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<mods:title id="2C17FF3234162523BD18D01F575F6043">Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)</mods:title>
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<taxonomicName LSID="081C3931-81B3-5C64-B075-68F6BF10CD2E" authority="Pholyotha & Panha" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira" status="sp. nov.">Sarika inferospira Pholyotha & Panha</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Geographic distribution of Sarika bocourti, S. hainesi, S. inferospira sp. nov., S. melanospira sp. nov. and S. pellosa sp. nov. based on examined specimens herein." figureDoi="10.3897/zookeys.976.53859.figure32" httpUri="https://binary.pensoft.net/fig/464414" pageId="0" pageNumber="1">, 32</figureCitation>
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Living snails of group II: Sarika hainesi group. A Sarika bocourti specimen CUMZ 7591 showing no left shell lobe (yellow arrow) B S. hainesi specimen CUMZ 7237 C S. bocourti specimen CUMZ 7591 D S. inferospira sp. nov. specimen CUMZ 7257 E S. melanospira sp. nov. paratype CUMZ 7243, and F S. pellosa sp. nov. paratype CUMZ 7517. Not to scale." figureDoi="10.3897/zookeys.976.53859.figure33" httpUri="https://binary.pensoft.net/fig/464415" pageId="0" pageNumber="1">, 33D</figureCitation>
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<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">, 38A, B</figureCitation>
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<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">, 39A, B</figureCitation>
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<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">, 40</figureCitation>
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Representative SEM images of the radula. A Sarika hainesi specimen CUMZ 7237 B S. bocourti specimen CUMZ 7597 C S. inferospira sp. nov. paratype CUMZ 7255 D S. melanospira sp. nov. paratype CUMZ 7243, and E S. pellosa sp. nov. specimen CUMZ 7251. Central tooth indicated by ' C'; Yellow colour indicates lateral teeth with the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure43" httpUri="https://binary.pensoft.net/fig/464425" pageId="0" pageNumber="1">, 43C</figureCitation>
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<taxonomicName id="9E1E7347D50B26BF7B629D9D2FADCC68" ID-CoL="7T3FH" LSID="081C3931-81B3-5C64-B075-68F6BF10CD2E" authority="Pholyotha & Panha" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira" status="sp. nov.">Sarika inferospira Pholyotha & Panha</taxonomicName>
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<figureCitation id="3E184209BCA8567E464054CFCFC51C18" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation id="77781654556AF5C6E884720BF35617BB" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Geographic distribution of Sarika bocourti, S. hainesi, S. inferospira sp. nov., S. melanospira sp. nov. and S. pellosa sp. nov. based on examined specimens herein." figureDoi="10.3897/zookeys.976.53859.figure32" httpUri="https://binary.pensoft.net/fig/464414" pageId="0" pageNumber="1">, 32</figureCitation>
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<figureCitation id="52846046B3718259A605E329179039C9" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Living snails of group II: Sarika hainesi group. A Sarika bocourti specimen CUMZ 7591 showing no left shell lobe (yellow arrow) B S. hainesi specimen CUMZ 7237 C S. bocourti specimen CUMZ 7591 D S. inferospira sp. nov. specimen CUMZ 7257 E S. melanospira sp. nov. paratype CUMZ 7243, and F S. pellosa sp. nov. paratype CUMZ 7517. Not to scale." figureDoi="10.3897/zookeys.976.53859.figure33" httpUri="https://binary.pensoft.net/fig/464415" pageId="0" pageNumber="1">, 33D</figureCitation>
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<figureCitation id="DE0E357F3D63FFA1A8E319BDA179F956" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">, 38A, B</figureCitation>
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<figureCitation id="ED5B68F0299C42CA0FB93378D644AC75" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">, 39A, B</figureCitation>
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<figureCitation id="B8C5DE4158E9BFF69117B16CAFFA5140" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">, 40</figureCitation>
|
||||
<figureCitation id="1FFBB6C0414915DABCA9A171FEE5F320" captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Representative SEM images of the radula. A Sarika hainesi specimen CUMZ 7237 B S. bocourti specimen CUMZ 7597 C S. inferospira sp. nov. paratype CUMZ 7255 D S. melanospira sp. nov. paratype CUMZ 7243, and E S. pellosa sp. nov. specimen CUMZ 7251. Central tooth indicated by ' C'; Yellow colour indicates lateral teeth with the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure43" httpUri="https://binary.pensoft.net/fig/464425" pageId="0" pageNumber="1">, 43C</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="28C5BBB5498C709F17719BE82C444068" pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph id="2440287A77E16516D3BD5A2147F2B059" pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph id="5BEB3CC9BEBA358C90C71420C806AE11" pageId="0" pageNumber="1">
|
||||
<emphasis id="4702CE06849589DE61254695869D990C" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
CUMZ 7254 (Fig.
|
||||
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38A</figureCitation>
|
||||
<figureCitation id="A13D502E06A9C6FCD3FD376E8317E8C6" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38A</figureCitation>
|
||||
, width 25.3 mm, height 11.5 mm).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
<emphasis id="8716859658BB66FAA4EDA57DD65C6612" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
CUMZ 7255 (two shells and 13 specimens preserved in ethanol; Fig.
|
||||
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38B</figureCitation>
|
||||
<figureCitation id="A22EF9ACAB438DF48B82B3FE43AFED6C" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38B</figureCitation>
|
||||
, width 24.8 mm, height 11.0 mm) CUMZ 7256 (four shells), 7257 (four specimens preserved in ethanol), NHMUK 20200285 (two shells).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="42BDDADE5CF3BABF186F0248866C353F" pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph id="16FD2668EEEF7298A6ED7BE6DC185C31" pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph id="ED1CA0117A5A69F118D9EBE23648A24D" pageId="0" pageNumber="1">
|
||||
Wat Tham Sai Thong, Nong Kung Si, Kalasin, Thailand,
|
||||
<geoCoordinate degrees="16" direction="north" minutes="50" orientation="latitude" precision="1" seconds="11.3" value="16.836473">16°50'11.3"N</geoCoordinate>
|
||||
<geoCoordinate id="D0DF3FC5AD418DB8EFBC52EB16B403E6" degrees="16" direction="north" minutes="50" orientation="latitude" precision="1" seconds="11.3" value="16.836473">16°50'11.3"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="103" direction="east" minutes="14" orientation="longitude" precision="1" seconds="18.7" value="103.238525">103°14'18.7"E</geoCoordinate>
|
||||
<geoCoordinate id="FF814737BEBF1C1623BD49474228A10B" degrees="103" direction="east" minutes="14" orientation="longitude" precision="1" seconds="18.7" value="103.238525">103°14'18.7"E</geoCoordinate>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Shell large, strongly depressed, very pale yellowish brown with shouldered body whorl. Animal with grey colour and four mantle lobes. Genitalia with a large straight epiphallic caecum, and triangular prism pilasters on inner penial sculpture. Spermatophore: tail filament near sperm sac with two spines and a series of several branching spines occurring continually to the middle region; middle region becoming smooth, spineless and then terminal part approximately half of its length with a series of branching spines.</paragraph>
|
||||
<caption doi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
|
||||
<subSubSection id="7E6A352DFAE4DA98D13EA6733E15F63B" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="F5757BD922215C2F8160D7B372C59D12" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="E693BDFB94B15C88EA1E3B03F102D56F" pageId="0" pageNumber="1">Shell large, strongly depressed, very pale yellowish brown with shouldered body whorl. Animal with grey colour and four mantle lobes. Genitalia with a large straight epiphallic caecum, and triangular prism pilasters on inner penial sculpture. Spermatophore: tail filament near sperm sac with two spines and a series of several branching spines occurring continually to the middle region; middle region becoming smooth, spineless and then terminal part approximately half of its length with a series of branching spines.</paragraph>
|
||||
<caption id="4575ED0EE020190BF5AEAA3CD59B909B" doi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
|
||||
<paragraph id="3739217F84FF34EF2069E92056A5E4EF" pageId="0" pageNumber="1">
|
||||
<emphasis id="7CF32F1BA3791590023C5B0FA39A1EC5" bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
|
||||
Spermatophore of
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
<taxonomicName id="0123BE461B75F28D19FD5CF4F4BD398C" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis id="96FAE1FFE0EE7D55A327FA31B59FBBA4" italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. specimen CUMZ 7257
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="AA4AB5CF04CECA158C4BC1BCD2649207" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
general view of spermatophore
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="853249093DCB78CF4B57D74BDB8F3186" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
head filament and
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C-E</emphasis>
|
||||
<emphasis id="F8D44FFC159E4E29A68B1F309E37F6A5" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
|
||||
tail filament showing
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="D442C46847ABEF7F97DC4891B54E9121" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
two spines located close to the sperm sac
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="E79E6E92954C93FC15248BEDDFCDFB1A" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
region with and without branching spines, and
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="FF3CCABCAE274EA51BA01A6EAAEAE6B1" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
branching spines on the tip region. Yellow line indicates the region with no spine.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
<subSubSection id="2DD1A71881D16430F14652348B200F0F" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="C20641ADB91594F232DD35256DFD4841" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="C2FB5BE9C6803B8678F9F227540265B3" pageId="0" pageNumber="1">
|
||||
<emphasis id="F681EFF39706E0E6B952AAF26EF7931F" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="4310F1ED6CE26F2A0D1402CF9681883B" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shell strongly depressed, large size (shell width up to 29.3 mm, shell height up to 13.9 mm) and thin. Surface smooth and polished; shell colour very pale yellowish brown. Whorls 6-
|
||||
<normalizedToken originalValue="6½">61/2</normalizedToken>
|
||||
<normalizedToken id="CB2137DAFA46FA093E903580357A6F8E" originalValue="6½">61/2</normalizedToken>
|
||||
, increasing regularly; body whorl large and shouldered. Spire slightly elevated; suture impressed. Aperture crescent-shaped and obliquely opened. Peristome simple. Columellar margin simple and slightly reflected near umbilicus. Umbilicus narrowly opened (Fig.
|
||||
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38A, B</figureCitation>
|
||||
<figureCitation id="B6CCCDB7D7F97C1C30176CE5A1EB1385" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Shells of Group II: Sarika hainesi group. A, B Sarika inferospira sp. nov. A holotype CUMZ 7254 and B paratype CUMZ 7255 C, D S. melanospira sp. nov. C holotype CUMZ 7258 and D paratype CUMZ 7243 E, F S. pellosa sp. nov. E holotype CUMZ 7249 and F paratype CUMZ 7517." figureDoi="10.3897/zookeys.976.53859.figure38" httpUri="https://binary.pensoft.net/fig/464420" pageId="0" pageNumber="1">38A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Genital organs</emphasis>
|
||||
<paragraph id="9C1F1FEF10753D49E02CA30A5B5AD4EE" pageId="0" pageNumber="1">
|
||||
<emphasis id="CBB07F263159975E3EE4281B4DEEA548" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="DAC408A2F23762DD7B783B59A58213B4" italics="true" pageId="0" pageNumber="1">Genital organs</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Atrium short. Penis cylindrical with thin penial sheath covering proximal penis. Inner sculpture of penis proximally more than ca. half of penial chamber with very finely longitudinal penial pilasters to nearly smooth surface, and then gradually transformed from small to large rhomboid pilasters with acute angle on top (triangular prism). Epiphallus cylindrical and approximately the as long as penis. Epiphallic caecum short, straight, diameter larger than epiphallus, and located near middle of epiphallus. Penial retractor muscle thin and attached at tip of epiphallic caecum. Flagellum long, slender and nearly one and half times of epiphallus length. Vas deferens thin tube connecting distal epiphallus and free oviduct (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">39A, B</figureCitation>
|
||||
<figureCitation id="DC4ECA1A885A707C55CD2C60B40C9681" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">39A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<paragraph id="2A44103B49060FCB7EA2919475CB8924" pageId="0" pageNumber="1">
|
||||
Vagina cylindrical, short, and approximately two-third of penis length. Dart apparatus large, long cylindrical and located on atrium at vagina and penis junction. Gametolytic sac enlarged and bulbous; gametolytic duct long cylindrical. Free oviduct cylindrical, approximately two and half times of vagina length (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">39A</figureCitation>
|
||||
<figureCitation id="13F217F27F9EDAE474717D389AD0D546" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Genitalia. A, B Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of the genital system and B internal structure of the penis C, D S. melanospira sp. nov. paratype CUMZ 7243 C general view of the genital system and D internal structure of the penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure39" httpUri="https://binary.pensoft.net/fig/464421" pageId="0" pageNumber="1">39A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<paragraph id="5D639EBEABD552403F926F7F318B311A" pageId="0" pageNumber="1">
|
||||
Spermatophore long and needle-shaped. Sperm sac enlarged and elongate-oval. Head filament was missing (incomplete spermatophore). Tail filament very long tube and region near sperm sac with two spines. Spine I simple and long. Spine II slightly longer and larger than spine I and branching into many small spinules. Continuously on tail filament with short branching spines arranged in a row, modified to longer branching spines arranged in several rows around middle region, and then become smooth and without spine (Fig.
|
||||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">40A</figureCitation>
|
||||
<figureCitation id="3325548C053764035B9D6E440B5E315F" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">40A</figureCitation>
|
||||
at yellow line). Terminal part of tail filament (more than ca. one-fourth of its length) with series of long branching spines arranged in opposite rows and tail filament tip with no spine (Fig.
|
||||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">40</figureCitation>
|
||||
<figureCitation id="94EE3FE3B2B5524E2F2B5D46596C09D4" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Spermatophore of Sarika inferospira sp. nov. specimen CUMZ 7257 A general view of spermatophore B head filament and C-E tail filament showing C two spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow line indicates the region with no spine." figureDoi="10.3897/zookeys.976.53859.figure40" httpUri="https://binary.pensoft.net/fig/464422" pageId="0" pageNumber="1">40</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<taxonomicName class="Bivalvia" family="Limidae" genus="Radula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radula" order="Limida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||||
<paragraph id="40F2BB5B9DFA8C69AE636C0FCD24BB02" pageId="0" pageNumber="1">
|
||||
<emphasis id="D8F3D1A72BB4F6D627FE30DDBD3E988E" bold="true" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="B9B6B27C8088CCA80BEE734199EA109F" class="Bivalvia" family="Limidae" genus="Radula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radula" order="Limida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="515563AD6B23CEA9C7A8D2416D1F8792" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Teeth with half row formula: 1-(13-14)-59. Central tooth symmetrical tricuspid; lateral teeth asymmetrical tricuspid; marginal teeth elongate bicuspid. Teeth shape is similar to that of
|
||||
<taxonomicName lsidName="S. resplendens" pageId="0" pageNumber="1" rank="species" species="resplendens">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. resplendens</emphasis>
|
||||
<taxonomicName id="2DD06CC897DD3EFBA8BF9CEBFC094788" lsidName="S. resplendens" pageId="0" pageNumber="1" rank="species" species="resplendens">
|
||||
<emphasis id="4D5120232A1C48FA55C8DCA24E72C628" italics="true" pageId="0" pageNumber="1">S. resplendens</emphasis>
|
||||
</taxonomicName>
|
||||
. Marginal teeth starting at approximately row number 13 or 14 (Fig.
|
||||
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Representative SEM images of the radula. A Sarika hainesi specimen CUMZ 7237 B S. bocourti specimen CUMZ 7597 C S. inferospira sp. nov. paratype CUMZ 7255 D S. melanospira sp. nov. paratype CUMZ 7243, and E S. pellosa sp. nov. specimen CUMZ 7251. Central tooth indicated by ' C'; Yellow colour indicates lateral teeth with the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure43" httpUri="https://binary.pensoft.net/fig/464425" pageId="0" pageNumber="1">43C</figureCitation>
|
||||
<figureCitation id="E57D4ADDE900A9348050C2356D2E2669" captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Representative SEM images of the radula. A Sarika hainesi specimen CUMZ 7237 B S. bocourti specimen CUMZ 7597 C S. inferospira sp. nov. paratype CUMZ 7255 D S. melanospira sp. nov. paratype CUMZ 7243, and E S. pellosa sp. nov. specimen CUMZ 7251. Central tooth indicated by ' C'; Yellow colour indicates lateral teeth with the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure43" httpUri="https://binary.pensoft.net/fig/464425" pageId="0" pageNumber="1">43C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">External features</emphasis>
|
||||
<paragraph id="790B1291DEEE9BADAD89509C5C8AB35B" pageId="0" pageNumber="1">
|
||||
<emphasis id="141C726A115908F16CEF0C94240A3A3F" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="B5B5CDD551F5287F6B5A0B2CF21228FE" italics="true" pageId="0" pageNumber="1">External features</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Animal with reticulated skin and body colour with dark grey above and creamy-grey below. Creamy-grey foot sole and dark creamy-grey caudal horn. Four mantle lobes well developed and pale grey colour. Left shell lobe absent (Fig.
|
||||
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Living snails of group II: Sarika hainesi group. A Sarika bocourti specimen CUMZ 7591 showing no left shell lobe (yellow arrow) B S. hainesi specimen CUMZ 7237 C S. bocourti specimen CUMZ 7591 D S. inferospira sp. nov. specimen CUMZ 7257 E S. melanospira sp. nov. paratype CUMZ 7243, and F S. pellosa sp. nov. paratype CUMZ 7517. Not to scale." figureDoi="10.3897/zookeys.976.53859.figure33" httpUri="https://binary.pensoft.net/fig/464415" pageId="0" pageNumber="1">33D</figureCitation>
|
||||
<figureCitation id="9DE49DE44C561A8B763742D327A163AD" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Living snails of group II: Sarika hainesi group. A Sarika bocourti specimen CUMZ 7591 showing no left shell lobe (yellow arrow) B S. hainesi specimen CUMZ 7237 C S. bocourti specimen CUMZ 7591 D S. inferospira sp. nov. specimen CUMZ 7257 E S. melanospira sp. nov. paratype CUMZ 7243, and F S. pellosa sp. nov. paratype CUMZ 7517. Not to scale." figureDoi="10.3897/zookeys.976.53859.figure33" httpUri="https://binary.pensoft.net/fig/464415" pageId="0" pageNumber="1">33D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="027E5CEC1340963018B25F587E026A84" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="7D66827FF59791C093E7B68E92495B66" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="F21BDAE256BABD11C27916BD363411D4" pageId="0" pageNumber="1">
|
||||
The specific epithet
|
||||
<taxonomicName lsidName="inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">inferospira</emphasis>
|
||||
<taxonomicName id="AB34F9AC65BBCE40121075AA88F8F48F" lsidName="inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis id="D7A82331276F0DD25B456A329B1D549A" italics="true" pageId="0" pageNumber="1">inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
is derived from the Latin word
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">infer</emphasis>
|
||||
<emphasis id="3EB743ACEAA75B705BF456DB559793F4" italics="true" pageId="0" pageNumber="1">infer</emphasis>
|
||||
meaning low and the Latin word
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">spira</emphasis>
|
||||
<emphasis id="58ED3E233F583E032358802682110B95" italics="true" pageId="0" pageNumber="1">spira</emphasis>
|
||||
meaning coil. It refers to the strongly depressed shell with low spire.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
<subSubSection id="6550065798DB2679D4CFDE8C72578CC7" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="B1E2D93F007F3B046670724B1DB257FD" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph id="D9207708EDEE31AC720265FB0D1F103F" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="C09EE2E6094CC9E63A9E8045B3A8AEEF" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis id="3A213006822D1D89CA4B133E47028736" italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is only known from sandstone habitats with dry dipterocarp forest at the type locality (Fig.
|
||||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Geographic distribution of Sarika bocourti, S. hainesi, S. inferospira sp. nov., S. melanospira sp. nov. and S. pellosa sp. nov. based on examined specimens herein." figureDoi="10.3897/zookeys.976.53859.figure32" httpUri="https://binary.pensoft.net/fig/464414" pageId="0" pageNumber="1">32</figureCitation>
|
||||
<figureCitation id="40450593F3745496C8E1FC6A1C8783B3" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Geographic distribution of Sarika bocourti, S. hainesi, S. inferospira sp. nov., S. melanospira sp. nov. and S. pellosa sp. nov. based on examined specimens herein." figureDoi="10.3897/zookeys.976.53859.figure32" httpUri="https://binary.pensoft.net/fig/464414" pageId="0" pageNumber="1">32</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="coi analysis">
|
||||
<paragraph pageId="0" pageNumber="1">COI analysis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="84168D870B86629523D2F24BA84AE4EE" pageId="0" pageNumber="1" type="coi analysis">
|
||||
<paragraph id="86D2A7475D88222EB105330A0B5B0BFD" pageId="0" pageNumber="1">COI analysis.</paragraph>
|
||||
<paragraph id="61AE3D1DAB3BA910CE2D750A28E28CEF" pageId="0" pageNumber="1">
|
||||
The ML and BI analyses revealed that the individuals of
|
||||
<taxonomicName lsidName="S. inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. inferospira</emphasis>
|
||||
<taxonomicName id="D46A07621E6401293C982C290FAD5587" lsidName="S. inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis id="243B6D74CD7DD07346AEDB9D9A40ED59" italics="true" pageId="0" pageNumber="1">S. inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. (n = 3) formed a monophyletic group with very strong support (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">1</figureCitation>
|
||||
<figureCitation id="470BF7C257B3E3D489038D6A5ED573FA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">1</figureCitation>
|
||||
; BS = 100%, PP = 1), sister group to
|
||||
<taxonomicName lsidName="S. melanospira" pageId="0" pageNumber="1" rank="species" species="melanospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. melanospira</emphasis>
|
||||
<taxonomicName id="D819B8601A2D6A837509713D363E6C1C" lsidName="S. melanospira" pageId="0" pageNumber="1" rank="species" species="melanospira">
|
||||
<emphasis id="678F425772BE356A9F194D5A348F9404" italics="true" pageId="0" pageNumber="1">S. melanospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. with only ML support (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">1</figureCitation>
|
||||
<figureCitation id="454717C34806D7DBD2A7FE34393230EE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">1</figureCitation>
|
||||
; BS = 78%). The mean intraspecific genetic distance of
|
||||
<taxonomicName lsidName="S. inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. inferospira</emphasis>
|
||||
<taxonomicName id="3D2560AFDA7A03085ACC9C6FCAB9726C" lsidName="S. inferospira" pageId="0" pageNumber="1" rank="species" species="inferospira">
|
||||
<emphasis id="AB9AD49233A55003D31F641B1BF3E627" italics="true" pageId="0" pageNumber="1">S. inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. was 0.1% (Table
|
||||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean intra-specific and inter-specific genetic divergences among species of Sarika from the mitochondrial COI gene sequences estimated by the K 2 P model. Taxa in bold are the new species described herein." httpUri="http://table.plazi.org/id/13BDBDF714F2FAA17ADB274D839659A1" pageId="0" pageNumber="1" tableUuid="13BDBDF714F2FAA17ADB274D839659A1">2</tableCitation>
|
||||
<tableCitation id="1C72297E2EB1FE78DCE83AA1BF6F8F09" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean intra-specific and inter-specific genetic divergences among species of Sarika from the mitochondrial COI gene sequences estimated by the K 2 P model. Taxa in bold are the new species described herein." httpUri="http://table.plazi.org/id/13BDBDF714F2FAA17ADB274D839659A1" pageId="0" pageNumber="1" tableUuid="13BDBDF714F2FAA17ADB274D839659A1">2</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
<subSubSection id="1E9968B93163205913021EAF484A49DB" pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph id="836AED9107E85A072382A02AC6FCBD34" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph id="F6B28469BAB6C55771F8D0A76CB4658B" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="233C516D8EDEC6B9E553FBF10FC22B8D" authorityName="Pholyotha & Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika inferospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="inferospira">
|
||||
<emphasis id="50AE575B8CE3C27F50E7A9EFBAA87B7D" italics="true" pageId="0" pageNumber="1">Sarika inferospira</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is distinguished from
|
||||
<taxonomicName lsidName="S. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. hainesi</emphasis>
|
||||
<taxonomicName id="280D028440DD6EFC9638225A232CB74A" lsidName="S. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||||
<emphasis id="2EB098F98960D9F0595A267D8D605BA3" italics="true" pageId="0" pageNumber="1">S. hainesi</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
<taxonomicName id="D723F8F73881FC221A712A3F8502B196" lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis id="98FBB4C682BB61B07A9FA3458A9D33F8" italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
</taxonomicName>
|
||||
by having a strongly depressed shape, shouldered body whorl, and spermatophore smooth or without spine on the middle part of tail filament.
|
||||
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika hainesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Sarika hainesi</emphasis>
|
||||
<taxonomicName id="1FBA5F49AA2E426545AA70847FDEA9E6" baseAuthorityName="Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika hainesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||||
<emphasis id="56FC96CF59D09481FEA389E10B227740" italics="true" pageId="0" pageNumber="1">Sarika hainesi</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
<taxonomicName id="7B3C6EC3C2AC7F4C162B0A02533BA1D0" lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis id="2AD113BF459EDADA5D486BAD64EA0180" italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
</taxonomicName>
|
||||
have a depressed shell with a higher spire and obtusely angulated body whorl. In addition, the tail filament of spermatophore contains a series of short branching spines more than ca. half of its length in
|
||||
<taxonomicName lsidName="S. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. hainesi</emphasis>
|
||||
<taxonomicName id="7F4A99BCF13E42C01D7F0FE6C52B353F" lsidName="S. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||||
<emphasis id="941754E49C202BB121D2EBE927A273EA" italics="true" pageId="0" pageNumber="1">S. hainesi</emphasis>
|
||||
</taxonomicName>
|
||||
and more than ca. two-thirds of its length in
|
||||
<taxonomicName lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
<taxonomicName id="3CC8FED646BB4918861BC5141912C3A2" lsidName="S. bocourti" pageId="0" pageNumber="1" rank="species" species="bocourti">
|
||||
<emphasis id="BFB26D3310273E53CD4F62DD1E2271D1" italics="true" pageId="0" pageNumber="1">S. bocourti</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
Loading…
Add table
Add a link
Reference in a new issue