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<mods:title id="6CE0FB6A3D36E40486941A2F7FCC33F5">Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)</mods:title>
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<mods:namePart>Pholyotha, Arthit</mods:namePart>
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<mods:affiliation>Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<treatment ID-GBIF-Taxon="168504529" LSID="urn:lsid:zoobank.org:act:B61773AB-0762-4B4D-A097-162ED9C43720" httpUri="http://treatment.plazi.org/id/B6073C8928CE513DB8B08369C717ED51" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="B6073C89-28CE-513D-B8B0-8369C717ED51" authority="Pholyotha & Panha" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi" status="sp. nov.">Sarika gratesi Pholyotha & Panha</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">Figs 49</figureCitation>
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<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">, 50A, B</figureCitation>
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<treatment id="B6073C8928CE513DB8B08369C717ED51" ID-GBIF-Taxon="168504529" LSID="urn:lsid:zoobank.org:act:B61773AB-0762-4B4D-A097-162ED9C43720" httpUri="http://treatment.plazi.org/id/B6073C8928CE513DB8B08369C717ED51" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Ariophantidae" scope_superOrder="Eupulmonata">
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<taxonomicName id="B7F7787216AE766623CCCA47F908A492" ID-CoL="7T3FD" LSID="B6073C89-28CE-513D-B8B0-8369C717ED51" authority="Pholyotha & Panha" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi" status="sp. nov.">Sarika gratesi Pholyotha & Panha</taxonomicName>
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<taxonomicNameLabel id="84B8B9325A3DFCD357B665C417B2F3C3" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="EFA4AEF372DAE8C32798FF6FE648150C" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">Figs 49</figureCitation>
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<figureCitation id="84B5F3C0B64BC1E33979A586E605B782" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">, 50A, B</figureCitation>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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<subSubSection id="C1D9F8CAA37F607871332F3B47E1AA4F" pageId="0" pageNumber="1" type="type material">
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<paragraph id="CC0811F396C3A7F6D8A16D6CE2FFFB0C" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="0279A264290816CABF5F9493DC98D614" pageId="0" pageNumber="1">
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<emphasis id="D2D40AD602236EB0FE678E6E719B19AC" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 7275 (Fig.
|
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<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A</figureCitation>
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||||
<figureCitation id="3B977817934C2C518D499D7F5C237DDE" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A</figureCitation>
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, width 16.5 mm, height 8.8 mm).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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<emphasis id="98646CDB80922AAB824F25ED80963469" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 7276 (10 shells; Fig.
|
||||
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50B</figureCitation>
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<figureCitation id="A8C981B116151E94DF0EF55937445E6F" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50B</figureCitation>
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, width 16.0 mm, height 8.3 mm), 7912 (30 shells), NHMUK 20200289 (two shells), SMF (two shells), ZRC.MOL.017031 (two shells).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Thailand-Northeastern.</emphasis>
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<subSubSection id="2F7257ECD553DF4FF340C7CDF6A1BD31" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="639FA184F9F267A19ACE7CCE300E2BE6" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="86120F15F7F60C50AE8667A914A40B39" pageId="0" pageNumber="1">
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<emphasis id="373199C9173FB67BF2983BB3B1457D0C" bold="true" pageId="0" pageNumber="1">Thailand-Northeastern.</emphasis>
|
||||
Dry dipterocarp forest at Phu Lan Kha, Nong Bua Daeng, Chaiyaphum,
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<geoCoordinate degrees="16" direction="north" minutes="00" orientation="latitude" precision="1" seconds="00.9" value="16.00025">16°00'00.9"N</geoCoordinate>
|
||||
<geoCoordinate id="F873FFE49300535B19E7A55244680ADB" degrees="16" direction="north" minutes="00" orientation="latitude" precision="1" seconds="00.9" value="16.00025">16°00'00.9"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="101" direction="east" minutes="52" orientation="longitude" precision="1" seconds="33.4" value="101.875946">101°52'33.4"E</geoCoordinate>
|
||||
<geoCoordinate id="20A0518952DB0FA278EDF2BA2399A246" degrees="101" direction="east" minutes="52" orientation="longitude" precision="1" seconds="33.4" value="101.875946">101°52'33.4"E</geoCoordinate>
|
||||
: CUMZ 7277.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="403072F6A23325DC0ABE02BF14675A11" pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph id="248F67914938F391AEDFBE72C68940D1" pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph id="FB133E5C9BD080E933F843E3F0FE3C67" pageId="0" pageNumber="1">
|
||||
The limestone outcrop with dry deciduous forest at Tham Phraya Nakarat (Cave), Chum Phae, Khon Kaen, Thailand,
|
||||
<geoCoordinate degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="30.3" value="16.808416">16°48'30.3"N</geoCoordinate>
|
||||
<geoCoordinate id="82D166FF1F3806B931F95AFF5C3AC92E" degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="30.3" value="16.808416">16°48'30.3"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="101" direction="east" minutes="57" orientation="longitude" precision="1" seconds="13.7" value="101.953804">101°57'13.7"E</geoCoordinate>
|
||||
<geoCoordinate id="4DF0B670A587E6055ADA04EAB9F7DCB1" degrees="101" direction="east" minutes="57" orientation="longitude" precision="1" seconds="13.7" value="101.953804">101°57'13.7"E</geoCoordinate>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Shell medium-sized, depressed to strongly depressed, and pale brown. Aperture irregular with peristome rather simple above then expanded middle with curved inside aperture and thickened below periphery.</paragraph>
|
||||
<caption doi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
|
||||
<subSubSection id="4E2F60B5BF4D10CFE068852BFA958F96" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="EEA6DE31ED4F95331D04338A8263DAB7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="817CCA46F6674AA32D9777707EB95298" pageId="0" pageNumber="1">Shell medium-sized, depressed to strongly depressed, and pale brown. Aperture irregular with peristome rather simple above then expanded middle with curved inside aperture and thickened below periphery.</paragraph>
|
||||
<caption id="F913CDB5B17AF688286871FFB01802B5" doi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
|
||||
<paragraph id="EB5176F3A054473793AAB030653F984A" pageId="0" pageNumber="1">
|
||||
<emphasis id="D77E46C7B7E517FB840630D67A6574FF" bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
|
||||
Geographic distribution of
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
<taxonomicName id="ED78F87C4302E8A365794921AA9AC4EA" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis id="C75F8673C075A9132FFE54152BAA38E4" italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. based on the specimens examined herein.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
<subSubSection id="545E8DE2BB1793110F4B4DE0E6663FAD" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="CEB65F250518B7AB18A850A4E300420C" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="F303087FD47D6A610FEA1FE692B20C4D" pageId="0" pageNumber="1">
|
||||
<emphasis id="CE731EFF874664E8CB8DC3C025CD45EE" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="828EDAA624F3CABEDD966A56545E8369" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shell depressed to strongly depressed or nearly flattened, medium-sized (shell width up to 17.4 mm, shell height up to 8.8 mm), rather thin, and slightly opaque. Shell surface smooth, polished and with thin growth lines; shell colour pale brown. Whorls 6-7, increasing regularly; body whorl large and well rounded. Spire little to moderately elevated; suture impressed. Aperture crescent-shaped and open obliquely; peristome irregular. Apertural lip at upper periphery simple; at periphery with invagination of triangular lip (beak-like); at below periphery rather thickened inside aperture and little expanded. Columellar margin straight, slightly thickened and expanded near umbilicus. Umbilicus narrowly opened (Fig.
|
||||
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A, B</figureCitation>
|
||||
<figureCitation id="3C2EC0CC872C2EF53C852CB5031CD77E" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
|
||||
<caption id="4DD7C56F564353203F5700F4457512BB" doi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
|
||||
<paragraph id="C96E15F01B51797597779F0BE84A9F2C" pageId="0" pageNumber="1">
|
||||
<emphasis id="734AFBE7050C9850225A1496B059428F" bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
|
||||
Shells of
|
||||
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
<taxonomicName id="D518ADF398EB1ADA7040CD03BB2C3454" authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="FD7DC74E9E34F19C1225DDDC32528AB3" italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
<emphasis id="30F5F4E9EAA155E70EE3AA796DEE148C" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||||
<taxonomicName id="CEA55146A42AC263A23052C17D36F598" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis id="C5CAADCA749C3F41240B4E8CA37E01F9" italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="51C906E14067D91DFAD8FB8552AC68AC" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
holotype CUMZ 7275 and
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="56D508234F4EC7665FFFD283B7AAD60B" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
paratype CUMZ 7276
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||||
<taxonomicName lsidName="S. pumicata" pageId="0" pageNumber="1" rank="species" species="pumicata">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. pumicata</emphasis>
|
||||
<emphasis id="BB55A95918D99FDE4313B12D14B52C0E" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||||
<taxonomicName id="52C9CBF80E43D8116EF2753527D35463" lsidName="S. pumicata" pageId="0" pageNumber="1" rank="species" species="pumicata">
|
||||
<emphasis id="CFA218B75E82611C1AF42BF8302B7C39" italics="true" pageId="0" pageNumber="1">S. pumicata</emphasis>
|
||||
</taxonomicName>
|
||||
syntype NHMUK 1893.2.4.980-981
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<taxonomicName lsidName="S. ochtogyra" pageId="0" pageNumber="1" rank="species" species="ochtogyra">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. ochtogyra</emphasis>
|
||||
<emphasis id="F8C1A394D91914E79BD5A51DBD5A3A10" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<taxonomicName id="196420DAA646EFB78A95E66CB4BCE8B6" lsidName="S. ochtogyra" pageId="0" pageNumber="1" rank="species" species="ochtogyra">
|
||||
<emphasis id="A6BD357D21A4F3BA172D3A1DFE8D2B25" italics="true" pageId="0" pageNumber="1">S. ochtogyra</emphasis>
|
||||
</taxonomicName>
|
||||
holotype SMF 227103/1
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<taxonomicName lsidName="S. benoiti" pageId="0" pageNumber="1" rank="species" species="benoiti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. benoiti</emphasis>
|
||||
<emphasis id="5D45DC2469E2BA8254786A3012D48A9D" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<taxonomicName id="A08494A8AF3CEFCAD1EAA074D0BE3147" lsidName="S. benoiti" pageId="0" pageNumber="1" rank="species" species="benoiti">
|
||||
<emphasis id="D361E7229FAC8129680C634817ADF957" italics="true" pageId="0" pageNumber="1">S. benoiti</emphasis>
|
||||
</taxonomicName>
|
||||
modified from
|
||||
<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="343 - 379" refId="B13" refString="Crosse, H, Fischer, P, 1863. Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues. Journal de Conchyliologie 11: 343 - 379" title="Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues." volume="11" year="1863">Crosse and Fischer (1863)</bibRefCitation>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
<taxonomicName lsidName="S. rex" pageId="0" pageNumber="1" rank="species" species="rex">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. rex</emphasis>
|
||||
<bibRefCitation id="C0D501D785CEAE019D911FDBCCA23DDC" author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="343 - 379" refId="B13" refString="Crosse, H, Fischer, P, 1863. Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues. Journal de Conchyliologie 11: 343 - 379" title="Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues." volume="11" year="1863">Crosse and Fischer (1863)</bibRefCitation>
|
||||
<emphasis id="051D95CE72690AEE45371C9C9D73F927" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
<taxonomicName id="E237EBAB034C9C8E956ED62F151EE2A6" lsidName="S. rex" pageId="0" pageNumber="1" rank="species" species="rex">
|
||||
<emphasis id="E51CDC0322A8D028E7AF4CCF92649901" italics="true" pageId="0" pageNumber="1">S. rex</emphasis>
|
||||
</taxonomicName>
|
||||
modified from
|
||||
<bibRefCitation author="Preston, HB" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="202 - 202" refId="B71" refString="Preston, HB, 1909. Descriptions of new species of Macrochlamys and Pseudodon from Siam. Proceedings of the Malacological Society of London 8: 202 - 202" title="Descriptions of new species of Macrochlamys and Pseudodon from Siam." volume="8" year="1909">Preston (1909)</bibRefCitation>
|
||||
<bibRefCitation id="C5C91C1713A2BA0F1B90E0F5E3EFCFD5" author="Preston, HB" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="202 - 202" refId="B71" refString="Preston, HB, 1909. Descriptions of new species of Macrochlamys and Pseudodon from Siam. Proceedings of the Malacological Society of London 8: 202 - 202" title="Descriptions of new species of Macrochlamys and Pseudodon from Siam." volume="8" year="1909">Preston (1909)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="2BA290B4D2752D4C4C6CA5B4B575DB74" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="4D6035132A7DD137F5D73B4D9686FF8B" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="624BDE8243FAE88BE67F1107E1F8535D" pageId="0" pageNumber="1">
|
||||
The specific name
|
||||
<taxonomicName lsidName="gratesi" pageId="0" pageNumber="1" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">gratesi</emphasis>
|
||||
<taxonomicName id="72B92CB84FA65B4461765D57947532D8" lsidName="gratesi" pageId="0" pageNumber="1" rank="species" species="gratesi">
|
||||
<emphasis id="64D704566C80DE2C164E13B2C43422D7" italics="true" pageId="0" pageNumber="1">gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
is named in honour to Admiral Chorchat Gra-tes of the Royal Thai Navy, who made possible many fieldtrips especially the remote islands areas in Thailand.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
<subSubSection id="ABDC3C7DDBA2B0F2B5935A8CBAA43F08" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="6F93CD3928E200BAC91A5768DB89779F" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph id="121727B6812222F24DED720AFDD3CB8F" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="CEA87B743C80311825F0BE10AB921BF8" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis id="B4A384F880BE374D8CEA1ED4F5C26E93" italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is currently known from the restricted area of the dry deciduous and dry dipterocarp forests in Khon Kaen and Chaiyaphum provinces (Fig.
|
||||
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">49</figureCitation>
|
||||
<figureCitation id="408C3ECCD812948856DEF3620E71A3D6" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">49</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="84DEE5309ACC11666048B6F843A634E4" pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph id="0BC162A2524B047583C964B31420C9DB" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph id="08B7B30FD389DD939EBF5681DAE35135" pageId="0" pageNumber="1">
|
||||
This new species is easy to distinguish from all known
|
||||
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
<taxonomicName id="FE86CC107A40EF7AD341E05CCCF549FD" authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="BAE5FCAAC01A677726289E5F291BE0CA" italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
</taxonomicName>
|
||||
as well as
|
||||
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
|
||||
<taxonomicName id="80A4A09D23790B4064A8F4279AE129AA" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="06EC959AE7D293E7A2EFA06B168E5470" italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
|
||||
</taxonomicName>
|
||||
species by its unique beak-like lip, while all other species in these two genera have simple to little thickened lips. Only,
|
||||
<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1863" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys aspides" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aspides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys aspides</emphasis>
|
||||
<taxonomicName id="D261170566535BD5C28C8B737AFF24FE" baseAuthorityName="Benson" baseAuthorityYear="1863" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys aspides" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aspides">
|
||||
<emphasis id="E4A91AA95E174720F0915274338D297A" italics="true" pageId="0" pageNumber="1">Macrochlamys aspides</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/00222936308681435" author="Benson, WH" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille, Serie II" pageId="0" pageNumber="1" refId="B7" refString="Benson, WH, 1863. Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318-323. https://doi.org/10.1080/00222936308681435" title="Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318 - 323." url="https://doi.org/10.1080/00222936308681435" year="1863">Benson 1863</bibRefCitation>
|
||||
<bibRefCitation id="1DAFAA0DC8562601973211712136016C" DOI="https://doi.org/10.1080/00222936308681435" author="Benson, WH" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille, Serie II" pageId="0" pageNumber="1" refId="B7" refString="Benson, WH, 1863. Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318-323. https://doi.org/10.1080/00222936308681435" title="Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318 - 323." url="https://doi.org/10.1080/00222936308681435" year="1863">Benson 1863</bibRefCitation>
|
||||
) from Myanmar tends to have a slightly thickened and expanded lip at below periphery, but
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
<taxonomicName id="945B8E51972F9B1F0EA0BB9BF9E18481" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis id="2CABF40CA9C3CFAD31060C1BAFD21499" italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. has a projecting and curved triangular shape lip at the periphery.
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
<taxonomicName id="96449511B338EE972D60C001EBE244AC" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
|
||||
<emphasis id="1A75898C2AD8EB54E6345F8C73035DC0" italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is identical to
|
||||
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
<taxonomicName id="CF0FF9EA21B23951CABFEEE7C5801B12" authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="849661C1F162905105FFEAFCC1A396F0" italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
</taxonomicName>
|
||||
because its shells are matched with
|
||||
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
<taxonomicName id="26108A8A9A627AE5650A740BE592A6CE" authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="0F71B3E5D0FB90EFC8B2F6771626E6B2" italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
|
||||
</taxonomicName>
|
||||
more than the Siam
|
||||
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
|
||||
<taxonomicName id="67A5BDA6AB1BBA2FE2B62A7F20F083B3" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="893CE61C38D94148780E5DD3DCB9D980" italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation author="Pholyotha, A" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="763 - 781" refId="B66" refString="Pholyotha, A, Sutcharit, C, Panha, S, 2018. The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae). Raffles Bulletin of Zoology 66: 763 - 781" title="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)." volume="66" year="2018">Pholyotha et al. 2018</bibRefCitation>
|
||||
<bibRefCitation id="E329CF03F3FB18230F3987ED009A5AF8" author="Pholyotha, A" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="763 - 781" refId="B66" refString="Pholyotha, A, Sutcharit, C, Panha, S, 2018. The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae). Raffles Bulletin of Zoology 66: 763 - 781" title="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)." volume="66" year="2018">Pholyotha et al. 2018</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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