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<document id="B60ED215EC7C5D2242DF818A1326FE2F" ID-DOI="10.11646/zootaxa.4969.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14895349" ID-ZooBank="CB871456-F1D3-40CB-A154-2525BD25F0F4" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1740000461591" checkinUser="plazi" docAuthor="Freyhof, Jörg, Kaya, Cüneyt, Abdullah, Younis Sabir &amp; Geiger, Matthias F." docDate="2021" docId="BB6C87E2FF93B237FF6BEC1EFED8FF4F" docLanguage="en" docName="zootaxa.4969.3.2.pdf" docOrigin="Zootaxa 4969 (3)" docSource="https://doi.org/10.11646/zootaxa.4969.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyptothorax daemon Freyhof, Kaya, Abdullah &amp; Geiger, 2021, new species" docType="treatment" docVersion="4" lastPageNumber="487" masterDocId="4755FF9AFF8EB214FFFCEE43FF8EFF9A" masterDocTitle="The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae)" masterLastPageNumber="491" masterPageNumber="453" pageNumber="482" updateTime="1740559170557" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="2067FFB3610994CFFA7A8B8DAA8FE036">The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae)</mods:title>
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<mods:namePart id="0E07A718D98009AE91F2C2941FA7401B">Kaya, Cüneyt</mods:namePart>
<mods:affiliation id="B87E7ABA09BA948847A63EE7D72C4FF4">Recep Tayyip Erdogan University, Faculty of Fisheries and Aquatic Sciences, 53100 Rize, Turkey.</mods:affiliation>
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<mods:namePart id="D88BE1A2834219C90D07D641A79B0905">Abdullah, Younis Sabir</mods:namePart>
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<taxonomicName id="F4C54D77FF93B209FF6BEC1EFE10FDE2" authorityName="Freyhof &amp; Kaya &amp; Abdullah &amp; Geiger" authorityYear="2021" box="[151,414,605,632]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="daemon" status="sp. nov.">
<emphasis id="01B1EAE6FF93B209FF6BEC1EFE10FDE2" bold="true" box="[151,414,605,632]" italics="true" pageId="29" pageNumber="482">Glyptothorax daemon</emphasis>
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(
<figureCitation id="ABFE2A71FF93B209FF63ECC2FE95FD01" box="[159,283,641,667]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStart-2="FIGURE 33" captionStart-3="FIGURE 34" captionStart-4="FIGURE 35" captionStart-5="FIGURE 36" captionStartId-0="29.[151,250,2014,2038]" captionStartId-1="30.[151,250,1141,1165]" captionStartId-2="30.[151,250,1904,1928]" captionStartId-3="31.[151,250,1080,1104]" captionStartId-4="32.[151,250,1828,1852]" captionStartId-5="34.[151,250,930,954]" captionTargetBox-0="[227,1360,793,1991]" captionTargetBox-1="[151,1436,417,1118]" captionTargetBox-2="[151,1436,1225,1879]" captionTargetBox-3="[151,1436,741,1057]" captionTargetBox-4="[151,1436,309,1805]" captionTargetBox-5="[151,1436,417,907]" captionTargetId-0="figure-229@29.[227,1360,790,1991]" captionTargetId-1="figure-132@30.[151,1436,417,1118]" captionTargetId-2="figure-158@30.[151,1436,1225,1879]" captionTargetId-3="figure-235@31.[151,1436,741,1057]" captionTargetId-4="figure-87@32.[151,1436,309,1805]" captionTargetId-5="figure-149@34.[151,1436,417,907]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="30" captionTargetPageId-3="31" captionTargetPageId-4="32" captionTargetPageId-5="34" captionText-0="FIGURE 31. Glyptothorax daemon, FFR 3928, holotype, 125 mm SL; Turkey: stream Dilektaşı." captionText-1="FIGURE 32. Glyptothorax daemon, FSJF 3962, paratypes; from top: 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River." captionText-2="FIGURE 33. Glyptothorax daemon, paratypes, from left: FSJF 3962, 77 mm SL, 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River." captionText-3="FIGURE 34. Glyptothorax daemon, FSJF 3962, paratypes; skin ornamentation; from left: 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River." captionText-4="FIGURE 35. Glyptothorax daemon, paratypes, from top: FFR 3926, 116 mm SL; Turkey: stream Dilektaşı; FSJF 3962, pale form, 78 mm SL; dark form, 110 mm SL; Iraq: Choman River." captionText-5="FIGURE 36. Glyptothorax daemon, FSJF 3677, 91 mm SL; Turkey: stream Çıratan." figureDoi-0="http://doi.org/10.5281/zenodo.14895413" figureDoi-1="http://doi.org/10.5281/zenodo.14895415" figureDoi-2="http://doi.org/10.5281/zenodo.14895417" figureDoi-3="http://doi.org/10.5281/zenodo.14895419" figureDoi-4="http://doi.org/10.5281/zenodo.14895421" figureDoi-5="http://doi.org/10.5281/zenodo.14895423" httpUri-0="https://zenodo.org/record/14895413/files/figure.png" httpUri-1="https://zenodo.org/record/14895415/files/figure.png" httpUri-2="https://zenodo.org/record/14895417/files/figure.png" httpUri-3="https://zenodo.org/record/14895419/files/figure.png" httpUri-4="https://zenodo.org/record/14895421/files/figure.png" httpUri-5="https://zenodo.org/record/14895423/files/figure.png" pageId="29" pageNumber="482">Fig. 3136</figureCitation>
)
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<paragraph id="337A36F4FF93B209FF6BEC80FF74FC9B" blockId="29.[151,1436,706,769]" pageId="29" pageNumber="482">
<materialsCitation id="83AD3CA9FF93B209FF6BEC80FF74FC9B" country="Turkey" latitude="37.6664" location="stream Dilektasi" longLatPrecision="6" longitude="44.1393" municipality="Dilektasi" pageId="29" pageNumber="482" specimenCount="1" stateProvince="Hakkari prov." typeStatus="holotype">
<emphasis id="01B1EAE6FF93B209FF6BEC80FE85FD47" bold="true" box="[151,267,707,733]" pageId="29" pageNumber="482">
<typeStatus id="EC7E8856FF93B209FF6BEC80FE88FD47" box="[151,262,707,733]" pageId="29" pageNumber="482" type="holotype">Holotype</typeStatus>
.
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FFR
<quantity id="F43D9B11FF93B209FEB1EC80FE7DFD47" box="[333,499,706,733]" metricMagnitude="3" metricUnit="m" metricValue="3.928125" pageId="29" pageNumber="482" unit="mm" value="3928125.0">3928, 125 mm</quantity>
SL;
<collectingCountry id="4BD27664FF93B209FDD7EC80FDF1FD47" box="[555,639,707,733]" name="Turkey" pageId="29" pageNumber="482">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF93B209FD77EC80FCA4FD47" box="[651,810,707,733]" country="Turkey" name="Hakkari" pageId="29" pageNumber="482">Hakkari prov.</collectingRegion>
:
<location id="361A602FFF93B209FCCAEC80FC60FD47" LSID="urn:lsid:plazi:treatment:BB6C87E2FF93B237FF6BEC1EFED8FF4F:361A602FFF93B209FCCAEC80FC60FD47" box="[822,1006,707,733]" country="Turkey" latitude="37.6664" longLatPrecision="6" longitude="44.1393" municipality="Dilektasi" name="stream Dilektasi" pageId="29" pageNumber="482" stateProvince="Hakkari prov.">stream Dilektaşı</location>
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<quantity id="F43D9B11FF93B209FC0AEC80FBB0FD47" box="[1014,1086,707,733]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="482" unit="km" value="16.0">16 km</quantity>
northeast Yüksekova
</locationDeviation>
,
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.
</materialsCitation>
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<paragraph id="337A36F4FF93B209FF6BE99DFBE1F86D" blockId="29.[151,1135,2014,2039]" box="[151,1135,2014,2039]" pageId="29" pageNumber="482">
<emphasis id="01B1EAE6FF93B209FF6BE99DFEAEF86C" bold="true" box="[151,288,2014,2038]" pageId="29" pageNumber="482">FIGURE 31.</emphasis>
<taxonomicName id="F4C54D77FF93B209FEDAE99CFDB3F86C" authority=", FFR" authorityName="FFR" box="[294,573,2015,2039]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF93B209FEDAE99CFD8DF86D" box="[294,515,2015,2039]" italics="true" pageId="29" pageNumber="482">Glyptothorax daemon</emphasis>
, FFR
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3928, holotype, 125 mm SL; Turkey: stream Dilektaşı.
</paragraph>
</caption>
<paragraph id="337A36F4FF90B20AFF3BEED4FC76FF4F" blockId="30.[151,1436,150,393]" pageId="30" pageNumber="483">
<materialsCitation id="83AD3CA9FF90B20AFF3BEED4FB71FF2B" box="[199,1279,150,177]" country="Iraq" latitude="36.6106" location="Choman" longLatPrecision="6" longitude="44.8381" municipality="Qubay Galala" pageId="30" pageNumber="483" specimenCount="1" stateProvince="Hakkari" typeStatus="paratype">
<emphasis id="01B1EAE6FF90B20AFF3BEED4FEC8FF2B" bold="true" box="[199,326,151,177]" pageId="30" pageNumber="483">
<typeStatus id="EC7E8856FF90B20AFF3BEED4FECFFF2B" box="[199,321,151,177]" pageId="30" pageNumber="483" type="paratype">Paratypes</typeStatus>
.
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FFR 3926, 9,
<quantity id="F43D9B11FF90B20AFE1BEED4FDE2FF2B" box="[487,620,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" metricValueMax="11.6" metricValueMin="7.3" pageId="30" pageNumber="483" unit="mm" value="94.5" valueMax="116.0" valueMin="73.0">73116 mm</quantity>
SL; FFR 3905, 10,
<quantity id="F43D9B11FF90B20AFCB5EED4FC41FF2B" box="[841,975,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="30" pageNumber="483" unit="mm" value="90.0" valueMax="110.0" valueMin="70.0">70110 mm</quantity>
SL; same data as holotype.
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<materialsCitation id="83AD3CA9FF90B20AFB03EED4FC7AFF4F" country="Iraq" latitude="36.6106" location="stream Choman" longLatPrecision="6" longitude="44.8381" municipality="Qubay Galala" pageId="30" pageNumber="483" specimenCount="1" stateProvince="Hakkari" typeStatus="paratype">
—FSJF 3962, 10,
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SL;
<collectingCountry id="4BD27664FF90B20AFE80EEF8FE23FF4F" box="[380,429,187,213]" name="Iraq" pageId="30" pageNumber="483">Iraq</collectingCountry>
:
<location id="361A602FFF90B20AFE44EEF8FDE7FF4F" LSID="urn:lsid:plazi:treatment:BB6C87E2FF93B237FF6BEC1EFED8FF4F:361A602FFF90B20AFE44EEF8FDE7FF4F" box="[440,617,187,213]" country="Iraq" latitude="36.6106" longLatPrecision="6" longitude="44.8381" municipality="Qubay Galala" name="stream Choman" pageId="30" pageNumber="483" stateProvince="Hakkari">stream Choman</location>
at
<collectingMunicipality id="D31EAC8EFF90B20AFD77EEF8FCA6FF4F" box="[651,808,187,213]" pageId="30" pageNumber="483">Qubay Galala</collectingMunicipality>
,
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.
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<paragraph id="337A36F4FF90B20AFF3BEE9DFBB7FE13" blockId="30.[151,1436,150,393]" pageId="30" pageNumber="483">
<emphasis id="01B1EAE6FF90B20AFF3BEE9DFDC7FF62" bold="true" box="[199,585,222,248]" pageId="30" pageNumber="483">Additional materials examined.</emphasis>
<materialsCitation id="83AD3CA9FF90B20AFDA9EE9CFE78FE87" country="Turkey" latitude="38.4028" location="stream Gumuskanat" longLatPrecision="6" longitude="41.7394" municipality="Gumuskanat" pageId="30" pageNumber="483" specimenCount="1" stateProvince="Bitlis prov.">
FFR 3902,
<quantity id="F43D9B11FF90B20AFD21EE9CFCEEFF63" box="[733,864,223,249]" metricMagnitude="0" metricUnit="m" metricValue="1.103" pageId="30" pageNumber="483" unit="mm" value="1103.0">1, 103 mm</quantity>
SL;
<collectingCountry id="4BD27664FF90B20AFC5CEE9CFC7AFF63" box="[928,1012,223,249]" name="Turkey" pageId="30" pageNumber="483">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF90B20AFBFFEE9CFB07FF63" box="[1027,1161,223,249]" country="Turkey" name="Bitlis" pageId="30" pageNumber="483">Bitlis prov.</collectingRegion>
:
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at
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,
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FFR 3904, 10,
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SL; FFR 3918, 9,
<quantity id="F43D9B11FF90B20AFBE3EF41FB29FE87" box="[1055,1191,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="11.5" metricValueMin="6.5" pageId="30" pageNumber="483" unit="mm" value="90.0" valueMax="115.0" valueMin="65.0">65115 mm</quantity>
SL;
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:
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northeast
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,
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FFR 3907, 5,
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SL; FFR 3920, 3,
<quantity id="F43D9B11FF90B20AFAB4EF64FF4DFEFF" metricMagnitude="-2" metricUnit="m" metricValue="8.95" metricValueMax="10.0" metricValueMin="7.9" pageId="30" pageNumber="483" unit="mm" value="89.5" valueMax="100.0" valueMin="79.0">79100 mm</quantity>
SL;
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:
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<quantity id="F43D9B11FF90B20AFD6EEF08FD42FEFF" box="[658,716,331,357]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="483" unit="km" value="3.0">
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southwest of
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,
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FSJF 3677, 2,
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SL;
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about
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east of
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,
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.
</paragraph>
<caption id="67BA667CFF90B20AFF6BEA36FF5BFB2B" ID-DOI="http://doi.org/10.5281/zenodo.14895415" ID-Zenodo-Dep="14895415" httpUri="https://zenodo.org/record/14895415/files/figure.png" pageId="30" pageNumber="483" startId="30.[151,250,1141,1165]" targetBox="[151,1436,417,1118]" targetPageId="30" targetType="figure">
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<emphasis id="01B1EAE6FF90B20AFF6BEA36FEADFB17" bold="true" box="[151,291,1141,1165]" pageId="30" pageNumber="483">FIGURE 32.</emphasis>
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<emphasis id="01B1EAE6FF90B20AFED7EA36FD85FB17" box="[299,523,1141,1165]" italics="true" pageId="30" pageNumber="483">Glyptothorax daemon</emphasis>
, FSJF
</taxonomicName>
3962, paratypes; from top: 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River.
</paragraph>
</caption>
<caption id="67BA667CFF90B20AFF6BE933FEBCF836" ID-DOI="http://doi.org/10.5281/zenodo.14895417" ID-Zenodo-Dep="14895417" httpUri="https://zenodo.org/record/14895417/files/figure.png" pageId="30" pageNumber="483" startId="30.[151,250,1904,1928]" targetBox="[151,1436,1225,1879]" targetPageId="30" targetType="figure">
<paragraph id="337A36F4FF90B20AFF6BE933FEBCF836" blockId="30.[151,1437,1904,1964]" pageId="30" pageNumber="483">
<emphasis id="01B1EAE6FF90B20AFF6BE933FEAFF812" bold="true" box="[151,289,1904,1928]" pageId="30" pageNumber="483">FIGURE 33.</emphasis>
<taxonomicName id="F4C54D77FF90B20AFED4E933FD88F812" box="[296,518,1904,1928]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="30" pageNumber="483" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF90B20AFED4E933FD88F812" box="[296,518,1904,1928]" italics="true" pageId="30" pageNumber="483">Glyptothorax daemon</emphasis>
</taxonomicName>
, paratypes, from left: FSJF 3962, 77 mm SL, 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River.
</paragraph>
</caption>
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<emphasis id="01B1EAE6FF91B20BFF3BEED5FD5AFF2A" bold="true" box="[199,724,150,177]" pageId="31" pageNumber="484">Material used in molecular genetic analysis.</emphasis>
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FSJF-DNA 2636-2637;
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FSJF-DNA 2673;
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FSJF-DNA 3337;
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FSJF-DNA 3338; G559F
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FSJF-DNA 3339;
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<collectingRegion id="F101F816FF91B20BFB43EF2CFACAFE13" box="[1215,1348,367,393]" country="Turkey" name="Bitlis" pageId="31" pageNumber="484">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="D31EAC8EFF91B20BFF6BEFD0FF67FE37" box="[151,233,403,429]" pageId="31" pageNumber="484">Çıratan</collectingMunicipality>
<quantity id="F43D9B11FF91B20BFF08EFD0FEBFFE37" box="[244,305,403,429]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="484" unit="km" value="3.0">
<locationDeviation id="22C82FD6FF91B20BFF08EFD0FEBFFE37" box="[244,305,403,429]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="484" unit="km" value="3.0">3 km</locationDeviation>
</quantity>
southwest of
<location id="361A602FFF91B20BFE24EFD7FDBAFE35" LSID="urn:lsid:plazi:treatment:BB6C87E2FF93B237FF6BEC1EFED8FF4F:361A602FFF91B20BFE24EFD7FDBAFE35" box="[472,564,404,431]" country="Turkey" latitude="38.3547" longLatPrecision="6" longitude="41.7814" municipality="Ciratan" name="Ucadim" pageId="31" pageNumber="484" stateProvince="Bitlis prov.">Üçadım</location>
,
<geoCoordinate id="56F15033FF91B20BFDBFEFD1FD10FE37" box="[579,670,402,429]" degrees="38.3547" direction="north" orientation="latitude" pageId="31" pageNumber="484" precision="5" value="38.3547">38.3547</geoCoordinate>
<geoCoordinate id="56F15033FF91B20BFD54EFD0FC8DFE37" box="[680,771,403,429]" degrees="41.7814" direction="east" orientation="longitude" pageId="31" pageNumber="484" precision="5" value="41.7814">41.7814</geoCoordinate>
(
<collectorName id="9E305322FF91B20BFCEAEFD0FC0CFE37" box="[790,898,403,429]" pageId="31" pageNumber="484">GenBank</collectorName>
accession numbers:
<accessionNumber id="2C96AB17FF91B20BFB89EFD1FEAAFE4B" isEnumeration="true" pageId="31" pageNumber="484">MW724518, MW724519, MW724520</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF91B20BFF3BEF99FBA6F9BF" pageId="31" pageNumber="484" type="diagnosis">
<paragraph id="337A36F4FF91B20BFF3BEF99FCF4FD57" blockId="31.[151,1437,150,717]" pageId="31" pageNumber="484">
<emphasis id="01B1EAE6FF91B20BFF3BEF99FECCFE6E" bold="true" box="[199,322,474,500]" pageId="31" pageNumber="484">Diagnosis.</emphasis>
<taxonomicName id="F4C54D77FF91B20BFEB5EF98FDB1FE6F" box="[329,575,475,501]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF91B20BFEB5EF98FDB1FE6F" box="[329,575,475,501]" italics="true" pageId="31" pageNumber="484">Glyptothorax daemon</emphasis>
</taxonomicName>
is distinguished from its congeners in the Euphrates and Tigris drainages by a combination of characters, none unique to the species.
<taxonomicName id="F4C54D77FF91B20BFCF0EFBCFB8EFD83" box="[780,1024,511,537]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF91B20BFCF0EFBCFB8EFD83" box="[780,1024,511,537]" italics="true" pageId="31" pageNumber="484">Glyptothorax daemon</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="F4C54D77FF91B20BFB04EFBCFA13FD82" box="[1272,1437,511,537]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF91B20BFB04EFBCFA13FD82" box="[1272,1437,511,537]" italics="true" pageId="31" pageNumber="484">G. armeniacus</emphasis>
</taxonomicName>
by having very indistinct or no yellow tips of the fins (vs. with distinct yellow tips), many shallow warts on the head giving the skin a wary appearance (vs. many minute tubercles and few warts). It is distinguished from
<taxonomicName id="F4C54D77FF91B20BFA7DEC04FF6EFD1F" authorityName="Coad" authorityYear="1981" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF91B20BFA7DEC04FF6EFD1F" italics="true" pageId="31" pageNumber="484">G. silviae</emphasis>
</taxonomicName>
by having many warts on the back and flank (vs. absent), well developed and numerous anteromedial striae in the thoracic adhesive apparatus (vs. absent or very short), and many black, dark-grey or dark-brown spots and/or blotches on the flank (flank with few scattered dark-brown spots).
</paragraph>
<caption id="67BA667CFF91B20BFF6BEA7BFE1DFBEE" ID-DOI="http://doi.org/10.5281/zenodo.14895419" ID-Zenodo-Dep="14895419" httpUri="https://zenodo.org/record/14895419/files/figure.png" pageId="31" pageNumber="484" startId="31.[151,250,1080,1104]" targetBox="[151,1436,741,1057]" targetPageId="31" targetType="figure">
<paragraph id="337A36F4FF91B20BFF6BEA7BFE1DFBEE" blockId="31.[151,1436,1080,1140]" pageId="31" pageNumber="484">
<emphasis id="01B1EAE6FF91B20BFF6BEA7BFEACFBCA" bold="true" box="[151,290,1080,1104]" pageId="31" pageNumber="484">FIGURE 34.</emphasis>
<taxonomicName id="F4C54D77FF91B20BFED5EA7BFDC5FBCA" authority=", FSJF" authorityName="FSJF" box="[297,587,1080,1104]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF91B20BFED5EA7BFD86FBCA" box="[297,520,1080,1104]" italics="true" pageId="31" pageNumber="484">Glyptothorax daemon</emphasis>
, FSJF
</taxonomicName>
3962, paratypes; skin ornamentation; from left: 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River.
</paragraph>
</caption>
<paragraph id="337A36F4FF91B20BFF3BEAE1FBA6F9BF" blockId="31.[151,1437,1186,2041]" pageId="31" pageNumber="484">
The new species is distinguished from
<taxonomicName id="F4C54D77FF91B20BFD85EAE0FD40FB26" box="[633,718,1187,1213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFD85EAE0FD40FB26" box="[633,718,1187,1213]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF91B20BFD20EAE0FC1AFB26" box="[732,916,1186,1213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF91B20BFD20EAE0FC1AFB26" box="[732,916,1186,1213]" italics="true" pageId="31" pageNumber="484">G. kurdistanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF91B20BFC35EAE0FB0DFB26" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[969,1155,1186,1213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF91B20BFC35EAE0FB0DFB26" box="[969,1155,1186,1213]" italics="true" pageId="31" pageNumber="484">G. steindachneri</emphasis>
</taxonomicName>
by having the dorsal and lateral head without tubercles (vs. many large, elongated, bony and striated tubercles in
<taxonomicName id="F4C54D77FF91B20BFBB3EA84FB2FFB7A" box="[1103,1185,1223,1249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFBB3EA84FB2FFB7A" box="[1103,1185,1223,1249]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF91B20BFB2FEA84FA05FB7A" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1235,1419,1222,1249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF91B20BFB2FEA84FA05FB7A" box="[1235,1419,1222,1249]" italics="true" pageId="31" pageNumber="484">G. steindachneri</emphasis>
</taxonomicName>
); a strongly elevated thoracic adhesive apparatus (vs. not or very slightly elevated in
<taxonomicName id="F4C54D77FF91B20BFBBAEAA8FB12FA9E" box="[1094,1180,1259,1285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFBBAEAA8FB12FA9E" box="[1094,1180,1259,1285]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
), 1.11.3 times longer than wide (vs.
<quantity id="F43D9B11FF91B20BFEC3EB4CFE3CFAB2" box="[319,434,1294,1321]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="31" pageNumber="484" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">
0.8
<emphasis id="01B1EAE6FF91B20BFE9EEB4CFEFEFAB2" box="[354,368,1295,1320]" italics="true" pageId="31" pageNumber="484"></emphasis>
1.1 in
</quantity>
<taxonomicName id="F4C54D77FF91B20BFE45EB4CFD9EFAB2" box="[441,528,1295,1321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFE45EB4CFD9EFAB2" box="[441,528,1295,1321]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
,
<quantity id="F43D9B11FF91B20BFDE3EB4CFD1FFAB2" box="[543,657,1294,1321]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.7779999999999998" pageId="31" pageNumber="484" unit="in" value="0.8" valueMax="0.9" valueMin="0.7">
0.7
<emphasis id="01B1EAE6FF91B20BFDBEEB4CFDDEFAB2" box="[578,592,1295,1320]" italics="true" pageId="31" pageNumber="484"></emphasis>
0.9 in
</quantity>
<taxonomicName id="F4C54D77FF91B20BFD65EB4CFCDDFAB2" box="[665,851,1294,1321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF91B20BFD65EB4CFCDDFAB2" box="[665,851,1294,1321]" italics="true" pageId="31" pageNumber="484">G. kurdistanicus</emphasis>
</taxonomicName>
), extending from the isthmus to the base of the last pectoral-fin ray or to the posterior limit of the pectoral-fin base (vs. to base of first or third branched pectoral-fin ray in
<taxonomicName id="F4C54D77FF91B20BFF49EB14FEE1FAEA" box="[181,367,1366,1393]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF91B20BFF49EB14FEE1FAEA" box="[181,367,1366,1393]" italics="true" pageId="31" pageNumber="484">G. kurdistanicus</emphasis>
</taxonomicName>
). The thoracic adhesive apparatus is well delineated at its posterior margin (poorly delineated in
<taxonomicName id="F4C54D77FF91B20BFF48EB38FE87FA0E" box="[180,265,1403,1429]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFF48EB38FE87FA0E" box="[180,265,1403,1429]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
), completely situated on the horseshoe shaped swelling (vs. extending beyond the swelling, usually onto the pectoral-fin base in
<taxonomicName id="F4C54D77FF91B20BFE5CEBDCFE7BFA22" box="[416,501,1439,1465]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF91B20BFE5CEBDCFE7BFA22" box="[416,501,1439,1465]" italics="true" pageId="31" pageNumber="484">G. cous</emphasis>
</taxonomicName>
). It is further distinguished from
<taxonomicName id="F4C54D77FF91B20BFC97EBDCFBA8FA22" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[875,1062,1438,1465]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF91B20BFC97EBDCFBA8FA22" box="[875,1062,1438,1465]" italics="true" pageId="31" pageNumber="484">G. steindachneri</emphasis>
</taxonomicName>
by having a short adipose-fin, its length 0.71.1 times (vs. 1.53.0) larger than the distance between the base of last dorsal-fin ray and the adipose-fin origin, the medial pit without striae (with striae), a blunt and roundish head, its length 2426% SL (vs. pointed, 2123), and 711 serrae on the inner margin of the pectoral-fin spine (vs. 1317).
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF91B236FF3BE86DFB6CFADC" lastPageId="34" lastPageNumber="487" pageId="31" pageNumber="484" type="description">
<paragraph id="337A36F4FF91B20BFF3BE86DFD8AF817" blockId="31.[151,1437,1186,2041]" pageId="31" pageNumber="484">
<emphasis id="01B1EAE6FF91B20BFF3BE86DFED7F9D2" bold="true" box="[199,345,1582,1608]" pageId="31" pageNumber="484">Description.</emphasis>
Morphometric data in
<tableCitation id="7E47034FFF91B20BFDA5E86DFD21F9D3" box="[601,687,1582,1609]" captionStart="TABLE 4" captionStartId="33.[151,239,151,175]" captionTargetPageId="33" captionText="TABLE 4. Morphometry of Glyptothorax daemon (holotype FFR 3928 and paratypes FFR 3905, FFR 3926, n=3; FSJF 3962, n=20). The calculations include the holotype." pageId="31" pageNumber="484">Table 4</tableCitation>
. Head depressed; body subcylindrical. Dorsal head profile straight, predorsal profile slightly convex: Profile rising from tip of snout to dorsal-fin origin, then almost straight, sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to end of caudal peduncle. Caudal-peduncle depth 1.62.1 times in its length. Anus and urogenital openings located below tip of adpressed pelvic fin. Skin of back and flank with small, roundish warts, sparsely set on head, densely set on flank and belly, warts larger and more distinct on belly. Lateral line complete and midlateral. Head broad, spade-shaped when viewed laterally. Snout blunt. Anterior and posterior nares large and separated only by base of nasal barbel. Bony elements of dorsal surface of head covered with thick skin, smooth, without tubercles, with many warts, slightly elongated on gill cover (
<figureCitation id="ABFE2A71FF91B20BFE5AE90DFE74F8F3" box="[422,506,1870,1897]" captionStart="FIGURE 34" captionStartId="31.[151,250,1080,1104]" captionTargetBox="[151,1436,741,1057]" captionTargetId="figure-235@31.[151,1436,741,1057]" captionTargetPageId="31" captionText="FIGURE 34. Glyptothorax daemon, FSJF 3962, paratypes; skin ornamentation; from left: 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River." figureDoi="http://doi.org/10.5281/zenodo.14895419" httpUri="https://zenodo.org/record/14895419/files/figure.png" pageId="31" pageNumber="484">Fig. 34</figureCitation>
). Eye ovoid, horizontal axis longest; located just below dorsal-head profile. Largest individual recorded
<quantity id="F43D9B11FF91B20BFE85E931FE58F817" box="[377,470,1906,1933]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="31" pageNumber="484" unit="mm" value="125.0">125 mm</quantity>
SL.
</paragraph>
<paragraph id="337A36F4FF91B234FF3BE9D5FB17FE87" blockId="31.[151,1437,1186,2041]" lastBlockId="32.[151,1437,151,285]" lastPageId="32" lastPageNumber="485" pageId="31" pageNumber="484">Barbels in four pairs. Maxillary barbel broad and thick, extending to, slightly in front of or beyond pectoral-fin base, velum at proximal part of barbel attached to head closer to posterior nare than to eye, many thick warts on outer base of velum, velum smooth. Nasal barbel broad, extending almost to anterior orbital margin. Inner mandibularbarbel extending to isthmus. Outer mandibular barbel extending to end of gill cover, not reaching pectoral-fin origin. Mouth inferior, premaxillary tooth band partially exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth appearing in single broad semilunate band. Dentary teeth in a single crescentic band, consisting of two separate halves tightly bound at midline.</paragraph>
<caption id="67BA667CFFAEB234FF6BE967FC9DF8FA" ID-DOI="http://doi.org/10.5281/zenodo.14895421" ID-Zenodo-Dep="14895421" httpUri="https://zenodo.org/record/14895421/files/figure.png" pageId="32" pageNumber="485" startId="32.[151,250,1828,1852]" targetBox="[151,1436,309,1805]" targetPageId="32" targetType="figure">
<paragraph id="337A36F4FFAEB234FF6BE967FC9DF8FA" blockId="32.[151,1436,1828,1889]" pageId="32" pageNumber="485">
<emphasis id="01B1EAE6FFAEB234FF6BE967FEAFF8A6" bold="true" box="[151,289,1828,1852]" pageId="32" pageNumber="485">FIGURE 35.</emphasis>
<taxonomicName id="F4C54D77FFAEB234FED5E967FD89F8A6" box="[297,519,1828,1852]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="32" pageNumber="485" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFAEB234FED5E967FD89F8A6" box="[297,519,1828,1852]" italics="true" pageId="32" pageNumber="485">Glyptothorax daemon</emphasis>
</taxonomicName>
, paratypes, from top: FFR 3926, 116 mm SL; Turkey: stream Dilektaşı; FSJF 3962, pale form, 78 mm SL; dark form, 110 mm SL; Iraq: Choman River.
</paragraph>
</caption>
<caption id="67BA667CFFAFB235FF6BEED4FD30FF49" pageId="33" pageNumber="486" startId="33.[151,239,151,175]" targetBox="[166,1399,227,1649]" targetIsTable="true" targetPageId="33" targetType="table">
<paragraph id="337A36F4FFAFB235FF6BEED4FD30FF49" blockId="33.[151,1436,151,212]" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FF6BEED4FE85FF35" bold="true" box="[151,267,151,175]" pageId="33" pageNumber="486">TABLE 4.</emphasis>
Morphometry of
<taxonomicName id="F4C54D77FFAFB235FE37EED4FD3BFF35" box="[459,693,151,175]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="486" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFAFB235FE37EED4FD3BFF35" box="[459,693,151,175]" italics="true" pageId="33" pageNumber="486">Glyptothorax daemon</emphasis>
</taxonomicName>
(holotype FFR 3928 and paratypes FFR 3905, FFR 3926, n=3; FSJF 3962, n=20). The calculations include the holotype.
</paragraph>
</caption>
<paragraph id="337A36F4FFAFB235FD11EEA0FAF9F9EA" pageId="33" pageNumber="486">
<table id="41C5C454FFAF4DEBFF5AEEA0FAF9F9EB" box="[166,1399,227,1649]" gridcols="6" gridrows="33" pageId="33" pageNumber="486">
<tr id="8DF534B6FFAF4DEBFF5AEEA0FAF9FF61" box="[166,1399,227,251]" gridrow="0" pageId="33" pageNumber="486" rowspan-0="1" rowspan-2="1" rowspan-5="1">
<th id="CE245DCAFFAF4DEBFD11EEA0FCC4FF61" box="[749,842,227,251]" gridcol="1" gridrow="0" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FD11EEA0FCC4FF61" bold="true" box="[749,842,227,251]" pageId="33" pageNumber="486">holotype</emphasis>
</th>
<th id="CE245DCAFFAF4DEBFBE3EEA0FA87FF61" box="[1055,1289,227,251]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FBE3EEA0FA87FF61" bold="true" box="[1055,1289,227,251]" pageId="33" pageNumber="486">holotype &amp; paratypes</emphasis>
</th>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEF48FAF9FEBE" box="[166,1399,267,292]" gridrow="1" pageId="33" pageNumber="486" rowspan-0="1" rowspan-1="1">
<td id="CE245DCAFFAF4DEBFC3EEF48FC77FEBE" box="[962,1017,267,292]" gridcol="2" gridrow="1" pageId="33" pageNumber="486">mean</td>
<td id="CE245DCAFFAF4DEBFBE3EF48FB0DFEBE" box="[1055,1155,267,292]" gridcol="3" gridrow="1" pageId="33" pageNumber="486">min</td>
<td id="CE245DCAFFAF4DEBFB26EF48FA87FEBE" box="[1242,1289,267,292]" gridcol="4" gridrow="1" pageId="33" pageNumber="486">max</td>
<td id="CE245DCAFFAF4DEBFAABEF48FAF9FEBE" box="[1367,1399,267,292]" gridcol="5" gridrow="1" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FAABEF48FAF9FEB9" bold="true" box="[1367,1399,267,291]" pageId="33" pageNumber="486">SD</emphasis>
</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEF77FAF9FED6" box="[166,1399,308,332]" gridrow="2" pageId="33" pageNumber="486" rowspan-2="1" rowspan-5="1">
<th id="CE245DCAFFAF4DEBFF5AEF77FD8CFED6" box="[166,514,308,332]" gridcol="0" gridrow="2" pageId="33" pageNumber="486">SL</th>
<td id="CE245DCAFFAF4DEBFD11EF77FCC4FED6" box="[749,842,308,332]" gridcol="1" gridrow="2" pageId="33" pageNumber="486">125</td>
<td id="CE245DCAFFAF4DEBFBE3EF77FB0DFED6" box="[1055,1155,308,332]" gridcol="3" gridrow="2" pageId="33" pageNumber="486">77</td>
<td id="CE245DCAFFAF4DEBFB26EF77FA87FED6" box="[1242,1289,308,332]" gridcol="4" gridrow="2" pageId="33" pageNumber="486">125</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEF1FFAF9FEEF" box="[166,1399,348,373]" gridrow="3" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEF1FFAF9FEEF" box="[166,1399,348,373]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FF5AEF1EFE6CFEEF" bold="true" box="[166,482,348,373]" pageId="33" pageNumber="486">In percent of standard length</emphasis>
</th>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEFC6FAF9FE07" box="[166,1399,389,413]" gridrow="4" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEFC6FD8CFE07" box="[166,514,389,413]" gridcol="0" gridrow="4" pageId="33" pageNumber="486">Head length</th>
<td id="CE245DCAFFAF4DEBFD11EFC6FCC4FE07" box="[749,842,389,413]" gridcol="1" gridrow="4" pageId="33" pageNumber="486">26.0</td>
<td id="CE245DCAFFAF4DEBFC3EEFC6FC77FE07" box="[962,1017,389,413]" gridcol="2" gridrow="4" pageId="33" pageNumber="486">25.1</td>
<td id="CE245DCAFFAF4DEBFBE3EFC6FB0DFE07" box="[1055,1155,389,413]" gridcol="3" gridrow="4" pageId="33" pageNumber="486">23.6</td>
<td id="CE245DCAFFAF4DEBFB26EFC6FA87FE07" box="[1242,1289,389,413]" gridcol="4" gridrow="4" pageId="33" pageNumber="486">26.3</td>
<td id="CE245DCAFFAF4DEBFAABEFC6FAF9FE07" box="[1367,1399,389,413]" gridcol="5" gridrow="4" pageId="33" pageNumber="486">0.8</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEFEEFAF9FE5F" box="[166,1399,429,453]" gridrow="5" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEFEEFD8CFE5F" box="[166,514,429,453]" gridcol="0" gridrow="5" pageId="33" pageNumber="486">Body depth at dorsalfin origin</th>
<td id="CE245DCAFFAF4DEBFD11EFEEFCC4FE5F" box="[749,842,429,453]" gridcol="1" gridrow="5" pageId="33" pageNumber="486">20.7</td>
<td id="CE245DCAFFAF4DEBFC3EEFEEFC77FE5F" box="[962,1017,429,453]" gridcol="2" gridrow="5" pageId="33" pageNumber="486">19.7</td>
<td id="CE245DCAFFAF4DEBFBE3EFEEFB0DFE5F" box="[1055,1155,429,453]" gridcol="3" gridrow="5" pageId="33" pageNumber="486">18.3</td>
<td id="CE245DCAFFAF4DEBFB26EFEEFA87FE5F" box="[1242,1289,429,453]" gridcol="4" gridrow="5" pageId="33" pageNumber="486">21.2</td>
<td id="CE245DCAFFAF4DEBFAABEFEEFAF9FE5F" box="[1367,1399,429,453]" gridcol="5" gridrow="5" pageId="33" pageNumber="486">0.8</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEF96FAF9FE77" box="[166,1399,469,493]" gridrow="6" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEF96FD8CFE77" box="[166,514,469,493]" gridcol="0" gridrow="6" pageId="33" pageNumber="486">Predorsal length</th>
<td id="CE245DCAFFAF4DEBFD11EF96FCC4FE77" box="[749,842,469,493]" gridcol="1" gridrow="6" pageId="33" pageNumber="486">35.5</td>
<td id="CE245DCAFFAF4DEBFC3EEF96FC77FE77" box="[962,1017,469,493]" gridcol="2" gridrow="6" pageId="33" pageNumber="486">35.5</td>
<td id="CE245DCAFFAF4DEBFBE3EF96FB0DFE77" box="[1055,1155,469,493]" gridcol="3" gridrow="6" pageId="33" pageNumber="486">34.1</td>
<td id="CE245DCAFFAF4DEBFB26EF96FA87FE77" box="[1242,1289,469,493]" gridcol="4" gridrow="6" pageId="33" pageNumber="486">36.9</td>
<td id="CE245DCAFFAF4DEBFAABEF96FAF9FE77" box="[1367,1399,469,493]" gridcol="5" gridrow="6" pageId="33" pageNumber="486">0.7</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEFBEFAF9FD8C" box="[166,1399,509,534]" gridrow="7" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEFBEFD8CFD8C" box="[166,514,509,534]" gridcol="0" gridrow="7" pageId="33" pageNumber="486">Postdorsal length</th>
<td id="CE245DCAFFAF4DEBFD11EFBEFCC4FD8C" box="[749,842,509,534]" gridcol="1" gridrow="7" pageId="33" pageNumber="486">57.3</td>
<td id="CE245DCAFFAF4DEBFC3EEFBEFC77FD8C" box="[962,1017,509,534]" gridcol="2" gridrow="7" pageId="33" pageNumber="486">56.5</td>
<td id="CE245DCAFFAF4DEBFBE3EFBEFB0DFD8C" box="[1055,1155,509,534]" gridcol="3" gridrow="7" pageId="33" pageNumber="486">53.1</td>
<td id="CE245DCAFFAF4DEBFB26EFBEFA87FD8C" box="[1242,1289,509,534]" gridcol="4" gridrow="7" pageId="33" pageNumber="486">59.5</td>
<td id="CE245DCAFFAF4DEBFAABEFBEFAF9FD8C" box="[1367,1399,509,534]" gridcol="5" gridrow="7" pageId="33" pageNumber="486">1.8</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEC65FAF9FDA5" box="[166,1399,550,575]" gridrow="8" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEC65FD8CFDA5" box="[166,514,550,575]" gridcol="0" gridrow="8" pageId="33" pageNumber="486">Preanal length</th>
<td id="CE245DCAFFAF4DEBFD11EC65FCC4FDA5" box="[749,842,550,575]" gridcol="1" gridrow="8" pageId="33" pageNumber="486">70.0</td>
<td id="CE245DCAFFAF4DEBFC3EEC65FC77FDA5" box="[962,1017,550,575]" gridcol="2" gridrow="8" pageId="33" pageNumber="486">69.8</td>
<td id="CE245DCAFFAF4DEBFBE3EC65FB0DFDA5" box="[1055,1155,550,575]" gridcol="3" gridrow="8" pageId="33" pageNumber="486">67.0</td>
<td id="CE245DCAFFAF4DEBFB26EC65FA87FDA5" box="[1242,1289,550,575]" gridcol="4" gridrow="8" pageId="33" pageNumber="486">72.1</td>
<td id="CE245DCAFFAF4DEBFAABEC65FAF9FDA5" box="[1367,1399,550,575]" gridcol="5" gridrow="8" pageId="33" pageNumber="486">1.4</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEC0CFAF9FDFD" box="[166,1399,591,615]" gridrow="9" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEC0CFD8CFDFD" box="[166,514,591,615]" gridcol="0" gridrow="9" pageId="33" pageNumber="486">Prepelvic length</th>
<td id="CE245DCAFFAF4DEBFD11EC0CFCC4FDFD" box="[749,842,591,615]" gridcol="1" gridrow="9" pageId="33" pageNumber="486">51.0</td>
<td id="CE245DCAFFAF4DEBFC3EEC0CFC77FDFD" box="[962,1017,591,615]" gridcol="2" gridrow="9" pageId="33" pageNumber="486">51.3</td>
<td id="CE245DCAFFAF4DEBFBE3EC0CFB0DFDFD" box="[1055,1155,591,615]" gridcol="3" gridrow="9" pageId="33" pageNumber="486">50.3</td>
<td id="CE245DCAFFAF4DEBFB26EC0CFA87FDFD" box="[1242,1289,591,615]" gridcol="4" gridrow="9" pageId="33" pageNumber="486">54.7</td>
<td id="CE245DCAFFAF4DEBFAABEC0CFAF9FDFD" box="[1367,1399,591,615]" gridcol="5" gridrow="9" pageId="33" pageNumber="486">1.0</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEC34FAF9FD29" box="[166,1399,631,691]" gridrow="10" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEC34FD8CFD29" box="[166,514,631,691]" gridcol="0" gridrow="10" pageId="33" pageNumber="486">Distance between pectoral and pelvic-fin origins</th>
<td id="CE245DCAFFAF4DEBFD11EC34FCC4FD29" box="[749,842,631,691]" gridcol="1" gridrow="10" pageId="33" pageNumber="486">34.5</td>
<td id="CE245DCAFFAF4DEBFC3EEC34FC77FD29" box="[962,1017,631,691]" gridcol="2" gridrow="10" pageId="33" pageNumber="486">32.8</td>
<td id="CE245DCAFFAF4DEBFBE3EC34FB0DFD29" box="[1055,1155,631,691]" gridcol="3" gridrow="10" pageId="33" pageNumber="486">18.8</td>
<td id="CE245DCAFFAF4DEBFB26EC34FA87FD29" box="[1242,1289,631,691]" gridcol="4" gridrow="10" pageId="33" pageNumber="486">37.1</td>
<td id="CE245DCAFFAF4DEBFAABEC34FAF9FD29" box="[1367,1399,631,691]" gridcol="5" gridrow="10" pageId="33" pageNumber="486">3.6</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEC80FAF9FD65" box="[166,1399,707,767]" gridrow="11" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEC80FD8CFD65" box="[166,514,707,767]" gridcol="0" gridrow="11" pageId="33" pageNumber="486">Distance between pelvic and anal-fin origins</th>
<td id="CE245DCAFFAF4DEBFD11EC80FCC4FD65" box="[749,842,707,767]" gridcol="1" gridrow="11" pageId="33" pageNumber="486">19.2</td>
<td id="CE245DCAFFAF4DEBFC3EEC80FC77FD65" box="[962,1017,707,767]" gridcol="2" gridrow="11" pageId="33" pageNumber="486">18.9</td>
<td id="CE245DCAFFAF4DEBFBE3EC80FB0DFD65" box="[1055,1155,707,767]" gridcol="3" gridrow="11" pageId="33" pageNumber="486">17.3</td>
<td id="CE245DCAFFAF4DEBFB26EC80FA87FD65" box="[1242,1289,707,767]" gridcol="4" gridrow="11" pageId="33" pageNumber="486">20.7</td>
<td id="CE245DCAFFAF4DEBFAABEC80FAF9FD65" box="[1367,1399,707,767]" gridcol="5" gridrow="11" pageId="33" pageNumber="486">0.9</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AED4CFAF9FCB2" box="[166,1399,783,808]" gridrow="12" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AED4CFD8CFCB2" box="[166,514,783,808]" gridcol="0" gridrow="12" pageId="33" pageNumber="486">Depth of caudal peduncle</th>
<td id="CE245DCAFFAF4DEBFD11ED4CFCC4FCB2" box="[749,842,783,808]" gridcol="1" gridrow="12" pageId="33" pageNumber="486">11.4</td>
<td id="CE245DCAFFAF4DEBFC3EED4CFC77FCB2" box="[962,1017,783,808]" gridcol="2" gridrow="12" pageId="33" pageNumber="486">10.8</td>
<td id="CE245DCAFFAF4DEBFBE3ED4CFB0DFCB2" box="[1055,1155,783,808]" gridcol="3" gridrow="12" pageId="33" pageNumber="486">9.7</td>
<td id="CE245DCAFFAF4DEBFB26ED4CFA87FCB2" box="[1242,1289,783,808]" gridcol="4" gridrow="12" pageId="33" pageNumber="486">12.4</td>
<td id="CE245DCAFFAF4DEBFAABED4CFAF9FCB2" box="[1367,1399,783,808]" gridcol="5" gridrow="12" pageId="33" pageNumber="486">0.6</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AED7BFAF9FCCA" box="[166,1399,824,848]" gridrow="13" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AED7BFD8CFCCA" box="[166,514,824,848]" gridcol="0" gridrow="13" pageId="33" pageNumber="486">Length of caudal peduncle</th>
<td id="CE245DCAFFAF4DEBFD11ED7BFCC4FCCA" box="[749,842,824,848]" gridcol="1" gridrow="13" pageId="33" pageNumber="486">18.6</td>
<td id="CE245DCAFFAF4DEBFC3EED7BFC77FCCA" box="[962,1017,824,848]" gridcol="2" gridrow="13" pageId="33" pageNumber="486">19.1</td>
<td id="CE245DCAFFAF4DEBFBE3ED7BFB0DFCCA" box="[1055,1155,824,848]" gridcol="3" gridrow="13" pageId="33" pageNumber="486">16.6</td>
<td id="CE245DCAFFAF4DEBFB26ED7BFA87FCCA" box="[1242,1289,824,848]" gridcol="4" gridrow="13" pageId="33" pageNumber="486">21.3</td>
<td id="CE245DCAFFAF4DEBFAABED7BFAF9FCCA" box="[1367,1399,824,848]" gridcol="5" gridrow="13" pageId="33" pageNumber="486">1.3</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AED23FAF9FCE2" box="[166,1399,864,888]" gridrow="14" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AED23FD8CFCE2" box="[166,514,864,888]" gridcol="0" gridrow="14" pageId="33" pageNumber="486">Dorsal-fin depth</th>
<td id="CE245DCAFFAF4DEBFD11ED23FCC4FCE2" box="[749,842,864,888]" gridcol="1" gridrow="14" pageId="33" pageNumber="486">16.1</td>
<td id="CE245DCAFFAF4DEBFC3EED23FC77FCE2" box="[962,1017,864,888]" gridcol="2" gridrow="14" pageId="33" pageNumber="486">17.9</td>
<td id="CE245DCAFFAF4DEBFBE3ED23FB0DFCE2" box="[1055,1155,864,888]" gridcol="3" gridrow="14" pageId="33" pageNumber="486">11.8</td>
<td id="CE245DCAFFAF4DEBFB26ED23FA87FCE2" box="[1242,1289,864,888]" gridcol="4" gridrow="14" pageId="33" pageNumber="486">20.6</td>
<td id="CE245DCAFFAF4DEBFAABED23FAF9FCE2" box="[1367,1399,864,888]" gridcol="5" gridrow="14" pageId="33" pageNumber="486">1.9</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEDCBFAF9FC3A" box="[166,1399,904,928]" gridrow="15" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEDCBFD8CFC3A" box="[166,514,904,928]" gridcol="0" gridrow="15" pageId="33" pageNumber="486">Adipose-fin length</th>
<td id="CE245DCAFFAF4DEBFD11EDCBFCC4FC3A" box="[749,842,904,928]" gridcol="1" gridrow="15" pageId="33" pageNumber="486">20.0</td>
<td id="CE245DCAFFAF4DEBFC3EEDCBFC77FC3A" box="[962,1017,904,928]" gridcol="2" gridrow="15" pageId="33" pageNumber="486">20.2</td>
<td id="CE245DCAFFAF4DEBFBE3EDCBFB0DFC3A" box="[1055,1155,904,928]" gridcol="3" gridrow="15" pageId="33" pageNumber="486">18.3</td>
<td id="CE245DCAFFAF4DEBFB26EDCBFA87FC3A" box="[1242,1289,904,928]" gridcol="4" gridrow="15" pageId="33" pageNumber="486">22.4</td>
<td id="CE245DCAFFAF4DEBFAABEDCBFAF9FC3A" box="[1367,1399,904,928]" gridcol="5" gridrow="15" pageId="33" pageNumber="486">1.1</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEDF3FAF9FC77" box="[166,1399,944,1005]" gridrow="16" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEDF3FD8CFC77" box="[166,514,944,1005]" gridcol="0" gridrow="16" pageId="33" pageNumber="486">Distance between last dorsal ray and adipose origin</th>
<td id="CE245DCAFFAF4DEBFD11EDF3FCC4FC77" box="[749,842,944,1005]" gridcol="1" gridrow="16" pageId="33" pageNumber="486">25.3</td>
<td id="CE245DCAFFAF4DEBFC3EEDF3FC77FC77" box="[962,1017,944,1005]" gridcol="2" gridrow="16" pageId="33" pageNumber="486">23.8</td>
<td id="CE245DCAFFAF4DEBFBE3EDF3FB0DFC77" box="[1055,1155,944,1005]" gridcol="3" gridrow="16" pageId="33" pageNumber="486">20.1</td>
<td id="CE245DCAFFAF4DEBFB26EDF3FA87FC77" box="[1242,1289,944,1005]" gridcol="4" gridrow="16" pageId="33" pageNumber="486">27.5</td>
<td id="CE245DCAFFAF4DEBFAABEDF3FAF9FC77" box="[1367,1399,944,1005]" gridcol="5" gridrow="16" pageId="33" pageNumber="486">1.7</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEDBEFAF9FB8F" box="[166,1399,1021,1045]" gridrow="17" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEDBEFD8CFB8F" box="[166,514,1021,1045]" gridcol="0" gridrow="17" pageId="33" pageNumber="486">Anal-fin base length</th>
<td id="CE245DCAFFAF4DEBFD11EDBEFCC4FB8F" box="[749,842,1021,1045]" gridcol="1" gridrow="17" pageId="33" pageNumber="486">11.8</td>
<td id="CE245DCAFFAF4DEBFC3EEDBEFC77FB8F" box="[962,1017,1021,1045]" gridcol="2" gridrow="17" pageId="33" pageNumber="486">12.0</td>
<td id="CE245DCAFFAF4DEBFBE3EDBEFB0DFB8F" box="[1055,1155,1021,1045]" gridcol="3" gridrow="17" pageId="33" pageNumber="486">11.0</td>
<td id="CE245DCAFFAF4DEBFB26EDBEFA87FB8F" box="[1242,1289,1021,1045]" gridcol="4" gridrow="17" pageId="33" pageNumber="486">13.4</td>
<td id="CE245DCAFFAF4DEBFAABEDBEFAF9FB8F" box="[1367,1399,1021,1045]" gridcol="5" gridrow="17" pageId="33" pageNumber="486">0.6</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEA66FAF9FBA7" box="[166,1399,1061,1085]" gridrow="18" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEA66FD8CFBA7" box="[166,514,1061,1085]" gridcol="0" gridrow="18" pageId="33" pageNumber="486">Pectoral-fin length</th>
<td id="CE245DCAFFAF4DEBFD11EA66FCC4FBA7" box="[749,842,1061,1085]" gridcol="1" gridrow="18" pageId="33" pageNumber="486">23.4</td>
<td id="CE245DCAFFAF4DEBFC3EEA66FC77FBA7" box="[962,1017,1061,1085]" gridcol="2" gridrow="18" pageId="33" pageNumber="486">23.9</td>
<td id="CE245DCAFFAF4DEBFBE3EA66FB0DFBA7" box="[1055,1155,1061,1085]" gridcol="3" gridrow="18" pageId="33" pageNumber="486">22.0</td>
<td id="CE245DCAFFAF4DEBFB26EA66FA87FBA7" box="[1242,1289,1061,1085]" gridcol="4" gridrow="18" pageId="33" pageNumber="486">25.1</td>
<td id="CE245DCAFFAF4DEBFAABEA66FAF9FBA7" box="[1367,1399,1061,1085]" gridcol="5" gridrow="18" pageId="33" pageNumber="486">0.8</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEA0EFAF9FBFF" box="[166,1399,1101,1125]" gridrow="19" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEA0EFD8CFBFF" box="[166,514,1101,1125]" gridcol="0" gridrow="19" pageId="33" pageNumber="486">Pelvic-fin length</th>
<td id="CE245DCAFFAF4DEBFD11EA0EFCC4FBFF" box="[749,842,1101,1125]" gridcol="1" gridrow="19" pageId="33" pageNumber="486">16.2</td>
<td id="CE245DCAFFAF4DEBFC3EEA0EFC77FBFF" box="[962,1017,1101,1125]" gridcol="2" gridrow="19" pageId="33" pageNumber="486">16.5</td>
<td id="CE245DCAFFAF4DEBFBE3EA0EFB0DFBFF" box="[1055,1155,1101,1125]" gridcol="3" gridrow="19" pageId="33" pageNumber="486">14.8</td>
<td id="CE245DCAFFAF4DEBFB26EA0EFA87FBFF" box="[1242,1289,1101,1125]" gridcol="4" gridrow="19" pageId="33" pageNumber="486">18.0</td>
<td id="CE245DCAFFAF4DEBFAABEA0EFAF9FBFF" box="[1367,1399,1101,1125]" gridcol="5" gridrow="19" pageId="33" pageNumber="486">0.7</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEA36FAF9FB17" box="[166,1399,1141,1165]" gridrow="20" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEA36FAF9FB17" box="[166,1399,1141,1165]" colspan="6" colspanRight="5" gridcol="0" gridrow="20" pageId="33" pageNumber="486">
<emphasis id="01B1EAE6FFAFB235FF5AEA36FE3BFB17" bold="true" box="[166,437,1141,1165]" pageId="33" pageNumber="486">In percent of head length</emphasis>
</th>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEADDFAF9FB2C" box="[166,1399,1182,1206]" gridrow="21" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEADDFD8CFB2C" box="[166,514,1182,1206]" gridcol="0" gridrow="21" pageId="33" pageNumber="486">Head depth at eye</th>
<td id="CE245DCAFFAF4DEBFD11EADDFCC4FB2C" box="[749,842,1182,1206]" gridcol="1" gridrow="21" pageId="33" pageNumber="486">41</td>
<td id="CE245DCAFFAF4DEBFC3EEADDFC77FB2C" box="[962,1017,1182,1206]" gridcol="2" gridrow="21" pageId="33" pageNumber="486">43.3</td>
<td id="CE245DCAFFAF4DEBFBE3EADDFB0DFB2C" box="[1055,1155,1182,1206]" gridcol="3" gridrow="21" pageId="33" pageNumber="486">39</td>
<td id="CE245DCAFFAF4DEBFB26EADDFA87FB2C" box="[1242,1289,1182,1206]" gridcol="4" gridrow="21" pageId="33" pageNumber="486">48</td>
<td id="CE245DCAFFAF4DEBFAABEADDFAF9FB2C" box="[1367,1399,1182,1206]" gridcol="5" gridrow="21" pageId="33" pageNumber="486">2.4</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEA85FAF9FB44" box="[166,1399,1222,1246]" gridrow="22" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEA85FD8CFB44" box="[166,514,1222,1246]" gridcol="0" gridrow="22" pageId="33" pageNumber="486">Snout length</th>
<td id="CE245DCAFFAF4DEBFD11EA85FCC4FB44" box="[749,842,1222,1246]" gridcol="1" gridrow="22" pageId="33" pageNumber="486">50</td>
<td id="CE245DCAFFAF4DEBFC3EEA85FC77FB44" box="[962,1017,1222,1246]" gridcol="2" gridrow="22" pageId="33" pageNumber="486">50.7</td>
<td id="CE245DCAFFAF4DEBFBE3EA85FB0DFB44" box="[1055,1155,1222,1246]" gridcol="3" gridrow="22" pageId="33" pageNumber="486">47</td>
<td id="CE245DCAFFAF4DEBFB26EA85FA87FB44" box="[1242,1289,1222,1246]" gridcol="4" gridrow="22" pageId="33" pageNumber="486">56</td>
<td id="CE245DCAFFAF4DEBFAABEA85FAF9FB44" box="[1367,1399,1222,1246]" gridcol="5" gridrow="22" pageId="33" pageNumber="486">1.8</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEAADFAF9FA9D" box="[166,1399,1262,1287]" gridrow="23" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEAADFD8CFA9D" box="[166,514,1262,1287]" gridcol="0" gridrow="23" pageId="33" pageNumber="486">Eye diameter</th>
<td id="CE245DCAFFAF4DEBFD11EAADFCC4FA9D" box="[749,842,1262,1287]" gridcol="1" gridrow="23" pageId="33" pageNumber="486">7</td>
<td id="CE245DCAFFAF4DEBFC3EEAADFC77FA9D" box="[962,1017,1262,1287]" gridcol="2" gridrow="23" pageId="33" pageNumber="486">9.1</td>
<td id="CE245DCAFFAF4DEBFBE3EAADFB0DFA9D" box="[1055,1155,1262,1287]" gridcol="3" gridrow="23" pageId="33" pageNumber="486">7</td>
<td id="CE245DCAFFAF4DEBFB26EAADFA87FA9D" box="[1242,1289,1262,1287]" gridcol="4" gridrow="23" pageId="33" pageNumber="486">13</td>
<td id="CE245DCAFFAF4DEBFAABEAADFAF9FA9D" box="[1367,1399,1262,1287]" gridcol="5" gridrow="23" pageId="33" pageNumber="486">1.3</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEB55FAF9FAB5" box="[166,1399,1302,1327]" gridrow="24" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEB55FD8CFAB5" box="[166,514,1302,1327]" gridcol="0" gridrow="24" pageId="33" pageNumber="486">Postorbital distance</th>
<td id="CE245DCAFFAF4DEBFD11EB55FCC4FAB5" box="[749,842,1302,1327]" gridcol="1" gridrow="24" pageId="33" pageNumber="486">47</td>
<td id="CE245DCAFFAF4DEBFC3EEB55FC77FAB5" box="[962,1017,1302,1327]" gridcol="2" gridrow="24" pageId="33" pageNumber="486">45.6</td>
<td id="CE245DCAFFAF4DEBFBE3EB55FB0DFAB5" box="[1055,1155,1302,1327]" gridcol="3" gridrow="24" pageId="33" pageNumber="486">40</td>
<td id="CE245DCAFFAF4DEBFB26EB55FA87FAB5" box="[1242,1289,1302,1327]" gridcol="4" gridrow="24" pageId="33" pageNumber="486">49</td>
<td id="CE245DCAFFAF4DEBFAABEB55FAF9FAB5" box="[1367,1399,1302,1327]" gridcol="5" gridrow="24" pageId="33" pageNumber="486">1.9</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEB7CFAF9FACD" box="[166,1399,1343,1367]" gridrow="25" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEB7CFD8CFACD" box="[166,514,1343,1367]" gridcol="0" gridrow="25" pageId="33" pageNumber="486">Maximum head width</th>
<td id="CE245DCAFFAF4DEBFD11EB7CFCC4FACD" box="[749,842,1343,1367]" gridcol="1" gridrow="25" pageId="33" pageNumber="486">79</td>
<td id="CE245DCAFFAF4DEBFC3EEB7CFC77FACD" box="[962,1017,1343,1367]" gridcol="2" gridrow="25" pageId="33" pageNumber="486">83.3</td>
<td id="CE245DCAFFAF4DEBFBE3EB7CFB0DFACD" box="[1055,1155,1343,1367]" gridcol="3" gridrow="25" pageId="33" pageNumber="486">79</td>
<td id="CE245DCAFFAF4DEBFB26EB7CFA87FACD" box="[1242,1289,1343,1367]" gridcol="4" gridrow="25" pageId="33" pageNumber="486">89</td>
<td id="CE245DCAFFAF4DEBFAABEB7CFAF9FACD" box="[1367,1399,1343,1367]" gridcol="5" gridrow="25" pageId="33" pageNumber="486">3.0</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEB24FAF9FAE5" box="[166,1399,1383,1407]" gridrow="26" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEB24FD8CFAE5" box="[166,514,1383,1407]" gridcol="0" gridrow="26" pageId="33" pageNumber="486">Interorbital distance</th>
<td id="CE245DCAFFAF4DEBFD11EB24FCC4FAE5" box="[749,842,1383,1407]" gridcol="1" gridrow="26" pageId="33" pageNumber="486">26</td>
<td id="CE245DCAFFAF4DEBFC3EEB24FC77FAE5" box="[962,1017,1383,1407]" gridcol="2" gridrow="26" pageId="33" pageNumber="486">28.6</td>
<td id="CE245DCAFFAF4DEBFBE3EB24FB0DFAE5" box="[1055,1155,1383,1407]" gridcol="3" gridrow="26" pageId="33" pageNumber="486">24</td>
<td id="CE245DCAFFAF4DEBFB26EB24FA87FAE5" box="[1242,1289,1383,1407]" gridcol="4" gridrow="26" pageId="33" pageNumber="486">33</td>
<td id="CE245DCAFFAF4DEBFAABEB24FAF9FAE5" box="[1367,1399,1383,1407]" gridcol="5" gridrow="26" pageId="33" pageNumber="486">2.2</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEBCCFAF9FA3D" box="[166,1399,1423,1447]" gridrow="27" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEBCCFD8CFA3D" box="[166,514,1423,1447]" gridcol="0" gridrow="27" pageId="33" pageNumber="486">Length of inner mandibular barbel</th>
<td id="CE245DCAFFAF4DEBFD11EBCCFCC4FA3D" box="[749,842,1423,1447]" gridcol="1" gridrow="27" pageId="33" pageNumber="486">22</td>
<td id="CE245DCAFFAF4DEBFC3EEBCCFC77FA3D" box="[962,1017,1423,1447]" gridcol="2" gridrow="27" pageId="33" pageNumber="486">25.5</td>
<td id="CE245DCAFFAF4DEBFBE3EBCCFB0DFA3D" box="[1055,1155,1423,1447]" gridcol="3" gridrow="27" pageId="33" pageNumber="486">21</td>
<td id="CE245DCAFFAF4DEBFB26EBCCFA87FA3D" box="[1242,1289,1423,1447]" gridcol="4" gridrow="27" pageId="33" pageNumber="486">30</td>
<td id="CE245DCAFFAF4DEBFAABEBCCFAF9FA3D" box="[1367,1399,1423,1447]" gridcol="5" gridrow="27" pageId="33" pageNumber="486">2.5</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEBF4FAF9FA55" box="[166,1399,1463,1487]" gridrow="28" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEBF4FD8CFA55" box="[166,514,1463,1487]" gridcol="0" gridrow="28" pageId="33" pageNumber="486">Length of outer mandibular barbel</th>
<td id="CE245DCAFFAF4DEBFD11EBF4FCC4FA55" box="[749,842,1463,1487]" gridcol="1" gridrow="28" pageId="33" pageNumber="486">45</td>
<td id="CE245DCAFFAF4DEBFC3EEBF4FC77FA55" box="[962,1017,1463,1487]" gridcol="2" gridrow="28" pageId="33" pageNumber="486">46.9</td>
<td id="CE245DCAFFAF4DEBFBE3EBF4FB0DFA55" box="[1055,1155,1463,1487]" gridcol="3" gridrow="28" pageId="33" pageNumber="486">42</td>
<td id="CE245DCAFFAF4DEBFB26EBF4FA87FA55" box="[1242,1289,1463,1487]" gridcol="4" gridrow="28" pageId="33" pageNumber="486">51</td>
<td id="CE245DCAFFAF4DEBFAABEBF4FAF9FA55" box="[1367,1399,1463,1487]" gridcol="5" gridrow="28" pageId="33" pageNumber="486">2.6</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AEB9CFAF9FA62" box="[166,1399,1503,1528]" gridrow="29" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AEB9CFD8CFA62" box="[166,514,1503,1528]" gridcol="0" gridrow="29" pageId="33" pageNumber="486">Length of maxillary barbel</th>
<td id="CE245DCAFFAF4DEBFD11EB9CFCC4FA62" box="[749,842,1503,1528]" gridcol="1" gridrow="29" pageId="33" pageNumber="486">73</td>
<td id="CE245DCAFFAF4DEBFC3EEB9CFC77FA62" box="[962,1017,1503,1528]" gridcol="2" gridrow="29" pageId="33" pageNumber="486">78.9</td>
<td id="CE245DCAFFAF4DEBFBE3EB9CFB0DFA62" box="[1055,1155,1503,1528]" gridcol="3" gridrow="29" pageId="33" pageNumber="486">72</td>
<td id="CE245DCAFFAF4DEBFB26EB9CFA87FA62" box="[1242,1289,1503,1528]" gridcol="4" gridrow="29" pageId="33" pageNumber="486">85</td>
<td id="CE245DCAFFAF4DEBFAABEB9CFAF9FA62" box="[1367,1399,1503,1528]" gridcol="5" gridrow="29" pageId="33" pageNumber="486">3.3</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AE84BFAF9F9BA" box="[166,1399,1544,1568]" gridrow="30" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AE84BFD8CF9BA" box="[166,514,1544,1568]" gridcol="0" gridrow="30" pageId="33" pageNumber="486">Length of nasal barbel</th>
<td id="CE245DCAFFAF4DEBFD11E84BFCC4F9BA" box="[749,842,1544,1568]" gridcol="1" gridrow="30" pageId="33" pageNumber="486">23</td>
<td id="CE245DCAFFAF4DEBFC3EE84BFC77F9BA" box="[962,1017,1544,1568]" gridcol="2" gridrow="30" pageId="33" pageNumber="486">27.4</td>
<td id="CE245DCAFFAF4DEBFBE3E84BFB0DF9BA" box="[1055,1155,1544,1568]" gridcol="3" gridrow="30" pageId="33" pageNumber="486">16</td>
<td id="CE245DCAFFAF4DEBFB26E84BFA87F9BA" box="[1242,1289,1544,1568]" gridcol="4" gridrow="30" pageId="33" pageNumber="486">32</td>
<td id="CE245DCAFFAF4DEBFAABE84BFAF9F9BA" box="[1367,1399,1544,1568]" gridcol="5" gridrow="30" pageId="33" pageNumber="486">3.4</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AE873FAF9F9D2" box="[166,1399,1584,1608]" gridrow="31" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AE873FD8CF9D2" box="[166,514,1584,1608]" gridcol="0" gridrow="31" pageId="33" pageNumber="486">Apparatus length</th>
<td id="CE245DCAFFAF4DEBFD11E873FCC4F9D2" box="[749,842,1584,1608]" gridcol="1" gridrow="31" pageId="33" pageNumber="486">54</td>
<td id="CE245DCAFFAF4DEBFC3EE873FC77F9D2" box="[962,1017,1584,1608]" gridcol="2" gridrow="31" pageId="33" pageNumber="486">57.8</td>
<td id="CE245DCAFFAF4DEBFBE3E873FB0DF9D2" box="[1055,1155,1584,1608]" gridcol="3" gridrow="31" pageId="33" pageNumber="486">53</td>
<td id="CE245DCAFFAF4DEBFB26E873FA87F9D2" box="[1242,1289,1584,1608]" gridcol="4" gridrow="31" pageId="33" pageNumber="486">62</td>
<td id="CE245DCAFFAF4DEBFAABE873FAF9F9D2" box="[1367,1399,1584,1608]" gridcol="5" gridrow="31" pageId="33" pageNumber="486">2.4</td>
</tr>
<tr id="8DF534B6FFAF4DEBFF5AE81BFAF9F9EB" box="[166,1399,1624,1649]" gridrow="32" pageId="33" pageNumber="486">
<th id="CE245DCAFFAF4DEBFF5AE81BFD8CF9EB" box="[166,514,1624,1649]" gridcol="0" gridrow="32" pageId="33" pageNumber="486">Apparatus width</th>
<td id="CE245DCAFFAF4DEBFD11E81BFCC4F9EB" box="[749,842,1624,1649]" gridcol="1" gridrow="32" pageId="33" pageNumber="486">47</td>
<td id="CE245DCAFFAF4DEBFC3EE81BFC77F9EB" box="[962,1017,1624,1649]" gridcol="2" gridrow="32" pageId="33" pageNumber="486">49.0</td>
<td id="CE245DCAFFAF4DEBFBE3E81BFB0DF9EB" box="[1055,1155,1624,1649]" gridcol="3" gridrow="32" pageId="33" pageNumber="486">45</td>
<td id="CE245DCAFFAF4DEBFB26E81BFA87F9EB" box="[1242,1289,1624,1649]" gridcol="4" gridrow="32" pageId="33" pageNumber="486">53</td>
<td id="CE245DCAFFAF4DEBFAABE81BFAF9F9EB" box="[1367,1399,1624,1649]" gridcol="5" gridrow="32" pageId="33" pageNumber="486">2.7</td>
</tr>
</table>
</paragraph>
<paragraph id="337A36F4FFAFB236FF3BE8E3FBD6FE13" blockId="33.[151,1437,1696,2010]" lastBlockId="34.[151,1436,151,393]" lastPageId="34" lastPageNumber="487" pageId="33" pageNumber="486">
Thoracic adhesive apparatus consisting of keratinised striae in an elongate oblong field extending from isthmus to almost posterior limit of pectoral-fin base (
<figureCitation id="ABFE2A71FFAFB235FD5DE887FD78F944" box="[673,758,1732,1758]" captionStart="FIGURE 33" captionStartId="30.[151,250,1904,1928]" captionTargetBox="[151,1436,1225,1879]" captionTargetId="figure-158@30.[151,1436,1225,1879]" captionTargetPageId="30" captionText="FIGURE 33. Glyptothorax daemon, paratypes, from left: FSJF 3962, 77 mm SL, 84 mm SL, 101 mm SL, 109 mm SL; Iraq: Choman River." figureDoi="http://doi.org/10.5281/zenodo.14895417" httpUri="https://zenodo.org/record/14895417/files/figure.png" pageId="33" pageNumber="486">Fig. 33</figureCitation>
); anterolateral edges of adhesive apparatus slightly convex, often almost straight; its width 1.11.3 times in its length; completely situated on a horse-shoe shaped swelling, associated with few, small warts at its lateral edge and without warts at its posterior edge. Anteromedial striae present and well developed. Narrow, spear-blade shaped medial pit on posterior half of thoracic adhesive apparatus. Dorsal fin located above anterior third of body, with 6 (10) branched rays; fin margin straight or slightly concave; spine short and straight, smooth on anterior and posterior margin; distal 1/3 poorly ossified and soft. Adipose fin with anterior margin straight or slightly convex and posterior margin roundish; its origin at vertical through or very slightly in front of anal-fin origin. Caudal fin with rounded lobes, lower lobe slightly longer than upper lobe and i,8+7,i (10) principal rays. Anal-fin base vertically opposite adipose-fin base. Anal fin with slightly convex anterior margin and straight or slightly concave or convex posterior margin; with 7 (8), or 6 (2) branched rays. Pelvic-fin origin at vertical through or slightly in front of or behind tip of adpressed dorsal fin. Pelvic fin with slightly convex anterior margin and I,5 (10) rays; tip of adpressed fin reaching anal-fin origin. Pectoral fin with I,8 (10) rays; posterior fin margin straight; anterior spine margin smooth, with many unculi and a honey-comp pattern on lower surface, inner margin with 711 serrae. Back anterior to adipose fin flat or slightly rounded, with a shallow keel in some individuals, expanded distal tips of neural spines not forming a series of bumps.
</paragraph>
<caption id="67BA667CFFACB236FF6BEDE1FC7DFC20" ID-DOI="http://doi.org/10.5281/zenodo.14895423" ID-Zenodo-Dep="14895423" box="[151,1011,930,954]" httpUri="https://zenodo.org/record/14895423/files/figure.png" pageId="34" pageNumber="487" startId="34.[151,250,930,954]" targetBox="[151,1436,417,907]" targetPageId="34" targetType="figure">
<paragraph id="337A36F4FFACB236FF6BEDE1FC7DFC20" blockId="34.[151,1011,930,954]" box="[151,1011,930,954]" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF6BEDE1FEAEFC20" bold="true" box="[151,288,930,954]" pageId="34" pageNumber="487">FIGURE 36.</emphasis>
<taxonomicName id="F4C54D77FFACB236FEDAEDE1FDCAFC20" authority=", FSJF" authorityName="FSJF" box="[294,580,930,954]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FEDAEDE1FD8DFC20" box="[294,515,930,954]" italics="true" pageId="34" pageNumber="487">Glyptothorax daemon</emphasis>
, FSJF
</taxonomicName>
3677, 91 mm SL; Turkey: stream Çıratan.
</paragraph>
</caption>
<paragraph id="337A36F4FFACB236FF3BEDABFB6CFADC" blockId="34.[151,1437,1000,2035]" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF3BEDABFEC1FB98" bold="true" box="[199,335,1000,1026]" pageId="34" pageNumber="487">Coloration.</emphasis>
In 70% ethanol: dorsal and lateral surfaces of head and body dark-grey to greyish brown, fading to pale-grey or beige on ventral surfaces of head and anterior belly and on pectoral and pelvic-fin bases. Dorsal and lateral surfaces of head and body with small irregular spots and / or small blotches slightly darker than background coloration. Spots smaller than eye diameter, blotches as large as eye or slightly larger. Latero-sensory pores same colour as surrounding tissue. A pale-grey blotch at dorsal-fin origin. Adipose fin with a pale-grey blotch behind origin and a pale-grey posterior margin. All other fins with a proximal dark-grey to blackish base, followed by a pale-grey band, then a dark-grey band and a hyaline or pale-grey margin; appearing as dark-grey or blackish fins with a whitish band and margin; whitish margin in caudal fin often absent or reduced to a large or small blotch on each lobe. Pattern in fins dissociated and lost in large individuals. Maxillary and nasal barbels grey or blackish dorsally, pale-grey ventrally and velum pale-grey or beige. Mandibular barbels beige or pale-grey.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFFACB236FF3BEB12FEE1FA14" pageId="34" pageNumber="487" type="etymology">
<paragraph id="337A36F4FFACB236FF3BEB12FEE1FA14" blockId="34.[151,1437,1000,2035]" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF3BEB12FEC0FAF1" bold="true" box="[199,334,1361,1387]" pageId="34" pageNumber="487">Etymology.</emphasis>
The specific epithet is latinised from the Greek adjective
<emphasis id="01B1EAE6FFACB236FC27EB13FBA3FAF1" box="[987,1069,1360,1387]" italics="true" pageId="34" pageNumber="487">δαίμων</emphasis>
usually interpreted as a ghost. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFFACB236FF3BEBDBFB58FA29" box="[199,1238,1432,1459]" pageId="34" pageNumber="487" type="distribution">
<paragraph id="337A36F4FFACB236FF3BEBDBFB58FA29" blockId="34.[151,1437,1000,2035]" box="[199,1238,1432,1459]" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF3BEBDBFEEFFA28" bold="true" box="[199,353,1432,1458]" pageId="34" pageNumber="487">Distribution.</emphasis>
Headwater streams in upper Tigris and Great Zab drainage in
<collectingCountry id="4BD27664FFACB236FBE7EBDBFBE5FA28" box="[1051,1131,1432,1458]" name="Turkey" pageId="34" pageNumber="487">Turkey</collectingCountry>
and
<collectingCountry id="4BD27664FFACB236FB5DEBDAFB5CFA29" box="[1185,1234,1433,1459]" name="Iraq" pageId="34" pageNumber="487">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFFACB236FF3BEBFFFDAEFA60" pageId="34" pageNumber="487" type="biology_ecology">
<paragraph id="337A36F4FFACB236FF3BEBFFFDAEFA60" blockId="34.[151,1437,1000,2035]" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF3BEBFFFEA4FA4C" bold="true" box="[199,298,1468,1494]" pageId="34" pageNumber="487">Habitat.</emphasis>
<taxonomicName id="F4C54D77FFACB236FECCEBFFFDAAFA4C" box="[304,548,1468,1494]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FECCEBFFFDAAFA4C" box="[304,548,1468,1494]" italics="true" pageId="34" pageNumber="487">Glyptothorax daemon</emphasis>
</taxonomicName>
is found in upland rivers and streams with fast current and a substrate composed predominantly of rocks and gravel.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFFACB237FF3BE846FED8FF4F" lastPageId="35" lastPageNumber="488" pageId="34" pageNumber="487" type="discussion">
<paragraph id="337A36F4FFACB237FF3BE846FED8FF4F" blockId="34.[151,1437,1000,2035]" lastBlockId="35.[151,1436,151,213]" lastPageId="35" lastPageNumber="488" pageId="34" pageNumber="487">
<emphasis id="01B1EAE6FFACB236FF3BE846FEB5F985" bold="true" box="[199,315,1541,1567]" pageId="34" pageNumber="487">Remarks.</emphasis>
The four populations of
<taxonomicName id="F4C54D77FFACB236FDA0E846FD57F984" box="[604,729,1540,1567]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FDA0E846FD57F984" box="[604,729,1540,1567]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
examined for this study are sorted into two molecular groups with a minimum K2P distance of 2.0% in their DNA barcode sequence. We were unable to distinguish fishes from these two molecular groups by the morphological characters examined and therefore treat them as conspecific. Indeed, this is a very surprising situation as all sequenced individuals of
<taxonomicName id="F4C54D77FFACB236FC39E832FB95F910" box="[965,1051,1649,1675]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFACB236FC39E832FB95F910" box="[965,1051,1649,1675]" italics="true" pageId="34" pageNumber="487">G. cous</emphasis>
</taxonomicName>
cluster as sister to one
<taxonomicName id="F4C54D77FFACB236FADDE832FA13F910" box="[1313,1437,1648,1675]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FADDE832FA13F910" box="[1313,1437,1648,1675]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
group with a K2P distance of 1.2% between them. While a K2P distance of 2.0% between populations of a fish species quite specialised for torrent headwaters is not a surprise, the close relationship of
<taxonomicName id="F4C54D77FFACB236FB8DE8FAFB65F948" box="[1137,1259,1720,1747]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FB8DE8FAFB65F948" box="[1137,1259,1720,1747]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
with
<taxonomicName id="F4C54D77FFACB236FAD4E8FAFAF2F948" box="[1320,1404,1721,1747]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFACB236FAD4E8FAFAF2F948" box="[1320,1404,1721,1747]" italics="true" pageId="34" pageNumber="487">G. cous</emphasis>
</taxonomicName>
is. Potentially,
<taxonomicName id="F4C54D77FFACB236FEE1E89EFE17F96C" box="[285,409,1756,1783]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FEE1E89EFE17F96C" box="[285,409,1756,1783]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
had hybridised with
<taxonomicName id="F4C54D77FFACB236FD7BE89EFD52F96C" box="[647,732,1757,1783]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFACB236FD7BE89EFD52F96C" box="[647,732,1757,1783]" italics="true" pageId="34" pageNumber="487">G. cous</emphasis>
</taxonomicName>
and at least some populations carry the mtDNA of
<taxonomicName id="F4C54D77FFACB236FADDE89EFA13F96C" box="[1313,1437,1756,1783]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FADDE89EFA13F96C" box="[1313,1437,1756,1783]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
as an indicator of a past introgressive hybridisation. Therefore, we would expect that fish identified as
<taxonomicName id="F4C54D77FFACB236FADEE942FAF6F880" box="[1314,1400,1793,1819]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFACB236FADEE942FAF6F880" box="[1314,1400,1793,1819]" italics="true" pageId="34" pageNumber="487">G. cous</emphasis>
</taxonomicName>
by morphological characters but having quite different mtDNA might be found in the future.Alternatively, both species are just very closely related. With only mitochondrial COI data available, the phylogenetic relationship between
<taxonomicName id="F4C54D77FFACB236FF6BE92EFE9AF81C" box="[151,276,1900,1927]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FF6BE92EFE9AF81C" box="[151,276,1900,1927]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FFACB236FEB0E92EFE2CF81C" box="[332,418,1901,1927]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFACB236FEB0E92EFE2CF81C" box="[332,418,1901,1927]" italics="true" pageId="34" pageNumber="487">G. cous</emphasis>
</taxonomicName>
cannot be resolved. We therefore regard it more appropriate and parsimonious to identify both populations as
<taxonomicName id="F4C54D77FFACB236FE8AE9D2FE7FF830" box="[374,497,1936,1963]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FE8AE9D2FE7FF830" box="[374,497,1936,1963]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
than to describe an additional species, only distinguished from
<taxonomicName id="F4C54D77FFACB236FB49E9D2FABEF830" box="[1205,1328,1936,1963]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="487" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFACB236FB49E9D2FABEF830" box="[1205,1328,1936,1963]" italics="true" pageId="34" pageNumber="487">G. daemon</emphasis>
</taxonomicName>
by a K2P distance of 2.0% without any additional support from morphology or (unstudied) nuclear DNA sequences. Only a study on nuclear DNA could exclude the possibility of an introgressive hybridisation and clarify the phylogeny of this group. We do not identify
<taxonomicName id="F4C54D77FFADB237FE14EED4FDECFF2B" box="[488,610,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="488" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FFADB237FE14EED4FDECFF2B" box="[488,610,151,177]" italics="true" pageId="35" pageNumber="488">G. daemon</emphasis>
</taxonomicName>
to be conspecific with
<taxonomicName id="F4C54D77FFADB237FC9EEED4FC38FF2A" box="[866,950,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="488" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FFADB237FC9EEED4FC38FF2A" box="[866,950,151,177]" italics="true" pageId="35" pageNumber="488">G. cous</emphasis>
</taxonomicName>
, as both species are very well distinguished morphologically.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,507 @@
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<mods:title id="4763C92895D891811536B782D15840C2">The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae)</mods:title>
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<mods:namePart id="C9511819E00C0161A996EB9733C6426A">Freyhof, Jörg</mods:namePart>
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<mods:namePart id="AA0A52E702C9C3A62D88142ACA1F1D6F">Kaya, Cüneyt</mods:namePart>
<mods:affiliation id="3CC5BED14F33E280FFB39E1E6D9D1846">Recep Tayyip Erdogan University, Faculty of Fisheries and Aquatic Sciences, 53100 Rize, Turkey.</mods:affiliation>
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<subSubSection id="7BDF657FFF94B20EFF6BEBECFD66FA53" box="[151,744,1455,1482]" pageId="26" pageNumber="479" type="nomenclature">
<paragraph id="337A36F4FF94B20EFF6BEBECFD66FA53" blockId="26.[151,744,1455,1517]" box="[151,744,1455,1482]" pageId="26" pageNumber="479">
<heading id="68328198FF94B20EFF6BEBECFD66FA53" bold="true" box="[151,744,1455,1482]" fontSize="11" level="1" pageId="26" pageNumber="479" reason="1">
<taxonomicName id="F4C54D77FF94B20EFF6BEBECFD66FA53" ID-CoL="3GJF4" authority="(Pietschmann, 1913)" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[151,744,1455,1482]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="479" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF94B20EFF6BEBECFD66FA53" bold="true" box="[151,744,1455,1482]" pageId="26" pageNumber="479">
<emphasis id="01B1EAE6FF94B20EFF6BEBECFE6DFA50" bold="true" box="[151,483,1455,1482]" italics="true" pageId="26" pageNumber="479">Glyptothorax steindachneri</emphasis>
(
<bibRefCitation id="57544B05FF94B20EFE0EEBECFD6EFA53" author="Pietschmann, V." box="[498,736,1455,1481]" pageId="26" pageNumber="479" pagination="93 - 95" refId="ref19664" refString="Pietschmann, V. (1913) Eine neue Glyptosternum - Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93-95." type="journal article" year="1913">Pietschmann, 1913</bibRefCitation>
)
</emphasis>
</taxonomicName>
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<subSubSection id="7BDF657FFF94B20EFF6BEB90FEADFA77" box="[151,291,1491,1517]" pageId="26" pageNumber="479" type="description">
<paragraph id="337A36F4FF94B20EFF6BEB90FEADFA77" blockId="26.[151,744,1455,1517]" box="[151,291,1491,1517]" pageId="26" pageNumber="479">
(
<figureCitation id="ABFE2A71FF94B20EFF63EB90FE95FA77" box="[159,283,1491,1517]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStart-2="FIGURE 30" captionStartId-0="27.[151,250,1498,1522]" captionStartId-1="28.[151,250,1041,1065]" captionStartId-2="28.[151,250,1607,1631]" captionTargetBox-0="[151,1436,705,1475]" captionTargetBox-1="[151,1436,345,1018]" captionTargetBox-2="[159,1436,1185,1551]" captionTargetId-0="figure-289@27.[151,1436,705,1475]" captionTargetId-1="figure-108@28.[151,1436,345,1018]" captionTargetId-2="figure-148@28.[151,1436,1126,1583]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="28" captionText-0="FIGURE 28. Glyptothorax steindachneri, from top: SMF 23678, 77 mm SL, Syria: Nahr Al-Khabur; SMF 28644, 112 mm SL, Syria: Der ez Zor; SMF 28694, 155 mm SL; 158 mm SL, Syria: Euphrates at Raqqa." captionText-1="FIGURE 29. Glyptothorax steindachneri, from left: SMF 23678, 77 mm SL, Syria: Nahr Al-Khabur; SMF 28644, 112 mm SL, 130 mm SL, Syria: Der ez Zor; SMF 28694, 158 mm SL, Syria: Euphrates at Raqqa." captionText-2="FIGURE 30. Glyptothorax steindachneri, not preserved, about 115 mm SL; Iraq: Qalah Chwalan River, 35.9150, 45.36788." figureDoi-0="http://doi.org/10.5281/zenodo.14895407" figureDoi-1="http://doi.org/10.5281/zenodo.14895409" figureDoi-2="http://doi.org/10.5281/zenodo.14895411" httpUri-0="https://zenodo.org/record/14895407/files/figure.png" httpUri-1="https://zenodo.org/record/14895409/files/figure.png" httpUri-2="https://zenodo.org/record/14895411/files/figure.png" pageId="26" pageNumber="479">Fig. 2830</figureCitation>
)
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</subSubSection>
<subSubSection id="7BDF657FFF94B20EFF6BE85BFD2BF9AA" box="[151,677,1560,1584]" pageId="26" pageNumber="479" type="reference_group">
<paragraph id="337A36F4FF94B20EFF6BE85BFD2BF9AA" blockId="26.[151,677,1560,1584]" box="[151,677,1560,1584]" pageId="26" pageNumber="479">
<treatmentCitationGroup id="13D511DAFF94B20EFF6BE85BFD2BF9AA" box="[151,677,1560,1584]" pageId="26" pageNumber="479">
<taxonomicName id="F4C54D77FF94B20EFF6BE85BFD09F9AA" ID-CoL="3GJ3Y" authority="Pietschmann, 1913" box="[151,647,1560,1584]" family="Sisoridae" genus="Glyptosternum" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="479" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF94B20EFF6BE85BFEA0F9AA" box="[151,302,1560,1584]" italics="true" pageId="26" pageNumber="479">Glyptosternum</emphasis>
s
<emphasis id="01B1EAE6FF94B20EFEC2E85BFE30F9AA" box="[318,446,1560,1584]" italics="true" pageId="26" pageNumber="479">teindachneri</emphasis>
<treatmentCitation id="B26410E5FF94B20EFE38E85BFD2BF9AA" author="Pietschmann, V." box="[452,677,1560,1584]" page="93" pageId="26" pageNumber="479" year="1913">
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</paragraph>
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<subSubSection id="7BDF657FFF94B20EFF6BE81DFCAFF893" pageId="26" pageNumber="479" type="materials_examined">
<paragraph id="337A36F4FF94B20EFF6BE81DFCAFF893" blockId="26.[151,1437,1630,2016]" pageId="26" pageNumber="479">
<emphasis id="01B1EAE6FF94B20EFF6BE81DFE08F9E2" bold="true" box="[151,390,1630,1656]" pageId="26" pageNumber="479">Material examined.</emphasis>
BMNH 1968.12.13.465-70, 3,
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SL; 1968.12.13.471, 3,
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SL;
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at
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,
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(photographs only).—
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,
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,
<quantity id="F43D9B11FF94B20EFBEBE8C1FBE8F906" box="[1047,1126,1666,1692]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="26" pageNumber="479" unit="mm" value="77.0">77 mm</quantity>
SL;
<collectingCountry id="4BD27664FF94B20EFB60E8C1FB55F906" box="[1180,1243,1666,1692]" name="Syria" pageId="26" pageNumber="479">Syria</collectingCountry>
:
<collectorName id="9E305322FF94B20EFB1AE8C1FA12F906" box="[1254,1436,1666,1692]" pageId="26" pageNumber="479">Nahr Al-Khabur</collectorName>
about
<quantity id="F43D9B11FF94B20EFF27E8E5FEAFF95A" box="[219,289,1702,1728]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="479" unit="km" value="16.0">16 km</quantity>
southeast of
<collectingRegion id="F101F816FF94B20EFE4EE8E5FDABF95A" box="[434,549,1702,1728]" country="Syria" name="Al Hasakah" pageId="26" pageNumber="479">al-Hasaka</collectingRegion>
, 36.43 40.87.—
<specimenCode id="63639E8FFF94B20EFD28E8E5FCD2F95A" box="[724,860,1702,1728]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="479">SMF 28644</specimenCode>
,
<specimenCode id="63639E8FFF94B20EFC9AE8E5FCFBF95A" box="[870,885,1702,1728]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="479">3</specimenCode>
,
<quantity id="F43D9B11FF94B20EFC7DE8E5FB9DF95A" box="[897,1043,1702,1728]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" metricValueMax="1.3" metricValueMin="1.1199999999999999" pageId="26" pageNumber="479" unit="mm" value="121.0" valueMax="130.0" valueMin="112.0">112130 mm</quantity>
SL;
<collectingCountry id="4BD27664FF94B20EFBBBE8E5FB08F95A" box="[1095,1158,1702,1728]" name="Syria" pageId="26" pageNumber="479">Syria</collectingCountry>
:
<location id="361A602FFF94B20EFB6CE8E5FA8EF95A" LSID="urn:lsid:plazi:treatment:BB6C87E2FF94B209FF6BEBECFC97FD83:361A602FFF94B20EFB6CE8E5FA8EF95A" box="[1168,1280,1702,1728]" country="Syria" name="Euphrates" pageId="26" pageNumber="479" stateProvince="Al Hasakah">Euphrates</location>
at Der ez Zor, 35.52 39.91.—
<specimenCode id="63639E8FFF94B20EFEC1E889FE49F97E" box="[317,455,1738,1764]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="479">SMF 28694</specimenCode>
,
<specimenCode id="63639E8FFF94B20EFE2EE889FE6CF97E" box="[466,482,1738,1764]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="479">4</specimenCode>
,
<quantity id="F43D9B11FF94B20EFE12E889FD0DF97E" box="[494,643,1738,1765]" metricMagnitude="-1" metricUnit="m" metricValue="1.51" metricValueMax="1.56" metricValueMin="1.46" pageId="26" pageNumber="479" unit="mm" value="151.0" valueMax="156.0" valueMin="146.0">146156 mm</quantity>
SL;
<collectingCountry id="4BD27664FF94B20EFD45E889FD76F97E" box="[697,760,1738,1764]" name="Syria" pageId="26" pageNumber="479">Syria</collectingCountry>
:
<collectorName id="9E305322FF94B20EFCFFE889FCFDF97E" box="[771,883,1738,1764]" pageId="26" pageNumber="479">Euphrates</collectorName>
at
<collectingRegion id="F101F816FF94B20EFC69E889FC51F97E" box="[917,991,1738,1764]" country="Syria" name="Ar Raqqah" pageId="26" pageNumber="479">Raqqa</collectingRegion>
, 35.93 38.99.—
<specimenCode id="63639E8FFF94B20EFB6DE889FA9DF97E" box="[1169,1299,1738,1764]" collectionCode="ZMH" pageId="26" pageNumber="479">ZMH 4431</specimenCode>
,
<specimenCode id="63639E8FFF94B20EFAE2E889FAA3F97E" box="[1310,1325,1738,1764]" collectionCode="ZMH" pageId="26" pageNumber="479">3</specimenCode>
,
<quantity id="F43D9B11FF94B20EFAC6E889FF4DF892" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.38" metricValueMin="1.0699999999999998" pageId="26" pageNumber="479" unit="mm" value="122.5" valueMax="138.0" valueMin="107.0">107138 mm</quantity>
SL;
<collectingCountry id="4BD27664FF94B20EFF05E8ADFEC3F892" box="[249,333,1774,1800]" name="Turkey" pageId="26" pageNumber="479">Turkey</collectingCountry>
:
<collectorName id="9E305322FF94B20EFEABE8ADFE49F892" box="[343,455,1774,1800]" pageId="26" pageNumber="479">Euphrates</collectorName>
at
<collectorName id="9E305322FF94B20EFE16E8ADFDDCF892" box="[490,594,1774,1800]" pageId="26" pageNumber="479">Kemalye</collectorName>
,
<geoCoordinate id="56F15033FF94B20EFDA1E8ADFD36F893" box="[605,696,1774,1801]" degrees="39.2625" direction="north" orientation="latitude" pageId="26" pageNumber="479" precision="5" value="39.2625">39.2625</geoCoordinate>
<geoCoordinate id="56F15033FF94B20EFD43E8ADFC90F893" box="[703,798,1774,1801]" degrees="38.5019" direction="east" orientation="longitude" pageId="26" pageNumber="479" precision="5" value="38.5019">38.5019</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF94B20EFF3BE951FC93F8B6" box="[199,797,1810,1836]" pageId="26" pageNumber="479" type="multiple">
<paragraph id="337A36F4FF94B20EFF3BE951FC93F8B6" blockId="26.[151,1437,1630,2016]" box="[199,797,1810,1836]" pageId="26" pageNumber="479">
<emphasis id="01B1EAE6FF94B20EFF3BE951FD5FF8B6" bold="true" box="[199,721,1810,1836]" pageId="26" pageNumber="479">Material used in molecular genetic analysis.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF94B20FFF3BE975FE6AFDA7" lastPageId="27" lastPageNumber="480" pageId="26" pageNumber="479" type="diagnosis">
<paragraph id="337A36F4FF94B20EFF3BE975FF6EF87A" blockId="26.[151,1437,1630,2016]" pageId="26" pageNumber="479">
<emphasis id="01B1EAE6FF94B20EFF3BE975FECCF8CA" bold="true" box="[199,322,1846,1872]" pageId="26" pageNumber="479">Diagnosis.</emphasis>
<taxonomicName id="F4C54D77FF94B20EFEBBE975FDF7F8CA" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[327,633,1846,1872]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="479" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF94B20EFEBBE975FDF7F8CA" box="[327,633,1846,1872]" italics="true" pageId="26" pageNumber="479">Glyptothorax steindachneri</emphasis>
</taxonomicName>
is distinguished from its congeners in the Euphrates and Tigris drainages by having a very long adipose-fin, its length is 1.53.0 times (vs.
<quantity id="F43D9B11FF94B20EFC87E919FC63F8EE" box="[891,1005,1882,1908]" metricMagnitude="-2" metricUnit="m" metricValue="2.2859999999999996" metricValueMax="3.048" metricValueMin="1.524" pageId="26" pageNumber="479" unit="in" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 in</quantity>
other species) larger than the distance between the base of last dorsal-fin ray and the adipose-fin origin; the medial pit of the thoracic adhesive apparatus with striae (vs. without striae), and there are 1317 serrae on the inner margin of the ossified pectoral fin-ray (vs. 714).
</paragraph>
<paragraph id="337A36F4FF95B20FFF3BEED4FE6AFDA7" blockId="27.[151,1437,151,681]" pageId="27" pageNumber="480">
<taxonomicName id="F4C54D77FF95B20FFF3BEED4FE71FF2B" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[199,511,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFF3BEED4FE71FF2B" box="[199,511,151,177]" italics="true" pageId="27" pageNumber="480">Glyptothorax steindachneri</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="F4C54D77FF95B20FFCA2EED4FB88FF2A" box="[862,1030,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF95B20FFCA2EED4FB88FF2A" box="[862,1030,151,177]" italics="true" pageId="27" pageNumber="480">G. armeniacus</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF95B20FFBEBEED4FBFEFF2A" box="[1047,1136,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFBEBEED4FBFEFF2A" box="[1047,1136,151,177]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF95B20FFB7DEED4FAB3FF2B" box="[1153,1341,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFB7DEED4FAB3FF2B" box="[1153,1341,151,177]" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
, and
<taxonomicName id="F4C54D77FF95B20FFA7DEED4FF6EFF4F" authorityName="Coad" authorityYear="1981" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF95B20FFA7DEED4FF6EFF4F" italics="true" pageId="27" pageNumber="480">G. silviae</emphasis>
</taxonomicName>
by the caudal-peduncle depth 1.82.5 times in its length (vs.
<quantity id="F43D9B11FF95B20FFC48EEF8FBA6FF4F" box="[948,1064,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="4.064" metricValueMin="2.794" pageId="27" pageNumber="480" unit="in" value="1.35" valueMax="1.6" valueMin="1.1">1.11.6 in</quantity>
<taxonomicName id="F4C54D77FF95B20FFBCEEEF8FB63FF4F" box="[1074,1261,187,213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFBCEEEF8FB63FF4F" box="[1074,1261,187,213]" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
), having many large, elongated, bony and striated tubercles on the dorsal and lateral head and predorsal back (vs. absent in
<taxonomicName id="F4C54D77FF95B20FFA7DEE9CFE94FE86" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF95B20FFA7DEE9CFE94FE86" italics="true" pageId="27" pageNumber="480">G. armeniacus</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF95B20FFED4EF40FE6FFE87" box="[296,481,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFED4EF40FE6FFE87" box="[296,481,259,285]" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
, and
<taxonomicName id="F4C54D77FF95B20FFDE3EF40FD05FE87" authorityName="Coad" authorityYear="1981" box="[543,651,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF95B20FFDE3EF40FD05FE87" box="[543,651,259,285]" italics="true" pageId="27" pageNumber="480">G. silviae</emphasis>
</taxonomicName>
), a slightly elevated thoracic adhesive apparatus (strongly elevated in
<taxonomicName id="F4C54D77FF95B20FFF6BEF64FEB2FEDA" box="[151,316,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF95B20FFF6BEF64FEB2FEDA" box="[151,316,295,321]" italics="true" pageId="27" pageNumber="480">G. armeniacus</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF95B20FFE8DEF64FE52FEDB" authorityName="Coad" authorityYear="1981" box="[369,476,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF95B20FFE8DEF64FE52FEDB" box="[369,476,295,321]" italics="true" pageId="27" pageNumber="480">G. silviae</emphasis>
</taxonomicName>
), 1.1
<emphasis id="01B1EAE6FF95B20FFDEAEF64FDAAFEDA" box="[534,548,295,320]" italics="true" pageId="27" pageNumber="480"></emphasis>
1.5 times longer than wide (vs.
<quantity id="F43D9B11FF95B20FFC7FEF64FC7AFEDB" box="[899,1012,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="27" pageNumber="480" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">
0.8
<emphasis id="01B1EAE6FF95B20FFC5AEF64FC3AFEDA" box="[934,948,295,320]" italics="true" pageId="27" pageNumber="480"></emphasis>
1.1 in
</quantity>
<taxonomicName id="F4C54D77FF95B20FFC06EF64FBC1FEDA" box="[1018,1103,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFC06EF64FBC1FEDA" box="[1018,1103,295,321]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
,
<quantity id="F43D9B11FF95B20FFBA1EF64FB40FEDB" box="[1117,1230,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.7779999999999998" pageId="27" pageNumber="480" unit="in" value="0.8" valueMax="0.9" valueMin="0.7">
0.7
<emphasis id="01B1EAE6FF95B20FFB7CEF64FB00FEDA" box="[1152,1166,295,320]" italics="true" pageId="27" pageNumber="480"></emphasis>
0.9 in
</quantity>
<taxonomicName id="F4C54D77FF95B20FFB28EF64FA02FEDB" box="[1236,1420,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFB28EF64FA02FEDB" box="[1236,1420,295,321]" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
), extending from the isthmus to the base of the last pectoral-fin ray or to the posterior limit of the pectoral-fin base (vs. to base of first or third branched pectoral-fin ray in
<taxonomicName id="F4C54D77FF95B20FFCECEF2CFC47FE13" box="[784,969,367,393]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFCECEF2CFC47FE13" box="[784,969,367,393]" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
). The thoracic adhesive apparatus is well delineated at its posterior margin (poorly delineated in
<taxonomicName id="F4C54D77FF95B20FFCE7EFD0FCFAFE36" box="[795,884,403,429]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFCE7EFD0FCFAFE36" box="[795,884,403,429]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
), completely situated on the horse-shoe shaped swelling (vs. extending beyond the swelling, usually onto the pectoral-fin base in
<taxonomicName id="F4C54D77FF95B20FFBD1EFF4FB0CFE4A" box="[1069,1154,439,465]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFBD1EFF4FB0CFE4A" box="[1069,1154,439,465]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
), the head is pointed and short, 2123% SL (vs. blunt and spade-shaped, 2428% SL in
<taxonomicName id="F4C54D77FF95B20FFC91EF98FB9AFE6E" box="[877,1044,475,501]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF95B20FFC91EF98FB9AFE6E" box="[877,1044,475,501]" italics="true" pageId="27" pageNumber="480">G. armeniacus</emphasis>
</taxonomicName>
, 2830% in
<taxonomicName id="F4C54D77FF95B20FFB56EF98FA8CFE6E" box="[1194,1282,475,501]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFB56EF98FA8CFE6E" box="[1194,1282,475,501]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
,
<quantity id="F43D9B11FF95B20FFAEEEF98FAF6FE6F" box="[1298,1400,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.62" metricValueMin="6.858" pageId="27" pageNumber="480" unit="in" value="28.5" valueMax="30.0" valueMin="27.0">2730 in</quantity>
<taxonomicName id="F4C54D77FF95B20FFA7DEF98FEA3FD83" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF95B20FFA7DEF98FEA3FD83" italics="true" pageId="27" pageNumber="480">G. kurdistanicus</emphasis>
</taxonomicName>
), and there are no or few small black spots on the flank (vs. many large blotches and/or spots in
<taxonomicName id="F4C54D77FF95B20FFA7DEFBCFF44FDA6" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFA7DEFBCFF44FDA6" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
). Size up to
<quantity id="F43D9B11FF95B20FFEA5EC60FE38FDA7" box="[345,438,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="480" unit="mm" value="190.0">190 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF95B20FFF3BEC05FE3EFD33" pageId="27" pageNumber="480" type="distribution">
<paragraph id="337A36F4FF95B20FFF3BEC05FE3EFD33" blockId="27.[151,1437,151,681]" pageId="27" pageNumber="480">
<emphasis id="01B1EAE6FF95B20FFF3BEC05FEEFFDFA" bold="true" box="[199,353,582,608]" pageId="27" pageNumber="480">Distribution.</emphasis>
<taxonomicName id="F4C54D77FF95B20FFE95EC04FD11FDFB" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[361,671,583,609]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFE95EC04FD11FDFB" box="[361,671,583,609]" italics="true" pageId="27" pageNumber="480">Glyptothorax steindachneri</emphasis>
</taxonomicName>
seems to be widespread in the Euphrates and Tigris. It is recorded from the Tigris at Mossul, the Euphrates in
<collectingCountry id="4BD27664FF95B20FFD87EC28FD37FD1F" box="[635,697,619,645]" name="Syria" pageId="27" pageNumber="480">Syria</collectingCountry>
, the Nahr Al-Khabur, a tributary of the Euphrates in
<collectingCountry id="4BD27664FF95B20FFAFEEC28FACEFD1F" box="[1282,1344,619,645]" name="Syria" pageId="27" pageNumber="480">Syria</collectingCountry>
, and the Lesser Zab River in
<collectingCountry id="4BD27664FF95B20FFE80ECCCFE23FD33" box="[380,429,655,681]" name="Iraq" pageId="27" pageNumber="480">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="67BA667CFF95B20FFF6BEB99FC7DF98C" ID-DOI="http://doi.org/10.5281/zenodo.14895407" ID-Zenodo-Dep="14895407" httpUri="https://zenodo.org/record/14895407/files/figure.png" pageId="27" pageNumber="480" startId="27.[151,250,1498,1522]" targetBox="[151,1436,705,1475]" targetPageId="27" targetType="figure">
<paragraph id="337A36F4FF95B20FFF6BEB99FC7DF98C" blockId="27.[151,1436,1498,1558]" pageId="27" pageNumber="480">
<emphasis id="01B1EAE6FF95B20FFF6BEB99FEAEFA68" bold="true" box="[151,288,1498,1522]" pageId="27" pageNumber="480">FIGURE 28.</emphasis>
<taxonomicName id="F4C54D77FF95B20FFED9EB99FDB2FA68" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[293,572,1498,1522]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFED9EB99FDB2FA68" box="[293,572,1498,1522]" italics="true" pageId="27" pageNumber="480">Glyptothorax steindachneri</emphasis>
</taxonomicName>
, from top: SMF 23678, 77 mm SL, Syria: Nahr Al-Khabur; SMF 28644, 112 mm SL, Syria: Der ez Zor; SMF 28694, 155 mm SL; 158 mm SL, Syria: Euphrates at Raqqa.
</paragraph>
</caption>
<subSubSection id="7BDF657FFF95B209FF3BE807FC97FD83" lastPageId="29" lastPageNumber="482" pageId="27" pageNumber="480" type="discussion">
<paragraph id="337A36F4FF95B208FF3BE807FD5CFEDB" blockId="27.[151,1437,1604,2027]" lastBlockId="28.[151,1437,151,321]" lastPageId="28" lastPageNumber="481" pageId="27" pageNumber="480">
<emphasis id="01B1EAE6FF95B20FFF3BE807FEB5F9C4" bold="true" box="[199,315,1604,1630]" pageId="27" pageNumber="480">Remarks.</emphasis>
The description by
<bibRefCitation id="57544B05FF95B20FFDDDE807FC8AF9C4" author="Pietschmann, V." box="[545,772,1604,1630]" pageId="27" pageNumber="480" pagination="93 - 95" refId="ref19664" refString="Pietschmann, V. (1913) Eine neue Glyptosternum - Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93-95." type="journal article" year="1913">Pietschmann (1913)</bibRefCitation>
is informative and gives some very useful characters. But no figures are provided and the
<typeStatus id="EC7E8856FF95B20FFDF0E82AFDE0F919" box="[524,622,1641,1667]" pageId="27" pageNumber="480" type="syntype">syntypes</typeStatus>
are missing in NMW, where hundreds of jars with Pietschmanns fish collection are still await sorting. Anja Palandacic (pers. comm., 2020) looked through the jars at our request but did not find the types of
<taxonomicName id="F4C54D77FF95B20FFE55E8F2FDEDF950" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[425,611,1712,1739]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFE55E8F2FDEDF950" box="[425,611,1712,1739]" italics="true" pageId="27" pageNumber="480">G. steindachneri</emphasis>
</taxonomicName>
. There is one jar (NMW) with three
<taxonomicName id="F4C54D77FF95B20FFC01E8F3FB1CF950" box="[1021,1170,1712,1738]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF95B20FFC01E8F3FB1CF950" box="[1021,1170,1712,1738]" italics="true" pageId="27" pageNumber="480">Glyptothorax</emphasis>
</taxonomicName>
specimens collected by Pietschmann at an unknown date (NMW 94378). They are labelled as coming from “
<collectingCountry id="4BD27664FF95B20FFBB0E897FB3DF974" box="[1100,1203,1748,1774]" name="Armenia" pageId="27" pageNumber="480">Armenia</collectingCountry>
” without any further information. These
<specimenCount id="25C3FD7DFF95B20FFE8AE8BBFDA5F888" box="[374,555,1784,1810]" count="3" pageId="27" pageNumber="480" type="generic">three specimens</specimenCount>
measure 122, 141 and
<quantity id="F43D9B11FF95B20FFCCAE8BBFC1AF889" box="[822,916,1784,1811]" metricMagnitude="-1" metricUnit="m" metricValue="2.19" pageId="27" pageNumber="480" unit="mm" value="219.0">219 mm</quantity>
in total length (
<quantity id="F43D9B11FF95B20FFBB0E8BBFA81F889" box="[1100,1295,1784,1811]" metricMagnitude="4" metricUnit="m" metricValue="8.9115185" pageId="27" pageNumber="480" unit="mm" value="8.9115185E7">89, 115, 185 mm</quantity>
SL) and are identified as
<taxonomicName id="F4C54D77FF95B20FFEDAE95EFEF5F8AC" box="[294,379,1821,1847]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF95B20FFEDAE95EFEF5F8AC" box="[294,379,1821,1847]" italics="true" pageId="27" pageNumber="480">G. cous</emphasis>
</taxonomicName>
. They do not agree with the character states given by
<bibRefCitation id="57544B05FF95B20FFC32E95FFB20F8AC" author="Pietschmann, V." box="[974,1198,1820,1846]" pageId="27" pageNumber="480" pagination="93 - 95" refId="ref19664" refString="Pietschmann, V. (1913) Eine neue Glyptosternum - Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93-95." type="journal article" year="1913">Pietschmann (1913)</bibRefCitation>
for
<taxonomicName id="F4C54D77FF95B20FFB27E95EFA1BF8AC" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1243,1429,1820,1847]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFB27E95EFA1BF8AC" box="[1243,1429,1820,1847]" italics="true" pageId="27" pageNumber="480">G. steindachneri</emphasis>
</taxonomicName>
. Pietschmann described
<taxonomicName id="F4C54D77FF95B20FFE5AE902FDEAF8C0" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[422,612,1856,1883]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFE5AE902FDEAF8C0" box="[422,612,1856,1883]" italics="true" pageId="27" pageNumber="480">G. steindachneri</emphasis>
</taxonomicName>
based on
<specimenCount id="25C3FD7DFF95B20FFD1CE902FCF8F8C1" box="[736,886,1857,1883]" count="2" pageId="27" pageNumber="480" type="generic" typeStatus="syntype">two syntypes</specimenCount>
of 150 and
<quantity id="F43D9B11FF95B20FFBF5E903FBE4F8C1" box="[1033,1130,1856,1883]" metricMagnitude="-1" metricUnit="m" metricValue="1.89" pageId="27" pageNumber="480" unit="mm" value="189.0">189 mm</quantity>
total length and it is very evident that NMW 94378 does not correspond to the types of
<taxonomicName id="F4C54D77FF95B20FFCA4E926FB9AF8E4" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[856,1044,1892,1919]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B20FFCA4E926FB9AF8E4" box="[856,1044,1892,1919]" italics="true" pageId="27" pageNumber="480">G. steindachneri</emphasis>
</taxonomicName>
. At Pietschmanns times,
<collectingCountry id="4BD27664FF95B20FFAC6E927FA12F8E4" box="[1338,1436,1892,1918]" name="Armenia" pageId="27" pageNumber="480">Armenia</collectingCountry>
was much larger than today and covered considerable parts of the upper Euphrates and Tigris drainage and we have no doubt that NMW 94378 was collected from that region. These fish make clear that Pietschmann did collect other
<taxonomicName id="F4C54D77FF95B20FFF26E993FEE1F870" box="[218,367,2000,2026]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF95B20FFF26E993FEE1F870" box="[218,367,2000,2026]" italics="true" pageId="27" pageNumber="480">Glyptothorax</emphasis>
</taxonomicName>
in the Euphrates and has compared different species that he had in hand when describing
<taxonomicName id="F4C54D77FF95B208FA7DE992FEA1FF2B" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lastPageId="28" lastPageNumber="481" order="Siluriformes" pageId="27" pageNumber="480" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF95B208FA7DE992FEA1FF2B" italics="true" lastPageId="28" lastPageNumber="481" pageId="27" pageNumber="480">G. steindachneri</emphasis>
</taxonomicName>
.
<bibRefCitation id="57544B05FF92B208FEC3EED4FE43FF2B" author="Coad, B. W." box="[319,461,151,177]" pageId="28" pageNumber="481" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
shortly mentioned that
<taxonomicName id="F4C54D77FF92B208FD21EED4FC17FF2B" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[733,921,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FD21EED4FC17FF2B" box="[733,921,151,177]" italics="true" pageId="28" pageNumber="481">G. steindachneri</emphasis>
</taxonomicName>
“may be a synonym of
<taxonomicName id="F4C54D77FF92B208FB51EED4FA8AFF2A" box="[1197,1284,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF92B208FB51EED4FA8AFF2A" box="[1197,1284,151,177]" italics="true" pageId="28" pageNumber="481">G. cous</emphasis>
</taxonomicName>
”. Indeed, we identify BMNH 1968 12.13.470-471, identified by
<bibRefCitation id="57544B05FF92B208FD1DEEF8FCE1FF4F" author="Coad, B. W." box="[737,879,187,213]" pageId="28" pageNumber="481" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
as
<taxonomicName id="F4C54D77FF92B208FC64EEF8FC61FF4E" box="[920,1007,187,213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF92B208FC64EEF8FC61FF4E" box="[920,1007,187,213]" italics="true" pageId="28" pageNumber="481">G. cous</emphasis>
</taxonomicName>
, as
<taxonomicName id="F4C54D77FF92B208FBDCEEF8FB52FF4F" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1056,1244,187,213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FBDCEEF8FB52FF4F" box="[1056,1244,187,213]" italics="true" pageId="28" pageNumber="481">G. steindachneri</emphasis>
</taxonomicName>
. But ZMH 4431 identified as
<taxonomicName id="F4C54D77FF92B208FED8EE9CFE46FF62" box="[292,456,223,249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF92B208FED8EE9CFE46FF62" box="[292,456,223,249]" italics="true" pageId="28" pageNumber="481">G. armeniacus</emphasis>
</taxonomicName>
by
<bibRefCitation id="57544B05FF92B208FE12EE9CFDF6FF63" author="Coad, B. W." box="[494,632,223,249]" pageId="28" pageNumber="481" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
are also identified as
<taxonomicName id="F4C54D77FF92B208FC99EE9CFB93FF63" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[869,1053,223,249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FC99EE9CFB93FF63" box="[869,1053,223,249]" italics="true" pageId="28" pageNumber="481">G. steindachneri</emphasis>
</taxonomicName>
by us.
<taxonomicName id="F4C54D77FF92B208FB95EE9CFA12FF63" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1129,1436,223,249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FB95EE9CFA12FF63" box="[1129,1436,223,249]" italics="true" pageId="28" pageNumber="481">Glyptothorax steindachneri</emphasis>
</taxonomicName>
is continuously listed for the fauna of
<collectingCountry id="4BD27664FF92B208FDBCEF40FDE3FE87" box="[576,621,259,285]" name="Iraq" pageId="28" pageNumber="481">Iraq</collectingCountry>
(for example
<bibRefCitation id="57544B05FF92B208FCF7EF40FC09FE87" author="Coad, B. W." box="[779,903,259,285]" pageId="28" pageNumber="481" pagination="1 - 16" refId="ref17777" refString="Coad, B. W. (2010) Freshwater fishes of Iraq. Sofia and Moscow (Pensoft): 275 pp., Pls. 1-16." type="book chapter" year="2010">Coad 2010</bibRefCitation>
) but no positive record of
<taxonomicName id="F4C54D77FF92B208FB53EF40FAE7FE87" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1199,1385,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FB53EF40FAE7FE87" box="[1199,1385,259,285]" italics="true" pageId="28" pageNumber="481">G. steindachneri</emphasis>
</taxonomicName>
was published and we are the first to clarify its identity.
</paragraph>
<caption id="67BA667CFF92B208FF6BEA52FC7CFBD7" ID-DOI="http://doi.org/10.5281/zenodo.14895409" ID-Zenodo-Dep="14895409" httpUri="https://zenodo.org/record/14895409/files/figure.png" pageId="28" pageNumber="481" startId="28.[151,250,1041,1065]" targetBox="[151,1436,345,1018]" targetPageId="28" targetType="figure">
<paragraph id="337A36F4FF92B208FF6BEA52FC7CFBD7" blockId="28.[151,1436,1041,1102]" pageId="28" pageNumber="481">
<emphasis id="01B1EAE6FF92B208FF6BEA52FE91FBB0" bold="true" box="[151,287,1041,1066]" pageId="28" pageNumber="481">FIGURE 29.</emphasis>
<taxonomicName id="F4C54D77FF92B208FED9EA51FDB2FBB0" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[293,572,1042,1066]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FED9EA51FDB2FBB0" box="[293,572,1042,1066]" italics="true" pageId="28" pageNumber="481">Glyptothorax steindachneri</emphasis>
</taxonomicName>
, from left: SMF 23678, 77 mm SL, Syria: Nahr Al-Khabur; SMF 28644, 112 mm SL, 130 mm SL, Syria: Der ez Zor; SMF 28694, 158 mm SL, Syria: Euphrates at Raqqa.
</paragraph>
</caption>
<caption id="67BA667CFF92B208FF6BE804FA00F9C5" ID-DOI="http://doi.org/10.5281/zenodo.14895411" ID-Zenodo-Dep="14895411" box="[151,1422,1607,1631]" httpUri="https://zenodo.org/record/14895411/files/figure.png" pageId="28" pageNumber="481" startId="28.[151,250,1607,1631]" targetBox="[159,1436,1185,1551]" targetPageId="28" targetType="figure">
<paragraph id="337A36F4FF92B208FF6BE804FA00F9C5" blockId="28.[151,1422,1607,1631]" box="[151,1422,1607,1631]" pageId="28" pageNumber="481">
<emphasis id="01B1EAE6FF92B208FF6BE804FEAEF9C5" bold="true" box="[151,288,1607,1631]" pageId="28" pageNumber="481">FIGURE 30.</emphasis>
<taxonomicName id="F4C54D77FF92B208FEDAE804FDB3F9C5" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[294,573,1607,1631]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FEDAE804FDB3F9C5" box="[294,573,1607,1631]" italics="true" pageId="28" pageNumber="481">Glyptothorax steindachneri</emphasis>
</taxonomicName>
, not preserved, about 115 mm SL; Iraq: Qalah Chwalan River, 35.9150, 45.36788.
</paragraph>
</caption>
<paragraph id="337A36F4FF92B208FF3BE8CEFD81F96A" blockId="28.[151,1436,1677,2028]" pageId="28" pageNumber="481">
Indeed,
<taxonomicName id="F4C54D77FF92B208FEDDE8CDFE54F93D" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[289,474,1677,1704]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF92B208FEDDE8CDFE54F93D" box="[289,474,1677,1704]" italics="true" pageId="28" pageNumber="481">G. steindachneri</emphasis>
</taxonomicName>
seems to be widespread in the Euphrates and Tigris drainage and we expect it to occur in all the countries in the catchment region. It might be continuously misidentified as
<taxonomicName id="F4C54D77FF92B208FB85E8F1FB5EF951" box="[1145,1232,1714,1740]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF92B208FB85E8F1FB5EF951" box="[1145,1232,1714,1740]" italics="true" pageId="28" pageNumber="481">G. cous</emphasis>
</taxonomicName>
, which is usually misidentified as
<taxonomicName id="F4C54D77FF92B208FEACE895FD86F975" box="[336,520,1749,1776]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="481" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF92B208FEACE895FD86F975" box="[336,520,1749,1776]" italics="true" pageId="28" pageNumber="481">G. kurdistanicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="337A36F4FF92B208FF3BE8BAFE7CF871" blockId="28.[151,1436,1677,2028]" pageId="28" pageNumber="481">
The most important characters given by
<bibRefCitation id="57544B05FF92B208FD79E8BAFCEBF889" author="Pietschmann, V." box="[645,869,1785,1811]" pageId="28" pageNumber="481" pagination="93 - 95" refId="ref19664" refString="Pietschmann, V. (1913) Eine neue Glyptosternum - Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93-95." type="journal article" year="1913">Pietschmann (1913)</bibRefCitation>
are: “Die Oberseite des Kopfes ist mit zahlreichen, länglichen, unregelmässigen Hautwarzen versehen, die Fettflosse sehr langgestreckt, ihre Basislänge grösser als die Entfernung ihres Ursprunges vom Ende der Dorsalbasis. Die Pektorale besitzt einen sehr starken, breiten, an seinem hinteren Rande mit ungefähr 17 bis 18 Dornen versehenen Stachel.“ [The upper side of the head has numerous, longish, irregular skin warts, the adipose fin is much elongated, its base being longer than the distance from its origin to the end of the dorsal-fin base. The pectoral fin possesses a strong, broad spine supplied with about 1718 thorns on its posterior margin.]
</paragraph>
<paragraph id="337A36F4FF93B209FF3BEED4FD9AFE37" blockId="29.[151,1437,151,537]" pageId="29" pageNumber="482">
Catfish identified here as
<taxonomicName id="F4C54D77FF93B209FE1FEED4FD12FF2B" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[483,668,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FE1FEED4FD12FF2B" box="[483,668,151,177]" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
have striated, elongated tubercles on the head, which might be called skin-warts (
<figureCitation id="ABFE2A71FF93B209FEE0EEF8FEE0FF4F" box="[284,366,187,213]" captionStart="FIGURE 29" captionStartId="28.[151,250,1041,1065]" captionTargetBox="[151,1436,345,1018]" captionTargetId="figure-108@28.[151,1436,345,1018]" captionTargetPageId="28" captionText="FIGURE 29. Glyptothorax steindachneri, from left: SMF 23678, 77 mm SL, Syria: Nahr Al-Khabur; SMF 28644, 112 mm SL, 130 mm SL, Syria: Der ez Zor; SMF 28694, 158 mm SL, Syria: Euphrates at Raqqa." figureDoi="http://doi.org/10.5281/zenodo.14895409" httpUri="https://zenodo.org/record/14895409/files/figure.png" pageId="29" pageNumber="482">Fig. 29</figureCitation>
) and this character state is only shared with
<taxonomicName id="F4C54D77FF93B209FC94EEF8FC30FF4E" box="[872,958,187,213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF93B209FC94EEF8FC30FF4E" box="[872,958,187,213]" italics="true" pageId="29" pageNumber="482">G. cous</emphasis>
</taxonomicName>
. Only in
<taxonomicName id="F4C54D77FF93B209FBD7EEF8FB68FF4F" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1067,1254,187,213]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FBD7EEF8FB68FF4F" box="[1067,1254,187,213]" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
, the adipose fin base is much longer than the distance between the adipose-fin origin and the end of the dorsal-fin base (adipose-fin length 1.53.0 times larger than the distance between base of last dorsal-fin ray and adipose-fin origin in
<taxonomicName id="F4C54D77FF93B209FA7DEF40FEA1FEDB" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FA7DEF40FEA1FEDB" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
vs.
<quantity id="F43D9B11FF93B209FEA2EF64FE41FEDB" box="[350,463,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.2859999999999996" metricValueMax="3.048" metricValueMin="1.524" pageId="29" pageNumber="482" unit="in" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 in</quantity>
all other species,
<quantity id="F43D9B11FF93B209FD65EF64FC84FEDB" box="[665,778,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.2859999999999996" metricValueMax="3.048" metricValueMin="1.524" pageId="29" pageNumber="482" unit="in" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 in</quantity>
<taxonomicName id="F4C54D77FF93B209FCEDEF64FCE9FEDA" box="[785,871,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF93B209FCEDEF64FCE9FEDA" box="[785,871,295,321]" italics="true" pageId="29" pageNumber="482">G. cous</emphasis>
</taxonomicName>
). In addition,
<taxonomicName id="F4C54D77FF93B209FBFBEF64FB4FFEDB" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1031,1217,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FBFBEF64FB4FFEDB" box="[1031,1217,295,321]" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
has more serrae on the pectoral spine than all other
<taxonomicName id="F4C54D77FF93B209FE01EF08FD1CFEFF" box="[509,658,331,357]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF93B209FE01EF08FD1CFEFF" box="[509,658,331,357]" italics="true" pageId="29" pageNumber="482">Glyptothorax</emphasis>
</taxonomicName>
examined for this study (1317 vs.
<quantity id="F43D9B11FF93B209FBD8EF08FBF6FEFF" box="[1060,1144,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.556" metricValueMin="1.778" pageId="29" pageNumber="482" unit="in" value="10.5" valueMax="14.0" valueMin="7.0">714 in</quantity>
other species). Therefore, we are very confident that our materials are correctly identified as
<taxonomicName id="F4C54D77FF93B209FC7FEF2CFBB3FE13" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[899,1085,367,393]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FC7FEF2CFBB3FE13" box="[899,1085,367,393]" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
despite the absence of
<typeStatus id="EC7E8856FF93B209FABFEF2CFAF0FE13" box="[1347,1406,367,393]" pageId="29" pageNumber="482">types</typeStatus>
or figures in the original description.
</paragraph>
<paragraph id="337A36F4FF93B209FF3BEFF4FC97FD83" blockId="29.[151,1437,151,537]" pageId="29" pageNumber="482">
The tubercles on the head are absent or very shallow and without surface structure in the three individuals of
<taxonomicName id="F4C54D77FF93B209FA7DEFF4FEA1FE6F" baseAuthorityName="Pietschmann" baseAuthorityYear="1913" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF93B209FA7DEFF4FEA1FE6F" italics="true" pageId="29" pageNumber="482">G. steindachneri</emphasis>
</taxonomicName>
from
<collectingCountry id="4BD27664FF93B209FE8FEF98FE4DFE6F" box="[371,451,475,501]" name="Turkey" pageId="29" pageNumber="482">Turkey</collectingCountry>
(ZMH 4431). As discussed under
<taxonomicName id="F4C54D77FF93B209FCBFEF98FC19FE6E" box="[835,919,475,501]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="482" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF93B209FCBFEF98FC19FE6E" box="[835,919,475,501]" italics="true" pageId="29" pageNumber="482">G. cous</emphasis>
</taxonomicName>
above, we expect that this skin ornamentation has been scraped-off by careless handling of these fishes.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,731 @@
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<mods:title id="1F2E425C943E13F7900537F1F5408D95">The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae)</mods:title>
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<subSubSection id="7BDF657FFF9DB207FF6BEBA0FD0DFA64" box="[151,643,1507,1534]" pageId="19" pageNumber="472" type="nomenclature">
<paragraph id="337A36F4FF9DB207FF6BEBA0FD0DFA64" blockId="19.[151,643,1507,1570]" box="[151,643,1507,1534]" pageId="19" pageNumber="472">
<heading id="68328198FF9DB207FF6BEBA0FD0DFA64" bold="true" box="[151,643,1507,1534]" fontSize="11" level="1" pageId="19" pageNumber="472" reason="1">
<taxonomicName id="F4C54D77FF9DB207FF6BEBA0FD0DFA64" ID-CoL="3GJD7" authority="(Berg, 1931)" baseAuthorityName="Berg" baseAuthorityYear="1931" box="[151,643,1507,1534]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="472" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF9DB207FF6BEBA0FD0DFA64" bold="true" box="[151,643,1507,1534]" pageId="19" pageNumber="472">
<emphasis id="01B1EAE6FF9DB207FF6BEBA0FE6DFA64" bold="true" box="[151,483,1507,1534]" italics="true" pageId="19" pageNumber="472">Glyptothorax kurdistanicus</emphasis>
(
<bibRefCitation id="57544B05FF9DB207FE0EEBA7FDF5FA64" author="Berg, L. S." box="[498,635,1508,1534]" pageId="19" pageNumber="472" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg, 1931</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF9DB207FF6BE84BFEADF9B8" box="[151,291,1544,1570]" pageId="19" pageNumber="472" type="description">
<paragraph id="337A36F4FF9DB207FF6BE84BFEADF9B8" blockId="19.[151,643,1507,1570]" box="[151,291,1544,1570]" pageId="19" pageNumber="472">
(
<figureCitation id="ABFE2A71FF9DB207FF63E84BFE92F9B8" box="[159,284,1544,1570]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStart-3="FIGURE 21" captionStart-4="FIGURE 22" captionStart-5="FIGURE 23" captionStart-6="FIGURE 24" captionStartId-0="20.[151,250,1664,1688]" captionStartId-1="21.[151,250,1116,1140]" captionStartId-2="21.[151,250,1981,2005]" captionStartId-3="22.[151,250,1051,1075]" captionStartId-4="22.[151,250,1909,1933]" captionStartId-5="23.[151,250,485,509]" captionStartId-6="23.[151,250,1496,1520]" captionTargetBox-0="[151,1436,181,1640]" captionTargetBox-1="[164,1424,561,1093]" captionTargetBox-2="[150,1438,1200,1957]" captionTargetBox-3="[151,1436,181,1027]" captionTargetBox-4="[150,1438,1135,1885]" captionTargetBox-5="[189,1398,181,461]" captionTargetBox-6="[151,1436,570,1473]" captionTargetId-0="figure-23@20.[151,1436,181,1640]" captionTargetId-1="figure-210@21.[164,1424,561,1093]" captionTargetId-2="figure-235@21.[150,1438,1200,1957]" captionTargetId-3="figure-23@22.[151,1436,181,1027]" captionTargetId-4="figure-52@22.[150,1438,1135,1885]" captionTargetId-5="figure-19@23.[189,1398,181,461]" captionTargetId-6="figure-49@23.[151,1436,570,1473]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="22" captionTargetPageId-5="23" captionTargetPageId-6="23" captionText-0="FIGURE 18. Glyptothorax kurdistanicus, ZIN 20780, type, 113 mm SL; Iran: Germau (or Germav). Drawings modified from Berg (1931)." captionText-1="FIGURE 19. Glyptothorax kurdistanicus, juveniles, from top: FSJF 3652, top: 77 mm SL, 69 mm SL; Iraq: Aw-e Shiler River." captionText-2="FIGURE 20. Glyptothorax kurdistanicus, adults, from top: FFR 3916, from top: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki." captionText-3="FIGURE 21. Glyptothorax kurdistanicus, juveniles, from left: FSJF 3652, from left: 66 mm SL, 69 mm SL, 77 mm SL; Iraq: Aw-e Shiler River." captionText-4="FIGURE 22. Glyptothorax kurdistanicus, adults, from top: FFR 3916, from left: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki." captionText-5="FIGURE 23. Glyptothorax kurdistanicus, skin ornamentation; from left: adults, FFR 3916, from left: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki." captionText-6="FIGURE 24. Glyptothorax kurdistanicus, from top: FSJF 3652, 77 mm SL; Iraq:Aw-e Shiler River; FSJF-DNA 3349; 120 mm SL; Iraq: Qalah Chwalan River." figureDoi-0="http://doi.org/10.5281/zenodo.14895387" figureDoi-1="http://doi.org/10.5281/zenodo.14895389" figureDoi-2="http://doi.org/10.5281/zenodo.14895391" figureDoi-3="http://doi.org/10.5281/zenodo.14895393" figureDoi-4="http://doi.org/10.5281/zenodo.14895395" figureDoi-5="http://doi.org/10.5281/zenodo.14895397" figureDoi-6="http://doi.org/10.5281/zenodo.14895399" httpUri-0="https://zenodo.org/record/14895387/files/figure.png" httpUri-1="https://zenodo.org/record/14895389/files/figure.png" httpUri-2="https://zenodo.org/record/14895391/files/figure.png" httpUri-3="https://zenodo.org/record/14895393/files/figure.png" httpUri-4="https://zenodo.org/record/14895395/files/figure.png" httpUri-5="https://zenodo.org/record/14895397/files/figure.png" httpUri-6="https://zenodo.org/record/14895399/files/figure.png" pageId="19" pageNumber="472">Fig. 1824</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF9DB207FF6BE80FFDFAF9FE" box="[151,628,1612,1637]" pageId="19" pageNumber="472" type="reference_group">
<paragraph id="337A36F4FF9DB207FF6BE80FFDFAF9FE" blockId="19.[151,628,1612,1637]" box="[151,628,1612,1637]" pageId="19" pageNumber="472">
<treatmentCitationGroup id="13D511DAFF9DB207FF6BE80FFDFAF9FE" box="[151,628,1612,1637]" pageId="19" pageNumber="472">
<treatmentCitation id="B26410E5FF9DB207FF6BE80FFDFAF9FE" author="Berg, L. S." box="[151,628,1612,1637]" page="1267" pageId="19" pageNumber="472" year="1931">
<taxonomicName id="F4C54D77FF9DB207FF6BE80FFDFAF9FE" ID-CoL="3GJ3D" authority="Berg, 1931: 1267" authorityName="Berg" authorityPageNumber="1267" authorityYear="1931" box="[151,628,1612,1637]" family="Sisoridae" genus="Glyptosternum" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="472" phylum="Chordata" rank="species" species="kurdistanicum">
<emphasis id="01B1EAE6FF9DB207FF6BE80FFE4AF9FE" box="[151,452,1612,1636]" italics="true" pageId="19" pageNumber="472">Glyptosternum kurdistanicum</emphasis>
<bibRefCitation id="57544B05FF9DB207FE36E80EFDFAF9FE" author="Berg, L. S." box="[458,628,1612,1637]" pageId="19" pageNumber="472" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg, 1931:1267</bibRefCitation>
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</paragraph>
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<subSubSection id="7BDF657FFF9DB200FF6BE8D1FE1AF815" lastPageId="20" lastPageNumber="473" pageId="19" pageNumber="472" type="materials_examined">
<paragraph id="337A36F4FF9DB207FF6BE8D1FAECF833" blockId="19.[151,1437,1682,1961]" pageId="19" pageNumber="472">
<emphasis id="01B1EAE6FF9DB207FF6BE8D1FE0EF936" bold="true" box="[151,384,1682,1708]" pageId="19" pageNumber="472">Material examined.</emphasis>
ZIN
<quantity id="F43D9B11FF9DB207FE41E8D0FDE3F937" box="[445,621,1683,1709]" metricMagnitude="4" metricUnit="m" metricValue="2.0780113" pageId="19" pageNumber="472" unit="mm" value="2.0780113E7">20780, 113 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FD5EE8D0FD5DF937" box="[674,723,1683,1709]" name="Iran" pageId="19" pageNumber="472">Iran</collectingCountry>
:
<collectingRegion id="F101F816FF9DB207FD22E8D0FCC1F937" box="[734,847,1683,1709]" country="Iran" name="Kordestan" pageId="19" pageNumber="472">Kurdistan</collectingRegion>
, at the village
<collectingMunicipality id="D31EAC8EFF9DB207FC0EE8D0FBC2F937" box="[1010,1100,1683,1709]" pageId="19" pageNumber="472">Germau</collectingMunicipality>
(or
<location id="361A602FFF9DB207FB86E8D0FB59F937" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FB86E8D0FB59F937" box="[1146,1239,1683,1709]" country="Iran" municipality="Germau" name="Germav" pageId="19" pageNumber="472" stateProvince="Kordestan">Germav</location>
).—
<specimenCode id="63639E8FFF9DB207FB03E8D0FAF2F937" box="[1279,1404,1682,1709]" collectionCode="FSJF" pageId="19" pageNumber="472">FSJF 3652</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FA7BE8D0FA19F937" box="[1415,1431,1683,1709]" collectionCode="FSJF" pageId="19" pageNumber="472">4</specimenCode>
,
<quantity id="F43D9B11FF9DB207FF6BE8F4FE81F94B" box="[151,271,1719,1745]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" metricValueMax="7.7" metricValueMin="4.9" pageId="19" pageNumber="472" unit="mm" value="63.0" valueMax="77.0" valueMin="49.0">4977 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FEB8E8F4FEFBF94B" box="[324,373,1719,1745]" name="Iraq" pageId="19" pageNumber="472">Iraq</collectingCountry>
:
<collectingCounty id="DA1B4E78FF9DB207FE82E8F4FD8DF94B" box="[382,515,1719,1745]" pageId="19" pageNumber="472">Aw-e Shiler</collectingCounty>
at
<collectingMunicipality id="D31EAC8EFF9DB207FDDFE8F4FD07F94B" box="[547,649,1719,1745]" pageId="19" pageNumber="472">Khewata</collectingMunicipality>
,
<geoCoordinate id="56F15033FF9DB207FD68E8F5FD61F94B" box="[660,751,1718,1745]" degrees="35.7509" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="35.7509">35.7509</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FD09E8F5FCDEF94B" box="[757,848,1718,1745]" degrees="45.4797" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="45.4797">45.4797</geoCoordinate>
(photographs only).—
<specimenCode id="63639E8FFF9DB207FBB0E8F4FB4CF94B" box="[1100,1218,1719,1745]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3906</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FB30E8F4FB52F94B" box="[1228,1244,1719,1745]" collectionCode="FFR" pageId="19" pageNumber="472">2</specimenCode>
,
<quantity id="F43D9B11FF9DB207FB14E8F5FAE0F94B" box="[1256,1390,1718,1745]" metricMagnitude="-2" metricUnit="m" metricValue="9.85" metricValueMax="12.2" metricValueMin="7.5" pageId="19" pageNumber="472" unit="mm" value="98.5" valueMax="122.0" valueMin="75.0">75122 mm</quantity>
SL; FFR 3924,
<quantity id="F43D9B11FF9DB207FEE8E898FE02F96F" box="[276,396,1755,1781]" metricMagnitude="0" metricUnit="m" metricValue="1.104" pageId="19" pageNumber="472" unit="mm" value="1104.0">1, 104 mm</quantity>
SL;
<collectingRegion id="F101F816FF9DB207FE3CE898FDCEF96F" box="[448,576,1755,1781]" country="Turkey" name="Bitlis" pageId="19" pageNumber="472">Bitlis prov.</collectingRegion>
: stream Çıratan
<quantity id="F43D9B11FF9DB207FD0FE898FCA5F96F" box="[755,811,1755,1781]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="472" unit="km" value="3.0">3 km</quantity>
southwest of
<location id="361A602FFF9DB207FC39E89FFBAFF96D" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FC39E89FFBAFF96D" box="[965,1057,1756,1783]" country="Iraq" county="Aw-e Shiler" latitude="37.6717" longLatPrecision="6" longitude="43.8628" municipality="Khewata" name="Ucadim" pageId="19" pageNumber="472" stateProvince="Bitlis">Üçadım</location>
,
<geoCoordinate id="56F15033FF9DB207FBD7E899FB08F96F" box="[1067,1158,1754,1781]" degrees="38.3547" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="38.3547">38.3547</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FB70E898FB67F96F" box="[1164,1257,1755,1781]" degrees="41.7814" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="41.7814">41.7814</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FAF5E898FAF3F96F" box="[1289,1405,1755,1781]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3916</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FA7BE899FA19F96F" box="[1415,1431,1754,1781]" collectionCode="FFR" pageId="19" pageNumber="472">5</specimenCode>
,
<quantity id="F43D9B11FF9DB207FF6BE8BCFE91F883" box="[151,287,1791,1817]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" metricValueMax="1.2" metricValueMin="0.84" pageId="19" pageNumber="472" unit="mm" value="102.0" valueMax="120.0" valueMin="84.0">84120 mm</quantity>
SL;
<collectingRegion id="F101F816FF9DB207FEAAE8BCFE78F883" box="[342,502,1791,1817]" country="Turkey" name="Hakkari" pageId="19" pageNumber="472">Hakkari prov.</collectingRegion>
: stream Eziki
<quantity id="F43D9B11FF9DB207FD6AE8BCFD41F883" box="[662,719,1791,1817]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="472" unit="km" value="6.0">6 km</quantity>
northeast Konak,
<geoCoordinate id="56F15033FF9DB207FC63E8BCFC74F883" box="[927,1018,1791,1817]" degrees="37.6717" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="37.6717">37.6717</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FBFEE8BCFBD0F883" box="[1026,1118,1791,1817]" degrees="43.8628" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="43.8628">43.8628</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FB82E8BCFB7AF883" box="[1150,1268,1791,1817]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3910</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FAFCE8BCFA80F883" box="[1280,1294,1791,1817]" collectionCode="FFR" pageId="19" pageNumber="472">1</specimenCode>
,
<quantity id="F43D9B11FF9DB207FAE1E8BCFAE2F883" box="[1309,1388,1791,1817]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="19" pageNumber="472" unit="mm" value="76.0">76 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FF6BE960FF65F8A7" box="[151,235,1827,1853]" name="Turkey" pageId="19" pageNumber="472">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF9DB207FF0AE960FE3AF8A7" box="[246,436,1827,1853]" country="Turkey" name="Diyarbakir" pageId="19" pageNumber="472">Diyarbakır prov.</collectingRegion>
: stream
<collectingCounty id="DA1B4E78FF9DB207FDECE960FDC8F8A7" box="[528,582,1827,1853]" pageId="19" pageNumber="472">Kara</collectingCounty>
<quantity id="F43D9B11FF9DB207FDB2E960FD1AF8A7" box="[590,660,1827,1853]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="472" unit="km" value="11.0">11 km</quantity>
northwest of
<collectingMunicipality id="D31EAC8EFF9DB207FCCCE960FCEBF8A7" box="[816,869,1827,1853]" pageId="19" pageNumber="472">Lice</collectingMunicipality>
,
<location id="361A602FFF9DB207FC93E960FC3FF8A7" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FC93E960FC3FF8A7" box="[879,945,1827,1853]" country="Turkey" county="Kara" latitude="38.5294" longLatPrecision="6" longitude="40.5455" municipality="Lice" name="Tigris" pageId="19" pageNumber="472" stateProvince="Diyarbakir">Tigris</location>
drainage,
<geoCoordinate id="56F15033FF9DB207FBDBE961FB0CF8A7" box="[1063,1154,1826,1853]" degrees="38.5294" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="38.5294">38.5294</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FB76E961FB68F8A7" box="[1162,1254,1826,1853]" degrees="40.5455" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="40.5455">40.5455</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FAFAE961FAF2F8A7" box="[1286,1404,1826,1853]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3921</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FA7BE960FA19F8A7" box="[1415,1431,1827,1853]" collectionCode="FFR" pageId="19" pageNumber="472">2</specimenCode>
,
<quantity id="F43D9B11FF9DB207FF6BE904FE9EF8FB" box="[151,272,1863,1889]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" metricValueMax="7.8" metricValueMin="6.4" pageId="19" pageNumber="472" unit="mm" value="71.0" valueMax="78.0" valueMin="64.0">6478 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FEBAE904FE14F8FB" box="[326,410,1863,1889]" name="Turkey" pageId="19" pageNumber="472">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF9DB207FE59E904FDA8F8FB" box="[421,550,1863,1889]" country="Turkey" name="Bitlis" pageId="19" pageNumber="472">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="D31EAC8EFF9DB207FD7EE904FD40F8FB" box="[642,718,1863,1889]" pageId="19" pageNumber="472">Oraniz</collectingMunicipality>
at
<location id="361A602FFF9DB207FD0DE904FCB6F8FB" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FD0DE904FCB6F8FB" box="[753,824,1863,1889]" country="Turkey" latitude="38.1386" longLatPrecision="6" longitude="42.4303" municipality="Oraniz" name="Ekinli" pageId="19" pageNumber="472" stateProvince="Bitlis">Ekinli</location>
,
<geoCoordinate id="56F15033FF9DB207FCB8E904FC11F8FB" box="[836,927,1863,1889]" degrees="38.1386" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="38.1386">38.1386</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FC5AE904FB8CF8FB" box="[934,1026,1863,1889]" degrees="42.4303" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="42.4303">42.4303</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FBDEE904FB16F8FB" box="[1058,1176,1863,1889]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3923</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FB5FE904FB3FF8FB" box="[1187,1201,1863,1889]" collectionCode="FFR" pageId="19" pageNumber="472">1</specimenCode>
,
<quantity id="F43D9B11FF9DB207FB43E904FA81F8FB" box="[1215,1295,1863,1889]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="472" unit="mm" value="70.0">70 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FAB9E904FA17F8FB" box="[1349,1433,1863,1889]" name="Turkey" pageId="19" pageNumber="472">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF9DB207FF6BE928FE82F81F" box="[151,268,1899,1925]" country="Turkey" name="Siirt" pageId="19" pageNumber="472">Siirt prov.</collectingRegion>
: stream
<collectingMunicipality id="D31EAC8EFF9DB207FE96E928FE23F81F" box="[362,429,1899,1925]" pageId="19" pageNumber="472">Kezer</collectingMunicipality>
at
<location id="361A602FFF9DB207FE2CE928FDC6F81F" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FE2CE928FDC6F81F" box="[464,584,1899,1925]" country="Turkey" latitude="37.9558" longLatPrecision="6" longitude="41.8561" municipality="Kezer" name="Koprubasi" pageId="19" pageNumber="472" stateProvince="Siirt">Köprübaşı</location>
,
<geoCoordinate id="56F15033FF9DB207FDA8E929FD21F81F" box="[596,687,1898,1925]" degrees="37.9558" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="37.9558">37.9558</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FD4BE929FC9DF81F" box="[695,787,1898,1925]" degrees="41.8561" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="41.8561">41.8561</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FCCFE929FC27F81F" box="[819,937,1898,1925]" collectionCode="FFR" pageId="19" pageNumber="472">FFR 3929</specimenCode>
,
<specimenCode id="63639E8FFF9DB207FC49E928FC4DF81F" box="[949,963,1899,1925]" collectionCode="FFR" pageId="19" pageNumber="472">1</specimenCode>
,
<quantity id="F43D9B11FF9DB207FC2EE928FBACF81F" box="[978,1058,1899,1925]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="19" pageNumber="472" unit="mm" value="63.0">63 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FBA5E928FB23F81F" box="[1113,1197,1899,1925]" name="Turkey" pageId="19" pageNumber="472">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF9DB207FB44E928FAC9F81F" box="[1208,1351,1899,1925]" country="Turkey" name="Sirnak" pageId="19" pageNumber="472">Şırnak prov.</collectingRegion>
: stream
<collectingMunicipality id="D31EAC8EFF9DB207FF6BE9CCFF6CF833" box="[151,226,1935,1961]" pageId="19" pageNumber="472">Ortasu</collectingMunicipality>
at
<location id="361A602FFF9DB207FEF8E9CCFED2F833" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FEF8E9CCFED2F833" box="[260,348,1935,1961]" country="Turkey" latitude="37.4411" longLatPrecision="6" longitude="42.7483" municipality="Ortasu" name="Baglica" pageId="19" pageNumber="472" stateProvince="Sirnak">Bağlıca</location>
,
<geoCoordinate id="56F15033FF9DB207FE94E9CCFE4CF833" box="[360,450,1935,1961]" degrees="37.4411" direction="north" orientation="latitude" pageId="19" pageNumber="472" precision="5" value="37.4411">37.4411</geoCoordinate>
<geoCoordinate id="56F15033FF9DB207FE35E9CCFDABF833" box="[457,549,1935,1961]" degrees="42.7483" direction="east" orientation="longitude" pageId="19" pageNumber="472" precision="5" value="42.7483">42.7483</geoCoordinate>
.—
<specimenCode id="63639E8FFF9DB207FDB8E9CCFD50F833" box="[580,734,1934,1961]" collectionCode="NMW" country="Austria" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="472">NMW 90584</specimenCode>
,
<quantity id="F43D9B11FF9DB207FD15E9CCFCECF833" box="[745,866,1935,1961]" metricMagnitude="0" metricUnit="m" metricValue="1.127" pageId="19" pageNumber="472" unit="mm" value="1127.0">1, 127 mm</quantity>
SL;
<collectingCountry id="4BD27664FF9DB207FC6BE9CCFC65F833" box="[919,1003,1935,1961]" name="Turkey" pageId="19" pageNumber="472">Turkey</collectingCountry>
:
<location id="361A602FFF9DB207FC0AE9CCFBF0F833" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9DB207FC0AE9CCFBF0F833" box="[1014,1150,1935,1961]" country="Turkey" name="Botan River" pageId="19" pageNumber="472">Botan River</location>
(photographs only).
</paragraph>
<caption id="67BA667CFF9AB200FF6BE8C3FE97F927" ID-DOI="http://doi.org/10.5281/zenodo.14895387" ID-Zenodo-Dep="14895387" httpUri="https://zenodo.org/record/14895387/files/figure.png" pageId="20" pageNumber="473" startId="20.[151,250,1664,1688]" targetBox="[151,1436,181,1640]" targetPageId="20" targetType="figure">
<paragraph id="337A36F4FF9AB200FF6BE8C3FE97F927" blockId="20.[151,1436,1664,1725]" pageId="20" pageNumber="473">
<emphasis id="01B1EAE6FF9AB200FF6BE8C3FEAEF902" bold="true" box="[151,288,1664,1688]" pageId="20" pageNumber="473">FIGURE 18.</emphasis>
<taxonomicName id="F4C54D77FF9AB200FEDBE8C3FDFDF902" authority=", ZIN" authorityName="ZIN" box="[295,627,1664,1688]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="473" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF9AB200FEDBE8C3FDB3F902" box="[295,573,1664,1688]" italics="true" pageId="20" pageNumber="473">Glyptothorax kurdistanicus</emphasis>
, ZIN
</taxonomicName>
20780, type, 113 mm SL; Iran: Germau (or Germav). Drawings modified from
<bibRefCitation id="57544B05FF9AB200FF6BE8E6FE9BF927" author="Berg, L. S." box="[151,277,1701,1725]" pageId="20" pageNumber="473" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="337A36F4FF9AB200FF3BE8A7FE1AF815" blockId="20.[151,1437,1764,2043]" pageId="20" pageNumber="473">
<emphasis id="01B1EAE6FF9AB200FF3BE8A7FD47F964" bold="true" box="[199,713,1764,1790]" pageId="20" pageNumber="473">Material used in molecular genetic analysis.</emphasis>
<materialsCitation id="83AD3CA9FF9AB200FD32E8A7FC33F8B8" accessionNumber="MW770717, MW770720, MW770728" country="Iraq" latitude="35.7509" location="Khewata" longLatPrecision="6" longitude="45.4797" municipality="Aw-e Shiler" pageId="20" pageNumber="473" specimenCount="1">
FSJF-DNA 2647;
<collectingCountry id="4BD27664FF9AB200FC6AE8A6FC49F965" box="[918,967,1765,1791]" name="Iraq" pageId="20" pageNumber="473">Iraq</collectingCountry>
:
<collectingMunicipality id="D31EAC8EFF9AB200FC33E8A7FBDDF964" box="[975,1107,1764,1790]" pageId="20" pageNumber="473">Aw-e Shiler</collectingMunicipality>
at
<location id="361A602FFF9AB200FB8DE8A7FB59F964" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9AB200FB8DE8A7FB59F964" box="[1137,1239,1764,1790]" country="Iraq" latitude="35.7509" longLatPrecision="6" longitude="45.4797" municipality="Aw-e Shiler" name="Khewata" pageId="20" pageNumber="473">Khewata</location>
,
<geoCoordinate id="56F15033FF9AB200FB1DE8A7FAB2F965" box="[1249,1340,1764,1791]" degrees="35.7509" direction="north" orientation="latitude" pageId="20" pageNumber="473" precision="5" value="35.7509">35.7509</geoCoordinate>
<geoCoordinate id="56F15033FF9AB200FABDE8A7FA12F965" box="[1345,1436,1764,1791]" degrees="45.4797" direction="east" orientation="longitude" pageId="20" pageNumber="473" precision="5" value="45.4797">45.4797</geoCoordinate>
(GenBank accession numbers:
<accessionNumber id="2C96AB17FF9AB200FE00E94BFC38F8B8" box="[508,950,1800,1826]" isEnumeration="true" pageId="20" pageNumber="473">MW770717, MW770720, MW770728</accessionNumber>
)
</materialsCitation>
.—
<materialsCitation id="83AD3CA9FF9AB200FC21E94BFEA9F8F1" accessionNumber="MW724522, MW724523, MW724524" country="Turkey" latitude="38.3547" location="Ucadim" longLatPrecision="6" longitude="41.7814" municipality="Ciratan" pageId="20" pageNumber="473" specimenCount="1" stateProvince="Bitlis">
FSJF-DNA 3335;
<collectingCountry id="4BD27664FF9AB200FB4FE94BFA89F8B8" box="[1203,1287,1800,1826]" name="Turkey" pageId="20" pageNumber="473">Turkey</collectingCountry>
:
<collectingRegion id="F101F816FF9AB200FAE9E94BFA17F8B9" box="[1301,1433,1800,1827]" country="Turkey" name="Bitlis" pageId="20" pageNumber="473">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="D31EAC8EFF9AB200FF1AE96FFEB6F8DC" box="[230,312,1836,1862]" pageId="20" pageNumber="473">Çıratan</collectingMunicipality>
<quantity id="F43D9B11FF9AB200FEC1E96FFEFDF8DC" box="[317,371,1836,1862]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="473" unit="km" value="3.0">3 km</quantity>
southwest of
<location id="361A602FFF9AB200FDFAE96EFDECF8D2" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9AB200FDFAE96EFDECF8D2" box="[518,610,1837,1864]" country="Turkey" latitude="38.3547" longLatPrecision="6" longitude="41.7814" municipality="Ciratan" name="Ucadim" pageId="20" pageNumber="473" stateProvince="Bitlis">Üçadım</location>
,
<geoCoordinate id="56F15033FF9AB200FD96E96FFD4BF8DD" box="[618,709,1836,1863]" degrees="38.3547" direction="north" orientation="latitude" pageId="20" pageNumber="473" precision="5" value="38.3547">38.3547</geoCoordinate>
<geoCoordinate id="56F15033FF9AB200FD35E96FFCAAF8DC" box="[713,804,1836,1862]" degrees="41.7814" direction="east" orientation="longitude" pageId="20" pageNumber="473" precision="5" value="41.7814">41.7814</geoCoordinate>
(GenBank accession numbers:
<accessionNumber id="2C96AB17FF9AB200FB87E96FFEAEF8F1" isEnumeration="true" pageId="20" pageNumber="473">MW724522, MW724523, MW724524</accessionNumber>
)
</materialsCitation>
.—
<materialsCitation id="83AD3CA9FF9AB200FEBBE913FE1EF815" accessionNumber="MW724521" country="Iraq" latitude="35.915" location="Bardbard" longLatPrecision="5" longitude="45.36772" municipality="Qal'ah Chwalan River" pageId="20" pageNumber="473" specimenCount="1">
FSJF-DNA 3349;
<collectingCountry id="4BD27664FF9AB200FDEAE912FDC9F8F1" box="[534,583,1873,1899]" name="Iraq" pageId="20" pageNumber="473">Iraq</collectingCountry>
:
<collectingMunicipality id="D31EAC8EFF9AB200FDAEE913FCC3F8F0" box="[594,845,1872,1898]" pageId="20" pageNumber="473">Qalah Chwalan River</collectingMunicipality>
at
<location id="361A602FFF9AB200FC93E913FC54F8F0" LSID="urn:lsid:plazi:treatment:BB6C87E2FF9DB20CFF6BEBA0FEE2F97B:361A602FFF9AB200FC93E913FC54F8F0" box="[879,986,1872,1898]" country="Iraq" latitude="35.915" longLatPrecision="5" longitude="45.36772" municipality="Qal'ah Chwalan River" name="Bardbard" pageId="20" pageNumber="473">Bardbard</location>
,
<geoCoordinate id="56F15033FF9AB200FC19E913FBCEF8F1" box="[997,1088,1872,1899]" degrees="35.9150" direction="north" orientation="latitude" pageId="20" pageNumber="473" precision="5" value="35.915">35.9150</geoCoordinate>
<geoCoordinate id="56F15033FF9AB200FBBBE913FB3EF8F1" box="[1095,1200,1872,1899]" degrees="45.36772" direction="east" orientation="longitude" pageId="20" pageNumber="473" precision="1" value="45.36772">45.36772</geoCoordinate>
(GenBank accession number:
<accessionNumber id="2C96AB17FF9AB200FF00E937FE06F815" box="[252,392,1908,1935]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW724521" pageId="20" pageNumber="473">MW724521</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF9AB201FF3BE9DBFBFBFD83" lastPageId="21" lastPageNumber="474" pageId="20" pageNumber="473" type="diagnosis">
<paragraph id="337A36F4FF9AB201FF3BE9DBFDC4FE37" blockId="20.[151,1437,1764,2043]" lastBlockId="21.[151,1437,151,537]" lastPageId="21" lastPageNumber="474" pageId="20" pageNumber="473">
<emphasis id="01B1EAE6FF9AB200FF3BE9DBFECCF828" bold="true" box="[199,322,1944,1970]" pageId="20" pageNumber="473">Diagnosis.</emphasis>
<taxonomicName id="F4C54D77FF9AB200FEBBE9DBFDF6F828" box="[327,632,1944,1970]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="473" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF9AB200FEBBE9DBFDF6F828" box="[327,632,1944,1970]" italics="true" pageId="20" pageNumber="473">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
is distinguished from its congeners in the Euphrates and Tigris drainages by having the thoracic adhesive apparatus 0.70.9 (1.0 in juveniles) times longer than wide (vs. 1.0
<quantity id="F43D9B11FF9AB200FAC8E9FFFAF9F84C" box="[1332,1399,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="20" pageNumber="473" unit="in" value="1.5">1.5 in</quantity>
<taxonomicName id="F4C54D77FF9AB200FA7DE9FFFE94F860" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="473" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF9AB200FA7DE9FFFE94F860" italics="true" pageId="20" pageNumber="473">G. armeniacus</emphasis>
</taxonomicName>
,
<quantity id="F43D9B11FF9AB200FED5E9A3FE15F860" box="[297,411,2016,2042]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="20" pageNumber="473" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">0.81.1 in</quantity>
<taxonomicName id="F4C54D77FF9AB200FE5EE9A3FE77F860" box="[418,505,2016,2042]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="473" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9AB200FE5EE9A3FE77F860" box="[418,505,2016,2042]" italics="true" pageId="20" pageNumber="473">G. cous</emphasis>
</taxonomicName>
), extending from the isthmus to the base of the first or third branched pectoral-fin ray (vs. to or beyond last pectoral-fin ray). It is further distinguished from
<taxonomicName id="F4C54D77FF9BB201FC28EED4FBF7FF2A" box="[980,1145,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF9BB201FC28EED4FBF7FF2A" box="[980,1145,151,177]" italics="true" pageId="21" pageNumber="474">G. armeniacus</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF9BB201FB53EED4FA8AFF2A" box="[1199,1284,151,177]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FB53EED4FA8AFF2A" box="[1199,1284,151,177]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
by having no or few very short anteromedial striae (vs. numerous and well developed), many small, often elongated warts on the dorsal and lateral head, back and flank (vs. with large, bony, striated and elongated tubercles in
<taxonomicName id="F4C54D77FF9BB201FB3FEE9CFA96FF62" box="[1219,1304,223,249]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FB3FEE9CFA96FF62" box="[1219,1304,223,249]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
; with small tubercles and few warts in
<taxonomicName id="F4C54D77FF9BB201FE37EF40FDFCFE86" box="[459,626,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF9BB201FE37EF40FDFCFE86" box="[459,626,259,285]" italics="true" pageId="21" pageNumber="474">G. armeniacus</emphasis>
</taxonomicName>
), an elevated thoracic adhesive apparatus (vs. not elevated in
<taxonomicName id="F4C54D77FF9BB201FAC9EF40FA02FE86" box="[1333,1420,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FAC9EF40FA02FE86" box="[1333,1420,259,285]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
), well delineated at its posterior margin (poorly delineated in
<taxonomicName id="F4C54D77FF9BB201FCBCEF64FC19FEDA" box="[832,919,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FCBCEF64FC19FEDA" box="[832,919,295,321]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
); completely situated on a horse-shoe shaped swelling (vs. reaching beyond swelling, usually onto the pectoral-fin base in
<taxonomicName id="F4C54D77FF9BB201FBE8EF08FBE2FEFE" box="[1044,1132,331,357]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FBE8EF08FBE2FEFE" box="[1044,1132,331,357]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
), and the caudal-peduncle depth 1.11.6 times in its length (vs.
<quantity id="F43D9B11FF9BB201FDC6EF2CFD22FE13" box="[570,684,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="4.953" metricValueMax="5.842" metricValueMin="4.064" pageId="21" pageNumber="474" unit="in" value="1.95" valueMax="2.3" valueMin="1.6">1.62.3 in</quantity>
<taxonomicName id="F4C54D77FF9BB201FD4FEF2CFC87FE12" box="[691,777,367,393]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF9BB201FD4FEF2CFC87FE12" box="[691,777,367,393]" italics="true" pageId="21" pageNumber="474">G. cous</emphasis>
</taxonomicName>
). See below for characters to distinguish this species from other
<taxonomicName id="F4C54D77FF9BB201FF24EFD0FEE3FE37" box="[216,365,403,429]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF9BB201FF24EFD0FEE3FE37" box="[216,365,403,429]" italics="true" pageId="21" pageNumber="474">Glyptothorax</emphasis>
</taxonomicName>
in the Middle East.
</paragraph>
<paragraph id="337A36F4FF9BB201FF3BEFF4FBFBFD83" blockId="21.[151,1437,151,537]" pageId="21" pageNumber="474">
Other characters useful for identification are: medial pit without striae; adipose-fin short, its length 0.61.0 times larger than the distance between the base of last dorsal-fin ray and adipose-fin origin; head blunt, spade-shaped, 2730% SL; few black or dark-brown blotches on flank. Size up to
<quantity id="F43D9B11FF9BB201FC15EFBCFBC8FD83" box="[1001,1094,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="474" unit="mm" value="127.0">127 mm</quantity>
SL.
</paragraph>
</subSubSection>
<caption id="67BA667CFF9BB201FF6BEA1FFF5BFB02" ID-DOI="http://doi.org/10.5281/zenodo.14895389" ID-Zenodo-Dep="14895389" httpUri="https://zenodo.org/record/14895389/files/figure.png" pageId="21" pageNumber="474" startId="21.[151,250,1116,1140]" targetBox="[164,1424,561,1093]" targetPageId="21" targetType="figure">
<paragraph id="337A36F4FF9BB201FF6BEA1FFF5BFB02" blockId="21.[151,1437,1116,1176]" pageId="21" pageNumber="474">
<emphasis id="01B1EAE6FF9BB201FF6BEA1FFEADFBEE" bold="true" box="[151,291,1116,1140]" pageId="21" pageNumber="474">FIGURE 19.</emphasis>
<taxonomicName id="F4C54D77FF9BB201FED1EA1FFDC8FBEE" box="[301,582,1116,1140]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF9BB201FED1EA1FFDC8FBEE" box="[301,582,1116,1140]" italics="true" pageId="21" pageNumber="474">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, juveniles, from top: FSJF 3652, top: 77 mm SL, 69 mm SL; Iraq: Aw-e Shiler River.
</paragraph>
</caption>
<caption id="67BA667CFF9BB201FF6BE9FEFE90F863" ID-DOI="http://doi.org/10.5281/zenodo.14895391" ID-Zenodo-Dep="14895391" httpUri="https://zenodo.org/record/14895391/files/figure.png" pageId="21" pageNumber="474" startId="21.[151,250,1981,2005]" targetBox="[150,1438,1200,1957]" targetPageId="21" targetType="figure">
<paragraph id="337A36F4FF9BB201FF6BE9FEFE90F863" blockId="21.[151,1437,1981,2041]" pageId="21" pageNumber="474">
<emphasis id="01B1EAE6FF9BB201FF6BE9FEFEAEF84F" bold="true" box="[151,288,1981,2005]" pageId="21" pageNumber="474">FIGURE 20.</emphasis>
<taxonomicName id="F4C54D77FF9BB201FEDAE9FEFDB5F84F" box="[294,571,1981,2005]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="474" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF9BB201FEDAE9FEFDB5F84F" box="[294,571,1981,2005]" italics="true" pageId="21" pageNumber="474">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, adults, from top: FFR 3916, from top: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki.
</paragraph>
</caption>
<caption id="67BA667CFF98B202FF6BEA58FEDBFBCD" ID-DOI="http://doi.org/10.5281/zenodo.14895393" ID-Zenodo-Dep="14895393" httpUri="https://zenodo.org/record/14895393/files/figure.png" pageId="22" pageNumber="475" startId="22.[151,250,1051,1075]" targetBox="[151,1436,181,1027]" targetPageId="22" targetType="figure">
<paragraph id="337A36F4FF98B202FF6BEA58FEDBFBCD" blockId="22.[151,1437,1051,1111]" pageId="22" pageNumber="475">
<emphasis id="01B1EAE6FF98B202FF6BEA58FEAFFBA9" bold="true" box="[151,289,1051,1075]" pageId="22" pageNumber="475">FIGURE 21.</emphasis>
<taxonomicName id="F4C54D77FF98B202FEDBEA58FDB0FBA9" box="[295,574,1051,1075]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="475" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF98B202FEDBEA58FDB0FBA9" box="[295,574,1051,1075]" italics="true" pageId="22" pageNumber="475">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, juveniles, from left: FSJF 3652, from left: 66 mm SL, 69 mm SL, 77 mm SL; Iraq: Aw-e Shiler River.
</paragraph>
</caption>
<caption id="67BA667CFF98B202FF6BE936FE90F82B" ID-DOI="http://doi.org/10.5281/zenodo.14895395" ID-Zenodo-Dep="14895395" httpUri="https://zenodo.org/record/14895395/files/figure.png" pageId="22" pageNumber="475" startId="22.[151,250,1909,1933]" targetBox="[150,1438,1135,1885]" targetPageId="22" targetType="figure">
<paragraph id="337A36F4FF98B202FF6BE936FE90F82B" blockId="22.[151,1437,1909,1969]" pageId="22" pageNumber="475">
<emphasis id="01B1EAE6FF98B202FF6BE936FEAEF817" bold="true" box="[151,288,1909,1933]" pageId="22" pageNumber="475">FIGURE 22.</emphasis>
<taxonomicName id="F4C54D77FF98B202FEDAE936FDB5F817" box="[294,571,1909,1933]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="475" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF98B202FEDAE936FDB5F817" box="[294,571,1909,1933]" italics="true" pageId="22" pageNumber="475">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, adults, from top: FFR 3916, from left: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki.
</paragraph>
</caption>
<caption id="67BA667CFF99B203FF6BEFA6FE65FDBB" ID-DOI="http://doi.org/10.5281/zenodo.14895397" ID-Zenodo-Dep="14895397" httpUri="https://zenodo.org/record/14895397/files/figure.png" pageId="23" pageNumber="476" startId="23.[151,250,485,509]" targetBox="[189,1398,181,461]" targetPageId="23" targetType="figure">
<paragraph id="337A36F4FF99B203FF6BEFA6FE65FDBB" blockId="23.[151,1436,485,545]" pageId="23" pageNumber="476">
<emphasis id="01B1EAE6FF99B203FF6BEFA6FEAEFE67" bold="true" box="[151,288,485,509]" pageId="23" pageNumber="476">FIGURE 23.</emphasis>
<taxonomicName id="F4C54D77FF99B203FED9EFA6FDB4FE67" box="[293,570,485,509]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FED9EFA6FDB4FE67" box="[293,570,485,509]" italics="true" pageId="23" pageNumber="476">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, skin ornamentation; from left: adults, FFR 3916, from left: 120 mm SL, 104 mm SL, 98 mm SL; Turkey: stream Eziki.
</paragraph>
</caption>
<caption id="67BA667CFF99B203FF6BEB9BFE6CF98F" ID-DOI="http://doi.org/10.5281/zenodo.14895399" ID-Zenodo-Dep="14895399" httpUri="https://zenodo.org/record/14895399/files/figure.png" pageId="23" pageNumber="476" startId="23.[151,250,1496,1520]" targetBox="[151,1436,570,1473]" targetPageId="23" targetType="figure">
<paragraph id="337A36F4FF99B203FF6BEB9BFE6CF98F" blockId="23.[151,1436,1496,1557]" pageId="23" pageNumber="476">
<emphasis id="01B1EAE6FF99B203FF6BEB9BFE91FA6B" bold="true" box="[151,287,1496,1521]" pageId="23" pageNumber="476">FIGURE 24.</emphasis>
<taxonomicName id="F4C54D77FF99B203FED9EB9AFDB4FA6B" box="[293,570,1497,1521]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FED9EB9AFDB4FA6B" box="[293,570,1497,1521]" italics="true" pageId="23" pageNumber="476">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, from top: FSJF 3652, 77 mm SL; Iraq: Aw-e Shiler River; FSJF-DNA 3349; 120 mm SL; Iraq: Qalah Chwalan River.
</paragraph>
</caption>
<subSubSection id="7BDF657FFF99B203FF3BE800FC7FF91B" pageId="23" pageNumber="476" type="distribution">
<paragraph id="337A36F4FF99B203FF3BE800FC7FF91B" blockId="23.[151,1437,1603,2025]" pageId="23" pageNumber="476">
<emphasis id="01B1EAE6FF99B203FF3BE800FEEFF9C7" bold="true" box="[199,353,1603,1629]" pageId="23" pageNumber="476">Distribution.</emphasis>
<taxonomicName id="F4C54D77FF99B203FE94E800FD15F9C7" box="[360,667,1603,1629]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FE94E800FD15F9C7" box="[360,667,1603,1629]" italics="true" pageId="23" pageNumber="476">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
seems to be endemic to the Tigris drainage where it has been found in mountain rivers in
<collectingCountry id="4BD27664FF99B203FE74E824FE57F91B" box="[392,473,1639,1665]" name="Turkey" pageId="23" pageNumber="476">Turkey</collectingCountry>
,
<collectingCountry id="4BD27664FF99B203FE18E824FD9FF91B" box="[484,529,1639,1665]" name="Iran" pageId="23" pageNumber="476">Iran</collectingCountry>
and
<collectingCountry id="4BD27664FF99B203FDBBE824FDFAF91B" box="[583,628,1639,1665]" name="Iraq" pageId="23" pageNumber="476">Iraq</collectingCountry>
down to the Lesser Zab drainage.
</paragraph>
</subSubSection>
<subSubSection id="7BDF657FFF99B20CFF3BE8C8FEE2F97B" lastPageId="24" lastPageNumber="477" pageId="23" pageNumber="476" type="discussion">
<paragraph id="337A36F4FF99B203FF3BE8C8FADFF85F" blockId="23.[151,1437,1603,2025]" pageId="23" pageNumber="476">
<emphasis id="01B1EAE6FF99B203FF3BE8C8FEB5F93F" bold="true" box="[199,315,1675,1701]" pageId="23" pageNumber="476">Remarks.</emphasis>
When
<bibRefCitation id="57544B05FF99B203FE75E8C8FD81F93F" author="Berg, L. S." box="[393,527,1675,1701]" pageId="23" pageNumber="476" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
described
<taxonomicName id="F4C54D77FF99B203FD7AE8C8FCB2F93F" box="[646,828,1675,1701]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FD7AE8C8FCB2F93F" box="[646,828,1675,1701]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
from the Lesser Zab, he was not aware of the existence of
<taxonomicName id="F4C54D77FF99B203FF4AE8ECFE82F953" box="[182,268,1711,1737]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FF4AE8ECFE82F953" box="[182,268,1711,1737]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
and therefore did not distinguish
<taxonomicName id="F4C54D77FF99B203FD70E8ECFCCBF953" box="[652,837,1711,1737]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FD70E8ECFCCBF953" box="[652,837,1711,1737]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
from
<taxonomicName id="F4C54D77FF99B203FC77E8ECFC6FF953" box="[907,993,1711,1737]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FC77E8ECFC6FF953" box="[907,993,1711,1737]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
. We found two molecular clades in the Lesser Zab, one of them, we identify as
<taxonomicName id="F4C54D77FF99B203FD9DE890FD36F977" box="[609,696,1747,1773]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FD9DE890FD36F977" box="[609,696,1747,1773]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
and the second as
<taxonomicName id="F4C54D77FF99B203FC6CE890FBC7F977" box="[912,1097,1747,1773]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FC6CE890FBC7F977" box="[912,1097,1747,1773]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
. The
<typeStatus id="EC7E8856FF99B203FB77E890FB35F977" box="[1163,1211,1747,1773]" pageId="23" pageNumber="476">type</typeStatus>
of
<taxonomicName id="F4C54D77FF99B203FB1FE890FA12F977" box="[1251,1436,1747,1773]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FB1FE890FA12F977" box="[1251,1436,1747,1773]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
(ZIN 20780,
<figureCitation id="ABFE2A71FF99B203FED4E8B4FEF9F88B" box="[296,375,1783,1809]" captionStart="FIGURE 18" captionStartId="20.[151,250,1664,1688]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure-23@20.[151,1436,181,1640]" captionTargetPageId="20" captionText="FIGURE 18. Glyptothorax kurdistanicus, ZIN 20780, type, 113 mm SL; Iran: Germau (or Germav). Drawings modified from Berg (1931)." figureDoi="http://doi.org/10.5281/zenodo.14895387" httpUri="https://zenodo.org/record/14895387/files/figure.png" pageId="23" pageNumber="476">Fig. 18</figureCitation>
) is not in a good condition, parts of the skin and the thoracic adhesive apparatus are lost, fins are broken, and the head is turned down. However, the diagnostic deep caudal peduncle and the short thoracic adhesive apparatus are well visible in the
<typeStatus id="EC7E8856FF99B203FDFAE97CFDB4F8C3" box="[518,570,1855,1881]" pageId="23" pageNumber="476">type</typeStatus>
. The thoracic adhesive apparatus extends from the isthmus to the base of the first or third branched pectoral-fin ray in ZIN 20780 as well as in our material of
<taxonomicName id="F4C54D77FF99B203FBD3E920FB66F8E7" box="[1071,1256,1891,1917]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FBD3E920FB66F8E7" box="[1071,1256,1891,1917]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
. In
<taxonomicName id="F4C54D77FF99B203FAE9E920FAE5F8E7" box="[1301,1387,1891,1917]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FAE9E920FAE5F8E7" box="[1301,1387,1891,1917]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
, the thoracic adhesive apparatus is extending from the isthmus to the base of the last pectoral-fin ray or to the posterior limit of the pectoral-fin base. Furthermore, the apparatus is elevated in
<taxonomicName id="F4C54D77FF99B203FC48E9E8FBE2F85F" box="[948,1132,1963,1989]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF99B203FC48E9E8FBE2F85F" box="[948,1132,1963,1989]" italics="true" pageId="23" pageNumber="476">G. kurdistanicus</emphasis>
</taxonomicName>
(vs. not in
<taxonomicName id="F4C54D77FF99B203FB17E9E8FACEF85F" box="[1259,1344,1963,1989]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FB17E9E8FACEF85F" box="[1259,1344,1963,1989]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="337A36F4FF99B20CFF3BE98CFADBFEDB" blockId="23.[151,1437,1603,2025]" lastBlockId="24.[151,1437,151,1761]" lastPageId="24" lastPageNumber="477" pageId="23" pageNumber="476">
We do not see the bony, striated, elongated tubercles on the head, back and flank diagnostic for
<taxonomicName id="F4C54D77FF99B203FB23E98CFABDF873" box="[1247,1331,1999,2025]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="476" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF99B203FB23E98CFABDF873" box="[1247,1331,1999,2025]" italics="true" pageId="23" pageNumber="476">G. cous</emphasis>
</taxonomicName>
. As these tubercles are very peculiar, we would expect
<bibRefCitation id="57544B05FF96B20CFD76EED4FC9EFF2B" author="Berg, L. S." box="[650,784,151,177]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
to mention them, what he did not. Mikhail Nazarkin (ZIN) examined the specimen and did not find tubercles on ZIN 20780 (pers. comm., 2020). While the tubercles might all have been scraped off in ZIN 20780, they also might have never existed, as in the other materials from the Lesser Zab examined by us. Indeed, the sole adult
<taxonomicName id="F4C54D77FF96B20CFD88EF40FCA5FE87" box="[628,811,259,285]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFD88EF40FCA5FE87" box="[628,811,259,285]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
collected from the Lesser Zab (
<figureCitation id="ABFE2A71FF96B20CFB75EF40FB57FE87" box="[1161,1241,259,285]" captionStart="FIGURE 24" captionStartId="23.[151,250,1496,1520]" captionTargetBox="[151,1436,570,1473]" captionTargetId="figure-49@23.[151,1436,570,1473]" captionTargetPageId="23" captionText="FIGURE 24. Glyptothorax kurdistanicus, from top: FSJF 3652, 77 mm SL; Iraq:Aw-e Shiler River; FSJF-DNA 3349; 120 mm SL; Iraq: Qalah Chwalan River." figureDoi="http://doi.org/10.5281/zenodo.14895399" httpUri="https://zenodo.org/record/14895399/files/figure.png" pageId="24" pageNumber="477">Fig. 24</figureCitation>
) agrees very well with the
<typeStatus id="EC7E8856FF96B20CFF05EF64FEA7FEDB" box="[249,297,295,321]" pageId="24" pageNumber="477">type</typeStatus>
in the general body shape also supporting our identification of this species as
<emphasis id="01B1EAE6FF96B20CFB69EF64FADBFEDB" box="[1173,1365,295,321]" italics="true" pageId="24" pageNumber="477">
<taxonomicName id="F4C54D77FF96B20CFB69EF64FADFFEDB" box="[1173,1361,295,321]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">G. kurdistanicus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BEF08FEB9FDA7" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
It is unclear, in which condition ZIN 20780 was, when Berg studied it. On the drawing, the head was already turned downwards, the pattern of black dots on the flank agrees well in the
<typeStatus id="EC7E8856FF96B20CFC0FEF2CFBADFE13" box="[1011,1059,367,393]" pageId="24" pageNumber="477">type</typeStatus>
and its drawing and
<bibRefCitation id="57544B05FF96B20CFAE8EF2CFA12FE13" author="Berg, L. S." box="[1300,1436,367,393]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
does not give the number of serrae on the inner margin of the ossified pectoral fin-ray. In ZIN 20780, the ossified ray is broken today. To us, it seems possible that ZIN 20780 was already in a poor condition when
<bibRefCitation id="57544B05FF96B20CFAEFEFF4FA12FE4B" author="Berg, L. S." box="[1299,1436,439,465]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
examined it but we are unable to demonstrate this. Potentially, he took some artistic freedom when preparing the drawing of ZIN 20780, resulting in complete fins, and a complete thoracic adhesive apparatus, which is incomplete in ZIN 20780.
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BEC04FE16FC53" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
<bibRefCitation id="57544B05FF96B20CFF3BEC04FEDDFDFB" author="Berg, L. S." box="[199,339,583,609]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
distinguish
<taxonomicName id="F4C54D77FF96B20CFE19EC04FD2CFDFB" box="[485,674,583,609]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFE19EC04FD2CFDFB" box="[485,674,583,609]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
from
<taxonomicName id="F4C54D77FF96B20CFD13EC04FC17FDFA" box="[751,921,583,609]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFD13EC04FC17FDFA" box="[751,921,583,609]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
, described by him in 1918, by the thoracic adhesive apparatus wider than long in
<taxonomicName id="F4C54D77FF96B20CFDBDEC28FD76FD1F" box="[577,760,619,645]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFDBDEC28FD76FD1F" box="[577,760,619,645]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
(vs. longer than wide in
<taxonomicName id="F4C54D77FF96B20CFBF6EC28FB20FD1E" box="[1034,1198,619,645]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFBF6EC28FB20FD1E" box="[1034,1198,619,645]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
), a character adopted by later authors (
<bibRefCitation id="57544B05FF96B20CFEA6ECCCFE54FD33" author="Kuru, M." box="[346,474,654,681]" pageId="24" pageNumber="477" refId="ref19167" refString="Kuru, M. (1975) Dicle-Firat, Kura-Aras, Van Golu Karadeniz Havzasi Tatlisularinda Yasayan Baliklarin (Pisces) Sistematik ve Zoocografik Yonden Incelenmesi. Docentlik Tezi. Ataturk Universitesi, Fen Fakultesi, Erzurum, 180 pp. [in Turkish]" type="book" year="1975">Kuru 1975</bibRefCitation>
,
<bibRefCitation id="57544B05FF96B20CFE1BECCCFD67FD33" author="Geldiay, R. &amp; Balik, S." box="[487,745,655,681]" pageId="24" pageNumber="477" pagination="1 - 519" refId="ref18248" refString="Geldiay, R. &amp; Balik, S. (1999) Turkiye Tatlisu Baliklari [Freshwater fishes of Turkey]. Ege. In: Universitesi Su Urunleri Fakultesi Yayinlari, Izmir, 46. Ders Kitabi Dizini, 16, pp. 1-519. [in Turkish]" type="journal article" year="1999">Geldiay &amp; Balık 1999</bibRefCitation>
,
<bibRefCitation id="57544B05FF96B20CFD0AECCCFC35FD33" author="Kaya, C. &amp; Turan, D. &amp; Unlu, E." box="[758,955,655,681]" pageId="24" pageNumber="477" pagination="545 - 562" refId="ref18890" refString="Kaya, C., Turan, D. &amp; Unlu, E. (2016) The latest status and distribution of fishes in upper Tigris River and two new records for Turkish freshwaters. Turkish Journal of Fisheries and Aquatic Sciences, 16, 545-562. https://doi.org/10.4194/1303-2712-v16_3_07" type="journal article" year="2016">
Kaya
<emphasis id="01B1EAE6FF96B20CFCC6ECCCFCE0FD33" box="[826,878,655,681]" italics="true" pageId="24" pageNumber="477">et al</emphasis>
. 2016
</bibRefCitation>
).
<bibRefCitation id="57544B05FF96B20CFC33ECCCFBD0FD33" author="Coad, B. W." box="[975,1118,655,681]" pageId="24" pageNumber="477" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
diagnosed
<taxonomicName id="F4C54D77FF96B20CFB1EECCCFA12FD33" box="[1250,1436,655,681]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFB1EECCCFA12FD33" box="[1250,1436,655,681]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
by having the thoracic adhesive apparatus wider than long or as wide as long. However, materials identified as
<taxonomicName id="F4C54D77FF96B20CFA7DECF0FEA3FD6B" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFA7DECF0FEA3FD6B" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
by
<bibRefCitation id="57544B05FF96B20CFEA6EC94FE69FD6B" author="Coad, B. W." box="[346,487,727,753]" pageId="24" pageNumber="477" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
originate from the Turkish Euphrates (ZMH 4430) and not from the Iranian Lesser Zab or even from the Tigris drainage, from where
<taxonomicName id="F4C54D77FF96B20CFD3EECB8FCF7FC8F" box="[706,889,763,789]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFD3EECB8FCF7FC8F" box="[706,889,763,789]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
was described. The two individuals (ZMH 4430) identified by
<bibRefCitation id="57544B05FF96B20CFED1ED5CFE34FCA3" author="Coad, B. W." box="[301,442,799,825]" pageId="24" pageNumber="477" pagination="291 - 295" refId="ref17741" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291-295." type="journal article" year="1981">Coad (1981)</bibRefCitation>
as
<taxonomicName id="F4C54D77FF96B20CFE1CED5CFD16FCA3" box="[480,664,799,825]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFE1CED5CFD16FCA3" box="[480,664,799,825]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
were identified by us as
<taxonomicName id="F4C54D77FF96B20CFC4DED5CFB88FCA2" box="[945,1030,799,825]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFC4DED5CFB88FCA2" box="[945,1030,799,825]" italics="true" pageId="24" pageNumber="477">G. cous</emphasis>
</taxonomicName>
, and most materials identified as
<taxonomicName id="F4C54D77FF96B20CFA7DED5CFF44FCC6" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFA7DED5CFF44FCC6" italics="true" pageId="24" pageNumber="477">G. cous</emphasis>
</taxonomicName>
from SMF was previously identified as
<taxonomicName id="F4C54D77FF96B20CFD73ED00FCC6FCC7" box="[655,840,835,861]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFD73ED00FCC6FCC7" box="[655,840,835,861]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
. Indeed, in
<taxonomicName id="F4C54D77FF96B20CFC31ED00FB0BFCC7" box="[973,1157,835,861]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFC31ED00FB0BFCC7" box="[973,1157,835,861]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF96B20CFB40ED00FA9FFCC6" box="[1212,1297,835,861]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFB40ED00FA9FFCC6" box="[1212,1297,835,861]" italics="true" pageId="24" pageNumber="477">G. cous</emphasis>
</taxonomicName>
the thoracic adhesive apparatus might be as wide as long and therefore these species are confused in the literature. The apparatus is reaching only to the base of the first or third branched pectoral-fin ray in
<taxonomicName id="F4C54D77FF96B20CFC28EDC8FB05FC3F" box="[980,1163,907,933]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFC28EDC8FB05FC3F" box="[980,1163,907,933]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
, a character state clearly different from
<taxonomicName id="F4C54D77FF96B20CFEC7EDECFE1FFC52" box="[315,401,943,969]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFEC7EDECFE1FFC52" box="[315,401,943,969]" italics="true" pageId="24" pageNumber="477">G. cous</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BED90FBCEFBE7" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
Except of the short thoracic adhesive apparatus,
<bibRefCitation id="57544B05FF96B20CFD27ED90FCEFFC77" author="Berg, L. S." box="[731,865,979,1005]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
distinguished
<taxonomicName id="F4C54D77FF96B20CFBFDED90FB36FC77" box="[1025,1208,979,1005]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFBFDED90FB36FC77" box="[1025,1208,979,1005]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
from
<taxonomicName id="F4C54D77FF96B20CFB05ED90FA13FC76" box="[1273,1437,979,1005]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFB05ED90FA13FC76" box="[1273,1437,979,1005]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
by the thoracic adhesive apparatus lacking pinnate striae (vs. present). In our material of
<taxonomicName id="F4C54D77FF96B20CFB7FEDB4FAB5FB8B" box="[1155,1339,1015,1041]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFB7FEDB4FAB5FB8B" box="[1155,1339,1015,1041]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
, pinnate striae occur while not as many as in
<taxonomicName id="F4C54D77FF96B20CFDB8EA58FD62FBAE" box="[580,748,1051,1077]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFDB8EA58FD62FBAE" box="[580,748,1051,1077]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
.
<bibRefCitation id="57544B05FF96B20CFD00EA58FCF1FBAF" author="Berg, L. S." box="[764,895,1051,1077]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931</bibRefCitation>
,
<figureCitation id="ABFE2A71FF96B20CFC71EA58FC41FBAF" box="[909,975,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1099,1123]" captionTargetBox="[433,1155,179,1073]" captionTargetId="figure-23@2.[433,1155,179,1073]" captionTargetPageId="2" captionText="FIGURE 1. Thoracic adhesive apparatus of Glyptothorax, showing structures defined and mentioned in this study: anterolateral striae (ALS); anteromedial striae (AMS); medial pit (MP) (G. daemon, FSJF 3962, 109 mm SL; based on Ng &amp; Kottelat 2016)." figureDoi="http://doi.org/10.5281/zenodo.14895351" httpUri="https://zenodo.org/record/14895351/files/figure.png" pageId="24" pageNumber="477">Fig. 1</figureCitation>
) shows that
<taxonomicName id="F4C54D77FF96B20CFB94EA58FAACFBAF" box="[1128,1314,1051,1077]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFB94EA58FAACFBAF" box="[1128,1314,1051,1077]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
also lacks anteromedial striae on the thoracic adhesive apparatus. However, no pinnate or anteromedial striae can be seen. Our materials show few and short anteromedial striae, much less than in
<taxonomicName id="F4C54D77FF96B20CFC68EA20FBB4FBE6" box="[916,1082,1123,1149]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFC68EA20FBB4FBE6" box="[916,1082,1123,1149]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BEAC4FE5FFA07" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
<bibRefCitation id="57544B05FF96B20CFF3BEAC4FEC0FB3B" author="Berg, L. S." box="[199,334,1159,1185]" pageId="24" pageNumber="477" pagination="1267 - 1270" refId="ref17533" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267-1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
gives the
<typeStatus id="EC7E8856FF96B20CFE43EAC4FE61FB3B" box="[447,495,1159,1185]" pageId="24" pageNumber="477">type</typeStatus>
locality „in
<collectingRegion id="F101F816FF96B20CFD85EAC4FD64FB3B" box="[633,746,1159,1185]" country="Iran" name="Kordestan" pageId="24" pageNumber="477">Kurdistan</collectingRegion>
, at the village Germau (or Germav), at the height of
<quantity id="F43D9B11FF96B20CFACFEAC5FA07FB3B" box="[1331,1417,1158,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="477" unit="m" value="1500.0">1500 m</quantity>
... Germau (or Germav, Germaw) is situated in latitude
<geoCoordinate id="56F15033FF96B20CFD1CEAE8FC92FB5F" box="[736,796,1195,1221]" degrees="36" direction="north" orientation="latitude" pageId="24" pageNumber="477" precision="55555" value="36.0">36°N</geoCoordinate>
southeast of Serdesht, on the western slope of the Sur-kei Range, in the basin of the river Bané, tributary to the Lesser Zab, which is tributary to the Tigris R.“.
<bibRefCitation id="57544B05FF96B20CFAF3EA8CFA12FB73" author="Coad, B. W." box="[1295,1436,1231,1257]" pageId="24" pageNumber="477" pagination="218 - 257" refId="ref17808" refString="Coad, B. W. (2014) Review of the Freshwater Catfishes of Iran (Order Siluriformes). Iranian Journal of Ichthyology, 1 (4), 218-257" type="journal article" year="2014">Coad (2014)</bibRefCitation>
stated that Germau is probably Garmab and Bané is probably Baneh. The village of Germab could not be located in gazetteers or on maps with a relevant longitude but Sar Dasht (
<geoCoordinate id="56F15033FF96B20CFCA3EB54FC4CFAAB" box="[863,962,1303,1329]" degrees="36" direction="north" minutes="09" orientation="latitude" pageId="24" pageNumber="477" precision="925" value="36.15">36°09N</geoCoordinate>
,
<geoCoordinate id="56F15033FF96B20CFC33EB55FBA3FAAB" box="[975,1069,1302,1329]" degrees="45" direction="east" minutes="28" orientation="longitude" pageId="24" pageNumber="477" precision="925" value="45.466667">45°28E</geoCoordinate>
) and Baneh (
<geoCoordinate id="56F15033FF96B20CFB39EB55FAA7FAAB" box="[1221,1321,1302,1329]" degrees="35" direction="north" minutes="59" orientation="latitude" pageId="24" pageNumber="477" precision="925" value="35.983334">35°59N</geoCoordinate>
,
<geoCoordinate id="56F15033FF96B20CFAC9EB55FA1DFAAB" box="[1333,1427,1302,1329]" degrees="45" direction="east" minutes="53" orientation="longitude" pageId="24" pageNumber="477" precision="925" value="45.883335">45°53E</geoCoordinate>
) are evident and the locality is between them and lies in the Iranian Lesser Zab drainage. We are not aware that any
<taxonomicName id="F4C54D77FF96B20CFF6BEB1CFEA2FAE3" box="[151,300,1375,1401]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF96B20CFF6BEB1CFEA2FAE3" box="[151,300,1375,1401]" italics="true" pageId="24" pageNumber="477">Glyptothorax</emphasis>
</taxonomicName>
has been recorded from the Iranian Lesser Zab and Esmaeili (pers. comm., 2012) was not aware of a recent record of the species.
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BEBE4FCFEF9B7" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
The distribution of
<taxonomicName id="F4C54D77FF96B20CFE62EBE4FDDBFA5B" box="[414,597,1447,1473]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFE62EBE4FDDBFA5B" box="[414,597,1447,1473]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
remains largely unclear and it could be endemic to the Tigris drainage from headwaters in
<collectingCountry id="4BD27664FF96B20CFECBEB88FE09FA7F" box="[311,391,1483,1509]" name="Turkey" pageId="24" pageNumber="477">Turkey</collectingCountry>
down to the Lesser Zab, in Iraqi
<collectingRegion id="F101F816FF96B20CFD08EB88FCECFA7F" box="[756,866,1483,1509]" country="Iran" name="Kordestan" pageId="24" pageNumber="477">Kurdistan</collectingRegion>
and
<collectingCountry id="4BD27664FF96B20CFC6BEB88FC46FA7F" box="[919,968,1483,1509]" name="Iran" pageId="24" pageNumber="477">Iran</collectingCountry>
.
<bibRefCitation id="57544B05FF96B20CFC2DEB88FAA3FA7F" author="Jouladeh-Roudbar, A. &amp; Ghanavi, H. R. &amp; Doadrio, I." box="[977,1325,1483,1509]" pageId="24" pageNumber="477" refId="ref18832" refString="Jouladeh-Roudbar, A., Ghanavi, H. R. &amp; Doadrio, I. (2020) Ichthyofauna from Iranian freshwater: annotated checklist, diagnosis, taxonomy, distribution and conservation assessment. Zoological Studies, 59, 21. https://doi.org/10.6620/ZS.2020.59-21." type="journal volume" year="2020">
Jouladeh-Roudbar
<emphasis id="01B1EAE6FF96B20CFB59EB88FB5BFA7F" box="[1189,1237,1483,1509]" italics="true" pageId="24" pageNumber="477">et al</emphasis>
. (2020)
</bibRefCitation>
show one
<taxonomicName id="F4C54D77FF96B20CFF6BEBACFEA2F993" box="[151,300,1519,1545]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="genus">
<emphasis id="01B1EAE6FF96B20CFF6BEBACFEA2F993" box="[151,300,1519,1545]" italics="true" pageId="24" pageNumber="477">Glyptothorax</emphasis>
</taxonomicName>
from the Sirvan identified as
<taxonomicName id="F4C54D77FF96B20CFD82EBACFCB9F993" box="[638,823,1519,1545]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFD82EBACFCB9F993" box="[638,823,1519,1545]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
, but this fish is quite different from those found in the Lesser Zab and resembles
<taxonomicName id="F4C54D77FF96B20CFE3CE850FDA5F9B7" authorityName="Coad" authorityYear="1981" box="[448,555,1555,1581]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF96B20CFE3CE850FDA5F9B7" box="[448,555,1555,1581]" italics="true" pageId="24" pageNumber="477">G. silviae</emphasis>
</taxonomicName>
from its general appearance.
</paragraph>
<paragraph id="337A36F4FF96B20CFF3BE874FEE2F97B" blockId="24.[151,1437,151,1761]" pageId="24" pageNumber="477">
We found
<taxonomicName id="F4C54D77FF96B20CFEC2E874FE79F9CB" box="[318,503,1591,1617]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFEC2E874FE79F9CB" box="[318,503,1591,1617]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF96B20CFDCCE874FD09F9CA" box="[560,647,1591,1617]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFDCCE874FD09F9CA" box="[560,647,1591,1617]" italics="true" pageId="24" pageNumber="477">G. cous</emphasis>
</taxonomicName>
sympatric in the Aw-e Shiler River which is a tributary of the Lesser Zab and
<taxonomicName id="F4C54D77FF96B20CFF06E818FE3BF9EF" authorityName=", Berg" authorityYear="1918" box="[250,437,1627,1653]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF96B20CFF06E818FE3BF9EF" box="[250,437,1627,1653]" italics="true" pageId="24" pageNumber="477">G. steindachneri</emphasis>
</taxonomicName>
is a third species also recorded from the Lesser Zab drainage.
<taxonomicName id="F4C54D77FF96B20CFB85E818FAC7F9EE" box="[1145,1353,1627,1653]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="cous">
<emphasis id="01B1EAE6FF96B20CFB85E818FAC7F9EE" box="[1145,1353,1627,1653]" italics="true" pageId="24" pageNumber="477">Glyptothorax cous</emphasis>
</taxonomicName>
and
<taxonomicName id="F4C54D77FF96B20CFA7DE818FEA1F903" authorityName=", Berg" authorityYear="1918" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="01B1EAE6FF96B20CFA7DE818FEA1F903" italics="true" pageId="24" pageNumber="477">G. steindachneri</emphasis>
</taxonomicName>
are widespread species found in middle-size to large rivers. But
<taxonomicName id="F4C54D77FF96B20CFC06E83CFB10F902" box="[1018,1182,1663,1689]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="armeniacus">
<emphasis id="01B1EAE6FF96B20CFC06E83CFB10F902" box="[1018,1182,1663,1689]" italics="true" pageId="24" pageNumber="477">G. armeniacus</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF96B20CFB57E83CFAABF903" box="[1195,1317,1663,1689]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="daemon">
<emphasis id="01B1EAE6FF96B20CFB57E83CFAABF903" box="[1195,1317,1663,1689]" italics="true" pageId="24" pageNumber="477">G. daemon</emphasis>
</taxonomicName>
,
<taxonomicName id="F4C54D77FF96B20CFACEE83CFA12F903" authorityName="Coad" authorityYear="1981" box="[1330,1436,1663,1689]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="silviae">
<emphasis id="01B1EAE6FF96B20CFACEE83CFA12F903" box="[1330,1436,1663,1689]" italics="true" pageId="24" pageNumber="477">G. silviae</emphasis>
</taxonomicName>
and potentially also
<taxonomicName id="F4C54D77FF96B20CFE8AE8E0FDA3F927" box="[374,557,1699,1725]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="477" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="01B1EAE6FF96B20CFE8AE8E0FDA3F927" box="[374,557,1699,1725]" italics="true" pageId="24" pageNumber="477">G. kurdistanicus</emphasis>
</taxonomicName>
are small species, which are highly rheophilic and they seem to inhabit smaller, fast flowing rivers.
</paragraph>
</subSubSection>
</treatment>
</document>