="[943,1291,1682,1706]" centered="true" fontSize="9" level="2" pageId="5" reason="3">
<taxonomicName id="8EFB4D6BFFB2710587C6F9401D1EF944" ID-CoL="625XL" authority="GEN. &amp; SP. INDET." box="[943,1291,1682,1706]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="family">
<emphasis id="7B8FEAFAFFB2710587C6F9401D1EF944" bold="true" box="[943,1291,1682,1706]" pageId="5">PHOCIDAE GEN. &amp; SP. INDET.</emphasis>
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</heading>
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</subSubSection>
<subSubSection id="01E16563FFB271078740F9081A5CFB56" lastPageId="7" pageId="5" type="materials_examined">
<paragraph id="494436E8FFB271068740F9081ADFFEF6" blockId="5.[809,1425,1754,1899]" lastBlockId="6.[163,779,197,280]" lastPageId="6" pageId="5">
<emphasis id="7B8FEAFAFFB271058740F9081C34F91E" box="[809,1057,1754,1776]" italics="true" pageId="5">Referred specimens:</emphasis>
<collectionCode id="2FEAAE2DFFB271058059F9091C65F91E" box="[1072,1136,1755,1776]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P42523, metatarsal
<collectionCode id="2FEAAE2DFFB271058112F9091D99F91E" box="[1403,1420,1755,1776]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" type="Museum">V</collectionCode>
, collected by J. Dixon;
<collectionCode id="2FEAAE2DFFB271058058F9281C65F8E1" box="[1073,1136,1786,1807]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P215759, metatarsal
<collectionCode id="2FEAAE2DFFB271058112F9281D99F8E1" box="[1403,1420,1786,1807]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" type="Museum">V</collectionCode>
, collected by D. Rawlings, 2002 (
<figureCitation id="D1C02A6DFFB2710580E6F8CA1CDAF8C0" box="[1167,1231,1816,1838]" captionStart="Figure 5" captionStartId="6.[163,241,1846,1868]" captionTargetBox="[324,1283,770,1805]" captionTargetId="figure-321@6.[323,1283,769,1807]" captionTargetPageId="6" captionText="Figure 5. Phocidae, gen. &amp; sp. indet., NMV P42523, right metatarsal V (A, B, C), and NMV P215759, left metatarsal V (D, E, F). A and D, lateral view. B and E, dorsal view. C and F, medial view." figureDoi="http://doi.org/10.5281/zenodo.5300489" httpUri="https://zenodo.org/record/5300489/files/figure.png" pageId="5">Fig. 5</figureCitation>
).
<collectionCode id="2FEAAE2DFFB27105808DF8CA1D34F8C3" box="[1252,1313,1816,1837]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P240711, partial interorbital region of cranium, collected by B. S. J. Francischelli, 2013 (
<figureCitation id="D1C02A6DFFB271058053F8871C6BF885" box="[1082,1150,1877,1899]" captionStart="Figure 6" captionStartId="7.[145,224,575,597]" captionTargetBox="[385,1185,195,534]" captionTargetId="figure-693@7.[385,1185,195,535]" captionTargetPageId="7" captionText="Figure 6. Phocidae, gen. &amp; sp. indet., NMV P246711, isolated interorbital region of cranium, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5300491" httpUri="https://zenodo.org/record/5300491/files/figure.png" pageId="5">Fig. 6</figureCitation>
).
<collectionCode id="2FEAAE2DFFB2710580FFF8841CC1F885" box="[1174,1236,1878,1899]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="5" type="Museum">NMV</collectionCode>
P41759 partial sacrum collected by C. Macrae, 1980 (
<figureCitation id="D1C02A6DFFB171068606FF171AA6FF35" box="[623,691,197,219]" captionStart="Figure 7" captionStartId="8.[164,242,1344,1366]" captionTargetBox="[325,1281,197,1303]" captionTargetId="figure-261@8.[323,1283,195,1305]" captionTargetPageId="8" captionText="Figure 7. Phocidae, gen. &amp; sp. indet., NMV P47759, isolated partial sacrum, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, anterior view. E, posterior view. F, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5300493" httpUri="https://zenodo.org/record/5300493/files/figure.png" pageId="6">Fig. 7</figureCitation>
).
<collectionCode id="2FEAAE2DFFB1710686A5FF141B1FFF35" box="[716,778,198,219]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
P160433 eight articulated thoracic vertebrae and five ribs, collected by
<collectionCode id="2FEAAE2DFFB17106850CFED11963FEF6" box="[357,374,259,280]" name="Tavera, Department of Geology and Geophysics" pageId="6">T</collectionCode>
. Flannery
<emphasis id="7B8FEAFAFFB171068587FED119ECFEF6" box="[494,505,259,280]" italics="true" pageId="6">c</emphasis>
. 1975 (
<figureCitation id="D1C02A6DFFB171068625FED01AAFFEF6" box="[588,698,258,280]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStartId-0="9.[144,222,1400,1422]" captionStartId-1="10.[162,240,1571,1593]" captionStartId-2="11.[145,223,1734,1756]" captionTargetBox-0="[385,1185,195,1360]" captionTargetBox-1="[405,1203,197,1530]" captionTargetBox-2="[226,1343,197,1692]" captionTargetId-0="figure-228@9.[385,1185,195,1360]" captionTargetId-1="figure-188@10.[403,1203,195,1531]" captionTargetId-2="figure-25@11.[225,1345,195,1694]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 8. Phocidae, gen. &amp; sp. indet., NMV P160433 articulated thoracic vertebrae and ribs in dorsal view." captionText-1="Figure 9. Phocidae, gen. &amp; sp. indet., NMV P160433, articulated thoracic vertebrae and ribs in right lateral view." captionText-2="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi-0="http://doi.org/10.5281/zenodo.5300495" figureDoi-1="http://doi.org/10.5281/zenodo.5300497" figureDoi-2="http://doi.org/10.5281/zenodo.5300499" httpUri-0="https://zenodo.org/record/5300495/files/figure.png" httpUri-1="https://zenodo.org/record/5300497/files/figure.png" httpUri-2="https://zenodo.org/record/5300499/files/figure.png" pageId="6">Figs 810</figureCitation>
).
</paragraph>
<caption id="1D846660FFB2710584FBF8CB1ACCF884" ID-DOI="http://doi.org/10.5281/zenodo.5300487" ID-Zenodo-Dep="5300487" httpUri="https://zenodo.org/record/5300487/files/figure.png" pageId="5" startId="5.[146,229,1817,1839]" targetBox="[150,756,1243,1777]" targetPageId="5">
<paragraph id="494436E8FFB2710584FBF8CB1ACCF884" blockId="5.[145,761,1817,1898]" pageId="5">
<emphasis id="7B8FEAFAFFB2710584FBF8CB1916F8C0" bold="true" box="[146,259,1817,1839]" pageId="5">Figure 4.</emphasis>
?
<taxonomicName id="8EFB4D6BFFB271058570F8CB1994F8C1" box="[281,385,1817,1839]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="5" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P16198, isolated upper left or lower right canine tooth, whitened with ammonium chloride. A, lingual view. B, labial view.
</paragraph>
</caption>
<paragraph id="494436E8FFB1710784CAFE901AB9FCE5" blockId="6.[163,780,321,711]" lastBlockId="7.[145,761,665,1882]" lastPageId="7" pageId="6">
<emphasis id="7B8FEAFAFFB1710684CAFE901957FEB9" box="[163,322,322,343]" italics="true" pageId="6">D e s c r i p t i o n:</emphasis>
N M
<collectionCode id="2FEAAE2DFFB1710685EBFE901981FEB9" box="[386,404,322,343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
P2 1 5 7 5 9 a n d N M
<collectionCode id="2FEAAE2DFFB1710686E5FE901A8BFEB9" box="[652,670,322,343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
P 4 2 5 2 3 (
<figureCitation id="D1C02A6DFFB1710684C5FEB218E3FE98" box="[172,246,352,374]" captionStart="Figure 5" captionStartId="6.[163,241,1846,1868]" captionTargetBox="[324,1283,770,1805]" captionTargetId="figure-321@6.[323,1283,769,1807]" captionTargetPageId="6" captionText="Figure 5. Phocidae, gen. &amp; sp. indet., NMV P42523, right metatarsal V (A, B, C), and NMV P215759, left metatarsal V (D, E, F). A and D, lateral view. B and E, dorsal view. C and F, medial view." figureDoi="http://doi.org/10.5281/zenodo.5300489" httpUri="https://zenodo.org/record/5300489/files/figure.png" pageId="6">Fig. 5</figureCitation>
) are both identified as fifth metatarsals due to the irregular, asymmetrical, transversely flattened proximal end.
<collectionCode id="2FEAAE2DFFB171068589FE4C1A35FE5D" box="[480,544,414,435]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
P215759 is a left metatarsal
<collectionCode id="2FEAAE2DFFB171068543FE6E1929FE3F" box="[298,316,444,465]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
and
<collectionCode id="2FEAAE2DFFB17106851CFE6E19A6FE3F" box="[373,435,444,465]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
P42523 is a right metatarsal
<collectionCode id="2FEAAE2DFFB1710684CAFE0918A0FE1E" box="[163,181,475,496]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
, both lacking the distal articular surfaces (
<figureCitation id="D1C02A6DFFB1710686D0FE081AE9FE1E" box="[697,764,474,496]" captionStart="Figure 5" captionStartId="6.[163,241,1846,1868]" captionTargetBox="[324,1283,770,1805]" captionTargetId="figure-321@6.[323,1283,769,1807]" captionTargetPageId="6" captionText="Figure 5. Phocidae, gen. &amp; sp. indet., NMV P42523, right metatarsal V (A, B, C), and NMV P215759, left metatarsal V (D, E, F). A and D, lateral view. B and E, dorsal view. C and F, medial view." figureDoi="http://doi.org/10.5281/zenodo.5300489" httpUri="https://zenodo.org/record/5300489/files/figure.png" pageId="6">Fig. 5</figureCitation>
). The absence of the distal articular surface may represent an immature condition. However, the majority of postcrania of mature phocids (based on cranial fusion and dental eruption) is usually unfused (pers. obsv.). Both specimens are identified as pinnipeds, based on a dorsoventrally expanded proximal end [e.g.
<taxonomicName id="8EFB4D6BFFB1710685EDFD631BE2FF35" authority="monachus (Hermann, 1779) NHMUK" authorityName="monachus (Hermann" authorityYear="1779" baseAuthorityName="Hermann" baseAuthorityYear="1779" box="[388,1015,197,710]" class="Mammalia" family="Phocidae" genus="Monachus" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="monachus">
<emphasis id="7B8FEAFAFFB1710685EDFD631A6AFD28" box="[388,639,689,710]" italics="true" pageId="6">Monachus monachus</emphasis>
(Hermann, 1779) NHMUK
</taxonomicName>
1958.11.26.1,
<taxonomicName id="8EFB4D6BFFB1710680DAFF171D3EFF35" box="[1203,1323,197,219]" class="Mammalia" family="Phocidae" genus="Phoca" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7B8FEAFAFFB1710680DAFF171CE8FF34" box="[1203,1277,197,218]" italics="true" pageId="6">Phoca</emphasis>
sp.
</taxonomicName>
<collectionCode id="2FEAAE2DFFB17106815FFF141DB4FF35" box="[1334,1441,198,219]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" type="Museum">NHMUK</collectionCode>
1951.3.2.3 and
<taxonomicName id="8EFB4D6BFFB171068794FF361CC2FF17" box="[1021,1239,228,249]" class="Mammalia" family="Otariidae" genus="Arctocephalus" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7B8FEAFAFFB171068794FF361CBEFF17" box="[1021,1195,228,249]" italics="true" pageId="6">Arctocephalus</emphasis>
sp.
</taxonomicName>
<collectionCode id="2FEAAE2DFFB17106808BFF361D37FF17" box="[1250,1314,228,249]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
C 33579]. Both
<collectionCode id="2FEAAE2DFFB171068715FED11BAEFEF6" box="[892,955,259,280]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
P215759 and
<collectionCode id="2FEAAE2DFFB171068000FED11CBDFEF6" box="[1129,1192,259,280]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
P42523 are referred to
<taxonomicName id="8EFB4D6BFFB171068731FEF31BD7FED9" box="[856,962,289,311]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="family">Phocidae</taxonomicName>
due to the shaft being more rounded (not dorsoventrally flattened) proximally approaching the base and having a plantar tubercle that (in dorsal view) is less than 90° proximal from the proximal distal plane. The lateral surface of the proximal end is mostly orientated laterally (similar to phocids, e.g.
<taxonomicName id="8EFB4D6BFFB171068703FE0B1CB9FE01" authority="NMV C" authorityName="NMV C" box="[874,1196,473,495]" class="Mammalia" family="Phocidae" genus="Hydrurga" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="leptonyx">
<emphasis id="7B8FEAFAFFB171068703FE0B1C5EFE00" box="[874,1099,473,494]" italics="true" pageId="6">Hydrurga leptonyx</emphasis>
NMV C
</taxonomicName>
23614 or
<taxonomicName id="8EFB4D6BFFB171068175FE0B1C48FDE3" authority=": Hendey 1972" authorityName=": Hendey" authorityYear="1972" class="Mammalia" family="Phocidae" genus="Homiphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="capensis">
<emphasis id="7B8FEAFAFFB171068175FE0B1BB0FDE3" italics="true" pageId="6">Homiphoca capensis</emphasis>
:
<bibRefCitation id="2D6A4B19FFB1710687DEFE2A1C48FDE3" author="Hendey QB &amp; Repenning CA" box="[951,1117,504,526]" pageId="6" pagination="79 - 98" refId="ref11500" refString="Hendey QB, Repenning CA. 1972. A Pliocene phocid from South Africa. Annals of the South African Museum 59: 79 - 98." type="journal article" year="1972">Hendey 1972</bibRefCitation>
</taxonomicName>
), thus differing from the otariid condition (
<taxonomicName id="8EFB4D6BFFB171068071FDC41CF8FDC2" box="[1048,1261,534,556]" class="Mammalia" family="Otariidae" genus="Arctocephalus" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7B8FEAFAFFB171068071FDC41CD7FDC5" box="[1048,1218,534,555]" italics="true" pageId="6">Arctocephalus</emphasis>
sp.
</taxonomicName>
<collectionCode id="2FEAAE2DFFB17106809FFDC51D20FDC2" box="[1270,1333,535,556]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
C33579) where the lateral surface of the base is deflected to face proximally. A slight depression on the medial surface of the base, directly medial to the plantar tubercle, is present in both specimens. In addition, there is a shallow fossa present on the lateral surface of the proximal end on the ventral side for both specimens.
<collectionCode id="2FEAAE2DFFB071078535FD6A198EFD23" box="[348,411,696,717]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="7" type="Museum">NMV</collectionCode>
P42523 is larger than
<collectionCode id="2FEAAE2DFFB0710786D3FD6A1AEDFD23" box="[698,760,696,717]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="7" type="Museum">NMV</collectionCode>
P215759 (Supporting Information, Appendix S2) and has a more ventrally developed proximal end.
</paragraph>
<caption id="1D846660FFB1710684CAF8E41B93F884" ID-DOI="http://doi.org/10.5281/zenodo.5300489" ID-Zenodo-Dep="5300489" httpUri="https://zenodo.org/record/5300489/files/figure.png" pageId="6" startId="6.[163,241,1846,1868]" targetBox="[324,1283,770,1805]" targetPageId="6">
<paragraph id="494436E8FFB1710684CAF8E41B93F884" blockId="6.[163,1443,1846,1898]" pageId="6">
<emphasis id="7B8FEAFAFFB1710684CAF8E4191EF8A2" bold="true" box="[163,267,1846,1868]" pageId="6">Figure 5.</emphasis>
<taxonomicName id="8EFB4D6BFFB17106857FF8E51963F8A3" box="[278,374,1847,1869]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="6" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P42523, right metatarsal V (A, B, C), and NMV P215759, left metatarsal V (D, E, F). A and D, lateral view. B and E, dorsal view. C and F, medial view.
</paragraph>
</caption>
<caption id="1D846660FFB0710784F8FDED1BC6FD9C" ID-DOI="http://doi.org/10.5281/zenodo.5300491" ID-Zenodo-Dep="5300491" httpUri="https://zenodo.org/record/5300491/files/figure.png" pageId="7" startId="7.[145,224,575,597]" targetBox="[385,1185,195,534]" targetPageId="7">
<paragraph id="494436E8FFB0710784F8FDED1BC6FD9C" blockId="7.[145,1425,575,626]" pageId="7">
<emphasis id="7B8FEAFAFFB0710784F8FDED18EEFDBA" bold="true" box="[145,251,575,597]" pageId="7">Figure 6.</emphasis>
<taxonomicName id="8EFB4D6BFFB07107856FFDED1972FDBB" box="[262,359,575,597]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="7" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P246711, isolated interorbital region of cranium, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, left lateral view.
</paragraph>
</caption>
<paragraph id="494436E8FFB0710784C0FCC61A5CFB56" blockId="7.[145,761,665,1882]" pageId="7">
<collectionCode id="2FEAAE2DFFB0710784C0FCC618FDFCC7" box="[169,232,788,809]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="7" type="Museum">NMV</collectionCode>
P240711 (
<figureCitation id="D1C02A6DFFB071078507FCC119A6FCC7" box="[366,435,787,809]" captionStart="Figure 6" captionStartId="7.[145,224,575,597]" captionTargetBox="[385,1185,195,534]" captionTargetId="figure-693@7.[385,1185,195,535]" captionTargetPageId="7" captionText="Figure 6. Phocidae, gen. &amp; sp. indet., NMV P246711, isolated interorbital region of cranium, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5300491" httpUri="https://zenodo.org/record/5300491/files/figure.png" pageId="7">Fig. 6</figureCitation>
) is the interorbital region of a phocid cranium. The nasal bones are inserted medially between the frontal bones, and the lack of a supraorbital process is diagnostic for
<taxonomicName id="8EFB4D6BFFB0710786E7FCBD192AFC4D" authority="(King, 1983 a)" baseAuthorityName="King" baseAuthorityYear="1983" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="7" phylum="Chordata" rank="family">
Phocidae (
<bibRefCitation id="2D6A4B19FFB0710784F3FC5C1926FC4D" author="King JE" box="[154,307,910,932]" pageId="7" refId="ref11560" refString="King JE. 1983 a. Seals of the world. London: Oxford University Press, 240." type="book" year="1983">King, 1983a</bibRefCitation>
)
</taxonomicName>
. The posterior extent of the nasal bone is preserved, along with an anterior section of the frontals. A small portion of cribriform plate and nasal turbinates of the ethmoid are preserved on the posteroventral surface; with a small section of the lateral interorbital region preserved on the dorsolateral surface of the frontal bone (
<figureCitation id="D1C02A6DFFB07107861AFB941AA0FBB2" box="[627,693,1094,1116]" captionStart="Figure 6" captionStartId="7.[145,224,575,597]" captionTargetBox="[385,1185,195,534]" captionTargetId="figure-693@7.[385,1185,195,535]" captionTargetPageId="7" captionText="Figure 6. Phocidae, gen. &amp; sp. indet., NMV P246711, isolated interorbital region of cranium, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5300491" httpUri="https://zenodo.org/record/5300491/files/figure.png" pageId="7">Fig. 6</figureCitation>
). The interorbital region must have been narrow relative to most other phocids (excluding phocins), based on the posterior tapering of the frontal bones.
</paragraph>
</subSubSection>
<subSubSection id="01E16563FFB0710984C0FB131A43F904" lastPageId="9" pageId="7" type="description">
<paragraph id="494436E8FFB0710784C0FB131A60F90E" blockId="7.[145,761,665,1882]" pageId="7">
NMV P41759 (
<figureCitation id="D1C02A6DFFB071078526FB131998FB38" box="[335,397,1217,1239]" captionStart="Figure 7" captionStartId="8.[164,242,1344,1366]" captionTargetBox="[325,1281,197,1303]" captionTargetId="figure-261@8.[323,1283,195,1305]" captionTargetPageId="8" captionText="Figure 7. Phocidae, gen. &amp; sp. indet., NMV P47759, isolated partial sacrum, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, anterior view. E, posterior view. F, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5300493" httpUri="https://zenodo.org/record/5300493/files/figure.png" pageId="7">Fig. 7</figureCitation>
) represents the anterior portion of the body of a sacrum, with partially incomplete sacral wings. The cranial articular surface and sacral crests are missing. Identified as a phocid due to sacral wings that are orientated dorsoventrally and laterally flared. Referral to
<taxonomicName id="8EFB4D6BFFB071078513FA8819C0FA9E" authorityName="Gray" authorityYear="1821" box="[378,469,1370,1392]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="7" phylum="Chordata" rank="tribe" tribe="Phocini">Phocini</taxonomicName>
can be excluded as the sacral wings are not orientated anteriorly [e.g.
<taxonomicName id="8EFB4D6BFFB0710786DAFAAA19EEFA43" authority="Linnaeus, 1758 NMV P" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Phocidae" genus="Phoca" kingdom="Animalia" order="Carnivora" pageId="7" phylum="Chordata" rank="species" species="vitulina">
<emphasis id="7B8FEAFAFFB0710786DAFAAA18F9FA42" italics="true" pageId="7">Phoca vitulina</emphasis>
Linnaeus, 1758 NMV P
</taxonomicName>
27683 and
<taxonomicName id="8EFB4D6BFFB071078613FA451A9BFA25" authority="(Erxleben, 1777) USNM" authorityName="USNM" baseAuthorityName="Erxleben" baseAuthorityYear="1777" class="Mammalia" family="Phocidae" genus="Pagophilus" kingdom="Animalia" order="Carnivora" pageId="7" phylum="Chordata" rank="species" species="groenlandicus">
<emphasis id="7B8FEAFAFFB071078613FA451952FA25" italics="true" pageId="7">Pagophilus groenlandicus</emphasis>
(Erxleben, 1777) USNM
</taxonomicName>
21532]. The dorsoventral orientation of the sacral wings is consistent across all other phocids. The promontory appears to be ovoid in shape (rather than spherical), but this may be due to the incomplete preservation of the dorsal surface. In dorsal view, the sacral foramina appear to be anteroposteriorly elongated.The auricular surface of the sacral wing appears to be concave and forms a fossa. This auricular fossa is present in most phocids and odobenids, but not in otariids.
</paragraph>
<paragraph id="494436E8FFB0710784C0F93B1B84FD02" blockId="7.[145,761,665,1882]" lastBlockId="7.[809,1425,664,1882]" pageId="7">
NMV P160433 (
<figureCitation id="D1C02A6DFFB071078537F93A19DEF910" box="[350,459,1768,1790]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStartId-0="9.[144,222,1400,1422]" captionStartId-1="10.[162,240,1571,1593]" captionStartId-2="11.[145,223,1734,1756]" captionTargetBox-0="[385,1185,195,1360]" captionTargetBox-1="[405,1203,197,1530]" captionTargetBox-2="[226,1343,197,1692]" captionTargetId-0="figure-228@9.[385,1185,195,1360]" captionTargetId-1="figure-188@10.[403,1203,195,1531]" captionTargetId-2="figure-25@11.[225,1345,195,1694]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 8. Phocidae, gen. &amp; sp. indet., NMV P160433 articulated thoracic vertebrae and ribs in dorsal view." captionText-1="Figure 9. Phocidae, gen. &amp; sp. indet., NMV P160433, articulated thoracic vertebrae and ribs in right lateral view." captionText-2="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi-0="http://doi.org/10.5281/zenodo.5300495" figureDoi-1="http://doi.org/10.5281/zenodo.5300497" figureDoi-2="http://doi.org/10.5281/zenodo.5300499" httpUri-0="https://zenodo.org/record/5300495/files/figure.png" httpUri-1="https://zenodo.org/record/5300497/files/figure.png" httpUri-2="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Figs 810</figureCitation>
) represents an articulated thoracic vertebral column, with eight thoracic vertebrae, four right and two left ribs preserved. The preserved thoracic vertebrae represent T7T14, based on the identification of the fifth vertebra from the anterior end of the column (T11) as the anticlinal vertebra.
</paragraph>
<paragraph id="494436E8FFB071078728FD261D69FB59" blockId="7.[809,1425,664,1882]" pageId="7">
Overall, the column is better preserved on the right lateral side, with the posterior vertebrae being the most complete. The column is eroded on the left posterolateral side. Most of T7 and T8 are missing. Both T9 and T12 are missing a large portion of the body on the left side. The spinous processes of all vertebrae, except T11 and T12, are not preserved, with T11 being the most complete. T11 (the anticlinal vertebrae) and T14 are the most complete vertebrae. While the entire column is articulated, it is contorted both mediolaterally and dorsoventrally relative to anatomical position. Descriptions and comparisons will be from the more complete right side and limited to pre-anticlinal (T7T10,
<figureCitation id="D1C02A6DFFB07107803BFB511CDAFB76" box="[1106,1231,1155,1177]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10AC</figureCitation>
), anticlinal (T11,
<figureCitation id="D1C02A6DFFB071078740FB701B9DFB59" box="[809,904,1186,1208]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10D</figureCitation>
) and post-anticlinal (T12T14,
<figureCitation id="D1C02A6DFFB071078099FB701D79FB59" box="[1264,1388,1186,1208]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10EF</figureCitation>
).
</paragraph>
<paragraph id="494436E8FFB071078728FB121CBAF98B" blockId="7.[809,1425,664,1882]" pageId="7">
The ribs of T9 and T10 are in articulation (
<figureCitation id="D1C02A6DFFB07107814DFB121D97FB3B" box="[1316,1410,1216,1238]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10A</figureCitation>
), with the heads situated within the anterior costal fovea and the posterior costal fovea (
<figureCitation id="D1C02A6DFFB07107808CFB2C1D5CFAFD" box="[1253,1353,1278,1300]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10C</figureCitation>
). Due to dorsal displacement of the ribs, the tubercles are not articulated with the costal fovea of the transverse process. The tubercle is not preserved on rib 9. The transverse processes of T8T10 are laterally separated from the anterior articular processes, supporting their identification as pre-anticlinal. On the transverse process, the mamillary process (preserved on T9) is clearly reduced, and the accessory process (preserved on T10) is underdeveloped, when contrasted to thoracic vertebrae later in the series. The posterior articular process on T7T10 faces ventrally.
</paragraph>
<paragraph id="494436E8FFB071088728F9BC1AA2F9EE" blockId="7.[809,1425,664,1882]" lastBlockId="8.[163,779,1453,1904]" lastPageId="8" pageId="7">
The anticlinal vertebra (T11) has an anterior articular process that contacts the anteromedial border of the mamillary process (
<figureCitation id="D1C02A6DFFB071078066F9791C65F92E" box="[1039,1136,1707,1729]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10B</figureCitation>
). The mamillary process projects anteriorly (
<figureCitation id="D1C02A6DFFB071078073F9181C95F931" box="[1050,1152,1738,1760]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="7">Fig. 10D</figureCitation>
). As the section of the transverse process posterior to the transverse fovea is not preserved, the condition of the accessory process is unknown.A rib is articulated to the anterior costal fovea, and the body of T10 appears to be missing a posterior costal fovea (
<figureCitation id="D1C02A6DFFBF7108852CFA7F19BFFA2C" box="[325,426,1453,1475]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="8">Fig. 10C</figureCitation>
). The anterior and posterior borders of the spinous process are nearly vertical. The posterior articular process faces ventrolaterally.
</paragraph>
<caption id="1D846660FFBF710884CDFA921C9FFA9A" ID-DOI="http://doi.org/10.5281/zenodo.5300493" ID-Zenodo-Dep="5300493" httpUri="https://zenodo.org/record/5300493/files/figure.png" pageId="8" startId="8.[164,242,1344,1366]" targetBox="[325,1281,197,1303]" targetPageId="8">
<paragraph id="494436E8FFBF710884CDFA921C9FFA9A" blockId="8.[163,1443,1344,1396]" pageId="8">
<emphasis id="7B8FEAFAFFBF710884CDFA921919FAB8" bold="true" box="[164,268,1344,1366]" pageId="8">Figure 7.</emphasis>
<taxonomicName id="8EFB4D6BFFBF7108857FFA921963FAB8" baseAuthorityName="King" baseAuthorityYear="1983" box="[278,374,1344,1366]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P47759, isolated partial sacrum, whitened with ammonium chloride. A, dorsal view. B, ventral view. C, right lateral view. D, anterior view. E, posterior view. F, left lateral view.
</paragraph>
</caption>
<paragraph id="494436E8FFBF710884D2F9DB1CCDF9B2" blockId="8.[163,779,1453,1904]" lastBlockId="8.[827,1443,1453,1904]" pageId="8">
The anterior articular process for the postanticlinal vertebrae (T12T14) are dorsomedially facing. This results in the anterior portion being more open than the more anterior thoracic vertebrae. The mammillary process becomes more developed moving down the column, with little separation from the anterior articular process in T13 and T14. None of the post-anticlinal vertebrae have ribs articulated or preserved. While the transverse fovea is present on T13, it is completely absent on T14 (
<figureCitation id="D1C02A6DFFBF71088600F8CF1ADCF8DC" box="[617,713,1821,1843]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="8">Fig. 10E</figureCitation>
). The accessory process is complete on both T13 and T14, but is relatively larger on T13 (
<figureCitation id="D1C02A6DFFBF71088584F8881A5BF881" box="[493,590,1882,1904]" captionStart="Figure 10" captionStartId="11.[145,223,1734,1756]" captionTargetBox="[226,1343,197,1692]" captionTargetId="figure-25@11.[225,1345,195,1694]" captionTargetPageId="11" captionText="Figure 10. Phocidae, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14." figureDoi="http://doi.org/10.5281/zenodo.5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="8">Fig. 10F</figureCitation>
). The accessory processes of both these vertebrae appear to be bifurcated, with a lateral and posterior facing tubercle. In ventral view, the posterior end of the body appears to be narrower than the anterior end. The posterior articular processes of the post-anticlinal thoracic vertebrae are ventrolaterally facing.
</paragraph>
<paragraph id="494436E8FFBF7109873AF9B71AB3F97B" blockId="8.[827,1443,1453,1904]" lastBlockId="9.[145,761,1479,1685]" lastPageId="9" pageId="8">
Several differences between representatives of
<taxonomicName id="8EFB4D6BFFBF71088752F9511BBDF977" authorityName="Gray" authorityYear="1825" box="[827,936,1667,1689]" class="Mammalia" family="Otariidae" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="family">Otariidae</taxonomicName>
(
<taxonomicName id="8EFB4D6BFFBF710887DAF9511CB2F977" authority="NMVC" authorityName="NMVC" box="[947,1191,1667,1689]" class="Mammalia" family="Otariidae" genus="Arctocephalus" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBF710887DAF9511C46F976" box="[947,1107,1667,1688]" italics="true" pageId="8">Arctocephalus</emphasis>
NMVC
</taxonomicName>
33579and
<taxonomicName id="8EFB4D6BFFBF71088172F9511B98F959" authority="USNM" authorityName="USNM" class="Mammalia" family="Otariidae" genus="Callorhinus" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBF71088172F9511DB6F976" box="[1307,1443,1667,1688]" italics="true" pageId="8">Callorhinus</emphasis>
USNM
</taxonomicName>
49439) and
<taxonomicName id="8EFB4D6BFFBF71088043F9701C8DF956" baseAuthorityName="King" baseAuthorityYear="1983" box="[1066,1176,1698,1720]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="family">Phocidae</taxonomicName>
(
<taxonomicName id="8EFB4D6BFFBF710880C2F9701D4FF956" authority="NMV C" authorityName="NMV C" box="[1195,1370,1698,1720]" class="Mammalia" family="Phocidae" genus="Phoca" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBF710880C2F9701CE1F959" box="[1195,1268,1698,1719]" italics="true" pageId="8">Phoca</emphasis>
NMV C
</taxonomicName>
27683 and
<taxonomicName id="8EFB4D6BFFBF7108871BF9131C63F938" authority="USNM" authorityName="USNM" box="[882,1142,1729,1750]" class="Mammalia" family="Phocidae" genus="Neomonachus" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBF7108871BF9131C0FF938" box="[882,1050,1729,1750]" italics="true" pageId="8">Neomonachus</emphasis>
USNM
</taxonomicName>
22543) support referral of NMV P160433 to
<taxonomicName id="8EFB4D6BFFBF71088043F90D1C83F91B" baseAuthorityName="King" baseAuthorityYear="1983" box="[1066,1174,1759,1781]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="family">Phocidae</taxonomicName>
. The accessory process of the post-anticlinal thoracic vertebrae is elongate, pronged and well separated from the transverse process, similar to
<taxonomicName id="8EFB4D6BFFBF71088079F8E91C40F8BE" box="[1040,1109,1851,1872]" class="Mammalia" family="Phocidae" genus="Phoca" kingdom="Animalia" order="Carnivora" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBF71088079F8E91C40F8BE" box="[1040,1109,1851,1872]" italics="true" pageId="8">Phoca</emphasis>
</taxonomicName>
. This contrasts with the thin and reduced accessory process seen in otariids, such as
<taxonomicName id="8EFB4D6BFFBE710984DBFA1519FAFA33" authority="Cuvier, 1826" authorityName="Cuvier" authorityYear="1826" box="[178,495,1479,1501]" class="Mammalia" family="Otariidae" genus="Arctocephalus" kingdom="Animalia" order="Carnivora" pageId="9" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBE710984DBFA151940FA32" box="[178,341,1479,1500]" italics="true" pageId="9">Arctocephalus</emphasis>
Cuvier, 1826
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFBE71098643FA1518DBFA12" authority="Gray, 1859" authorityName="Gray" authorityYear="1859" class="Mammalia" family="Otariidae" genus="Callorhinus" kingdom="Animalia" order="Carnivora" pageId="9" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBE71098643FA151AA0FA32" box="[554,693,1479,1500]" italics="true" pageId="9">Callorhinus</emphasis>
Gray, 1859
</taxonomicName>
. Lastly, the vertical condition of the spinous process on the anticlinal vertebrae of NMV P160433 appears to be most similar to phocids (NMV C27683, USNM 22543), which have a much steeper angle than otariids (NMV C33579, USNM49439) and odobenids (USNM396932), which have a shallower angle.
</paragraph>
<caption id="1D846660FFBE710984F9FAAA1CE1FA60" ID-DOI="http://doi.org/10.5281/zenodo.5300495" ID-Zenodo-Dep="5300495" box="[144,1268,1400,1422]" httpUri="https://zenodo.org/record/5300495/files/figure.png" pageId="9" startId="9.[144,222,1400,1422]" targetBox="[385,1185,195,1360]" targetPageId="9">
<paragraph id="494436E8FFBE710984F9FAAA1CE1FA60" blockId="9.[144,1268,1400,1422]" box="[144,1268,1400,1422]" pageId="9">
<emphasis id="7B8FEAFAFFBE710984F9FAAA18EDFA63" bold="true" box="[144,248,1400,1422]" pageId="9">Figure 8.</emphasis>
<taxonomicName id="8EFB4D6BFFBE7109856DFAAA1971FA60" baseAuthorityName="King" baseAuthorityYear="1983" box="[260,356,1400,1422]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="9" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P160433 articulated thoracic vertebrae and ribs in dorsal view.
</paragraph>
</caption>
<paragraph id="494436E8FFBE7109855AF9031A43F904" blockId="9.[307,598,1745,1770]" box="[307,598,1745,1770]" pageId="9">
<taxonomicName id="8EFB4D6BFFBE7109855AF9031A43F904" authority="GRAY, 1869" authorityName="GRAY" authorityYear="1869" box="[307,598,1745,1770]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="9" phylum="Chordata" rank="subFamily" subFamily="Monachinae">MONACHINAE GRAY, 1869</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="01E16563FFBE7109856DF92C1A74F884" pageId="9" type="nomenclature">
<paragraph id="494436E8FFBE7109856DF92C1A93F8F8" blockId="9.[260,646,1789,1814]" box="[260,646,1789,1814]" pageId="9">
<heading id="120C8184FFBE7109856DF92C1A93F8F8" box="[260,646,1789,1814]" centered="true" fontSize="9" level="2" pageId="9" reason="3">
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<emphasis id="7B8FEAFAFFBE7109856DF92C1A93F8F8" bold="true" box="[260,646,1789,1814]" pageId="9">MONACHINAE GEN. &amp; SP. INDET.</emphasis>
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<paragraph id="494436E8FFBE710984F8F8E41A74F884" blockId="9.[145,759,1846,1899]" pageId="9">
<emphasis id="7B8FEAFAFFBE710984F8F8E4196DF8A2" box="[145,376,1846,1868]" italics="true" pageId="9">Referred specimen:</emphasis>
NMV P232849, isolated right humerus, collected by R. Wilkie (
<figureCitation id="D1C02A6DFFBE7109866DF8871A44F884" box="[516,593,1877,1899]" captionStart="Figure 11" captionStartId="12.[163,243,1812,1834]" captionTargetBox="[188,1416,196,1770]" captionTargetId="figure-27@12.[187,1418,195,1772]" captionTargetPageId="12" captionText="Figure 11. Monachinae, gen. &amp; sp. indet., NMV P232849, isolated right humerus. A, posterior view. B, lateral view. C, anterior view. D, anteromedial view. E, posteromedial view. F, proximal view." figureDoi="http://doi.org/10.5281/zenodo.5300501" httpUri="https://zenodo.org/record/5300501/files/figure.png" pageId="9">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01E16563FFBE710D8740FA1A1D14FED8" lastPageId="13" pageId="9" type="description">
<paragraph id="494436E8FFBE71098740FA1A1D64F8C0" blockId="9.[809,1425,1479,1900]" pageId="9">
<emphasis id="7B8FEAFAFFBE71098740FA1A1BD1FA33" box="[809,964,1480,1501]" italics="true" pageId="9">Description:</emphasis>
NMV P232849 (
<figureCitation id="D1C02A6DFFBE710980CDFA151CEBFA32" box="[1188,1278,1479,1501]" captionStart="Figure 11" captionStartId="12.[163,243,1812,1834]" captionTargetBox="[188,1416,196,1770]" captionTargetId="figure-27@12.[187,1418,195,1772]" captionTargetPageId="12" captionText="Figure 11. Monachinae, gen. &amp; sp. indet., NMV P232849, isolated right humerus. A, posterior view. B, lateral view. C, anterior view. D, anteromedial view. E, posteromedial view. F, proximal view." figureDoi="http://doi.org/10.5281/zenodo.5300501" httpUri="https://zenodo.org/record/5300501/files/figure.png" pageId="9">Fig. 11</figureCitation>
) is a right humerus. The most anterior extent of the deltopectoral crest is missing, as is the lateral epicondyle. NMV P232849 represents a monachine phocid on the basis of a deltopectoral crest that curves gradually down the shaft, and the lack of an entepicondylar foramen. The lesser tubercle is slightly damaged anteriorly and posteriorly. The lesser tubercle, greater tubercle and the head are the same level. The bicipital groove is narrow and shallow, lacking a deep insertion for the m. supraspinatus. The head is rounded, but compressed proximodistally in both posterior and lateral views.
</paragraph>
<paragraph id="494436E8FFBE710A8728F8E51C6AF8C3" blockId="9.[809,1425,1479,1900]" lastBlockId="10.[827,1443,1632,1899]" lastPageId="10" pageId="9">The deltopectoral crest is anteroposteriorly deep, tapering towards the distal end of the shaft. The lateral surface posterior to the crest (the fossa for m. brachialis) is shallow. A deltopectoral overhang usually forms the anterior border of the m. brachialis fossa. The absence of an overhang suggests a more weakly developed m. brachialis. The deltoid tuberosity of NMV P232849 is laterally bulbous and large relative to the small size of the deltopectoral crest. The medial surface of the shaft is concave, smoothly transitioning into the medial epicondyle. Despite the lack of a preserved lateral epicondylar crest, it appears that the olecranon fossa is shallow. What is preserved of the trochlea on the medial side is small, but there may be a substantial portion missing due to the lack of the lateral epicondyle. The coronoid fossa is deep relative to the size of the medial epicondyle, and a visible outline is present between the trochlea and the epicondyles.</paragraph>
<caption id="1D846660FFBD710A84CBF9F11D5EF9D7" ID-DOI="http://doi.org/10.5281/zenodo.5300497" ID-Zenodo-Dep="5300497" box="[162,1355,1571,1593]" httpUri="https://zenodo.org/record/5300497/files/figure.png" pageId="10" startId="10.[162,240,1571,1593]" targetBox="[405,1203,197,1530]" targetPageId="10">
<paragraph id="494436E8FFBD710A84CBF9F11D5EF9D7" blockId="10.[162,1355,1571,1593]" box="[162,1355,1571,1593]" pageId="10">
<emphasis id="7B8FEAFAFFBD710A84CBF9F1191FF9D6" bold="true" box="[162,266,1571,1593]" pageId="10">Figure 9.</emphasis>
<taxonomicName id="8EFB4D6BFFBD710A857FF9F11963F9D7" baseAuthorityName="King" baseAuthorityYear="1983" box="[278,374,1571,1593]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="10" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P160433, articulated thoracic vertebrae and ribs in right lateral view.
</paragraph>
</caption>
<paragraph id="494436E8FFBD710D873AF8E419A0FCED" blockId="10.[827,1443,1632,1899]" lastBlockId="13.[145,762,197,1905]" lastPageId="13" pageId="10">
The fossa for the m. triceps brachii caput mediale in NMV P232849 is deep in comparison to the majority of phocids. Other taxa with a deep m. triceps brachii caput mediale are the phocine
<taxonomicName id="8EFB4D6BFFBA710D85A9FF361AECFF17" authorityName="USNM" baseAuthorityName="Erxleben" baseAuthorityYear="1777" box="[448,761,228,249]" class="Mammalia" family="Phocidae" genus="Pagophilus" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="groenlandicus">
<emphasis id="7B8FEAFAFFBA710D85A9FF361AECFF17" box="[448,761,228,249]" italics="true" pageId="13">Pagophilus groenlandicus</emphasis>
</taxonomicName>
(NHMUK 1938.12.10.1), the extinct monachine
<taxonomicName id="8EFB4D6BFFBA710D84F8FEF31A8CFED9" authority="Dewaele et al., 2018 b" authorityName="Dewaele" authorityYear="2018" box="[145,665,289,311]" class="Mammalia" family="Phocidae" genus="Auroraphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="atlantica">
<emphasis id="7B8FEAFAFFBA710D84F8FEF3198DFED8" box="[145,408,289,310]" italics="true" pageId="13">Auroraphoca atlantica</emphasis>
<bibRefCitation id="2D6A4B19FFBA710D85C9FEF31A8CFED9" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[416,665,289,311]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D8663FEF01A28FED8" box="[522,573,289,311]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
</taxonomicName>
(USNM 181419) and the extinct lobodontin
<taxonomicName id="8EFB4D6BFFBA710D8674FE92195EFE9A" authority="de Muizon, 1981" authorityName="de Muizon" authorityYear="1981" class="Mammalia" family="Phocidae" genus="Piscophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="pacifica">
<emphasis id="7B8FEAFAFFBA710D8674FE921AECFEBB" box="[541,761,320,341]" italics="true" pageId="13">Piscophoca pacifica</emphasis>
<bibRefCitation id="2D6A4B19FFBA710D84F8FE8C195EFE9A" author="de Muizon C" box="[145,331,350,372]" pageId="13" pagination="1 - 88" refId="ref11955" refString="de Muizon C. 1981. Les vertebres fossiles de la Formation Pisco (Perou). Premiere paretie: deux nouveaux Monachinae (Phocidae, Mammalia) du Pliocene de Sud-Sacaco. Memoire de l'Institut Francais d'Etudes Andines 50: 1 - 88." type="journal article" year="1981">de Muizon, 1981</bibRefCitation>
</taxonomicName>
(MNHN.F.SAS564), but the condition is not as well developed as in NMV P232849. The proximal level of the lesser tubercle is level with the head (when resting on the medial epicondyle), similar to stem-Lobodontini (
<bibRefCitation id="2D6A4B19FFBA710D85E2FE0B1A33FE00" author="Hendey QB &amp; Repenning CA" box="[395,550,473,495]" pageId="13" pagination="79 - 98" refId="ref11500" refString="Hendey QB, Repenning CA. 1972. A Pliocene phocid from South Africa. Annals of the South African Museum 59: 79 - 98." type="journal article" year="1972">Hendey, 1972</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D865DFE0B1AE6FE00" author="de Muizon C" box="[564,755,473,495]" pageId="13" pagination="1 - 88" refId="ref11955" refString="de Muizon C. 1981. Les vertebres fossiles de la Formation Pisco (Perou). Premiere paretie: deux nouveaux Monachinae (Phocidae, Mammalia) du Pliocene de Sud-Sacaco. Memoire de l'Institut Francais d'Etudes Andines 50: 1 - 88." type="journal article" year="1981">de Muizon, 1981</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D84F8FE2A19D1FDE3" author="Amson E &amp; de Muizon C" box="[145,452,504,526]" pageId="13" pagination="523 - 548" refId="ref10210" refString="Amson E, de Muizon C. 2014. A new durophagous phocid (Mammalia: Carnivora) from the late Neogene of Peru and considerations on monachine seals phylogeny. Journal of Systematic Palaeontology 12: 523 - 548." type="journal article" year="2014">Amson &amp; de Muizon, 2014</bibRefCitation>
) and
<taxonomicName id="8EFB4D6BFFBA710D8661FE2A195FFDC2" authority="Slater &amp; Helgen, 2014" authorityName="Slater &amp; Helgen" authorityYear="2014" class="Mammalia" family="Phocidae" genus="Neomonachus" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBA710D8661FE2A1ABCFDE3" box="[520,681,504,525]" italics="true" pageId="13">Neomonachus</emphasis>
Slater &amp; Helgen, 2014
</taxonomicName>
(USNM 395999 and USNM 22543). This differs from extant and extinct
<taxonomicName id="8EFB4D6BFFBA710D8651FDE71AB6FDA5" authorityName="Gray" authorityYear="1821" box="[568,675,565,587]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="subFamily" subFamily="Phocinae">Phocinae</taxonomicName>
, extant
<taxonomicName id="8EFB4D6BFFBA710D84F8FD86190BFD84" baseAuthorityName="Gray" baseAuthorityYear="1869" box="[145,286,596,618]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="tribe" tribe="Lobodontini">Lobodontini</taxonomicName>
,
<taxonomicName id="8EFB4D6BFFBA710D8543FD861A08FD87" authority="monachus" authorityName="monachus (Hermann" authorityYear="1779" box="[298,541,596,617]" class="Mammalia" family="Phocidae" genus="Monachus" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="monachus">
<emphasis id="7B8FEAFAFFBA710D8543FD861A08FD87" box="[298,541,596,617]" italics="true" pageId="13">Monachus monachus</emphasis>
</taxonomicName>
,
<taxonomicName id="8EFB4D6BFFBA710D8642FD861932FD66" authority="Tavani, 1941" authorityName="Tavani" authorityYear="1941" class="Mammalia" family="Phocidae" genus="Pliophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="etrusca">
<emphasis id="7B8FEAFAFFBA710D8642FD861AEDFD87" box="[555,760,596,617]" italics="true" pageId="13">Pliophoca etrusca</emphasis>
Tavani, 1941
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFBA710D850BFDA11A4EFD66" authority="Gray, 1827" authorityName="Gray" authorityYear="1827" box="[354,603,627,648]" class="Mammalia" family="Phocidae" genus="Mirounga" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="genus">
<emphasis id="7B8FEAFAFFBA710D850BFDA119C1FD66" box="[354,468,627,648]" italics="true" pageId="13">Mirounga</emphasis>
Gray, 1827
</taxonomicName>
, in which the lesser tubercle greatly extends proximal to the level of the head (see:
<bibRefCitation id="2D6A4B19FFBA710D8500FD621A57FD28" author="Berta A &amp; Kienle S &amp; Bianucci G &amp; Sorbi S" box="[361,578,688,710]" pageId="13" pagination="889144" refId="ref10338" refString="Berta A, Kienle S, Bianucci G, Sorbi S. 2015. A reevaluation of Pliophoca etrusca (Pinnipedia, Phocidae) from the Pliocene of Italy: phylogenetic and biogeographic implications. Journal of Vertebrate Paleontology 35: e 889144." type="journal article" year="2015">
Berta
<emphasis id="7B8FEAFAFFBA710D85D1FD6219FBFD2B" box="[440,494,688,709]" italics="true" pageId="13">et al</emphasis>
., 2015
</bibRefCitation>
). Like most of
<taxonomicName id="8EFB4D6BFFBA710D84F8FD1C1949FD0A" authority=", NMV P" authorityName="NMV P" box="[145,348,718,740]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="family">Phocidae, NMV P</taxonomicName>
232849 has a greater tubercle level with the head proximally.
</paragraph>
<caption id="1D846660FFBC710B84F8F9141C51F881" ID-DOI="http://doi.org/10.5281/zenodo.5300499" ID-Zenodo-Dep="5300499" httpUri="https://zenodo.org/record/5300499/files/figure.png" pageId="11" startId="11.[145,223,1734,1756]" targetBox="[226,1343,197,1692]" targetPageId="11">
<paragraph id="494436E8FFBC710B84F8F9141C51F881" blockId="11.[145,1425,1734,1903]" pageId="11">
<emphasis id="7B8FEAFAFFBC710B84F8F9141913F935" bold="true" box="[145,262,1734,1756]" pageId="11">Figure 10.</emphasis>
<taxonomicName id="8EFB4D6BFFBC710B8578F9141964F932" box="[273,369,1734,1756]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="11" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, gen. &amp; sp. indet., NMV P160433, close-ups of articulated thoracic vertebrae and ribs. A, right lateral view of thoracic vertebrae 9 and 10 and right rib 9, showing articulated ribs. B, dorsal view of thoracic vertebrae 10 and 11, showing the differing associations of the mamillary process with the anterior articular process. C, lateralventral view of thoracic vertebrae 9 and 10 with ribs in articulation with anterior and posterior costal fovea, and 11 with rib only in articulation with the anterior costal fovea. D, right lateral view of thoracic vertebrae 11 (anticlinal vertebrae). E, right lateral view of thoracic vertebrae 13 and 14. F, dorsal view of thoracic vertebrae 13 and 14.
</paragraph>
</caption>
<caption id="1D846660FFBB710C84CAF8C61BAAF8A9" ID-DOI="http://doi.org/10.5281/zenodo.5300501" ID-Zenodo-Dep="5300501" httpUri="https://zenodo.org/record/5300501/files/figure.png" pageId="12" startId="12.[163,243,1812,1834]" targetBox="[188,1416,196,1770]" targetPageId="12">
<paragraph id="494436E8FFBB710C84CAF8C61BAAF8A9" blockId="12.[163,1442,1812,1863]" pageId="12">
<emphasis id="7B8FEAFAFFBB710C84CAF8C6190EF8C7" bold="true" box="[163,283,1812,1834]" pageId="12">Figure 11.</emphasis>
<taxonomicName id="8EFB4D6BFFBB710C854EF8C619B9F8C4" box="[295,428,1812,1834]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="12" phylum="Chordata" rank="subFamily" subFamily="Monachinae">Monachinae</taxonomicName>
, gen. &amp; sp. indet., NMV P232849, isolated right humerus. A, posterior view. B, lateral view. C, anterior view. D, anteromedial view. E, posteromedial view. F, proximal view.
</paragraph>
</caption>
<paragraph id="494436E8FFBA710D84C0FCDE19F6FBDB" blockId="13.[145,762,197,1905]" pageId="13">
The compressed condition of the head of NMV P232849 is similar to that seen in
<taxonomicName id="8EFB4D6BFFBA710D8665FCF81AECFCD1" authority="monachus" authorityName="monachus (Hermann" authorityYear="1779" box="[524,761,810,831]" class="Mammalia" family="Phocidae" genus="Monachus" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="monachus">
<emphasis id="7B8FEAFAFFBA710D8665FCF81AECFCD1" box="[524,761,810,831]" italics="true" pageId="13">Monachus monachus</emphasis>
</taxonomicName>
(NHMUK 1894.7.27.1),
<taxonomicName id="8EFB4D6BFFBA710D85F5FC9B1A91FCB1" baseAuthorityName="Gray" baseAuthorityYear="1844" box="[412,644,841,863]" class="Mammalia" family="Phocidae" genus="Ommatophoca" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="rossii">
<emphasis id="7B8FEAFAFFBA710D85F5FC9B1A91FCB1" box="[412,644,841,863]" italics="true" pageId="13">Ommatophoca rossii</emphasis>
</taxonomicName>
(NHMUK 1965.8.2.1) and
<taxonomicName id="8EFB4D6BFFBA710D8536FCBA1935FC72" authority="Dewaele et al., 2018 b" authorityName="Dewaele" authorityYear="2018" class="Mammalia" family="Phocidae" genus="Virginiaphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="magurai">
<emphasis id="7B8FEAFAFFBA710D8536FCBA1A93FC93" box="[351,646,872,893]" italics="true" pageId="13">Virginiaphoca magurai</emphasis>
<bibRefCitation id="2D6A4B19FFBA710D86F8FCBA1935FC72" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D84F8FC5518D1FC75" box="[145,196,902,924]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
</taxonomicName>
(USNM 639750). There is little division between the head and lesser tubercle, such as in
<taxonomicName id="8EFB4D6BFFBA710D84F8FC16190AFC37" box="[145,287,964,985]" class="Mammalia" family="Phocidae" genus="Virginiaphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="magurai">
<emphasis id="7B8FEAFAFFBA710D84F8FC16190AFC37" box="[145,287,964,985]" italics="true" pageId="13">V. magurai</emphasis>
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFBA710D850FFC161AA2FC37" box="[358,695,964,985]" class="Mammalia" family="Phocidae" genus="Australophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="changorum">
<emphasis id="7B8FEAFAFFBA710D850FFC161AA2FC37" box="[358,695,964,985]" italics="true" pageId="13">Australophoca changorum</emphasis>
</taxonomicName>
. The narrow, shallow bicipital groove is a feature not found in extant
<taxonomicName id="8EFB4D6BFFBA710D8568FBD3199AFBF9" box="[257,399,1025,1047]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="subFamily" subFamily="Monachinae">Monachinae</taxonomicName>
, but shared with
<taxonomicName id="8EFB4D6BFFBA710D8631FBD319C8FBDB" authority="(Valenzuela-Toro et al., 2016)" baseAuthorityName="Valenzuela-Toro" baseAuthorityYear="2016" class="Mammalia" family="Phocidae" genus="Australophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="changorum">
<emphasis id="7B8FEAFAFFBA710D8631FBD31AECFBF8" box="[600,761,1025,1046]" italics="true" pageId="13">A. changorum</emphasis>
(
<bibRefCitation id="2D6A4B19FFBA710D84F0FBF219C1FBD8" author="Valenzuela-Toro AM &amp; Pyenson ND &amp; Gutstein CS &amp; Suarez ME" box="[153,468,1056,1078]" pageId="13" pagination="101 - 115" refId="ref12703" refString="Valenzuela-Toro AM, Pyenson ND, Gutstein CS, Suarez ME. 2016. A new dwarf seal from the late Neogene of South America and the evolution of pinnipeds in the southern hemisphere. Papers in Palaeontology 2: 101 - 115." type="journal article" year="2016">
Valenzuela-Toro
<emphasis id="7B8FEAFAFFBA710D8531FBF2199CFBDB" box="[344,393,1056,1077]" italics="true" pageId="13">et al</emphasis>
., 2016
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="494436E8FFBA710D84C0FBEC1AEFFA0D" blockId="13.[145,762,197,1905]" pageId="13">
The anteroposteriorly wide deltopectoral crest is similar to the state in
<taxonomicName id="8EFB4D6BFFBA710D85DAFB8F1A1BFB7F" authority="(Valenzuela-Toro et al., 2016)" baseAuthorityName="Valenzuela-Toro" baseAuthorityYear="2016" class="Mammalia" family="Phocidae" genus="Australophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="changorum">
<emphasis id="7B8FEAFAFFBA710D85DAFB8F1AEDFB9C" box="[435,760,1117,1138]" italics="true" pageId="13">Australophoca changorum</emphasis>
(
<bibRefCitation id="2D6A4B19FFBA710D84F3FBAE1A14FB7C" author="Valenzuela-Toro AM &amp; Pyenson ND &amp; Gutstein CS &amp; Suarez ME" box="[154,513,1148,1170]" pageId="13" pagination="101 - 115" refId="ref12703" refString="Valenzuela-Toro AM, Pyenson ND, Gutstein CS, Suarez ME. 2016. A new dwarf seal from the late Neogene of South America and the evolution of pinnipeds in the southern hemisphere. Papers in Palaeontology 2: 101 - 115." type="journal article" year="2016">
Valenzuela-Toro
<emphasis id="7B8FEAFAFFBA710D851BFBAE19BEFB7F" box="[370,427,1148,1169]" italics="true" pageId="13">et al</emphasis>
., 2016
</bibRefCitation>
)
</taxonomicName>
. The fossa for the m. brachialis is much deeper in multiple fossil phocid taxa (
<bibRefCitation id="2D6A4B19FFBA710D84BAFB6B19DEFB21" author="Dewaele L &amp; Lambert O &amp; Louwye S" box="[211,459,1209,1231]" pageId="13" pagination="3024" refId="ref10802" refString="Dewaele L, Lambert O, Louwye S. 2017 b. On Prophoca and Leptophoca (Pinnipedia, Phocidae) from the Miocene of the North Atlantic realm: redescription, phylogenetic affinities and paleobiogeographic implications. PeerJ 5: e 3024." type="journal article" year="2017">Dewaele et al., 2017b</bibRefCitation>
,
<bibRefCitation id="2D6A4B19FFBA710D85BEFB6B1A0AFB21" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[471,543,1209,1231]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">2018b</bibRefCitation>
), as it is for extant phocines. NMV P232849 lacks the deltopectoral overhang that is present in extant phocines, the extinct phocine
<taxonomicName id="8EFB4D6BFFBA710D853BFAC71A09FAA7" authority="(van Beneden, 1876) (Dewaele et al., 2017 b)" baseAuthorityName="van Beneden" baseAuthorityYear="1876" class="Mammalia" family="Phocidae" genus="Leptophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="proxima">
<emphasis id="7B8FEAFAFFBA710D853BFAC71A51FAC5" box="[338,580,1301,1323]" italics="true" pageId="13">Leptophoca proxima</emphasis>
(van Beneden, 1876) (
<bibRefCitation id="2D6A4B19FFBA710D8499FAE61A07FAA4" author="Dewaele L &amp; Lambert O &amp; Louwye S" box="[240,530,1332,1354]" pageId="13" pagination="3024" refId="ref10802" refString="Dewaele L, Lambert O, Louwye S. 2017 b. On Prophoca and Leptophoca (Pinnipedia, Phocidae) from the Miocene of the North Atlantic realm: redescription, phylogenetic affinities and paleobiogeographic implications. PeerJ 5: e 3024." type="journal article" year="2017">
Dewaele
<emphasis id="7B8FEAFAFFBA710D8503FAE619B3FAA7" box="[362,422,1332,1353]" italics="true" pageId="13">et al</emphasis>
., 2017b
</bibRefCitation>
)
</taxonomicName>
and the extinct monachines
<taxonomicName id="8EFB4D6BFFBA710D854CFA801A03FA86" authorityName=": Hendey" authorityYear="1972" box="[293,534,1362,1384]" class="Mammalia" family="Phocidae" genus="Homiphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="capensis">
<emphasis id="7B8FEAFAFFBA710D854CFA801A03FA86" box="[293,534,1362,1384]" italics="true" pageId="13">Homiphoca capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFBA710D863AFA8018E2FA69" class="Mammalia" family="Phocidae" genus="Virginiaphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="magurai">
<emphasis id="7B8FEAFAFFBA710D863AFA8018E2FA69" italics="true" pageId="13">Virginiaphoca magurai</emphasis>
</taxonomicName>
. A variation of the deltopectoral overhang is present in
<taxonomicName id="8EFB4D6BFFBA710D8546FA4218CBFA2A" authority="(Dewaele et al., 2018 b)" baseAuthorityName="Dewaele" baseAuthorityYear="2018" class="Mammalia" family="Phocidae" genus="Auroraphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="atlantica">
<emphasis id="7B8FEAFAFFBA710D8546FA421A2CFA4B" box="[303,569,1424,1445]" italics="true" pageId="13">Auroraphoca atlantica</emphasis>
(
<bibRefCitation id="2D6A4B19FFBA710D8622FA4218C0FA2A" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D86DEFA421AFEFA4B" box="[695,747,1424,1445]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
)
</taxonomicName>
, with a distinct separation of the feature on the lateral surface present on the proximal and distal end.
</paragraph>
<paragraph id="494436E8FFBA710D84C0FA3E1D14FED8" blockId="13.[145,762,197,1905]" lastBlockId="13.[809,1424,197,311]" pageId="13">
The medial side of the diaphysis is concave in posterior view, a feature present in multiple extant and fossil monachines, such as
<taxonomicName id="8EFB4D6BFFBA710D8674F9FB1997F9B3" authority="(Berta et al., 2015)" baseAuthorityName="Berta" baseAuthorityYear="2015" class="Mammalia" family="Phocidae" genus="Pliophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="etrusca">
<emphasis id="7B8FEAFAFFBA710D8674F9FB1AEDF9D1" box="[541,760,1577,1599]" italics="true" pageId="13">Pliophoca etrusca</emphasis>
(
<bibRefCitation id="2D6A4B19FFBA710D84F3F99A196CF9B3" author="Berta A &amp; Kienle S &amp; Bianucci G &amp; Sorbi S" box="[154,377,1607,1629]" pageId="13" pagination="889144" refId="ref10338" refString="Berta A, Kienle S, Bianucci G, Sorbi S. 2015. A reevaluation of Pliophoca etrusca (Pinnipedia, Phocidae) from the Pliocene of Italy: phylogenetic and biogeographic implications. Journal of Vertebrate Paleontology 35: e 889144." type="journal article" year="2015">
Berta
<emphasis id="7B8FEAFAFFBA710D8485F99A1937F9B3" box="[236,290,1608,1629]" italics="true" pageId="13">et al</emphasis>
., 2015
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="8EFB4D6BFFBA710D85AEF99A19ACF992" authority="(de Muizon &amp; Hendey, 1980)" baseAuthorityName="de Muizon &amp; Hendey" baseAuthorityYear="1980" class="Mammalia" family="Phocidae" genus="Homiphoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="species" species="capensis">
<emphasis id="7B8FEAFAFFBA710D85AEF99A1ADDF9B3" box="[455,712,1608,1629]" italics="true" pageId="13">Homiphoca capensis</emphasis>
(
<bibRefCitation id="2D6A4B19FFBA710D86B4F99A19BAF995" author="de Muizon C &amp; Hendey QB" pageId="13" pagination="91 - 128" refId="ref12068" refString="de Muizon C, Hendey QB. 1980. Late tertiary seals of the South Atlantic Ocean. Annals of the South African Museum 82: 91 - 128." type="journal article" year="1980">de Muizon &amp; Hendey, 1980</bibRefCitation>
)
</taxonomicName>
, as well as multiple North Atlantic species (
<bibRefCitation id="2D6A4B19FFBA710D8536F9571A49F975" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[351,604,1669,1691]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D85A5F95419EAF974" box="[460,511,1669,1691]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
). The medial diaphysis smoothly transitions into the medial epicondyle, unlike the more abrupt transition observed in other
<taxonomicName id="8EFB4D6BFFBA710D850BF93319D8F919" box="[354,461,1761,1783]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="family">Phocidae</taxonomicName>
. The medial epicondyle of NMV P232849 is relatively small compared to other
<taxonomicName id="8EFB4D6BFFBA710D84F8F8CC1933F8DA" box="[145,294,1822,1844]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="subFamily" subFamily="Monachinae">Monachinae</taxonomicName>
(
<bibRefCitation id="2D6A4B19FFBA710D8553F8CC1A12F8DD" author="de Muizon C" box="[314,519,1822,1844]" pageId="13" pagination="1 - 88" refId="ref11955" refString="de Muizon C. 1981. Les vertebres fossiles de la Formation Pisco (Perou). Premiere paretie: deux nouveaux Monachinae (Phocidae, Mammalia) du Pliocene de Sud-Sacaco. Memoire de l'Institut Francais d'Etudes Andines 50: 1 - 88." type="journal article" year="1981">de Muizon, 1981</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D8670F8CD1AE4F8DA" author="Berta A &amp; Kienle S &amp; Bianucci G &amp; Sorbi S" box="[537,753,1822,1844]" pageId="13" pagination="889144" refId="ref10338" refString="Berta A, Kienle S, Bianucci G, Sorbi S. 2015. A reevaluation of Pliophoca etrusca (Pinnipedia, Phocidae) from the Pliocene of Italy: phylogenetic and biogeographic implications. Journal of Vertebrate Paleontology 35: e 889144." type="journal article" year="2015">
Berta
<emphasis id="7B8FEAFAFFBA710D8601F8CD1A8BF8DD" box="[616,670,1822,1844]" italics="true" pageId="13">et al</emphasis>
., 2015
</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D84F8F8EF19C8F8BD" author="Valenzuela-Toro AM &amp; Pyenson ND &amp; Gutstein CS &amp; Suarez ME" box="[145,477,1853,1875]" pageId="13" pagination="101 - 115" refId="ref12703" refString="Valenzuela-Toro AM, Pyenson ND, Gutstein CS, Suarez ME. 2016. A new dwarf seal from the late Neogene of South America and the evolution of pinnipeds in the southern hemisphere. Papers in Palaeontology 2: 101 - 115." type="journal article" year="2016">
Valenzuela-Toro
<emphasis id="7B8FEAFAFFBA710D8533F8EC199BF8BC" box="[346,398,1853,1875]" italics="true" pageId="13">et al</emphasis>
., 2016
</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D8585F8EF1AFCF8BD" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[492,745,1853,1875]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D8630F8EC1A99F8BC" box="[601,652,1853,1875]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
). The shallow olecranon fossa is a common feature of fossil and extant
<taxonomicName id="8EFB4D6BFFBA710D807EFF171C91FF35" box="[1047,1156,197,219]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="13" phylum="Chordata" rank="family">Phocidae</taxonomicName>
, but is not necessarily exclusive to them (
<bibRefCitation id="2D6A4B19FFBA710D806AFF311CEBFF17" author="Dewaele L &amp; Lambert O &amp; Louwye S" box="[1027,1278,227,249]" pageId="13" pagination="3024" refId="ref10802" refString="Dewaele L, Lambert O, Louwye S. 2017 b. On Prophoca and Leptophoca (Pinnipedia, Phocidae) from the Miocene of the North Atlantic realm: redescription, phylogenetic affinities and paleobiogeographic implications. PeerJ 5: e 3024." type="journal article" year="2017">Dewaele et al., 2017b</bibRefCitation>
,
<bibRefCitation id="2D6A4B19FFBA710D8160FF311D47FF17" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[1289,1362,227,249]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">2018b</bibRefCitation>
). The coronoid fossa is deep relative to other monachines (
<bibRefCitation id="2D6A4B19FFBA710D8758FEF31BFAFED8" author="de Muizon C" box="[817,1007,289,311]" pageId="13" pagination="1 - 88" refId="ref11955" refString="de Muizon C. 1981. Les vertebres fossiles de la Formation Pisco (Perou). Premiere paretie: deux nouveaux Monachinae (Phocidae, Mammalia) du Pliocene de Sud-Sacaco. Memoire de l'Institut Francais d'Etudes Andines 50: 1 - 88." type="journal article" year="1981">de Muizon, 1981</bibRefCitation>
;
<bibRefCitation id="2D6A4B19FFBA710D8794FEF31CE4FED9" author="Dewaele L &amp; Peredo CM &amp; Meyvisch P &amp; Louwye S" box="[1021,1265,289,311]" pageId="13" pagination="172437" refId="ref10889" refString="Dewaele L, Peredo CM, Meyvisch P, Louwye S. 2018 b. Diversity of late Neogene Monachinae (Carnivora, Phocidae) from the North Atlantic, with the description of two new species. Royal Society Open Science 5: 172437." type="journal article" year="2018">
Dewaele
<emphasis id="7B8FEAFAFFBA710D800FFEF01C83FED8" box="[1126,1174,289,311]" italics="true" pageId="13">et al</emphasis>
., 2018b
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,325 @@
<document ID-DOI="http://dx.doi.org/10.3897/nl.43.49128" ID-GBIF-Dataset="f13774bd-fbef-4608-97e7-8841a21254f9" ID-GBIF-Taxon="163640124" ID-Pensoft-Pub="2367-5365-43-61" ID-Pensoft-UUID="8571D264804B582DB5C6826EB1F605EB" ID-ZooBank="AFE0F1E99E4B4E559A6473FA955F1895" LSID="urn:lsid:zoobank.org:act:55E14266-F078-455C-9CDC-1041F696AE45" ModsDocID="2367-5365-43-61" checkinTime="1586965389506" checkinUser="pensoft" docAuthor="Alipanah, Helen, Malm, Tobias &amp; Asselbergs, Jan" docDate="2020" docId="C152A15F4A4453CB8A363F2DF2D7554E" docLanguage="en" docName="NotaLepidopt 43: 61-76" docOrigin="Nota Lepidopterologica 43" docSource="http://dx.doi.org/10.3897/nl.43.49128" docTitle="Sitochroa urmiensis Alipanah 2020, sp. nov." docType="treatment" docUuid="55E14266-F078-455C-9CDC-1041F696AE45" docUuidSource="ZooBank" docVersion="3" id="8571D264804B582DB5C6826EB1F605EB" lastPageNumber="61" masterDocId="8571D264804B582DB5C6826EB1F605EB" masterDocTitle="A new species of Sitochroa Huebner, 1825 (Lepidoptera, Crambidae, Pyraustinae) from Iran, with taxonomic notes on the genus" masterLastPageNumber="76" masterPageNumber="61" pageId="0" pageNumber="61" updateTime="1643439810771" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of Sitochroa Huebner, 1825 (Lepidoptera, Crambidae, Pyraustinae) from Iran, with taxonomic notes on the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alipanah, Helen</mods:namePart>
<mods:affiliation>Iranian Research Institute of Plant Protection (IRIPP), Agricultural Research, Education and Extension Organization (AREEO), Tehran; e-mail: alipanah @ iripp. ir; halipanah @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">halipanah@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Malm, Tobias</mods:namePart>
<mods:affiliation>Department of Zoology, Swedish Museum of Natural History, Stockholm, Sweden; email: tobias. malm @ nrm. se</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asselbergs, Jan</mods:namePart>
<mods:affiliation>Neerland 20, 4614 GD Bergen op Zoom, The Netherlands; e-mail: jef. asselbergs @ hetnet. nl</mods:affiliation>
</mods:name>
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<mods:title>Nota Lepidopterologica</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>43</mods:number>
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<mods:start>61</mods:start>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ZooBank">AFE0F1E99E4B4E559A6473FA955F1895</mods:identifier>
<mods:identifier type="Pensoft-UUID">8571D264804B582DB5C6826EB1F605EB</mods:identifier>
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<treatment ID-GBIF-Taxon="163640124" LSID="urn:lsid:zoobank.org:act:55E14266-F078-455C-9CDC-1041F696AE45" httpUri="http://treatment.plazi.org/id/C152A15F4A4453CB8A363F2DF2D7554E" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="C152A15F-4A44-53CB-8A36-3F2DF2D7554E" authority="Alipanah" authorityName="Alipanah" authorityYear="2020" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa urmiensis" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="urmiensis" status="sp. nov.">Sitochroa urmiensis Alipanah</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adult males of Sitochroa spp. A, B Sitochroa urmiensis Alipanah, sp. nov., holotype, C-F S. palealis (Denis &amp; Schiffermueller). A, C, E Upper side, B, D, F Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61">Figs 1A, B</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">, 2A-C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of Sitochroa spp. A, C S. urmiensis Alipanah, sp. nov., holotype, male, B, D S. straminealis (Hampson), female. A, B Frontal view, C, D Lateral view." figureDoi="10.3897/nl.43.49128.figure3" httpUri="https://binary.pensoft.net/fig/399934" pageId="0" pageNumber="61">, 3A, C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="materials_examined">
<paragraph pageId="0" pageNumber="61">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="61">
Holotype, ♂, Iran, West
<normalizedToken originalValue="Āzarbāijān">Azarbaijan</normalizedToken>
Prov[ince]: Orumiyeh,
<normalizedToken originalValue="Ghāsemlu">Ghasemlu</normalizedToken>
, 10.vi.1975,
<normalizedToken originalValue="Abāi">Abai</normalizedToken>
leg. (genitalia preparation No. HA-2457, HMIM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
In external appearance, especially in wing pattern,
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
is close to
<taxonomicName genus="S." lsidName="S. palealis" pageId="0" pageNumber="61" rank="species" species="palealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. palealis</emphasis>
</taxonomicName>
(in populations with weakly expressed wing patterns) and less so to
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
. However, the medial line of the hindwing is more visible in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. compared with
<taxonomicName genus="S." lsidName="S. palealis" pageId="0" pageNumber="61" rank="species" species="palealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. palealis</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adult males of Sitochroa spp. A, B Sitochroa urmiensis Alipanah, sp. nov., holotype, C-F S. palealis (Denis &amp; Schiffermueller). A, C, E Upper side, B, D, F Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61">1A-F</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Adults of Sitochroa spp. A, B S. malekalis (Amsel) comb. nov., female, C S. straminealis (Hampson), female. A, C Upper side, B, D Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure8" httpUri="https://binary.pensoft.net/fig/399939" pageId="0" pageNumber="61">8C, D</figureCitation>
). There is no noticeable difference among the three above mentioned species in the shape of the head and the labial palpi (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of Sitochroa spp. A, C S. urmiensis Alipanah, sp. nov., holotype, male, B, D S. straminealis (Hampson), female. A, B Frontal view, C, D Lateral view." figureDoi="10.3897/nl.43.49128.figure3" httpUri="https://binary.pensoft.net/fig/399934" pageId="0" pageNumber="61">3A-D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head. A, B Loxostege farsalis Amsel syn. nov., holotype, male, C, D Sitochroa palealis (Denis &amp; Schiffermueller), male. A, C Frontal view, B, D Lateral view." figureDoi="10.3897/nl.43.49128.figure5" httpUri="https://binary.pensoft.net/fig/399936" pageId="0" pageNumber="61">5A-D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="61">
Based on the male genitalia, the new species is more similar to
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
than to any other known species of the genus (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A-F</figureCitation>
). These two species resemble each other mainly in the shape of the distal half of the valvae; the width of the uncus in relation to the width of the valvae; the rounded sinus between the prongs of the clasper; the presence of an inwardly pointed trapezoidal blade-like structure above the clasper; and a thorn-like or needle-shaped distal process in the phallus. Furthermore, in both species the cornuti consist of a bundle of elongate thorns and a finger-shaped process that is wavy in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. whereas it is straight in
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, C, D, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="61">
However, the male genitalia of these two species differ as follows: the tip of the uncus is semi-triangular in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. and dome-shaped in
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, D</figureCitation>
); the clasper prongs in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. are longer than those of
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, B, D, E</figureCitation>
); the internal prong of the clasper in
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
is clearly curved inward while it is straight in the new species (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, B, D, E</figureCitation>
); the medially directed process at the internal base of the clasper is dagger-shaped and long in
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
, but is a pointed triangular, deeply serrate and relatively short structure in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, B, D, E</figureCitation>
); the juxtal arms in the new species are slightly longer and narrower than in
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
; the saccus in
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
sp. nov. is slightly narrower than that of
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
and the anterior tip is nearly pointed (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 1.</emphasis>
Adult males of
<taxonomicName authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
<taxonomicName authorityName="Alipanah" authorityYear="2020" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa urmiensis" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa urmiensis</emphasis>
</taxonomicName>
Alipanah, sp. nov., holotype,
<emphasis bold="true" pageId="0" pageNumber="61">C-F</emphasis>
<taxonomicName genus="S." lsidName="S. palealis" pageId="0" pageNumber="61" rank="species" species="palealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. palealis</emphasis>
</taxonomicName>
(Denis &amp;
<normalizedToken originalValue="Schiffermüller">Schiffermueller</normalizedToken>
).
<emphasis bold="true" pageId="0" pageNumber="61">A, C, E</emphasis>
Upper side,
<emphasis bold="true" pageId="0" pageNumber="61">B, D, F</emphasis>
Underside. Scale bars: 10.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
Male genitalia of
<taxonomicName authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="61">A-C</emphasis>
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
Alipanah, sp. nov., holotype, genitalia preparation No. HA-2457, HMIM,
<emphasis bold="true" pageId="0" pageNumber="61">D-F</emphasis>
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
(Hampson), genitalia preparation No. HA-2436, HMIM.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description of male.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Head</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of Sitochroa spp. A, C S. urmiensis Alipanah, sp. nov., holotype, male, B, D S. straminealis (Hampson), female. A, B Frontal view, C, D Lateral view." figureDoi="10.3897/nl.43.49128.figure3" httpUri="https://binary.pensoft.net/fig/399934" pageId="0" pageNumber="61">3A, C</figureCitation>
). Frons conical, covered with pale brown scales, white laterally just next to compound eyes (most of the frontal scales removed); vertex erected, with pale yellowish-cream scales (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of Sitochroa spp. A, C S. urmiensis Alipanah, sp. nov., holotype, male, B, D S. straminealis (Hampson), female. A, B Frontal view, C, D Lateral view." figureDoi="10.3897/nl.43.49128.figure3" httpUri="https://binary.pensoft.net/fig/399934" pageId="0" pageNumber="61">3A</figureCitation>
); labial palpus porrect, 1.80 times horizontal diameter of compound eye, first segment covered with cream scales, second and third segments covered with cream scales ventrally, brown dorsally, apical segment as second except for brown scales dorso-laterally and slightly hanging; maxillary palpus slender, porrect and covered with brown scales; proboscis covered with dark brown scales, paler basally; antennae covered with cream scales dorsally, with a single considerably long seta present on dorsal surface of basal segment; collar yellowish-cream; thorax and tegula yellowish-cream; abdomen cream (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adult males of Sitochroa spp. A, B Sitochroa urmiensis Alipanah, sp. nov., holotype, C-F S. palealis (Denis &amp; Schiffermueller). A, C, E Upper side, B, D, F Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61">1A</figureCitation>
).
</paragraph>
<caption doi="10.3897/nl.43.49128.figure3" httpUri="https://binary.pensoft.net/fig/399934" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
Head of
<taxonomicName authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="61">A, C</emphasis>
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
Alipanah, sp. nov., holotype, male,
<emphasis bold="true" pageId="0" pageNumber="61">B, D</emphasis>
<taxonomicName genus="S." lsidName="S. straminealis" pageId="0" pageNumber="61" rank="species" species="straminealis">
<emphasis italics="true" pageId="0" pageNumber="61">S. straminealis</emphasis>
</taxonomicName>
(Hampson), female.
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
Frontal view,
<emphasis bold="true" pageId="0" pageNumber="61">C, D</emphasis>
Lateral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Forewing</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adult males of Sitochroa spp. A, B Sitochroa urmiensis Alipanah, sp. nov., holotype, C-F S. palealis (Denis &amp; Schiffermueller). A, C, E Upper side, B, D, F Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61">1A, B</figureCitation>
). Forewing length 15 mm; costa straight, slightly convex towards apex, rounded apically, termen obliquely convex. Ground colour glossy yellowish-cream with pale green tint, with veins and terminal line suffused pale-brownish, a hardly visible pale brown patch near apex, an obscure discoidal spot, and a hardly visible sinuate postmedial line fading away towards dorsum. Underside pale greyish-yellow, with brown costal margin, brown veins and terminal line, a brown sinuate postmedial line thickened towards costa and fading away towards dorsum, an oval pale brown spot near apex, a brown discoidal spot, and a short brown antemedial spot.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Hindwing</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adult males of Sitochroa spp. A, B Sitochroa urmiensis Alipanah, sp. nov., holotype, C-F S. palealis (Denis &amp; Schiffermueller). A, C, E Upper side, B, D, F Underside. Scale bars: 10.0 mm." figureDoi="10.3897/nl.43.49128.figure1" httpUri="https://binary.pensoft.net/fig/399932" pageId="0" pageNumber="61">1A, B</figureCitation>
). Greyish-yellow, paler than forewing, veins and terminal line with pale-brownish suffusion, a relatively wide medial line extending to two-thirds of the wing, and an obscure oval pale brown spot just next to apex. Underside ground colour as that of forewing, veins and terminal line brown, pale brown medial line narrowing and fading away towards dorsum, with small discoidal spot. Fringes of both fore- and hindwings glossy yellowish-cream with very fine light brown medial line.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A-C</figureCitation>
). Uncus relatively wide, rounded apically, with some relatively long apical hairs dorso-laterally; tegumen stout, scobinate dorsally; valva broad, rounded apically, with large, arched, clasper adorned with two ventrally directed thorn-like processes, external process serrate ventrally, sinus between prongs of clasper rounded, with deeply serrate, pointed, triangular and relatively large process at internal base of clasper directed medially, and a trapezoidal blade-like structure above clasper pointed internally; juxta deeply V-shaped; saccus elongated triangular (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, B</figureCitation>
); phallus with ventral thorn-like, straight and relatively long sclerotized process distally, cornuti comprising a bundle of elongate, heavily sclerotized spines and a finger-shaped and wavy sclerotized process (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Sitochroa spp. A-C S. urmiensis Alipanah, sp. nov., holotype, genitalia preparation No. HA- 2457, HMIM, D-F S. straminealis (Hampson), genitalia preparation No. HA- 2436, HMIM." figureDoi="10.3897/nl.43.49128.figure2" httpUri="https://binary.pensoft.net/fig/399933" pageId="0" pageNumber="61">2A, C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">Iran: West Azarbaijan Province (Orumiyeh or Urmia).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="61">The new species is named from the type locality, Urmia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="biology_ecology">
<paragraph pageId="0" pageNumber="61">Biology.</paragraph>
<paragraph pageId="0" pageNumber="61">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="remarks">
<paragraph pageId="0" pageNumber="61">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="61">
The studied male of
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
was collected in 1975. Although several collecting trips have been made by the first author and her colleagues in HMIM to Urmia and Ghasemlu since then, no
<taxonomicName authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa</emphasis>
</taxonomicName>
specimens have been collected again from that locality till now. In spite of the superficial similarities among the single examined
<taxonomicName genus="S." lsidName="S. urmiensis" pageId="0" pageNumber="61" rank="species" species="urmiensis">
<emphasis italics="true" pageId="0" pageNumber="61">S. urmiensis</emphasis>
</taxonomicName>
and some other
<taxonomicName authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sitochroa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitochroa" order="Lepidoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Sitochroa</emphasis>
</taxonomicName>
species, the differences in the male genitalia are considerable and the species can easily be distinguished.
</paragraph>
</subSubSection>
</treatment>
</document>