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<document id="4E4058B2F517E616CFF5FEE2C2E9B98E" ID-DOI="10.1206/0003-0082(2004)459<0001:ACONDP>2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5382708" IM.bibliography_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1630295385096" checkinUser="carolina" docAuthor="SPARKS, JOHN S. & DUNLAP, PAUL V." docDate="2004" docId="03C7A2794C77FFBEBAF5FCC6FB44E722" docLanguage="en" docName="N3459.pdf" docOrigin="American Museum Novitates 3459 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2004)459%3C0001%3AACONDP%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.4:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="4" docTitle="Leiognathus robustus SPARKS & DUNLAP 2004, new species" docType="treatment" docUuid="82735940-419C-4A60-AA82-3BECF5789B1B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="9" masterDocId="FFFEDA014C72FFB6BA5EFF93FFE2E51A" masterDocTitle="A Clade of Non-Sexually Dimorphic Ponyfishes (Teleostei: Perciformes: Leiognathidae): Phylogeny, Taxonomy, and Description of a New Species" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1729786802594" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="DE0311378A70F28F1EB600B514E5E56D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F6F601A45B156E4D79C9A1FA9117308D">
|
||||
<mods:title id="EC8D604BF312C9B514A02A46EBE22A7A">A Clade of Non-Sexually Dimorphic Ponyfishes (Teleostei: Perciformes: Leiognathidae): Phylogeny, Taxonomy, and Description of a New Species</mods:title>
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</mods:titleInfo>
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<mods:name id="8283F975752BA1BCCA03535A764C3295" type="personal">
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<mods:roleTerm id="5055475E7C88B65FB206C90C0CD1FADD">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="131E0AB0F90794343645EC3F7224A904">SPARKS, JOHN S.</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="150C62BBAC5B2F0ACD364B1073C5B30F" type="personal">
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<mods:roleTerm id="DE026C193BE1AF93CE0D580CC2C2B67E">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="9872D288D424C91B29D0E5F86AE9A8BB">DUNLAP, PAUL V.</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="52D1FF2449976E7F945D0D1CEBD04719">text</mods:typeOfResource>
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<mods:titleInfo id="99C59E7B2473AD36A1CF61C348B2AD27">
|
||||
<mods:title id="85A0A4DF647124DC69391352B5C782FB">American Museum Novitates</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="8446FA68A037DFB5E18D89055086FAC8">
|
||||
<mods:date id="83A48EDC5909AC824D8769DAEB5C65AA">2004</mods:date>
|
||||
<mods:detail id="E838937761D76CCAF82808A432B5EC34" type="pubDate">
|
||||
<mods:number id="C8FA8FFC35A2BFC113700956FBF26C31">2004-10-28</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="4B4979F950F42FD290AD577F543E9DC1" type="volume">
|
||||
<mods:number id="BDEFB3CEE06566515605CBBA86F3885F">3459</mods:number>
|
||||
</mods:detail>
|
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<mods:detail id="45382B669B6824F65BED1417EC4C10B2" type="issue">
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<mods:number id="45AF2ABF8FD2491E0CA531B9D295F02A">1</mods:number>
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<mods:url id="3F001F325F7CC2BF4FA7207E0DDC2E43">http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2004)459%3C0001%3AACONDP%3E2.0.CO%3B2</mods:url>
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</mods:location>
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<mods:classification id="DD2B55634B3F6D20D865FBC16A673342">journal article</mods:classification>
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<mods:identifier id="83FC8C6C930F3816880A2D8C5AD7AC40" type="DOI">10.1206/0003-0082(2004)459<0001:ACONDP>2.0.CO;2</mods:identifier>
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<mods:identifier id="E953D110D4F441EB13C52BBFE2E10D77" type="ISSN">0003-0082</mods:identifier>
|
||||
<mods:identifier id="11B90C10B874139AF53DFF2E03071AF4" type="Zenodo-Dep">5382708</mods:identifier>
|
||||
<mods:identifier id="11A91119B2D55DA7093798A058EB1C55" type="ZooBank">8A7446E5-53BB-4F88-B048-2A66D44C87F3</mods:identifier>
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</mods:mods>
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<treatment id="03C7A2794C77FFBEBAF5FCC6FB44E722" LSID="urn:lsid:zoobank.org:act:82735940-419C-4A60-AA82-3BECF5789B1B" httpUri="http://treatment.plazi.org/id/03C7A2794C77FFBEBAF5FCC6FB44E722" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
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<subSubSection id="C37440E44C77FFB3BAF5FCC6FDB3E676" box="[171,593,853,877]" pageId="5" pageNumber="6" type="nomenclature">
|
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<paragraph id="8BD1136F4C77FFB3BAF5FCC6FDB3E676" blockId="5.[171,593,853,877]" box="[171,593,853,877]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B91ACF7D4C77FFB3BAF5FCC6FE5BE677" bold="true" box="[171,441,853,877]" pageId="5" pageNumber="6">
|
||||
<taxonomicName id="4C6E68EC4C77FFB3BAF5FCC6FE50E677" ID-CoL="3SYPG" authority="SPARKS & DUNLAP, 2004" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[171,434,853,877]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="B91ACF7D4C77FFB3BAF5FCC6FE50E677" bold="true" box="[171,434,853,877]" italics="true" pageId="5" pageNumber="6">Leiognathus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A22972064C77FFB3BB9DFCC6FDB3E676" box="[451,593,853,876]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C77FFB3BB1DFCE7FE20E693" box="[323,450,884,905]" pageId="5" pageNumber="6" type="description">
|
||||
<paragraph id="8BD1136F4C77FFB3BB1DFCE7FE20E693" blockId="5.[323,450,884,905]" box="[323,450,884,905]" pageId="5" pageNumber="6">
|
||||
<heading id="D099A4034C77FFB3BB1DFCE7FE20E693" box="[323,450,884,905]" centered="true" fontSize="36" level="3" pageId="5" pageNumber="6" reason="7">
|
||||
<figureCitation id="13550FEA4C77FFB3BB1DFCE7FE20E693" box="[323,450,884,905]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="9.[149,187,1696,1717]" captionStartId-1="10.[159,197,826,847]" captionStartId-2="11.[149,187,1134,1155]" captionStartId-3="12.[150,188,1394,1415]" captionTargetBox-0="[125,1224,191,1673]" captionTargetBox-1="[125,1229,187,805]" captionTargetBox-2="[124,651,188,1112]" captionTargetBox-3="[125,1229,187,1373]" captionTargetId-0="figure-10@9.[123,1227,187,1674]" captionTargetId-1="figure-17@10.[125,1229,187,805]" captionTargetId-2="figure-13@11.[123,651,187,1113]" captionTargetId-3="figure-15@12.[125,1229,187,1373]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="Fig. 3. Leiognathus robustus, holotype, UMMZ 242144, 183.4 mm SL, adult male; Singapore. A. External anatomy, illustrating general pigmentation pattern, and absence of transparent flank or opercular" captionText-1="Fig. 4. Leiognathus robustus, holotype, UMMZ 242144, adult male, 183.4 mm SL, Singapore." captionText-2="Fig. 5. Left lateral view of head of: (A) Leiognathus robustus, holotype, UMMZ 242144, adult male, 183.4 mm SL; (B) Leiognathus equulus, UMMZ 238805, adult male, 173.4 mm SL. Arrow indicates nuchal spine." captionText-3="Fig. 6. Comparative radiographs of similarly sized (A) Leiognathus robustus, holotype, UMMZ 242144, adult male, 183.4 mm SL, and (B) Leiognathus equulus, UMMZ 238805, adult male, 173.4 mm SL." figureDoi-0="http://doi.org/10.5281/zenodo.5382714" figureDoi-1="http://doi.org/10.5281/zenodo.5382718" figureDoi-2="http://doi.org/10.5281/zenodo.5382720" figureDoi-3="http://doi.org/10.5281/zenodo.5382722" httpUri-0="https://zenodo.org/record/5382714/files/figure.png" httpUri-1="https://zenodo.org/record/5382718/files/figure.png" httpUri-2="https://zenodo.org/record/5382720/files/figure.png" httpUri-3="https://zenodo.org/record/5382722/files/figure.png" pageId="5" pageNumber="6">Figures 3–6</figureCitation>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C77FFB3BAC6FC0DFECEE199" pageId="5" pageNumber="6" type="materials_examined">
|
||||
<paragraph id="8BD1136F4C77FFB3BAC6FC0DFEFCE6EA" blockId="5.[123,652,926,1742]" pageId="5" pageNumber="6">
|
||||
<materialsCitation id="3B0619324C77FFB3BAC6FC0DFEFCE6EA" collectingDate="2001-07-16" collectorName="H. H. Ng" country="Singapore" location="Singapore" pageId="5" pageNumber="6" specimenCount="1" typeStatus="holotype">
|
||||
<typeStatus id="54D5ADCD4C77FFB3BAC6FC0DFEECE6AF" box="[152,270,926,949]" pageId="5" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
|
||||
: UMMZ
|
||||
<quantity id="4C96BE8A4C77FFB3BB21FC0DFDB8E6AF" box="[383,602,926,949]" metricMagnitude="5" metricUnit="m" metricValue="2.421441834" pageId="5" pageNumber="6" unit="mm" value="2.421441834E8">242144, 183.4 mm</quantity>
|
||||
SL, adult male;
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||||
<collectingCountry id="F37953FF4C77FFB3BB58FC2FFE62E6C9" box="[262,384,956,979]" name="Singapore" pageId="5" pageNumber="6">Singapore</collectingCountry>
|
||||
: fish market;
|
||||
<collectorName id="269B76B94C77FFB3B87EFC2FFD64E6C9" box="[544,646,956,979]" pageId="5" pageNumber="6">H.H. Ng</collectorName>
|
||||
,
|
||||
<date id="FFD035AF4C77FFB3BA25FC4AFEF8E6EA" box="[123,282,985,1008]" pageId="5" pageNumber="6" value="2001-07-16">
|
||||
<collectingDate id="EF94CC474C77FFB3BA25FC4AFEF8E6EA" box="[123,282,985,1008]" pageId="5" pageNumber="6" value="2001-07-16">16 July 2001</collectingDate>
|
||||
</date>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BD1136F4C77FFB3BAC6FC65FECEE199" blockId="5.[123,652,926,1742]" pageId="5" pageNumber="6">
|
||||
<materialsCitation id="3B0619324C77FFB3BAC6FC65FEC4E152" collectingDate="2002-07" collectorName="H. H. Ng" country="Singapore" location="Singapore" pageId="5" pageNumber="6" specimenCount="1" typeStatus="paratype">
|
||||
<typeStatus id="54D5ADCD4C77FFB3BAC6FC65FEF4E117" box="[152,278,1014,1037]" pageId="5" pageNumber="6" type="paratype">PARATYPES</typeStatus>
|
||||
: AMNH 233607,
|
||||
<specimenCount id="9D68D8E64C77FFB3BBA7FC65FDDAE117" box="[505,568,1014,1037]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
|
||||
,
|
||||
<quantity id="4C96BE8A4C77FFB3B817FC65FF4AE131" metricMagnitude="-1" metricUnit="m" metricValue="1.6789999999999998" pageId="5" pageNumber="6" unit="mm" value="167.9">167.9 mm</quantity>
|
||||
SL, male;
|
||||
<collectingCountry id="F37953FF4C77FFB3BB6CFB87FE4EE131" box="[306,428,1044,1067]" name="Singapore" pageId="5" pageNumber="6">Singapore</collectingCountry>
|
||||
: fish market;
|
||||
<collectorName id="269B76B94C77FFB3B80CFB87FF43E152" pageId="5" pageNumber="6">H.H. Ng</collectorName>
|
||||
,
|
||||
<date id="FFD035AF4C77FFB3BAF1FBA2FEC4E152" box="[175,294,1073,1096]" pageId="5" pageNumber="6" value="2002-07">
|
||||
<collectingDate id="EF94CC474C77FFB3BAF1FBA2FEC4E152" box="[175,294,1073,1096]" pageId="5" pageNumber="6" value="2002-07">July 2002</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B0619324C77FFB3BB6AFBA2FECAE199" collectingDate="2002-07" collectorName="H. H. Ng" country="Singapore" location="Singapore" pageId="5" pageNumber="6" specimenCount="1" typeStatus="paratype">
|
||||
UMMZ 240362,
|
||||
<specimenCount id="9D68D8E64C77FFB3BBA1FBA2FDD9E152" box="[511,571,1073,1096]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
|
||||
,
|
||||
<quantity id="4C96BE8A4C77FFB3B817FBA2FF4AE17F" metricMagnitude="-1" metricUnit="m" metricValue="1.655" pageId="5" pageNumber="6" unit="mm" value="165.5">165.5 mm</quantity>
|
||||
SL, female;
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||||
<collectingCountry id="F37953FF4C77FFB3BB60FBDDFE5AE17F" box="[318,440,1102,1125]" name="Singapore" pageId="5" pageNumber="6">Singapore</collectingCountry>
|
||||
: fish market;
|
||||
<collectorName id="269B76B94C77FFB3B80DFBDDFF43E199" pageId="5" pageNumber="6">H.H. Ng</collectorName>
|
||||
,
|
||||
<date id="FFD035AF4C77FFB3BAEEFBFFFECAE199" box="[176,296,1132,1155]" pageId="5" pageNumber="6" value="2002-07">
|
||||
<collectingDate id="EF94CC474C77FFB3BAEEFBFFFECAE199" box="[176,296,1132,1155]" pageId="5" pageNumber="6" value="2002-07">July 2002</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C77FFB3BAC6FB1AFD9BE307" pageId="5" pageNumber="6" type="diagnosis">
|
||||
<paragraph id="8BD1136F4C77FFB3BAC6FB1AFD9BE307" blockId="5.[123,652,926,1742]" pageId="5" pageNumber="6">
|
||||
DIAGNOSIS:
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||||
<taxonomicName id="4C6E68EC4C77FFB3BB7FFB1AFDFFE1BA" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[289,541,1161,1184]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="robustus">
|
||||
<emphasis id="B91ACF7D4C77FFB3BB7FFB1AFDFFE1BA" box="[289,541,1161,1184]" italics="true" pageId="5" pageNumber="6">Leiognathus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from the only other species of leiognathid known to possess a nonsexually dimorphic LOS,
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<taxonomicName id="4C6E68EC4C77FFB3BB73FB72FE4AE1E2" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[301,424,1249,1272]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="equulus">
|
||||
<emphasis id="B91ACF7D4C77FFB3BB73FB72FE4AE1E2" box="[301,424,1249,1272]" italics="true" pageId="5" pageNumber="6">L. equulus</emphasis>
|
||||
</taxonomicName>
|
||||
, by the absence of an occipital hump (vs. pronounced hump), the presence of a mildly sloping predorsal profile (vs. strongly curved, creating the image of an arched back), frontal and lateral ethmoid ossifications that project anterodorsally and extend well anterior of the orbit to form a distinct preorbital protuberance (vs. slight bulge above orbit), and a nuchal spine that is not exposed in lateral view (vs. exposed and projecting, particularly distally).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C77FFBEBAC6F9B7FF7EE321" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
|
||||
<paragraph id="8BD1136F4C77FFB3BAC6F9B7FCBEE1BA" blockId="5.[123,652,926,1742]" lastBlockId="5.[699,1228,193,1741]" pageId="5" pageNumber="6">
|
||||
DESCRIPTION: Selected proportional measurements and meristic data are presented in table 2. A deepbodied and robust
|
||||
<taxonomicName id="4C6E68EC4C77FFB3B87CF9CCFF49E389" authorityName="Lacepede" authorityYear="1802" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B91ACF7D4C77FFB3B87CF9CCFF49E389" italics="true" pageId="5" pageNumber="6">Leiognathus</emphasis>
|
||||
</taxonomicName>
|
||||
, which grows to a large size (>
|
||||
<quantity id="4C96BE8A4C77FFB3B877F9EFFD69E389" box="[553,651,1660,1683]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="6" unit="mm" value="180.0">180 mm</quantity>
|
||||
SL) (figs. 3, 4). Body laterally compressed. No pronounced supraoccipital (= predorsal) hump (fig. 5A). Lateral snout outline mildly concave. Strong preorbital protuberance due to hypertrophy and protrusion of frontal and lateral ethmoid ossifications (fig. 6A). Predorsal head profile mostly straight to mildly curved; back not strongly arched. Nuchal spine not protruding, and distal tip not exposed (figs. 3A, 4, 5A). Nuchal spine with distinct median keel. Two short and stout postnasal spines present on lateral ethmoid, located posterior to nasal foramina and just rostrodorsal of orbit. Postnasal spines followed posteriorly by welldeveloped supraorbital ridges; ridges converge posteriorly. Dorsal and ventral profiles about evenly curved. Dorsalfin origin about midway between pelvicfin and analfin origins. Analfin origin at about level of vertical through seventh dorsalfin spine. Eye large. Caudal peduncle slender and shallow. Mouth small and terminal in position, directed slightly downward when protruded. Caudal margin of maxilla exposed, reaching to level of vertical through anterior margin of orbit. Anterior nasal pore round, posterior foramen crescentshaped, partially encircling anterior pore. Lower preopercular margin weakly serrate. Vertebral count: 9 precaudal + 14 caudal = 23. Neural and hemal spines of vertebral centrum PU4 expanded and bladelike (fig. 6A). Fifteen or 16 stout and triangular outer ceratobranchial gill rakers arrayed along lower limb (= ceratobranchial one) of first gill arch.
|
||||
</paragraph>
|
||||
<paragraph id="8BD1136F4C77FFB0B886FB34FE5EE0B2" blockId="5.[699,1228,193,1741]" lastBlockId="6.[124,653,1366,1741]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||||
<emphasis id="B91ACF7D4C77FFB3B886FB34FCF7E1A4" box="[728,789,1191,1214]" italics="true" pageId="5" pageNumber="6">Fins:</emphasis>
|
||||
Dorsal fin with VIII spines and 16 branched rays. First dorsalfin spine greatly reduced in length, yet relatively robust. Second through fourth dorsalfin spines elongate and robust, second spine longest. Third and fourth dorsalfin spines serrate along anterior margin, ‘‘lock’’ into groove on preceding spine when erect. Dorsalfin spines five through eight feeble, shorter than second through fourth spines. Anal fin with III spines and 14 branched rays. First analfin spine very short. Second and third analfin spines robust and elongate, second spine longest. Third analfin spine serrate on anterior margin, ‘‘locks’’ into groove on posterior margin of second spine when erect. Spinous dorsal and anal fins with asquamate basal sheath, creating furrow into which fins may retract. Pelvic fins short, not reaching first anal spine when adducted. Eight upper and seven lower branched caudalfin rays. Nineteen total pectoralfin rays.
|
||||
</paragraph>
|
||||
<caption id="DF1143E74C74FFB0BAC8FBFFFDF4E03B" ID-DOI="http://doi.org/10.5281/zenodo.5382712" ID-Zenodo-Dep="5382712" httpUri="https://zenodo.org/record/5382712/files/figure.png" pageId="6" pageNumber="7" startId="6.[150,188,1132,1153]" targetBox="[137,1215,187,1111]" targetPageId="6">
|
||||
<paragraph id="8BD1136F4C74FFB0BAC8FBFFFDF4E03B" blockId="6.[124,1229,1132,1313]" pageId="6" pageNumber="7">
|
||||
Fig. 2. Single optimal tree of leiognathid relationships recovered by combined analysis of mitochondrial (16S and COI) nucleotide characters.
|
||||
<emphasis id="B91ACF7D4C74FFB0B822FB14FD69E186" box="[636,651,1159,1180]" italics="true" pageId="6" pageNumber="7">L</emphasis>
|
||||
. =
|
||||
<taxonomicName id="4C6E68EC4C74FFB0B8E8FB14FCD5E186" authorityName="Lacepede" authorityYear="1802" box="[694,823,1159,1180]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B91ACF7D4C74FFB0B8E8FB14FCD5E186" box="[694,823,1159,1180]" italics="true" pageId="6" pageNumber="7">Leiognathus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<emphasis id="B91ACF7D4C74FFB0B919FB14FCB8E186" box="[839,858,1159,1180]" italics="true" pageId="6" pageNumber="7">G</emphasis>
|
||||
. =
|
||||
<taxonomicName id="4C6E68EC4C74FFB0B9DBFB14FC24E186" box="[901,966,1159,1180]" class="Actinopterygii" family="Leiognathidae" genus="Gazza" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B91ACF7D4C74FFB0B9DBFB14FC24E186" box="[901,966,1159,1180]" italics="true" pageId="6" pageNumber="7">Gazza</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<emphasis id="B91ACF7D4C74FFB0B988FB14FC01E186" box="[982,995,1159,1180]" italics="true" pageId="6" pageNumber="7">S</emphasis>
|
||||
. =
|
||||
<taxonomicName id="4C6E68EC4C74FFB0BE50FB14FBBCE186" authorityName="Gistel" authorityYear="1848" box="[1038,1118,1159,1180]" class="Actinopterygii" family="Leiognathidae" genus="Secutor" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B91ACF7D4C74FFB0BE50FB14FBBCE186" box="[1038,1118,1159,1180]" italics="true" pageId="6" pageNumber="7">Secutor</emphasis>
|
||||
</taxonomicName>
|
||||
. Numbers above branches represent Bremer support and numbers below branches represent Jackknife resampling percentages (>50%). Letters at nodes correspond to clades discussed in text: clade A =
|
||||
<taxonomicName id="4C6E68EC4C74FFB0BE6EFB2FFB2BE1CB" authorityName="T.N.Gill" authorityYear="1893" box="[1072,1225,1212,1233]" class="Actinopterygii" family="Leiognathidae" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="family">Leiognathidae</taxonomicName>
|
||||
; clade B = leiognathids with nonsexually dimorphic LOS; clade C = leiognathids bearing sexually dimorphic LOS. LOS features characteristic of members of recovered clades (Sparks and Dunlap, in review) are indicated on the topology.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD1136F4C74FFB0BAC7FA3CFEE4E389" blockId="6.[124,653,1366,1741]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B91ACF7D4C74FFB0BAC7FA3CFEF3E0DC" box="[153,273,1455,1478]" italics="true" pageId="6" pageNumber="7">Dentition:</emphasis>
|
||||
Multiple rows of small, closely set, elongate and moderately recurved, villiform teeth present in both upper and lower jaws. Four to five closely set rows anteriorly and one to two rows posteriorly in upper jaw. Three to four rows anteriorly and one to two rows posteriorly in lower jaw. Lips fleshy and rugose.
|
||||
</paragraph>
|
||||
<paragraph id="8BD1136F4C74FFB0BAC7F90AFCDDE31A" blockId="6.[124,653,1366,1741]" lastBlockId="6.[700,1229,1366,1741]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B91ACF7D4C74FFB0BAC7F90AFECFE3AA" box="[153,301,1689,1712]" italics="true" pageId="6" pageNumber="7">Squamation:</emphasis>
|
||||
Body scales small and cycloid. Head and opercular region naked. Breast naked, asquamate region extending to pectoralfin base. Lateral line arched and complete. Pored scales in lateral line number 60–65. Pores well developed. Pelvic axillary scale well developed and elongate. All fins asquamate.
|
||||
</paragraph>
|
||||
<paragraph id="8BD1136F4C74FFB0B887F995FB2FE3D7" blockId="6.[700,1229,1366,1741]" pageId="6" pageNumber="7">PIGMENTATION IN PRESERVATION: Body ground coloration gray to grayishblue dorsal of midline, and pale yellow, white, or golden ventral of midline. Iridescent golden patches present to varying degree along lateral midline. Cheek and opercular region iridescent silvery to golden. Head above orbit and nape</paragraph>
|
||||
<paragraph id="8BD1136F4C75FFB1B908FD97F963E750" blockId="7.[145,1665,516,586]" pageId="7" pageNumber="8">
|
||||
TABLE 2
|
||||
<emphasis id="B91ACF7D4C75FFB1BB8AFD8CFADFE728" bold="true" box="[468,1341,543,562]" pageId="7" pageNumber="8">
|
||||
Morphometric and Meristic Data of
|
||||
<taxonomicName id="4C6E68EC4C75FFB1B919FD8CFBFAE728" authority="SPARKS & DUNLAP, 2004" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[839,1048,543,562]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="B91ACF7D4C75FFB1B919FD8CFBFAE728" bold="true" box="[839,1048,543,562]" italics="true" pageId="7" pageNumber="8">Leiognathus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="A22972064C75FFB1BE78FD8CFB7EE728" box="[1062,1180,543,562]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
|
||||
, and
|
||||
<taxonomicName id="4C6E68EC4C75FFB1BE8BFD8CFADFE728" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[1237,1341,543,562]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="equulus">
|
||||
<emphasis id="B91ACF7D4C75FFB1BE8BFD8CFADFE728" bold="true" box="[1237,1341,543,562]" italics="true" pageId="7" pageNumber="8">L. equulus</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
Measurements (mm) are percentage standard length (SL) or percentage head length (HL), unless noted otherwise. H indicates count corresponding to
|
||||
<typeStatus id="54D5ADCD4C75FFB1BC77FDA4F99FE750" box="[1577,1661,567,586]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD1136F4C7AFFBEBA22FF52FF30E61F" blockId="8.[124,654,193,1741]" pageId="8" pageNumber="9">grayish to grayishbrown. Snout dusky to blackish; appears spotty due to concentrated melanophores. Nasal region and lips pale yellow. Gular region with iridescent silvery or golden patches. Chest and belly white or pale gray. Caudal peduncle and base of caudal fin iridescent silvery or golden. Dorsal, anal, pectoral, and pelvic fins whitish to yellow. Dorsal fin with black pigment distally. Pectoralfin axil blackish due to concentration of melanophores, surrounded by large iridescent silver patch. Pectoralfin base silvery. Caudal fin yellowish to light brown, dorsal and ventral rays with black pigment, especially proximal to base. Caudal fin with prominent black terminal band. Pelvic axillary scale and body along analfin base silvery or golden. Pores of lateral line scales edged dorsally and ventrally with melanophores.</paragraph>
|
||||
<paragraph id="8BD1136F4C7AFFBEBAC7FC9FFF7EE321" blockId="8.[124,654,193,1741]" pageId="8" pageNumber="9">LIGHT ORGAN SYSTEM (LOS) (fig. 3): Sexual dimorphism of the light organ and associated structures is not detected. The light organ of males is not enlarged compared to the similarly sized conspecific female and does not exhibit any apparent shape dimorphism. The light organ is a comparatively simple, dorsoventrally compressed, doughnutshaped structure surrounding the esophagus (fig. 3B). Neither the dorsal nor ventral lobes of males are hypertrophied compared to the conspecific female examined. Likewise, associated structures of the LOS (e.g., clearing of the gasbladder lining, modifications of the integument) do not exhibit any sexually dimorphic attributes. As in all other leiognathids, the posteroventral margin of the gasbladder chamber and the small, thin anteroventral patch separating the light organ from the gasbladder are transparent. This ‘window’ is sparsely ‘peppered’ with iridescent silvery and bluish chromatophores. No lateral clearing of the silvery gasbladder lining is evident (fig. 3B). Externally sexually dimorphic features of the LOS (i.e., male specific transparent patches or stripes in the opercular region or on the flanks) are absent (figs. 3A, 4).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C7AFFBEBAC7F9D2FD1EE4BF" pageId="8" pageNumber="9" type="materials_examined">
|
||||
<paragraph id="8BD1136F4C7AFFBEBAC7F9D2FD1EE4BF" blockId="8.[124,654,193,1741]" lastBlockId="8.[700,1229,193,568]" pageId="8" pageNumber="9">
|
||||
DISTRIBUTION:
|
||||
<location id="8EB145B44C7AFFBEBB13F9D2FE41E342" LSID="urn:lsid:plazi:treatment:03C7A2794C77FFBEBAF5FCC6FB44E722:8EB145B44C7AFFBEBB13F9D2FE41E342" box="[333,419,1601,1624]" country="Singapore" county="Indo-Pacific" municipality="Given" name="Known" pageId="8" pageNumber="9">Known</location>
|
||||
at this time only from market specimens purchased in
|
||||
<collectingCountry id="F37953FF4C7AFFBEB81FF9CDFF51E389" name="Singapore" pageId="8" pageNumber="9">Singapore</collectingCountry>
|
||||
.
|
||||
<collectingMunicipality id="6BB589154C7AFFBEBA98F9EFFEEFE389" box="[198,269,1660,1683]" pageId="8" pageNumber="9">Given</collectingMunicipality>
|
||||
that the
|
||||
<collectingCountry id="F37953FF4C7AFFBEBBD9F9EFFE1FE389" box="[391,509,1660,1683]" name="Singapore" pageId="8" pageNumber="9">Singapore</collectingCountry>
|
||||
fleet fishes throughout much of the
|
||||
<collectingCounty id="62B06BE34C7AFFBEBBF3F90AFDD9E3AA" box="[429,571,1689,1712]" pageId="8" pageNumber="9">IndoPacific</collectingCounty>
|
||||
basin, we are unable to report the collection locality of the
|
||||
<typeStatus id="54D5ADCD4C7AFFBEB94EFF52FCA0E5C2" box="[784,834,193,216]" pageId="8" pageNumber="9">type</typeStatus>
|
||||
series. The overall similarity of
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB8E2FF4CFCA5E5EC" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[700,839,223,246]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="robustus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB8E2FF4CFCA5E5EC" box="[700,839,223,246]" italics="true" pageId="8" pageNumber="9">L. robustus</emphasis>
|
||||
</taxonomicName>
|
||||
to
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB925FF4CFC1EE5EC" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[891,1020,223,246]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="equulus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB925FF4CFC1EE5EC" box="[891,1020,223,246]" italics="true" pageId="8" pageNumber="9">L. equulus</emphasis>
|
||||
</taxonomicName>
|
||||
, a common and widespread species, or to other large leiognathid species such as
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB983FE8AFB60E42A" baseAuthorityName="Valenciennes" baseAuthorityYear="1835" box="[989,1154,281,304]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="dussumieri">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB983FE8AFB60E42A" box="[989,1154,281,304]" italics="true" pageId="8" pageNumber="9">L. dussumieri</emphasis>
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEBEEBFE8AFCC1E454" baseAuthorityName="Lacepede" baseAuthorityYear="1803" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="fasciatus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEBEEBFE8AFCC1E454" italics="true" pageId="8" pageNumber="9">L. fasciatus</emphasis>
|
||||
</taxonomicName>
|
||||
, suggest that
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB987FEA4FB80E454" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[985,1122,311,334]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="robustus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB987FEA4FB80E454" box="[985,1122,311,334]" italics="true" pageId="8" pageNumber="9">L. robustus</emphasis>
|
||||
</taxonomicName>
|
||||
may be easily misidentified, and therefore may traditionally have been overlooked in collections.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37440E44C7AFFBEB887FE3FFB44E722" pageId="8" pageNumber="9" type="etymology">
|
||||
<paragraph id="8BD1136F4C7AFFBEB887FE3FFB44E722" blockId="8.[700,1229,193,568]" pageId="8" pageNumber="9">
|
||||
ETYMOLOGY: Named in reference to the robust nature and large size of the species compared to all congeners except
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEBE7AFE74FB42E4E4" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[1060,1184,487,510]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="equulus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEBE7AFE74FB42E4E4" box="[1060,1184,487,510]" italics="true" pageId="8" pageNumber="9">L. equulus</emphasis>
|
||||
</taxonomicName>
|
||||
, its sister taxon, and
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB9C8FD97FBC1E701" baseAuthorityName="Lacepede" baseAuthorityYear="1803" box="[918,1059,516,539]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="fasciatus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB9C8FD97FBC1E701" box="[918,1059,516,539]" italics="true" pageId="8" pageNumber="9">L. fasciatus</emphasis>
|
||||
</taxonomicName>
|
||||
. The specific epithet,
|
||||
<taxonomicName id="4C6E68EC4C7AFFBEB942FDB2FC62E722" authorityName="SPARKS & DUNLAP" authorityYear="2004" box="[796,896,545,568]" class="Actinopterygii" family="Leiognathidae" genus="Leiognathus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="robustus">
|
||||
<emphasis id="B91ACF7D4C7AFFBEB942FDB2FC62E722" box="[796,896,545,568]" italics="true" pageId="8" pageNumber="9">robustus</emphasis>
|
||||
</taxonomicName>
|
||||
, is used as an adjective.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue