Add updates up until 2024-08-26 14:55:35

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<document id="D4DD69CF7B114D03A8F6D5D65121BBF5" ID-DOI="10.3897/zookeys.1210.125963" ID-ZooBank="150A50A5-7C0A-41E1-B983-08517F57F7E7" ID-publisher-id="125963" URI-arpha="33525710-9122-55AF-87C2-83F6E31A224B" URI-zoobank="http://zoobank.org/150A50A5-7C0A-41E1-B983-08517F57F7E7" article-type="research-article" checkinTime="1724683991666" checkinUser="pensoft" docAuthor="Jin, Qing, Li, Jiali, Jiang, Jibao &amp; Qiu, Jiangping" docDate="2024" docId="21FE19A90A4251EF9FC664467170CB31" docLanguage="en" docName="ZooKeys 1210: 247-271" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/125963/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Amynthas taoyuanensis Qiu &amp; Jin 2024, sp. nov." docType="treatment" docUuid="99EAAAC1-F761-4082-A96E-645F864F631E" docUuidSource="ZooBank" docVersion="1" dtd-version="3.0" lastPageNumber="271" masterDocId="33525710912255AF87C283F6E31A224B" masterDocTitle="Four new earthworm species of the genera Amynthas and Metaphire (Oligochaeta, Megascolecidae) from Hunan and Anhui provinces, China" masterLastPageNumber="271" masterPageNumber="247" pageNumber="247" updateTime="1724684060960" updateUser="pensoft">
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<mods:title id="5ADBCC30E5C26721CD285AF289021C1D">Four new earthworm species of the genera Amynthas and Metaphire (Oligochaeta, Megascolecidae) from Hunan and Anhui provinces, China</mods:title>
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<mods:name id="93C8A6E3596C9F28B1BBE4D44A784D8B" type="personal">
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<mods:namePart id="72F78D8E02183A740EA7265F3805B2E5">Jin, Qing</mods:namePart>
<mods:nameIdentifier id="3E07E9A59BB860D9146BA954B6F29F08" type="ORCID">0000-0001-7731-3150</mods:nameIdentifier>
<mods:affiliation id="A01BD85CC1A0F7146AFDB73CE1497DB0">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China &amp; Shanghai Urban Forest Research Station, State Forestry Administration, Shanghai, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="F33AF134590BCDF3DC88CDED9D49026D">Li, Jiali</mods:namePart>
<mods:affiliation id="916BC0C763769CA90E8031A8F4258F8C">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China &amp; Shanghai Urban Forest Research Station, State Forestry Administration, Shanghai, China</mods:affiliation>
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<mods:namePart id="23F9443B98B57A0664D8FFE881DC5459">Jiang, Jibao</mods:namePart>
<mods:affiliation id="1A570BCD35F8A1E0941D14CA0AEB7D98">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China &amp; Shanghai Urban Forest Research Station, State Forestry Administration, Shanghai, China</mods:affiliation>
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<mods:namePart id="C41A788E190588D9B260BA76FA14B94E">Qiu, Jiangping</mods:namePart>
<mods:affiliation id="8BCFC706EFD9B7345A1BE1434307129F">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China &amp; Shanghai Urban Forest Research Station, State Forestry Administration, Shanghai, China</mods:affiliation>
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<mods:date id="1C7C15C24CBB566CF35ABE385DDC6D92">2024</mods:date>
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<subSubSection id="F93D9E3F5718DC2AA4BBC3CC388EB991" type="nomenclature">
<paragraph id="5FB2B89B53C25D0EDF2360FBF66A725E">
<taxonomicName id="760265D90C9227D043EF343AC7CFC61A" authority="Qiu &amp; Jin" authorityName="Qiu &amp; Jin" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov." subClass="Oligochaeta">
<emphasis id="8BE65BB1101369D97D7362DA483894F6" italics="true">Amynthas taoyuanensis</emphasis>
Qiu &amp; Jin
</taxonomicName>
<taxonomicNameLabel id="BE71FB18DA4C230E5B888F6C27DD11C2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6A7F793EA7C7532485F84D2842C1B111" type="description">
<paragraph id="2950B6F21B2726A42BE6FF9D2F1AE5CA">
<figureCitation id="3F5BDE63C107ECE8970BE8510413B936" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG6AG" type="material">
<paragraph id="5E407F05051C36821DB6E484716A77AF">
<heading id="C3BBCD8229D721D2E798316DD5EFC1A4" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="BFE01A869DE1B521923DCB8395D40FBC">
<materialsCitation id="1CACA107A7E962107C7ABDCCEB2740E8" collectingDate="2019-05-17" collectorName="JB Jiang &amp; JL Li &amp; BY Yin" country="China" elevation="201" latitude="25.3017" location="Yongzhou City" longLatPrecision="1" longitude="111.63432" specimenCount="1" stateProvince="Hunan" typeStatus="holotype">
<emphasis id="44299DFF079FDE466413736356BC8A26" bold="true">
<emphasis id="21EDBF2A9522E967130FD188F96CBEF4" bold="true" italics="true">
<typeStatus id="1AC07A5353A701D02E64178A00F727B1" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
• 1 clitellate (P 1 CJHUSH 190517069 N 8-01 A),
<collectingCountry id="CDE677A5E50607D9DB44EA1984FA4ECA" name="China">China</collectingCountry>
,
<collectingRegion id="7FAD442841269574CE215C21A0EE8B56" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="E9B84D1EBCA0291766D14143609AAE43" LSID="urn:lsid:plazi:treatment:21FE19A90A4251EF9FC664467170CB31:E9B84D1EBCA0291766D14143609AAE43" country="China" latitude="25.3017" longLatPrecision="1" longitude="111.63432" name="Yongzhou City" stateProvince="Hunan">Yongzhou City</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EU6AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.634320,25.301700]}">
<geoCoordinate id="51C551B471C44EE774E1AA2FA95385D8" degrees="25.30170" direction="north" orientation="latitude" precision="1" value="25.3017">25.30170 ° N</geoCoordinate>
,
<geoCoordinate id="22390526116B706942DD64956CCEB952" degrees="111.63432" direction="east" orientation="longitude" precision="1" value="111.63432">111.63432 ° E</geoCoordinate>
</named-content>
),
<quantity id="5E4B7F5FA04FFF0D267CE176ED876266" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">
<elevation id="A75D5AF7CD7ADC50BE6CCCE3000530DE" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">201 m</elevation>
</quantity>
elevation, brown soil under weeds in dry farmland,
<date id="7D59D6C309DA69A7ACDF4F4262E6881E" value="2019-05-17">
<collectingDate id="233B70448760388ED8C0A99A613F4776" value="2019-05-17">17 May 2019</collectingDate>
</date>
,
<collectorName id="AD4DDE9DF116473DB5EE484A67E7B25E">JB Jiang</collectorName>
,
<collectorName id="D540DAEA17E366D8B8F9F4772387C68A">JL Li</collectorName>
and
<collectorName id="3A8C8FD2A9BEA10BD42DC35D03C5CC77">BY Yin</collectorName>
</materialsCitation>
.
<emphasis id="4D5DF965DBEBA8473AF169335A16EEEB" bold="true">
<emphasis id="83647BAC8A411D256868BDE67CAF71D0" bold="true" italics="true">
<typeStatus id="485915F9C49DFA16A6FE7AD7C64C77CF" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
7 clitellates in total •
<materialsCitation id="9F75E341C3D1A3EC3FB5CFF4696EC0F2" collectingDate="2019-05-17" collectorName="JB Jiang &amp; JL Li &amp; Yin" country="China" elevation="201" latitude="25.3017" location="Yongzhou City" longLatPrecision="1" longitude="111.63432" specimenCount="1" stateProvince="Hunan" typeStatus="paratype">
3 clitellates (P 1 CJHUSH 190517069 N 8-01 B),
<collectingCountry id="D52AFB808D41E8E1C01255F7B9AAAF76" name="China">China</collectingCountry>
,
<collectingRegion id="036BC9AFDCAADD01DE1C72FAAABDB49E" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="7417C43AD5EDA2FAFA202BDDB77B960E" LSID="urn:lsid:plazi:treatment:21FE19A90A4251EF9FC664467170CB31:7417C43AD5EDA2FAFA202BDDB77B960E" country="China" latitude="25.3017" longLatPrecision="1" longitude="111.63432" name="Yongzhou City" stateProvince="Hunan">Yongzhou City</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.634320,25.301700]}">
<geoCoordinate id="46B0369622F50984FF68AAF5DF85EF12" degrees="25.30170" direction="north" orientation="latitude" precision="1" value="25.3017">25.30170 ° N</geoCoordinate>
,
<geoCoordinate id="6B2D218DC30CCB8403933E005FF67710" degrees="111.63432" direction="east" orientation="longitude" precision="1" value="111.63432">111.63432 ° E</geoCoordinate>
</named-content>
),
<quantity id="4FC53D541A8983F661C7AF82488887E5" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">
<elevation id="69669C274912AF0EF8F4B32FF6C5AAAE" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">201 m</elevation>
</quantity>
elevation, brown soil under weeds in dry farmland,
<date id="1C59A837FC2EF2491501DA2DC231D17F" value="2019-05-17">
<collectingDate id="906568EE4E5CE55663C0EB6731984687" value="2019-05-17">17 May 2019</collectingDate>
</date>
,
<collectorName id="3829E4738770EA7CE576BBC7E37FF7B8">JB Jiang</collectorName>
,
<collectorName id="3865F389820A1AB0872134E9CCB4D013">JL Li</collectorName>
and BY
<collectorName id="D12D5F76C81ECB4C35FEC3992EB0802C">Yin</collectorName>
</materialsCitation>
<materialsCitation id="761A137A507342A88105ACDCB2280FA6" collectingDate="2019-05-26" collectionCode="R" collectorName="JB Jiang &amp; JL Li &amp; Wang" country="China" elevation="55" latitude="27.63491" location="Liling City" longLatPrecision="1" longitude="113.54768" specimenCode="R 8-04" specimenCount="1" stateProvince="Hunan" typeStatus="paratype">
1 clitellates (P 1 CJHUSH 190526781
<specimenCode id="A909D3BE55027EF84B1D35EBE4A7DBB2" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile">R 8-04</specimenCode>
),
<collectingCountry id="9A33512A3A791FD43C977EA90F3CD72B" name="China">China</collectingCountry>
,
<collectingRegion id="56EE5AAECAD163B15BBE193692F987E7" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="EA3108E400F48470D1BBA69AE9D692E6" LSID="urn:lsid:plazi:treatment:21FE19A90A4251EF9FC664467170CB31:EA3108E400F48470D1BBA69AE9D692E6" country="China" latitude="27.63491" longLatPrecision="1" longitude="113.54768" name="Liling City" stateProvince="Hunan">Liling City</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[113.547680,27.634910]}">
<geoCoordinate id="7B78AC37A16B6ED43E795A2157DCBE08" degrees="27.63491" direction="north" orientation="latitude" precision="1" value="27.63491">27.63491 ° N</geoCoordinate>
,
<geoCoordinate id="992ACD24F87BEF9649953BEAB2017B75" degrees="113.54768" direction="east" orientation="longitude" precision="1" value="113.54768">113.54768 ° E</geoCoordinate>
</named-content>
),
<quantity id="2A06482AE02B3B71627D29BA04111A7C" metricMagnitude="1" metricUnit="m" metricValue="5.5" unit="m" value="55.0">
<elevation id="3072FC2DEE391AB34BF745A458496BC8" metricMagnitude="1" metricUnit="m" metricValue="5.5" unit="m" value="55.0">55 m</elevation>
</quantity>
elevation, yellow soil under weeds in woodland,
<date id="7ED1814DA132633BEE6FC635F5529C64" value="2019-05-26">
<collectingDate id="0AE388044C06879686F1DB297BAEA321" value="2019-05-26">26 May 2019</collectingDate>
</date>
,
<collectorName id="1B07141E9B2EF3E30AE8DA4556F76882">JB Jiang</collectorName>
,
<collectorName id="082C48AD117905F9BA802A5BAEC649BA">JL Li</collectorName>
and Y
<collectorName id="EC6260BE9A881F65AFEC94CDF6970034">Wang</collectorName>
</materialsCitation>
<materialsCitation id="5D41BA7DA2E25BA0C4751B372A489ECC" collectingDate="2016-05-02" collectorName="JB Jiang &amp; J Sun &amp; Dong &amp; Zheng" country="China" elevation="66" latitude="28.70779" location="Changde City" longLatPrecision="1" longitude="111.38225" specimenCount="1" stateProvince="Hunan" typeStatus="paratype">
2 clitellates (HU 201601-03),
<collectingCountry id="F0FE60A74670C45B9DFE988B5967A062" name="China">China</collectingCountry>
,
<collectingRegion id="9B1F62FAD4651108DCFB6360E852FF8F" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="F4ACE6F68E2FAB03CDC7D9B2CCADB63D" LSID="urn:lsid:plazi:treatment:21FE19A90A4251EF9FC664467170CB31:F4ACE6F68E2FAB03CDC7D9B2CCADB63D" country="China" latitude="28.70779" longLatPrecision="1" longitude="111.38225" name="Changde City" stateProvince="Hunan">Changde City</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.382250,28.707790]}">
<geoCoordinate id="E1D60CBE08D48B66CAF5C2247432F40E" degrees="28.70779" direction="north" orientation="latitude" precision="1" value="28.70779">28.70779 ° N</geoCoordinate>
,
<geoCoordinate id="F7D7326D32AE8570FD12921EC374A403" degrees="111.38225" direction="east" orientation="longitude" precision="1" value="111.38225">111.38225 ° E</geoCoordinate>
</named-content>
),
<quantity id="C1C97831FAC1380AA45D423A8BAEC3BE" metricMagnitude="1" metricUnit="m" metricValue="6.6" unit="m" value="66.0">
<elevation id="3E7130BA18238587B3584FC7A4E3D6B5" metricMagnitude="1" metricUnit="m" metricValue="6.6" unit="m" value="66.0">66 m</elevation>
</quantity>
elevation, brown soil under moss and fern in farmland,
<date id="A770DD85317A3240021A189666B6A7EE" value="2016-05-02">
<collectingDate id="C42D544E57C854076579D396D9E9B9D3" value="2016-05-02">2 May 2016</collectingDate>
</date>
,
<collectorName id="5241BC41732EBC7E0B4EAFC7A11A4281">JB Jiang</collectorName>
,
<collectorName id="67D2935232BEA55AEDDE0B48EE423D14">J Sun</collectorName>
, Y
<collectorName id="9F9CEE3BDECDF00E20E99063EB17A27D">Dong</collectorName>
and Y
<collectorName id="6648C3F8F21949FC2C6BF856DA0D8AF0">Zheng</collectorName>
</materialsCitation>
<materialsCitation id="F902BA64DAA1A6024776025AEA13E58B" collectingDate="2016-05-02" collectorName="JB Jiang &amp; J Sun &amp; Dong &amp; Zheng" country="China" elevation="69" latitude="28.65267" location="Changde City" longLatPrecision="1" longitude="111.34672" specimenCount="1" stateProvince="Hunan" typeStatus="paratype">
1 clitellates (HU 201602-04),
<collectingCountry id="39FC72060796F84FCF245C77B62ECE63" name="China">China</collectingCountry>
,
<collectingRegion id="A001C9F792310825D87DA912D6E7FD5E" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="5FD6A6EFF80918B95DE0744C27DA3152" LSID="urn:lsid:plazi:treatment:21FE19A90A4251EF9FC664467170CB31:5FD6A6EFF80918B95DE0744C27DA3152" country="China" latitude="28.65267" longLatPrecision="1" longitude="111.34672" name="Changde City" stateProvince="Hunan">Changde City</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.346720,28.652670]}">
<geoCoordinate id="D783B7090BE92ABCA73E82262C4E2375" degrees="28.65267" direction="north" orientation="latitude" precision="1" value="28.65267">28.65267 ° N</geoCoordinate>
,
<geoCoordinate id="D59F9999C1457BA3FB38536761FC0F13" degrees="111.34672" direction="east" orientation="longitude" precision="1" value="111.34672">111.34672 ° E</geoCoordinate>
</named-content>
),
<quantity id="FFBDA6E18918A13B528EDF80C7F1D1AC" metricMagnitude="1" metricUnit="m" metricValue="6.9" unit="m" value="69.0">
<elevation id="2FFA42455F843E9B71F9D302B9B90881" metricMagnitude="1" metricUnit="m" metricValue="6.9" unit="m" value="69.0">69 m</elevation>
</quantity>
elevation, reddish brown soil under moss and fern in farmland,
<date id="5B7D4B52AA92598C2818801096A98194" value="2016-05-02">
<collectingDate id="843593998AC68EED44C7399E41C3ABA0" value="2016-05-02">2 May 2016</collectingDate>
</date>
,
<collectorName id="659D6EF248DA37D63AC8968D7EF54C10">JB Jiang</collectorName>
,
<collectorName id="ED6E666B5E872D22E50F5EF528C98542">J Sun</collectorName>
, Y
<collectorName id="A1D42A72340B764C52F2D15162F98DAC">Dong</collectorName>
and Y
<collectorName id="C9949CFC9C5DB6C88BA7FB750FB9A0FD">Zheng</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4ABG" type="diagnosis">
<paragraph id="805C9FF0EDD9008CB0FC41093B4F550E">
<heading id="0D9924071E3E3D891544E759EC86B5EE" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="09A50DD092729F14FE11278ABBEE88E6">
Size small to medium. Spermathecal pores in 5 / 68 / 9, separated by 1 / 3 of body circumference. Male pores in
<collectionCode id="56233A03AB1D78F89A08E214C2926A2C">XVIII</collectionCode>
, separated by 1 / 3 of body circumference, each on the top of a slightly raised, circular porophore. Spermathecae four pairs in
<collectionCode id="F2B9C21178A8820C7F8336164E1F0FE0" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
<collectionCode id="46F556F4994C8F74CFA34F00250B7391">IX</collectionCode>
, ampulla heart- or rod-shaped, duct is thick and ~ 1 / 3 of ampulla. Diverticulum is ~ 3 / 4 of main pouch (duct and ampulla together), terminal 3 / 4 dilated into bag-shaped chamber intestinal caeca are simple. Prostate glands are degenerated.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECBBG" type="description">
<paragraph id="CF34C21E24006FEFA94FF9F1D2590616">
<heading id="B88F915F0D706211FEB0F57E659CF9BA" reason="title">External characters.</heading>
</paragraph>
<paragraph id="342043FD4F2388A2A502A6F239749BF7">
No pigmentation on dorsal and ventral. Dimensions
<quantity id="FF1208AC64371E3780DCE92B2A9A1F54" metricMagnitude="-2" metricUnit="m" metricValue="8.05" metricValueMax="12.0" metricValueMin="4.1" unit="mm" value="80.5" valueMax="120.0" valueMin="41.0">41120 mm</quantity>
by
<quantity id="1D828F1E5EB9A25673703B82695AF07A" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.54.5 mm</quantity>
at clitellum, segments 49112. Annulus present on X XIII. The dorsal midline is clearly visible and green-brown. First dorsal pore of all examined individuals in 10 / 11. Prostomium 1 / 2 epilobous. Clitellum annular, grey, in XIV 2 / 3
<collectionCode id="F48F99D58612EF3E4FFF53364B9CB640">XVI</collectionCode>
, smooth, setae numbering 8 at XIV (P 1 CJHUSH 190517069 N 8-01 and HU 201602-04), 11 at XV (HU 201602-04), 13 at
<collectionCode id="41F7F63DB9858A4B6EC3710F1A5D8D8C">XVI</collectionCode>
(P 1 CJHUSH 190517069 N 8-01) or 14 at
<collectionCode id="1DEAFB47FA8B192E6E1E8E98EDD315DD">XVI</collectionCode>
(P 1 CJHUSH 190526781
<collectionCode id="FD24C86DB0F7F68557B49D8503F43C92" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
8-04), dorsal pores visible on clitellum. Setae numbering 3242 at III, 4048 at
<collectionCode id="5BEA490F854A2A175BBE49431244B94C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
, 4450 at
<collectionCode id="727CDAA3B8F04C3AD04AC39489E46CCA">VIII</collectionCode>
, 5058 at
<collectionCode id="3883FF05E734D5CF8F7B92FBE7C2777D">XX</collectionCode>
, 5662 at XXV; 1314 between male pores; 1819 (
<collectionCode id="1047FFB11E080E6F1BB2809973E08B0A" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
), 1821 (
<collectionCode id="70F72621431A4A418F0F7EC28FA7E395" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
), 1921 (
<collectionCode id="22CABA0923EA8DA59B556A81DF7A0B51">VII</collectionCode>
) and 1922 (
<collectionCode id="02B8430C04772EF9A420D32C8EFDFE3F">VIII</collectionCode>
) between spermathecal pores; setal formula, aa = 1.01.4 ab, zz = 1.01.4 zy. Male pores one pair in
<collectionCode id="D149768B58B75E594486B312D1982CA6">XVIII</collectionCode>
, separated by 1 / 3 body circumference, each on the top of a slightly raised, circular porophore. Two pairs of circular medium-sized flat-topped papillae present after the setae ring in
<collectionCode id="E1054ED9F7719FF3F005E5FCF339A18E">XVI</collectionCode>
and
<collectionCode id="E5DC980345D76C480A183E3EC039EA4F">XVII</collectionCode>
, the interval is ~ 1 / 3 of body circumference. Two pairs of circular medium-sized flat-topped papillae present before the setae ring on
<collectionCode id="58F3AFAC059EC27F54ADF431CDCC12AF">XVII</collectionCode>
and
<collectionCode id="552352F7AE6E081808DAAE9D3874826E">XVIII</collectionCode>
, the interval is ~ 1 / 9 of body circumference (Fig.
<figureCitation id="8054B3623213F4CEAF5BA2E527595D9F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">3 A</figureCitation>
). Sometimes a similar papilla presents in the center of
<collectionCode id="7F0781E68E97C8509286E5D614C21EC0">XVII</collectionCode>
(specimen HU 201601-03) (Fig.
<figureCitation id="C76B68BACA39DB2729E16D752D155465" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">3 D</figureCitation>
). Sometimes another similar papilla presents in the center of
<collectionCode id="8D40AD0F861FDEA46DE680284864233F">XIX</collectionCode>
(specimen HU 201602-04) (Fig.
<figureCitation id="19158FCB5042CD25BF65294A8DDF1851" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">3 E</figureCitation>
). Female pore single in XIV, oval, milky white. Spermathecal pores four pairs in 5 / 68 / 9, ventral, small eye-like, milky white porophore in center, separated by 1 / 3 body circumference. A pair of oval medium-sized flat-topped papillae after the setae ring between the spermathecal pores in
<collectionCode id="FAA0229EED630D7B626DE5605A26EB71">VII</collectionCode>
, the interval is ~ 1 / 4 of body circumference. Sometimes another pair of similar papillae present in the same position of
<collectionCode id="E86C061905179F157DA3434EA0DF8C03" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
(specimen HU 201601-03 and HU 201602-04). Sometimes another pair of similar papillae present in the same position of
<collectionCode id="6398FB99334A841922F54DE0774B5C7B">VIII</collectionCode>
(specimen P 1 CJHUSH 190526781
<collectionCode id="7252232359D14DD63ADC55CBB8009E06" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
8-04).
</paragraph>
<caption id="EAE9E413736D9471F66FA1EA2000FF77" ID-DOI="10.3897/zookeys.1210.125963.figure3" ID-arpha="1AA4B78A-79FD-522B-B014-42511A9399FB" httpUri="https://binary.pensoft.net/fig/1121080" startId="F3">
<paragraph id="68237ABB972AFE00EA81FD40C1E73DF6">
<label id="1D21A51912FABE50EF60432FAEDE12D5">Figure 3.</label>
</paragraph>
<paragraph id="7D056BCB046779A1614870F3F1F8F41A">
<taxonomicName id="5BC603442DEA98194550B422F85EB4D5" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="2C945E2898542219A76AED00EC8911AB" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CFDFD5CFA67661020F2C72A96626FC65" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE88ED2C71EB936883282A2BC31524CE" bold="true">A</emphasis>
ventral view showing spermathecal pores, and male pores
<emphasis id="498D792145C3D7F999B342C689013167" bold="true">B</emphasis>
intestinal caeca
<emphasis id="EA9004686AAC2A5F6200D515ED612648" bold="true">C</emphasis>
spermathecae
<emphasis id="2F477FFBBE403307C3F494BAB8045A25" bold="true">D</emphasis>
male pores of paratypes (HU 201601-03)
<emphasis id="29BB6E33A83D44089B00CB51E81FB711" bold="true">E</emphasis>
male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWCBG" type="description">
<paragraph id="60C3DF465F4CEE60AC95086E079136D6">
<heading id="2C68FAEEF7CA2B850C7B47272F7E1633" reason="title">Internal characters.</heading>
</paragraph>
<paragraph id="26BF30E2F015B86E946A4DEA0A8AF3DE">
Septa 5 / 67 / 8 thick and muscular, 10 / 1112 / 13 slightly thickened, 8 / 9 and 9 / 10 absent. Gizzard spherical, in
<collectionCode id="91DD9B0563749173AC1FFC18B41618F8">IX</collectionCode>
X. Intestine enlarged distinctly from XV. Intestinal caeca paired in XXVII, extending anteriorly to XXIV, simple, smooth on both sides (Fig.
<figureCitation id="08C034C1DCA318A0DCAD929AA54AE0EC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">3 B</figureCitation>
). Four esophageal hearts in XI XIII, well developed. Male sexual system holandric, testis sacs two pairs, in X and XI, well developed, left and right lobes separated on the ventral side. Seminal vesicles two pairs, extending in XI and XII, well developed, left and right lobes separated on the ventral side. Prostate glands degenerated, prostatic duct U-shaped inserting in
<collectionCode id="11B52202AF4692E16073CCB769DA75C2">XVIII</collectionCode>
, several small lumpy accessory glands in center of ventral
<collectionCode id="E27FBFC520446CC65250DFE1835E3FC2">XVI</collectionCode>
,
<collectionCode id="810C2FA3FDBA931FB246D6C5534AA54A">XVII</collectionCode>
, and
<collectionCode id="5353D32120691F304DE8C95B9B4FFC57">XVIII</collectionCode>
; the position is consistent with the position of the ventral mastoid on the body surface. Spermathecae four pairs in
<collectionCode id="4E739763B248768785B388C92F11C6D9" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
<collectionCode id="C39E0D631E127C623F05AE6C009C2DE1">IX</collectionCode>
, ampulla oval-shaped, ~
<quantity id="BA8BA6778618C07D5282B808CEA859E8" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" unit="mm" value="1.2" valueMax="1.8" valueMin="0.6">0.61.8 mm</quantity>
long in
<typeStatus id="A0C005CB9AB0674093FBDEF90C1A4251" type="holotype">holotype</typeStatus>
; ampulla duct is thick and ~ 1 / 2 of ampulla. Diverticulum is ~ 2 / 3 of main pouch (duct and ampulla together), terminal 3 / 4 dilated into bag-shaped chamber. Two pairs of large lumpy accessory glands present on the ventrum of
<collectionCode id="CB7FD72726148379A0A94F6C29683756">VII</collectionCode>
and
<collectionCode id="5BD7B839AED4658105E440EA5A9DB4F8">VIII</collectionCode>
(Fig.
<figureCitation id="0621D70FF5AECF8C4A662EFB268281AA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. taoyuanensis sp. nov. A ventral view showing spermathecal pores, and male pores B intestinal caeca C spermathecae D male pores of paratypes (HU 201601-03) E male pores of paratypes (HU 201602-04). Abbreviations: sp, spermathecal pores; mp, male pores." figureDoi="10.3897/zookeys.1210.125963.figure3" httpUri="https://binary.pensoft.net/fig/1121080">3 C</figureCitation>
). Sometimes pair of large lumpy accessory glands present on the ventrum of
<collectionCode id="9393CFE2040B00C8DC4EC948BC62FD27">VII</collectionCode>
(specimen HU 201601-03 and HU 201602-04).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEDBG" type="etymology">
<paragraph id="16A5A916BC9F050FA31ABBC18B31E6C1">
<heading id="301558501383B5321066B1E622CD7268" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C67C4789D155D8BDA2E790EE148B1B7E">
The species is named after its
<typeStatus id="72F20FD675DC41383BE87A540B8BEFBA">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJDBG" type="remarks">
<paragraph id="1AC01B6E38A357539081DCE6878867F3">
<heading id="D6A22CC1BDA24319F3252C577B236771" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="71FAA033D281CEDEBA2F742474FCDF1B">
<taxonomicName id="B463C3548724345C1133D7350EEFB76D" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="B437A7E6E86602B9E4F54EC5FE1D1B00" italics="true">Amynthas taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C677FD94897CCBC6A4A73D2092CFE512" rank="species">sp. nov.</taxonomicNameLabel>
, with four pairs of spermathecal pores in 5 / 68 / 9, also belongs to the
<taxonomicName id="232A77EAC6F4BBE3839E83E64B589A05" baseAuthorityName="Kingberg" baseAuthorityYear="1867" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="corticis">
<emphasis id="58309D64C27BE20E565F14097BEAA99B" italics="true">Amynthas corticis</emphasis>
</taxonomicName>
group.
<taxonomicName id="0843BBDF037A556424F86C41EAE10A1E" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="B9473747515917525EB2D0E9320A71DE" italics="true">Amynthas taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F34744FEC6A0739CD8FBFA4CD3474156" rank="species">sp. nov.</taxonomicNameLabel>
is close to
<taxonomicName id="A0DF5583052611DA04B4A6C2B948E175" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="xiangtanensis" status="sp. nov.">
<emphasis id="E1FFBC8EE307DF2F42F1093037B3614D" italics="true">Amynthas xiangtanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EA03A8A5B5603FD37F651EAFE0FC5C3" rank="species">sp. nov.</taxonomicNameLabel>
in setae number, the position and characteristics of spermathecal pores and male pores, and the simple intestinal caeca. However, the differences between the two new species are body size, pigmentation, the position of the first dorsal pore, the position of clitellum, the position of papillae within spermathecal pores and male pore region, the existence or nonexistence of prostate glands, and the characteristics of the spermathecae and diverticulum (Table
<tableCitation id="74488F6AEABD72280F321F0B1637B7BB" captionStart="Table 2" captionStartId="T2" captionText="Table 2. A comparison of characters of A. xiangtanensis sp. nov., A. taoyuanensis sp. nov., and similar species of the Amynthas corticis group. Abbreviations: sp, spermathecal pores, mp, male pores.">2</tableCitation>
).
</paragraph>
<paragraph id="0BD94E23C61762BD640B61A5DB5D6ACB">
The new species is similar to
<taxonomicName id="A7FAC7DA6382D2306734278CDC4336E1" authority="(Chen, 1938)" baseAuthorityName="Chen" baseAuthorityYear="1938" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="homosetus">
<emphasis id="C866757FED79BE558B63F2A29BFEDA3A" italics="true">Amynthas homosetus</emphasis>
(
<bibRefCitation id="977B8A81C22C8A39E51B94A345955892" author="Chen Y" issue="10" journalOrPublisher="Contributions from the Biological Laboratory of the Science Society of China, Zoological Series" pagination="375-427" refId="B2" refString="Chen Y (1938) Oligochaeta from Hainan, Kwangtung. Contributions from the Biological Laboratory of the Science Society of China, Zoological Series 12 (10): 375 - 427." title="Oligochaeta from Hainan, Kwangtung." volume="12" year="1938">Chen, 1938</bibRefCitation>
)
</taxonomicName>
in terms of body size, position and characteristics of spermathecal pores and male pores, and simple intestinal caeca. Whereas, the new species differs from
<taxonomicName id="2DE67CD08810D74617B2C08C151C937F" baseAuthorityName="Chen" baseAuthorityYear="1938" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="homosetus">
<emphasis id="73402B71CFCA665725B2BE0639BF3F93" italics="true">A. homosetus</emphasis>
</taxonomicName>
in being unpigmented, the first dorsal pore in 10 / 11, clitellum in XIV 2 / 3
<collectionCode id="CBC9AF7134790BA4C8C23F8C75C28676">XVI</collectionCode>
, 13 or 14 setae between male pores, the ventral distance of spermathecal pores is 1 / 3 C, papillae paired after setae in
<collectionCode id="BDF1607088ABBAE72321EC0FE04D1E8E" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
,
<collectionCode id="A7A53716D820722486890B448F4F448B">VII</collectionCode>
, or
<collectionCode id="ACC5CD546FEB7CE22681C3CC8334EE71">VIII</collectionCode>
, two paired or more papillae in
<collectionCode id="AF03DCD27927BE47498570B31D42E56B">XVI</collectionCode>
,
<collectionCode id="E54214ACFDA80E64DC492E2E5FE5E3AC">XVII</collectionCode>
,
<collectionCode id="D871B41CA3AB03D14223B6B5B7F85038">XVIII</collectionCode>
, or
<collectionCode id="DC1C2BEEC655D3530293D1507E430C5E">XIX</collectionCode>
, prostate glands degenerated, spermathecae ~
<quantity id="72AE775007EB7335C18F9D099C7AA958" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" unit="mm" value="1.2" valueMax="1.8" valueMin="0.6">0.61.8 mm</quantity>
long, ampulla oval-shaped, duct is thick and ~ 1 / 2 of ampulla, diverticulum terminal 3 / 4 dilated into bag-shaped seminal chamber, two paired large lumpy accessory glands on
<collectionCode id="EEB225BFF796AE6AB63834324E2DEB6F">VII</collectionCode>
or
<collectionCode id="AA0A1D9D011F56D1E21FFF98FDAABAA8">VIII</collectionCode>
.
<taxonomicName id="05769CAFDBD8667941163E9D7D2E077D" baseAuthorityName="Chen" baseAuthorityYear="1938" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="homosetus">
<emphasis id="B20137807136C8BDAA335C408A41CC6E" italics="true">Amynthas homosetus</emphasis>
</taxonomicName>
is dark chocolate-colored anteriorly and grey on other parts of dorsum, grey on the ventrum, the first dorsal pore in 12 / 13, clitellum without setae in XIV
<collectionCode id="CC9F9141751CBC8BECBA3BE0F2D08E40">XVI</collectionCode>
, nine setae between male pores, the ventral distance of spermathecal pores is 1 / 4 C, papillae invisible within spermathecal pores and male pore regions, prostate glands developed in
<collectionCode id="5DCB8673486B740E829B86DDA1D0F07B">XVI</collectionCode>
XXI, spermathecae heart-shaped, diverticulum seminal chamber ovoid (Table
<tableCitation id="6C6223AC357B2B12B8AF611F7A1AA3B6" captionStart="Table 2" captionStartId="T2" captionText="Table 2. A comparison of characters of A. xiangtanensis sp. nov., A. taoyuanensis sp. nov., and similar species of the Amynthas corticis group. Abbreviations: sp, spermathecal pores, mp, male pores.">2</tableCitation>
).
</paragraph>
<paragraph id="C8A701C713CEDC4125B4EB864E4730D8">
<taxonomicName id="8567BB4E04078A850E4D73FF45E63C93" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="D1BF2BD97F9D3048A7EED162BBF9C7D9" italics="true">Amynthas taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB0B1F042297A6A9819BE3DCC6611E23" rank="species">sp. nov.</taxonomicNameLabel>
is also close to
<taxonomicName id="94D75185933B98ADF86A078134FD02CA" authority="Qiu &amp; Sun, 2012" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="2D25AF58C9C18895F937D11DD6406A86" italics="true">Amynthas stricosus</emphasis>
Qiu &amp;
<bibRefCitation id="F03A2240FBD41504CBF6C756240BC706" DOI="10.11646/zootaxa.3458.1.8" author="Sun J &amp; Jiang JB &amp; Qiu JP" issue="1" journalOrPublisher="Zootaxa" pagination="149-158" refId="B32" refString="Sun J, Jiang JB, Qiu JP (2012) Four new species of the Amynthas corticis group (Oligochaeta: Megascolecidae) from Hainan Island, China. Zootaxa 3458 (1): 149 - 158. https: // doi. org / 10.11646 / zootaxa. 3458.1. 8" title="Four new species of the Amynthas corticis group (Oligochaeta: Megascolecidae) from Hainan Island, China." volume="3458" year="2012">Sun, 2012</bibRefCitation>
</taxonomicName>
by body size, pigmentation, setae number, the characteristics of spermathecal pores and male pores, the ventral distance of male pores, simple intestinal caeca. However, the new species is distinguished from
<taxonomicName id="C617B43A0F9BDB47987FE552D19C719A" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="7AD4A15B956349A8E6D165188A7D2E62" italics="true">A. stricosus</emphasis>
</taxonomicName>
by the first dorsal pore, the position and existence or nonexistence setae of clitellum, and the ventral distance of spermathecal pores;
<taxonomicName id="F8F795E9EAF06E1034C937C61DFEA0E4" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="A1D27310117B071D98969780E121DB33" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FD857F5233936E49973F3D54150BEE5" rank="species">sp. nov.</taxonomicNameLabel>
has paired papillae after setae in
<collectionCode id="F4A7F1EFC32A037CEE2FDEF0185024CE" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
,
<collectionCode id="F295875CF6BAF68AB40C4327E4C255A3">VII</collectionCode>
, or
<collectionCode id="6B864F348CD8A1ED19150D95E6592CB9">VIII</collectionCode>
, but
<taxonomicName id="9D7DF983948D73574D27D3B85EFE672E" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="6C7F8B58340AF2356164DE945539D60E" italics="true">A. stricosus</emphasis>
</taxonomicName>
has no papillae; in addition,
<taxonomicName id="A9B46743C7D5CD2F71C1C9C6C3A5D5B4" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="6393451B739004A9172F3BD5551738CC" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="156EE83CED5165EDA7B831652BEACEA7" rank="species">sp. nov.</taxonomicNameLabel>
has two paired papillae or more in
<collectionCode id="66CDFFAA9CAE15A35555164F6A3996F9">XVI</collectionCode>
,
<collectionCode id="EA7BD84D9786098AAB42A23FB6362F88">XVII</collectionCode>
,
<collectionCode id="5A6178B20CF8755221FD5207589D262F">XVIII</collectionCode>
, or
<collectionCode id="AF039169A4034E602B45B6A7DC10A85D">XIX</collectionCode>
, but
<taxonomicName id="5FBB55AAEDD0A20272F15137C671805A" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="1F0754AE3DE732AC5AF4EED1E565175A" italics="true">A. stricosus</emphasis>
</taxonomicName>
has a single or paired after setae in
<collectionCode id="4C4162F1008A856755B126F410D6327E">XVII</collectionCode>
,
<collectionCode id="636CB46DECDD74FDF0CA99B8B06FAB9A">XIX</collectionCode>
, and
<collectionCode id="82150DD65E0657F53D1AC559F2A5992E">XX</collectionCode>
; prostate glands of
<taxonomicName id="47B5633B3F36962C1E4754958F616846" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="73955DA98249A93BF921B2C52BB2ACCC" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1A1BC669791929C3FCC05EB7E4D6803" rank="species">sp. nov.</taxonomicNameLabel>
are degenerated with several lumpy accessory glands in
<collectionCode id="1DB0D4023A6EB3CCCA8C47B232F8DFEA">XVI</collectionCode>
,
<collectionCode id="BCD00C57E97877B982C5E9F337E55992">XVII</collectionCode>
, and
<collectionCode id="48CA22E4C7BB887BF3599AF55EB46A96">XVIII</collectionCode>
, while developed in
<collectionCode id="FED37202769162F2D42A031B8E983016">XVI</collectionCode>
<collectionCode id="119FA29DF4A78257E38D7F0D65C0C74A">XX</collectionCode>
with accessory glands invisible in
<taxonomicName id="AD3EF6F39188B95FCE921159F4893FA8" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="F08251F27EBEE3BDF4FFEE3D0052249E" italics="true">A. stricosus</emphasis>
</taxonomicName>
; furthermore, spermathecae of
<taxonomicName id="0E91EDAC893AD4AB9C58513D9EE723C5" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="41745F143C3C4A47965A2C3BF61791E6" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CE48C2C8A99F06C301DDAC2F4FFFFFD" rank="species">sp. nov.</taxonomicNameLabel>
are ~
<quantity id="06FDE4ED11F814E63084442534F388A9" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" unit="mm" value="1.2" valueMax="1.8" valueMin="0.6">0.61.8 mm</quantity>
long, ampulla oval-shaped, duct is thick and ~ 1 / 2 of ampulla, whereas ~
<quantity id="BC16231AD7F9C34CDA85790CBFE95CBB" metricMagnitude="-3" metricUnit="m" metricValue="1.6" unit="mm" value="1.6">1.6 mm</quantity>
long, ampulla heart-shaped, gradually slender duct as long as ampulla in
<taxonomicName id="C0A870E377C8BFB14423A2C180863680" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="286D4235F5B5337910793C54458B56FF" italics="true">A. stricosus</emphasis>
</taxonomicName>
; diverticulum of
<taxonomicName id="D191C39863121F51C349B75572CDD197" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="05AFF1D8637D507D3AF6A60F53623B95" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4F3A770601EB58F924B41A4086E5DE42" rank="species">sp. nov.</taxonomicNameLabel>
is ~ 3 / 4 of main pouch, terminal 3 / 4 dilated into bag-shaped seminal chamber, but as long as main pouch, slender, terminal 2 / 5 dilated into band-shaped seminal chamber in
<taxonomicName id="DC700F6C42274D05A0341A9DCFE7248B" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="89D5BFCD2F1C3CCBF3FC3CD310625E2B" italics="true">A. stricosus</emphasis>
</taxonomicName>
; two paired large lumpy accessory glands on
<collectionCode id="C6C45D253DCA6D13487BE4BDFEA656D7">VII</collectionCode>
or
<collectionCode id="37D8BDCD6555BC5D2C252D7E39F28D63">VIII</collectionCode>
in
<taxonomicName id="C7ADA50391579DF1BC81E9E92C11DF65" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="E719AEC0A149979A8D847198CA992BBE" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2811C517558A4F60C555ACC0BD7CBEE0" rank="species">sp. nov.</taxonomicNameLabel>
, but no accessory glands in
<taxonomicName id="D9A70901DC0304B803BD32CA343B731C" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="stricosus">
<emphasis id="EC4B630AC40B900CBC3DBF49BAB526D8" italics="true">A. stricosus</emphasis>
</taxonomicName>
(Table
<tableCitation id="821F5B6E2E50D9E23D26065455BE12CA" captionStart="Table 2" captionStartId="T2" captionText="Table 2. A comparison of characters of A. xiangtanensis sp. nov., A. taoyuanensis sp. nov., and similar species of the Amynthas corticis group. Abbreviations: sp, spermathecal pores, mp, male pores.">2</tableCitation>
).
</paragraph>
<paragraph id="61E51993A8830A1FF10A04563B6EC2DF">
According to body size, pigmentation, setae ventrally in clitellum, the position and characteristics of spermathecal pores and male pores, simple intestinal caeca, and the characteristics of spermathecae,
<taxonomicName id="82C7045B8B4A686E4DC9CDF3047F7230" authorityName="Jin &amp; Li &amp; Jiang &amp; Qiu" authorityYear="2024" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="taoyuanensis" status="sp. nov.">
<emphasis id="495EF2B30D681AEA00BF5FBDAF3DC47D" italics="true">A. taoyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A31D0FCC9C04B9256EAC0FD97724B0E6" rank="species">sp. nov.</taxonomicNameLabel>
is somewhat similar to
<taxonomicName id="A1509F1585D1D438539A10770523AA9B" authority="Qiu &amp; Sun, 2012" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="genitalis">
<emphasis id="C99D339329E3DF193E4511C0625A5F50" italics="true">Amynthas genitalis</emphasis>
Qiu &amp;
<bibRefCitation id="DCA41F794B48C88D5D7BFBC4CFA71F37" DOI="10.11646/zootaxa.3458.1.8" author="Sun J &amp; Jiang JB &amp; Qiu JP" issue="1" journalOrPublisher="Zootaxa" pagination="149-158" refId="B32" refString="Sun J, Jiang JB, Qiu JP (2012) Four new species of the Amynthas corticis group (Oligochaeta: Megascolecidae) from Hainan Island, China. Zootaxa 3458 (1): 149 - 158. https: // doi. org / 10.11646 / zootaxa. 3458.1. 8" title="Four new species of the Amynthas corticis group (Oligochaeta: Megascolecidae) from Hainan Island, China." volume="3458" year="2012">Sun, 2012</bibRefCitation>
</taxonomicName>
. However, the new species is characterized by the first dorsal in 10 / 11, clitellum in XIV 2 / 3
<collectionCode id="8D15498D5C17B6264DD3C95823762E65">XVI</collectionCode>
, 1922 setae between spermathecal pores (
<collectionCode id="35A1B7C5F4758758A90F0CFE36888D5D">VIII</collectionCode>
), papillae paired within spermathecal pore region, two paired or more papillae in
<collectionCode id="C8AC2BFCD84A3CED5FCA03B4FAD825C7">XVI</collectionCode>
,
<collectionCode id="A1ABC26269B8E53876AD92F387748A0D">XVII</collectionCode>
,
<collectionCode id="3677C26D5BE52509528300784903319C">XVIII</collectionCode>
, or
<collectionCode id="7D5C7C0BD026A139DFA4418CC86AC6AE">XIX</collectionCode>
, prostate glands degenerated with several lumpy accessory glands in
<collectionCode id="7FA83B912479801B58313CA02188A3D5">XVI</collectionCode>
,
<collectionCode id="8C982E31E02944D4D019C3E3A98FD090">XVII</collectionCode>
, and
<collectionCode id="F34F8722F38FAE61275782100345B90F">XVIII</collectionCode>
, diverticulum ~ 3 / 4 of main pouch, terminal 3 / 4 dilated into bag-shaped seminal chamber, two paired large lumpy accessory glands on
<collectionCode id="71DFC2004980851CEA7F11BEC0964A6E">VII</collectionCode>
or
<collectionCode id="B788E52C99B4EFCC0A822DAAA694523A">VIII</collectionCode>
; whereas
<taxonomicName id="D2B0F79B160461572D0BA1ADAF7290DE" authorityName="Qiu &amp; Sun" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="genitalis">
<emphasis id="8CA922B4EE7E904A4E0A6FB03A121579" italics="true">A. genitalis</emphasis>
</taxonomicName>
has 811 setae between spermathecal pores (
<collectionCode id="06A7334C48648D8034C4F52000BAE683">VIII</collectionCode>
), no papillae within spermathecal pore region, paired papillae before setae annulet in
<collectionCode id="4B3EA3C33B0A4996FCE1CEA15AF4E8CC">XVIII</collectionCode>
and
<collectionCode id="9FA98E27D5646146D788C7B89F096C0D">XIX</collectionCode>
, after setae annulet in
<collectionCode id="EFAE03CED29783618333438CBAA30327">XVII</collectionCode>
and
<collectionCode id="C31CD8CF5A94F00BF56F045A6E7690F6">XVII</collectionCode>
, prostate glands developed with accessory glands invisible in
<collectionCode id="2A21BB03289F546F689316F2E80909D0">XVII</collectionCode>
<collectionCode id="90A2E491B4D11EA3B27F4709BEAF3A8B">XX</collectionCode>
, diverticulum longer than main pouch, slender, terminal 0.29 dilated into rod-shaped seminal chamber, no accessory glands (Table
<tableCitation id="BC36506D58211081B5432DFC8947B502" captionStart="Table 2" captionStartId="T2" captionText="Table 2. A comparison of characters of A. xiangtanensis sp. nov., A. taoyuanensis sp. nov., and similar species of the Amynthas corticis group. Abbreviations: sp, spermathecal pores, mp, male pores.">2</tableCitation>
).
</paragraph>
<paragraph id="AB735BC8EC0E2167B64E04E2E97F7124">
Another similar species with four pairs of spermathecal pores in 5 / 68 / 9 is
<taxonomicName id="811C20CE5B4C7D16DB2183C62EE68250" authority="Yuan &amp; Jiang, 2019" authorityName="Yuan &amp; Jiang" authorityYear="2019" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" phylum="Annelida" rank="species" species="recavus">
<emphasis id="E1E49CBB2E9830A85E6721D2B0EA8D3C" italics="true">Amynthas recavus</emphasis>
<bibRefCitation id="E10C7AB0B9719997CC57900637A05D40" DOI="10.1080/00222933.2019.1680760" author="Yuan Z &amp; Dong Y &amp; Jiang JB &amp; Qiu JP" refId="B39" refString="Yuan Z, Dong Y, Jiang JB, Qiu JP (2019) Three new species of earthworms belonging to the genera Amynthas and Metaphire (Oligochaeta: Megascolecidae) from Yunnan, China. Journal of Natural History 53: 31 - 32, 1961 - 1974. https: // doi. org / 10.1080 / 00222933.2019. 1680760" year="2019">Yuan &amp; Jiang, 2019</bibRefCitation>
</taxonomicName>
. The two species share some similarities, such as the position of male pores and simple intestinal caeca. However, the two species are distinguished by body size, pigmentation, the first dorsal pore position, setae number, the ventral distance of spermathecal pores, existence or nonexistence of papillae within spermathecal pore region, the characteristics of male pores and papillae within male pore region, prostate glands, the characteristics of spermathecae and diverticulum, and the existence or nonexistence of accessory glands (Table
<tableCitation id="67C10A3A6437EA51033549D2A8024BA3" captionStart="Table 2" captionStartId="T2" captionText="Table 2. A comparison of characters of A. xiangtanensis sp. nov., A. taoyuanensis sp. nov., and similar species of the Amynthas corticis group. Abbreviations: sp, spermathecal pores, mp, male pores.">2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,562 @@
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<mods:title id="6FAA67AC6409F5F05ABA2D203374D8C6">Two new species and a new record of the ant genus Meranoplus Smith, 1853 (Hymenoptera, Formicidae, Myrmicinae) from Thailand</mods:title>
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<mods:end id="789DAF1AEBCB35FCAA8343D826179591">227</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3E38B0BEE6F83C8598E3999C38C5C9FF">journal article</mods:classification>
<mods:identifier id="1E45CA09EB1FF0AE5F87C3C34FD87574" type="DOI">10.3897/zookeys.1210.125990</mods:identifier>
<mods:identifier id="0F13D2BBC60E965B658C23FC52D64BB9" type="ZooBank">B158276E-7AC6-40AF-8713-0881D42F3953</mods:identifier>
</mods:mods>
<treatment id="5D5203100B0054B796CE127B6CAFD2F2" ID-arpha="5D520310-0B00-54B7-96CE-127B6CAFD2F2" ID-zoobank="https://zoobank.org/752DDF38-72E5-4325-9B3D-4C70BEEFA1DF" LSID="urn:lsid:zoobank.org:act:752DDF38-72E5-4325-9B3D-4C70BEEFA1DF" LSID-ZBK="urn:lsid:zoobank.org:act:752DDF38-72E5-4325-9B3D-4C70BEEFA1DF" httpUri="http://treatment.plazi.org/id/5D5203100B0054B796CE127B6CAFD2F2">
<subSubSection id="A962E35A6C58899BDDB3CE3185D72215" type="nomenclature">
<paragraph id="56EA98E0ADEE1508DAE8D5275D93AC18">
<taxonomicName id="42536B95EA48993CF82E8670BAE908EA" authority="Yodprasit &amp; Jaitrong" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="9D8C27E8F63AE09B7154F317F3F13CFE" italics="true">Meranoplus siamensis</emphasis>
Yodprasit &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="603D899386D672B0A453340945E75DE2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8765F5375476CC1A29539455B3115B2F" type="description">
<paragraph id="E28344CC3A955B44D4EA3BCA87073CEC">
<figureCitation id="DBA65E24152F1BBECC903051B6961140" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Meranoplus siamensis sp. nov. (holotype, THNHM - I- 00027303) A head in full-face view B body in profile view C promesonotal shield in dorsal view D body in dorsal view." figureDoi="10.3897/zookeys.1210.125990.figure2" httpUri="https://binary.pensoft.net/fig/1121261">Figs 2 A D</figureCitation>
,
<figureCitation id="CC2D58E3CE515C4D5C56526A88CEF4BF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 H I</figureCitation>
,
<figureCitation id="9D2263FE46918C26FAFAB8AE256A40B3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Characters used in key A Meranoplus siamensis (non-type worker, THNHM - I 00028942) B, F M. castaneus (non-type, THNHM - I- 00027323) C M. laeviventris (non-type, THNHM - I- 00027329) D, E M. bellii (lectotype, CASENT 0908933) G, H M. dlusskyi (in Zryanin 2015) I M. levis (holotype, CASENT 0902025) A eye B, C petiole and postpetiole in profile D, F, G, I mesosoma in profile E gaster in dorsal view H head in profile." figureDoi="10.3897/zookeys.1210.125990.figure6" httpUri="https://binary.pensoft.net/fig/1121266">6 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBOAE" type="Types">
<paragraph id="7FFEFD87962EDE143C747346A6C50CA7">
<heading id="F21BC303DEABD72EFA3E56482C1F12AA" reason="title">Types.</heading>
</paragraph>
<paragraph id="AA02A8B01B41730838EFE59580FF9896">
<materialsCitation id="7E8EC7D108FA93C893707DF82E3F8EEF" collectingDate="2003-10-18" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" latitude="13.2837" location="Kasetsart Sriracha Campus" longLatPrecision="7" longitude="100.9238" municipality="Sri Racha District" specimenCount="1" stateProvince="Chon Buri" typeStatus="holotype">
<emphasis id="7C330958D6269B8DA33B3AA60937FCED" bold="true" italics="true">
<typeStatus id="C3D70F782569370EB22FDFE688BF2968" type="holotype">Holotype</typeStatus>
</emphasis>
: • worker (
<abbrev content-type="institution" id="ABBRID0EMOAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="92905F894182BD764304B7B86AEFAEA2">THNHM</collectionCode>
</abbrev>
- I- 00027303,
<abbrev content-type="institution" id="ABBRID0EROAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="96E81AD9A3779BC05FBA65239C688EE0">THNHM</collectionCode>
</abbrev>
), eastern
<collectingCountry id="0185137CD9EC9FB140222B34A5F8BF6A" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="E055859A7D30D60675027EA3C0792722" country="Thailand" name="Chon Buri">Chonburi Province</collectingRegion>
,
<collectingMunicipality id="278713ADD59048D57C321FA8E8B99AA7">Sri Racha District</collectingMunicipality>
,
<location id="C1917C33F90F99E8C22AE905F0044876" LSID="urn:lsid:plazi:treatment:5D5203100B0054B796CE127B6CAFD2F2:C1917C33F90F99E8C22AE905F0044876" country="Thailand" latitude="13.2837" longLatPrecision="7" longitude="100.9238" municipality="Sri Racha District" name="Kasetsart Sriracha Campus" stateProvince="Chon Buri">Kasetsart Sriracha Campus</location>
, dry evergreen forest,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.923800,13.283700]}">
<geoCoordinate id="6FE7C99EE5BC25ACA21D70D66DCC3CE7" degrees="13.2837" direction="north" orientation="latitude" precision="5" value="13.2837">13.2837 ° N</geoCoordinate>
,
<geoCoordinate id="073C3C749789EFEFD64FFDFCD9B8C167" degrees="100.9238" direction="east" orientation="longitude" precision="5" value="100.9238">100.9238 ° E</geoCoordinate>
</named-content>
,
<date id="565865280399DE4554A9F7631DC17E86" value="2003-10-18">
<collectingDate id="B92E4865A1724C7386182FFE8FB64C14" value="2003-10-18">18 October 2003</collectingDate>
</date>
,
<collectorName id="F8E11725C17E8DC515A3010B54945878">W. Jaitrong</collectorName>
leg., TH 03 - WJT- 313.
</materialsCitation>
</paragraph>
<paragraph id="48A548B7A2C96B6338E1D06B4FBF1064">
<materialsCitation id="6846AD7B95C37658821D613C2B55E4CC" collectingDate="2003-10-18" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" latitude="13.2837" location="Kasetsart Sriracha Campus" longLatPrecision="7" longitude="100.9238" municipality="Sri Racha District" specimenCount="6" specimenCount-worker="6" stateProvince="Chon Buri" typeStatus="paratype">
<emphasis id="D595E17DFF15DEC7FD904D6E402EB9F0" bold="true" italics="true">
<typeStatus id="3629CAEB5C0EF2BA2F1437387E7CC96E" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="CB555E082B88AB64C56F1FCB3BBDEB37" count="6" type="worker">6 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EFPAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="4F26B52B99851324EECF0561E7C6CE54">THNHM</collectionCode>
</abbrev>
- I- 00027304 to
<abbrev content-type="institution" id="ABBRID0EKPAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="BE2B7E09BD5A9D3CBFB1307B7F456E5A">THNHM</collectionCode>
</abbrev>
- I- 0002730,
<abbrev content-type="institution" id="ABBRID0EPPAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">THNHM</abbrev>
- I- 00027310,
<abbrev content-type="institution" id="ABBRID0EUPAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="0709E88D359EAEF48A3C89CAE3E9CBB7">THNHM</collectionCode>
</abbrev>
- I- 00028942), same data as holotype
</materialsCitation>
<materialsCitation id="C2FA36B53B5B72F63E083E1B130F359E" collectionCode="THNHM" country="Thailand" specimenCount="4" specimenCount-worker="4" stateProvince="Chon Buri" typeStatus="paratype">
<specimenCount id="587144632BED83A54460304AEDF12480" count="4" type="worker">4 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EZPAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="95D1D7B25EBF1DBE776E962649614FB1">THNHM</collectionCode>
</abbrev>
- I- 00027309,
<abbrev content-type="institution" id="ABBRID0E5PAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="D1BF4190A35756C9EDC50A014477DE0B">THNHM</collectionCode>
</abbrev>
- I- 00027311 to
<abbrev content-type="institution" id="ABBRID0EDQAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="FBB19A8FE41B5972E5BEF7026C72FCBE">THNHM</collectionCode>
</abbrev>
- I- 00027313)
</materialsCitation>
<materialsCitation id="4D3B2088798641CC63C45DC96BB3CD4C" collectingDate="2003-04-19" collectionCode="THNHM" collectorName="A. Suwanasri" country="American Samoa" latitude="13.2837" location="Kasetsart Sriracha Campus" longLatPrecision="7" longitude="100.9238" municipality="Sri Racha District" specimenCount="3" specimenCount-worker="3" stateProvince="Chon Buri" typeStatus="paratype">
<specimenCount id="6D02DF4FB6FBB3D2F8946D007F1A9500" count="3" type="worker">3 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EIQAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="9B802BA3D6F80FA1198560FFEB67BD19">THNHM</collectionCode>
</abbrev>
- I- 00027314), same locality as holotype, but
<date id="C56FCF5E5447DB066D79F490BE31756A" value="2003-04-19">
<collectingDate id="25F0FF6BBCF979E5926496543A5F0DD3" value="2003-04-19">19. IV. 2003</collectingDate>
</date>
,
<collectorName id="746F8CDF9BF6E2579498B7C8C7CD72C3">A. Suwanasri</collectorName>
leg.,
<collectingCountry id="07EE45E6910D38CA6BBA9C24D767FF26" name="American Samoa">AS</collectingCountry>
190403-01. The
<typeStatus id="8A147EF7D50B733446AF968822F210EC" type="paratype">paratypes</typeStatus>
are deposited in
<abbrev content-type="institution" id="ABBRID0ENQAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">THNHM</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESQAE" type="type material">
<paragraph id="0B71340DEC70756C967DD82614755BEB">
<heading id="728953DE81E9B24EFDB931D1664C4CEC" reason="title">Non-type material examined.</heading>
</paragraph>
<paragraph id="CD1D1F0D9BD103D5C573484E697BAADF">
<emphasis id="F3511F8B03AF0873518CDC47D17327AD" bold="true">Central</emphasis>
: •
<materialsCitation id="747B5F5022F0E0CDFBFBFE0874B5BC74" collectingDate="2002-06-01" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Ban Rai District" latitude="15.1225" location="Western" longLatPrecision="7" longitude="99.2755" municipality="Kaen Ma Kurd Village" specimenCount="22" specimenCount-worker="22" stateProvince="Uthai Thani">
<specimenCount id="B6F5B89F7B337D36720C12A9E1C67EBA" count="1" type="worker">1 worker</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0E1QAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="CB02CB8D79BBA28FA0A4832E7B14FFD9">THNHM</collectionCode>
</abbrev>
- I- 00028919,
<abbrev content-type="institution" id="ABBRID0E6QAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="8102C2CC4F947B57527B8CFBDAC0E2F3">THNHM</collectionCode>
</abbrev>
), central
<collectingCountry id="7DBF63EEE446D20C4F6B461D08FE1B68" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="224F3DF268EE5E67B1ADC722E756FAB9" country="Thailand" name="Uthai Thani">Uthai Thani Province</collectingRegion>
,
<collectingCounty id="8A10E02C350346237BDAC39E46E7AD04">Ban Rai District</collectingCounty>
,
<collectingMunicipality id="BAE564063BBE23B9617FF1DE1F95E79C">Kaen Ma Kurd Village</collectingMunicipality>
, dry dipterocarp forest,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[99.275500,15.122500]}">
<geoCoordinate id="B88B4BC6EAC50A9AB2BFC393DC9268DD" degrees="15.1225" direction="north" orientation="latitude" precision="5" value="15.1225">15.1225 ° N</geoCoordinate>
,
<geoCoordinate id="84A2583170E393658F4023F5B2FC4867" degrees="99.2755" direction="east" orientation="longitude" precision="5" value="99.2755">99.2755 ° E</geoCoordinate>
</named-content>
,
<date id="12BF953DE873B4DC7A59EC58DE72F60A" value="2002-06-01">
<collectingDate id="581595C78D1C3D6D7C515C6644F82607" value="2002-06-01">1 June 2002</collectingDate>
</date>
,
<collectorName id="B5BDE265AFFFBB37235C5E674090FCFD">W. Jaitrong</collectorName>
leg., TH 02 - WJT- 039;
<specimenCount id="9461949BCE52F22A7DE3090D946B3E9C" count="21" type="worker">21 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EMRAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="E6068805660D89E4FAE83078BA964DB5">THNHM</collectionCode>
</abbrev>
- I- 00028920 to
<abbrev content-type="institution" id="ABBRID0ERRAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="4DE947A8CDF282B53131FACAEF800FF6">THNHM</collectionCode>
</abbrev>
- I- 00028940,
<abbrev content-type="institution" id="ABBRID0EWRAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="86AFCA2F8B705C50D42C87E9CAB8922E">THNHM</collectionCode>
</abbrev>
), same data locality.
<emphasis id="604224165876A23EEDA9792C8F14C1F9" bold="true">
<location id="1CB1DD20942CA46B916C46118C6752E1" LSID="urn:lsid:plazi:treatment:5D5203100B0054B796CE127B6CAFD2F2:1CB1DD20942CA46B916C46118C6752E1" country="Thailand" county="Ban Rai District" latitude="15.1225" longLatPrecision="7" longitude="99.2755" municipality="Kaen Ma Kurd Village" name="Western" stateProvince="Uthai Thani">Western</location>
</emphasis>
</materialsCitation>
: •
<materialsCitation id="907D10140BEE339D4AAC1DA2E6439215" collectingDate="2019-05-25" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Ban Chong Keab" municipality="Sai Yok District" specimenCount="3" specimenCount-worker="3" stateProvince="Kanchanaburi">
<specimenCount id="07FB66115316C834722549F9CA062431" count="3" type="worker">3 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0E4RAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="D27C02242FD1365003C8D2CE6A1266F7">THNHM</collectionCode>
</abbrev>
- I- 00028941,
<abbrev content-type="institution" id="ABBRID0ECSAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="B663781BB1FF0A552FCAE834F1709F7F">THNHM</collectionCode>
</abbrev>
),
<collectingRegion id="CD547B7C08711E3512F8CE95663FA68E" country="Thailand" name="Kanchanaburi">Kanchanaburi Province</collectingRegion>
,
<collectingMunicipality id="1F55CA9FBD969DC28AF7CB8D3FAF8BF9">Sai Yok District</collectingMunicipality>
,
<location id="D1E5FCDA3542DBF9D9B487513A326AF7" LSID="urn:lsid:plazi:treatment:5D5203100B0054B796CE127B6CAFD2F2:D1E5FCDA3542DBF9D9B487513A326AF7" country="Thailand" municipality="Sai Yok District" name="Ban Chong Keab" stateProvince="Kanchanaburi">Ban Chong Keab</location>
, dry dipterocarp forest,
<date id="62347A4F5B7FB8A8ACC0E9164527EA59" value="2019-05-25">
<collectingDate id="F0E625EAC2BDE6E3B5C9BAC5B890F5C9" value="2019-05-25">25 May 2019</collectingDate>
</date>
,
<collectorName id="E17483DCED9831F71176FCF302BF105F">W. Jaitrong</collectorName>
leg.
<emphasis id="CA97F03F6838BDF11D93B0C7D4684655" bold="true">Northeastern</emphasis>
</materialsCitation>
: •
<materialsCitation id="311D914302A336127A8D782014299228" collectingDate="2022-06-05" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Wang Nam Kheao District" latitude="14.5031" location="Eastern" longLatPrecision="7" longitude="101.9368" municipality="Sakaerat Environmental Research Station" specimenCount="7" specimenCount-worker="7" stateProvince="Nakhon Ratchasima">
<specimenCount id="A6043A997BE2C003682E0368AC25E51E" count="7" type="worker">7 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EJSAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="7B8FCCCBCA9911236E936A5A4A80D654">THNHM</collectionCode>
</abbrev>
- I- 00027315,
<abbrev content-type="institution" id="ABBRID0EOSAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="FB740A32054395AAC0E969AE9E88816B">THNHM</collectionCode>
</abbrev>
),
<collectingRegion id="4C4E9081C0EF8463A0318F61A2C4385F" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima Province</collectingRegion>
,
<collectingCounty id="6321F31CE6C1D78DAA6F44BC46D6B5D0">Wang Nam Kheao District</collectingCounty>
,
<collectingMunicipality id="8DECE19CFDA9A3ED24D40716301DBC24">Sakaerat Environmental Research Station</collectingMunicipality>
(ERS), dry dipterocarp forest,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.936800,14.503100]}">
<geoCoordinate id="76905675AECEAA8B17B0970EC6BA7B03" degrees="14.5031" direction="north" orientation="latitude" precision="5" value="14.5031">14.5031 ° N</geoCoordinate>
,
<geoCoordinate id="9BDC52ED7A882DF8863387640C0D8F11" degrees="101.9368" direction="east" orientation="longitude" precision="5" value="101.9368">101.9368 ° E</geoCoordinate>
</named-content>
,
<date id="AF5DBA6014E253A85A8848A6CD27970F" value="2022-06-05">
<collectingDate id="0A81A328BCC97A0A0B1F1E37031285D1" value="2022-06-05">5 June 2022</collectingDate>
</date>
,
<collectorName id="8B5B48F2888D03678D020BEEF1A481F1">W. Jaitrong</collectorName>
leg., TH 22 - WJT- 264 (
<abbrev content-type="institution" id="ABBRID0E2SAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="02847F7B19ABC621AECC4ACFC429E224">THNHM</collectionCode>
</abbrev>
);
<emphasis id="E3DEC30D02BDBF2FE441A495DD0DA15F" bold="true">
<location id="0E8005E6192499CD7B7E6259A40FDC96" LSID="urn:lsid:plazi:treatment:5D5203100B0054B796CE127B6CAFD2F2:0E8005E6192499CD7B7E6259A40FDC96" country="Thailand" county="Wang Nam Kheao District" latitude="14.5031" longLatPrecision="7" longitude="101.9368" municipality="Sakaerat Environmental Research Station" name="Eastern" stateProvince="Nakhon Ratchasima">Eastern</location>
</emphasis>
</materialsCitation>
: •
<materialsCitation id="AC0906B106512D993DDC4D2101C88CE3" collectingDate="2003-04-06" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Tha Takiab District" specimenCount="4" specimenCount-worker="4" stateProvince="Chachoengsao">
<specimenCount id="79784F496D89EC050EAA52DD62A89A40" count="4" type="worker">4 workers</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0ECTAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="4A52C504F7FE73727EB5672CA9243444">THNHM</collectionCode>
</abbrev>
- I- 00027316,
<abbrev content-type="institution" id="ABBRID0EHTAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="DFDC26F7D58A82538EB6772A79CFD6AF">THNHM</collectionCode>
</abbrev>
),
<collectingRegion id="6EC37D806723B524BC2404594136B4BD" country="Thailand" name="Chachoengsao">Chachoengsao Province</collectingRegion>
,
<location id="C475022FD7800CCCCD481F83A28C819F" LSID="urn:lsid:plazi:treatment:5D5203100B0054B796CE127B6CAFD2F2:C475022FD7800CCCCD481F83A28C819F" country="Thailand" name="Tha Takiab District" stateProvince="Chachoengsao">Tha Takiab District</location>
,
<date id="9CCAB1168F203F312FCB7BCFA310E459" value="2003-04-06">
<collectingDate id="3BF4BAE5E80B2908E28868B49AD5B6D4" value="2003-04-06">6 April 2003</collectingDate>
</date>
,
<collectorName id="8E68F9BB39A3F2B412C9E1CD696D3A27">W. Jaitrong</collectorName>
leg., WJT 260403-01
</materialsCitation>
.
</paragraph>
<caption id="A4C8D2A4276A4E6958402EE791F9F667" ID-DOI="10.3897/zookeys.1210.125990.figure2" ID-arpha="633C77BD-38D3-5061-991A-B86AEEC63BF5" httpUri="https://binary.pensoft.net/fig/1121261" startId="F2">
<paragraph id="5CDCAA9A9EC6A34774A9B4579C17DD4A">
<label id="8E03FDF823AC7746DCEA2CB55BD5F516">Figure 2.</label>
</paragraph>
<paragraph id="5F9A8089561523BAE5C498DA0CAB77FC">
<taxonomicName id="E7CBC362C7EA203893A138AA37E22F51" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="23E733581B8AE402450DD05613CDC213" italics="true">Meranoplus siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F12F101BA78C24AF7801198CDC39E9" rank="species">sp. nov.</taxonomicNameLabel>
(holotype,
<abbrev content-type="institution" id="ABBRID0EVKBI" xlink_title="Natural History Museum of the National Science Museum, Thailand">THNHM</abbrev>
- I- 00027303)
<emphasis id="F2D33E9113F2B645132A44AF759392FC" bold="true">A</emphasis>
head in full-face view
<emphasis id="E13810050E1544252C6FC20A11929F3F" bold="true">B</emphasis>
body in profile view
<emphasis id="C674461EC0DFE0E0A252835875F8D9B5" bold="true">C</emphasis>
promesonotal shield in dorsal view
<emphasis id="8BB84BD485D9F50C63F53EFDDE660F93" bold="true">D</emphasis>
body in dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMTAE" type="description">
<paragraph id="0A9D9F0513DE06E85BDC5ED68E8AF59D">
<heading id="72EBDFE0AF7131B33F4D30E463A566CA" reason="title">Measurements and indices.</heading>
</paragraph>
<paragraph id="325B4C39502DBB8F518EF38F15EEB56E">
<typeStatus id="68CE5BA508604682117F6F1FA003C8B0" type="holotype">Holotype</typeStatus>
worker:
<abbrev id="ABBRID0ESTAE" xlink_title="Head length">HL</abbrev>
0.63,
<abbrev id="ABBRID0EWTAE" xlink_title="Head width">HW</abbrev>
0.65,
<abbrev id="ABBRID0E1TAE" xlink_title="Mesosomal length">ML</abbrev>
0.62,
<abbrev id="ABBRID0E5TAE" xlink_title="Length of promesonotal shield">PML</abbrev>
0.48,
<abbrev id="ABBRID0ECUAE" xlink_title="Pronotal width">PW</abbrev>
0.63,
<abbrev id="ABBRID0EGUAE" xlink_title="Scape length">SL</abbrev>
0. 0.45,
<abbrev id="ABBRID0EKUAE" xlink_title="Total length">TL</abbrev>
2.58,
<abbrev id="ABBRID0EOUAE" xlink_title="Cephalic index">CI</abbrev>
102.38,
<abbrev id="ABBRID0ESUAE" xlink_title="Pronotum index">PMI</abbrev>
130.21,
<abbrev id="ABBRID0EWUAE" xlink_title="Scape index">SI</abbrev>
69.77.
<typeStatus id="1783B84942CF73D3C6E105AA0DAF0ACE" type="paratype">Paratype</typeStatus>
workers (
<emphasis id="FE421344EF6533CB60805DB54D48E2C5" italics="true">n</emphasis>
= 5):
<abbrev id="ABBRID0E3UAE" xlink_title="Head length">HL</abbrev>
0.600.65,
<abbrev id="ABBRID0EAVAE" xlink_title="Head width">HW</abbrev>
0.630.66,
<abbrev id="ABBRID0EEVAE" xlink_title="Mesosomal length">ML</abbrev>
0.600.65,
<abbrev id="ABBRID0EIVAE" xlink_title="Length of promesonotal shield">PML</abbrev>
0.460.51,
<abbrev id="ABBRID0EMVAE" xlink_title="Pronotal width">PW</abbrev>
0.620.65,
<abbrev id="ABBRID0EQVAE" xlink_title="Scape length">SL</abbrev>
0.430.48,
<abbrev id="ABBRID0EUVAE" xlink_title="Total length">TL</abbrev>
2.562.82,
<abbrev id="ABBRID0EYVAE" xlink_title="Cephalic index">CI</abbrev>
100106.45,
<abbrev id="ABBRID0E3VAE" xlink_title="Pronotum index">PMI</abbrev>
127136.26,
<abbrev id="ABBRID0EAWAE" xlink_title="Scape index">SI</abbrev>
67.7271.97.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEWAE" type="description">
<paragraph id="0197843C044D46479B6C6E9D7B60CC7A">
<heading id="7376F8D87419E83637A372972371734A" reason="title">Description of worker.</heading>
</paragraph>
<paragraph id="BC29D0DAA98BF619CACFC47B04597413">
<emphasis id="4E8E9D4A6854E88CFC8C3D610F65F592" bold="true" italics="true">Head</emphasis>
in full-face view subquadrate, slightly shorter than broad, with sides broadly convex, posterior margin distinctly convex, posterolateral corner bluntly angulate. Antennal scapes short, reaching level of posterior margin of compound eyes, apical half incrassate; antennal segment II slender, longer than each of segments III VI, and almost as long as III + IV + V; segment VI broader than each of segments II V. Clypeus roughly subrectangular, shorter than broad, its anterior margin feebly concave medially, while posterior clypeal margin almost straight. Mandibles subtriangular, masticatory margin with four teeth. Compound eyes large and convex when seen in full-face view, located laterally and well behind mid-length of head, with 8 or 9 ommatidia along longest axis, each facet hexagonal (Fig.
<figureCitation id="95E5C9C6264F7A881FDF70AA280C8511" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Characters used in key A Meranoplus siamensis (non-type worker, THNHM - I 00028942) B, F M. castaneus (non-type, THNHM - I- 00027323) C M. laeviventris (non-type, THNHM - I- 00027329) D, E M. bellii (lectotype, CASENT 0908933) G, H M. dlusskyi (in Zryanin 2015) I M. levis (holotype, CASENT 0902025) A eye B, C petiole and postpetiole in profile D, F, G, I mesosoma in profile E gaster in dorsal view H head in profile." figureDoi="10.3897/zookeys.1210.125990.figure6" httpUri="https://binary.pensoft.net/fig/1121266">6 A</figureCitation>
). Frontal lobes broad, its anterior corners right angled and lateral margin almost straight (Fig.
<figureCitation id="C0ABE8C4A4462D1CD8E67BA658AD2006" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 H</figureCitation>
). Frontal carinae long reaching posterolateral corners of head.
</paragraph>
<paragraph id="9CC1926907F7A29F57EC934CBEDB38ED">
<emphasis id="2626EB6C38972F2B6A8D45852D86342E" bold="true" italics="true">Mesosoma</emphasis>
in dorsal view, promesonotal shield distinctly shorter than broad, its lateral margin convex, serrate, margined and slightly overhanging mesosoma; lateral and posterior portions of promesonotal shield with translucent fins; posterior margin of promesonotal shield sinuate and distinctly concave; anterolateral corners of promesonotal shield bluntly angulate and posterolateral corners of promesonotal shield roundly angulate; promesonotal shield with two pairs of fenestrae laterally; metanotal groove absent. Declivity of propodeum almost invisible from above, mostly overhung by posterior margin of promesonotal shield (propodeal spines are visible in profile). Mesosoma in profile subquadrate, dorsal outline weakly convex, lateral face of mesosoma relatively flat; lateral portion of pronotum subtriangular; metapleuron not clearly demarcated from mesopleuron and lateral face of propodeum. Propodeal spines long and acute, longer than wide at its base, located at middle of propodeal declivity length.
</paragraph>
<paragraph id="DCC3A7CF532E82472CB08B109B5C40B7">
<emphasis id="3C4D83E759045507C9EACFD28A08B478" bold="true" italics="true">Petiole</emphasis>
in profile subtriangular. Subpetiolar process low, its ventral outline weakly convex, with small anterior denticle.
<emphasis id="C26FC4C0AD7485FE786C87F07AF9D077" bold="true">Postpetiole</emphasis>
in profile subquadrate, shorter than high; in dorsal view, distinctly shorter than broad, anterior margin weakly convex, posterior margin distinctly convex; dorsum of postpetiole somewhat flat, marginated with distinct ridge, posterior face convex.
<emphasis id="EC6BD9CC3310330B0F131B7FFDBD6728" bold="true">Gaster</emphasis>
about as large as head and mesosoma combined; first gastral tergite largest, in dorsal view, its anterior margin distinctly concave.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXAE" type="Sculpture">
<paragraph id="41A5AD7525DD6F3050E60415F100D57E">
<heading id="F025B50FB237C13E9FCB3B7387B576A6" reason="title">Sculpture.</heading>
</paragraph>
<paragraph id="36609F367F66BBCEB80EF88491B745FE">Mandibles striate, shiny. Antennal scapes superficially striate. Head dorsally sparsely reticulate-rugulose laterally, while median portion weakly sculptured; half posterior portion of antennal scrobes shagreened mixed with few transverse ridges. Promesonotal shield more weakly sculptured than dorsum of head, with median portion smooth, shiny, and lacking any rugae; in profile, upper one-third portion of pronotum shagreened, while lower two-third portion with sparse irregular ridges; upper one-third portion of mesopleuron shagreened, lower two-thirds longitudinal weakly striate; metapleuron, and lateral faces of propodeum somewhat smooth and shiny. Propodeum declivity shagreened. Petiole smooth and shiny, postpetiole somewhat smooth but posterior face of postpetiole scabrous. First gastral tergite superficially shagreened with smooth and shiny interspaces.</paragraph>
</subSubSection>
<subSubSection id="SECID0EOXAE" type="Pilosity and coloration">
<paragraph id="FDD269B1C6D96319840A30B5E64C8848">
<heading id="73FD6EB39F769CC50697E6BA09EBF2A8" reason="title">Pilosity and coloration.</heading>
</paragraph>
<paragraph id="F671CF166760EA74AD6ECABC4D9808F1">Dorsa of head and mesosoma with dense erect hairs mixed with sparse longer hairs; antennae with dense suberect hairs; in profile, lower two-thirds of pronotum with sparse suberect hairs; lower one-third of mesopleuron and metapleuron with sparse suberect hairs; area around propodeal spiracle with sparse suberect hairs; femora and tibiae with numerous long outstanding hairs as well; petiole with weakly sparse erect hairs on its anterior face and dorsum; postpetiole with dense long erect hairs, except anterior face without hairs; gaster with dense long erect hairs. Body mainly reddish brown; mandibles, antennae, legs, and tip of gaster yellowish brown.</paragraph>
</subSubSection>
<subSubSection id="SECID0ETXAE" type="distribution">
<paragraph id="3330F6505C73C414AB1635DA82236622">
<heading id="F9C903083BF635A8576E07AA22E68579" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="D394BCDB622408B70F901C6680F82643">
<collectingCountry id="EED82E839898C29066FBA2059CEBC9CF" name="Thailand">Thailand</collectingCountry>
(
<collectingRegion id="5BA110005FEF96DB147FCE20E961F6C2" country="Thailand" name="Uthai Thani">Uthai Thani</collectingRegion>
,
<collectingRegion id="468F491908DEDCD9DF5E9ECD5BD46A3C" country="Thailand" name="Chon Buri">Chonburi</collectingRegion>
,
<collectingRegion id="928B023BB2B2499A86DA72E8B24D11F5" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
and
<collectingRegion id="64B2A9D87D656713D38ADA9246DC2EC5" country="Thailand" name="Kanchanaburi">Kanchanaburi</collectingRegion>
Provinces, Fig.
<figureCitation id="17E00BB8B9D252C35321E264C4E55F98" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Meranoplus Thai species in Thailand." figureDoi="10.3897/zookeys.1210.125990.figure4" httpUri="https://binary.pensoft.net/fig/1121263">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4XAE" type="etymology">
<paragraph id="06BA138320A334DB589A3B845731CD32">
<heading id="50E3A6CB4F8F616BDF45A5FF1CDAC82A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="B676F321C37FFD9FE6A32B3807998A67">
The specific name is after
<collectingCountry id="5E82B882251E22B084B51C9C2D932A9E" name="Thailand">Thailand</collectingCountry>
where the
<typeStatus id="AA31E090850ABB3E8BCF4C11D642CA99">type</typeStatus>
locality is located;
<collectingCountry id="F9CB97864B53F80482CDC93D1F706532" name="Thailand">Thailand</collectingCountry>
was called “
<collectingCountry id="71D69D4E650EA7411A7F5322F6515670" name="Thailand">Siam</collectingCountry>
” in the past.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECYAE" type="habitat">
<paragraph id="DAC8889A37ED3A892482542600EFEE8C">
<heading id="8F833778ED9D10FCA5D5E1B52F609CCE" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="B24AD16D99B5691933540B4E73E59F20">
This species can be found in dry evergreen and dry dipterocarp forests. The specimens collected from northeastern
<collectingCountry id="176DB1C7321276F91DBB06244CB0040B" name="Thailand">Thailand</collectingCountry>
(colony code TH 22 - WJT- 264) nested in the soil. Workers moved slowly on the ground.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHYAE" type="Differential diagnosis">
<paragraph id="6E64FBD4D43E7C17BA9F7447BAA7BC7B">
<heading id="2CF3C5D30D176295120F6D8CBCC7BC22" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="07ED1906E23AA4A63CDDFC6830FA8AE6">
<taxonomicName id="6EE7BFE62DEA174A119C05117B632764" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="438613B95F77CD02EB112905FF1EA331" italics="true">Meranoplus siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7693456896032B20E12241EAE7D9375D" rank="species">sp. nov.</taxonomicNameLabel>
is a small species that is most similar to
<taxonomicName id="B1B8A4150E5724047C5D47D7EF054B64" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi" status="sp. nov.">
<emphasis id="3365C165042C9ACC3523D933867DB09D" italics="true">Meranoplus tanomtongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="692EBAF83DBC8760D5523B8BD9B33049" rank="species">sp. nov.</taxonomicNameLabel>
in general appearance, having a pair of fenestrae along each lateral margin of the promesonotal shield, and having a subrectangular postpetiole when seen in profile. However,
<taxonomicName id="49EB0C775205F1C64B5196DB80CBB339" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="3488191FB889826C3114447101F22AB8" italics="true">M. siamensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="960779883C41747E7EE71EE23EADE957" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="C169C345EA4D5FE0F6E8FDF4D5365474" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
by: 1) anterior corners of frontal lobes right angled and lateral margin almost straight (round and lateral margin weakly convex in
<taxonomicName id="37BEB037FCC118B81364F2B25D633456" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="D45A376386D67242570C1DF970A8A023" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
, see Figs
<figureCitation id="C900374FDAB46B6665CD18E0766100B9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 E, H</figureCitation>
for comparison); 2) compound eyes with 8 or 9 ommatidia along longest axis, each facet hexagon (each facet round or elliptical in
<taxonomicName id="520F0AFC48FA0B8AF070AB160E6D7B6A" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="890B998F22AC02E09A66883812097E93" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
, see Figs
<figureCitation id="33112B9773602C1FB036B82E39EFE084" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 G</figureCitation>
,
<figureCitation id="EAF4D0F100D565C039D51027C79F5AA8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Characters used in key A Meranoplus siamensis (non-type worker, THNHM - I 00028942) B, F M. castaneus (non-type, THNHM - I- 00027323) C M. laeviventris (non-type, THNHM - I- 00027329) D, E M. bellii (lectotype, CASENT 0908933) G, H M. dlusskyi (in Zryanin 2015) I M. levis (holotype, CASENT 0902025) A eye B, C petiole and postpetiole in profile D, F, G, I mesosoma in profile E gaster in dorsal view H head in profile." figureDoi="10.3897/zookeys.1210.125990.figure6" httpUri="https://binary.pensoft.net/fig/1121266">6 A</figureCitation>
for comparison); 3) dorsum of head weakly sculptured (dorsum of head entirely and distinctly reticulate in
<taxonomicName id="C8AB28AC33A3CC3FBA8FFB282D7BCBE6" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="C29C125C21BBAB698FB61A086B8E7325" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
, see Fig.
<figureCitation id="66F4B648BFB7864C3C626C74BD0F3BDE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 F, I</figureCitation>
for comparison); 4) dorsum of postpetiole somewhat flat, marginated with distinct ridge (shallowly concave, marginated with distinct ridge in
<taxonomicName id="6F7DF996FD1AD3A108B69E472326843B" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="877D5E3B1D3C641921A2C0C342B9E82C" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
); 5) entire head with dense short hairs mixed with sparse longer hairs (hairs along head margin clearly longer than hairs on middle of head in
<taxonomicName id="60EE2A878B1264973036512FBE3CE4F1" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanomtongi">
<emphasis id="96E8E7CE26A186560B1B877F635278E5" italics="true">M. tanomtongi</emphasis>
</taxonomicName>
, see Fig.
<figureCitation id="E545E7BE4F284ECA9D4AB75ACB3AA0B0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters used in key A, B, E Meranoplus tanomtongi (holotype, THNHM - I 00028903) F, G M. tanomtongi (non-type worker, THNHM - I- 00028904) C, D M. bicolor (non-type, THNHM - I- 00027270) H M. siamensis (holotype, THNHM - I- 00027303) I M. siamensis (non-type worker, THNHM - I- 00028942) A, C petiole and postpetiole in profile B, D petiole and postpetiole in dorsal view E, H anterior corners of frontal lobes F, I head in full-face view G eye." figureDoi="10.3897/zookeys.1210.125990.figure5" httpUri="https://binary.pensoft.net/fig/1121265">5 F, I</figureCitation>
for comparison).
</paragraph>
<paragraph id="302C3A398B1E829FEA36B0707F8EDC91">
The type series of
<taxonomicName id="B8F737C361D2250C0D3369C26CC4BA99" authorityName="Yodprasit &amp; Buddhakala &amp; Tasen &amp; Jaitrong" authorityYear="2024" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="915618B2E3887F3BED6E146BF94BAA2F" italics="true">M. siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0C6430CF761A275DD2ED130FDA0BD091" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to the non-type specimens from Central
<collectingCountry id="2EAA1F75D4245BF0358486A02C2EEFD9" name="Thailand">Thailand</collectingCountry>
(TH 02 - WJT- 039). However, the two colonies have some variations: 1) compound eyes with 9 ommatidia along longest axis in the
<typeStatus id="0EEA239F49797A4C1807310490F61D7E">type</typeStatus>
series (8 ommatidia in colony no. TH 02 - WJT- 039); 2) promesonotal shield shorter than broad in the
<typeStatus id="0B0165E3CD78BEC9D084972A6692D440">type</typeStatus>
series (almost as long as broad in colony no. TH 02 - WJT- 039); 3) posterior half of head with sparse and weak reticulation in the
<typeStatus id="6CCCE7957194F50AFF3EDC3602D484C4">type</typeStatus>
series (dense distinct reticulations in colony no. TH 02 - WJT- 039); 4) propodeal declivity somewhat shagreened in the
<typeStatus id="E64052533F837DE6B4E471D4036A25AA">type</typeStatus>
series (smooth and shiny in colony no. TH 02 - WJT- 039); 5) first gastral tergite superficially shagreened with smooth and shiny interspaces in the
<typeStatus id="DFE17F99F30987C595BCDA7177B0294E">type</typeStatus>
series (distinctly shagreened in colony no. TH 02 - WJT- 039). These characters are not clear enough to distinguish the two populations.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,635 @@
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<mods:title id="F9568824122E820DF308346D60851273">Morphological and molecular data warrant the description of a new species of the genus Scutiger (Anura, Megophryidae) from the Central Himalaya</mods:title>
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<mods:namePart id="641853EF445C5BC6925769063F76513A">Hofmann, Sylvia</mods:namePart>
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<mods:affiliation id="C4563C5AAF0B2E4E77E741FFA74A301B">Leibniz Institute for the Analysis of Biodiversity Change, Museum Koenig, 53113 Bonn, Germany &amp; Helmholtz-Centre for Environmental Research GmbH UFZ, Department Conservation Biology, 04318 Leipzig, Germany</mods:affiliation>
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<mods:namePart id="798A95CB41A405F39D9F563C8033E7DC">Jablonski, Daniel</mods:namePart>
<mods:nameIdentifier id="2427F5AA4D0733D152117309C739CBC7" type="ORCID">0000-0002-5394-0114</mods:nameIdentifier>
<mods:affiliation id="651336791BB18A27F7E2A1280C4FE6E9">Department of Zoology, Comenius University in Bratislava, Bratislava, Slovakia</mods:affiliation>
</mods:name>
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<mods:namePart id="5E5A29D05A8089A5B9D2834D1B4AC9B3">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="39BF908E54A0E1E0C492FA4D1BE7AA1C">General and Systematic Zoology, Institute of Biosciences, University of Rostock, 18055 Rostock, Germany</mods:affiliation>
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<mods:date id="6C013EC79A6193D34CEF000EA27FEF21">2024</mods:date>
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<mods:number id="0706FE2F8171A4314E4EAEC565789725">2024-08-26</mods:number>
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<subSubSection id="221E00DFAEAD37C638C46045AEEB5556" type="nomenclature">
<paragraph id="2AEDBEDB63F7A1870128275991C23CF8">
<taxonomicName id="0C965FBDEA7B45EE38E48D667D831C2F" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba" status="sp. nov.">
<emphasis id="C7FA5D29E8D121438BB62E2575DFA45A" italics="true">Scutiger kanjiroba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E65D80F2C0AE1934E9BEB9FFC3CBCC6E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="E85FDEC46AF6E5EF8869BF1998928AFB" type="description">
<paragraph id="5EC9E29110ABF43DD8EF2F54C4DF9D20">
<figureCitation id="00FC8ADEB57253866F7FE1C241FD293A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scutiger kanjiroba sp. nov. holotype NHME A 0576 / 99 in preservative (credit: Morris Flecks) A dorsal view of body B ventral view C dorsolateral view of body D ventral view of left foot." figureDoi="10.3897/zookeys.1210.127106.figure5" httpUri="https://binary.pensoft.net/fig/1120959">Fig. 5</figureCitation>
<xref id="C6C81288CA4DDFC4F1C0A13937C8B804" ref-type="supplementary-material" rid="S1">; Suppl. material 1: figs S 3, S 7, tables S 5, S 6 Suggested common name: Kanjiroba lazy toad</xref>
</paragraph>
</subSubSection>
<subSubSection id="D0050C2A3841CCF0714914452F70DF47" type="reference_group">
<paragraph id="49F2C5B121926C44FE1295658A440B0A">
<treatmentCitationGroup id="A8E0EC4CF8FD866F9B7A9E7660DB0323">
<treatmentCitation id="7E36DA1ED475C049324BF25F1E86F7A2">
<taxonomicName id="07AA5C0F64AE402CBC2EDA898F63E144" ID-CoL="6XYHC" authorityName="Dubois" authorityYear="1974" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B1DD065C623CD19EA5556EED4396C210" italics="true">Scutiger nepalensis</emphasis>
</taxonomicName>
:
<bibRefCitation id="8E43CA9CD74F3E1A16D1264B8DC0C820" DOI="10.1038/s41598-017-03395-4" author="Hofmann S &amp; Stoeck M &amp; Zheng Y &amp; Ficetola FG &amp; Li JT &amp; Scheidt U &amp; Schmidt J" etAl="et al." firstAuthor="Hofmann" issue="1" journalOrPublisher="Scientific Reports" pagination="3308" refId="B10" refString="Hofmann S, Stoeck M, Zheng Y, Ficetola FG, Li JT, Scheidt U, Schmidt J (2017) Molecular Phylogenies indicate a Paleo-Tibetan Origin of Himalayan Lazy Toads (Scutiger). Scientific Reports 7 (1): 3308. https://doi.org/10.1038/s41598-017-03395-4" title="Molecular Phylogenies indicate a Paleo-Tibetan Origin of Himalayan Lazy Toads (Scutiger)." volume="7" year="2017">Hofmann et al. (2017)</bibRefCitation>
</treatmentCitation>
, partim.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMBAG" type="type material">
<paragraph id="C14F8EDE73C838A7FD5DC08F8485589E">
<heading id="D47E8E5B9A049569AFB248BFB7E5EA8D" reason="title">Type material.</heading>
</paragraph>
<paragraph id="AEFC8AC6417085715F8F07C355ECB62D">
<materialsCitation id="DEB410059D6538F0E70FA98F5BC7B0D9" collectingDate="1997-05-31" collectionCode="NHME" country="Nepal" county="Dolpa district" elevation="2600" latitude="29.016666" location="Juphal" longLatPrecision="1229" longitude="82.78333" municipality="Tripurakot" specimenCode="A 0576" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="9207F1573C0E8647B08F137C126320DD" bold="true" italics="true">
<typeStatus id="4A0BAF000CDD05A12C9502A5A563E40F" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="32275E16D480E2D05086B59DF0A6CA94" name="Nepal">Nepal</collectingCountry>
<specimenCount id="3A8C2DBAA7A4964DED65CF2DE94208B3" count="1" type="male">1 ♂</specimenCount>
;
<abbrev id="ABBRID0EXBAG" xlink_title="Natural history museum">
<collectionCode id="688728331368515F49FB131FB82A9C58" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="C7D036917BEC9E791695F39DE4B8A2F0" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 0576</specimenCode>
/ 99; adult; collected at
<location id="38B985577E382CEC64C7389BF0979C92" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:38B985577E382CEC64C7389BF0979C92" country="Nepal" county="Dolpa district" latitude="29.016666" longLatPrecision="1229" longitude="82.78333" municipality="Tripurakot" name="Juphal">Juphal</location>
in the direction to
<collectingMunicipality id="AE9E9E0BCA44075B102D0E7421DFFA42">Tripurakot</collectingMunicipality>
,
<collectingCounty id="95DD44E9AF5BCD3B2D81FB58000A049A">Dolpa district</collectingCounty>
,
<collectingCountry id="A4CD952B765CBB34F7E1929D234F1724" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5BAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.783333,29.016667]}">
<geoCoordinate id="A8BC3C226B421DF46064BC50C37C33D2" degrees="29" direction="north" minutes="01" orientation="latitude" precision="925" value="29.016666">29 ° 01 ' N</geoCoordinate>
,
<geoCoordinate id="5D482429130A9A413745022B589EB082" degrees="82" direction="east" minutes="47" orientation="longitude" precision="925" value="82.78333">82 ° 47 ' E</geoCoordinate>
</named-content>
,
<elevation id="E44E4606B35FCF5EB449A28E91DBC95F" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<quantity id="3B813073D8A1F1F051FA29B164520FA6" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</quantity>
a. s. l.
</elevation>
) on
<date id="3FD6D80083177723BAA23FBF67B4AC34" value="1997-05-31">
<collectingDate id="84ADC990935AA1F5B8B8C5E7F9271FED" value="1997-05-31">31 / 05 / 1997</collectingDate>
</date>
by Marco Fischer; deposited in the
<abbrev id="ABBRID0EDCAG" xlink_title="Natural history museum">
<collectionCode id="6E25ED4EB4545AD59C80B37429FE5F52" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
</materialsCitation>
.
<materialsCitation id="D1D9115512F5645A415BB3B86DB7995C" country="Nepal" location="Nepal" specimenCount="1" typeStatus="paratype">
<emphasis id="C80A9CF5E71698883E1B88FCD7C6D3FC" bold="true" italics="true">
<typeStatus id="FC3A4FB988A69DA146FBB03834C88080" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="1785E5FF03E08A49BA596F0D3369931C" name="Nepal">Nepal</collectingCountry>
</materialsCitation>
<materialsCitation id="7450EA415237A01E6D9C46BFD2C50817" collectingDate="1997-06-03" collectingDateMax="1999-07-08" collectingDateMin="1997-06-03" collectionCode="NHME" country="Nepal" county="Marco Fischer" elevation="3400" latitude="29.35" location="Talphi" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" specimenCode="A 0570, A 1247, A 0574, A 1248, A 1249, A 1250, A 1251, A 1253, A 1254, A 2014" specimenCount="9" specimenCount-female="3" specimenCount-juvenile="5" specimenCount-subadult="1" typeStatus="paratype">
<specimenCount id="5997639054166E0B80FBBBB44E5ECD1F" count="1" type="female">1 ♀</specimenCount>
;
<abbrev id="ABBRID0EKCAG" xlink_title="Natural history museum">
<collectionCode id="84AAA16CEC76EC863ADCA9AA3F0CC26D" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="D15D6765704DAD42CE1DF775E679528F" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 0570</specimenCode>
/ 99; adult; and
<specimenCount id="385DB64652B2E364FE3879374E1AE6F2" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0EOCAG" xlink_title="Natural history museum">
<collectionCode id="45E34E7F9CCF613B4AE59AF40A5E28B3" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="B030CF96E0A3800F0B1AE2D16FE17E13" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1247</specimenCode>
/ 05; collected at
<location id="A4979B86A751EF0C6A2F113EA9BE61EE" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:A4979B86A751EF0C6A2F113EA9BE61EE" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Talphi">Talphi</location>
,
<location id="72D2C4B9DF10FB7E999EA134D9856C3C" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:72D2C4B9DF10FB7E999EA134D9856C3C" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="CC83190D910CB0A8AA503FF13AEA50E0" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVCAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.333333,29.300000]}">
<geoCoordinate id="4CE79640DC6AA037E51CEC0CE20A1DFA" degrees="29" direction="north" minutes="18" orientation="latitude" precision="925" value="29.3">29 ° 18 ' N</geoCoordinate>
,
<geoCoordinate id="188E7FCBB0B65904125B6A5AEBA8BC39" degrees="82" direction="east" minutes="20" orientation="longitude" precision="925" value="82.333336">82 ° 20 ' E</geoCoordinate>
</named-content>
,
<elevation id="6C49087E0DAE49635309F26D3DBBAD9F" metricMagnitude="3" metricUnit="m" metricValue="3.4" unit="m" value="3400.0">
<quantity id="BC79BB2D70F1E6F79CDCA67A1333D483" metricMagnitude="3" metricUnit="m" metricValue="3.4" unit="m" value="3400.0">3400 m</quantity>
a. s. l.
</elevation>
) on
<date id="615374A2FE88F4922A6EDFC6F7FC80A4" value="1997-06-16">
<collectingDate id="BBFE07A82C25ADF9E2B4442652C20827" value="1997-06-16">16 / 06 / 1997</collectingDate>
</date>
by Marco Fischer; deposited in the
<abbrev id="ABBRID0E1CAG" xlink_title="Natural history museum">
<collectionCode id="E4A4379D2142AD4166DF2BD3E886CB26" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="78D2FF9A97F290854A63CCAB7761C472" count="1" type="female">1 ♀</specimenCount>
;
<abbrev id="ABBRID0E5CAG" xlink_title="Natural history museum">
<collectionCode id="FBACFB9BBF769ED9023A24D7FCBDD5EA" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="80786773B33D50A98E3D04BA4A823D9C" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 0574</specimenCode>
/ 99; adult; collected at Kaigaon,
<location id="7ABB2BF705AFCF443B337996AD8DC603" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:7ABB2BF705AFCF443B337996AD8DC603" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="001DB8DEDAE5FA6D24FCDDC51F46CD0D" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFDAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.583333,29.100000]}">
<geoCoordinate id="C3A6E1DD7533D5688B021A14F91F2A5C" degrees="29" direction="north" minutes="06" orientation="latitude" precision="925" value="29.1">29 ° 06 ' N</geoCoordinate>
,
<geoCoordinate id="BF9E371813EA060FBDFCBCA170541A70" degrees="82" direction="east" minutes="35" orientation="longitude" precision="925" value="82.583336">82 ° 35 ' E</geoCoordinate>
</named-content>
,
<elevation id="9AA24E60FC089564EB80BFE4C33A1ED6" metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">
<quantity id="DFD4D5B6A73383D02D8CD18ED245595E" metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">3600 m</quantity>
a. s. l.
</elevation>
) on
<date id="F14E8A8201632372A08A7E4C1B6F865A" value="1997-06-03">
<collectingDate id="84BE485288DDA2E95AD07F580C136C29" value="1997-06-03">03 / 06 / 1997</collectingDate>
</date>
by Marco Fischer; deposited in the
<abbrev id="ABBRID0EKDAG" xlink_title="Natural history museum">
<collectionCode id="D3E3BB19F1FE56128BE16A94858AE365" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="DC9F445DB7DB31C934361A32A5C47542" count="1" type="female">1 ♀</specimenCount>
;
<abbrev id="ABBRID0EODAG" xlink_title="Natural history museum">
<collectionCode id="5150952FBAC1D4FAB5A95B5A6E2D6D9B" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="4078FA8EAA0F97B03EBF016513A4C9BE" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1248</specimenCode>
/ 05, adult, collected at
<location id="496E7187955F35C498CE6D192BB5F463" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:496E7187955F35C498CE6D192BB5F463" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Dhauli Lake">Dhauli Lake</location>
,
<location id="AA24E3CDC8814CC2D64A2CAC132A6EEE" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:AA24E3CDC8814CC2D64A2CAC132A6EEE" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="6367C11C98FF2AB27EDC6318DEF1A4B3" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVDAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.383333,29.350000]}">
<geoCoordinate id="B2F14B7E4802FF626F3C73AB265941D4" degrees="29" direction="north" minutes="21" orientation="latitude" precision="925" value="29.35">29 ° 21 ' N</geoCoordinate>
,
<geoCoordinate id="F99691D2116E0CAD239590CDCBE90CB4" degrees="82" direction="east" minutes="23" orientation="longitude" precision="925" value="82.38333">82 ° 23 ' E</geoCoordinate>
</named-content>
,
<elevation id="8103B374EC8DEAF32F3865847387FAE1" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">
<quantity id="44B1F62FE35840AA73B3758950FFE0AF" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">4400 m</quantity>
a. s. l.
</elevation>
) on
<date id="4E1219F7815E0EA59116D55E34F5522C" value="1999-07-05">
<collectingDate id="E34A228FE71B6685268998A1917DE07A" value="1999-07-05">05 / 07 / 1999</collectingDate>
</date>
by Marco Fischer, deposited in the
<abbrev id="ABBRID0E1DAG" xlink_title="Natural history museum">
<collectionCode id="BF4A09830C2DEAA4D892B9E400EB4863" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="31298F0123951CACB73BEBAFE19F21F0" count="1" type="subadult">1 subadult</specimenCount>
;
<abbrev id="ABBRID0E5DAG" xlink_title="Natural history museum">
<collectionCode id="640A3C0A19795E65978BEA1F9B53882C" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="6D5415B272709C24A12735394D9CD491" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1249</specimenCode>
/ 05; collected at
<location id="84ECA287212304F78CDD13939D1C08E6" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:84ECA287212304F78CDD13939D1C08E6" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Dhauli Lake">Dhauli Lake</location>
,
<location id="ABFF1DB8B0D1223AF00F5C05FA559872" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:ABFF1DB8B0D1223AF00F5C05FA559872" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="C60846C367B17ACDA8D96CF48822DA34" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFEAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.383333,29.350000]}">
<geoCoordinate id="0619E32F9C6F3668BCCA0D66F663FF67" degrees="29" direction="north" minutes="21" orientation="latitude" precision="925" value="29.35">29 ° 21 ' N</geoCoordinate>
,
<geoCoordinate id="EC42C86647C855830B73331BC4A3170F" degrees="82" direction="east" minutes="23" orientation="longitude" precision="925" value="82.38333">82 ° 23 ' E</geoCoordinate>
</named-content>
,
<elevation id="E7BA696FAFB9C336166EB99BADFFC4AC" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">
<quantity id="83AD69D09A9559A62DFE79DFACDDCC25" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">4400 m</quantity>
a. s. l.
</elevation>
) on
<date id="683FBFCF7DC85139614EDC99419B7DEE" value="1999-07-05">
<collectingDate id="8E3462088207E0971412E354D5837FE7" value="1999-07-05">05 / 07 / 1999</collectingDate>
</date>
by Marco Fischer, deposited in the
<abbrev id="ABBRID0EKEAG" xlink_title="Natural history museum">
<collectionCode id="944FA9FDDB55C2C5FF7AEF67DD9C2AEC" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
. 1 tadpole;
<abbrev id="ABBRID0EOEAG" xlink_title="Natural history museum">
<collectionCode id="F982B3A68112C0EEEA5E9D553E89FE4D" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="F2797152F263711AB2F217EC504F259B" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1250</specimenCode>
/ 05; collected at
<location id="9F3FDF60F41C496091C95FEA6F82C65B" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:9F3FDF60F41C496091C95FEA6F82C65B" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Dhauli Lake">Dhauli Lake</location>
,
<location id="C8F26323B631CF42983A219B4286F3F9" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:C8F26323B631CF42983A219B4286F3F9" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="C1A5D001441EBB2A1926CCF2F371EA30" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVEAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.383333,29.350000]}">
<geoCoordinate id="C9ACE49338B27EB9ACF11801829A53E8" degrees="29" direction="north" minutes="21" orientation="latitude" precision="925" value="29.35">29 ° 21 ' N</geoCoordinate>
,
<geoCoordinate id="327D173CFDC3BC32F1CB332CB7FD258A" degrees="82" direction="east" minutes="23" orientation="longitude" precision="925" value="82.38333">82 ° 23 ' E</geoCoordinate>
</named-content>
,
<elevation id="088ADCD71E1EF4B6B29BC6AA05660A2E" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">
<quantity id="7F58AE7B5E88121ED58AA19ADF047CF7" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">4400 m</quantity>
a. s. l.
</elevation>
) on
<date id="9988E8EE1B0181BC55A97A6C57ABA279" value="1999-07-05">
<collectingDate id="59B963B8C39E4CF41FC6C75791A79C30" value="1999-07-05">05 / 07 / 1999</collectingDate>
</date>
by Marco Fischer, deposited in the
<abbrev id="ABBRID0E1EAG" xlink_title="Natural history museum">
<collectionCode id="1CF879F2D32F861304577A314BB73CCF" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="E1778E7129C4B6692734AE09E9EF2678" count="2" type="juvenile">2 juveniles</specimenCount>
;
<abbrev id="ABBRID0E5EAG" xlink_title="Natural history museum">
<collectionCode id="92B3BB1A36CB011B47185F03B830C0F9" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="3A3A4E0F950911F767E47F207CA658D3" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1251</specimenCode>
/ 05 and A 1252 / 05; collected at
<location id="89A6163C4A8B881741660D54CDE5218C" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:89A6163C4A8B881741660D54CDE5218C" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Dhauli Lake">Dhauli Lake</location>
,
<location id="FFDD6FF4D045FEEAD8A07A5A0F23EED2" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:FFDD6FF4D045FEEAD8A07A5A0F23EED2" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="A8D83575783914F6838091D8A2C7F16B" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFFAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.383333,29.350000]}">
<geoCoordinate id="2C68868481A30780A6D419B40426F91C" degrees="29" direction="north" minutes="21" orientation="latitude" precision="925" value="29.35">29 ° 21 ' N</geoCoordinate>
,
<geoCoordinate id="987D37B311B062A167DB6A939821A91B" degrees="82" direction="east" minutes="23" orientation="longitude" precision="925" value="82.38333">82 ° 23 ' E</geoCoordinate>
</named-content>
,
<elevation id="2926595D5F3D4B03023D9B3AD108E8D6" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">
<quantity id="FD862CDCF1519C453FCDB169627753DB" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">4400 m</quantity>
a. s. l.
</elevation>
) on
<date id="BB53DC20F786EF857197B19695D26719" value="1999-07-05">
<collectingDate id="73AB0E5941FE72381D527960D32D320A" value="1999-07-05">05 / 07 / 1999</collectingDate>
</date>
by Marco Fischer, deposited in the
<abbrev id="ABBRID0EKFAG" xlink_title="Natural history museum">
<collectionCode id="AF0901C3103C5915D93107E3D110F9F7" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="AF9CDCA4BDADEC75E121A15DF5D84D04" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0EOFAG" xlink_title="Natural history museum">
<collectionCode id="D53C6A60AD2DD43D89290BB2E22CCE52" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="49127E56144CF6EAB38698D5B6D4ED5A" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1253</specimenCode>
/ 05; collected at Bumra [Bumramadichaur],
<location id="1CE778A5574A05E518DFAADB1D95481D" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:1CE778A5574A05E518DFAADB1D95481D" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="156B3DFDFBA2C5923AA338E8E4CD57B6" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVFAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.133333,29.383333]}">
<geoCoordinate id="B889DDE75F4A6EBE308D54634B4F5716" degrees="29" direction="north" minutes="23" orientation="latitude" precision="925" value="29.383333">29 ° 23 ' N</geoCoordinate>
,
<geoCoordinate id="A5C0398095FFB463DF34C7A04BA4DA8C" degrees="82" direction="east" minutes="08" orientation="longitude" precision="925" value="82.13333">82 ° 08 ' E</geoCoordinate>
</named-content>
) on
<date id="624BCCB254AC03A837F1B120B9DCF694" value="1999-06-22">
<collectingDate id="EEB4C3682D587EB1C76F20F0E5A527D8" value="1999-06-22">22 / 06 / 1999</collectingDate>
</date>
by Ulrich Scheidt, deposited in the
<abbrev id="ABBRID0E1FAG" xlink_title="Natural history museum">
<collectionCode id="5281202A483C3D369C9C5BBEED12ED04" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
.
<specimenCount id="C3AD4F3C9B942F03926B503633454942" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0E5FAG" xlink_title="Natural history museum">
<collectionCode id="AE9D8D6A4F7A1E18968A5F87C3461FAF" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="D7BD4805D48C42D8F82F29D52688A54E" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 1254</specimenCode>
/ 05; collected at
<location id="EEA3EF0BBA3F08492320833623466D86" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:EEA3EF0BBA3F08492320833623466D86" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Maharigaon">Maharigaon</location>
,
<location id="F095C0C2B4C7CEAF3B762A748DACBD3E" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:F095C0C2B4C7CEAF3B762A748DACBD3E" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Jumla district">Jumla district</location>
,
<collectingCountry id="B4AEB641E6BD5C2463155261E38034D6" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFGAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.383333,29.333333]}">
<geoCoordinate id="421F618B5F9EEA5B5CF6C9C61126C61A" degrees="29" direction="north" minutes="20" orientation="latitude" precision="925" value="29.333334">29 ° 20 ' N</geoCoordinate>
,
<geoCoordinate id="F2C059680511FD3D4FD5D2ED3B04D30D" degrees="82" direction="east" minutes="23" orientation="longitude" precision="925" value="82.38333">82 ° 23 ' E</geoCoordinate>
</named-content>
) on
<date id="516616C63BA6D7DCCA49B347AD8BE035" value="1999-07-08">
<collectingDate id="D5246394B39D63F0550ECDA38B044822" value="1999-07-08">08 / 07 / 1999</collectingDate>
</date>
by
<location id="C71CFB74E60617B945182DAA5D7CA7C5" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:C71CFB74E60617B945182DAA5D7CA7C5" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Andreas Weigel">Andreas Weigel</location>
, deposited in the
<abbrev id="ABBRID0EKGAG" xlink_title="Natural history museum">
<collectionCode id="E81FFF6E7FCF5B6B778083C1536F111D" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
. 1 metamorp;
<abbrev id="ABBRID0EOGAG" xlink_title="Natural history museum">
<collectionCode id="60EABBB1D405B3C7C60C199691D9401A" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
<specimenCode id="1EDAA82032B2134A0C753EFBE6AA4329" collectionCode="NHME" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">A 2014</specimenCode>
/ 13; collected near
<location id="A33131573DCCE0B6D5301CC5A0889887" LSID="urn:lsid:plazi:treatment:B71333991BE25E99897C955D6E7DB6A5:A33131573DCCE0B6D5301CC5A0889887" country="Nepal" county="Marco Fischer" latitude="29.35" longLatPrecision="1227" longitude="82.15" municipality="Jumla district" name="Khari La">Khari La</location>
,
<collectingMunicipality id="04D8C0595F7CEBA8C547AB815722805A">Jumla district</collectingMunicipality>
,
<collectingCountry id="61B00AD915A35D5AE3D397A9DF109930" name="Nepal">Nepal</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVGAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[82.150000,29.350000]}">
<geoCoordinate id="436DA5DF7D09ACCF41537E614CCF502F" degrees="29" direction="north" minutes="21" orientation="latitude" precision="925" value="29.35">29 ° 21 ' N</geoCoordinate>
,
<geoCoordinate id="04682E0B509DD4B08F14EC81B96B76B1" degrees="82" direction="east" minutes="09" orientation="longitude" precision="925" value="82.15">82 ° 09 ' E</geoCoordinate>
</named-content>
,
<elevation id="7BE71518342AB9BC2E55E63054965984" metricMagnitude="3" metricUnit="m" metricValue="3.285" unit="m" value="3285.0">
<quantity id="DA98C0580A40CA19B9637891C0021A47" metricMagnitude="3" metricUnit="m" metricValue="3.285" unit="m" value="3285.0">3285 m</quantity>
a. s. l.
</elevation>
) in 1999 by
<collectingCounty id="6A92FD385A4770126E7215C4EF719CD6">Marco Fischer</collectingCounty>
, deposited in the
<abbrev id="ABBRID0E1GAG" xlink_title="Natural history museum">
<collectionCode id="EBC1144E956089A321490DC3AD3BE02F" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5GAG" type="diagnosis">
<paragraph id="86D123C4F99E865A280F1A291F0FCA5A">
<heading id="F9A081DFB4B4847A6B485912A0FE4AAC" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B8DE90A284B9D55135163F3FDA1D1101">
<taxonomicName id="5C76E96D9E2C6F30C657CF5BE73CE56C" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba" status="sp. nov.">
<emphasis id="A82795F59F96FF93D91364268A5B2C62" italics="true">Scutiger kanjiroba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="989D25E5E1FCE761BFCF38E5429D7270" rank="species">sp. nov.</taxonomicNameLabel>
is assigned to the genus
<taxonomicName id="3979407DC3A758BFE52451389DB9EC25" authorityName="Theobald" authorityYear="1868" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="EE18BA212393F26B33B283BEFFEC7813" italics="true">Scutiger</emphasis>
</taxonomicName>
based in the above-mentioned morphological features and by its molecular phylogenetic position relative to other
<taxonomicName id="1E85310E32F7382BE9D15EB2098A9D8E" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="F201A210D693331D5DA0246811C1E1BF" italics="true">Scutiger</emphasis>
species.
</taxonomicName>
It can be separated from all other species of the genus
<taxonomicName id="81FFD6D87BE4E6371D335668E2913C04" authorityName="Theobald" authorityYear="1868" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="49A76EE4F4A91682F979B8C3AD0EBBEA" italics="true">Scutiger</emphasis>
</taxonomicName>
by the combination of the following characters (for character states in other species see section Morphological comparison, above): (1) moderate body size,
<abbrev id="ABBRID0EEIAG" xlink_title="Snout vent length">SVL</abbrev>
adult male
<quantity id="99E8A9DB0077884C6A89F92B7FC9BB3F" metricMagnitude="-2" metricUnit="m" metricValue="5.54" unit="mm" value="55.4">55.4 mm</quantity>
(
<emphasis id="C083DB93C34793F433E363BF4F1711AB" italics="true">n</emphasis>
= 1), adult female
<quantity id="972A6255E82515E7B2FA733018413A62" metricMagnitude="-2" metricUnit="m" metricValue="6.045" metricValueMax="6.67" metricValueMin="5.42" unit="mm" value="60.45" valueMax="66.7" valueMin="54.2">54.2-66.7 mm</quantity>
(
<emphasis id="D9CBF53D034F8FEFDB5D0F32F3876170" italics="true">n</emphasis>
= 3); (2) vomerine and maxillary dentation absent; (3) vocal sac absent; (4) subarticular tubercles absent; (5) presence of numerous small, densely arranged spines on the surface of fingers I, II and inner surface of finger III of males in breeding condition; (6) relative length of fingers 3&gt; 4&gt; 2&gt; 1; (7) spines on (inner) (fore) arms absent; (8) toes rudimentary webbed; (9) relative length of toes 4&gt; 3&gt; 5&gt; 2&gt; 1; (10) a pair of pectoral glands and a pair of axillary glands; (11) spines on pectoral and axillary glands present in males; (12) belly without spines; (13) tubercles / warts on dorsal and lateral surfaces of body present; (14) dorsal tubercles with black spines of males in breeding condition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMIAG" type="description">
<paragraph id="600CDFD78E0CDC377A0E35C618C6F3F4">
<heading id="FE597815AA8C16367CFB6F11DFD65392" reason="title">
Description of
<typeStatus id="DE3E8E767B7C384A851FA37B02514893" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="773092C9DD63D34FFA62A375343E8C0A">
(Fig.
<figureCitation id="12AF038F38E5540AAC35C4AE88F5321F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scutiger kanjiroba sp. nov. holotype NHME A 0576 / 99 in preservative (credit: Morris Flecks) A dorsal view of body B ventral view C dorsolateral view of body D ventral view of left foot." figureDoi="10.3897/zookeys.1210.127106.figure5" httpUri="https://binary.pensoft.net/fig/1120959">5</figureCitation>
): Measurements are provided in Suppl. material
<xref id="5E66902F42C379A623EB38276F13C6BF" ref-type="supplementary-material" rid="S1">1</xref>
: table S 6; all morphological measurements in mm. Adult male
<abbrev id="ABBRID0E1IAG" xlink_title="Snout vent length">SVL</abbrev>
55.4; head moderate, wider than length (
<abbrev id="ABBRID0E5IAG" xlink_title="head width">
<collectionCode id="207C8DC22F7CE7E843D79C14C761E697">HW</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0ECJAG" xlink_title="head length">HL</abbrev>
1.3); snout short (
<abbrev id="ABBRID0EGJAG" xlink_title="snout length">SL</abbrev>
7.9) and rounded, slightly protruding beyond jaw; canthus rostralis obvious; nostril dorsolateral, closer to tip of snout than eyes (
<collectionCode id="DAC9CA819ABF23B625FD485467E4AC71">SND</collectionCode>
3.4,
<abbrev id="ABBRID0EKJAG" xlink_title="eyelid-naris distance">END</abbrev>
4.0); loreal region slightly oblique and concave; internarial distance larger than distance from anterior margin of eye to nostril (
<abbrev id="ABBRID0EOJAG" xlink_title="internarial distance">
<collectionCode id="49636C51DA0F2BC26DA41A3B5E4204B9" country="USA" lsid="urn:lsid:biocol.org:col:15485" name="Indiana University" type="Herbarium">IND</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0ESJAG" xlink_title="eyelid-naris distance">
<collectionCode id="828C56829857589AD5670DF243B824B9">END</collectionCode>
</abbrev>
1.4); eye moderately in size (
<collectionCode id="57544D61AC05A9619FA45F3FE8905F49">EHD</collectionCode>
/
<abbrev id="ABBRID0EWJAG" xlink_title="head length">HL</abbrev>
0.4); pupil vertical; interorbital space flat; tympanum and tympanic ring absent; supratympanic ridge moderately thick, from posterior part of upper eyelids to shoulder; tongue oval; choanae oval, widely separated, visible when viewed from below; vomerine and maxillary teeth absent; vocal sac absent.
</paragraph>
<caption id="885286FA0DDC377283CC9E6BB3E08CDF" ID-DOI="10.3897/zookeys.1210.127106.figure5" ID-arpha="F60C8235-79D7-5294-8160-71323FC7F367" httpUri="https://binary.pensoft.net/fig/1120959" startId="F5">
<paragraph id="D2155080F74B2525EACCA72E634C5879">
<label id="D41993CC4071BD13627AA1A8EC6750A2">Figure 5.</label>
</paragraph>
<paragraph id="8C7B35D436C5613D4A70A0E3F9538A7F">
<taxonomicName id="52EF7D2C10C2AF7C12EE428F943AC181" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba" status="sp. nov.">
<emphasis id="C90E260234019942D4BBBC7BB3987E7D" italics="true">Scutiger kanjiroba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E21D427EB0CB6AB6BB45646CE30CA6A8" rank="species">sp. nov.</taxonomicNameLabel>
holotype
<abbrev id="ABBRID0ENKAG" xlink_title="Natural history museum">NHME</abbrev>
A 0576 / 99 in preservative (credit: Morris Flecks)
<emphasis id="62F9FECB6D244A6C402E5DBCB498CDCA" bold="true">A</emphasis>
dorsal view of body
<emphasis id="7F8E5F024F111792F942C83C8CF86682" bold="true">B</emphasis>
ventral view
<emphasis id="C61B2BEE19FCCCFEB0EE7CE8853A79FB" bold="true">C</emphasis>
dorsolateral view of body
<emphasis id="88712CC15E7037F91DA819C707EB9A5D" bold="true">D</emphasis>
ventral view of left foot.
</paragraph>
</caption>
<paragraph id="FE09EA217940E7BC50313C96B4BE9561">
Forelimbs strong; forearm of median length (
<abbrev id="ABBRID0EALAG" xlink_title="forearm length">
<collectionCode id="7A0D75B00AB00EA8FABC05F6B83A17C2">FAL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EELAG" xlink_title="Snout vent length">
<collectionCode id="5BC8825A8D69853695D1A39AE65C4D1F">SVL</collectionCode>
</abbrev>
0.25) and of similar length as the hand (
<abbrev id="ABBRID0EILAG" xlink_title="hand length">
<collectionCode id="F17ABC2C17C7B7BAC82B215C51FEF8E9" country="Germany" lsid="urn:lsid:biocol.org:col:15866" name="Martin-Luther-Universität" type="Herbarium">HAL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EMLAG" xlink_title="Snout vent length">
<collectionCode id="CB32C57CB3262F26850AA78FCD1D9C4A">SVL</collectionCode>
</abbrev>
0.24), without spines; fingers slender, without webbing and lateral dermal fringes; all fingertips rounded, not dilated; relative finger lengths: 3&gt; 4&gt; 2&gt; 1; subarticular tubercles absent; inner metacarpal tubercle flat and distinct, lager than the outer, indistinct metacarpal tubercle; nuptial spines on dorsal and lateral surface of first and second fingers, and on inner side of third finger.
</paragraph>
<paragraph id="A302CB96B227736999E0019294E8A82B">
Hindlimbs robust, moderate in length (
<abbrev id="ABBRID0ESLAG" xlink_title="tibia (shank) length">
<collectionCode id="8BE779DB498E520A04E2E690E6772015">TIBL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EWLAG" xlink_title="Snout vent length">
<collectionCode id="B2118E669631F3B1DE2FF78164B88D7C">SVL</collectionCode>
</abbrev>
= 0.38), thighs slightly shorter than shanks
<abbrev id="ABBRID0E1LAG" xlink_title="femur (thigh) length">
<collectionCode id="7D086790E52CDF45025F6944222AAC66">FEL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E5LAG" xlink_title="tibia (shank) length">
<collectionCode id="60A9291A9F511CE50A4662A26799F3D9">TIBL</collectionCode>
</abbrev>
= 1.1), heels do not touch when folded at right angles to the body (see also Fig.
<figureCitation id="9443F032BEB077E26DA286761776D5EA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scutiger kanjiroba sp. nov. holotype NHME A 0576 / 99 in preservative (credit: Morris Flecks) A dorsal view of body B ventral view C dorsolateral view of body D ventral view of left foot." figureDoi="10.3897/zookeys.1210.127106.figure5" httpUri="https://binary.pensoft.net/fig/1120959">5 A</figureCitation>
); foot longer than shank (
<abbrev id="ABBRID0EGMAG" xlink_title="foot length">
<collectionCode id="B4477A7CB227E88F7B52C9E01BC55428">FOL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EKMAG" xlink_title="tibia (shank) length">
<collectionCode id="E765E97833D130F821F86176AFFD2D58">TIBL</collectionCode>
</abbrev>
= 1.1); tips of toes round; toes not webbed and without lateral fringes; toes relative lengths 4&gt; 3&gt; 5&gt; 2&gt; 1; subarticular tubercles absent; moderately large inner metatarsal tubercle, outer metatarsal tubercle indistinct.
</paragraph>
<paragraph id="337E97EE5EEBD2F0E1A54A6A46564F86">Dorsal and lateral skin with distinct tubercles in preservative, each tubercle with creamy coloured keratinised tip; tubercles with small, keratinised tips present below and on supratympanic fold and on shank; forehead and surfaces of lower arm relatively smooth; throat and belly surface smooth; a pair of pectoral glands and a pair of axillary glands present on chest, pectoral glands only slightly larger than axillary glands, all covered by tiny spines.</paragraph>
</subSubSection>
<subSubSection id="SECID0EPMAG" type="Sexual dimorphism">
<paragraph id="6F8D23A27D36CC50230387BC79F0C881">
<heading id="4FA79C7B447836B8C354576A04D3C046" reason="title">Sexual dimorphism.</heading>
</paragraph>
<paragraph id="AD4840B580DD496CBD2805D702903616">In males, a pair of pectoral glands and a pair of axillary glands present on breast (absent in females), all of them covered by spines in breeding season.</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMAG" type="Colour in preservative">
<paragraph id="BDAEAE8DE019CF4C0D2CDC0A21633D02">
<heading id="D49B77EF5AB5049146174A2B7824AA3B" reason="title">Colour in preservative.</heading>
</paragraph>
<paragraph id="B4135778CECFC58E9FE9BCE1B130C3B6">The background colouration of the dorsum fades to grey, tubercles with creamy white point, dorsal surface of finger I, II and inner surface of finger III pale brown; ventral belly pale greyish; ventral limbs, ventral chest, and ventral head greyish pale brown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EZMAG" type="Tadpole">
<paragraph id="AC661E06A75958B7248E92CE1B3B9679">
<heading id="ABC0CBE7138D1164B0226BDC292589B0" reason="title">Tadpole.</heading>
</paragraph>
<paragraph id="2ACBCA928352F4EF5253149A0F2C4AD6">
One tadpole (
<abbrev id="ABBRID0E6MAG" xlink_title="Natural history museum">
<collectionCode id="C8899E9BC73C8EC58BE5BF81D8009726" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" type="">NHME</collectionCode>
</abbrev>
A 1250 / 05) of
<taxonomicName id="675C972C812DF41DCEE7AF014FD59A66" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba" status="sp. nov.">
<emphasis id="208DEDB61242A6DDFC4B732F0D4666F1" italics="true">Scutiger kanjiroba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5B084A4C9CAF0AF54F16E40028AE3545" rank="species">sp. nov.</taxonomicNameLabel>
was collected at stage 36 (
<bibRefCitation id="56AA0D6B22825EF9026FDD5F06ED184F" author="Gosner KL" journalOrPublisher="Herpetologica" pagination="183-190" refId="B9" refString="Gosner KL (1960) A simplified table for staging anuran embryos and larvae with notes on identification. Herpetologica 16: 183 - 190. https: // www. jstor. org / stable / 3890061" title="A simplified table for staging anuran embryos and larvae with notes on identification." url="https://www.jstor.org/stable/3890061" volume="16" year="1960">Gosner, 1960</bibRefCitation>
) (Suppl. material
<xref id="CE1A6D3764CC9161BC340B47B7BBC571" ref-type="supplementary-material" rid="S1">1</xref>
: fig. S 7);
<abbrev id="ABBRID0ESNAG" xlink_title="labial tooth row formula">
<collectionCode id="CD2E97CD939EDE7B75B9735AFDE0CBB6">LTRF</collectionCode>
</abbrev>
: 1: 2 + 2 / 2 + 2: 1.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWNAG" type="distribution">
<paragraph id="098E5810BB19FB796CB3FC3E1B04C5F7">
<heading id="81226F257932D66E3DE6BED0289BC687" reason="title">Distribution and ecological notes.</heading>
</paragraph>
<paragraph id="278567F2710705D4CD89257304C7091F">
<taxonomicName id="CE90C865F67881E86B28365A48C511D8" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba">
<emphasis id="EA3EF48ADDBAD5D25B6509E943DF2219" italics="true">Scutiger kanjiroba</emphasis>
</taxonomicName>
is a forest-dwelling species that is currently known from high-montane areas of the Kanjiroba Himal at altitudes between ~ 3300 and
<quantity id="0F02C86A7584D5394843C0BB0ED6375A" metricMagnitude="3" metricUnit="m" metricValue="4.4" unit="m" value="4400.0">4400 m</quantity>
a. s. l. in west
<collectingCountry id="724171A7D55A7C1D275059153E6DA04C" name="Nepal">Nepal</collectingCountry>
(Fig.
<figureCitation id="0628918BB233289709986D1F7B2CF28F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overview of the Himalaya-Tibet orogenic area and occurrences of Scutiger species. Holotype localities are indicated by symbols with a central black dot. Some species lack the type locality, others could only be approximated due to imprecise information in the original description." figureDoi="10.3897/zookeys.1210.127106.figure1" httpUri="https://binary.pensoft.net/fig/1120955">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELOAG" type="etymology">
<paragraph id="D44E528401385A8D0B511BF3C7E2DB9E">
<heading id="2819BD7C280F7729C366CBB13504B671" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="58275F0C9D089B41C27679C20ED1F117">
The specific epithet
<taxonomicName id="56C6D193B526EFCDE0D92FCCE0E66E58" authorityName="Hofmann &amp; Jablonski &amp; Schmidt" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="kanjiroba">
<emphasis id="12C3736A34FB5F34C9C16A0BDBFE7B8A" italics="true">kanjiroba</emphasis>
</taxonomicName>
is a geographical proper name referring to the Kanjiroba Himal in
<collectingCountry id="09EDFD029D49E9290E4CC3501AD1C9F4" name="Nepal">Nepal</collectingCountry>
. The southern area of the Kanjiroba Himal is so far the only known occurrence region of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,480 @@
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<mods:title id="C27CD045BF85E8F3846F6832072782A9">Two new species and a new record of the ant genus Meranoplus Smith, 1853 (Hymenoptera, Formicidae, Myrmicinae) from Thailand</mods:title>
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<mods:namePart id="9D7E02EDEEA3456E1D671093C447DB53">Yodprasit, Kuntima</mods:namePart>
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<mods:namePart id="4EB017680FF16FAA6D036E9A39AF969B">Buddhakala, Nopparat</mods:namePart>
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<mods:affiliation id="4E34D70B9E186C711160B1C3F8E48FD8">Division of Biology, Faculty of Science and Technology, Rajamangala University of Technology Thanyaburi, Pathum Thani, 12120 Thailand &amp; Office of Natural Science Research, National Science Museum, 39 Moo 3, Khlong 5, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
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<treatment id="FC6BDA712E96530F85B97DC52E813717" ID-arpha="FC6BDA71-2E96-530F-85B9-7DC52E813717" LSID="urn:lsid:plazi:treatment:FC6BDA712E96530F85B97DC52E813717" httpUri="http://treatment.plazi.org/id/FC6BDA712E96530F85B97DC52E813717">
<subSubSection id="30FBD0CE26C6F155939CDFF5884773C5" type="nomenclature">
<paragraph id="BA6532F04891C7E44F6DD5BB328CA494">
<taxonomicName id="16B6ABA499A0A2A50C11D0E78A279C3C" authority="Schodl, 1998" authorityName="Schodl" authorityYear="1998" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="8939BBD9F203B70CA2C26B8625EBA3A3" italics="true">Meranoplus malaysianus</emphasis>
<bibRefCitation id="EEBC5FDFDBD2F46272FABF1144D9351F" author="Schödl S" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien Serie B für Botanik und Zoologie" pagination="361-394" refId="B26" refString="Schodl S (1998) Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae). Annalen des Naturhistorischen Museums in Wien Serie B fur Botanik und Zoologie 100: 361 - 394." title="Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae)." volume="100" year="1998">Schödl, 1998</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3D25969AA44579A6BC85BDF19086196D" type="description">
<paragraph id="E96F7F26EE9D0F53B3D4A64813E61DE1">
<figureCitation id="4BA520FD4B1C9A53F009B72AA6E12089" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Meranoplus malaysianus A, C, E holotype (CASENT 0902029) B, D, F non-type worker (THNHM - I- 00028943) A, B head in full-face view C, D promesonotal shield in dorsal view E, F body in profile." figureDoi="10.3897/zookeys.1210.125990.figure1" httpUri="https://binary.pensoft.net/fig/1121260">Figs 1 A F</figureCitation>
,
<figureCitation id="B1B5BBC724E83D12DDCD27FCDAD044EF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Characters used in key A, B Meranoplus levis (holotype, CASENT 0902025) C, D M. mucronatus (non-type, THNHM - I- 00027335) E, H M. borneensis (paratype, CASENT 0902030) F, I M. malaysianus (non-type, THNHM - I- 00028943) G M. periyarensis (in Bharti and Akbar 2014) A eye B C head in profile D, E, F, G promesonotal shield in dorsal view H, I petiole and postpetiole in profile." figureDoi="10.3897/zookeys.1210.125990.figure7" httpUri="https://binary.pensoft.net/fig/1121267">7 F I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="83489838666F1C9639C94730C3948ADC" type="reference_group">
<paragraph id="7174C900E00515F8A7C399B3B00F78DE">
<treatmentCitationGroup id="130A047327C837598007BA69644A18C5">
<treatmentCitation id="19A720C17CA2BF54357BD94E3C23EC8A" author="Schödl S" page="385" year="1998">
<taxonomicName id="92BB15FBA4881462050ED8C2F1B858EF" authority="Schodl, 1998: 385" authorityName="Schodl" authorityPageNumber="385" authorityYear="1998" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="38E67B03A1F6B3FA11A2B0190E4FAB27" italics="true">Meranoplus malaysianus</emphasis>
<comment id="E56F5620395F77FEFD329B60E60D574A">
<bibRefCitation id="4F5599C9F7F74DC913A0A0EFB5056B46" author="Schödl S" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien Serie B für Botanik und Zoologie" pagination="361-394" refId="B26" refString="Schodl S (1998) Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae). Annalen des Naturhistorischen Museums in Wien Serie B fur Botanik und Zoologie 100: 361 - 394." title="Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae)." volume="100" year="1998">Schödl, 1998: 385</bibRefCitation>
, figs 4, 18, 32 (workers and queen).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="810C4C30D7CE79807566A7F9B20EF2CA">
<treatmentCitationGroup id="A6CE890A04C53CE5103AEA61BF164D91">
<treatmentCitation id="3A49430C4FE45BA46BA0A60BE5CF76EB">
<taxonomicName id="B755A1CACDBB6856AF6F2D6229EE0197" authorityName="Schodl" authorityYear="1998" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="3C033C454EAB7E257FAD05A3CB2E1E15" italics="true">Meranoplus malaysianus</emphasis>
</taxonomicName>
<comment id="448A50D2CCDA51A530E6D45D1B0D9FD0">
:
<treatmentCitation id="622ADFA50FF6794E797EB507D3EE1AD0" author="Pfeiffer M &amp; Mezger D &amp; Hosoishi S &amp; Bakhtiar EY &amp; Kohout RJ" page="47" year="2011">
<bibRefCitation id="0F2B62035DD691DAF655329A2DE8DF2F" DOI="10.20362/am.004002" author="Pfeiffer M &amp; Mezger D &amp; Hosoishi S &amp; Bakhtiar EY &amp; Kohout RJ" etAl="et al." firstAuthor="Pfeiffer" journalOrPublisher="Asian Myrmecology" page="47" pagination="9-58" refId="B24" refString="Pfeiffer M, Mezger D, Hosoishi S, Bakhtiar EY, Kohout RJ (2011) The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4: 958. https://doi.org/10.20362/am.004002" title="The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list." volume="4" year="2011">Pfeiffer et al. 2011: 47</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="97522C31B1DDB191BDEC273B6B065E7B" author="Satria R &amp; Herwina H" page="83" year="2020">
<bibRefCitation id="8889581F014831B9023EFE48FF8157CC" DOI="10.2991/absr.k.200807.020" author="Satria R &amp; Herwina H" firstAuthor="Satria" journalOrPublisher="Advances in Biological Sciences Research" page="83" pagination="82-84" refId="B25" refString="Satria R, Herwina H (2020) New distribution record of ants species (Hymenoptera: Formicidae) to the fauna of Sumatra Island, Indonesia. Advances in Biological Sciences Research 10: 8284. https://doi.org/10.2991/absr.k.200807.020" title="New distribution record of ants species (Hymenoptera: Formicidae) to the fauna of Sumatra Island, Indonesia." volume="10" year="2020">Satria and Herwina 2020: 83</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKFAE" type="Types">
<paragraph id="8DC371096F381A92133B8FFEAE1C1EB7">
<heading id="657AEDB1FC529812CCD37987076ADB99" reason="title">Types.</heading>
</paragraph>
<paragraph id="CB1BABA28FBA18772FA7112FED4E980D">
<materialsCitation id="E3FAAE75C4A68A7EA25B92E0BF33562D" collectingDate="1973-10-08" collectionCode="BMNH" collectorName="B. Bolton" country="Malaysia" location="West" specimenCode="CASENT 0902029" specimenCount="1" stateProvince="Kuala Lumpur" typeStatus="holotype">
<emphasis id="EA745ABAEFEADD848547055C4F777D14" bold="true" italics="true">
<typeStatus id="6018B0247EB6B40AE8B22B6553120AAA" type="holotype">Holotype</typeStatus>
</emphasis>
• (
<named-content id="91D13F6FF12BAB03EA30BBCE15330C11" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
<collectionCode id="EFEE13360C8B9D598A33D727139FE010">BMNH</collectionCode>
</named-content>
,
<ext-link id="B31C2DFAFA36C127576351C9E1B2F2E9" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0902029" xlink_type="simple">
<specimenCode id="A5814B5A7625275EFD1AB5A93CFFB4D2" collectionCode="CASENT">CASENT 0902029</specimenCode>
</ext-link>
, images examined),
<location id="5D6B820155EE23AFF052ADB66FC3496A" LSID="urn:lsid:plazi:treatment:FC6BDA712E96530F85B97DC52E813717:5D6B820155EE23AFF052ADB66FC3496A" country="Malaysia" name="West" stateProvince="Kuala Lumpur">West</location>
<collectingCountry id="38D464106A935D47B7B5D228FDE4F0A5" name="Malaysia">Malaysia</collectingCountry>
(Malaya),
<collectingRegion id="36444064C82FA3D5D290454A2BF90857" country="Malaysia" name="Kuala Lumpur">Kuala Lumpur</collectingRegion>
,
<date id="8937AA47B70E81C89B1250D913529E94" value="1973-10-08">
<collectingDate id="DCE2E5D670D340FBB242CFC9C973BC4C" value="1973-10-08">8 October 1973</collectingDate>
</date>
,
<collectorName id="8E91280376E3DED7FF2FC5598767AEA3">B. Bolton</collectorName>
</materialsCitation>
.
<materialsCitation id="A1D355D661643961386C7027667D43F0" collectingDate="1973-10-08" collectionCode="BMNH, NHMW" collectorName="B. Bolton" country="Malaysia" location="West" specimenCount="11" specimenCount-queen="1" specimenCount-worker="10" stateProvince="Kuala Lumpur" typeStatus="paratype">
<emphasis id="1F602AAA15B0DC9B7460D6D566AF01E3" bold="true" italics="true">
<typeStatus id="BE2F19BE277B47F27807E334EDFD671B" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="A4492CF199247E90228B319E83A5222E" count="10" type="worker">10 workers</specimenCount>
and
<specimenCount id="A213D6F73D66B78DE9E5E32ACE2C428C" count="1" type="queen">1 queen</specimenCount>
, same locality data as
<typeStatus id="BF82C155365F1E249747412571055134" type="holotype">holotype</typeStatus>
(
<named-content id="90548DBAC76EAC72ED6842E3ED30A2D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
<collectionCode id="67E797BFA2C16F2C5BE44D64D13A303E">BMNH</collectionCode>
</named-content>
,
<named-content id="E97D33D7E0677FA8737DFAD9E44D22A2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
<collectionCode id="E56AEDBE0B7B703E89554EE76EE8DF18" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="B475B18BE95E593812BDC9D5FDCCD040" collectingDate="1973-10-08" collectionCode="NHMB, NHMW" collectorName="K. Lumpur &amp; Bolton" country="Malaysia" determinerName="B. Bolton" location="Malaysia" specimenCount="3" specimenCount-worker="3" typeStatus="paratype">
<specimenCount id="E1DAC6B8D19B59EB536489CD7DCC3DCB" count="3" type="worker">3 workers</specimenCount>
,
<collectingMethod id="6B6F697929F97AC18D367B0D42DB2129">Berlese funnel</collectingMethod>
,
<collectingCountry id="4524409DD41BD2F1FFB122252A6AAD12" name="Malaysia">Malaysia</collectingCountry>
(MALAYA),
<collectorName id="9F3C672C28171941630D4CCE69E9C11D">K. Lumpur</collectorName>
,
<date id="61D6F809A8FFA6626EF218D5775F8DBB" value="1973-10-08">
<collectingDate id="377EB04AD6FD775756C8E289B375B9BB" value="1973-10-08">8 October 1973</collectingDate>
</date>
, B.
<collectorName id="999E63524BEBB38CC335F38C9B10C48B">Bolton</collectorName>
,
<taxonomicName id="87F08BDBA4ECFE6F23DCEF7A0A28FA15" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5F050C6C55A217674259C9A8CA2EB091" italics="true">Meranoplus</emphasis>
sp.
</taxonomicName>
det.
<determinerName id="250B703A6F0B54369EBB67F3A4AF0D33">B. Bolton</determinerName>
, 1974 (
<named-content id="D809BDC80D8DE6F550946E1DFFF0B311" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-basel" xlink_title="Naturhistorisches Museum, Basel, Switzerland">
<collectionCode id="D0AC6289E2516DC3D73123F6477F3B20" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" type="">NHMB</collectionCode>
</named-content>
,
<named-content id="74A06E30E8BE5FDD78CF6FC9F5178482" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
<collectionCode id="0CD0A1AA33AD48436E0723A90D5EAF6B" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="7EC5DB49AD6A522B387DA086E3F418C8" collectingDate="1994-11" collectionCode="BMNH, NHMW" collectorName="Neg. Sembilan Pasoh For. Res. &amp; M. Brendell &amp; K. Jackson &amp; S. Lewis" country="Malaysia" location="Malaysia" specimenCount="5" specimenCount-worker="5" typeStatus="paratype">
<specimenCount id="EBD656B3F663001C4B47CD700D906D48" count="5" type="worker">5 workers</specimenCount>
,
<collectingCountry id="B331C58955A1180FF7ADB2DFA06E9F86" name="Malaysia">Malaysia</collectingCountry>
<collectorName id="8F656D50D5C494771D72717ADAE9DB19">Neg. Sembilan Pasoh For. Res.</collectorName>
<date id="92709978081F40AE5A05123D2AF7F751" value="1994-11">
<collectingDate id="1FBC3FE1090E05A1542321FF991B53FA" value="1994-11">November 1994</collectingDate>
</date>
, litt (= litter?) sample
<collectorName id="50B2ECBC33BEA21A57C87CF0D49A9770">M. Brendell</collectorName>
,
<collectorName id="DA1E2F743910F5CC57B69048A8B0F873">K. Jackson</collectorName>
,
<collectorName id="EFF4BEAA72A28E1B34CC23FD9C7AD045">S. Lewis</collectorName>
(
<named-content id="76943853217295B6415F0109C8787A6B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London, United Kingdom">
<collectionCode id="6CC2760513BCD2C1D7D2B6D567BDBF01">BMNH</collectionCode>
</named-content>
,
<named-content id="D11F149653C92B479FB58B32BF0A10D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
<collectionCode id="8E38EB8BD55D9F02378C878518B45564" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="C0F8250A53550C043AC2CF864C606232" collectionCode="MCZC, NHMW" collectorName="Damm. &amp; Depok" country="Malaysia" location="Museum of Comparative Zoology" specimenCount="6" specimenCount-male="1" specimenCount-worker="4" typeStatus="paratype">
<specimenCount id="0832C349626987F35D2ACD491E93B446" count="4" type="worker">4 workers</specimenCount>
, 2 queens,
<specimenCount id="214CD32D4D521EDB9B4D43C1AE83CF55" count="1" type="male">1 male</specimenCount>
(head missing),
<collectorName id="A0DEA399269B3F57715F6FBB9834859C">Damm.</collectorName>
,
<collectorName id="6206A8AE018E89CB537E34D7E5D4C892">Depok</collectorName>
7.1. [
<specimenCount id="209E3419CEB3185A115E75A3E4150063" count="1" type="generic">1 ex.</specimenCount>
30.111.] 1923,
<named-content id="F012C9C8D355288D6909C33E708C3D46" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Cambridge, USA">MCZC</named-content>
<location id="D5EE284312C1964A2E9749987DC54DC4" LSID="urn:lsid:plazi:treatment:FC6BDA712E96530F85B97DC52E813717:D5EE284312C1964A2E9749987DC54DC4" country="Malaysia" name="Museum of Comparative Zoology">Museum of Comparative Zoology</location>
(
<named-content id="123A011227E381F1DD9B224282E2FF1E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-comparative-zoology" xlink_title="Museum of Comparative Zoology, Cambridge, USA">
<collectionCode id="1DD7E106A3F2FD4CFD0AC9F9FB686B9C">MCZC</collectionCode>
</named-content>
,
<named-content id="14A1CBA93B41CB2F6F751A178CF08304" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
<collectionCode id="E102D06FE1E17D73F823463AFC693539" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="6FC0D225FB66574A76140511DF3A8454" collectionCode="NHMW" collectorName="Kebun Raya &amp; Bogor &amp; Janaury &amp; F. Ito" country="Indonesia" location="Indonesia" specimenCount="3" specimenCount-worker="3" typeStatus="paratype">
<specimenCount id="50B6AFB5C28C95F3BB5FE5F3AD912D13" count="3" type="worker">3 workers</specimenCount>
, F 192-387,
<collectorName id="DD8F4C143E5BE02FE5AD1B3F85448E71">Kebun Raya</collectorName>
,
<collectorName id="69AF87E08546EE1D49417F5D5D47395A">Bogor</collectorName>
, W-Java,
<collectingCountry id="C1B1D38FEE20F53B07CAE27E4A5ACD06" name="Indonesia">Indonesia</collectingCountry>
, 1131
<collectorName id="226D7933873704E2DC571BECF97BD0CB">Janaury</collectorName>
1992,
<collectorName id="F6AD61A56BBF74A4D3294B941D3385ED">F. Ito</collectorName>
(
<named-content id="CDFC501E73ADDD3DAB38AE84E9392BE1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
<collectionCode id="829843A4BD9E1E7CF1333D7EAC5E3A64" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2HAE" type="type material">
<paragraph id="C29EC9EFCB3FE06211B8A5BD8EB59B88">
<heading id="30F222DE509AD59D51971D48EC02F961" reason="title">Non-type material examined.</heading>
</paragraph>
<paragraph id="F97B825049A3C01B3DB503CB0D9E5878">
<emphasis id="37982CB806A1D71FF58B866C5ED4210D" bold="true">Southern</emphasis>
: •
<materialsCitation id="7F25B858B2BAE140DBCAE32B5EF38B81" collectingDate="2002-07-27" collectionCode="THNHM" collectorName="N. Noon-anant" location="Thung Tam Sao Subdistrict" municipality="Hat Yai District" specimenCount="1" specimenCount-worker="1" stateProvince="Songkhla">
<specimenCount id="07715687E4D2B52776185278B4352B59" count="1" type="worker">1 worker</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EDIAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="4DAB3D4B01FF56F3412AA7D5E5796C1E">THNHM</collectionCode>
</abbrev>
- I- 00028943,
<abbrev content-type="institution" id="ABBRID0EIIAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="265BEBFFFD77121A4650F05B90E004E6">THNHM</collectionCode>
</abbrev>
),
<collectingRegion id="5804AD7058C051A3445F9563F7180CD1" country="Thailand" name="Songkhla">Songkhla Province</collectingRegion>
,
<collectingMunicipality id="E6FEDC06E64DC448CD6BBDB3E88873FC">Hat Yai District</collectingMunicipality>
,
<location id="4B0680B5CDECCA7B89EA9BE0D5915503" LSID="urn:lsid:plazi:treatment:FC6BDA712E96530F85B97DC52E813717:4B0680B5CDECCA7B89EA9BE0D5915503" municipality="Hat Yai District" name="Thung Tam Sao Subdistrict" stateProvince="Songkhla">Thung Tam Sao Subdistrict</location>
, evergreen rain forest,
<date id="597B2D5C14E39D63236C2B219C4F239F" value="2002-07-27">
<collectingDate id="389F3C438E98A6FB65014C273C88EC84" value="2002-07-27">27 July 2002</collectingDate>
</date>
,
<collectorName id="5C6982A239BCAC643C23180721C0F208">N. Noon-anant</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8EA87095B0A0C5D19A85CB62C54A8CED" collectingDate="2001-09-25" collectionCode="THNHM" collectorName="R. Poonjampa" latitude="5.8455" location="Lo Chood Subdistrict" longLatPrecision="7" longitude="101.8756" municipality="Wang District" specimenCount="1" specimenCount-worker="1" stateProvince="Narathiwat">
<specimenCount id="E7745A0D3E34FE680E2CEB161F9662CE" count="1" type="worker">1 worker</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0ENIAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="E417265057115D1493A784C919454F8A">THNHM</collectionCode>
</abbrev>
- I- 00027334,
<abbrev content-type="institution" id="ABBRID0ESIAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="EF1D3605EB947EF222EC03D97100BBFA">THNHM</collectionCode>
</abbrev>
),
<collectingRegion id="40257D3433E805E56D3A44024C815C61" country="Thailand" name="Narathiwat">Narathiwat Province</collectingRegion>
,
<collectingMunicipality id="8A093F5FCC3717DEF3B07C233F1BB043">Wang District</collectingMunicipality>
,
<location id="360DC17A63F172D9FEE2EEC6395337AD" LSID="urn:lsid:plazi:treatment:FC6BDA712E96530F85B97DC52E813717:360DC17A63F172D9FEE2EEC6395337AD" latitude="5.8455" longLatPrecision="7" longitude="101.8756" municipality="Wang District" name="Lo Chood Subdistrict" stateProvince="Narathiwat">Lo Chood Subdistrict</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1IAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.875600,5.845500]}">
<geoCoordinate id="1F1EDA1EC78F5EBBBC264BE9E1C11954" degrees="5.8455" direction="north" orientation="latitude" precision="5" value="5.8455">5.8455 ° N</geoCoordinate>
,
<geoCoordinate id="D0F509FE6D3F8153107BC55D754E8EDF" degrees="101.8756" direction="east" orientation="longitude" precision="5" value="101.8756">101.8756 ° E</geoCoordinate>
</named-content>
,
<date id="C65571F49743B56081F69688D4765EB0" value="2001-09-25">
<collectingDate id="9D8A58ED615A33286FA2CB466CD3256D" value="2001-09-25">25 September 2001</collectingDate>
</date>
,
<collectorName id="D6D862219E2582ADED3E6A95E641E044">R. Poonjampa</collectorName>
leg., hand collecting
</materialsCitation>
<materialsCitation id="61456437A9465273A2F03BEB6F1AE962" collectingDate="2001-09-25" collectionCode="AMK" collectorName="S. Hasin" latitude="5.8455" location="Lo Chood Subdistrict" longLatPrecision="7" longitude="101.8756" municipality="Wang District" specimenCount="1" specimenCount-worker="1" stateProvince="Narathiwat">
<specimenCount id="E94DA709AD0A95B6DAC53253AD0E118F" count="1" type="worker">1 worker</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0E6IAE" xlink_title="Ant Museum, Faculty of Forestry, Kasetsart University, Thailand">
<collectionCode id="EB3E58A68EF4DF819A7D50E1F20CB8E9">AMK</collectionCode>
</abbrev>
), same locality and date,
<collectorName id="74EF7C68AA22F778A5B5FE4252568584">S. Hasin</collectorName>
leg., general collection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEJAE" type="description">
<paragraph id="8933817A7F71241EDBB108FDCE7F0F77">
<heading id="199B491CE465E24A21DB88164D2600F7" reason="title">Measurements and indices.</heading>
</paragraph>
<paragraph id="8297A5704019EB70961030334A93A2B8">
Workers (
<emphasis id="3457A4CADEB8CB7F0104E810A93A5FA7" italics="true">n</emphasis>
= 2):
<abbrev id="ABBRID0EMJAE" xlink_title="Head length">HL</abbrev>
0.680.69,
<abbrev id="ABBRID0EQJAE" xlink_title="Head width">HW</abbrev>
0.680.70,
<abbrev id="ABBRID0EUJAE" xlink_title="Mesosomal length">ML</abbrev>
0.65,
<abbrev id="ABBRID0EYJAE" xlink_title="Length of promesonotal shield">PML</abbrev>
0.450.49,
<abbrev id="ABBRID0E3JAE" xlink_title="Pronotal width">PW</abbrev>
0.65,
<abbrev id="ABBRID0EAKAE" xlink_title="Scape length">SL</abbrev>
0.44,
<abbrev id="ABBRID0EEKAE" xlink_title="Total length">TL</abbrev>
2.792.84,
<abbrev id="ABBRID0EIKAE" xlink_title="Cephalic index">CI</abbrev>
99104,
<abbrev id="ABBRID0EMKAE" xlink_title="Pronotum index">PMI</abbrev>
133142,
<abbrev id="ABBRID0EQKAE" xlink_title="Scape index">SI</abbrev>
6364.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUKAE" type="diagnosis">
<paragraph id="CA7E668B8D77C09D09D24C08270B23A8">
<heading id="C5376D43817A0BDAFEA789DEC3DA5DFA" reason="title">Diagnosis of worker.</heading>
</paragraph>
<paragraph id="0862E21595611B72F45AE54694A47202">
Small species (
<abbrev id="ABBRID0E1KAE" xlink_title="Head width">HW</abbrev>
<quantity id="F0072A6D5E3855A1638CD2BF756833E9" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.0" metricValueMin="6.8" unit="mm" value="0.69" valueMax="0.7" valueMin="0.68">0.680.70 mm</quantity>
,
<abbrev id="ABBRID0E5KAE" xlink_title="Total length">TL</abbrev>
<quantity id="C422617E6DC8233B14501C6A6E5ECA77" metricMagnitude="-3" metricUnit="m" metricValue="2.815" metricValueMax="2.84" metricValueMin="2.79" unit="mm" value="2.815" valueMax="2.84" valueMin="2.79">2.792.84 mm</quantity>
). Promesonotal shield shorter than broad and distinctly margined, broadly transparent at sides, overhanging lateral face of mesosoma; anterior corners of pronotum almost right angles; lateral margins of pronotum parallel, slightly sinuate. Petiole in profile tapered, the crest obliquely and narrowly truncate. Postpetiole nodiform, almost as long as high and roundly convex dorsal outline. Dorsa of head and promesonotal shield densely reticulate-rugulose; lateral portion of pronotum reticulate-rugulose; mesopleuron, metapleuron, and lateral faces of propodeum sparsely reticulate-rugulose, with smooth interspaces. Anterior face of petiole smooth and shiny, dorsum and lateral faces rugulose, posterior face smooth. First gastral tergite smooth, with an occasional faint shagreening around piliferous punctures.
</paragraph>
<paragraph id="AA0C50E658EE23652181E9B35ABAAB9E">
The Thai specimens agreed well with the
<typeStatus id="0DDF6F51DEF604B092CD424C9D190BBD" type="holotype">holotype</typeStatus>
(
<ext-link id="3518E2767E1453A25DD79BB3BAB31B89" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0902029" xlink_type="simple">
<collectionCode id="008C30E1D789DB9844D43996B3187886">CASENT</collectionCode>
0902029
</ext-link>
) in structure, sculpturing, and pilosity. However, body color of the specimens collected from
<collectingCountry id="81301E3F8DE36DE37E722F779C72BA52" name="Thailand">Thailand</collectingCountry>
are reddish brown, while the
<typeStatus id="7D7D95DD79BAA882B3835D25AD1D8CC6" type="holotype">holotype</typeStatus>
is paler, yellow (Fig.
<figureCitation id="1274F64F5DBAFCC2F8A0B06A7EF156A7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Meranoplus malaysianus A, C, E holotype (CASENT 0902029) B, D, F non-type worker (THNHM - I- 00028943) A, B head in full-face view C, D promesonotal shield in dorsal view E, F body in profile." figureDoi="10.3897/zookeys.1210.125990.figure1" httpUri="https://binary.pensoft.net/fig/1121260">1</figureCitation>
). Reticulations on dorsum of head rather denser and smaller than that on the
<typeStatus id="CF2C8EDCAEA86524FC60D4591F6EA446" type="holotype">holotype</typeStatus>
(see Fig.
<figureCitation id="37D920DD3323E687B874AE28C52196A5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Meranoplus malaysianus A, C, E holotype (CASENT 0902029) B, D, F non-type worker (THNHM - I- 00028943) A, B head in full-face view C, D promesonotal shield in dorsal view E, F body in profile." figureDoi="10.3897/zookeys.1210.125990.figure1" httpUri="https://binary.pensoft.net/fig/1121260">1 A, B</figureCitation>
for comparison).
</paragraph>
<caption id="353F7970220696A3D1C135C4E5FBC19F" ID-DOI="10.3897/zookeys.1210.125990.figure1" ID-arpha="745F23DE-FA43-5B5D-A86E-294C4BFFE41F" httpUri="https://binary.pensoft.net/fig/1121260" startId="F1">
<paragraph id="E7C31E846ED66B203FD6CAFC5777D90D">
<label id="309C47DBA9B60BF7530C245CB7135851">Figure 1.</label>
</paragraph>
<paragraph id="8A4A534352CE8C16309A088A3F726C22">
<taxonomicName id="0E4B2EA2EC12B4D8B944B0E81BA24EBC" authorityName="Schodl" authorityYear="1998" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="C693796315B22EB56F2234D930C44477" italics="true">Meranoplus malaysianus</emphasis>
</taxonomicName>
<emphasis id="DA5F2BA4F5B216821C0EFD29390B1B9C" bold="true">A, C, E</emphasis>
holotype (
<ext-link id="80CABE0CB3C173E8493F487E03DAA2E9" ext-link-type="uri" xlink_href="http://www.antweb.org/specimen/CASENT0902029" xlink_type="simple">CASENT 0902029</ext-link>
)
<emphasis id="938D3C746AF1A43224EADDA516C1436A" bold="true">B, D, F</emphasis>
non-type worker (
<abbrev content-type="institution" id="ABBRID0ESJBI" xlink_title="Natural History Museum of the National Science Museum, Thailand">THNHM</abbrev>
- I- 00028943)
<emphasis id="B8CD7B1D1884542BBB49AA60BB0A4736" bold="true">A, B</emphasis>
head in full-face view
<emphasis id="0A8F7A3816A7CC974D35DC99A5E3CFEE" bold="true">C, D</emphasis>
promesonotal shield in dorsal view
<emphasis id="4DDAAEA73DE10386996BE8BAE4C03671" bold="true">E, F</emphasis>
body in profile.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERLAE" type="distribution">
<paragraph id="A41030CE846C4AB23D2794C06CBAF0D3">
<heading id="1B251637A8A8818D595E7B20B9681DF0" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2710871C76529E6194B381FF44EFB585">
<collectingCountry id="CF89C9100A4EE7240953FB07C0A7694E" name="Thailand">Thailand</collectingCountry>
(
<collectingRegion id="7F5226755BA3A4BE665C1734819EC897" country="Thailand" name="Songkhla">Songkhla</collectingRegion>
and
<collectingRegion id="7D7AAD08063AD9EEE0D3BCCDEF27DA3C" country="Thailand" name="Narathiwat">Narathiwat</collectingRegion>
, new record, Fig.
<figureCitation id="E020DEC96FFB5339BD8415D0F5CEE974" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Meranoplus Thai species in Thailand." figureDoi="10.3897/zookeys.1210.125990.figure4" httpUri="https://binary.pensoft.net/fig/1121263">4</figureCitation>
),
<collectingCountry id="B05490E0B9FEA193E8B530598C90D877" name="Malaysia">Malaysia</collectingCountry>
(West
<collectingCountry id="437F1C711643E1075B83FF0905D3A6C0" name="Malaysia">Malaysia</collectingCountry>
,
<collectingRegion id="DCC110587D1ECEE737B3651B7A5CE481" country="Malaysia" name="Sabah">Sabah</collectingRegion>
,
<collectingCountry id="18F210825A94AAC7F71F738CA7906C70" name="Malaysia">Sarawak</collectingCountry>
),
<collectingCountry id="BE792E231BBA8BE43727FFFB5803A726" name="Singapore">Singapore</collectingCountry>
,
<collectingCountry id="E9DE74B0DE849DC5B95CB91A7A838E93" name="Indonesia">Indonesia</collectingCountry>
(
<collectingRegion id="53D6F6076B3E3A9DED88B20BB87F6502" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2LAE" type="habitat">
<paragraph id="6CEE29847A60079AA560D199D7BE4A4A">
<heading id="BCC7D12DDDA7BF6BB418DDEB649C7123" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="E3AEF6E9832F19EBA4652CE8D637088C">
<specimenCount id="46D4E406AAAF72CEF4E5CDAD3E0944DB" count="2" type="generic">Two specimens</specimenCount>
from
<collectingRegion id="636A3A367C64E4DEAD29BC508CC2655A" country="Thailand" name="Narathiwat">Narathiwat Province</collectingRegion>
were collected from the ground in a disturbed area near a lowland evergreen forest, near the Thai Malay border. A specimen (
<abbrev content-type="institution" id="ABBRID0EBMAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="58E217761B060864673B7AC78D65CEB1">THNHM</collectionCode>
</abbrev>
- I- 00028943,
<abbrev content-type="institution" id="ABBRID0EGMAE" xlink_title="Natural History Museum of the National Science Museum, Thailand">
<collectionCode id="6B5C030B705E7658290276B3CA1032A1">THNHM</collectionCode>
</abbrev>
) from
<collectingRegion id="3F11E86DA639E0104F77D1221807B86F" country="Thailand" name="Songkhla">Songkhla Province</collectingRegion>
was collected in a primary evergreen forest.
</paragraph>
</subSubSection>
</treatment>
</document>