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<document id="BD21D274275C0F9B23B60E4983F46C63" ID-DOI="10.11646/phytotaxa.367.1.10" ID-ISSN="1179-3163" ID-Zenodo-Dep="13704396" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1725583144942" checkinUser="felipe" docAuthor="Suetsugu, Kenji, Suleiman, Monica, Anthony, Florina &amp; Tsukaya, Hirokazu" docDate="2018" docId="E97A87AEFFA6FFDAFF75FAC6FECD2B30" docLanguage="en" docName="phytotaxa.367.1.10.pdf" docOrigin="Phytotaxa 367 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.367.1.10" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Aphyllorchis maliauensis Suetsugu, M. Suleiman &amp; Tsukaya 2018, sp. nov." docType="treatment" docVersion="1" lastPageNumber="86" masterDocId="1543FFD6FFA6FFDBFFFDFF91FFDC2E25" masterDocTitle="Aphyllorchis maliauensis (Orchidaceae), a new species from the Maliau Basin, Sabah, Borneo" masterLastPageNumber="90" masterPageNumber="85" pageNumber="85" updateTime="1742296480746" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="71E25E138582F20421C543654BEED246">
<mods:title id="3B9975D5CCAA544A36DA8A68077C54BA">Aphyllorchis maliauensis (Orchidaceae), a new species from the Maliau Basin, Sabah, Borneo</mods:title>
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<mods:namePart id="7262524A4047C1701132A8FE15684E12">Suetsugu, Kenji</mods:namePart>
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<mods:namePart id="21178F12B4EB8BB12760DE52C845E77B">Suleiman, Monica</mods:namePart>
<mods:affiliation id="4B4F95822AC0F856F7D9AD116C47E2DC">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
</mods:name>
<mods:name id="ECFB634274743D07D6687217F98BC604" type="personal">
<mods:role id="3E3EDBBB2E81E9ABAEF241137DCDE51D">
<mods:roleTerm id="D5D796FF2C0F3D022349CF9ABC1A1A79">Author</mods:roleTerm>
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<mods:namePart id="843729C34DE888663B6152E18DAF0964">Anthony, Florina</mods:namePart>
<mods:affiliation id="C4A8210B2FD395AECB472F3B2C7A6215">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:name id="2A591DA4A98D63A35657368FAF1F0141" type="personal">
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<mods:namePart id="0134767B7F3142427689E4A245B503CE">Tsukaya, Hirokazu</mods:namePart>
<mods:affiliation id="F8A0BA6CB00231117B33D36FE9ABEADE">Department of Biological Sciences, Faculty of Science, The University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 0033, Japan &amp; Bio-Next Project, Exploratory Research Center on Life and Living Systems, National Institutes of Natural Sciences, Yamate Build. # 3, 5 - 1, Higashiyama, Myodaiji, Okazaki, Aichi 444 - 8787, Japan</mods:affiliation>
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<mods:title id="9421A7CF69390437BB506A8665FA0459">Phytotaxa</mods:title>
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<mods:part id="AE1F87C68FDD31A4E0EA87E6A87B5C4D">
<mods:date id="2A87D5A0E480A06062F531E57E10FE2A">2018</mods:date>
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<mods:number id="E71037D47E1BE54A02756CEFB1C49A9D">2018-08-30</mods:number>
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<mods:number id="8749CA43E973F423C1117F0223CF6BF8">367</mods:number>
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<mods:number id="68B9284118AFFC2EBDFFBEA4DE164903">1</mods:number>
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<mods:start id="F6945BF1566C1F8F91B645D17236F889">85</mods:start>
<mods:end id="5D0D099350100B2695BB4EE8FF83E37C">90</mods:end>
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<mods:url id="C32271DB2407BA3E7E319DC1D419CB7D">http://dx.doi.org/10.11646/phytotaxa.367.1.10</mods:url>
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<mods:identifier id="15FD3F62007F2097A0082EA140B0C5B5" type="DOI">10.11646/phytotaxa.367.1.10</mods:identifier>
<mods:identifier id="0DE67FCB64C08303583E70A8CD2EEA03" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="A48556CAC9DE53B050FFDC42C4EFC0FF" type="Zenodo-Dep">13704396</mods:identifier>
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<treatment id="E97A87AEFFA6FFDAFF75FAC6FECD2B30" LSID="urn:lsid:plazi:treatment:E97A87AEFFA6FFDAFF75FAC6FECD2B30" httpUri="http://treatment.plazi.org/id/E97A87AEFFA6FFDAFF75FAC6FECD2B30" lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85">
<subSubSection id="29C96533FFA6FFDBFF75FAC6FC472B48" box="[136,923,1367,1389]" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="616C36B8FFA6FFDBFF75FAC6FC472B48" blockId="0.[136,1451,1367,1497]" box="[136,923,1367,1389]" pageId="0" pageNumber="85">
<taxonomicName id="A6D34D3BFFA6FFDBFF75FAC6FD062B48" ID-CoL="8WR5P" authority="Suetsugu, M. Suleiman &amp; Tsukaya" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[136,730,1367,1389]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="maliauensis" status="sp. nov.">
<emphasis id="53A7EAAAFFA6FFDBFF75FAC6FE582B48" bold="true" box="[136,388,1367,1389]" italics="true" pageId="0" pageNumber="85">Aphyllorchis maliauensis</emphasis>
Suetsugu, M. Suleiman &amp; Tsukaya
</taxonomicName>
,
<emphasis id="53A7EAAAFFA6FFDBFD1EFAC9FCF12B48" box="[739,813,1368,1389]" italics="true" pageId="0" pageNumber="85">
<taxonomicNameLabel id="489457D1FFA6FFDBFD1EFAC9FCF12B48" box="[739,813,1368,1389]" pageId="0" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F9E82A3DFFA6FFDBFCC4FAC9FC4F2B48" box="[825,915,1368,1389]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[136,229,1995,2016]" captionStartId-1="2.[136,229,1987,2008]" captionStartId-2="3.[136,229,1979,2000]" captionTargetBox-0="[177,1411,1322,1967]" captionTargetBox-1="[164,1406,193,1923]" captionTargetBox-2="[164,1388,192,1870]" captionTargetId-0="figure-514@1.[177,1411,1322,1967]" captionTargetId-1="figure-16@2.[164,1423,187,1959]" captionTargetId-2="figure-14@3.[164,1423,187,1951]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Aphyllorchis maliauensis (from the holotype). A. Flowering plant. B. Flower. Photographed by Kenji Suetsugu." captionText-1="FIGURE 2. Aphyllorchis maliauensis (from the holotype).A. Flowering plant. Bar = 3cm. Drawings by Kumi Hamasaki." captionText-2="FIGURE 3. Aphyllorchis maliauensis (from the holotype).A. Flower and ovary. B. Flower. C. Lip and column. D. Dorsal sepal. E. Lateral sepal. F. Lateral petal. G. Lip. H. Column. I. Anther cap. A–B. Scale bar = 1 cm. C–H = 5 mm. I = 1 mm. Drawings by Kumi Hamasaki." figureDoi-0="http://doi.org/10.5281/zenodo.13704398" figureDoi-1="http://doi.org/10.5281/zenodo.13704404" figureDoi-2="http://doi.org/10.5281/zenodo.13704406" httpUri-0="https://zenodo.org/record/13704398/files/figure.png" httpUri-1="https://zenodo.org/record/13704404/files/figure.png" httpUri-2="https://zenodo.org/record/13704406/files/figure.png" pageId="0" pageNumber="85">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA6FFDBFF75FAEDFC812BFC" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="616C36B8FFA6FFDBFF75FAEDFC812BFC" blockId="0.[136,1451,1367,1497]" pageId="0" pageNumber="85">
<collectingCounty id="880D4E34FFA6FFDBFF75FAEDFF652BB4" box="[136,185,1404,1425]" pageId="0" pageNumber="85">Type</collectingCounty>
:—
<materialsCitation id="D1BB3CE5FFA6FFDBFF28FAEDFE722BFC" collectingDate="2017-08-15" collectionCode="BORH" collectorName="Tsukaya &amp; Suetsugu &amp; Anthony" country="Malaysia" elevation="525" latitude="4.6969447" location="Maliau Basin Conservation Area, from Agathis Camp" longLatPrecision="21" longitude="116.91111" pageId="0" pageNumber="85" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="19C47628FFA6FFDBFF28FAEDFE8A2BB4" box="[213,342,1404,1425]" name="Malaysia" pageId="0" pageNumber="85">MALAYSIA</collectingCountry>
.
<collectingRegion id="A317F85AFFA6FFDBFE9CFAEDFE432BB4" box="[353,415,1404,1425]" country="Malaysia" name="Sabah" pageId="0" pageNumber="85">Sabah</collectingRegion>
:
<location id="640C6063FFA6FFDBFE57FAEDFC642BB4" LSID="urn:lsid:plazi:treatment:E97A87AEFFA6FFDAFF75FAC6FECD2B30:640C6063FFA6FFDBFE57FAEDFC642BB4" box="[426,952,1404,1425]" country="Malaysia" latitude="4.6969447" longLatPrecision="21" longitude="116.91111" name="Maliau Basin Conservation Area, from Agathis Camp" pageId="0" pageNumber="85" stateProvince="Sabah">Maliau Basin Conservation Area, from Agathis Camp</location>
(
<geoCoordinate id="04E7507FFFA6FFDBFC35FAEDFBE92BB4" box="[968,1077,1404,1425]" degrees="4" direction="north" minutes="41" orientation="latitude" pageId="0" pageNumber="85" precision="15" seconds="49" value="4.6969447">N4°41’49”</geoCoordinate>
;
<geoCoordinate id="04E7507FFFA6FFDBFBBCFAEDFB1C2BB4" box="[1089,1216,1404,1425]" degrees="116" direction="east" minutes="54" orientation="longitude" pageId="0" pageNumber="85" precision="15" seconds="40" value="116.91111">E116°54’40”</geoCoordinate>
,
<quantity id="A62B9B5DFFA6FFDBFB31FAEDFAD72BB4" box="[1228,1291,1404,1425]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="0" pageNumber="85" unit="m" value="525.0">
<elevation id="EAFED18BFFA6FFDBFB31FAEDFAD72BB4" box="[1228,1291,1404,1425]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="0" pageNumber="85" unit="m" value="525.0">525 m</elevation>
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alt.) to
<location id="640C6063FFA6FFDBFAA6FAEDFF2A2B90" LSID="urn:lsid:plazi:treatment:E97A87AEFFA6FFDAFF75FAC6FECD2B30:640C6063FFA6FFDBFAA6FAEDFF2A2B90" country="Malaysia" county="Type" latitude="4.6969447" longLatPrecision="21" longitude="116.91111" municipality="Maliau Basin Conservation Area" name="Ginseng Camp" pageId="0" pageNumber="85" stateProvince="Sabah">Ginseng Camp</location>
(N4°44’44”; E116°55’05”, 680 m alt.), Maliau Basin, Sabah, Borneo, Malaysia,
<date id="156D1078FFA6FFDBFC0FFA31FB552B90" box="[1010,1161,1440,1461]" pageId="0" pageNumber="85" value="2017-08-15">
<collectingDate id="0529E990FFA6FFDBFC0FFA31FB552B90" box="[1010,1161,1440,1461]" pageId="0" pageNumber="85" value="2017-08-15">15 August 2017</collectingDate>
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,
<emphasis id="53A7EAAAFFA6FFDBFB6CFA31FF262BFC" italics="true" pageId="0" pageNumber="85">
<collectorName id="CC26536EFFA6FFDBFB6CFA31FB3D2B90" box="[1169,1249,1440,1461]" pageId="0" pageNumber="85">Tsukaya</collectorName>
,
<collectorName id="CC26536EFFA6FFDBFB17FA31FA9D2B90" box="[1258,1345,1440,1461]" pageId="0" pageNumber="85">Suetsugu</collectorName>
&amp;
<collectorName id="CC26536EFFA6FFDBFAA6FA31FA772B90" box="[1371,1451,1440,1461]" pageId="0" pageNumber="85">Anthony</collectorName>
TSA-1
</emphasis>
(
<typeStatus id="BE68881AFFA6FFDBFEFBFA55FE832BFC" box="[262,351,1476,1497]" pageId="0" pageNumber="85" type="holotype">holotype</typeStatus>
:
<collectionCode id="07C2AE7DFFA6FFDBFE95FA55FE722BFC" box="[360,430,1476,1497]" pageId="0" pageNumber="85">BORH</collectionCode>
</materialsCitation>
;
<materialsCitation id="D1BB3CE5FFA6FFDBFE4AFA55FC8E2BFC" box="[439,850,1476,1497]" collectingDate="2017-08-15" collectionCode="KYO" collectorName="Tsukaya &amp; Suetsugu &amp; Anthony" country="Malaysia" elevation="525" latitude="4.6969447" location="Maliau Basin Conservation Area, from Agathis Camp" longLatPrecision="21" longitude="116.91111" pageId="0" pageNumber="85" specimenCount="1" stateProvince="Sabah" typeStatus="isotype">
<typeStatus id="BE68881AFFA6FFDBFE4AFA55FDDC2BFC" box="[439,512,1476,1497]" pageId="0" pageNumber="85" type="isotype">isotype</typeStatus>
:
<collectionCode id="07C2AE7DFFA6FFDBFDF4FA55FDE32BFC" box="[521,575,1476,1497]" pageId="0" pageNumber="85">KYO</collectionCode>
, a flower in spirit collection
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA6FFDBFF75F99BFD322862" pageId="0" pageNumber="85" type="diagnosis">
<paragraph id="616C36B8FFA6FFDBFF75F99BFD322862" blockId="0.[136,1452,1546,2075]" pageId="0" pageNumber="85">
<taxonomicName id="A6D34D3BFFA6FFDBFF75F99BFE7F2806" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[136,419,1546,1571]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA6FFDBFF75F99BFE7F2806" box="[136,419,1546,1571]" italics="true" pageId="0" pageNumber="85">Aphyllorchis maliauensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="A6D34D3BFFA6FFDBFDC8F99BFC162806" authority="Reichenbach (1876: 57)" authorityName="sensu Reichenbach" authorityPageNumber="57" authorityYear="1876" box="[565,970,1546,1571]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA6FFDBFDC8F99BFD6A2806" box="[565,694,1546,1571]" italics="true" pageId="0" pageNumber="85">A. montana</emphasis>
<bibRefCitation id="05424B49FFA6FFDBFD40F99BFC162806" author="Reichenbach, H. G." box="[701,970,1546,1571]" pageId="0" pageNumber="85" pagination="1 - 98" refId="ref2424" refString="Reichenbach, H. G. (1876) Orchideae Roezlianae novae seu criticae. Linnaea 41: 1 - 98." type="journal article" year="1876">Reichenbach (1876: 57)</bibRefCitation>
</taxonomicName>
but differs in having larger narrowly ovate to lanceolate sepals and larger lanceolate lateral petals.
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA6FFDAFF40F9C3FBEE2EF8" lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85" type="description">
<paragraph id="616C36B8FFA6FFDAFF40F9C3FBEE2EF8" blockId="0.[136,1452,1546,2075]" lastBlockId="1.[136,1452,160,1301]" lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85">
Terrestrial, mycoheterotrophic herbs. Inflorescence
<quantity id="A62B9B5DFFA6FFDBFCFBF9C3FC58284E" box="[774,900,1618,1643]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="0" pageNumber="85" unit="cm" value="90.0" valueMax="100.0" valueMin="80.0">80–100 cm</quantity>
tall, dark brownish purple,
<quantity id="A62B9B5DFFA6FFDBFB3DF9C3FAC2284E" box="[1216,1310,1618,1643]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="0" pageNumber="85" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
in diameter, bearing 20–30 scale-like leaves. Rhizome condensed, producing, numerous, horizontally or downwardly elongate roots. Rachis
<quantity id="A62B9B5DFFA6FFDBFEDEF90BFE4A2896" box="[291,406,1690,1715]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="0" pageNumber="85" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 cm</quantity>
, glabrous to subglabrous, with c. 30 flowers; floral bracts (linear-)lanceolate, acute, 15–25 ×
<quantity id="A62B9B5DFFA6FFDBFF75F92FFF3B28F2" box="[136,231,1726,1751]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="0" pageNumber="85" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
. Pedicel and ovary slightly tilted upwards or downwards,
<quantity id="A62B9B5DFFA6FFDBFC94F92FFC3E28F2" box="[873,994,1726,1751]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="0" pageNumber="85" unit="mm" value="28.0" valueMax="32.0" valueMin="24.0">24–32 mm</quantity>
long,
<quantity id="A62B9B5DFFA6FFDBFBDBF92FFBB528F2" box="[1062,1129,1726,1751]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="85" unit="mm" value="2.0">2 mm</quantity>
, pubescent. Flowers opening widely. Dorsal sepal ivory, often with purple veins, cymbiform, narrowly ovate to lanceolate, entirely pubescent on the dorsal side, margin entire, 17.0–18.0 ×
<quantity id="A62B9B5DFFA6FFDBFDC1F897FD18293A" box="[572,708,1798,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="0" pageNumber="85" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.7–4.9 mm</quantity>
, acute at the apex; lateral sepal ivory, narrowly ovate to lanceolate, entirely pubescent on the dorsal side, margin entire or minutely denticulate, 17.0–18.0 ×
<quantity id="A62B9B5DFFA6FFDBFBADF8BBFB0B2966" box="[1104,1239,1834,1859]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="0" pageNumber="85" unit="mm" value="4.3" valueMax="4.4" valueMin="4.2">4.2–4.4 mm</quantity>
, slightly outwardly curved and acute at the apex. Lateral petals ivory with purple veins, lanceolate pubescent on the base of the dorsal side, margin entire or minutely denticulate, 16.0–16.5 ×
<quantity id="A62B9B5DFFA6FFDBFD2CF8E3FC8029AE" box="[721,860,1906,1931]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="0" pageNumber="85" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
, slightly outcurved and acute at the apex. Labellum ivory white, often with maroon marking; hypochile trapezoid,
<quantity id="A62B9B5DFFA6FFDBFCAFF807FC07298A" box="[850,987,1942,1967]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="0" pageNumber="85" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
× c.
<quantity id="A62B9B5DFFA6FFDBFBE5F807FBAC298A" box="[1048,1136,1942,1967]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="85" unit="mm" value="2.5">2.5 mm</quantity>
wide, with well developed, erect, obliquely triangular, obtuse to acute side lobes, 2.3–2.5 ×
<quantity id="A62B9B5DFFA6FFDBFC9CF82BFC3529F6" box="[865,1001,1978,2003]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="0" pageNumber="85" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
when flattened; epichile
<quantity id="A62B9B5DFFA6FFDBFAF4F82BFA7029F6" box="[1289,1452,1978,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.1199999999999999" metricValueMin="1.0699999999999998" pageId="0" pageNumber="85" unit="mm" value="10.95" valueMax="11.2" valueMin="10.7">10.7–11.2 mm</quantity>
×
<quantity id="A62B9B5DFFA6FFDBFF5DF84FFEF429D2" box="[160,296,2014,2039]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.4" metricValueMin="6.2" pageId="0" pageNumber="85" unit="mm" value="6.3" valueMax="6.4" valueMin="6.2">6.2–6.4 mm</quantity>
when flattened, trilobed, sidelobes c. 4 ×
<quantity id="A62B9B5DFFA6FFDBFCFAF84FFC9729D2" box="[775,843,2014,2039]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="85" unit="mm" value="2.0">2 mm</quantity>
, rounded, midlobe narrowly triangular
<quantity id="A62B9B5DFFA6FFDBFAF4F84FFA7029D2" box="[1289,1452,2014,2039]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.1199999999999999" metricValueMin="1.0699999999999998" pageId="0" pageNumber="85" unit="mm" value="10.95" valueMax="11.2" valueMin="10.7">10.7–11.2 mm</quantity>
×
<quantity id="A62B9B5DFFA6FFDBFF5DF793FEF5263E" box="[160,297,2050,2075]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="0" pageNumber="85" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
, thickened, erose at the apex. Column pale yellow, incurved, ca.
<quantity id="A62B9B5DFFA6FFDBFBF5F793FB84263E" box="[1032,1112,2050,2075]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="0" pageNumber="85" unit="mm" value="12.0">12 mm</quantity>
long, with slightly protruding or bearing a hook-shaped appendage on ventral side; stigma oblong concave; anther ovate in upper view, 2.0–
<quantity id="A62B9B5DFFA7FFDAFAABFF31FA702E9C" box="[1366,1452,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="86" unit="mm" value="2.2">2.2 mm</quantity>
long, apex obtuse; pollinia 2, soft and mealy, devoid of caudicles. Capsule not seen.
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA7FFDAFF40FF79FD782F6C" pageId="1" pageNumber="86" type="materials_examined">
<paragraph id="616C36B8FFA7FFDAFF40FF79FD782F6C" blockId="1.[136,1452,160,1301]" pageId="1" pageNumber="86">
<emphasis id="53A7EAAAFFA7FFDAFF40FF79FD832F24" bold="true" box="[189,607,232,257]" pageId="1" pageNumber="86">Additional specimen examined:—</emphasis>
<materialsCitation id="D1BB3CE5FFA7FFDAFDA2FF79FD7C2F6C" collectingDate="2015-08-12" collectionCode="BORH" collectorName="Tsukaya &amp; Okada &amp; Hayashi &amp; Suleiman" country="Malaysia" elevation="312" latitude="5.019167" location="Imbak Canyon Conservation Area, from Kuli Station" longLatPrecision="21" longitude="117.040276" municipality="Imbak Canyon Conservation Area" pageId="1" pageNumber="86" specimenCount="1" stateProvince="Sabah">
<collectingCountry id="19C47628FFA7FFDAFDA2FF79FD152F24" box="[607,713,232,257]" name="Malaysia" pageId="1" pageNumber="86">Malaysia</collectingCountry>
.
<collectingRegion id="A317F85AFFA7FFDAFD27FF79FCFF2F24" box="[730,803,232,257]" country="Malaysia" name="Sabah" pageId="1" pageNumber="86">Sabah</collectingRegion>
:
<location id="640C6063FFA7FFDAFCCEFF79FA772F24" LSID="urn:lsid:plazi:treatment:E97A87AEFFA6FFDAFF75FAC6FECD2B30:640C6063FFA7FFDAFCCEFF79FA772F24" box="[819,1451,232,257]" country="Malaysia" latitude="5.019167" longLatPrecision="21" longitude="117.040276" municipality="Imbak Canyon Conservation Area" name="Imbak Canyon Conservation Area, from Kuli Station" pageId="1" pageNumber="86" stateProvince="Sabah">Imbak Canyon Conservation Area, from Kuli Station</location>
(
<geoCoordinate id="04E7507FFFA7FFDAFF6CFE9DFEFE2F00" box="[145,290,268,293]" degrees="05" direction="north" minutes="06" orientation="latitude" pageId="1" pageNumber="86" precision="15" seconds="48" value="5.113333">N05°06’48’’</geoCoordinate>
;
<geoCoordinate id="04E7507FFFA7FFDAFED2FE9DFE612F00" box="[303,445,268,293]" degrees="117" direction="east" minutes="02" orientation="longitude" pageId="1" pageNumber="86" precision="15" seconds="25" value="117.040276">E117°02’25</geoCoordinate>
’,
<quantity id="A62B9B5DFFA7FFDAFE32FE9DFDC72F00" box="[463,539,268,293]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="1" pageNumber="86" unit="m" value="312.0">
<elevation id="EAFED18BFFA7FFDAFE32FE9DFDC72F00" box="[463,539,268,293]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="1" pageNumber="86" unit="m" value="312.0">312 m</elevation>
</quantity>
.) to N05°01’09’’; E117°02’01’’, along the slope trail,
<date id="156D1078FFA7FFDAFB78FE9DFAE12F00" box="[1157,1341,268,293]" pageId="1" pageNumber="86" value="2015-08-12">
<collectingDate id="0529E990FFA7FFDAFB78FE9DFAE12F00" box="[1157,1341,268,293]" pageId="1" pageNumber="86" value="2015-08-12">12 August 2015</collectingDate>
</date>
,
<emphasis id="53A7EAAAFFA7FFDAFAB7FE9DFDE82F6C" italics="true" pageId="1" pageNumber="86">
<collectorName id="CC26536EFFA7FFDAFAB7FE9DFA742F00" box="[1354,1448,268,293]" pageId="1" pageNumber="86">Tsukaya</collectorName>
,
<collectorName id="CC26536EFFA7FFDAFF75FEA1FF092F6C" box="[136,213,304,329]" pageId="1" pageNumber="86">Okada</collectorName>
,
<collectorName id="CC26536EFFA7FFDAFF1CFEA1FE9D2F6C" box="[225,321,304,329]" pageId="1" pageNumber="86">Hayashi</collectorName>
, &amp;
<collectorName id="CC26536EFFA7FFDAFE94FEA1FE0D2F6C" box="[361,465,304,329]" pageId="1" pageNumber="86">Suleiman</collectorName>
TOH-21
</emphasis>
(
<collectionCode id="07C2AE7DFFA7FFDAFDB9FEA1FD442F6C" box="[580,664,304,329]" pageId="1" pageNumber="86">BORH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA7FFDAFF40FEC5FB3A2CF0" pageId="1" pageNumber="86" type="discussion">
<paragraph id="616C36B8FFA7FFDAFF40FEC5FF0D2C60" blockId="1.[136,1452,160,1301]" pageId="1" pageNumber="86">
<emphasis id="53A7EAAAFFA7FFDAFF40FEC5FEFA2F48" bold="true" box="[189,294,339,365]" pageId="1" pageNumber="86">
Notes:
<emphasis id="53A7EAAAFFA7FFDAFEF7FEC2FEFA2F48" box="[266,294,339,365]" italics="true" pageId="1" pageNumber="86">—</emphasis>
</emphasis>
<taxonomicName id="A6D34D3BFFA7FFDAFEDBFEC5FD9D2F48" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[294,577,340,365]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">Aphyllorchis maliauensis</taxonomicName>
is similar to
<taxonomicName id="A6D34D3BFFA7FFDAFD2EFEC5FC882F48" authorityName="sensu Reichenbach" authorityYear="1876" box="[723,852,340,365]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFD2EFEC5FC882F48" box="[723,852,340,365]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
but differs in the shape and length of the dorsal sepal (narrowly ovate to lanceolate, 17.0–18.0 ×
<quantity id="A62B9B5DFFA7FFDAFD86FEE9FCDA2FB4" box="[635,774,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="1" pageNumber="86" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.7–4.9 mm</quantity>
, acute at the apex vs. oblong to obovate, 9–13 ×
<quantity id="A62B9B5DFFA7FFDAFAB8FEE9FA7B2FB4" box="[1349,1447,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="86" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, rounded to obtuse at the apex), lateral sepal (narrowly ovate to lanceolate, 17.0–18.0 ×
<quantity id="A62B9B5DFFA7FFDAFBA6FE0DFB392F90" box="[1115,1253,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="1" pageNumber="86" unit="mm" value="4.3" valueMax="4.4" valueMin="4.2">4.2–4.4 mm</quantity>
, acute at the apex vs. (elliptic-) oblong, 9–12 ×
<quantity id="A62B9B5DFFA7FFDAFE21FE51FDE12FFC" box="[476,573,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="86" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, obtuse at the apex), and lateral petals (lanceolate, 16.0–16.5 ×
<quantity id="A62B9B5DFFA7FFDAFAE1FE51FA7B2FFC" box="[1308,1447,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="1" pageNumber="86" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
, slightly out curving and acute at the apex vs. falcately linear-oblong 8.0–11.0 ×
<quantity id="A62B9B5DFFA7FFDAFBDAFE75FB6E2FD8" box="[1063,1202,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="1" pageNumber="86" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 mm</quantity>
, rounded to obtuse at the apex) and the colour of the tepals (ivory white often with purple veins vs. yellowish white;
<bibRefCitation id="05424B49FFA7FFDAFB4BFD99FA5F2C04" author="Seidenfaden, G." box="[1206,1411,520,545]" pageId="1" pageNumber="86" pagination="1 - 195" refId="ref2571" refString="Seidenfaden, G. (1978) Orchid genera in Thailand VI. Neottioideae Lindl. Dansk Botanisk Arkiv 32: 1 - 195." type="journal article" year="1978">Seidenfaden 1978</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFA73FD99FF192C60" author="Su, H. J." pageId="1" pageNumber="86" pagination="749 - 751" refId="ref2627" refString="Su, H. J. (2000) Aphyllorchis. In: Huang, T. C., Boufford, D. E., Hsieh, C. F., Kuoh, C. S., Ohashi, H. &amp; Su, H. J. (Eds.) Flora of Taiwan, 2 nd ed., 5. Department of Botany, National Taiwan University, Taipei, pp. 749 - 751." type="book chapter" year="2000">Su 2000</bibRefCitation>
).
</paragraph>
<paragraph id="616C36B8FFA7FFDAFF40FDC1FB3A2CF0" blockId="1.[136,1452,160,1301]" pageId="1" pageNumber="86">
In addition,
<taxonomicName id="A6D34D3BFFA7FFDAFEB5FDC1FE322C4C" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[328,494,592,617]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFEB5FDC1FE322C4C" box="[328,494,592,617]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
also differs clearly from the taxa that are now considered synonyms of
<taxonomicName id="A6D34D3BFFA7FFDAFADFFDC1FA782C4C" authorityName="sensu Reichenbach" authorityYear="1876" box="[1314,1444,592,617]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFADFFDC1FA782C4C" box="[1314,1444,592,617]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
, i.e.,
<taxonomicName id="A6D34D3BFFA7FFDAFF47FDE5FDE42CA8" authority="Ames (1908: 48)" authorityName="Ames" authorityPageNumber="48" authorityYear="1908" box="[186,568,628,653]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="benguetensis">
<emphasis id="53A7EAAAFFA7FFDAFF47FDE5FEB02CA8" box="[186,364,628,653]" italics="true" pageId="1" pageNumber="86">A. benguetensis</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFE8EFDE5FDE42CA8" author="Ames, O." box="[371,568,628,653]" pageId="1" pageNumber="86" pagination="17 - 257" refId="ref2015" refString="Ames, O. (1908) Studies in the orchid flora of the Philippines. In: Ames, O. (Aut.) Orchidaceae, illustrations and studies of the family Orchidaceae 2. Ames Botanical Laboratory, North Easton, pp. 17 - 257." type="book chapter" year="1908">Ames (1908: 48)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A6D34D3BFFA7FFDAFDB8FDE5FC332CA8" authority="Schlechter (1906: 299)" authorityName="Schlechter" authorityPageNumber="299" authorityYear="1906" box="[581,1007,628,653]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="borneensis">
<emphasis id="53A7EAAAFFA7FFDAFDB8FDE5FD032CA8" box="[581,735,628,653]" italics="true" pageId="1" pageNumber="86">A. borneensis</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFD15FDE5FC332CA8" author="Schlechter, R." box="[744,1007,628,653]" pageId="1" pageNumber="86" pagination="295 - 310" refId="ref2541" refString="Schlechter, R. (1906) Neue Orchidaceen der Flora des Monsun - Gebietes. Bulletin de l'Herbier Boissier, ser. 2, 4: 295 - 310" type="journal article" year="1906">Schlechter (1906: 299)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A6D34D3BFFA7FFDAFC01FDE5FA7B2CA8" authority="Reichenbach (1886: 345)" authorityName="Reichenbach" authorityPageNumber="345" authorityYear="1886" box="[1020,1447,628,653]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="odoardii">
<emphasis id="53A7EAAAFFA7FFDAFC01FDE5FBA02CA8" box="[1020,1148,628,653]" italics="true" pageId="1" pageNumber="86">A. odoardii</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFB78FDE5FA7B2CA8" author="Reichenbach, H. G." box="[1157,1447,628,653]" pageId="1" pageNumber="86" pagination="343 - 346" refId="ref2446" refString="Reichenbach, H. G. (1886) Odoardi Beccarii novitiae Orchidaceae Papuanae describuntur. Botanisches Centralblatt 28: 343 - 346." type="journal article" year="1886">Reichenbach (1886: 345)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A6D34D3BFFA7FFDAFF75FD09FE3F2C94" authority="Hooker (1890: 117)" authorityName="Hooker" authorityPageNumber="117" authorityYear="1890" box="[136,483,664,689]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="prainii">
<emphasis id="53A7EAAAFFA7FFDAFF75FD09FF292C94" box="[136,245,664,689]" italics="true" pageId="1" pageNumber="86">A. prainii</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFF02FD09FE3F2C94" author="Hooker, J. D." box="[255,483,664,689]" pageId="1" pageNumber="86" pagination="1 - 198" refId="ref2287" refString="Hooker, J. D. (1890) Orchidaceae. In: Hooker, J. D. (Ed.) Flora of British India, vol. 6. Reeve, London, pp. 1 - 198." type="book chapter" year="1890">Hooker (1890: 117)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A6D34D3BFFA7FFDAFE0CFD09FC562C94" authority="Fukuyama (1934: 431)" authorityName="Fukuyama" authorityPageNumber="431" authorityYear="1934" box="[497,906,664,689]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="purpurea">
<emphasis id="53A7EAAAFFA7FFDAFE0CFD09FDA52C94" box="[497,633,664,689]" italics="true" pageId="1" pageNumber="86">A. purpurea</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFD7FFD09FC562C94" author="Fukuyama, N." box="[642,906,664,689]" pageId="1" pageNumber="86" pagination="429 - 442" refId="ref2224" refString="Fukuyama, N. (1934) Studia orchidacearum Japonicarum III. Botanical Magazine Tokyo 48: 429 - 442. https: // doi. org / 10.15281 / jplantres 1887.48.429" type="journal article" year="1934">Fukuyama (1934: 431)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A6D34D3BFFA7FFDAFC65FD09FA852C94" authority="Hayata (1911: 344)" authorityName="Hayata" authorityPageNumber="344" authorityYear="1911" box="[920,1369,664,689]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="tanegashimensis">
<emphasis id="53A7EAAAFFA7FFDAFC65FD09FBAE2C94" box="[920,1138,664,689]" italics="true" pageId="1" pageNumber="86">A. tanegashimensis</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFB86FD09FA852C94" author="Hayata, B." box="[1147,1369,664,689]" pageId="1" pageNumber="86" pagination="1 - 471" refId="ref2256" refString="Hayata, B. (1911) Material for a flora of Formosa. Journal of the College of Science, Imperial University of Tokyo 30: 1 - 471." type="journal article" year="1911">Hayata (1911: 344)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="A6D34D3BFFA7FFDAFA6EFD09FDB82CF0" authority="Rolfe ex Downie (1925: 415)" authorityName="Downie" authorityPageNumber="415" authorityYear="1925" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="unguiculata">
<emphasis id="53A7EAAAFFA7FFDAFA6EFD09FED22CF0" italics="true" pageId="1" pageNumber="86">A. unguiculata</emphasis>
Rolfe ex
<bibRefCitation id="05424B49FFA7FFDAFE86FD2DFDB82CF0" author="Downie, D. G." box="[379,612,700,725]" pageId="1" pageNumber="86" pagination="404 - 423" refId="ref2188" refString="Downie, D. G. (1925) Contributions to the flora of Siam. Bulletin of Miscellaneous Information 1925: 404 - 423. https: // doi. org / 10.2307 / 4115102" type="journal article" year="1925">Downie (1925: 415)</bibRefCitation>
</taxonomicName>
], all of which are morphologically similar to
<taxonomicName id="A6D34D3BFFA7FFDAFBA0FD2DFB032CF0" authorityName="sensu Reichenbach" authorityYear="1876" box="[1117,1247,700,725]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFBA0FD2DFB032CF0" box="[1117,1247,700,725]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="29C96533FFA7FFDAFF40FD71FECD2B30" pageId="1" pageNumber="86" type="distribution">
<paragraph id="616C36B8FFA7FFDAFF40FD71FB0C2DD0" blockId="1.[136,1452,160,1301]" pageId="1" pageNumber="86">
<emphasis id="53A7EAAAFFA7FFDAFF40FD71FCEE2CDC" bold="true" box="[189,818,735,761]" pageId="1" pageNumber="86">
Distribution, phenology, and conservation status:
<emphasis id="53A7EAAAFFA7FFDAFCEBFD4EFCEE2CDC" bold="true" box="[790,818,735,761]" italics="true" pageId="1" pageNumber="86">—</emphasis>
</emphasis>
Fewer than 10 individuals of
<taxonomicName id="A6D34D3BFFA7FFDAFB70FD71FA702CDC" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[1165,1452,736,761]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFB70FD71FA702CDC" box="[1165,1452,736,761]" italics="true" pageId="1" pageNumber="86">Aphyllorchis maliauensis</emphasis>
</taxonomicName>
were observed at two locations in the wet understory of a kerangas forest in the Maliau Basin Conservation Area, which is dominated by
<taxonomicName id="A6D34D3BFFA7FFDAFEBEFCB9FE2D2D64" box="[323,497,808,833]" pageId="1" pageNumber="86">
<emphasis id="53A7EAAAFFA7FFDAFEBEFCB9FE452D64" box="[323,409,808,833]" italics="true" pageId="1" pageNumber="86">Agathis</emphasis>
species
</taxonomicName>
and dipterocarps. In addition, only one individual of
<taxonomicName id="A6D34D3BFFA7FFDAFBB7FCB9FB332D64" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[1098,1263,808,833]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFBB7FCB9FB332D64" box="[1098,1263,808,833]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
was observed in the Imbak Canyon Conservation Area,
<collectingRegion id="A317F85AFFA7FFDAFDBBFCDDFD512D40" box="[582,653,844,869]" country="Malaysia" name="Sabah" pageId="1" pageNumber="86">Sabah</collectingRegion>
, under a lowland dipterocarp forest.
<taxonomicName id="A6D34D3BFFA7FFDAFBCFFCDDFA932D40" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[1074,1359,844,869]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFBCFFCDDFA932D40" box="[1074,1359,844,869]" italics="true" pageId="1" pageNumber="86">Aphyllorchis maliauensis</emphasis>
</taxonomicName>
flowers in mid-August in both populations. Furthermore, it is likely that
<taxonomicName id="A6D34D3BFFA7FFDAFC93FCE1FBCA2DAC" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[878,1046,880,905]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFC93FCE1FBCA2DAC" box="[878,1046,880,905]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
is distributed more widely because it seems to have sometimes been misidentified as the more widespread species,
<taxonomicName id="A6D34D3BFFA7FFDAFBEDFC05FB4E2D88" authorityName="sensu Reichenbach" authorityYear="1876" box="[1040,1170,916,941]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFBEDFC05FB4E2D88" box="[1040,1170,916,941]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
, which has similar gross morphology. For example, the
<taxonomicName id="A6D34D3BFFA7FFDAFE20FC29FD4E2DF4" box="[477,658,952,977]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="53A7EAAAFFA7FFDAFE20FC29FDB72DF4" box="[477,619,952,977]" italics="true" pageId="1" pageNumber="86">Aphyllorchis</emphasis>
sp.
</taxonomicName>
recorded as
<taxonomicName id="A6D34D3BFFA7FFDAFCE0FC29FC412DF4" authorityName="sensu Reichenbach" authorityYear="1876" box="[797,925,952,977]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFCE0FC29FC412DF4" box="[797,925,952,977]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
by
<bibRefCitation id="05424B49FFA7FFDAFC38FC29FB752DF4" author="Beaman, T. E. &amp; Wood, J. J. &amp; Beaman, R. S. &amp; Beaman, J. H." box="[965,1193,952,977]" pageId="1" pageNumber="86" refId="ref2115" refString="Beaman, T. E., Wood, J. J., Beaman, R. S. &amp; Beaman, J. H. (2001) Orchids of Sarawak. Natural History Publications, Kota Kinabalu, 584 pp." type="book" year="2001">
Beaman
<emphasis id="53A7EAAAFFA7FFDAFBDAFC29FB832DF4" box="[1063,1119,952,977]" italics="true" pageId="1" pageNumber="86">et al.</emphasis>
(2001
</bibRefCitation>
;
<collectingCountry id="19C47628FFA7FFDAFB49FC29FACB2DF4" box="[1204,1303,952,977]" name="Malaysia" pageId="1" pageNumber="86">Sarawak</collectingCountry>
, Borneo) and
<bibRefCitation id="05424B49FFA7FFDAFF75FC4DFECA2DD0" author="Wood, J. J." box="[136,278,988,1013]" pageId="1" pageNumber="86" refId="ref2932" refString="Wood, J. J. (2003) Orchids of Borneo, vol. 4, revised classification and selection of species. Sabah Society and Kew Publishing, Sabah, 314 pp." type="book" year="2003">Wood (2003</bibRefCitation>
;
<collectingRegion id="A317F85AFFA7FFDAFEDCFC4DFEB42DD0" box="[289,360,988,1013]" country="Malaysia" name="Sabah" pageId="1" pageNumber="86">Sabah</collectingRegion>
, Borneo) appears identical to
<taxonomicName id="A6D34D3BFFA7FFDAFD4AFC4DFC802DD0" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[695,860,988,1013]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFD4AFC4DFC802DD0" box="[695,860,988,1013]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
, judging from their photographs.
</paragraph>
<paragraph id="616C36B8FFA7FFDAFF40FB91FECD2B30" blockId="1.[136,1452,160,1301]" pageId="1" pageNumber="86">
However, the original descriptions and subsequent literature clearly indicate that there is little morphological variation in
<taxonomicName id="A6D34D3BFFA7FFDAFEE4FBB5FD022A18" authority="sensu Reichenbach (1876)" authorityName="sensu Reichenbach" authorityYear="1876" box="[281,734,1060,1085]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFEE4FBB5FE342A18" box="[281,488,1060,1085]" italics="true" pageId="1" pageNumber="86">A. montana sensu</emphasis>
<bibRefCitation id="05424B49FFA7FFDAFE09FBB5FD022A18" author="Reichenbach, H. G." box="[500,734,1060,1085]" pageId="1" pageNumber="86" pagination="1 - 98" refId="ref2424" refString="Reichenbach, H. G. (1876) Orchideae Roezlianae novae seu criticae. Linnaea 41: 1 - 98." type="journal article" year="1876">Reichenbach (1876)</bibRefCitation>
</taxonomicName>
, despite its extensive distribution, which includes
<collectingCountry id="19C47628FFA7FFDAFACFFBB5FA742A18" box="[1330,1448,1060,1085]" name="Sri Lanka" pageId="1" pageNumber="86">Sri Lanka</collectingCountry>
,
<collectingRegion id="A317F85AFFA7FFDAFF75FBD9FF0B2A44" box="[136,215,1096,1121]" country="India" name="Assam" pageId="1" pageNumber="86">Assam</collectingRegion>
, southern
<taxonomicName id="A6D34D3BFFA7FFDAFEB1FBD9FE542A44" box="[332,392,1096,1121]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="89" phylum="Tracheophyta" rank="genus">India</taxonomicName>
, the
<collectingCountry id="19C47628FFA7FFDAFE42FBD9FD9C2A44" box="[447,576,1096,1121]" name="Philippines" pageId="1" pageNumber="86">Philippines</collectingCountry>
, Borneo,
<collectingCountry id="19C47628FFA7FFDAFD50FBD9FCCF2A44" box="[685,787,1096,1121]" name="Thailand" pageId="1" pageNumber="86">Thailand</collectingCountry>
,
<collectingCountry id="19C47628FFA7FFDAFCDDFBD9FCA92A44" box="[800,885,1096,1121]" name="Taiwan" pageId="1" pageNumber="86">Taiwan</collectingCountry>
,
<collectingCountry id="19C47628FFA7FFDAFC7CFBD9FC1D2A44" box="[897,961,1096,1121]" name="Japan" pageId="1" pageNumber="86">Japan</collectingCountry>
and
<collectingCountry id="19C47628FFA7FFDAFC04FBD9FBBA2A44" box="[1017,1126,1096,1121]" name="Indonesia" pageId="1" pageNumber="86">Indonesia</collectingCountry>
(
<bibRefCitation id="05424B49FFA7FFDAFB8BFBD9FA972A44" author="Reichenbach, H. G." box="[1142,1355,1096,1121]" pageId="1" pageNumber="86" pagination="343 - 346" refId="ref2446" refString="Reichenbach, H. G. (1886) Odoardi Beccarii novitiae Orchidaceae Papuanae describuntur. Botanisches Centralblatt 28: 343 - 346." type="journal article" year="1886">Reichenbach 1886</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFAAAFBD9FF1F2AA0" author="Hooker, J. D." pageId="1" pageNumber="86" pagination="1 - 198" refId="ref2287" refString="Hooker, J. D. (1890) Orchidaceae. In: Hooker, J. D. (Ed.) Flora of British India, vol. 6. Reeve, London, pp. 1 - 198." type="book chapter" year="1890">Hooker 1890</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFF32FBFDFE542AA0" author="Schlechter, R." box="[207,392,1132,1157]" pageId="1" pageNumber="86" pagination="295 - 310" refId="ref2541" refString="Schlechter, R. (1906) Neue Orchidaceen der Flora des Monsun - Gebietes. Bulletin de l'Herbier Boissier, ser. 2, 4: 295 - 310" type="journal article" year="1906">Schlechter 1906</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFE6EFBFDFDCB2AA0" author="Ames, O." box="[403,535,1132,1157]" pageId="1" pageNumber="86" pagination="17 - 257" refId="ref2015" refString="Ames, O. (1908) Studies in the orchid flora of the Philippines. In: Ames, O. (Aut.) Orchidaceae, illustrations and studies of the family Orchidaceae 2. Ames Botanical Laboratory, North Easton, pp. 17 - 257." type="book chapter" year="1908">Ames 1908</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFDDEFBFDFD6F2AA0" author="Hayata, B." box="[547,691,1132,1157]" pageId="1" pageNumber="86" pagination="1 - 471" refId="ref2256" refString="Hayata, B. (1911) Material for a flora of Formosa. Journal of the College of Science, Imperial University of Tokyo 30: 1 - 471." type="journal article" year="1911">Hayata 1911</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFD3DFBFDFC872AA0" author="Downie, D. G." box="[704,859,1132,1157]" pageId="1" pageNumber="86" pagination="404 - 423" refId="ref2188" refString="Downie, D. G. (1925) Contributions to the flora of Siam. Bulletin of Miscellaneous Information 1925: 404 - 423. https: // doi. org / 10.2307 / 4115102" type="journal article" year="1925">Downie 1925</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFC9BFBFDFBC32AA0" author="Fukuyama, N." box="[870,1055,1132,1157]" pageId="1" pageNumber="86" pagination="429 - 442" refId="ref2224" refString="Fukuyama, N. (1934) Studia orchidacearum Japonicarum III. Botanical Magazine Tokyo 48: 429 - 442. https: // doi. org / 10.15281 / jplantres 1887.48.429" type="journal article" year="1934">Fukuyama 1934</bibRefCitation>
,
<bibRefCitation id="05424B49FFA7FFDAFBD6FBFDFB012AA0" author="Roy, M. &amp; Watthana, S. &amp; Stier, A. &amp; Richard, F. &amp; Vessabutr, S. &amp; Selosse, M." box="[1067,1245,1132,1157]" pageId="1" pageNumber="86" refId="ref2470" refString="Roy, M., Watthana, S., Stier, A., Richard, F., Vessabutr, S. &amp; Selosse, M. (2009) Two mycoheterotrophic orchids from Thailand tropical dipterocarpacean forests associate with a broad diversity of ectomycorrhizal fungi. BMC biology 7: 51. https: // doi. org / 10.1186 / 1741 - 7007 - 7 - 51" type="journal volume" year="2009">
Roy
<emphasis id="53A7EAAAFFA7FFDAFB9CFBFDFB462AA0" box="[1121,1178,1132,1157]" italics="true" pageId="1" pageNumber="86">et al.</emphasis>
2009
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,
<bibRefCitation id="05424B49FFA7FFDAFB1AFBFDFF1F2A8C" author="Aravindhan, V. &amp; Mahendran, G. &amp; Rajendran, A. &amp; Bai, N." pageId="1" pageNumber="86" pagination="153 - 155" refId="ref2065" refString="Aravindhan, V., Mahendran, G., Rajendran, A. &amp; Bai, N. (2013) Aphyllorchis montana Rchb. f. (Orchidaceae), a new distributional record for Peninsular India. Annals of Plant Sciences 2: 153 - 155." type="journal article" year="2013">
Aravindhan
<emphasis id="53A7EAAAFFA7FFDAFA8EFBFDFA702AA0" box="[1395,1452,1132,1157]" italics="true" pageId="1" pageNumber="86">et al.</emphasis>
2013
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,
<bibRefCitation id="05424B49FFA7FFDAFF2DFB01FE6A2A8C" author="Rao, A. &amp; Kumar, V." box="[208,438,1168,1193]" pageId="1" pageNumber="86" pagination="527 - 530" refId="ref2390" refString="Rao, A. &amp; Kumar, V. (2015) Record of holomycotrophic Aphyllorchis montana HG Reichenbach (Orchidaceae) from Manipur, India. Pleione 9: 527 - 530." type="journal article" year="2015">Rao &amp; Kumar 2015</bibRefCitation>
,
<figureCitation id="F9E82A3DFFA7FFDAFE3EFB01FDD52A8C" box="[451,521,1168,1193]" captionStart="FIGURE 4" captionStartId="4.[136,229,918,939]" captionTargetBox="[152,1436,187,890]" captionTargetId="figure-16@4.[151,1436,187,891]" captionTargetPageId="4" captionText="FIGURE 4. Aphyllorchis montana in Okinawa Prefecture, Japan on 25 August 2009 (Obara KS217, TNS).A. Flowering plant. B. Flower. Photographed by Yuji Obara." figureDoi="http://doi.org/10.5281/zenodo.13704408" httpUri="https://zenodo.org/record/13704408/files/figure.png" pageId="1" pageNumber="86">Fig. 4</figureCitation>
). Because the morphological differences between
<taxonomicName id="A6D34D3BFFA7FFDAFBB5FB01FB322A8C" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[1096,1262,1168,1193]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFBB5FB01FB322A8C" box="[1096,1262,1168,1193]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A6D34D3BFFA7FFDAFAD5FB01FA772A8C" authorityName="sensu Reichenbach" authorityYear="1876" box="[1320,1451,1168,1193]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="53A7EAAAFFA7FFDAFAD5FB01FA772A8C" box="[1320,1451,1168,1193]" italics="true" pageId="1" pageNumber="86">A. montana</emphasis>
</taxonomicName>
are clear and stable,
<taxonomicName id="A6D34D3BFFA7FFDAFE8FFB25FDC42AE8" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[370,536,1204,1229]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFE8FFB25FDC42AE8" box="[370,536,1204,1229]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
should be treated as an independent species rather than as an infraspecific taxon of
<emphasis id="53A7EAAAFFA7FFDAFF5BFB49FEF32AD4" box="[166,303,1240,1265]" italics="true" pageId="1" pageNumber="86">
<taxonomicName id="A6D34D3BFFA7FFDAFF5BFB49FEF72AD4" authorityName="sensu Reichenbach" authorityYear="1876" box="[166,299,1240,1265]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="montana">A. montana</taxonomicName>
.
</emphasis>
Conducting additional surveys for
<taxonomicName id="A6D34D3BFFA7FFDAFD44FB49FC822AD4" authorityName="Suetsugu, M. Suleiman &amp; Tsukaya" authorityYear="2018" box="[697,862,1240,1265]" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="maliauensis">
<emphasis id="53A7EAAAFFA7FFDAFD44FB49FC822AD4" box="[697,862,1240,1265]" italics="true" pageId="1" pageNumber="86">A. maliauensis</emphasis>
</taxonomicName>
during the flowering season would reveal its precise distribution.
</paragraph>
</subSubSection>
</treatment>
</document>