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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.666.9191" ID-PMC="PMC5534527" ID-Pensoft-Pub="1313-2970-666-1" ID-Pensoft-UUID="C43EEB41A94B930FFE439D1FAD29FF9C" ID-PubMed="28769631" ID-Zenodo-Dep="579453" ID-ZooBank="B3C377E8BBB14F328AECA2C22D1E625A" ModsDocID="1313-2970-666-1" checkinTime="1491551283800" checkinUser="pensoft" docAuthor="Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. &amp; Seidel, Matthias" docDate="2017" docId="C10912D48A4817302A62CB157EBA769F" docLanguage="en" docName="ZooKeys 666: 1-349" docOrigin="ZooKeys 666" docPubDate="2017-04-06" docSource="http://dx.doi.org/10.3897/zookeys.666.9191" docTitle="Eremophygus Ohaus 1910" docType="treatment" docVersion="6" id="C43EEB41A94B930FFE439D1FAD29FF9C" lastPageNumber="1" masterDocId="C43EEB41A94B930FFE439D1FAD29FF9C" masterDocTitle="Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies" masterLastPageNumber="349" masterPageNumber="1" pageNumber="1" updateTime="1668164364999" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jameson, Mary L.</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
<mods:nameIdentifier type="email">maryliz.jameson@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garner, Beulah H.</mods:namePart>
<mods:affiliation>Natural History Museum, Insects Division, Department of Life Sciences, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Audibert, Cedric</mods:namePart>
<mods:affiliation>Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew B. T.</mods:namePart>
<mods:affiliation>Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-04-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>666</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>349</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.666.9191</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.666.9191</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-666-1</mods:identifier>
<mods:identifier type="ZooBank">B3C377E8BBB14F328AECA2C22D1E625A</mods:identifier>
<mods:identifier type="Pensoft-UUID">C43EEB41A94B930FFE439D1FAD29FF9C</mods:identifier>
<mods:identifier type="Zenodo-Dep">579453</mods:identifier>
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<treatment ID-GBIF-Taxon="128344249" LSID="urn:lsid:plazi:treatment:C10912D48A4817302A62CB157EBA769F" httpUri="http://treatment.plazi.org/id/C10912D48A4817302A62CB157EBA769F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="C10912D4-8A48-1730-2A62-CB157EBA769F" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eremophygus Ohaus, 1910</taxonomicName>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype female of Eremophygus calvus Gutierrez from UCCC. A Dorsal habitus B Specimen labels." figureDoi="10.3897/zookeys.666.9191.figure20" httpUri="https://binary.pensoft.net/fig/130448" pageId="0" pageNumber="1">Figs 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Eremophygus lasiocalinus Ohaus holotype male from ZMHB. A Dorsal habitus B Lateral habitus C Specimen labels, mouthparts, and male genitalia D Male genitalia, lateral view E Parameres, dorsal view." figureDoi="10.3897/zookeys.666.9191.figure21" httpUri="https://binary.pensoft.net/fig/130449" pageId="0" pageNumber="1">, 21</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Holotype male of Eremophygus leo Gutierrez from UCCC. A Dorsal habitus B Specimen labels, mouthparts, and male genitalia." figureDoi="10.3897/zookeys.666.9191.figure22" httpUri="https://binary.pensoft.net/fig/130450" pageId="0" pageNumber="1">, 22</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Eremophygus pachyloides Ohaus holotype female from ZMHB. A Dorsal habitus B Lateral habitus C Specimen labels, mouthparts, and egg." figureDoi="10.3897/zookeys.666.9191.figure23" httpUri="https://binary.pensoft.net/fig/130451" pageId="0" pageNumber="1">, 23</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Eremophygus philippii Ohaus holotype male from ZMHB. A Dorsal habitus B Lateral habitus C Parameres, dorsal view D Male genitalia, lateral view E Specimen labels." figureDoi="10.3897/zookeys.666.9191.figure24" httpUri="https://binary.pensoft.net/fig/130452" pageId="0" pageNumber="1">, 24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ohaus" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus philippii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippii">
<emphasis italics="true" pageId="0" pageNumber="1">Eremophygus philippii</emphasis>
</taxonomicName>
Ohaus, 1910.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="species">
<paragraph pageId="0" pageNumber="1">Species.</paragraph>
<paragraph pageId="0" pageNumber="1">6 species; length 14-15 mm.</paragraph>
<paragraph pageId="0" pageNumber="1">
Rarity of specimens in collections as well as possible paraphyly with the genera
<taxonomicName authorityName="Solier" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Oogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oogenius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Oogenius</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Gutierrez" authorityYear="1951" class="Insecta" family="Scarabaeidae" genus="Microogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microogenius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Microogenius</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Mondaca &amp; Valencia" authorityYear="2016" class="Insecta" family="Scarabaeidae" genus="Peruquime" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peruquime" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peruquime</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Lasiocala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiocala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lasiocala</emphasis>
</taxonomicName>
hampers our understanding of the biodiversity of this group. Species in the genus
<taxonomicName authorityName="Ohaus" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eremophygus</emphasis>
</taxonomicName>
are distributed in the altiplano of Bolivia, Argentina, Peru, and Chile.
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
described two species in the genus (
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="0" pageNumber="1" pagination="106 - 125" refId="B104" refString="Gutierrez, AR, 1951. Notas sobre Scarabaeidae neotropicos II (Coleopt. Lamellic.). Anales de la Sociedad Cientifica Argentina 151: 106 - 125" title="Notas sobre Scarabaeidae neotropicos II (Coleopt. Lamellic.)." volume="151" year="1951">
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
1951
</bibRefCitation>
,
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="1" pagination="207 - 227" refId="B105" refString="Gutierrez, AR, 1952. Notas sobre Scarabaeidae neotropicos (III) (Coleoptera). Revista Chilena de Entomologia 2: 207 - 227" title="Notas sobre Scarabaeidae neotropicos (III) (Coleoptera)." volume="2" year="1952">1952</bibRefCitation>
), discussed the genus (
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="0" pageNumber="1" pagination="9 - 35" refId="B102" refString="Gutierrez, AR, 1949. Notas sobre Scarabaeidae Neotropicos (Coleoptera Lamellicornia). Anales de la Sociedad Cientifica Argentina 148: 9 - 35" title="Notas sobre Scarabaeidae Neotropicos (Coleoptera Lamellicornia)." volume="148" year="1949">
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
1949
</bibRefCitation>
,
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="0" pageNumber="1" pagination="52 - 75" refId="B103" refString="Gutierrez, AR, 1950. Scarabaeidae del Norte de Chile. Anales de la Sociedad Cientifica Argentina 149 (2): 52 - 75" title="Scarabaeidae del Norte de Chile." volume="149" year="1950">1950</bibRefCitation>
,
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="0" pageNumber="1" pagination="106 - 125" refId="B104" refString="Gutierrez, AR, 1951. Notas sobre Scarabaeidae neotropicos II (Coleopt. Lamellic.). Anales de la Sociedad Cientifica Argentina 151: 106 - 125" title="Notas sobre Scarabaeidae neotropicos II (Coleopt. Lamellic.)." volume="151" year="1951">1951</bibRefCitation>
,
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="1" pagination="207 - 227" refId="B105" refString="Gutierrez, AR, 1952. Notas sobre Scarabaeidae neotropicos (III) (Coleoptera). Revista Chilena de Entomologia 2: 207 - 227" title="Notas sobre Scarabaeidae neotropicos (III) (Coleoptera)." volume="2" year="1952">1952</bibRefCitation>
), and provided the most recent key to species (
<bibRefCitation author="Gutierrez, AR" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="1" pagination="207 - 227" refId="B105" refString="Gutierrez, AR, 1952. Notas sobre Scarabaeidae neotropicos (III) (Coleoptera). Revista Chilena de Entomologia 2: 207 - 227" title="Notas sobre Scarabaeidae neotropicos (III) (Coleoptera)." volume="2" year="1952">
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
1952
</bibRefCitation>
), yet he did not discuss the
<normalizedToken originalValue="group’s">group's</normalizedToken>
relationships or context within the
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Rutelinae">Rutelinae</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Some species in the genus lack the independently movable claws that are diagnostic of
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Rutelinae">Rutelinae</taxonomicName>
(that is, the apex of meso- and metatarsomere 5 lack a longitudinal slit, a character suite shared with cyclocephaline rhinoceros beetles [
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
:
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
]). One species,
<taxonomicName authorityName="Martinez" authorityYear="1960" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus pereirai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">Eremophygus pereirai</emphasis>
</taxonomicName>
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
(from Jujuy, Argentina), was transferred to the dynastine tribe
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
and the genus
<taxonomicName class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala</emphasis>
</taxonomicName>
by
<bibRefCitation author="Martinez, A" journalOrPublisher="Frey Tutzing bei Muenchen" pageId="0" pageNumber="1" pagination="263 - 274" refId="B208" refString="Martinez, A, 1975b. Cyclocephala sudamericanas nuevas o poco conocidas (Col. Scarabaeidae Dynastinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Muenchen 26: 263 - 274" title="Cyclocephala sudamericanas nuevas o poco conocidas (Col. Scarabaeidae Dynastinae). Entomologische Arbeiten aus dem Museum G." volume="26" year="1975 b">
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
(1975b)
</bibRefCitation>
who compared the toothless maxillary galea of
<taxonomicName lsidName="E. pereirai" pageId="0" pageNumber="1" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">E. pereirai</emphasis>
</taxonomicName>
to the similar maxilla in
<taxonomicName authorityName="Martinez" authorityYear="1965" class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala zischkai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala zischkai</emphasis>
</taxonomicName>
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
from Bolivia (
<bibRefCitation author="Martinez, A" journalOrPublisher="Ciencia" pageId="0" pageNumber="1" pagination="131 - 133" refId="B201" refString="Martinez, A, 1960. Una nueva especie de Eremophygus (Col., Scarab., Rutelin.). Ciencia 20: 131 - 133" title="Una nueva especie de Eremophygus (Col., Scarab., Rutelin.)." volume="20" year="1960">
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
1960
</bibRefCitation>
,
<bibRefCitation author="Martinez, A" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="0" pageNumber="1" pagination="63 - 74" refId="B202" refString="Martinez, A, 1965. Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63 - 74" title="Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae)." volume="179" year="1965">1965</bibRefCitation>
).
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.91390" author="Endroedi, S" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="1" pagination="319 - 321" refId="B64" refString="Endroedi, S, 1977. Alissonotum piceum besucheti subsp. n. (Col. Melolonthidae, Dynastinae). Revue Suisse de Zoologie 84: 319 - 321, DOI: https://doi.org/10.5962/bhl.part.91390" title="Alissonotum piceum besucheti subsp. n. (Col. Melolonthidae, Dynastinae)." url="https://doi.org/10.5962/bhl.part.91390" volume="84" year="1977">
<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
(1977)
</bibRefCitation>
agreed with the tribal transfer and also compared
<taxonomicName lsidName="E. pereirai" pageId="0" pageNumber="1" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">E. pereirai</emphasis>
</taxonomicName>
to
<taxonomicName lsidName="C. zischkai" pageId="0" pageNumber="1" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="1">C. zischkai</emphasis>
</taxonomicName>
, considering these species distinctive enough so that, together, they could warrant subgeneric status within
<taxonomicName class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Martinez" authorityYear="1960" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus pereirai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">Eremophygus pereirai</emphasis>
</taxonomicName>
(as
<taxonomicName lsidName="C. pereirai" pageId="0" pageNumber="1" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">C. pereirai</emphasis>
</taxonomicName>
) was later included in the key to world
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
and
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
(
<bibRefCitation author="Endroedi, S" journalOrPublisher="Junk Publisher, Dordrecht" pageId="0" pageNumber="1" refId="B65" refString="Endroedi, S, 1985. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht" title="The Dynastinae of the World. Dr. W." year="1985">
<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
1985
</bibRefCitation>
).
<bibRefCitation author="Krajcik, M" journalOrPublisher="X Supplement 5, Plzen" pageId="0" pageNumber="1" refId="B156" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Plzen" title="Checklist of the world Scarabaeoidea. Animma." year="2012">Krajcik (2012)</bibRefCitation>
included
<taxonomicName lsidName="E. pereirai" pageId="0" pageNumber="1" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">E. pereirai</emphasis>
</taxonomicName>
under
<taxonomicName class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala</emphasis>
</taxonomicName>
following
<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
.
<taxonomicName authorityName="Martinez" authorityYear="1965" class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala zischkai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala zischkai</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. pereirai" pageId="0" pageNumber="1" rank="species" species="pereirai">
<emphasis italics="true" pageId="0" pageNumber="1">C. pereirai</emphasis>
</taxonomicName>
have male parameres that are formed from two, laterally articulated plates, a character associated with
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
and not
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
(male parameres are fused into a single plate that is not laterally articulating). This represents another example of genera historically considered to be part of
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
(e.g.,
<taxonomicName class="Insecta" family="Dynastidae" genus="Peltonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peltonotus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peltonotus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Ohaus" authorityYear="1912" class="Insecta" family="Dynastidae" genus="Acrobolbia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Acrobolbia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Acrobolbia</emphasis>
</taxonomicName>
Ohaus) that were later transferred to
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
. This highlights the need for phylogenetic analyses including
<taxonomicName authorityName="Ohaus" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Eremophygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremophygus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eremophygus</emphasis>
</taxonomicName>
to broadly sample taxa from
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
and
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
to resolve the tribal, and thus the subfamilial, placement of this genus.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Diagnostic characters have greatly diminished reliability because of overlap with
<emphasis italics="true" pageId="0" pageNumber="1">
<taxonomicName authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Lasiocala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiocala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Lasiocala</taxonomicName>
,
<taxonomicName authorityName="Solier" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Oogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oogenius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Oogenius</taxonomicName>
</emphasis>
,
<taxonomicName authorityName="Mondaca &amp; Valencia" authorityYear="2016" class="Insecta" family="Scarabaeidae" genus="Peruquime" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peruquime" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peruquime</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Gutierrez" authorityYear="1951" class="Insecta" family="Scarabaeidae" genus="Microogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microogenius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Microogenius</emphasis>
</taxonomicName>
, and
<taxonomicName class="Insecta" family="Dynastidae" genus="Cyclocephala" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyclocephala</emphasis>
</taxonomicName>
and should be used with great caution: dorsal surface often with long, tawny setae; apex of labrum extends beyond clypeal apex, visible in dorsal view; antenna 9- or 10-segmented (9-segmented according to
<bibRefCitation author="Mondaca, J" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 10" refId="B217" refString="Mondaca, J, Valencia, G, 2016. Nuevo genero y especie de Rutelini (Coleoptera: Scarabaeidae: Rutelinae) de los Andes peruanos. Insecta Mundi 473: 1 - 10" title="Nuevo genero y especie de Rutelini (Coleoptera: Scarabaeidae: Rutelinae) de los Andes peruanos." volume="473" year="2016">Mondaca and Valencia [2016]</bibRefCitation>
); lateral edge of mandibles rounded and without reflexed teeth; apex of clypeus varies from rounded to subtrapezoidal; pronotum with apical bead complete medially, laterally, and basally; lateral edge of protibia with three rounded teeth; apex of fourth metatarsomere lacking spiniform attenuation; base of metatibia nearly straight, lacking distinct notch; apex of meso- and metatibia with many spinules, and; mesosternum not appreciably produced beyond the mesometasternal suture. In some species (e.g.,
<taxonomicName lsidName="E. lasiocalinus" pageId="0" pageNumber="1" rank="species" species="lasiocalinus">
<emphasis italics="true" pageId="0" pageNumber="1">E. lasiocalinus</emphasis>
</taxonomicName>
Ohaus), the protarsal claw is enlarged and deeply split; the meso- and metatarsal claws may be deeply split or simple; the unguitractor plate of meso- and metatarsus is subcylindrical with 2 or 3 setae; and the apex of tarsomere 5 (meso- and metatarsus) with 2 weak, longitudinal slits at apex (a character that is not shared by most other
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
; instead it is more common in the
<taxonomicName lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Melolonthinae">Melolonthinae</taxonomicName>
and
<taxonomicName authorityName="MacLeay" authorityYear="1819" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
). Larvae, natural history, and sister-group relationships are not known.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Twelve new species and fifty-three new provincial distribution records of Aleocharinae rove beetles of Saskatchewan, Canada (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart>Larson, David J.</mods:namePart>
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classification Animalia Coleoptera Staphylinidae
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<paragraph pageId="48" pageNumber="93">
<taxonomicName authority="Klimaszewski &amp; Webster" class="Insecta" family="Staphylinidae" genus="Gnypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnypeta minuta" order="Coleoptera" pageId="48" pageNumber="93" phylum="Arthropoda" rank="species" species="minuta">Gnypeta minuta Klimaszewski &amp; Webster</taxonomicName>
</paragraph>
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(for diagnosis and illustrations, see
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="ZooKeys" pageId="51" pageNumber="96" pagination="11 - 84" title="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution." url="10.3897/zookeys.2.4" volume="2" year="2008 c">Klimaszewski et al. 2008c</bibRefCitation>
)
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</subSubSection>
<subSubSection pageId="48" pageNumber="93" type="distribution">
<paragraph pageId="48" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="48" pageNumber="93">
<table pageId="48" pageNumber="93">
<tr pageId="48" pageNumber="93">
<td colspan="1" pageId="48" pageNumber="93" rowspan="1">NBSK</td>
</tr>
<tr pageId="48" pageNumber="93">
<td colspan="1" pageId="48" pageNumber="93" rowspan="1">SaskatchewanABDLC</td>
</tr>
<tr pageId="48" pageNumber="93">
<td colspan="1" pageId="48" pageNumber="93" rowspan="1">
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="ZooKeys" pageId="51" pageNumber="96" pagination="11 - 84" title="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution." url="10.3897/zookeys.2.4" volume="2" year="2008 c">Klimaszewski et al. 2008c</bibRefCitation>
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="49" pageNumber="94" title="Checklist of beetles (Coleoptera) of Canada and Alaska." year="2013">Bousquet et al. 2013</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="94" type="natural history">
<paragraph pageId="49" pageNumber="94">
<pageBreakToken pageId="49" pageNumber="94" start="start">Natural</pageBreakToken>
history.
</paragraph>
<paragraph pageId="49" pageNumber="94">
In SK, one female was captured in June from sandy-clay river bank. In NB, two specimens were captured in June, one from under debris on muddy soil near a small pool in a silver maple forest, and the other from under debris on clay and sand mix at river margin (
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)
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<taxonomicName authority="rhadamanthus (Esper, [1789])" authorityName="rhadamanthus (Esper" authorityYear="1789" baseAuthorityName="Esper, [1789]" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) rhadamanthus subsp. rhadamanthus" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subGenus="Zygaena" subSpecies="rhadamanthus">Zygaena (Zygaena) rhadamanthus rhadamanthus (Esper, [1789])</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="reference_group">
<paragraph pageId="4" pageNumber="127">
<taxonomicName authorityName="Esper" authorityYear="1789" class="Insecta" family="Zygaenidae" genus="Sphinx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphinx rhadamanthus" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="rhadamanthus">Sphinx rhadamanthus</taxonomicName>
Esper, [1789], Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen 2 (Supplement): pl. 40, figs 1, 2; 1794, ibidem 2: 13. Type-locality. France: Gard (Languedoc),
<normalizedToken originalValue="Nîmes">Nimes</normalizedToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="127">
<taxonomicName authorityName="Verity" authorityYear="1920" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena rhadamanthus subsp. pyrenaea" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subSpecies="pyrenaea">Zygaena rhadamanthus pyrenaea</taxonomicName>
Verity, 1920, The
<normalizedToken originalValue="Entomologist’s">Entomologist's</normalizedToken>
Record and Journal of Variation 32: 161 (
<taxonomicName authorityName="Efetov &amp; Hofmann &amp; Tarmann &amp; Tremewan" authorityYear="2014" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="genus">Zygaena</taxonomicName>
). Type-locality. France:
<normalizedToken originalValue="Pyrénées-Orientales">Pyrenees-Orientales</normalizedToken>
, between Prades and Mont Canigou, La Trancada
<normalizedToken originalValue="d’Ambouilla">d'Ambouilla</normalizedToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="127">
<taxonomicName authorityName="Aistleitner" authorityYear="1990" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena rhadamanthus subsp. boixolsis" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subSpecies="boixolsis">Zygaena rhadamanthus boixolsis</taxonomicName>
Aistleitner, 1990, Nachrichten des Entomologischen Vereins Apollo (N.F.) 11: 79. Type-locality. Spain:
<normalizedToken originalValue="Lérida">Lerida</normalizedToken>
, Collado de Boixols, 1,300-1,400 m. [Paratypes 4 ♂, 4; Topotypes 4 ♂, 1 ♀ examined.]
<emphasis bold="true" pageId="4" pageNumber="127">Syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="distribution and taxonomic notes">
<paragraph pageId="4" pageNumber="127">Distribution and taxonomic notes.</paragraph>
<paragraph pageId="4" pageNumber="127">
Based on a description by
<bibRefCitation author="Oberthuer, C" journalOrPublisher="Etudes de Lepidopterologie comparee" pageId="8" pageNumber="131" refId="B28" refString="Oberthuer, C, 1910. . Etudes de Lepidopterologie comparee 4" volume="4" year="1910">
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
(1910
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: 591), Verity provided the name
<taxonomicName authorityName="Verity" authorityYear="1920" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subsp. pyrenaea" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subSpecies="pyrenaea">
<emphasis italics="true" pageId="4" pageNumber="127">pyrenaea</emphasis>
</taxonomicName>
for the population of
<taxonomicName baseAuthorityName="Esper" baseAuthorityYear="1789" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena rhadamanthus" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="rhadamanthus">
<emphasis italics="true" pageId="4" pageNumber="127">Zygaena rhadamanthus</emphasis>
</taxonomicName>
occurring at La Trancada
<normalizedToken originalValue="d’Ambouilla">d'Ambouilla</normalizedToken>
(
<normalizedToken originalValue="Pyrénées-Orientales">Pyrenees-Orientales</normalizedToken>
, between Prades and Mont Canigou).
<bibRefCitation author="Leraut, P" journalOrPublisher="NAP Editions, Verrieres-le Buisson" pageId="7" pageNumber="130" refId="B20" refString="Leraut, P, 2012. Volume 3: Moths of Europe, Zygaenids, Pyralids 1 and Brachodids. NAP Editions, Verrieres-le Buisson" title="Volume 3: Moths of Europe, Zygaenids, Pyralids 1 and Brachodids." year="2012">Leraut (2012</bibRefCitation>
: 98) considered this taxon to be a 'simple
<normalizedToken originalValue="synonym’">synonym'</normalizedToken>
of the nominotypical
<taxonomicName authority="rhadamanthus" authorityName="rhadamanthus (Esper" authorityYear="1789" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena rhadamanthus subsp. rhadamanthus" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subSpecies="rhadamanthus">
<emphasis italics="true" pageId="4" pageNumber="127">Zygaena rhadamanthus rhadamanthus</emphasis>
</taxonomicName>
, a placement with which we agree even though
<bibRefCitation author="Hofmann, A" journalOrPublisher="Harley Books, Colchester" pageId="7" pageNumber="130" refId="B17" refString="Hofmann, A, Tremewan, WG, 1996. A systematic Catalogue of the Zygaeninae (Lepidoptera: Zygaenidae). Harley Books, Colchester" title="A systematic Catalogue of the Zygaeninae (Lepidoptera: Zygaenidae)." year="1996">Hofmann and Tremewan (1996</bibRefCitation>
: 131) had considered it to be a valid subspecies with which the taxon
<taxonomicName authorityName="Aistleitner" authorityYear="1990" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena rhadamanthus subsp. boixolsis" order="Lepidoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="rhadamanthus" subSpecies="boixolsis">
<emphasis italics="true" pageId="4" pageNumber="127">Zygaena rhadamanthus boixolsis</emphasis>
</taxonomicName>
Aistleitner, 1990, had been placed as a synonym. The latter is now automatically transferred as a synonym of the nominotypical subspecies. Specimens from La Trancada
<normalizedToken originalValue="d’Ambouilla">d'Ambouilla</normalizedToken>
lack the white scaling (griseoid) on the ground colour of the forewings; moreover, a red abdominal cingulum is absent, as found in the nominotypical subspecies.
</paragraph>
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</treatment>
</document>

97
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<mods:title id="5D9F743113D344CA913CCFE6929639F4">Chapter 7: Linnaean Plant Names and their Types (part A)</mods:title>
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<emphasis id="325E2AE85DEA9682232572CDE3595BB7" bold="true" pageNumber="299">Anthericum alooides</emphasis>
Linnaeus
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,
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<emphasis id="981A469800AADA21389185067282F048" italics="true" pageNumber="299">Species Plantarum</emphasis>
1
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.
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<paragraph id="00FF40383FA8AF83088CC612CCD74BD0" pageNumber="299">&quot;Habitat ad Cap. b. Spei.&quot; RCN: 2450.</paragraph>
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<paragraph id="721B247610B37EC2ECA1D897E2C99A7D" pageNumber="299">Type not designated.</paragraph>
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<paragraph id="A6DE1F4D134926BE896C6E972A08CF70" pageNumber="299">Original material: [icon] in Dillenius, Hort. Eltham. 2: 312, t. 232, f. 299. 1732.</paragraph>
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<paragraph id="9F16F1E91F84608D225571685DC81DB1" pageNumber="299">
Current name:
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<emphasis id="0DAD237D9C71039275BA7D79E3BD66B7" italics="true" pageNumber="299">
<emphasis id="DAAE0B72581BD19A25DE7AD4F2FBDF23" bold="true" italics="true" pageNumber="299">Bulbine alooides</emphasis>
</emphasis>
(L.) Willd.
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(
<taxonomicName id="6494F3197FC4615644D773A21F17E9D9" class="Liliopsida" family="Liliaceae" kingdom="Plantae" order="Liliales" phylum="Tracheophyta" rank="family">Liliaceae</taxonomicName>
/
<taxonomicName id="EFA26D1CE08343C549CB2773EEDF98C0" authorityName="de Jussieu" authorityYear="1789" class="Magnoliopsida" family="Asphodelaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="family">Asphodelaceae</taxonomicName>
).
</paragraph>
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<paragraph id="48F3A90B058F1A15A1AF14AF43E86918" pageNumber="299">
<emphasis id="5285E568DE03901B4FC6083B06D76C82" italics="true" pageNumber="299">Note:</emphasis>
Wijnands (
<emphasis id="D4603203CB719CB1176A43D914255B67" italics="true" pageNumber="299">Bot. Commelins</emphasis>
: 129. 1983) stated that the Dillenius plate &quot;might serve as the type&quot;, but his uncertainty prevents this being accepted as a formal typification. Kativu (in
<emphasis id="3EC7A49E0DEAE59B19266D962D318DBE" italics="true" pageNumber="299">Kirkia</emphasis>
16: 48. 1997) indicated Clifford material (BM) as the
<typeStatus id="AE7D481C8643F4543BB48B08EE621EF8">holotype</typeStatus>
but this appears to be a Gronovius collection now excluded from the Clifford herbarium, and not original material for the name. Subsequently, Van Jaarsveld &amp; Forster (in Eggli,
<emphasis id="B3E9453C08D9FCBF6B4123C14DFF3727" italics="true" pageNumber="299">Ill. Handb. Succ. Pl., Monocots</emphasis>
: 233. 2001) indicated the Dillenius plate as type but as this choice was published after 1 Jan 2001, their omission of the phrase &quot;designated here&quot; or an equivalent (Art. 7.11) means that the choice is not effective.
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<emphasis id="7BD4DC31FFFDFFBE11D2CF73FEB1FAAE" bold="true" box="[151,327,1290,1315]" pageId="3" pageNumber="419">Type material.</emphasis>
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, ‘
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,
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,
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,
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–
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,
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</date>
; E-Y: 1060, from under stones, leg.
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(
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)
</materialsCitation>
.
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<typeStatus id="961BBE81FFFDFFBE13A5CF56FCA7FACA" box="[736,849,1327,1351]" pageId="3" pageNumber="419" type="paratype">Paratypes</typeStatus>
:
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,
<specimenCount id="5FA6CBAAFFFDFFBE12D0CF56FC23FACB" box="[917,981,1327,1351]" pageId="3" pageNumber="419" type="female">3 ♀♀</specimenCount>
, 7 spec., the same data as holotype (
<specimenCount id="5FA6CBAAFFFDFFBE1436CF56FA6AFACB" box="[1395,1436,1327,1351]" pageId="3" pageNumber="419" type="male">1 ♂</specimenCount>
<specimenCount id="5FA6CBAAFFFDFFBE11D2CF2AFF21FAE7" box="[151,215,1363,1387]" pageId="3" pageNumber="419" type="female">2 ♀♀</specimenCount>
, 7 spec.
<collectionCode id="2FB198E6FFFDFFBE107CCF2AFE7CFAE6" box="[313,394,1363,1387]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="419">TMSA</collectionCode>
,
<specimenCount id="5FA6CBAAFFFDFFBE10D1CF2AFE48FAE7" box="[404,446,1363,1387]" pageId="3" pageNumber="419" type="female">1 ♀</specimenCount>
<collectionCode id="2FB198E6FFFDFFBE1083CF2AFDE4FAE6" box="[454,530,1363,1387]" pageId="3" pageNumber="419">RBSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFDFFB81182CF0FFBF3F87B" lastPageId="5" lastPageNumber="421" pageId="3" pageNumber="419" type="description">
<paragraph id="491F0023FFFDFFBE1182CF0FFB3AFA03" blockId="3.[151,1437,1290,1999]" box="[199,1228,1398,1423]" pageId="3" pageNumber="419">
<emphasis id="7BD4DC31FFFDFFBE1182CF0FFEAFFA02" bold="true" box="[199,345,1398,1423]" pageId="3" pageNumber="419">Description.</emphasis>
Body length males
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, females
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,
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.
</paragraph>
<paragraph id="491F0023FFFDFFBE1182CFE3FC0FF992" blockId="3.[151,1437,1290,1999]" pageId="3" pageNumber="419">Elytra with sparse regular rows of setae on odd intervals, setae fine, slightly bent backward, about equally long as width of one interval, 8–10 on the length of one interval, at apical part some setae placed also on even intervals. Pronotum, head and rostrum with similar fine setae, half as long as elytral ones and semi-appressed, those setae on pronotum directed transversely, on head with rostrum directed longitudinally.</paragraph>
<paragraph id="491F0023FFFDFFBE1182CC52FDA2F906" blockId="3.[151,1437,1290,1999]" pageId="3" pageNumber="419">
Rostrum (
<figureCitation id="D19B1CA6FFFDFFBE1073CC53FE8CF9CE" box="[310,378,1578,1603]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="3" pageNumber="419">Fig. 2</figureCitation>
) 1.09–1.15 × as wide as long, base 1.11–1.15 × wider than apex; rostrum and head moderately shiny, irregularly punctured, distance of punctures smaller than their diameter, basal part of vertex and space behind eyes without punctures, smooth.
</paragraph>
<paragraph id="491F0023FFFDFFBE1182CCEFFAEBF896" blockId="3.[151,1437,1290,1999]" pageId="3" pageNumber="419">
Antennal scape (
<figureCitation id="D19B1CA6FFFDFFBE10C2CCEFFE3CF922" box="[391,458,1686,1711]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="3" pageNumber="419">Fig. 1</figureCitation>
) regularly curved at midlength, evenly enlarged at apical third, at apex 0.7–0.8 × as wide as club. Funicle segment 1 conical, 2.1–2.2 × as long as wide and 2.2–2.5 × as long as segment 2, which is 1.2–1.3 × as long as wide; segment 3 isodiametric; segment 4 1.1 × as wide as long; segment 5 1.2 × as wide as long; segment 6 1.3 × as wide as long; segment 7 1.4–1.5 × as wide as long; club 1.5 × as long as wide.
</paragraph>
<paragraph id="491F0023FFFDFFBE1182CD5EFE1EF842" blockId="3.[151,1437,1290,1999]" pageId="3" pageNumber="419">
Pronotum (
<figureCitation id="D19B1CA6FFFDFFBE100FCD5FFE66F8B2" box="[330,400,1830,1855]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="3" pageNumber="419">Fig. 1</figureCitation>
) 1.07–1.11 × as wide as long, widest at midlength or before midlength, with strongly regularly rounded sides, anterior border narrower than posterior border. Disc strongly shiny, punctured, smooth between punctures; punctation variable, punctures ranging from very small, fine and deeper with long interspaces (longer than puncture diameter) to larger, shallower and with interspaces shorter than puncture diameter. Punctures on disc mostly variably-sized.
</paragraph>
<caption id="1DDF50ABFFFAFFB911D2CCEFFC08F921" box="[151,1022,1686,1708]" httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="4" pageNumber="420" startId="4.[151,250,1686,1708]" targetBox="[312,1284,211,1605]" targetPageId="4">
<paragraph id="491F0023FFFAFFB911D2CCEFFC08F921" blockId="4.[151,1022,1686,1708]" box="[151,1022,1686,1708]" pageId="4" pageNumber="420">
<emphasis id="7BD4DC31FFFAFFB911D2CCEFFEE5F921" bold="true" box="[151,275,1686,1708]" pageId="4" pageNumber="420">FIGURE 1.</emphasis>
Dorsal habitus of the male holotype of
<taxonomicName id="8EA07BA0FFFAFFB913E1CCEFFC48F926" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[676,958,1686,1707]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="endroedyi" status="sp. nov.">
<emphasis id="7BD4DC31FFFAFFB913E1CCEFFC48F926" box="[676,958,1686,1707]" italics="true" pageId="4" pageNumber="420">Basothorhynchus endroedyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E7614AFFFAFFB91280CCEFFC08F921" box="[965,1022,1686,1708]" pageId="4" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="491F0023FFFAFFB91182CCAAFC34F89D" blockId="4.[151,1437,1747,2024]" pageId="4" pageNumber="420">
Elytra (
<figureCitation id="D19B1CA6FFFAFFB91058CCAAFE97F961" box="[285,353,1747,1772]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="4" pageNumber="420">Fig. 1</figureCitation>
) in males 1.48–1.53 ×, in females 1.34–1.43 × as long as wide. Striae constituted by slender punctures, space between punctures not deepened; intervals matt, rough.
</paragraph>
<paragraph id="491F0023FFFAFFB91182CD62FABBF865" blockId="4.[151,1437,1747,2024]" pageId="4" pageNumber="420">Protarsi in both sexes shorter and broader than meso- and metatarsi; all tarsi in males shorter and more robust than those in females. Male protarsi with segment 2 twice as wide as long; segment 3 1.2–1.3 × as wide as long and onychium equally long as segment 3; meso- and metatarsi with segment 2 1.7–1.8 × as wide as long; segment 3 1.3 × as wide as long and onychium 1.3 × as long as segment 3. Female protarsi with segment 2 1.7–1.8 × as wide as long; segment 3 1.3 × as wide as long and onychium 1.3 × as long as segment 3; meso- and metatarsi with segment 2 1.5–1.6 × as wide as long; segment 3 1.1–1.2 × as wide as long and onychium 1.4 × as long as segment 3.</paragraph>
<caption id="1DDF50ABFFFBFFB811D2CCE3FD27F880" httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="5" pageNumber="421" startId="5.[151,264,1690,1712]" targetBox="[224,1370,263,1601]" targetPageId="5">
<paragraph id="491F0023FFFBFFB811D2CCE3FD27F880" blockId="5.[151,1436,1690,1805]" pageId="5" pageNumber="421">
<emphasis id="7BD4DC31FFFBFFB811D2CCE3FEBCF93D" bold="true" box="[151,330,1690,1712]" pageId="5" pageNumber="421">FIGURES 2–11.</emphasis>
Structural details of
<taxonomicName id="8EA07BA0FFFBFFB81368CCE2FCBDF93D" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[557,843,1691,1712]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="421" phylum="Arthropoda" rank="species" species="endroedyi" status="sp. nov.">
<emphasis id="7BD4DC31FFFBFFB81368CCE2FCBDF93D" box="[557,843,1691,1712]" italics="true" pageId="5" pageNumber="421">Basothorhynchus endroedyi</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFBFFB81213CCE3FC65F93D" bold="true" box="[854,915,1690,1712]" pageId="5" pageNumber="421">
<taxonomicNameLabel id="60E7614AFFFBFFB81213CCE3FC65F93D" box="[854,915,1690,1712]" pageId="5" pageNumber="421" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11).
</paragraph>
</caption>
<paragraph id="491F0023FFFBFFB81182CD36FE12F83D" blockId="5.[151,1436,1871,2038]" pageId="5" pageNumber="421">
Penis (
<figureCitation id="D19B1CA6FFFBFFB81057CD36FEA2F8E5" box="[274,340,1871,1896]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="5" pageNumber="421">Fig. 7</figureCitation>
) dorsally widest at apical third, basally evenly tapered with sides basally straight; apically tapered distinctly with sides of apical part rounded, tip small, subtriangular; in profile slender, distinctly irregularly curved, tip elongated, evenly pointed.
</paragraph>
<paragraph id="491F0023FFFBFFB81182CDC2FBF3F87B" blockId="5.[151,1436,1871,2038]" pageId="5" pageNumber="421">
Spermatheca (
<figureCitation id="D19B1CA6FFFBFFB81023CDC2FE40F859" box="[358,438,1979,2004]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="5" pageNumber="421">Fig. 11</figureCitation>
) with cornu wide, robust, regularly curved; ramus perpendicular to body of spermatheca, globular, isodiametric; collum not differentiated; tip of spermatheca truncated.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFF8FFBB1182CAEEFB1BFEBB" pageId="6" pageNumber="422" type="etymology">
<paragraph id="491F0023FFF8FFBB1182CAEEFB1BFEBB" blockId="6.[151,1437,151,310]" pageId="6" pageNumber="422">
<emphasis id="7BD4DC31FFF8FFBB1182CAEEFEB8FF3D" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="422">Etymology.</emphasis>
The newly described species is named in memory of the late Sebastian Endrödy-Younga (
<date id="3D1E26E3FFF8FFBB1411CAE1FF25FF5C" pageId="6" pageNumber="422" value="1934" valueMax="1999">1934– 1999</date>
), (Sebestyén Endrődy in his original name and surname), an originally Hungarian coleopterist who spent the majority of his life in
<collectingCountry id="31B740B3FFF8FFBB10D4CAA3FDD4FF7E" box="[401,546,218,243]" name="South Africa" pageId="6" pageNumber="422">South Africa</collectingCountry>
and who collected important and extensive material of different
<taxonomicName id="8EA07BA0FFF8FFBB1446CAA3FA88FF7E" authorityName="" authorityYear="1758" box="[1283,1406,218,243]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="422" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
in
<collectingCountry id="31B740B3FFF8FFBB11D2CA85FEDAFE98" box="[151,300,252,277]" name="South Africa" pageId="6" pageNumber="422">South Africa</collectingCountry>
, and also from less visited parts, like the high altitude areas of
<collectingCountry id="31B740B3FFF8FFBB154FCA85FB9EFE98" box="[1034,1128,252,277]" name="Lesotho" pageId="6" pageNumber="422">Lesotho</collectingCountry>
. Material collected by him constitutes a major foundation for our knowledge of the mainly terricolous beetles of
<collectingCountry id="31B740B3FFF8FFBB1513CB64FB1FFEBB" box="[1110,1257,285,310]" name="South Africa" pageId="6" pageNumber="422">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

404
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<paragraph id="491F0023FFFFFFBC11D2CAAEFE17FF7F" blockId="1.[151,481,215,242]" box="[151,481,215,242]" pageId="1" pageNumber="417">
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<emphasis id="7BD4DC31FFFFFFBC11D2CAAEFE9AFF7C" bold="true" box="[151,364,215,241]" italics="true" pageId="1" pageNumber="417">Basothorhynchus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E7614AFFFFFFBC1031CAA1FE17FF7F" box="[372,481,216,242]" pageId="1" pageNumber="417" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="01BA53A8FFFFFFBC11D2CB61FC79FEA3" box="[151,911,280,302]" pageId="1" pageNumber="417" type="materials_examined">
<paragraph id="491F0023FFFFFFBC11D2CB61FC79FEA3" blockId="1.[151,911,280,302]" box="[151,911,280,302]" pageId="1" pageNumber="417">
<emphasis id="7BD4DC31FFFFFFBC11D2CB61FED4FEA3" bold="true" box="[151,290,280,302]" pageId="1" pageNumber="417">
<typeStatus id="961BBE81FFFFFFBC11D2CB61FF3AFEA3" box="[151,204,280,302]" pageId="1" pageNumber="417">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8EA07BA0FFFFFFBC106CCB61FDB5FEA0" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[297,579,280,301]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="endroedyi" status="sp. nov.">
<emphasis id="7BD4DC31FFFFFFBC106CCB61FDB5FEA0" box="[297,579,280,301]" italics="true" pageId="1" pageNumber="417">Basothorhynchus endroedyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E7614AFFFFFFBC130FCB61FD6CFEA0" box="[586,666,280,301]" pageId="1" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFFFFBC11D2CB23FB91FDE2" pageId="1" pageNumber="417" type="diagnosis">
<paragraph id="491F0023FFFFFFBC11D2CB23FB91FDE2" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
<emphasis id="7BD4DC31FFFFFFBC11D2CB23FEE7FEFE" bold="true" box="[151,273,346,371]" pageId="1" pageNumber="417">Diagnosis.</emphasis>
Small-sized genus with dorsal surface of body glabrous; rostrum evenly tapered apicad with straight sides, posteriorly continuous with head; base of epifrons as wide as space between anterior margins of eyes, apex narrower than half width of rostrum; frons glabrous, smooth; epistome slightly carinate; scrobes placed dorsally; scape short, not reaching anterior border of pronotum in repose; elytron without humeral callus; all femora with distinct tooth; tibiae slender at basal third and here abruptly enlarged inside, apical two thirds of tibiae subparallelsided, outer edge straight; metatibiae lacking corbel; claws free; tegmen bearing two parameres; female sternite VIII with plate about as long as apodeme; plate narrowly subtriangular, apically pointed, with two longitudinal sclerites at distal three quarters, apical margin sclerotised, and basal margin ill-defined.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFFFFBF1182C803FC46FC45" lastPageId="2" lastPageNumber="418" pageId="1" pageNumber="417" type="description">
<paragraph id="491F0023FFFFFFBC1182C803FD88FD1F" blockId="1.[151,1437,346,2027]" box="[199,638,634,659]" pageId="1" pageNumber="417">
<emphasis id="7BD4DC31FFFFFFBC1182C803FEAFFD1E" bold="true" box="[199,345,634,659]" pageId="1" pageNumber="417">Description.</emphasis>
Body length 2.3–3.0 mm.
</paragraph>
<paragraph id="491F0023FFFFFFBC1182C8E7FC95FCE7" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
Body integument dark brownish to blackish (
<figureCitation id="D19B1CA6FFFFFFBC138BC8E7FCE7FD3A" box="[718,785,670,695]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="1" pageNumber="417">Fig. 1</figureCitation>
). Antennae and legs paler, rusty brownish, covered with short, fine, semi-erect setae; only clubs slightly darker, finely and densely setose. Dorsal side of body glabrous, lacking appressed scales or setae, with inconspicuous fine, slender erect setae on elytra and shorter semi-erect setae on pronotum and head plus rostrum. Head and rostrum glabrous on ventral side; sternum densely covered by oval greyish appressed scales; abdominal ventrites covered with short, fine, sparse, semi-appressed setae and several individual appressed scales in lateral parts of ventrites 1 and 2..
</paragraph>
<paragraph id="491F0023FFFFFFBC1182C90FFC6BFAEF" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
Head wide and short, slightly enlarged posteriad, vertex flat, shorter than half length of eye; ventrally unpunctured, smooth. Eyes moderately large, subdorsal, hardly prominent from outline of head; laterally subcircular, not reaching dorsal border of head. Rostrum (
<figureCitation id="D19B1CA6FFFFFFBC125FC9C7FC8DFC5A" box="[794,891,958,983]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="1" pageNumber="417">Figs 2, 3</figureCitation>
) short, 1.1–1.2 × as wide as long, evenly tapered apicad, at base 1.1 × as wide as at apex; dorsal border convex and ventral border straight in profile. Epifrons widest at base, conspicuously tapered apicad, with faintly concave sides, distinctly carinate laterally, at base as wide as space between anterior margins of eyes, at apical half slightly narrower than half width of rostrum in the same place, extending apically to level of anterior border of scrobes, median carina present along apical half; rostrum posteriorly continuous with head. Frons glabrous, smooth, yellowish-brown, moderately shiny, not separated posteriorly from epifrons, with five to six prominent pairs of stout setae apically. Epistome emarginate at apex, carinate, separated from frons by a keel. Scrobes placed dorsally, in dorsal view fully visible in apical third of rostrum as deep, oval fossae, broadly reniform; furrow-shaped, with conspicuous dorsal and ventral edges, and directed towards eye in lateral view, ventral margin terminated slighty above ventral margin of eye. Venter of rostrum shallowly depressed on the whole surface, covered with semi-appressed sparse setae. Mandibles small, projecting anteriorly, lacking scales, with three laterally placed setae.
</paragraph>
<paragraph id="491F0023FFFFFFBC1182CF16FDD7FA42" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
Antennae (
<figureCitation id="D19B1CA6FFFFFFBC1004CF17FE76FA0A" box="[321,384,1390,1415]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="1" pageNumber="417">Fig. 1</figureCitation>
) moderately short and robust. Scape not reaching anterior border of pronotum in repose, 1.1 × longer than funicle; funicle 7-segmented, segment 1 long and conical (the longest), segments 3–7 isodiametric to wider than long; club broadly oval.
</paragraph>
<paragraph id="491F0023FFFFFFBC1182CFA2FC05F9B7" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
Pronotum (
<figureCitation id="D19B1CA6FFFFFFBC1001CFA3FE72FA7E" box="[324,388,1498,1523]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="1" pageNumber="417">Fig. 1</figureCitation>
) 1.1 × wider than long, with distinctly rounded sides; disc regularly convex without any keel, furrow or depression; base slightly arched. Anterior border of pronotum laterally straight without ocular lobes or setae. Procoxal cavities contiguous, round, placed at midlength of pronotum.
</paragraph>
<paragraph id="491F0023FFFFFFBC1182CC3FFD28F9D2" blockId="1.[151,1437,346,2027]" box="[199,734,1606,1631]" pageId="1" pageNumber="417">Scutellum triangular, very small, hardly visible.</paragraph>
<paragraph id="491F0023FFFFFFBC1182CC13FE16F8F2" blockId="1.[151,1437,346,2027]" pageId="1" pageNumber="417">
Elytra (
<figureCitation id="D19B1CA6FFFFFFBC105FCC13FEACF90E" box="[282,346,1642,1667]" captionStart="FIGURE 1" captionStartId="4.[151,250,1686,1708]" captionTargetBox="[312,1284,211,1605]" captionTargetId="figure@4.[302,1287,195,1672]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of the male holotype of Basothorhynchus endroedyi sp. n." httpUri="https://zenodo.org/record/2625549/files/figure.png" pageId="1" pageNumber="417">Fig. 1</figureCitation>
) oval, 1.3–1.5 × longer than wide; humeri weakly rounded, lacking a distinct callus; base slightly wider than base of pronotum, distinctly arched, widest at midlength; apex broadly rounded; sides slightly convex, slope of declivity hardly overhanging apices of elytra in dorsal view. Elytra 10-striate; striae punctate, intervals flat. Mesocoxae semiglobular, mesosternal process narrow, about as wide as 1/5 of mesocoxal width. Metacoxae semiglobular, metaventral process obtuse, narrower than transverse diameter of metacoxa. Tergite VII (
<figureCitation id="D19B1CA6FFFFFFBC146ACC83FA85F89E" box="[1327,1395,1786,1811]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="1" pageNumber="417">Fig. 4</figureCitation>
) of males short and wide, apically V-shaped concave with straight sides; tergite VIII (
<figureCitation id="D19B1CA6FFFFFFBC1568CD67FB86F8BA" box="[1069,1136,1822,1847]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="1" pageNumber="417">Fig. 4</figureCitation>
) of males well sclerotised, bowl-shaped, apically weakly rounded. Tergite VII and VIII of females larger than those in males, weakly sclerotised, apically rounded.
</paragraph>
<paragraph id="491F0023FFFFFFBF1182CDF2FD6BFE90" blockId="1.[151,1437,346,2027]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="418" pageId="1" pageNumber="417">
Legs moderately robust. Femora (
<figureCitation id="D19B1CA6FFFFFFBC1301CDF3FD71F82E" box="[580,647,1930,1955]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="1" pageNumber="417">Fig. 5</figureCitation>
) swollen, all femora with distinct tooth equal in both sexes, almost as large as half the width of femora at this level, concave at inner edge. Procoxae subglobular, protibiae (
<figureCitation id="D19B1CA6FFFFFFBC1454CDD7FAA2F84A" box="[1297,1364,1966,1991]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="1" pageNumber="417">Fig. 5</figureCitation>
) short and robust, laterally straight, slender at basal third and abruptly enlarged distad, inside with inner edge slightly concave and with three to six small brownish teeth, apex rounded and densely fringed by brown bristles, mucronate. Meso- and metatibiae weakly enlarged outside. Metatibial apex oval, glabrous fringed externally and internally with dense brown bristles. Tarsi with segment 3 bilobed, distinctly wider than segment 1 and 2; onychium long; claws free, weakly divaricate.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CB5EFE33FE5D" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
Abdomen (
<figureCitation id="D19B1CA6FFFCFFBF1001CB5EFE71FECD" box="[324,391,295,320]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 6</figureCitation>
) 1.2–1.3 × longer than wide, with rounded sides; ventrite 1 twice as long as ventrite 2, behind metacoxa equally long as ventrite 2; ventrite 2 slightly shorter than ventrites 3 and 4 combined; ventrite 5 of males subtrapezoidal, short, shallowly concave at apex, ventrite 5 of females subtriangular, longer than in males, convex, apically rounded. Suture between ventrites 1 and 2 indistinctly sinuose, fine and narrow; all other sutures straight, deep and moderately wide.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CBA5FD51FD95" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182CBA5FE4CFE78" bold="true" box="[199,442,476,501]" pageId="2" pageNumber="418">Sexual dimorphism.</emphasis>
Males distinguishable from females by the more slender elytra, the wider and shorter tarsi, and the shape and concavity of ventrite 5.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182C85DFEBEFD7D" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182C85DFE63FDB0" bold="true" box="[199,405,548,573]" pageId="2" pageNumber="418">Male terminalia.</emphasis>
Penis (
<figureCitation id="D19B1CA6FFFCFFBF10B3C85DFDB6FDB1" box="[502,576,548,573]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 7</figureCitation>
) short, well sclerotised, temones 1.4–1.5 × longer than body of penis; endophallus long, with two moderately long, basally connected sclerites and one short transverse arched sclerite placed distally. Tegmen (
<figureCitation id="D19B1CA6FFFCFFBF10F0C815FE0FFD09" box="[437,505,620,645]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 8</figureCitation>
) slender, parameres distinct, fused at base and distinctly shorter than diameter of ring, manubrium 1.4 × as long as diameter of ring, 0.8 × as long as temones. Spiculum gastrale moderately short, anteriorly curved and enlarged to form a narrow plate, posteriorly with fused basal arms; hemisternites very slender, curved.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182C885FC46FC45" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182C885FE5EFC98" bold="true" box="[199,424,764,789]" pageId="2" pageNumber="418">Female terminalia.</emphasis>
Sternite VIII (
<figureCitation id="D19B1CA6FFFCFFBF130EC885FD78FC99" box="[587,654,764,789]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 9</figureCitation>
) with plate subtriangular, widest near base, 2.4–2.6 × as long as wide, apically sharply pointed and bearing two separate tufts of setae, with two longitudinal sclerites at apical three quarters; apodeme longer than sternite VIII, widest at midlength, terminating just at midlength of plate, at base enlarged to small, triangular translucent plate. Gonocoxites (
<figureCitation id="D19B1CA6FFFCFFBF120DC91EFC6DFC0D" box="[840,923,871,896]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 10</figureCitation>
) flat, long, tapered apically, with long apical styli bearing apical setae. Spermatheca (
<figureCitation id="D19B1CA6FFFCFFBF1322C9F5FD4FFC29" box="[615,697,908,933]" captionStart="FIGURES 2–11" captionStartId="5.[151,264,1690,1712]" captionTargetBox="[224,1370,263,1601]" captionTargetId="figure@5.[151,1436,193,1681]" captionTargetPageId="5" captionText="FIGURES 2–11. Structural details of Basothorhynchus endroedyi sp. n. 2. head with rostrum, dorsal view; 3. head with rostrum, lateral view; 4. tergite VII and VIII in male; 5. femur and tibia of anterior right leg; 6. abdominal ventrites of female; 7. penis, dorsal and lateral view; 8. tegmen, dorsal view; 9. Sternite VIII of female, dorsal view; 10. ovipositor, dorsal view; 11. spermatheca. Scale bars: 1 mm (6), 0.5 mm (2–5, 7–11)." httpUri="https://zenodo.org/record/2625552/files/figure.png" pageId="2" pageNumber="418">Fig. 11</figureCitation>
) U-shaped with regularly curved wide cornu, elongated corpus, short ramus (= gland entrance) and indistinct collum (= duct entrance).
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFCFFBF1182C9ADFD5FFBD5" pageId="2" pageNumber="418" type="etymology">
<paragraph id="491F0023FFFCFFBF1182C9ADFD5FFBD5" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182C9ADFEB8FC60" bold="true" box="[199,334,980,1005]" pageId="2" pageNumber="418">Etymology.</emphasis>
The new genus is derived from the word Basotho and Latin
<emphasis id="7BD4DC31FFFCFFBF1541C9ADFB9CFC61" box="[1028,1130,980,1004]" italics="true" pageId="2" pageNumber="418">rhynchus</emphasis>
. Basotho is a Bantu ethnic group whose ancestors have lived in southern Africa since around the fifth century. Most Basotho people today live in
<collectingCountry id="31B740B3FFFCFFBF11F1CE65FEB5FBB8" box="[180,323,1052,1077]" name="South Africa" pageId="2" pageNumber="418">South Africa</collectingCountry>
and
<collectingCountry id="31B740B3FFFCFFBF103CCE65FE25FBB8" box="[377,467,1052,1077]" name="Lesotho" pageId="2" pageNumber="418">Lesotho</collectingCountry>
(
<bibRefCitation id="2D317DD2FFFCFFBF10A7CE65FD36FBB9" author="Gardner, P. &amp; Scott, A. &amp; Rohan, M. S. &amp; Shackleton, A." box="[482,704,1052,1077]" pageId="2" pageNumber="418" refId="ref4510" refString="Gardner, P., Scott, A., Rohan, M. S. &amp; Shackleton, A. (1999) Encyklopedie Zemepis sveta [Czech edition of Encyklopedia of World Geography]. Columbus, Praha, 512 pp." type="book" year="1999">
Gardner
<emphasis id="7BD4DC31FFFCFFBF1303CE64FD80FBB9" box="[582,630,1052,1076]" italics="true" pageId="2" pageNumber="418">et al</emphasis>
. 1999
</bibRefCitation>
).
<emphasis id="7BD4DC31FFFCFFBF1391CE65FCB7FBB9" box="[724,833,1052,1076]" italics="true" pageId="2" pageNumber="418">Rhynchus</emphasis>
is the Latin form of the Greek word
<emphasis id="7BD4DC31FFFCFFBF1599CE65FAB4FBB9" box="[1244,1346,1052,1076]" italics="true" pageId="2" pageNumber="418">rhynkhos</emphasis>
, having a snout, bill, beak. The name is of male gender.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFCFFBF1182CE1DFAF2FB2D" pageId="2" pageNumber="418" type="distribution">
<paragraph id="491F0023FFFCFFBF1182CE1DFD6FFBF0" blockId="2.[151,1437,151,1977]" box="[199,665,1124,1149]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182CE1DFE96FBF0" bold="true" box="[199,352,1124,1149]" pageId="2" pageNumber="418">Distribution.</emphasis>
Known only from
<collectingCountry id="31B740B3FFFCFFBF1372CE1DFD63FBF0" box="[567,661,1124,1149]" name="Lesotho" pageId="2" pageNumber="418">Lesotho</collectingCountry>
.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CEFEFAF2FB2D" blockId="2.[151,1437,151,1977]" box="[199,1284,1159,1185]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182CEFEFE65FB2D" bold="true" box="[199,403,1159,1184]" pageId="2" pageNumber="418">Species included.</emphasis>
The genus is described as monotypic, including only the species
<taxonomicName id="8EA07BA0FFFCFFBF152ACEF0FB0BFB2D" authorityName="Borovec" authorityYear="2019" box="[1135,1277,1160,1184]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="endroedyi">
<emphasis id="7BD4DC31FFFCFFBF152ACEF0FB0BFB2D" box="[1135,1277,1160,1184]" italics="true" pageId="2" pageNumber="418">B. endroedyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFCFFBF1182CED5FCB7F834" pageId="2" pageNumber="418" type="description">
<paragraph id="491F0023FFFCFFBF1182CED5FB16FAF5" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
<emphasis id="7BD4DC31FFFCFFBF1182CED5FE4EFB48" bold="true" box="[199,440,1196,1221]" pageId="2" pageNumber="418">Taxonomic position.</emphasis>
The newly described genus belongs to the subfamily Entiminae, as defined by
<bibRefCitation id="2D317DD2FFFCFFBF1470CED5FED2FB65" author="Marvaldi, A. E. &amp; Lanteri, A. A. &amp; Rio, N. &amp; Oberprieler, R. G." pageId="2" pageNumber="418" pagination="503 - 522" refId="ref4634" refString="Marvaldi, A. E., Lanteri, A. A., Guadalupe del Rio, N. &amp; Oberprieler, R. G. (2014) 3.7. 5 Entiminae Schoenherr, 1823. In: Leschen, R. A. B. &amp; Beutel, R. G. (Eds.), Handbook of Zoology, Arthropoda: Insecta; Coleoptera, Beetles. Uol. 3. Morphology and Systematics (Phytophaga). Walter de Gruyter, Berlin / Boston, pp. 503 - 522." type="book chapter" year="2014">
Marvaldi
<emphasis id="7BD4DC31FFFCFFBF11D2CEA8FF3FFB65" box="[151,201,1232,1256]" italics="true" pageId="2" pageNumber="418">et al</emphasis>
. (2014)
</bibRefCitation>
, and here to the Oosomini, based on its free claws (this character distinguishes it from the
<taxonomicName id="8EA07BA0FFFCFFBF1466CEB6FEB0FA81" authority="Marshall, 1942" authorityName="Marshall" authorityYear="1942" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Embrithini">
Embrithini
<bibRefCitation id="2D317DD2FFFCFFBF11D2CE8DFEB0FA81" author="Marshall, G. A. K." box="[151,326,1268,1293]" pageId="2" pageNumber="418" pagination="1 - 26" refId="ref4591" refString="Marshall, G. A. K. (1942) On some East African Otiorrhynchinae (Col., Curcul.). Annals and Magazine of Natural History, Series 11, 9 (49), 1 - 26." type="journal article" year="1942">Marshall, 1942</bibRefCitation>
</taxonomicName>
), trisetose mandibles (distinguishing it from the
<taxonomicName id="8EA07BA0FFFCFFBF1228CE8DFB2AFA80" authority="Schoenherr, 1826" authorityName="Schoenherr" authorityYear="1826" box="[877,1244,1268,1293]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Otiorhynchini">Otiorhynchini Schoenherr, 1826</taxonomicName>
), dorsally placed scrobes (distinguishing it from the
<taxonomicName id="8EA07BA0FFFCFFBF136ACF6EFC62FABD" authority="Schoenherr, 1826" authorityName="Schoenherr" authorityYear="1826" box="[559,916,1303,1328]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Brachyderini">Brachyderini Schoenherr, 1826</taxonomicName>
and
<taxonomicName id="8EA07BA0FFFCFFBF1297CF6EFB15FABD" authority="Sharp, 1891" authorityName="Sharp" authorityYear="1891" box="[978,1251,1303,1328]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Sciaphilini">Sciaphilini Sharp, 1891</taxonomicName>
), lack of ocular lobes and vibrissae (distinguishing it from the Tanyrhynchini Schoenherr, 1826 and
<taxonomicName id="8EA07BA0FFFCFFBF1507CF45FA79FAD8" authority="Lacordaire, 1863" authorityName="Lacordaire" authorityYear="1863" box="[1090,1423,1340,1365]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Tanymecini">Tanymecini Lacordaire, 1863</taxonomicName>
), and lack of a protruding humeral callus (distinguishing it from the
<taxonomicName id="8EA07BA0FFFCFFBF12C7CF26FB25FAF5" authority="Schoenherr, 1823" authorityName="Schoenherr" authorityYear="1823" box="[898,1235,1375,1400]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Polydrusini">Polydrusini Schoenherr, 1823</taxonomicName>
).
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CFFDFCC0F985" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
Oosomini were never redefined after
<bibRefCitation id="2D317DD2FFFCFFBF132CCFFDFCE8FA11" author="Marshall, G. A. K." box="[617,798,1412,1437]" pageId="2" pageNumber="418" pagination="1 - 26" refId="ref4591" refString="Marshall, G. A. K. (1942) On some East African Otiorrhynchinae (Col., Curcul.). Annals and Magazine of Natural History, Series 11, 9 (49), 1 - 26." type="journal article" year="1942">Marshall (1942)</bibRefCitation>
split them into Oosomini with free claws and
<taxonomicName id="8EA07BA0FFFCFFBF1466CFFDFA6AFA10" authorityName="Marshall" authorityYear="1942" box="[1315,1436,1412,1437]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="tribe" tribe="Embrithini">Embrithini</taxonomicName>
with fused claws. This tribe does not seem to be monophyletic and a further study is much needed. Oosomini contains fewer genera than is listed in
<bibRefCitation id="2D317DD2FFFCFFBF1302CFB5FC46FA69" author="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C." box="[583,944,1484,1509]" pageId="2" pageNumber="418" refId="ref4398" refString="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C. (1999) A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) (excepting Scolytidae and Platypodidae). Entomopraxis, S. C. P., Barcelona, 315 pp." type="book" year="1999">Alonso-Zarazaga &amp; Lyal (1999)</bibRefCitation>
because some genera were later transferred to other tribes (for example
<bibRefCitation id="2D317DD2FFFCFFBF1095CF88FCC5F985" author="Borovec, R. &amp; Oberprieler, R. G." box="[464,819,1519,1544]" pageId="2" pageNumber="418" pagination="365 - 378" refId="ref4451" refString="Borovec, R. &amp; Oberprieler, R. G. (2013) Afrophloeus, a new genus of African weevils of the tribe Embrithini (Coleoptera: Curculionidae: Entiminae), with description of a new species and notes on the composition of Embrithini. Zootaxa, 3693 (3), 365 - 378." type="journal article" year="2013">Borovec &amp; Oberprieler (2013))</bibRefCitation>
.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CC6DFB8CF8C0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
Endemic to
<collectingCountry id="31B740B3FFFCFFBF1012CC6DFE18F9A0" box="[343,494,1556,1581]" name="South Africa" pageId="2" pageNumber="418">South Africa</collectingCountry>
,
<taxonomicName id="8EA07BA0FFFCFFBF10BACC6DFD37F9A1" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[511,705,1556,1580]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFCFFBF10BACC6DFD37F9A1" box="[511,705,1556,1580]" italics="true" pageId="2" pageNumber="418">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFCFFBF1389CC6DFCD6F9A0" bold="true" box="[716,800,1556,1581]" pageId="2" pageNumber="418">
<taxonomicNameLabel id="60E7614AFFFCFFBF1389CC6DFCD6F9A0" box="[716,800,1556,1581]" pageId="2" pageNumber="418" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from genera of Oosomini comprising large-sized species by the following characters: small size (maximum length 3.0 mm), rostrum posteriorly continuous with head, base of epifrons as wide as space width between anterior margins of eyes, epistome slightly emarginate, scape not reaching anterior border of pronotum in repose, dentate femora, tibiae short and robust, abruptly enlarged to inner side at basal third and body glabrous lacking appressed vestiture. Other genera of Oosomini have larger species (over
<quantity id="8E58ADC6FFFCFFBF13F3CCBEFCE6F96D" box="[694,784,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="418" unit="mm" value="5.8">5.8 mm</quantity>
long), covered by appressed scales, rostrum posteriorly separated from head by a transverse sulcus or shallow groove, base of epifrons distinctly narrower than width between anterior margins of eyes, epistome conspicuous (except in
<taxonomicName id="8EA07BA0FFFCFFBF12C9CD69FBF4F8A5" authorityName="Schönherr" authorityYear="1826" box="[908,1026,1808,1832]" class="Insecta" family="Curculionidae" genus="Phlyctinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFCFFBF12C9CD69FBF4F8A5" box="[908,1026,1808,1832]" italics="true" pageId="2" pageNumber="418">Phlyctinus</emphasis>
</taxonomicName>
), scape in repose exceeding anterior border of pronotum, adentate femora, tibiae long, slender, slightly sinuose on inner edge.
</paragraph>
<paragraph id="491F0023FFFCFFBF1182CD20FCB7F834" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="418">
By its small size,
<taxonomicName id="8EA07BA0FFFCFFBF10CBCD21FDA6F8FD" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[398,592,1880,1904]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFCFFBF10CBCD21FDA6F8FD" box="[398,592,1880,1904]" italics="true" pageId="2" pageNumber="418">Basothorhynchus</emphasis>
</taxonomicName>
can be compared only with three Oosomini genera:
<taxonomicName id="8EA07BA0FFFCFFBF15DBCD21FF24F818" authority="Marshall, 1953" authorityName="Marshall" authorityYear="1953" class="Insecta" family="Curculionidae" genus="Holcolaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFCFFBF15DBCD21FADCF8FD" box="[1182,1322,1880,1904]" italics="true" pageId="2" pageNumber="418">Holcolaccus</emphasis>
Marshall, 1953
</taxonomicName>
,
<taxonomicName id="8EA07BA0FFFCFFBF11A5CD05FDEFF818" authority="Schoenherr, 1826" authorityName="Schoenherr" authorityYear="1826" box="[224,537,1916,1941]" class="Insecta" family="Curculionidae" genus="Oosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFCFFBF11A5CD05FEBEF81E" box="[224,328,1916,1939]" italics="true" pageId="2" pageNumber="418">Oosomus</emphasis>
Schoenherr, 1826
</taxonomicName>
and
<taxonomicName id="8EA07BA0FFFCFFBF1310CD05FC70F818" authority="Marshall, 1926" authorityName="Marshall" authorityYear="1926" box="[597,902,1916,1941]" class="Insecta" family="Curculionidae" genus="Periderces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFCFFBF1310CD05FD3BF819" box="[597,717,1916,1940]" italics="true" pageId="2" pageNumber="418">Periderces</emphasis>
Marshall, 1926
</taxonomicName>
. From all these three genera
<taxonomicName id="8EA07BA0FFFCFFBF159FCD05FA6AF819" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[1242,1436,1916,1940]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFCFFBF159FCD05FA6AF819" box="[1242,1436,1916,1940]" italics="true" pageId="2" pageNumber="418">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFCFFBF11D2CDD9FF10F834" bold="true" box="[151,230,1952,1977]" pageId="2" pageNumber="418">
<taxonomicNameLabel id="60E7614AFFFCFFBF11D2CDD9FF10F834" box="[151,230,1952,1977]" pageId="2" pageNumber="418" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
may be distinguished according to the following key:
</paragraph>
</subSubSection>
<subSubSection id="01BA53A8FFFDFFBE11D2CAEEFD69FBFD" pageId="3" pageNumber="419" type="key">
<key id="3D315712FFFDFFBE11D2CAEEFA6DFD0F" pageId="3" pageNumber="419">
<keyStep id="F2541886FFFDFFBE11D2CAEEFA6DFED1" pageId="3" pageNumber="419">
<paragraph id="491F0023FFFDFFBE11D2CAEEFA6DFE89" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2CAEEFA6DFE89" pageId="3" pageNumber="419">
1. Rostrum at least 1.5 × as long as wide, widest at apical portion with sides rounded around scrobes. Metatibial apex with wide, sparsely squamose corbels. Sternite VIII of females with plate wider than long, apodeme 3.3–6.2 × longer than plate. Size
<quantity id="8E58ADC6FFFDFFBE1434CACCFED8FF6A" metricMagnitude="-3" metricUnit="m" metricValue="3.6999999999999997" metricValueMax="5.1" metricValueMin="2.3" pageId="3" pageNumber="419" unit="mm" value="3.6999999999999997" valueMax="5.1" valueMin="2.3">2.3– 5.1 mm</quantity>
...................................................................................................................................................................................
<taxonomicName id="8EA07BA0FFFDFFBE158ACA96FA6DFE89" ID-CoL="8HNFV" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1826" box="[1231,1435,239,261]" class="Insecta" family="Curculionidae" genus="Oosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE158ACA96FADEFE88" box="[1231,1320,239,261]" italics="true" pageId="3" pageNumber="419">Oosomus</emphasis>
Schoenherr
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491F0023FFFDFFBE11D2CB74FA6DFED1" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2CB74FA6DFED1" pageId="3" pageNumber="419">- Rostrum wider than long or at most 1.2 × as long as wide, widest at base and tapered anteriad, not rounded around scrobes. Metatibial apex simple (= without corbels). Sternite VIII of females with plate distinctly longer than wide, apodeme short, about same length as plate............................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="F2541886FFFDFFBE11D2CB1CFA6DFD81" pageId="3" pageNumber="419">
<paragraph id="491F0023FFFDFFBE11D2CB1CFA6AFE38" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2CB1CFA6AFE38" pageId="3" pageNumber="419">
2. Body dorsally and ventrally glabrous. Scrobes dorsally conspicuous, pit-shaped, laterally directed towards eyes. Antennal scape short and robust, 4.9–5.3 × as long as wide at apex. Tibiae slender at basal third and then abruptly enlarged. Size 2.3–3.0 mm..............................................................................
<taxonomicName id="8EA07BA0FFFDFFBE15E9CBE6FAA4FE38" ID-CoL="8KGBR" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[1196,1362,415,437]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFDFFBE15E9CBE6FAA4FE38" box="[1196,1362,415,437]" italics="true" pageId="3" pageNumber="419">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFDFFBE141DCBE6FA6AFE38" bold="true" box="[1368,1436,415,437]" pageId="3" pageNumber="419">
<taxonomicNameLabel id="60E7614AFFFDFFBE141DCBE6FA6AFE38" box="[1368,1436,415,437]" pageId="3" pageNumber="419" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="491F0023FFFDFFBE11D2CBC4FA6DFD81" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2CBC4FA6DFD81" pageId="3" pageNumber="419">- Body dorsally and ventrally squamose. Scrobes not visible from above, laterally not continuing behind antennal insertion or curved downwords. Antennal scape long and slender, 9.1–10.7 × as long as wide at apex. Tibiae slender and long, slightly sinuate at inner side...................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="F2541886FFFDFFBE11D2C86CFA6DFD0F" pageId="3" pageNumber="419">
<paragraph id="491F0023FFFDFFBE11D2C86CFA6DFDCA" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2C86CFA6DFDCA" pageId="3" pageNumber="419">
3. Scrobes very narrow and not continuing backwards. Antennal scape shortly exceeding anterior border of pronotum. Epifrons narrow (about as wide as half rostral width). Femora adentate. Size
<quantity id="8E58ADC6FFFDFFBE122CC84BFC2AFDCA" box="[873,988,562,583]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.8" metricValueMin="2.5" pageId="3" pageNumber="419" unit="mm" value="3.15" valueMax="3.8" valueMin="2.5">2.5–3.8 mm</quantity>
...................
<taxonomicName id="8EA07BA0FFFDFFBE158DC848FA6DFDCA" ID-CoL="8HKKB" authority="Marshall" authorityName="Marshall" authorityYear="1953" box="[1224,1435,561,583]" class="Insecta" family="Curculionidae" genus="Holcolaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE158DC848FAB6FDCA" box="[1224,1344,561,583]" italics="true" pageId="3" pageNumber="419">Holcolaccus</emphasis>
Marshall
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491F0023FFFDFFBE11D2C836FA6DFD0F" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="419">
<keyLead id="F251A316FFFDFFBE11D2C836FA6DFD0F" pageId="3" pageNumber="419">
- Scrobes continuing backwards and curved downwards. Antennal scapes not reaching posterior margin of eyes. Epifrons wide (occuping most rostral surface). Femora with large tooth. Size 4.0–
<quantity id="8E58ADC6FFFDFFBE1221C814FC5AFD0F" box="[868,940,621,642]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="419" unit="mm" value="4.8">4.8 mm</quantity>
.........................
<taxonomicName id="8EA07BA0FFFDFFBE1591C815FA6DFD0F" ID-CoL="8HP8F" authority="Marshall" authorityName="Marshall" authorityYear="1926" box="[1236,1435,620,642]" class="Insecta" family="Curculionidae" genus="Periderces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE1591C815FAC9FD0F" box="[1236,1343,620,642]" italics="true" pageId="3" pageNumber="419">Periderces</emphasis>
Marshall
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="491F0023FFFDFFBE1182C8D1FD69FBFD" blockId="3.[151,1436,679,1136]" pageId="3" pageNumber="419">
By its general appearance, small size and glabrous, cylindrical body,
<taxonomicName id="8EA07BA0FFFDFFBE12A8C8D1FB59FD4D" ID-CoL="8KGBR" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[1005,1199,680,704]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFDFFBE12A8C8D1FB59FD4D" box="[1005,1199,680,704]" italics="true" pageId="3" pageNumber="419">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFDFFBE15FCC8DEFAFDFD4D" bold="true" box="[1209,1291,679,704]" pageId="3" pageNumber="419">
<taxonomicNameLabel id="60E7614AFFFDFFBE15FCC8DEFAFDFD4D" box="[1209,1291,679,704]" pageId="3" pageNumber="419" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
resembles a group of genera known from high elevation mountains of
<collectingCountry id="31B740B3FFFDFFBE1235C8B4FC4AFD68" box="[880,956,717,741]" name="Kenya" pageId="3" pageNumber="419">Kenya</collectingCountry>
, such as
<taxonomicName id="8EA07BA0FFFDFFBE157FC8B5FA6EFD69" ID-CoL="8HBWB" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[1082,1432,716,741]" class="Insecta" family="Curculionidae" genus="Abarypeithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE157FC8B5FB38FD69" box="[1082,1230,716,740]" italics="true" pageId="3" pageNumber="419">Abarypeithes</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE1598C8B5FA6EFD69" author="Hustache, A." box="[1245,1432,716,741]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8EA07BA0FFFDFFBE11D2C889FDB2FC85" ID-CoL="8HCBJ" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[151,580,751,776]" class="Insecta" family="Curculionidae" genus="Afrotroglorrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE11D2C889FE8AFC85" box="[151,380,752,776]" italics="true" pageId="3" pageNumber="419">Afrotroglorrhynchus</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE10CEC896FDB2FC85" author="Hustache, A." box="[395,580,751,776]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8EA07BA0FFFDFFBE1313C889FC35FC85" ID-CoL="8HKK6" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[598,963,751,776]" class="Insecta" family="Curculionidae" genus="Hobarypeithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE1313C889FD0DFC85" box="[598,763,752,776]" italics="true" pageId="3" pageNumber="419">Hobarypeithes</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE124FC896FC35FC85" author="Hustache, A." box="[778,963,751,776]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8EA07BA0FFFDFFBE1293C889FA93FC85" ID-CoL="8HNGZ" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[982,1381,751,776]" class="Insecta" family="Curculionidae" genus="Opseobarypeithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE1293C889FB56FC85" box="[982,1184,752,776]" italics="true" pageId="3" pageNumber="419">Opseobarypeithes</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE15EAC896FA93FC85" author="Hustache, A." box="[1199,1381,751,776]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8EA07BA0FFFDFFBE11D2C96DFE4DFCA1" ID-CoL="8HNQF" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[151,443,788,813]" class="Insecta" family="Curculionidae" genus="Palaepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE11D2C96DFF09FCA1" box="[151,255,788,812]" italics="true" pageId="3" pageNumber="419">Palaepus</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE104DC96DFE4DFCA1" author="Hustache, A." box="[264,443,788,813]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
. The first genus is listed in
<taxonomicName id="8EA07BA0FFFDFFBE13BEC96DFBFEFCA1" ID-CoL="8JS3N" authority="Sharp, 1891" authorityName="Sharp" authorityYear="1891" box="[763,1032,788,813]" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="tribe" tribe="Sciaphilini">Sciaphilini Sharp, 1891</taxonomicName>
, the others in
<taxonomicName id="8EA07BA0FFFDFFBE15E9C96DFF24FCDD" ID-CoL="8JRZV" authority="Lacordaire, 1863" authorityName="Lacordaire" authorityYear="1863" class="Insecta" family="Curculionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="tribe" tribe="Peritelini">Peritelini Lacordaire, 1863</taxonomicName>
. From all these genera
<taxonomicName id="8EA07BA0FFFDFFBE1096C941FD63FCDD" ID-CoL="8KGBR" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[467,661,824,848]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFDFFBE1096C941FD63FCDD" box="[467,661,824,848]" italics="true" pageId="3" pageNumber="419">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFDFFBE13DEC94EFD1FFCDD" bold="true" box="[667,745,823,848]" pageId="3" pageNumber="419">
<taxonomicNameLabel id="60E7614AFFFDFFBE13DEC94EFD1FFCDD" box="[667,745,823,848]" pageId="3" pageNumber="419" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
can be easily distinguished not only by the free claws (it must be pointed out that some of these Kenyan genera have claws connected at their short bases), but mainly by the rostrum lacking laterally prominent pterygiae, the short scape, femora with a distinct tooth, the inner side of the tibiae sinuose and denticulate, and tibiae mucronate. In contrast, all genera from equatorial mountains have a rostrum with laterally prominent pterygiae, long scapes, adentate femora, inner sides of tibiae straight and not denticulate and tibiae amucronate.
<taxonomicName id="8EA07BA0FFFDFFBE1365C995FD14FB89" ID-CoL="8KGBR" authority="Borovec, 2019" authorityName="Borovec" authorityYear="2019" box="[544,738,1004,1028]" class="Insecta" family="Curculionidae" genus="Basothorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7BD4DC31FFFDFFBE1365C995FD14FB89" box="[544,738,1004,1028]" italics="true" pageId="3" pageNumber="419">Basothorhynchus</emphasis>
</taxonomicName>
<emphasis id="7BD4DC31FFFDFFBE13ACC995FCCFFB88" bold="true" box="[745,825,1004,1029]" pageId="3" pageNumber="419">
<taxonomicNameLabel id="60E7614AFFFDFFBE13ACC995FCCFFB88" box="[745,825,1004,1029]" pageId="3" pageNumber="419" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
is most similar to the genus
<taxonomicName id="8EA07BA0FFFDFFBE1538C994FA6EFB89" ID-CoL="8HMXX" authority="Hustache, 1939" authorityName="Hustache" authorityYear="1939" box="[1149,1432,1004,1029]" class="Insecta" family="Curculionidae" genus="Neomias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE1538C994FB29FB89" box="[1149,1247,1005,1028]" italics="true" pageId="3" pageNumber="419">Neomias</emphasis>
<bibRefCitation id="2D317DD2FFFDFFBE15A3C995FA6EFB89" author="Hustache, A." box="[1254,1432,1004,1029]" pageId="3" pageNumber="419" pagination="175 - 272" refId="ref4554" refString="Hustache, A. (1939) Mission scientifique de l'Omo, Coleoptera. XX. Curculionidae. Memoires du Museum national d'histoire naturelle, New Series, 9, 175 - 272, 1 pl." type="journal article" year="1939">Hustache, 1939</bibRefCitation>
</taxonomicName>
, known from the mountains of
<collectingCountry id="31B740B3FFFDFFBE10ABCE76FDA6FBA5" box="[494,592,1039,1064]" name="Ethiopia" pageId="3" pageNumber="419">Ethiopia</collectingCountry>
,
<collectingCountry id="31B740B3FFFDFFBE1319CE69FD5EFBA5" box="[604,680,1040,1064]" name="Kenya" pageId="3" pageNumber="419">Kenya</collectingCountry>
,
<collectingCountry id="31B740B3FFFDFFBE13F1CE76FCFDFBA5" box="[692,779,1039,1064]" name="Uganda" pageId="3" pageNumber="419">Uganda</collectingCountry>
and
<collectingCountry id="31B740B3FFFDFFBE1206CE76FC5DFBA5" box="[835,939,1039,1064]" name="Tanzania" pageId="3" pageNumber="419">Tanzania</collectingCountry>
. This genus also has a rostrum with straight sides in dorsal view, without distinct pterygiae. But in all other characters
<taxonomicName id="8EA07BA0FFFDFFBE129ECE4CFBCBFBC1" ID-CoL="8HMXX" authorityName="Hustache" authorityYear="1939" box="[987,1085,1077,1100]" class="Insecta" family="Curculionidae" genus="Neomias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE129ECE4CFBCBFBC1" box="[987,1085,1077,1100]" italics="true" pageId="3" pageNumber="419">Neomias</emphasis>
</taxonomicName>
shares the same distinguishing characters as
<taxonomicName id="8EA07BA0FFFDFFBE1068CE21FE37FBFD" ID-CoL="8HBWB" authorityName="Hustache" authorityYear="1939" box="[301,449,1112,1136]" class="Insecta" family="Curculionidae" genus="Abarypeithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="7BD4DC31FFFDFFBE1068CE21FE37FBFD" box="[301,449,1112,1136]" italics="true" pageId="3" pageNumber="419">Abarypeithes</emphasis>
</taxonomicName>
and similar genera.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Wilson, Trevor C.</mods:namePart>
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<treatment ID-GBIF-Taxon="159244365" LSID="urn:lsid:plazi:treatment:C109F0C89FC154B29578E07A2ADE6287" httpUri="http://treatment.plazi.org/id/C109F0C89FC154B29578E07A2ADE6287" lastPageNumber="16" pageId="15" pageNumber="16">
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<taxonomicName LSID="C109F0C8-9FC1-54B2-9578-E07A2ADE6287" authority="(Gamble) A. J. Paton" authorityName="Paton &amp; Mwanyambo &amp; Govaerts &amp; Smitha &amp; Suddee &amp; Phillipson &amp; Wilson &amp; Forster &amp; Culham" authorityYear="2019" baseAuthorityName="Gamble" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus argenteus" order="Lamiales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="argenteus" status="comb. nov.">Coleus argenteus (Gamble) A.J.Paton</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="16">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="reference_group">
<paragraph pageId="15" pageNumber="16">
<taxonomicName authorityName="Gamble" authorityYear="1924" class="Magnoliopsida" family="Lamiaceae" genus="Anisochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anisochilus argenteus" order="Lamiales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="argenteus">Anisochilus argenteus</taxonomicName>
Gamble, Fl. Madras: 1127. 1924. Type: India, Madras, Pulney Hills, Kodaikanal Ghat new road, 29 Dec. 1898, Bourne 1441C (lectotype: K; isolectotype: K, designated by Suddee &amp; Paton (2009).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="distribution">
<paragraph pageId="15" pageNumber="16">Distribution.</paragraph>
<paragraph pageId="15" pageNumber="16">S. India.</paragraph>
</subSubSection>
</treatment>
</document>